Description of a new species and new faunistic records of the genus Anomala Samouelle (Coleoptera, Scarabaeidae, Rutelinae) from China and neighboring regions

Abstract

Anomala xiongi Wang & Zorn spec. nov. is described from Yunnan, China. This new species can be distinguished from all other known Anomala species by conspicuous long erect setae rising from the primary costae of elytra. Additionally , ten new distributional records of Anomala species from China and neighboring countries are provided.

Full text

147
ISSN 0005-805X
2021
71 (1): 147 – 159
© 2021 The Authors
Description of a new species and new faunistic records of the genus
Anomala S (Coleoptera, Scarabaeidae, Rutelinae) from China
and neighboring regions
With 51 figures
F-L W 1 and C Z 2, 3
1 Engineering Research Center for Forest and Grassland Disaster Prevention and Reduction, Mianyang Normal University,
166 MianxingWest Road, Mianyang, Sichuan 621000, P. R. China. – 531609816@qq.com.
2 Rostocker Straße 1a, 17179 Gnoien, Germany. – czorn70@hotmail.com
3 Corresponding author
Published on 2021–06–30
DOI:10.21248/contrib.entomol.71.1.147-159
Abstract
Anomala xiongi W & Z spec. nov. is described from Yunnan, China. This new species can be distinguished
from all other known Anomala species by conspicuous long erect setae rising from the primary costae of elytra. Addi-
tionally, ten new distributional records of Anomala species from China and neighboring countries are provided.
Key words
Oriental region, Palaearctic region, Asia, beetles, Insecta, Coleoptera, taxonomy, new distributional records
Zusammenfassung
Anomala xiongi W & Z spec. nov. wird aus Yunnan, China, beschrieben. Diese neue Art unterscheidet sich von
allen anderen bekannten Anomala-Arten durch lang abstehende Haare auf den primären Rippen der Elytren. Darüber
hinaus werden neue Funde von zehn Anomala-Arten aus China und angrenzenden Ländern präsentiert.
Nomenclatural act
Anomala xiongi W & Z spec. nov. – urn:lsid:zoobank.org:act:F10AD66D-BEF8-41E2-B22D-42D0492D9E95
Introduction
Anomala S, 1819 is the most species-rich
genus in subfamily Rutelinae, with approximately 1300
described species (K 2012). So far, nearly 220
species have been recorded in China (K 2012;
Z & B 2016; Z et al. 2017). During the
past decade, eight Anomala species were described from
China (Z 2011; P 2015; H & W
2019; Z 2019). Based on an examination of recently
collected Anomala material from southern China and
neighboring regions, we describe one new species from
China and present new distributional records for ten
known species.

W, F-L & Z, C.: Description of a new species and new faunistic records of the genus Anomala S
148
Material and Methods
Morphological terminology used in this study is based
on Z (2007). The body length was measured from
the apex of the clypeus to the apex of the pygidium. Type
specimens of the new species described in this paper bear
the following labels: 1) printed type designation label:
“HOLOTYPE (or PARATYPE) / Anomala xiongi new
species, det. Wang & Zorn 2019”; 2) red “Holotype” or
“Paratype” label; 3) collecting data labels. The collect-
ing data of the type material are cited verbatim from the
labels, individual labels are separated by a slash (/). All
photographs were edited in Adobe Photoshop® CC 2018.
We use the following abbreviations for collections:
AHPC Collection Andreas Hetzel, Münster, Germany
CCPC Collection Chang-Chin Chen, Tianjin, China
CWNU China West Normal University, Nanchong,
China
CZGG Collection Carsten Zorn, Gnoien, Germany
CZMZ Collection Ming-Zhi Zhao, Guangzhou, China
FLWC Collection Fa-Lei Wang, Chongqing, China
HNHM Hungarian Natural History Museum, Budapest,
Hungary
MFNB Museum für Naturkunde Berlin, Leibniz-Institut
für Evolutions- und Biodiversitätsforschung,
Berlin, Germany
MSPC Collection Matthias Seidel, Hamburg, Germany
MNHN Museum national d’Histoire naturelle Paris,
France
MYTC Mianyang Normal University, Sichuan, China
NMEG Naturkundemuseum Erfurt, Erfurt, Germany
NMPC National Museum (National History), Prague,
Czech Republic
PLPC Collection Paul Lago, Oxford, Mississippi, USA
PPPC Collection Petr Pacholátko, Brno, Czech
Republic
SNSD Senckenberg Naturhistorische Sammlungen
Dresden, Dresden, Germany
SYSU The Museum of Biology, Sun Yat-sen Univer-
sity, Guangzhou, China
ZMH Zoologisches Museum Hamburg, Centrum für
Naturkunde, Germany
Description of the new species
Anomala xiongi W & Z, spec. nov.
urn:lsid:zoobank.org:act:F10AD66D-BEF8-41E2-B22D-42D0492D9E95
(Figs 1–12)
Type locality: China, Yunnan, Lincang, Yunxian
Cou nty.
Type material: Holotype:  (MYTC), China, Yunnan,
Lincang, Yunxian County, Manwan, Shuibatou /
24°70'17"N, 100°36'31"E, alt. 1791 m, VII-VIII.2018,
Zi-Chun Xiong leg. Paratypes: 8 , 8  (MYTC),
same data as the holotype; 3  (MYTC), China,
Yunnan, Lincang, Yunxian County, Manwan, Shuibatou
/ 24°70'17"N, 100°36'31"E, alt. 1791 m, VII-VIII.2017,
Zi-Chun Xiong leg.; 3 , 3  (FLWC), China,
Yunnan, Lincang, Yunxian County, Manwan, Shuibatou
/ 24°70'17"N, 100°36'31"E, alt. 1791 m, VII-VIII.2019,
Zi-Chun Xiong leg.; 4  (MYTC), China, Yunnan,
Lincang, Yunxian County, Manwan, Shuibatou /
24°70'17"N, 100°36'31"E, alt. 1791 m, VII-VIII.2019,
Zi-Chun Xiong leg.; 1 , 1 (MYTC), China, Yunnan,
Dehong, Yingjiang County, Mangyun / 24°45'55"N,
98°51'57"E, alt. 943 m, VII-VIII.2019, Shao-Fu Chen
leg.; 3 , 2  (CZMZ), China: Yunnan, Lincang,
Cangyuan County, Nangunhe River, Wengding Preserve
Station, 1800 m, 2019.V.1-6, Yu-Chen Zheng leg.; 1 ,
2  (CZMZ), China: Yunnan, Lincang, Yunxian
County, Manwan, Shuibatou Village, 2014.VII-VII, light
trap, Zi-Chun Xiong leg.; 1 (CCPC), China, Yunnan,
Lincang, Yongde, Yalian, Damaidi Village, 2012.V.8,
X. D. Yang leg.
Description (male, holotype): Length: 13.4 mm; width:
6.5 mm; body elongate ovoid, moderately convex.
Coloration (Fig. 1): Body light brown except for head and
legs; head medium brown; anterior margin of clypeus,
frontoclypeal suture, ocular canthus and posterolateral
portion of vertex blackened; antenna light brown; prono-
tum with small, vague brown spots near middle of each
side; anterior and posterior margin blackened; scutellum
with blackened margin; elytra with all margins narrowly
blackened except for the suture with a broader blackened
area reaching the subsutural interstice; pygidium with
a vague, blurry blackish patch near middle of anterior
margin; membrane of posterior margin of each abdomi-
nal ventrite dark brown; legs predominantly red-brown,
with the distal portion of tarsomeres and tibiae blackened,
major part of femurs light brown. Head. Clypeus subtrap-
ezoidal, disc transversely punctate, anterior margin
rounded, weakly reflexed; frons with deep punctures,
which are rugose anteriorly and separated posteriorly.
Ratio of interocular width/head width approximately
0.55; antenna long, with pale hairs, antennal club length
equals combined length of antennomeres 2-6.
Thorax: Pronotum approximately 1.8 times as wide
as long; disc with vague and discontinuous impressed
midline; with moderately dense, moderately fine, deep,
irregular punctation, somewhat denser and smaller
laterally; anterior angles acute and protruding, posterior
angles subrectangular, rounded off; sides of pronotum
weakly converging anteriad in middle, slightly sinuate
in anterior half, subparallel in posterior half; lateral
margins with several long setae; anterior marginal line
complete, with several short setae near anterior angles
which gradually become more distant toward middle;
basal marginal line complete and distinct, ending near
posterior angles.
Scutellum triangular, with the sides slightly curved;
punctures denser and smaller laterally than those at disc;
lateral margins smooth; posterior corner sharp.

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Elytra approximately 1.5 times longer than wide; elytral
surface smooth and shining, striate, strial punctures
large and deep; primary striae regular, sulcate, punc-
tures sometimes adjacent to each other; subsutural
interstice with irregular punctation anteriorly and a
regular secondary stria in the posterior 2/3; second
interstice with single secondary stria anteriorly which
is irregularly doubled in the posterior 3/4; third inter-
stice with somewhat irregular secondary stria; primary
costae and lateral margin with long erect setae, setae
of lateral margin distinctly more numerous than those
of primary costae; umbones prominent, apico-sutural
angle subrectangular, slightly rounded; epipleuron
broad near humerus, narrow, ending in posterior third,
flat; lateral margins with numerous short yellow setae,
sparse in anterior and posterior fourth, dense in elytral
mid length.
Ventral thoracic surface with dense, round, large punc-
tures, and dense, moderately long semierect setosity;
disc with very fine, sparse punctures, almost glabrous.
Pygidium (Fig. 9) triangular, convex; disc with moder-
ately coarse, somewhat transverse punctures, indistinct
along anterior margin, partly confluent laterally; poste-
rior marginal line complete; with very sparse long, erect
setae.
Legs: Protibia tridentate, apical and middle tooth rather
long, basal tooth short, obtuse; with a rather long inner
spur; mesotibia weakly fusiform; metatibia very fusiform;
outer surface smooth with sparse fine punctures with a
single oblique row of rigid spines; with two strong termi-
nal spurs, the upper one 2 times longer than the lower;
all tarsi rather long and slender, inner protarsal claw not
clefted; metatarsus much longer than metatibia; outer
mesotarsal claw clefted, inner mesotarsal claw and meta-
tarsal claws long and simple.
Abdominal ventrites (Fig. 10) with rather sparse, trans-
verse fine punctures in the middle, punctures become
gradually denser and larger laterad; with a transverse
row of moderately long, yellow setae, interrupted in
the middle; with several additional setae laterally; sides
carinate.
Aedeagus: parameres simple, short; ventral plate short,
hook-shaped in lateral view. As in Figs 3-6.
Female (Fig. 2): Body slightly stouter; sides of prono-
tum more strongly curved, subparallel in posterior half;
blackish markings much more extended compared to
males: pronotum with small black spot near middle of
each lateral margin and with two large trapezoidal spots
covering the middle of anterior disc, encompassing also
the black anterior margin; basal margin of pronotum
blackish; major part of central elytral disc and humeral
umbones black; protarsus slightly shorter than in male;
inner protarsal claw and outer mesotarsal claw apically
clefted; erect setae of elytra somewhat longer than in
male.
Variability of body size: Male body length: 12.5-
14.3 mm, body width: 5.6-6.7 mm; female body length:
13.4-14.9 mm, body width: 6.1-7.2 mm.
Diagnosis: Anomala xiongi W & Z, new species
can be distinguished from all other Anomala species by
conspicuous long erect setae rising from the primary
costae of elytra (Figs 1, 2). This new species is most simi-
lar to A. fusitibia L, 1992, described from Yunnan and
found in Laos and Thailand (see below). But, in addition
to the aforementioned setae, A. xiongi also differs from
A. fusitibia in the following characters: protibia triden-
tate, inner protarsal claw not cleft in male (Fig. 7).
Etymology: The new species is named after Zi-Chun
Xiong (Yunxian County, Yunnan), who collected the
majority of the type material, including the holotype.
New distributional records
Anomala ahrensi Z, 2011
Distribution: Vietnam, China (new country record):
Yunnan.
Specimens examined: CHINA: Yunnan: 1 , 1 
(FLWC), Jinping County, Honghe Pref., V-VI.2018,
Yun-Chuan Xu leg.; 1 (CCPC), Biaoshuiyan, Ma’andi,
Jinping County, Honghe Pref., 2010.V.13 X. D. Yang leg.;
1  (CCPC), Luobodi Village, Ma’andi, Jinping County,
Honghe Pref., 2010.V.17, X. D. Yang leg.
Remarks: Anomala ahrensi was previously known only
from northern Vietnam (Z 2011, Z et al. 2017).
Herein, we newly report this species from Yunnan, China.
Anomala anguliceps A, 1917
(Figs 13, 22–24)
Distribution: Cambodia; China: Fujian, Guangdong,
Guangxi, Sichuan, Yunnan, Zhejiang, Hunan (new
Chinese provincial record); Laos; Myanmar; Thailand;
Vietnam.
Specimens examined: CHINA: Guangdong: 4 ,
5  (FLWC ), Fenghuang, Fuyong, Baoan, Shen-
zhen, 18.V.2019, Ming-Yu Zhou leg. Hunan. 1 
(FLWC), Zhongzhou, Changsha, 09.VI.2011, Hao Xu
leg. Sichuan: 1  (FLWC), Luozong Mount, Puge
County, VI.2016, Chao Zhou leg.; 1 , 2  (FLWC),
Mujiashanzhai, Emei Mount, Leshan, 20-21.VII.2018,
Lu Qiu leg.; 1 , 4  (CWNU), Hekou, Jiangyou,
Sichuan, 16-17.VII.2009, Juan Li & Xia Yan leg.; 2 ,
4  (CWNU), Wudu, Jiangyou, Sichuan, alt. 436 m,
9.VII.2009, Ai-Ming Shi leg.; 1  (CWNU), Meiguiyuan,
Jiange, Sichuan, alt. 542 m, 7.VII.2009, Yong-Fu Wang &

W, F-L & Z, C.: Description of a new species and new faunistic records of the genus Anomala S
150
Ru-Yi Li leg.; 1  (CWNU), Shiqiao, Jianyang, Sichuan,
8.VII.2009, Ai-Ming Shi leg.; 1  (CWNU), Chenghua
District, Chengdu, Sichuan, 12.VIII.2017, Si-Yu Chen
leg.; 1  (CWNU), Mianyang, Sichuan, 7.VII.2017,
Yu-Shuang Zhao leg.; 2 , 3  (CWNU), Qian-
feng, Sichuan, 26.VI.2016, Xiang He leg. Yunnan: 2 
(FLWC), Manjiu, Manwan, Yunxian County, Lincang,
alt. 2000 m, 20.V.2015, Fa-Lei Wang leg.; 12 , 12 
(FLWC), Manjiu, Manwan, Yunxian County, Lincang,
alt. 2,000 m, VII.2017, Zi-Chun Xiong leg.; 11 ,
12  (FLWC), Manjiu, Manwan, Yunxian County,
Lincang, alt. 2000, VII-VIII.2018, Zi-Chun Xiong leg.;
1  (FLWC), Baihualing, Longyang, Baoshan, VII.2014,
Huang-Shun Xi leg.; 1  (FLWC), Ailao Mount, Xinping
County, Yuxi, VI.2014, Huang-Shun Xi leg.; 2 
(FLWC), Baihualing, Longyang, Baoshan, V.2016, Zhi-
Wei Dong leg.; 1  (FLWC), Tongbiguan, Yingjiang
County, Dehong, 2.VI.2018, alt. 1345 m, Lu Qiu leg.;
3 , 5  (FLWC), Hulukou, Xima, Yingjiang County,
Dehong, VI-VII.2018, Wei-Zong Yang leg.; 1  (FLWC),
Mangyun, Pingyuan, Yingjiang County, Dehong, 15-30.
VI.2019, Gui-Chang Liu leg.; 1  (FLWC), Tongbiguan,
Yingjiang County, Dehong, 2.VI.2018, alt. 1345 m,
Lu Qiu leg.; 2  (FLWC), Nabang, Yingjiang County,
Dehong, VIII.2018, Lu Qiu leg.; 1  (FLWC), Meizihu,
Simao, Puer, 9-11.V.2018, Lu Qiu leg.; 1  (FLWC),
Yingjiang County, Dehong, 15-30.VI.2018, Peng Wang
leg.; 1  (FLWC), Zhenyuan County, Puer, VII.2018,
Lu Qiu leg.
Remarks: This is a widely distributed species in South
East Asia, which occurs in China, Cambodia, Laos,
Myanmar, Vietnam and Thailand (A 1917;
P 2014; Z & B 2016; Z et al.
2017). Z & B (2016) reported this species from
the Chinese provinces of Fujian, Guangdong, Guangxi,
Sichuan, Yunnan and Zhejiang. Herein, Hunan is a new
provincial record for this species in China.
Anomala anthracina A, 1912
(Figs 14, 25–28)
Distribution: Bhutan; India: Sikkim; China (new country
record): Xizang, Yunnan; Nepal (new country record).
Type material: Syntypes. 1  (BMNH), Type /  /
Kurseong, Sikkim. R. P. Verschraeghen. 1911-218. /
Kurseong Inde Verschraeghen 1904. 1908 / Anomala
anthracina, Arrow Type [Arrow’s handwriting]; 2 
(BMNH), Kurseong, Sikkim. R. P. Verschraeghen.
1911-218. / Kurseong Inde Verschraeghen 1904. 1908;
1  (BMNH), Co-type / MUSEUM PARIS SIKKIM
HARMAND 1886 / Anomala anthracina Arr. [Arrow’s
handwriting]; 1  (BMNH), Cotype / MUSEUM
PARIS SIKKIM DARJEELING HARMAND 1890; 1 
(BMNH), Mungphu / Atkinson Coll. 92-3 / Anomala
anthracina, Arrow [Arrow’s handwriting].
Other specimens examined: CHINA: Xizang: 1 
(SYSU), Tibet, Bomi County, nr. 318 national road,
30°03'54"N 95°11'23"E, 2300 m, 2018.VI.19, S. S. Wang
& Z. L. Liang leg. Yunnan: 6 , 13  (FLWC), Gong-
shan County, 30.V.2018, Zheng Zhou leg.; 1  (FLWC),
Hulukou, Xima, Yingjiang County, VI-VII.2018, Wei-
Zong Yang leg.; 1  (FLWC), Mangyun, Pingyuan,
Yingjiang County, VIII.2019, Gui-Chang Liu leg. INDIA:
1 , 1  (CZGG), W Sikkim Pemayangste env., 1900-
2080 m, 15-17.V.1998 leg. Fabrici & Ahrens; 1  (CZGG),
India (Arunachal Pradesh), Tawang Ugelling, 19.VI.2014,
K. Krause lgt.; 1  (CZGG), India W. Bengal, Darjeeling
env. 1800-2500 m, 8-10.V.1991, St. Jakl lgt. NEPAL: 1 ,
1  (CZGG), Nepal east, Taplejung, 23-24.V.2013, lgt.
E. Kučera.
Remarks: Anomala anthracina was formerly known only
from Bhutan and India (Sikkim) (A 1912a; A
1917; F 1975; S 1997; Z & B
2016). Here, it is reported from Xizang and Yunnan,
China, and Nepal for the first time.
Anomala bilobata A, 1912
(Figs 15, 29–32)
Distribution: India; Myanmar; Nepal; China (new
country record): Yunnan.
Specimens examined: CHINA: Yunnan: 7 , 11 
(FLWC), Hulukou, Xima, Yingjiang County, Dehong,
VI-VII.2018, Wei-Zong Yang leg.; 1  (FLWC), Yingji-
ang township, Yingjiang County, Dehong, VI.2018, Yang
Yu leg. NEPAL: 1  (NMEG), Nepal, Prov: Narayani
vic. Chandranigahaui 27°07'25"N, 85°21'11"E, 130 m,
02.V.2003, leg. A. Weigel HF/LF; 1  (NMEG), Nepal,
Mechi/Taplejung, 9 km S Taplejung, Angbung Kabeli
Khola (bridge), 450 m, 27°17'11"N, 87°43'21"E, 03.V.2003,
leg. A. Weigel.
Remarks: Anomala bilobata was originally described
from Bengal, India, and several localities in Myanmar
(A 1912b) and subsequently reported from Nepal
(Z & B 2016). It is here reported from Yunnan
for the first time, representing a new country record for
China.
Anomala callifera O, 1937
(Figs 16, 33–35)
Distribution: Myanmar; China (new country record):
Yunnan.
Specimens examined: CHINA: Yunnan: 1  (CZGG),
China, W Yunnan prov., mts. 60 km E Tengchong, 2200 m,
19-22.V.2006, S. Murzin & I. Shokhin leg.; 1  (CZGG),
China, Yunnan prov., Gaoligongshan mts. 90 km W of

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Baoshan, 26-28.V.1995, lgt. S. Bečvář; 19 , 28 
(FLWC), Dulongjiang, Gongshan County, 30.V.2018,
Zheng Zhou leg.; 1  (FLWC), Tongbiguan, Yingjiang
County, Dehong, 2.VI.2018, alt. 1345, Lu Qiu leg.; 2 
(FLWC), Hulukou, Xima, Yingjiang County, Dehong,
V-VII.2018, Wei-Zong Yang leg.; 1  (FLWC), Mangyun,
Pingyuan, Yingjiang County, Dehong, V-VII.2018, Bao-
Ying Jin leg.
Remarks: Anomala callifera was previously known only
from Myanmar (O 1914a, 1937). In this article, we
for the first time provide records from Yunnan, China.
Anomala cantorioides P, 1959
(Figs 17, 36–39)
Distribution: Vietnam; China (new country record):
Yunnan; Laos (new country record); Myanmar (new
country record); Thailand (new country record).
Type material: Holotype.  (MNHN), YEN-BAI Tonkin
/ Guy Babault / MUSÈUM PARIS 19 GUY BABAULT /
Ohaus determ. Anomala cantori Hope [Ohaus’s hand-
writing] / Anomala cantorioides n. sp. Type. [Paulian’s
handwriting] R. Paulian det. / HOLOTYPE / MNHN EC
1704.
Other specimens examined: China: Yunnan: 2 
(CZGG), China, Yunnan, Xishuangbanna, Mengla,
1000 m, VI-VIII.2014, local collector lgt.; 1  (CZMZ),
Yunnan, South of Mengla, 2015.IX.10, leg. Huang; 1 
(CCPC), China, Yunnan, Mengla, Mohan 1100 m,
2018.V.13-18, leg. C. C. Chen; 1  (NMEG), China,
S-Yunnan, Xishuangbanna, 23 km NW Jinghong, vic.
Na Ban, 21°09,49E 100°39,92N, 730 m, 20.V.2008, leg.
A. Weigel. Laos: 2  (CZGG), Laos central, 70 km
NE Vientiane, Ban Phabat env., N18°16.1 E103°10.9,
150 m, 27.IV.-1.V.1997, M. Štrba & R. Hergovits leg.;
2  (CZGG), Laos Luangphabang province, Mt. Phou
Phakhao, Namtap vill., June 7-11.2008, local collector
leg.; 1  (PPPC), Laos, 4.V.1997. Louangnamtha pr.,
Louang Namtha, 600 m, 21°09'N 101°19'E, Vít Kubán
leg. Myanmar: 1  (BMNH) Upper Burma Mt. Popa
600-1000 m, X.-.XI.37, leg. G. Heinrich. Thailand:
1  (PLPC), Thailand: Nan Province, Amphur
Chiang Klang, 8 km N of Chiang Klang on Hwy 1080,
19°21'N 100°51'E; 277 m, lights; 20 April 2003, AV-5,
Vitheepradit, Prommi, Setaphan; 1  (CZGG), Thai-
land, Chiang Dao, 600 m, 19°24'N 98°55'E, 10-16.V.1991,
Vít Kubáň leg.; 1  (CZGG), North Thailand, Chiang
Dao, 10-20.VII.2004, R. Kocina lgt.; 1  (CZGG),
Thailand N., Mae Hong Son env., Thai/Burma border,
1300 m, 17-21.VI.1993, J. Schneider lgt. Vietnam: 1 
(CZGG), NW Vietnam, Ninh Binh Prov., Dong Tam
vill., Cuc Phuong N. P., 20°15'N 105°43'E, 03.V.2019,
h=90 m, D. Spiridonov leg.
Remarks: This species was previously known only from
the type locality “Yen Bai” in northern Vietnam (P
1959). Here we report new records for China, Laos,
Myanmar, and Thailand.
Anomala fusitibia L, 1992
(Figs 18, 40–42)
Distribution: China; Laos (new country record);
Thailand (new country record).
Specimens examined: CHINA: Yunnan: 1  (FLWC),
Manwan, Yunxian County, Lincang, VII.2017, Zi-Chun
Xiong leg.; 1 , 1  (FLWC), Manwan, Yunxian County,
Lincang, VI-VII.2018, Zi-Chun Xiong leg.; 1  (FLWC),
Xingguo, Pingyuan, Yingjiang County, Dehong,
30.V.2018, Bao-Ying Jin leg.; 1  (FLWC), Tongbiguan,
Yingjiang County, Dehong, 2.VI.2018, Lu Qiu leg.; 4 ,
5  (FLWC), Xima, Yingjiang County, Dehong, 15-30.
VI.2018, Peng Wang leg.; 1  (FLWC), Zhenyuan County,
Puer, VII.2018, Lu Qiu leg.; 2 , 4  (FLWC), Hulu-
kou, Xima, Yingjiang County, Dehong, VI-VII.2018,
Wei-Zong Yang leg.; 2 , 3  (FLWC), Xima, Yingji-
ang County, Dehong, 15-30.VI.2018, Wei-Zong Yang
leg.; 2 , 1  (FLWC), Menglun Reservoir, Mengla
County, Xishuangbanna, alt. 620 m, 15-17.V.2018, Jian-
Yue Qiu & Hao Xu leg.; 1  (FLWC), Nabang, Yingjiang
County, Dehong, VIII.2018, Lu Qiu leg.; 1  (FLWC),
Mount Jinuo, Jinghong, 13-15.V.2018, Lu Qiu leg.; 2 
(CZGG, NMPC), China: Yunnan prov., Gaolingong Mts.
NNR, Baihualin vill., 1535 m, 25°17.7'N 98°48.1'E, at
light in village, J. Hájek & J. Růžička leg. LAOS: 6 
(CZGG), Laos-N, Xieng Khuang prov., Nong Haet,
19°30'N 104°03'E, 29-31.V.2001, D. Hauck leg.; 1 
(NMPC), Laos, Phongsaly prov. 21°41'N, 102°06'E,
Phongsaly env., 6-17.V.2004, ~1500 m, Vít Kubáň leg.;
1  (CZGG), Laos-NE, Hua Phan prov., 25 km SE Vieng
Xai (by road), Ban Kangpabong env., 20°19'N 104°25'E,
14-18.V.2001, D. Hauck leg.; 1  (PPPC), Laos, Louang-
namtha prov., Namtha-MuangSing, 1200 m, 21°09'N
101°19'E, 5-31.V.1997, Vít Kubáň leg. THAILAND:
1  (CZGG), Thailand bor., Chiang Dao env., 17-19.
IV.1995, leg. M. Snižek; 1 , 1  (CZGG), Thailand
bor., Chiang Dao env., 21.V.-4.VI.1995, leg. M. Snižek;
1  (PLPC), Thailand: Chiang Mai Prov., Doi Inthanon
N. P., Nam Tok, Sob Ab, 18°31.775'N 98°36.488'E, 543 m,
UV pan trap, 17 May 2004, Vitheepradit & Prommi leg.;
1  (PLPC), Thailand: Chiang Mai Prov., Doi Intha-
non N. P., Nam Tok, Sob Ab, 18°31'N 98°36'E, 543 m,
17. May 2004, Vitheepradit & Prommi leg.; 1  (CZGG),
Thai-N, Chiang Mai prov., Ban Sanpakia, 1400 m,
1-19.V.1998, Ivo Martinů leg.; 1 , 3  (CZGG), Thai-
land, Mae Hong Son prov., Soppong, 1500 m, 19°27'N,
98°20'E, 7-12.V.1996, lgt. S. Bečvář; 2  (CZGG),
Thai-N, Nan prov., Doi Pukha N.P. Headq., 1500 m,
19°13'N 101°07'E, 22-26.IV.1999, D. Hauck leg.; 1 , 1 
(CZGG), Thai-N, Mae Hong Son prov., SE Soppong,

W, F-L & Z, C.: Description of a new species and new faunistic records of the genus Anomala S
152
1500 m, 19°27'N 98°20'E, 23-27.V.1999, D. Hauck leg.;
4  (CZGG, PLPC), Thailand, Phitsanulok prov.,
Phu Hin Rongkla N.P., UV pan trap at guest house,
1400 m, 17°00'N 100°59'E, 5. May 2003, leg. Vitheepra-
dit, Prommi, and Ferro; 1  (CZGG), Thailand: Chiang
Mai prov., Doi Inthanon N.P., Nam Tok Sob Ab, 543 m,
18°31'N 98°36'E, 17. May 2004, leg. Vitheepradit &
Prommi; 4  (PLPC), Thailand, Loei Province, Na
Haew Nat. Park, 978 m, light at guest house, 24.IV.2003,
leg. Vitheepradit, Prommi, Setaphan.
Remarks: This species was described by L (1992) from
Yunnan, China and subsequently also reported from
the “Oriental Region” without further details in Z
& B (2016). Herein, a considerable number of
specimens were examined; those from Laos and Thailand
represent new country records.
Anomala kuatuna (M, 1955)
(Figs 19, 43–45)
Distribution: China: Fujian, Guangdong, Guangxi, Zheji-
ang, Guizhou (new Chinese provincial record), Hunan
(new Chinese provincial record).
Specimens examined: CHINA: Fujian: 1  (FLWC),
Sanming, VII.2014, Bi-Huang Zhan leg.; 1  (FLWC),
Mangdang Mount, Nanping, 5.VII.2009, no leg.; 1 ,
2  (FLWC), Damo Mount, Sanming, 15.VI.2018,
Xin Liu leg. Guangdong: 3  (FLWC), Houzidong
Mount, Wengyuan County, Shaoguan, 18-19.V.2018,
Jian-Wei Chen leg. Guangxi: 2 , 6  (FLWC), Dayao
Mount, Jinxiu County, Laibin, Guangxi, China, VII.2015,
Bao-Sheng Su leg.; 1  (FLWC), Ziyuan County, Guilin,
Guangxi, China, VII-VIII.2019, Wen Jin leg. Guizhou: 1 
(FLWC), Shunchangxiang, Shuicheng County, Shuicheng,
Liupanshui, China, alt. 1100 m, 22.V.2019, Gui-Qiang
Huang leg. Hunan: 1 , 1  (FLWC), Mount Yangming,
Yongzhou, Hunan, China, alt. 900 m, 1.VII.2019, Yu-Tang
Wang & Yi-Ting Zhong leg. Zhejiang: 1 , 1  (FLWC),
Kuocang Mount, Taizhou, Zhejiang, China, VII.2015,
Zhao-Yang Tang leg.
Remarks: This species is currently only known from
China. Previously it was recorded from Fujian, Guang-
dong and Guangxi (M 1955; L 1993; L
2002b; Z 2011; Z & B 2016). Here we
newly report this species from Guizhou and Hunan.
Anomala siamensis (N, 1891)
(Figs 20, 46–49)
Distribution: Thailand; China (new country record):
Yunnan; Laos (new country record).
Type material: Syntypes[?]. 1  (MFNB), Coll. Nonfried.
Siam / Aprosterna siamensis Type Nonfr. [Ohaus’s
handwriting] / Anomala siamensis Bang-Pain Type
A. F. Nonfried [possibly Nonfried’s handwriting]; 1 
(MFNB), Siam Nonfried / Heteroplia n. sp. / Aprosterna
siamensis Cotype Nonfr. [Ohaus’s handwriting].
Note: It is not entirely certain that these two specimens
from Ohaus’s collection are real syntypes of Aprosterna
siamensis N, 1891 because they do not bear
Nonfried’s type label (compare with the label shown in
H et al. 1990).
Other specimens examined: CHINA: Yunnan: 1 
(FLWC), Menglun, Xishuangbanna, 15.IV.2019,
Zi-Chun Xiong leg.; LAOS: 1  (CZGG), Laos N W,
Nam Tha NPA, 65 km nw Luang Nam Tha, 1050 m,
8-15.IV.2010, S. Murzin leg.; 1  (PPPC), LAOS centr
27.IV.-1.V.1997. 70 km NE Vientiane, Ban Phabat
env., 150 m N18°16.1'E 103°10.9'E. Jendek & O. Sausa
leg.; 1  (PPPC), S Laos Attapu, Nong Lom (lake),
18-30.IV.1999, 800 m, 15°02'N, 106°35'E, M. Strba leg.
THAILAND: 1  (PLPC), THAILAND: Phayao Prov.
Naresuan University; lights on campus; 19. April 2003;
L-467 Vitheepradit, Prommi, Setaphan; 1  (AHPC),
Thailand, 9.11.1992 12 km SW Pak-Chong 130 km NO
Bangkok Mischobstplantage, 400 m Lichtfang, Thielen
leg.; 1  (PLPC), Thailand, Ubon Ratchathani Prov.,
Phu Chong Na Yoi National Park, Kaeng Ka Loa, 11.
April 2004, 182 m, 14°26.178'N 105°16.631E, Sites &
Vitheepradit leg.; 3 , 1  (CZGG), NW Thailand,
Mae Hong Son, Pan Huai Po, 1600 m, 9-16.V.1991, leg.
P. Pachlátko; 1  (CZGG), NW Thailand, Chom Thong,
24-27.IV.1991, leg. Pacholátko.
Remarks: P (1959) misidentified Anomala anguli-
ceps as A. siamensis, but later Z et al. (2017) corrected
this identification based on the male genitalia illustrated
in P (1959). Also, Y (1989) misidentified
A. anguliceps as A. siamensis when reporting this species
from Guizhou, China. Both species mentioned above
were also confused by L (1987, 1992, 2002b), who
consequently described A. corneola L, 2002, which
was subsequently synonymized with A. anguliceps by
P (2014).
In fact, Anomala siamensis is similar to A. bilobata
A, 1912 and A. fissilabris A, 1912 rather than
A. anguliceps. Both A. bilobata and A. fissilabris have a
bilobed clypeus in males, which is the same in males of
A. siamensis, while the clypeal front margin is straight in
A. anguliceps. Anomala siamensis was previously known
only from Thailand, here this species is newly recorded
from China and Laos.

153
CONTRIBUTIONS TO ENTOMOLOGY : BEITRÄGE ZUR ENTOMOLOGIE — 71 (1) 147–159
Anomala xanthoptera B, 1851
(Figs 21, 50–52)
Distribution: India; Nepal; Pakistan; China (new country
record): Yunnan.
Type material: Syntype of Anomala prasinicollis B,
1891 [junior subjective synonym of Anomala xanthop-
tera]. 1  (MFNB), PUNJAB Kulu Distr. / Euchlora
prasinicollis Cotype Bts. [Ohaus’s handwriting] / Anom-
ala prasinicollis Bates [Bates’s handwriting].
Other specimens examined: CHINA: Yunnan: 1 ,
2  (FLWC), Nabang, Yingjiang County, Dehong,
17.II.2016, Hao Xu & Jian-Yue Qiu leg. INDIA: 34 ,
8  (CZGG, ZMH), Indien, Bhimtal, F. Smetacek
leg., Museum Hamburg, Eing. Nr. 3/1977; 1  (SMTD),
Sikkim India. NEPAL: 1  (CZGG), Nepal, Naray-
ani Prov. Sauraha Ufer Rapti River 180 m, 27°34'80"N,
84°29'49"E, 18.IV.2000, leg. A. Skale, LF [collected at
light]; 1  (CZGG), Nepal-Himalaya, Myagdi distr.,
Babichor-Beni, 16.III.1994, 900-1000 m, leg. D. Ahrens.
PAKISTAN: 1  (HNHM), Pakistan, Kashmir, Himalaya,
Mts. 20 km S Muzaffarabad, Nara village, 750 m, 73°29'E,
34°01'N, 20.V.1998, leg. M. László & G. Ronkay.
Remarks: This species was previously known from India,
Nepal and Pakistan (B 1891; A 1917; Z &
B 2016), and is here newly recorded in China based
on three specimens collected from Yunnan.
Anomala zornella P, 2015
Distribution: Laos; Vietnam; China (new record):
Yunnan.
Specimens examined: CHINA: Yunnan: 12 , 4 
(FLWC), Jinping County, Honghe, V-VI.2018, Yun-
Chuan Xu leg.; 3  (FLWC), Lvchun County, Honghe,
20.IV.2019, Zi-Chun Xiong leg.
Remarks: Anomala zornella was originally described
from Laos, Myanmar and Vietnam (P 2015).
Here, we newly record this species from Yunnan, China.
Specimens from both Jinping County and Lvchun County
have reddish legs, which are also present in the speci-
mens collected from Laos and Sa Pa, Vietnam, whereas
specimens from Tam Dao, Vietnam, have greenish leg.
(P 2015; Z et al. 2017).
Discussion
The species described in this work, i.e. Anomala xiongi
sp. nov., is easily recognized by the conspicuous elytral
setae. Except for the pygidium and propygidium, a
sometimes present row of setae along the elytral margin,
and isolated tactile setae on head and pronotum, distinct
setation of the dorsal surface is a rather uncommon char-
acter among most genera of Anomalini (M
1957). However, dorsal setation is found in the major-
ity of species of the subtribe Anisopliina, in Isopliina,
in several genera of the subtribe Popilliina, and,
within the subtribe Anomalina, in many species of the
genera Anoplanomala A, Asiopertha M,
Blitopertha R, Cyriopertha R, Epectinaspis
B, Exomala R, Hontoriella E,
Phyllopertha S, Proagopertha R,
Pseudotrigonocnemis K, and Trigonocnemis
K. In the genus Anomala, setose elytra are a rarely
observed character, which has incurred special attention
in some taxonomic research (F et al. 2013; L
1996; M 1994). Dorsally setose elytra are present
in Asian and New World Anomala species but not in
Afrotropical species and seem to appear in clusters of
otherwise unrelated species-group, e. g. A. hirsutula-
group (L 1996; M 1957; M 1994),
A. trapezifera-group (F et al. 2015). Notably, the
trapezifera-group comprises species with and without
dorsal setation (F et al. 2015), which demon-
strates the plasticity of this character. The presence
of microscopic hairs found in the elytral and prono-
tal punctures of many Anomala species suggests that a
distinctly setose dorsal face evolves by enlargement of
an already existing structure, which makes it more likely
to appear independently in unrelated groups.
In Asian Anomala species, setose elytra are only found
in the 21 species of the hirsutula-group, which occur in
southern China, Laos, Myanmar, Thailand, and Vietnam
(L 1996, M 1957, M 1994, 1996;
K 2008), and also in A. dayaoshanensis W, 2020
(China: Guangxi), A. graminea O, 1905 (southern
China, Vietnam), A. herbacea Z, 2017 (Vietnam),
A. inopinata O, 1914 (Philippines), A. obesa
C, 1869 (Philippines), A. ordinata Z, 2017
(Vietnam), A. sordidula S, 1881 (West Malaysia),
A. xiongi sp. n., and A. yunnana (F, 1886)
(southern China, Vietnam). Except for the species
within the hirsutula-group and for A. graminea and
A. yunnana, which both share a very similar apomorph
structure of the aedeagus, none of the other species seem
to be related to each other. The elytral setae in Asian
Anomala are usually grayish to yellowish, short and
adpressed to semierect. They are confined to the irreg-
ular punctures and never seem to be present in strial
punctures. Anomala xiongi sp. n. is the only known
Anomala species with long erect setae confined to the
primary costae of elytra. However, external morphology
and shape of aedeagus suggest that the closest relatives
of A. xiongi sp. nov. are species without dorsal setosity.
Thus, this character is suitable for species diagnosis,
but seems to have limited significance for higher level
systematics in Anomalini.

W, F-L & Z, C.: Description of a new species and new faunistic records of the genus Anomala S
154
Acknowledgements
We wish to express our sincere gratitude to Zi-Chun
Xiong (Yunxian, Yunnan) for collecting the majority
of the type material of A. xiongi W & Z new
species, as well as to Shao-Fu Chen (Yingjiang, Yunnan),
Andreas Hetzel (Münster), Paul K. Lago (Oxford,
Mississippi), Petr Pacholátko (Brno), Ai-Min Shi, Yun-
Chun Li, and Yuan-Yuan Ma (CWNU), Matthias Seidel
(Hamburg), Chang-Chin Chen (Tianjin), and Ming-Zhi
Zhao (Guangzhou, Guangdong) for providing mate-
rial from their collections for this study. Moreover, we
are very grateful to the collection managers and cura-
tors Ottó Merkl † (HNHM), Antoine Mantilleri, Olivier
Montreuil and Thierry Deuve (MNHN), Matthias Hart-
mann (NMEG), Martin Fikáček, Jiří Hájek and Lukáš
Sekerka (NMPC), Olaf Jäger (SNSD), Johannes Frisch,
Bernd Jäger and Joachim Willers (MFNB), and Martin
Husemann and Matthias Seidel (ZMH) for providing
access to the collections of their institutions. We also
sincerely thank Hao Xu and Jian-Yue Qiu (MYTC), Lu
Qiu (Southwest University, Chongqing), Zhi-Wei Dong
(Kunming, Yunnan), Gui-Qiang Huang (Liupanshui
Normal University, Guizhou), Zhi-Hao Qi (Wuyishan,
Fujian), Zhao-Yang Tang (Shenzhen, Guangdong),
Huang-Shun Xi (Shanghai, Shanghai), Yun-Chuan Xu
(Yuxi, Yunnan), Zheng Zhou (Chuxiong, Yunnan), and
Chao Zhou (Chengdu, Sichuan) for collecting Anomala
specimens for this study.
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W, F-L & Z, C.: Description of a new species and new faunistic records of the genus Anomala S
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Figs 1–12: Anomala xiongi W & Z, new species – 1, 3–7, 9, 10. holotype, male. 2, 8, 11. paratype, female. 1, 2. habitus.
3-6. male genitalia. 7, 8. protibial and protarsal. 9. pygidium. 10, 11. abdominal ventrites. 12. provenance label of holotype.
1, 2, 4. dorsal view. 3. ventral view. 5. lateral view from left. 6. lateral view from right.

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Figs 13–21: Male habitus of Anomala spp., male. – 13. A. anguliceps (Emei Mount, Sichuan). 14. A. anthracina (Gongshan County,
Yunnan). 15. A. bilobata (Yingjiang County, Yunnan). 16. A. callifera (Gongshan County, Yunnan). 17. A. cantorioides (Mengla,
Yunnan). 18. A. fusitibia (Yingjiang County, Yunnan). 19. A. kuatuna (Jinxiu County, Guangxi). 20. A. siamensis (Menglun County,
Yunnan). 21. A. xanthoptera (Yingjiang County, Yunnan).

W, F-L & Z, C.: Description of a new species and new faunistic records of the genus Anomala S
158
Figs 22–52: Aedeagus of Anomala spp., male. – 22–24. A. anguliceps (Emei Mount, Sichuan). 25–28. A. anthracina (Gongshan
County, Yunnan). 29–32. A. bilobata (Yingjiang County, Yunnan). 33–35. A. callifera (Gongshan County, Yunnan).
36-39. A. cantorioides (Mengla, Yunnan). 40–42. A. fusitibia (Yingjiang County, Yunnan). 43–45. A. kuatuna (Jinxiu County,
Guangxi). 46–49. A. siamensis (Menglun County, Yunnan). 50–52. A. xanthoptera (Yingjiang County, Yunnan). 22, 25, 29, 33, 36,
40, 43, 46, 50. dorsal view. 23, 26, 30, 34, 37, 41, 44, 47, 51. ventral view. 24, 28, 31, 35, 38, 42, 45, 48, 52. lateral view from right.
28, 39. lateral view from left. 32, 49. frontal view.

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Fig. 53: Distribution map of
Anomala xiongi W & Z,
new species.