Annotated list of taxonomic novelties published in "Fungi Fenniae Exsiccati. Samling af Finska svampar" issued by P.A. Karsten between 1861 and 1870

Abstract

New taxa and new combinations published by P. A. Karsten in “Fungi Fenniae Exsiccati” in the second half of the 19th century are listed and annotated, supplemented by copies of the labels with the original descriptions or new combinations. References, citations, and the synonymy are corrected when necessary. The nomenclature of some taxa is discussed in more detail. The new combinations Byssonectria deformis, Calycellina leucella, Calycellina microspis, Echinosphaeria canescens, Hymenoscyphus suspectus, Keissleriella culmifida, Lachnellula abietis, Leiosphaerella lycopodina, Paraleptosphaeria praetermissa, Pezoloma ciliiferum, Rhodotarzetta euchroa, Sydowiella depressula, and Tephrocybe cessans and the replacement name Niptera bispora are introduced.

Full text

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Schlechtendalia 40, 2023
Annotated list of taxonomic novelties published in “Fungi Fenniae
Exsiccati. Samling af Finska svampar” issued by P.A. Karsten between
1861 and 1870
James K. MITCHELL & Konstanze BENSCH
Abstract: Mitchell, J. K. & Bensch, K. 2023: Annotated list of taxonomic novelties published in “Fungi Fenniae Exsiccati.
Samling af Finska svampar” issued by P.A. Karsten between 1861 and 1870. Schlechtendalia 40: 280–347.
New taxa and new combinations published by P. A. Karsten in “Fungi Fenniae Exsiccati” in the second half of the 19th
century are listed and annotated, supplemented by copies of the labels with the original descriptions or new combinations.
References, citations, and the synonymy are corrected when necessary. The nomenclature of some taxa is discussed in more
detail. The new combinations Byssonectria deformis, Calycellina leucella, Calycellina microspis, Echinosphaeria canescens,
Hymenoscyphus suspectus, Keissleriella culmifida, Lachnellula abietis, Leiosphaerella lycopodina, Paraleptosphaeria
praetermissa, Pezoloma ciliiferum, Rhodotarzetta euchroa, Sydowiella depressula, and Tephrocybe cessans and the
replacement name Niptera bispora are introduced.
Zusammenfassung: Mitchell, J. K. & Bensch, K. 2023: Annotierte Liste taxonomischer Neuheiten publiziert in “Fungi
Fenniae Exsiccati. Samling af Finska svampar”, herausgegeben von P. A. Karsten zwischen 1861 und 1870. Schlechtendalia
40: 280–347.
Neue Taxa und Kombinationen publiziert von P. A. Karsten in “Fungi Fenniae Exsiccati” in der zweiten Hlfte des 19.
Jahrhunderts werden aufgelistet und annotiert, ergnzt durch Kopien der Etiketten mit den Originalbeschreibungen oder
Kombinationen. Referenzangaben, Zitate und die Synonymie werden korrigiert falls notwendig. Die Nomenklatur einiger
Taxa wird detaillierter besprochen. Die neuen Kombinationen Byssonectria deformis, Calycellina leucella, Calycellina
microspis, Echinosphaeria canescens, Hymenoscyphus suspectus, Keissleriella culmifida, Lachnellula abietis, Leiosphaerella
lycopodina, Paraleptosphaeria praetermissa, Pezoloma ciliiferum, Rhodotarzetta euchroa, Sydowiella depressula, und
Tephrocybe cessans und der neue Name Niptera bispora werden eingefhrt.
Keywords: Fungi, nomenclature, exsiccatae.
Published online 23 Aug. 2023
Introduction
Petter Adolf Karsten, the “Father of Finnish mycology” and the pioneer of fungal microscopy
(https://www.luomus.fi/en/botanical-and-mycological-collections), was born on 16 Feb. 1834 in
Merimasku and died on 22 March 1917 in Forssa. His research work included a vast number of
specimens representing all groups of fungi and was largely based on material he collected locally at
Tammela, southern Finland where he worked as a lecturer at the Mustiala Agriculture Institute (now
part of Häme Polytechnic) from 1865 until 1908.
His exsiccata “Fungi Fenniae Exsiccati. Samling af Finska svampar” comprises 998 specimens issued
in ten centuries between 1861 and 1870 (nos. 332 and 660 apparently not having been issued). Most
are specimens collected near Tammela, but an additional significant source of material was an
expedition Karsten had participated in in 1861 to the Kola Peninsula in Russia funded by the Societas
pro Fauna et Flora Fennica (Sennikov & Kozhin 2018). The standard abbreviation according to
IndExs, the Index of Exsiccatae (http://indexs.botanischestaatssammlung.de/) is “Karsten, Fungi Fenn.
Exs.” Karsten’s collection is preserved at the Botanical Museum of Finland (H), which bought it from
him in 1905 for 10.000 Mk (Elfving 1921: 17).
Details of this exsiccata are provided in Pfister (1985) but are scant. This is due largely to the fact that
little has been published about this set. Despite being among Karsten’s earliest scientific works, even
detailed biographies typically mention only the start and end dates of this ten-century exsiccata
(Elfving 1921, Hintikka 1935, 1950). It is not known how many copies there are or where they are all
held, but in addition to the listed copies at FH, UPS, H, and BM and a partial copy at NY can be added
a copy at K, possibly an additional partial copy in K in Plowright’s herbarium, and at least a partial
copy in S in Rehm’s herbarium. The listing of a possible set at BPI by Pfister has been confirmed to be
an error; records at BPI confused this set with Kari’s Fungi Exsiccati Fennici. The dates given by
Pfister are based on the title pages of the ten centuries, but there has been some uncertainty about the
accuracy of the date given for the first century. This uncertainty has risen largely from the fact that

281
Saelan, in an overview of Finnish botanical literature, listed Centuries I–IV as having been issued in
1865–6 (Saelan 1867: 104). Saelan later contradicted this, listing Century I as having been issued in
1861 as indicated on the title page (Saelan 1916: 575). Verification of the publication dates has been
complicated by the fact that no announcements or advertisements for Fungi Fenniae Exsiccati were
known to have been published; the only known mentions in contemporary scientific contexts were a
report on the 1867 International Exposition in Paris that stated that Century V was being sold by
Wasenius & Comp. in Helsinki for 25 Mk (Kornhuber 1869: 279) and Karsten’s (1869: 101) mention
in the introduction to Monographia Pezizarum Fennicarum that Century VII had been published and
was available for 20 Mk from Wasenius & Co. in Helsinki. In both cases, the century mentioned is
supposed to have already been available for purchase for years by the time of publication, limiting the
usefulness of these mentions for confirming the title page dates. It has come to our attention, though,
that announcements were published in local Finnish newspapers. In particular, Century I was
announced as being available from Frenckellska Bokhandeln and A. W. Edgren for 4 Rbls in the 25
May 1861 (p. 4) and 28 May 1861 (p. 4) editions of Åbo Underrättelser, confirming its publication in
1861 (https://digi.kansalliskirjasto.fi/sanomalehti/ binding/365664?term=Samling%20af%20finska%
20svampar&page=4). The publication dates of some later centuries are likewise confirmed by these
announcements: Century II was announced for 16 Mk in the 31 Oct. 1865 edition of Helsingfors
Dagblad (p. 4, https://digi.kansalliskirjasto.fi/sanomalehti/ binding/688848?term=P.%20A.%20
Karsten&page=4), Century III was announced for 18 Mk in the 21 Nov. 1865 edition of Helsingfors
Tidningar (p. 4, https://digi.kansalliskirjasto.fi/sanomalehti/ binding/516909?term=P.%20A.%20
Karsten&page=4), and Century VII was announced in the 28 Dec. 1867 edition of Helsingfors
Dagblad (p. 4, https://digi.kansalliskirjasto.fi/sanomalehti/binding/ 436528?term=Fungi&term=
Fenniae&term=exsiceati&page=4).
While extremely helpful for verification of publication dates, the announcements in local newspapers
also raise questions about the purpose and intended audience of this exsiccata. By most biographic
accounts of Karsten (e.g., Elfving 1921, Hintikka 1935, 1950), mycology was in its infancy in Finland
while this set was being issued. There was only one notable mycologist in Finland, William Nylander,
and not yet much interest at the amateur level. The choice to advertise this set (at least at first) solely
in local newspapers and not in scientific publications would seem to indicate that Karsten intended the
set to be used primarily by Finns, perhaps for the purpose of sparking interest in mycology in his
country. If the 16 advertisements placed by Wasenius & Comp. in February and March of 1866
offering Centuries II and III for sale in Helsingfors Tidningar are anything to go by, this effort was
less than successful. This would seem to be borne out also by the fact that only one set is at present
known to still be in Finland.
An almost complete set of “Fungi Fenniae Exsiccati” (only one label missing, no. 316) preserved in
FH was the base for the present nomenclatural study and reassessment of names; a scan of the missing
label was provided by staff at H. While possessing an almost complete set of labels, the FH set
presents several irregularities. Foremost among these is the fact that many specimens are missing. This
has been noted on many packets, most dating to the re-curation of the set in 1975, when the packets
and sheets specimens were stored in were replaced with archival materials. In addition to the empty
packets, many packets contain replacements. These are generally indicated by a handwritten slip in the
packet in Karsten’s hand containing a taxon name and collecting information. In at least one case (no.
932, Schmitzomia elevata) though, the specimen in the packet is apparently wrong (the specimen is on
a monocot and not Symphoricarpos, as indicated on the packet) and lacks any other indication of
replacement. While these factors complicate use of this set (researchers may wish to consult one of the
other sets as well), the substituted material in some cases seems to represent candidate type material of
taxa not included in the original issue of the set (e.g., FH 00785331, Sphaeria hydrophila P. Karst.
substituted for no. 335, Hysterium pinastri var. juniperinum) or issued later in the exsiccata (e.g., FH
00785019, Sphaerella caulicola substituted for no. 20, Sphaeria leptidea Fr.). These substitutions
would seem to point to this copy of Fungi Fenniae Exsiccati having been made up of leftover material
and labels rather than being one of the original sets presumably prepared by Karsten. The archives of
the Farlow Herbarium were searched for information on the provenance of this set to no avail. While
Karsten corresponded directly with both William Gilson Farlow and his student, Roland Thaxter, very
little of this correspondence has been preserved, and no mention of the set is made in the remaining
letters.

282
According to ICN, Art. 30.7 (Ex. 13), descriptions on labels accompanying distributed specimens
(exsiccatae) issued prior to 1953 are effectively published. This means that (contrary to widely held
belief) valid names published on printed labels of distributed exsiccatae must be taken into
consideration with regard to priority issues of taxa. Later publications in books or journals (possibly
with repeated descriptions) are later isonyms (Art. 6.3, Note 2) that have to be disregarded (Braun
2018).
Another portion of the ICN relevant to this exercise is Art. 41.5–8, the portion concerning
requirements post-1953 for valid publication of new combinations, names at new rank, and
replacement names. While originally vaguely worded (Greuter 1998), the clarified and added rules in
the St. Louis (Art. 33.3–6) and later codes require that authors publishing new combinations, names at
new state, and replacement names cite the place of original publication of the previously validly
published name their proposed nomenclatural novelty is based on, and not later publications. This
requirement does include a handful of exceptions (outlined in Art. 41.8) which allow citation of a later
or earlier work to be treated as a correctable error, not requiring any additional nomenclatural acts, but
these exceptions are limited. This has resulted in a fairly large number of invalidly published
combinations proposed after 1953 which have not widely been realized to be invalidly published. This
has been recognized by some authors (e.g., Redhead et al. 2011, Ahti et al. 2015, Parra et al. 2017),
but remains relatively unknown with many people apparently satisfied to take the entries in Index
Fungorum or MycoBank at face value. This rule is particularly relevant in the case of names originally
published in exsiccatae as publication of these names frequently precedes publication elsewhere by
years. The situation is made worse by the widespread misunderstanding that printed labels in
exsiccatae do not constitute valid publication and researchers’ limited access to exsiccatae labels for
verification.
In the particular case of Fungi Fenniae Exsiccati, this is a significant issue. Some names published in
this exsiccata precede publication in a later work by up to 13 years (Puccinia lysimachiae). Most
authors, perhaps ultimately following Saccardo’s treatment in his Sylloge Fungorum, have cited as the
place of publication of these new names Karsten’s Mycologia Fennica, consisting of four parts
published from 1871–9, entirely after the publication of Fungi Fenniae Exsiccati had concluded. This
work contains extensive references to the specimens issued in Fungi Fenniae Exsiccati, as do many of
Karsten’s publications from the era. Citation as basionym a later work when Karsten references a label
from Fungi Fenniae Exsiccati constituting valid publication is therefore not correctable under Art.
41.8(a). For an error in citation to be correctable in most cases, designation of or full and direct
reference to a type is required. Not all the labels Karsten issued in this exsiccata and cited in later
works bear a description or diagnosis, however, and so some name first mentioned in Fungi Fenniae
Exsiccati are in fact to be cited as validly published in a later work. The fact that only eight partial or
full copies of this exsiccata are known to exist and individual labels bear neither the name of the
exsiccata or Karsten’s name makes it difficult for researchers to obtain the information required to
verify whether their citations of basionyms or replaced synonyms are correct when dealing with names
Karsten cited in his journal articles as appearing in Fungi Fenniae Exsiccati.
In the course of this project, 14 "names" in current use were found to have been invalidly published
and are validated here. This is done with the help of Hermann Voglmayr (Leiosphaerella lycopodina),
Uwe Lindemann (Byssonectria deformis), Jan Hengstmengel (Hymenoscyphus suspectus), Andrew
Miller (Echinosphaeria canescens), Ron Bronckers (Rhodotarzetta euchroa), Hans-Otto Baral
(Niptera bispora), and Gerard Verkleij (Paraleptosphaeria praetermissa). The remaining
combinations validated here were originally proposed by authors now deceased.
The following list contains taxonomic novelties published in Karsten’s exsiccata, together with copies
of the original labels.
Annotated list of taxa
All scanned labels of “P.A. Karsten, Fungi Fenniae Exsiccati. Samling af Finska svampar” added to the
following list are based on a nearly complete set of this exsiccata deposited at the Farlow Herbarium (FH) of
Harvard University. Specimen barcodes are provided for all specimens in the figure legends.
Cent. 1
Phragmidium incrassatum f. rosarum (Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 4, 1861.
≡ Phragmidium incrassatum a. rosarum Rabenh., Deutschl. Krypt.-Fl. 1: 32, 1844.

283
≡ Phragmidium incrassatum var. rosarum Wallr., Fl. Crypt. Germ. 2: 188, 1833, nom. illeg., Art. 52.1.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 4 (fig.
1). The combination was also proposed by Fuckel (1861: 8).
Fig. 1 (FH 00785003)
Tubercularia vulgaris f. betulae (Wallr.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 14, 1861.
≡ Tubercularia vulgaris var. betulae Wallr., Fl. Crypt. Germ. 2: 238, 1833.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 14 (fig.
2). It is very probably a synonym of Nectria cinnabarina.
Fig. 2 (FH 00785013)
Aecidium cornutum f. sorbi (Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 28, 1861.
≡ Aecidium cornutum d. sorbi Rabenh., Deutschl. Krypt.-Fl. 1: 20, 1844.
≡ Caeoma cylindrites Link, in Willd., Sp. Pl. 6(2): 64, 1825, nom. illeg., Art. 52.1.
= Aecidium cornutum J.F. Gmel., Syst. Nat. Ed. 1791 2(2): 1472, 1792, nom. sanct. [Pers., Syn. Meth.
Fung. 1: 205].
≡ Gymnosporangium cornutum (J.F. Gmel.) Arthur, Mycologia 1(6): 240, 1909.
[Full synonymy, see https://www.mycobank.org/MB/541831]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 28 (fig.
3).
Fig. 3 (FH 00785027)
Aecidium compositarum f. tussilaginis (J.F. Gmel.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 39, 1861.
≡ Aecidium tussilaginis J.F. Gmel., Syst. Nat. Ed. 1791 2(2): 1473, 1792, nom. sanct. [Pers., Syn.
Meth. Fung. 1: 209].
= Puccinia poarum Nielsen, Bot. Tidsskr. Ser. 3 2: 34, 1879.
[Full synonymy, see https://www.mycobank.org/MB/194454]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 39 (fig.
4). The current name for this species is not compliant with Art. 11.4; a combination in Puccinia should
likely be proposed for Aecidium tussilaginis, or Puccinia poarum should be conserved or protected.
Fig. 4 (FH 00785038)
Phragmidium incrassatum f. ruborum (Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 60, 1861.
≡ Phragmidium incrassatum b. ruborum Rabenh., Deutschl. Krypt.-Fl. 1: 32, 1844.
≡ Phragmidium incrassatum var. ruborum Wallr., Fl. Crypt. Germ. 2: 189, 1833, nom. illeg., Art.
52.1.

284
≡ Puccinia mucronata var. rubi Pers., Neues Mag. Bot. 1: 118 & T. 3 fig. 5b, 1794, nom. sanct. [Pers.,
Syn. Meth. Fung. 1: 230].
≡ Phragmidium rubi (Pers.) G. Winter, Rabenh. Krypt.-Fl. Ed. 2, Band 1 Abth. 1(Lief. 4): 230, 1881
[1884].
[Full synonymy, see https://www.mycobank.org/MB/210476]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 60 (fig.
5). The combination was also proposed by Fuckel (1861: 8).
Fig. 5 (FH 00785059)
Uredo candida f. cruciferarum (Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 78, 1861.
≡ Uredo candida a. cruciferarum Rabenh., Deutschl. Krypt.-Fl. 1: 13, 1844.
≡ Uredo cruciferarum DC., in Lam. & DC., Fl. Franç. 2: 596, 1805, nom. illeg., Art. 52.1.
= Aecidium candidum Pers. ex. J.F. Gmel., Syst. Nat. Ed. 1791 2(2): 1473, 1792.
≡ Uredo candida (Pers. ex. J.F. Gmel.) Pers., Syn. Meth. Fung. 1: 223, nom. sanct. [Pers., l.c.].
≡ Albugo candida (Pers. ex. J.F. Gmel.) Roussel, Fl. Calvados Ed. 2: 47, 1806.
[Full synonymy, see https://www.mycobank.org/MB/122113]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 78 (fig.
6).
Fig. 6 (FH 00785077)
Uredo fulva f. sonchorum (Wallr.) P. Karst., Fungi Fenn. Exs., Cent. 1: no. 83, 1861.
≡ Erysibe fulva var. sonchorum Wallr., Fl. Crypt. Germ. 2: 188, 1833.
≡ Uredo fulva a. sonchorum (Wallr.) Rabenh., Deutschl. Krypt.-Fl. 1: 12, 1844.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs.
83 (fig. 7).
Fig. 7 (FH 00785082)
Cent. 2
Coniophora puteana (Schumach.) P. Karst. (as ‘puteanea’), Fungi Fenn. Exs., Cent. 2: no. 135, 1865
[Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 370, 1868].
≡ Thelephora puteana Schumach., Enum. Plant. 2: 397, 1803, nom. sanct. [Fr., Syst. Mycol. 1: 448,
1821].
[Full synonymy, see https://www.mycobank.org/MB/204571]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 135 (fig.
8).
Fig. 8 (FH 00785134)

285
Cyathicula coronata (Bull.) P. Karst., Fungi Fenn. Exs., Cent. 2: no. 146, 1865.
≡ Peziza coronata Bull., Herb. France 9: pl. 416, fig. 4, 1789, nom. sanct. [Fr., Syst. Mycol. 2: 120,
1822].
[Full synonymy, see https://www.mycobank.org/MB/144480]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 146 (fig.
9). The combination was also proposed by Rehm (1893: 740).
Fig. 9 (FH 00785145)
Tubercularia vulgaris f. aesculi (Opiz) P. Karst., Fungi Fenn. Exs., Cent. 2: no. 156, 1865.
≡ Tubercularia aesculi Opiz, in Corda, Icon. fung. 1: 4, tab. 1, fig. 77, 1837.
= Sphaeria cinnabarina Tode, Fung. Mecklenb. Sel. 2: 9, 1791, nom. sanct. [Fr., Syst, Mycol. 2(2):
412, 1823].
≡ Nectria cinnabarina (Tode) Fr., Summa Veg. Scand. 2: 388, 1849.
[Full synonymy, see https://www.mycobank.org/MB/249710]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 156 (fig.
10).
Fig. 10 (FH 00785155)
Cent. 3
Cantharellus aurantiacus var. fuscolutescens P. Karst., Fungi Fenn. Exs., Cent. 3: no. 229, 1865.
Notes: This is a new variety name validly published on the label of Karsten, Fungi Fenn. Exs. 229 (fig.
11).
Fig. 11 (FH 00785228)
Peziza omphalodes f. confluens (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 3: no. 256, 1865.
≡ Peziza confluens Pers., Observ. Mycol. 2: 81, 1800 [1799].
= Pyronema omphalodes (Bull.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 319, 1870.
[Full synonymy, see https://www.mycobank.org/MB/195122]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 311 (fig.
12).
Fig. 12 (FH 00785255)
Tubercularia vulgaris f. sambuci (Corda) P. Karst., Fungi Fenn. Exs., Cent. 3: no. 258, 1865.
≡ Tubercularia sambuci Corda, Icon. Fung. 1: 4, tab. 1, fig. 69, 1837.
≡ Tubercularia vulgaris l. sambuci (Corda) Rabenh., Deutschl. Krypt.-Fl. 1: 54, 1844.
= Nectria cinnabarina (Tode) Fr., Summa Veg. Scand. 2: 388, 1849.
[Full synonymy, see https://www.mycobank.org/MB/249710]

286
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 258 (fig.
13).
Fig. 13 (FH 00785257)
Sordaria coprophila f. hirtula P. Karst., Fungi Fenn. Exs., Cent. 3: no. 284, 1865, nom. inval., Art.
35.2.
= Sordaria winteri P. Karst. (as ‘Winterii’), Bidrag Knnedom Finlands Natur Folk 23: [251], 1873.
= Podospora appendiculata (Auersw. ex Niessl) Niessl, Hedwigia 22(10): 156, 1883.
Notes: This is an invalidly published name as the epithet is not definitely associated with the name of
the species on the label of Karsten, Fungi Fenn. Exs. 284 (fig. 14). Later, it was included in the
circumscription of the new species Sordaria winteri P. Karst. 1873. Karsten (1873: 52) first published
the name Sordaria breviseta with a detailed description but as a combination based on Malinvernia
breviseta Fuckel. The latter one is however an illegitimate name as its circumcription includes the
name Sphaeria pauciseta Ces. On page [251] Karsten (1873) corrected the name to ‘winteri’ stating
that it is not identical with Fuckel’s name. Lundqvist (1972: 129–132) lectotypified Sordaria winteri
and placed it in synonymy with Podospora appendiculata (Auersw. ex Niessl) Niessl.
Fig. 14 (FH 00785283)
Cent. 4
Trametes fibrillosa (P. Karst.) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 311, 1866.
≡ Polyporus fibrillosus P. Karst., Sydv. Finl. Polypor.: 30, 1859.
= Pycnoporellus fulgens (Fr.) Donk, Persoonia 6(2): 216, 1971.
[Full synonymy, see https://www.mycobank.org/MB/322113]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 311 (fig.
15). The name was lectotypified by Lowe (1956), see also Niemelä (1980).
Fig. 15 (FH 00785310)
Tremella albida f. betulae P. Karst., Fungi Fenn. Exs., Cent. 4: no. 317, 1866.
= Exidia candida Lloyd, Mycol. Not. 5(44): 620, 1916.
[Full synonymy, see https://www.mycobank.org/MB/205540]
Notes: This is a new form name validly published on the label of Karsten, Fungi Fenn. Exs. 317 by
provision of the host (“björk”, Betula sp.) (fig. 16). A lectotype (H 6048760) was designated by Spirin
et al. (2017: 5).

287
Fig. 16 (FH 00785315)
Peziza resinae var. stipitulata P. Karst., Fungi Fenn. Exs., Cent. 4: no. 324, 1866, nom. inval., Art.
38.1(a).
= Zythia resinae (Ehrenb.) P. Karst., Meddeland. Soc. Fauna Fl. Fenn. 14: 104, 1887 [1888].
[Full synonymy, see https://www.mycobank.org/MB/146209].
Notes: This is a nomen nudum, invalidly published as a description is lacking (fig. 17). Later, it was
mentioned by Karsten (1885: 154) with the brief diagnosis “variat stipitata” under Tromera resinae,
but without denoting a rank or definitely associating the epithet “stipitulata” with the name Tromera
resinae, a fact overlooked by Mitchell et al. (2021). The name was validly published as Tromera
resinae var. stipitulata Sacc. in 1889 by reference to Karsten’s (l.c.) diagnosis (Saccardo 1889: 469).
Fig. 17 (FH 01093952)
Cenangium furfuraceum (Roth) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 326, 1866.
≡ Peziza furfuracea Roth, Catal. Bot. 2: 257, 1800, nom. sanct. [Fr., Syst. Mycol. 2: 76, 1822]
≡ Encoelia furfuracea (Roth) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 218, 1871.
[Full synonymy, see https://www.mycobank.org/MB/202507]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 326 (fig.
18). The combination is often attributed to De Notaris (1864: 384), but the species is given there as
Dermatea furfuracea although listed in the genus Cenangium.
Fig. 18 (FH 00785323)
Lachnella alboviolascens f. caraganae P. Karst., Fungi Fenn. Exs., Cent. 4: no. 329, 1866.
= Lachnella alboviolascens (Alb. & Schwein.) Fr., Summa Veg. Scand. 2: 365, 1849.
Notes: This is a new form name validly published on the label of Karsten, Fungi Fenn. Exs. 329 by
provision of the host (Caragana arborescens) (fig. 19). Donk (1959) lists the name as synonym of
Lachnella alboviolascens.
Fig. 19 (FH 00785326)
Phacidium pini f. juniperi P. Karst., Fungi Fenn. Exs., Cent. 4: no. 339, 1866.
≡ Phacidium juniperi (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 257, 1870 [1871].
≡ Coccomyces juniperi (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 254, 1871.
≡ Colpoma juniperi (P. Karst.) Dennis, Kew Bull. 12(3): 401, 1957.

288
[Full synonymy, see https://www.mycobank.org/MB/295446]
Notes: This is a new form name validly published on the label of Karsten, Fungi Fenn. Exs. 339 by
provision of the host (“enbarr”, needles of Juniperus sp.). (fig. 20).
Fig. 20 (FH 00785335)
Pseudovalsa schistostroma (Duby ex Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 344, 1866.
≡ Sphaeria schistostroma Duby ex Rabenh., Fungi Eur. Exs., Cent. 1: no. 53, 1859.
[Full synonymy, see https://www.mycobank.org/MB/202825]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 344 (fig.
21).
Fig. 21 (FH 00785340)
Gnomonia gnomon (Tode) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 348, 1866.
≡ Sphaeria gnomon Tode, Fung. Mecklenb. Sel. 2: 50, 1791, nom. sanct. [Fr., Syst. Mycol. 2: 517,
1822].
[Full synonymy, see https://www.mycobank.org/MB/120553]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 348 (fig.
22). Karsten attributes the combination to Cesati & De Notaris (1863: 232), but those authors used
instead the illegitimate (Art. 52.1) replacement name Gnomonia vulgaris, likely to avoid a perceived
tautonym. The combination was also proposed by Schröter in Cohn (1897: 390). The type species of
the genus Gnomonia was lecto- and epitypified in Sogonov et al. (2005).
Fig. 22 (FH 00785344)
Gnomonia curva (Wallr.) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 349, 1866.
≡ Sphaeria curva Wallr., Fl. Crypt. Germ 2: 803, 1833.
≡ Stromatostysanus curvus (Wallr.) U. Braun, Sydowia 45(1): 88, 1993.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 349 (fig.
23). The combination was also proposed by Fuckel (1870: 123).
Fig. 23 (FH 00785345)
Cryptosphaeria stricta (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 353, 1866.
≡ Sphaeria stricta Pers., Syn. Meth. Fung. 1: 59, 1801, nom. sanct. [Fr., Syst. Mycol. 2: 474, 1823].
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 353 (fig.
24).
Fig. 24 (FH 00785349)

289
Valsa sorbi f. padi P. Karst., Fungi Fenn. Exs., Cent. 4: no. 355, 1866.
≡ Valsa padi (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 134, 1873.
≡ Eutypella padi (P. Karst.) Sacc., Syll. Fung. 1: 147, 1882.
[Full synonymy, see https://www.mycobank.org/MB/150300]
Notes: This is a new form name validly published on the label of Karsten, Fungi Fenn. Exs. 355 (fig.
25) by provision of the host (“hgg”, Prunus padus).
Fig. 25 (FH 00785351)
Phoma excentrica P. Karst. (as ‘excentricum’), Fungi Fenn. Exs., Cent. 4: no. 364, 1866.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 364
(fig. 26).
Fig. 26 (FH 00785360)
Uncinula adunca f. salicum (Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 367, 1866.
≡ Erysiphe adunca bb. salicum Rabenh. (as ‘Erysibe’), Deutschl. Krypt.-Fl. 1: 236, 1844.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 367 (fig.
27).
Fig. 27 (FH 00785363)
Phyllosticta trientalis (Lasch ex Rabenh.) P. Karst., Fungi Fenn. Exs., Cent. 4: no. 373, 1866.
≡ Depazea trientalis Lasch ex Rabenh., Deutschl. Krypt.-Fl. 1: 138, 1844.
≡ Septoria trientalis (Lasch ex Rabenh.) Sacc., Bull. Soc. Mycol. France 5(4): 121, 1890 [1889].
[Full synonymy, see https://www.mycobank.org/MB/208628]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 373 (fig.
28).
Fig. 28 (FH 00785369)
Ustilago segetum f. polygoni P. Karst., Fungi Fenn. Exs., Cent. 4: no. 394, 1866.
≡ Caeoma utriculosum Nees, Syst. Pilze: 14, tab. 1, fig. 6, 1816.
= Sphacelotheca hydropiperis (Schumach.) de Bary, Vergl. Morph. Biol. Pilze: 187, 1884.
[Full synonymy, see https://www.mycobank.org/MB/227738]
Notes: This is a new replacement name validly published on the label of Karsten, Fungi Fenn. Exs.
394 (fig. 29).

290
Fig. 29 (FH 00785390)
Cent. 5
Heyderia cucullata (Batsch) P. Karst., Fungi Fenn. Exs., Cent. 5: no. 447, 1866.
≡ Helvella cucullata Batsch (as ‘Elvela’), Elench. Fung. Continuatio Prima: 189, 1786.
≡ Geoglossum cucullatum (Batsch) Fr., Elench. Fung. 1: 233, 1828, nom. sanct. [Fr., l.c.].
[Full synonymy, see https://www.mycobank.org/MB/511214]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 447 (fig.
30). The combination was also proposed by Bacyk & Van Vooren in Bacyk (2005: 42).
Fig. 30 (FH 00785443)
Cyathicula calycina (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 5: no. 460, 1866 [Enum. Fung.
Lapponia: 207, 1866].
≡ Peziza calycina Pers., Syn. Meth. Fung. 2: 653, 1801 [Schumach., Enum. Pl. 2: 424, 1803], nom.
sanct. [Fr., Syst. Mycol. 2: 91, 1822].
≡ Helvella calyciformis Batsch (as ‘Elvela’), Elench. Fung. Continuatio Prima: 195, 1786.
≡ Lachnellula calyciformis (Batsch) Dharne, Phytopathol. Z. 53(2): 124, 1964.
[Full synonymy, see https://www.mycobank.org/MB/332845]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 460 (fig.
31).
Fig. 31 (FH 00785456)
Ascobolus carbonarius P. Karst., Fungi Fenn. Exs., Cent. 5: no. 463, 1866 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 202, 1870 [1871].
[Full synonymy, see https://www.mycobank.org/MB/145251]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 463
(fig. 32).
Fig. 32 (FH 00785459)
Gnomonia coryli (Batsch) P. Karst., Fungi Fenn. Exs., Cent. 5: no. 472, 1866 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 13: 222, 1873].

291
≡ Sphaeria coryli Batsch, Elench. Fung. Continuatio Secunda: 131, 1789, nom. sanct. [Fr., Syst.
Mycol. 2: 436, 1823].
≡ Mamianiella coryli (Batsch) Höhn., Ber. Deutsch. Bot. Ges. 35(8): 637, 1917.
[Full synonymy, see https://www.mycobank.org/MB/179634]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 472 (fig.
33). The combination was also proposed by Auerswald in Gonnermann & Rabenhorst (1869: 23).
Fig. 33 (FH 00785468)
Gnomonia chamaemori (Fr.) P. Karst., Fungi Fenn. Exs., Cent. 5: no. 473, 1866 [Bidrag Kännedom
Finlands Natur Folk 23: 123, 1873; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 222, 1873].
≡ Sphaeria chamaemori Fr., Syst. Mycol. 2(2): 519, 1823, nom. sanct. [Fr., l.c.].
≡ Gnomoniopsis chamaemori (Fr.) Berl., Icon. Fung. 1(3): 93, 1893 [1894].
[Full synonymy, see https://www.mycobank.org/MB/209402]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 473 (fig.
34).
Fig. 34 (FH 00785469)
Hypopteris graminis (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 5: no. 483, 1866.
≡ Sphaeria graminis Pers., Ann. Bot. (Usteri) 15: 18, 1795, nom. sanct. [Fr., Syst. Mycol. 2(2): 434,
1823].
≡ Phyllachora graminis (Pers.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 216, 1870.
[Full synonymy, see https://www.mycobank.org/MB/200927]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 483 (fig.
35).
Fig. 35 (FH 00785479)
Epicoccum scabrum var. tritici P. Karst., Fungi Fenn. Exs., Cent. 5: no. 489, 1866.
≡ Epicoccum tritici Oerst., Sygd. Plant: 110, 113, 1863, nom. inval., Art. 36.1(a).
Notes: This is a new variety name validly published on the label of Karsten, Fungi Fenn. Exs. 489 (fig.
36) by indirect reference to Ørsted’s (1863: 110 & 113) figures.
Fig. 36 (FH 00785485)
Puccinia graminis f. calthae P. Karst. (as ‘gracilis’), Fungi Fenn. Exs., Cent. 5: no. 495, 1866.
Notes: This is a new form name validly published on the label of Karsten, Fungi Fenn. Exs. 495 (fig.
37). While Karsten cites the species name as ‘Puccinia gracilis’, the citation he gives leads to Corda’s
(1840: 11) treatment of Puccinia graminis Pers.

292
Fig. 37 (FH 00785491)
Cent. 6
Agaricus sororiatus P. Karst., Fungi Fenn. Exs., Cent. 6: no. 507, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 9: 339, 1868].
= Pluteus leoninus (Schaeff.) P. Kumm., Führer Pilzk.: 98, 1871.
[Full synonymy, see https://www.mycobank.org/MB/150582]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 507
(fig. 38).
Fig. 38 (FH 00785503)
Hydnum gracilipes P. Karst., Fungi Fenn. Exs., Cent. 6: no. 521, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 9: 362, 1868].
≡ Hydnellum gracilipes (P. Karst.) P. Karst., Meddeland. Soc. Fauna Fl. Fenn. 5: 41, 1879.
[Full synonymy, see https://www.mycobank.org/MB/100891]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 521
(fig. 39). Kõljalg & Renvall (2000) confirmed the placement of this species in the genus Hydnellum
and designated a lectotype (H 6006461).
Fig. 39 (FH 00785517)
Peziza perforata P. Karst., Fungi Fenn. Exs., Cent. 6: no. 529, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 112, 1869; Bidrag Kännedom Finlands Natur Folk 19: 56, 1871], nom. illeg., Art. 53.1,
homonym, non Peziza perforata Saut. 1841.
≡ Geopyxis perforata Sacc., Syll. Fung. 8: 73, 1889.
= Tarzetta catinus (Holmsk.) Korf & J.K. Rogers, Phytologia 21 (4): 206, 1971.
[Full synonymy, see https://www.mycobank.org/MB/324436]
Notes: This is an illegitimate name introduced on the label of Karsten, Fungi Fenn. Exs. 529 (fig. 40).
Saccardo (1889) introduced a replacement name for it. A lectotype (H 6033946) was designated in
Huhtinen (1994) and identified as Tarzetta catinus.

293
Fig. 40 (FH 00785525)
Peziza abundans P. Karst., Fungi Fenn. Exs., Cent. 6: no. 532, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 124, 1869; Bidrag Kännedom Finlands Natur Folk 19: 67, 1871].
≡ Trichophaea abundans (P. Karst.) Boud., Hist. Classif. Discomyc. Eur.: 61, 1907.
[Full synonymy, see https://www.mycobank.org/MB/119754]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 532
(fig. 41).
Fig. 41 (FH 00785528)
Peziza radians P. Karst., Fungi Fenn. Exs., Cent. 6: no. 533, 1866 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 200, 1869], nom. illeg., Art. 53.1, homonym, non Peziza radians Saut. 1841.
≡ Lachnea radians P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 251, 1870 [1871].
≡ Lachnum radians (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 177, 1871.
≡ Trichopeziza karstenii Sacc., Syll. Fung. 8: 417, 1889.
[Full synonymy, see https://www.mycobank.org/MB/174733]
Notes: This is an illegitimate name introduced on the label of Karsten, Fungi Fenn. Exs. 533 (fig. 42).
Saccardo (1889) introduced a replacement name for it and Dennis (1963: 360) designated a lectotype
(H 6039294).
Fig. 42 (FH 00785529)
Peziza impudicella P. Karst., Fungi Fenn. Exs., Cent. 6: no. 534, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 185, 1869].
≡ Encoelia impudicella (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 253, 1870 [1871],
nom. inval., Art. 35.1.
≡ Encoelia impudicella (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 219, 1871.
≡ Cenangium impudicellum (P. Karst.) P. Karst., Acta Soc. Fauna Fl. Fenn. 2(6): 145, 1885.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 534
(fig. 43).
Fig. 43 (FH 00785530)

294
Peziza limosella P. Karst., Fungi Fenn. Exs., Cent. 6: no. 538, 1866 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 152, 1869].
≡ Ombrophila violacea var. limosella (P. Karst.) P. Karst. (as ‘var. γ. limosella’), Bidrag Kann dom
Finlands Natur Folk 19: 88, 1871.
≡ Ombrophila limosella (P. Karst.) Rehm, Ascomyceten Fasc. 11: no. 508, 1879.
[Full synonymy, see https://www.mycobank.org/MB/210212]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 538
(fig. 44).
Fig. 44 (FH 00785534)
Peziza lutiseda P. Karst., Fungi Fenn. Exs., Cent. 6: no. 539, 1866 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 152, 1869].
[Full synonymy, see https://www.mycobank.org/MB/512515]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 539
(fig. 45). Dennis (1956: 105) cited as the ‘typus’ of this species a specimen collected in 1867 (H
6003814), the year after the original publication, and this is thus not original material. This must be
taken to be a neotypification, but it should be noted that original material does exist, should it be found
that this material does not agree with Karsten’s original concept.
Fig. 45 (FH 00785535)
Peziza echinospora P. Karst. (as ‘echinispora’), Fungi Fenn. Exs., Cent. 6: no. 541, 1866 [Not. Sällsk.
Fauna Fl. Fenn. Förh. 10: 115, 1869; Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 226, 1870 [1871]; Bidrag
Kännedom Finlands Natur Folk 19: 54, 1871].
[Full synonymy, see https://www.mycobank.org/MB/221954].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 541
(fig. 46). Both molecular and morphological characters support P. echinospora as well-separated
species (Hansen et al. 2002). A copy of the exsiccatae preserved in H (H 6039047) was chosen there
as lectotype of the species.
Fig. 46 (FH 00785537)
Peziza ovulispora P. Karst., Fungi Fenn. Exs., Cent. 6: no. 543, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 167, 1869].
≡ Patellaria ovulispora (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 251, 1870 [1871].
= Durella melanochlora (Sommerf.) Rehm, Hedwigia 21(8): 114, 1882.
[Full synonymy, see https://www.mycobank.org/MB/237935].

295
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 543
(fig. 47).
Fig. 47 (FH 00785539)
Peziza cortiseda P. Karst., Fungi Fenn. Exs., Cent. 6: no. 544, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 149, 1869].
≡ Helotium uliginosum subsp. cortisedum (P. Karst.) P. Karst. (as ‘H. uliginosum * H. cortisedum’),
Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 236, 1870 [1871].
≡ Helotium uliginosum var. cortisedum (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk
19: 122, 1871.
≡ Hymenoscyphus cortisedus (P. Karst.) Dennis, Persoonia 3(1): 75, 1964 [1963].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 544
(fig. 48).
Fig. 48 (FH 00785540)
Peziza ciliifera P. Karst., Fungi Fenn. Exs., Cent. 6: no. 546, 1866 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 153, 1869].
≡ Pezoloma ciliiferum (P. Karst.) Korf (as ‘ciliifera’), Phytologia 21(4): 205, 1971, nom. inval., Art.
41.8.
≡ Pezoloma ciliiferum (P. Karst.) Korf ex J.K. Mitch. & Bensch, comb. nov., MB 848924.
[Basionym: Peziza ciliifera P. Karst., Fungi Fenn. Exs., Cent. 6: no. 546, 1866.]
Syntypes: [Finland,] Mustiala, på fuktig mark [moist ground], Sept. [September], [Karsten, Fungi
Fenn. Exs. 546] (e.g., H 6039308, UPS F-128814, FH 00785542).
[Full synonymy, see https://www.mycobank.org/MB/848924]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 546
(fig. 49). The combination Pezoloma ciliiferum was proposed by Korf (1971: 205), who referred to
Monographia Pezizarum Fennicarum (Karsten 1869: 153) as the place of valid publication of Peziza
ciliifera when making the combination in Pezoloma. Because Korf did not cite a type or make a full
and direct citation to one and Karsten (l.c.) clearly cites his earlier publication in Fungi Fenniae
Exsiccati, none of the provisions of Art. 41.8 apply and this combination is invalidly published; we
validate it here.
Fig. 49 (FH 00785542)
Peziza caudata P. Karst., Fungi Fenn. Exs., Cent. 6: no. 547, 1866 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 144, 1869].
≡ Hymenoscyphus caudatus (P. Karst.) Dennis, Persoonia 3(1): 76, 1964 [1963].
[Full synonymy, see https://www.mycobank.org/MB/332330].

296
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 547
(fig. 50).
Fig. 50 (FH 00785543)
Peziza stemmata P. Karst., Fungi Fenn. Exs., Cent. 6: no. 548, 1866 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 154, 1869].
[Full synonymy, see https://www.mycobank.org/MB/224996].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 548
(fig. 51). Dixon (1974: 100) examined original material in H and FH and found this to be a species of
Mollisia.
Fig. 51 (FH 00785544)
Peziza sauciella P. Karst., Fungi Fenn. Exs., Cent. 6: no. 549, 1866 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 185, 1869].
?= Hyphodiscus hymeniophilus (P. Karst.) Baral, Z. Mykol. 59(1): 7, 1993.
[Full synonymy, see https://www.mycobank.org/MB/363707].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 549
(fig. 52). A lectotype (H 6052782) was designated by Carpenter (1981: 252, as ‘holotype’).
Fig. 52 (FH 00785545)
Ascobolus myriadeus P. Karst., Fungi Fenn. Exs., Cent. 6: no. 552, 1866.
[Full synonymy, see https://www.mycobank.org/MB/654383].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 552
(fig. 53). Karsten’s original concept of the species encompassed apothecia with asci containing
anywhere from 8 to 64 spores at maturity, but he soon (1871a: 81 & 1871b: 208) decided to restrict his
concept to those with only 24–32 spores per ascus. According to Doveri (2007: 65 & 67) this species
in the sense of Saccardo is a synonym of Thelebolus caninus and in the sense of Boudier is a synonym
of Thelebolus crustaceus, but according to de Hoog et al. (2005: 66) the polysporous portions of
Karsten’s original concept are synonyms of a broadly circumscribed Thelebolus stercoreus. These
latter authors apparently studied the copy of Fungi Fenn. Exs. no. 552 in K(M), listing it as an isotype
and lectotype of different portions of the original concept of Ascobolus myriadeus. In any case, they
did not include a statement indicating designation in that publication and so did not achieve a
lectotypification (Art. 7.11).

297
Fig. 53 (FH 00785548)
Sphaeria linnaeae P. Karst. [as ‘linneae’], Fungi Fenn. Exs., Cent. 6: no. 562, 1866, nom. illeg., Art.
53.1, homonym, non Sphaeria linnaeae Ehrenb. 1818.
Notes: This is an illegitimate name published on the label of Karsten, Fungi Fenn. Exs. 562 (fig. 54).
Later, Karsten (1873a: 188) listed the specimen under the name Venturia dickiei, though he did not
mention the name he assigned it on the label.
Fig. 54 (FH 00785558)
Sphaeria obducens var. minor P. Karst., Fungi Fenn. Exs., Cent. 6: no. 564, 1866, nom. inval., Art.
38.1(a).
≡ Sphaeria obducens b. minor Fr., Syst. Mycol. 2(2): 456, 1823, nom. inval., Art. 38.1(a).
Notes: This is an invalidly published variety name, a nomen nudum (fig. 55). It is a reference to
Fries’s above infraspecific taxon, but this was likewise a nomen nudum.
Fig. 55 (FH 00785560)
Sphaerella lycopodina P. Karst., Fungi Fenn. Exs., Cent. 6: no. 569, 1866 [Bidrag Kännedom Finlands
Natur Folk 23: 184, 1873].
≡ Leiosphaerella lycopodina (P. Karst.) Jaklitsch & Voglmayr, Fungal Diversity 52(1): 82, 2012, nom.
inval., Art. 41.8 (Shenzhen).
≡ Leiosphaerella lycopodina (P. Karst.) Jaklitsch & Voglmayr, comb. nov. MB 848925
[basionym: Sphaerella lycopodina, P. Karst., Fungi Fenn. Exs., Cent. 6: no. 569, 1866].
[Full synonymy, see https://www.mycobank.org/MB/848925].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 569
(fig. 56). Aptroot (2006: 124) mentioned a lectotype (in H) without indicating where it was designated.
Jaklitsch & Voglmayr (2012) transferred the species in the genus Leiosphaerella, however, did not cite
a type or make a full and direct citation to one and Karsten (1873) clearly cites his earlier publication
in Fungi Fenniae Exsiccati. Therefore, none of the provisions of Art. 41.8 apply and the combination
is invalidly published. It is validated herein.
Fig. 56 (FH 00785565)
Polysaccum boreale P. Karst., Fungi Fenn. Exs., Cent. 6: no. 570, 1866 [Enum. Fung. Lapponia: 203,
1866].
= Pisolithus arhizus (Scop.) Rauschert, Z. Pilzk. 25(2): 50, 1959.

298
[Full synonymy, see https://www.mycodb.fr/fiche.php?genre=Pisolithus&espece=arhizus;
https://www.mycobank.org/MB/276857].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 570
(fig. 57).
Fig. 57 (FH 00785566)
Puccinia lysimachiae P. Karst., Fungi Fenn. Exs., Cent. 6: no. 590, 1866 [Bidrag Kännedom Finlands
Natur Folk 31: 27, 1879; Bidrag Kännedom Finlands Natur Folk 39: 49, 1884].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 590
(fig. 58). In the protologue, Karsten (l.c.) also cited ‘Uredo lysimachiae’ which is an indirect reference
to the combination Uredo lysimachiae (Schltdl.) Spreng. based on Caeoma lysimachiae which was
given as synonym in Karsten (1879). However, he added a brief separate description for Puccinia
lysimachiae, suggesting that he intended to introduce a new species for the teleomorphic state (Art.
F.8.1).
Fig. 58 (FH 00785586)
Puccinia angelicae P. Karst., Fungi Fenn. Exs., Cent. 6: no. 591, 1866, nom. inval., Art. 38.1(a).
Notes: This is an invalidly published name. Since Karsten did not list even an author on the label of
Fungi Fenn. Exs. 591 (fig. 59), it is not clear whether this was intended as a combination or as a new
species. Karsten (1879: 36) seemed to clarify that he considered it to be Puccinia angelicae
(Schumach.) Fuckel, published nine years earlier (Fuckel 1870: 52), but four years after the issuance
of the sixth century of Fungi Fenniae Exsiccati. Puccinia karstenii Lindr. (1901), originally published
as ‘Karsteni’ is based on Puccinia angelicae sensu Karsten, Bidrag Kännedom Finlands Natur Folk
31: 36, 1879, non (Schumach.) Fuckel.
Fig. 59 (FH 00785587)
Tilletia serpens P. Karst., Fungi Fenn. Exs., Cent. 6: no. 599, 1866.
≡ Ustilago serpens (P. Karst.) B. Lindeb., Symb. Bot. Upsal. 16(2): 133, 1959.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 599
(fig. 60). The species was lectotypified (TUB HUV 10432) and revisited by Kruse et al. (2018).
Fig. 60 (FH 00785595)

299
Cent. 7
Lentinus dentatus (Fr.) P. Karst., Fungi Fenn. Exs., Cent. 7: no. 611, 1867.
≡ Agaricus cochleatus var. dentatus Fr., Syst. Orb. Veg.: 78, 1825.
≡ Agaricus dentatus Pers., in Hoffm., Abbild. Schwämme 3: [ii], 1793, nom. illeg., Art. 53.1,
homonym, non Agaricus dentatus L. 1753.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 611 (fig.
61). The combination was also proposed by Quélet (1886a: 151; 1886b: 449). Probably a synonym of
Lentinellus micheneri (Berk. & M.A. Curtis) Pegler (1983).
Fig. 61 (FH 00785606)
Polyporus euporus P. Karst., Fungi Fenn. Exs., Cent. 7: no. 618, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 9: 360, 1868].
= Junghuhnia nitida (Pers.) Ryvarden, Persoonia 7(1): 18, 1972.
[Full synonymy, see https://www.mycobank.org/MB/315983]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 618
(fig. 62). Lowe (1956) designated the copy preserved in UPS as lectotype.
Fig. 62 (FH 00785613)
Polyporus hians P. Karst., Fungi Fenn. Exs., Cent. 7: no. 619, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
9: 360, 1868].
= Anomoporia bombycina (Fr.) Pouzar, Česká Mykol. 20(3): 172, 1966.
[Full synonymy, see https://www.mycobank.org/MB/326232]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 619
(fig. 63). The specimen preserved in UPS was designated as lectotype in Lowe (1956).
Fig. 63 (FH 00785614)
Peziza virella P. Karst., Fungi Fenn. Exs., Cent. 7: no. 627, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 188, 1869].
≡ Amicodisca virella (P. Karst.) Huhtinen, Karstenia 34: 5, 1994.
[Full synonymy, see https://www.mycobank.org/MB/362258]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 627
(fig. 64). Huhtinen (1994) designated the copy of the exsiccatae preserved in H as lectotype (H
6003237) and transferred the name into the genus Amicodisca.
Fig. 64 (FH 00785622)

300
Peziza deformis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 628, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 119, 1869; Bidrag Kännedom Finlands Natur Folk 19: 47, 1871].
≡ Byssonectria deformis (P. Karst.) U. Lindem. & M. Vega, Z. Mykol. 81 (2): 383, 2015, nom. inval.,
Art. 41.8.
≡ Byssonectria deformis (P. Karst.) U. Lindem. & M. Vega, comb. nov. MB 849034.
[Basionym: Peziza deformis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 628, 1867.]
Lectotype: [Finland,] Mustiala, ad terram argillaceam [clay soil], 4 Juli 1866, P.A. Karsten 3013 (H
6038822, designated by Khare & Tewari 1978: 2115).
[Full synonymy, see https://www.mycobank.org/MB/849034]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 628
(fig. 65). The combination Byssonectria deformis was proposed by Lindemann & Vega in Lindemann
et al. (2015: 383), who referred to Monographia Pezizarum Fennicarum (Karsten 1869: 119) as the
place of valid publication of Peziza deformis when making the combination in Byssonectria. Because
Lindemann and Vega did not cite a type or make a full and direct citation to one and Karsten (l.c.)
clearly cites his earlier publication in Fungi Fenniae Exsiccati, none of the provisions of Art. 41.8
apply and this combination is invalidly published; it is validated here.
Fig. 65 (FH 00785623)
Ditiola conformis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 629, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 223, [1871] 1870].
= Femsjonia peziziformis (Lév.) P. Karst. (as ‘pezizaeformis’), Bidrag Ka nedom Finlands Natur Folk
25: 352, 1876.
[Full synonymy, see https://www.mycobank.org/MB/147491]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 629
(fig. 66).
Fig. 66 (FH 00785624)
Peziza nobilis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 635, 1867.
= Pseudombrophila merdaria (Fr.) Brumm., Libri Bot. 14: 45, 1995.
[Full synonymy, see https://www.mycobank.org/MB/413971; https://www.mycodb.fr/fiche.php?
genre=Pseudombrophila&espece=merdaria].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 635
(fig. 67). It was lectotypified (H 6057295) and reduced to synonymy with Pseudombrophila merdaria
by Brummelen (1995).
Fig. 67 (FH 00785630)
Peltidium oocardii var. lignarium P. Karst. (as ‘lignaria’), Fungi Fenn. Exs., Cent. 7: no. 636, 1867.
≡ Peziza oocardii var. lignaria (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 128, 1869.
≡ Peltidium oocardii f. lignarium (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 232,
1870 [1871].
≡ Humaria oocardii var. lignaria (P. Karst.) Sacc. (as ‘ligniaria’), Syll. Fung. 8: 144, 1889.

301
= Adelphella babingtonii (Berk. & Broome) Pfister, Matočec & I. Kušan, Mycol. Montenegr. 11: 14,
2009 [2008].
[Full synonymy, see https://www.mycobank.org/MB/538132].
Notes: This is a new variety name validly published on the label of Karsten, Fungi Fenn. Exs. 636 (fig.
68).
Fig. 68 (FH 00785631)
Peziza discreta P. Karst., Fungi Fenn. Exs., Cent. 7: no. 637, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 146, 1869].
≡ Calycina discreta (P. Karst.) Kuntze, Revis. Gen. Pl. 3 (3): 448, 1898.
[Full synonymy, see https://www.mycobank.org/MB/246531]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 637
(fig. 69). A lectotype (H 6039291) was designated by Carpenter (1981: 209).
Fig. 69 (FH 00785632)
Peziza conscripta P. Karst., Fungi Fenn. Exs., Cent. 7: no. 638, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 134, 1869].
≡ Helotium conscriptum (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 236, 1870 [1871].
≡ Helotium sublenticulare subsp. conscriptum (P. Karst.) P. Karst. (as ‘* conscriptum’), Bidrag
Kännedom Finlands Natur Folk 19: 119, 1871.
≡ Hymenoscyphus conscriptus (P. Karst.) Korf, in Kobayasi et al., Rep. (Annual) Inst. Ferment. Res.
Osaka. 3: 55, 1967.
[Full synonymy, see https://www.mycobank.org/MB/332333; https://www.mycodb.fr/fiche.php?
genre=Hymenoscyphus&espece=conscriptus].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 638
(fig. 70). Dennis (1956: 84) discusses a ‘typus’ of this species, but there are two specimens in
Karsten’s herbarium (H 6039403 and H 6039392) that are broadly consistent with the information
Dennis gives. There may be additional text on the packets that clarifies the situation.
Fig. 70 (FH 00785633)
Peziza complicata P. Karst., Fungi Fenn. Exs., Cent. 7: no. 642, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 166, 1869].
≡ Mollisia complicata (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 192, 1871.
≡ Cyathicula complicata (P. Karst.) Dennis, Kew Bull. 30(2): 348, 1975.
[Full synonymy, see https://www.mycobank.org/MB/312398]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 642
(fig. 71). Carpenter (1981: 48) designated a lectotype (H 6039312).

302
Fig. 71 (FH 00785637)
Peziza hydrophila P. Karst., Fungi Fenn. Exs., Cent. 7: no. 643, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 163, 1869].
≡ Niptera palustris subsp. hydrophila (P. Karst.) P. Karst. (as ‘* hydrophila’), Not. Sa lsk. Fauna Fl.
Fenn. Förh. 11: 247, 1870 [1871].
≡ Mollisia palustris subsp. hydrophila (P. Karst.) P. Karst. (as ‘* hydrophila’), Bidrag Kann dom
Finlands Natur Folk 19: 196, 1871.
≡ Mollisia hydrophila (P. Karst.) Sacc., Syll. Fung. 8: 345, 1889.
[Full synonymy, see https://www.mycobank.org/MB/145883]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 643
(fig. 72).
Fig. 72 (FH 00785638)
Peziza excelsior P. Karst., Fungi Fenn. Exs., Cent. 7: no. 644, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 165, 1869].
≡ Niptera palustris subsp. excelsior (P. Karst.) P. Karst. (as ‘****** excelsior’), Not. Sa lsk. Fauna Fl.
Fenn. Förh. 11: 247, 1870 [1871].
≡ Belonopsis excelsior (P. Karst.) Rehm, Rabenh. Krypt.-Fl. Ed. 2, Band 1 Abth. 3(Lief. 36): 572,
1891 [1896].
[Full synonymy, see https://www.mycobank.org/MB/142008].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 644
(fig. 73). A lectotype (H 6039317) was designated by Dennis (1950: 183).
Fig. 73 (FH 00785639)
Peziza caricinella P. Karst., Fungi Fenn. Exs., Cent. 7: no. 645, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 166, 1869].
≡ Coronellaria caricinella (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 248, 1870
[1871] [Bidrag Kännedom Finlands Natur Folk 19: 185, 1871].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 645
(fig. 74).
Fig. 74 (FH 00785640)

303
Peziza microspis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 647, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 178, 1869].
≡ Calycellina microspis (P. Karst.) Dennis, Trans. Bot. Soc. Edinburgh 45(3): 213, 1989 [1988], nom.
inval., Art. 41.8.
≡ Calycellina microspis (P. Karst.) Dennis, Kew Bull. 45(2): 293, 1990, nom. inval., Art. 41.8.
≡ Calycellina microspis (P. Karst.) Dennis ex J.K. Mitch. & Bensch, comb. nov. MB 848926.
[Basionym: Peziza microspis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 647, 1867.]
Syntypes: [Finland,] Runsala, på blad af Phragm. [leaves of Phragmites], Aug. [August], [P. A.
Karsten, Fungi Fenn. Exs. 647] (e.g., H 6007765, UPS s.n., FH 00785642, S F7848, S F7851).
[Full synonymy, see https://www.mycobank.org/MB/848926]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 647
(fig. 75). The combination Calycellina microspis was proposed twice by Dennis (1989: 213 & 1990:
293), who referred to Monographia Pezizarum Fennicarum (Karsten 1869: 178) as the place of valid
publication of Peziza microspis when making the combination in Calycellina. Because Dennis did not
cite a type or make a full and direct citation to one and Karsten (l.c.) clearly cites his earlier
publication in Fungi Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this combination
is invalidly published in both places; it is validated here. Dennis (1956: 181) gave as ‘typus’ a
specimen in Karsten’s herbarium at H collected on 1 September 1868 in Runsala on rotting leaves of
Phragmites. As this was collected the year after the publication of Peziza microspis, this has to be
considered a neotypification. According to Baral et al. (2020: 1678), however, this material cannot
currently be located in H, and may be lost. In any case, original material is still in existence (see
above) and a lectotype can be chosen from among the extant material.
Fig. 75 (FH 00785642)
Peziza scrupulosa P. Karst., Fungi Fenn. Exs., Cent. 7: no. 648, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 178, 1869].
≡ Olla scrupulosa (P. Karst.) Svrček, Česká Mykol. 40(4): 216, 1986.
[Full synonymy, see https://www.mycobank.org/MB/131040].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 648
(fig. 76).
Fig. 76 (FH 00785643)
Peziza granulosella P. Karst., Fungi Fenn. Exs., Cent. 7: no. 649, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 180, 1869].
≡ Cistella granulosella (P. Karst.) Nannf., Nova Acta Regiae Soc. Sci. Upsal. Ser. 4, 8(2): 270, 1932.
[Full synonymy, see https://www.mycobank.org/MB/266483]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 649
(fig. 77).
Fig. 77 (FH 00785644)

304
Peziza tenebrosa P. Karst., Fungi Fenn. Exs., Cent. 7: no. 650, 1867.
≡ Peziza rosae subsp. tenebrosa (P. Karst.) P. Karst. (as ‘* Peziza tenebrosa’), Not. Sllsk. Fauna Fl.
Fenn. Förh. 10: 177, 1869.
≡ Trochila rosae f. tenebrosa (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 248, 1870
[1871], nom. inval., Art. 35.2 & 41.1.
≡ Mollisia rosae var. tenebrosa (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 209,
1871, nom. inval., Art. 35.2.
[Full synonymy, see https://www.mycobank.org/MB/569703].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 650
(fig. 78).
Fig. 78 (FH 00785645)
Peziza leucella P. Karst., Fungi Fenn. Exs., Cent. 7: no. 651, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 179, 1869].
≡ Calycellina leucella (P. Karst.) Dennis, Beitr. Kryptogamenfl. Schweiz 15 (1): 29, 1977, nom. inval.
Art. 41.8.
≡ Calycellina leucella (P. Karst.) Dennis ex J.K. Mitch. & Bensch, comb. nov. MB 848927.
[Basionym: Peziza leucella P. Karst., Fungi Fenn. Exs., Cent. 7: no. 651, 1867.]
Lectotype: [Finland,] Mustiala, på al- och björkblad [leaves of Alnus and Betula], Sept. [September],
[Karsten, Fungi Fenn. Exs. 651] (H 6039287, designated by Raschle 1979: 671).
[Full synonymy, see https://www.mycobank.org/MB/848927]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 651
(fig. 79). The combination Calycellina leucella was proposed by Dennis in Müller (1977: 29), who
referred to Monographia Pezizarum Fennicarum (Karsten 1869: 179) as the place of valid publication
of Peziza leucella when making the combination in Calycellina. Because Dennis did not cite a type or
make a full and direct citation to one and Karsten (l.c.) clearly cites his earlier publication in Fungi
Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this combination is invalidly
published; it is validated here.
Fig. 79 (FH 00785646)
Peziza eurotioides P. Karst., Fungi Fenn. Exs., Cent. 7: no. 652, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 190, 1869].
≡ Helotium eurotioides (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 241, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 146, 1871].
≡ Unguiculella eurotioides (P. Karst.) Nannf., Trans. Brit. Mycol. Soc. 20 (3–4): 194, 1936.
[Full synonymy, see https://www.mycobank.org/MB/252057; https://www.mycodb.fr/fiche.php?
genre=Unguiculella&espece=eurotioides]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 652
(fig. 80).
Fig. 80 (FH 00785647)

305
Peziza crispula P. Karst., Fungi Fenn. Exs., Cent. 7: no. 653, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 181, 1869].
≡ Helotium crispulum (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 241, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 161, 1871].
≡ Urceolella crispula (P. Karst.) Boud., Hist. Classif. Discomyc. Eur.: 130, 1907.
[Full synonymy, see https://www.mycobank.org/MB/120257]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 653
(fig. 81). It was lectotypified (H 6051484) in Raschle (1977: 206).
Fig. 81 (FH 00785648)
Peziza xylita P. Karst., Fungi Fenn. Exs., Cent. 7: no. 654, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 190, 1869].
≡ Cistella xylita (P. Karst.) Nannf., Nova Acta Regiae Soc. Sci. Upsal. Ser. 4, 8 (2): 271, 1932.
[Full synonymy, see https://www.mycobank.org/MB/282206]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 654
(fig. 82).
Fig. 82 (FH 00785649)
Ascobolus polysporus P. Karst., Fungi Fenn. Exs., Cent. 7: no. 655, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 208, 1871]; non Ascobolus polysporus Auersw. 1868.
≡ Pezizula polyspora (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 82, 1871.
≡ Thelebolus polysporus (P. Karst.) Otani & Kanzawa, Trans. Mycol. Soc. Japan 11 (2): 45 (1970).
[Full synonymy, see https://www.mycobank.org/MB/324535]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 655
(fig. 83).
Fig. 83 (FH 00785650)
Ascobolus punctiformis P. Karst., Fungi Fenn. Exs., Cent. 7: no. 655, 1867.
≡ Ascobolus polysporus subsp. punctiformis (P. Karst.) P. Karst. (as ‘* Asc. punctiformis’), Not.
Sällsk. Fauna Fl. Fenn. Förh. 11: 209, 1871.
≡ Pezizula polyspora subsp. punctiformis (P. Karst.) P. Karst. (as ‘* punctiformis’), Bidrag Kann dom
Finlands Natur Folk 19: 82, 1871.
= Thelebolus microsporus (Berk. & Broome) Kimbr., in Kobayasi et al., Rep. (Annual) Inst. Ferment.
Res. Osaka. 3: 50, 1967.

306
[Full synonymy, see https://www.mycobank.org/MB/340022]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 655
(fig. 83).
Ditangium P. Karst., Fungi Fenn. Exs., Cent. 7: no. 656, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh. 11:
213, 1870 [1871].
[Full synonymy, see https://www.mycobank.org/MB/17519]
Notes: This is a new genus name validly published on the label of Karsten, Fungi Fenn. Exs. 656 (fig.
84). See Malysheva et al. (2019, https://link.springer.com/article/10.1007/s11557-019-01485-3).
Ditangium insigne P. Karst., Fungi Fenn. Exs., Cent. 7: no. 656, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 213, 1870 [1871].
= Ditangium cerasi (Schumach.) Costantin & L.M. Dufour, Nouv. Fl. Champ. Ed. 1: 207, 1891.
[Full synonymy, see https://www.mycobank.org/MB/492544]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 656
(fig. 84). Malysheva et al. (2019: 759) lectotypified (H 6060993) the species and synonymized it with
Ditangium cerasi.
Fig. 84 (FH 00785651)
Ascobolus hirtellus P. Karst., Fungi Fenn. Exs., Cent. 7: no. 657, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 210, 1871].
[Full synonymy, see https://www.mycobank.org/MB/487364]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 657
(fig. 85). According to Brummelen (1967) this species may be one of the many forms of Lasiobolus
pilosus; Bezerra & Kimbrough (1975) treated it as a nomen dubium.
Fig. 85 (FH 00785652)
Ascobolus versicolor P. Karst., Fungi Fenn. Exs., Cent. 7: no. 659, 1867 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 203, 1871; Bidrag Kännedom Finlands Natur Folk 19: 79, 1871].
≡ Saccobolus versicolor (P. Karst.) P. Karst., Acta Soc. Fauna Fl. Fenn. 2 (6): 123, 1885.
[Full synonymy, see https://www.mycobank.org/MB/238189]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 659
(fig. 86). See https://ascomycete.org/de/2000-Pezizales/Art/Saccobolus-versicolor?mapid=521.
Fig. 86 (FH 00785654)

307
Patellaria pitya P. Karst. (as ‘pithya’), Fungi Fenn. Exs., Cent. 7: no. 661, 1867.
≡ Tympanis pitya (P. Karst.) P. Karst. (as ‘pithya’), Not. Sllsk. Fauna Fl. Fenn. Förh. 11: 218 & 252,
1870 [1871] [Bidrag Kännedom Finlands Natur Folk 19: 228, 1871].
[Full synonymy, see https://www.mycobank.org/MB/169656]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 661
(fig. 87).
Fig. 87 (FH 00785655)
Peziza acutipila P. Karst., Fungi Fenn. Exs., Cent. 7: no. 662, 1867 [Not. Sällsk. Fauna Fl. Fenn. Förh.
10: 195, 1869].
≡ Albotricha acutipila (P. Karst.) Raitv., Scripta Mycol. 1: 40, 1970.
[Full synonymy, see https://www.mycobank.org/MB/308446]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 662
(fig. 88).
Fig. 88 (FH 00785656)
Nectria dacrymycella (Nyl.) P. Karst., Fungi Fenn. Exs., Cent. 7: no. 667, 1867 [Bidrag Kännedom
Finlands Natur Folk 23: 216, 1873].
≡ Sphaeria dacrymycella Nyl., Flora 46(21): 322, 1863.
≡ Nectriella dacrymycella (Nyl.) Rehm, Ascomyceten Fasc. 5: no. 232, 1874.
[Full synonymy, see https://www.mycobank.org/MB/120293]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 667 (fig.
89).
Fig. 89 (FH 00785661)
Nectria dacrymycella var. sceptri (P. Karst.) P. Karst., Fungi Fenn. Exs., Cent. 7: no. 667, 1867.
≡ Nectria sceptri P. Karst., Enum. Fung. Lapponia: 213, 1866.
≡ Charonectria sceptri (P. Karst.) Lowen, in Rossman et al., Stud. Mycol. 42: 181, 1999.
[Full synonymy, see https://www.mycobank.org/MB/460464]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 667 (fig.
89). For full synonymy see Rossman et al. (1999: 181).
Sphaeria helvellae P. Karst., Fungi Fenn. Exs., Cent. 7: no. 674, 1867.
≡ Sphaeronaemella helvellae (P. Karst.) P. Karst., Hedwigia 23(2): 18, 1884.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 674
(fig. 90). A lectotype (FH 00290654) was designated by Upadhyay (1981: 98, as ‘holotype’).

308
Fig. 90 (FH 00290654)
Pilobolus lentiger var. conformis P. Karst. (as ‘lentigerus’), Fungi Fenn. Exs., Cent. 7: no. 678, 1867.
Notes: This is a new variety name validly published on the label of Karsten, Fungi Fenn. Exs. 678 (fig.
91).
Fig. 91 (FH 00785671)
Pilobolus intermedius (Coem.) P. Karst. (as ‘intermedium’), Fungi Fenn. Exs., Cent. 7: no. 678, 1867
[Bidrag Kännedom Finlands Natur Folk 32: 71, 1879].
≡ Pilobolus oedipus var. intermedius Coem. (as ‘intermedia’), Bull. Acad. Roy. Sci. Belgique Sér. 2,
16(7): 71, 1863.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 678 (fig.
91).
Cent. 8
Agaricus cessans P. Karst., Fungi Fenn. Exs., Cent. 8: no. 703, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 219, 1870 [1871].
≡ Collybia cessans (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 32: 163, 1879.
≡ Tephrocybe cessans (P. Karst.) M.M. Moser, Kleine Krypt.-Fl. Band 2b, Teil 2: 116, 1967, nom.
inval., Art. 41.8.
≡ Tephrocybe cessans (P. Karst.) M.M. Moser ex J.K. Mitch. & Bensch, comb. nov. MB 848928.
[Basionym: Agaricus cessans P. Karst., Fungi Fenn. Exs., Cent. 8: no. 703, 1868.]
Syntypes: [Finland,] Mustiala, på granbarr [needles of Picea abies], Oct. [October] 1867, [P. A.
Karsten, Fungi Fenn. Exs. 703] (e.g., H 6019037, UPS s.n., FH 00785696)
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 703
(fig. 92). The combination Tephrocybe cessans was proposed by Moser (1967: 116), who referred to
the first part of Symbolae ad Mycologiam Fennicam (Karsten 1870: 219) as the place of valid
publication of Agaricus cessans (cited as ‘Collybia cessans’) when making the combination in
Tephrocybe. Because Moser did not cite a type or make a full and direct citation to one and Karsten
(l.c.) clearly cites his earlier publication in Fungi Fenniae Exsiccati, none of the provisions of Art.
41.8 apply and this combination is invalidly published; it is validated here.
Fig. 92 (FH 00785696)

309
Cyphella punctiformis (Fr.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 714, 1868 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 221, 1870 [1871].
≡ Peziza punctiformis Fr., Syst. Mycol. 2(1): 105, 1822, nom. sanct. [Fr., l.c.].
≡ Flagelloscypha punctiformis (Fr.) Agerer, Sydowia 27: 246, 1975 [1973].
[Full synonymy, see https://www.mycobank.org/MB/314163]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 714 (fig.
93).
Fig. 93 (FH 00785707)
Cyphella alboviolascens (Alb. & Schwein.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 715, 1868 [Not.
Sällsk. Fauna Fl. Fenn. Förh. 11: 221, 1870 [1871].
≡ Peziza alboviolascens Alb. & Schwein. (as ‘albo-violascens’), Consp. Fung. Lusat.: 322, 1805, nom.
sanct. [Fr., Syst. Mycol. 2(1): 96, 1822].
≡ Lachnella alboviolascens (Alb. & Schwein.) Fr., Summa Veg. Scand. 2: 365, 1849.
[Full synonymy, see https://www.mycobank.org/MB/414590]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 715 (fig.
94). A lectotype was designated in Donk (1959).
Fig. 94 (FH 00785708)
Cyphella filicina P. Karst., Fungi Fenn. Exs., Cent. 8: no. 717, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 220, 1870 [1871].
≡ Nochascypha filicina (P. Karst.) Agerer, Mitt. Bot. Staatssamml. München 19: 268, 1983.
[Full synonymy, see https://www.mycobank.org/MB/108274]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 703
(fig. 95). Agerer (1983: 268) examined the specimen preserved in H (H 6005775) and called it the
holotype of that species. It should however rather be regarded as lectotype (see Art. 9.10.)
Fig. 95 (FH 00785710)
Cyphella abieticola P. Karst., Fungi Fenn. Exs., Cent. 8: no. 718, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 221, 1870 [1871], nom. illeg., Art. 53.1, homonym, non Cyphella abieticola P. Crouan & H.
Crouan, 1867.
≡ Chaetocypha abieticola Kuntze, Revis. Gen. Pl. 2: 847, 1891.
≡ Flagelloscypha abieticola (Kuntze) W.B. Cooke, Beih. Sydowia 4: 59, 1961.
[Full synonymy, see https://www.mycobank.org/MB/330897]
Notes: This is an illegitimate species name published on the label of Karsten, Fungi Fenn. Exs. 718
(fig. 96).

310
Fig. 96 (FH 00785711)
Peziza semiimmersa P. Karst., Fungi Fenn. Exs., Cent. 8: no. 724, 1868, nom. inval., Art. 38.1(a).
≡ Peziza semiimmersa P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 117, 1869 [Not. Sällsk. Fauna
Fl. Fenn. Förh. 11: 227, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 254, 1871].
≡ Sepultariella semiimmersa (P. Karst.) Van Vooren, U. Lindem. & Healy (as ‘semi-immersa’),
Ascomycete.org 9(4): 118, 2017.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 724 (fig.
97). It was validly published in Karsten (1869).
Fig. 97 (FH 00785717)
Peziza umbilicata P. Karst., Fungi Fenn. Exs., Cent. 8: no. 729, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 226, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 55, 1871], nom. illeg., Art.
53.1, homonym, non Peziza umbilicata Pers., 1822.
≡ Discina umbilicata Sacc., Syll. Fung. 8: 100, 1889.
≡ Galactinia umbilicata (Sacc.) Boud., Hist. Classif. Discomyc. Eur.: 48, 1907.
≡ Plicaria umbilicata (Sacc.) Bánhegyi, Borbásia 1(3–7): 85, 1939.
= Sporotrichum fulvum Fr., Syst. Mycol. 3(2): 418, 1832, nom. sanct. [Fr., l.c.].
≡ Chromelosporium fulvum (Fr.) McGinty, Hennebert & Korf, in Hennebert & Korf, Mycologia
67(2): 216, 1975.
[Full synonymy, see https://www.mycobank.org/MB/311086]
Notes: This is an illegitimate species name published on the label of Karsten, Fungi Fenn. Exs. 729
(fig. 98). Dissing & Korf (1978: 59) selected as lectotype no. 729 in the copy of Fungi Fenniae
Exsiccati held at K [K(M) 38214] and established the synonymy with Peziza ostracoderma, the
teleomorph of Chromelosporium fulvum.
https://ascomycete.org/de/2000-Pezizales/Art/Peziza-ostracoderma?mapid=449.
Fig. 98 (FH 00785722)
Peziza vitreola P. Karst., Fungi Fenn. Exs., Cent. 8: no. 731, 1868, nom. inval., Art. 38.1(a).
≡ Peziza vitreola P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 180, 1869.
≡ Helotium hyalinum subsp. vitreolum (P. Karst.) P. Karst. (as ‘* H. vitreolum’), Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 240, 1870 [1871].
≡ Helotium vitreolum (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 145, 1871.
≡ Hyaloscypha vitreola (P. Karst.) Boud., Icon. Mycol. Sér. 1(Livr. 2): [4], 1904.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 731 (fig.
99). It was validly published in Karsten (1869).
Fig. 99 (FH 00785724)

311
Peziza ventosa P. Karst., Fungi Fenn. Exs., Cent. 8: no. 733, 1868, nom. inval., Art. 38.1(a).
≡ Peziza ventosa P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 157, 1869.
≡ Trochila ventosa (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 244, 1870 [1871].
≡ Mollisia ventosa (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 188, 1871.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 733 (fig.
100). It was validly published in Karsten (1869) and later transferred to Mollisia (see also Dennis
1950).
Fig. 100 (FH 00785726)
Belonidium litoreum P. Karst., Fungi Fenn. Exs., Cent. 8: no. 737, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 213, 1870 [1871].
≡ Belonidium litoreum subsp. proteus P. Karst. (as ‘* proteus’), Not. Sa lsk. Fauna Fl. Fenn. Förh. 11:
242, 1870 [1871], nom. inval., Art. 26.2.
≡ Belonidium campanula var. litoreum (P. Karst.) P. Karst. (as ‘var. β. litoreum’), Bidrag Kan edom
Finlands Natur Folk 19: 163, 1871.
≡ Belonioscypha vexata var. litorea (P. Karst.) Rehm (as ‘littorea’), Rabenh. Krypt.-Fl. Ed. 2, Band 1
Abth. 3(Lief. 39): 746, 1893 [1896].
= Cyathicula culmicola (Desm.) S.E. Carp. & Dumont, Mycologia 70(6): 1231, [1978] 1979.
[Full synonymy, see https://www.mycobank.org/MB/247602]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 737
(fig. 101).
Fig. 101 (FH 00785730)
Peziza conformata P. Karst., Fungi Fenn. Exs., Cent. 8: no. 738, 1868, nom. inval., Art. 38.1(a).
≡ Peziza conformata P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 149, 1869.
≡ Helotium conformatum (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 236, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 124, 1871].
≡ Ciboria conformata (P. Karst.) Svrček, Česká Mykol. 36(3): 152, 1982.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 738 (fig.
102). It was validly published in Karsten (1869) and later transferred to Ciboria.
Fig. 102 (FH 00785731)
Peziza suspecta (Nyl.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 740, 1868.
≡ Peziza virgultorum var. suspecta Nyl., Observ. Peziz. Fenn.: 38, 1868.
≡ Hymenoscyphus suspectus (Nyl.) Hengstm., Persoonia 12(4): 489, 1985, nom. inval., Art. 41.8.
≡ Hymenoscyphus suspectus (Nyl.) Hengstm., comb. nov. MB848929.
[Basionym: Peziza virgultorum var. suspecta Nyl., Observ. Peziz. Fenn.: 38, 1868.]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 740 (fig.
103). Century 8 of Fungi Fenniae Exsiccati was issued at least after 1 October 1868 (the collecting
date of no. 701), after Nylander’s Observationes Circa Pezizas Fenniae was issued in September
1868. The combination Hymenoscyphus suspectus was proposed by Hengstmengel (1985: 489), who
cited as basionym the publication of Peziza suspecta on the label of Fungi Fenn. Exs. 740 when
making the combination in Hymenoscyphus. Because Hengstmengel did not cite a type or Latin

312
diagnosis or make a full and direct citation to either and Karsten indirectly references Nylander’s
earlier name and publication on the label, none of the provisions of Art. 41.8 apply and this
combination is invalidly published; it is validated here.
Fig. 103 (FH 00785733)
Peziza byssacea P. Karst., Fungi Fenn. Exs., Cent. 8: no. 741, 1868.
≡ Allophylaria byssacea (P. Karst.) P. Karst., Acta Soc. Fauna Fl. Fenn. 2(6): 131, 1885.
[Full synonymy, see https://www.mycobank.org/MB/543351]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 741
(fig. 104).
Fig. 104 (FH 00785734)
Peziza clavuliformis P. Karst., Fungi Fenn. Exs., Cent. 8: no. 742, 1868.
≡ Allophylaria clavuliformis (P. Karst.) P. Karst., Acta Soc. Fauna Fl. Fenn. 2(6): 131, 1885.
[Full synonymy, see https://www.mycobank.org/MB/212945]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 742
(fig. 105). A lectotype (H 6039309) was designated by Verkley (1999: 143).
Fig. 105 (FH 00785735)
Peziza nanella P. Karst., Fungi Fenn. Exs., Cent. 8: no. 743, 1868 [Not. Sällsk. Fauna Fl. Fenn. Förh.
11: 216, 1870 [1871].
≡ Ombrophila nanella (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 243, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 90, 1871].
[Full synonymy, see https://www.mycobank.org/MB/569644]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 743
(fig. 106).
Fig. 106 (FH 00785736)
Peziza fallens P. Karst., Fungi Fenn. Exs., Cent. 8: no. 744, 1868.
≡ Trochila fallens (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 217, 1870 [1871].
≡ Trochila cinerea subsp. fallens (P. Karst.) P. Karst. (as ‘** Tr. fallens’), Not. Sllsk. Fauna Fl. Fenn.
Förh. 11: 244, 1870 [1871].
≡ Mollisia cinerea subsp. fallens (P. Karst.) P. Karst. (as ‘** M. fallens’), Bidrag Kann dom Finlands
Natur Folk 19: 191, 1871.

313
≡ Mollisia fallens (P. Karst.) Sacc., Syll. Fung. 8: 332, 1889.
[Full synonymy, see https://www.mycobank.org/MB148100]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 744
(fig. 107).
Fig. 107 (FH 00785737)
Peziza paradoxa P. Karst., Fungi Fenn. Exs., Cent. 8: no. 746, 1868, nom. inval., Art. 38.1(a).
≡ Peziza paradoxa P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 155, 1869.
≡ Clibanites paradoxus (P. Karst.) P. Karst. (as ‘paradoxa’), Bidrag Knnedom Finlands Natur Folk
19: 167, 1871.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 746 (fig.
108). It was validly published in Karsten (1869) and later transferred to Clibanites.
Fig. 108 (FH 00785739)
Peziza graminea P. Karst., Fungi Fenn. Exs., Cent. 8: no. 747, 1868.
≡ Niptera palustris subsp. graminea (P. Karst.) P. Karst. (as ‘***** N. graminea’), Not. Sällsk. Fauna
Fl. Fenn. Förh. 11: 247, 1870 [1871].
≡ Mollisia graminea (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 199, 1871.
≡ Belonopsis graminea (P. Karst.) Sacc. & P. Syd., Syll. Fung. 16: 753, 1902.
[Full synonymy, see https://www.mycobank.org/MB/235827]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 747
(fig. 109). Dennis (1950: 182) designated a lectotype (H, [Finland,] Ml. [Mustiala], ad Calamagr.
[Calamagrostis] (?), 27 Sept. [September] 68 [1868], P.A. Karsten (as ‘Peziza junciseda’ but later
annotated ‘Belonopsis graminea (Karst.) Sacc. & Syd.’).
Fig. 109 (FH 00785740)
Peziza juncinella P. Karst., Fungi Fenn. Exs., Cent. 8: no. 748, 1868, nom. inval., Art. 38.1(a).
≡ Peziza junciseda subsp. juncinella P. Karst. (as ‘* Peziza juncinella’), Not. Sllsk. Fauna Fl. Fenn.
Förh. 10: 164, 1869.
≡ Niptera palustris subsp. juncinella (P. Karst.) P. Karst. (as ‘**** N. juncinella’), Not. Sllsk. Fauna
Fl. Fenn. Förh. 11: 247, 1870 [1871].
≡ Mollisia junciseda subsp. juncinella (P. Karst.) P. Karst. (as ‘* juncinella’), Bidrag Knnedom
Finlands Natur Folk 19: 198, 1871.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 748 (fig.
110). It was validly published at infraspecific rank in Karsten (1869). According to Dennis (1950:
182) it closely resembles Belonopsis junciseda but differs in having larger asci and spores.
Fig. 110 (FH 00785741)

314
Peziza personata P. Karst., Fungi Fenn. Exs., Cent. 8: no. 749, 1868, nom. inval., Art. 38.1(a).
≡ Peziza personata P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 154, 1869.
≡ Aleuriella personata (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 243, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 186, 1871].
Notes: This is an invalid species name published without description on the label of Karsten, Fungi
Fenn. Exs. 749 (fig. 111). It was validly published in Karsten (1869) and later transferred to
Aleuriella. A lectotype (UPS, Fungi Fenn. Exs. 749) was designated by Huhtinen 1994: 8).
Fig. 111 (FH 00785742)
Peziza aridula P. Karst., Fungi Fenn. Exs., Cent. 8: no. 750, 1868, nom. inval., Art. 38.1(a).
≡ Peziza aridula P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 148, 1869.
≡ Gorgoniceps aridula (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 247, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 185, 1871].
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 750 (fig.
112). It was validly published in Karsten (1869) and later transferred to Gorgoniceps.
Fig. 112 (FH 00785743)
Peziza sordidula P. Karst., Fungi Fenn. Exs., Cent. 8: no. 751, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 10: 170, 1869].
≡ Trochila sordidula (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 244, 1870 [1871].
≡ Mollisia sordidula (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 193, 1871.
≡ Pseudohelotium sordidulum (P. Karst.) Huhtinen, Karstenia 34: 6, 1994.
[Full synonymy, see https://www.mycobank.org/MB/362259]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 751
(fig. 113). A lectotype (H 6046681) was designated by Huhtinen (1994: 6).
Fig. 113 (FH 00785744)
Tympanis atra P. Karst., Fungi Fenn. Exs., Cent. 8: no. 755, 1868, nom. illeg. superfl., Art. 52.1.
≡ Lecanidion atrum Rabenh., Deutschl. Krypt.-Fl. 1: 342, 1844, nom. illeg. superfl., Art. 52.1.
≡ Lichen atratus Hedw., Descr. Micr.-Anal. Musc. Frond. 2: 61, tab. 21, fig. A, 1789.
≡ Patellaria atrata (Hedw.) Fr., Syst. Mycol. 2(1): 160, 1822, nom. sanct. [Fr., l.c.].
Notes: This is a replacement name validly but illegitimately published on the label of Karsten, Fungi
Fenn. Exs. 755 (fig. 114).
Fig. 114 (FH 00785747)
Tympanis bacillifera P. Karst., Fungi Fenn. Exs., Cent. 8: no. 756, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 252, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 229, 1871].
≡ Patellaria bacillifera (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 235, 1873.

315
≡ Pragmopora bacillifera (P. Karst.) Rehm, Rabenh. Krypt.-Fl. Ed. 2, Band 1 Abth. 3(Lief. 33): 340,
1890 [1896].
[Full synonymy, see https://www.mycobank.org/MB/433791]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 756
(fig. 115).
Fig. 115 (FH 00785748)
Ascobolus cinerellus P. Karst., Fungi Fenn. Exs., Cent. 8: no. 760, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 206, 1871].
≡ Peziza cinerella (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 51, 1871, nom.
illeg., Art. 53.1, homonym, non Peziza cinerella P. Crouan & H. Crouan, 1867.
[Full synonymy, see https://www.mycobank.org/MB/210561]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 760
(fig. 116).
Fig. 116 (FH 00785752)
Peziza fuscella P. Karst., Fungi Fenn. Exs., Cent. 8: no. 761, 1868.
≡ Trochila fuscella (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 246, 1870 [1871]
[Bidrag Kännedom Finlands Natur Folk 19: 248, 1871].
≡ Hysteropezizella fuscella (P. Karst.) Nannf., Nova Acta Regiae Soc. Sci. Upsal. Ser. 4, 8(2): 115,
1932.
[Full synonymy, see https://www.mycobank.org/MB/265634]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 761
(fig. 117).
Fig. 117 (FH 00785753)
Ascobolus crenulatus P. Karst., Fungi Fenn. Exs., Cent. 8: no. 763, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 202, 1871; Bidrag Kännedom Finlands Natur Folk 19: 77, 1871].
[Full synonymy, see https://www.mycobank.org/MB/248895]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 763
(fig. 118).
Fig. 118 (FH 00785755)

316
Peziza fiscella P. Karst., Fungi Fenn. Exs., Cent. 8: no. 764, 1868, nom. inval., Art. 38.1(a).
≡ Peziza fiscella P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 154, 1869.
≡ Vibrissea filisporia f. fiscella (P. Karst.) A. Sánchez, J. Agric. Univ. Puerto Rico 51(1): 87, 1967.
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 764 (fig.
119). It was validly published in Karsten (1869) and later reduced to a form of Vibrissea filisporia.
Fig. 119 (FH 00785756)
Stictis emergens P. Karst., Fungi Fenn. Exs., Cent. 8: no. 765, 1868 [Not. Sällsk. Fauna Fl. Fenn. Förh.
11: 254, 1870 [1871].
≡ Mollisia emergens (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 194, 1871.
≡ Diplonaevia emergens (P. Karst.) B. Hein, Sydowia 36: 84, 1983.
[Full synonymy, see https://www.mycobank.org/MB/115144]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 765
(fig. 120).
Fig. 120 (FH 00785757)
Lophium decipiens P. Karst., Fungi Fenn. Exs., Cent. 8: no. 767, 1868 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 261, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 23: 246, 1873].
≡ Mytilinidion decipiens (P. Karst.) Sacc. (as ‘Mytilidion’), Michelia 1(1): 55, 1877.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 767
(fig. 121).
Fig. 121 (FH 00785759)
Schmitzomia polycocca P. Karst., Fungi Fenn. Exs., Cent. 8: no. 768, 1868 [Bidrag Kännedom
Finlands Natur Folk 19: 241, 1871; see also Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 236, 1873].
≡ Stictis polycocca (P. Karst.) P. Karst., Acta Soc. Fauna Fl. Fenn. 2(6): 166, 1885.
[Full synonymy, see https://www.mycobank.org/MB/139830]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 768
(fig. 122).
Fig. 122 (FH 00785760)

317
Lophium laeviusculum P. Karst., Fungi Fenn. Exs., Cent. 8: no. 771, 1868 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 261, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 23: 246, 1873].
= Lophium mytilinellum Fr., Syst. Mycol. 2(2): 534, 1823, nom. sanct. [Fr., l.c.].
≡ Mytilinidion mytilinellum (Fr.) H. Zogg, Beitr. Kryptogamenfl. Schweiz 11(3): 106 & 108, 1962.
[Full synonymy, see https://www.mycobank.org/MB/120959]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 771
(fig. 123). Lohman (1939: 363) said of a specimen in Karsten’s herbarium (H 6057286) that it “may
be considered the type.” Because it is not clear that he definitely accepted it as such, he did not
achieve a lectotypification.
Fig. 123 (FH 00785763)
Hypoxylon reticulatum P. Karst., Fungi Fenn. Exs., Cent. 8: no. 773, 1868 [Bidrag Kännedom
Finlands Natur Folk 23: 43, 1873; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 236, 1873].
= Sphaeria hydnicola Schwein., J. Acad. Nat. Sci. Philadelphia 5(1): 14, 1825, nom. sanct. [Fr., Syst.
Mycol. 3(Index): 167, 1832].
≡ Nemania serpens var. hydnicola (Schwein.) Y.M. Ju & J.D. Rogers, Nova Hedwigia 74(1–2): 109,
2002.
[Full synonymy, see https://www.mycobank.org/MB/477332]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 773
(fig. 124). A lectotype [K(M) 68871] was designated by Granmo et al. (1999: 71).
Fig. 124 (FH 00785765)
Anthostoma fennicum P. Karst., Fungi Fenn. Exs., Cent. 8: no. 774, 1868.
≡ Phaeosperma fennicum (P. Karst.) P. Karst. (as ‘fennica’), Bidrag Knnedom Finlands Natur Folk
23: 54, 1873.
= Pseudovalsaria ferruginea (Nitschke) Rappaz, Mycol. Helv. 7(1): 159, 1995.
[Full synonymy, see https://www.mycobank.org/MB/363314]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 774
(fig. 125). Ju et al. (1996: 442) designated a lectotype (H 6052387).
Fig. 125 (FH 00785766)
Hypoxylon depauperatum P. Karst., Fungi Fenn. Exs., Cent. 8: no. 776, 1868, nom. inval., Art.
38.1(a).
≡ Hypoxylon pauperatum P. Karst., Enum. Fung. Lapponia: 211, 1866 [Bidrag Kännedom Finlands
Natur Folk 23: 42, 1873].
= Entoleuca mammata (Wahlenb.) J.D. Rogers & Y.M. Yu, Mycotaxon 59: 446, 1996.

318
[Full synonymy, see https://www.mycobank.org/MB/415807]
Notes: This is an invalid species name published on the label of Karsten, Fungi Fenn. Exs. 776 (fig.
126). It is probably just a lapsus calami for Hypoxylon pauperatum, published two years earlier, as the
collecting information is the same and Karsten (1873a: 42) later included this collection under that
name with no mention of the ‘depauperatum’ spelling.
Fig. 126 (FH 00785768)
Valsa sorbi subsp. padi (P. Karst.) P. Karst. (as ‘* V. padi’), Fungi Fenn. Exs., Cent. 8: no. 781, 1868.
≡ Valsa sorbi f. padi P. Karst., Fungi Fenn. Exs., Cent. 4: no. 355, 1866.
≡ Eutypella padi (P. Karst.) Sacc., Syll. Fung. 1: 147, 1882.
[Full synonymy, see https://www.mycobank.org/MB/150300].
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 781 (fig.
127).
Fig. 127 (FH 00785773)
Diatrypella pulvinata subsp. intermedia P. Karst. (as ‘* intermedia’), Fungi Fenn. Exs., Cent. 8: no.
782, 1868 [Bidrag Kännedom Finlands Natur Folk 23: 152, 1873].
≡ Diatrypella intermedia (P. Karst.) Sacc., Syll. Fung. 1: 204, 1882.
[Full synonymy, see https://www.mycobank.org/MB/218975].
Notes: This is a new subspecies name validly published on the label of Karsten, Fungi Fenn. Exs. 782
(fig. 128).
Fig. 128 (FH 00785774)
Calosphaeria ciliatula (Fr.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 787, 1868 [Bidrag Kännedom
Finlands Natur Folk 23: 156, 1873].
≡ Sphaeria ciliatula Fr., Syst. Mycol. 2(2): 406, 1823, nom. sanct. [Fr., l.c.].
[Full synonymy, see https://www.mycobank.org/MB/193344].
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 787 (fig.
129).
Fig. 129 (FH 00785779)
Calosphaeria obvoluta P. Karst., Fungi Fenn. Exs., Cent. 8: no. 788, 1868 [Bidrag Kännedom
Finlands Natur Folk 23: 158, 1873; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 239, 1873].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 788
(fig. 130).
Fig. 130 (FH 00785780)

319
Valsa millepunctata (Grev.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 789, 1868.
≡ Cryptosphaeria millepunctata Grev., Fl. Edin.: 360, 1824.
= Sphaeria eunomia Fr., Syst. Mycol. 2(2): 377, 1823, nom. sanct. [Fr., l.c.].
≡ Cryptosphaeria eunomia (Fr.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 212, 1870.
[Full synonymy, see https://www.mycobank.org/MB/355867].
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 789 (fig.
131). The combination was later also proposed by Nitschke (1870: 161).
Fig. 131 (FH 00785781)
Sporormia stercoris (DC.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 794, 1868.
≡ Sphaeria stercoris DC., in Lam. & DC., Fl. Franç. 2: 294, 1805, nom. sanct. [Fr., Elench. Fung. 2:
104, 1828].
≡ Sphaeria stercoraria β. stercoris (DC.) Fr., Syst. Mycol. 2(2): 456, 1823, nom. sanct. [Fr., l.c.].
≡ Bombardioidea stercoris (DC.) N. Lundq., Acta Univ. Upsal., Symb. Bot. Upsal. 20(1): 281, 1972.
[Full synonymy, see https://www.mycobank.org/MB/309833].
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 794 (fig.
132). The combination was also later proposed by Pirotta (1878: 142).
Fig. 132 (FH 00785786)
Lasiosphaeria canescens (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 8: no. 796, 1868 [Bidrag
Kännedom Finlands Natur Folk 23: 162, 1873].
≡ Sphaeria canescens Pers., Observ. Mycol. 1: 67, 1796, nom. sanct. [Fr., Syst. Mycol. 2(2): 448,
1823].
≡ Echinosphaeria canescens (Pers.) A.N. Mill. & Huhndorf, Mycol. Res. 108(1): 31, 2004, nom.
inval., Art. 41.8.
≡ Echinosphaeria canescens (Pers.) A.N. Mill. & Huhndorf, comb. nov. MB 848930.
[Basionym: Sphaeria canescens Pers., Observ. Mycol. 1: 67, 1796.]
[Full synonymy, see https://www.mycobank.org/MB/848930]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 796 (fig.
133). The combination Echinosphaeria canescens was proposed by Miller & Huhndorf (2004: 31),
who referred to Synopsis Methodica Fungorum (Persoon 1801: 72) as the place of valid publication of
Sphaeria canescens when making the combination in Echinosphaeria. Because Miller & Huhndorf did
not cite a type or make a full and direct citation to one and Persoon (l.c.) clearly cites his earlier
publication in Observationes Mycologicae (Persoon 1796: 67), none of the provisions of Art. 41.8
apply and this combination is invalidly published; it is validated here.
Fig. 133 (FH 00785788)
Cent. 9
Tremella juniperina P. Karst., Fungi Fenn. Exs., Cent. 9: no. 812, 1869 [Bidrag Kännedom Finlands
Natur Folk 37: 197, 1882], nom. illeg., Art. 53.1, homonym, non Tremella juniperina L. 1753.
≡ Tremella karstenii Hauerslev, Mycotaxon 72: 480, 1999.
[Full synonymy, see https://www.mycobank.org/MB/461017]

320
Notes: This is an illegitimate name introduced on the label of Karsten, Fungi Fenn. Exs. 812 (fig. 134).
A lectotype (H 6006127) was designated by Torkelsen (1978: 435).
Fig. 134 (FH 00785804)
Peziza euchroa P. Karst., Fungi Fenn. Exs., Cent. 9: no. 816, 1869 [(as ‘euchlora’), Not. Sllsk. Fauna
Fl. Fenn. Förh. 11: 228, 1870 [1871].
≡ Rhodotarzetta euchroa (P. Karst.) Bronckers, Ascomycete.org 11(4): 143, 2019, nom. inval., Art.
41.8.
≡ Rhodotarzetta euchroa (P. Karst.) Bronckers, comb. nov. MB 849032.
[Basionym: Peziza euchroa P. Karst., Fungi Fenn. Exs., Cent. 9: no. 816, 1869.]
Lectotype: [Finland,] Pellinsuo, Aug. [August] 1869, P. A. Karsten 3010 (H 6035382, designated by
Khare 2003: 450).
[Full synonymy, see https://www.mycobank.org/MB/849032]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 816
(fig. 135). The combination Rhodotarzetta euchroa was proposed by Bronckers (2019: 143), who
referred to the first part of Symbolae ad Mycologiam Fennicam (Karsten 1870: 228) as the place of
valid publication of Peziza euchroa when making the combination in Rhodotarzetta. Because
Bronckers did not cite a type or make a full and direct citation to one and Karsten (l.c.) clearly cites his
earlier publication in Fungi Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this
combination is invalidly published; it is validated here.
Fig. 135 (FH 00785808)
Peziza sylvatica P. Karst., Fungi Fenn. Exs., Cent. 9: no. 819, 1869 [Not. Sällsk. Fauna Fl. Fenn. Förh.
11: 226, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 52, 1871], nom. illeg., Art. 53.1,
homonym, non Peziza sylvatica R. Ludw. 1859.
[Full synonymy, see https://www.mycobank.org/MB/166548]
Notes: This is an illegitimate name introduced on the label of Karsten, Fungi Fenn. Exs. 819 (fig. 136).
Fig. 136 (FH 00785811)
Helotium procerum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 821, 1869 [Bidrag Kännedom Finlands
Natur Folk 19: 130, 1871; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 233, 1873].
[Full synonymy, see https://www.mycobank.org/MB/221042]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 821
(fig. 137).

321
Fig. 137 (FH 00785813)
Trochila atrata (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 822, 1869 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 244, 1870 [1871].
≡ Peziza atrata Pers., Syn. Meth. Fung. 2: 669, 1801, nom. sanct. [Fr., Syst. Mycol. 2(1): 148, 1822].
≡ Pyrenopeziza atrata (Pers.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 294, 1870.
[Full synonymy, see https://www.mycobank.org/MB/244643]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 822 (fig.
138).
Fig. 138 (FH 00785814)
Trochila thallophila P. Karst., Fungi Fenn. Exs., Cent. 9: no. 823, 1869 [Not. Sällsk. Fauna Fl. Fenn.
Förh. 11: 246, 1870 [1871].
≡ Unguiculariopsis thallophila (P. Karst.) W.Y. Zhuang, Mycotaxon 32: 62, 1988.
[Full synonymy, see https://www.mycobank.org/MB/133804]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 823
(fig. 139). A lectotype (H 6051985) was designated by Zhuang (1988: 63, as ‘holotype’).
Fig. 139 (FH 00785815)
Sphaeria hirtula P. Karst., Fungi Fenn. Exs., Cent. 9: no. 825, 1869 [Bidrag Kännedom Finlands Natur
Folk 23: 96, 1873].
≡ Chaetomastia hirtula (P. Karst.) Berl., Icon. Fung. 1(1): 39, 1890.
[Full synonymy, see https://www.mycobank.org/MB/431495]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 825
(fig. 140).
Fig. 140 (FH 00785817)
Lachnea P. Karst., Fungi Fenn. Exs., Cent. 9: no. 826, 1869, nom. inval., Art. 38.1(a).
≡ Peziza sect. Lachnea P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 123, 192, 1869.
≡ Lachnea (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 249, 1870 [1871], nom. illeg.,
Art. 53.1, homonym, non Lachnaea L. 1753 (Thymelaeaceae).
Notes: This is an invalid new genus name, a nomen nudum with no direct or indirect reference to a
potential basionym. It was later validated.
Lachnea brunneola (Desm.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 826, 1869, nom. inval., Art. 35.1.
≡ Peziza brunneola Desm., Pl. Crypt. N. France Ed. 1, Fasc. 24: no. 1156, 1842 [Ann. Sci. Nat., Bot.
Sér. 2, 17: 96, 1842].

322
≡ Brunnipila brunneola (Desm.) Baral, in Læssøe & Petersen, Index Fungorum 369: 1, 2018.
[Full synonymy, see https://www.mycobank.org/MB/555351]
Notes: This is an invalid new combination as the generic name was not yet validly published at that
time (fig. 141).
Fig. 141 (FH 00785818)
Peziza myrtillina P. Karst., Fungi Fenn. Exs., Cent. 9: no. 827, 1869.
≡ Pezicula myrtillina (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 165, 1871.
[Full synonymy, see https://www.mycobank.org/MB/181634]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 827
(fig. 142). Eriksson (1970: 52) designated a lectotype [K(M), Fungi Fenn. Exs. 827 ex BM].
Fig. 142 (FH 00785819)
Lachnea albotestacea (Desm.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 829, 1869, nom. inval., Art.
35.1.
≡ Peziza albotestacea Desm. (as ‘albo-testacea’), Ann. Sci. Nat., Bot. Sér. 2, 19: 368, 1843.
≡ Albotricha albotestacea (Desm.) Raitviir, Scripta Mycol. 1: 40, 1970.
[Full synonymy, see https://www.mycobank.org/MB/308447]
Notes: This is an invalid new combination as the generic name was not yet validly published at that
time (fig. 143).
Fig. 143 (FH 00785821)
Sphaeria perfidiosa P. Karst., Fungi Fenn. Exs., Cent. 9: no. 830, 1869.
≡ Gnomonia perfidiosa (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 119, 1873.
[Full synonymy, see https://www.mycobank.org/MB/847870]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 830
(fig. 144).
Fig. 144 (FH 00785822)
Niptera graminis (Desm.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 831, 1869 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 247, 1870 [1871].
≡ Peziza graminis Desm., Pl. Crypt. N. France Ed. 1, Fasc. 22: no. 1066, 1841 [Ann. Sci. Nat., Bot.
Sér. 2, 15: 133, 1842].
[Full synonymy, see https://www.mycobank.org/MB/653459]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 831 (fig.
145).
Fig. 145 (FH 00785823)

323
Helotium chrysophthalmum (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 832, 1869 [Not. Sällsk.
Fauna Fl. Fenn. Förh. 11: 242, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 155, 1871].
≡ Peziza chrysophthalma Pers. (as ‘chrysopthalma’), Mycol. Eur. 1: 259, 1822.
≡ Lachnellula chrysophthalma (Pers.) P. Karst., Meddeland. Soc. Fauna Fl. Fenn. 11: 138, 1884.
[Full synonymy, see https://www.mycobank.org/MB/179219]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 832 (fig.
146).
Fig. 146 (FH 00785824)
Phacidium epilobii P. Karst., Fungi Fenn. Exs., Cent. 9: no. 833, 1869.
≡ Trochila epilobii (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 250, 1871.
≡ Naeviopsis epilobii (P. Karst.) B. Hein, Willdenowia, Beih. 9: 71, 1976.
[Full synonymy, see https://www.mycobank.org/MB/318472]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 833
(fig. 147). A lectotype (UPS, Fungi Fenn. Exs. 833) was designated by Hein (1976: 71).
Fig. 147 (FH 00785825)
Orbilia rubella (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 834, 1869 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 248, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 102, 1871].
≡ Peziza rubella Pers., Syn. Meth. Fung. 2: 635, 1801, nom. sanct. [Fr., Syst. Mycol. 2(1): 141, 1822].
[Full synonymy, see https://www.mycobank.org/MB/219340]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 834 (fig.
148).
Fig. 148 (FH 00785826)
Lachnea callimorpha P. Karst., Fungi Fenn. Exs., Cent. 9: no. 835, 1869, nom. inval., Art. 35.1.
≡ Lachnea callimorpha P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 250, 1870 [1871].
≡ Lachnum callimorphum (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 19: 173, 1871.
[Full synonymy, see https://www.mycobank.org/MB/225135]
Notes: This is an invalid new combination as the generic name was not yet validly published at that
time (fig. 149). It was later validated.
Fig. 149 (FH 00785827)
Helotium cyathoideum (Bull.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 836, 1869 [Not. Sällsk. Fauna
Fl. Fenn. Förh. 11: 237, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 136, 1871], nom.
illeg., Art. 53.1, homonym, non Helotium cyathoideum De Not. 1864.
≡ Peziza cyathoidea Bull., Herb. France 9(104): Pl. 416, fig. 3, 1789, nom. sanct. [Fr., Syst. Mycol.
2(1): 124, 1822].

324
≡ Cyathicula cyathoidea (Bull.) Thüm., Fungi Austr. Exs., Cent. 12: no. 1115, 1874.
[Full synonymy, see https://www.mycobank.org/MB/207893]
Notes: This is a new combination validly but illegitimately published on the label of Karsten, Fungi
Fenn. Exs. 836 (fig. 150).
Fig. 150 (FH 00785828)
Helotium abietis P. Karst., Fungi Fenn. Exs., Cent. 9: no. 837, 1869 [Bidrag Kännedom Finlands
Natur Folk 19: 154, 1871; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 234, 1873].
≡ Lachnellula abietis (P. Karst.) Dennis, Persoonia 2(1): 183, 1962, nom. inval., Art. 41.8.
≡ Lachnellula abietis (P. Karst.) Dennis ex J.K. Mitch. & Bensch, comb. nov. MB 848931.
[Basionym: Helotium abietis P. Karst., Fungi Fenn. Exs., Cent. 9: no. 837, 1869.]
Lectotype: [Finland,] Mustiala, på grenar af Gran [branches of Picea abies], våren [spring] 1869, [P.
A. Karsten, Fungi Fenn. Exs. 837] (H 6003820, designated by Dharne 1965: 139).
[Full synonymy, see https://www.mycobank.org/MB/848931]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 837
(fig. 151). The combination Lachnellula abietis was proposed by Dennis (1962: 183), who referred to
the first part of Mycologia Fennica (Karsten 1871a: 154) as the place of valid publication of Helotium
abietis when making the combination in Lachnellula. Because Dennis did not cite a type or make a
full and direct citation to one and Karsten (l.c.) clearly cites his earlier publication in Fungi Fenniae
Exsiccati, none of the provisions of Art. 41.8 apply and this combination is invalidly published; we
validate it here.
Fig. 151 (FH 00785829)
Propolis sorbina P. Karst., Fungi Fenn. Exs., Cent. 9: no. 839, 1869.
≡ Karstenia sorbina (P. Karst.) Fr. ex P. Karst., Acta Soc. Fauna Fl. Fenn. 2(6): 166, 1885.
[Full synonymy, see https://www.mycobank.org/MB/180185]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 839
(fig. 152).
Fig. 152 (FH 00785831)
Tympanis amphibola (A. Massal.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 840, 1869 [Not. Sällsk.
Fauna Fl. Fenn. Förh. 11: 252, 1870 [1871]; Bidrag Kännedom Finlands Natur Folk 19: 230, 1871].
≡ Pragmopora amphibola A. Massal., Framm. Lichenogr.: 13, 1855.
[Full synonymy, see https://www.mycobank.org/MB/225873]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 840 (fig.
153).
Fig. 153 (FH 00785832)

325
Phacidium arctostaphyli P. Karst., Fungi Fenn. Exs., Cent. 9: no. 843, 1869 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 256, 1870 [1871], nom. illeg., Art. 52.1.
≡ Stictis phacidioides Fr., Syst. Mycol. 2(1): 198, 1822, nom. sanct. [Fr., l.c.].
≡ Marthamyces phacidioides (Fr.) Minter, Mycotaxon 87: 51, 2003.
[Full synonymy, see https://www.mycobank.org/MB/488897]
Notes: This is a superfluous replacement name validly published on the label of Karsten, Fungi Fenn.
Exs. 843 (fig. 154). The epithet was presumably changed to avoid a perceived tautonym.
Fig. 154 (FH 00785835)
Phacidium macrosporum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 844, 1869 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 256, 1870 [1871].
≡ Niptera bispora Baral, Acta Univ. Upsal., Symb. Bot. Upsal. 36(2): 194, 2014, nom. inval., Art.
41.8.
≡ Niptera bispora Baral, nom. nov. MB 849033.
[Replaced synonym: Phacidium macrosporum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 844, 1869.]
Syntypes: [Finland,] Mustiala, på blad af Car. vesic. [leaves of Carex vesicaria], Nov. [November],
[P.A. Karsten, Fungi Fenn. Exs. 844] (e.g., H 6052013, UPS s.n., FH 00785836, S F86697)
[Full synonymy, see https://www.mycobank.org/MB/849033]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 844
(fig. 155). The replacement name Niptera bispora was proposed by Baral in Eriksson (2014: 194),
who referred to the first part of Symbolae ad Mycologiam Fennicam (Karsten 1870: 256) as the place
of valid publication of Phacidium macrosporum when proposing the replacement name in Niptera.
Because Baral did not cite a type or make a full and direct citation to one and Karsten (l.c.) clearly
cites his earlier publication in Fungi Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and
this replacement name is invalidly published; it is validated here.
Fig. 155 (FH 00785836)
Trochila diminuens P. Karst., Fungi Fenn. Exs., Cent. 9: no. 851, 1869 [Bidrag Kännedom Finlands
Natur Folk 19: 248, 1871].
≡ Hysteropezizella diminuens (P. Karst.) Nannf., Nova Acta Regiae Soc. Sci. Upsal. Ser. 4 8(2): 114,
1932.
[Full synonymy, see https://www.mycobank.org/MB/263221]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 851
(fig. 156).
Fig. 156 (FH 00785843)

326
Sphaeria praetermissa P. Karst., Fungi Fenn. Exs., Cent. 9: no. 852, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 89, 1873].
≡ Paraleptosphaeria praetermissa (P. Karst.) Gruyter, Aveskamp & Verkley, in Gruyter et al., Stud.
Mycol. 75: 21, 2012 [2013], nom. inval., Art. 41.8.
≡ Paraleptosphaeria praetermissa (P. Karst.) Gruyter, Aveskamp & Verkley, comb. nov. MB
848932.
[Basionym: Sphaeria praetermissa P. Karst., Fungi Fenn. Exs., Cent. 9: no. 852, 1869.]
Syntypes: [Finland,] Mustiala, på Hallonörten [Rubus idaeus or Rubus odoratus], 17 Oct. [October]
1866, [P.A. Karsten, Fungi Fenn. Exs. 852] (e.g., H 6051811, UPS s.n., FH 00785844)
[Full synonymy, see https://www.mycobank.org/MB/848932]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 852
(fig. 157). The combination Paraleptosphaeria praetermissa was proposed by Gruyter et al. (2012:
121), who referred to the second part of Mycologia Fennica (Karsten 1873a: 89) as the place of valid
publication of Sphaeria praetermissa when making the combination in Paraleptosphaeria. Because
Gruyter et al. did not cite a type or make a full and direct citation to one and Karsten (l.c.) clearly cites
his earlier publication in Fungi Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this
combination is invalidly published; it is validated here.
Fig. 157 (FH 00785844)
Sphaeria emergens P. Karst., Fungi Fenn. Exs., Cent. 9: no. 853, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 93, 1873], nom. illeg., Art. 53.1, homonym, non Sphaeria emergens Schwein. 1832.
≡ Zignoella emergens Sacc., Michelia 1(3): 346, 1878.
≡ Keissleriella emergens (Sacc.) S.K. Bose, Phytopathol. Z. 41(2): 191, 1961.
[Full synonymy, see https://www.mycobank.org/MB/332719]
Notes: This is an illegitimate name published on the label of Karsten, Fungi Fenn. Exs. 853 (fig. 158).
Aptroot (1998: 149) designated a ‘lectotype’ (H 6052586), but the specimen so designated was
collected in 1870, the year after the publication of the name Sphaeria emergens P. Karst., and so is not
original material; this typification must be corrected to designation of a neotype. Original material
does still exist, though, so this type may be superseded.
Fig. 158 (FH 00785845)
Ceratostoma depressulum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 854, 1869.
≡ Gnomonia depressula (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 121, 1873.
≡ Sydowiella depressula (P. Karst.) M.E. Barr, Mycol. Mem. 7: 159, 1978.
≡ Sydowiella depressula (P. Karst.) M.E. Barr ex J.K. Mitch. & Bensch, comb. nov. MB 848933.
[Basionym: Ceratostoma depressulum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 854, 1869.]
Syntypes: [Finland,] Mustiala, på Rub. idaeus [Rubus idaeus], [P. A. Karsten, Fungi Fenn. Exs. 854]
(e.g., H 6052027, UPS s.n., FH 00785846, S F109228).
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 854
(fig. 159). The combination Sydowiella depressula was proposed by Barr (1978: 159), who referred to
the publication of Gnomonia depressula in the second part of Mycologia Fennica (Karsten 1873a:
121) as the basionym when making the combination in Sydowiella. Because Barr did not cite a type or
make a full and direct citation to one and Karsten (l.c.) clearly cites his earlier publication in Fungi

327
Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this combination is invalidly
published; we validate it here.
Fig. 159 (FH 00785846)
Cucurbitaria pulveracea P. Karst., Fungi Fenn. Exs., Cent. 9: no. 855, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 62, 1873].
[Full synonymy, see https://www.mycobank.org/MB/216186]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 855
(fig. 160).
Fig. 160 (FH 00785847)
Calosphaeria idaeicola P. Karst., Fungi Fenn. Exs., Cent. 9: no. 856, 1869.
≡ Gnomonia idaeicola (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 126, 1873.
≡ Gnomoniopsis idaeicola (P. Karst.) D.M. Walker, in Walker et al., Mycologia 102(6): 1490, 2010.
[Full synonymy, see https://www.mycobank.org/MB/515711]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 856
(fig. 161). Walker et al. (2010: 1492) examined original material of this species that they considered a
‘holotype’ (FH 00785848). Because of a lack of a statement indicating designation in that publication,
this cannot be corrected to ‘lectotype’ and considered a lectotypification (Art. 7.11). As such, the
‘epitype’ they designated (BPI 879157) does not have a designated holotype, lectotype, or neotype
which it supports, and has no standing.
Fig. 161 (FH 00785848)
Valsa ribesia P. Karst., Fungi Fenn. Exs., Cent. 9: no. 857, 1869 [Bidrag Kännedom Finlands Natur
Folk 23: 138, 1873].
[Full synonymy, see https://www.mycobank.org/MB/206791]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 857
(fig. 162).

328
Fig. 162 (FH 00785849)
Hysterographium conjungens P. Karst., Fungi Fenn. Exs., Cent. 9: no. 858, 1869 [Not. Sällsk. Fauna
Fl. Fenn. Förh. 11: 259, 1870 [1871].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 858
(fig. 163). A lectotype (H 6052569) was designated by Lohman (1939: 358).
Fig. 163 (FH 00785850)
Ceratostoma subrostratum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 859, 1869.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 859
(fig. 164).
Fig. 164 (FH 00785851)
Anthostoma microsporum P. Karst., Fungi Fenn. Exs., Cent. 9: no. 860, 1869.
≡ Phaeosperma microsporum (P. Karst.) P. Karst. (as ‘microspora’), Bidrag Kan edom Finlands
Natur Folk 23: 54, 1873 [Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 240, 1873].
≡ Camaropella microspora (P. Karst.) S.K. Huang & K.D. Hyde, in Huang et al., Fungal Diversity
111: 474, 2021.
[Full synonymy, see https://www.mycobank.org/MB/558249]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 860
(fig. 165). A lectotype (H 6052040) was designated by Nannfeldt (1972: 364, as ‘holotype’).
Fig. 165 (FH 00785852)
Sphaeria crustacea P. Karst., Fungi Fenn. Exs., Cent. 9: no. 865, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 95, 1873], nom. illeg., Art. 53.1, homonym, non Sphaeria crustacea Sowerby 1803.
≡ Zignoella crustacea Sacc., Syll. Fung. 2: 220, 1883.

329
≡ Chaetosphaeria crustacea (Sacc.) Réblová & W. Gams, Czech Mycol. 51(1): 12, 1999.
Notes: This is an illegitimate name published on the label of Karsten, Fungi Fenn. Exs. 865 (fig. 166).
Réblová and Gams (1999: 17) designated a lectotype (H 6034088).
Fig. 166 (FH 00785857)
Gnomonia alniella P. Karst., Fungi Fenn. Exs., Cent. 9: no. 867, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 119, 1873; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 223, 1873].
≡ Apiognomonia alniella (P. Karst.) Höhn., Hedwigia 62: 49, 1920 [1921].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 867
(fig. 167).
Fig. 167 (FH 00785859)
Sphaerella macularis (Fr.) Auersw., in Gonn. & Rabenh., Mycol. Eur. 5–6: 18, 1869 [Karsten, Fungi
Fenn. Exs., Cent. 9: no. 868, 1869].
≡ Sphaeria macularis Fr., Observ. Mycol. 1: 186, 1815, nom. sanct. [Fr., Syst. Mycol. 2(2): 502,
1823].
≡ Venturia macularis (Fr.) E. Müll. & Arx, Ber. Schweiz. Bot. Ges. 60: 366, 1950.
[Full synonymy, see https://www.mycobank.org/MB/307715]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 868 (fig.
168). It was, however, published in the same year by Auerswald in Gonnermann & Rabenhorst (1869:
18) and it is unclear which has priority.
Fig. 168 (FH 00785860)
Gnomonia comari P. Karst., Fungi Fenn. Exs., Cent. 9: no. 869, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 122, 1873].
≡ Gnomonia chamaemori subsp. comari (P. Karst.) P. Karst. (as ‘Gnomonia Chamaemori * Gn.
Comari’), Not. Sa lsk. Fauna Fl. Fenn. Förh. 13: 222, 1873.
≡ Gnomoniopsis comari (P. Karst.) Sogonov, in Sogonov et al., Stud. Mycol. 62: 47, 2008, nom.
inval., Art. 41.8.
≡ Gnomoniopsis comari (P. Karst.) Sogonov, in Walker et al., Mycologia 102(6): 1487, 2010.
[Full synonymy, see https://www.mycobank.org/MB/848238]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 869
(fig. 169). A lectotype [K(M) 157111] was designated by Walker et al. (2010: 1489). The combination
Gnomoniopsis comari was proposed by Sogonov in Sogonov et al. (2008: 47), who referred to the
publication of Gnomonia comari in the second part of Mycologia Fennica (Karsten 1873a: 122) as the
basionym when making the combination in Gnomoniopsis. Because Sogonov did not cite a type or
make a full and direct citation to one and Karsten (l.c.) clearly cites his earlier publication in Fungi

330
Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this combination is invalidly
published. The name was inadvertently validated by Walker et al. (2010: 1487) when they listed the
name, again citing Karsten’s later Latin description as the place of valid publication of Gnomonia
comari, but also designated a lectotype, giving the name all necessary components for publication of a
new species and thus satisfying Art. 41.8(d).
Fig. 169 (FH 00785861)
Nectria cinnabarina subsp. tiliae P. Karst. (as ‘* tiliae’), Fungi Fenn. Exs., Cent. 9: no. 870, 1869
[Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 241, 1873].
≡ Nectria cinnabarina var. tiliae (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 213,
1873.
Notes: This is a new subspecies name validly published on the label of Karsten, Fungi Fenn. Exs. 870
(fig. 170).
Fig. 170 (FH 00785862)
Sphaeria salicaria P. Karst., Fungi Fenn. Exs., Cent. 9: no. 871, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 95, 1873].
[Full synonymy, see https://www.mycobank.org/MB/179930]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 871
(fig. 171).
Fig. 171 (FH 00785863)
Diaporthe alniella P. Karst., Fungi Fenn. Exs., Cent. 9: no. 872, 1869.
≡ Enchnoa alniella (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 198, 1873.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 872
(fig. 172).
Fig. 172 (FH 00785864)
Diaporthe transversalis P. Karst., Fungi Fenn. Exs., Cent. 9: no. 873, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 113, 1873].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 873
(fig. 173).

331
Fig. 173 (FH 00785865)
Sphaeria corylina P. Karst., Fungi Fenn. Exs., Cent. 9: no. 874, 1869.
≡ Otthia corylina (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 59, 1873.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 874
(fig. 174).
Fig. 174 (FH 00785866)
Sphaeria pygmaea P. Karst., Fungi Fenn. Exs., Cent. 9: no. 875, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 93, 1873].
≡ Gongromeriza pygmaea (P. Karst.) Réblová, Stud. Mycol. 103(1): 199, 2022.
[Full synonymy, see https://www.mycobank.org/MB/846731]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 875
(fig. 175).
Fig. 175 (FH 00785867)
Hypomyces bombycinus P. Karst. (as ‘bombacinus’), Fungi Fenn. Exs., Cent. 9: no. 877, 1869, nom.
inval., Art. 38.1(a).
Notes: This is an invalidly published name, a nomen nudum (fig. 176). Apparently intended as the
publication of a teleomorph, as Karsten attributed the name to himself and listed Institale bombycina
as the ‘forma conidiifera’. Karsten (1873a: 210 & 1873b: 243) later introduced the combination
Hypomyces bombycinus (Fr. ex Weinm.) P. Karst. with the statements ‘perithecium nondum inventa’
and ‘fungus ascophorus ignotus,’ suggesting that he changed his mind about finding a teleomorph.
Rogerson & Samuel (1993: 248–9) suggest that this is the modified subiculum of the species they call
Hypomyces chrysostomus (≡ Sporophagomyces chrysostomus).
Fig. 176 (FH 00785869)
Hypocrea atrata P. Karst., Fungi Fenn. Exs., Cent. 9: no. 878, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 207, 1873; Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 244, 1873].
= Trichoderma citrinoviride Bissett, Canad. J. Bot. 62(5): 926, 1984, nom. prot.
[Full synonymy, see https://www.mycobank.org/MB/107345]

332
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 878
(fig. 177). Samuels (2014: 936–7) proposed the conservation of Trichoderma citrinoviride against a
number of teleomorph-typified names but did not include Hypocrea atrata explicitly due to the
mistaken assumption that it was first published in the second part of Mycologia Fennica by Karsten
(1873a: 207), making the name appear to be an illegitimate superfluous renaming of Hypocrea
repanda Fuckel. As the proposal was modified from conservation to protection, Trichoderma
citrinoviride has to be treated as conserved against any competing listed or unlisted synonyms (Art.
F.2.1).
Fig. 177 (FH 00785870)
Cucurbitaria rubicola P. Karst., Fungi Fenn. Exs., Cent. 9: no. 879, 1869.
≡ Fenestella princeps subsp. rubicola (P. Karst.) P. Karst. (as ‘Fenestrella princeps * F. rubicola’),
Bidrag Kännedom Finlands Natur Folk 23: 63, 1873.
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 879
(fig. 178).
Fig. 178 (FH 00785871)
Amphisphaeria sapinea P. Karst., Fungi Fenn. Exs., Cent. 9: no. 880, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 56, 1873].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 880
(fig. 179).
Fig. 179 (FH 00785872)
Diplodia sapinea (Fr.) P. Karst., Fungi Fenn. Exs., Cent. 9: no. 880, 1869.
≡ Sphaeria sapinea Fr., Syst. Mycol. 2(2): 491, 1823, nom. sanct. [Fr., l.c.].
[Full synonymy, see https://www.mycobank.org/MB/146913]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 880 (fig.
179). It was later also proposed by Fuckel (1870: 393).
Rebentischia P. Karst., Fungi Fenn. Exs., Cent. 9: no. 881, 1869 [Bidrag Kännedom Finlands Natur
Folk 23: 14, 1873].
Notes: This is a new genus name validly published on the label of Karsten, Fungi Fenn. Exs. 881 (fig.
180).
Fig. 180 (FH 00785873)

333
Rebentischia pomiformis P. Karst., Fungi Fenn. Exs., Cent. 9: no. 881, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 97, 1873].
= Rebentischia massalongoi (Mont.) Sacc. (as ‘Massalongi’), Consp. Gen. Pyrenomyc: 12, 1875.
[Full synonymy, see https://www.mycobank.org/MB/533330]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 881
(fig. 180). Boonmee et al. (2014: 287) examined original material of this species that they considered a
‘holotype’ (FH 00785873; the reference to the ‘Herbarium of F. Theissen’ is apparently an error).
Because of a lack of a statement indicating designation in that publication, this cannot be corrected to
‘lectotype’ and considered a lectotypification (Art. 7.11).
Pleospora eleocharidis P. Karst. (as ‘heleocharidis’), Fungi Fenn. Exs., Cent. 9: no. 882, 1869 [Bidrag
Kännedom Finlands Natur Folk 23: 72, 1873].
[Full synonymy, see https://www.mycobank.org/MB/529176]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 882
(fig. 181).
Fig. 181 (FH 00785874)
Sphaeria fennica P. Karst., Fungi Fenn. Exs., Cent. 9: no. 883, 1869.
≡ Melanconis fennica (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 75, 1873.
[Full synonymy, see https://www.mycobank.org/MB/847901]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 883
(fig. 182).
Fig. 182 (FH 00785875)
Lophiostoma crenatum subsp. viburni P. Karst. (as ‘* viburni’), Fungi Fenn. Exs., Cent. 9: no. 885,
1869.
Notes: This is a new subspecies name validly published on the label of Karsten, Fungi Fenn. Exs. 885
(fig. 183).
Fig. 183 (FH 00785877)
Lophiostoma crenatum subsp. rosae P. Karst. (as ‘* rosae’), Fungi Fenn. Exs., Cent. 9: no. 886, 1869.
Notes: This is a new subspecies name validly published on the label of Karsten, Fungi Fenn. Exs. 886
(fig. 184).
Fig. 184 (FH 00785878)

334
Lasiosphaeria heterostoma P. Karst., Fungi Fenn. Exs., Cent. 9: no. 887, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 162, 1873].
≡ Echinosphaeria heterostoma (P. Karst.) Huhndorf & A.N. Mill., in Miller et al., Mycologia 106(3):
509, 2014.
[Full synonymy, see https://www.mycobank.org/MB/805973]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 887
(fig. 185). Miller et al. (2014: 509) examined original material of this species that they considered a
‘holotype’ (H 6034100). Because of a lack of a statement indicating designation in that publication,
this cannot be corrected to ‘lectotype’ and considered a lectotypification (Art. 7.11).
Fig. 185 (FH 00785879)
Leptosphaeria ptarmicae P. Karst., Fungi Fenn. Exs., Cent. 9: no. 888, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 102, 1873].
[Full synonymy, see https://www.mycobank.org/MB/197145]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 888
(fig. 186).
Fig. 186 (FH 00785880)
Calosphaeria consobrina P. Karst., Fungi Fenn. Exs., Cent. 9: no. 890, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 158, 1873].
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 890
(fig. 187).
Fig. 187 (FH 00785882)
Sphaeria holmskjoldii P. Karst. (as ‘Holmskjöldi’), Fungi Fenn. Exs., Cent. 9: no. 891, 1869 [(as
‘Holmskjoeldii’), Bidrag Kann dom Finlands Natur Folk 23: 91, 1873].
≡ Leptosphaeria holmskjoldii (P. Karst.) Chi Y. Chen & W.H. Hsieh (as ‘holmskjoeldii’), in Chen et
al., I.M.I. Descript. Fungi Bact. 154(1535): [1], 2002.
[Full synonymy, see https://www.mycobank.org/MB/521594]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 891
(fig. 188).
Fig. 188 (FH 00785883)

335
Cucurbitaria chevalieri P. Karst., Fungi Fenn. Exs., Cent. 9: no. 892, 1869.
≡ Teichospora chevalieri (P. Karst.) P. Karst. (as ‘Chevalierii’), Bidrag Kan edom Finlands Natur
Folk 23: 66, 1873.
[Full synonymy, see https://www.mycobank.org/MB/847906]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 892
(fig. 189).
Fig. 189 (FH 00785884)
Amphisphaeria pusiola P. Karst., Fungi Fenn. Exs., Cent. 9: no. 893, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 57, 1873].
[Full synonymy, see https://www.mycobank.org/MB/225752]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 893
(fig. 190). A lectotype (H 6057316) was designated by Aptroot (1995: 108). According to Aptroot
(l.c.) this is a species of Chaetosphaeria.
Fig. 190 (FH 00785885)
Cucurbitaria pomiformis P. Karst., Fungi Fenn. Exs., Cent. 9: no. 894, 1869.
≡ Teichospora pomiformis (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 67, 1873.
[Full synonymy, see https://www.mycobank.org/MB/847907]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 894
(fig. 191).
Fig. 191 (FH 00785886)
Sphaerella canescens P. Karst., Fungi Fenn. Exs., Cent. 9: no. 895, 1869.
≡ Venturia chlorospora var. canescens (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk
23: 190, 1873.
?= Venturia chlorospora (Ces.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 957, 1870.
[Full synonymy, see https://www.mycobank.org/MB/179512]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 895
(fig. 192).
Fig. 192 (FH 00785887)

336
Leptosphaeria affinis P. Karst., Fungi Fenn. Exs., Cent. 9: no. 896, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 102, 1873].
[Full synonymy, see https://www.mycobank.org/MB/162560]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 896
(fig. 193).
Fig. 193 (FH 00785888)
Sphaerella proximella P. Karst., Fungi Fenn. Exs., Cent. 9: no. 897, 1869 [Bidrag Kännedom Finlands
Natur Folk 23: 177, 1873].
[Full synonymy, see https://www.mycobank.org/MB/151084]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 897
(fig. 194). A lectotype (H 6052048) was designated by Corlett & Smith (1978: 2822). Aptroot (2006:
167) states that this is morphologically indistinguishable from Davidiella allicina (≡ Cladosporium
allicinum), but Bensch et al. (2012: 52) indicate that Aptroot’s (l.c.) concept of this species is mixed.
Fig. 194 (FH 00785889)
Sphaerella chamaemori P. Karst., Fungi Fenn. Exs., Cent. 9: no. 899, 1869 [Bidrag Kännedom
Finlands Natur Folk 23: 173, 1873].
≡ Venturia chamaemori (P. Karst.) Arx, Acta Bot. Neerl. 6(3): 340, 1957.
[Full synonymy, see https://www.mycobank.org/MB/307713]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 899
(fig. 195).
Fig. 195 (FH 00785891)
Cent. 10
Lachnum hystriculum P. Karst., Fungi Fenn. Exs., Cent. 10: no. 921, 1870 [Bidrag Kännedom
Finlands Natur Folk 19: 182, 1871].
≡ Trichopezizella nidulus var. hystricula (P. Karst.) J.H. Haines, Mycologia 66(2): 222, 1974.
[Full synonymy, see https://www.mycobank.org/MB/218499]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 921
(fig. 196). A lectotype (H 6060182) was designated by Haines (1974: 223) from material not issued in
the set.

337
Fig. 196 (FH 00785913)
Peziza laeticolor P. Karst., Fungi Fenn. Exs., Cent. 10: no. 924, 1870 [Bidrag Kännedom Finlands
Natur Folk 19: 66, 1871].
≡ Spooneromyces laeticolor (P. Karst.) T. Schumach. & J. Moravec, Nordic J. Bot. 9(4): 427, 1989.
[Full synonymy, see https://www.mycobank.org/MB/125912]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 924
(fig. 197).
Fig. 197 (FH 00785916)
Allophylaria (P. Karst.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 927, 1870, nom. inval., Art. 41.1.
≡ Peziza sect. Allophylaria P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 10: 103, 147, 1869.
≡ Allophylaria (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 243, 1870 [1871], nom.
inval., Art. 41.1.
≡ Allophylaria (P. Karst.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 234, 1873.
[Full synonymy, see https://www.mycobank.org/MB/141]
Notes: Invalid name at new rank, no direct or indirect reference to the basionym or description (fig.
198).
Fig. 198 (FH 00785919)
Allophylaria eucrita (Nyl.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 927, 1870, nom. inval., Art. 35.1.
≡ Peziza eucrita Nyl., Observ. Peziz. Fenn.: 47, 1868.
≡ Allophylaria eucrita (Nyl.) P. Karst., Not. Sällsk. Fauna Fl. Fenn. Förh. 11: 243, 1870 [1871], nom.
inval., Art. 35.1, 41.1.
= Pezicula abietina (Auersw.) U. Braun & Bensch, Schlechtendalia 38: 194, 2021.
[Full synonymy, see https://www.mycobank.org/MB/839521]
Notes: This is an invalid new combination as the generic name was not yet validly published at that
time (fig. 198). While often credited to Karsten, authorship of Peziza eucrita should instead be
credited to Nylander since he did not ascribe the validating description to Karsten (Art. 46.5).
Additionally, despite ascribing the name to Karsten on page 47, Nylander (1868) indicated in the
footnote on page 48 that, finding the name Karsten proposed to be aesthetically displeasing, he
substituted one of his own choosing without Karsten’s knowledge.
Crumenula ribis (Fr.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 929, 1870 [Not. Sällsk. Fauna Fl.
Fenn. Förh. 11: 251, 1870 [1871].
≡ Cenangium ribis Fr., Syst. Mycol. 2: 179, 1822, nom. sanct. [Fr., l.c.].
≡ Godronia ribis (Fr.) Seaver, Mycologia 37(3): 339, 1945.
[Full synonymy, see https://www.mycobank.org/MB/286718]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 929 (fig.
199).

338
Fig. 199 (FH 00785921)
Schmitzomia luzulae subsp. junci P. Karst. (as ‘* junci’), Fungi Fenn. Exs., Cent. 10: no. 931, 1870
[Bidrag Kännedom Finlands Natur Folk 19: 238, 1871].
= Stictis graminicola Lasch, Klotzschii Herb. Viv. Mycol., Ed. Nova, Cent. 8: no. 713, 1858.
[Full synonymy, see https://www.mycobank.org/MB/240798]
Notes: This is a new subspecies name validly published on the label of Karsten, Fungi Fenn. Exs. 931
(fig. 200).
Fig. 200 (FH 00785923)
Schmitzomia elevata P. Karst., Fungi Fenn. Exs., Cent. 10: no. 932, 1870 [Bidrag Kännedom Finlands
Natur Folk 19: 240, 1871].
≡ Stictis elevata (P. Karst.) P. Karst., Acta Soc. Fauna Fl. Fenn. 2(6): 166, 1885.
[Full synonymy, see https://www.mycobank.org/MB/219951]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 932
(fig. 201).
Fig. 201 (FH 00785924)
Trochila trifolii (Biv.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 933, 1870 [Bidrag Kännedom
Finlands Natur Folk 19: 246, 1871].
≡ Ascobolus trifolii Biv., Stirp. Rar. Sicilia 4: 27, [1816] 1818, nom. sanct. [Fr., Syst. Mycol. 2: 165,
1822].
≡ Pseudopeziza trifolii (Biv.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 290, 1870.
[Full synonymy, see https://www.mycobank.org/MB/159987]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 933 (fig.
202). This combination is sometimes attributed to De Notaris (1864: 358), but he merely says that
Ascobolus trifolii is a Trochila without definitely associating the epithet with the new genus.
Fig. 202 (FH 00785925)
Valsa prunastri var. betulae (Sommerf.) P. Karst. (as ‘β Betulae’), Fungi Fenn. Exs., Cent. 10: no.
937, 1870.
≡ Sphaeria prunastri var. betulae Sommerf. (as ‘β. Betulae’), Suppl. Fl. Lapp.: 208, 1826.
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 937 (fig.
203).

339
Fig. 203 (FH 00785929)
Delitschia chaetomioides P. Karst., Fungi Fenn. Exs., Cent. 10: no. 939, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 60, 1873].
[Full synonymy, see https://www.mycobank.org/MB/176788]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 939
(fig. 204). A lectotype (K, Fungi Fenn. Exs. 939) was designated by Luck-Allen & Cain (1975: 1834).
Fig. 204 (FH 00785931)
Rhynchostoma P. Karst., Fungi Fenn. Exs., Cent. 10: no. 950, 1870 [Bidrag Kännedom Finlands
Natur Folk 23: 7, 1873].
[Full synonymy, see https://www.mycobank.org/MB/4734]
Notes: This is a new genus name validly published on the label of Karsten, Fungi Fenn. Exs. 950 (fig.
205).
Fig. 205 (FH 00785942)
Rhynchostoma minutum P. Karst., Fungi Fenn. Exs., Cent. 10: no. 950, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 58, 1873].
[Full synonymy, see https://www.mycobank.org/MB/166496]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 950
(fig. 205). Mathiassen (1993: 147) designated a lectotype (H 6052127).
Ixodopsis P. Karst., Fungi Fenn. Exs., Cent. 10: no. 955, 1870 [Bidrag Kännedom Finlands Natur Folk
23: 6, 1873].
≡ Sordaria Ces. & De Not., Comment. Soc. Crittog. Ital. 1(4): 225, 1863, nom. cons.
[Full synonymy, see https://www.mycobank.org/MB/5061]
Notes: This is a new genus name validly published on the label of Karsten, Fungi Fenn. Exs. 955 (fig.
206). When Sordaria was conserved with Sordaria fimicola as type, it became superfluous since they
share the same type.
Fig. 206 (FH 00785947)

340
Ixodopsis fimicola (Roberge ex Desm.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 955, 1870 [Bidrag
Kännedom Finlands Natur Folk 23: 50, 1873].
≡ Sphaeria fimicola Roberge ex Desm., Ann. Sci. Nat., Bot. Sér. 3, 11: 353, 1849.
≡ Sordaria fimicola (Roberge ex Desm.) Ces. & De Not., Comment. Soc. Crittog. Ital. 1(4): 226,
1863.
[Full synonymy, see https://www.mycobank.org/MB/147988]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 955 (fig.
206).
Venturia chlorospora (Ces.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 957, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 189, 1873].
≡ Sphaeria chlorospora Ces., in Rabenh., Fungi Eur. Exs., Cent. 1: no. 48, 1859.
[Full synonymy, see https://www.mycobank.org/MB/199040]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 957 (fig.
207).
Fig. 207 (FH 00785949)
Sphaerella vulgaris P. Karst., Fungi Fenn. Exs., Cent. 10: no. 960, 1870 [Bidrag Kännedom Finlands
Natur Folk 23: 168, 1873], nom. illeg., Art. 52.1.
[Full synonymy, see https://www.mycobank.org/MB/499880]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 961
(fig. 208) by reference to the validating descriptions of three previously published species (Sphaeria
sparsa Wallr., Sphaeria maculiformis Pers., and Sphaeria conglomerata Wallr.), the epithet of one of
which ought to have been used instead. Each of these species had already been legitimately combined
into Sphaerella, making this name even more superfluous. Sphaerella vulgaris was considered a
replacement name for Sphaeria maculiformis by Aptroot (2006: 126), but it is not clear that it was
Karsten’s intent to found Sphaerella vulgaris on Sphaeria maculiformis in particular. Without a
typification, its application is unclear.
Fig. 208 (FH 00785952)
Leptosphaeria culmifida P. Karst., Fungi Fenn. Exs., Cent. 10: no. 961, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 103, 1873].
≡ Keissleriella culmifida (P. Karst.) S.K. Bose, Phytopathol. Z. 41(2): 188, 1961, nom. inval., Art.
41.8.
≡ Keissleriella culmifida (P. Karst.) S.K. Bose ex J.K. Mitch. & Bensch, comb. nov. MB 848934.
[Basionym: Leptosphaeria culmifida P. Karst., Fungi Fenn. Exs., Cent. 10: no. 961, 1870.]
Syntypes: [Finland,] Mustiala, på timotejstrån [Phleum pratense straw], Okt. [October] [P. A. Karsten,
Fungi Fenn. Exs. 961] (e.g., H 6052142, UPS s.n., FH 00785953)
[Full synonymy, see https://www.mycobank.org/MB/848934]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 961
(fig. 209). While currently in use (e.g., Tanaka et al. 2015), the name Keissleriella culmifida has not
yet been validly published. Bose (1961: 188) referred to the second part of Mycologia Fennica
(Karsten 1873a: 103) as the place of valid publication of Leptosphaeria culmifida when making the
combination in Keissleriella. Because Bose did not cite a type and Karsten clearly cites his earlier
publication in Fungi Fenniae Exsiccati, none of the provisions of Art. 41.8 apply and this combination
is invalidly published; we validate it here.

341
Fig. 209 (FH 00785953)
Leptosphaeria coccodes P. Karst., Fungi Fenn. Exs., Cent. 10: no. 963, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 103, 1873].
?= Lophiostoma semiliberum (Desm.) Ces. & De Not., Comment. Soc. Crittog. Ital. 1(4): 220, 1863.
[Full synonymy, see https://www.mycobank.org/MB/249985]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 963
(fig. 210). Aptroot (1998: 147) designated a lectotype (H 6052120).
Fig. 210 (FH 00785955)
Leptosphaeria rustica P. Karst., Fungi Fenn. Exs., Cent. 10: no. 964, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 105, 1873].
= Leptosphaeria doliolum (Pers.) Ces. & De Not., Comment. Soc. Crittog. Ital. 1(4): 234, 1863.
[Full synonymy, see https://www.mycobank.org/MB/161618]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 964
(fig. 211).
Fig. 211 (FH 00785956)
Sphaerella innumerella P. Karst., Fungi Fenn. Exs., Cent. 10: no. 965, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 182, 1873].
[Full synonymy, see https://www.mycobank.org/MB/174943]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 965
(fig. 212). Aptroot (2006: 109) considered this species a synonym of Mycosphaerella punctiformis
based on a morphological examination of original material he considered a ‘holotype’ (H 6048756).
Because of a lack of a statement indicating designation in that publication, this cannot be corrected to
‘lectotype’ and considered a lectotypification (Art. 7.11).
Fig. 212 (FH 00785957)
Ceratostoma dispersum P. Karst., Fungi Fenn. Exs., Cent. 10: no. 966, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 116, 1873].
= Barbatosphaeria barbirostris (Dufour) Réblová, Mycologia 99(5): 727, 2008 [2007].

342
[Full synonymy, see https://www.mycobank.org/MB/244875]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 966
(fig. 213). Réblová (2008: 730) examined original material of this species that she considered a
‘holotype’ (UPS F-639913). Because of a lack of a statement indicating designation in that
publication, this cannot be corrected to ‘lectotype’ and considered a lectotypification (Art. 7.11).
Fig. 213 (FH 00785958)
Rhynchostoma exasperans P. Karst., Fungi Fenn. Exs., Cent. 10: no. 967, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 58, 1873].
[Full synonymy, see https://www.mycobank.org/MB/166406]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 967
(fig. 214).
Fig. 214 (FH 00785959)
Lophiostoma duplex P. Karst., Fungi Fenn. Exs., Cent. 10: no. 970, 1870 [Bidrag Kännedom Finlands
Natur Folk 23: 86, 1873].
= Lophiotrema nucula (Fr.) Sacc., Michelia 1(3): 338, 1878.
[Full synonymy, see https://www.mycobank.org/MB/148201]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 970
(fig. 215).
Fig. 215 (FH 00785962)
Sphaeria lonicerina P. Karst., Fungi Fenn. Exs., Cent. 10: no. 971, 1870 [Bidrag Kännedom Finlands
Natur Folk 23: 89, 1873].
= Leptosphaeria holmskjoldii (P. Karst.) Chi Y. Chen & W.H. Hsieh (as ‘holmskjoeldii’), in Chen et
al., I.M.I. Descript. Fungi Bact. 154(1535): [1], 2002.
[Full synonymy, see https://www.mycobank.org/MB/163000]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 971
(fig. 216).
Fig. 216 (FH 00785963)

343
Sphaeria deflectens P. Karst., Fungi Fenn. Exs., Cent. 10: no. 972, 1870.
≡ Teichospora deflectens (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 67, 1873.
[Full synonymy, see https://www.mycobank.org/MB/655688]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 972
(fig. 217).
Fig. 217 (FH 00785964)
Leptosphaeria dolioloides (Auersw.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 978, 1870 [Bidrag
Kännedom Finlands Natur Folk 23: 106, 1873].
≡ Nodulosphaeria dolioloides Auersw., in Rabenh., Fungi Eur. Exs., Cent. 6: no. 547, 1863.
[Full synonymy, see https://www.mycobank.org/MB/249313]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 978 (fig.
218).
Fig. 218 (FH 00785970)
Anthostoma foedans P. Karst., Fungi Fenn. Exs., Cent. 10: no. 982, 1870.
≡ Phaeosperma foedans (P. Karst.) P. Karst., Bidrag Kännedom Finlands Natur Folk 23: 55, 1873.
= Pseudovalsaria ferruginea (Nitschke) Rappaz, Mycol. Helv. 7(1): 159, 1995.
[Full synonymy, see https://www.mycobank.org/MB/363314]
Notes: This is a new species name validly published on the label of Karsten, Fungi Fenn. Exs. 982
(fig. 219). A lectotype [K(M) 190799] was designated by Ju et al. (1996: 442).
Fig. 219 (FH 00785974)
Sphaerella gangraena (Fr. ex Duby) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 983, 1870 [Bidrag
Kännedom Finlands Natur Folk 23: 185, 1873].
≡ Sphaeria gangraena Fr. ex Duby (as ‘gangrena’), Bot. Gall. 2: 695, 1830, nom. sanct. [Fr., Syst.
Mycol. 3(index): 165, 1832].
≡ Telimenella gangraena (Fr. ex Duby) Petr., Sydowia 1(1–3): 79, 1947.
[Full synonymy, see https://www.mycobank.org/MB/291340]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 983 (fig.
220).
Fig. 220 (FH 00785975)
Cryptospora tessella (Pers.) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 985, 1870 [Bidrag Kännedom
Finlands Natur Folk 23: 78, 1873].
≡ Sphaeria tessella Pers., Syn. Meth. Fung. 1: 48, 1801, nom. sanct. [Fr., Syst. Mycol. 2(2): 393,
1823].
≡ Allantoporthe tessella (Pers.) Petr., Hedwigia 62: 289, 1921.
[Full synonymy, see https://www.mycobank.org/MB/280069]

344
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 985 (fig.
221).
Fig. 221 (FH 00785977)
Pleospora herbarum subsp. pisi (Sowerby) P. Karst., Fungi Fenn. Exs., Cent. 10: no. 991, 1870.
≡ Sphaeria pisi Sowerby, Col. Fig. Engl. Fung. 3: [77], 1803, nom. sanct. [Fr., Syst. Mycol. 2(2): 509,
1823].
= Stemphylium vesicarium (Wallr.) E.G. Simmons, Mycologia 61(1): 9, 1969.
[Full synonymy, see https://www.mycobank.org/MB/339660]
Notes: This is a new combination validly published on the label of Karsten, Fungi Fenn. Exs. 991 (fig.
222).
Fig. 222 (FH 00785983)
Puccinia ptarmicae P. Karst., Fungi Fenn. Exs., Cent. 10: no. 999, 1870, nom. inval., Art. 38.1(a).
≡ Puccinia ptarmicae P. Karst., Bidrag Kännedom Finlands Natur Folk 31: 41, 1879.
[Full synonymy, see https://www.mycobank.org/MB/214169]
Notes: This name was originally invalidly published as a nomen nudum on the label of Karsten, Fungi
Fenn. Exs. 999 (fig. 223) but later validated.
Fig. 223 (FH 00785991)
Literature
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the Teloschistaceae (lichenized ascomycetes). Graphis Scripta 27(1–2): 37–41.
Agerer, R. 1983: Typusstudien an cyphelloiden Pilzen IV. Lachnella Fr. s. l. Mitteilungen der Botanischen
Staatssammlung München 19: 163–334.
Aptroot, A. 1998: A world revision of Massarina (Ascomycota). Nova Hedwigia 66(1–2): 89–162.
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Addresses of the authors
James K. Mitchell, Farlow Herbarium and Library of Cryptogamic Botany, Department of Organismic
and Evolutionary Biology, Harvard University, Cambridge, Massachusetts 02138, USA.
(E-mail: jmitchell@fas.harvard.edu)
Konstanze Bensch, Westerdijk Fungal Biodiversity Institute, P.O. Box 85167, NL-3508 AD Utrecht, The
Netherlands.
(E-mail: k.bensch@wi.knaw.nl)