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... Some CE4 enzymes are metal ion-dependent (typically Zn2+ or Co2+) and all contain a conserved region called the NodB homology domain (Correia, 2010;Hoell et al., 2010;Nakamura et al., 2017;Sista Kameshwar and Qin, 2018 and excellent host organisms for the production of recombinant enzymes for application in biotechnology owing to their ability to effectively secrete large amounts of proteins and metabolites to extracellular culture media. In terms of industrial enzyme production, current industrial production is reported to be 100 g/L homologous protein in the cultivation medium from some fungi (Kour et al., 2019;Siddiqui, 2016). ...