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Checklist of California Paleogene–Neogene marine Mollusca since Keen and Bentson (1944)

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PaleoBios
Title
Checklist of California Paleogene–Neogene marine Mollusca since Keen and Bentson (1944)
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https://escholarship.org/uc/item/7t50309r
Journal
PaleoBios, 38(0)
ISSN
0031-0298
Authors
Groves, Lindsey T.
Squires, Richard L.
Publication Date
2021-02-06
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LINDSEY T. GROVES & RICHARD L. SQUIRES (2021). Checklist of California
Paleogene–Neogene marine Mollusca since Keen and Bentson (1944).
"Body fossils are obviously amongst the most powerful environmental indicators to be found in sedimen-

Andrew P. Miall (1984: p. 41) in Principles of sedimentary basin analysis, Springer-Verlag, New York.
Cover: A representative of each class of Cenozoic California mollusks. A) bivalve Patinopecten healeyi (Arnold, 1906). B) gas-
tropod Turritella infragranulata Gabb (1864). C) chiton Mopalia muscosa (Gould, 1846). D) scaphopod Dentalium neohex-
agonum Sharp and Pilsbry in Pilsbry and Sharp (1897). E) nautiloid Aturia myrlae M.A. Hanna (1927). Scale bars=10 mm.
Citation: Groves, L.T. and R.L. Squires. 2021. Checklist of California Paleogene–Neogene marine Mollusca since Keen and
Bentson (1944). PaleoBios, 38. ucmp_paleobios_52082.
PaleoBios 38:1–360, February 6, 2021
PaleoBios
OFFICIAL PUBLICATION OF THE UNIVERSITY OF CALIFORNIA MUSEUM OF PALEONTOLOGY
Citation: Groves, L.T. and R.L. Squires. 2021. Checklist of California Paleogene–Neogene marine Mollusca since Keen and Bentson
(1944). PaleoBios, 38. ucmp_paleobios_52082.
Permalink: https://escholarship.org/uc/item/7t50309r
Copyright: Published under Creative Commons Attribution-NonCommercial-ShareAlike 4.0 International (CC-BY-NC-SA) license.
Checklist of California Paleogene–Neogene marine Mollusca since Keen and
Bentson (1944)
LINDSEY T. GROVES1* and RICHARD L. SQUIRES2, 3
1Natural History Museum of Los Angeles County, Malacology Department, Los Angeles, CA 90007; lgroves@nhmorg
2Professor Emeritus, California State University, Northridge, Department of Geological Sciences, CA
3Research Associate, Natural History Museum of Los Angeles County, Invertebrate Paleontology Department, Los Angeles,
California, 90007; richardsquires@csunedu
This checklist is a sequel to the one published by A. Myra Keen and Herdis Bentson in 1944 and is an alphabet-
ical listing of California marine Paleogene–Neogene mollusk species 
in the published literature spanning the interval of 1944 through 2020. The original data are given for each
species and subspecies of bivalves, gastropods, chitons (polyplacophorans), scaphopods, and cephalopods.
Where detected, formation names, ages, taxonomy, systematics, and specimen disposition were corrected. A
total of 559 genera and 1,698 species/subspecies were tabulated, with the bivalves and gastropods being the
most abundant taxa. Bivalve and gastropod diversity steadily built up during the Paleocene, nearly doubled
during the Eocene warm time, declined greatly during the cool time of the Oligocene, rebounded to its high-
est peak in the Miocene and then declined slightly during the Pliocene. The other classes represented only
minor faunal components. Chitons were only reported from the Pliocene, and cephalopods had their highest
diversity during the Paleocene and Eocene. Bivalve genera having the highest diversity are: Glycymeris, Ma-
coma, Mactromeris, Nuculana, and Tellina. Gastropod genera having the highest diversity are Cancellaria and
Turritella. It is very likely that the Turritellaovernamed.
The high point of research productivity of published reports on California Paleogene-Neogene mollusks was
in 1990–1994, and a        Menlo Park
USGS Paleontology and Stratigraphy facility. An overall decline in productivity has continued, with noticeable
upticks in 2003 and in recent years.
Keywords: Bivalves, gastropods, chitons, scaphopods, cephalopods, taxonomy, biodiversity, fossil
DEDICATION
This work is dedicated to Drs. Angeline Myra Keen
(1905–1986) (Fig. 1A) and Herdis Bentson (1914–2006)
(Fig. 1B), authors of    
Tertiary Marine Mollusca.” Much has been written about
Myra Keen and her life summarized in a memorial by
Ellen Moore (USGS Menlo Park) (1988a). This memorial
contains a selected bibliography, as well as a selected
biography, about her. However, since very little has
been written about Herdis Bentson, her legacy has been
obscured. Thanks to her son Dr. Bentson H. McFarland
(of Seattle, Washington) and to several online referenc-
es, we are able to provide her with some long overdue
recognition.
Myra Keen
She was a world-renowned American malacologist
and molluscan paleontologist, who was born in Colorado
Springs, Colorado on May 23, 1905. She attended grad-
uate school at Stanford University (M.S. degree in 1931)
and then attended the University of California at Berke-
ley (Ph.D. in 1934), with a dissertation in psychology.
Soon thereafter, she became interested in geology and
paleontology    
Department at Stanford, where she taught geology and
paleontology courses. Eventually, she also became inter-
ested in malacology of the west coast of North America,
and several of her graduate students at Stanford Univer-
sity became prominent malacologists and paleontologists
including Carole S. Hickman, James H. McLean, Judy T.
Smith, and Eugene V. Coan. She passed away at the age
of 80 on January 4, 1986 in Santa Rosa, California.
Herdis Bentson
Herdis Bentson was an American invertebrate pale-
ontologist, who later became interested in petroleum
geology. She was born October 24, 1914 in Alameda,
California to Olaf R. and Agatha E. Bentson immigrants
*author for correspondence
2 PALEOBIOS, VOLUME 38, FEBRUARY 2021
from Denmark. After attending public schools in Alam-
        
family to do so, at the University of California, Berkeley
where she completed her education (B.A. 1935, M.A.
paleontology 1937, Ph.D. paleontology 1941). She was
elected an associate member of the American Museum of

of Paleontology at the University of California, Berkeley
(UCMP), and was the second woman to earn a PhD in
paleontology from UCMP. Her doctoral thesis was on
the stratigraphy and fauna of the Capay Eocene of Sac-
ramento Valley, northern California.
Bentson started working in 1940 for the Standard Oil
Company of California (now Chevron) then located in San

Oiler” company magazine, which included a photograph
of her giving a presentation summarizing her thesis.
She became an executive with Standard Oil, where she
focused on petroleum exploration and retired in 1974,
    
UCMP and at Standard Oil, Herdis described four species
of fossil gastropods of the genus Exilia and six species
of fossil corals. Five species of fossils were named in her
honor: three gastropods; a scaphopod; and a radiolarian
(see below). As a tribute to her fossil collecting prowess
Miller and Downs (1950      
Miss Herdis Bentson found the only nautiloid known
from the Eocene Capay shale of California,” a specimen
they referred to as Eutrephoceras sp. (hypotype UCMP
11412). Herdis married William H. McFarland in 1948
and they had one child, son Bentson H. McFarland. In
retirement, Bentson was active in conservation organi-
zations in California, maintained contact with numerous
relatives in Denmark as well as in the United States, and
compiled a history focusing on her family’s migration to
and their experiences in the Western Hemisphere. She
passed away at the age of 92 on September 8, 2006.
BIBLIOGRAPHY OF HERDIS BENTSON
Bentson, H., 1940. A systematic study of the fossil gas-
tropod Exilia. University of California Publications Bulletin
of the Department of Geological Sciences 25(5):199–238,
pls. 1–3.
Bentson, H. 1943. Eocene (Capay) corals from Califor-
nia. Journal of Paleontology 17(3):289–297, pls. 50–51.
Keen, A.M. and H. Bentson. 1944. Check list of Cali-
fornia Tertiary marine Mollusca. Geological Society of
America Special Papers
-
lection types at the University of California. Journal of
Paleontology 28(3):297–309, pl. 29.
TAXA DESCRIBED BY HERDIS BENTSON
Gastropoda
Exilia clarki 
8, 12, 15, 17, 19, 20. Type Locality: CASG 183, Cowlitz
Fm., Eocene, west bank Cowlitz River, Lewis Co., Wash-
ington. Type Material:
 

20), locality UCMP 7161; paratype UCMP 12141 (pl. 2,

Exilia llajasensis 
Type Locality: UCMP A3042, Llajas Fm., Eocene,
Simi Valley, Ventura Co., California (as Los Angeles Co.,
California in UCMP online database). Type Material:
Holotype UCMP 12144 (pl. 2  
 UCMP 3296 [as Aliso Can-
yon (=Devil Canyon), Los Angeles Co., California UCMP
in online database].
Exilia potreroensis Bentson, 1940:210–211, pl. 1,
 Type Locality: UCMP A1349, Domengine Fm.,
Eocene, Potrero Hills, Solano Co., California. Type Mate-
rial:
Figure 1. Images of Dr. Angeline Myra Keen (A, photo taken and
provided by the late Dr. James H. McLean), and Dr. Herdis Bent-
son (B, photo provided by Bentson McFarland).
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 3
Exilia simiensis   
6, 7, 22. Type Locality: UCMP 3296, Llajas Fm., Eocene,
Simi Valley, Ventura Co. (as Aliso Canyon [=Devil Canyon],
as Los Angeles Co., California in UCMP online database).
Type Material:

      
A3041 (as Los Angeles Co., California in UCMP online
database).
Cnidaria
Caryophillia? capayensis Bentson, 1943:291–292,
    Type Locality: UCMP A1313, Capay
Fm., Eocene, south bank of Smith Canyon, west of Tan-
cred Station (now abandoned) in Capay Valley, Yolo Co.,
California. Type Material: Holotype UCMP 30442 (pl.

Turbonolia capayensis Bentson, 1943:296, pl. 51,
Type Locality: UCMP A1325, Capay Fm., Eocene,
from fossiliferous boulders stratigraphically above local-
ity UCMP A1313, south bank of Smith Canyon west of
Tancred Station (now abandoned) in Capay Valley, Yolo
Co., California. Type Material: Holotype UCMP 30457

Flabellum clarki 
15. Type Locality: UCMP A1800, Capay Fm, Eocene, in
north-south canyon in foothills north of Putah Creek
west-northwest of Winters, Yolo Co., California. Type
Material:
Trochocyathus (Aplocyathus) durhami Bentson
    Type Locality: UCMP
A1800 [=localities UCMP A3072 and A3359], Capay For-
mation, Eocene, north of Putah Creek west-northwest of
Winters, just above stream bottom, Yolo Co., California.
Type Material:

      
UCMP A3359.
Trochocyathus nomlandi Bentson, 1943:293, pl. 50,
Type Locality: UCMP A1800 [=lo-
calities UCMP A1315 and A3359], Capay Fm., Eocene,
in north-south canyon in foothills of Putah Creek west-
northwest of Winters, Yolo Co., California. Type Material:



Madracis wellsi  
14. Type Locality: UCMP A1314, Capay Fm., Eocene,
above locality UCMP A1313, south bank of Smith Canyon
west of Tancred Station (now abandoned) in Capay Valley,
Yolo County, California. Type Material: Holotype UCMP

SPECIES NAMED AFTER HERDIS BENTSON
Cantharus bentsonae Turner, 1940:518; New name
for Cantharus cowlitzensis
preoccupied. [Gastropoda]. Type Locality: UCMP 7164,
Cowlitz Fm., Eocene, Stillwater, Lewis Co., Washington.
Type Material: Holotype UCMP 33208. Remarks: Not
Cantharus (Eocantharus) cowlitzensis Clark, 1938b:714–

Exilia bentsonae   
5–11. [Gastropoda]. Type Locality: USGS 15280, Middle
member, Keasey Fm., Oligocene, upper Nehalem River
Basin, northwestern Oregon. Type Material: Holotype
 



paratypes USNM 251406a–m.
Gemmula bentsonae 
14, 17. [Gastropoda]. Type Locality: UCMP A1802,
Quimper Sandstone, lower Oligocene, on beach north of
Woodman’s Station, Discovery Bay, Jefferson Co., Wash-
ington. Type Material: Holotype UCMP 35454 (pl. 14,

35457. Remarks: In the genus Procerapex,  Hickman
(1976a: p. 85).
Tristylospyris (Tristylospyris) herdisae Campbell
Type
Locality: UCMP A3465, Miocene, below the top of the
Monterey Fm. (Valmonte Diatomite), Miocene, Peck Park
Canyon, Palos Verdes Hills, Los Angeles Co., California.
Type Material: Holotype UCMP 49596.
Cadulus (Cadulus) mcfarlandae Emerson, 1957:986–
Type Locality: UCMP
   
Creek, southwest of Dunns Peak, Vaca Valley, Solano Co.,
California. Type Material: Holotype UCMP 37590 (pl.

INTRODUCTION
Marine mollusks (bivalves, gastropods, chitons, scaph-
opods, and cephalopods) represent collectively a major
component of fossil-invertebrate marine assemblages
found in California. Keen and Bentson (1944) compiled
     -
   
4 PALEOBIOS, VOLUME 38, FEBRUARY 2021

of 1941. Their list included coverage for new species up
to June 16, 1943. Their total number apparently included

explicitly make that clear. This checklist serves as the
sequel to their compilation and begins with their clos-
ing date of June, 1944 and ends, 77 years later, in 2021.
Extensive compilations like Keen and Bentson (1944),
as well as this one, are not common in the literature,
but they are very useful to paleontologists, geologists,
evolutionary biologists, and malacologists. The term
 
in format, but a more descriptive term, for their work
    
Keen and Bentson (1944

Paleocene Series through the Pliocene Series. The term is
antiquated, and its usage is now discouraged (Gradstein
et al. 2012: p. 856). Although no replacement single-word
term has been proposed, Paleogene-Neogene is used in

Keen and Bentson (1944) and Keen (1968) summa-
rized the history of paleontological research on west-
coast Paleogene-Neogene mollusks and concluded that
1) without economic incentives, paleontological studies
tend to languish, and 2) there is a need for training new
paleontologists who are competent in stratigraphy and
also have a thorough grounding in the often neglected
discipline of systematics. These two conclusions are still
relevant today.
Addicott (1972a, 1973a), who focused on the trends of
west-coast reports dealing only with Neogene (Miocene
and Pliocene) mollusks, noted four periods of heightened
   
(1840–1850), two intermediate phases associated with
petroleum exploration, and a phase associated with aca-
demic interest in deciphering earth history (1960–1969).
Keen and Bentson (1944: p. 18) were concerned
about the proper curation of the very important type
specimens found and used by W.M. Gabb, who, starting
-
stantial fossil material of west-coast marine mollusks.
Yet, to date, the type specimens are still stored in three
separate institutions and many still need proper curation.
Ideally, high-resolution-digital images of them should
be made available for online-research purposes. Keen
and Bentson (1944: p. 18) were prescient also in their
concern about the constant looming possibility of tenu-
ous commitment of institutions in regard to the proper
care and long-term curation of type specimens. Most

changes in researchers’ interests and priorities, and
constant redistribution of available space for research.
Storage of collections can also be an issue. In the last
several decades, certain collections (e.g., USC, CIT, UCLA,
and CSUN) have been transferred to LACMIP. The USGS
collection of the Paleontology and Stratigraphy Branch,
Menlo Park, California was shut down as well, and the
collections were transferred to UCMP, after a short pe-
riod of considerable uncertainty about the fate of these
specimens. The impending closure of the Menlo Park
facility was announced in 1995–1996, which followed an
all-time high point of research productivity in 1990–1994
of 40 published reports of California Paleogene-Neogene
mollusks. Collections at Stanford University were trans-
ferred to CASG in 1972. Since this high point, there has
been pronounced decline in productivity, with an uptick
in 2003 and in recent years (Fig. 2).
   -
ments in the way paleontological studies have been
conducted since 1944. The concepts of plate tectonics,
suspect terranes, greenhouse/icehouse paleoclimates,
    
paleoenvironmental/sedimentological analyses of as-
semblages, and recognition of ecomorphic variation in
species, have all changed the way fossils are now studied.
On-going advancements in DNA-based studies of modern
-
tion system of mollusks. The establishment of planktonic
foraminifera and calcareous nannofossil biozones, as
well as the usage of paleomagnetic stratigraphy, which
Figure 2. Number of California Paleogene–Neogene publica-

0
5
10
15
20
25
30
35
40
45
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 5
is largely dependent upon fossils for age control, have
  
molluscan fossils. Computers, high-resolution digital
cameras, and scanning-electron microscopes have helped
greatly in systematic studies. Digitization of data is con-
stantly changing the way paleontologic studies are being
conducted, whether by Google Earth© images, online-
literature access, online-viewing of type specimens,
and online viewing of modern-day representatives of
invertebrate organisms. Access of online-digital images
of primary type specimens of Paleogene-Neogene mol-
lusks, which began a few years ago, will only become
better in the future. CASG has already done so for many
of their fossil-mollusk types, and UCMP and LACMIP have
begun doing so.
Undoubtedly, despite our efforts to verify all of the
specimen information listed in this present report, errors
will be found. Sincere apologies in advance to all users
of this compilation
MATERIALS AND METHODS
This current list is based on an exhaustive review of
  
previously named species and new species described
since the work of Keen and Bentson (1944). Publications
issued between June of 1941 and the cover date of 1944

are listed here. Figured and/or undescribed species over-
looked or missed by Keen and Bentson (1944), including


Paleogene–Neogene chitons (polyplacophorans) and

Known data for each entry include original generic as-
signment, reference, publication, formation, age, locality
information, locality number, and type number(s). Any
updates as to formation, age, taxonomic changes, and
   
Visits to USNM, CASG, UCMP, and SDSNH by the senior
author between 1995 and 2005 were made in order to
verify data.
T.A. Conrad named numerous new species from
California in 1853 through early 1857. He provided de-
scriptions of them, and in some cases, subsequently gave

In early 1857 and continuing for a few years, Conrad
wrote additional papers and used his original names and

the dates of the original description. As a result, there
has been considerable and on-going confusion in the
literature regarding the dates when he published the
original names. In this present report, the links between

The specimens used for the cover of this present pa-
per are also listed here as entries within the checklist,
except for the chiton and scaphopod because they are of
Pleistocene age or younger.
Explanation of Format
Sample entry: Acila (Truncacila) decisa (Conrad,
1855b)a. [Nucula]b. Squires 1988cc. NHMLAC CS 398d:17,
e, Eocenef. Lockwood Valley, Sawmill
Mtn. quad. (1943), Ventura Cog. CSUN 979 [=LACMIP
16081]h. Hypotype LACMIP 7724i. Remarks: j [Nucu-
lidae] k.
a). Genera are listed alphabetically, and the species
within each genus are listed also alphabetically, without
regard to any alphabetical order of the subgenera. Genus,
subgenus (if used), species, subspecies (if cited), and
original author preserving the exact formulation used;
an asterisk following any word indicates a misspelling
which is corrected in the remarks section. In the citation
of publications by G D. Hanna, it is useful to note that
 G, just like that, no period. This causes
no end of confusion among editors and bibliographers”
(Miller 1962: p. 5). Parentheses around an author’s name
indicates that the species was assigned to another genus
in the original description. New species, new subspecies,
or n. nom. are indicated with ‘n. sp., ‘n. subsp.’, or ‘n.
nom.’ following the author and listed in bold type. Type
specimens from the same locality are mentioned within
a single entry for the species. Type specimens not from
the same locality are mentioned in successive separate
entries. b). Original generic and subgeneric assignment
for species transferred to another genus. c). Cited author
and date of publication. There are some cases where
there is more than one author for a surname (e.g., E.J.
Moore and R.C. Moore), but there will be no reason
for any confusion if one uses the year of a publication
along with the author’s last name, when searching the
bibliography for a particular reference. d). Abbreviated
reference (see ‘Publications Abbreviations’ section).
Complete citations are listed in the ‘References Cited’
section. e). Formation and member (if indicated) cited by
the original author. A question mark following a forma-
tion entry indicates that the original author was unsure
of the correct formational assignment. If a formation is
considered incorrect by the authors, an equivalent and/
or correct formation name follows in brackets. Subse-
quent entries for the same species and same formation
6 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Drexel University, Philadelphia, PA; BMNH (= NHMUK),
The Natural History Museum, London, England; CASG,
California Academy of Sciences, Geology, San Francisco,
CA; CASIZ, California Academy of Sciences, Inverte-
brate Zoology, San Francisco, CA; CC, John D. Cooper
Center, Santa Ana, CA; CGS, California Geological Survey
(collections now at UCMP and CASG); CIT, California
Institute of Technology, Pasadena, CA (collections now
at LACMIP); CSMB, California State Mining Bureau, Sac-
ramento, CA (collections now at UCMP and CASG); CSUN,
California State University, Northridge, CA (collections
now at LACMIP); GZG, Geowissenschaftliches Museum
der Universitä Göttingen, Germany; IGPS, Institute of
Geology and Paleontology, Tohoku University, Sendai,
Japan; IU, Indiana University, Bloomington, IN; KU, Uni-
versity of Kansas, Lawrence, KS; LACM, Natural History
Museum of Los Angeles County, CA, Malacology Depart-
ment; LACMIP, Natural History Museum of Los Angeles
County, CA, Invertebrate Paleontology Department;
LSU, Louisiana State University, Baton Rouge, LA; MCZ,
Museum of Comparative Zoology, Harvard University,
Cambridge, MA; MPM, Milwaukee Public Museum, WI;
NMB, Naturhistorisches Museum, Basel, Switzerland;
NZGS, New Zealand Geological Survey, Lower Hutt, NZ;
PRI, Paleontological Research Institution, Ithaca, NY;
SAMO, Santa Monica Mountains National Recreation
Area, Calabasas, CA; SBMNH, Santa Barbara Museum of
Natural History, CA; SCCM, Santa Cruz City Museum of
Natural History, CA; SDSNH, San Diego Society of Natural
History, San Diego, CA; SDSU, California State University,
San Diego, CA (collections now at SDSNH); SHKM, Saito
Ho-on Kai Museum, Sendai, Japan; SU, Stanford Univer-
sity, Palo Alto, CA (collections now at CASG, if type desig-
nation is unknown, specimens will be designated as SU/
CASG); UC, University of Chicago, IL; UCLA, University
of California, Los Angeles (collections now at LACMIP);
UCMP, University of California, Museum of Paleontology,
Berkeley, CA; UCR, University of California, Riverside, CA;
UCSB, University of California, Santa Barbara, CA (type
specimens now at SBMNH); USC, University of Southern
California, Los Angeles, CA (collections now at LACMIP);
USGS, United States Geological Survey, Menlo Park, CA
(collections now at UCMP); USNM, National Museum of
Natural History, Smithsonian Institution, Washington,
DC; WHC, West Hills College, Coalinga, CA.
Text Abbreviations
aff.,auct., of authors;
cf., compared to Co., County; , on the authority of
(published statement); Fm., Formation; Is., island; Ids.,
by the same author(s), but from a different locality, do
not re-list the formation name and age. f). The geologic
age is as cited by original author. If necessary a corrected
age follows the original citation in brackets. Geologic
epochs/series are generalized (e.g., Eocene rather than
early to middle Eocene). For some species or subspecies,
the geologic range has been published inexactly as two
epoch/series names, separated with a slash (e.g., Oli-
gocene/Miocene). g). The original locality description
is given in general terms in order to be in adherence to
new regulations per the Paleontological Resources Pro-
    
of fossil localities on federal lands. In addition, detailed
information concerning private land is generalized in
order to protect the privacy of land owners. Thus, the
locality description consists of brief geographic and/or
geologic description, topographic quadrangle (with date
if cited), and county. Locality descriptions published by
the individual author(s) may differ from descriptions
in institutional records. Supplemental data are given,
when necessary, in brackets by the authors. h). Institu-
tional locality number (when known). Locality numbers
listed in brackets may not have been part of the original
publication and have been added by the authors. i).
Type-specimen designation and institutional number
(when known). Type numbers listed in brackets may not
have been part of the original publication and have been
added by the authors. Type specimens include: Holotype;
paratype(s); lectotype; paralectotype(s); syntype(s); ne-
otype; neoholotype (an invalid term), and hypotype(s).
j). Remarks (where necessary). Updates to the taxonomy
were not included unless a reliable reference was avail-
able. If a species has been transferred to another genus,
the most recent transfer (if there is more than one) is so
 Ficus cooperiana,
 genus -
cus,  Squires (2014a).” If there are any 
   
under the updated genus name (e.g., under ).
k). Family assignments [shown in brackets] are derived
from modern literature: bivalves from Coan et al. (2000)
and/or Coan and Valentich-Scott (2012); gastropods
from Bouchet et al. (2017); chitons from Vendrasco et
al. (2012); scaphopods from Steiner and Kabat (2001);
nautiloids from Teichert and Moore (1964); and coleoids
from Doyle et al. (1994).
Institutional Abbreviations
ABDSP, Anza-Borrego Desert State Park, Borrego
Springs, CA; ANSP, Academy of Natural Sciences of
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 7
islands; Mtn(s)., mountain(s); n. nom., new name; n.
sp., new species; n, subsp., new subspecies; n. var.,
new variety; ph(s)., photo(s); PR, photorevised; quad.,
quadrangle; yd(s)., yard(s).
Literature-Cited Abbreviations
Acta Pala. Polon., Acta Palaeontologica Polonica;
Amer. Conch., American Conchologist; AAPG, American
Association of Petroleum Geologists; AAPG Bull., Bulletin
of the American Association of Petroleum Geologists;
AAPG Mem., Memoirs of the AAPG; AAPG Misc. Pub.,
Miscellaneous Publication of the AAPG; AMN, American
Museum Novitates; BAIM, Bulletin of the Auckland Insti-
tute and Museum; BAP, Bulletins of American Paleontolo-
gy; BSCAS, Bulletin of the Southern California Academy of
Sciences; Bull. BM(NH), Bulletin of the British Museum
of Natural History; Bull. Mizunami Foss. Mus., Bulletin
of the Mizunami Fossil Museum; Bull. SCPS, Bulletin of
the Southern California Paleontological Society; Calif.
Geol., California Geology; CDM Bull., California Divi-
sion of Mines Bulletin; CDMG Bull., California Division
of Mines and Geology Bulletin; CMPUM, Contributions
from the Museum of Paleontology, University of Michigan;
Earth Sci., Earth Science; Geo-Mar. Lett., Geo-Marine
Letters; GSA Bull., Geological Society of America Bulletin;
GSA Mem., Geological Society of America Memoirs; GSA
Sp.Pap., Geological Society of America Special Papers;
Handb. Palaoz., Handbuch der Paläozoologie (Wenz
1938–1944); Index Foss. N.A., Index Fossils of North
America (Shimer and Shrock 1944); Indo-Pac. Moll.,
 Inv. Foss., Invertebrate Fossils
(Moore, Lalicker, and Fischer 1952); Inv. Paleo., Inver-
tebrate Paleontology (Easton 1960); Jour. Moll. Studies,
Journal of Molluscan Studies; Jour. Paleo., Journal of
Paleontology; Jour. Res. USGS, Journal of Research of
the United States Geological Survey; LACM CS, Los An-
geles County Museum, Contributions in Science; Malac.
Rev., Malacolgical Review; MAPS Digest, Mid-American
Paleontology Society Digest; Min. Info. Serv., Mineral In-
formation Service; MNHN Mém., Mémoires du Muséum
National d’Histoire Naturelle, Paris; Mol. Phylo. Evol.,
Molecular Phylogenetics and Evolution; MWNAN, Mon-
ographs of the Western North American Naturalist; NPS,
National Park Service; NHFOC Mem., Natural History
Foundation of Orange County Memoirs; NHMLAC CS,
Natural History Museum of Los Angeles County Contri-
butions in Science; OCF, Orange County Fossils (Stadum
1973); OPCAS, Occasional Papers of the California Acad-
emy of Sciences; Palaeont, Palaeontology; Pal. Amer.,
Paleontographica Americana; Paleo.3, Palaeogeography,
Palaeoclimatology, Palaeoecology; Pr.CAS, Proceedings of
the California Academy of Sciences; Pr. USNM, Proceed-
ings of the United States National Museum; Rev. Mex.
Cien. Geol., Revista Mexicana de Ciencias Geológicas;
SBMNH Mem., Memoirs of the Santa Barbara Museum
of Natural History; Sci. Rept. Tohoku Univ., Science Re-
ports of Tohoku University; SCMA, Santa Cruz Museum
Association; SCPS Bull., Southern California Paleonto-
logical Society Bulletin; SDSNH Mem., San Diego Society
of Natural History Memoirs; SDSNH Occas. Pap., San
Diego Society of Natural History Occasional Paper; SEPM,
Society of Economic Paleontologists and Mineralogists
(now=Society of Sedimentary Geology); Treatise Inv.
Paleo., Treatise of Invertebrate Paleontology; Tr. Proc.
Pal. Soc. Japan, Transactions and Proceedings of the
Paleontological Society of Japan; Tr. SDSNH, Transactions
of the San Diego Society of Natural History; TSGP, Tulane
Studies in Geology and Paleontology; UCPG, University of
California Publications, Geology or Geological Sciences;
USGS Bull., United States Geological Survey Bulletin;
USGS OFR, United States Geological Survey Open-File
Report; USGS PP, United States Geological Survey Profes-
sional Paper; USGS SIR, United States Geological Survey
WSM Ann. Rept., Annual
Report of the Western Society of Malacologists.
DISCUSSION
Stratigraphic Considerations
It is important to emphasize that the original strati-
graphic information is reported for each species. Some of
these stratigraphic names are invalid, however, because
they were not formally named according to the rules of
the North American Stratigraphic Code (Oriel et al. 1983).
The checklist was not intended to serve, however, as a
way to update or revise the nomenclature of the forma-
tions/members associated with each species. To do so
would be beyond the scope of this present work and a
daunting task in its own regard. Nevertheless, in a few
cases, especially concerning Paleocene/Eocene strata,
however, it was imperative to provide some updated in-
formation in order to obtain, at the very least, the general
age of a given species. Quote marks are used for these
invalid stratigraphic names.

outcrops throughout California, but this broad term can
only be used near its type locality area in the vicinity of
the town of Martinez, Contra Costa County. The age of
the Cerros Shale at UCMP 1817, in the Urruttia Creek
area of the San Joaquin Valley, Fresno County, California,
8 PALEOBIOS, VOLUME 38, FEBRUARY 2021
has been inconsistently referred to as Paleocene or
Paleocene/Eocene, but the report of Squires (1988b)
indicates that the age at this locality is actually early
     -
fornia has been referred to as Paleocene or Eocene, but
the report of Squires (1999b), which is the most detailed

ranges from late Paleocene to early Eocene in age in
southern California. More detailed stratigraphic studies

Shale (a.k.a. Rose Canyon Fm.) and the La Jolla Fm. are
imprecise and abandoned stratigraphic terms so will be
used in quotation marks. Both terms have the possibility
of being synonymous and can also be equivalent, to one
or more (see Kennedy and Moore 1971) of the following
valid middle Eocene stratigraphic terms: Mount Soledad
Fm., Delmar Fm., Torrey Sandstone, Ardath Shale, Scripps
Fm., and Friars Fm. (in ascending stratigraphic order).
   
similar fashion but should only be used for the southern
Tehachapi Mountains in Kern County.
The upper Oligocene to lower Miocene Vaqueros
Formation is an example of a stratigraphic unit that ex-

Vaqueros Valley, Monterey County, central California.
This formation has been reported from as far north as
Contra Costa County to as far south as Orange county in
southern California. The formation was named by Hamlin
(1904) on the basis of fossils, which is not in accordance
with the rules of stratigraphic nomenclature. In addition,
his description of the formation was brief and did not
1943)

lithologic units. In this present report, the name Vaqueros
is used without quote marks, but the validity of mapping
the Vaqueros Formation as a continuous stratigraphic
unit in a geologically dynamic and complex coastal area,
extending from central to southern California, remains
open to inquiry.
The Miocene Monterey Fm. is another example of a
stratigraphic unit that has been reported at many locales
in western California, but its type section, located near
the town of Monterey in Monterey County, central Cali-
fornia, is unsatisfactory because it cannot be measured or
examined in any unbroken sequence (Bramlette 1946).
In Los Angeles and Orange counties in southern Califor-
   
association with various other stratigraphic units and
subunits. Additional examples are the, Imperial Fm.,
Latrania Fm., Latrania Sand member, and Saddleback
Valley limestone. These stratigraphic terms have a range
in geologic age from the Miocene to Pliocene. There is a
need for a revision of all of them, and their names need to
be formalized. These informal and/or invalid names are
used in the checklist because that is how the original au-
thors listed them. This present checklist is not the forum
to even attempt to sort out this stratigraphic confusion.
The southern California Pliocene Fernando Fm. has
     
ever been designated (Davis 1998). A few workers have
restricted its use to only the Los Angeles basin (Davis
1998, Swanson 2014). To date, no formal replacement
name has been proposed, thus in this present report

similar age Pico Fm. (Pliocene) is here restricted to
the Ventura basin (Swanson 2014, Swanson and Irvine
2015), including northern Los Angeles County (Squires
2012). Based on detailed stratigraphic studies (Swanson
and Irvine 2015) and on cosmogenic 26A/10Be isochron
burial dating by Rood et al. (2020), this formation is time
transgressive and youngs westwardly from late Pliocene,
in the Valencia area in the eastern part of the basin, to
early Pleistocene to the west. Groves (1991a, b) reported
also that the lower Saugus [=Pico] in the Santa Susana
Mountains (Valencia area) is Plio-Pleistocene age. Buczek
et al. (2020) reported that the Pico Fm. in the Valencia
area might range into the early Pleistocene.
The Careaga Sandstone, which crops out in the Santa
Marina area of Santa Barbara County in southern Cali-
fornia, has been historically regarded as of late Pliocene
age. Based on strontium-isotope data (Buczek et al. 2020)
and
as dictated by the International Commission of Stratig-
raphy (see a series of online charts initiated by Cohen
et al. 2009), the age of the Careaga Sandstone and its
rich molluscan content has now been assigned an early
Pleistocene age. Such an age would normally exclude
any molluscan fauna from being included in this current
checklist. However, because this new information was
obtained late in the editorial-review process and because
the molluscan data was already part of the checklist, it is
useful to-
pretation of new techniques in geochemical studies might
change the understanding of the dynamic and complex
relationships among the factors of time, tectonics, sed-
imentary environments, and localized shallow-marine
water temperatures.
Faunal Observations
The total number of listed molluscan genera is 559,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 9
and is comprised of 227 bivalves, 310 gastropods, 11
chitons, three scaphopods, and eight cephalopods. The
numbers of named species/subspecies per geological

1. Molluscan species/subspecies diversity in California
Paleogene–Neogene strata steadily built up during the
Paleocene, nearly doubled in the Eocene, declined greatly
in the Oligocene, rebounded to its highest peak in the
Miocene, and then declined slightly during the Pliocene
(Fig. 3). Keen and Bentson (1944: p. 7) also noticed this
Pliocene decline in diversity and attributed it to the
reduction in the extensiveness of the marine basins of
deposition during the Pliocene.
The most diverse bivalve genera (i.e., those having
20 or more species/subspecies) are listed here, with
their number of species/subspecies: Glycymeris (20),
Macoma (27), Mactromeris (29), Nuculana (25), and
Tellina (42). Keen and Bentson (1944) reported that in
Tertiary strata of California the bivalve genera Pecten,
Ostrea, and Spisula each consist of more than 20 species,
but these genera based on the updated taxonomy used in
the present study, are represented only by the following
number of species/subspecies: PectenOstrea” (5),
and Spisula (0).
The most diverse gastropod genera (i.e., those hav-
ing 20 or more species/subspecies) are listed here with
their number of species/subspecies: Cancellaria (29) and
Turritella (62). The latter genus has a greater number
of species than any other molluscan genus listed in this
present study. Turritella subspecies, furthermore, make
up 60 percent of all the gastropod subspecies and it is
very likely that the Turritella species/subspecies complex
has been overnamed.
There are a few species whose geologic age is referred

are predominantly found in the Dip Creek area of Lake
Nacimiento, San Luis Obispo County, California. These
1986) and
     
Cretaceous mollusk species became extinct at the end of
the Cretaceous. This observation is in accord with what
Squires (2018d) found in his study of Late Cretaceous

No Paleocene mollusk species, and only a very few
Eocene mollusk species, can be documented as ranging
into the Miocene. In contrast to the numerous Eocene,
Miocene, and Pliocene mollusk species, there are relative-
ly few Oligocene species, especially gastropods, known
from California (Fig. 3). There was a profound change
in the biodiversity of the shallow-marine mollusks in
California at or near the end of the Eocene. This change,
which was detected by early workers (e.g., Keen and
Bentson 1944Hickman 2003,
Nesbitt 2003, Oleinik and Marincovich 2003, Squires
2003) affected the entire northeast  
was the result of a turnover associated with the transi-
tion from a warm, subtropical climate to a cooler climate
beginning near the end of the Eocene.
0
100
200
300
400
500
600
K/Pg Pal Eoc Olig Mio Plio
No. of named mollusk spp. & subspp.
Figure 3. Number of named Paleogene–Neogene mollusk spe-
cies and subspecies of California in the literature from 1944 to
mid-2020. Data tabulated per epoch, starting with the Creta-
ceous/Paleocene boundary (K/Pg).
Table 1. Numerical listing, per geologic stage, of California
Paleogene–Neogene mollusk species and subspecies described

-
tions: Ch, chitons; Sc, scaphopods; Bi, bivalves; Ga, gastropods;
Ce, cephalopods.
Time Ch Sc Bi Ga Ce total per
stage
Pliocene 18 0 222 190 0 430
Miocene 0 5 268 258 2 534
Oligocene 0 0 24 15 0 39
Eocene 0 6 198 240 4 448
Paleocene 0 0 84 87 5 176
K/Pg 0 0 5 5 0 10
total per group 18 11 801 795 11 1637
10 PALEOBIOS, VOLUME 38, FEBRUARY 2021
CHECKLIST
MOLLUSCA 
BIVALVIA 
Acanthocardia (Schedocardia) brewerii (Gabb, 1864). [Cardium]. Givens 1974. UCPG 
Fm., Eocene. East Valley, Reyes Peak quad. (1943
ed.), Ventura Co. UCR 4741. Hypotype UCR 4741/4. [Cardiidae].
Acanthocardia brewerii (Gabb, 1864). [Cardium]. Deméré et al. 1979. SEPM 
Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hy-
potype SDSNH 20779. Remarks: In the subgenus Schedocardia, Moore (2003a: p. 3). [Cardiidae].
Acanthocardia (Schedocardia) brewerii (Gabb, 1864). [Cardium]. Squires 1984. NHMLAC 
Llajas Fm., Eocene. Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype
LACMIP 6583. [Cardiidae].
Acanthocardia (Schedocardia) brewerii (Gabb, 1864). [Cardium]. Squires 1987. NHMLAC 
Fm., Eocene. North side of side canyon to Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN
359 [=LACMIP 16101]. Hypotype LACMIP 7529. [Cardiidae].
Acanthocardia (Schedocardia) cf.  (Gabb, 1864). [Cardium]. Throckmorton 1988a. USGS OFR 88-59:46, pl.
Throckmorton 1988b
Midway quad., Alameda Co. UCMP D8157. UCMP unnumbered hypotype lost,  C. Throckmorton (personal
communication, 2002). [Cardiidae].
Acanthocardia (Schedocardia) brewerii (Gabb, 1864). [Cardium]. Squires 1991c. SEPM 
21. Maniobra Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 671 [=LACMIP 16076].
Hypotype LACMIP 11485. [Cardiidae].
Acanthocardia (Shedocardia)* brewerii (Gabb, 1864). [Cardium]. Squires 1994b. SEPM 
6. Upper Matilija Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23688. Hypotype LACMIP 11631.
Remarks: Error for Schedocardia. [Cardiidae].
Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23710. Hypotype LACMIP
11664.
Acanthocardia (Schedocardia) brewerii brewerii (Gabb, 1864) [Cardium]. Moore 2003a. USGS PP 1228-F [electronic
format] (family Cardiidae):3, 4, Holotype (of Cardium modes-
tum Conrad, 1855b) USNM 1864. [Cardiidae].
Paleocene. Simi Valley area, Ventura Co.]. CASG 66545. Hypotype CASG 66545.01.
Acanthocardia (Schedocardia) breweri (Gabb, 1864). [Cardium]. Squires 2008. NHMLAC 
Fm., Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP
13448. [Cardiidae].
Acanthocardia (Schedocardia) brewerii hartleyensis (Clark and Woodford, 1927). [Cardium]. Moore (2003a). USGS
1228-F [electronic format] (family Cardiidae):4–5, 
Contra Costa Co. UCMP 3609. Holotype UCMP 31277. [Cardiidae].
Acanthocardia (Acanthocardia) kirkensis* (Clark, 1918). [Cardium]. Moore 2003a. USGS PP 1228-F [electronic for-
mat] (family Cardiidae):2, pl. 3Cardiumkirkensis). Kirker Tuff, Oligocene. West of Kirker's Creek, Mt.
Diablo quad., Contra Costa Co. UCMP 2033. Holotype UCMP 11165. Remarks: Error for kirkerensis. In the genus
Ciliatocardium,  Kafanov (2002: p. 6). [Cardiidae].
Acanthocardia (Acanthocardia) reedi (Loel and Corey, 1932). [Cardium]. Moore 2003a. USGS PP 1228-F [electronic
format] (family Cardiidae):1, La Panza Mtns.,
San Luis Obispo Co., UCMP A504. Holotype UCMP 31772. Remarks: In the genus Ciliatocardium,  Kafanov
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 11
(2002: p. 9). [Cardiidae].
Acanthocardia (Agnocardia) sorrentoensis (M.A. Hanna, 1927). [Cardium]. Squires -
cene. Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 22312. Hypotype LACMIP 12752. [Cardiidae].
Acanthocardia (Agnocardia) sorrentoensis (M.A. Hanna, 1927). [Cardium]. Moore 2003a. USGS PP 1228-F [electronic
format] (family Cardiidae):2–3, San Diego Co. UCMP 3975.
Holotype UCMP 31000. [Cardiidae].
Acesta (Acesta) hamlini (Dall, 1900a). [Lima]. Moore 1987. USGS Fm.”
(Repetto Fm. in locality description), Pliocene. Third St. tunnel, Los Angeles, Los Angeles Co. USGS 3426. Holotype
USNM 495186. [Limidae].
Acesta (Plicacesta) haseltinei (Dickerson, 1914a). [Lima]. Moore 1987. USGS 10,
Hills, Camulos quad., Ventura Co. SU 61933. Holotype
(of Lima perrini Waring, 1917) SU/CASG 303 [=CASG 61933.01]. [Limidae].
....., , Mt. Diablo quad., Contra Costa Co. UCMP 1540.
Holotype UCMP 11676.
Acesta (Acesta) multiradiata (Gabb, 1869b). [Lima]. Moore 1987. USGS 
Tejon Fm., Eocene. [Southeast of Lower Lake, Lake Co. USGS 4192. Hypotype (of Stanton 1896   
USNM 157826. [Limidae].
....., Gabb 1869b, pl. 33
Co. MCZ 1055. Holotype MCZ 15016 [=MCZ 108547], missing, F. Collier (personal communication, 1996).
Acharaxdunnensis (Palmer, 1923). [SolemyaMoore 1983. USGS PP 1228-
Near Vacaville, banks of Ulatis Creek, just south of Dunn's Peak, Vaca Valley, Solano Co. UCMP 3573. Holotype
UCMP 30690. [Solemyidae].
Acharax johnsoni (Dall, 1891). [Solemya]. Moore 1983. USGS PP 1228-
Rio Dell side of Eel River, near Scotia-Rio Dell 
53043. [Solemyidae].
Acharax aff. johnsoni (Dall, 1891). [Solemya]. Moore 1983. USGS PP Fernando
Fm.,” Pliocene. Standard Oil Co. Baldwin 73, Montebello Field, Los Angeles Co. USGS 13864. Hypotype (of Wood-
ring 1938
Acharax cf.  (Dall, 1891). [Solemya]. Powell et al. 2019b. PaleoBios-
cene. Caldecott Tunnel excavation, Berkeley Hills, Oakland Hills, Alameda Co. UCMP IP13003. Hypotype UCMP
no. 410421. [Solemyidae].
Acila castrensis (Hinds, 1843). [Nucula]. Woodring et al. 1940 [1941]. 
Fm. (Acila zone), Pliocene. North Dome, Kettleman Hills, Kings Co. USGS 12353. Hypotypes (7) USNM 495718.
Remarks: In the subgenus Truncacila,  Coan et al. (2000: p. 75). [Nuculidae].
Acila castrensis (Hinds, 1843). [Nucula]. G D. Hanna and Hertlein 1943
Pliocene. Elsmere Canyon, Los Angeles Co. CASG 710. Hypotype CASG 5727. Remarks: In the subgenus Truncacila,
 Coan et al. (2000: p. 75). [Nuculidae].
Acila castrensis (Hinds, 1843). [Nucula]. Cummings et al. 1962. CDMG   
(Pomponio Member), Pliocene. Purisima Creek, Half Moon Bay quad., San Mateo Co. 645-3M. Hypotype deter-
mined here as missing. Remarks: In the subgenus Truncacila,  Coan et al. (2000: p. 75). [Nuculidae].
Acila (Truncacila) castrensis (Hinds, 1843). [Nucula]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):147–148, pl. 27,
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP
305. Hypotypes LACMIP [4447–4450]. [Nuculidae].
Acila castrensis (Hinds, 1843). [Nucula]. Perry 1977. Foss. Santa Cruz 
12 PALEOBIOS, VOLUME 38, FEBRUARY 2021
[East side of Rodeo Gulch Rd., Laurel quad.], Santa Cruz Co. [SCCM 18. Hypotype SCCM 21010]. Remarks: In the
subgenus Truncacila,  Coan et al. (2000: p. 75). [Nuculidae].
Acila (Truncacila) castrensis (Hinds, 1843). [Nucula]. Moore 1983. USGS      
Diego Fm., Pliocene. [San Diego Co. LACMIP 305]. Hypotype (of Hertlein and Grant 1972
4448. [Nuculidae].
Acila (Truncacila) dalli (Arnold, 1908). [Nucula]. Moore 1983. USGS Fm.,
Eocene/Oligocene. On the headwaters of south fork of Waddell Creek, Big Basin, Santa Cruz Co. [SU 115]. Holo-
type USNM 165452. [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Moore 1968. SDSNH 
Shale,” Eocene. Rose Canyon, San Diego Co. [SDSNH 2366. Hypotype SDSNH 4164]. [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Givens 1974. UCPG   
West of Thorn Meadows, South Fork of Piru Creek area, Lockwood Valley quad. (1943), Ventura Co. UCR 4675.
Hypotype UCR 4675/231. [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Moore 1983. USGS 
Eocene. Five km north of La Jolla, San Diego Co. [UCMP 5062]. Neotype (of Schenck, 1936: p. 55) UCMP 31132
[=holotype of Acila lajollaensis M.A. Hanna 1927]. [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b), unnamed form, of Vokes. [Nucula]. Moore 1983. USGS PP 1228-A:A10,
pl. 1, Canyon at end of Eocene ridge, Cholame quad., Kings Co.
UCMP A1165. Hypotype UCMP 15747 (not UCMP 15477 as in text and pl. caption). [Nuculidae].
Acilia* cf.  (Conrad, 1855b). [Nucula]. Zinsmeister 1983a. SEPM 
Fm., Paleocene. Simi Hills, Ventura Co. [UCR 6666]. Hypotype UCR 6666/1 determined here as missing. Remarks:
In the subgenus Truncacila,  Moore (1983: p. A10). Error for Acila. [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Squires 1984. NHMLAC 
El Toro Canyon, Santa Susana quad., Los Angeles Co. CSUN 467 [=LACMIP 16079]. Hypotype LACMIP 6564 (not
LACMIP 6554 as in pl. caption). [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Squires 1987. NHMLAC 
Eocene. Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype
LACMIP 7504. [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Squires 1988c. NHMLAC 
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 979 [=LACMIP 16081]. Hypotype LACMIP 7724.
[Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Throckmorton 1988a. USGS 
in Throckmorton 1988bFrom roadcut on west side of
Tesla Rd., Midway quad., Alameda Co. UCMP D8151. UCMP unnumbered hypotypes lost,  C. Throckmorton
(personal communication, 2002). [Nuculidae].
Acila (Trunacila)* decisa (Conrad, 1855b). [Nucula]. Squires 1991c. SEPM 
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype
LACMIP 11479. Remarks: Error for  [Nuculidae].
Acila (Truncacila) decisa (Conrad, 1855b). [Nucula]. Squires 1999b. MHMLAC 
Susana Fm., Eocene. North side Simi Valley, Santa Susana quad. (1951, PR 1967), Ventura Co. CSUN 959 [=LACMIP
17189]. Hypotype LACMIP 12693. [Nuculidae].
Acila cf.  Howe, 1922. Powell et al. 2019b. PaleoBios
Tunnel excavation, Berkeley Hills, Oakland, Alameda Co. UCMP IP 13003. Hypotype UCMP 410923. [Nuculidae].
Acila (Truncacila) muta Clark, 1918. Addicott 1973b. USGS PP 
Member), Oligocene. South side of road along Carneros Creek, Carneros Rocks quad., Kern Co. USGS M4471.
Hypotype USNM 646517. [Nuculidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 13
.....,  tributary to Zemorra Creek, Carneros Rocks quad., Kern Co.
USGS M4466. Hypotype USNM 646518. [Nuculidae].
Acila (Truncacila) muta Clark, 1918. Moore 1983. USGS PP       Ramon Fm.,
Miocene. Near Oakland/Antioch bridge, Contra Costa Co. UCMP 1131. Holotype UCMP 11196. [Nuculidae].
Acila (Truncacila) aff.  G D. Hanna, 1924. Weaver and Kleinpell 1963
Upper Gaviota Fm., Eocene. West of Arroyo El Bulito, western Santa Ynez Mtns., Lompoc quad., Santa Barbara
Co. UCMP B7023. Hypotype SU 9243 [=CASG 70128]. [Nuculidae].
Acila (Acila) semirostrata (Grant and Gale, 1931). [Nucula]. Moore 1983  25.
north side of Santa Clara Valley, approximately on
the boundary line between Ventura and Los Angeles counties. SDSNH 442. Holotype SDSNH 370. [Nuculidae].
Acila shumardi (Dall, 1909). [Nucula]. Phillips et al. 1974. SEPM 
Eocene/Oligocene. Monocline Ridge, Fresno Co. [UCMP D6528]. Hypotype UCMP 37161. Remarks: In the sub-
genus Truncacila,  Moore (1983: p. A10). [Nuculidae].
Monocline Ridge, Fresno Co. [UCMP D6537]. Hypotype UCMP 37162.
Acila shumardi (Dall, 1909). [Nucula]. Watkins 19747E (cast). Kreyenhagen Fm., Oligocene.
Monocline Ridge, Monocline Ridge quad. (1955), Fresno Co. [UCMP D6528]. Hypotype UCMP 37161. Remarks:
In the subgenus Truncacila, Moore (1983: p. A10). [Nuculidae].
Ridge quad. (1955), Fresno Co. [UCMP D6537]. Hypotype UCMP 37162.
Acila sp. Perry 1977. Foss. Santa Cruz Co., p. 6, Lorenzo
River, Santa Cruz Co. [SCCM 37. Hypotype SCCM 21001]. [Nuculidae].
Acila  Acila Zone”), Pliocene. North
Dome, Kettleman Hills, Kings Remarks: Determined here as Acila (Truncacila) castrensis
(Hinds, 1843). [Nuculidae].
Acila (Truncacila) sp. Weaver and Kleinpell, 1963. UCPG 43:195, pl
Cañada de Cojo, southwestern Santa Ynez Mtns., Lompoc-Pt. Conception quad., Santa Barbara Co., UCMP B6930.
Hypotype SU 9245 [=CASG 68842]. [Nuculidae].
Acila (Truncacila) sp., not (T.) decisa (Conrad, 1855b). [Nucula]. Smith 1975b. SEPM Symposium vol., p. 479, pl.
. 3. Lodo Fm., Paleocene. Lowermost sand above the Cretaceous, Chounet Ranch quad., Fresno Co. CASG
31318a. Hypotype CASG 55758. [Nuculidae].
Acila (Truncacila) sp. B. Schenck 1936. Weaver and Kleinpell, 1963-
ota Fm., Eocene. Divide on east side of Cañada de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP B7013.
Hypotype SU 9244 [=CASG 68843]. [Nuculidae].
Acutostrea gaviota Squires, 2018a62–69, 78–79. Gaviota Fm., Eocene. Divide
between Cuarta Canyon and next canyon west, Santa Ynez Range, Santa Barbara Co. LACMIP 7289. Hypotypes
LACMIP 14678–14681, 14686. [Ostreidae].
…..,  Barbara Co. LACMIP 7261 (not 7262 as in pl. caption).
Hypotype LACMIP 14682.
Range, Santa Barbara Co. LACMIP 7202. Hypotype LACMIP 14683.

Acutostrea idriaensis idriaensis (Gabb, 1869b). [Ostrea]. Moore 1987. USGS PP   
from Gabb 1869Gabb 1869
     Gabb 1869, pl. 34,     Gabb 1869,  
103a). Domengine Fm., Eocene. East of Hacienda at the New Idria, Priest Valley quad., San Benito Co. MCZ 1041.
14 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Lectotype (of Stewart 1930: p. 127) MCZ 15048 [=MCZ 108500], paralectotypes MCZ 15020, 15048 [=MCZ
108497–108499]. [Ostreidae].
.....,  Pozo, Santa Cruz Is., California Channel Ids. [CASG 66752].
Holotype (of Ostrea haleyi Hertlein 1933) CASG 5526 [=CASG 66752.02]. Remarks: Phygraea haleyi (Hertlein,
1933),  Squires (2018a: p. 16–18). [Gryphaeidae].
Acutostrea idriaensisOstrea]. Squires et al. 1988. SEPM 
Eocene. Near head of northeast-trending ravine, Inyokern SE quad. (1972), El Paso Mtns., Kern Co. USGS M9068.
Hypotype USNM 424894. [Ostreidae].
Acutostrea idriaensis idriaensis (Gabb, 1869b). [Ostrea]. Squires 1999c. NHMLAC CS 
Sandstone, Eocene. On south bank of North Fork of Matilija Creek, west of junction of Hwy. 33, Matilija quad.
(1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963]. Hypotypes LACMIP 12457, 12458. [Ostreidae].
Acutostrea idriaensis (Gabb, 1869b). [Ostrea]. Squires 2018a. NHMLAC CS 526:4–5, 
Mouth of Soledad Canyon, San Diego Co. LACMIP 5240. Hypotype LACMIP 14649. [Ostreidae].
…..,  cliff, Torrey Pines State Beach, San Diego Co. LACMIP 199. Hypo-
types LACMIP 14650, 14652, 14653, 14461.
…..,  Beach, San Diego Co. LACMIP 25829. Hypotype LAC-
MIP 14651.

25201 as in caption). Hypotypes LACMIP 14654, 14667.

Hypotype LACMIP 14655.
     
LACMIP 41295. Hypotypes LACMIP 14658, 14660, 14664, 14665.


4899. Hypotypes LACMIP 14462, 14633.
        
25210. Hypotype LACMIP 14666.
Acutostrea idriaensis fettkei (Weaver, 1912). [Ostrea
Fm., Paleocene. East of Lower Lake village, Lake Co. UCMP 790. Holotype (of Ostrea buwaldana Dickerson, 1914a)
UCMP 11719. Remarks: Plicatula ostreiformis Stanton, 1896 (family Plicatulidae)=Ostrea buwaldana Dickerson,
1914a,  Squires (2018a: p. 4). [Ostreidae].
-
tion) south of Soledad Valley, La Jolla quad., San Diego Co. UCMP 3981. Holotype (of Ostrea crandalli M.A. Hanna,
1927) UCMP 31072.
Acutostreamiguelensis (Hertlein, 1928a). [Ostrea-
ros Fm., Oligocene/Miocene. East of Crook Pt., San Miguel Is., California Channel Ids. CASG 1163. Holotype CASG
4121, paratype CASG 4122. [Ostreidae].
Acutostrea simiensis (Zinsmeister, 1983b). [Ostrea-

Ventura Co. UCR 6899. Holotype UCR 6899/101, paratype UCR 6899/102. [Ostreidae]. Remarks: Phygraea haleyi
(Hertlein, 1933),  Squires (2018a: p. 16).
Adulamcknighti (M.A. Hanna, 1927). [Lithophaga
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 15
Fm., Eocene. In seacliff south of of mouth of Soledad Valley, La Jolla quad., San Diego Co. UCMP 5062. Holotype
UCMP 31151. [Mytilidae].
Aeidimytilus adamsianus (Dunker, 1857). [Mytilus
4, 5. San Diego Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Hypotype
LACMIP [4473]. Remarks: In the genus Brachidontes,  Coan and Valentich-Scott (2012: p. 112). [Mytilidae].
Aequipecten andersoni (Arnold, 1906). [Pecten-

Remarks: In the genus Pacipecten,  Moore (1984: p. B45). [Pectinidae].


6890. Hypotype USNM 166577.

Aequipecten andersoni (Arnold, 1906) [Pecten-
terey Fm. (Altamira Shale Member), Miocene. Bluff Cove-Malaga Cove area, slope above beach at Bluff Cove,
Palos Verdes Hills, Los Angeles Co. USGS 13123. Hypotype USNM 497081. Remarks: In the genus Pacipecten,
 Moore (1984: p. B45). [Pectinidae].
Aequipecten andersoni (Arnold, 1906). [Pecten
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 649178. Remarks: In the genus Paci-
pecten,  Moore (1984: p. B45). [Pectinidae].
Aequipecten andersoni andersoni (Arnold, 1906). [Pecten
Miocene. Sandstone ledges, Domengine Ranch quad., Fresno Co. UCMP D1078. Hypotype UCMP 36617. Remarks:
In the genus Pacipecten,  Moore (1984: p. B45). [Pectinidae].

Ridge quad., Kings Co. UCMP D1067. Hypotype UCMP 36618.
Aequipecten andersoni (Arnold, 1906). [Pecten
Miocene. West side of Vine Hill Rd., Santa Cruz Co. [SCCM 27. Hypotype SCCM 21005]. Remarks: In the genus
Pacipecten,  Moore (1984: p. B45). [Pectinidae].
Aequipecten cf.  (Arnold, 1906), scallop [Pecten

Hypotype to be donated to LACMIP.
Aequipecten andersoni subandersoni (Loel and Corey, 1932). [Pecten    
 
Fresno Co. UCMP D1070. Hypotype UCMP 36619. Remarks: In the genus Pacipecten,  Moore (1984: p. B45).
[Pectinidae].
Aequipecten circularis subsp. bramkampi
Fm. [=Latrania Fm.], Pliocene. Oyster reef on ridge running west from east side of meander of stream in badland
area, northwest side of Carrizo Mtn., Imperial Co. UCMP A1268. Holotype UCMP 30035, paratype UCMP 30045.
Remarks: Determined here as Argopecten ventricosus bramkampi (Durham, 1950). [Pectinidae].
Aequipecten circularis eldridgei (Arnold, 1906). [Pecten
10–13. San Joaquin Fm. (Cascajo Conglomerate Member), Pliocene. North of Oyster Hill, South Dome, Kettleman
Hills, Kings Co. USGS 12658 [=USGS 12793 and 12794). Hypotypes USNM 495346–495349. Remarks: Deter-
mined here as Argopecten ventricosus eldridgei (Arnold, 1906). [Pectinidae].
Aequipecten circularis impostor (G D. Hanna, 1924). [Pecten]. Woodring et al. 1940 [1941]. USGS PP 195:91 pl. 13,
Pecten Zone), Pliocene. North of Arroyo Robador and south of Hanford Rd., North
Dome, Kettleman Hills, Kings Co. USGS 12364. Hypotype USNM 495729, 495730, 495732–495735. Remarks:
16 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Determined here as Argopecten ventricosus impostor (G D. Hanna, 1924). [Pectinidae].
Aequipecten discus (Conrad, 1857d). [Pecten
Santa Margarita Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 287 [=LACMIP 16085]. Hypo-
type UCLA 58210 [=LACMIP 10592]. Remarks: In the genus Pacipecten,  Moore (1984: p. B46). [Pectinidae].
Aequipecten impostor (G D. Hanna, 1924). [Pecten
Pliocene. Bed outcropping along the crest of hill, Reef Ridge quad., Kings Co. UCMP B6539. Hypotype UCMP 36620.
Remarks: Determined here as Argopecten ventricosus impostor (G D. Hanna, 1924). [Pectinidae].
Aequipecten raymondi (Clark, 1915). [Pecten
Uppermost Aequipecten and Balanus bed which crops out on top of a low, narrow ridge, Domengine Ranch quad.,
Fresno Co. UCMP D1179. Hypotypes UCMP 36622, 36623. Remarks: Junior synonym of Pacipecten discus (Con-
rad, 1857d),  Moore (1984: p. B47). [Pectinidae].

of Garza Creek, Reef Ridge quad., Kings Co. UCMP D1086. Hypotype UCMP 36621.
Aequipecten raymondi brionianus (Trask, 1922). [Pecten 
(McClure Member), Miocene. From basal conglomerate beds on the north bank of Garza Creek, Reef Ridge quad.,
Kings Co. UCMP D1086. Hypotype UCMP 36624. Remarks: Junior synonym of Pacipecten discus (Conrad, 1857d),
 Moore (1984: p. B47). [Pectinidae].
Aequipecten cf.  (Anderson and Martin, 1914) [Pecten]. Woodring et al. 1946. USGS PP 207:27, pl.
28, Fig. 10 (clay peel). Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon
area, west side of George F. Canyon, Palos Verdes Hills, Los Angeles Co. USGS 13127. Hypotype USNM 497082.
Remarks: In the genus Chlamys,  Moore (1984: p. B22). [Pectinidae].
Aequipecten

cf. Aequipecten sp., scallop. . Foss. Palos Verdes Hills, p
Monterey Fm., Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to LACMIP.
Aligena diegoana
10–11, 13. San Diego Fm., Pliocene. West side of next gully east of LACMIP 305 at the same elevation, San Diego

[Lasaeidae].




Aligena (Aligena) diegoana   
Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Holotype LACMIP 4547. [Lasaeidae].

305A [=LACMIP 16862]. Paratypes LACMIP 4548, 4551.
Americardia          
Canyon, Imperial Co. Hypotype probably at SDSU (current whereabouts unknown). [Cardiidae].
Amiantis callosa (Conrad, 1837). Squires 2012. [Cytherea
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14360. [Veneridae].
Amiantis (Anderson, 1905). [Cytherea]. Addicott 1973b. USGS PP 791:35, pl. 7, 
Olcese Sand, Miocene. Twelve mi. N30°E west,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 17
Kern Co. USGS 6627. Hypotype USNM 646592. [Veneridae].
    
UCMP B1599. Hypotype UCMP 14090.
Amiantis  mathewsonii (Gabb, 1866). [Chione
Gaviota Fm., Eocene. Near junction of Yridisis Creek with El Jaro Creek, Lompoc quad., Santa Barbara Co. UCMP
B7007. Hypotype SU 9300 [=CASG 70133]. Remarks: In the genus Amiantis,  Addicott (1973b: p. 35). [Ven-
eridae].
Amiantis mathewsoni (Gabb, 1866). [Chione
Sandstone Member), Miocene. On northeast-trending ridge on south side of Media Agua Creek, Las Yeguas Ranch
quad., Kern Co. USGS M3280. Hypotype USNM 646588. [Veneridae].
       
646589, 646590.

Ranch quad., Kern Co. USGS M3772. Hypotype USNM 646591.
Amiantis stadleri (Clark, 1915). [Pitaria-
ton, La Costa Valley quad. (1953), Alameda Co. SU 3252. Hypotype SU/CASG 8456. Remarks: Junior synonym of
Amiantis callosa (Conrad, 1837),  Coan et al. (2000: p. 381). [Veneridae].
Amiantis
Co. Hypotype SU 8457 [=CASG 69962]. [Veneridae].
Amiantis-
ber), Miocene. About 30–40 ft. above unnamed stream, Carneros Rocks quad., Kern Co. USGS M3578. Hypotypes
USNM 646584, 646595, 646596. [Veneridae].
Amusium lompocensis (Arnold, 1906). [Pecten-
cia, Miocene. East of Trabuco Canyon, Orange Co. USGS M3179. Hypotype USNM 489565 (not USNM 647084 as
in pl. caption). [Pectinidae].
Amusium lompocensis (Arnold, 1906). [Pecten
Fm., Miocene. Six km south of Lompoc, Santa Barbara Co. Holotype CASG 81. [Pectinidae].

3903]. Paratype USNM 164852.
Amussiopecten lompocensis (Arnold, 1906) [Pecten-
terey Fm. (Saddleback Valley Limestone member), Miocene. Laguna Hills, San Juan Capistrano quad., Orange Co.
SDSNH 4521 (not SDSNH 4512 as published). Hypotype SDSNH 83232. [Pectinidae].
Amussiopecten vanvlecki (Arnold, 1907). [Pecten]. Masuda 1971. Tr. Proc. Pal. 
b. Monterey Fm. (Saltos Shale Member), Miocene. North of Santa Barbara Canyon Ranch, Fox Mtn. quad., Santa
Barbara Co. USGS M2418. Hypotype USNM 646495. [Pectinidae].

Barbara Canyon, Fox Mtn. quad., Santa Barbara Co. USGS M2417. Hypotype USNM 646498.


Amussiopecten vanvlecki (Arnold, 1907) [Pecten
Fm. (Buttonbed Sand Member) (of Stinemeyer et al. 1959: pp. 8, 13), Miocene. Temblor Range, Kern Co. USGS
M3988. Hypotype USNM 646793b (not USNM 646793 as in pl. caption). [Pectinidae].
18 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Amussiopecten vanvlecki (Arnold, 1907). [Pecten
(Saltos Shale Member), Miocene. Near Santa Barbara Canyon Ranch, Santa Barbara Co. USGS M2418. Hypotype
USNM 646495. [Pectinidae].
Amussiopecten vanvlecki (Arnold, 1907). [Pecten
1. Vaqueros Fm., Oligocene/Miocene. 3 km south of Santa Ynez, west end Santa Ynez Range, Lompoc quad., Santa
Barbara Co. USGS 4478. Holotype USNM 165305, paratype USNM 165306. [Pectinidae].
Amussiopecten vanvlecki (Arnold, 1907) [Pecten-
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4312. Hypotype SDSNH 83232. [Pectinidae].
Amussiopecten vanvlecki (Arnold, 1907) [Pecten-
cene. Rancho Mission Viejo Sendero Marketplace site, south side San Juan Creek, Orange Co. [Hypotype at CC].
[Pectinidae].
Anadara (Larkinia) camuloensis (Osmont, 1905). [Arca

UCMP 1637. Holotype UCMP 12006. Remarks: Junior synonym of Anadara (Larkinia) multicostata (G.B. Sowerby
I, 1833b), Coan and Valentich-Scott (2012: p. 179.) [Arcidae].
Anadara camuloensis (Osmont, 1905). [Arca]. Nicol 1950Rein-
hart 1943Pliocene. North of Camulos, Ventura Co. Holotype UCMP 12006. Remarks:
Junior synonym of Anadara (Larkinia) multicostata (G.B. Sowerby I, 1833b),  Coan and Valentich-Scott (2012:
p. 179). [Arcidae].
Anadara (Anadara) carrizoensis
[=Latrania Fm.], Miocene/Pliocene. South slope of Carrizo Mtn., crest of ridge 1120 ft. northeast of mouth of
Alverson Canyon, Imperial Co. CIT 738 [=LACMIP 9855]. Holotype LACMIP 4069 [ex CIT 3248]. [Arcidae].
Anadara devincta (Conrad, 1849a). [Arca
Reef Ridge, west side of Zapato Canyon, Kreyenhagen Hills quad., Fresno Co. USGS 14392a. Hypotype USNM
498745. [Arcidae].
Anadara devincta (Conrad, 1849a). [Arca
collected along the creek in Crow Canyon, Hayward quad., Alameda Co. UCMP A4570. Hypotype UCMP 34125.
[Arcidae].

Anadara devincta var. montereyana (Osmont, 1905). [Arca
Miocene. In trail cut 50 ft. west of UCMP 4931, Contra Costa Co. UCMP A4935. Hypotype UCMP 34127. Remarks:
Anadara (Anadara) montereyana,  Moore (1983: p. A39). [Arcidae].
Anadara formosa (G.B. Sowerby I, 1833b). [Arca]. Deméré and Rugh, 2006. Foss. Treas. Anza-Borrego Desert, p. 57,

Remarks: Anadara carrizoensis Reinhart 1943,  S. Rugh (personal communication, 2019). [Arcidae].
Anadara (Cunearca) hamelini (Wiedey, 1928). [Arca
Fm., Miocene. Near junction of Little Sespe River with Big Sespe River Ventura Co. SDSNH 407. Holotype SDSNH
18. Remarks: In the genus Scapharca,  Moore (1983: p. A45). [Arcidae].
Anadara (Anadara) cf. lakei (Wiedey, 1928). [Arca
Mtn. Silt, Miocene. Caliente quad., Kern Co. Hypotype SU 7346 [=CASG 72027]. [Arcidae].
    
SDSNH 19.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 19
Anadara (Anadara) lakei (Wiedey, 1928). [Arca
Fm., Miocene. West of Syncline Hill, west of Simmler, La Panza quad., San Luis Obispo Co. SU 432. Holotype SD-
SNH19. [Arcidae].
Anadara (Anadara) mediaimpressa (Clark, 1918). [Arca (Scapharca)]. Moore 1983. USGS PP 1228-A:A38–A39, pl. 7,

Co. UCMP 1131. Holotype UCMP 11174. [Arcidae].
Anadara (Anadara) mediaimpressa mediaimpressa (Clark, 1918). [Arca (Scapharca)]. Reinhart 1943. GSA Sp.Pap.
47:39, pl. 4, Oligocene. Near town of Walnut Creek, Contra Costa
Co. UCMP 1131. Holotype UCMP 11174, paratype UCMP 11172. [Arcidae].
Anadara (Anadara) mediaimpressa submontereyana (Clark, 1918). [Arca (Scapharca)]. Reinhart 1943. GSA Sp.Pap.

11186. Remarks: Anadara (Anadara) submontereyana,  Moore (1983: p. A37). [Arcidae].
Anadara (Anadara microdonta (Conrad, 1855b). [Arca (Scapharca)]. Reinhart 1943. GSA Sp.Pap. 47:46, 47, pl. 6,
Remarks:
Anadara (Anadara microdonta (Conrad, 1855b), Moore (1983: A46) who considered it an indeterminate
species. [Arcidae].
Anadara cf.  (Conrad, 1855b). [Arca
2. Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 148 [=LACMIP 16102]. Hypotype
UCLA 58207 [=LACMIP 10579]. Remarks:Anadara (Anadara microdonta (Conrad, 1855b), Moore (1983:
p, A46) who considered it an indeterminate species. [Arcidae].
Anadara (Anadara) montereyana (Osmont, 1905). [Arca 
 Reinhart (1943: p. 48).
SU 1166. Holotype UCMP 11925. [Arcidae].
Anadara montereyana (Osmont, 1905) (not Conrad as in pl. caption). [Arca
Oursan Fm., Miocene. Northeast of Irvington, La Costa Valley quad. (1953), Alameda Co. SU 3248. Hypotype SU
8435 [=CASG 69963]. [Arcidae].
Anadara aff. (Anadara) montereyana (Osmont, 1905). [Arca
Castaic Fm., Miocene. Bottom of small canyon tributary to Haskell Canyon, Los Angeles Co. CIT 1671 [=LACMIP
2556]. [Hypotype LACMIP 4071]. [Arcidae].
Anadara (Anadara) montereyana (Osmont, 1905). [Arca
Fm., Miocene. Selby, Contra Costa Co. SU 1166. Holotype UCMP 11925. [Arcidae].
Anadara sp. cf. 
Miocene. Reata Glen, northeast of San Juan Capistrano, Orange Co. [Hypotype at CC]. [Arcidae].
Anadara (Scaphacra obispoana obispoana (Conrad, 1857d). [Arca]. Reinhart 1943. GSA Sp. Pap. 47:70–72, pl. 10,

[=CASG 72030]. Remarks: In the genus Scapharca Moore (1983: p. A44). [Arcidae].

UCMP 32473.
Anadara obispoana (Conrad, 1857d). [Arca]. Hall 1958 Fm., Miocene. Northeast
of Irvington, Niles quad. (1953.), Alameda Co. SU 3233. Hypotype SU 8436 [=CASG 69964]. Remarks: In the
genus Scapharca Moore (1983: p. A44). [Arcidae].

Anadara obispoana (Conrad,1857d). [Arca
Near Felton and Scott's Valley, Santa Cruz Co. Hypotype determined herein as missing. Remarks: In the genus
ScapharcaMoore (1983: p. A44). [Arcidae].
20 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Anadara obispoana (Conrad, 1857d). [Arca
Fm. (Hames Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M1680. Hypotype
USNM 254352. Remarks: In the genus ScapharcaMoore (1983: p. A44). [Arcidae].
Anadara obispoana (Conrad, 1857d). [Arca
  
Range, San Luis Obispo Co. USGS M1680. Hypotype USNM 254352. Remarks: In the genus Scapharca
Moore (1983: p. A44).[Arcidae].
Anadara (Scapharca obispoana perdisparis (Wiedey, 1928). [Arca
8. Monterey Fm., Miocene. On old road, near Zayante railroad station, Santa Cruz Co. SU 443. Paratype SU 5163
[=CASG 71524]. Remarks: In the genus Scapharca Moore (1983: p. A44). [Arcidae].
Anadara (Anadara) osmonti (Dall, 1909). [Arca 
Miocene. Barker's Ranch, Kern Co. Hypotype CASG 5274. [Arcidae].


  
Devils Den, Kern Co. Holotype UCMP 11927.
Anadara (Anadara) osmonti (Dall, 1909). [Arca
         
Anadara microdonta Conrad of Osmont 1905). UCMP 11927. [Arcidae].
Anadara cf. (Dall, 1909) [Arca]. Powell et al. 2019b. PaleoBios 36: cover (as cf. Anadara osmonti), 7, 8,

Hypotype UCMP 412699. [Arcidae].
Anadara (Esmerarca) reinharti (Lowe, 1935). [Arca     
[=Latrania Fm.], Pliocene. Near Travertine Point, Oasis quad., Imperial Co. (not San Diego Co. as in locality de-
scription). UCMP A1705. Hypotype UCMP 39868, missing,  Powell (2008: p. 6). [Arcidae].
Anadara (Cunearca) rivulata (Wiedey, 1928). [Arca]. Reinhart 1943. GSA Sp. -
blor Fm., Miocene. Near the Kern River, Caliente quad., Kern Co. UCMP 2716. Hypotypes UCMP 32440, 32441.
Remarks: In the genus Scapharca,  Moore (1983: p. A45). [Arcidae].

432. Holotype SDSNH 20.
Anadara (Cunearca) santaclarana (Loel and Corey, 1932). [Arca
12, 13. Vaqueros Fm., Miocene. Ridge west of mouth of Wiley Canyon, Ventura Co. UCMP A252. Holotype UCMP
31765. Remarks: In the subgenus Scapharca,  Moore (1983: p. A44). [Arcidae].

Anadara (Larkinia) santana santana (Loel and Corey, 1932). [Arca
Fm., Miocene. West side of Plano Trabuco opposite south end of hill on plain, Santa Ana Mtns., Orange Co. UCMP
6128. Holotype UCMP 31768. Remarks: In the genus Larkinia,  Moore (1983: p. A42). [Arcidae].
Anadara santana (Loel and Corey, 1932). [Arca
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. UCLA 4268 [=LACMIP 24268]. Hypotype
UCLA 58225 [=LACMIP 10578]. Remarks: In the genus Larkinia,  Moore (1983: p. A42). [Arcidae].
Anadara (Larkinia) santana weddlei (Loel and Corey, 1932). [Arca]. Reinhart 1943. GSA Sp. Pap. 47:64–65, pl. 8,

quad., Monterey Co. UCMP 3675. Holotype UCMP 31771. Remarks: In the genus Larkinia,  Moore (1983: p.
A43). [Arcidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 21
Anadara (Anadara strongi (Loel and Corey, 1932). [Arca    
Vaqueros Fm., Miocene. On spur in large turn in Laguna Canyon (west side of canyon), San Joaquin Hills, Orange
Co. UCMP A527. Lectotype (of Reinhart 1943: p. 42) UCMP 31762. [Arcidae].
Anadara (Anadara strongi (Loel and Corey, 1932). [Arca
14. Upper Vaqueros Fm., Miocene. On spur in large turn in Laguna Canyon (west side of canyon), San Joaquin
Hills, Orange Co. UCMP A527. Lectotype (of Reinhart 1943: p. 42) UCMP 31762, paralectotype UCMP 31762 (on
same sample as lectotype). [Arcidae].
Anadara submontereyana (Clark, 1918). [Arca (Scapharca)
Temblor Fm. (Phacoides Sand member) (of Stinemeyer et al. 1959: pp. 12, 13), Miocene. Las Yeguas Ranch quad.,
Temblor Range, Kern Co. USGS M3772. Hypotype USNM 646519. [Arcidae].
Anadara (Anadara) submontereyana (Clark, 1918). [Arca (Scapharca)]. Addicott 1973b. USGS PP 791:7, 23–24, pl. 1,

Kern Co. USGS M3978. Hypotypes USNM 646519–646521. [Arcidae].
 southeast side of ridge paralleling Media Agua Creek about 200 ft. above dirt road, Las Yeguas
Ranch quad., Kern Co. USGS M3772. Hypotype USNM 646522, not USNM 646519 as in Addicott 1972b, pl. 1, Fig. 7.
Anadara (Anadara) submontereyana (Clark, 1918). [Arca (Scapharca)
12. San Ramon Fm., Miocene. South of town of Walnut Creek, Concord quad., Contra Costa Co. UCMP 52. Holotype
UCMP 11186. [Arcidae].
Anadara thauma Maury, 1925. [Scapharca (Scapharca
Fm. (Latrania Sand Member) [=Latrania Fm.], Miocene. Coyote Mtns., Imperial Co. LACMIP 9855. [Hypotype
LACMIP 8275]. [Arcidae].
Anadara (Anadara) topangensis
Miocene. On Mulholland Dr. near Laurel Canyon Rd., Los Angeles Co. UCLA 391 [=LACMIP 20391]. Holotype UCLA
[UCMP in text and pl. caption] 3258 [=LACMIP 10095]. [Arcidae].
Anadara (Anadara) topangensis Reinhart, 1943. Tweet et al. 2014. MWNAN 7:76, . 7. Topanga Canyon Fm., Mio-
cene. Santa Monica Mtns. National Recreation Area, Los Angeles Co. [LACMIP 20391]. Holotype LACMIP 100095
(ex UCLA 3258). [Arcidae].
Anadara trilineata (Conrad, 1857a) of Arnold. [Arca
10, 19–21. San Joaquin Fm. (Acila Zone), Pliocene. Arroyo Vadoso, North Dome, Kettleman Hills, Kings Co. USGS
12330. Hypotypes USNM 495717, 495720–495722. Remarks:
Conrad, 1857c. [Arcidae].

Co. USGS. 12340. Hypotype USNM 495723.

USNM 495724, 495725.

Neverita Zone), Pliocene. Butte east of Lower Rd.,
North Dome, Kettleman Hills, Kings Co. USGS 12499. Hypotype USNM (slab with Neverita reclusiana [=Glossau-
lax), Olivella cf.  [=Callianax], Anadara trilineata, and Volsella fragments) determined here as missing.
Anadara (Anadara) trilineata trilineata (Conrad, 1857a). [Arca  
9. Merced Fm., Pliocene [=Wilson Grove Fm., Miocene/Pliocene]. Near West Mark Creek, Sonoma Co. SU 1218.
Hypotype (of Anadara schizotoma (Dall, 1898)) CASG 5276. Remarks:-
lustrated in Conrad, 1857c. [Arcidae].


22 PALEOBIOS, VOLUME 38, FEBRUARY 2021
5315 [=CASG 72043].
-
potype SU 5314 [=CASG 72044].
Anadara trilineata (Conrad, 1857a) of Arnold. [Arca]. Woodring and Bramlette 1950 [1951]. USGS PP 222:82, pl.

Hypotype USNM 560028. Remarks:
....., pl. 

560052.

USNM 560099.
Anadara trilineata     Arca]. Woodring et al. 1940 [1941]. USGS PP
Pecten Zone), Pliocene. Near divide between Arroyo Vadoso and Arroyo
Largo, North Dome, Kettleman Hills, Kings Co. USGS 12331. Hypotype USNM 495750. Remarks: Supplemental

Neverita Zone), Pliocene. East fork of Arroyo Toreido, North Dome, Kettle-
man Hills, Kings Co. USGS 12508. Hypotypes USNM 495329, 495331.

495332.
Anadara trilineata    Arca]. Woodring et al. 1940 [1941]. USGS PP
Pseudocardium Zone), Pliocene. North branch of Arroyo Doble-
gado, southeast of El Rascador, North Dome, Kettleman Hills, Kings Co. USGS 12406. Hypotype USNM 495784.
Remarks: 

12400. Hypotype USNM 495788.
Anadara trilineata (Conrad, 1857a). [Arca
Remarks:
Line drawing, source specimen unknown. [Arcidae].
Anadara trilineata (Conrad, 1857a). [Arca
Pleasanton, La Costa Valley quad. (1953), Alameda Co. SU 3237. Hypotype SU. 8438 [=CASG 69953]. Remarks:


Anadara trilineata (Conrad, 1857a). [Arca
(Pomponio Member), Pliocene. Near mouth of Purisima Creek, Half Moon Bay quad., San Mateo Co. 700-1M. Hy-
potype determined here as missing. Remarks:
[Arcidae].
Anadara (Anadara) trilineata (Conrad, 1857a). [Arca
Fm., Pliocene. Along the west bank of Eel River, Scotia quad., Humboldt Co. UCMP B7919. Hypotype UCMP 37757.
Remarks:
Anadara (Anadara) trilineata (Conrad, 1857a). [Arca]. Durham and Addicott 1965. USGS PP 524-A:
2 (cast). Pancho Rico Fm., Miocene/Pliocene. Ridge between Moody Canyon and Pine Valley, San Ardo quad.,
Monterey Co. USGS M996. Hypotype USNM 649090. Remarks:
Conrad, 1857c.Arcidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 23
Anadara (Anadara) trilineata (Conrad, 1857a). [Arca
Juaquin [sic] Fm., Pliocene. Locality unknown, possibly Kings Co. Hypotype current whereabouts unknown. Re-
marks: 

Anadara (Anadara) trilineata (Conrad, 1857a). [Arca       
[San Diego Fm.], Pliocene. Reynard Way, San Diego, San Diego Co. [SDSNH 430. Hypotypes SDSNH 4121, 4122].
Remarks:
Anadara trilineata (Conrad, 1857a). [Arca
Cut and pipeline trench on north side of Arastradero Rd., Palo Alto quad., Santa Clara Co. USGS M1715. Hypotypes
USNM 650961, 650962. Remarks:
Anadara trilineata (Conrad, 1857a). [Arca
4, 6. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305.
Hypotypes LACMIP [4459–4461]. Remarks: 
[Arcidae].

San Diego Co. UCLA 312 [=LACMIP.20312]. Hypotype UCLA 48611 [=LACMIP 9288].
Anadara trilineata (Conrad, 1857a). [Arca
13. San Joaquin Fm. (upper Mya Zone), Pliocene. Kettleman Hills, Kings Co. Hypotypes IU 13833, 13839, 13868,
all missing , R. Alexander (personal communication, 1999). Remarks: 
illustrated in Conrad, 1857c.[Arcidae].
Acila Zone), Pliocene. Kettleman Hills, Pliocene. Hypotypes
IU 13828, 13829, 13846, 13848, 13869–13871.
 Pecten Zone), Pliocene.
Kettleman Hills, Kings Co. Hypotypes IU 13815–13823,13825,13826, 13832, 13834, 13838, 13841–13845,
13847, 13872, 13878.
Neverita Zone), Pliocene. Kettleman Hills, Kings Co. Hypotype
IU 13827, 13831, 13835, 13879.
Pseudocardium Zone), Pliocene. Kettleman Hills,
Kings Co. Hypotypes IU 13836, 13837, 13840, 13880, 13881.
Siphonalia Zone), Pliocene. Kettleman Hills, Kings Co. Hypotypes IU
13824, 13882.

Anadara trilineata (Conrad, 1857a). [Arca]. Perry 1977. Foss. Santa Cruz Co., p. 
Santa Cruz Co. Hypotype determined here as missing. Remarks:
in Conrad, 1857c. [Arcidae].
Anadara trilineata (Conrad, 1857a). [Arca
Jacalitos Fm. (of Fowkes 1982: p. 31), Miocene/Pliocene. North Dome, Kettleman Hills, Kings Co. WHC hypotype
Remarks: 
Anadara (Arca) trilineata). Road cut just northwest of the Atlas Asbestos gate, near
Remarks:
207) from different localities, therefore the actual locality is unknown.
Anadara (Arca) trilineata
Anadara (Anadara) trilineata (Conrad, 1857a). [Arca
24 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Diego Fm., Pliocene. San Ysidro quad. (1943)], San Diego Co. LACMIP 305 Hypotype (of Hertlein and Grant 1972,
Remarks: 


Anadara trilineata (Conrad, 1857a). [Arca     
Capitola City Beach and New Brighton State Beach areas, Santa Cruz Co. [CASG 66173. Hypotype CASG 66173.01].
Remarks: 
Anadara trilineata
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14338. Remarks: 
illustrated in Conrad, 1857c. [Arcidae].
Anadara (Anadara) trilineata subsp. calcarea (Grant and Gale, 1931). [Arca]. Reinhart 1943. GSA Sp.Pap. 47:61–62,

61956.01]. [Arcidae].
Anadara trilineata calcarea (Grant and Gale, 1931). [Arca]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):156, pl.

Hypotypes CASG [=CASG 70450]. [Arcidae].
Anadara (Anadara) trilineata calcarea (Grant and Gale, 1931). [Arca]. Moore 1983. USGS PP 1228-A:A41, pl. 8,

61956.01]. [Arcidae].
Anadara trilineata (Conrad, 1857a) of Arnold, var. cf. canalis (Conrad, 1857a). [Arca]. Woodring et al. 1940 [1941].
Cerastoderma-bearing sandstone), Pliocene. Branch of Arroyo
Doblegado, North Dome, Kettleman Hills, Kings Co. USGS 12706. Hypotype USNM 495344. Remarks: Considered
a subspecies of Anadara trilineata rather than a variety,  Adegoke (1969: p. 91). [Arcidae].
....., pl. 31Siphonalia zone), Pliocene. Southeast slope of Las Paredes, North Dome, Kettleman
Hills, Kings Co. USGS 12826. Hypotype USNM 495815.
Anadara (Anadara) trilineata canalis (Conrad, 1857a). [Arca
Purisima Fm., Pliocene. Pebble bed near Capitola, Santa Cruz Co. SU 1203. Hypotype SU/CASG 5304 determined
here as missing. [Arcidae].
Anadara (Anadara) trilineata canalis (Conrad, 1857a). [Arca]. Durham and Addicott 1965. USGS PP 524-A:A13, pl.

quad., Monterey Co. USGS M982. Hypotype USNM 649099. [Arcidae].
Anadara (Anadara) trilineata canalis (Conrad, 1857a). [Arca-

CASG 5304, lost,  Moore (1983: p. A41). [Arcidae].
Anadara       
Remarks: Determined here as Anadara trilineata
(Conrad, 1857a). [Arcidae].
Anadara
Posa area, Santa Cruz Is., California Channel Ids. [Hypotype SBMNH 60208]. Remarks: Determined here as indet.
cardiid sp. [Cardiidae].

Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 1827 [=SBMNH 60207]. Remarks: Determined
here as indet. cardiid sp.[Cardiidae]
Anadara (ArcaNeverita Zone”), Plio-
Remarks: Determined here as Anadara trilineata (Conrad, 1857a). [Arcidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 25


Anapteris wardi (M.A. Hanna, 1927). [Corbula-
cene. Sea cliff north of La Jolla, San Diego Co. UCMP 5062 (=UCR 4922). Holotype UCMP 31146, paratype UCMP
31147. [Corbulidae].
camera lucida drawing). Ardath Shale, Eocene. East wall of Rose Canyon, San Diego Co. UCR 4952. Hypo-
type UCR 4952/471.
Angulararca californiensis  
from Grant and Gale 1931, as Arca terminumbonis). Pico Fm., Pliocene. Elesmere [sic] Canyon, Los Angeles Co.
[SDSNH 207]. Holotype SDSNH 80 [ex paratype of Arca terminumbonis], paratype SDSNH 82. Remarks: Arca
(Arca) terminumbonis Grant and Gale, 1931, Moore (1983: p. A33). [Arcidae].
Angulararca terminumbonis (Grant and Gale, 1931). [Arca]. Noda 1966b. Saito Ho-on Kai Mus. Res. Bull. 35:16,
from Reinhart 1943, as Arca terminumbonis). Pico Fm., Pliocene. Elesmere [sic]
Canyon, Los Angeles Co. [SDSNH 207]. Holotype SDSNH 79. Remarks: In the genus/subgenus Arca (Arca), 
Moore (1983: p. A33). [Arcidae].
AnodontiaAnodontia (Wagner and Schilling, 1923). [Phacoides]. Moore 1988b. USGS PP 1228-D:D16, pl.

[Lucinidae].
Anomia hannai
Val Celico, a little west of Pleyto, Monterey Co. SU 449. Holotype SU 514 [=CASG 71545]. [Anomiidae].
Anomia inconspicua
of town of Walnut Creek Concord quad., Contra Costa Co. UCMP 1131. Holotype UCMP 11205. [Anomiidae].
Anomia inornatus (Gabb, 1864). [Placunanomia         
Domengine Fm., Eocene. Corral Hollow, Alameda Co. Lectotype (of Moore 1987:C9) ANSP 4442. [Anomiidae].
Anomia mcgoniglensis
-
types UCR 4865/150-4865/151. [Anomiidae].
Anomia mcgoniglensis
Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 438 [=LACMIP 22775]. Hypotype UCLA 59280
[=LACMIP 10560]. [Anomiidae].
Anomiamcgoniglensis
Group in locality description), Eocene. Point on the north side of Soledad Valley formed by McConigle Canyon
joining Soledad Valley, San Diego Co. UCMP 5080 [not 3981 as in text]. Syntypes UCMP 31011–31012 (as 31011

Anomia mcgoniglensis  
Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17849. Hypotype LACMIP 13442. [Anomiidae].
Anomia peruviana
Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Hypotype LACMIP 4531.
[Anomiidae].

Anomia peruviana
member) [=Latrania Fm.], Miocene/Pliocene. [Siltstones cropping out on front of Indio Hills near Willis Palms,
Thousand Palms quad., Riverside Co. UCMP A1417. Hypotype UCMP 37531. [Anomiidae].
Anomia peruviana d’Orbigny, 1846. Powell et al. 2019a. USGS SIR 2019-5021:3, 30, pl. 2
26 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Anomia subcostata-
trania Fm.], Miocene/Pliocene. Carrizo Creek, Colorado Desert, Imperial Co. Lectotype (of Moore 1987: p. C10)
USNM 1865, hypotype USNM 13345a (holotype,Keen and Bentson (1944: p. 25)). Remarks: Supplemental
descriptions in Conrad, 1855b, 1857b, and 1857e. [Anomiidae].


Anomia subcostata
Deguynos Fm., Pliocene. Painted Gorge, Imperial Co. SDSNH 4981]. Hypotype SDSNH 97260. Remarks: Supple-
mental descriptions in Conrad, 1855b, 1857b, amd 1857e. [Anomiidae].
Anomia vaquerosensis 
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 383 [=LACMIP 16105]. Hypotype
UCLA 58217 [=LACMIP 10594]. [Anomiidae].
Anomia

of the Moreno Fm., Midway quad., Alameda Co. UCMP D8150. UCMP unnumbered hypotype lost,  C. Throck-
morton (personal communication, 2002). [Anomiidae].
Apolymetis cf.  (Hanley, 1844). [Tellina-
chegoin Fm. (Siphonalia zone), Pliocene. Southeast slope of Las Paredes, North Dome, Kettleman Hills, Kings Co.
USGS 12826. Hypotype USNM 495817. Remarks: In the genus/subgenus Psammotreta (Leporimetis),  Coan
and Valentich-Scott (2012: p. 617). [Tellinidae].
Apolymetis cf.  (Loel and Corey, 1932). [Macoma
2. Gaviota Fm., Eocene. Ridge between Las Varas Canyon and Dos Pueblos Canyon Goleta quad., Santa Barbara
Co. UCMP B6978. Hypotype SU 9201 (not SU 9321 as in text) [=CASG 70135]. Remarks: Junior synonym of
Heteromacoma rostellata (Clark, 1918),  Addicott (1973b: p. 34). [Tellinidae].
Arca (Arca)

8366]. [Arcidae].
Arca (Arca)
side Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 965 [=LACMIP 16111). Holotype LACMIP
8365. [Arcidae].
Arca (Arca) givensi      
Ventura Co. UCR 4667. Holotype UCR 4667/131, paratypes UCR 4667/132, 4667/133. [Arcidae].
Arca (Arca) hawleyi Reinhart, 1943. Weaver and Kleinpell 1963, Eo-
cene. South of Buelton, east side of Nojoqui Canyon, Santa Barbara Co. UCMP B6932. Hypotypes SU. 9249, 9250
[=CASG 70136, 70137]. [Arcidae].
Arca (Arca) hawleyi
locality description), Eocene. East side of Nojoqui Creek, north of Gaviota Pass, Santa Barbara Co. SU 834. Holo-
type CASG 5343 [=CASG 72046]. [Arcidae].
Arca (Arca) leptogrammica
Miocene. Nipomo quad. (1952), San Luis Obispo Co. UCLA 4180 [=LACMIP 24180]. Holotype UCLA 34997 [=LAC-

Arca (Arca) leptogrammica    
(Phoenix Member) (of Hall, 1962: p. 49), Miocene. Nipoma [sic] quad. (1952), San Luis Obispo Co. UCLA 4180 [=
LACMIP 24180]. Holotype UCLA 34977 [=LACMIP 9228]. [Arcidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 27
Arca cf.  (G.B. Sowerby I, 1833). [Byssoarca
[=Latrania Fm.], Pliocene. Near Travertine Point, Oasis quad., Imperial Co. (not San Diego Co. as in locality de-
scription). UCMP A1707 [=UCMP A1705]. Hypotype UCMP 39867. [Arcidae].
 (G.B. Sowerby I, 1833). [Byssoarca-
mens). Latrania Fm. (Andrade Member), Miocene. Ocotillo Canyon, Imperial Co. [ABDSP 5014.7. Hypotype ABDSP
I9846]. [Arcidae].
Arca (Arca) santamariensis-
cene. Fugler Pt. asphalt beds, southeast of Santa Maria, Santa Barbara Co. [LACMIP 16112]. Holotype CIT 1381
[=LACMIP 4072]. [Arcidae].
Arca santamariensis
Fm. (Cebada Member), Pliocene. Fugler Pt., brachiopod bed, Santa Barbara Co. USGS 14770. Hypotype USNM
560097. [Arcidae].
Arca (Arca) santamariaensis
Member), Pliocene. Asphalt beds at Fugler Pt., 10.2 km southeast (not southwest as in text) of Santa Maria, Santa
Barbara Co. LACMIP 16112. Holotype LACMIP 4072 (ex-CIT 1381). [Arcidae].
Arca cf. 
        
78403, 78404. [Arcidae].
Arca santana
at SDSNH (current whereabouts unknown). Remarks: Ex F. and M. Grouard (Silverado, CA) collection. In the
genus Larkinia,  Moore (1983: p. A42). [Arcidae].
Arca (Arca) sisquocensis
Fugler Pt. asphalt beds, near Santa Maria, Santa Barbara Co. [LACMIP 16112]. Holotype CIT 1382 [=LACMIP
4073]. [Arcidae].
Arca (Arca) sisquocensis Reinhart, 1937. Hertlein and Grant 1972. SDSNH Mem. 2(2B):
31–33. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP
305. Hypotypes LACMIP [4457, 4458]. [Arcidae].
Arca (Arca) sisquocensis
member), Pliocene. Fugler Pt. asphalt beds near Santa Maria, Santa Barbara Co. [LACMIP 16112]. Holotype LAC-
MIP 4073 (ex CIT 1382). [Arcidae].
Arca (Arca) terminumbonis
Pliocene. Near SDSNH 202, Elsmere Canyon, Los Angeles Co. SDSNH 207. Holotype SDSNH 79. [Arcidae].
Arca (Arca) terminumbonis
[=Pico Fm.], Pliocene. Near Elsmere Canyon, San Fernando quad., Los Angeles Co. SDSNH 207. Holotype SDSNH
79. [Arcidae].
Arca trilineata         
Fm. (Arca Zone), Pliocene. Kettleman Hills, Fresno Co. CASG 26805. Hypotypes CASG 5922, 5922A. Remarks:
Anadara,  Moore (1983: p. A40).
[Arcidae].
Arca (Arca) trilineata
       
4665. Hypotype USNM 165534. Remarks:
the genus Anadara,  Moore (1983: p. A40). [Arcidae].
Arca (Anadara) trilineata
Pliocene. Near Millbrae, San Mateo Co. CASG 32670. [Hypotype CASG 8343]. Remarks: Supplemental description
Anadara,  Moore (1983: p. A40). [Arcidae].
28 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Arca trilineata      
source specimen unknown. Remarks:
genus Anadara,  Moore (1983: p. A40). [Arcidae].
Arca
quad., Sonoma Co. UCMP B2062. Hypotype UCMP 32932. [Arcidae].
Arca (Arca  
Alverson Canyon, Carrizo Mtns. [=Coyote Mtn.], Imperial Co. SDSNH 347B. Hypotype SDSNH 1604, determined
here as missing. [Arcidae].
Arca (Arca
to Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4667. Hypotype UCR 4667/131. Remarks:
Arca (Arca) givensi n. sp.,  Squires (1991a: p. 70, 71. [Arcidae].
Arcinella californica (Dall, 1903a). [Echinochama]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p. 62,

[Chamidae].
Arcinella cf.  (Dall,1903a). [Echinochama
[=Latrania Fm.], Pliocene. Near Desert Shores and Travertine Rock, Imperial Co. (not San Diego Co. as in locality
description). USGS M9779 [=USGS M9778]. Hypotype USNM 486908. [Chamidae].
Arcinella californica (Dall, 1903a) [Echinochama
Latrania Fm. (Andrade Member), Miocene. Alverson Canyon, Imperial Co. ABDSP 5005.7. Hypotype ABDSP I959.
[Chamidae].
Arcopsis (Arcopsis) eba (M.A. Hanna, 1927). [Arca
Canyon Shale,” Eocene. At the junction of San Clemente and Rose canyons, La Jolla quad., San Diego Co. UCMP
3990. Holotype UCMP 31047. [Noetiidae].
Arcopsis (Arcopsis) eba (M.A. Hanna, 1927). [Arca
Canyon Shale,” Eocene. On the east side of canyon in bottom of Rose Creek, La Jolla quad., San Diego Co. UCMP
3990. Holotype UCMP 31047. [Noetiidae].
Arcopsis
17917. Hypotype LACMIP 14337. [Noetiidae].
Argopecten abietis abbotti (Hertlein and Grant, 1972). [Chlamys
5. San Diego Fm., Pliocene. Loma Portal, San Diego, San Diego Co. SDSNH 716. Holotype SDSNH 4334. Remarks:
SDSNH 716 was proven to be of Pleistocene age (Deméré 1981: p. 3). [Pectinidae].
Argopecten circularis bramkampi (Durham, 1950). [Aequipecten]. Moore 1984. USGS PP 1228-B:B35–B36, pl. 9,

UCMP A1268. Holotype UCMP 30035. Remarks: Determined here as Argopecten ventricosus bramkampi (Dur-
ham, 1950). [Pectinidae].
Argopecten circularis eldridgei (Arnold, 1906). [Pecten
Fm., Miocene/Pliocene. McKittrick District, Kern Co., USGS 3878. Holotype USNM 164580. Remarks: Determined
here as Argopecten ventricosus eldridgei (Arnold, 1906). [Pectinidae].
Argopecten circularis imposter (G D. Hanna, 1924). [Pecten-
per Etchegoin Fm., Pliocene. Near Zapato Creek, Coalinga quad., Fresno Co. UCMP 2991. Holotype UCMP 11089.
Remarks: Determined here as Argopecten ventricosus impostor (G D. Hanna, 1926). [Pectinidae].
Argopectencrassiradiatus (Clark, 1915). [Pecten      
Miocene. Oyster Pt., Riggs Canyon, Mt. Diablo quad. Contra Costa Co. UCMP 467 (not UCMP 367 as cited in text).
Holotype UCMP 11584. [Pectinidae].
Argopecten deserti (Conrad, 1855b). [Pecten]. Moore 1984. USGS PP 1228-B
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 29
[=Latrania Fm.], Miocene/Pliocene. Carrizo Creek, Colorado Desert, Imperial Co. Holotype USNM
by Moore 1984, however, based on careful observation, USNM 8398 is not the holotype]. [Pectinidae].
Argopecten deserti (Conrad, 1855b). [Pecten]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p. 59,

SDSNH 12003. [Pectinidae].
Argopecten deserti (Conrad, 1855b). [Pecten -
mens). Deguynos Fm. (Mud Hills Member), Pliocene. Fish Creek/Vallecitos Creek, San Diego Co. ABDSP 5089.
Hypotype ABDSP I 10197. [Pectinidae].
Argopecten ericellus (Hertlein, 1929b). [Pecten   

Argopecten invalidus G D. Hanna, 1924). [Pecten

Argopecten invalidus (G D. Hanna, 1924). [Pecten
Pliocene. Southwest of Valencia, Oat Mtn. quad. (1952, PR 1969), Los Angeles Co. LACMIP 15719. Hypotypes
LACMIP 13344,13345. [Pectinidae].
Argopecten invalidus (G D. Hanna, 1924). [Pecten
Newhall area, Los Angeles Co. LACMIP 9659. Hypotypes LACMIP 14342–14344. [Pectinidae].
Argopecten sp. cf.  (Arnold, 1906). [Pecten
Fm. (Latrania Sand Member) [=Latrania Fm.], Miocene/Pliocene. Whitewater/Supercreek area, Riverside Co.
UCR 5035. Hypotype UCR 5035/2 (not UCR 5035/11 as pl. caption). [Pectinidae].
Argopecten mendenhalli (Arnold, 1906). [Pecten]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p.

[Pectinidae].
Argopecten subdolis (Hertlein, 1925b). [Pecten

[Pectinidae].
Argopecten subventricosus (Dall, 1898). [Pecten
Holotype USNM 61246. Remarks: Junior synonym of Argopecten ventricosus (G.B. Sowerby II, 1842),  Coan
et al. (2000: p. 235). [Pectinidae].
Argopecten sverdrupi (Durham, 1950). [Pecten-
trania Sand Member) [=Latrania Fm.], Miocene/Pliocene. Whitewater/Super Creek area, Riverside Co. UCR 5042.
Hypotype UCR 5042/22 (not UCR 5042/23 as in pl. caption). [Pectinidae].
Argopecten sp. Smith 1991a. USGS. Middle fork of Vineyard
Canyon, San Miguel quad., Monterey Co. USGS 16833. Hypotype USNM 334998. Remarks: Described as Lyropecten
terrysmithae n. sp.,  Powell et al. (2020: p. 2–7). [Pectinidae].
-
 
(as Pecten (Lyropecten) estrellanus (Conrad, 1857a). Remarks: Described as Lyropecten terrysmithae n. sp., 
Powell et al. (2020: p. 2–7). [Pectinidae].
Assorted marine invertebrate fossils. Rugh 2001. Quarterly Dose (IDEC Pharmaceuticals Corp.) 21(3):8, unnum-

San Diego Co. SDHHM 4572. Hypotype SDSNH 158089. Remarks: Barbatia morsei (Gabb, 1864),  S. Rugh
(personal communication, 2019).
Atrina alamedensis (Yates in Cooper 1888). [Pinna

30 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Y474 [=SBNHM 35621].
Atrina bicuneata (Nomland, 1917b). [Pinna
Miocene. Sixteen km northeast of Coalinga, Fresno Co. UCMP 2284. Holotype UCMP 11307. [Pinnidae].
Atrina stephensi
Miocene/Pliocene. Coyote Mtn., Imperial Co. SDSNH 53]. Holotype SDSNH
Atrina triangularis-
stone at bottom of narrow gully on east side of Runkle Canyon, Simi Hills, Ventura Co. UCR 6832. Holotype UCR
6832/100 (not UCR 6832/1 as in text). [Pinnidae].
Atrina venturensis (Yates in Cooper, 1888). [Pinna

 Coan and Scott 1990:405]
UCSB Y3096 [=SBNHM 35622].
Atrina           
Ventura Co. [UCR 6832]. Hypotype UCR 6832/1. Remarks: Atrina triangularis n. sp.,  Zinsmeister (1983b:
p. 1286). [Pinnidae].
Axinopsida serricata (Carpenter, 1864a). [Cyptodon]. Hertlein and Grant, 1972. SDSNH Mem. 2 (2B):257–258, pl.

San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 4653. [Thyasiridae].
Barbatia (Cucullaearca) biloba (Weaver, 1905). [Arca]. Squires et al., 1988   
(peel). Goler Fm. (1972), El Paso Mtns., Kern
Co. USGS M9067. Hypotype USNM 424893. Remarks: Junior synonym of Cucullaea mathewsonii Gabb, 1864, 
Moore (1983: p. A47). [Arcidae].
Barbatia (Cucullaearca) cliffensis
Shale, Eocene. South of the mouth of Soledad Valley in ocean cliff, La Jolla quad., San Diego Co. UCMP 5062. Lec-
totype (of Reinhart 1943: p. 32) UCMP 31077, paratypes [=paralectotypes] UCMP 31076, 31081 (all three listed
as syntypes in UCMP online database). [Arcidae].
Barbatia (Cucullaearca) cliffensis
Shale, Eocene. In sea cliff south of mouth of Soledad Valley, La Jolla quad., San Diego Co. UCMP 5062. Lectotype
(of Reinhart 1943: p. 32) UCMP 31077, paratype [=paralectotype] UCMP 31081 (all listed as syntypes in UCMP
online database). [Arcidae].
Barbatia (Cucullaearca) cliffensis
Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 828 [=LACMIP 16141]. Hypotype LACMIP
7507. [Arcidae].
Barbatia (Cucullaearca) cliffensis

Hypotype LACMIP 7507. [Arcidae].
Barbatia cf. (Cucullaearca) cliffensis
Eocene. Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LAC-
MIP 7725. [Arcidae].
Barbatia (Cucullaearca) cliffensis
North side of Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP
13436. [Arcidae].
Barbatia (Fugleria) illota (G.B. Sowerby I, 1833). [Byssoarca]. Hertlein and Grant, 1972. SDSNH Mem. (2B):157–158,
   
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 31
Diego Co. LACMIP 305. Hypotypes LACMIP [4462–4465]. [Arcidae].
Barbatia (Fugleria) illota (G.B. Sowerby I, 1833). [Byssoarca
San Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. [LACMIP 305]. Hypotype (of Hertlein and Grant

Barbatia (Fugleria) illota (G.B. Sowerby I, 1833). Byssoarca
Fm., Pliocene. A mile-long cut west of Oso Parkway near Cabot Rd., Orange Co. [SDSNH 4080]. SDSNH hypotype
determined here as missing. [Arcidae].
Barbatia (Obliqarca) morsei Gabb, 1864. Reinhart 19434. Unknown Fm., Eocene.
Near San Diego, San Diego Co. Lectotype (of Reinhart 1943:31) UCMP 11984 (listed as holotype in UCMP online
database) [paralectotype UCMP 4447]. Remarks: In the subgenus Barbatia,  Moore (1983: p. A34). [Arcidae].
Barbatia (Barbatia) morsei
San Diego, San Diego Co. Lectotype (of Reinhart 1943: p. 31) UCMP 11984 (listed as holotype in UCMP online
database). [Arcidae].
Barbatia (Barbatia) cf.  () morsei  
Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=UCLA 7210, =LACMIP 16114]. Hypo-
type LACMIP 7506. [Arcidae].
Barbatia (Barbatia) morsei
south bank of North Fork of Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP
16963]. LACMIP hypotype 12456. [Arcidae].
Barbatia (Fugleria) pseudoillota    
Pliocene. Fugler Pt. asphalt beds, vicinity of Santa Maria, Santa Barbara Co. [=LACMIP 16112]. Holotype CIT
1383 [=LACMIP 4075]. [Arcidae].
Barbatia pseudoillota  
Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., below 4 ft. fossil bed, southeast of brachiopod bed, Santa
Barbara Co. USGS 14769. Hypotype USNM 560098. Remarks: In the subgenus Fugleria,  Moore (1983: p.
A35). [Arcidae].
Barbatia (Fugleria) pseudoillota

Santa Maria, Santa Barbara Co. [=LACMIP 16112. Holotype CIT 1383 (=LACMIP 4075)]. [Arcidae].
Barbatia (Fugleria) pseudoillota
Fm. (Cebada Member), Pliocene. Asphalt beds at Fugler Pt., southeast of Santa Maria, Santa Barbara Co. [LACMIP
16112]. Holotype LACMIP 4075 (ex-CIT 1381). [Arcidae].
Barbatia
Barbatia morsei)). Domengine Fm. [as Tejon
Fm.], Eocene. West of the San Joaquin Coal mine, west of Coalinga, Fresno Co. Hypotype USNM 165635. [Arcidae].
Barbatia
PR 1969), Ventura Co. LACMIP 25837. Hypotype LACMIP 12748. [Arcidae].
BarbatiaBarbatia sp.]. Imperial Fm. [=Latrania
Fm.], Pliocene. Near Travertine Point, Oasis quad., Imperial Co. (not San Diego Co. as in description). UCMP A1707
[=UCMP A1705]. Hypotype UCMP 39871, missing,  Powell (2008: p. 6) [Arcidae].
Basterotia (Basterotella) hertleini Durham, 1950. Hertlein and Grant 1972. SDSNH Mem. 2(2B): 241–242, pl. 57,
    quad. (1953), San Diego Co. LACMIP 305C [=LAMCIP 16817].
Hypotype LACMIP [4635]. [Sportellidae].
Bornia (Temblorniana) frankiana
Diego Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Holotype LACMIP
32 PALEOBIOS, VOLUME 38, FEBRUARY 2021
[4624]. [Lasaeidae].
Bornia (Temblorniana) frankiana

San Diego Co. LACMIP 305C [=LACMIP 16817]. Holotype LACMIP 4624. [Lasaeidae].
Bornia (Tembloriana) triangulata (Anderson and Martin, 1914). [Donax]. Keen 1943. Tr. SDSNH 10(2):39, pl. 3,

SU 2121. Hypotypes SU 7523, 7523a [=CASG 71994]. [Lasaeidae].
Bornia (Tembloriana) triangulata (Anderson and Martin, 1914). [Donax]. Moore 1992. USGS PP 1228-E:E8, pl. 3,
  
quad., Kern Co. CASG 65. Holotype CASG 130 [=CASG 65.02]. [Lasaeidae].
BotulaPecten Zone),
Pliocene. North Dome, Kettleman Hills, Kings Co. USGS 12335. Hypotype USNM 495739. [Mytilidae].
Brachidontes (Aeidimytilus) adamsianus (Dunker, 1857). [Mytilus
11. San Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. [LACMIP 305C (=LACMIP 16817)]. Hypotype

BrachidontesBrachidontes altiobliquus (Nelson, 1925). [Modiolus]. Moore 1983. USGS PP 1228-A:A67–A68, pl.

Holotype UCMP 30587. [Mytilidae].
Brachidontes (Brachidontes) cooperi Moore, 1983 (new name for Septifer dichotomus Cooper, 1894, preoccupied).

13735. Holotype CASG 613. [Mytilidae].
Brachidontes cowlitzensis (Weaver and Palmer, 1922). [Modiolus]. Weaver and Kleinpell 1963. UCPG 43:197, pl. 29

Santa Barbara Co. UCMP B6966. Hypotype SU 9255 [=CASG 70139]. [Mytilidae].
Brachidontes (Brachidontes) cowlitzensis (Weaver and Palmer, 1922). [Modiolus]. Squires 1984. NHMLAC CS 350:42,

Hypotype LACMIP 6567. [Mytilidae].
Brachidontes (Brachidontes) cowlitzensis (Weaver and Palmer, 1922). [Modiolus]. Squires 1987. NHMLAC CS 388:56,

16080]. Hypotype LACMIP 7510. [Mytilidae].
Brachidontes (Brachidontes) cowlitzensis (Weaver and Palmer, 1922). [Modiolus]. Squires 1988c. NHMLAC CS 398:18,

Hypotype LAMCIP 7727. [Mytilidae].
Brachidontes (Brachidontes) cowlitzensis (Weaver and Palmer, 1922). [Modiolus]. Throckmorton 1988a. USGS OFR
   
        C.
Throckmorton (personal communication, 2002). [Mytilidae].
Brachidontes (Brachidontes) cowlitzensis (Weaver and Palmer, 1922). [Modiolus]. Squires 1994b. SEPM Guidebook
-
potype LACMIP 11630. [Mytilidae].

LACMIP. 11643.

11656.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 33
Brachidontes () cowlitzensis (Weaver and Palmer, 1922). [Modiolus
Fm., Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17850. Hypotype LACMIP
13439. [Mytilidae].
BrachidontesBrachidontes dichotomus (Gabb, 1864). [Septifer
6, 7. Tejon Fm., Eocene. Ft. Tejon, Tejon quad., Kern Co. Holotype UCMP 11993. [Mytilidae].
Brachidontes (Aeidimytilus) gabbi (Clark, 1915). [Modiolus     
8. San Pablo Fm., Miocene. Mt. Diablo quad., Contra Costa Co. UCMP 2040. Holotype UCMP 11550. [Mytilidae].
BrachidontesBrachidontes kirkerensis (Clark, 1918). [Modiolus

Holotype UCMP 11121. [Mytilidae].
Brachidontes lawsoni (Nelson, 1925). [Modiolus
Paleocene. East side Panoche Pass Rd. near the junction of Panoche and Silver creeks, Tumey Hills quad., Fresno
Co. SU 2073 [=UCMP A9717]. Hypotype UCMP 14206. Remarks: Provisionally in the genus Brachidontes
Moore (1983: p. A68). [Mytilidae].
BrachidontesBrachidontes lawsoni (Nelson, 1925). [Modiolus 
5. Martinez Fm. [=Santa Susana Fm.], Paleocene. Simi Hills, Camulos quad., Ventura Co. UCMP 4004. Holotype
UCMP 30651. [Mytilidae].
Brachidontes (Scolimytilus margaritana (Nomland, 1917b). [Septifer]. Moore 1983. USGS PP 1228-A:A69, pl. 17,
Santa Margarita Fm., Miocene. Northeast of Coalinga, Coalinga quad. Fresno Co. UCMP 2276. Holotype
UCMP 11310. [Mytilidae].
BrachidontesScolimytilus multiradiatus (Gabb, 1866). [Modiola
12, 17. Martinez Fm., Paleocene. Martinez, Contra Costa Co. Holotype ANSP 4482. [Mytilidae].
Brachidontes (Scolimytilus subconvexus (Trask, 1922). [Modiolus

BrachidontesBrachidontes susanaensis (Nelson, 1925). [Septifer
2. Martinez Fm. [=Santa Susana Fm.], Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCMP 3776. Holotype
UCMP 30501. [Mytilidae].
Bruetiatraski (Nelson, 1925). [Diplodonta         
(Martinez Fm. in pl. caption), Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCMP 3768. Holotype UCMP
30579. [Ungulinidae].
Callista (Macrocallista) andersoni (Dickerson, 1915). [Macrocallista
Fm., Eocene. Lion Canyon quad. (1943), Ventura Co. UCR 4731. Hypotype UCR 4731/101. [Veneridae].
Callista andersoni (Dickerson, 1915). [Macrocallista
Shale (Circle B Sandstone Member), Eocene. Upper Sespe Creek, Ventura Co. CSUN 322 [=LACMIP 16669]. Hy-
potype LACMIP 11645. [Veneridae].
Callista conradiana (Gabb, 1864). [Tapes
Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype
SDSNH 20765. [Veneridae].
Callista conradiana (Gabb, 1864). [Tapes
Eocene. Upper Sespe Creek, Ventura Co. LACMIP 20581. Hypotype LACMIP 11665. [Veneridae].
Callista (Macrocallista) domenginica
Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP
7537. [Veneridae].
Callista (Macrocallista) domenginica
34 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13451. [Veneridae].
Callista (Costacallista) hornii (Gabb, 1864). [Meretrix
Northeast of Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4723. Hypotype UCR 4723/61. [Ven-
eridae].
Callista (Costacallista) cf. () hornii (Gabb, 1864). [Meretrix]. Squires 1984. 
Fm., Eocene. Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 465 [=LACMIP 16149]. Hypotype LACMIP
6588. [Veneridae].
Callista hornii (Gabb, 1864). [Meretrix
(Circle B Sandstone Member), Eocene. Upper Sespe Creek, Ventura Co. CSUN 322 [=LACMIP 16669]. Hypotype
LACMIP 11644. [Veneridae].
Callista (Costacallista) hornii vokesi
Eocene. South portion of crest of Parson's Peak, Coalinga quad., Fresno Co. UCMP 672. Holotype UCMP 11667.
[Veneridae].
Callista (Costacallista) hornii vokesi
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 978 [=LACMIP 16150]. Hypotype LACMIP 7733.
[Veneridae].
Callista (Costacallista) hornii vokesi 

Road, Midway quad., Alameda Co. UCMP D8152. UCMP unnumbered hypotype lost,  C. Throckmorton (per-
sonal communication, 2002). [Veneridae].
Callista (Costacallista) hornii vokesi
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 675 [=LACMIP 16151]. Hypotype
LACMIP 11486. [Veneridae].
Callithaca tenerrima (Carpenter in Gould and Carpenter, 1857). [Tapes
Pico Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14358. [Veneridae].
Callocardia (Nitidavenus) tejonensis (Waring, 1914). [Isocardia
Fm., Eocene. Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6589. [Veneridae].
Callocardia (Nitidavenus) tejonensis (Waring, 1914). [Isocardia

Hypotype LACMIP 7538. [Veneridae].
Calva (Calva) baptisia
Taliaferro 1944:512–517), uppermost Cretaceous/lowermost Paleocene. South side Lake Nacimiento, east side
of Narrows of Dip Creek, Lime Mtn. quad., San Luis Obispo Co. UCLA 6526 [=LACMIP 26526]. Holotype LACMIP
7559, paratypes LACMIP 7560, 8234, 8235. [Veneridae].
Calva cf.  (Gabb, 1864). [Venus-
most Cretaceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co.
UCLA 6526 [=LACMIP 26526]. Hypotypes LACMIP 7559, 7560. Remarks: Calva (Calva) baptista n. sp.,  Saul
and Popenoe (1992: p. 24). [Veneridae].
Calyptogena (Phreagena) lasia (Woodring, 1938). [Phreagena]. Cox et al. 1969b. Treatise Invert. Paleo. N6(2):N664,
Woodring 1938 tunnel, Los Angeles,
Los Angeles Co. USGS 3426. Paratype USNM 496098. Remarks: Junior synonym of Calyptogena (Calyptogena)
gibbera Crickmay, 1929,  Squires (1991b: p. 74). [Vesicomyidae].

Holotype USNM 496097].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 35
Calyptogena (Calyptogena) lasia (Woodring, 1938). [Phreagena
Towsley Fm., Pliocene. Between Tapo and an unnamed canyon west of Salt Canyon, north side of Santa Susana
Mtns., Val Verde quad. (1952), Ventura Co. LACMIP 21363. Hypotypes LACMIP 8401, 8402. Remarks: Junior
synonym of Calyptogena (Calyptogena) gibbera Crickmay, 1929,  Squires (1991b: p. 74). [Vesicomyidae].

Cardiomya israelskyi (M.A. Hanna, 1927). [Cuspidaria-
cinity of Park Canyon, San Guillermo quad. (1943), Ventura Co. UCR 4700. Hypotype UCR 4700/2. [Cuspidariidae].
Cardiomya aff.  (M.A. Hanna, 1927). [Cuspidaria
Eocene. Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 512d [=LACMIP 16161]. Hypotype
LACMIP 6594. [Cuspidariidae].
Cardita n. sp. aff. 
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2859. Hypotype USNM
254370. [Carditidae].
Cardita n. sp. aff. -

Luis Obispo Co. USGS M2859. Hypotype USNM 254370. [Carditidae].
Cardita californica Dall, 1903b. [Venericardia (Cyclocardia)]. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):176,
Re-
marks: In the genus Cyclocardia,  Coan et al. (2000: p. 303). [Carditidae].
Cardita (Cyclocardia) californica Dall, 1903b. [Venericardia (Cyclocardia)] Cummings et al. 1962. CDMG Bull. 181:204,
Balanus cf. aqulia Pilsbry as in pl. caption). Purisima Fm. (Pomponio Member), Pliocene. Tunitas
Creek, San Mateo Co. Locality 653M. Hypotypes determined here as missing. Remarks: In the genus Cyclocardia,
 Coan et al. (2000: p. 303). [Carditidae].
Cardita (Cardita) superioris
McCray wells (Oil Canyon), Santa Susana quad., Ventura Co. SU 2696 [=CASG 61913]. Holotype SU/CASG 143
[=CASG 61913.06]. [Carditidae].
Cardiidae, indeterminate. Powell et al. 2019a. USGS SIR.

Cardium (Schedocardia) brewerii
Fm., Eocene. Live Oak Creek east of the mouth of Grapevine Canyon, Kern Co. CASG 244. Hypotype CASG 5701.
Remarks: In the genus/subgenus Acanthocardia (Schedocardia),  Moore (2003a: p. 3). [Cardiidae].
Cardium (Trachycardium) brewerii brewerii
Undifferentiated Sacate/Gaviota fms., Eocene. In small gully just east of Nojoqui Creek, Santa Ynez Mtns., Lompoc
quad., Santa Barbara Co. UCMP B6960. Hypotype SU 9288 [=CASG 70141]. Remarks: Label in CASG collection
Acan-
thocardia (Schedocardia),  Moore (2003a: p. 3). [Cardiidae].
   
Canyon, Goleta quad., Santa Barbara Co. UCMP B6934. Hypotype SU 9289 [=CASG 70142]. Remarks: Label in

Cardium (Trachycardium) brewerii robusta
Middle Gaviota Fm., Eocene. East of gorge in Gaviota Canyon, Lompoc quad., Santa Barbara Co. UCMP B7010.
Holotype SU.9290, missing. Remarks: In the genus/subgenus Acanthocardia (Schedocardia),  Moore (2003a:
p. 4). [Cardiidae].
Cardium linteum
Cardium cooperi Gabb, 1864)). Tejon Fm., Eocene. [Coalinga District,
Fresno Co. USGS 4617. Hypotype USNM 165637]. Remarks: In the genus Nemocardium,  Moore (2003a: p.
36 PALEOBIOS, VOLUME 38, FEBRUARY 2021
14). [Cardiidae].
Cardium (Nemocardium) linteum
Sandstone, Eocene. Nojoqui Creek area, west of Nojoqui creek, Lompoc quad., Santa Barbara Co. UCMP B6919.
Hypotype SU 9291 [=CASG 70143]. Remarks: In the genus Nemocardium,  Moore (2003a: p. 14). [Cardiidae].
Cardium (Cerastoderma) meekianum
Pliocene. On the Mad River Rd., near the mouth of Canon Creek, Humboldt Co. UCMP A4234. [Hypotype UCMP
34177]. Remarks: In the genus Clinocardium,  Moore (2003a: p. 20). [Cardiidae].
Cardium (Dallocardia) quadragenarium Conrad, 1837. Hertlein and Grant 1972. SDSNH Mem. 2(2B):259–260, pl.
     
LACMIP 305. Hypotype LACMIP 4654. Remarks: In the genus Dallocardia,  Coan and Valentich-Scott (2012:
p. 413). [Cardiidae].

Cardium Fowkes 1982. Coalinga Guidebook, p. 21, unnumbered Arnold 1909 [1910]
3; Arnold and Anderson 1910 as Cardium cooperi Gabb, 1864)]. Domengine Fm.
[=Tejon Fm.], Eocene. Fresno Co. No locality number
given. [Hypotype USNM 165637]. [Cardiidae].

 Cardium meekianum Gabb, 1866)]. Etchegoin Fm., Pliocene/Miocene. Southeast of
Coalinga, Fresno Co. Hypotype USNM 165542.
Cardium
Posa area, Santa Cruz Is. California Channel Ids. Hypotype UCSB 8772 [=SBMNH 60182]. Remarks: Determined
here as indet. venerid sp. [Veneridae}
Cerastoderma cf.  (Gabb, 1866). [Cardium
9. San Joaquin Fm. (Cerastoderma-bearing sandstone), Pliocene. West branch of Arroyo Doblegado, North Dome,
Kettleman Hills, Kings Co. USGS 12706. Hypotype USNM 495345. Remarks: Clinocardium meekianum (Gabb,
1866),  Kafanov (1998: p. 37). [Cardiidae].
Pseudocardium Zone), Pliocene. North branch of Arroyo Torcido, east of Double
Hill, North Dome, Kettleman Hills, Kings Co. USGS 12400. Hypotype USNM 495796.
Cerastoderma cf. 
3. Etchegoin Fm. (Littorina zone), Pliocene. Arroyo Mellado, south of Los Jinetes, North Dome, Kettleman Hills,
Kings Co. USGS 12392. Hypotype USNM 495784. Remarks: Clinocardium meekianum (Gabb, 1866),  Kafanov
(1998: p. 37). [Cardiidae].
Burrow of Chaceia ovoidea (Gould, 1851). [Pholas
in part], Miocene (possibly the Temblor and Santa Margarita fossils were slightly mixed in locs. 28485). One mi.
southeast of Oil City, Fresno Co. CASG 28485 (this locality represents at least two different time periods, one of
which is probably Pliocene). Hypotype CASG 54825.01. Remarks: Chaceia sp. aff.  (Gould, 1851), 
 Chaceia fulcherae n. sp.,
 Kennedy (1993: p. 399).
Chaceia fulcherae
Temblor Reef, Jasper Canyon, Fresno Co. CASG 1841 [=CASG 54826]. Paratype CASG 54826.01. [Pholadidae].

54825]. Holotype CASG 54825.01.

    
LACMIP 11444, 11445.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 37
Chaceia ovoidea (Gould, 1851). [Pholas
Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14373. Remarks: Undetermined species of Penitella,  G.L.
Kennedy (personal communication, 2019). [Pholadidae].
ChaceiaAdegoke 1966. The Veliger 9(2):234, pl-
ocene. San Luis Obispo area, San Luis Obispo Co. UCMP A3415. Hypotypes UCMP 12197, 15547, 31409, 31410,
32522, 32523, 34092, all determined here as missing. Remarks: Penitella sp. indet.,  Kennedy (1974: p. 37).
[Pholadidae].
Chama (Chama) arcana-

12, 15 as Chama pellucida Broderip, 1835) LACMIP. 4534. [Chamidae].
Chama pellucida
San Joaquin Fm. (Pecten zone), Pliocene. Arroyo Pequeño and Arroyo Finito areas, North Dome, Kettleman Hills,
Kings Co. USGS 12343. Hypotypes USNM 495744 and 495753. Remarks: Chama arcana Bernard, 1976,  Coan
et al. (2000: p. 309). [Chamidae].



Chama pellucida  
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotype
LACMIP 4534. Remarks: Chama arcana Bernard, 1976,  Coan et al. (2000: p. 309). [Chamidae].
Chama pellucida
SDSNH 4080. Hypotype SDSNH 75918]. Remarks: Chama arcana Bernard, 1976,  Coan et al. (2000: p. 309).
[Chamidae].
Chama piruensis
Los Angeles Co. CSUN 822 [=LACMIP 16163]. Holotype LACMIP 7520. [Chamidae].



Chama (ChamaMoore, 1988b4, 5. Vaqueros Fm., Oligocene/Miocene. [Torrey
Canyon, Oak Ridge, 
[Chamidae].
Chama
Travertine Rock area, Imperial Co. (not San Diego Co. as in description). USGS M9779 [=USGS M9778]. Hypotype
USNM 486906. [Chamidae].
Chlamys (Argopecten) abietis abbotti Hertlein and Grant, 1972 (n. subsp.). SDSNH Mem. 2 (2B): 196–197, pl. 34,

Holotype SDSNH 4334. Remarks: Valid subspecies in the genus Argopecten,  Moore (1984: p. B42). SDSNH
716 proven to be of Pleistocene age (Deméré 1981: p. 3). [Pectinidae].

Chlamys bartschi (Arnold, 1906). [Pecten
South side of Light-House [sic] Pt., Crescent City, Del Norte Co. USGS. 3352. Holotype USNM 164841. [Pectinidae].
Chlamys (Leptopecten) bellilamellata (Arnold, 1906). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):203,
38 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Remarks: Valid species in the genus Leptopecten,  Moore (1984: p. B43); junior synonym of Leptopecten
latiauratus (Conrad, 1837),  Coan et al. (2000: p. 237). [Pectinidae].


Chlamys (Vertipecten) bowersi (Arnold, 1906). [Pecten

Santa Barbara Co. Holotype UCMP 12075 [=Vertipecten nevadensisRemarks: In the genus Vertipecten, 
Moore (1984: p. B62). [Pectinidae].
Chlamys branneri (Arnold, 1906). [Pecten
Unnamed strata (of Dibblee 1966), Miocene. Tuff Hill, near Stanford University, San Mateo Co. Holotype SU/CASG
358 [=CASG 61953.01]. [Pectinidae].
Chlamys calkinsi (Arnold, 1906). [Pecten
Eocene. Sisar Valley, Santa Paula quad., Ventura Co. Holotype UCMP 12076. [Pectinidae].
Chlamys calkinsi (Arnold, 1906). [Pecten
Eocene. Upper Sespe creek, Ventura Co. CSUN 74 [=LACMIP 16671]. Hypotype LACMIP 11658. [Pectinidae].
Chlamys (Argopecten) callida (Hertlein, 1925b). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):198–199,
Remarks:
Argopecten callidus (Hertlein, 1925b),  Moore (1984: p. B38). [Pectinidae].
....., pl. 
Chlamys (Lyropecten) cerrosensis (Gabb, 1866). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):209–211, pl.

Hypotype LACMIP 4505. Remarks: In the genus Lyropecten,  Moore (1984: p. B55). [Pectinidae].

Chlamys (Argopecten) circularis (G.
B. Sowerby I, 1835b). [Pecten 
Diego Fm., Pliocene. San Diego, San Diego Co. CASG 1137. Hypotype CASG 72544. Remarks: Junior synonym of
Argopecten ventricosus (G.B. Sowerby II, 1842),  Coan et al. (2000: p. 235). [Pectinidae].


Pliocene. Northwest side of Carrizo Mtn., Imperial Co. UCMP A1267. Paratypes UCMP 34031, 34032. [Pectinidae].
Chlamys cf. 
Pliocene. Vicinity of Desert Shores and Travertine Rock, Imperial Co. (not San Diego Co. as in description). USGS
M9778. Hypotype USNM 486903. [Pectinidae].
Chlamys (Lyropecten) crassicardo (Conrad, 1857a). [Pecten]. Cox et al. 1969a. Treatise Invert. Paleo. N6(1):N358,

Hypotype USNM 154154]. Remarks: In the genus Lyropecten,  Moore (1984: p. B51). [Pectinidae].
Chlamys crassicardo-
sibly at SDSNH (current whereabouts unknown). Remarks: Ex Grouard collection. In the genus Lyropecten, 
Moore (1984: p. B51). [Pectinidae].
Chlamys (Lyropecten) crassicardo (Conrad, 1857a). [Pecten
Remarks:
In the genus Lyropecten,  Moore (1984: p. B51). [Pectinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 39
Chlamys durhami
Ranch quad., Fresno Co. UCMP D1099. Holotype UCMP 36615. Remarks: Junior synonym of Chlamys hodgei
(Hertlein, 1925a),  Moore (1984: p. B21). [Pectinidae].
Chlamys egregius (Nomland, 1917a). [Pecten
Near Gualala School, Plantation quad., Sonoma Co. UCMP B2062. Hypotype UCMP 32915. [Pectinidae].
Chlamys egregius (Nomland, 1917a). [Pecten]. Moore 1984. USGS PP 1228-B
Fm., Pliocene. Near Zapato Creek, Coalinga quad., Fresno Co. UCMP 2991. Holotype UCMP 11090. [Pectinidae].
Chlamys n. sp., aff.  (Nomland, 1917a). [Pecten]. Powell et al. 2019a, USGS SIR 2019-5021:3 (as Chlamys
aff Chlamys aff. 
Sonoma Co. CASG 54135. Hypotypes CASG 78407, 78408. [Pectinidae].
Chlamys (Argopecten) ericellus (Hertlein, 1929b). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):199, pl.
Remarks: In
the genus Argopecten,  Moore (1984: p. B37). [Pectinidae].
Chlamys (Lyropecten) estrellana (Conrad, 1857a). [Pallium]. Cox et al. 1969a. Treatise Invert. Paleo. N6(1):N357,

Remarks: Supplemental descriptions
Lyropecten estrellanus (Conrad, 1857a),  Moore
(1984: p. B53). [Pectinidae]

Obispo Co. Cotype [=syntype] USNM 13317].
Chlamys etchegoini (Anderson, 1905). [Pecten          
Joaquin Fm. (Pecten Zone), Pliocene. East fork of Surprise Canyon, North Dome, Kettleman Hills, Kings Co. USGS
12322. Hypotype USNM 495731. Remarks: Swiftopecten parmeleei etchegoini (Anderson, 1905),  Moore
(1984: p. B60). [Pectinidae].
Siphonalia Zone), Pliocene. East end of La Muralla, North Dome, Kettleman Hills,
Kings Co. USGS 12434. Hypotype USNM 495816.
Chlamys (Argopecten) hakei (Hertlein, 1925b). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):199–200, pl.
Remarks: In
the genus Argopecten,  Moore (1984: p. B39). [Pectinidae].
Chlamys (Chlamys) hastata (G.B. Sowerby II, 1842). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):188–189,
-
tinidae].

Chlamys (Chlamys) hastata (G.B. Sowerby II, 1842). [Pecten ]. Vedder and Moore 1976. AAPG Misc. Pub. 24:123, 134,

235601. [Pectinidae].
Chlamys cf.  (G.B. Sowerby II, 1842). [Pecten

Chlamys (Chlamys) hastata ellisi Hertlein and Grant, 1972 (n. subsp.). 
2, 3; Diego Co., San Ysidro quad. (1943), San Diego Co.
LACMIP 305. Holotype LACMIP 4483, paratype LACMIP 4484 Re-
marks: Junior synonym of Chlamys hastata (G.B. Sowerby II, 1842),  Coan et al. (2000: p. 233). [Pectinidae].
Chlamys (Chlamys) hastata ellisi
Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Holotype LACMIP 4483. Remarks: Junior
synonym of Chlamys hastata (G.B. Sowerby II, 1842),  Coan et al. (2000: p. 233). [Pectinidae].
40 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Chlamys (Chlamys) hastata hericius (Gould, 1850b). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):189–190,
Remarks:
Junior synonym of Chlamys hastata (G.B. Sowerby II, 1842),  Coan et al. (2000: p. 233). [Pectinidae].
Chlamys (Chlamys) hastata hericius (Gould, 1850b). [Pecten
Fm., Pliocene. Orange Co. SDSNH4080. Hypotype SDSNH 75916. Remarks: Junior synonym of Chlamys hastata
(G.B. Sowerby II, 1842),  Coan et al. (2000: p. 233). [Pectinidae].
Chlamys hertleini (Loel and Corey, 1932). [Pecten
Hill sand” (=Jewett Sand of Addicott 1965b: p. C104), Miocene. Pyramid Hill, Kern Co. UCLA MB2000 [=LACMIP
16168]. Hypotype UCLA 39412 [=LACMIP 8428]. [Pectinidae].
Chlamys hertleini (Loel and Corey, 1932). [Pecten
Hill sand (=Jewett Sand of Addicott 1965b), Miocene. Pyramid Hill, Kern Co. UCLA MB2000 [=LACMIP 16168].
Hypotype UCLA 33491 [=LACMIP 11039]. [Pectinidae].
      Ostrea biostrome, Woody quad. (1952), Kern Co. UCMP
B1654. Hypotype UCMP 32254.
Chlamys cf.  (Loel and Corey, 1932). [Pecten 
Basal Skooner Gulch Fm., Miocene. South of Pt. Arena, Mendocino Co. USGS M2638. Hypotypes USNM 649592,
649595. [Pectinidae].
Chlamys hertleini (Loel and Corey, 1932). [Pecten
Oligocene/Miocene. Lompoc quad., Santa Barbara Co. UCMP A316. Holotype UCMP 31725. [Pectinidae].
Chlamys hodgei (Hertlein, 1925a). [Pecten -
cene. Humphreys quad. (1932, RP 1946), Los Angeles Co. CIT 1663 [=LACMIP 2555]. [Hypotype LACMIP 4094].
[Pectinidae].
Chlamys hodgei (Hertlein, 1925a). [Pecten
Wildhorse Canyon, King City quad., Monterey Co. UCMP A906. Hypotype UCMP 14585. [Pectinidae].
of Reliz Canyon, Reliz Canyon quad., Monterey Co. USGS M902. Hypotype USNM 254388.
Chalmys hodgei (Hertlein, 1925a). [Pecten
Fm., Miocene. Coalinga region, Fresno Co. SU F-6 [=CASG 61843]. Holotype SU 20 [=CASG 61843.01], paratype
SU 21 [=CASG 61843.02]. [Pectinidae].
Chlamys durhami Adegoke, 1969)
UCMP 36615.
Chlamys hodgei (Hertlein, 1925a). [Pecten]. Powell et al. 2019a. USGS SIR 2019-5021:3 (as Chlamys cf. ), 30,

Chlamys (Argopecten) invalida (G D. Hanna, 1924). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):200–201,
Remarks:
In the genus Argopecten,  Moore (1984: p. B37). [Pectinidae].


Hypotype CASG 55071.
Chlamys (Chlamys) jordani (Arnold, 1903). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):191–192, pl. 30,

Remarks: Junior synonym of Chlamys rubida (Hinds, 1845),  Coan et al. (2000: p. 234). [Pectinidae].
Chlamys (Argopecten) jordani in pl. caption). Balboa Park, San Diego, San Diego Co. LACMIP
107. Hypotypes LACMIP 4487, 4488].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 41
Chlamys (Argopecten) jordani
CASG 72556.
Chlamys (Chlamys) cf. () jordani (Arnold, 1903). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):191–192,
Remarks: Junior
synonym, of Chlamys rubida (Hinds, 1845),  Coan et al. (2000: p. 234). [Pectinidae].
Chlamys (Leptopecten) latiaurata (Conrad, 1837). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):203–204,
   
20312]. Hypotype UCLA 48612 (=LACMIP 9293). Remarks: Leptopecten latiauratus,  Coan et al. (2000: p.
237). [Pectinidae].
Chlamys (Macrochlamis) magnolia (Conrad, 1857d). [Pecten]. Stadum 1973 Fm.,
Miocene. [Ojai Valley, Ventura Co. UCLA 1311 [=LACMIP 21311]. Hypotype UCLA 49513 [=LACMIP 10611]. Re-
marks: In the genus Macrochlamis,  Moore (1984: p. B56). [Pectinidae].
Chlamysmediacostata mediacostata (G D. Hanna, 1926). [Pecten
Imperial Fm. [=Latrania Fm.], Miocene/Pliocene. Alverson Canyon, near source, Coyote Mtn., Imperial Co. CASG
681. Holotype CASG 1830. [Pectinidae].
Chlamys cf.  (Reeve, 1853). [Pecten
Gulch, Miocene. South of Pt. Arena, Mendocino Co. USGS M2638. Hypotype USNM 649593. [Pectinidae].
Chlamys (Vertipecten) nevadanus (Conrad, 1855b). [Pecten
Miocene (Cold Creek Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22487 [=UCLA
2487]. Hypotype LACMIP 13083 [=UCLA 57937]. Remarks: Junior synonym of Vertipecten kernensis (Hertlein,
1925a),  Moore (1984: p. B64). [Pectinidae].
Chlamys (Chlamys) opuntia (Dall, 1898). [Pecten]. Hertlein and Grant, 1972. SDSNH Mem. 2 (2B):192–193, pl. 30,

   
LACMIP 4489.
Chlamys (Chlamys) opuntia (Dall, 1898). [Pecten
5. Unnamed strata, Pliocene. Southeast of Wilson Cove, San Clemente Is., California Channel Ids. USGS M6503.
Hypotype USNM 235582. [Pectinidae].
Chlamys (Chlamys) opuntia (Dall, 1898). [Pecten
strata, Pliocene. San Clemente Is., San Clemente Is. North quad., California Channel Ids. UCLA 6316 [=LACMIP
26316]. Hypotype UCLA 38797 [=LACMIP 10835]. [Pectinidae].
Chlamys (Chlamys) opuntia (Dall, 1898). [Pecten

[=LACMIP 16173]. Hypotype UCLA 59403 [=LACMIP 8316]. [Pectinidae].
Chlamys opuntia (Dall, 1898). [Pecten-
Remarks: Keen and Bentson 1944: p. 91cited USNM
107752 and 7938 as syntypes but 7938 is not labeled as such in USNM collection. [Pectinidae].
Chlamys opuntia (Dall,1898). [Pecten

Chlamys parmeleei (Dall, 1898). [Pecten]. Woodring and Bramlette 1950 [1951]. USGS PP 222:83, 
Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. Alex Clark locality. Hypotype USNM 560101.
Remarks: In the genus Swiftopecten,  Moore (1984: p. B59). [Pectinidae].
Chlamys parmeleei (Dall, 1898). [Pecten
(Tahana Member), Pliocene. Sea cliff south of mouth of Pomponio Creek, San Gregorio quad., San Mateo Co. Lo-
cality F467-7M. Hypotype determined here as missing. Remarks: In the genus Swiftopecten,  Moore (1984:
42 PALEOBIOS, VOLUME 38, FEBRUARY 2021
p. B59). [Pectinidae].
Chlamys (Swiftopecten) parmeleei (Dall, 1898). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):206–208,

Remarks: In the genus Swiftopecten,  Moore (1984: p. B59). [Pectinidae].

305. Hypotypes LACMIP 4494–4503.
Chlamys (Swiftopecten) parmeleei (Dall, 1898). [Pecten
Pliocene. Orange Co. [SDSNH 4080]. Hypotype determined here as missing. Remarks: In the genus Swiftopecten,
 Moore (1984: p. B59). [Pectinidae].
Chlamys proavus (Arnold, 1906). [Pecten-
cene. Between headwaters of San Lorenzo River and Pescadero Creek, just west of Santa Cruz Co., Santa Cruz
quad., San Mateo Co. SU 2694. Holotype USNM 164930. [Pectinidae].
Chlamys (Chlamys) rubida (Hinds, 1845). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):193–195, pl. 35,


4490.
Chlamys (Chlamys) rubida (Hinds, 1845). [Pecten
1. Unnamed strata, Pliocene. San Clemente Is., California Channel Ids. USGS M6501. Hypotype USNM 235589.
[Pectinidae].
Chlamys (Chlamys) rubida jordani (Arnold, 1903). [Pecten
Fm., Pliocene. Tustin quad., Orange Co. USGS M2130. Hypotype USNM 644945. Remarks: Junior synonym of
Chlamys rubida (Hinds, 1845),  Coan et al. (2000: p. 234). [Pectinidae].
Chlamys sanctiludovici (Anderson and Martin, 1914). [Pecten
Margarita Fm., Miocene. La Panza quad., San Luis Obispo Co. Holotype CASG 107, paratype CASG 108. [Pectinidae].
Chlamys sespeensis (Arnold, 1906). [Pecten     
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns, Ventura Co. CSUN 36 [=LACMIP 16173 ]. Hypotype
UCLA 58197 [=LACMIP 10591]. [Pectinidae].
Chlamys sespeensis (Arnold, 1906) of Loel and Corey 1932. Moore 1984, 3. Vaqueros
Fm., Oligocene/Miocene. South Mtn. near Santa Paula, Piru quad., Ventura Co. UCMP A244]. Hypotypes (of Loel
and Corey 1932
Chlamys sespeensis hydei (Arnold, 1906). [Pecten
Fm., Oligocene/Miocene. Lynch's Mtn., Monterey Co. Holotype SU/CASG 10 [=CASG 61859.01]. [Pectinidae].
Chlamys (Argopecten) subdola (Hertlein, 1925b). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):201–202, pl.
Remarks:
Argopecten subdolus (Hertlein, 1925b),  Moore (1984: p. B37). [Pectinidae].


Chlamys swifti (Bernardi, 1858). [Pecten (Pallium)
Hypotype possibly at SDSNH (current whereabouts unknown). Remarks: Ex Grouard collection. Swiftopecten
parmeleei parmeleei (Dall, 1898),  Moore (1984: p. B59). [Pectinidae].
Chione (Chione) allisoni
Fm., Pliocene. San Diego Co. SDSU 223 [SDSNH 5063]. Paratypes SDSNH 4321, 4322. Remarks: Junior synonym
of Chione (Chione) californiensis (Broderip in Broderip and G.B. Sowerby I, 1835),  Coan et al. (2000: p. 369).
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 43
[Veneridae].

CASG 70422].

Chione (Chionopsis) coalingensis
Pliocene. Domengine Ranch quad., Fresno Co. UCMP D1104. Holotype UCMP 36692, paratype UCMP 36693.
[Veneridae].
Chione (Anomalocardia) fernandoensis-
nando Fm.” [=Pico Fm.], Pliocene. Elsmere Canyon, San Fernando quad., Los Angeles Co.. UCMP 1735. Paratype
UCMP 11921. [Veneridae].

4a, b (as Venus (Chione) securis var. fernandoensis) SDSNH 159.
Chione (Anomalocardia) fernandoensis
Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14357. [Veneridae].
Chione (Anomalocardia) hannai Bramkamp, [MS]. Parker 1949
Fm. [=Latrania Fm.], Pliocene. San Jacinto quad., Imperial Co. UCMP A1407. Holotype UCMP 36087. [Veneridae].
Chione (Securella)
Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Paratype LACMIP




Chione (Anomalocardia) latilaminosa
6, 11. Temblor Fm., Miocene. Kern River, Kern Co. UCMP 2713. Hypotype UCMP 33063. [Veneridae].
Chione (Lirophora) aff.  (d'Orbigny, 1846). [Venus
Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, west side of George
F. Canyon, Palos Verdes Hills, Los Angeles Co. USGS 13127. Hypotype USNM 497088. [Veneridae].
Chione peabodyi-
Remarks: Junior synonym of Chione
(Chione) californiensis (Broderip, in Broderip and G.B. Sowerby I 1835),  Coan et al. (2000: p. 369). [Veneridae].
Chione richthofeni
Rincon Fm., Miocene. Upper Cañada Garanon, west central Santa Rosa Is., California Channel Ids. UCSB SR64-4P
[=LACMIP 16420]. Hypotype UCSB M1 [=LACMIP 6321]. [Veneridae].
Chione cf. 
7. Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 150 [=LACMIP 16167]. Hypotype
UCLA 58208 [=LACMIP 10598]. [Veneridae].
Chione cf. richthofeni    

Chione cf. -
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4312. Hypotype SDSNH 83248. [Veneridae].
Chione securis (Shumard, 1858). [Venus]. Manning and Ogle 1950. B. Falor Fm., Pliocene.
44 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Near the bridge crossing Boulder Creek, adjacent to the Wiggins Ranch, Humboldt Co. UCMP A4233. Hypotype
UCMP 34170. [Veneridae].
Chione semiplicata
UCMP A2569. Hypotype UCMP 30674. Remarks: In the subgenus Chionopsis,  Hall (1958: p. 55). [Veneridae].
Chione (Gnidiella) semiplicata
Santa Margarita Fm., Miocene. Northeast of Coalinga, Fresno Co. UCMP 2283. Holotype UCMP 11318. Remarks:
In the subgenus Chionopsis,  Hall (1958: p. 55). [Veneridae].

Chione (Chionopsis) simplicata*-
west of Pleasanton, Dublin quad. (1953), Alameda Co. SU 3229. Hypotype SU 8462 [=CASG 69959]. Remarks:
Error for semiplicata. [Veneridae].
Chione (Gnidiella) temblorensis (Anderson, 1905). [Venus
   
[Veneridae].
Chione (Chionopsis) temblorensis (Anderson, 1905). [Venus
Sand, Miocene. Caliente quad. (1914), Kern Co. UCMP B1599. Hypotype UCMP 36564. [Veneridae].
Chione temblorensis (Anderson, 1905). [Venus
(as Chione sp. in text). Vaqueros Fm., Miocene. Sea cliff north of Pozo Anchorage, Santa Cruz Is., California Chan-
nel Ids. UCSB RB65-6 [=LAMCIP 16421]. Hypotype UCSB M2 [=LACMIP 6303]. [Veneridae].
Chione temblorensis (Anderson, 1905). [Venus
Topanga Canyon Rd., Los Angeles Co. UCLA 2632 [=LACMIP 22632]. Hypotype UCLA 49515 [=LACMIP 10614].
[Veneridae].
Chione temblorensisVenus]. Shapiro 1998. AAPG Guidebook MP 45:129, pl. 1 (pl. 2 in cap-

[=SBMNH 60183]. Remarks: Determined here as indet. Carditidae sp. [Carditidae].
Chione vickeryi  Chione cf. ). Briones Fm., Mio-
cene. East of Irvington, La Costa Valley quad. (1953), Alameda Co. SU 3252. Hypotype SU 8463 [=CASG 69946].
[Veneridae].
Chione

Remarks:Clinocardium sp., Macoma
sp., and Protothaca sp. [=Leukoma sp.]. [Veneridae].
Ciliatocardium kirkerense* (Clark, 1918). [Cardium]. Kafanov 2002. Bull. Mizunami Foss. Mus. 29:6, pl. 3, 
2. Kirker Tuff, Oligocene. West of Kirker’s Creek, Diablo Quadangle, Contra Costa Co. [UCMP 2033]. Holotype
UCMP 11165. Remarks: In the genus Acanthocardia Moore (2003a: p. 2). Error for[Cardiidae].
Ciliatocardium reedi (Loel and Corey, 1932). [Cardium
Vaqueros Fm., Oligocene/Miocene. Anderson Creek, La Panza Mtns., San Luis Obispo Co. [UCMP A504]. Holotype
UCMP 31772. Remarks: In the genus Acanthocardia,  Moore (2003a: p. 1). [Cardiidae].
Claibornites diegoensis (Dickerson, 1916). [Lucina
San Guillermo quad. (1943), Ventura Co. UCR 4752. Hypotype UCR 4752/41. [Lucinidae].
Claibornites diegoensis (Dickerson, 1916). [Lucina
Side canyon to Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242].
Hypotype UCLA 59281 [=LACMIP 10561]. [Lucinidae].
Claibornites diegoensis (Dickerson, 1916). [Lucina
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 45
Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=UCLA. 7210 =LACMIP 16114]. Hypo-
type LACMIP. 7518. [Lucinidae].

CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 10561 (not LACMIP 59281 as in pl. caption) [=UCLA 59281].
[Lucinidae].
Claibornites (Claibornites) diegoensis (Dickerson, 1916). [Lucina

2226. Holotype UCMP 11788. [Lucinidae].


Claibornites diegoensis (Dickerson, 1916). [Lucina
Eocene. Simi Valley, Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype UCLA 59281 [=LACMIP 7729]. [Lucinidae].
Claibornites diegoensis (Dickerson, 1916). [Lucina
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 987 [=LACMIP 16175]. Hypotype LACMIP 7729.
[Lucinidae].
Claibornites diegoensis (Dickerson, 1916). [Lucina-
mere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13445. [Lucinidae].
Claibornites (Codalucina) muirensis (Dickerson, 1914a). [Phacoides
6, 8. Martinez Fm., Paleocene. Near Muir Station, Concord quad., Contra Costa Co. UCMP 243. Holotype UCMP
11682. [Lucinidae].
Claibornites (Codalucina) turneri (Stanton, 1896). [Lucina]. Moore 1988b. USGS PP    
Martinez Fm., Paleocene. Lower Lake, Lake Co. Syntypes USNM 108971. [Lucinidae].
Clementia (Egesta) brioniana (Trask, 1922). [Venus
Shale Member), Miocene. Tar Peak, Reef Ridge quad., Kings Co. UCMP D1207. Hypotype UCMP 36721. [Veneridae].
Clementia (Egesta) conradiana (Anderson, 1905). [Venus
Miocene. Domengine Ranch quad., Fresno Co. UCMP B8584. Hypotype UCMP 36719. [Veneridae].
Clementia conradiana (Anderson, 1905). [Venus
Old Topanga Canyon Rd. area, Topanga Canyon, Santa Monica Mtns., Los Angeles Co. Line drawing, source speci-
men unknown. Remarks:
Clementia martini (Clark, 1915). [Venus   
Topanga Canyon Rd. area, Topanga Canyon, Santa Monica Mtns., Los Angeles Co. Line drawing, source specimen
unknown. Remarks:
Clementia (Egesta) pertenuis (Gabb, 1866). [Venus
Venus pertenuis Gabb 1866)). [Vaqueros Fm.],
Miocene. [Coalinga District, Fresno Co. USGS 4631. Hypotype USNM 165597]. [Veneridae].
Clementia pertenuis (Gabb, 1866). [Venus
caption), Miocene. Southeast of Pleasanton, La Costa Valley quad. (1953), Alameda Co. SU 2739. Hypotype SU
8459 [=CASG 69947]. Remarks: In the subgenus Egesta,  Woodring (1926 [1927]: p. 40);  Adegoke (1969:
p. 148);  Addicott (1973b: p. 36). [Veneridae].
Clementia (Egesta) pertenuis (Gabb, 1866). [Venus
(Wygal Sandstone Member), Miocene. Media Agua Creek area, Las Yeguas Ranch quad., Kern Co. USGS M3280.
Hypotype USNM 646597. [Veneridae].

9432. Hypotype USNM 646598.
46 PALEOBIOS, VOLUME 38, FEBRUARY 2021

646600.

Clementia (Egesta) pertenuis (Gabb, 1866). [Venus
Topanga Canyon Rd., Los Angeles Co. UCLA 2630 [=LACMIP 22630]. Hypotype UCLA 49514 [=LACMIP 10613].
[Veneridae].
Clementia (Egesta) pertenuis (Gabb, 1866). [Venus]. Albi 1991. Bull. SCPS 23(5-6):50-53,   
Miocene. Old Topanga Canyon Rd. area, Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 12589.
Line drawings, source specimens unknown. Remarks: Fig. 1 [
Woodring 1926 [1927]
Remarks: May have been

Clementia pertenuis (Gabb, 1866) [Venus
North side San Juan Creek, Orange Co. [Hypotype at CC]. [Veneridae].
Clementia pertenuis (Gabb) conradiana (Anderson, 1905). [Venus]. Bereskin and Edwards 1969. AAPG/SEPM Guide-

[=LACMIP 16422]. Hypotype UCSB M3 [=LACMIP 6304]. [Veneridae].
Clinocardium blandum (Gould, 1851). [Cardium-
cene. Fortuna quad., Humboldt Co. UCMP B7875. Hypotype UCMP 37771. Remarks: In the genus Keenocardium,
 Herrera et al. (2015: pp. 99–101). [Cardiidae].
ClinocardiumClinocardium dickersoni (Clark, 1918). [Cardium]. Moore 2003a. USGS PP 1228-F [electronic
Cardium dickerson
Southwest of Walnut Creek, Concord quad., Contra Costa Co. UCMP 1131. Holotype UCMP 11207. [Cardiidae].
Clinocardium lispum
quad. (1950), Humboldt Co. UCMP B7643. Holotype UCMP 14152, paratype UCMP 37774. [Cardiidae].
Clinocardium meekianum (Gabb, 1866). [Cardium 
Fm., Pliocene. Ft. Ross quad., Sonoma Co. UCMP B2155. Hypotype UCMP 12529.
Clinocardium cf.  (Gabb, 1866). [Cardium-

Clinocardium meekianum (Gabb, 1866). [Cardium
Pliocene. Santa Cruz Co. SCCM 1. Hypotype SCCM 21012. [Cardiidae].
Clinocardium meekianum (Gabb, 1866). [Cardium
Santa Cruz Co. CASG66084. Hypotype CASG 66084.04. [Cardiidae].
Clinocardium meekianum (Gabb, 1866). [Cardium
9–11. Wildcat Group, Pliocene. Eagle Prairie, Humboldt Co. Holotype ANSP 4497. [Cardiidae].
, 6. Etchegoin Fm., Pliocene. Kettleman Hills area, San Joaquin Valley, Kings Co. USGS 12400. Holotype
(of Clinocardium meekianum myrae Adegoke, 1969) USNM 495796.
Clinocardium meekianum myrae      
Pliocene. Garza Creek, Reef Ridge quad., Kings Co. UCMP D1145. Paratype UCMP 36688. Remarks: Junior syn-
onym of Clinocardium meekianum (Gabb, 1866),  Kafanov (1998: p. 37), valid subspecies,  Moore (2003a:
p. 21). [Cardiidae].

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 47
Clinocardium (Clinocardium) meekianum myrae Adegoke, 1969. Moore 2003a. USGS PP 1228-F [electronic format]

Kings Co. UCMP D1145]. Paratype UCMP 36688 (not 36668 as in pl. caption). Remarks: Junior synonym of
Clinocardium meekianum (Gabb, 1866),  Kafanov (1998: p. 37). [Cardiidae].
Clinocardium nuttalli (Conrad, 1837). [Cardium
(as Cardium (Clinocardium) nuttalli in pl. caption). San Diego Fm., Pliocene. Bluff with fossiliferous concretions
in clay quarry, Balboa Park, San Diego, San Diego Co. LACMIP 107. Hypotype LACMIP 4655. [Cardiidae].
Clinocardium praeblandum   
West end of Las Trampas Ridge, near Walnut Creek, Contra Costa Co. UCMP 1262. Holotype UCMP 14836, para-
type UCMP 14835. [Cardiidae].

Clinocardium (Clinocardium) praeblandum Keen, 1954. Moore 2003a. USGS PP 1228-F [electronic format] (family

Creek, Concord quad., Contra Costa Co. UCMP 1262. Holotype UCMP 14836. [Cardiidae].
Clinocardium pristinum
(probably Neroly Fm.), Miocene. Southwest part of Shell Ridge, near Walnut Creek, Concord quad., Contra Costa
Co. [UCMP 1943]. Holotype UCMP 14838, paratype UCMP 14837. [Cardiidae].
Clinocardium aff. 
quad., Humboldt Co. UCMP B7643. Hypotype UCMP 37774. Remarks: Clinocardium lispum n. sp.,  Roth and
Talmadge (1975: p. 3–5); in the subgenus Keenocardium,  Moore (2003a: p. 19). [Cardiidae].
Clinocardium

Remarks:Chione sp., Macoma sp.,
and Protothaca sp. [=Leukoma sp.]. [Cardiidae].
Codakia (Epilucina) californica (Conrad, 1837). [Lucina]. Moore 1988b. USGS PP 1228-D:D11, pl
Diego Fm., Pliocene. West of the mouth of Rose Canyon, south slope of Mt. Soledad, San Diego Co. SDSNH 80].
Hypotype (of Hertlein and Grant 1972Remarks: In the genus Epilucina,
Coan et al. (2000: p. 263),  Squires (2012: p. 76). [Lucinidae].
Codakia distinguenda (Tryon, 1872). [Lucina]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p. 601,
        
[Lucinidae].
Codakia distinguenda (Tryon, 1872). [Lucina     
[=Latrania Fm.], Pliocene. Vicinity of Travertine Point, Oasis quad., Imperial Co. (not San Diego Co. as in locality
description). UCMP A1707 [=UCMP A1705]. Hypotype UCMP 39876. [Lucinidae].
Codakia sp. cf.  (Linnaeus, 1758). [Venus
Fm. (Latrania Sand member) [=Latrania Fm.], Miocene. Coyote Mtns., Imperial Co. UCR 7267. [Hypotype UCR
7267/4]. [Lucinidae].
Compsomyax cf.  (Carpenter, 1864b). [Clementia]. Woodring et al. 1940 [1941]. USGS PP 195:33, pl.
Acila Zone), Pliocene. North side of Arroyo Hondo, North Dome, Kettleman Hills, Kings
Co. USGS 12340. Hypotype USNM 495715. [Veneridae].
Compsomyax subdiaphana (Carpenter, 1864b). [Clementia]. Hertlein and Grant, 1972. SDSNH Mem. 2(2B):269–270,

-
MIP 4688.
Compsomyax subdiaphana (Carpenter, 1864b). [Clementia 
48 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pliocene. Newhall area, Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14359. [Veneridae].
Compsomyax subdiaphana (Carpenter, 1864b) subsp. gibbosus (Gabb, 1869a). [Saxidomus]. Faustman 1964. UCPG

[Veneridae].
Compsomyax       
southwest of Trinidad Head, Humboldt Co. CASG 33223. Hypotype CASG 4284 (with Patinopecten caurinus).
[Veneridae].
Conchocele bisecta (Conrad, 1849a). [Venus-
onwheel Fm., Eocene. Wagonwheel Mtn., Emigrant Hill quad. (1953, PR 1973), Kern Co. CSUN 1580 [=LACMIP
16886]. Hypotypes LACMIP 12357–12358. Remarks: Renamed Conchocele bathyaulax Hickman, 2015, 
Hickman (2015: p. 10). [Thyasiridae].

LACMIP 12359. Remarks: Hryniewicz et al. (2017: p. 719) suggested that this species was is Maorithyas folgeri

Corbicula jestesi Squires, 1999c (n. sp
quad. (1952, PR 1967), Ventura Co. CSUN 1445 (=LACMIP 24259). Holotype LACMIP 12464, paratypes LACMIP
12465–12468. [Corbiculidae].
Corbicula cf.  Anderson and G D. Hanna, 1925. Squires and Deméré 1991. SEPM Guidebook 68:184,

Corbis washingtoniana  
Fm., Eocene. Coyote Canyon, Pt. Conception quad., Santa Barbara Co. UCMP B7027. Hypotypes SU 9286, 9287
[=CASG 70147, 70148]. Remarks: Epilucina washingtoniana (Clark, 1925),  Squires and Gring (1996: p. 66).
[Lucinidae].
Corbula (Varicorbula) capayana-
ker Peak quad. (1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7541. [Corbulidae].
Corbula complicata  
Reef Ridge, southeast of Big Tar Canyon, Reef Ridge quad., Kings Co. USGS 14495. Hypotype USNM 498708.
[Corbiculidae].

14503. Hypotype USNM 498709.
Corbula (Caryocorbula) dickersoni   
Eocene. Tributary to Alamo Creek above the mouth of Wagon Rd. Canyon, San Guillermo quad. (1943), Ventura
Co. UCR 4707. Hypotype UCR 4707/32. [Corbiculidae].
Corbula dickersoni
Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype
SDSNH 20780 (not SDSNH 20781 as in pl. caption). Remarks: In the subgenus Caryocorbula,  Squires (1987:
p. 71). [Corbiculidae].
Corbula (Caryocorbula) dickersoni
Fm., Eocene. Side canyon to Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115].
Hypotype LACMIP 6592. [Corbiculidae].
Corbula (Caryocorbula) dickersoni
Fm., Eocene. Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 807 [=LACMIP 16191]. Hypo-
type LACMIP 7539. [Corbiculidae].
Corbula (Caryocorbula) dickersoni

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 49
LACMIP 7539. [Corbiculidae].
Corbula aff. Corbula (Carycorbula) dickersoni Weaver and Palmer, 1922. Squires et al. 1988. SEPM Guidebook 58:185,

Kern Co. USGS M9067. Hypotype USNM 424897. [Corbiculidae].
Corbula cf. C() dickersoni Weaver and Palmer, 1922. Squires 1991c. 
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 676 [=LACMIP 16192]. Hypotype
LACMIP 11488. [Corbiculidae].
Corbula (Caryocorbula) dickersoni-
sana Fm., Eocene. South side of Simi Valley, Calabasas quad. (1952, PR 1967), Ventura Co. CSUN 1344 [=LACMIP
17192]. Hypotype LACMIP 12702. [Corbiculidae].
Corbula (Caryocorbula) dickersoni
Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13455.
[Corbiculidae].
Corbula (Caryocorbula) domenginica

Hypotypes UCR 4865/160-4865/161. [Corbiculidae].
Corbula (Varicorbula) gibbiformis Grant and Gale, 1931. Hertlein and Grant 1972. SDSNH Mem. 2(2B):323–324, pl.
Corbula gibbiformis). San Diego Fm., Pliocene. Near mouth of Rose Canyon, San Diego, La Jolla
quad., San Diego Co. UCLA 2359 [=LACMIP. 22359]. Hypotype UCLA [48636 (=LACMIP 9319)]. [Corbiculidae].
Corbula hornii
Canyon, west of San Marcos Pass, Santa Barbara Co. Hypotype SU 9258 (not SU 9323 as in text) [=CASG 70149].
[Corbiculidae].
Corbula (Caryocorbula) hornii
of Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4717. Hypotype UCR 4717/301. [Corbiculidae].
Corbula (Lentidium) luteola
5, 6, 15 (as Corbula luteola). San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San
Diego Co. LACMIP 305. Hypotypes LACMIP [4827, 4828]. Remarks: In the genus Caryocorbula,  Coan and
Valentich-Scott (2012: p. 870). [Corbiculidae].
Corbula parilis
Reef Ridge quad., Kings Co. USGS 14494. Hypotypes USNM 498710, 498711. [Corbiculidae].
Corbula (Caryocorbula) parilis
Meadows Campground, Lion Canyon quad. (1943), Ventura Co. UCR 4680. Hypotype UCR 4680/101. [Corbiculidae].
Corbula (Caryocorbula) parilis
Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP
7540. [Corbiculidae].
Corbula (Caryocorbula) parilis    
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 974 [=LACMIP 16193]. Hypotype LACMIP 7734.
[Corbiculidae].
Corbula (Caryocorbula) parilis Gabb, 1864. Squires 1999b.
North side of Simi Valley, bulldozer-generated exposure now under houses, Santa Susana quad. (1951, PR 1969),
Ventura Co. CSUN 958 [=LACMIP 17190]. Hypotype LACMIP. 12703. [Corbiculidae].
Corbula n. sp. aff.           

member of the Moreno Fm., Midway quad., Alameda Co. UCMP D8150. UCMP unnumbered hypotype lost,  C.
50 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Throckmorton (personal communication, 2002). [Corbiculidae].
Corbula rosecanyonensis
Eocene. Rose Canyon, San Diego Co. SDSNH 2363. Hypotype SDSNH 4166]. [Corbiculidae].
Corbula

Crassadoma cf.  (Gray, 1825). [Lima
Fm., Saddleback Valley Limestone member, Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capist-
rano quad. SDSNH 4312. Hypotype SDSNH 140354. [Pectinidae].
Crassatella branneri (Waring, 1914). [Crassatellites
Susana Fm., Paleocene. Simi Hills, Ventura Co. [UCR 6825]. Hypotype UCR 6825/1. Remarks: Junior synonym of
of Crassatella compacta Gabb, 1869b,  Moore (2003a: p. 1). [Crassatellidae].
Crassatella claytonensis (Dickerson, 1914a). [Crassatellites]. Moore 2003a. USGS 1228-F [electronic format] (family

1558. Syntype UCMP 11669 (not UCMP 11670 as in text). [Crassatellidae].
Crassatella collina
Sacate/Gaviota fms., Eocene. Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6954. Hypotypes SU 9271–


Lompoc quad., Santa Barbara Co. UCMP B7006. Hypotypes SU 9275, 9276.

Co. UCMP B6987. Hypotype SU 9274.
Crassatella collina-
cene. Upper Sespe Creek, Ventura Co. LACMIP 23727. Hypotype LACMIP 11663. [Crassatellidae].
Crassatella collina Conrad, 1857d. Shapiro 1998. AAPG Q. Pozo
Fm., Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8718 [=SBMNH 60180].
Remarks: Determined here as indet. bivalve internal mold. [Crassatellidae].
Crassatella compacta
1, 2. Lodo Fm., Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073
[=CASG 61666 (=UCMP A9717)]. Hypotype SU 10234 [=CASG 61666.05]. [Crassatellidae].
Crassatella compacta
17. Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. [UCR 4569]. Hypotype UCR 4569/38. [Crassatellidae].
Crassatella compacta Gabb, 1869b. Moore 2003a. USGS 1228-F [electronic format] (family Crassatellidae):1, pl. 2,

(of Crassatellites branneri Waring, 1914) CASG 160 [=CASG 61919.03]. [Crassatellidae].
Crassatella granti (Wiedey, 1928). [Spisula-
cene. Cajon Pass, Hesperia quad., San Bernardino Co. CIT 568 [=LACMIP 7673]. Hypotype CIT 3899 [=LACMIP
4116]. [Crassatellidae].
 
498962, 498963.
Crassatella granti (Wiedey, 1928). [Spisula]. Moore 2003a. USGS 1228-F [electronic format] (family Crassatelli-

SU 200. Holotype SDSNH 40. [Crassatellidae].
Crassatella lillisi (Dickerson, 1916). [Crassatellites]. Moore 2003a. USGS 1228-F [electronic format] (family
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 51

Urruttia Canyon and Salt Creek, Coalinga quad., Fresno Co. UCMP 1817. Holotype UCMP 11797. [Crassatellidae].
Crassatella lincolnensisCrassatellites]. Weaver and Kleinpell 1963. UCPG 43:199–200, pl. 33,

B6957. Hypotype SU 9278 [=CASG 70158]. [Crassatellidae].
Crassatella lomitensis (Oldroyd, 1924). [Crassatellites
Fm., Pliocene. East bluff of upper Newport Bay, Orange Co. UCR 4915. Hypotype UCR 4915/273. Remarks: Junior
synonym of  (Carpenter, 1864b),  Coan et al. (2000: p. 293). In the genus Hybolophus

Crassatella meganosensis (Clark and Woodford, 1927). [Crassatellites]. Moore 2003a. USGS PP 1228-F [electronic

Costa Co. UCMP 3159. Holotype UCMP 31283, paratype UCMP 31284. [Crassatellidae].
Crassatella (Crassatellites) mulates M.A. Hanna, 1927. Moore 2003a. USGS 1228-F [electronic format] (family Cras-
satellidae):6–7, pl. Jolla quad., San Diego Co. UCMP 5062.
Holotype UCMP 31006. [Crassatellidae].
Crassatella stewartvillensis (Dickerson, 1914a). [Crassatellites]. Moore 2003a. USGS PP 1228-F [electronic format]

Costa Co. UCMP 1540. Holotype UCMP 11668. [Crassatellidae].
Crassatella studleyi (Dickerson, 1914a). [Crassatellites]. Moore 2003a. USGS 1228-F [electronic format] (family

Mt. Diablo quad., Contra Costa Co. UCMP 1540. Holotype UCMP 32756. [Crassatellidae].
Crassatella unioides-
leocene. Simi Hills, Ventura Co. [UCR 6834]. Hypotype UCR 6834/2. [Crassatellidae].
Crassatella uvasana-
cene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH.
20729. [Crassatellidae].
Crassatella uvasana
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotypes LACMIP
6576–6579, 6581,6582. [Crassatellidae].

caption) [=LACMIP 16203]. Hypotype LACMIP 6580.
Crassatella uvasana
Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP
7528. [Crassatellidae].
Crassatella uvasana
Valley, Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 6577. [Crassatellidae].
Crassatella uvasana

Hypotype LACMIP 6577]. [Crassatellidae].
Crassatella uvasana uvasana Conrad, 1855b. Moore 2003a. USGS 1228-F [electronic format] (family Crassatelli-

Canyon Shale,” Eocene. Rose Canyon, San Diego Co. [CASG 66541]. Syntypes (of Astarte semidentata Cooper, 1894)
CASG 618, 618a–c [=CASG 66541.01–66541.04]. [Crassatellidae].

(of Crasatella grandis Gabb, 1864) UCMP 31449.
52 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Crassatella uvasana Conrad, 1855b. Squires 2008,
San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13447. [Crassatellidae].
Crassatella weaveri (Clark, 1938). [Crassatellites]. Moore 2003a. USGS PP 1228-F [electronic format] (family Crassatel-
Crassatella). Kreyenhagen Fm. (Markley Sandstone member), Eocene. Sandstone cliff on
northeast bank of Pleasants Creek, Napa quad., Solano Co. UCMP A1297. Holotype UCMP 30831. [Crassatellidae].
Crassatella   
Pass Rd. near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP loc. A4657
and A9717]. Hypotype UCMP 14201. [Crassatellidae].
Crassatellites brainerdi Dickerson, 1914b.” Moore 2003a. USGS 1228-F [electronic format] (family Crassatelli-

Co. UCMP 2233. Holotype UCMP 11757. [Crassatellidae].
Crassatellites uvasana (Conrad, 1855b). [Crassatella
Crassatella uvasana Conrad). Tejon Fm., Eocene. Ft. Tejon, Kern
Co. Neotype (of Stewart 1930: p. 142), ANSP 4461, missing,  K. Estes-Smargianni (personal communication,
2019). Remarks: In the genus Crassatella,  Moore (2003a: p. 4). [Crassatellidae].
Crassinella branneri (Arnold, 1903). [Astarte
25, 28, 29. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP
305. Hypotypes LACMIP 4535, 4536. Remarks: Junior synonym of  (C.B. Adams, 1852), 
Coan et al. (2000: p. 294). [Crassatellidae].
Crassinella cf. 
Fm. (Altamira Shale Member), Miocene. West side of George F. Canyon, Palos Verdes Hills, Los Angeles Co. USGS
13127. Hypotype USNM 497083. Remarks: Junior synonym of  (C.B. Adams, 1852),  Coan
et al. (2000: p. 294). [Crassatellidae].
Crassostrea ashleyi (Hertlein, 1934). [Ostrea
3. Temblor Fm., Oligocene/Miocene. Domengine Ranch quad., Fresno Co. UCMP D1075. Holotype UCMP 36646,
paratype UCMP 36647 (both of Ostrea arnoldi Adegoke, 1969). [Ostreidae].

-
type (of Ostrea hertleini Adegoke, 1969) UCMP 36654).

canyons, Caliente Range, McKittrick quad., San Luis Obispo Co. UCLA 512 [=LACMIP 20512]. Holotype (of Ostrea
altatemblorensis Grant and Eaton in Eaton et al., 1941) UCLA 8970 [=LACMIP 8953].
Crassostrea eldridgei ynezana (Loel and Corey, 1932). [Ostrea]. Addicot, 1973b. USGS PP 791: 7, 
Temblor Fm. (Wygal Sandstone Member), Miocene. Aqua Media Creek, Las Yeguas Ranch quad., Kern Co. USGS
M3280. Hypotypes USNM 646533, 646534. Remarks: In the genus/subgenus Pycnodonte (Pycnodonte) (family
Gryphaeidae),  Moore (1987: p. C18). [Gryphaeidae].
Crassostreaenglekyi (Hertlein, 1928a). [Ostrea        
Vaqueros Fm., Oligocene/Miocene. South of San Augustine Canyon, east of dike near top of hill, Santa Rosa Is.,
California Channel Ids. CASG 1156. Holotype CASG 4116. [Ostreidae].
Crassostrea eucorrugata (Hertlein, 1934). [Ostrea 
  
LACMIP 21751]. Holotype (of Ostrea cierboensis Grant and Eaton in Eaton et al., 1941) UCLA 8980 [=LACMIP
8968]. [Ostreidae].

Crassostrea tectonica
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 53
Eocene. Piru Creek, Los Angeles Co. LACMIP 40028. Holotype LACMIP 14687 (both valves), 14688. [Ostreidae].
-
MIP 14689.
-
MIP 40214]. Paratypes LACMIP 14690, 14697.

in pl. caption). Hypotype LACMIP 14691.

23732. Hypotype LACMIP 14693.

40246. Hypotype LACMIP 14694.

41097. Hypotype LACMIP 14695.
-
types LACMIP 14696, 14699.

40252. Hypotype LACMIP 14698.
Crassostrea titan (Conrad, 1853). [Ostrea
Margarita Fm.], Miocene. Taylor Canyon quad., Santa Barbara Co. UCLA L1766. Hypotype LSU I-10716]. ]. Re-
marks:
Crassostrea titan (Conrad, 1853). [Ostrea]. Addicott 1972b. SEPM Symposium vol., p. 15-
garita Fm., Miocene. Fellows quad., Temblor Range, Kern Co. USGS M3787. Hypotype USNM 646820. Remarks:

Crassostrea titan (Conrad, 1853). [Ostrea
Robles Fm., Pliocene. Near Atascadero, San Luis Obispo Co. USGS M4621. Hypotype USNM 647010. Remarks: :

from Miocene Santa Margarita Fm.,  Addicott and Galehouse (1973: p. 511). [Ostreidae].
Crassostrea titan (Conrad, 1853). [Ostrea       

USNM 164987. Remarks:
(1857c). [Ostreidae].



Ostrea panzana Conrad, 1857d) USNM 13338. Remarks: Numbered the same as Ostrea subjecta Conrad, 1857d
in USNM collection.

Valley, Los Angeles Co. Holotype (of Ostrea subjecta Conrad 1857d) USNM 13338. Remarks: Numbered the same
as Ostrea panzana Conrad, 1857d in USNM collection.
   Ostrea-Pecten  
quad., Fresno Co. UCMP D701. Holotype UCMP 36661, paratype UCMP 36662 (both of Ostrea titan andersoni
Adegoke, 1969).

54 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Gulch, Cuyama Valley, Santa Barbara Co. UCLA 1745 [=LACMIP 21745]. Holotype (of Ostrea titan prior Grant and
Eaton in Eaton et al., 1941) UCLA 8991 [=LACMIP 8977].

Santa Barbara Co. UCLA 1795 [=LACMIP 21795]. Holotype (of Ostrea ligminuta Grant and Eaton in Eaton et al.,
1941) UCLA 8985 [=LACMIP 8971].
Crassostrea titan (Conrad, 1853) [Ostrea
Miocene. Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP22632 [=UCLA 2623]. Hypotype LACMIP
13078 [=UCLA 57967]. Remarks:
Conrad (1857c). [Ostreidae].
Crassostrea titan (Conrad, 1853) [Ostrea     
Margarita Fm. Godwin Canyon, Pine Mtn., Ventura Co. LACMIP 17749. Hypotypes LACMIP 8272, 8282, 8283,
Remarks: Supplementary descriptions in Conrad (1854,


Crassostrea titan (Conrad, 1853) [Ostrea]. Stadum and Finger 2016, 37. Monterey
Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Cap-
istrano quad. SDSNH 4312. Hypotype SDSNH 83238. Supplementary descriptions in Conrad (1854, 1857c) and

Crassostrea vaquerosensis (Loel and Corey, 1932). [Ostrea
3, 7. Temblor Fm. (Aqua Sand Member) (of Stinemeyer et al. 1959: pp. 9, 13), Miocene. Las Yeguas Ranch quad.,
Temblor Range, Kern Co. USGS M4464. Hypotypes USNM 646787, 646791. Remarks: Supplementary descrip-
  Crassostrea
 Moore (1987: p. C38). [Ostreidae].
Crassostreavaquerosensis (Loel and Corey, 1932). [Ostrea

CASG 6789, 6790. [Ostreidae].
Crenella decussata (Montagu, 1808). [Mytilus
Fm., Pliocene. [Newport Beach, Orange Co. LACMIP 471]. Hypotype LACMIP 5747. [Mytilidae].

Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotype LACMIP
4479. Remarks: Junior synonym of Crenella decussata (Montagu, 1808),  Coan et al. (2000: p. 166). [Mytilidae].


Remarks: Junior synonym of Crenella decussata (Montagu, 1808),  Coan et al. (2000: p. 166). [Mytilidae].
Indeterminate CrenellaPaleoBios-
vation, Berkeley Hills, Oakland, Alameda Co. UCMP IP 13003. Hypotype UCMP 218709. [Mytilidae].
Crenomytilusarnoldi (Clark, 1918). [Mytilus
(Wygal Sandstone Member), Oligocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotype USNM 646511.
Remarks: In the genus/subgenus Mytilus (Mytilus Moore (1983: p. A6). [Mytilidae].

646512.

646540.
Crenomytilus cf.  (Clark, 1918). [Mytilus

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 55
Co. SU 1756. Hypotype CASG 60373, determined here as missing. Remarks: In the genus/subgenus Mytilus
(Mytilus Moore (1983: p. A62). [Mytilidae].
Crenomytilus cf.  (Arnold, 1909 [1910]). [Mytilus]. Powell et al. 2019a. USGS SIR 2019-5021:3, 29,

[Mytilidae].
Crenomytilus expansus (Arnold, 1907). [Mytilus     
(Wygal Sandstone Member), Miocene. Basal beds of Vaqueros Fm. [=Temblor Fm.] near Carneros Spring, Kern
Co. USGS 9427. Hypotype USNM 646549. Remarks: In the genus/subgenus Mytilus (Crenomytilus),  Moore
(1983: p. A63). [Mytilidae].
Crenomytilus expansus (Arnold, 1907). [Mytilus
Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Cap-
istrano quad. SDSNH 4312. Hypotype SDSNH 83246. Remarks: In the genus/subgenus Mytilus (Crenomytilus),
 Moore (1983: p. A63). [Mytilidae].
Crepella* columbiana 
Member), Pliocene. Tunitas, Woodside quad., San Mateo Co. Locality 653M. Hypotype determined here as miss-
ing. Remarks: In the genus Solamen,  Coan et al. (2000: p. 171). Error for Crenella. [Mytilidae].
Cryptomya cf.  (Conrad, 1837). [Sphaenia]. Woodring and Bramlette 1950 [1951]. USGS PP 222:91, pl.

14872. Hypotype USNM 560017. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphaenia
Fm. (San Gregorio Member), Pliocene. Near mouth of Tunitas Creek, San Gregorio quad., San Mateo Co. Locality
482-3M. Hypotype determined here as missing. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphaenia

[Myidae].
Cryptomya californica (Conrad, 1837). [Sphenia-

Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotypes CASG, determined here as missing. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphaenia]. Hertlein and Grant 1972. SDSNH Mem. (2B):319–320, pl. 55,

LACMIP 305. Hypotypes LACMIP 4823–4825. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphenia-

Fm., Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. CASG hypotypes determined here as missing. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphenia
Fm., Pliocene. Santa Cruz Co. SCCM hypotype determined here as missing. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphenia]. Yancey, 1978. Pr.CAS 41(15
Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D3688. Hypotype USNM 251804. [Myidae].
Cryptomya californica (Conrad, 1837). [Sphenia -
cene. Capitola, New Brighton section, wave-cut platform (recent), Santa Cruz Co. CASG66084. Hypotype CASG
66084.09. [Myidae].
Cryptomya californica magna
          
CASG 70443, 70448. Remarks: Junior synonym of Cryptomya californica (Conrad, 1837),  Coan et al. (2000:
p. 474). [Myidae].
56 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Cryptomya cf.  
Joaquin Fm. (Acila Zone), Pliocene. West fork of Arroyo Conchoso east of El Dombo, North Dome, Kettleman Hills,
Kings Co. USGS 12351. Hypotype USNM 495852. Remarks: Junior synonym of Cryptomya californica (Conrad,
1837),  Coan et al. (2000: p. 474). [Myidae].
Cubitostrea californiana
Canyon, Los Angeles Co. LACMIP 40485. Holotype LACMIP 14700, hypotype LACMIP 14701 [Ostreidae].

LACMIP 14702, 14707, paratypes LACMIP 14703, 14705, 14706.
Cucullaea mathewsonii
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype
SU/CASG 10232. [Cucullaeidae].
Cucullaea mathewsonii
Martinez, Contra Costa Co. Lectotype (of Stewart 1930: p. 78) ANSP 4559, paralectotype ANSP 4559 (also num-
bered ANSP 6125,  Moore (1983: pl. 10 caption). [Cucullaeidae].
Cucullaea mathewsonii  
Cretaceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA
6525 [=LACMIP 26525]. Hypotype LACMIP 7555. [Cucullaeidae].
Cucullaea morani
Fm.], Paleocene. Llajas Canyon, east of McCray wells, Camulos quad., Simi Valley Ventura Co. Holotype SU/CASG
166, three syntypes in CASG collection do not match the specimens illustrated in Moore (1983) [CASG 61916.01,
9363, 9820]. [Cucullaeidae].
Cucullaria morani (Waring, 1914). [Cucullaea
Fm., Paleocene. Simi Hills, Ventura Co. [UCR 6683]. Hypotype UCR. 6838/2 (not UCR 6683/1 as in pl. caption).
[Cucullaeidae].
Cuneocorbula torreyensis (M.A. Hanna, 1927). [Corbula]. Squires 1999cSand-
stone, Eocene. Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1445 (=LACMIP 24259). Hypotype LACMIP
12478. [Corbulidae].
Cuspidaria (Cardiomya) pectinata (Carpenter, 1864b). [Neaera]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):342–

4883. Remarks: In the genus Cardiomya,  Coan et al. (2000: p. 556). [Cuspidariidae].
Cyathodonta pedroana
Angeles Co. LACMIP 7757. Hypotype LACMIP 14374. [Thraciidae].
Cyclinella elevata (Gabb, 1864). [Dosinia
Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4726. Hypotype UCR 4726/21. [Veneridae].
Cyclocardia californica (Dall, 1903b). [Venericardia (Cyclocardia)]. Woodring and Bramlette 1950 [1951]. USGS PP

Co. USGS 14878. Hypotype USNM 560036. [Carditidae].

Hypotypes (2) USNM 560054.

Co. USGS 14608. Hypotype USNM 560103.

USGS 14611. Hypotype USNM 560139.
Cyclocardia californica (Dall, 1903b). [Venericardia (Cyclocardia)]. Durham and Addicott 1965. USGS PP 524-A:A13,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 57
     
USNM 649113. [Carditidae].

649114.
Cyclocardia (Cyclocardia) kirkerensis (Clark, 1918). [Cardium
Kirker Tuff, Oligocene. Kirker Creek area, Mt. Diablo quad., Contra Costa Co. UCMP 2033. Holotype UCMP 11165.
[Carditidae].
Cyclocardia (Cyclocardia) montereyana (Arnold, 1908). [Venericardia]. Moore 1992. USGS PP 1228-E:E13, pl. 7,

165464. [Carditidae].
Cyclocardia occidentalis (Conrad, 1855a). [Cardita]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):230–231, pl. 43,
-
MIP 22359]. Hypotypes UCLA [=LACMIP 9299–9301]. Remarks: Supplemental description in Conrad (1855b).
Junior synonym of Cyclocardia monilicosta (Gabb, 1861),  Coan et al. (2000: p. 303). [Carditidae].
Cyclocardia occidentalis (Conrad, 1855a). [Cardita]. Squires and White 1983. 
Saugus Fm. [=Pico Fm.], Pliocene. Northern Simi Valley, Santa Susana quad. (1951, PR 1969),Ventura Co. CSUN
725 [ LACMIP 29227]. Hypotype UCLA 59409 [=LACMIP 8322]. Remarks: Supplemental description in Conrad,
1855b. Junior synonym of Cyclocardia monilicosta (Gabb, 1861),  Coan et al. (2000: p. 303). [Carditidae].
Cyclocardia (Cyclocardia) occidentalis (Conrad, 1855a). [Cardita
San Diego Fm., Pliocene. San Diego, San Diego Co. LACMIP 16202. Hypotype (of Hertlein and Grant, 1972 pl. 43,
Remarks: Supplemental description in Conrad, 1855b. Junior synonym
of Cyclocardia monilicosta (Gabb, 1861),  Coan et al. (2000: p. 303). [Carditidae].


Cyclocardia occidentalis (Conrad, 1855a). [Cardita

CSUN 725 (=LACMIP 29227). Hypotype UCLA 59409 (LACMIP 8322)]. Remarks: Supplemental description in
Conrad, 1855b. Junior synonym of Cyclocardia monilicosta (Gabb, 1861),  Coan et al. (2000: p. 303). [Carditidae].
Cyclocardia occidentalis (Conrad, 1855a). [Cardita
Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14355. Remarks: Supplemental description in
Conrad, 1855b. Junior synonym of Cyclocardia monilicosta (Gabb, 1861),  Coan et al. (2000: p. 303). [Carditidae].
Cyclocardia ventricosa (Gould, 1850a). [Cardita
8. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305.
Hypotype LACMIP 4537. [Carditidae].
Cyclocardia (Cyclocardia) ventricosa (Gould, 1850a). [Cardita
San Diego Fm., Pliocene. San Ysidro quad. (1943], San Diego Co.[LACMIP 305]. Hypotype (of Hertlein and Grant

Carditid cf. Cyclocardia ventricosa      
northeast of San Juan Capistrano, Orange Co. Monterey Fm., Miocene. [Hypotype at CC]. [Carditidae].
Cyclocardia sp. Powell et al. 2019b. PaleoBios
Berkeley Hills, Oakland Hills, Alameda Co. UCMP IP13001. Hypotype UCMP 410563. [Carditidae].
Cyclopecten lompicoensis (Arnold, 1906). [Pecten
Miocene. Head of Lompico Creek, Santa Cruz quad., Santa Cruz Co. Holotype USNM 164931. [Propeamussiidae].
Cyclopectenmartinezensis (Gabb, 1869b). [Pecten]. Moore 1984. 9, 10. Martinez
Fm., Paleocene. Martinez, Contra Costa Co. [MCZ 1043]. Lectotype (of Stewart 1930: p. 117) MCZ 15026a [=MCZ
58 PALEOBIOS, VOLUME 38, FEBRUARY 2021
108502], paralectotype MCZ 15026 [=MCZ 108503]. [Propeamussiidae].
Cyclopecten (Cyclopecten) pernomus (Hertlein, 1935). [Pecten]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):213–214,

4507, 4508. [Propeamussiidae].
Cyclopecten sp. cf.  (Hertlein, 1935). [Pecten
Fm. (Latrania Sand Member) [=Latrania Fm.], Miocene/ Pliocene. Willis Palm area, Riverside Co. UCMP A1407.
Hypotype UCMP 37533. [Propeamussiidae].
Cyrtopleura (Scobinopholas) costata (Linneaus, 1758). [Pholas-
perial Fm. [=Latrania Fm.], Pliocene. Carrizo Creek (locality B), Imperial Co. SDSNH 51. Hypotype SDSNH 4277.
[Propeamussidae].
Delectopecten alternilineatus (Clark, 1918). [Pecten (Pseudomuseum)]. Moore 1984. USGS PP 1228-B:B15–B16, pl.

[Pectinidae].
Delectopecten harfordus (Davis, 1913). [Pecten        
Fm., Miocene. Ten km north of Port Harford (=Port San Luis), San Luis Obispo Co. Syntypes SU/CASG 406, 408
[=CASG 61963.01–61963.03], paratype CASG 9819.
Delectopecten lillisi (Hertlein, 1934). [Pecten
Eocene/Oligocene. Stanislaus Co. CASG 1874 (not CASG 184 as in text). Holotype CASG 6062, paratype CASG
6063. [Pectinidae].
Delectopecten peckhami-
terey Fm., Miocene. Southeast of Pinole (not Pinnole as in pl. caption), Contra Costa Co. USGS 2593. Hypotype

Delectopecten peckhami (Gabb, 1869a). [Pecten  
Tumey Fm., Eocene/Oligocene. Monocline Ridge, Fresno Co. [UCMP D6534]. Hypotype UCMP 37154. [Pectinidae].
Delectopecten peckhami (Gabb, 1869a). [Pecten-
gocene. Monocline Ridge, Monocline Ridge quad., Fresno Co. [UCMP D6534]. Hypotype UCMP 37154. [Pectinidae].
Delectopecten peckhami


Delactopecten peckhami 
San Juan Capistrano, Orange Co. Monterey Fm., Miocene. [Hypotype at CC]. [Pectinidae].
Delectopecten pedroanus (Trask, 1856). [Plagiostoma]. Valpey 1975. Earth 
Puente Fm., Miocene. Near Elsereno [sic], Los Angeles Co. Hypotype current whereabouts unknown. Remarks:
Possibly equal to Delectopecten peckhami (Gabb) of Arnold 1906,  Moore (1984: p. B15). [Pectinidae].
Delectopecten vancouverensis fernandoensis (Hertlein, 1925a). [Pecten]. Moore 1984. USGS PP 1228-B:B16–B17,

Orange Ave., 230 m north of Willow St., Long Beach, Los Angeles Co. (Ventura Co. in text). SU 155 [=CASG 61840].
Holotype SU/CASG 16 [=CASG 61840.01]. Remarks: Junior synonym of Delectopecten vancouverensis (Whiteaves,
1893),  Coan et al. (2000: p. 227). [Pectinidae].
Delectopecten vancouverensis fernandoensis (Hertlein, 1925a). [Pecten]. Anonymous 2001. Calif. Geol. 55(4):26, left
-
dall Park subway tunnel], Los Angeles, Los Angeles Co. [LACMIP 16847. Hypotype LACMIP 13015.]. Remarks:
Junior synonym of Delectopecten vancouverensis (Whiteaves, 1893),  Coan et al. (2000: p. 227). [Pectinidae].
Delectopecten

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 59
Delectopecten
Creek, Ventura Co. LACMIP 23690. Hypotype LACMIP 11634. [Pectinidae].
Dendostrea angelica (Rocheburne, 1895). [Ostrea
(Latraina Sand Member) [=Latrania Fm.], Miocene / Pliocene. Super Creek area, Riverside Co. UCR 8595. Hypotype
LACMIP 7164 [=LACMIP 7869]. Remarks: Provisionally in the genus Dendostrea Moore (1987: p. C26).
[Ostreidae].
Dendostrea angermanni (Hertlein and Jordan, 1927). [Ostrea
6. Imperial Fm. [=Latrania Fm.], Miocene/Pliocene. Imperial Co. Hypotype UCMP 37535 determined here as
missing. [Ostreidae].
Dendostrea vespertina (Conrad, 1854). [Ostrea      
(Latrania Sand Member) [=Latrania Fm.], Miocene/Pliocene. Willis Palm area, Riverside Co. CIT 187 [=LACMIP
16204]. Hypotype LACMIP 7163 [=LACMIP 7868] determined here as missing. Remarks: Supplemental descrip-

Dendostrea vespertina (Conrad, 1854). [Ostrea

Co. Lectotype (of Woodring 1938: p. 43) ANSP 13366, paralectotypes ANSP 13366a, 13366d, 13366e. Remarks:




(of Ostrea vespertina sequens Arnold, 1910) USNM 165545. Remarks: Myrakeena veatchii (Gabb, 1866), 
Squires et al. (2006: p. 10).
Dendostrea? vespertina (Conrad, 1854). [Ostrea]. Peska 1992p. 34. Imperial Fm.
[=Latrania Fm.], Miocene/Pliocene. Coyote Mtns., Imperial Co. Remarks: Supplemental descriptions in Conrad
Line drawing, source specimen unknown. [Ostreidae].
Dendostrea vespertina (Conrad, 1854). [Ostrea
Ostera vespertina Conrad, 1854)). Saugus Fm. [=Pico Fm.], Pliocene.
Browns Canyon, Los Angeles Co. [CSUN 241D (not CSUN 757, =LACMIP 16171)]. Hypotype UCLA 59406 (=LACMIP
8319)]. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 10). [Ostreidae].
Dermatomya tenuiconcha (Dall, 1913). [Poromya
6, 7, 10, 11, 13. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co.
LACMIP 305. Hypotype LACMIP 4882. [Poromyidae].
cf. Dinocardium
Miocene. [ABDSP 5151.1. Hypotype ABDSP I 10691]. [Cardiidae].
Diodus tenuis (Gabb, 1864). [Cyprinella-

on west side of Tesla Rd., Midway quad., Alameda Co. UCMP D8147. UCMP unnumbered hypotypes lost,  C.
Throckmorton (personal communication, 2002). [Corbiculidae].
Diodus tenuis (Gabb, 1864). [Cyprinella

the top of the shale member of the Moreno Fm., Midway quad., Alameda Co. UCMP D8150. UCMP unnumbered
hypotype lost,  C. Throckmorton (personal communication, 2002). [Corbiculidae].
Diplodonta (Diplodonta) buwaldana   
9, 10. Round Mt. Silt, Miocene. Near Barker's ranch house, Kern Co. Holotype CASG 111, paratype CASG 112.
[Ungulinidae].
Diplodonta (Felaniella) cornea (Reeve, 1850). [Lucina]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):253–254, pl.
60 PALEOBIOS, VOLUME 38, FEBRUARY 2021

LACMIP 305. Hypotypes LACMIP 4650, 4651. Remarks: In the genus Felaniella,  Moore (1988b: p. D29);
junior synonym of Diplodonta sericata (Reeve, 1850),Coan et al. (2000: p. 271). [Ungulinidae].
Diplodonta (Diplodonta) cretaceaLucina

Diablo, Contra Costa Co. Holotype missing,Stewart (1930: p. 7). [Ungulinidae].
-

Diplodonta (Felaniella) harfordi Anderson, 1905. Susuki 1992Cold Creek
Member), Miocene. Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22635 [=UCLA 2635]. Hypotype
LACMIP 13074 [=UCLA 57993]. Remarks: In the genus Felaniella,Moore (1988b: p. D29). [Ungulinidae].
Diplodonta (Diplodonta) orbella (Gould, 1851). [Lucina]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):252–253, pl.

Diego Co. LACMIP 305. Hypotypes LACMIP 4648, 4649]. [Ungulinidae].
Diplodonta cf. orbella (Gould, 1851). [Lucina
possibly at SDSNH (current whereabouts unknown). Remarks: Ex Grouard collection. [Ungulinidae].
Diplodonta (Diplodonta) orbella (Gould, 1851). [Lucina
San Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305]. Hypotypes (of Hertlein and Grant



Diplodonta (Zemysina)
Paleocene. Large concretions on west side of Meier Canyon, Calabasas Quad., Simi Hills, Ventura Co. UCR 6682.
Holotype UCR 6682/100 (not UCR 6682/2 as in text). [Ungulinidae].
Diplodonta (Zemysina)    

6682/100. [Ungulinidae].
Diplodonta (Diplodonta) polita (Gabb, 1864). [?Mysia
Fm., Paleocene. Near Martinez and at Clayton, near the coal mines (from Division B), Contra Costa Co. Holotype
UCMP 11990. [Ungulinidae].
Diplodonta (Diplodonta) stephensoni

1131. Holotype UCMP 11171. [Ungulinidae].
Diplodonta (Diplodonta) unisulcatus (Vokes, 1939). [Taras
17. Domengine Fm., Eocene. South part of crest of Parson's Peak, Coalinga quad. [=Joaquin Rocks quad.], Fresno
Co. UCMP 672. Holotype UCMP 15636. [Ungulinidae].
Diplodonta
Diplodonta parilis Conrad, 1848)). Etchegoin
Fm., Miocene. Southeast of Coalinga, Fresno Co. [Hypotype USNM 165484]. [Ungulinidae].
Diplodonta (ZemysinaSusana Fm., Paleocene.
Simi Hills, Ventura Co. [UCR 6682]. Hypotype UCR 6682/100 (not UCR 6682/2 as in pl. caption). Remarks: Dip-
lodonta (Zemysina) n. sp.,  Zinsmeister (1983b: p. 1287). [Ungulinidae].
Divalinga (Divalinga) eburnea (Reeve, 1850). [Lucina
Imperial Fm. [=Latrania Fm.], Miocene/Pliocene. From two small washes in Alverson Canyon, at base of Coyote

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 61
Divalinga cf.  (Reeve, 1850). [LucinaDuvalinga sp. cf.
). Imperial Fm. [=Latrania Fm.], Pliocene. Vicinity of Desert Shores and Travertine Rock, Imperial Co.
(not San Diego Co. as in locality description). USGS M9778. Hypotype USNM 486912. [Lucinidae].
Divaricella cumulata (Gabb, 1864). [Lucina
Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype
SDSNH 20778. [Lucinidae].
Divaricella cumulata (Gabb, 1864). [Lucina
canyon east of Chivo Canyon, north side of Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP
7430. Hypotype LACMIP 12751. [Lucinidae].
DivaricellaEgracina cumulata (Gabb, 1864). [Lucina 

258. [Lucinidae].

Divaricella cf.  (Reeve, 1850). [Lucina
peels). Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, west side of
George F. Canyon, Palos Verdes Hills, Los Angeles Co. USGS 13127. Hypotypes USNM 497084–497086. Remarks:
In the genus Divalinga,Moore (1988b: p. D22). [Lucinidae].
Donax (Serrula) gouldii
sic] River plain, San Diego Co. UCLA 294
[=LACMIP 20294]. Hypotype UCLA 48631 [=LACMIP 9314]. [Donacidae].

4808, 4809.
Donax latus          

Idria, San Benito Co. Holotype MCZ 15028 [=MCZ 108506]. [Donacidae].
Donax scrippsensis M.A. Hanna, 1927. Moore 2003b. USGS PP 1228-G [electronic format] (family Donacidae):1–2,
   
5089. Holotype UCMP 30992. [Donacidae].
Donax triangulata Anderson and Martin, 1914. Moore 2003b. USGS PP 1228-G [electronic format] (family Donaci-
dae): of Barker’s ranch house, Kern
Co. CASG 65. Holotype CASG 130 [=CASG 65.02]. [Donacidae].
Donax
Co. SU 2121. Hypotype SU 7524 [=CASG 71998]. [Donacidae].
Dosinia longidens
Canyon, Hayward quad., Alameda Co. UCMP A4565. Hypotypes UCMP 34148–34150. [Veneridae].
Dosinia (Dosinidia) margaritana
Mtn. Silt, Miocene. Caliente quad., Kern Co. SU 2121. Hypotypes SU. 7525, 7525b [=CASG 71999]. [Veneridae].
Dosinia margaritana
Pozo Fm., Paleocene or Cañada Fm., Eocene. Santa Cruz Is., California Channel Ids. Line drawing, source specimen
unknown. Remarks: Determined here as Venericardia sp. (family Cardiidae). [Veneridae].
Dosinia margaritana Wiedey var. projecta Loel and Corey, 1932. Bereskin and Edwards 1969. AAPG/SEPM Guide-

Ids. UCSB FA-JK65-102 [=LACMIP 16405]. Hypotype UCSB M5 [=LACMIP 6302]. [Veneridae].
Dosinia margaritana projecta
62 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Fm. (Carneros Sand Member) (of Stinemeyer et al. 1959: pp. 9, 13), Miocene. Packwood Creek quad., Temblor
Range, Miocene. USGS M2633. Hypotype USNM 646786. [Veneridae].
Dosinia margaritana projecta  
Monterey Fm. (Saltos Shale Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS
M5025. Hypotype USNM 254369. [Veneridae].
Dosinia margaritana projecta

area, La Panza Range, San Luis Obispo Co. USGS M5025. Hypotype USNM 254369. [Veneridae].
Dosinia mathewsonii
-
potype USNM 165596]. [Veneridae].
Dosinia (Dosinidia) mathewsonii
Oligocene. North side of Mt. Diablo, Contra Costa Co.] UCMP A4660. Hypotype UCMP 11149A. Remarks: Junior
synonym of Dosinia whitenyi (Gabb, 1866),  Addicott (1973b: p. 36). [Veneridae].
Dosinia merriami Clark, 1915. Addicott 1972b. SEPM Symposium vol., p. 12, pl. 
Sand Member) (of Stinemeyer et al. 1959: pp. 8, 13), Miocene. Shale Pt. quad., Temblor Range, Kern Co. USGS
M3571. Hypotype USNM 646810. [Veneridae].
Dosinia merriami
Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M5025. Hypotype USNM 254364. [Veneridae].
Dosinia merriami

Obispo Co. USGS M5025. Hypotype USNM 254364. [Veneridae].
Dosinia ponderosa (Gray, 1838), var. [Artemis
6. Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. Alex Clark locality. Hypotype USNM
560105. [Veneridae].
  
14614. Hypotypes USNM 560145, 560146.
Dosinia aff.  (Gray, 1838). [Artemis
Fm. (Altamira Shale Member), Miocene. George F. Canyon–Miraleste Canyon area, west side of George F. Canyon,
Palos Verdes Hills, Los Angeles Co. USGS 13127. Hypotypes USNM 497089, 497090. [Veneridae].
Dosinia cf.  (Gray, 1838). [Artemis
Rico Fm., Miocene/Pliocene. Adelaida quad., Monterey Co. USGS M2047. Hypotype USNM 649086. [Veneridae].
Dosinia ponderosa (Gray, 1838). [Artemis
Pliocene. Near the U.S./Mexico boundary, San Diego Co. [SDSNH 430. Hypotype SDSNH 4151]. [Veneridae].
Dosinia ponderosa (Gray, 1838). [Artemis
3.16E. Latrania Fm., Miocene/Pliocene. [Painted Gorge, Coyote Mtn., Imperial Co. SDSNH 44]. Hypotype SDSNH
53797. [Veneridae].
Dosinia ponderosa (Gray, 1838). [Artemis
area, Los Angeles Co. LACMIP 7725. Hypotype LACMIP 14362. [Veneridae].
Dosinia (Dosinia) ponderosa diegoana

and paratype CASG 70446. Remarks: Possible junior synonym of Dosinia ponderosa (Gray, 1838). [Veneridae].


GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 63
....., 

Dosinia santana-
loged. Remarks: Ex Grouard collection. [Veneridae].
Dosinia cf. -
trania Fm.], Pliocene. Near Travertine Point, Oasis quad., Imperial Co. (not San Diego Co. as in description). UCMP
A1707 [=UCMP A1705]. Hypotype UCMP 39877. [Veneridae].
Dosinia sobrantensis
Hayward quad., Alameda Co. UCMP A4565. Holotype UCMP 34151, paratype UCMP 34152. [Veneridae].
Dosinia (Dosinidia) whitneyi (Gabb, 1866). [Chione
Miocene. Near Martinez, Contra Costa Co. Holotype UCMP 11999 [ex CGS. 272]. [Veneridae].
-
type UCMP 34153.
Dosinia (Dosinia) whitneyi (Gabb, 1866). [ChioneDosinia
mathewsoni Gabb” of Clark, 1918). San Ramon Fm., Oligocene. Near town of Walnut Creek, Contra Costa Co. UCMP
1131. Hypotype UCMP 33734. [Veneridae].
-
lotype UCMP 11999. [Veneridae].
Dosinia whitneyi (Gabb, 1866). [Chione       
(Phacoides Sand member) (of Stinemeyer et al. 1959: pp. 12, 13), Miocene. Las Yeguas Ranch quad., Temblor
Range, Kern Co. USGS M3280. Hypotype USNM 646579. [Veneridae].
Dosinia (Dosinia) whitneyi (Gabb, 1866). [Chione
Sandstone (=Temblor Fm., Wygal Sandstone Member), Miocene. Kern Co. USGS 9432. Hypotype USNM 646576.
[Veneridae].

Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotypes USNM 646577–646579, 646593.

M3772. Hypotype USNM 646594.
Dosinia (Dosinidia
of Mt. Diablo, Contra Costa Co. UCMP A4660. Hypotypes UCMP 30012, 30013. [Veneridae].
DosiniaWeaver and Kleinpell, 1963. UCPG 43:205
above Sacate-Gaviota contact in narrow gorge of eastern branch of Cañada del Corral, Los Olivos quad., Santa
Barbara Co. UCMP B6982. Hypotype SU 9313 [=CASG 70161]. [Veneridae].
Dosinia
Co. CIT 2093 [=LACMIP 11853]. Hypotypes LACMIP 4136, 4137. [Veneridae].
Dosinidia margaritana
Fm., Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8714 [=SBMNH 60184].
Remarks: Determined here as indet. pitarid sp. [Veneridae].
Dosiniopsis stewartvillensis Clark and Woodford, 1927. Shimer and Shrock 1944. Index Foss. N.A., p. 427, pl. 170,

Diablo, Contra Costa Co. UCMP 3159. Holotype UCMP 31287]. [Veneridae].
Ensis myrae
64 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 4811 (within Spisula (Mactromeris)
hemphilli (Dall, 1894), hypotype 4820). [Pharidae].
Ensis   

Eomiltha turneri (Stanton, 1896). [Lucina
Paleocene. Panoche Pass Rd. near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073
[=UCMP A9717]. Hypotype SU 10230 [=CASG 61666.15]. Remarks: In the genus/subgenus Claibornites (Coda-
lucina),Moore (1988b: p. D20). [Lucinidae].
Epilucina californica (Conrad, 1837). [Lucina
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14350. [Lucinidae].
Epilucina washingtoniana (Clark, 1925). [Corbis
Wagonwheel Fm., Eocene. Wagonwheel Mtn., Emigrant Hill quad. (1953, PR 1973), Kern Co. CSUN 1580 [=LACMIP
16886]. Hypotypes LACMIP 12354–12356. [Lucinidae].
Epilucina
Imperial Co. Hypotype probably at SDSU (current whereabouts unknown). [Lucinidae].
Eucrassatella cf.  (Reeve, 1842). [Crassatella
Fm. [=Latrania Fm.], Pliocene. Near Travertine Point, Oasis quad., Imperial Co. (not San Diego co. as in descrip-
tion). UCMP A1707 [=UCMP A1705]. Hypotype UCMP 39874, missing,  Powell 2008: p. 6. [Crassatellidae].
Eucrassatella subgibbosa (G D. Hanna, 1926). [Crassatellites]. Stanton 1966. 9–12.
Castaic Fm., Miocene. Humphreys quad. (1932, PR 1946), Los Angeles Co. CIT 1663 [=LACMIP 2555]. Hypotypes
LACMIP 4118–4121. Remarks: Eucrassatella subgibbosus,  Moore (2003a: p. 13). [Crassatellidae].
Eucrassatella subgibbosa (G D. Hanna, 1926). [Crasaitellites]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego
          
SDSNH 23122. Remarks: Eucrassatella subgibbosus,  Moore (2003a: p. 13). [Crassatellidae].
Eucrassatella subgibbosus (G D. Hanna, 1926). [Crassatellites]. Moore 2003a. USPS PP 1228-F [electronic format]

area, Coyote Mtn., Imperial Co. CASG 682. Holotype CASG 1818 [=CASG 682.02]. [Crassatellidae].
Eucrassatella subgibbosus (G D. Hanna, 1926). [Crassatellites
Imperial Fm. [=Latrania Fm.], Pliocene. Alverson Canyon area, Coyote Mtn., Imperial Co.] CASG 682. Holotype
CASG 682.2. [Crassatellidae].
Euvola keepi (Arnold, 1906). [Pecten
Fm.], Miocene/Pliocene. Carrizo Creek district, San Diego Co. Holotype SU/CASG 5 [=CASG 61887.01]. [Pectinidae].
Euvola keepi (Arnold, 1906). [Pecten
Latrania Fm., Miocene/Pliocene. Alverson Canyon, Imperial Co. SDSNH 347]. Hypotype SDSNH 12030. [Pectinidae].
Euvola keepi (Arnold, 1906). [Pecten
Member) [=Latrania Fm.], Miocene. Coyote Mtns., Imperial Co. Hypotype (holotype as per CASG online database)
SU/CASG 5 [=CASG 61887.01]. [Pectinidae].
Euvola stearnsii (Dall, 1878a). [not 1874 as in pl. caption]. [Pecten 
2.4. San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexican boundary, San Diego Co. LACMIP 305.
Hypotype LACMIP 14851. [Pectinidae].
Felaniella (Felaniella) cornea (Reeve, 1850). [Lucina
Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305]. Hypotype (of Hertlein and Grant 1972,
Remarks: Junior synonym of Diplodonta sericata (Reeve, 1850),Coan
et al. (2000: p. 271). [Ungulinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 65
Felaniella (Felaniella) harfordi (Anderson, 1905). [Diplodonta
Temblor Fm. (Wygal Sandstone Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS
M3280. Hypotypes USNM 646550, 646551. [Ungulinidae].
Felaniella harfordi (Anderson, 1905). [Diplodonta
Oligocene/Miocene. [Grimes Canyon, Oak Ridge, Ventura Co. UCMP A355]. Hypotype (of Loel and Corey 1932,


Syntypes CASG 62, 63 [=CASG 66748.01, 66748.02].
Felicia phrear (Woodring, 1938). [Limopsis]. Moore 1983. USGS PP 1228-A:A59, pl. 13-
ocene. Shell Oil Company Reyes 101  
USNM 496077. [Limopsidae].

Co. [USGS 13892]. Paratype USNM 496079.
Fimbria susanensis
of Meier Canyon, south side Simi Valley, Santa Susana quad. (1903., reprinted 1924), Ventura Co. UCMP 7009.
Holotype UCMP 38568. [Fimbriidae].
Fimbria susanensis
In a small gulley, in sandy shale about 100 m north of UCMP 3759 [=2134 m south of BM 961 at Santa Susana well,

Fimbria      
(1958), Los Angeles Co. CSUN 830 [=LACMIP 16166]. Hypotype LACMIP 7519. [Fimbriidae].
Flabellipecten carrizoensis (Arnold, 1906). [Pecten-
rial Fm. [=Latrania Fm.], Miocene/Pliocene. Head of Garnet Canyon, San Diego Co. Holotype SU/CASG 11 [=CASG.
61861.01]. [Pectinidae].
Flabellipecten hawleyi (Hertlein, 1925a). [Pecten
Oligocene/Miocene. Santa Inez [sic] Mtns., Santa Barbara Co. SU 860 [=CASG 61841]. Holotype SU/CASG 19
[=CASG 61841.01]. [Pectinidae].
Flabellipectenmerriami (Arnold, 1906). [Pecten
Pico Fm., Pliocene. San Felician Creek, near Piru, Ventura Co. Holotype UCMP 12086. Remarks: Junior synonym
of Patinopecten caurinus (Arnold, 1906),  Coan et al. (2000: p. 240). [Pectinidae].
Flabellipecten stearnsii (Dall, 1878a). [Pecten
Fm., Pliocene. Well at San Diego, San Diego Co. Holotype USNM 7942. Remarks: In the genus Leopecten, 
Waller (2007: pp. 937–938); in the genus Euvola,  Coan and Valentich-Scott (2012: p. 270). [Pectinidae].
Florimetis biangulataScrobicularia
2. Pancho Rico Fm., Miocene/Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M1929. Hypotype
USNM 649097. Remarks: Junior synonym of Psammotreta (Leporimetis) obesa (Deshayes, 1855),  Coan and
Valentich-Scott (2012: p. 617). [Tellinidae].
Florimetis biangulata Scrobicularia     
Margarita Fm., Miocene. Comanche Pt., Tejon Hills, Kern Co. USGS M1619. Hypotype USNM 649165. Remarks:
Junior synonym of Psammotreta (Leporimetis) obesa (Deshayes, 1855),  Coan and Valentich-Scott (2012: p.
617). [Tellinidae].
Florimetis biangulataScrobicularia]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):298, pl.

20312]. Hypotypes UCLA [48629, 48630 (=LACMIP 9312, 9313)]. Remarks: Junior synonym of Psammotreta
(Leporimetis) obesa (Deshayes, 1855),  Coan and Valentich-Scott (2012: p. 617). [Tellinidae].
66 PALEOBIOS, VOLUME 38, FEBRUARY 2021
 
Humboldt Co. Hypotype current whereabouts unknown.


Remarks: Macoma nasuta (Conrad 1837),  Bedrossian (1974: p. 178).
Gari (Gobraeus) cylindrica (Dickerson, 1914a). [Psammobia
 
Syntype UCMP 11677. [Psammobiidae].
Gari (Gobraeus) diegoensis (M.A. Hanna, 1927). [Psammobia]. Moore 2003b. USGS PP 1228-G [electronic format]

UCMP 31019. [Psammobiidae].
Gari edentulaSiliquaria
Domengine Ranch quad., Fresno Co. UCMP D1095. Hypotype UCMP 36737. Remarks: Junior synonym of Gari
(Gobraeus) fucata (Hinds 1845),  Coan et al. (2000: p. 426). [Psammobiidae].
Gari cf. 
Susana quad. (1951, PR 1969), Ventura Co. CSUN 471d [=LACMIP 16240]. Hypotype LACMIP 6587. Remarks:
Gari texta Gabb, 1864,  Squires (2001: p. 17). [Psammobiidae].
Gari
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7535. [Psammo-
biidae].
Gari (Gobraeus) eoundulata Vokes, 1939. Moore 2003b. USGS PP 1228-G [electronic format] (family Psammobi-

outcrop, Fresno Co. UCMP A820. Holotype UCMP 15707. [Psammobiidae].
Gari hornii (Gabb, 1864). [Tellina
Nojoqui Creek area, on west side of Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6920. Hypotype
SU 9242 (not SU 9322 as in text) [=CASG 70175]. Remarks: In the subgenus Gobraeus,  Moore (2003b: p.
1). [Psammobiidae].
Gari hornii (Gabb, 1864). [Tellina]. Deméré et al. 1979 Mission Valley Fm.,
Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH
20782. Remarks: In the subgenus Gobraeus,  Moore (2003b: p. 1). [Psammobiidae].
Gari (Gobraeus) hornii hornii (Gabb, 1864). [Tellina]. Moore 2003b. USGS PP 1228-G [electronic format] (family

ANSP 4453. [Psammobiidae].
Gari texta  
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 232 [=LACMIP 16075]. Hypotype LACMIP 7534. Remarks:
In the subgenus Gobraeus,  Moore (2003b: p. 4). [Psammobiidae].
Gari texta
Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 (=LACMIP 22312). Hypotype LACMIP 12755. Remarks:
In the subgenus Gobraeus,  Moore (2003b: p. 4). [Psammobiidae].
Gari (Gobraeus) texta Gabb, 1864. Moore 2003b. USGS PP 1228-G [electronic format] (family Psammobiidae):4, pl.


Hypotype LACMIP 14367. [Psammobiidae].
Gastrochaenadubitata
Susana quad. (1951, PR 1969), Ventura Co. CSUN 548 (=LACMIP 15450). Hypotype LACMIP 12758. [Remarks:
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 67
In the genus Rocellaria,  Carter et al. (2008:p. 110). [Gastrochaenidae].
Gibbolucina (Eomiltha) gyrata (Gabb, 1864). [Dosinia-
gine Fm. (Tejon Fm. in pl. caption), Eocene. Southeast of Mt. Diablo, Contra Costa Co. Holotype UCMP 11986.
[Lucinidae].
Gibbolucina (Eomiltha packi (Dickerson, 1916). [Lucina-
gine Fm. (Tejon Fm. in pl. caption), Parson's Peak area, Coalinga quad. [=Joaquin Rocks quad.], Fresno Co. UCMP
672. Holotype UCMP 11787. [Lucinidae].
Glans subquadrata (Carpenter, 1864b). [Lazaria
16, 18, 19, 22, 30. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co.
LACMIP 305. Hypotypes LACMIP 4538–4540]. Remarks: Renamed Cardita carpenteri by Lamy (1922) and ac-
cepted as valid under ICZN Art. 59; Glans carpenteri (Lamy, 1922); both  Coan et al. (2000: p. 305). [Carditidae].
Glans (Glans) subquadrata (Carpenter, 1864b). [Lazaria
Diego Fm., Pliocene. San Ysidro quad. (1943)], San Diego Co. [LACMIP 305]. Hypotype LACMIP 4539 (of Hertlein
Remarks: Renamed Cardita carpenteri by Lamy (1922) and accepted as valid
under ICZN Art. 59; Glans carpenteri (Lamy, 1922); both  Coan et al. (2000: p. 305). [Carditidae].
Glans (Centrocardita) veneriformis (Gabb, 1864). [Cardita]. Moore 1992. USGS PP 1228   
 of Martinez, Contra Costa Co. Holotype ANSP. 4381, paratype ANSP 4381a
[invalid number]. [Carditidae].
Glossus (Meiocardia) susukii-
bra Fm., Eocene. Northeastern Orocopia Mtns., Canyon Spring NW quad. (1963), Riverside Co. CSUN. 662 [=UCLA

10569]. [Glossidae].
Glossus (Meiocardia) susukii     
Maniobra Fm., Eocene. Canyon Spring NW quad. (1963), Riverside Co. CSUN 662 (=LACMIP 23779). Holotype
UCLA 48427 [=LACMIP 10568]. [Glossidae].
Glycymeris branneri
Miocene. On south and west slopes of Pyramid Hill, 4 or 5 mi. due north of the Power Plant in Kern River and

            
32221. Hypotype CASG 8565.
Glycymeris (Glycymeris) branneri        
Fm., Oligocene/Miocene. Mindego Creek, Santa Cruz quad., San Mateo Co. [SU 12]. Holotype USNM 165455.
[Glycymerididae].
Glycymeris (Glycymeris) buwaldi

Glycymeris coalingensis           
Northwest of Pleasanton, Dublin quad. (1953), Alameda Co. SU 3229. Hypotypes SU 8431, 8432 [=CASG 69949,
Remarks: Junior syn-
onym of Glycymeris (Axinola) grewingki Dall (1909),  Hertlein and Grant (1972: p. 159);  Moore (1983:
p. A58); junior synonym of Glycymeris (Axinola) septentrionalis (Middendorff, 1849),  Coan et al. (2000: p.
143). [Glycymerididae].


Glycymeris coalingensis
 Glycymeris       
68 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Remarks: Junior synonym of Glycymeris (Axinola) grewingki Dall, 1909,  Hertlein and Grant (1972:p. 159);
 Moore (1983:p. A58); junior synonym of Glycymeris (Axinola) septentrionalis (Middendorff, 1849),  Coan
et al. (2000: p. 143). [Glycymerididae].

Glycymeris in caption). East of Coalinga Sulphur Baths, west of Coalinga, Fresno Co.




Glycymeris aff.  (Weaver, 1912). [Pectunculus
9, 11. Middle Gaviota Fm., Eocene. Hills near junction of Yridisis Creek with El Jaro Creek, Lompoc quad., Santa
Barbara Co. UCMP B7007. Hypotypes SU 9251, 9252 [=CASG 70176, 70177]. [Glycymerididae].
Glycymeris (Tucetona) evermanni

Co. CASG 711. Holotype CASG 964. [Glycymerididae].
Glycymeris (Glycymeris) aff. () fresnoensis
Eocene. East side of Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4661. Hypotype UCR
4661/91. Remarks: In the subgenus Tucetona,  Moore (1983:p. A55). [Glycymerididae].
Glycymeris (Tucetona) fresnoensis
(Cerros Shale member), Paleocene [Eocene]. Vicinity of where Urruttia Canyon enters Salt Creek, Coalinga quad.,
Fresno Co. UCMP 1817. Holotype UCMP 11795. [Glycymerididae].
Glycymeris cf.  (Reeve, 1843). [Pectunculus
Fm., Miocene. Elderberry Canyon, Los Angeles Co. CIT 2075 [=LACMIP 16244. Hypotypes LACMIP 4077, 4259.
[Glycymerididae].
Glycymeris (Glycymeris) gigantea (Reeve, 1843). [Pectunculus      
Imperial Fm. [=Latrania Fm.], Pliocene. Near Travertine Rock, Imperial Co. (not San Diego Co. as in description).
USGS M9778. Hypotype USNM 486898. [Glycymerididae].
Glycymeris gigantea (Reeve, 1843) [Pectunculus
Latrania Fm., Miocene. Garnet Canyon, Imperial Co. ABDSP 5132.5. Hypotype ABDSP I 10684. [Glycymerididae].
Glycymeris grewingki
(Pseudocardium Zone), Pliocene. North branch of Arroyo Doblegado, southeast of El Rascador, North Dome,
Kettleman Hills, Kings Co. USGS 12406. Hypotypes USNM 495792, 495793. Remarks: In the subgenus Axinola,
 Moore (1983: p. A58). [Glycymerididae].
Siphonalia Zone), Pliocene. Arroyo Pino, north of Bullwheel Ridge, North Dome,
Kettleman Hills, Kings Co. USGS 12815. Hypotypes USNM 495827, 495828.
Glycymeris grewingki Dall, 1909. Weaver and Meyer 1969. AAPG/SEPM Guidebook, p. 101,  -
tato Harbor Fm., Pliocene. North side of Potato Harbor, Santa Cruz Is., California Channel Ids. UCSB GM65-21
[=LACMIP 16425]. Hypotype UCSB M7 [=LACMIP 6308]. Remarks: In the subgenus Axinola,  Moore (1983:
p. A58). [Glycymerididae].
Glycymeris (Axinola) grewingki
19, 22, 24. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP
305. Hypotypes LACMIP 4466, 4467. [Glycymerididae].
Glycymeris (Glycymerita) major (Stanton, 1896). [Pectunculus 
10. Martinez Fm., Paleocene. Southeast of Lower Lake, Lake Co. USGS 4192. Syntype USNM 157830 (erroneously
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 69
cited by Keen and Bentson (1944:96) as USNM 157827). Remarks: In the genus Glycymerita,  Squires (2010:
p. 909). [Glycymerididae].
Glycymeris major (Stanton, 1896). [Pectunculus-
sana Fm., Paleocene. Simi Hills, Ventura Co. UCR6901. Hypotype UCR 6901/3 (not UCR 6891/5 as in pl. caption).
Remarks: In the genus Glycymerita,  Squires (2010: p. 909). [Glycymerididae].
Glycymeris (Glycymerita) major meganosensis Clark and Woodford, 1927. Moore 1983. USGS PP 1228-A:A53–A54,

Holotype UCMP 31289. [Glycymerididae].
Glycymeris (Tucetona maccrayi
Oil Canyon, Ventura Co. SU. 8 [=CASG 61913]. Holotype SU/CASG 167 [=CASG 61913.04]. Remarks: In the genus
Glycymerita,  Squires (2010: p. 909). [Glycymerididae].
Glycymeris meganosensis-
cene/Eocene. Mt. Diablo quad., Contra Costa Co. [LACMIP 7173. Hypotype LACMIP 4078]. Remarks: Glycymeris
(Glycymerita) major meganosensis,  Moore (1983: p. A53). [Glycymerididae].
Glycymeris (Tucetona) multicostata (G.B. Sowerby I, 1833a). [Pectunculus]. Powell 2008. USGS SIR 2008-5155:8–9,

description). USGS M9779 [=USGS M9778]. Hypotype USNM 486905. [Glycymerididae].
Glycymeris perrini
Ridge, Kettleman Plain quad., Kings Co. USGS 14482. Hypotype USNM 498734. Remarks: In the subgenus Tuce-
tona,  Moore (1983: p. A55). [Glycymerididae].
Glycymeris (Glycymeris() perrini
Eocene. Thorn Pt. area, Lion Canyon quad. (1943), Ventura Co. UCR 4697. Hypotype UCR 4697/301. Remarks:
In the subgenus Tucetona,  Moore (1983: p. A55). [Glycymerididae].
Glycymeris perrini Dickerson, 1916. Deméré et al, 1979. SEPM Guidebook 4:98, pl. 1, 
Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH
20764. Remarks: In the subgenus Tucetona,  Moore (1983: p. A55). [Glycymerididae].
Glycymeris (Tucetona) perrini
Fm., Eocene. Parson's Peak area, Coalinga quad., Fresno Co. UCMP 672. Holotype UCMP 11792. [Glycymerididae].
Glycymeris (Axinola) profunda (Dall, 1878a). [Axinea]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):160–161, pl. 27,
Remarks:
Valid species,  Moore (1983: p. A58); junior synonym of Glycymeris (Axinola) septentrionalis (Middendorff,
1849),  Coan et al. (2000:p. 143). [Glycymerididae].
     
Hypotypes LACMIP 4468, 4469.

Glycymeris (Glycymerita reefensis
Big Tar Canyon, Cholame quad., Kings Co. UCMP A1165. Holotype UCMP 15571. [Glycymerididae].
Glycymeris (Glycymeris) rosecanyonensis

UCMP 3990. Holotype UCMP 30989. [Glycymerididae].
Glycymeris (Glycymeris) rosecanyonensis
Fm., Eocene. Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 489 [=LACMIP 16103]. Hypotype LACMIP
6565. [Glycymerididae].
Glycymeris (Glycymeris) rosecanyonensis
70 PALEOBIOS, VOLUME 38, FEBRUARY 2021
-
type LACMIP 7508. [Glycymerididae].
Glycymeris () rosecanyonensis-
mere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13437. [Glycymeri-
didae].
Glycymeris (Tucetona) ruckmani Dickerson, 1915. [Glycimeris   
9. Tejon Fm., Eocene. West side of Grapevine Creek, Tejon quad., Kern Co. UCMP 458. Holotype UCMP 11051.
[Glycymerididae].
Glycymeris sagittata (Gabb, 1864). [Axinaea (Limopsis?)      
Sandstone, Eocene. Reef Ridge, Reef Ridge quad., Fresno Co. 14497a. Hypotype USNM 498733. Remarks: In the
subgenus Glycymerita,  Moore (1983: p. A54). [Glycymerididae].
Glycymeris sagittata (Gabb, 1864). [Axinaea (Limopsis?)]. Weaver and Kleinpell 1963. UCPG 43:196–197, pl. 28,
 
B6955. Hypotype SU 9254 [=CASG 70178]. Remarks: In the subgenus Glycymerita,  Moore (1983: p. A54).
[Glycymerididae].
Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6954. Hypotype SU 9253
[=CASG 70179].
Glycymeris sagittata (Gabb, 1864). [Axinaea (Limopsis?)
Mission Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Resevoir, San Diego Co. SDSNH 2962. Hypotype
SDSNH 20763. Remarks: In the subgenus Glycymerita,  Moore (1983: p. A54.) [Glycymerididae].
Glycymeris (Glycymerita) sagittata (Gabb, 1864). [Axinaea (Limopsis?)]. Moore 1983. USGS PP 1228-A: A54–A55,
 
[cited as holotype in Stewart (1930: p. 72); Keen and Bentson (1944: p. 31); and Richards (1968: p. 82)] ANSP
4422. [Glycymerididae].
Glycymeris (Glycymerita) sagittata (Gabb, 1864). [Axinaea (Limopsis?)
Llajas Fm., Eocene. Devil Canyon area, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype
LACMIP 6566. [Glycymerididae].
Glycymeris (Glycymerita) sagittata (Gabb,1864). [Axinaea (Limopsis?)
Juncal Fm., Eocene. Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 807 [=LACMIP 16191].
Hypotype LACMIP 7509. [Glycymerididae].
Glycymeris (Glycymerita) sagittata (Gabb, 1864). [Axinea (Limopsis?)

Hypotype LACMIP 7726. [Glycymerididae].
Glycymeris (Glycymerita) sagittataAxinaea (Limopsis?)]. Squires 1991c. SEPM Guidebook 68:222,

(=LACMIP 23779). Hypotype LACMIP 11480. [Glycymerididae].
Glycymeris (Glycymerita) sagittata (Gabb, 1864). [Axinaea (Limopsis?)
Juncal Fm., Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype
LACMIP 13438. [Glycymerididae].
Glycymeris (Tucetona septentrionalis (Middendorff, 1948). [Pectunculus]. Moore, 1983. USGS PP 1228-A:A57, pl.


Glycymeris (Glycymerita tecolotensis   

[Glycymerididae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 71
Glycymeris (Tucetona) tenuimbricata Clark, 1918. [Glycimeris
24, 25 (as Glycymeris (Tucetona) teniumbricata
Creek], Contra Costa Co. UCMP 1131. Holotype UCMP 11183. [Glycymerididae].
Glycymeris veatchii major (Stanton, 1896). [Petunculus]. Smith 1945
[=Santa Susana Fm.], Paleocene. South side Simi Valley, on crest of Simi Hills, Ventura Co. LACMIP 7089. Hypotype
LACMIP 4083]. Remarks: In the genus Glycymerita, 
11.3). [Glycymerididae].
Glycymeris (Tucetona) verticordia
 
Holotype CASG 776. [Glycymerididae].
Glycymeris (Glycymeris) viticola
Fm., Eocene. Near Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4723. Hypotype UCR 4723/141.
[Glycymerididae].
Glycymeris (Glycymeris) viticola

CASG 777. [Glycymerididae].
Glycymeris whaleyi  

72121]. Remarks: Provisionally in the subgenus Glycymeris Moore (1983: p. A53). [Glycymerididae].

8584 [=CASG 32221.01], paratype CASG 8585 [=CASG 32221.02].
Glycymeris (Glycymeris whayleyi
(Saltos Shale Member), Miocene. Caliente Mtn. quad., San Luis Obispo Co. Holotype CASG 8584 [=CASG. 32221.01].
[Glycymerididae].
Glycymeris

-
Remarks:
therefore the exact locality is unknown.
  

Glycymeris (Glycymeris-
ceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA 6528
[=LACMIP 26528]. Hypotype LACMIP 7557. [Glycymerididae].
Glycymeris

unnumbered hypotype lost,  C. Throckmorton (personal communication, 2002). [Glycymerididae].
Glycymeris
Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8734 [=SBMNH 60168]. [Glycymerididae].
Glycymerita major (Stanton, 1896). [Petunculus
Fm., Paleocene. Near junction of Silver Creek with Panoche Creek, NE end of Tumey Hills, Fresno Co. LACMIP
7044. Hypotype LACMIP 13628, 13629. [Glycymerididae].

Hypotype (of Glycymeris veatchii major
72 PALEOBIOS, VOLUME 38, FEBRUARY 2021
    
Angeles Co. LACMIP 22334. Hypotypes LACMIP 13630, 13631

[=LACMIP 41602]. Hypotype LACMIP 13632.


Fresno Co. LACMIP 4845. Hypotype LACMIP 13633.

Glycymerita  
Topanga Rd. area, Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 12589. Line drawings, source
specimen unknown. [Glycymerididae].
Glyptoactis domenginica (Vokes, 1939). [Venericardia    
Fm., Eocene. Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6575. [Carditidae].
Glyptoactis (Glyptoactis) domenginica (Vokes, 1939). [Venericardia
Juncal Fm., Eocene. Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 828 [=LACMIP 16141].
Hypotype LACMIP 7526. [Carditidae].
Glyptoactis (Claibornicardia) domenginica (Vokes, 1939). [Venericardia]. Squires 1988a. SEPM Guidebook 58:110,
-
type LACMIP 6575. [Carditidae].
Glyptoactis (Claibornicardia) domenginica (Vokes, 1939). [Venericardia
  
[Carditidae].
Glyptoactis (Claibornicardia) domenginica (Vokes, 1939). [Venericardia]. Squires 1991c. SEPM Guidebook 68:222,

16082]. Hypotype LACMIP 11483. [Carditidae].
Glyptoactis (Claibornicardia) domenginica (Vokes, 1939). [Venericardia]. Moore 1992. USGS PP 1228-E:E27–E28, pl.
13. Domengine Fm., Eocene. Coalinga quad., Fresno Co. UCMP A1219. Holotype UCMP 15611. [Carditidae].

15612, 15613.
Glyptoactis (Claibornicardia) keenae (Verástegui, 1953). [Venericardia
6, 10, 12, 15. Basal Lodo Fm., Paleocene. Near junction of Panoche and Silver creeks, Panoche quad., Fresno Co.
CASG 2073. Holotype (of Venericardia () argentea Verástegui, 1953) SU/CASG 7992 [=CASG 66535.01],
holotype SU/CASG 7995 [=CASG 66535.02]. [Carditidae].

Glyptoactis (Claibornicardia) marksi
Fm. (Live Oak member of Verástegui, 1953), Eocene. East side of Live Oak Canyon, Kern Co. CASG 183. Holotype
SU/CASG 8021 [=CASG 6536.01]. [Carditidae].
Glyptoactis (Claibornicardia) sandiegoensis (M.A. Hanna, 1927). [Cardita
110. Juncal Fm., Eocene Whitaker Peak quad. (1958), Los Angeles Co. CSUN 844 [=LACMIP 16119]. Hypotype
LACMIP 7527. [Carditidae].
Glyptoactis (Claibornicardia) sandiegoensis (M.A. Hanna, 1927). [Cardita]. Squires 1988a. SEPM Guidebook 58:110,
       
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 73
LACMIP 7527. [Carditidae].
Glyptoactis (Claibornicardia) sandiegoensis (M.A. Hanna, 1927). [Cardita]. Moore 1992. USGS PP 1228-E:E26–E27,

San Diego Co. UCMP 5062. Syntypes UCMP 30980, 30982. [Carditidae].
Glyptoactis (Claibor-
nicardia) mcmastersi Verástegui, 1953) CASG 8011 [=CASG 66534.01].
Glyptoactis (Claibornicardia) sandiegoensis (M.A. Hanna, 1927). [Cardita]. Squires 1994b. SEPM Guidebook 74:54,
     
11662. [Carditidae].
Glyptoactis (Claibornicardia) sandiegoensis (M.A. Hanna, 1927). [Cardita 
45. Juncal Fm., Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype
LACMIP 13446. [Carditidae].
Glyptoactis

Gobraeus edentulus (Gabb, 1869a). [Siliquaria]. Hertlein and Grant 1972. SDSNH. Mem
15. San Diego Fm., Pliocene. Near mouth of Rose Canyon, La Jolla quad., San Diego Co. UCLA 2359 [=LACMIP
22359]. Hypotypes UCLA [=LACMIP 9315, 9316]. Remarks: Junior synonym of Gari (Gobraeus) fucata (Hinds,
1845),  Coan et al. (2000: p. 426). [Psammobiidae].
Gregariella chenui (Récluz, 1842). [Mytilus (Modiola)]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):168–169, pl. 41,

305. Hypotype LACMIP 4477. [Mytilidae].

Gryphaeostrea aviculiformis (Anderson, 1905). [Ostrea-
gine Fm., Eocene. West and north of Coalinga, Fresno Co. CASG 66747. Holotype CASG 21 [=CASG 66747.01].
[Gryphaeidae].
Gryphaeostrea aviculiformis (Anderson, 1905). [Ostrea
Ostrea aviculiformis Anderson 1905). Domengine Fm., Eocene.
Alcalde Hills, west of Coalinga, Fresno Co. CASG 66747. Holotype CASG 66747.01 [ex CASG 21]. [Gryphaeidae].

Halonanus hornii hornii (Gabb, 1864). [Arca    
Fm., Eocene. Parson's Peak, north of Coalinga, Fresno Co. UCMP 672. Hypotype UCMP 32544. Remarks: In the
genus/subgenus Pachecoa (Pachecoa),  Moore (1983: p. A48). [Arcidae].

Hawaiarca mediaimpressa s.s. (Clark, 1918). [Arca (Scapharca)]. Noda 1986. Paleo. Soc. Japan Sp. Pap. 29:64, pl.

1174. Remarks: In the genus/subgenus Anadara (Anadara),  Moore (1983: p. A38). [Arcidae].
Here (Here)  (Weaver and Kleinpell, 1963). [Lucina

Barbara Co. UCMP B6955. Holotype SU/CASG 9285 [CASG 70181]. [Lucinidae].
Here excavata (Carpenter, 1857b). [Lucina
(Wygal Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotypes USNM 646545–
646547. [Lucinidae].
Here excavata (Carpenter, 1857b). [Lucina
74 PALEOBIOS, VOLUME 38, FEBRUARY 2021
area, Los Angeles Co. LACMIP 17920. Hypotype LACMIP 14351. [Lucinidae].
Here (Here) excavata (Carpenter, 1857b). [Lucina

....., as Lucina (Here) richtofeni Gabb, 1869)). Miocene [=Pliocene,
Pico Co. Lectotype (of Stewart 1930: p. 182) ANSP
4492], missing,  K. Estes-Smargiassi (personal communication, 2019). [Lucinidae].
Here (Here) excavata (Carpenter, 1857b). [Lucina





Lucina richthofeni Gabb, 1866) ANSP 4492.

UCMP A510. Hypotype (of Phacoides richthofeni
Here (Here) hannai (Clark, 1938). [Lucina  
Fm., Eocene. Northeast bank of Pleasants Creek, Napa quad., Solano Co. UCMP A1297. Holotype UCMP 30841,
paratype UCMP 30842. [Lucinidae].
Heteromacoma rostellata (Clark, 1918). [Metis
Fm. (Phacoides Sand member) (of Stinemeyer et al. 1959:12, 13), Miocene. Las Yeguas Ranch quad., Temblor
Range, Kern Co. USGS M3280. Hypotypes USNM 646564, 646568. Remarks: In the genus Leporimetis,  Moore
(2003b: p. 52). [Tellinidae].
Heteromacoma rostellata (Clark, 1918). [Metis

Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotypes USNM 646563, 646564,
646567, 646568. Remarks: In the genus Leporimetis,  Moore (2003b: p. 52). [Tellinidae].

   
M3772. Hypotype USNM 646566.

646582.
Hiatella arctica (Linneaus, 1767). [Mya    
17, 18. San Diego Fm., Pliocene, San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotypes LACMIP 4829, 4830.
[Hiatellidae].
Hilgardiaparkei (Anderson and G D. Hanna, 1925). [Leda
7. Tejon Fm., Eocene. Live Oak Creek, Tejon quad., Kern Co. CASG 244. Holotype CASG 782 (determined here as
missing), paratype CASG 783. [Nuculanidae].
Hilgardiaparkei (Anderson and G D. Hanna, 1925). [Leda
Sandstone
Hypotype LACMIP 7505. [Nuculanidae].
Hilgardia parkei (Anderson and G D. Hanna, 1925). [Leda
Eocene. Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 26624. Hypotype LACMIP
12747. [Nuculanidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 75
Hinnites benedicti-
nent ledge-forming Ostrea-Pecten
paratype UCMP 36635. Remarks: Determined here as in the genus Crassadoma,. [Pectinidae].
Hinnites benedicti
From approximately 2 m-thick oyster bed, Domengine Ranch quad., Fresno Co. (Kern Co. in locality description).
UCMP D702. Holotype UCMP 36638. Remarks: Determined here as in the genus Crassadoma. [Pectinidae].
Hinnites cf. 
8. Etchegoin Fm. (Siphonalia Zone), Pliocene. El Campo, North Dome, Kettleman Hills, Kings Co. USGS 12440.
Hypotypes USNM 495810, 495812. Remarks: Junior synonym of Crassadoma gigantea (Gray, 1825),  Coan
et al. (2000: p. 238). [Pectinidae].

Hinnites giganteus (Gray, 1825). [Lima
Remarks: Crassa-
doma gigantea (Gray, 1825), Coan et al. (2000: p. 238). [Pectinidae].
Hinnites giganteus (Gray, 1825). [Lima
Robles Fm., Pliocene. Near Atascadero, San Luis Obsipo Co. USGS M4621. Hypotypes USNM 647003 (with Balanus
sp.), 647008. Remarks: In the genus Crassadoma,  Coan et al. (2000: p. 238). [Pectinidae].
Hinnites giganteus (Gray, 1825). [Lima       
Unnamed strata, Pliocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS M6501.
Hypotypes USNM 235599, 235600. Remarks: Crassadoma gigantea (Gray, 1825),  Coan et al. (2000:p. 238).
[Pectinidae].
Hinnites giganteus (Gray, 1825). [Lima
Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2857. Hypotype USNM 254386.
Remarks: Crassadoma gigantea (Gray, 1825), Coan et al. (2000: p. 238). [Pectinidae].
Hinnites giganteus (Gray, 1825). [Lima      
  
Luis Obispo Co. USGS M2857. Hypotype USNM 254386. Remarks: Crassadoma gigantea (Gray, 1825),  Coan
et al. (2000: p. 238). [Pectinidae].
Hinnites giganteus (Gray, 1825). [Lima
Margarita Fm., Miocene. Salinas Valley, San Luis Obispo Co. Holotype (lectotype,  Keen and Bentson (1944:
p. 54) USNM 13336, paratype USNM 13336a (both of Hinnites crassa Conrad 1857d). Remarks: Crassadoma
gigantea (Gray, 1825),  Coan et al. (2000: p. 238). [Pectinidae].
Hinnites multirugosus var. crassiplicatus (Gale, 1928). [Pecten
Castaic Fm., Miocene. Humphreys quad. (1932, PR 1946), Los Angeles Co. CIT 2069 [=LACMIP 2558]. Hypotype
LACMIP 4095. Remarks: Junior synonym of Crassadoma gigantea (Gray, 1825),  Coan et al. (2000: p. 238).
[Pectinidae].
Hinnites multirugosus subsp. crassiplicatus (Gale, 1928). [Pecten
Margarita Fm., Miocene. Prominent ledge-forming Ostrea-Pecten  
UCMP D702. Hypotypes UCMP 36633, 36634. Remarks: Junior synonym of Crassadoma gigantea (Gray, 1825),
 Coan et al. (2000: p. 238). [Pectinidae].
Humilaria perlaminosa (Conrad, 1855a) of Arnold (1907) [Mercenaria]. Addicott 1969. Pr.CAS 37(3):63, 68, 73,
   
[Veneridae].
Idasola bakeri (Dickerson, 1914a). [Modiolus
Paleocene. South of Stewartville, Mt. Diablo quad., Contra Costa Co. UCMP 1556. Holotype UCMP 11686. [Mytilidae].

76 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8743 [=SBMNH 60173]. Remarks: Determined
here as Pycnodonte sp. [Gryphaeidae].

Santa Cruz Mtns., Santa Cruz Co. Hypotype current whereabouts unknown. Remarks: Determined here as Op-
penheimopecten sanctaecruzensis (Arnold, 1906). [Pectinidae].
Indet solenid (not Ficus   
Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8713 [=SBMNH 60186].
Remarks: Determined here as indet. solenid. [Solenidae].

Cañada Posa area, Santa Cruz Is., California Channel Ids. [Hypotypes SBMNH 60170–60172]. Remarks: All de-
termined here as l Pycnodonte sp. [Gryphaeidae].

Paleocene/Eocene. Cañada Posa area, Santa Cruz Is., California Channel Is. [Hypotype SBMNH 60187]. Remarks:
Determined here as indet. teredinid. [Teredinidae].

North of Iverson Pt., Saunders Reef quad., Mendocino Co. UCMP D7149. Hypotype UCMP 14440. Remarks: De-
termined here as possibly a corbulid.
Irusella lamellifera (Conrad, 1837). [Venus
Diego Fm., Pliocene. Lowest Pecten healeyi bed, from just south of a ravine which is immediately south of the
-
potype UCLA [=LACMIP 9306]. [Veneridae].
Isognomon (Isognomon) clarkiPedalion
Torrey Sandstone, Eocene. Hillside exposure (now mostly covered), Del Mar quad. (1967), San Diego Co. SDSNH
3282. Hypotypes SDSNH 35235, 35236. [Isognomonidae].
Isognomon (Isognomon) clarkiPedalion    
Coldwater Sandstone, Eocene. Upper Sespe Creek, Ventura Co. CSUN 252 [=LACMIP 16670]. Hypotype LACMIP
11657. [Isognomonidae].
Isognomon (Isognomon) panzana (Loel and Corey, 1932). [Pedalion
6. Vaqueros Fm., Oligocene/Miocene. San Luis Obispo Co. UCMP A498. Holotype UCMP 31780. [Isognomonidae].
Isognomon (Isognomon) panzana (Loel and Corey, 1932). [Pedalion
1–6. Vaqueros Fm., Miocene. West side of Plano Trabuco, Santa Ana Mtns., Orange Co. LACMIP 16667 (not LAC-
MIP 27006 as in pl. caption). Hypotypes LACMIP 12371–12374 (not LACMIP 12251–12254 as in pl. caption).
[Isognomonidae].
Isognomon-
leocene/Eocene. Bryon quad., Contra Costa Co. UCMP 3609. Hypotype UCMP 31301. Remarks: Equivalent to
Pedalion
Isognomon-
tura Co. UCR 4752. Hypotype UCR 4752/61. Remarks: Isognomon (Isognomon) clarki Squires
(1989b: p. 279). [Isognomonidae].
Isognomon          
Remarks: Isognomon (Isognomon)
clarki Squires (1989b: p. 279). [Isognomonidae].
Jouannetia        -
cene/Eocene. On top of ridge at middle of north edge of Contra Costa Co. UCMP 3158. Hypotype UCMP 31340.
[Pholadidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 77
Jupiteria taphria (Dall, 1896b). [Leda      
area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14336. Remarks: In the genus Saccella,  Coan and
Valentich-Scott (2012: p. 94). [Nuculanidae].
Katherinella (Katherinella) angustifrons (Conrad, 1849a). [Venus    

CASG 101112, 101113, 101115–101118].
Kellia catacta Anderson and G D. Hanna, 1925. 7. Tejon Fm., Eocene. Live
Oak Creek, Tejon quad., Kern Co. CASG 244. Holotype CASG 275. [Lasaeidae].
Kellia lajollaensis
Hanna 1927: p. 256), Eocene. San Clemente Canyon, where the canyon enters Rose Canyon, San Diego Co. UCMP
3991. Holotype UCMP 31036. [Lasaeidae].
Kellia laperousii (Deshayes, 1839). [Chironia    
19. San Diego Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 4623. Remarks:
Valid species,  Moore (1992: p. E7); junior synonym of Kellia suborbicularis (Montagu, 1803),  Coan et al.
(2000: p. 323). [Lasaeidae].
Kellia laperousii (Deshayes, 1839). [Chironia

19) LACMIP 4623. Remarks: Valid species,  Moore (1992: p. E7); junior synonym of Kellia suborbicularis
(Montagu, 1803),  Coan et al. (2000: p. 323). [Lasaeidae].
Kellia uvasana (Dickerson, 1915). [Corbula
Live Oak Canyon, Tejon quad., Kern Co. CASG 244. Holotype CASG 276 [=CASG 244.04]. Hypotype (of Anderson

Laevicardium (Cerastoderma) meekianum (Gabb, 1866). [Cardium

UCMP 37626. Remarks: In the genus Clinocardium,  Moore (2003a: p. 20). [Cardiidae].
Larkinia camuloensis (Osmont, 1905). [Arca       
Pliocene. Between Salt and Tapo canyons, Ventura Co. UCLA 1363 [=LACMIP 21363]. Hypotype UCLA 49525
[=LACMIP 10624]]. Remarks: Junior synonym of Anadara (Larkinia) multicostata (G.B. Sowerby I, 1833b), 
Coan and Valentich-Scott (2012: p. 179). [Arcidae].
Larkinia camuloensis (Osmont, 1905). [Arca
Fm. in text), Pliocene. 1.5 km north of Camulos, Ventura Co. Holotype UCMP 12006. Remarks: Junior synonym of
Anadara (Larkinia) multicostata (G.B. Sowerby I, 1833b),  Coan and Valentich-Scott (2012: p. 179). [Arcidae].
Larkinia santana santana (Loel and Corey, 1932). [Arca
Vaqueros Fm., Miocene. West side of Plano Trabuco, Santa Ana Mtns., Orange Co. UCMP 6128. Holotype UCMP
31768. [Arcidae].
Larkinia santana weddlei (Loel and Corey, 1932). [Arca
Fm., Miocene. Junipero Serra quad., Monterey Co. UCMP 3675. Holotype UCMP 31771. [Arcidae].
Ledina duttonae (Vokes, 1939). [Jupiteria]. Moore 1983. USGS PP 1228
Member), Paleocene/Eocene [Eocene]. Vicinity where Urruttia Canyon enters Salt Creek, Coalinga quad., Fresno
Co. UCMP 1817. Holotype UCMP 15564. [Nuculanidae].
Ledina duttonae (Vokes, 1939). [Jupiteria-
cene. Inyokern SE quad. (1972), El Paso Mtns., Kern Co. USGS M9177. Hypotype USNM 424892. [Nuculanidae].
Ledina fresnoensis (Dickerson, 1916). [Leda 
Springs Creek, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4662. Hypotype UCR 4662/101. [Nuculanidae].
Ledina fresnoensis (Dickerson, 1916). [Leda
78 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Shale Member), Paleocene [Eocene]. Vicinity where Urruttia Canyon enters Salt Creek, Coalinga quad., Fresno
Co. UCMP 1817. Holotype UCMP 11790. [Nuculanidae].
Leochlamyserici (Wiedey, 1928). [Pecten
Oligocene/Miocene. Southwest of Santa Paula, Ventura Co. SU 406. Holotype SDSNH 27. [Pectinidae].
Leopecten stearnsii (Dall, 1878a). [Pecten
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14347. [Pectinidae].
Leporimetis dombei (Hanley, 1844). Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):53, pl. 7,
Leporimentis dombi (Hanley, 1844). Latrania Fm., Miocene/ Pliocene. West side of Coyote Mtn., Impe-
Remarks: In the genus/subgenus
Psammotreta (Leporimetis),  Coan and Valentich-Scott (2012: p. 617). [Tellinidae].
Leporimetis obesa (Deshayes, 1855). [Tellina
area, Los Angeles Co. LACMIP 17921. Hypotype LACMIP 14366. Remarks: In the genus/subgenus Psammotreta
(Leporimetis),  Coan and Valentich-Scott (2012: p. 617). [Tellinidae].
Leporimetis rostellata (Clark, 1918). Moore 2003b. [Metis]. USGS PP 1228-G [electronic format] (family Tellini-

Contra Costa Co. UCMP 1131. Holotype UCMP 11120. [Tellinidae].
Leptopecten andersoni (Arnold, 1906). [Pecten
Fm. (Gould Shale Member), Miocene. Las Yeguas Ranch quad., Temblor Range, Kern Co. USGS M3282. Hypotype
USNM 646802. Remarks: In the genus Pacipecten,  Moore (1984: p. B45). [Pectinidae].

quad., Temblor Range, Kern Co. USGS M3987. Hypotype USNM 646809.
Leptopecten andersoni (Arnold, 1906). [Pecten      
Monterey Fm. (Saltos Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M5026.
Hypotypes USNM 254358, 254368. Remarks: In the genus Pacipecten,  Moore (1984: p. B45). [Pectinidae].
Leptopecten andersoni (Arnold, 1906). [Pecten

area, La Panza Range, San Luis Obispo Co. USGS M5026. Hypotypes USNM 254358, 254368. Remarks: In the
genus Pacipecten,  Moore (1984: p. B45). [Pectinidae].
Leptopecten bellilamellatus (Arnold, 1906). [Pecten
Remarks: Junior
synonym of Leptopecten latiauratus (Conrad, 1837),  Coan et al. (2000: p. 237). [Pectinidae].
Leptopecten discus (Conrad, 1857d). [Pecten
6,7, 9–12. Unnamed strata, Miocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS
M3743b. Hypotypes USNM 235577–235581. Remarks: In the genus Pacipecten,  Moore (1984: p. B46).
[Pectinidae].

13335. [Pectinidae].
Leptopecten discus (Conrad, 1857d). [Pecten
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M3568. Hypotype USNM
254385. Remarks: In the genus Pacipecten,  Moore (1984: p. B46). [Pectinidae].
Leptopecten discus (Conrad, 1857d). [Pecten
   
Range, San Luis Obispo Co. USGS M3568. Hypotype USNM 254385. Remarks: In the genus Pacipecten,  Moore
(1984: p. B46). [Pectinidae].
Leptopecten pabloensis (Conrad, 1857c). [Pecten
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 79
2. Santa Margarita Fm., Miocene. Atascadero quad., San Luis Obispo Co. USGS M3934. Hypotypes USNM 235574,
235575. Remarks: In the genus Pacipecten,  Moore (1984: p. B47). [Pectinidae].
Leptopecten (Leptopecten) palmeri (Dall, 1897). [Pecten-
rial Fm. (Latrania Sand Member) [=Latrania Fm.], Miocene/Pliocene. Willis Palm area, Riverside Co. UCR 4777.
Hypotype UCR 4777/80 (not UCR 4777/11 as in pl. caption). [Pectinidae].
Leptopecten
Miocene. Carneros Rocks quad., Temblor Range, Kern Co. USGS M3983. Hypotype USNM 646784. [Pectinidae].
Leptopecten 
Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM 646527. [Pectinidae].
Lima (Limaria) orcutti Hertlein and Grant, 1972 (
Diego Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Paratype LACMIP
   Remarks: In the genus Limaria,  Moore (1987: p. C16).
[Limidae].
    



4522, 4523].
Lima vedderi
Juan Capistrano quad. (1949), Orange Co. USGS M3220. Holotype USNM 240060. Remarks: In the subgenus
Lima,  Moore (1987: p. C13). [Limidae].

Lima (Lima) vedderi
East bank of Aliso Creek, San Juan Capistrano quad. (1949.), Orange Co. USGS M3220 (not USGS 3220 as in local-
ity description). Holotype USNM 240060. [Limidae].
Lima (Lima) n. sp. aff. () tetrica
  
Hypotype USGS 235596. [Limidae].
Lima
Orange Co. SDSNH 4312. Hypotype SDSNH 83247. [Limidae].
Lima        
member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312. Hypotype
SDSNH 140182. [Limidae].
Indeterminate LimaPaleoBios-
tion, Berkeley Hills, Oakland, Alameda Co. UCMP IP 13001. Hypotype UCMP 412479. [Limidae].
Limatula aff.  (Montagu, 1808). [Pecten

496096. Remarks: Junior synonym of Limatula saturna Bernard, 1978,  Coan et al. (2000: p. 207). [Limidae].
Limaria (Limaria) clarki (Nelson, 1925). [Lima
Susana Fm.], Paleocene. Simi Valley, Santa Susana quad., Ventura Co. UCMP 3752. Holotype UCMP 30494. [Limidae].
Limaria (Limaria) orcutti (Hertlein and Grant, 1972). [Lima]. Moore 1987. USGS PP 1228-C:
8. San Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305C [=LACMIP 16817]. Holotype
LACMIP 4509, paratype LACMIP 4511. [Limidae].
80 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Limaria sp., cf.  (Hertlein and Grant, 1972). [Lima
Pliocene. Newhall area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14339. [Limidae].
Limaria
member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312. Hypotype
SDSNH 121368. [Limidae].
LimeaIsolimea claytonensis (Dickerson, 1914a). [Lima
Martinez Fm., Paleocene. Mt. Diablo quad., Contra Costa Co. UCMP 1592. Syntypes UCMP 11720, 11721. [Limidae].
Limopsis (Limopsis) marysvillensis (Dickerson, 1913). [Glycymeris
Fm., Eocene. East side of Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4656. Hypotype
UCR 4656/101. [Limopsidae].
Limopsis (Limopsis) marysvillensis (Dickerson, 1913). [Glycymeris
10. Meganos Fm. (Marysville Claystone Member) (not Capay Fm. as in locality description), Paleocene/Eocene.
Marysville (Sutter) Buttes, Marysville (Sutter) Buttes quad., Sutter Co. UCMP 1853. Holotype UCMP 11766. [Li-
mopsidae].
Lithophaga (Lithophaga) clarki
Eocene. Soledad Valley, La Jolla quad., San Diego Co. UCMP 3981. Holotype UCMP 31031. [Mytilidae].
Lithophaga sp. Watkins 1990b. Palaios-
rial Co. [MPM 90088]. Hypotype MPM 28229 (extracted from Gastrochaenolites Ostreaheermanni).
[Mytilidae].
Sole-
nastrea fairbanksi). [Mytilidae].
Lituyapecten dilleri (Dall, 1901a). [Pecten]. Addicott 1981. GSA Sp. Pap. 184:34, pl. 5. Wildcat Group, Pliocene.
Thompkins Hill, Humboldt Co. USGS M7258. Hypotype USNM 252514. [Pectinidae].
Lituyapecten dilleri (Dall, 1901a). [Pecten
Eel River, Humboldt Co. Lectotype (of Moore 1984: p. B88) USNM 164846. [Pectinidae].
Lituyapecten falorensis (MacNeil, 1961). [Pecten
Pliocene. About 16 km southeast of Blue Lake, Humboldt Co. UCMP A4233. Holotype UCMP 34172. [Pectinidae].
Lituyapecten purisimaensis (Arnold, 1906). [Pecten]. Moore, 4. Purisima
Fm., Pliocene. North of mouth of Pescadero Creek, Santa Cruz quad., San Mateo Co. Holotype SU/CASG 3 [=CASG
61852.01]. [Pectinidae].
Lituyapecten turneri (Arnold, 1906). [Pecten
Pliocene. Arroyo San Antonio, Tomales Bay, Marin Co. Holotype SU/CASG 430 [=CASG 61949.02], paratype SU/
CASG 430A [=CASG 61949.03]. [Pectinidae].
Lopha vespertina (Conrad, 1854). [Ostrea
Pliocene. Fish Creek area, Fish Creek Mtns., Imperial Co. [MPM 90015]. Hypotypes MPM 28212–28217. Remarks:

the genus Dendostrea Moore (1987: p. C25). [Ostreidae].

 beds].

 beds].

 beds].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 81

 beds].
 with
encrusting cheilostome bryozoa).
 with
balanid barnacle).
-
pertina with attached  (MPM 28224), with Gastrochaenolites (MPM 28225), with Entobia borings
(MPM 28226), and with Caulostrepsis borings MPM. 28227, 10F.
Lucina (Lucinoma) acutilineata
Temblor Fm., Miocene. South of Oil City, Fresno Co. CASG 27276. Hypotype CASG 5730. [Lucinidae].
Lucina (Lucinoma) acutilineata  
Phacoides acutilineatus (Conrad)). [Vaqueros Fm.],
Miocene. Coalinga District, Fresno Co. USGS 4803. Hypotype USNM 165564]. [Lucinidae].
Lucina (Myrtea) acutilineata
Fm. [=Purisima Fm.], Pliocene. Near Princeton, San Mateo Co. UCMP B4791. Hypotype UCMP 37624. Remarks:
Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266). [Lucinidae].
Lucina cf. acutilineata Conrad, 1849a. Stadum 1984b. NHFOC Mem. 1:92, pl. 2   

Lucina (Lucinoma) annulata
Diego Fm., Pliocene. Southwestern San Diego Co, San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes
LACMIP 4643, 4644. Remarks: Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266). [Lucinidae].
Lucina (Lucinoma) annulata

12953]. Hypotype UCLA 59408 [=LACMIP 8321]. Remarks: Lucinoma annulatum (Reeve, 1850),  Coan et al.
(2000: p. 266). [Lucinidae].
Lucina (Lucina) annulata
Co. SDSNH 4080]. SDSNH hypotype determined here as missing. Remarks: Lucinoma annulatum (Reeve, 1850),
 Coan et al. (2000: p. 266). [Lucinidae].
Lucina (Lucinoma) annulata

CSUN 757) (=LACMIP 12953). Hypotype UCLA 59408 (=LACMIP 8321)]. Remarks: Lucinoma annulatum (Reeve,
1850),  Coan et al. (2000: p. 266). [Lucinidae].
Lucina annulatum
Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. [Hypotype LACMIP 14852]. Remarks:
Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266). [Lucinidae].
LucinaLucina bramkampi
Member), Paleocene [Eocene]. Near where Urruttia Canyon enters Salt Creek, Coalinga quad. (=Joaquin Rocks
quad.), Fresno Co. UCMP 1817. Holotype UCMP 15629. [Lucinidae].
Lucina (Epilucina) californica  
11, 16. San Diego Fm., Pliocene. West of the mouth of Rose Canyon, San Diego Co. SDSNH 80. Hypotype SDSNH
[4348]. Remarks: In the genus Epilucina,Coan et al. (2000: p. 263). [Lucinidae].
Lucina (Lucina diabloi (Dickerson, 1914a). [Phacoides
Martinez Fm., Paleocene. Mt. Diablo quad., Contra Costa Co. UCMP 340. Holotype UCMP 11681. [Lucinidae].
82 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Lucina (Myrtea) aff. -
per Gaviota Fm., Eocene. Coyote Canyon, Pt. Conception quad., Santa Barbara Co. UCMP B7027. Hypotype SU.
9284 [=CASG 70180]. Remarks: Epilucina washingtoniana (Clark, 1925),  Squires and Gring (1996: p. 66).
[Lucinidae].

Co. UCMP B6962. Hypotype SU 9283 [=CASG 70182].
Lucina (Here)
Gaviota fms., Eocene. Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6955. Holotype SU 9285. Remarks:
In the genus Here,Moore (1988: p. D13). [Lucinidae].
Lucina (Here) excavata
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes
LACMIP 4636–4638. Remarks: In the genus Here,Coan et al. (2000: p. 263). [Lucinidae].
Lucina (Here) excavata temblorensis
Miocene. Uppermost fossiliferous sandstone ledge of the Temblor Fm., Domengine Ranch quad., Fresno Co. UCMP
D1074. Holotype UCMP 36676. Remarks:Lucina (Here) excavata temblorensis Adegoke, Moore
(1988: p. D23). Species seems to be a venerid,Moore (1988: p. D23). [Lucinidae].
Lucina (Here) excavata temblorensis
Fm., Oligocene/Miocene. Uppermost fossiliferous sandstone ledge of the Temblor Fm., Domengine Ranch quad.,
Fresno Co. (Kern Co. in text) UCMP D1074. Holotype UCMP 36676. Remarks: Species seems to be a venerid,
Moore (1988: p. D23). [Lucinidae].
Lucina gaylordi (Wagner and Schilling, 1923). [Phacoides

321. Holotype CASG 6176. [Lucinidae].
Lucina (Lucinisca) menuda (Keen, 1943). [Lucinisca
Silt, Miocene. Caliente quad., Kern Co. SU 2121. Holotype SU/CASG 7526 [=CASG 72004]. [Lucinidae].
Lucina nasuta-
tion), Eocene. Martinez, Contra Costa Co. Holotype ANSP 4465 [ANSP. 4475,Richards (1968: p. 68)], missing,
 K. Estes-Smargiassi (personal communication, 2019).[Lucinidae].
Lucina (Lucinisca) nuttalli
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes
LACMIP 4639, 4640. Remarks: In the genus Lucinisca,Coan et al. (2000: p. 263). [Lucinidae].
Lucina (Lucinisca) nuttalli
Co. SDSNH 4080. Hypotype SDSNH 75917. Remarks: In the genus Lucinisca,Coan et al. 2000:263. [Lucinidae].
Lucina (Lucinisca) nuttallii nuttallii
Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes (of Hertlein and Grant, 1972, pl.
Remarks: In the genus Lucinisca,Coan et al. (2000: p. 263). [Lucinidae].
Lucina (Lucinisca) nuttalli antecedens (Arnold, 1907). [Phacoides]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):246,
9, 13, 14. San Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes
LACMIP 4641. 4642. Remarks: Junior synonym of Lucinisca nuttalli (Conrad, 1837),Coan et al. (2000: p.
263). [Lucinidae].
Lucina (Lucinisca) nuttallii antecedens (Arnold, 1907). [Phacoides
7.8, 10.11. San Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes (of Hertlein
 Remarks: Junior synonym of Lucinisca nuttalli
(Conrad, 1837),Coan et al. (2000: p. 263). [Lucinidae].
LucinaLucina quadrata (Dickerson, 1914a). [Phacoides-
tinez Fm., Paleocene. Near Lower Lake, Lake Co. UCMP 784. Holotype UCMP 11683. [Lucinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 83
Lucina richthofeni-
cene. Hills just north of Kern River Kern Co. CASG 65. Hypotype CASG 5737. Remarks: Junior synonym of Here
excavata (Carpenter, 1857),Coan et al. (2000: p. 263). [Lucinidae].
Lucina (Parvilucina) tenuisculpta intensa (Dall, 1903b). [Phacoides (Parvilucina)]. Hertlein and Grant 1972. SDSNH
Lucina (Parvilucina) intensa in pl. captions). San Diego Fm., Pliocene.
San Diego well, Balboa Park, San Diego, San Diego Co. Syntype CASG 698D, determined here as missing. Remarks:
Junior synonym of Parvilucina approximata (Dall, 1901b),Coan et al. (2000: p. 265). [Lucinidae].
Lucina (Parvilucina) intensa in pl. caption). West side of next gully east of LACMIP 305 at the
same elevation. LACMIP 305A [=LACMIP 16862]. Hypotype LAMCIP 4645.
Lucina (Parvilucina) intensa in pl. caption). Knox Ranch hill, near U.S./Mexico boundary, San
Diego Co. LACMIP 318. Hypotype LACMIP 4646.
Lucina
El Paso Mtns., Kern Co. USGS M9178. Hypotype USNM 424895. [Lucinidae].
   
Panoche Hills, Fresno Co. Hypotype current whereabouts unknown. Remarks: Now Nymphalucina panochensis,
 Kiel (2013: pp. 259, 260).
Lucinisca? brabbi Powell, 2001. (n. sp.). PaleoBios
Rico Creek, Spreckles quad., Monterey Co. SU 1957 [=CASG 68587]. Holotype CASG 68586, paratypes CASG
68587.01, 68587.02. [Lucinidae].
Lucinisca menuda-
cene. Caliente quad., Kern Co. SU 2121. Holotype SU 7526 [=CASG 72004]. Remarks: In the genus/subgenus
Lucina (Lucinisca),Moore (1988b: p. D9). [Lucinidae].
Lucinisca nuttalli (Conrad, 1837). [Lucina]. Addicott 1965b. Mi-
ocene. Comanche Pt., Tejon Hills, Kern Co. USGS M1619. Hypotype USNM 649166. [Lucinidae].
Lucinisca nuttalli (Conrad, 1837). [Lucina 
Pliocene. Balboa Park, San Diego, San Diego Co. [SDSNH 29. Hypotype SDSNH 4138]. [Lucinidae].
Lucinisca nuttalli (Conrad, 1837). [Lucina
area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14352. [Lucinidae].
Lucinisca nuttallii antecedens (Arnold, 1907). [Phacoides] Woodring and Bramlette 1950 [1951]. USGS PP 222:86,

USGS 14622. Hypotype USNM 560140. Remarks: Junior synonym of Lucinisca nuttalli (Conrad, 1837),Coan
et al. (2000: p. 263]. [Lucinidae].

Lucinisca nuttallii nuttallii (Conrad, 1837). [LucinaLucina nuttallii nuttallii). Imperial
Fm. [=Latrania Fm.], Pliocene. Near Travertine Rock, Imperial Co. (not San Diego Co. as in locality description).
USGS M9779 [=USGS M9778]. Hypotype USNM 486910. [Lucinidae].
LuciniscaLuci-
nisca
Fm. (of Dibblee 1968), Pliocene. Panorama Hills quad., Temblor Range, Kern Co. USGS M1059. Hypotype USNM
646814. [Lucinidae].
Lucinoma acutilineata (Conrad, 1849a). [Lucina        
Miocene. Reef Ridge, west side of Zapato Canyon, Kreyenhagen Hills quad., Fresno Co. USGS 14392. Hypotype
USNM 498747. Remarks: Lucinoma acutilineatum (Conrad, 1849a),  Coan et al. (2000: p. 266). [Lucinidae].
Lucinoma acutilineata (Conrad, 1849a). [Lucina        
84 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Temblor Fm. (Phacoides Sand member) (of Stinemeyer et al. 1959:12, 13), Miocene. Las Yeguas Ranch quad.,
Temblor Range, Kern Co. USGS M3280. Hypotypes USNM 646537, 646541. Remarks: Lucinoma acutilineatum
(Conrad, 1849a),  Coan et al. (2000: p. 267). [Lucinidae].
Lucinoma acutilineata (Conrad, 1849a). [Lucina

Sandstone Member), Miocene. South side of Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280.
Hypotypes USNM 646537, 646541, 646542. Remarks: Lucinoma acutilineatum (Conrad, 1849a),  Coan et
al. (2000: p. 267). [Lucinidae].
Lucinoma cf.  (Conrad, 1849a). [Lucina
Etchegoin Fm. (Pseudocardium Zone), Pliocene. North branch of Arroyo Torcido, east of Double Hill, North Dome,
Kettleman Hills, Kings Co. USGS 12400. Hypotype USNM 495789. Remarks: Lucinoma acutilineatum (Conrad,
1848),  Coan et al. (2000: p. 267). [Lucinidae].
Lucinoma cf.  (Reeve, 1850). [Lucina]. Woodring and Bramlette 1950 [1951]. USGS PP 222:86, pl. 19,
. Careaga Fm. (Graciosa Member), Pliocene. Casmalia Hills, northeast of Shuman, Santa Barbara Co. USGS
14612. Hypotype USNM 560142. Remarks: Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266).
[Lucinidae].
Lucinoma annulata (Reeve, 1850). [Lucina
(Pomponio Member), Pliocene. San Mateo Co. Locality 711M. Hypotype determined here as missing. Remarks:
Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266). [Lucinidae].
Lucinoma annulata (Reeve, 1850). [Lucina
Pliocene. Near the south base of Mt. Soledad, San Diego Co. [SDSNH 365. Hypotype SDSNH 4156]. Remarks:
Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266). [Lucinidae].
Lucinoma annulata (Reeve, 1850). [Lucina
Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotypes (of Hertlein and Grant, 1972, pl. 46,
Remarks: Lucinoma annulatum (Reeve, 1850),  Coan et al. (2000: p. 266).
[Lucinidae].
Lucinoma annulata (Reeve, 1850). [Lucina
Cruz Co. CASG 66062. Hypotype CASG 66062.02. Remarks: Lucinoma annulatum (Reeve, 1850),  Coan et al.
(2000: p. 266). [Lucinidae].
Lucinoma annulatum (Reeve, 1850). [Lucina
area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14353. [Lucinidae].
aff. LucinomaPaleoBios
Berkeley Hills, Oakland, Alameda Co. UCMP IP 13005. Hypotype UCMP 218741. [Lucinidae].
Lutrariatraskei

Lyropecten catalinae (Arnold, 1906). [Pecten 
Breccia, Miocene. Santa Catalina Is., California Channel Ids. Holotype SU/CASG 30 [=CASG 61825.01], paratype
SU/CASG 561 [=CASG 6966]. [Pectinidae].
Lyropecten catalinae (Arnold, 1906). [Pecten
Miocene. Santa Catalina Is. California Channel Ids. Holotype SU 30 [=CASG 61825.01] (as Pecten (Lyropecten)
estrellanus Conrad var. catalinae Arnold, 1906). [Pectinidae].

Catalina Is., California Channel Ids. UCMP D-8793. Hypotypes UCMP 37384–37386.
 Fm. [=Castaic Fm.,  Squires et al. (2006:13, 15], Miocene. Santa Clara Valley north of
Newhall and the Santa Susana Mtns., Mint Canyon quad., Los Angeles Co. UCMP D-8261. Hypotype UCMP 16098.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 85
Lyropecten catalinae (Arnold, 1906). [Pecten
Pliocene. Southwest of Valencia, Oat Mtn. quad. (1952 [PR 1969]), Los Angeles, Co. LACMIP 17778. Hypotypes
LACM 13346–13348. [Pectinidae].
Lyropecten catalinae (Arnold, 1906). [Pecten
area, Los Angeles Co. LACMIP 7752. Hypotype LACM 14345. [Pectinidae].
Lyropecten cerrosensis (Gabb, 1866). [Pecten
1. Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., below brachiopod bed, Santa Barbara Co. USGS 14771.
Hypotype USNM 560138. [Pectinidae].
Lyropecten cerrosensis (Gabb, 1866). [Pecten
Rico Fm., Miocene/Pliocene. Priest Valley quad., Monterey Co. UCMP A911. Hypotype UCMP 36469. [Pectinidae].
Lyropecten cerrosensis (Gabb, 1866). [Pecten
Mission Viejo area, Orange Co. Line drawing, source specimen unknown. [Pectinidae].
Lyropecten cerrosensis (Gabb, 1866). [Pecten
Miocene/Pliocene. Salinas Valley, San Ardo quad., Monterey Co. UCMP A911. Hypotype (of Durham and Addicott


U.S./Mexico boundary, southwest of Goat Canyon, Imperial Beach quad., San Diego Co. LACMIP 305-C [=LACMIP.

Lyropecten crassicardo (Conrad, 1857a). [Pallium
Fm., Miocene. Comanche Pt., Tejon Hills, Kern Co. USGS M1619. Hypotype USNM 649164. [Pectinidae].
Lyropecten crassicardo (Conrad, 1857a). [Pallium
Fm. (Buttonbed Sand Member) (of Stinemeyer et al. 1959: p. 8, 13), Miocene. Shale Pt. quad., Temblor Range,
Kern Co. USGS M3988. Hypotype USNM 646796. [Pectinidae].
       
USNM 646812.
Lyropecten crassicardo (Conrad, 1857a). [Pallium]. Vedder and Moore 1976. AAPG Misc. Pub. 24:121, 122, 130, pl.
-
type UCR 7957/1. [Pectinidae].

Hypotype UCLA 59266 [=LAMCIP 10844].
 Is., California Channel Ids. LACMIP 1194. Hypotype LACMIP 6272.
Lyropecten crassicardo (Conrad, 1857a). [Pallium    
inside back cover. Santa Margarita Fm., Miocene. Camatti Ranch, San Luis Obispo Co. USGS. M2023. Hypotype
USNM 254389. [Pectinidae].
Lyropecten crassicardo (Conrad, 1857a). [Pallium

Ranch, San Luis Obispo Co. USGS M2023. Hypotype USNM 254389. [Pectinidae].
Lyropecten crassicardo (Conrad, 1857a). [Pallium       
Margarita Fm., Miocene. Tejon Hills, Caliente quad., Kern Co. UCMP 3029. Holotype (of Pecten nomlandi Hertlein,
1931) UCMP 11308. [Pectinidae].
Pecten
vickeryi Trask 1922) SU/CASG 26 [=CASG 61833.01].
Pecten ricei Trask, 1922) UCMP 12364.
86 PALEOBIOS, VOLUME 38, FEBRUARY 2021

ANSP 30745a.
Pecten ham-
iltoni Arnold, 1906) USNM 164845.
Swiftopecten
adekunbiana Adegoke, 1969) UCMP 36639.

(of Pecten holwayi Clark, 1915) UCMP 11580.
Lyropecten crassicardo (Conrad, 1857a). [Pallium        
Fm., Miocene. Alum Rock Canyon, Santa Clara Co. Holotype and paratype (of Pecten (Lyropecten) crassicardo var.
hamiltoni Arnold, 1906) USNM 164845. [Pectinidae].
       Pecten-oyster biostrome, north of Coalinga along Domengine
Creek, Domengine Ranch quad., Fresno Co. UCMP D8791. Hypotype UCMP 37381.
  
1984:B52) ANSP 30745a.
 


determined here as missing.
....., pl. 24Pecten bed, Coalinga quad., Fresno Co. UCMP D1088. Holotype
(of Swiftopecten adekunbiana Adegoke, 1969) UCMP 36639.

Co. UCMP 3535. Holotype UCMP 12364 (of Pecten (Lyropecten) ricei Trask 1922).

(of Pecten (Lyropecten)

Is., California Channel Ids. UCLA 5899 [=LACMIP 25899]. Hypotype UCLA 59266 [=LACMIP 10844].

Is., California Channel Ids. LACMIP 1194. Hypotype LACMIP 5877.

UCMP 3029. Holotype (of Pecten crassicardo biformatis Nomland, 1917) UCMP 11308 (not 11318 as in pl. caption).
-
way, San Juan Capistrano quad., Orange Co. LACMIP 1189. Hypotype LACMIP 5876.
Pecten
(Lyropecten) vickeryi Trask, 1922) SU 26 [=CASG 61833.01].

Costa Co. UCMP 1632. Holotype (of Pecten (Pallium) holwayi Clark, 1915)). UCMP 11580.
-
istrano quad., Orange Co. UCLA 635 [=LACMIP 20635]. Hypotype UCLA 45105 [=LACMIP 11429].

Co. UCMP 1910. Hypotype UCMP 16099.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 87

Is., California Channel Ids. LACMIP 1193. Hypotype LACMIP 5875.
Lyropecten crassicarso (Conrad, 1857d) [Pallium
Monterey Fm. (Saddleback Valley Limestone member), Miocene. Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4521. Hypotypes SDSNH 98671, 98672. [Pectinidae].
Lyropecten crassicardo nomlandi (Hertlein, 1931). [Pecten]. Adegoke 1969. 
Fm. (McClure Member), Miocene. Tar Peak area, Kings Co. UCMP D1207. Hypotype UCMP 36626. Remarks: Junior
synonym of Lyropecten crassicardo (Conrad, 1857a),  Moore (1984: p. B51). [Pectinidae].
Lyropecten crassicardo vaughani (Arnold, 1906). [Pecten
6. Temblor Fm. (Carneros Sand Member) (of Stinemeyer et al. 1959: pp. 9, 13), Miocene. Temblor Range, Kern
Co. USGS 4941. Hypotype USNM 646790. Remarks: Lyropecten vaughani (Arnold, 1906),  Moore (1984: p.
B50). [Pectinidae].
Lyropecten estrellanus (Conrad, 1857a). [Pallium   
Pancho Rico Fm., Miocene/Pliocene. Adelaida quad., Monterey Co. USGS M2050. Hypotype USNM 649085. Re-
marks: 
Lyropecten estrellanus (Conrad, 1857a). [Pallium
Miocene. Domengine Creek, Polvadero Gap quad., Kings Co. UCMP B4350. Hypotype UCMP. 36627. Remarks:

Lyropecten estrellanus (Conrad, 1857a). [Pallium
Re-
marks: 
Lyropecten estrellanus (Conrad, 1857a). [Pallium 
Margarita Fm., Miocene. San Juan River area, San Luis Obispo Co. USGS 4944. Hypotype USNM 647533 (not
holotype USNM 13317 as in text). Remarks: -
lustrated in Conrad (1857c). [Pectinidae].
Lyropecten estrellanus (Conrad, 1857a). [Pallium  
Fm., Miocene/Pliocene. Northeast of San Lucas, Monterey Co. USGS M903. Hypotype USNM 254349. Remarks:

Lyropecten estrellanus (Conrad, 1857a). [Pallium
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2872. Hypotype USNM
254378. Remarks: 
[Pectinidae].
Lyropecten estrellanus (Conrad, 1857a). [Pallium-

San Luis Obispo Co. USGS M2872. Hypotype USNM 254378. Remarks: Supplemental descriptions in Conrad

Lyropecten estrellanus (Conrad, 1857a). [Pallium
Margarita Fm. (TamiosomaRemarks: Supplemental descriptions

Lyropecten-
Pecten estrellanus Conrad 1857a)). Santa Margarita Fm., Miocene.
Tamiosoma Zone above Big Blue Fm., Fresno Co. [Hypotype USNM 165616]. [Pectinidae].
Lyropecten estrellanus (Conrad, 1857a). [Pallium-
garita Fm., Miocene. Estrella Valley, San Luis Obispo Co. Lectotype of Woodring 1938:33; neoholotype (invalid
term) of Tucker-Rowland 1938: p. 4) USNM 13317. Remarks: Supplemental descriptions in Conrad (1857c,

88 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Lyropecten estrellanus (Conrad, 1857a). [Pallium

Co. USGS 4944. Hypotype USNM 647533. Remarks: Supplemental descriptions in Conrad (1857c, 1857d) and


60983.


Lyropecten estrellanus (Conrad, 1857a). [Pallium]. Powell 2007. PaleoBios
Fm., Miocene. Cosio Knob quad., Monterey Co. UCMP A8968. Hypotype UCMP 54535. Remarks: Supplemental

Lyropecten estrellanus (Conrad, 1857a). [Pallium]. Powell et al., 2020. PaleoBios
Fm., Miocene. Pancho Rico Canyon, Monterey Co. CASG 78559. Hypotype CASG 78560. Remarks: Supplemental

Lyropecten magnolia (Conrad, 1857d). [Pecten
Miocene. Coffee Canyon, Kern Co. UCR 1185. Hypotype UCR 1185/6 (not UCR 1185-b as in pl. caption). Remarks:
In the genus Macroclamis,  Moore (1984: p. B56). [Pectinidae].
Lyropecten magnolia (Conrad, 1857d). [Pecten

Ventura Co. UCMP A326. Hypotype UCMP 31734. Remarks: In the genus Macroclamis,  Moore (1984: p. B56).
[Pectinidae].
....., pl. 2, fgs. 4, 5. Cuyama area, Caliente Range, sec. 11, T10N, R25W, Cuyama quad., San Luis Obipo Co. USGS M3520.
 J.T. Smith
(personal communication, 1999).
Remarks: Not Lyropecten magnoliaJ.T. Smith
(personal communication, 1999).
Lyropecten magnolia (Conrad, 1857d). [Pecten 
Santa Barbara Co. Hypotype determined here as missing. Remarks: In the genus Macroclamis,  Moore (1984:
p. B56). [Pectinidae].
Lyropecten miguelensis (Arnold, 1906). [Pecten
Fm. (Carneros Sand Member) (of Stinemeyer et al.1959: pp. 9, 13), Miocene. Carneros Rocks quad., Temblor
Range, Kern Co. USGS M3981. Hypotype USNM 646789. [Pectinidae].
Lyropecten miguelensis (Arnold, 1906). [Pecten
Fm., Oligocene/Miocene. San Miguel Is., California Channel Ids. Holotype UCMP 12079. [Pectinidae].
Lyropecten miguelensis (Arnold, 1906). [Pecten]. Smith 1991a. USGS PP 1391:61
(of Hill et al. 1958: pp. 2991–2993), Miocene. West of Santa Barbara Canyon, Fox Mtn. quad., Santa Barbara Co.
USGS M2424. Hypotype USNM 335952. [Pectinidae].
 Avila and Weaver 1969:49–51), Miocene. Southeast coast, west of East Pt., Santa Rosa
Is. East quad., Santa Rosa Isd., California Channel Ids. UCSB SR67-28. Hypotype UCSB 4675 [=SBMNH 60212].
-
nel Ids. Hypotype UCSB 1844 [=SBNHM 80167].
-
potype (of Pecten miguelensis
[=LACMIP 6300].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 89
-
nel Ids. CASG 1156. Hypotype (of Pecten (Lyropecten) miguelensis
CASG 4143.

  
Hypotype UCSB 1912 [=SBMNH 60213].
Lyropecten miguelensis (Arnold, 1906). [Pecten]. Shapiro 1998. AAPG Guidebook MP 45:129, pl. 1 (pl. 2 in caption),

6527 [=SBMNH 60178, 60179]. [Pectinidae].
Lyropecten pretiosus (Hertlein, 1925b). [Pecten
(Diorite breccia member), Oligocene/Miocene. Cañada Posa, southwestern Santa Cruz Is., California Channel Ids.
USGS M8022. Hypotypes USNM 335000, 335001. [Pectinidae].

California Channel Ids. UCMP A311. Holotype UCMP 31737 (of Pecten (Lyropecten) miguelensis submiguelensis
Loel and Corey, 1932).

Ids. UCSB SR67-53 [=LACMIP 16404]. Hypotype (of Pecten miguelensis Arnold 1906 of Avila and Weaver 1969,

Lyropecten submiguelensis (Loel and Corey, 1932). [Pecten

31737. [Pectinidae].
Lyropecten submiguelensis (Loel and Corey, 1932). [Pecten

Santa Rosa Is., California Channel Is. UCMP A311. Holotype UCMP 31737. [Pectinidae].
Lyropecten terminus (Arnold, 1906). [Pecten]. Durham and Addicott 1965. USGS PP 524-A:A13, pl
Pancho Rico Fm., Miocene/Pliocene. Walker Canyon, Wunpost quad., Monterey Co. USGS M1935. Hypotypes
USNM 649083, 649084. [Pectinidae].


Lyropecten terminus (Arnold, 1906). [Pecten

Hypotype USNM 649088. [Pectinidae].
Lyropecten terminus (Arnold, 1906). [Pecten
Pliocene. Terraces on east side of Salinas Valley between King City and San Lucas, Monterey Co.,  Durham
and Addicott (1965: p. A15). Holotype UCMP 11662. [Pectinidae].
Lyropecten terminus (Arnold, 1906). [Pecten
Fm., Miocene/Pliocene. Cholame Hills, east side Salinas Valley, Monterey Co. Holotype UCMP 11622. [Pectinidae].

UCMP 37391.
Lyropecten terrysmithae Powell et al., 2020 (n.sp). PaleoBios-
cene. Cholame Hills, between Indian Valley and Portuguese Canyon, Stockdale Mtn. quad., Monterey Co. CASG
28473. Holotype CASG 5900, paratypes CASG 78557, 78558.
       
90 PALEOBIOS, VOLUME 38, FEBRUARY 2021
CASG 78554–78556, 78563.
Lyropecten tiburonensis
[=Latrania Fm.], Miocene. Carrizo Mtn. quad., Imperial Co. UCR 7267. Hypotype UCR 7267/3. [Pectinidae].

Co. UCR 5042. Hypotype UCR 5042/4.

quad., Imperial Co. [UCMP 738]. Hypotype (of Pecten subnodosus G.B. Sowerby I, 1835 of G.D. Hanna 1926, pl.

Lyropecten vaughani (Arnold, 1906). [Pecten
Fm., Oligocene/Miocene. Ojai Valley, Ventura Co. Holotype SU/CASG 9 [=CASG 61858.01]. [Pectinidae].
Lyropecten vaughani (Arnold, 1906). [Pecten]. Smith    
Fm., Oligocene/Miocene. Ojai Valley (exact locality unknown), Ventura Co. Holotype SU 9 [=CASG 61858.01].
[Pectinidae].
Lyropecten   
Miocene/Pliocene. Barrett Canyon, Coyote Mtns., Imperial Co. ABDSP 4121. Hypotype ABDSP I 14396. Remarks:
Specimen on exhibit at the Anza-Borrego Desert State Park visitor center,  L. Gilbert (personal communica-
tion, 2019). [Pectinidae].
Macoma (Macoma) acolasta
San Diego Fm., Pliocene. Balboa Park, San Diego, San Diego Co. UCLA 1386 [=LACMIP 21386]. Hypotypes UCLA
[48626, 48627 [=LACMIP 9309, 9310]. Remarks: In the subgenus Psammacoma,  Coan et al. (2000: p. 414).
[Tellinidae].
Macoma addicotti
Alto quad., Santa Clara Co. CASG 44020. Holotype CASG 55956 [=CASG 44020.01]. Remarks: Possibly Macoma
nasuta (Conrad, 1837),  Coan et al. (2000). [Tellinidae].

241063 as in text).
Macoma addicotti
CASG 66095. Hypotype CASG 66095.01]. Remarks: Possibly Macoma nasuta (Conrad, 1837),  Coan et al.
(2000: p. 420). [Tellinidae].
Macoma (Rexithaerus addicotti Nikas, 1977. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

[=CASG 44020.01]. Remarks: Possibly Macoma nasuta (Conrad, 1837),  Coan et al. (2000: p. 420). [Tellinidae].
        
Fm. (Upper Mya Zone), Pliocene. Las Alturas, Middle Dome, Kettleman Hills, Kings Co. USGS 12697. Hypotypes
USNM 495301–495303. [Tellinidae].
Macoma Zone), Pliocene. South slope of east end of El Serrijon, North Dome,
Kettleman Hills, Kings Co. USGS 12424. Hypotypes USNM 495830–495833.

Fm., Miocene/Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M979. Hypotype USNM 649092.
Remarks: Macoma (Macoma Nomland, 1917,  Moore (2003b: p. 37). [Tellinidae].
 Nomland 1917a, small var. cf.  Stewart in Woodring et al. 1940 [1941]. USGS PP 195:93,
Mya layer below Pecten Zone, between Pecten and Neverita Zones), Pliocene.
West of Arroyo Raso, North Dome, Kettleman Hills, Kings Co. USGS 12380. Hypotype USNM 495766. [Tellinidae].
Macoma  Stewart in Woodring, et al., 1940 [1941]). Adegoke 
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 91
Fm., Pliocene. Reef Ridge quad., Kings Co. UCMP D1193. Hypotype UCMP 36761, determined here as missing.
[Tellinidae].
Macoma cf.  Stewart in Woodring et al., 1940 [1941]. Addicott 1972b. SEPM Symposium vol., p. 15,

USGS M1059. Hypotype USNM 646815. [Tellinidae].
Macoma (Macoma Stewart in Woodring et al., 1940 [1941]. Moore 2003b. USGS PP 1228-G [electronic
Littorina Zone), Miocene/ Pliocene. North Dome
of] Kettleman Hills, [Kings Co.] [USGS 14096a]. Holotype USNM. 495853. [Tellinidae].
Tellinid Macoma albaria
San Juan Capistrano, Orange Co. Monterey Fm., Miocene. [Hypotype at CC]. [Tellinidae].
Macoma (Rexithaerus andersoni Clark, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

11557. [Tellinidae].
Macoma arctata (Conrad, 1849a). [Tellina      
Northeast of Irvington, La Costa Valley quad. (1943), Alameda Co. SU 3248. Hypotype SU 8465 [ CASG 69939].
Remarks: Provisionally in the subgenus Psammacoma Moore (2003b: p. 39). [Tellinidae].
Macoma arctata (Conrad, 1849a). [Tellina
(Phacoides Sand member) (of Stinemeyer et al. 1959: pp. 12, 13), Miocene. Carneros Rocks quad., Temblor Range,
Kern Co. USGS M3578. Hypotype USNM 646785. Remarks: Provisionally in the subgenus Psammacoma
Moore (2003b: p. 39). [Tellinidae].
Macoma arctata (Conrad, 1849a). [Tellina
Sandstone Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotypes USNM
646571, 646572. Remarks: Provisionally in the subgenus Psammacoma Moore (2003b: p. 39). [Tellinidae].
Macoma arctata (Conrad, 1849a). [Tellina  
Fm. (Hames Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M2044. Hypotype
USNM 254350. Remarks: Provisionally in the subgenus Psammacoma Moore (2003b: p. 39). [Tellinidae].

USGS M3814. Hypotype USNM 254365.
Macoma arctata (Conrad, 1849a). [Tellina
    
Range, San Luis Obispo Co. USGS M2044. Hypotype USNM 254350. Remarks: Provisionally in the subgenus
Psammacoma Moore (2003b: p. 39). [Tellinidae].
....., pl. 1
Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M3814. Hypotype USNM. 254365.
Macoma arctata (Conrad, 1849a). [Tellina        
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 338 [=LACMIP 16277]. Hypotype
UCLA 58214 [=LACMIP 10595]. Remarks: Provisionally in the subgenus Psammacoma Moore (2003b: p.
39). [Tellinidae].
Macoma astori-
boldt Co. UCMP B7925. Hypotype UCMP 37781. [Tellinidae].
Macoma brota Dall var. liparia
(Tahana Member), Pliocene. Pomponio Creek, San Gregorio quad., San Mateo Co. locality 470-1M. Hypotype
determined here as missing. Remarks: Macoma (Macoma) liparia Dall, 1916b,  Coan et al. (2000: p. 410).
[Tellinidae].
Macoma brota lipara
92 PALEOBIOS, VOLUME 38, FEBRUARY 2021
(reworked from Purisima and deposited in late Pleistocene sediments). Near Santa Cruz Pt., Santa Cruz quad.
(1954), Santa Cruz Co. USGS M1691. Hypotype USNM 649312. Remarks: Macoma (Macoma) liparia Dall, 1916b,
 Coan et al. (2000: p. 410). [Tellinidae].
Macoma calcarea (Gmelin, 1791). [Tellina
(San Gregorio Member), Pliocene. San Gregorio quad., San Mateo Co. Locality 672M. Hypotype determined here
as missing. Remarks: Junior synonym of Macoma (Macoma) elimata Dunnill and Coan, 1968,  Hertlein and
Grant (1972: p. 291). [Tellinidae].
Macoma calcarea (Gmelin 1791). [Tellina      
Fortuna quad., Humboldt Co. UCMP B7875. Hypotype UCMP 37782. Remarks: Junior synonym of Macoma (Ma-
coma) elimata Dunnill and Coan, 1968,  Hertlein and Grant 1972: p. 291. [Tellinidae].
Macoma copelandi
Sand Member) (of Stinemeyer et al.1959:8, 13), Miocene. Shale Pt. quad., Temblor Range, Kern Co. USGS M3987.
Hypotype USNM 646799. Remarks: In the subgenus Rexithaerus,  Moore (2003b: p. 40). [Tellinidae].
Macoma (Rexithaerus) copelandi Wiedey, 1928. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-
      
Holotype SDSNH 37. [Tellinidae].
Macoma (Rexithaerus diabloensis Clark, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Holotype UCMP 11560. [Tellinidae].
Macoma (Macoma) elimata Dunnill and Coan, 1968. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):291–292, pl.

Hypotype UCLA [48628 [=LACMIP 9311]. [Tellinidae].
Macoma cf.  Carpenter, 1864b. Woodring et al. 1940 [1941]. USGS PP 195:92, pl. 35, 
Fm. (Patinopecten Zone), Pliocene. Near Arroyo Doblegado, southeast of Los Paredes, North Dome, Kettleman
Hills, Kings Co. USGS 12426. Hypotype USNM 495838. Remarks: In the subgenus Rexithaerus,  Coan et al.
(2000: p. 415). [Tellinidae].
Macoma (Rexithaerus) indentata
3. San Diego Fm., Pliocene. Near the U.S./Mexico boundary, San Diego Co. LACMIP 319. Hypotype LACMIP 4785.
[Tellinidae].

LACMIP 4786.
Macoma aff.  Carpenter, 1864b. Powell et al. 2019a. USGS SIR 2019-5021:3 (as Macoma aff.
Macoma aff. 
Sonoma Co., CASG 54135, Hypotype CASG 78411. [Tellinidae].
Macoma aff.  Carpenter subsp. tenuirostris
Rio Dell Fm., Pliocene. Scotia, Humboldt Co. UCMP A4539. Hypotype UCMP 37784. Remarks: Junior synonym
of Macoma (Rexithaerus) indentata Carpenter, 1864b,  Coan et al. (2000: p. 415). [Tellinidae].
Macoma (Rexithaerus) indentata tenuirostris Dall, 1900b. Hertlein and Grant 1972. SDSNH Mem. 2(2B):297, pl. 41,
     
305. Hypotype LACMIP 4787. Remarks: Junior synonym of Macoma (Rexithaerus) indentata Carpenter, 1864b,
 Coan et al. (2000: p. 415). [Tellinidae].
  sic] River plain, San Diego Co. SDSNH 331.
Hypotype SDSNH 4357.

Macoma inquinata (Deshayes, 1855) subsp. arnheimi
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 93
2. Rio Dell Fm., Pliocene. Fortuna quad., Humboldt Co. UCMP B7875. Hypotype UCMP 37780. Remarks: Junior
synonym of Macoma inquinata (Deshayes, 1855),  Coan et al. (2000: p. 419). [Tellinidae].
Macoma (Macoma) inquinata (Deshayes, 1855) [Tellina]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):292–293,

[Tellinidae].
Macoma inquinata (Deshayes, 1855). [Tellina
Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D5924. Hypotype USNM 251798. [Tellinidae].
Macoma (Macoma Nomland, 1917a. Moore 2003b. USGS PP 1228-G [electronic format] (family
 Fm. (Mya japonica Zone), Miocene/Pliocene. Garza Creek area, Kings Co.
UCMP 2965. Holotype UCMP 11092. [Tellinidae].
Macoma lorenzoensis (Arnold, 1908). Moore 2003b. USGS PP 1228-G [electronic format] (family

Santa Cruz Co. Holotype USNM 165439. [Tellinidae].
Macoma (Macoploma) medioamericana
6, 8. San Diego Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 478]. [Tellinidae].
Macoma (Macoploma) medioamericana Olsson, 1942. Moore 2003b. USGS PP 1228-G [electronic format] (family


Macoma (Macoma menkeni G.D. Hanna and Hertlein, 1938. Moore 2003b. USGS PP 1228-G [electronic format]

7, at depth of 1223 to 1240 ft, Kern Co. CASG. 28682. Holotype CASG 6900. [Tellinidae].
Macoma cf.  (Conrad, 1837). [Tellina 
Joaquin Fm. (Pecten Zone), Pliocene. Vicinity of Arroyo Vadosa and Arroyo Largo, North Dome Kettleman Hills,
Kings Co. USGS 12331. Hypotype USNM 495748. [Tellinidae].
Neverita Zone), Pliocene. South of Arroyo Doblegado, North Dome, Kettleman
Hills, Kings Co. USGS 12635. Hypotype USNM 495327.
Macoma nasuta (Conrad, 1837). [Tellina

Hypotype USNM 165515. [Tellinidae].
Macoma nasuta (Conrad, 1837). [Tellina-
ing, source specimen unknown. [Tellinidae].
Macoma nasuta (Conrad, 1837). [Tellina 
Pleasanton, Niles quad. (1943), Alameda Co. SU 3232. Hypotype SU 8466 [=CASG 69938]. [Tellinidae].
Macoma nasuta (Conrad, 1837). [Tellina
(Pomponio Member), Pliocene. Purisima Creek, Half Moon Bay quad., San Mateo Co. Locality 631M. Hypotype
determined here as missing. [Tellinidae].
Macoma (Macoma) nasuta (Conrad, 1837). [Tellina]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):293–294, pl.

72539]. [Tellinidae].
Macoma nasuta (Conrad, 1837). [Tellina]. Bedrossian 1974. Calif. Geol. 27(8):178, 
Pliocene [Wilson Grove Fm., Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotype CASG determined here
as missing. [Tellinidae].
Macoma nasuta (Conrad, 1837). [Tellina     
Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D3688, Hypotype USNM 251799. [Tellinidae].
94 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Macoma nasuta (Conrad, 1837). [Tellina-
ity of Capitola, Santa Cruz Co. CASG 66137. Hypotype CASG 66137.01]. [Tellinidae].
Macoma nasuta (Conrad, 1837). [Tellina
area, Los Angeles Co. LACMIP 17916. Hypotype LACMIP 14365. [Tellinidae].
Macoma (Rexithaerus nasuta jacalitosana Arnold, 1909 [1910]. Moore 2003b. USGS PP 1228-G [electronic format]
Macoma (Rexithaerus jacalitosana). Etchegoin Fm., Miocene/Pliocene.
Jasper Creek just above Jacalitos Creek, Coalinga quad., Fresno Co. USGS 4765. Holotype USNM 156613. Remarks:
Junior synonym of Macoma nasuta (Conrad, 1837),  Coan et al. (2000: p. 420). [Tellinidae].
Macoma nasuta kelseyi
Fm. (Graciosa Member), Pliocene. Graciosa Ridge, Santa Barbara Co. USGS 14614. Hypotype USNM 560143. Re-
marks: Valid subspecies provisionally in the subgenus Rexithaerus Moore (2003b: p. 47); junior synonym
of Macoma nasuta (Conrad, 1837),  Coan et al. (2000: p. 420). [Tellinidae].
Macoma nasuta forma kelseyi
Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650984. Remarks: Valid subspecies provisionally
in the subgenus Rexithaerus Moore (2003b: p. 47); junior synonym of Macoma nasuta (Conrad, 1837), 
Coan et al. (2000: p. 420). [Tellinidae].
Macoma (Macoma) nasuta kelseyi
San Diego, Pliocene. Balboa Park, San Diego, San Diego Co. SDSNH 29. Hypotype SDSNH 4354. Remarks: Valid
subspecies provisionally in the subgenus Rexithaerus Moore (2003b: p. 47); junior synonym of Macoma
nasuta (Conrad, 1837),  Coan et al. (2000: p. 420). [Tellinidae].


Macoma nasuta kelseyi
Vinicty of Capitola City Beach Soquel quad., Santa Cruz Co. SCCM. 39. Hypotype SCCM 21013]. Remarks: Valid
subspecies provisionally in the subgenus Rexithaerus Moore (2003b: p. 47); junior synonym of Macoma
nasuta (Conrad, 1837),  Coan et al. (2000: p. 420). [Tellinidae].
Macoma (Macoma) nasuta kelseyi
Orange Co. SDHNM 4080. SDSNH hypotype determined here as missing. Remarks: Valid subspecies provision-
ally in the subgenus Rexithaerus Moore (2003b: p. 47); junior synonym of Macoma nasuta (Conrad, 1837),
 Coan et al. (2000: p. 420). [Tellinidae].
Macoma (Rexithaerus nasuta kelseyi Dall, 1900c. Moore 2003b. USGS PP 1228-G [electronic format] (family

  Remarks:
Junior synonym of Macoma nasuta (Conrad, 1837),  Coan et al. (2000: p. 420). [Tellinidae].
Macoma oldroydi (Wiedey, 1928). [Tellina
Creek Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP22645 (=UCLA 2645). Hypotype
LACMIP 13082 [=UCLA 58009]. Remarks: In the genus Tellina and provisionally in the subgenus Eurytellina
Moore (2003b: p. 14) [Tellinidae].
Macoma pabloensis Clark, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-
            
Holotype UCMP 11563. [Tellinidae].
Macoma (Rexithaerus) panzana Wiedey, 1928. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Co. SDSNH 432. Holotype SDSNH 38. [Tellinidae].
Macoma pittsburgensis (Clark, 1925). [Tellina      
Tumey Fm., Eocene/Oligocene. Monocline Ridge, Fresno Co. UCMP D6542. Hypotype UCMP 37167. [Tellinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 95
Macoma pittsburgensis (Clark, 1925). [Tellina     
Oligocene. T16S, R13E, Monocline Ridge quad., Fresno Co. UCMPD6542. Hypotype UCMP 37167. [Tellinidae].
Macoma rosa
(1951, PR 1969), Ventura Co. CSUN 489 [=LACMIP 16103]. Hypotype LACMIP 6586. Remarks: Provisionally in
the subgenus Rexithaerus Moore (2003b: p. 44). [Tellinidae].
Macoma rosa
South side of Simi Valley, Calabasas quad. (1952, PR 1967), Ventura Co. CSUN 1342 [=LACMIP 17187]. Hypotype
LACMIP 12700. Remarks: Provisionally in the subgenus Rexithaerus Moore (2003b: p. 44). [Tellinidae].
Macoma rosa M.A. Hanna, 1927. Ekdale and Ekdale. 2018. Paleo3
near Interstate 5/805 merge, San Diego, San Diego Co. SDSHN 5581. Hypotype SDSNH 112104. (Tellinidae].
Macoma (Rexithaerus rosa M.A. Hanna, 1927. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-
 Jolla, La Jolla quad., San
Diego Co. UCMP 3993. Holotype UCMP 31094. [Tellinidae].
Macoma cf.  (Conrad, 1837). [Tellina
Fm. (Pecten Zone), Pliocene. South end of El Caballete, Middle Dome, Kettleman Hills, Kings Co. USGS 12787.
Hypotype USNM 495776. Remarks: In the subgenus Rexithaerus,  Coan et al. (2000: p. 417). [Tellinidae].
Macoma secta (Conrad, 1837). [Tellina
Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650975. Remarks: In the subgenus Rexithaerus,
 Coan et al. (2000: p. 417). [Tellinidae].
Macoma (Rexithaerus) secta (Conrad, 1837). [Tellina
Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14364. [Tellinidae].
Macoma sespeensis
Concord quad., Contra Costa Co. UCMP 516. Hypotype UCMP 34138. Remarks: Heteromacoma rostellata (Clark,
1918),  Addicott (1973b: p. 34). [Tellinidae].
-
totype UCMP 34139. Remarks: Heteromacoma rostellata (Clark, 1918),  Addicott 1973b: p. 34. [Tellinidae].
Macoma (Rexithaerus sespeensis Loel and Corey, 1932. Moore 2003b. USGS PP 1228-G[electronic format] (family

Sespe Creek, Santa Clara Valley, Ventura Co. UCMP A322. Holotype UCMP 31840. [Tellinidae].
Macoma (Macoma sheridani Vokes, 1939. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):34,

UCMP 15703. [Tellinidae].
Macoma (Rexithaerus vanvlecki Arnold, 1909 [1910]. Moore 2003b. USGS PP 1228-G [electronic format] (family
    
USNM 165576. [Tellinidae].
Macoma viticola
Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4726. Hypotype UCR 4726/12. [Tellinidae].
Macomaviticola Anderson and G D. Hanna, 1925. Moore 2003b. USGS PP 1228-G [electronic format]

[Tellinidae].
Macoma wilcoxi Hall and Ambrose, 1916. Moore 2003b. USGS PP 1228-G [electronic format] (fam-

Alameda Co. Holotype SU 506 [=CASG 69805]. [Tellinidae].
Macoma (Conrad, 1837). [Tellina]. Addicott 1969. Pr.CAS 37(3):63, 66, 70, 72, 73, pl.
96 PALEOBIOS, VOLUME 38, FEBRUARY 2021
M1715. Hypotype USNM
650970. Remarks: Macoma addicotti n. sp.,  Nikas (1977: pp. 434–437); in the subgenus Rexithaerus
Moore (2003b: p. 46). [Tellinidae].
MacomaTrophon Zone”),


Macoma
strata), Miocene. Sisquoc Ranch, Santa Barbara Co. USGS 14963. Hypotype USNM 560038. [Tellinidae].
Macoma
and Elizabeth Lake canyons, Los Angeles Co. CIT 277 [=LACMIP 7776]. Hypotypes LACMIP 4127, 4128. [Tellinidae].
Macoma-
cene [=Wilson Grove Fm.], Miocene/Pliocene. Santa Rosa Valley, Sonoma Co. Hypotype CASG, determined here
as missing. [Tellinidae].
Macoma

Remarks:Chione sp., Clinocardium
sp. and Protothaca sp. [=Leukoma sp.]. [Tellinidae].
Macoma
Co., SDSNH 6871. Hypotype SDSNH 139003. [Tellinidae].
Macoma sp. indeterminate. Powell et al. 2019a. USGS SIR 2019-5021:3, 31, pl. 3, . 2, 3. Wilson Grove Fm., Mio-
cene. North of , Sonoma Co., CASG 54135, Hypotype CASG 78412 [Tellinidae].
Macrocallista andersoni  

32977 [not in UCMP online catalog], UCMP paratype missing,  D. Haasl (personal communication, 2006).
[Veneridae].

B6934. Hypotype SU 9296 [=CASG 70183].
Macrocallista conradiana (Gabb, 1864). [Tapes
Tejon Fm., Eocene. Live Oak Creek, Kern Co. CASG 244. Hypotype CASG 5623. [Veneridae].
Macrocallista furlongi       
Paleocene. Simi Hills, Ventura Co. UCR6683. Hypotype UCR 6683/20. [Veneridae].
Macrocallista hornii (Gabb, 1864). [Meretrix
Sacate/Gaviota fms., Eocene. Nojoqui Creek area, Los Olivos quad., Santa Barbara Co. UCMP B6933. Hypotype SU
9297 [=CASG 70184]. [Veneridae].
Santa Barbara Co. UCMP 3170. Hypotype SU 9298[=CASG 70185].
Macrocallista horni (Gabb, 1864). [Meretrix
[= possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH 2356. Hypotype SDSNH 4150]. [Veneridae].
Macrocallista cf.  (Linneaus, 1758). [Venus-
terey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, Palos Verdes Hills, Los
Angeles Co. USGS 13127. Hypotype USNM 497091. [Veneridae].
Macrocallista pittsburgensis (Dall, 1900c). [Callista       

5667. [Veneridae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 97
Macrocallista pittsburgensis (Dall, 1900c). [Callista
Tumey Fm., Eocene/Oligocene. Monocline Ridge, Fresno Co. UCMP D6546. Hypotype UCMP 14512]. [Veneridae].
Macrocallista pittsburgensis (Dall, 1900c). [Callista
Oligocene. Monocline Ridge, Monocline Ridge quad., Fresno Co. Hypotype UCMP 37163. [Veneridae].

Macrocallista stantoni

Macrocallista stantoni-
cene. Simi Hills, Ventura Co. [UCR 6684]. Hypotype UCR 6684/22 (not CASG 4928 as in pl. caption). [Veneridae].
Macrocallista

Macrochlamis magnolia (Conrad, 1857d). [Pecten
Fm. (Agua Sand Member) (of Stinemeyer et al. 1959: pp. 9, 13), Miocene. Packwood Creek quad., Temblor Range,
Kern Co. USGS M2631. Hypotype USNM 646792. [Pectinidae].
Macrochlamis magnolia (Conrad, 1857d). [Pecten
(Pyramid Hill Sand Member), Miocene. Kern River area, Kern Co. UCR 1185. Hypotype UCR 1185/6 (not 1185-b
as in pl. caption). Remarks:Macroclamis magnolia ojaiensis n. subsp.,  Smith (1991a: p. 74). [Pectinidae].
Macrochlamis magnolia (Conrad, 1857d). [Pecten]. Squires and Filewicz (eds.) 1983. SEPM Guidebook 35:182, un-
Big Mtn. Oil Field, Santa Susana Mtns., Ventura Co. LACMIP17098.
Hypotype LACMIP. 12721. [Pectinidae].
Macrochlamis magnolia (Conrad, 1857d). [Pecten
Fm., Oligocene/Miocene. Santa Inez [sic] Mtns., Santa Barbara Co. Lectotype (of Keen and Bentson 1944: p. 89)
USNM 13325. [Pectinidae].

as cited in text). Holotype (of Pecten (Lyropecten) vaughani emigdioensis Wagner and Schilling 1923) UCMP
11417. [Pectinidae].
Macrochlamis magnolia (Conrad, 1857d). [Pecten

north side Simi Valley, Ventura Co. LACMIP17098. Hypotype LACMIP 12721. [Pectinidae].
Macrochlamis magnolia magnolia (Conrad, 1857d). [Pecten
Lyropecten magnolia)). Vaqueros
Fm., Oligocene/Miocene. Caliente Range, Salisbury Canyon quad., San Luis Obispo Co. USGS M3520. Hypotypes
USNM 335003, 647534. [Pectinidae].

quad., Santa Barbara Co. UCMP D8794. Hypotype UCMP 37387.

Remarks: Hypotype lost and never assigned a LACMIP type number.

3A. Hypotype CASG 61039.
Macrochlamis magnolia ojaiensis       
(Pyramid Hill Sand Member), Miocene. North of Poso Creek-Coffee Creek, Pine Mtn. quad., Kern Co. UCR 1185.
Hypotype (of Macrochlamis magnolia

98 PALEOBIOS, VOLUME 38, FEBRUARY 2021
of mouth of Lion Canyon, Matilija quad., Ventura Co. UCMP A326. Hypotype (of Pecten (Lyropecten) magnolia

ex SU 22363].

Hypotype UCLA 59265 [=LACMIP 11428].
ex SU 22366].
?“Macrochlamis magnolia ojaiensis Smith, 1991a (n.subsp.) USGS PP
Oligocene. San Emigdio Mtns., Kern Co. UCMP 3234. Holotype (of Pecten (Lyropecten) vaughani emigdioensis

Mactracoalingensis
(Siphonalia Zone), Pliocene. South side of El Tolete, North Dome, Kettleman Hills, Kings Co. USGS 14104. Hypo-
type USNM 495821. Remarks: Provisionally in the genus Mactromeris Moore (2003a: p. 30). [Mactridae].
Mactra (Harvella) elegans G.B. Sowerby I, 1825” of Yates (1903). Moore 2003a. USGS 1228-F [electronic format]

CASG. 66543. Hypotype CASG 2828 [=CASG 66543.01) Remarks:
Chione temblorensis var.,  Keen and Bentson (1944: p. 64). [Mactridae].
Mactra sectoris
Ridge quad., Kings Co. UCMP D1060. Hypotype UCMP 36733. Remarks: Provisionally in the genus Mactromeris
 Moore (2003a: p. 27). [Mactridae].
Mactromeris abbotti (Wiedey, 1928). [Spisula]. Moore 2003a. USGS PP 128-F [electronic format] (family Mactri-

SU 440 [=SDSNH 440]. Holotype SDSNH. [Mactridae].
Mactromeris acutirostrata (Packard, 1916). [Spisula]. Moore 2003a. USGS 1228-F [electronic format] (family Mac-

463. Holotype UCMP 33377. [Mactridae].
Mactromerisaequilateralis (Waring, 1917). [Spisula]. Moore 2003a. USGS 1228-F [electronic format] (family Mac-

quad., Ventura Co. CASG. 61927. Holotype SU/CASG 154 [=CASG 61927.01]. [Mactridae].
Mactromeris albaria albaria (Packard, 1916). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic format] (family
   

Mactromeris albaria selbyensis (Packard, 1916). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family

Holotype UCMP 11483. [Mactridae].


Mactromeris (Mactromeris beali (Hall and Ambrose, 1916). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic
Mactromeris beali in pl. caption]. Monterey Fm. (as Jewett Sand in
pl. caption), Miocene. Pleasanton quad., Alameda Co. [CASG 66544]. Holotype SU/CASG 507 [=CASG 66544.01].
[Mactridae].
Mactromeris bisculpturata (Anderson and G D. Hanna, 1925). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic
format] (family Mactridae):9, pl Co. CASG 244. Holotype CASG
794 [=CASG 244.04]. [Mactridae].
Mactromeris brevirostrata (Packard, 1916). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 99
-
boldt Co. UCMP 1875. Holotype UCMP 11489. [Mactridae].
Mactromeris catilliformis lenticularis (Gabb, 1866). [Hemimactra]. Moore 2003a. USGS PP 228-F [electronic format]

Stewart 1930:208, 209) ANSP 4549. [Mactridae].
Mactromeris capayana (Vokes, 1939). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family Mactri-
sic] Fm. in pl. caption, Cerros (as Caerros [sic] in pl. caption) Shale
Member), Paleocene [Eocene]. Vicinity of where Urruttia Canyon enters Salt Creek, Coalinga quad., Fresno Co.
UCMP 1817. Syntypes UCMP 15714, 15715 (syntype UCMP 15715 (not UCMP 11715 as in pl. caption)). Remarks:
Species seems to be a tellinid,  Moore (2003a: p. 8). [Mactridae].
Mactromeriscoalingensis (Arnold, 1910). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic format] (family

Waltham Creek Valley, south of Coalinga, Fresno Co. Holotype USNM 165513. [Mactridae].
Mactromerisgaviotaensis (Conrad, 1857d). [Mactra?]. Moore 2003a. USGS PP 1228-F [electronic format] (family

13310. Remarks: Error for gabiotensis, but is likely a misprint in original description as the type locality is listed
as Gaviote Pass. [Mactridae].
Mactromeris markleyensis (Clark, 1938). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic format] (family Mac-

Napa quad., Solano Co. UCMP A1297. Holotype UCMP 30852. [Mactridae].
Mactromeris mercedensis (Packard, 1916). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family

Co. Holotype UCMP 11467. Remarks: Junior synonym of Mactromeris catilliformis (Conrad, 1867),  Coan et
al. (2000: p. 454). [Mactridae].
Mactromerismerriami (Packard, 1916). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family Mac-

672. Holotype UCMP 11484 (not UCMP 11467 as in pl. caption), paratype UCMP 11485. [Mactridae].
Mactromeris merriami (Packard, 1916). [Spisula
Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 26617. Hypotypes LACMIP 12753, 12754. [Mactridae].
Mactromeris (Mactromeris montereyana (Arnold, 1908). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic
Mactromeris montereyana in pl. caption). Monterey Fm., Miocene.
Love Creek 
[Mactridae].
Mactromeris mossbeachensis (Glen, 1959). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family

UCMP B4790. Holotype UCMP 37643. [Mactridae].
Mactromeris muliniaformis (Wagner and Schilling, 1923). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic for-
Mactromeris mulineaformis in pl. caption]. Pleito Fm., Oligocene.
Devil's Kitchen, Mt. Pinos quad., Kern Co. UCMP 3203. Holotype UCMP 11423. [Mactridae].
Mactromeris occidentalisHemimactra]. Moore 2003a. USGS PP 1228-F [electronic format] (fam-

ANSP 4545. [Mactridae].
Mactromeris ramonensis attenuata (Clark, 1918). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (fam-
ily Mactridae):11, pl. 6, . 8, 10, 11. San Ramon Fm., MioceneSobrante Ridge area, Contra Costa Co. UCMP
1167. Holotype UCMP 11119. [Mactridae].
Mactromeris ramonensis ramonensis (Packard, 1916). [Spisula]. Moore 2003a. USGS 1228-F [electronic format]
100 PALEOBIOS, VOLUME 38, FEBRUARY 2021

area, Concord quad., Contra Costa Co. UCMP 1687. Holotype UCMP 11118. [Mactridae].
Mactromeris rushi (Wagner and Schilling, 1923). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (fam-

Kern Co. UCMP 3179. Holotype UCMP 11422. Remarks: Figures 6 and 9 on pl. 6 do not match those of Wagner

MactromerisMactromeris sectoris (Anderson and Martin, 1914). [Mactra]. Moore 2003a. USGS PP 1228-F [elec-
Mactra? sectoris in pl. caption]. Jewett (as Jewitt
[sic] in text and pl. caption) Sand, Miocene. South and west slopes of Pyramid Hill, Kern Co. CASG 69. Holotype
CASG 131 [=CASG 69.01], paratype CASG 132 [=CASG 69.02]. [Mactridae].
Mactromeris subquadrata (Clark, 1925). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family Mac-

Costa Co. UCMP 1131. Holotype UCMP 30344. [Mactridae].
Mactromeris tejonensis (Packard, 1916). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic format] (family Mac-

[Mactridae].
Mactromeris (Mactromeris trampasensis (Packard, 1916). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic

Creek, Concord quad., Contra Costa Co. UCMP 1227 (not UCMP 1237 as in text). Holotype UCMP 11499. [Mactridae].
Mactromeris weaveri Packard in Dickerson, 1914a.” Moore 2003a. USGS PP 1228-F [electronic format] (family Mac-
Spisula weaveri” in pl. caption, image reversed in pl.). Martinez Fm., Paleocene. Near
Stewartville, Mt. Diablo quad., Contra Costa Co. UCMP 1556. Holotype UCMP 11714 (syntype in text). Remarks:
Not a mactrid, but actually an Arctica,”  Smith (1975b: p. 470). [Mactridae].
Maorithyas folgeri (Wagner and Schilling, 1923) [Thyasira
13. San Emigdio Fm., Oligocene. Devil’s Kitchen, Kern Co. UCMP 3195. Holotype UCMP 11434.
Marcia angustifrons (Conrad, 1849a). [Venus
Near Lawson Hill and Oursan Ridge, Contra Costa Co. UCMP A4931. Hypotype UCMP 34154. [Veneridae].
Marcia (Mercimonia) bunkeri (M.A. Hanna, 1927). [Dosinia
Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7142; =LACMIP
22312]. Hypotype UCLA 59282 [=LACMIP 10562]. [Veneridae].
Martesia meganosensis      
Paleocene/Eocene. Northeast of Mt. Diablo, Byron quad., Contra Costa Co. UCMP 3577. Holotype UCMP 31297.
[Pholadidae].

Martesia (Paramartesia) tolkieni    
Fm., Eocene [Paleocene]. Vicinity of Ortigalito and Little Panoche creeks, west side of San Joaquin Valley, Merced
 
163804a, 163804c, d. [Pholadidae].

Hypotype SDSNH 4292 [peel of USNM 163804b].
Martesia

1981), Los Angeles Co. LACMIP 16869 [=CSUN 1590). Hypotype LACMIP 7944. [Pholadidae].
Martesia        
Lower Lake Lake Co. UCMP 784. Hypotype UCMP 10607. Remarks: Teredina sp.,  G.L. Kennedy (personal
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 101
communication, 2019). [Pholadidae].
 Kennedy 1974. SDSNH Mem. 8: frontispiece. Lodo Fm., Eocene [Paleocene]. Vicinity of Ortigalito and Little
Panoche creeks, west side of San Joaquin Valley, Merced Co. USGS. 5712. Hypotype USNM 163804. [Pholadidae].
Megapitaria squalida (G.B. Sowerby I, 1835a). [Cytherea]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):268–269,
pl. 50
[Veneridae].
Megapitaria
Pliocene. Painted Gorge, Coyote Mtn., Imperial Co. SDSNH 44]. Hypotype SDSNH 23150. [Veneridae].
Meretrix dibbleei       
Gaviota fms., Eocene. Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6954. Holotype SU 9294 [=CASG
70187]. [Veneridae].

Meretrix stantoni-
cene. East side Panoche Pass Rd., near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU
2073 [=UCMP A9717, =UCMP. 4657]. Hypotypes UCMP 14204, 14205. [Veneridae].
Meretrix
Meretrix ovalis (Gabb 1864)). Domengine Fm.,
Eocene. Northwest of Coalinga, Fresno Co. [Hypotype USNM 165629]. [Veneridae].
Metis
Metis aff. alta (Conrad 1837)). Vaqueros Fm.,

Microcallista (Costacallista) conradiana (Gabb, 1864). [Tapes
7. Undifferentiated Sacate/Gaviota fms., Eocene. Nojoqui Creek, Santa Ynez Mtns., Lompoc quad., Santa Barbara
Co. UCMP B6960. Hypotype SU 9299 [=CASG 70198]. [Veneridae].
Milneria minima (Dall, 1871). [Ceropsis
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotype
LACMIP 4546. [Carditidae].
Milneria minima (Dall, 1871). [Ceropsis-

2, 14) LACMIP 4546. [Carditidae].
Miltha (Miltha jacalitosana (Arnold, 1909 [1910]). [Paphia     
Etchegoin Fm., Miocene/Pliocene. Jasper Creek area, Coalinga quad., Fresno Co. USGS 4765. Holotype USNM
165587. [Lucinidae].
Miltha (Miltha meganosensis (Clark and Woodford, 1927). [Phacoides]. Moore 1988b. USGS PP 1228-D:D19, pl.
  
31303, paratype UCMP 31304. [Lucinidae].
. 8. Deer Valley, Mt. Diablo quad., Contra Costa Co. UCMP 3152. Paratype UCMP 31305.
Miltha packi (Dickerson, 1916). [Lucina
Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotypes
SDSNH. 20767, 20768. [Lucinidae].
Miltha packi (Dickerson, 1916) [Lucina
Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN. 362 [=LACMIP 16273]. Hypotype LACMIP 7517.
Remarks: In the genus/subgenus Gibbolucina (EomilthaMoore (1988: p. D21). [Lucinidae].
102 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Miltha packi (Dickerson, 1916). [Lucina
Whitaker Peak area, Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype LACMIP 7571. Remarks: In the
genus/subgenus Gibbolucina (EomilthaMoore (1988: p. D21).
Miltha packi (Dickerson, 1916). [Lucina     
Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13444. [Lucinidae].
Remarks: In the genus/subgenus Gibbolucina (EomilthaMoore (1988: p. D21).
Miltha (Miltha) parsoni
Fm.], Paleocene. Las Virgenes Canyon, Simi Hills, Camulos quad., Ventura Co. SU 2697 [=CASG 61667]. Holotype
SU/CASG 150 [=CASG 61667.19]. [Lucinidae].
Miltha sanctaecrucis (Arnold, 1909 [1910]). [Phacoides
63-18. Vaqueros Fm., Miocene. Ventura Co. CASG 27409. Hypotype CASG 5740. [Lucinidae].
Miltha sanctaecrucis (Arnold, 1909 [1910]). [Phacoides
Fm., Miocene. Kern River, Kern Co. USGS 6622. Hypotype USNM 498743. [Lucinidae].
     
498746.
Miltha sanctaecrucis (Arnold, 1909 [1910]). [Phacoides
peel). Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, Palos Verdes
Hills, Los Angeles Co. USGS 13127. Hypotype USNM 497087. [Lucinidae].
Miltha (Miltha) sanctaecrucis (Arnold, 1909 [1910]). [Phacoides
5. Temblor Fm. (Wygal Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3579. Hypotype
USNM 646543. [Lucinidae].

9432. Hypotype USNM 646544.
Miltha santaecrucis (Arnold, 1909 [1910]). [Phacoides]. Stadum 1973. -
cene. Topanga Canyon, Los Angeles Co. UCLA 2488 [=LACMIP 22488]. Hypotype UCLA 49518 [=LACMIP 10617].
[Lucinidae].
Miltha cf.  (Arnold, 1909 [1910]). [Phacoides]. Addicott 1979. SEPM Guidebook 14:47–48, pl. 1,

Range, Monterey Co. SU 1756. Hypotype CASG 60371, determined here as missing. [Lucinidae].
Miltha (Miltha) sanctaecrucis (Arnold, 1909 [1910]). [Phacoides
7. Vaqueros Fm., Oligocene/Miocene. Carrizo Creek, La Panza Mtns., San Luis Obispo Co. UCMP A501. Hypotype


165569.
Miltha xantusi (Dall, 1905). [Phacoides  
Pliocene. Balboa Park, San Diego, San Diego Co. SDSNH 1965. Hypotype SDSNH 4176. [Lucinidae].
Miltha xantusi (Dall, 1905). [Phacoides
Los Angeles Co. LACMIP 17934. Hypotype LACMIP 14354. [Lucinidae].
Miltha (Miltha) xantusi (Dall, 1905). [Phacoides]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):250–251, pl. 45,
 
[Lucinidae].

72538].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 103
Miltha (Miltha) xantusi (Dall, 1905). [Phacoides
2. Pico Fm., Pliocene. Holser Canyon, Los Angeles Co. SDSNH 217a. Hypotypes (of Grant and Gale 1931, pl. 14,

-

Miltha xantusi (Dall, 1905). [Phacoides      
Fm.], Pliocene. Near Travertine Rock, Imperial Co. (not San Diego Co. as in locality description). USGS M9778.
Hypotype USNM 486911. [Lucinidae].
Miltha

Miodonticus prolongatus (Carpenter, 1864b). [Miodon]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):233–234, pl.
      
[Carditidae].
Miodonticus prolongatus (Carpenter, 1864b). [Miodon]. 12–14, 16. San
Diego Fm., Pliocene. San Diego, San Diego Co. LACMIP 323. Hypotypes (of Hertlein and Grant 1972, pl. 
1, 4) LACMIP 4541, 4544. [Carditidae].
Modiolus altiobliquus-
cene. Simi Hills, Ventura Co. UCR 6673. Hypotype UCR 6673/7. Remarks: Provisionally in the genus/subgenus
BrachidontesBrachidontes Moore (1983: p. A67). [Mytilidae].
Modiolus bakeri-
leocene. Simi Hills, Ventura Co. [UCR 7321]. Hypotype UCR 7321/1. Remarks: In the genus Idasola Moore
(1983:A78). [Mytilidae].
Modiolus? (Modiolus?) clarki Moore, 1983 (new name for Mytilus merriami Clark, 1915, preoccupied). USGS PP

Modiolus (Modiola) contracta    

Modiolus directus-
ber), Pliocene. Pomponio Creek, San Gregorio quad., San Mateo Co. locality 469-3M. Hypotype determined here
as missing. Remarks: Junior synonym of Modiolus (Modiolusia) rectus (Conrad, 1837),  Coan et al. (2000: p.
186). [Mytilidae].
 (Gould, 1850) [Mytilus (Modiola
Pliocene. Near Capitola beach, Santa Cruz Co. SCCM 1. Hypotype SCCM 9681.02. Remarks: Junior synonym of
Modiolus (Modiolusia) rectus Conrad, 1837,  Coan et al. (2000: p. 186). [Mytilidae].
Modiolus gabbi
La Costa Valley quad. (1953), Alameda Co. SU 2739. Hypotype SU 8446 [=CASG 69940]. Remarks: In the genus/
subgenus Brachidontes (Aeidimytilus),  Moore (1983: p. A69). [Mytilidae].
Modiolus (Modiolus) lagunanus-
ros Fm., Oligocene/Miocene. San Joaquin Hills, Corona quad., Orange Co. UCMP A543. Holotype UCMP 31788.
[Mytilidae].
ModiolusModiolus meganosensis-
ganos Fm., Paleocene/Eocene. Mt. Diablo quad., Contra Costa Co. UCMP. 3155. Holotype UCMP 31363. [Mytilidae].
Modiolus (Modiolus merriami-
cene. Concord quad., Contra Costa Co. UCMP 333. Holotype UCMP 11892. [Mytilidae].
Modiolus (Modiolus pittsburgensis Clark, 1918. Moore, 4 (cast, CASG).
104 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Kirker Tuff, Oligocene. Vicinity of Pittsburg, north of Mt. Diablo, Mt. Diablo quad., Contra Costa Co. UCMP 1895.
Holotype UCMP 11122. [Mytilidae].
Modiolus rectus
Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. [Hypotype LACMIP 4474. Remarks: In the subgenus
Modiolusia,  Coan et al. 2000: p. 186. [Mytilidae].
Modiolus rectus
CASG 66068. Hypotype CASG 66068.01]. Remarks: In the subgenus Modiolusia,  Coan et al. 2000: p. 186.
[Mytilidae].
Modiolus sacculifer (Berry, 1953b). [Volsella
2–4. San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305.
Hypotypes LACMIP 4475, 4476. [Mytilidae].
Modiolus (Modiolus) veronensis veronensis
Miocene. Pleasanton quad., Alameda Co. UCMP 3529. Holotype UCMP 12373. [Mytilidae].
Modiolus veronensis temblorensis
Reef Ridge quad., Fresno Co. UCMP D1065. Holotype UCMP 36606, paratype UCMP 36607. [Mytilidae].
Modiolus (Modiolus) veronensis temblorensis
Temblor Fm., Miocene. Reef Ridge quad., Kings Co. UCMP D1065. Holotype UCMP 36606. [Mytilidae].
Modiolus (Modiolus) ynezianus
Oligocene/Miocene. Southeast of Lompoc, Santa Barbara Co. USGS 4504. Holotype USNM 165324. [Mytilidae].

Ridge quad., Fresno Co. UCMP D1065. Holotype UCMP 36601, paratype UCMP 36602. [Mytilidae].
Modiolus (Modiolus) ynezianus garzaensis
Temblor Fm., Miocene. Reef Ridge quad., Kings Co. UCMP D1065. Holotype UCMP 36601. [Mytilidae].
Modiolus-
ity of Gaviota Pass, western Santa Ynez Mtns., Los Olivos quad., Santa Barbara Co. UCMP B6998. Hypotype SU
9256 [=CASG 70191]. [Mytilidae].
Mulina* (Pseudocardium       
Remarks: Determined here as
Pseudocardium densatum (Conrad, 1857a). Error for Mulinia. [Mactridae].
Mulinia densata Conrad, 1857a. G D. Hanna and Hertlein 1943. 
Fm., Pliocene. [Mulinia Zone], Kettleman Hills, Kings Co. CASG 26820. Hypotypes CASG 5743, 5743A. Remarks:
Pseudocardium densatum (Conrad, 1857a),  Moore (2003a: p. 4). [Mactridae].
Musculus (Musculus) stalderi (Martin, 1914). [Modiolus
Fm., Pliocene. Vicinity of Ferndale, Humboldt Co. UCMP 1859. Holotype UCMP 12354. Remarks: Junior synonym
of Musculus niger (Gray, 1824),  Coan et al. (2000: p. 169). [Mytilidae].
Mya arenaria
Co. UCMP 1224. Hypotype UCMP 15007. [Myidae].
Mya Zone], Pliocene. North Dome, Kettleman Hills, Kings Co. USGS 12480.
Mya cf. ) USNM 495304.

Mya japonica” Zone, Kettleman Hills, Kings Co. UCMP A3167.
Hypotypes UCMP 30801, 30802.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 105
Mya arenaria
Pliocene. North of Sarah Canyon, Wunpost quad., Monterey Co. USGS M1934. Hypotype USNM 649093. [Myidae].
Mya (Arenomya) arenaria 
Reef Ridge quad., Kings Co. UCMP B6527. Hypotype UCMP 36786. [Myidae].
Mya (Arenomya) arrosis
Vicinity of Tice Valley, Contra Costa Co. UCMP 1687. Hypotype UCMP 15003. [Myidae].


15006.
Mya cf. 
Mya Zone), Pliocene. Las Alturas, Middle Dome, Kettleman Hills, Kings Co. USGS 12684. Hypotype USNM 495305.
Remarks: Mya (Arenomya) arenaria Linneaus, 1758,  MacNeil (1965: p. G33). [Myidae].
Remarks:
Mya (Arenomya) arenaria Linneaus, 1758,  MacNeil (1965: p. G33).

Remarks: Mya (Arenomya) arenaria Linneaus, 1758,  MacNeil (1965: p. G33).
. (upper oolite between Neverita Zone and Cascajo Conglomerate Member), Pliocene.
North of El Campo, North Dome, Kettleman Hills, Kings Co. USGS 12841. Hypotype USNM 495339.
     Pseudocardium-Anadara Zone), Pliocene. North of La Escudilla, Middle Dome,
Kettleman Hills, Kings Co. USGS 12640. Hypotype USNM 495357.
Mya Arenomya) dickersoni
Fm., Miocene. Near town of Rodeo, Contra Costa Co. UCMP 1617. Holotype UCMP 11554, hypotypes UCMP 15001,
15002.
Mya (Arenomya) dickersoni 
Domengine Ranch quad., Fresno Co. UCMP D703. Hypotype UCMP 36781. [Myidae].
Mya (Arenomya) fujieiMya cf.  MacNeil (n. sp)).
Mya dickersoni
Clark, 1915) UCMP 11553 (not UCMP 11533 as in pl. caption). [Myidae].
Mya (Arenomya) fujiei
Oil Canyon, Joaquin Rocks quad., Fresno Co. UCMP B7085. Hypotype UCMP 36779. [Myidae].

Mya (Arenomya) fujiei
3). Temblor Fm., Miocene. Garza Creek area, Reef Ridge quad., Kings Co. UCMP D1059. Hypotype UCMP 36780.
[Myidae].

UCMP B7085. Hypotype UCMP 36779.
Mya japonica
[Mya Zone], Pliocene. North Dome, Kettleman Hills, Fresno Co. Hypotypes CASG 5925, 5925A. Remarks: Junior
synonym of Mya arenaria Linneaus, 1758,  Coan et al. (2000: p. 470).[Myidae].
Mya (Arenomya) japonica
Ridge quad., Kings Co. UCMP D1200. Hypotype UCMP 36787. Remarks: Junior synonym of Mya arenaria Lin-
neaus, 1758,  Coan et al. (2000: p. 470). [Myidae].
106 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Mya (Arenomya) macneili
Ridge quad., Kings Co. UCMP D1120. Paratypes UCMP 36783–36785. [Myidae].

MyaMya Zone”), Pliocene. North

Myadesmaeocenica (Clark, 1938). [Mesodesma]. Moore, 2003a. USGS PP 1228-F [electronic format] (family Mes-
odesmatidae):1, 
[Mesodesmatidae].
 (Hall and Ambrose, 1916) [Mesodesma      
Alameda Creek, Alameda Co. CASG61804]. Holotype CASG 61804.01. [Myadesmatidae, not Mesodesmatidae,
 Saul (1989: p. 200)].
Myrakeena cf.  (Rochebrune, 1895). [Ostrea
Fm. [=Latrania Fm.], Pliocene. Near Travertine Rock, Imperial Co. (not San Diego Co. as in locality description).
USGS M9778. Hypotype USNM 486902. [Ostreidae].
Myrakeena veatchii (Gabb, 1866). [Ostrea
Pliocene. Near Valencia, Oat Mtn. quad. (1952, PR 1969), Los Angeles Co. LACMIP 17777. Hypotypes LACMIP
13339, 13341–13343. [Ostreidae].

Myrakeena veatchii (Gabb, 1866). [Ostrea
area, Los Angeles Co. LACMIP 9659. Hypotype LACMIP 14340. [Ostreidae].

Myrtea (Myrtea) taffana (Dickerson, 1916). [Phacoides
Fm. (Cerros Shale Member), Eocene. Parson's peak area, Coalinga quad. [=Joaquin Rocks quad.], Fresno Co. UCMP
672. Holotype UCMP 11789. [Lucinidae].
Mysella tumida Carpenter, 1864b. [Tellimya 
2–5. San Diego Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotypes LACMIP 4625, 4626.
Remarks: In the genus Kurtiella,  Coan and Valentich-Scott (2012: p. 517). [Lasaeidae].

LACMIP 4627, 4628.
Mysella (Rochefortia) tumida Carpenter, 1864b. [Tellimya  
Diego Fm., Pliocene. Near U.S./Mexico boundary, southwest of Goat Canyon, Imperial Beach quad., San Diego Co.
Remarks:
In the genus Kurtiella,  Coan and Valentich-Scott (2012: p. 517). [Lasaeidae].
Mytella inezensis (Conrad, 1857d). [Mytilus
Keen and Bentson 1944: p. 72), Eocene/Oligocene. Santa Ynez Mtns. and Santa Ynez River, Santa Barbara Co.
Syntypes USNM 13319. [Mytilidae].
Mytilus (Mytilus arnoldi
Pittsburg, Contra Costa Co. UCMP 78. Holotype UCMP 11153. [Mytilidae].
Mytilus (Mytilus ascia Gabb, 1864. Moore, 1983. USGS PP 1228-A
Kern Co. IP12930. Holotype UCMP 11989. [Mytilidae].
Mytilus coalingensis
Fm. (Siphonalia Zone), Pliocene. Arroyo Estrecho, southwest of Broken Hill, North Dome, Kettleman Hills, Kings
Co. USGS 12812. Hypotypes USNM 495818, 495819. Remarks: In the subgenus Crenomytilus, Moore (1983:
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 107
p. A64). [Mytilidae].
Mytilus (Crenomytilus) coalingensis-
goin Fm., Miocene/Pliocene. At northwest end of Anticline Ridge, northeast of Coalinga, Coalinga quad.], Fresno
Co. USGS 4656. Holotype USNM 165551, paratype USNM 165557. [Mytilidae].
Mytilus coalingensis-
ange Co. [SDSNH 4080]. SDSNH hypotype determined here as missing. Remarks: In the subgenus Crenomytilus,
 Moore (1983: p. A64). [Mytilidae].
Mytilus coalingensis
Saddleback Valley, Orange Co. SDSNH 4312. Hypotype SDSNH 83246. Remarks: In the subgenus Crenomytilus,
 Moore (1983: p. A64). [Mytilidae].
Mytilus (Crenomytilus) coalingensis sternbergi Hertlein and Grant, 1972 (n.subsp.). SDSNH Mem. 2(2B):163–164,
Remarks:
Mytilus (Crenomytilus) sternbergi Hertlein and Grant, 1972,  Moore (1983: p. A64). [Mytilidae].



Mytilus condoni  
South quad., San Mateo Co. UCMP D3364. Hypotype UCMP 10928. [Mytilidae].
Mytilus condoni         
Twelvemile Creek area, San Mateo Co. UCMP D3364. Hypotype USNM 251800. [Mytilidae].
         
10928. [Mytilidae].
Mytilus cf.    
(Upper Mya Zone), Pliocene. Las Alturas, Middle Dome, Kettleman Hills, Kings Co. USGS 12685. Hypotype USNM
495293. Remarks: Probably Mytilus trossulus Gould, 1850b,  Coan et al. 2000: p. 159. [Mytilidae].
Mytilus edulis-
boldt Co. UCMP A4233. Hypotype UCMP 34176]. Remarks: Probably Mytilus trossulus Gould, 1850,  Coan
et al. (2000: p. 159). [Mytilidae].
Mytilus (Crenomytilus) expansus Arnold, 1907. Moore 1983. USGS., Oli-
gocene/Miocene. Southwest of Piru, Piru quad., Ventura Co. USGS 3897]. Holotype USNM 164968. [Mytilidae].
Mytilus expansus
Fm., Miocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. UCSB RB65-6 [=LACMIP 16421]. Hypo-
type UCSB M8 [=LACMIP 6294]. Remarks: In the subgenus Crenomytilus,  Moore (1983: p. A63). [Mytilidae].
Mytilus expansus
Fm., Miocene. Cañada Posa area, Santa Cruz Is., California Channel Is. Hypotype UCSB 1851 [=SBMNH 60169].
Remarks: In the subgenus Crenomytilus,  Moore 1983: p. A63. [Mytilidae].
Mytilus highoohiae
quad. (1943), Humboldt Co. UCMP A4234. Holotype UCMP 35518 [= UCMP 12506]. Remarks: Junior synonym
of Mytilus condoni Dall, 1890b,  Addicott (1974b: p. 355);  Moore (1983: p. A61). [Mytilidae].
Mytilus highoohiae       
1950:22–25), Pliocene. Blue Lake quad. (1943), Humboldt Co. UCMP A4234. Holotype UCMP 35518 [=UCMP
12506]. Remarks: Junior synonym of Mytilus condoni Dall, 1890b,  Addicott (1974b: p. 355);  Moore
(1983: p. A61). [Mytilidae].
108 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Mytilus (Crenomytilus) kewi
Miocene/Pliocene. Fresno Co. UCMP 2680. Holotype UCMP 12061. [Mytilidae].
Mytilus (Mytilus loeli-
cene. Vaqueros Creek, Junipero Serra quad., Monterey Co. SU 200. Holotype SU/CASG 515. [Mytilidae].
Mytilus (Crenomytilus) mathewsonii

[Mytilidae].
Mytilus aff.  Grewingk, new species. Manning and Ogle 1950. CDMG Bull. 148:24 [as ],
 Man-
dra 1949) UCMP 35523. Remarks: Mytlius condoni Dall, 1890b,  Addicott (1974b: p. 355); in the subgenus
Plicatomytilus,  Moore (1983: p. A65). Error for . [Mytilidae].
-
vington, La Costa Valley quad. (1953), Alameda Co. SU 3248. Hypotype SU/CASG 8445 [=CASG 69943]. Remarks:
Mytilus (Plicatomytilus) gratacapi n. sp.,  Allison and Addicott (1976: p. 10). [Mytilidae].
  
Creek area, New Idria quad., San Benito Co. USGS M3614. Hypotypes USNM 647269 [as USNM 647064 in Alison
and Addicott 1976] and 647270. Remarks: In the subgenus Plicatomytilus,  Moore (1983: p. A65). [Mytilidae].
Mytilus (Plicatomytilus) Grewingk, 1850. Allison and Addicott
3, 9. Temblor Fm., Miocene. Silver Creek area, Griswold Hills, New Idria quad., San Benito Co. USGS M3614. Hy-
potypes USNM 647058, 647059, 647269 [as USNM 647064 in Allison and Addicott 1976: pl. 1, 

Mytilus (Crenomytilus perrini
(Neroly Sandstone), Miocene. Near town of Rodeo, Napa quad., Contra Costa Co. UCMP 1617. Holotype UCMP
11548. [Mytilidae].
Mytilus (Crenomytilus schencki

Mytilus (Crenomytilus) sternbergi
Diego Fm., Pliocene. San Diego, San Diego Co. LACMIP 107. Holotype LACMIP 4470. [Mytilidae].
Mytilus trampasensis    
(1915), Contra Costa Co. UCMP A2568. Hypotype UCMP 30675. Remarks: In the subgenus Crenomytilus, 
Moore (1983: p. A63). [Mytilidae].
Mytilus (Crenomytilus) trampasensis
(Neroly Fm.) (Cierbo Sandstone in locality register), Miocene. South of Pittsburg, Contra Costa Co. UCMP 402.
Syntype UCMP 11545. [Mytilidae].

Mytilus
Remarks:
206) as being from two localities, the exact locality of Mytilus of Fowkes 1982 is unknown. [Mytilidae].


Mytilus
Topatopa Mtns., Ventura Co. CSUN 412 [=LACMIP 16284]. Hypotype UCLA 58224 [=LACMIP 10580]. [Mytilidae].
Mytilus
Miocene. Kern River area, Kern Co. USGS M1591. Hypotype USNM 647077. [Mytilidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 109
Nanaochlamys (Arnold, 1906). [Pecten
San Gregorio Creek area, Santa Cruz quad., San Mateo Co. CASG 68153. Holotype SU/CASG 6 [=CASG 61853.01].
[Pectinidae].
Nayadina (Exputens) alexi (Clark, 1934). [Exputens
Fm., Eocene. Los Gatos Creek area, Coalinga quad., Fresno Co. UCMP A1007. Holotype UCMP 32386. Remarks:
Junior synonym of Nayadina (Exputens) llajasensis (Clark, 1934),  Squires (1990b: p. 309). [Malleidae].
Nayadina (Exputens) llajasensis (Clark, 1934). [Exputens
Hot Springs Creek area, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4659. Hypotype UCR 4659/81. [Malleidae].
Nayadina (Exputens) llajasensis (Clark, 1934). [Exputens
13. Llajas Fm., Eocene. Las Llajas Canyon Simi Valley quad. [=Santa Susana quad.], Ventura Co. UCMP 7004. Ho-
lotype UCMP 32391. [Malleidae].
Nayadina (Exputens) llajasensis (Clark, 1934). [Exputens     
Fm., Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242].
Hypotypes LACMIP 6569, 6570. [Malleidae].
Nayadina (Exputens) llajasensis (Clark, 1934). [Exputens
30, 32, 33, 46. Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana quad. (1969), northern Simi Valley, Ventura
Co. UCMP 7004. Holotype UCMP 32391, paratypes UCMP 32389, 32390, 32392, 32393. [Malleidae].

   
UCMP A1007. Holotype UCMP 32386, paratypes (of Exputens alexi Clark 1934) UCMP 32384, 32385.
Neaeromya (Orbitella) compressa (Dall, 1899b). [Erycina
Orbitella com-
pressa)), determined here as missing. [Lasaeidae].
Nemocardium (Keenaea) (Carpenter, 1864b). [Cardium]. Hertlein and Grant 1972. SDSNH Mem.

4347. [Cardiidae].

7154].
Nemocardium linteum (Conrad, 1855b). [Cardium-
stone, Eocene. Reef Ridge, Kettleman Plain quad., Kings Co. USGS 14483. Hypotype USNM 498720. [Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium-
yon Fm.” [=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH 2363. Hypotype SDSNH 4161].
[Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium]. Zinsmeister 1983a. SEPM
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6806]. Hypotype UCR. 6806/5. [Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium-
cene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype
LACMIP 6584. [Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium-
-
potype LACMIP 7530. [Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LACMIP 7731.
[Cardiidae].
110 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Nemocardium linteum (Conrad, 1855b). [Cardium
Fm., Eocene. Inyokern SE quad. (1972), El Paso Mtns., Kern Co. USGS M9067. Hypotype USNM 424896. [Cardiidae].
Nemocardium (Nemocardium) linteum (Conrad, 1855b) [Cardium]. Moore 2003a. USGS PP 1228-F [electronic format]

Cardium dalli Dickerson 1913) UCMP 11756. [Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium
Fm., Eocene. North side Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 26111 (= CSUN
965). Hypotype LACMIP 12698. [Cardiidae].
Nemocardium linteum (Conrad, 1855b). [Cardium-
mere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13449. [Cardiidae].
Nemocardium
Remarks: Determined here as Nemocardium linteum (Conrad,
1855b). [Cardiidae].

Remarks: Determined here as Nemocardium linteum (Conrad, 1855b).
Nemodon morani (Waring, 1917). [Cucullaea
uppermost Cretaceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo
Co. UCLA. 6525 [=LACMIP 26525]. Hypotype LACMIP 7556. Remarks: In the genus Cucullaea,  Moore (1983:
p. A47). [Cucullaeidae].

Eocene. North Fork of Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963].
Holotype LACMIP 12459, paratypes LACMIP 12460, 12461, 12463. [Trapezidae].
32. Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1445 [=LACMIP 24259]. Paratype LACMIP 12462.
Netastoma squiresi
North side of Simi Valley, Ventura Co. LACMIP 12648. Holotype LACMIP 8405. [Pholadidae].
Netastoma squiresi
side Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 967 [=LACMIP 12648]. Holotype LACMIP
8405. [Pholadidae].
Netastomella rostrata (Valenciennes, 1846). [Pholas]. Addicott and Galehouse 1973. Jour. Res. USGS 1(5):510, 513,

647004. Remarks: Netastoma rostratum (Valenciennes, 1846),  Coan et al. (2000: p. 502). [Pholadidae].
Nototeredo-

quad. (1952, PR 1981), Los Angeles Co. LACMIP 16869 [=CSUN 1590). Hypotype LACMIP 7945. [Teredinidae].
Nucula
ridge north of Sobrante Ridge, southeast of west fork of Bear Creek, Concord quad., Contra Costa Co. UCMP 331.
Holotype UCMP 11184. [Nuculidae].
Nucula (Ennucula) balboana
Diego Fm., Pliocene. San Diego well, Balboa Park, San Diego, San Diego, Co. CASG 12099. Holotype CASG [=CASG
Remarks: Valid species 
Moore (1983: p. A8); junior synonym of Ennucula tenuis (Montagu, 1808),  Coan et al. (2000: p. 76). [Nuculidae].
Nucula (Leionucula) balboana
Fm., Pliocene. Well in Balboa Park, San Diego, San Diego Co. CASG 12099. Holotype CASG 59684. Remarks: Valid
species,  Moore (1983: p. A8). Junior synonym of Ennucula tenuis (Montagu, 1808),  Coan et al. (2000: p.
76). [Nuculidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 111
Nucula (Ennucula) birchi  
Miocene. Caliente quad., Kern Co. SU 2121 [=CASG 71992]. Holotype SU 7527 [=CASG 72008], paratypes SU
Remarks: In the subgenus Leionucula,
 Moore (1983: p. A7). [Nuculidae].
….., [SDSNH 5422 (ex
Nucula (Leionucula) birchi
Silt, Miocene. Caliente quad., Kern Co. SU 2121. Holotype SU/CASG 7527 [=CASG 72008]. [Nuculidae].
Nucula (Nucula) capayensis  
Capay Valley quad., Yolo Co. UCMP A1319. Holotype UCMP 30196. [Nuculidae].
Nucula (Nucula) cooperi Dickerson, 1913. Moore 1983. USGS PP 1228-A:A6
Marysville [Sutter] Buttes, Marysville [Sutter] Buttes quad., Sutter, Co. UCMP 1853. Lectotype (as holotype in pl.
caption) (of Moore 1983: p. A6) UCMP 11754a. [Nuculidae].
Nucula (Lamellinucula) exigua G.B. Sowerby I, 1833. Hertlein and Grant 1972. SDSNH Mem. 2(2B):146–147, pl. 27,

Hypotype UCLA 48610 [=LACMIP 9287]. [Nuculidae].
Nucula (Lamellinucula) exigua
Pliocene. Near the U.S./Mexico boundary, San Diego Co. UCLA312 [=LACMIP 20312]. Hypotype (of Hertlein and

Nucula (Lamellinucula) kennedyi-
sana Fm., Paleocene. First concretionary horizon above the Las Virgenes Sandstone, Thousand Oaks quad., Simi
Hills, Ventura Co. UCR 6683. Holotype UCR 6683/300, paratype UCR 6683/301. [Nuculidae].
Nucula (Leionucula) postangulata-

Nucula (Nucula) vitis
Eocene. Near mouth of Grapevine Canyon, Tejon quad., Kern Co., CASG 245. Holotype CASG 788. [Nuculidae].
Nucula (Lamellinucula
Simi Hills, Ventura Co. [UCR 6683]. Hypotype UCR 6683/301 (not UCR 6683/12 as in pl. caption). Remarks:
Nucula (Lamellinucula) kennedyi n. sp.,  Zinsmeister (1983b: p. 1284). [Nuculidae].
Nuculana (Saccella) alaeformis (Gabb, 1869b). [Corbula-
Leda
alaeformis)) USNM 157829. [Nuculanidae].

Nuculana (Saccella) alaeformis (Gabb, 1869b). [Corbula
Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR6683. Hypotype UCR 6683/28 (not UCR 6682/28 as in
pl. caption). [Nuculanidae].
Nuculana (Saccella) cahillensis (Arnold, 1908). [Leda

57, west of Woodside, Santa Cruz quad., San Mateo Co. Holotype SU/CASG 5368 (not SU/CASG 5386 as in text)
[=CASG 71658]. [Nuculanidae].
Nuculana (Saccella) chaneyi Vokes, 1939. Moore 1983. USGS. Avenal Sand-
stone, Eocene. East side of Big Tar Canyon, Cholame quad., Kings Co. UCMP A1165 and A976. Syntypes UCMP
15745, 15746. [Nuculanidae].
Nuculana chehalisensis (Weaver, 1912). [Leda
Contra Costa Co. UCMP A4928. Hypotype UCMP 34120. Remarks: In the subgenus Saccella,  Moore (1983:p.
A18. [Nuculanidae].
112 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Nuculana (Saccella denominata (G D. Hanna, 1924). [Leda-
tinez Fm., Paleocene. South of Stewartville, Mt. Diablo quad., Contra Costa Co. UCMP 1556. Holotype (of Leda
milleri Dickerson, 1914a [=Leda denominata G D. Hanna 1924: p. 169) UCMP 11663, new name for Leda milleri
Dickerson, 1914a, preoccupied]. [Nuculanidae].
Nuculana (Nuculana elongorostrata (Clark, 1918). [Leda
Tuff, Oligocene. Sobrante Ridge, on west fork of Bear Creek, Contra Costa Co. UCMP 2755. Holotype UCMP 11206.
[Nuculanidae].
Nuculana (Saccella fabata (Nelson, 1925). [Leda
[=Santa Susana Fm.], Paleocene. Simi Valley, Santa Susana quad., Ventura Co. UCMP 3752. Holotype UCMP 30717.
[Nuculanidae].
Nuculana (Saccella) furlongi (Trask, 1922). [Leda
Fm., Miocene. Concord quad., Contra Costa Co. UCMP 15. Holotype UCMP 12362. [Nuculanidae].
Nuculana furlongi (Trask, 1922). [LedaNuculana forlongi).

101109]. [Nuculanidae].
Nuculana (Saccella) gabbi (Gabb, 1869b). [Leda
Saccella gabbii (Gabb 1864)). Martinez Fm., Eocene. Martinez, Contra
Costa Co. Lectotype (of Stewart 1930: p. 56) ANSP 4476]. Remarks: Should be cited as Nuculana (Saccella) gabbi
Conrad (ms.) of Gabb 1869b (not 1864). See Groves and Squires 2018: p. 49 regarding Leda protexta
an explanation of the complicated naming history of this species. Type material missing,  K. Estes-Smargiassi
(personal communication, 2019). [Nuculanidae].
Nuculana (Saccella) gabbii (Gabb, 1869b). [Leda  -
ferentiated Sacate/Gaviota fms., Eocene. Nojoqui Creek area, Lompoc quad., Santa Barbara Co. UCMP B6957.
Hypotype SU 9246 [=CASG 70197]. Remarks: Should be cited as Nuculana (Saccella) gabbi Conrad (ms.) of Gabb
1869 (not 1864). See Groves and Squires 2018: p. 49 regarding Leda protexta
complicated naming history of this species. [Nuculanidae].
Nuculana (Saccella) gabbii (Gabb, 1869b). [Leda
Guillermo quad. (1943), Ventura Co. UCR 4743. Hypotype UCR 4743/301. Remarks: Should be cited as Nuculana
(Saccella) gabbi Conrad (ms.) of Gabb 1869b, (not 1864). See Groves and Squires 2018: p. 49 regarding Leda
protexta
Nuculana gabbii (Gabb, 1869b). [Leda]. Deméré et al. 1979. SEPM Guidebook 4:98, pl. 
Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH
20770. Remarks: In the subgenus Saccella,  Moore (1983: p. A16). Remarks: Should be cited as Nuculana
(Saccella) gabbi Conrad (ms.) of Gabb 1869b (not 1864). See Groves and Squires (2018: p. 49) regarding Leda
protextahistory of this species. [Nuculanidae].
Nuculana (Saccella) gabbii (Gabb, 1869b). [Leda
Fm., Eocene [=probably Vine Hill Fm., Paleocene]. Martinez, Contra Costa Co. Lectotype (of Stewart, 1930: p. 56)
ANSP 4476, hypotype ANSP. 4476a. Remarks: See Weaver (1953: p. 28) for probable occurrence of this species
in the area of the type locality. Should be cited as Nuculana (Saccella) gabbi Conrad (ms.), of Gabb 1869b, (not
1864). See Groves and Squires (2018: p. 49) regarding Leda protexta
naming history of this species. [Nuculanidae].
Nuculana (Calorhadia) gabbii (Gabb, 1869b). [Leda
Susana Fm., Paleocene. Simi Hills, Ventura Co. [UCR 6684]. Hypotype UCR 6684/19 (not UCR 6684/1 as in pl.
caption). Remarks: In the subgenus Saccella,  Moore (1983: p. A16). Should be cited as Nuculana (Saccella)
gabbi Conrad (ms.) of Gabb 1869b (not 1864). See Groves and Squires (2018: p. 49) regarding Leda protexta
Gabb, for an explanation of the complicated naming history of this species. [Nuculanidae].
Nuculana (Saccella) gabbii (Gabb, 1869b). [Leda
Santa Susana quad., Los Angeles Co. CSUN 467 [=LACMIP 16079]. Hypotype UCLA 59279 [=LACMIP 10559].
Remarks: Should be cited as Nuculana (Saccella) gabbi Conrad (ms.), of Gabb 1869b (not 1864). See Groves
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 113
and Squires (2018: p. 49) regarding Leda protexta
of this species. [Nuculanidae].
Nuculana (Saccella) gabbi (Gabb, 1869b). [Leda

Remarks: UCMP unnumbered hypotype lost , C. Throckmorton (personal communication, 2002). Should be
cited as Nuculana (Saccella) gabbi Conrad (ms.) of Gabb 1869b (not 1864). See Groves and Squires (2018: p. 49)
regarding Leda protexta
Nuculana (Saccella) gabbii (Gabb, 1869b). [Leda
Fm., Eocene. Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 967 (=LACMIP 12648). Hypotype LACMIP
12694. Remarks: Should be cited as Nuculana (Saccella) gabbi Conrad (ms.) of Gabb 1869b (not 1864). See
Groves and Squires (2018: p. 49) regarding Leda protexta
history of this species. [Nuculanidae].
Nuculana (Saccella) hindsii (Hanley, 1860). [Leda
Fm. (Cebada Member), Pliocene. Graciosa Ridge southeast of Orcutt, Santa Barbara Co. USGS 14648]. Hypotype
Saccella redondoensis (Burch, 1944)) USNM 560096.
[Nuculanidae].
Nuculana (Saccella) hondana Vokes, 1939. Givens 1974. UCPG 
Creek, Topatopa Mtns. Quad. (1943), Ventura Co. UCR 4662. Hypotype UCR 4662/12. [Nuculanidae].
Nuculana (Saccella) hondana
Member), Paleocene [Eocene]. Vicinity of where Urruttia Canyon enters Salt Creek, Coalinga quad., Fresno Co.
UCMP 1817. Holotype UCMP 15561. [Nuculanidae].
Nuculana (Nuculana) aff. () leonina (Dall, 1896a). [Leda]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):149,
Nuculana aff. ). San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro
quad. (1943), San Diego Co. LACMIP 305. Hypotype LACMIP [4451]. [Nuculanidae].
Nuculana (Saccella) markleyensis (Clark, 1918). [Leda   

UCMP 11272. [Nuculanidae].
Nuculana ochsneri (Anderson and Martin, 1914). [Ledaochosneri in pl.
caption]. Temblor Fm., Miocene. Reef Ridge quad., Fresno Co. UCMP D1056. Hypotype UCMP 33505. Remarks:
In the subgenus Saccella,  Moore (1983: p. A19). [Nuculanidae].
Nuculana (Saccella) ochsneri (Anderson and Martin, 1914). [Leda
24, 25. Round Mtn. Silt, Miocene. North bank of Kern River, Caliente quad., Kern Co. CASG 68. Holotype CASG
103 [=CASG 68.01]. [Nuculanidae].
Nuculana (Saccella) orcutti (Arnold, 1907). [Leda
Fm., Pliocene. South of Guadelupe [sic], Santa Barbara Co. USGS 4473. Holotype USNM 165271. [Nuculanidae].
Nuculana (Saccella packardi (Dickerson, 1914a). [Leda
Fm., Paleocene. Near Lower Lake, Lake Co. UCMP 784. Holotype UCMP 11725. [Nuculanidae].
Nuculana (Saccella) cf.  () parkei coosensis Turner, 1938. Givens and Kennedy 1976. Jour. Paleo. 50(5):969,

4865. Hypotype UCR 4865/140. Remarks: Provisionally in the genus Hilgardia Moore (1983: p. A23).
[Nuculanidae].
Nuculana (Nuculana) pernula
Upper Newport Bay, Orange Co. UCR 4915. Hypotype UCR 4915/317, determined here as missing. [Nuculanidae].
Nuculana (Costelloleda powersi (Dickerson, 1914a). [Yoldia   
5. Martinez Fm., Paleocene. Near Stewartville, Mt. Diablo quad., Contra Costa Co. UCMP 1556. Holotype UCMP
11724. [Nuculanidae].
114 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Nuculana (Saccella) pulchrisinuosa (Clark, 1918). [Leda
Ramon Fm., Miocene. Near mouth of Markley Canyon, Mount Diablo quad., Contra Costa Co. UCMP 3081. Holo-
type UCMP 11109. [Nuculanidae].
Nuculana (Saccella ramonensis (Clark, 1918). [Leda]. Moore San Ramon

Nuculana taphria (Dall, 1896b). [Leda-
noma Co. UCMP B2062. Hypotype UCMP 31559. Remarks: In the genus Saccella,  Coan and Valentich-Scott
(2012: p. 74). [Nuculanidae].
Nuculana taphria (Dall, 1896b). [Leda
Ranch quad., Fresno Co. UCMP B6532. Hypotype UCMP 33331. Remarks: In the genus Saccella,  Coan and
Valentich-Scott (2012: p. 74). [Nuculanidae].
Nuculana (Saccella) taphria (Dall, 1896b). [Leda]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):150–151, pl. 27,
   
Co. LACMIP 305. Hypotypes LACMIP 4452–4456. Remarks: In the genus Saccella,  Coan and Valentich-Scott
(2012: p. 74). [Nuculanidae].
Nuculana taphria (Dall, 1896b). [Leda     
Miocene. Near Elsereno [sic], Los Angeles Co. Hypotype current whereabouts unknown. Remarks: In the genus
Saccella,  Coan and Valentich-Scott (2012: p. 74). [Nuculanidae].
Nuculana (Saccella) taphria (Dall, 1896b). [Leda 

e, g) SDSNH 77 [=SDSNH 4167]. Remarks: In the genus Saccella,  Coan and Valentich-Scott (2012: p. 74).
[Nuculanidae].
Nuculana temblorensis (Anderson and Martin, 1914). [Yoldia]. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):170,
Remarks: In the genus Yoldia, 
Moore (1983: p. A23). [Yoldiidae].
Nuculana (Saccella) uvasana (Dickerson, 1915). [Leda
Eocene. Live Oak Creek, Tejon quad., Kern Co. CASG 244. Holotype CASG 250, paratype CASG 251. [Nuculanidae].
Nuculana washingtonensis (Weaver, 1916). [Leda
Tumey Fm., Eocene/Oligocene. Monocline Ridge, Fresno Co. [UCMP D6533]. Hypotype UCMP 37165. Remarks:
In the subgenus Saccella,  Moore (1983: p. A17). [Nuculanidae].

Macoma pittsburgensis) (peel). Monocline Ridge, Fresno Co. UCMP D6522. Hypotype UCMP 14052.
Nuculana washingtonensis (Weaver, 1916). [Leda-
gen Fm., Oligocene. Monocline Ridge, Monocline Ridge quad., Fresno Co. UCMPD6533. Hypotype UCMP 37165.
Remarks: In the subgenus Saccella,  Moore (1983: p. A17). [Nuculanidae].
. 8D [peel, CASG]. Monocline Ridge, Monocline Ridge quad., Fresno Co. UCMP D6555]. Hypotype UCMP 37166.
Nuculana-
angle, Alameda Co. UCMP A4565. Hypotype UCMP 34122. [Nuculanidae].
Nuculana
Bulito, Pt. Conception quad., Santa Barbara Co. UCMP B6994. Hypotype SU 9247 [=CASG 70196]. [Nuculanidae].
Nuculana Leda
 Remarks: Determined here as Nuculana (Saccella) washingtonensis (Weaver,
1916). [Nuculanidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 115
Nuculana (Saccella-
istrano, Orange Co. Monterey Fm., Miocene. [Hypotype at CC]. [Nuculanidae].
Nuculana (Scaeoleda
      , C.
Throckmorton (personal communication, 2002). [Nuculanidae].
Nuttallia alata (Gabb, 1866). [Gari]. Moore 2003b. USGS PP 1228-G [electronic format] (family Psammobiidae):8–9,
Nuttalia alata in pl. caption]. San Pablo Fm., Miocene. East of Kirker Creek, north of Mt. Diablo,

Nuttallia howardi (Dickerson, 1915). [Tellina]. Moore 2003b. USGS PP 1228-G [electronic format] (family Psam-
Nuttalia howardi in pl. caption]. Tejon Fm., Eocene. Live Oak Canyon, Tejon quad.,
Kern Co. CASG 244. Holotype [CASG. 271 not UCMP 271 as cited]. [Psammobiidae].
Nuttallia jamesi-
cene [=Wilson Grove Fm., Miocene/Pliocene]. Near town of Trenton Sebatsopol quad. (1954), Sonoma Co. CASG
54164. Holotype CASG 54330 [=CASG 54164.01], paratypes CASG 54331, 54332 [=CASG 54164.02, 54164.03],



Nuttallia salina
Rico Fm., Miocene. Lynch Canyon, Salinas Valley, San Ardo quad., Monterey Co. USGS M2283. Holotype USNM

Nuttallia salina Roth and Guruswami-Naidu, 1978. Moore 2003b. USGS PP 1228-G [electronic format] (family
Nuttalia salina in pl. caption]. Pancho Rico Fm., Miocene. Lynch Canyon,
San Ardo quad., Monterey Co. USGS. M2283. Holotype USNM 242417. [Psammobiidae].
Nymphalucina panochensis Kiel , 2013 (Cima Sand-
stone lens, Paleocene. Cold-seep deposit, Escarpado Canyon, Panoche Hills, west side San Joaquin Valley. San
Benito Co. [no locality number given]. Paratype GZG.INV.88316. [Lucinidae].


Odontogryphaea haleyi (Hertlein, 1933). [Ostraea
Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4670. Hypotypes UCR 4670/32–4670/34.
Remarks: In the genus Phygraea,  Squires (2018a: p. 16). [Gryphaeidae].
Oppenheimopecten coalingensis (Arnold, 1906). [Pecten
Fm., Pliocene. Kreyenhagen Ranch on Zapata Chino Creek, Fresno Co. Lectotype (of Moore 1984: p. B76) SU/
CASG. [Pectinidae].
Oppenheimopecten juanensis (Grant and Stevenson, 1948). [Pecten]. Moore 1984. USGS PP 1228-B:B75–B76, pl. 33,

10441 [=LACMIP. 9137]. [Pectinidae].
Oppenheimopecten sanctaecruzensis (Arnold, 1906). [Pecten
Fm., Oligocene. West side of Salt Creek, Kern Co. UCMP 3231. Holotype (of Pecten dickersoni Wagner and Schil-
ling, 1923) UCMP 11416. [Pectinidae].

Cruz Co. SU 111. Holotype SU/CASG 360 [=CASG 61947.01].
Ostera* vespertina

116 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Hypotype UCLA 59406 [=LACMIP 8319]. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006:
p. 9). Error for Ostrea. [Ostreidae].
Ostrea altatemblorensis Grant and Eaton in
Miocene. Caliente Range, San Luis Obispo Co. UCLA 512 [=LACMIP 20512]. Holotype UCLA 8970 [=LACMIP 8953],
Remarks: Junior synonym
of Crassostrea ashleyi (Hertlein, 1934),  Moore (1987: p. C39). [Ostreidae].

Ostrea angelica
Fm., Pliocene. San Diego, San Diego Co. UCLA 303 [=LACMIP 20303]. Hypotype UCLA 4008 [=LACMIP 9294].
Remarks: Provisionally in the genus Dendostrea Moore (1987: p. C26). [Ostreidae].
Ostrea arnoldi
Domengine Ranch quad., Fresno Co. UCMP D1075. Holotype UCMP 36646, paratypes UCMP 36647, 36648. Re-
marks: Junior synonym of Crassostreaashleyi (Hertlein, 1934),  Moore (1987: p. C39). [Ostreidae].
Ostrea ashleyi
(1906), Kern Co. UCMP B1643. Hypotype UCMP 36565. Remarks: Provisionally in the genus Crassostrea
Moore (1987:p. C39). [Ostreidae].
Ostrea ashleyi
quad., Fresno Co. UCMP D1075. Hypotype UCMP 36651. Remarks: Provisionally in the genus Crassostrea
Moore (1987: p. C39). [Ostreidae].
Ostrea atwoodiPseudocardium
Zone), Pliocene. Arroyo Largo, North Dome, Kettleman Hills, Kings Co. USGS 12504. Hypotype USNM 495783.
Remarks: In the subgenus Ostrea, Moore (1987: p. C28). [Ostreidae].

12400. Hypotype USNM 495795.
Ostrea atwoodi          
Miocene/Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M1939. Hypotype USNM 649094.
Remarks: In the subgenus Ostrea,  Moore (1987: p. C28). [Ostreidae].
Ostrea atwoodi
Pliocene. Near Atascadero, San Luis Obispo Co. USGS M4621. Hypotypes USNM 416485, 647009. Remarks: Label

missing.. In the subgenus Ostrea,  Moore (1987: p.C28). [Ostreidae].
Ostrea (Ostrea) atwoodi

649094. [Ostreidae].
Ostrea bourgeoisii
Range, San Luis Obispo Co. UCLA 1820 [=LACMIP 21820]. Hypotype UCLA 8975 [=LACMIP 8967]. Remarks:
Provisionally in the genus Striostrea Moore (1987: p. C34). [Ostreidae].
Ostrea bourgeoisii 
Miocene. Kirker's Pass, west side of Markley Canyon, Mt. Diablo quad., Contra Costa Co. UCMP 2040. [Hypotype
Remarks: Provisionaly in the genus Striostrea Moore (1987:
p. C34). [Ostreidae].
Ostrea cierboensis Grant and Eaton in Eaton et al., 1941 Fm., Miocene.
Caliente Range, San Luis Obispo Co. UCLA 1751-
ured paratypes UCLA 8981, 8982] [=LACMIP 8969, 8970]. Remarks: Junior synonym of Crassostreaeucorrugata
(Hertlein, 1934),  Moore (1987: p. C39). [Ostreidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 117
Ostrea eldridgei-
cene. Ventura Co. CASG 27409. Hypotype CASG 5964. Remarks: Provisionally in the genus/subgenus Pycnodonte
(Crenostrea Moore (1987: p. C22). [Gryphaeidae].
Ostrea eldridgei  
Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 48 [=LACMIP 16297]. Hypotype UCLA 58199
[=LACMIP 10589]. Remarks: Provisionally in the genus/subgenus Pycnodonte Crenostrea Moore (1987:
p. C22). [Gryphaeidae].
Ostrea eldridgei-
fre Breccia, Miocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8764 [=SBMNH
60174]. Remarks: Provisionally in the genus/subgenus Pycnodonte (Crenostrea Moore (1987: p. C22).
[Gryphaeidae].
Ostrea eldridgei ynezana
Ostrea biostrome, Woody quad. (1952), Kern Co. UCMP B1654. Hypotype UCMP 36551. Remarks: A valid full
species in the genus/subgenus Pycnodonte (Pycnodonte),  Moore (1987: p. C18). [Gryphaeidae].
Ostrea eldridgei ynezana
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 53 [=LACMIP 16298]. Hypotype
UCLA 58203 [=LACMIP 10590]. Remarks: A valid full species in the genus/subgenus Pycnodonte (Pycnodonte),
 Moore (1987: p. C18). [Gryphaeidae].
Ostrea englekyi-
queros Fm., Miocene. Upper Cañada Posa, Santa Cruz Is., California Channel Ids. UCSB JK-FA65-106A [=LACMIP
16428]. Hypotype UCSB M10 [=LACMIP 6295]. Remarks: Provisionally in the genus Crassostrea Moore

Ostrea erici
(diatomaceous strata), Miocene. Graciosa Ridge near Orcutt, Santa Barbara Co. USGS 14903. Hypotype USNM
560035. Remarks: In the genus/subgenus Pycnodonte (Pycnodonte),  Moore (1987: p. C19). [Gryphaeidae].


Ostrea cf.   
Gregorio Member), Pliocene. Near mouth of Purisima Creek, Half Moon Bay quad., San Mateo Co. 696M. Hypo-
type determined here as missing. Remarks: Error for erici. In the genus/subgenus Pycnodonte (Pycnodonte),
 Moore (1987: p. C19). [Gryphaeidae].
Ostrea erici Hertlein, 1929a. Hertlein 4. San Diego Fm.,
Pliocene. San Diego Co. LACMIP 124. Hypotype LACMIP 4524. Remarks: In the genus/subgenus Pycnodonte
(Pycnodonte),  Moore (1987: p. C19). [Gryphaeidae].

Ostrea freudenbergi
Miocene. Domengine Ranch quad., Fresno Co. UCMP D1075. Hypotype UCMP 36653. Remarks: Provisionally in
the genus Striostrea Moore (1987: p. C34). [Ostreidae].
Ostrea haleyi
(1958), Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype LACMIP 7514. Remarks: In the genus Phygraea,
 Squires (2018a: p. 16). [Gryphaeidae].
Ostrea haleyi
Remarks:
In the genus Phygraea,  Squires (2018a: p. 16). [Gryphaeidae].
Ostrea hertleini   
118 PALEOBIOS, VOLUME 38, FEBRUARY 2021
-
types UCMP 36655, 36656. Remarks: Junior synonym of Crassostreaashleyi (Hertlein, 1934),  Moore (1987:
p. C39). [Ostreidae].
Ostrea howelli-
cene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 39 [=LACMIP 16301]. Hypotype UCLA 58198 [=LACMIP
10586]. Remarks: In the genus/subgenus Pycnodonte (Pycnodonte

UCLA 58193–58195 [=LACMIP 10583–10585].

58192 [=LACMIP 10582].

58202 [= LACMIP 10587].

58190 [=LACMIP 10581].
     
UCLA 58209 [=LACMIP 10588].
Ostrea idriaensis
Susana quad. (1951, PR 1969), Ventura Co. CSUN 458 [=LACMIP 16280]. Hypotype LACMIP 6572. Remarks: In
the genus Acutostrea,  Squires (2018a: p. 4). [Ostreidae].
Ostrea idriaensis Gabb, 1869b. Shapiro 1998. AAPG Guidebook MP 45:129, 
Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Is. Hypotype UCSB 8777 [=SBMNH 60175]. Re-
marks: In the genus Acutostrea,  Moore (1987: p. C31). [Ostreidae].
Ostrea iridescensOs-
trea iradescens]. Latrania Fm., Miocene/Pliocene. Painted Gorge, Coyote Mtn., Imperial Co. SDSNH 43. Hypotype
SDSNH 23101. Remarks: A synonym of Striostrea prismatica (Gray, 1825),  Coan and Kabat (2012: p. 303).
[Ostreidae].
Ostrea ligminuta Grant and Eaton in
Caliente Range, San Luis Obispo Co. UCLA 1795 [=LACMIP 21795]. Holotype UCLA 8985 [=LACMIP 8971], un-
Remarks: Junior synonym of Crassostrea titan (Conrad, 1853),
 Moore (1987: p. C36). [Ostreidae].

      
11042)].

Ostrea (Agerostrea) megodon
7. San Diego Fm., Pliocene. La Jolla quad., San Diego Co. UCLA 2420 [=LACMIP 22420]. Hypotypes UCLA [48614,
48650 [=LACMIP 9296, 9297)]. Remarks: Provisionally in the genus Ostreola Moore (1987: p. C29). In the
genus 
Ostrea simiensis-
cene. Concretions with abundant oyster debris, Calabasas quad., Simi Hills, Ventura Co. UCR 6899. Holotype
UCR 6899/102 (not UCR 6899/1 as in text), paratypes UCR 6899/100, 6899/101. Remarks: Junior synonym of
Phygraea haleyi (Hertlein, 1933),  Squires (2018a: p. 16). [Gryphaeidae].
Ostrea stewarti       
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 216 [=LACMIP 16299]. Hypotype LACMIP 7515. Remarks:
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 119
Provisionally in the genus/subgenus PycnodonteCrenostrea Moore (1987: p. C22). [Gryphaeidae].
Ostrea cf.   
Peak quad. (1958), Los Angeles Co. CSUN 364 [=LACMIP 16306]. Hypotype LACMIP 7516. Remarks: In the genus
Phygraea,  Squires (2018a: p. 18). [Gryphaeidae].
Ostrea tayloriana
Oligocene. Gaviota quad., Santa Barbara Co. UCMP B7030. Hypotype SU 9259 [=CASG 70204]. Remarks: Nomen
dubium,  Squires (2018a: p. 1).
Ostrea titan
Luis Obispo Co. UCLA 1766 [=LACMIP 21766]. Hypotype UCLA 8995 [=LACMIP 8979]. Remarks: Supplemen-
Crassostrea, 
Moore (1987: p. C35). [Ostreidae].
Ostrea titan Conrad, 1853. G.D. Hanna and Hertlein 1943. 
Miocene. Fresno Co. CASG 1282. Hypotype CASG 5961. Remarks: Supplementary descriptions in Conrad (1854,
In the genus Crassostrea,  Moore (1987: p. C35). [Ostreidae].
Ostrea titan

missing and presumed lost,  Moore (1987: p. C36). Remarks: Supplementary descriptions in Conrad (1854,
Crassostrea,  Moore (1987: p. C35). [Ostreidae].
Ostrea titan
of Twitchell Reservoir, Cuyama Valley, San Luis Obispo Co. Hypotype determined herea as missing. In the genus
Crassostrea, Moore (1987: p. C35). Remarks:
illustrated in Conrad (1857c). [Ostreidae].
Ostrea titan       
[Locality and source-specimen deposition institution unknown]. Remarks: Supplementary descriptions in
   Crassostrea,  Moore (1987: p.
C35). [Ostreidae].
Ostrea titan
Pt., Tejon Hills, Kern Co. USGS M1702. Hypotype USNM 649163. Remarks: Supplementary descriptions in Con-
Crassostrea,  Moore (1987: p. C35).
[Ostreidae].
Ostrea titan titan
ledge-forming Ostrea-Pecten 
Remarks:
genus Crassostrea,  Moore (1987: p. C35). [Ostreidae].
Ostrea titan  

p. 17). Santa Margarita Fm., Miocene. Tamiosoma Zone above Big Blue, Fresno Co. Hypotype USNM 165617].
Remarks:
genus Crassostrea,  Moore (1987: p. C35). [Ostreidae].
Ostrea titan
Miocene. Diablo Range, Fresno Co. [Hypotype SBMNH 60214]. Remarks: Supplementary descriptions in Con-
Crassostrea,  Moore (1987: p. C35).
[Ostreidae].
Ostrea titan andersoni
ledge-forming Ostrea-PectenRemarks: Junior synonym
of Crassostrea titan (Conrad, 1853),  Moore (1987: p. C36). [Ostreidae].

120 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Ostrea titan eucorrugata (Hertlein, 1934). [Ostrea
Miocene. Shell Peak, NE of Twitchell Reservoir, Cuyama Valley, San Luis Obispo Co. Hypotype determined here as
missing. Remarks: Full species provisionally in the genus Crassostrea Moore (1987: p. C39). [Ostreidae].
Ostrea titan subtitanMonterey Fm., Miocene.
Valley, Orange Co. SDSNH. 4312. Hypotype SDSNH 83238. Remarks: Junior synonym
of Crassostrea titan (Conrad, 1853),  Moore (1987: p. C36). [Ostreidae].
Ostrea titan var. prior Grant and Eaton in 
Miocene. Branch Canyon, Cuyama Valley, Santa Barbara Co. UCLA 1745 [=LACMIP 21745]. Holotype UCLA 8991
[=LACMIP 8977]. Remarks: Junior synonym of Crassostrea titan (Conrad, 1853),  Moore (1987: p. C36).
[Ostreidae].

Ostrea vaquerosensis
Ostrea biostrome, Woody quad. (1952), Kern Co. UCMP B1659. Hypotype UCMP 36552. Remarks: Provisionally
in the genus Crassostrea Moore (1987: p. C38). [Ostreidae].
Ostrea veatchii
Remarks: In the genus Myrakeena,
 Squires et al. (2006: p. 7). [Ostreidae].
Ostrea vespertinaPecten
Zone), Pliocene. southeast of El Bulto, North Dome, Kettleman Hills, Kings Co. USGS 12384. Hypotype USNM
495752. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9). [Ostreidae].
Ostrea vespertina
Near Camulos, Ventura Co. CASG 725. Hypotype CASG 5912. Remarks: Myrakeena veatchii (Gabb, 1866), 
Squires et al. (2006: p. 9). [Ostreidae].
Ostrea vespertina  

SDSNH 116]. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9). [Ostreidae].
Ostrea vespertina
Fm. (undifferentiated), Pliocene. Casmalia Hills, northeast of Shuman, Santa Barbara Co. USGS 14626. Hypotype
USNM 560102. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9). [Ostreidae].
Ostrea vespertina
[=Latrania Fm.], Pliocene. Painted Gorge area, Imperial Co. CASG 33277A. Hypotype CASG 12715]. Remarks:

four hypotypes at CASG, according to the online database. They cannot be matched, however, to any of the known
hypotypes and are thusly noted for the record as CASG 70278–70281. [Ostreidae].


Ostrea vespertina Conrad, 1853. Frazier 1968. Earth Sci. San
Diego Co. Hypotype current wherebouts unknown. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et
al. (2006: p. 9). [Ostreidae].
Ostrea vespertina     
Diego Fm., Pliocene. Tijuana River area, San Diego Co. CASG 36555. Hypotypes CASG. 72560, 72561]. Remarks:
Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9). [Ostreidae].

[=LACMIP 9295].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 121

Ostrea vespertina
Robles Fm., Pliocene. Near Atascadero, San Luis Obispo Co. USGS M4621. Hypotypes USNM 646995–646997,
647000, 647002, 647006, 647007. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9).
[Ostreidae].
Ostrea vespertina
Pliocene. Brown's Canyon, Los Angeles Co. UCLA 22517 [=LACMIP 22517]. Hypotype UCLA 49522 [=LACMIP
10621]]. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9). [Ostreidae].
Ostrea vespertina
Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2872. Hypotype USNM 254384. Remarks:
Determined here as Myrakeena veatchii (Gabb, 1866). [Ostreidae].
Ostrea vespertina

M2872. Hypotype USNM 254384. Remarks: Determined here as Myrakeena veatchii (Gabb, 1866). [Ostreidae].
Ostrea vespertina
Remarks: Determined here
as Myrakeena veatchii (Gabb, 1866). [Ostreidae].
Ostrea vespertina
SDSNH4080. SDSNH hypotype determined here as missing. Remarks: Myrakeena veatchii (Gabb, 1866), 
Squires et al. (2006: p. 9). [Ostreidae].
Ostrea vespertina
Pt. Fm. [=Vaqueros Fm.], Miocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype SBMNH
60176. Remarks: Determined here as Ostrea sp. [Ostreidae].
Ostrea vespertina Conrad loeli Hertlein, 1928a. Avila and Weaver 1969. AAPG/
8a, b. Rincon Fm., Miocene. East of Cañada Tecolote, north coast Santa Rosa Is., California Channel Ids. UCSB
SR67-51 [=LACMIP 16429]. Hypotype UCSB M11 [=LACMIP 6319]. Remarks: PycnodontePycnodonte loeli,
 Moore (1987: p. C21). [Gryphaeidae].
Ostrea vespertina sequens
Joaquin Fm. (Upper Mya Zone), Pliocene. La Ceja, North Dome, Kettleman Hills, Kings Co. USGS 12326. Hypotypes
USNM 495295–495297. Remarks: Myrakeena veatchii (Gabb, 1866), Squires et al. (2006: p. 9). [Ostreidae].
           
495298, 495299.
Mya and Acila Zone), Pliocene. Branch of Arroyo Seco, North
Dome, Kettleman Hills, Kings Co. USGS 12547. Hypotypes USNM 351764 (not USNM. 495307a as in pl. caption),
495308–495311.
Ostrea vespertina sequens
-
mined here as missing. Remarks: Myrakeena veatchii (Gabb, 1866),  Squires et al. (2006: p. 9). [Ostreidae].
Ostrea weaveri
Near Lower Lake, Lake Co. UCMP 781. Holotype UCMP 32624. [Ostreidae].
Ostrea wiedeyi Hertlein, 1928a. 
San Miguel Is., California Channel Ids. Specimen cited by Stenzel as being in the CASG collection of the San Fran-
cisco Academy of Sciences but could not be located and is determined here as presumably missing. Remarks:
Provisionally in the genus/subgenus PycnodontePycnodonte    Moore (1987: p.
C20). [Gryphaeidae].
122 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Ostrea wiedeyi
Saddleback Valley, Orange Co. SDSNH4312. Hypotype SDSNH 83234. Remarks: Provisionally in the genus/sub-
genus Pycnodonte (Pycnodonte Moore (1987: p. C20). [Gryphaeidae].
Ostrea sp. -
fornia Borderland, west of San Diego, San Diego Co. Remarks: Hypotype determined here as probably lost. This
oyster is Ostrea palmula Carpenter, 1857c, currently in the genus Saccostrea,  Moore (1987: p. C32). [Ostreidae].
Ostrea
Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13440. [Ostreidae].
Ostrea
Hypotype UCMP 34129. [Ostreidae].
Ostrea n. sp Susana Fm., Paleocene. Simi Hills, Ven-
tura Co. UCR.6899. Hypotype UCR 6899/101 (not UCR 6899/1 as in pl. caption). Remarks: Ostrea simiensis n.
sp.,  Zinsmeister 1983b; Phygraea haleyi (Hertlein, 1933),  Squires (2018a: p. 16). [Gryphaeidae].
Ostrea


   Ostrea atwoodi Mulina (Pseudocardium)
Zone”) [as Etchegoin Fm.], Miocene/Pliocene. North Dome, Kettleman Hills, Kings Co. Hypotype USNM 165531].
Ostrea-

Remarks: Determined here as Crassostrea titan (Conrad, 1853). [Ostreidae].
Ostrea     
Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to LACMIP. [Ostreidae].
Ostreola conchaphila (Carpenter, 1857b). [Ostrea       

Remarks: In the genus Ostrea,  Coan and Valentich-Scott (2012: p. 236). [Ostreidae].
Ostreolaventurana (Loel and Corey, 1932). [Ostrea
Vaqueros Fm., Oligocene/Miocene. Alisal Creek, Lompoc quad., Santa Barbara Co. UCMP A321. Holotype UCMP
31751, paratype UCMP 10035. [Ostreidae].
-
ity and hypotype deposition unknown. Remarks: Determined here as Pycnodonte heermanni (Conrad 1855a).
[Ostreidae].

Miocene/Pliocene. Fish Creek Wash, Imperial Co. Hypotypes disposition unknown. Remarks: Determined here
as Pycnodonte heermanni (Conrad, 1855a). [Ostreidae].
Pachecoa (Pachecoa) hornii (Gabb, 1864). [Arca
Eocene. Ft. Tejon, Kern Co. Paratype ANSP 4460a. Paratype missing,  K. Estes-Smargiassi (personal commu-
nication, 2019). [Arcidae].
Pachecoa (Pachecoa) hornii elusa (Clark and Woodford, 1927). [Arca]. Moore 1983. USGS PP 1228-A:A48, pl. 10,
    

Pachydesma crassatelloides (Conrad, 1837). [Cytherea (Trigonella)]. Woodring and Bramlette 1950 [1951]. USGS
    
Co. USGS 14752. Hypotype USNM 560147. Remarks: Junior synonym of Tivela (Pachydesma) stultorum (Mawe,
1823),  Coan et al. (2000: p. 379). [Veneridae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 123
Pachydesma inezana Conrad   . 5. Upper Gaviota Fm.,
Eocene. Cañada de Santa Anita area, Lompoc quad., Santa Barbara Co. UCMP B7021. Hypotype SU 9293 [=CASG
70203]. [Veneridae].

Pachypernagoniglensis (M.A. Hanna, 1927). [Perna
Ardath Shale, Eocene. La Jolla quad., San Diego Co. UCMP 4233. Holotype UCMP 30912. [Isognomonidae].
Pachypernajoaquinensis (Vokes, 1939). [Pedalion  
Fm., Eocene. Near San Joaquin coal mine, Coalinga quad., Fresno Co. UCMP 7177. Holotype UCMP 15577. [Isog-
nomonidae].
Pacipecten andersoni andersoni (Arnold, 1906) (not Anderson as in pl. caption). [Pecten]. Moore, 1984. USGS PP

[Pectinidae].
Pacipecten andersoni (Arnold, 1906). [Pecten
Fm. (Saddleback Valley Limestone member). Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Cap-
istrano quad. SDSNH 4312. Hypotype SDSNH 140181. [Pectinidae].
Pacipecten andersoni barkerianus (Arnold, 1906). [Pecten-

 Moore (1984: p. B45). [Pectinidae].

Chlamys barkeriana) SU/CASG 6588.
Pacipecten andersoni subandersoni (Loel and Corey, 1932). [Pecten
6. Vaqueros Fm., Oligocene/Miocene. San Luis Obispo Co. UCMP A572. Holotype UCMP 31741. [Pectinidae].
Pacipecten bilineatus (Clark, 1915). [Pecten
Near town of Rodeo, Napa quad., Contra Costa Co. UCMP 1632. Holotype UCMP 11579. [Pectinidae].
Pacipecten discus (Conrad, 1857d). [Pecten   
Group, Miocene. South side of Mt. Diablo, Mt. Diablo quad., Contra Costa Co. UCMP 1492. Holotype (of Pecten
raymondi Clark, 1915) UCMP 11581, paratype (of Pecten raymondi Clark, 1915) UCMP 12371. [Pectinidae].
Pecten (Pecten)
andersoni gonicostatus Trask, 1922) UCMP 12370.
Pecten (Pecten) cier-
boensis Clark, 1915) UCMP 11585.
Pleasanton quad., Alameda Co. UCMP 3532. Holotype (of Pecten (Pecten) raymondi brionianus Trask,
1922) UCMP 12368.

Holotype USNM 13335.
Pacipecten discus (Conrad, 1857d). [Pecten

Pacipecten pabloensis (Conrad, 1857c) of Arnold. [Pecten

and presumed lost,  Moore (1984: p. B48) SU/CASG 302, 302a [=CASG 69655]. [Pectinidae].
Pacipecten tolmani (Hall and Ambrose, 1916). [Pecten
Miocene. Vicinity of Welch Creek, Pleasanton quad., Alameda Co. CASG. 61866. Holotype SU/CASG 23 [=CASG
61866.01]. [Pectinidae].
124 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pacipecten weaveri (Clark, 1915). [Pecten
San Pablo Bay area, Napa quad., Contra Costa Co. UCMP 407. Syntypes UCMP 11575, 11576. [Pectinidae].
Palliolum (Delectopecten) pechami* (Gabb, 1869a). [Pecten
Vida Member), Miocene. Hypotype current whereabouts unknown. Remarks: Ex Chapman College. In the genus
Delectopecten,  Moore (1984: p. B15). Error for [Pectinidae].
Pandora (Pandorella) bilirata
-
MIP 305. Hypotypes LACMIP 4835, 4836]. Remarks: Supplemantal description in Conrad (1855b). [Pandoridae].
Pandora cf.  (Carpenter, 1864b). [Kennerlia]. Woodring and Bramlette 1950 [1951]. USGS PP 222:91, pl. 8,

USGS 15020. Hypotype USNM 560040. Remarks: In the subgenus Pandorella  
[Pandoridae].
Pandora punctata
Fm. (Graciosa Member), Pliocene. Graciosa Ridge, near Orcutt, road cut, Santa Barbara Co. USGS 14614. Hypotype
USNM 560150. Remarks: Heteroclidus punctatus,  Coan and Valentich-Scott (2012: p. 964). [Pandoridae].
Pandora cf.  
(Pseudocardium Zone), Pliocene. Arroyo Conchoso, North Dome, Kettleman Hills, Kings Co. USGS 12524. Hypotype
USNM 495786. Remarks: Heteroclidus punctatus,  Coan and Valentich-Scott (2012: p. 964). [Pandoridae].
Pandora (Heteroclidus) punctata
San Diego Fm., Pliocene. Balboa Park, San Diego, San Diego Co. UCLA 1386 [=LACMIP 21386]. Hypotypes UCLA
48638, 48639 [=LACMIP 9321, 9322]. Remarks: Heteroclidus punctatus,  Coan and Valentich-Scott (2012:
p. 964). [Pandoridae].
Beach, San Diego Co. SDSNH 21.
Hypotype SDSNH 4344.
Pandora punctata
Cruz Co. CASG66163. Hypotype CASG 66163.01]. Remarks: Heteroclidus punctatus,  Coan and Valentich-Scott
(2012: p. 964). [Pandoridae].
Pandora (Heteroclidus) punctata
area, Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14373. Remarks: Heteroclidus punctatus,  Coan and
Valentich-Scott (2012: p. 964). [Pandoridae].


Panomya beringiana
Humboldt Co. UCMP B7879. Hypotype UCMP 37793. Remarks: Junior synonym of Panomya priapus (Tilesius
1822),  Coan et al. (2000: p. 489). [Hiatellidae].
Panomya cf. 
Diego Fm., Pliocene. San Diego, San Diego Co. CASG 1418. Hypotype CASG 70444. Remarks: Junior synonym of
Panomya priapus (Tilesius 1822),  Coan et al. (2000: p. 489). [Hiatellidae].
Panope (Panope) abrupta (Conrad, 1849a). [Mya
San Diego Fm., Pliocene. Balboa Park, San Diego, San Diego Co. SDSNH 1965. Hypotype SDSNH 4120. Remarks:
In the genus Panopea,  Coan et al. (2000: p. 490). Pliocene through Recent specimens of Panopea are inferred
here as Panopea generosa Gould, 1850,  P. Valentich-Scott (personal communication, 2019 as interpreted from
Vadopalas et al. 2010). [Hiatellidae].
Panopea abrupta (Conrad, 1849a). [Mya
Latrania Fm., Miocene/Pliocene. Alverson Canyon, Imperial Co. SDSNH 345. Hypotype SDSNH 12101. [Hiatellidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 125
Panopea abrupta (Conrad, 1849a). [Mya
(Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4312. Hypotype SDSNH 83237. [Hiatellidae].
Panopea abrupta (Conrad, 1849a). [Mya
area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14371. Remarks: Pliocene through Recent specimens
of Panopea are inferred to be  Gould, 1850, P. Valentich-Scott (personal communication, 2019 as
interpreted from Vadopalas et al. 2010). [Hiatellidae].
Panope cf.  Gould, 1850. Woodring et al. 1940 [1941]. USGS PP 195:chart opposite p. 38, pl
San Joaquin Fm. (Pecten zone), Pliocene. Near Arroyo Pequeño and Arroyo Finito, North Dome, Kettleman Hills,
Kings Co. USGS 12343. Hypotype USNM 495754. Remarks: Pliocene through Recent specimens of Panopea are
inferred to be  Gould, 1850,  P. Valentich-Scott (personal communication, 2019 as interpreted from
Vadopalas et al. 2010). [Hiatellidae].
Panope generosa
Line drawing, source specimen unknown. Remarks: Junior synonym of Panopea abrupta (Conrad, 1849a), 
Coan et al. (2000: p. 490). [Hiatellidae].
Panope (Panope) generosa         
Remarks: Junior synonym
of Panopea abrupta (Conrad 1849a),  Coan et al. (2000: p. 490). Miocene specimens of Panopea are inferred
to be   (Conrad, 1849a),  P. Valentich-Scott (personal communication, 2019 as interpreted from
Vadopalas et al. 2010). [Hiatellidae].
Panopea generosa        
Diego Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 4831. Remarks: Junior
synonym of Panopea abrupta (Conrad, 1849a),  Coan et al. (2000: p. 490). Miocene specimens of Panopea are
inferred to be  (Conrad, 1849a),  P. Valentich-Scott (personal communication, 2019 as interpreted
from Vadopalas et al. 2010). [Hiatellidae].
Panopea generosa
Cruz Co. CASG 66079. Hypotype CASG 66079.01]. Remarks: Junior synonym of Panopea abrupta (Conrad 1849a),
 Coan et al. (2000: p. 490). Miocene specimens of Panopea are inferred to be  (Conrad, 1849a), 
P. Valentich-Scott (personal communication, 2019 as interpreted from Vadopalas et al. 2010). [Hiatellidae].
Panopea ramonensis (Clark, 1925). [Panope        
(Wygal Sandstone Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypo-
type USNM 646612. [Hiatellidae].
Panopea ramonensis (Clark, 1925). [Panope 
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 338 [=LACMIP 16277]. Hypotype
UCLA 58215 [=LACMIP 10956]. [Hiatellidae].
Panope tenuis
Garza Creek area, Reef Ridge quad., Kings Co. UCMP B6518. Hypotype UCMP 36777. Remarks: Junior synonym
of Panopea abrupta (Conrad, 1849a),  Coan et al. (2000: p. 490). Miocene specimens of Panopea are inferred
to be   (Conrad, 1849a),  P. Valentich-Scott (personal communication, 2019 as interpreted from
Vadopalas et al. 2010). [Hiatellidae].
Panopea tenuis
Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH
4312. Hypotype SDSNH 140185. Remarks: Junior synonym of Panopea abrupta (Conrad, 1849),  Coan et al.
(2000: p. 490). Miocene specimens of Panopea are inferred to be  (Conrad, 1849a),  P. Valentich-
Scott (personal communication, 2019 as interpreted from Vadopalas et al. 2010). [Hiatellidae].
Panopea
Remarks: Determined here as Panopea abrupta (Conrad, 1849a) based on
the inference that Miocene specimens of Panopea are  (Conrad, 1849a), P. Valentich-Scott (personal
communication, 2019 as interpreted from Vadopalas et al. 2010). [Hiatellidae].
126 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Panopea generosa (Gould, 1850)). W-K Hill, north of Coalinga, Fresno Co. Hypotype USNM
165556]. Remarks: Determined here as Panopea abrupta (Conrad, 1849a) based on the inference that Miocene
specimens of Panopea are  (Conrad, 1849a),  P. Valentich-Scott (personal communication, 2019 as
interpreted from Vadopalas et al. 2010).
Parvamussium alaskensis (Dall, 1871). [Pecten
Fm., Pliocene. Fourth and Broadway, Los Angeles, Los Angeles Co. UCMP 3030. Holotype UCMP 11084, paratype
UCMP 11779 (both of Pecten calamitus M.A. Hanna 1924). Remarks: Parvamussium alaskense,  Coan et al.
(2000: p. 251). [Propeamussiidae].
Parvamussium lajollaense (M.A. Hanna, 1927). [Pecten-
dath Shale, Eocene Soledad Valley area, La Jolla Quad., San Diego Co. UCMP 5062. Syntypes UCMP 31106, 31108.
[Propeamussiidae].
Parvamussium mideocenicum (Vokes, 1939). [Propeamussium
Fm. (Cerros Shale Member), Paleocene [Eocene]. Vicinity of where Urruttia Canyon enters Salt Creek, Coalinga
quad., Fresno Co. UCMP 1817. Syntype UCMP 15584. [Propeamussiidae].
Parvamussium riversi (Arnold, 1906). [Pecten
Fm., Pliocene. Santa Monica Canyon, Los Angeles Co. Holotype SU/CASG 33 [=CASG 61829.01]. Remarks: Junior
synonym of Parvamussium alaskense (Dall, 1871),  Coan et al. (2000:p. 251). [Propeamussiidae].
Parvamussium stanfordense (Arnold, 1906). [Pecten  
Fm., Eocene. Los Trancos Creek, near Stanford University, San Mateo Co. USGS M1492. Hypotype (topotype) (of


Salt Creek, Coalinga quad., Fresno Co. UCMP 1817. Syntype (of Parvamussium mideocenicum Vokes, 1939, pl. 3,


CASG 12 [=CASG 61862.01].
Parvamussium
Mtn. quad. (1943), Ventura Co. CSUN 983 [=LACMIP 16110]. Hypotype LACMIP 7728. [Propeamussiidae].
Parvamussium
(1995), Los Angeles Co. LACMIP 17850. Hypotype LACMIP 13441. [Propeamussiidae].
Parvilucina (Parvilucina) tenuisculpta intensa (Dall, 1874). [Lucina
17, 22. San Diego Fm., Pliocene. Near U.S./Mexico boundary, San Diego Co. LACMIP 318. Hypotype (of Hertlein and
Remarks: Junior synonym of Parvilucina approximata (Dall, 1901b),
Coan et al. (2000: p. 265). [Lucinidae].

4645.
Parvilucina (Parvilucina) tenuisculpta tenuisculpta (Carpenter, 1864b). [Lucina]. Moore 1988b. USGS PP 1228-D:D14-
-
type (of Phacoides tenuisculptaRemarks: Junior synonym of Parvilucina
approximata (Dall, 1901b),  Coan et al. (2000: p. 265). [Lucinidae].
Patinopecten calaverasensis (Hall, 1958). [Pecten
(Briones Fm. in locality description), Miocene La Costa Valley quad. (1953), Contra Costa Co. SU 3252. Holotype
SU/CASG 8442. [Pectinidae].
Patinopecten caurinus (Gould, 1850). [Pecten
(of Ogle 1953:24), Pliocene/Pleistocene. Dredged from Eel Submarine Canyon (40.6°N, 124.5°W), Humboldt Co.
USGS M5212. Hypotype USNM 647086. [Pectinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 127
Patinopecten dilleri (Dall, 1901a), var. [Pecten]. Woodring and Bramlette 1950 [1951]. USGS PP 222:84, 85, pl. 9,
      
USGS 14878. Hypotypes USNM. 560030–560032. [Pectinidae].

USNM 560053.
Patinopecten (Lituyapecten) falorensis     

A4233. Holotype UCMP 34172 (right valve), 34171 (left valve), hypotype UCMP 31333. Remarks: In the genus
Lituyapecten,  Moore (1984: p. B86). [Pectinidae].


70096], but unable to match specimens here or below.

Patinopecten haywardensis (Lutz, 1951). [Pecten]. Moore 1984. USGS 
Sobrante Fm., Miocene. Near junction of Hollis Creek and Eden Creek, Hayward quad., Alameda Co. UCMP A4566.
Holotype UCMP 34133 (not UCMP 341133 as in pl. caption), paratype UCMP 34134. [Pectinidae].
Patinopecten healeyi (Arnold, 1906). [Pecten

USNM 560136, 560137. [Pectinidae].
Patinopecten healeyi (Arnold, 1906). [Pecten
Pliocene. Northeast Bank, southern California Borderland, 230 km west of San Diego, San Diego Co. Hypotype
determined here as presumably lost. [Pectinidae].
Patinopecten healeyi (Arnold, 1906). [Pecten
8. Unnamed strata, Pliocene. Near Wilson Cove, San Clemente Is., California Channel Ids. USGS M6503. Hypotypes
UNSM. 235590–235593, 235595. [Pectinidae].

Patinopecten healeyi (Arnold, 1906). [Pecten

5219, 5238–5246. [Pectinidae].



5227, 5228, 5250, 5252–5255.


CASG hypotypes determined here as missing.




235595.

128 PALEOBIOS, VOLUME 38, FEBRUARY 2021
-
potypes USNM 249742, 249743, 249745.
El Toro, San Juan Capistrano quad. (1949),
Orange Co. USGS M2096. Hypotypes USNM 249744, 249751, 249755.

Orange Co. USGS M2753. Hypotypes USNM 249746–249748, 249750.

Hypotypes LACMIP 5229, 5233.

122. Hypotypes LACMIP 5230, 5236, 5237, 5247–5249, 5251.

5234, 5235.

USGS M2098. Hypotype USNM 249749.


Patinopecten healeyi (Arnold, 1906). [Pecten


5228, 5250, 5253, 5254. [Pectinidae].


    




5229.

Patinopecten healeyi (Arnold, 1906). [Pecten]. Groves 1991b. Amer. Conch. 
Pliocene. Santa Susana Mtns., Ventura Co. CSUN 725 [=LACMIP 29227. Hypotype LACMIP 11614. [Pectinidae].
Patinopecten healeyi (Arnold, 1906). [Pecten
Pecten (Patinopecten) healeyi sanclementensis Susuki and Stadum,
1978)). Saugus Fm. [=Pico Fm.] Pliocene. North side Simi Valley, Ventura Co. CSUN725 [=LACMIP 29227]. Hypo-
type UCLA 59405 [=LACMIP 8318]. [Pectinidae].
Patinopecten healeyi (Arnold, 1906). [Pecten
area, Los Angeles Co. LACMIP 9659. Hypotype LACMIP 14348. [Pectinidae].
Patinopecten healeyi
Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305B (not 305 as published) [=LACMIP
16817]. Hypotype LACMIP. 14854. [Pectinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 129
Patinopecten healeyi (Arnold, 1906). [Pecten]. Specimen illustrated on the front cover of the present paper. Pico
Fm., Pliocene. Gillibrand Quarry, north side Simi Valley, Ventura Co. LACMIP 40725. Hypotype LACMIP 14870.
[Pectinidae].
Patinopecten lohri (Hertlein, 1928b). [Pecten        
Etchegoin Fm. (Patinopecten zone), Pliocene. North Dome, Kettleman Hills, Kings Co. USGS 12428a. Hypotype
USNM 495839. [Pectinidae].


       
495842.
Patinopecten lohri (Hertlein, 1928b). [Pecten]. Woodring and Bramlette 1950 [1951]. USGS PP 222:83–84, pl. 7,

Barbara Co. USGS 14965. Hypotype USNM 560016. [Pectinidae].

Patinopecten lohri (Hertlein, 1928b). [Pecten
Rico Fm., Miocene/Pliocene. Long Valley, Monterey Co. Hypotype SU 9753 [=CASG 70324]. [Pectinidae].

Patinopecten lohri (Hertlein, 1928b). [Pecten
Coalinga quad., Fresno Co. UCMP D1166. Hypotype UCMP 36632. [Pectinidae].
Patinopecten lohri (Hertlein, 1928b). [Pecten 
Pliocene. Los Gatos Creek area, Fresno Co. USGS M3782. Hypotype USNM 647087. Remarks: Label in USNM

Patinopecten lohri (Hertlein, 1928b). [Pecten
Miocene/Pliocene. Metz quad., Monterey Co. UCMP A3161. Hypotype UCMP 36467. [Pectinidae].
Patinopecten lohri (Hertlein, 1928b). [Pecten      
(Tinaquaic Sandstone Member), Pliocene. Near Foxen Canyon, Santa Barbara Co. Holotype UCMP 12081. [Pec-
tinidae].
Patinopecten lohri (Hertlein, 1928b). [Pecten
3. Sisquoc Fm. (Tinaquaic Sandstone Member), Pliocene. Santa Barbara Co. Holotype UCMP 12081. [Pectinidae].
Patinopecten lohri (Hertlein, 1928b). [Pecten
Sandstone Member), Pliocene. Santa Barbara Co. Holotype UCMP 12801. [Pectinidae].
Patinopecten owenii (Arnold, 1906). [Pecten
Remarks: Junior synonym of Patinopecten lohri
(Hertlein, 1928b),  Moore (1984: p. B80). [Pectinidae].
....., p. 30 (as Patinopecten
Pecten (Patinopecten) oweni)). Etchegoin Fm., Miocene.
Santa Barbara Co. Hypotype UCMP 12081].
Patinopecten purisimaensis (Arnold, 1906). [Pecten
Pliocene. Annapolis quad., Sonoma Co. UCMP B2063. Hypotype UCMP 31558. Remarks: In the genus Lituyapec-
ten,  Moore (1984: p. B87). [Pectinidae].
Patinopecten (Lituyapecten) purisimaensis (Arnold, 1906). [Pecten
1, 3. Purisima Fm., Pliocene. San Gregorio Lagoon area, San Mateo Co. UCMP A4417. Hypotype UCMP 15181.
130 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Remarks: In the genus Lituyapecten,  Moore (1984: p. B87). [Pectinidae].
Patinopecten turneri (Arnold, 1906). [Pecten
Pliocene. Sebatsopol quad., Sonoma Co. UCMP A6894. Hypotypes UCMP 35533, 35534. Remarks: In the genus
Lituyapecten,  Moore (1984: p. B87). [Pectinidae].
Pecten andersoni
Miocene. Kern River, Kern Co. CASG 2023. Hypotypes CASG 5739, 5739A. Remarks: In the genus Pacipecten, 
Moore (1984: p. B45). [Pectinidae].
Pecten (Plagioctenium) andersoni Arnold, 1906. Shimer and Shrock, 1944. Index Foss. N.A., p. 407,  
Fresno Co.
USGS 4859. Hypotype USNM 165583]. Remarks: In the genus Pacipecten,  Moore (1984: p. B45). [Pectinidae].

Cruz Co. USGS 4803. Holotype USNM 164932].
Pecten andersoni
Remarks: In the genus Pacipecten,  Moore
(1984: p. B45) [Pectinidae].
Pecten bellus (Conrad, 1857a). [Janira
Harbor Fm., Pliocene. Vicinity of Potato Harbor, Santa Cruz Is., California Channel Ids. UCSB GM65-21 [=LACMIP
16425]. Hypotype UCSB M12 [=LACMIP 6307]. Remarks: Supplemental description in Conrad (1857c). Euvola
bella,  Waller (2007: p. 940). [Pectinidae].
Pecten (Pecten) bellus (Conrad, 1857a). [Janira

Remarks: Supplemental description in Conrad (1857c). Euvola bella,  Waller (2007: p. 940). [Pectinidae].
 Keen
and Bentson 1944: p. 82]. CASG 526b (all paratypes of Pecten hemphilli Dall, 1878a) determined here as missing.

4480.
Pecten (Pecten) bellus (Conrad, 1857a). [Janira

Remarks: Supplemental description in Conrad (1857c). Euvola bella,  Waller (2007: p. 940). [Pectinidae].
Pecten (Pecten) bellus (Conrad, 1857a). [Janira
1, 3, 5. Unnamed strata, Pliocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS
M6501. Hypotypes USNM 235597, 235598. Remarks: Supplemental description in Conrad (1857c). Euvola bella,
 Waller (2007: p. 940). [Pectinidae].

56316].
Pecten (Pecten) bellus (Conrad, 1857a). [Janira
strata, Pliocene. San Clemente Is. North quad., San Clemente Is., California Channel Ids. UCLA 6316 [=LACMIP
26316]. Hypotype UCLA 38786 [=LACMIP 10833]. Remarks: Supplemental description in Conrad (1857c). Eu-
vola bella,  Waller (2007: p. 940). [Pectinidae].
Pecten bellus (Conrad, 1857a). [Janira].
South side of Santa Clara River, between Piru and Fillmore, Ventura Co. SCPS 103B. Line drawing, source specimen
unknown. Remarks: Meals’ collection now at LACMIP. Remarks: Supplemental description in Conrad (1857c).
Euvola bella,  Waller (2007: p. 940). [Pectinidae].
Pecten (Pecten) bellus (Conrad, 1857a). [Janira].     2.
Saugus Fm. [=Pico Fm.], Pliocene. Northern Simi Valley, Santa Susana quad. (1951, PR1969), Ventura Co. CSUN
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 131
725 [=LACMIP 29227]. Hypotype UCLA 59404 [=LACMIP. 8317]. Remarks: Supplemental description in Conrad
(1857c). Euvola bella,  Waller (2007: p. 940). [Pectinidae].
Pecten (Pecten) bellus (Conrad, 1857a). [Janira
Fm., Pliocene. San Diego, San Diego Co. Holotype (of Pecten (Pecten) hemphilli Dall, 1878) USNM 7943. Remarks:
Supplemental description in Conrad (1857c). Euvola bella,  Waller (2007: p. 940). [Pectinidae].
Pecten (Pecten) bellus (Conrad, 1857a). [Janira

CSUN725 (=LACMIP 29227). Hypotype UCLA 59404 (=LACMIP 8317)]. Remarks: Euvola bella,  Waller (2007:
p. 940). [Pectinidae].
Pecten (Pecten) bellus hemphilli          
Remarks: Junior synonym
of Pecten (Pecten) bellus (Conrad, 1857a),  Hertlein and Grant 1972: p. 175,  Moore (1984: p. B68). Euvola
bella,  Waller (2007: p. 940). [Pectinidae].
Pecten cf. beringianus    
Orange Co. SDSNH 4080. SDSNH hypotype determined here as missing. Remarks: Chlamys behringiana (Mid-
dendorff 1849),  Coan et al. (2000: p. 233). [Pectinidae].
Pecten (Vertipecten) bowersi-
ros Fm., Miocene. Temblor Range, Kern Co. Hypotype CASG 7739. Remarks: In the genus Vertipecten,  Moore
(1984: p. B62). [Pectinidae].
Pecten (Chlamys) branneri Arnold, 1906 (n. sp.). USGS PP 47:55. Vaqueros Fm., Miocene. Tuff Hill, near Stanford
ex: SU, ex: CIT 4668]. Remarks: In the genus
Chlamys,  Moore (1984: p. B19). [Pectinidae].
Pecten branneri  
Santa Clara Co. Hypotype CASG 12314. Hypotype determined here as missing. Remarks: In the genus Chlamys,
 Moore (1984: p. B19). [Pectinidae].
Pecten caurinus
96 ftms., southwest of Trinidad Head, Humboldt Co. CASG 33223. Hypotype CASG 4284 (with Compsomyax sp.).
Remarks: In the genus Patinopecten,  Moore (1984: p. B83). [Pectinidae].
Pecten (Lyropecten) cerrosensis
Pliocene. Chula Vista, San Diego Co. SDSNH2287. Hypotype SDSNH 4119]. Remarks: In the genus Lyropecten,
 Moore (1984: p. B55). [Pectinidae].
Pecten coalingaensis Arnold, 1906. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):
(Pecten coalingaensis Zone), Pliocene. North Dome, Kettleman Hills, Kings Co. CASG 698. Hypotype CASG 5946.
Remarks: In the genus Oppenheimopecten,  Moore (1984: p. B76). [Pectinidae].
Pecten coalingaensis
      
[=LACMIP 9136]. Remarks: In the genus Oppenheimopecten,  Moore (1984: p. B76). [Pectinidae].
Pecten (Pecten) coalingaensis
Reef Ridge quad., Kings Co. UCMP B6539. Hypotype UCMP 36613. Remarks: In the genus Oppenheimopecten,
 Moore (1984:p. B76). [Pectinidae].

Pecten coalingensis*
(Pecten 
495727. Remarks: In the genus Oppenheimopecten,  Moore (1984: p. B76). Error for coalingaensis. [Pectinidae].

132 PALEOBIOS, VOLUME 38, FEBRUARY 2021


Hypotype USNM 495743.

Pecten coalingaensis).
Pecten coalingensis*

Fm.], Pliocene. Zapato Creek area, Fresno Co. Hypotype USNM 165530. Remarks: In the genus Oppenheimopecten,
 Moore (1984: p. B76). Error for coalingaensis. [Pectinidae].
Pecten (Patinopecten) aff.   ( ) coosensis

UCMP 37616. Remarks: May be Lituyapecten dilleri (Dall, 1901),  Moore (1984: p. B88). [Pectinidae].
Pecten (Lyropecten) crassicardo (Conrad, 1857a). [Pallium]. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):172,
Remarks: In the genus
Lyropecten,  Moore (1984: p. B51). [Pectinidae].
Pecten crassicardo (Conrad, 1857a). [Pallium]. Bereskin and Edwards 1969. AAPG/SEPM Guidebook, p. 77, pl. 32,
Lyropecten in text]. Rincon Fm., Miocene. Upper Cañada Posa, Santa Cruz Is., California Channel Ids. UCSB
RB65-59 [=LACMIP 16430]. Hypotype UCSB M13 [=LACMIP 6301]. Remarks: Lyropecten submiguelensis (Loel
and Corey, 1932) ,  Smith 1991a: p. 64, [=Lyropecten pretiosus (Hertlein, 1925)],  Smith (1991a: p.
64). [Pectinidae].
Pecten crossicardo* (Lyropecten) (Conrad, 1857a). [Pallium]. Fowkes 1982. Coalinga Guidebook, p. 199, unnumbered
 north of Coalinga, Fresno Co. WHC
Remarks: In the genus Lyropecten,  Moore (1984: p. B51). Error for crassicardo. [Pectinidae].
Pecten (Patinopecten) dilleri
area, Scotia quad., Humboldt Co. UCMP B7644. Hypotype UCMP 37760. Remarks: In the genus Lituyapecten, 
Moore (1984: p. B88). [Pectinidae].
Pecten (Lituyapecten) dilleri
San Diego Fm., Pliocene. San Diego, San Diego Co. UCLA 309 [=LACMIP 20309]. Hypotype UCLA 8072 [=LACMIP
9292]. Remarks: In the genus Lituyapecten,  Moore (1984: p. B88). [Pectinidae].
Pecten eldridgei (Aequipecten sircularis [sic] eldridgei
Fm.], Pliocene. South of McKittrick, Kern Co., Hypotypes Kern County Schools Collection, determined here as
missing. Remarks: Determined here as Argopecten ventricosus eldridgei (Arnold, 1906). [Pectinidae].
Pecten (Lyropecten) estrellanus (Conrad, 1857a). [Pallium
63-7. Santa Margarita Fm., Miocene. Cholame Hills, Monterey Co. CASG 28473. Hypotype CASG 5900. Remarks:
In the genus Lyropecten,  Moore (1984: p. B53); Argopecten sp.,  Smith (1991a: p. 56). Described as Lyro-
pecten terrysmithae n. sp.,  Powell et al. (2020: pp. 2–7). [Pectinidae].
Pecten (Lyropecten) estrellanus (Conrad, 1857a). [Pallium]. Shimer and Shrock 1944. Index Foss. N.A., p. 407, pl.

Monterey Co. USGS 3586. Hypotype USNM 164851]. Remarks: Supplemental descriptions in Conrad (1857c, d)
Lyropecten,  Moore (1984: p. B53). [Pectinidae].
Pecten (Lyropecten) estrellanus (Conrad, 1857a). [Pallium
Miocene/Pliocene. Line drawing, source specimen unknown. Remarks: Supplemental descriptions in Conrad
Lyropecten,  Moore (1984: p. B53). [Pectinidae].
Pecten (Lyropecten) estrellanus (Conrad, 1857a). [Pallium     -

 Keen and Bentson (1944: p. 85). Remarks:
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 133
Lyropecten,
 Moore (1984: p. B53). [Pectinidae].
Pecten estrellanus (Lyropecten) (Conrad, 1857a). [Pallium]. Fowkes 1982. Coalinga Guidebook, p. 199, unnumbered

Remarks:
In the genus Lyropecten,  Moore (1984: p. B53). [Pectinidae].
Pecten (Lyropecten) estrellanus terminus Arnold, 1906. Shimer and Shrock 1944. Index Foss. N.A., p. 407, pl. 161,
Fm., Miocene. Terraces on east side of Sali-
nas Valley or in tributary canyons to the valley between King City and San Lucas, Monterey Co. Holotype UCMP
11662]. Remarks: Lyropecten terminus Arnold, 1906,  Moore (1984: p. B54). [Pectinidae].
Pecten etchegoini
Pliocene. CASG. 2032. Hypotype CASG 5945 (designated as lectotype of Pecten wattsi Arnold 1906, which was lost
 G D. Hanna and Hertlein (1943: p. 176), from approximately same
locality). Remarks: Swiftopecten parmeleei etchegoini Anderson, 1905,  Moore (1984: p. B60). [Pectinidae].
Pecten (Swiftopecten) etchegoini Anderson subsp. wattsi
Rio Dell Fm., Pliocene. Scotia quad., Humboldt Co. UCMP B7642. Hypotype UCMP 37762. Remarks: Nanochlamys
nutteri (Arnold, 1906),  Moore (1984: p. B31). [Pectinidae].
Pecten hastatus
Creek area, Ft. Ross quad., Sonoma Co. UCMP B2156. Hypotype UCMP 31239. Remarks: Chlamys hastata hastata
(G.B. Sowerby II, 1842),  Moore (1984: p. B22). [Pectinidae].
Pecten (Patinopecten) haywardensis
Miocene. Near junction of Hollis Creek and Eden Creek, Hayward quad., Alameda Co. UCMP A4566. Plastoholo-
type UCMP 34133, plastoparatype UCMP 34134. Remarks: In the genus Patinopecten,  Moore (1984: p. B80).
[Pectinidae].
Pecten (Patinopecten) haywardensis calaverasensis
 

but cannot match to these entries]. Remarks: Treated as a valid full species in the genus Patinopecten,  Moore
(1984: p. B80). [Pectinidae].

paratype LACMIP 9138. [ex UCLA 46025].
Pecten (Patinopecten) healeyi Arnold, 1906 (n. sp.). USGS PP 47:103. Purisima Fm., Pliocene. San Gregorio Creek,
ex UCLA 46020]. Remarks: In the genus Pati-
nopecten,  Moore (1984: p. B1). [Pectinidae].
Pecten (Patinopecten) healeyi
Remarks: In the
genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi-

Remarks: In the genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi Arnold, 1906. Cummings et al. 1962. CDMG Bull. 181:200
(Tahana Member), Pliocene. Pomponio Creek area, San Gregorio quad., San Mateo Co. locality 470-2M. Hypotype
determined here as missing. Remarks: In the genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
Pecten aff. 
Member), Pliocene. Lobitos Creek area, Half Moon Bay quad., San Mateo Co. locality 639M. Hypotype determined
here as missing. Remarks: In the genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
134 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pecten (sp. healeyi
San Diego Co. Hypotype current whereabouts unknown. Remarks: In the genus Patinopecten,  Moore (1984:
p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi
Remarks: In the genus Patinopecten,
 Moore (1984: p. B81). [Pectinidae].
Pecten healeyi
Pliocene. North side Potato Harbor, Santa Cruz Is., California Channel Ids. UCSB GM65-21 [=LACMIP 16425]. Hy-
potype UCSB M14 [=LACMIP 6306]. Remarks: In the genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi
Re-
marks: In the genus Patinopecten,  Moore (1984:p. B81). There are two hypotypes listed in the CASG online
database, but neither one can be matched to the specimens listed below; thus they are listed for the record as
CASG 72546 and 72553. [Pectinidae].

Hypotype SDSNH 4336.
Pecten expansus Dall, 1878a]. San Diego, San Diego Co. Hypotype CASG 527 deter-
mined here as missing.



Pecten (Patinopecten) healeyi
Canyon, San Diego Co. UCLA 2359 [=LACMIP 22359]. Hypotype UCLA 49521 [=LACMIP 10620]. Remarks: In
the genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi

B) UCLA 1950 [=LACMIP 9291]. Remarks: In the genus Patinopecten,  Moore (1984: p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi Arnold, 1906 Fm., Pliocene.
Orange Co. SDSNH 4080. Hypotype SDSNH 75915. [Pectinidae].
Pecten (Patinopecten) healeyi sanclementensis
18, 19. Unnamed strata, Pliocene. San Clemente Is. North quad., San Clemente Is., California Channel Ids. UCLA
 
38792, 38793 [=LACMIP 10825, 10826]. Remarks: Junior synonym of Patinopecten healeyi (Arnold, 1906), 
Moore (1984: p. B81). [Pectinidae].
Pecten (Patinopecten) healeyi sanclementensis Susuki and Stadum, 1978. Squires and White 1983. SEPM Guidebook

Ventura Co. CSUN 725 [=LACMIP 29227]. Hypotype UCLA 59405 [=LACMIP 8318]. Remarks: Junior synonym
of Patinopecten healeyi (Arnold, 1906),  Moore (1984: p. B81). [Pectinidae].
Pecten hemphilli
Fm. (Cebada Member), Pliocene. Casmalia Hills, southeast of Waldorf, Santa Barbara Co. USGS 14631. Hypotype
USNM 560100. Remarks: Junior synonym of Pecten (Pecten) bellus (Conrad, 1857a),  Hertlein and Grant
(1972: p. 175);  Moore (1984: p. B68). [Pectinidae].

USGS 4474. Hypotype USNM 560135.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 135
Pecten (Pecten) hemphilli

San Diego Co. [Holotype USNM 7943]. Remarks: Junior synonym of Pecten (Pecten) bellus (Conrad, 1857a), 
Moore (1984: p. B68). [Pectinidae].
Pecten hemphilli

LACMIP1219]. Hypotypes (3) current whereabouts unknown. Remarks: Junior synonym of Pecten (Pecten) bel-
lus (Conrad, 1857a), Moore (1984: p. B68). [Pectinidae].
Pecten (Propeamusium) interradiatus
28. Kreyenhagen Fm., Eocene. Canoas Creek, Kreyenhagen Ranch, Kings Co. CASG 899. Hypotype CASG 5733.
Remarks: In the genus Propeamussium,Moore (1984: p. B7). [Propeamussidae].
Pecten (Pecten) juanensis
Fm., Miocene. Sandstone fossil reef on east side of valley of San Juan Creek, San Luis Obispo Co. UCLA 435 [=LAC-
MIP 20435]. Holotype UCLA 10441 [=LACMIP 9137]. Remarks: In the genus Oppenheimopecten, Moore
(1984: p. B75). [Pectinidae].

61716]. Paratype CASG 7729 [=CASG 61716.01].
Creek, San Luis Obispo Co. UCLA 435 [=LACMIP 20435]. Paratype USNM 560368.
Pecten (Pecten) cf.   (P.) lecontei
strata, Pliocene. San Clemente Island North quad., San Clemente Is., California Channel Ids. UCLA 6316 [=LACMIP
26316]. Hypotype UCLA 38787 [=LACMIP 10832]. [Pectinidae].
Pecten (Patinopecten) lohri-
litos Fm., Miocene Waltham Creek southwest of Coalinga, Fresno Co. Hypotype CASG 7734, determined here as
missing. Remarks: In the genus Patinopecten,Moore (1984: p. B80). [Pectinidae].
Pecten lohri
area, San Mateo Co. CASG 28689. [Hypotype CASG 9812]. Remarks: In the genus Patinopecten,Moore (1984:
p. B80). [Pectinidae].
Pecten (Patinopecten) lohri
Fm. [=Purisima Fm.], Pliocene. Near Moss Beach, San Mateo Co. CASG 29579. Hypotype CASG 10498 [=CASG
69970]. Remarks: In the genus Patinopecten,Moore (1984: p. B80). [Pectinidae].
Pecten (Patinopecten) lohri
Fm. (Pomponio Member), Pliocene. El Corte de Madera Creek La Honda quad., San Mateo Co. 726M. Hypotype
determined here as missing. Remarks: In the genus Patinopecten,Moore (1984: p. B80). [Pectinidae].
Pecten (Amussiopecten) lompocensis 
Unnamed strata, Miocene. San Clemente Is. North quad., San Clemente Is., California Channel Ids. UCLA 6317
[=LACMIP 26317]. Hypotypes UCLA 38788, 38789, 38895–38897 [=LACMIP 10827–10831]. Remarks: In the
genus Amusium,Moore (1984: p. B66). [Pectinidae].
Pecten (Amussiopecten) lompocensis
Miocene. Saddleback Valley, Orange Co. SDSNH 4312. Hypotypes SDSNH 83232, 83233]. Remarks: In the genus
Amusium,Moore (1984: p. B66). [Pectinidae].
Pecten magnolia
  Remarks: In the genus Macrochlamis,
Moore (1984: p. B56). [Pectinidae].
Pecten miguelensis
Miocene. Cañada Garanon, Santa Rosa Is., California Channel Is. UCSB SR67-53 [=LACMIP 16404]. Hypotype UCSB
M15 [=LACMIP 6320]. Remarks: Lyropecten submiguelensis (Loel and Corey, 1932) auct.,  Smith (1991a:p.
136 PALEOBIOS, VOLUME 38, FEBRUARY 2021
69); [=Lyropecten pretiosus (Hertlein, 1925),Smith (1991a:p. 69). [Pectinidae].
Pecten miguelensis Arnold, 1906. Shapiro 1997. Santa Cruz Is., California Channel Ids. Geol. Guide, p. 40 (as Lyropecten
miguelensis
drawing, source specimen unknown. Remarks: In the genus Lyropecten,Moore (1984: p. B49). [Pectinidae].
Pecten miguelensis (supervariant) Arnold, 1906. Bereskin and Edwards 1969. AAPG/SEPM Guidebook, p. 74, pl.
Lyropecten in text). Rincon Fm., Miocene. Cañada de los Sauces area, Santa Cruz Is., California
Channel Is. UCSB E8 [=LACMIP 16406]. Hypotype UCSB M16 [=LACMIP 6300]. Remarks: In the genus Lyropecten,
Moore (1984: p. B49). [Pectinidae].
Pecten miguelensis submiguelensis Loel and Corey, 1932. Bereskin and Edwards 1969. AAPG/SEPM Guidebook, p.
Lyropecten in text). Vaqueros Fm., Miocene. Near Pt., Santa Cruz Is., California Channel Ids.
UCSB DWW-16 [=LACMIP 16407]. Hypotype UCSB M17 [=LACMIP 6298]. Remarks: Vertipecten bowersi (Arnold,
1906),  Smith (1991a: p. 77). [Pectinidae].
Pecten miguelensis submiguelensis Loel and Corey, 1932. Avila and Weaver 1969. AAPG/ SEPM Guidebook, p. 59, pl.

UCSB M18 [=LACMIP 6299]. Remarks: Lyropecten submiguelensis,  Moore (1984: p. B50). [Pectinidae].
Pecten oregonensis Howe, 1922 var. Manning and Ogle, 1950. CDM Bull. 148:23 [as Pecten caurinus Gould orego-
nensis Howe var.], pls. 6, 7. Falor Fm., Pliocene. Humboldt Co. UCMP A4233. Hypotype UCMP 34172. Remarks:
Patinopecten (Lituyapecten) falorensis n.sp, MacNeil (1961: p. 234). [Pectinidae].
Pecten pabloensis
the town of Rodeo, Contra Costa Co. CASG 27420. Hypotype CASG 6081]. Remarks: In the genus Pacipecten,
Moore (1984: p. B47). [Pectinidae].
Pecten (Pseudamusium) peckhami

Remarks: In the genus Delectopecten,Moore (1984: p. B15). [Pectinidae].
Pecten (Delectopecten) pedroanus (Trask, 1856). [Plagiostoma]. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):174,
Remarks:
Nomen dubium and probably equivalent to Delectopecten peckhami (Gabb, 1869), Moore (1984: p. B15).
[Pectinidae].
Pecten (Pseudamusium) pedroanus (Trask, 1856). [Plagiostoma]. Shimer and Shrock 1944. Index Foss. N.A., p. 407,
Pecten peckhami
Remarks: Nomen dubium and
probably equivalent to Delectopecten peckhami (Gabb, 1869a),Moore (1984: p. B15). [Pectinidae].
Pecten (Patinopecten) purisimaensis Arnold, 1906 (n. sp.). USGS PP 47:105. Purisima, Miocene/ Pliocene. Ano
Remarks: In the
genus Patinopecten,Moore (1984: p. B87). [Pectinidae].
Pecten (Patinopecten) purisimaensis Arnold, 1906. Glen
. Near Princeton, San Mateo Co. UCMP B4791. Hypotype UCMP 37617.
Remarks: In the genus Patinopecten,Moore (1984: p. B87). [Pectinidae].
Pecten (Patinopecten) purisimaensis
Fm. (Tahana Member), Pliocene. Near Pomponio Creek, San Gregorio quad., San Mateo Co. 469-5M. Hypotype
determined here as missing. Remarks: In the genus Patinopecten,Moore (1984: p. B87). [Pectinidae].
Pecten raymondi brionianus
Monica Mtns., Los Angeles Co. USNM unknown. USNM hypotype determined here as missing. Remarks: Junior
synonym of Pacipecten discus (Conrad, 1857d),Moore (1984: p. B47). [Pectinidae].
Pecten (Pecten) raymondi brionianus
of Pleasanton, Niles quad. (1953), Alameda Co. SU 3232. Hypotype SU 8440 [=CASG 69924]. Remarks: Junior
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 137
synonym of Pacipecten discus (Conrad, 1857d),Moore (1984: p. B47). [Pectinidae].
Pecten santaecruzensisPhacoi-
des Sand member) (of Stinemeyer et al. 1959:12, 13), Miocene. Carneros Rocks quad., Temblor Range, Kern Co.
USGS M4467. Hyoptype USNM 646535. Remarks: In the genus Oppenheimopecten,Moore (1984: p. B75).
[Pectinidae].
Pecten (Pecten) sanctaecruzensis

Carneros Rocks quad., Kern Co. USGS M4467. Hypotypes USNM 646535, 646536. Remarks: In the genus Op-
penheimopecten,Moore (1984: p. B75). [Pectinidae].
Pecten sanctaecruzensis
Fm. (upper sandstone member), Oligocene. The Indians, Junipero Serra quad., Santa Lucia Range, Monterey Co.
SU/CASG 1758. Hypotypes CASG. 60364, 60369, 60374. CASG hypotypes determined here as missing. Remarks:
In the genus Oppenheimopecten,Moore (1984: p. B75). [Pectinidae].
Pecten (Chlamys) cf.   (C.) sespeensis
Fm., Oligocene. Vicinity of Cañada Omentero, Santa Barbara Co. UCMP A4646. Plastotypes UCMP 33448, 33449.
Remarks: Pecten (Vertipecten) yneziana yneziana Arnold, 1907,  Weaver and Kleinpell (1963:p. 198); Verti-
pecten yneziana (Arnold, 1907),  Smith (1991a: p. 85). [Pectinidae].
Pecten (Chlamys) sespeensis
Pecten sespeensis in pl. captions)). Vaque-
ros Fm., Miocene. Near South Mtn., Ventura Co.]. UCMP A244. Hypotypes UCMP 31720, 31721. Remarks: In the
genus Chlamys,Moore (1984: p. B20). [Pectinidae].

Ventura Co.]. UCMP A252. Hypotype UCMP 31723].
UCMP B7015.
Hypotype SU 9261 [=CASG 70223]. Remarks: Vertipecten yneziana (Arnold, 1907),  Smith (1991a: p. 86).

Rafael Mtns. quad., Santa Barbara Co. UCMP B6940. Hypotype SU 9260 [=CASG 70202].
Pecten (sp. stearnsii
San Diego Co. Hypotype current whereabouts unknown. Remarks. In the genus Flabellipecten,Moore (1984:
p. B72). [Pectinidae].
Pecten (Pecten) stearnsi
Remarks: In the genus Flabellipecten,
Moore (1984: p. B72). [Pectinidae].
Pecten (Flabellipecten) stearnsii
   Re-
marks: In the genus Flabellipecten,Moore (1984: p. B72). [Pectinidae].

Pecten (Flabellipecten) stearnsii
San Diego Co. UCLA 2420 [=LACMIP 22420). Hypotype UCLA 49520 [=LACMIP 10619]. Remarks: In the genus
Flabellipecten,Moore (1984: p. B72). [Pectinidae].
Pecten (Flabellipecten) stearnsii
Orange Co. SDSNH 4080. SDSNH hypotype determined here as missin. Remarks: In the genus Flabellipecten,
Moore (1984: p. B72). [Pectinidae].
Pecten. Purpuratus subdolis
Diego, San Diego Co. Hypotype current whereabouts unknown. Remarks: In the genus Argopecten,Moore
138 PALEOBIOS, VOLUME 38, FEBRUARY 2021
(1984:p. B37). [Pectinidae].
Pecten (Argopecten) subdolus
Remarks: In the genus
Argopecten,Moore (1984: p. B37). [Pectinidae].
Pecten (Pallium) swiftii Bernardi var. etchegoini

Hypotype UCMP 37614. Remarks: Swiftopecten swiftii parmeleei (Dall, 1898),  Masuda (1972:p. 401); Swifto-
pecten parmeleei etchegoini, Moore (1984: p. B60). [Pectinidae].
Pecten (Pallium) swifti nutteri Arnold, 1906. Shimer and Shrock 1944. Index-
 Pecten nutteri Arnold 1906)). Etchegoin Fm., Pliocene.
Coalinga District, Fresno Co. USGS 4712. Hypotype USNM 165549. Remarks: Nanaochlamys nutteri (Arnold,
1906), Moore (1984:p. B31). [Pectinidae].
Pecten (Pallium) swifti nutteri  
ing, source specimen unknown. Remarks: Nanaochlamys nutteri (Arnold, 1906),  Moore (1984: p. B31).
[Pectinidae].
Pecten tolmani-
ton, La Costa Valley quad. (1953), Alameda Co. SU 3252. Hypotype SU 8441 [=CASG 69923]. Remarks: In the
genus Pacipecten,Moore (1984: p. B46). [Pectinidae].
Pecten vanvlecki -
queros Fm., Miocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. UCSB RB65-40 [=LACMIP 16431.
Hypotypes UCSB M19, M20 [=LACMIP 6296, 6297]. Remarks: In the genus Amussiopecten,Moore (1984:
p. B69). [Pectinidae].
Pecten (Oppenheimopecten) vogdesi Arnold, 1906. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):181–182, pl. 29,
 
Remarks: In the genus Oppenheimopecten,Moore (1984: p. B77); in the genus Euvola,  Coan et al. (2000:
p. 225). [Pectinidae].


Sacate/Gaviota fms., Eocene. Lompoc Quad., Santa Barbara Co. UCMP B6963. Paratype SU 9268 [=CASG 70208].
Remarks: In the genus Vertipecten,Moore (1984: p. B61). [Pectinidae].

Rafael Mtn. quad., Santa Barbara Co. UCMP B6940. Holotype SU/CASG 9265, paratypes SU/CASG 9266, 9267.
Pecten yneziana subyneziana
Sandstone (of Weaver and Kleinpell, 1963: pp. 83, 91, 92), Eocene. Along Camino Cielo Rd., San Rafael Mtn., Santa
Brabara Co. UCMP B6940. Holotype SU 9265 [=CASG 7021, as Pecten (Vertipecten) yneziana yneziana in online
database]. Remarks: In the genus Vertipecten,Moore (1984: p. B61). [Pectinidae].
Pecten yneziana yneziana       
Sacate/Gaviota fms., Eocene. Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6955. Hypotype SU/CASG
9263. Remarks: In the genus Vertipecten,Moore (1984: p. B61). [Pectinidae].

Lompoc quad., Santa Barbara Co. UCMP B7033. Hypotype SU/CASG 9264.

SU/CASG 9262.
Pecten. Edinger 1960. KCCB 10:2, 12   
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 139
Kern Co. Hypotypes Kern County Schools Collection, determined here as missing. Remarks: Determined here
as Vertipecten sp. [Pectinidae].
Pecten sp. Redman et al. 2007. Palaios
Imperial Co. Hypotype probably at SDSU (current whereabouts unknown). [Pectinidae].
  
Obispo Co. UCMP A742. UCMP hypotype 12160 (pectinid fragment encrusted with bryozoan colony Canopeum
sp.). [Pectinidae].

Santa Monica Mtns., Los Angeles Co. Hypotype SAMO 245. Remarks: Determined here as Vertipecten kernensis
(Hertlein, 1925a). [Pectinidae].
Pegophysema edentuloides (Verrill, 1870). [Loripes]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p.
  
Hypotype SDSNH 17701. [Lucinidae].
Pegophysema edentuloides (Verrill, 1870). [Loripes
[=Latrania Fm.], Pliocene. Vicinity of Travertine Point, Oasis quad., Imperial Co. (not San Diego Co. as in locality de-
scription). UCMP A1707 [=UCMP A1705]. Hypotype UCMP 39875, missing, Powell (2008: p. 12). [Lucinidae].
Pelecyora aequilateralis (Gabb, 1869b). [Venus
Eocene. Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 (=LACMIP 16963). Hypotype LACMIP 12469,
12470. [Veneridae].
Pelecyora gabbi (Arnold, 1909 [1910]). [Meretrix
Vicinity of Thorn Meadows, Lion Canyon quad. (1943), Ventura Co. UCR 4683. Hypotypes UCR 4683/701,
4683/702. [Veneridae].
Penitella conradi
Diego Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP. 305C [= LACMIP 16817]. Hypotype LACMIP
4832. [Pholadidae].
Penitella durhami
Fm., Oligocene. Walnut Creek quad., Contra Costa Co. UCMP A7762. Holotype UCMP 30101, paratypes UCMP

Penitella durhami
Contra Costa Co. UCMP A4296 (not UCMP A7762 corrected in text). Holotype UCMP 30108. [Pholadidae].

Penitella lorenzana (Clark, 1925). [Pholadidea]. Adegoke 1967d. PCAS 35(1):11–12, 
Oligocene. Near town of Walnut Creek, Concord quad., Contra Costa Co. UCMP 1131. Holotype UCMP 30323,
paratype UCMP 33491. [Pholadidae].
Penitella lorenzana (Clark, 1925). [Pholadidea
Oligocene. Near town of Walnut Creek, Contra Costa Co. UCMP 1131. Holotype UCMP 30323, paratype UCMP
33491. [Pholadidae].

Penitella penita (Conrad, 1837). [Pholas
Diego Fm., Pliocene. Near U.S./Mexico boundary, San Diego Co. SDSNH 417. Hypotype SDSNH 4356. [Pholadidae].

UCLA [48637 [=LACMIP 9320].

140 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Penitella turnerae
Vicinity of Guadalupe, Santa Barbara Co. USGS 4473. Hypotype USNM 189803. [Pholadidae].
Periploma cf. 
Canyon area, Los Angeles Co. CIT 1671 [=LACMIP 2556]. Hypotype LACMIP 4149]. Remarks: In the subgenus
Circoploma,  Coan et al. (2000: p. 542). [Periplomatidae].
Periploma stenopa
Fm., Pliocene. San Diego Co. LAMCIP 305A [=LACMIP 16862]. Hypotypes LACMIP 4837, 4838 [Periplomatidae].
Periploma
quad. (1943), Ventura Co. UCR 4732. Hypotype UCR 4732/2. [Periplomatidae].
Periploma
 C.
Throckmorton (personal communication, 2002). [Periplomatidae].
Perna (Perna) montana

as stated,  Conrad (1857d: p. 189). Santa Buenaventura, Santa Barbara Co. [=Ventura, Ventura Co.) Holotype
USNM 13329. Remarks: Determined here as a nomen dubium. [Mytilidae].
Petricola (Rupellaria) carditoides Conrad, 1837. [Saxicava]. Hertlein and Grant 1972. SDSNH Mem. 2 (2B):283, 284,

SDSNH 4346. [Petricolidae].
Cast Merced” Fm., Pliocene. Moss Beach, San Mateo Co.
CASG 34437. Hypotype CASG 54827. [Pholadidae].

Reef, Jasper Canyon, Fresno Co. CASG 1841. Hypotype CASG 54826 [this specimen is now a paratype of Chaceia
fulcherae Kennedy, 1993,  Kennedy (1993: p. 399). [Pholadidae].
Pholadidea ovoidea (Gould, 1851), short var. [Pholas]. Woodring et al. 1940 [1941]. USGS PP 195: chart opposite p.
Pecten zone), Pliocene. Arroyo Conshoso, North Dome, Kettleman Hills, Kings
Co. USGS 12374. Hypotype USNM 495749. Remarks: Pholadopsis pectinata Conrad, 1849,  Kennedy (1974:
p. 69);  label with USNM specimen.[Pholadidae].
Pholadidea penita (Conrad, 1837). [Pholas    
Sisquoc Fm. (diatomaceous strata), Miocene. Santa Barbara Co. USGS 4472. Hypotype USNM 560039. Remarks:
Penitella turnerae Evans and Fisher, 1966,  label with USNM specimen; Kennedy (1974: p. 54). [Pholadidae].

Hypotype USNM 560106. Remarks: Penitella turnerae Evans and Fisher, 1966;  label with USNM specimen;
 Kennedy (1974: p. 54).
Pholadomya (Bucardiomya) givensi
Spring Canyon, Topatopa Mtn. quad., Santa Barbara Co. UCR 4662. Holotype UCR 4662/110. [Pholadomyidae].
....., Hot Spring Canyon, Ventura Co. UCR 4659. Paratype UCR 4659/10
....., Hot Spring Canyon, Ventura Co. UCR 4664. Paratypes UCR 4664/5–4664/8.
Pholadomya (Bucardiomya) givensi      
Eocene. Los Angeles Co. CSUN. 816 [=LACMIP 16336]. Hypotype LACMIP 7543. [Pholadomyidae].
Pholadomya (Pholadomya) kernensis     
Miocene. North of Poso Creek, Kern Co. SU 438. Holotype SU/CASG 437 [=5166 on CASG label] [=CASG 71527].
[Pholadomyidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 141
Pholadomya (Bucardiomya) mounti
Paleocene. Simi Hills, Ventura Co. UCR 6899. Paratype UCR 6899/4. [Pholadomyidae].
-
type UCR 6898/1.
....., North slope Simi Hills, Ventura Co. UCR 4575. Paratypes UCR 4575/33, 4575/34.
....., North slope Simi Hills, Ventura Co. UCR 6602. Paratypes UCR 6602/4.
....., North slope Simi Hills, Ventura Co. UCR 6816. Paratype UCR 6816/2.
....., Vine Hill Fm., Paleocene. Pacheco Rd., Contra Costa Co. UCR 7481. Paratype UCR 7481/1.
Pholadomya mounti
Paleocene. Simi Hills, Ventura Co. UCR 6898. Paratype UCR 6898/1. [Pholadomyidae].
Pholadomya (Pholadomya) murrayensis  -

[Pholadomyidae].
Pholadomya (Pholadomya) nasuta
Paleocene. Simi Hills, Ventura Co. UCR 6813. Hypotype UCR 6813/4. [Pholadomyidae].
Pholadomya nasuta-
cene. Simi Hills, Ventura Co. UCR 6813 (not UCR 6814 as in pl. caption). Hypotype UCR 6813/4 (not UCR 6814/4
as in pl. caption). [Pholadomyidae].
Pholadomya
Vicinity of Cañada de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP B6922. Hypotype SU 9269 (not SU
9326 as it text) [=CASG 70214]. [Pholadomyidae].
Pholadomya (Pholadomya-
topa Mtn. quad. (1943), Ventura Co. UCR 4662. Hypotype UCR 4662/110. Remarks: Pholadomya (Bucardiomya)
givensi n. sp.,  Zinsmeister (1978a:p. 235). [Pholadomyidae].
Pholadopsis pectinataPecten

Pholadidea ovoidea)) USNM 189804, 189805, 495749. [Pholadidae].
Phygraea badgeri
Derrydale Creek, Upper Sespe Creek area, Ventura Co. LACMIP 23724. Holotype LACMIP 14731, paratypes LAC-
MIP 14732–14734. [Gryphaeidae].


14735–14736.

Phygraea haleyi (Hertlein, 1933). [Ostrea137–139. Pozo Fm., Pale-
ocene. Cañada del Pozo, Santa Cruz Is., California Channel Ids. CASG 5526. Holotype CASG 66752.02. [Gryphaeidae].

Ids. LACMIP 7890. Hypotype LACMIP 14708.


142 PALEOBIOS, VOLUME 38, FEBRUARY 2021
14713.
  
LACMIP 14711, 14714.

Phygraea stewarti (M.A. Hanna, 1927). [Ostrea
173. Llajas Fm., Eocene. Llajas Canyon, north side of Simi Valley, Ventura Co. LACMIP 22312. Hypotypes LACMIP
14715–14719, 14721–14729. [Gryphaeidae].

14720.


UCMP 30921.
Pinna (Pinna) barrowsi   -
leocene. Just south of Arroyo del Hambre Creek, Concord quad., Contra Costa Co. UCMP 1547. Holotype UCMP
11728. [Pinnidae].
Pinna (Pinna) latrania
Fm.], Miocene/Pliocene. North side of Coyote Mtn., Imperial Co. USGS 3922. Holotype USNM 324593. [Pinnidae].
Pinna cf. 
Pliocene. Travertine Point area, Oasis quad., Imperial Co. (not San Diego Co. as in locality description). UCMP
A1705. Hypotype UCMP 39869, missing,  Powell (2008: p. 6). [Pinnidae].
Pinna (Pinna) lewisi
Wells [= Oil Canyon], Susana quad., Ventura Co. SU 2696. Holotype SU/CASG 5194 [=CASG 69886]. [Pinnidae].
Pinna lewisi Waring, 1917. Squires is in type
section of formation., Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 475 [=LACMIP 16337]. Hypotype
LACMIP 6568. [Pinnidae].
Pinna llajasensis 
Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 458 [=LACMIP 16280]. Holotype UCLA 59196 [=LACMIP
10549]. [Pinnidae].
Pinna llajasensis         
Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 458 [=LACMIP 16280]. Holotype UCLA 59196 [=LACMIP
10549]. [Pinnidae].
Pinna (Pinna) mendenhalli-
trania Fm.], Miocene/Pliocene. Garnet Canyon, Coyote Mtn., Imperial Co. USGS 3922. Holotype USNM 324593a.
[Pinnidae].
Pinna (Pinna) stocktoni
Oligocene/Miocene. Near Wiley Canyon, Piru quad., Ventura Co. UCMP A252. Syntypes UCMP 31778, 31779.
[Pinnidae].
Pinna (Pinna        
Remarks: Pinna
llajasensis n. sp.,  Squires (1983a: p. 359). [Pinnidae].
Pitar addicotti-
basas quad., Simi Hills, Ventura Co. UCR 7319 (not UCR 7819 as in pl caption). Holotype UCR 7319/100 (not UCR
7819/100 as in pl. caption or UCR. 7319/1 as in text). [Veneridae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 143

Pitar (Lamelliconcha) avenalensis
Meadows area, Lion Canyon quad. (1943), Ventura Co. UCR 4679. Hypotypes UCR 4679/30, 4679/31. [Veneridae].
Pitar (Lamelliconcha) avenalensis
Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13453. [Veneridae].
Pitar behri
sandstone ledge of upper Temblor Fm. below McLure Shale contact, Reef Ridge quad., Kings Co. UCMP D1051.
Hypotype UCMP 14997. [Veneridae].
Pitar (Katherinella) cf.   () californica (Clark, 1918). [Callocardia (Agrippoma)]. Addicott 1973b. USGS PP 791:7,
    
quad., Kern Co. USGS M3280. Hypotype USNM 646586. [Veneridae].
Pitar (Lamelliconcha) dickersoni Givens, 1974 (Eocene. Near
Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. CASG 244. Holotype CASG 262, paratype CASG 263,
hypotype UCR 4721/171. Remarks: Givens (1974) proposed the new name Pitar (Lamelliconcha) dickersoni
for the species Meretrix tejonensis Dickerson (1915: p. 53 [in Stewart
(1930, p. 236), Dickerson’s nameis an objective synonym of Pitar uvasanus (Conrad) of Eocene age from the
Tejon Formation. [Veneridae].

Pitar (Macrocallista) horni (Gabb, 1864). [Meretrix
Macrocallista hornii)). Tejon Fm., Eocene. Ft. Tejon, Kern Co.
Lectotype (of Stewart 1930: p. 243) ANSP 4454]. [Veneridae].
Pitarjoaquinensis  
Ridge, Reef Ridge quad., Kings Co. USGS 14497a. Hypotype USNM 498736. Remarks: In the genus/subgenus
Pitar (Lamelliconcha),  Squires (1987: p, 69). [Veneridae].
Pitar (Lamelliconcha) joaquinensis
Fork of Piru Creek and near Thorn Meadows, Lockwood Valley quad. (1943), Ventura Co. UCR 4673. Hypotype
UCR. 4673/201. [Veneridae].
Pitar (Lamelliconcha) joaquinensis
Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6591. [Veneridae].
Pitar (Lamelliconcha) joaquinensis
Eocene. Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype
LACMIP 7536. [Veneridae].
Pitar (Lamelliconcha) joaquinensis       

[Veneridae].
Pitar (Lamelliconcha) joaquinensis
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 974 [=LACMIP 16193]. Hypotype LACMIP 7732.
[Veneridae].
Pitar (Lamelliconcha) joaquinensis   
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype
LACMIP 11487. [Veneridae].
Pitar (Lamelliconcha) joaquinensis
Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13452. [Veneridae].
Pitar kelleyi
144 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Goleta quad., Santa Barbara Co. UCMP B6979. Paratype SU/CASG 9302 [=CASG 70217]. [Veneridae].

....., pl. 36 Olivos quad., Santa Barbara Co.
UCMP B6933. Paratype SU/CASG 9303 [=CASG 70218].
Pitar lascrucensis
Western Santa Ynez Mtns., Lompoc quad., Santa Barbara Co. UCMP B7019. Holotype SU 9304 [=CASG 70219],
paratype SU 9305 [=CASG 70220]. [Veneridae].
  
Paratype SU 9306 [=CASG 70221].
Pitar (Pitar) cf.   ( ) lascrucensis
Fm., Eocene. Near Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4732. Hypotype UCR 4732/101.
[Veneridae].
Pitar lorenzanus (Clark, 1918). [Pitaria
Lompoc quad., Santa Barbara Co. UCMP B7006. Hypotype SU 9307 [=CASG 70222]. [Veneridae].
Pitar mulinoidus
El Jaro Creek, Lompoc quad., Santa Barbara Co. UCMP B7000. Holotype SU 9308 [=CASG 70224], paratype SU
9309 [=CASG 70225]. [Veneridae].
Pitar murphyi-
cene. Calabasas quad., Simi Hills, Ventura Co. UCR 6806. Holotype UCR 6806/142 (not UCR 6806/141 as in text),
paratypes UCR 6806/141, 6806/143–6806/144. [Veneridae].
Pitar (Lamelliconcha) soledadensis (M.A. Hanna, 1927). [Pitaria-
cal Fm., Eocene. Alamo Creek area, San Guillermo quad. (1943), Ventura Co. UCR 4705. Hypotype UCR 4705/1.
[Veneridae].
Pitar simiensis (Nelson, 1925). [Callocardia
Fm., Paleocene. Simi Hills, Ventura Co. [UCR 4569]. Hypotype UCR 4569/5. [Veneridae].
Pitar tejonensisMeretrix
Sandstone, Eocene. Brush Peak area on Camino Cielo, San Rafael Mtn. quad., Santa Barbara Co. UCMP B6946.
Hypotype SU 9310 [=CASG 70226]. [Veneridae].
Pitar uvasana (Conrad, 1855b). [Meretrix
Pitar uvasanus)). Tejon Fm., Eocene. Ft. Tejon, Kern Co. Neotype (of Stewart
1930: p. 236) ANSP. 4457]. [Veneridae].
Pitar uvasana (Conrad, 1855b). [Meretrix-
ing, source specimen unknown. [Veneridae].
Pitar (Calpitaria) uvasanus (Conrad, 1855b). [Meretrix
Eocene. Devil Canyon area, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6590. [Veneridae].
Pitar (Calpitaria) uvasanus (Conrad, 1855b). [Meretrix]. Squires 1988a. SEPM Guidebook-
 . 12i). Llajas Fm., Eocene. Simi Valley, Ventura Co. CSUN 371 [=LACMIP 16115].
Hypotype LACMIP 6590. [Veneridae].
Pitar (Calpitaria) uvasanus (Conrad, 1855b). [Meretrix-
cene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13454.
[Veneridae].
Pitar (Calpitaria) uvasanas coquillensis
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 145
Eocene. Reef Ridge, Reef Ridge quad., Fresno Co. USGS 14494. Hypotype USNM 498735. [Veneridae].
Pitar uvasana coquillensis
South side of Simi Valley, Calabasas quad. (1952, PR 1967), Ventura Co. CSUN 1344 [=LACMIP 17192]. Hypotype
LACMIP 12701. [Veneridae].
Pitar uvasana coquillensis
22312. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 22312]. Hypo-
types LACMIP 12756, 12757. [Veneridae].
Pitar


Pitar (Lamelliconcha
(1932, PR 1946), Los Angeles, Co. CIT 1663 [=LACMIP 2555]. [Hypotype LACMIP 4143]. [Veneridae].
Pitar-
joqui Creek area, Lompoc quad., Santa Barbara Co. UCMP B6958. Hypotype SU/CASG 9312. [Veneridae].
Pitar
Creek, Carneros Rocks quad., Kern Co. USGS M3978. Hypotype USNM 646587. [Veneridae].
Pitar     
 C.
Throckmorton (personal communication, 2002). [Veneridae].
          
Paleocene. Midway quad., Alameda Co. UCMP D8147. UCMP unnumbered hypotype lost, , C. Throckmorton
(personal communication, 2002).
Pitar sp. Shapiro 1998. AAPG Guidebook Strata unknown, [Miocene]. Cañada
Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8825 [=SBMNH 60185]. Remarks: Determined
here as Mactromeris cf.  (Gabb, 1866). [Mactridae].
Pitar        
Ventura Co. UCR 7319. Hypotype UCR 7319/100 (not UCR 6893/2 as in pl. caption). Remarks: Pitar addicotti
n. sp.,  Zinsmeister 1983b: p. 1288. [Veneridae].
Pitar
Co. UCR 6806]. Hypotype UCR 6806/141 (not UCR 6806/15 as in pl. caption). Remarks: Pitar murphyi n. sp.,
 Zinsmeister (1983b: p. 1288). [Veneridae].
Placunanomia californica
Reef Ridge quad., Kings Co. UCMP B6537. Hypotype UCMP 36644. [Anomiidae].
Placunanomia californica
Fm., Miocene/Pliocene. South end of Kettleman Hills, Lost Hills quad., Kern Co. USGS 4715. Holotype USNM
165546. [Anomiidae].
Placunanomia granti     
Miocene. West of Arlington Canyon, Santa Rosa Is., California Channel Ids. CASG 1150. Holotype CASG 4139.
[Anomiidae].
Placuanomia hannibali
Pliocene. Southwest of Valencia, Oat Mtn. quad. (1952, PR 1969), Los Angeles Co. LACMIP 17777. Hypotypes
LACMIP 13349, 13350. [Anomiidae].
Placunanomia

146 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Plagiocardium (Schedocardia) breweri (Gabb, 1864). [Cardium  
Avenal Sandstone, Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS 14492. Hypotype USNM 498721. Re-
marks: In the genus/subgenus Acanthocardia (Schedocardia),  Squires (1984: p. 49; 1987: p. 65);  Moore
(2003a: p. 3). [Cardiidae].
Platyodon clarki
Walnut Creek, Concord sheet, Contra Costa Co. UCMP 1942. Holotype UCMP 11555, paratypes UCMP 32516.

Platyodon pecki
Arnold Industrial Hwy. (St. Rte. 4), Walnut Creek quad. Contra Costa Co. UCMP A7762. Holotype UCMP 15471,

Pleurolucina        
of Travertine Rock, Imperial Co. (not San Diego Co. as in locality description). USGS. M9778. Hypotype USNM
486901. [Lucinidae].
Plicatula juncalensis Squires, 1987 (n. sp.). NHMLAC 
Whitaker Peak quad. (1958.), Los Angeles Co. CSUN 362 [=LACMIP 16273. Holotype LACMIP 7513. [Plicatulidae].
Plicatula juncalensis
Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Holotype LACMIP
7513. [Plicatulidae].
Plicatula lapidicina
Paleocene. Quarry Canyon area, Topanga quad. (1952, PR 1981), east-central Santa Monica Mtns., Los Angeles
Co. LACMIP 26725. Holotype LACMIP 11278. [Plicatulidae].
....., Vicinity of Quarry and Trailer canyons, Topanga quad. (1952, PR 1981), east-central Santa Monica Mtns., Los

Plicatula ostreiformis
Lower Lake, Lake Co. USGS 4192. Syntypes USNM 157838. [Plicatulidae].

Plicatula ostreiformis      
Martinez Fm., Paleocene. Vicinity of Herndon Creek near Lower Lake, Lower Lake quad. (1975), Lake Co. LACMIP
7047 [=UCMP 790]. Hypotypes LACMIP 8063–8065. [Plicatulidae].
Ostrea
buwaldana Dickerson, 1914a) UCMP 11719.
Plicatula simiensis-
cene. Northeast of Meier Canyon, Santa Susana quad. (1951, PR 1969), south side Simi Valley, Ventura Co. UCMP
3754. Holotype UCMP 154004. [Plicatulidae].
Plicatula trailerensis
Paleocene. Trailer Canyon area, Topanga quad. (1952, PR 1981), east-central Santa Monica Mtns., Los Angeles
Co. LACMIP 10508. Holotype LACMIP 11280. [Plicatulidae].
Pododesmus cf. Pmacrochisma (Deshayes, 1839). [Anomia]. Stanton 1966. Jour. Paleo. 40(1): 28, pl. 5, . 7. Castaic
Fm., Miocene. Dry Canyon Dam area, Los Angeles Co. CIT 1670 [=LACMIP 7751. Hypotype LACMIP 4092]. Re-
marks: In the subgenus Monia,  Coan et al. (2000: p. 256). [Anomiidae].
Pododesmus macrochisma (Deshayes, 1839). [Anomia]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):225–226, pl.

Co. LACMIP 305. Hypotypes LACMIP 4532, 4533. Remarks: In the subgenus Monia,Coan et al. (2000: p.
256). [Anomiidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 147

9298].
Pododesmus (Monia) macrochisma (Deshayes, 1839). [Anomia]. Vedder and Moore 1976. AAPG Misc. Pub. 24:123,128,
 Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS
M6501. Hypotype USNM 235576. [Anomiidae].
Pododesmus (Monia) macroschisma (Deshayes, 1839). [Anomia]. Squires and White 1983. SEPM Guidebook 35:222,

Co. CSUN 725 [=LACMIP 29227]. Hypotype UCLA 59407 [=LACMIP 8320]. [Anomiidae].
Pododesmus macrochisma (Deshayes, 1839). [Anomia
Pliocene. Orange Co. SDSNH 4080. Hypotype determined here as missing. Remarks: In the subgenus Monia,
Coan et al. (2000: p. 256). [Anomiidae].
Pododesmus (Monia) macrochisma (Deshayes, 1839). [Anomia-
chegoin Fm., Miocene/Pliocene. Coalinga area, Fresno Co. USGS 4767. Hypotype (of Arnold 1909 [1910], pl. 14,

Pododesmus macrochisma (Deshayes, 1839). [Anomia]. Squires 2012. NHMLAC CS 520:76, . 24. Pico Fm., Pliocene.
Newhall area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14349. [Anomiidae].
Pododesmus cf.  (Deshayes, 1839) [Anomia]. Powell et al. 2019a. USGS SIR 2019-5021:3, 29. 30, pl.
   
78429. Remarks: In the subgenus Monia,Coan et al. (2000: p. 256). [Anomiidae].
Porterius woodfordi (M.A. Hanna, 1927). [Barbatia
Fm., Eocene. Near mouth of Soledad Valley, La Jolla quad., San Diego Co. UCMP 5062. Holotype UCMP 31062.
[Parallelodontidae].
Portlandia (Portlandia) markleyensis (Clark, 1938). [Yoldia-
enhagen Fm. (Markley Sandstone Member), Eocene Pleasants Creek area Napa quad., Solano Co. UCMP. A1297.
Holotype UCMP 30833. [Yoldiidae].
Portlandia (Portlandia) mortuasuensis (Clark and Woodford, 1927). [Yoldia]. Moore 1983. USGS PP 1228-A:A29,

3577. Holotype UCMP. 31337. [Yoldiidae].
Portlandia (Portlandia) mosesi (Palmer, 1923). [Yoldia
Fm., Eocene. Ualtis Creek, just south of Dunn's Peak, Vaca Valley, Napa quad., Solano Co. UCMP 3573. Holotype
UCMP 30704. [Yoldiidae].
Portlandia (Portlandia) packardi (Clark, 1918). [Malletia
Tuff, Oligocene. West of Kirker's Creek, Mt. Diablo quad., Contra Costa Co. UCMP 2033. Holotype UCMP 11154.
[Yoldiidae].
Portlandia (Portlandia) rosa (M.A. Hanna, 1927). [Leda
Eocene. Rose Creek, east of La Jolla, La Jolla quad., San Diego Co. UCMP 3993. Holotype UCMP 31090. [Yoldiidae].
Pristes oblongus

Co. LACMIP 305. Hypotypes LACMIP 4629–4634. Remarks: In the genus/subgenus Thecodonta (Pristes), 
Moore (1992: p. E9). [Lasaeidae].
 
Eocene. Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype
LACMIP 7512. [Propeamussidae].
Propeamussium interradiatum (Gabb, 1869b). [Pecten
Fm., Eocene. Reef Ridge, Dark Hole quad., Fresno Co. USGS 14563. Hypotype USNM 498738. [Propeamussiidae].
148 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Propeamussium (Propeamussium) interradiatum (Gabb, 1869b). [Pecten 
6, 11. Kreyenhagen Fm., Eocene. Near Idria, San Benito Co. USGS 5742 or 5743 (both cited in text). Hypotypes
USNM 646450, 646451. [Propeamussiidae].
Propeamussium interradiatum (Gabb, 1869b). [Pecten
Fm., Eocene/Oligocene. East of New Idria, Priest Valley quad., San Benito Co. Lectotype (of Stewart, 1930:124)
ANSP 4652 (not ANSP 4442 as in Stewart 1930: p. 124). [Propeamussiidae].
Propeamussium interradiatum (Gabb, 1869b). [Pecten
Dell Shale, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23686. Hypotype LACMIP 11633. [Propeamussiidae].
Propeamussium (Parvamussium) stanfordensis (Arnold, 1906). [Pecten]. Addicott 1971. Veliger 13(3):228, 229,

Remarks: In the genus Parvamussium,  Moore (1984: p. B10). [Propeamussiidae].

USNM 646446.
Propeamussium stanfordensis (Arnold, 1906). [Pecten
6, 7. Gallaway Fm., Oligocene. North of Iverson Pt., Saunders Reef quad., Mendocino Co. UCMP D7149. Hypotypes
UCMP 14438, 14439. Remarks: In the genus Parvamussium,  Moore (1984: p. B10). [Propeamussiidae].
Propeamussium

Protothaca staleyi (Gabb, 1866). [Dosinia
2, 4. Careaga Fm. (Graciosa Member), Pliocene. Purisima Hills, Harris area, Santa Barbara Co. USGS 14752. Hy-
potype USNM 560148. Remarks: In the genus Leukoma,  P. Valentich-Scott (personal communication, 2019).
[Veneridae].
Graciosa Ridge, southeast of Orcutt, Santa Barbara Co. USGS 14618. Hypotype USNM 560149.
Protothaca staleyi (Gabb, 1866). [Dosinia
Pliocene. Palo Alto quad., Santa Clara Co. USGS M1715. Hypotypes USNM 650965, 650966. Remarks: In the
genus Leukoma,  P. Valentich-Scott (personal communication, 2019). [Veneridae].

Protothaca staleyi (Gabb, 1866). [Dosinia
Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M5034. Hypotype USNM 254373.
Remarks: In the genus Leukoma,  P. Valentich-Scott (personal communication, 2019). [Veneridae].
Protothaca staleyi (Gabb, 1866). [Dosinia 

Obispo Co. USGS M5034. Hypotype USNM 254373. Remarks: In the genus Leukoma,  P. Valentich-Scott (per-
sonal communication, 2019). [Veneridae].
Protothaca staleyi (Gabb, 1866). [Dosinia
Santa Cruz Co. CASG 66111. Hypotype CASG 66111.01]. Remarks: In the genus Leukoma,  P. Valentich-Scott
(personal communication, 2019). [Veneridae].
Protothaca staleyi hannibali (Howe, 1922). [Paphia-
cene. Coalinga quad., Fresno Co. UCMP B6545. Hypotype UCMP 36714. Remarks: In the genus Leukoma,  P.
Valentich-Scott (personal communication, 2019). [Veneridae].
Protothaca staminea (Conrad, 1837) [Venus
 
Valley, Sonoma Co. Hypotype CASG determined here as missing. Remarks: In the genus Leukoma,  Coan and
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 149
Valentich-Scott (2012: p. 778). [Veneridae].
Protothaca tenerrima (Carpenter in Gould and Carpenter, 1857). [Tapes]. Cummings et al. 1962. CDMG Bull. 181:204,

Co.. 645-1M. Hypotype determined here as missing. Remarks: In the genus Callithaca,  Coan and Valentich-
Scott (2012: p. 756.) [Veneridae].
Protothaca (Callithaca) tenerrima (Carpenter in Gould and Carpenter, 1857). [Tapes]. Hertlein and Grant 1972. SDSNH
Mem. 2(2B):277–278, pl. LACMIP 305A [=LACMIP 16862]. Hypo-
type LACMIP 4733. Remarks: In the genus Callithaca,  Coan and Valentich-Scott (2012: p. 756). [Veneridae].

473].
3. Cabrillo Canyon, Balboa Park, San Diego, San Diego Co. CASG 1402. Hypotype CASG [=CASG 72558].

Hypotype LACMIP 4735.
Protothaca (Callithaca) tenerrima alta (Waterfall, 1929). [Paphia]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):278–

Hypotype LACMIP 4736. Remarks: Junior synonym of Protothaca (Callithaca) tenerrima (Carpenter in Gould
and Carpenter, 1857),  Coan et al. (2000: p. 376); in the genus Callithaca,  Coan and Valentich-Scott (2012:
p. 756). [Veneridae].
Protothaca

Remarks:Chione sp., Clinocardium
sp., and Macoma sp. [Veneridae].
Psammotreta (Florimetis) biangulata (Carpenter, 1856a). [Scrobicularia]. Stanton 1966. Jour. Paleo. 40(1):30–31,

Hypotype CIT 2093 [=LACMIP 4129]. Remarks: Junior synonym of Psammotreta (Leporimetis) obesa (Deshayes,
1855),  Coan and Valentich-Scott (2012: p. 617). [Tellinidae].
Psammotreta viridotincta (Carpenter, 1856b) [Scrobicularia    
Latrania Fm., Miocene. Andrade Canyon, Imperial Co. [ABDSP 5051.2]. [Hypotype ABDSP I 10987]. Remarks:
Tellina ochracea Carpenter (1864a),  S. Rugh (personal communication, 2019). [Tellinidae].
Psephidia ovalis
Diego Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Hypotypes LACMIP
4737, 4738]. Remarks: In the genus Nutricola,  Coan et al. (2000: p. 383). [Veneridae].
Psephidia stephensae
Diego Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Holotype LACMIP 4739paratypes LACMIP
4740, 4741. Remarks: Junior synonym of Nutricola cymata (Dall, 1913),  Coan et al. (2000: p. 383). [Veneridae].


Psephidia
between Neverita Zone and Cascajo Conglomerate Member], Pliocene. North of El Campo, North Dome, Kettle-

Pseudocardium densatum (Conrad, 1857a) of Arnold. [Mulinia]. Woodring et al. 1940 [1941]. USGS PP 195:93–94,
Pseudocardium-bearing sandstone), Pliocene. South Dome, , Kettleman Hills, Kings
Co. USGS 12641. Hypotype USNM 495361. [Mactridae].
 Siphonalia Zone), Pliocene. Arroyo Pino, North Dome, Kettleman Hills, Kings Co.
USGS 12814. Hypotype USNM 495857.
150 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pseudocardium densatum (Conrad, 1857a). [Mulinia
Southeast of Pleasanton, La Costa Valley quad. (1943), Alameda Co. SU 2739. Hypotype SU 8450 [=CASG 69925].
[Mactridae].
Pseudocardium aff.  (Conrad, 1857a). [Mulinia

UCMP 37647. [Mactridae].
Pseudocardium densatum (Conrad, 1857a). [Mulinia]. Cox et al. 1969b. Treatise Invert. Paleo. N6(2):N601, Fig. E94

504. Hypotype UCMP 11449]. Hypotype missing,  Keen and Bentson 1944: p. 70, however specimen located
at UCMP June 2002,  LTG (senior author). [Mactridae].
-
cality unknown. Hypotypes UCMP 11441,11447.
Pseudocardium densatum densatum (Conrad, 1857a). [Mulinia]. Moore 2003a. USGS PP 1228-F [electronic format]

66542.01. [Mactridae].
Pseudocardium densatum (Conrad, 1857a) of Arnold, vars. [Mulinia]. Woodring et al. 1940 [1941]. USGS PP 195:93–94,
-
man Hills, Kings Co. USGS 12645 and 12802. Hypotypes USNM. 495350, 495351. [Mactridae].
Pseudocardium densatum (Conrad, 1857a) of Arnold, medium-sized var. [Mulinia]. Woodring et al. 1940 [1941].
  Pseudocardium-Anadara Zone), Pliocene. Arroyo Culebrino
area, Middle Dome, Kettleman Hills, Kings Co. USGS 12719. Hypotype USNM 495355. [Mactridae].
 
495356.
Pseudocardium densatum (Conrad, 1857a) of Arnold, short var. [Mulinia]. Woodring et al. 1940 [1941]. USGS PP
Pseudocardium-bearing sandstone), Pliocene. South Dome, Kettleman
Hills, Kings Co. USGS 12641. Hypotypes USNM 495362, 495363.[Mactridae].
Pseudocardium densatum (Conrad, 1857a) of Arnold, var. cf. gabbii Rémond, 1863. [Cardium]. Woodring et al. 1940
Pseudocardium Zone), Pliocene.
Ridge El Perno area, North Dome, Kettleman Hills, Kings Co. USGS 12460. Hypotypes USNM 495798–495806.
Remarks: Junior synonym of Pseudocardium densatum (Conrad, 1857a),  Moore (2003a: p. 4). [Mactridae].
Pseudocardium-bearing conglomerate), Pliocene. Middle Dome, Kettleman Hills,
Kings Co. USGS 12671. Hypotype USNM 495358.

Pseudocardium densatum (Conrad, 1857a) of Arnold, var. cf. gabbii Rémond, 1863. Woodring and Bramlette,1950

Harris, Santa Barbara Co. USGS 14752. Hypotype USNM 560144. Remarks: Junior synonym of Pseudocardium
densatum (Conrad, 1857),  Moore (2003a: p. 4). [Mactridae].
Pseudocardium densatum minor (Arnold, 1909 [1910]). [Mulinia
Temblor Fm., Miocene. Reef Ridge, Garza Canyon area, Reef Ridge quad., Kings Co. USGS 14399. Hypotype USNM
498744. [Mactridae].
Pseudocardium densatum minor (Arnold, 1909 [1910]). [Mulinia]. Moore 2003a. USGS PP 1228-F [electronic for-

Holotype USNM 165601. [Mactridae].
Pseudocardium meganosensis (Clark and Woodford, 1927). [Spisula]. Moore 2003a. USGS PP 1228-F [electronic for-

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 151


Pseudocardiumorthomorpha (Grant and Gale, 1931). [Mactra]. Moore 2003a. USGS PP 1228-F [electronic format]

1, depth 2794 ft. (852 m), near Tipton, Tulare Co. [SDSNH 4413]. Holotype SDSNH 193. [Mactridae].
Pseudocardium pabloensis (Packard, 1916). [Mulinia
of Irvington, La Costa Valley quad. (1943), Alameda Co. SU 3242. Hypotype SU 8451 [=CASG 69926]. [Mactridae].
Pseudocardium pabloensis (Packard, 1916). [Mulinia]. Moore 2003a. USGS 1228-F [electronic format] (family Mac-

Holotype UCMP 11442. [Mactridae].
Pseudocardium panzanum (Loel and Corey, 1932). [Mulinia]. Moore 2003a. USGS PP 1223-F [electronic format]
   
Carrizo Plains, La Panza Mtns., San Luis Obispo Co. UCMP A505. Holotype UCMP 31792 (not USNM as in pl. cap-
tion). [Mactridae].
Pseudocardium Fowkes       1909 [1910],
Mulinia densata (Conrad, 1857a)),
Etchegoin Fm., Miocene. Kings Co. Hypotype USNM 165531. Remarks: Determined here as Pseudocardium den-
satum (Conrad, 1857a). [Mactridae].
Re-
marks: Determined here as Pseudocardium densatum (Conrad, 1857a).

Remarks: Determined here as Pseudocardium densatum (Conrad, 1857a).
Pseudocardium

Pteria berryi
below the top of ridge along north bank of Garza Creek, Reef Ridge quad., Kings Co. UCMP D1063. Holotype UCMP
36611, paratype UCMP 36612. [Pteriidae].
Pteria berryi 
Ridge, Fresno Co. UCMP D1063. Holotype UCMP 36611. [Pteriidae].
Pteria clarki      
Eocene. Near Tecuya Creek, Tejon quad., Kern Co. UCMP 460. Hypotype UCMP 15487. [Pteriidae].


Hypotypes LACMIP 12749, 13480.

32623.
Pteria hertleini
East side of Vaqueros Creek, Junipero Serra quad., Monterey Co. SU 200 [=CASG 71528]. Holotype SU 5160
[=CASG 71528]. [Pteriidae].
Pteria howei
Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCMP 3767. Holotype UCMP 30515. Remarks: Figured by

152 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pteriahowei  
quad., Ventura Co. UCMP 3783. Paratype UCMP 30516. [Pteriidae].
Pteria jordani
area, south of Calabasas, Santa Monica Mtns., , Calabasas quad., Los Angeles Co. SU 425 [=SDSNH 425,  Keen
and Bentson (1944: p. 104]. Holotype SDSNH 25. [Pteriidae].
Pteria pellucida (Gabb, 1864). [Avicula]. Weaver and Kleinpell 1963. UCPG 43:197, pl
Sandstone, Eocene. Along., Santa Barbara Co.
UCMP B6940. Hypotype SU 9257 [=CASG. 70230]. Remarks: Pteria clarki,  Squires (2014b: p. 227). [Pteriidae].
Pteria pellucida (Gabb, 1864). [Avicula 
Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4714. Hypotype UCR 4714/6. Remarks: Pteria clarki,
 Squires (2014b: p. 227). [Pteriidae].
Pteria pellucida (Gabb, 1864). [Avicula
Near Martinez, Contra Costa Co. CGS 198]. Lectotype (of Vokes 1939:50) UCMP 11983. Remarks: Pteria clarki,
 Squires (2014b: p. 227). [Pteriidae].
Pteria pellucida (Gabb, 1864). [Avicula-
royo, Santa Susana quad. (1951, PR 1969). Ventura Co. CSUN 548 [=LACMIP 15450]. Hypotype LACMIP 12749.
Remarks: Pteria clarki,  Squires (2014b: p. 227). [Pteriidae].
Pteria rositae       
Vicinity of San Augustine Canyon, Santa Rosa Is., California Channel Ids. CASG 1156. Holotype CASG 4142. [Pte-
riidae]. Remarks: Junior synonym of Pteria sterna, Coan et al. (2000: p. 193). [Pteriidae].
Pteria sterna (Gould, 1851) [Avicula
Fm., Miocene. Andrade Canyon, Imperial Co. ABDSP5050.3. Hypotype ABDSP I 10967. [Pteriidae].
Pteria    
Eocene. Deer Valley, Mt. Diablo quad., Contra Costa Co. UCMP 3152. Hypotype UCMP 31321. Remarks: Also
Pteria” sp., 
Pteria
quad., Santa Monica Mtns., Los Angeles Co. LACMIP 17868. Hypotype LACMIP 13478. [Pteriidae].
Pulvinites californica     Pulvinites
 in pl. caption]. [Santa Susana Fm.], Paleocene. Meier Canyon, Simi Hills, Ventura Co. UCR 6821. Holotype

Pulvinites californica
Fm., Paleocene. Meier Canyon, Simi Hills, Ventura Co. UCR 6821. Holotype UCR 6821/19. [Pulvinitidae].
Pulvinites californicaPul-
 in pl. caption]. Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. [UCR 6821]. Hypotype UCR
6821/20 (not UCR 7342/1 as in pl. caption). [Pulvinitidae].
 Zinsmeister, 1978b. (6 (as Pulvinites californica in pl.
caption). Santa Susana Fm., Paleocene. Near Meier and Runkle canyons, Ventura Co. UCR 6900. Holotype UCR.
7342/1 (not UCR 6900/101 as in text), paratype UCR 6900/102. [Pulvinitidae].




(of Pedalion sp.). UCMP 30513. [Pulvinitidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 153

Pulvinites
californica in pl. caption). Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 7342. Holotype UCR 7342/1
(not UCR 6900/4 as in pl. caption). [Pulvinitidae].
PycnodonteCrenostrea eldridgei (Arnold, 1907). [Ostrea
Vaqueros Fm., Oligocene/Miocene. East of Grimes Canyon, near Filmore, Piru quad. Ventura Co. Holotype USNM

Pycnodonte (Phygraea) cf.  Squires and Demetrion, 1990. Squires 2001. NHMLAC CS

LACMIP 7405. Hypotype LACMIP 12750. Remarks: Phygraea stewarti (M.A. Hanna 1927),  Squires (2018:
p. 18). [Gryphaeidae].
Pycnodonte (Pycnodonte heermanni (Conrad, 1855a). [Ostrea
Imperial Fm. Latrania Sand member [=Latrania Fm.], Miocene/Pliocene. Garnet Hill area, Riverside Co. CIT 192
[=LACMIP 16347]. Hypotype LACMIP 7162 [=LACMIP 7867], determined here as missing. Remarks: Supple-
mental description in Conrad (1855b). [Gryphaeidae].
Pycnodonte (Pycnodonte heermanni (Conrad, 1855a). [Ostrea

Syntypes (as holotype in text) ANSP 13367, 13367b. Remarks: Supplemental description in Conrad (1855b).
[Gryphaeidae].
Pycnodonte heermani [sic] (Conrad, 1855a). [Ostrea]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p.

Co. SDSNH. 360. Hypotype SDSNH 23331. Remarks: Supplemental description in Conrad (1855b).[Gryphaeidae].
Pycnodonte (Pycnodonte howelli (Wiedey, 1928). [Ostrea
7. Vaqueros Fm., Oligocene/Miocene (Temblor Fm. in locality description). Near Sespe Creek, Mt. Pinos quad.,
Ventura Co. SU 446 [251 on specimen label]. Syntypes CASG. 2981, 2982. [Gryphaeidae].
Pycnodonte cf. P (Wiedey, 1928) [Ostrea]. Stadum and Finger 2016. NHMLAC CS-
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad., SDSNH 4312. Hypotype SDSNH 83234. [Gryphaeidae].
PycnodontePycnodonte loeli (Hertlein, 1928a). [Ostrea
Vaqueros Fm., Oligocene/Miocene. Santa Rosa Is., California Channel Ids. CASG 1153. Holotype CASG 4117.
[Gryphaeidae].
Pycnodonte (Phygraea) sp., aff. Pycnodonte (Phygraea) Squires and Demetrion, 1990. Squires 1999b. NHM-

(1952, PR 1967), Ventura Co. CSUN 1343 [=LACMIP 17186]. Hypotypes LACMIP 12696, 12697. Remarks: In the
genus Phygraea,  Squires (2018a: p. 17). [Gryphaeidae].
Pycnodonte (Crenostrea stewarti (M.A. Hanna, 1927). [Ostrea 

Co. UCMP 5062. Syntypes UCMP 30921, 30922. Remarks: In the genus Phygraea,  Squires (2018a: p. 18).
[Gryphaeidae].
Pycnodonte (Pycnodonte wiedeyi (Hertlein, 1928a). [Ostrea
1, 4, 5, 8. Vaqueros Fm., Oligocene/Miocene. Vicinity of San Augustine Canyon, Santa Rosa Is., California Channel
Ids. CASG 1154. Holotype CASG 4129, paratype CASG 4130. [Gryphaeidae].
Pycnodonte (Pycnodonte) yneziana (Loel and Corey, 1932). [Ostrea]. Moore 1987. USGS PP 1228-C:C18–C19, pl.
        
[Gryphaeidae].
Raetaplicatella longior (Grant and Gale, 1931). [Anatina]. Moore 2003a. USGS PP 1228-F [electronic format] (family
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
Raeta transmontana (Conrad, 1857a). [Lutraria]. Moore 2003a. USGS 1228-F [electronic format] (family Mactri-

[Mactridae].
Saccella cellulita (Dall, 1896a). [Leda
Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., brachiopod bed, Santa Barbara Co. USGS 14770. Hypotype
USNM 560095. Remarks: In the genus/subgenus Nuculana (Jupiteria),  Coan et al. (2000: p. 88). [Nuculanidae].
Saccella ochsneri (Anderson and Martin, 1914). [Leda  
Temblor Fm., Miocene. Kern River about, Caliente quad., Kern Co. CASG Holotype CASG 103 [=CASG 68.01],
paratype CASG 105 [=CASG 68.04]. Remarks: In the genus/subgenus Nuculana (Saccella),  Moore (1983: p.
B19). [Nuculanidae].
Saccella orcutti (Arnold, 1907). [Leda]. Woodring and 7, 8,
13. Sisquoc Fm. (diatomaceous strata), Miocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14902.
Hypotypes (4) USNM 560023. Remarks: In the genus/subgenus Nuculana (Saccella),  Moore (1983: p. B20).
[Nuculanidae].

Saccella redondoensis (Burch, 1944). [Nuculana]. Woodring and Bramlette 1950 [1951]. USGS PP 222:81, pl. 16,
   
Hypotype USNM 560096. Remarks: Junior synonym of Nuculana (Jupiteria) penderi (Dall and Bartsch, 1910),
 Coan et al. (2000: p. 89). [Nuculanidae].
Saccella taphria (Dall, 1896b) [Leda       
Careaga Fm. (Graciosa Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14614. Hypotype
USNM 560134. [Nuculanidae].
Saccella taphria (Dall, 1896b). [Leda       
Pliocene. Balboa Park, San Diego, San Diego Co. SDSNH 1965. Hypotypes SDSNH 4167, 4168]. [Nuculanidae].
Sanguinolaria alata (Gabb, 1866). [Gari (Psammocola)
Southeast of Pleasanton, La Costa Valley quad. (1943), Alameda Co. SU 2739. Hypotype SU 8467 [=CASG 69934].
Remarks: In the genus Nuttallia,  Moore (2003b: p. 8). [Psammobiidae].
Sanguinolaria cf. 
(Patinopecten Zone), Pliocene. Vicinity of Arroyo Doblegado, southeast of Las Paredes, North Dome, Kettleman
Hills, Kings Co. USGS 12426. Hypotype USNM 495847. Remarks: In the genus Nuttallia,  Coan et al. (2000:
p. 429). [Psammobiidae].
Sanguinolaria cf. 
Castaic Canyon, Los Angeles Co. CIT 2093 [=LACMIP 11853. Hypotype LACMIP 4126]. Remarks: In the genus
Nuttallia,  Coan et al. (2000: p. 429). [Psammobiidae].
Saulella undulifera (Gabb, 1869b). [Tellina
Paleocene. Simi Hills, Ventura Co. UCLA5996 [=LACMIP 25996]. Hypotype (not holotype as in text) UCLA 59255
[=LACMIP 10984]. [Tellinidae].

Saulella undulifera (Gabb, 1869b). [Tellina

4551], missing,  K. Estes-Smargiassi (personal communication, 2019). [Tellinidae].
Saulella undulifera (Gabb, 1869b). [Tellina
Eocene. North side Simi Valley, Santa Susana quad. (1951., PR 1969), Ventura Co. LACMIP 26610 [=CSUN 967].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 155
Hypotype LACMIP 12699. [Tellinidae].
Saxidomus sp. aff.  Conrad, 1837. Moore San Diego Fm.,
Pliocene. Balboa Park, San Diego, San Diego Co. SDSNH 1965. Hypotype SDSNH 4157]. [Veneridae].
Saxidomus nuttalli  
Pliocene. Near U.S./Mexico boundary, San Diego Co. UCLA 312 [=LACMIP 20312]. Hypotype UCLA 48622 [=LAC-
MIP 9305]. [Veneridae].
Saxidomus nuttalli Conrad, 1837. Fowkes 1982. Coalinga Guidebook, p. 33 (as Saxidomus), p. 206, unnumbered

Jacalitos Fm., Miocene. At northwest end of Anticline Ridge, 6 mi. north-northeast of Coalinga, Fresno Co. Hypo-
type USNM 165529. [Veneridae].
Saxidomus nuttalli
Fm.], Pliocene. North side of Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 725 [=LACMIP
29227]. Hypotype UCLA 59410 [=LACMIP 8323]. [Veneridae].
Saxidomus nuttali Conrad, 1837. 
Angeles Co. LACMIP 17921. Hypotype LACMIP 14361. [Veneridae].
Saxidomus nuttalli latus Stewart in-
goin Fm., Pliocene. Coalinga quad., Fresno Co. UCMP B6546. Hypotype UCMP 36701. Remarks: Junior synonym
of Saxidomus nuttalli Conrad, 1837,  Coan et al. (2000: p. 386). [Veneridae].
Saxidomus nuttalli latus Stewart in Woodring et al., 1940 [1941]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):272,

Hypotype CASG 70447. Remarks: Junior synonym of Saxidomus nuttalli Conrad, 1837,  Coan et al. (2000: p.
386). [Veneridae].
Saxidomus vaquerosensis-
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4312. Hypotype SDSNH 180184. [Veneridae].
Saxolucina sp. or a variant of Eomiltha turneri (Stanton, 1896). [Lucina]. Smith 1975b. SEPM Symposium vol., p.

SU 2073 [=UCMP A9717]. Hypotype SU 10233 [=CASG 61666.16]. [Lucinidae].
Scapharca (Cunearca) hamelini (Wiedey, 1928). [Arca
Fm., Miocene. Near junction of Little Sespe River with Big Sespe River, Ventura Co. SDSNH 407. Holotype SDSNH
18. [Arcidae].
Scapharca (Scapharca obispoana obispoana (Conrad, 1857d). [Arca]. Moore 1983. USGS PP 1228A:A44, pl. 9,

1931:955) USNM 13330. [Arcidae].
ScapharcaScapharca obispoana pedisparis (Wiedey, 1928). [Arca]. Moore 1983. USGS PP 1228-A:A44, 
6. Monterey Fm., Miocene. Vicinity of Zayante railroad station, Santa Cruz Co. SDSNH 443. Holotype SDSNH 23.
[Arcidae].
Scapharcaobispoana perdisparis (Wiedey, 1928). [Arca      

101095]. [Arcidae].
Scapharca (Cunearca) rivulata (Wiedey, 1928). [Arca
Temblor Fm. (Santos Shale Member) (as Monterey Fm. in text), Miocene. Near Simmler, La Panza quad., San Luis
Obispo Co. SU 432. Holotype SDSNH 432. Remarks: The number 432 was used independently by SU and SDSNH
for the same locality and is not an error in cataloguing (Kesler Randall, personal communication, 2020). [Arcidae].
Scapharca (Cunearca) santaclarana (Loel and Corey, 1932). [Arca]. Moore 1983. USGS PP 1228-A:A44–A45, pl. 9,
156 PALEOBIOS, VOLUME 38, FEBRUARY 2021

UCMP 31765. [Arcidae].
Schizothaerus cf.  (Conrad, 1837). [Lutraria (Cryptodon)]. Woodring et al. 1940 [1941]. USGS PP 195:94
(as Saxidomus nuttalliAcila Zone), Pliocene. El Perro, North Dome, Kettleman
Hills, Kings Co. USGS 12368. Hypotype USNM 495726. Remarks: In the genus Tresus, Coan et al. (2000: p.
463). [Mactridae].
Siphonalia Zone), Pliocene. Los Jinetes, North Dome, Kettleman Hills, Kings Co. USGS
12393. Hypotype USNM 495829.
Schizothaerus nuttalli (Conrad, 1837). [Lutraria (Cryptodon)Schizo-
thaerus cf. 
Vicente Creek, San Mateo Co. UCMP B4790 (not UCMP B4807 as in text). Hypotype UCMP 37648. Remarks: In
the genus Tresus,  Coan et al. (2000: p. 463). [Mactridae].
Schizothaerus pajaroanus (Conrad, 1857d). [Venus
Pliocene. Annapolis quad., Sonoma Co. UCMP B2055. Hypotype UCMP 12559. Remarks: In the genus Tresus, 
Adegoke (1969: p. 123). [Mactridae].
Schizothaerus pajaroanus (Conrad, 1857d). [Venus       
Purisima Fm. (Pomponio Member), Pliocene. Tunitas Rd. area, Woodside quad., San Mateo Co. 657M. Hypotype
determined here as missing. Remarks: In the genus Tresus,  Adegoke (1969: p. 123) [Mactridae].
Securella cryptolineata (Clark, 1918). [Chione

UCMP 1131. Holotype UCMP 11178, holotype (of Chione mediostriata Clark 1918) UCMP 11181, hypotypes
UCMP 36090–36095. [Veneridae].
Securella cf.  (Clark, 1918). [Chione
(Wygal Sandstone Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypo-
type USNM 646583. [Veneridae].
Securella elsmerensis (English, 1914a). [Chione
from Grant and Gale 1931, pl. 16 Co. SDSNH 202.
Lectotype (of Parker 1949:590) SDSNH 151, hypotype SDSNH 152. [Veneridae].

 Keen and Bentson (1944: p. 36); 
Parker (1949: p. 590).
….., Jacalitos Fm. [Etchegoin Fm. as per UCMP online database], Pliocene. Coalinga quad., Fresno Co. UCMP 2655.

Securella juanensis (Loel and Corey, 1932). [Chione
Fm., Miocene. La Panza quad., San Luis Obispo Co. UCMP A505. Holotype UCMP 31803. [Veneridae].
Securella margaritana (Anderson and Martin, 1914). [Chione
Margarita Fm., Miocene. Domengine Ranch quad., Fresno Co. UCMP D1102. Hypotype UCMP 36696. [Veneridae].
Securella panza (Anderson and Martin, 1914). [Chione

72123, 72124]. [Veneridae].
Chione val-
entinei Wiedey, 1928) SU 6956 [=CASG 72125].
Securella panza (Anderson and Martin, 1914). [Chione
Fm., Miocene. Garza Creek area, Reef Ridge quad., Kings Co. UCMP B6511. Hypotype UCMP 36695. [Veneridae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 157
Securella securis (Shumard, 1858). [Venus-
cene. Humboldt Co. Hypotype UCMP 31961. [Veneridae].
Semele ashleyi-
cene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Paratype LACMIP 4790,
Remarks: Junior synonym of Semele (Semele) rubropicta Dall, 1871,
 Coan et al. (2000: p. 432). [Semelidae].



Semele (Semele) ashleyi Hertlein and Grant, 1972. Moore 2003b. USGS PP 1228-G [electronic format] (family Semeli-
sic], San Diego Co. Holotype LACMIP 4789.
Remarks: Junior synonym of Semele (Semele) rubropicta Dall, 1871,  Coan et al. (2000: p. 432). [Semelidae].
Semele diabloi Dickerson, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Semelidae):6, pl. 11,
 Fm.”, Eocene. Mt. Diablo quad., Contra Costa Co. UCMP 469. Holotype UCMP 11654. [Semelidae].
Semele (Semele) fausta Nomland, 1917a. Moore 2003b. USGS PP 1228-G [electronic format] (family Semelidae):2,
Semele fausata). Etchegoin Fm., Miocene/Pliocene. Zapato Creek, Fresno Co. UCMP 2991. Holo-
type UCMP 11102. [Semelidae].
Semele (Semele) gayi Arnold, 1908. Moore 2003b. USGS PP 1228-G [electronic format] (family Semelidae):1, pl. 11,

Cruz quad., San Mateo Co. Holotype USNM 165435. [Semelidae].
Semele morani Anderson and Martin, 1914. Moore 2003b. USGS PP 1228-G [electronic format] (family Semelidae):6,

Semele cf. Acila
Zone), Pliocene Arroyo Hondo, North Dome, Kettleman Hills, Kings Co. USGS 12340. Hypotype USNM 495719.
Remarks: Semele rubropicta Dall 1871, Adegoke (1969: p. 125). [Semelidae].
Semele rubropicta

USNM 165517. [Semelidae].
Semele cf. 
Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. Alex Clark locality. Hypotype USNM 560104. Re-
marks: Semele rubropicta, Adegoke 1969: p. 125. [Semelidae].
Semele rubropicta
Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817. Hypotype LACMIP 4806. [Se-
melidae].

SemeleSiphona-
lia Zone), Pliocene. Arroyo Robador, east of Cerro Ultimo, North Dome, Kettleman Hills, Kings Co. USGS 12413.
Hypotype USNM 495824. [Semelidae].
Semele
quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Hypotype LACMIP 4805. Remarks: Probably a
juvenile of Semele ashleyi n. sp.,  Hertlein and Grant (1972: p. 300). [Semelidae].
Septifer bifurcatus (Conrad, 1837). [Mytilus
12. San Diego Fm., Pliocene. West of Goat Canyon, San Ysidro quad. (1953), San Diego Co. SDSNH 47 [=SDSNH
5047]. Hypotype SDSNH [4319. [Mytilidae].
158 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Septifer (Septifer) bifurcatus (Conrad, 1857d). [Mytilus  . 7, 9. San
Diego Fm., Pliocene. West of Goat Canyon, San Ysidro quad. (1953), San Diego Co. SDSNH 47 [= SDSNH 5047].

Septifer (Septifer) coalingensis
Fm., Miocene/Pliocene. Near Coalinga, Fresno Co. USGS 4634. Holotype USNM 165580. [Mytilidae].
Septifer (Septifer) elegans
North side of Simi Valley, Ventura Co. CASG 322 [=CASG 61913]. Neotype (of Moore 1983: p. A70) SU 322 [=CASG
61913.07, as syntype in CASG collection]. [Mytilidae].
Septifer (Septifer) cf. () elegans
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 364 [=LACMIP 16306]. Hypotype LACMIP 7511. [Mytilidae].
Septifer Fowkes 1982. Coalinga Guidebook, p. 21, unnumbered ig. Domengine Fm., Eocene. West of Coalinga, Fresno

Septifer
southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH 20769. [Mytilidae].
Siliqua cf.  (Broderip and G.B. Sowerby I, 1829). [Solen

UCMP B4790. Hypotype UCMP 37639. [Pharidae].
Siliqua lucida (Conrad, 1837). [Solecurtus
Fm., Pliocene. Reynard Wy., San Diego, San Diego Co. SDSNH 43. Hypotype SDSNH 4350. [Pharidae].
Siliqua lucida (Conrad, 1837). [Solecurtus]. Moore 2003a. USGS 1228-F [electronic format] (family Pharidae):3–4,


Siliqua cf.  (G.B. Sowerby I, 1839). [Solen]. Woodring and Bramlette 1950 [1951]. USGS PP 222:91, pl. 7,

15008. Hypotype USNM 560018. Remarks: Junior synonym of Siliqua alta (Broderip and G.B. Sowerby I, 1829),
 Coan et al. (2000: p. 448). [Pharidae].

Siliqua oregonia
quad., Humboldt Co. UCMP B7908. Hypotype UCMP 37788. [Pharidae].
SiliquaSiphonalia Zone), Pliocene.
Arroyo Torcido, North Dome, Kettleman Hills, Kings Co. USGS 14108. Hypotype USNM 495823. [Pharidae].
Siliqua sp. Bedrossian 1974, lower specimen.
]. Santa Rosa Valley, Sonoma Co. Hypotype CASG
determined here as missing. [Pharidae].
Simomactra falcata brioniana (Trask, 1922). [Spisula]. Moore 2003a. USGS 1228-F [electronic format] (family Mac-
Simomactra falcata barioniana in pl. caption). Briones Sandstone, San Pablo Group,
Miocene. South of Walnut Creek Concord quad., Contra Costa Co. UCMP 3522. Holotype UCMP 12380. [Mactridae].
Solemya ventricosa, auct-
boldt Co. Hypotype not located at CASG where Talmadge deposited numerous other specimens. [Solemyidae].
Solendomenginicus Vokes, 1939. Moore 2003a. USGS 1228-F [electronic format] (family Solenidae):3–4, pl. 14,

15711. [Solenidae].
Solen aff. 
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 159
Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM 646557.
[Solenidae].
Solen perriniMya
Zone), Pliocene. Las Alturas, Middle Dome, Kettleman Hills, Kings Co. USGS 12685. Hypotype USNM 495294.
Remarks: Junior synonym of Solen (Ensisolen) sicarius Gould, 1850,  Coan et al. (2000: p. 445). [Solenidae].
Neverita Zone and Cascajo Conglomerate Member), Pliocene.
North of El Campo, North Dome, Kings Co. USGS 12841. Hypotype USNM 495338.
Solen rosaceus
Pliocene. Mt. Soledad, San Diego Co. Hypotype SDSNH 5006 [=SDSNH 4349]. Remarks: Determined by P. Don
Vito and T. Deméré (SDSNH) to be Solena sp. from SDSNH 8, an undetermined Eocene fm. from the base of Mt.
Soledad. [Solenidae].
Solen sicarius      
Pliocene. South slope of Mt. Soledad, San Diego Co. SDSNH 4. Hypotype SDSNH 5005 [=SDSNH 4351]. Remarks:
In the subgenus Ensisolen,  Coan et al. (2000: p. 445). [Solenidae].
Solen sicarius-
tola Beach], Santa Cruz Co. SCCM1. Hypotype SCCM 9685.02]. Remarks: In the subgenus Ensisolen,  Coan
et al. (2000: p. 445). [Solenidae].
Solen sicarius
Co. CASG 66175. Hypotype CASG 66175.01]. Remarks: In the subgenus Ensisolen,  Coan et al. (2000: p. 445).
[Solenidae].
Solen (Ensisolen) sicarius Fm., Pliocene. Newhall area,
Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14368. [Solenidae].
Solen sp., cf. 
Angeles Co. LACMIP 17917. Hypotype LACMIP 14369. Remarks: Junior synonym of Solen (Ensisolen) sicarius
Gould, 1850,  Coan et al. (2000: p. 445).
Solena (Eosolen) coosensis
Valley quad. (1943), Ventura Co. UCR 4676. Hypotype UCR 4676/22. Remarks: Junior synonym of Solena (Eo-
solen) novacularis (Anderson and G D. Hanna, 1925),  Squires (1987: p. 67). [Solenidae].
Solena coosensis
Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH 20781.
Remarks: Solena (Eosolen) novacularis (Anderson and G D.Hanna, 1925),  Squires (1987: p. 67). [Solenidae].
Solena (Plectosolen) gravidus gravidus (Clark, 1918). [Solen]. Moore 2003a. USGS PP 1228-F [electronic format]

Contra Costa Co. UCMP 1131. Holotype UCMP 11133. [Solenidae].
Solena (Plectosolen) gravidus baileyi (Loel and Corey, 1932). [Solen]. Moore 2003a. USGS PP 1228-F [electronic

end of South Mtn., Santa Paula quad., Ventura Co. UCMP A244. Holotype UCMP 31831. [Solenidae].
Solena (Eosolen lorenzanus (Wagner and Schilling, 1923). [Solen]. Moore 2003a. USGS PP 1228-F [electronic

Muddy Creek, Mt. Pinos quad., Kern Co. UCMP 3212. Holotype UCMP 11421. [Solenidae].
Solena (Eosolen) novacularis (Anderson and G D.Hanna, 1925). [Solen]. Squires 1984. NHMLAC CS 350:50, Fig. 12d.
Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP
7242]. Hypotype LACMIP 6585. [Solenidae].
Solena (Eosolen) novacularis (Anderson and G D. Hanna, 1925). [Solen
115. Juncal Fm., Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype
160 PALEOBIOS, VOLUME 38, FEBRUARY 2021
LACMIP 7532. [Solenidae].
Solena (Eosolen) novacularis (Anderson and G D. Hanna, 1925). [Solen]. Squires 1988a. SEPM Guidebook 58:110, pl.

LACMIP 6585. [Solenidae].
Solena (Eosolen) novacularis (Anderson and G D. Hanna, 1925). [Solen]. Moore, 2003b. USGS PP 1228-F [electronic

Co. CASG 792. Holotype CASG 882 [=CASG 792.01]. [Solenidae].
Solena (Eosolen) perrini (Clark, 1915). [Solen]. Moore 2003a. USGS 1228-F [electronic format] (family Soleni-
dae):9–10, Walnut Creek, Contra Costa Co. UCMP 1478. Holo-
type UCMP 11564. Remarks: Junior synonym of Solen (Ensisolen) sicarius Gould, 1850,  Coan et al. (2000:
p. 445). [Solenidae].
Solena (Eosolen) stantoni (Weaver, 1905). [Solen
Susana Fm., Paleocene. Simi Hills, Ventura Co. [UCR 6665]. Hypotype UCR 6665/1 (not UCR 6834/3 as in pl.
caption). [Solenidae].
Solena (Eosolen) stantoni (Weaver, 1905). [Solen
Santa Susana Fm., Paleocene. Upper Pulga Canyon, Palisades Highlands, east-central Santa Monica Mtns., Topanga
quad. (1952, PR 1981), Los Angeles Co. LACMIP 16897. Hypotype LACMIP 7943. [Solenidae].
Solena (Eosolen) stantoni (Weaver, 1905). [Solen]. Moore 2003a. USGS 1228-F [electronic format] (family Soleni-

Concord quad., Contra Costa Co. UCMP 532. Holotype UCMP 11941. [Solenidae].
Solena (Eosolen) subverticulata Vokes, 1939. Moore 2003a. USGS 1228-F [electronic format] (family Solenidae):8,

Holotype UCMP 15713. [Solenidae].
Sphenia cf.  (Valenciennes, 1846). [Corbula]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):321–322, pl. 56,

305. Hypotype LACMIP 4826. [Myidae].
Spisula (Stereomactra) abcissa (Gabb, 1866). [Schizodesma

UCMP 1608. Hypotype UCMP 11472. Remarks: In the genus Stereomactra,  Moore (2003a: p. 31). [Mactridae].

ANSP 4548].
Spisula cf.  (Conrad, 1848). [Mactra
(Wygal Sandstone Member), MioceneCarneros Rocks quad., Kern Co. USGS M3578. Hypotypes USNM 646552,
646553. Remarks: In the genus Mactromeris,  Moore (2003a: p. 16). [Mactridae].

Spisula albaria coosensis
(as Spisula labaria coosensis
USNM 650971–650973, 650985. Remarks: In the genus Mactromeris,  Moore (2003a: p. 18). [Mactridae].
Spisula albaria coosensis Howe, 1922. Perry 1977. Foss. Pliocene.
Near Capitola Beach, Santa Cruz Co. SCCM 1. Hypotype SCCM 9686. Remarks: In the genus Mactromeris, 
Moore (2003a: p. 18). [Mactridae].
Spisula albaria coosensis-
tocene. Twelvemile Creek area, San Mateo Co. USGS M5754. Hypotype USNM 251803. Remarks: In the genus
Mactromeris,  Moore (2003a: p. 18). [Mactridae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 161
Spisula albaria var. selbyensis
Near junction of Norris Creek with Crow Creek, Hayward quad., Alameda Co. UCMP A4565. Hypotypes UCMP
34143, 34144. Remarks: Mactromeris albaria selbyensis (Packard, 1916),  Moore (2003a: p. 18). [Mactridae].
Spisula bisculpturata
San Guillermo quad. (1943), Ventura Co. UCR 4743. Hypotype UCR 4743/701. Remarks: In the genus Mactrom-
eris,  Moore (2003a:p. 9). [Mactridae].
Spisula bisculpturata
Mission Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962.
Hypotype SDSNH 20766. Remarks: In the genus Mactromeris,  Moore (2003a: p. 9). [Mactridae].
Spisula (Hemimactra) brevirostrata-
cene. Scotia quad., Humboldt Co. UCMP B7642. Hypotype UCMP 37790. Remarks: In the genus Mactromeris,
 Moore (2003a: p. 21). [Mactridae].
Spisula catilliformis
Contra Costa Co. UCMP 305. Hypotype UCMP 34145. Remarks: In the genus Mactromeris,  Coan et al. (2000:
p. 454). [Mactridae].

Spisula (Mactromeris) catilliformis
5. San Diego Fm., Pliocene. Balboa Park, San Diego, San Diego Co. LACMIP 107. Hypotype LAMCIP 4812. Remarks:
In the genus Mactromeris,  Coan et al. (2000: p. 454). [Mactridae].

4813.
Spisula coosensis
Santa Cruz Co. CASG66151. Hypotype CASG 66151.01]. Remarks: Mactromeris albaria coosensis (Howe 1922),
 Moore (2003a: p. 18). [Mactridae].
Spisula aff.  (Clark, 1925) [Mulinia]. Powell et al. 2019b. PaleoBios-
cene. Caldecott Tunnel excavation, Berkeley Hills, Oakland Hills, Alameda Co. UCMP IP13004. Hypotype UCMP
410439. [Mactridae].
Spisula (Mactromeris) cf. () falcata (Gould, 1850). [Mactra]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):314,
Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 4814.
Remarks: In the genus Simomactra,  Coan et al. (2000: p. 458). [Mactridae].
Spisula gaviotaensis* (Conrad, 1857d). [Mactra
Fm., Eocene. East of Tecolote Canyon, Goleta quad., Santa Barbara Co. UCMP B6979. Hypotype SU 9314 [=CASG
70233]. Remarks: Error for gabiotensis, but likely a misprint in original description as the type locality is listed
as Gaviote Pass. Provisionally in the genus Mactromeris Moore (2003a: p. 26). [Mactridae].
Spisula (Hemimactra) hemphilla* (Dall, 1894). [Mactra

UCMP 37642 determined herea as missing. Remarks: Error for In the genus Mactromeris,  Moore
(2003a: p. 23). [Mactridae].
Spisula (Mactromeris) hemphillii (Dall, 1894). [Mactra]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):315–316,

Diego Co. LACMIP 305. Hypotypes LACMIP 4815, 4816, 4819. Remarks: In the genus Mactromeris,  Coan et
al. (2000). [Mactridae].

Ensis myrae Berry 1953a, hypotype LACMIP 4811)].

162 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Spisula (Mactromeris) mercedensis
San Diego Fm., Pliocene. Balboa Park, San Diego, San Diego Co. LACMIP 107. Hypotype LACMIP 4821. Remarks:
This specimen is not from the San Diego Fm. According to accession records, it is from the Johnson collection and
is from the San Benito River area, San Benito Co. [=LACMIP 2960] Junior synonym of Mactromeris catilliformis
(Conrad, 1867),  Coan et al. (2000: p. 454). [Mactridae].
Spisula merriami
Ridge, southeast of Big Tar Canyon, Reef Ridge quad., Kings Co. USGS 14495. Hypotype USNM 498724. Remarks:
Provisionally in the genus Mactromeris Moore (2003a: p. 25). [Mactridae].
Spisula merriami
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 828 [=LACMIP 16141]. Hypotype LACMIP 7531. Remarks:
Provisionally in the genus Mactromeris Moore (2003a: p. 25). [Mactridae].
Spisula (Hemimactra) mossbeachensis

B4790. Holotype UCMP 37643, paratype UCMP 37644. Remarks: In the genus Mactromeris,  Moore (2003a:p.
19). [Mactridae].
Spisula packardi
East of Tecolote Canyon, Goleta quad., Santa Barbara Co. UCMP B6979. Hypotypes SU 9315, 9316 [=CASG 70238,
70239]. Remarks: In the genus Pseudocardium,  Moore (2003a: p. 3). [Mactridae].
Spisula (Mactromeris) cf.  () planulata (Conrad, 1837). [Mactra]. Hertlein and Grant 1972. SDSNH mem.
Spisula (Mactromeris) cf. M. (M.) planulata]. San Diego Fm., Pliocene. Near U.S./
Mexico boundary, San Diego Co. UCLA 294 [=LACMIP 20294]. Hypotype UCLA 48634 [=LACMIP 9317]. Remarks:
In the genus Simomactra,  Coan et al. (2000: p. 458). [Mactridae].
Spisula ramonensis
Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM 646555.
Remarks: In the genus Mactromeris,  Moore (2003a: p. 10). [Mactridae].
Spisula cf. 
Sandstone Member), Miocene. Vicinity of Zemorra Creek, Carneros Rocks quad., Kern Co. USGS M3979. Hypotype
USNM 646562. Remarks: In the genus Mactromeris,  Moore (2003a: p. 11). [Mactridae].
Spisula (Hemimactra) voyi (Gabb, 1866). [Callista-
cene. Scotia quad., Humboldt Co. UCMP B7871. Hypotype UCMP 37789. Remarks: In the genus Mactromeris,
 Moore (2003a: p. 21). [Mactridae].
Spisula
Carneros Rocks quad., Kern Co. USGS M3579. Hypotype USNM 646548. [Mactridae].
Spisula
[=Wilson Grove Fm.], Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotype CASG, determined here as
missing. [Mactridae].
Spisula
-
numbered hypotype lost,  C. Throckmorton (personal communication, 2002). [Mactridae].
Spondylus bostrychites

UCMP 32279, 32280. [Spondylidae].
Spondylus bostrychites-
trania Fm., Miocene/Pliocene. Alverson Canyon, Imperial Co. SDSNH 2995. Hypotype SDSNH 23240. [Spondylidae].
Spondylus carlosensis        
Eocene. Nojoqui Canyon, Santa Barbara Co. UCMP B6932. Hypotype SU 9270 [=CASG 70234]. [Spondylidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 163
Spondylus carlosensis
quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 6571. [Spondylidae].
Spondylus carlosensis Anderson, 1905. Moore 1987. USGS PP 1228-C:C6–C7, pl. 1
West and north of Coalinga, Fresno Co. CASG 66738]. Holotype CASG 56 [=CASG 66738.01]. [Spondylidae].
Spondylus cf. 

11481. Remarks: Spondylus carlosensis Anderson, 1905,  Squires (1999b: p. 24). [Spondylidae].
Spondylus carlosensis
Eocene. South side Simi Valley, Calabasas quad. (1952, PR 1967), Ventura Co. CSUN 1343 [=LACMIP 17186].
Hypotype LACMIP 12695. [Spondylidae].
Spondylus carlosensis
San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13443. [Spondylidae].
Spondylus cliffensis
possibly Ardath Shale], Eocene. South of mouth of Soledad Valley, La Jolla quad., San Diego Co. UCMP 5062. Ho-
lotype UCMP 31025. [Spondylidae].
“Spondylus estrellanus
Estrella Valley, San Luis Obispo Co. Holotype (hypotype) USNM 13312. Remarks: Supplemental descriptions
Lyropecten,Moore (1987: p. C8).
[Pectinidae].
Spondylus perrini
Fm. (Beechers Bay Member), Miocene. East central Santa Rosa Is., California Channel Ids. UCSB SR64-9 [=LACMIP
16434]. Hypotype UCSB M23 [=LACMIP 6322]. [Spondylidae].
Spondylus perrini
Miocene. Sespe Creek area, Topatopa Mtns.,Ventura Co. CSUN 125 [=LACMIP 16363]. Hypotype UCLA 58205
[=LACMIP 10593]. [Spondylidae].
Spondylus perrini  
Reef,” Saddleback Valley, Orange Co. SDSNH 4312. Hypotype SDSNH 83235]. [Spondylidae].
Spondylus perrini
Oligocene/Miocene. Wiley Canyon, Piru quad., Ventura Co. SDSNH 437. Holotype SDSNH 28, holotype (of Spon-
dylus inezana Wiedey, 1928) SDSNH 29. [Spondylidae].
Spondylus scotti
(Saddleback Valley Limestone member), Miocene. North of Aliso Parkway, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4520 [error for SDSNH 5421]. Hypotype SDSNH 83255. [Spondylidae].
Spondylus sp. aff. 
Southwestern Coyote Mtns., Imperial Co. J.T. Smith locality 88JS9. No hypotype nor museum data given. [Spon-
dylidae].
Spondylus victoriae
Sand Member) [=Latrania Fm.], Miocene/Pliocene. East of Whitewater River, San Gorgonio Pass, Riverside Co.
UCMP A3188. Hypotype UCMP 37532. Remarks: Junior synonym of Spondylus princeps Broderip, 1833,
Moore (1987: p. C8). [Spondylidae].
Spondylus
and Fish creeks, Los Angeles Co. CIT 2104 [=LACMIP 7775]. [Hypotype LACMIP 4102]. [Spondylidae].
Stereomactra abcissa (Gabb, 1869). [Schizodesma]. Moore 2003a. USGS 1228-F [electronic format] (family Mac-

ANSP 4548. [Mactridae].
164 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Striostreaappressa (Gabb, 1869b). [Ostrea

1940:135) or Tejon Fm. (of Gabb 1869b: pp. 203, 204), Oligocene/Miocene. Vicinity of Salt Creek, southwest of
Round Valley, Mendocino Co. [MCZ 1062]. Paratype (syntype in text) MCZ 15013 [=MCZ 112951]. [Ostreidae].
Striostrea bourgeoisii bourgeoisii (Rémond, 1863). [Ostrea

(of Clark, 1915, pl. 43) UCMP 11566. [Ostreidae].
Striostrea bourgeoisii perrini (Hall and Ambrose, 1916). [Ostrea

Striostrea subtitan (Loel and Corey, 1932). [Ostrea
Fm., Oligocene/Miocene. Northwest of mouth of Lion Canyon, Ventura quad., Ventura Co. UCMP A326. Holotype
UCMP 31750. [Ostreidae].
Striostrea tayloriana (Gabb, 1866). [Ostrea
Oligocene. San Marcos Pass, near Santa Barbara, Santa Ynez quad., Santa Barbara Co. Holotype UCMP 12005.
Remarks: Nomen dubium,  Squires (2018a: p. 15). [Ostreidae].
Striostrea tayloriana (Gabb, 1866). [Ostrea-
stone, Eocene. Upper Sespe Creek, Ventura Co. CSUN 252 [=LACMIP 16670]. Hypotype LACMIP 11659. Remarks:
Nomen dubium,  Squires (2018a: p. 15). [Ostreidae].
Strophocardia megastropha (Gray, 1825). [Venericardia
Fm. [=Latrania Fm.], Pliocene. Near Travertine Rock, Imperial Co. (not San Diego Co. as in locality description).
USGS M9778. Hypotype USNM 486907. [Carditidae].
Swiftopecten adekunbiana 
Oyster-pecten bed, Coalinga quad., Fresno Co. UCMP D1088. Holotype UCMP 36639. Remarks: Junior synonym
of Lyropecten crassicardo (Conrad, 1857a),Smith (1991: p. 52). [Pectinidae].
Swiftopecten hamlini (Arnold, 1906). [Pecten]. Moore 1984. USGS PP 1228-B:B60, pl. 
Fm., Miocene. Head of Slacks Canyon, Priest Valley quad., Monterey Co. USGS 3851. Holotype USNM 164844.
[Pectinidae].
Swiftopecten parmeleei (Dall, 1898). [Pecten
area, Los Angeles Co. LACMIP. Hypotype LACMIP 14346. [Pectinidae].
Swiftopecten parmeleei (Dall, 1898). [Pecten

Swiftopecten parmeleei parmeleei (Dall, 1898). [Pecten


quad., Fresno Co. USGS 4712]. Holotype (of Swiftopecten wattsi (Arnold, 1906)  determined here as missing.

Swiftopecten parmeleei etchegoini (Anderson, 1905). [Pecten    
Etchegoin Fm., Pliocene. Zapata (Zapato) Chino Creek area, Mt. Diablo Range, Fresno Co. CASG 66740. Holotype
CASG 55 [=CASG 66740.01]. [Pectinidae].
Swiftopecten morani (Arnold 1906)) USNM
164929.
Swiftopecten swifti (Bernardi, 1858). [Pecten

70407]. Remarks: Swiftopecten parmeleei parmeleei (Dall, 1898),  Moore (1984: p. B59). [Pectinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 165
Swiftopecten swifti (Bernardi, 1858). [Pecten   
Remarks:
Swiftopecten parmeleei (Dall, 1898),  Moore (1984: p. B59). [Pectinidae].
Swiftopecten swifti parmeleei (Dall, 1898). [Pecten]. Masuda 1972. Trans. Proc. Paleo. Soc. Japan 84:401–402, pl.
Remarks:
Swiftopecten parmeleei (Dall, 1898),  Moore (1984: p. B59). [Pectinidae].

16862. Hypotypes LACMIP 4999–5001].

Swiftopecten, 2. Santa Margarita Fm., Miocene. From oyster
and pecten-bearing bed, Coalinga quad., Fresno Co. UCMP D1088. Hypotype UCMP 36639. Remarks: Swiftopecten
adekumbiana n. sp.,  Adegoke (1969: p. 104). Junior synonym of Lyropecten crassicardo (Conrad, 1857a),
Smith (1991a: p. 52). [Pectinidae].
Tagelus clarki
Orange Co. Line drawing, source specimen unknown. [Solecurtidae].
Taras cf.  (Conrad, 1848). [Loripes
Fm. (Patinopecten zone), Pliocene. Vicinity of La Aleta, North Dome, Kettleman Hills, Kings Co. USGS 12711. Hy-
potype USNM 495843. Remarks: In the genus Felaniella,Moore (1988: p. D29). [Ungulinidae].
Taras parilis (Conrad, 1848). [Loripes
  Diplodonta parilis (Conrad 1848)). Etchegoin Fm., Pliocene.
Coalinga District, Fresno Co. USGS 4806. Hypotype USNM 165484]. Remarks: In the genus Felaniella,Moore
(1988: p. D29). [Ungulinidae].
aff. Tehamatea n. sp. Powell et al. 2019b. PaleoBios-
vation, Berkeley Hills, Oakland, Alameda Co. UCMP IP 13001. Hypotype UCMP 410562. [Lucinidae].
Tellina aragonia  
Kings Co. UCMP D1138. Hypotype UCMP 36768. Remarks: Provisionally in the subgenus Peronidia Moore
(2003b: p. 21). [Tellinidae].
Tellina (PeronidiaagaroniaTellina aff.  (Peronidia
agaronia
CASG 101099, 101100]. [Tellinidae].
Tellina cf.        
(Siphonalia Zone), Pliocene. Vicinity of La Aleta, North Dome, Kettleman Hills, Kings Co. USGS 12430. Hypotype
USNM 495822. Remarks: In the subgenus Peronidia,  Coan et al. (2000: p. 404). [Tellinidae].
Tellina (Peronidia) bodegensis   
San Diego Fm., Pliocene. San Diego Co. LACMIP 305A [=LACMIP 16862]. Hypotype LACMIP 4779. [Tellinidae].

[=LACMIP 9308].
Tellina (Cadella californica Gabb, 1864. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):11–12,

Costa Co. Type material missing,  Stewart (1930: p. 7). [Tellinidae].
Tellina (Moerella) carpenteri
San Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hy-
potypes LACMIP [4775, 4777]. Remarks: In the subgenus Angulus,  Coan et al. (2000: p. 400). [Tellinidae].
....., pl. 
166 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Tellina (Angulus) carpenteri Dall, 1900b. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):2–3,


Tellina castacana
Guillermo quad. (1943), Ventura Co. UCR 4743. Hypotype UCR 4743/502. Remarks: Provisionally in the subge-
nus Angulus Moore (2003b: p. 6). [Tellinidae].
Tellina castacana
Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hy-
potype SDSNH 20772. Remarks: Provisionally in the subgenus Angulus Moore (2003b: p. 6). [Tellinidae].
Tellina (Angulus castacana Anderson and G D. Hanna, 1925. Moore 2003b. USGS PP 1228-G [electronic format]

[Tellinidae].
Tellina congesta
Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M1680. Hypotype USNM 254351.
Remarks: Provisionally in the subgenus Cadella Moore (2003b: p. 12). [Tellinidae].
Tellina congesta

Obispo Co. USGS M1680. Hypotype USNM. 254351. Remarks: Provisionally in the subgenus Cadella Moore
(2003b: p. 12). [Tellinidae].
Tellina (Cadella congesta Conrad, 1855b. Moore, 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):12,

Tellina (Megangulus crooki Nelson, 1925. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

UCMP 3776. Holotype UCMP 30523. [Tellinidae].
Tellina (Angulus diabloensis Clark, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):8–9,

1478. Holotype UCMP 11531. [Tellinidae].
Tellina domenginensis
North Fork of Matilija Creek, Matilija quad. 1952, PR 1967), Ventura Co. CSUN 1445 [=LACMIP 24259). Hypotype
LACMIP 12473. Remarks: In the genus Tellina and provisionally in the subgenus Angulus Moore (2003b:
p. 7). [Tellinidae].
Fork of Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP
16963]. Hypotypes LACMIP 12474–12476.
Tellina (Angulus domenginensis Vokes, 1939. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

UCMP 15694. [Tellinidae].
Tellina (Oudardia) emacerata
Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotype USNM 646560. Remarks:
Provisionally in the subgenus Oudardia Moore (2003b: p. 19). [Tellinidae].
Tellina (Phyllodina englishi Clark, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):23–24,

Tellina (Eurytellina hannibali Clark, 1915. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

11532. [Tellinidae].
Tellina (Eurytellina herndonensis Dickerson, 1914a. Moore 2003b. USGS PP 1228-G [electronic format] (family
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 167

[Tellinidae].
Tellina (Tellinella) idae   
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1953), San Diego Co. LACMIP 305. Hypotype
LACMIP 4771. [Tellinidae].
Tellina (Tellinella) idae
Angeles Co. LACMIP 17920. Hypotype LACMIP 14367. [Tellinidae].
Tellina joaquinensis
Eocene. North Fork of Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1445. Hypotypes LACMIP
12471, 12472. Remarks: In the genus Tellina and provisionally in the subgenus Angulus Moore (2003b:
p. 4). [Tellinidae].
Tellina (Angulusjoaquinensis Arnold, 1909 [1910]. Moore 2003b. USGS PP 1228-G [electronic format] (family

Coalinga quad., Fresno Co. USGS 4801. Holotype USNM 165619. [Tellinidae].
Tellina (Angulus jollaensis Dickerson, 1916. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

2226. Holotype UCMP 33252. [Tellinidae].
Tellina (Megangulus kewi Dickerson, 1914a. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-
 
[Tellinidae].
Tellina lebecki
Matilija Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23688. Hypotype LACMIP 11632. Remarks:
Provisionally in the subgenus Phyllodina Moore (2003b: p. 22). [Tellinidae].
Tellina (Phyllodina lebecki Anderson and G D. Hanna, 1925. Moore 2003b. USGS PP 1228-G [electronic format]
-
type CASG 244 [=CASG 70903]. [Tellinidae].
Tellina (Angulus longa Gabb, 1864. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):5–6, pl.

Costa Co. Paratype MCZ 15037, missing  Moore (2003b: p. 5). [Tellinidae].
Tellina lorenzoensis
Lorenzo River, Santa Cruz Co. SCCM 37. Hypotype SCCM 21003. Remarks: Provisionally in the subgenus Eurytel-
lina Moore (2003b: p. 14). [Tellinidae].
Tellina cf.      
(diatomaceous strata), Miocene. Gato Ridge, Santa Barbara Co. USGS 15032. Hypotype USNM 560037. Remarks:
Tellina lutea alternidentata Broderip and G.B. Sowerby I, 1829,  Coan (1971: p. 8); Tellina (Meganulus) lutea
Wood, 1828,  Coan et al. (2000: p. 403). [Tellinidae].
Tellina cf. 
Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650967. Remarks: Tellina lutea alternidentata
Broderip and G.B. Sowerby I, 1829, Coan (1971: p. 8); Tellina (Meganulus) lutea Wood, 1828,  Coan et al.
(2000: p. 403). [Tellinidae].
Tellina (Eurytellina martinezensis Weaver, 1905. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Holotype UCMP 11876. [Tellinidae].
Tellina nevadensis
Rio Bravo Ranch quad. (1954), Kern Co. USGS M1599. Hypotype USNM 649168. Remarks: Junior synonym of
Tellina (Olcesia) piercei (Arnold, 1909 [1910]),  Addicott (1973b: p. 31); Moore (2003b: p. 18). [Tellinidae].
168 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Tellina (Olcesia ocoyana Conrad, 1855b. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):19,
 Keen and Bentson
(1944: p. 115). Holotype missing,  Keen and Bentson (1944: p. 115);  Moore (2003b: p. 19). [Tellinidae].
Tellina oldroydi
(Pecten Zone), Pliocene. North DomeKettleman Hills, Kings Co. USGS locality 12331. Hypotype USNM 495751.
Remarks: Hypotype now a paratype of Tellina (Angulus woodringi Adegoke, 1969,  Moore (2003b: p. 9).
[Tellinidae].
 Fm. (Siphonalia Zone), Pliocene. Arroyo Pino, north of Bullwheel Ridge, North Dome,
Kettleman Hills, Kings Co. USGS 12815. Hypotype USNM 495825. Remarks: Hypotype now a paratype of Tellina
(Angulus woodringi Adegoke 1969,  Moore (2003b: p. 9).
Tellina oldroydi
Ranch quad., Fresno Co. UCMP D1101. Hypotype UCMP 36767. Remarks: In the genus Tellina and provisionally
in the subgenus Eurytellina Moore (2003b: p. 14). [Tellinidae].
Tellina (Eurytellina oldroydi Wiedey, 1928. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Calabasas quad., Los Angeles Co. SU 425. Holotype SDSNH 36. [Tellinidae].
Tellina (Angulus packardi Dickerson, 1914a. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):4,

Tellinaperrini Dickerson, 1914a. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

[Tellinidae].
Tellina (Olcesia) piercei (Arnold, 1909 [1910]). [Macoma      
Temblor Fm. (Wygal Sandstone Member), Miocene. Temblor Creek, Carneros Rocks quad., Kern Co. USGS M3978.
Hypotype USNM 646558. [Tellinidae].


646574, 646575 (hypotype 646575 determined here as lost).
Tellina piercei (Arnold, 1909 [1910]). [Macoma
Fm. (Saltos Shale Member), Miocene. Shell Creek, La Panza Range, San Luis Obispo Co. USGS M2869. Hypotype
USNM 254355. Remarks: In the subgenus Olcesia,  Moore (2003b: p. 18). [Tellinidae].
Tellina piercei (Arnold, 1909 [1910]). [Macoma-

Range, San Luis Obispo Co. USGS M2869. Hypotype USNM 254355. Remarks: In the subgenus Olcesia,  Moore
(2003b: p. 18). [Tellinidae].
Tellina (Olcesia) piercei (Arnold, 1909 [1910]). Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Holotype (of Tellina nevadensis Anderson and Martin, 1914) CASG 124. [Tellinidae].
Tellina praecuta Clark, 1918. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Co. UCMP 14. Holotype UCMP 11166. [Tellinidae].
Tellina aff.  Linneaus, 1758. Lutz 1951. Vicinity of
Crow Creek near Castro Valley, Alameda Co. UCMP A4932. Plastotype UCMP 34137. [Tellinidae].
Tellinaremondii Gabb, 1864. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):27,

31451 [as UCMP 3145 in text]. [Tellinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 169
Tellina (Macaliopsis () rosa (M.A. Hanna, 1927). [Macoma]. Squires 1987. NHMLAC CS 388:68,

LACMIP 7533. [Tellinidae].
Tellina (Cadella) salmonea (Carpenter, 1864b). [Maera]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):286–287, pl.

LACMIP 305. Hypotypes LACMIP 4772–4774. Remarks: Junior synonym of Tellina (Cadella) nuculoides (Reeve,
1854),  Coan et al. (2000: p. 403). [Tellinidae].
Tellina (Angulus scrippsensis M.A. Hanna, 1927. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-

Holotype UCMP 31022. [Tellinidae].
Tellina (Angulus simiensis Nelson, 1925. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):6,

UCMP 3778. Holotype UCMP 30522, paratype UCMP 30521. [Tellinidae].
Tellina cf. 
Eocene. Vicinity of Cañada Omentaro, Gaviota quad., Santa Barbara Co. UCMP B7011. Hypotype SU 9317 [=CASG
70243]. Remarks: Tellinasoledadensis M.A. Hanna 1927,  Moore (2003b: p. 27). [Tellinidae].
Tellina soledadensis-
ley quad. (1943), Ventura Co. UCR 4678. Hypotype UCR 4678/16. Remarks: Tellinasoledadensis
M.A.Hanna, 1927,  Moore (2003b: p. 27). [Tellinidae].
Tellina soledadensis M.A. Hanna, 1927. Moore 2003b. USGS PP 1228-G [electronic format] (family

3975. Syntype 31368. [Tellinidae].

as in text)].
Tellina soledadensis M.A. Hanna, 1927. Ekdale and Ekdale 2018. Paleo.3 493: . 5B. Scripps Fm., Eocene. Torrey
Pines Science Center, San Diego Co. [SDSNH 4552]. Hypotype SDSNH 80165. [Tellinidae].
Tellinasutterensis Dickerson, 1913. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):30, pl.
   
Holotype UCMP 11767. [Tellinidae].
Tellina tehachapi

Co. UCMP B6941. Hypotype SU 9318 [=CASG 70255]. Remarks: Tellinatehachapi Anderson and G
D. Hanna, 1925,  Moore (2003b: p. 28). [Tellinidae].
Tellinatehachapi Anderson and G D. Hanna, 1925. Moore 2003b. USGS PP 1228-G [electronic format]

CASG 878. [Tellinidae].
Tellina (Tellinella) tenuilineata  
Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotype USNM 646569. [Tellinidae].

Hypotype USNM 646570.
Tellina (Tellinella) tenuilineata Clark, 1918. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellini-
         
11130. [Tellinidae].
Tellina cf.     
Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotype USNM 646585. [Tellinidae].
170 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Tellina
ft., San Guillermo quad. (1943), Ventura Co. UCR 4743. Hypotype UCR 4743/6. [Tellinidae].
Tellina
14. Mission Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH
2962. Hypotype SDSNH. 20771. [Tellinidae].
Tellina

UCMP unnumbered hypotype lost,  C. Throckmorton (personal communication, 2002). [Tellinidae].
Tellina undulifera

UCMP 3783. Hypotype UCMP 30512. Remarks: In the subgenus Saulella,  Moore (2003b: p. 9]. [Tellinidae].
Telliina undulifera
unknown. Remarks: In the subgenus Saulella,  Moore (2003b: p. 9). [Tellinidae].
Tellina” n.gen. undulifera Gabb, 1869b. Zinsmeister 1983a. SEPM Guidebook 
Paleocene. Simi Hills, Ventura Co. UCR6848. Hypotype UCR 6848/100 (not UCR 6848/1 as in pl. caption). Re-
marks: Tellina (Saulella) undulifera Gabb, 1869b, Moore (2003b: p. 9). [Tellinidae].
Tellina (Saulella) undulifera Gabb, 1869b. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):9–10,

59255 [=LACMIP 10984]. [Tellinidae].

Tellina
(Wygal Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotype USNM 646561.
[Tellinidae].
Tellina (Angulus) vorbei M.A. Hanna, 1927. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):1–2,

UCMP 5074. Holotype UCMP 30984. [Tellinidae].
Tellina sp. cf.  of Ekdale and Ekdale, 2018. Paleo.3, 
San Diego Co. SDSNH 5598. Hypotype SDSNH 103702. [Tellinidae].

Tellinawilsoni Anderson and Martin, 1914. Moore 2003b. USGS PP 1228-G [electronic format] (fam-

Holotype CASG 127. [Tellinidae].
Tellina woodringi
quad., Kings Co. UCMP D1193. Holotype UCMP 36771. Remarks: In the genus Tellina and provisionally in the
subgenus Angelus Moore (2003b: p. 9). [Tellinidae].

Tellina (Angelus woodringi Adegoke, 1969. Moore 2003b. USGS PP 1228-G [electronic format] (family Tellinidae):9,

Tellina
Cielo Rd. west of San Marcos Pass at Sacate-"Coldwater" contact, San Rafael Mtn. quad., Santa Barbara Co. UCMP
B6941. Hypotype SU 9176 (not SU 9320 as published) [=CASG 70242] [Tellinidae].

B6994. Hypotype SU 9319 [=CASG 70254]. [Tellinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 171
Tellina
Clara Co. USGS M1715. Hypotype USNM 650983. [Tellinidae].
Tellina sp. Throckmorton 1988a. USGS OFR 88-59:52
Tesla Fm., Eocene. Midway quad., Alameda Co. UCMP D8154. UCMP
Tellina sp. Ekdale and Ekdale, 2018. Paleo. 3 
Co. SDSNH 4147. Hypotype SDSNH 65080. [Tellinidae].
Tellina
Los Angeles Co. LACMIP 17850. Hypotype LACMIP 13450. [Tellinidae].
-

Teredo      
quad., Los Angeles Co. CSUN 524 [=LACMIP 16379]. Hypotype LACMIP 6593. [Teredinidae].
Teredo
(1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype LACMIP 7542. [Teredinidae].

Los Angeles Co. LACMIP 17850. Hypotype LACMIP 13456. [Teredinidae].
          
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 987 [=LACMIP 16175]. Hypotype LACMIP 7735.
[Teredinidae].
Thecodonta (Pristes) oblongus (Carpenter, 1864b). [Pristes
Diego Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305. Hypotype (of Hertlein and Grant 1972,
Remarks: In the genus Pristes,  Coan et al. (2000: p. 327). [Lasaeidae].
Thracia aff.               
Inyokern SE quad. (1972), El Paso Mtns., Kern Co. USGS M9067. Hypotype USNM 424898. Remarks: In the sub-
genus Cetothrax,  Coan et al. (2000: p. 531). [Thraciidae].
Thracia dilleri
of Agua Caliente, Lompoc quad., Santa Barbara Co. UCMP B6995. Hypotype SU/CASG 9219 (not CASG 9325 as
published) [=CASG 70247]. [Thraciidae].
     
B7007D. Hypotype SU 9217 (not CAS 9324 as published) [=CASG 70248].
Thracia (Thracia) dilleri
(1943), Ventura Co. UCR 4731. Hypotype UCR 4731/81. [Thraciidae].

referred to as Pico Fm. (Pliocene) but most likely is Castaic Fm. (Miocene). Silt beds south of Humphrey railroad
      
paratypes LACMIP 4840–4877. Remarks: Junior synonym of Thracia (Homoeodesma) trapezoides Conrad, 1849a,
 Coan et al. (2000: p. 532. [Thraciidae].


Thracia trapezoides
quad., Sonoma Co. UCMP A9410. Hypotype UCMP 15779. Remarks: In the subgenus Homoeodesma,  Coan et
al. (2000: p. 532). [Thraciidae].
172 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Thracia trapezoides-
ponio Member), Pliocene, Purisima Creek area, Half Moon Bay quad., San Mateo Co. Locality 645-1M. Hypotype
determined here as missing. Remarks: In the subgenus Homoeodesma,  Coan et al. (2000: p. 532). [Thraciidae].
Thracia trapezoides
area, Ferndale quad., Humboldt Co. UCMP B7954. Hypotype UCMP 37765. Remarks: In the genus Homoeodesma,
 Coan et al. (2000: p. 532). [Thraciidae].
Thyasira bisecta (Conrad, 1849a). [Venus
Fm., [=Purisima Fm.], Pliocene. Near the town of Princeton, San Mateo Co. UCMP B4791. Hypotype UCMP 37623.
Remarks: In the genus Conchocele,  Coan et al., (2000: p. 277). (Thyasiridae).
Thyasira cf. bisecta (Conrad, 1849a). [Venus
Tumey Fm., Eocene/Oligocene. Monocline Ridge, Fresno Co. UCMP D6536]. Hypotypes UCMP 37157, 37158.
Remarks: In the genus Conchocele,  Coan et al., (2000: p. 277). [Thyasiridae].
Thyasira cf. bisecta (Conrad, 1849a). [Venus
Fm., Oligocene. Monocline Ridge, Monocline Ridge quad. (1955), Fresno Co. [UCMP D6536]. Hypotypes UCMP
37157, 37158. Remarks: In the genus Conchocele,  Coan et al. (2000: p. 277). [Thyasiridae].
Thyasira (Conchocele) folgeri       
Emigdio Fm., Eocene. Devil's Kitchen area Kern Co. UCMP 3195. Holotype UCMP 11434. Remarks: Possibly in
the genus Maorithyas, 
p. 348). [Thyasiridae].
Thyasira gouldii (Philippi, 1845). [Lucina
Diego Fm., Pliocene. Southwestern San Diego Co., San Ysidro quad. (1943), San Diego Co. ACMIP 305. Hypotype
LACMIP 4652. Remarks: Junior synonym of  (Montagu 1803), Coan et al. (2000: p. 275).
[Thyasiridae].
....., pl. 43. UCLA 294 [=LACMIP 20294]. Hypotype UCLA 48620
[=LACMIP 9303].
Thyasira (Thyasira) gouldi (Philippi, 1845). [Lucina
Fm., Pliocene. San Ysidro quad. (1943), San Diego Co. LACMIP 305]. Hypotype (of Hertlein and Grant 1972, pl. 43,
Remarks: Junior synonym of ,  Coan et al. (2000: p. 275). [Thyasiridae].
Thyasira


quad., Fresno Co. UCMP D6534. Hypotype UCMP 37156. [Thyasiridae].
Tivela diabloensis-
ton, Alameda Co. SU 3232. Hypotype SU 8452 [=CASG 69983]. Remarks: In the subgenus Pachydesma,  Clark
(1915: p. 462). [Veneridae].
Tivela (Pachydesma) gabbi        
quad. (1915), Contra Costa Co. UCMP A2569. Hypotype UCMP 30673. [Veneridae].
Tivela stanfordia  
Costa Valley quad. (1943), Alameda Co. SU 3244. Holotype SU 8454 [=CASG 69986, paratypes SU 8453, 8455
[=CASG 69984, 69985]. [Veneridae].
Tivela (Pachydesma) stultorum (Mawe, 1823). [Donax]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):266–267, pl.

48621 [=LACMIP 9304]. [Veneridae].

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 173
Tivela (Pachydesma) weaveri (Packard in Dickerson, 1914a). [Spisula]. Squires and Deméré 1991. SEPM Guidebook

Trachycardium (Trachycardium) arcumbona (Wiedey, 1928). [Cardium]. Moore 2003a. USGS PP 1228-F [electronic

quad., San Luis Obispo Co. SU 432. Holotype SDSNH 31. [Cardiidae].
Trachycardium cf. Trachycardium quadragenarium (Conrad, 1837). [Cardium]. Woodring et al. 1940 [1941]. USGS
Pecten zone), Pliocene. Surprise Canyon, North Dome, Kettleman Hills,
Kings Co. USGS 12322. Hypotype USNM 495777. Remarks: In the genus Dallocardia,  Coan and Valentich-
Scott (2012: p. 413). [Cardiidae].
Trachycardium (Dallocardia) quadragenarium (Conrad, 1837). [Cardium
31. Pico Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14356. Remarks: In the
genus Dallocardia,  Coan and Valentich-Scott (2012: p. 413). [Cardiidae].
Trachycardium (Dallocardia) quadragenarium fernandoensis (Arnold, 1907). [Cardium]. Moore 2003a. USGS 1228-F

Fm., Pliocene. Elsmere Canyon, Los Angeles Co. Holotype USNM 164947. Remarks: Junior synonym of Trachy-
cardium quadragenarium (Conrad, 1837),  Coan et al. (2000: p. 362); in the genus Dallocardia,  Coan and
Valentich-Scott (2012: p. 413). [Cardiidae].
Trachycardium sagaseri
quad., Kings Co. UCMP D1134. Holotype UCMP 36686. Remarks: Possibly in the genus Dallocardia. [Cardiidae].

Trachycardium (Dallocardia) sagaseri Adegoke, 1969. Moore 2003a. USGS 1228-F [electronic format] (family

quad., Kings Co. UCMP D1134. Holotype UCMP 36686. Remarks: Possibly in the genus Dallocardia. [Cardiidae].
Trachycardium vaquerosensis (Arnold, 1908). [Cardium

Trachycardium (Trachycardium) vaquerosensis (Arnold, 1908). [Cardium]. Moore 2003a. USGS PP 1228-F [elec-
  
Creek and Alpine Creek, Santa Cruz quad., San Mateo Co. Holotype USNM 165467 (not USNM 105467 as in pl.
caption). [Cardiidae].
Trachycardium (Trachycardium) woodringi (Loel and Corey, 1932). [Cardium]. Moore 2003a. USGS PP 1228-F [elec-

Eastern foothills of La Panza Mtns., La Panza quad., San Luis Obispo Co. UCMP A495. Holotype UCMP 31774 (not
UCMP 31714 as in pl. caption). [Cardiidae].
Trachycardium          
Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312.
Hypotype SDSNH 93837. [Cardiidae].
Transenella joaquinensis
Round Mtn. Silt, Miocene. Caliente quad., Kern Co. SU 2121. Hypotypes SU 7528, 7528a [=CASG 72013]. [Veneridae].
Transenella tantilla (Gould, 1853). [Venus  
Diego Fm., Pliocene. San Ysidro quad. (1953), San Diego Co. LACMIP 305C [=LACMIP 16817]. Hypotype LACMIP
4768. Remarks: In the genus Nutricola,  Coan et al. (2000: p. 384). [Cardiidae].

.....,      Co. UCLA 1386 [=LACMIP 21386]. Hypotype UCLA 48624
[=LACMIP 9307].
174 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Transenella tantila (Gould, 1853). [Venus
Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D3386. Hypotypes USNM 251801, 251802. Remarks:
In the genus Nutricola,  Coan et al. (2000: p. 384). [Cardiidae].
Transenella
Kings Co. UCMP D9939. Hypotype UCMP 37442. Remarks: Determined here as in the genus Nutricola. [Cardiidae].
Tresus nuttallii (Conrad, 1837). [Lutraria (Cryptodon)]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):318, pl. 54,

LACMIP 305. Hypotype LAMCIP 4822. [Mactridae].
-
MIP 9318]. Remarks: Specimen actually from Pleistocene Bird Rock terrace overlying San Diego Fm.,  G.L.
Kennedy (personal communication, 2001).
Tresus nuttallii (Conrad, 1837). [Lutraria (Cryptodon)]. 
Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14370. [Mactridae].
Tresus pajaroanus (Conrad, 1857d). [Venus       
Canoas Creek, Reef Ridge quad., Kings Co. UCMP D1154. Hypotype UCMP 36736. [Mactridae].
Tresus pajaroanus (Conrad, 1857d). [Venus     
Southwest of Escalona Gulch, Santa Cruz Co. CASG66088. Hypotype CASG 66088.01]. [Mactridae].
Tresus pajaroanus (Conrad, 1857d) [Venus

Tresus saxadomoides (Clark, 1918). [Spisula
-
type UCMP 11268. [Mactridae].
Trigoniocardia aff.  (d'Orbigny, 1842). [Cardium
24, 25 (clay peel). Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area,
Palos Verdes Hills, Los Angeles Co. USGS 13127. Hypotypes USNM 497092, 497093. Remarks: Determined here
as Americardia biangulata (Broderip and G.B. Sowerby I, 1829).[Cardiidae].
cf. Trigonicardia
Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to LACMIP. [Cardiidae].

lower). Monterey Fm., Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated
to LACMIP. [Pectinidae].

Fm., Pliocene [=Wilson Grove Fm.), Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotype CASG determined
here as missing. Possibly a cardiid.
Venericardia (Leuroactis) alisoensis       
Fm., Eocene. West side of Aliso Canyon, Los Angeles Co. UCMP 7019]. Holotype UCMP 30176. Remarks: Junior
synonym of Venericardia () aragonia joaquinensis (Vokes, 1939),  Squires (1984: p. 46; 1987: p. 63);
 Moore (1992: p. E23). [Carditidae].
Venericardia () aragonia diabloensis

32799. [Carditidae].

Venericardia () aragonia joaquinensis (Vokes, 1939). [Megacardita]. Squires 1988a. SEPM Guidebook

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 175
LACMIP 7525. [Carditidae].
Venericardia () aragonia joaquinensis (Vokes, 1939). [Megacardita]. Saul 1983b. SEPM Guidebook 35:76, pl.

37433. [Carditidae].
Venericardia () aragonia joaquinensis (Vokes, 1939). [Megacardita
10p. Llajas Fm., Eocene. Thousand Oaks quad. (1950, PR 1967), Ventura Co. UCMP 7193 [=CSUN 702; =UCLA
7030]. Hypotype UCMP 37433. [Carditidae].
Venericardia () aragonia joaquinensis (Vokes, 1939). [Megapitaria
108. Juncal Fm., EoceneWhitaker Peak quad. (1958), Los Angeles Co. CSUN 815 [=LACMIP 16401]. Hypotype
LACMIP 7525. [Carditidae].
Venericardia () aragonia joaquinensis (Vokes, 1939). [Megacardita]. Moore 1992. USGS PP 1228-E:E23–

Fresno Co. UCMP A1022]. Hypotype UCMP 15615 [=paratype of Venericardia () vallecitosensis (Vokes,
1939),  Keen and Bentson (1944: p. 65]. [Carditidae].
Veneri-
cardia (Leuroactis) alisoensis Verástegui, 1953) UCMP 30176.

UCMP 15617.
of Verástegui 1953: p. 50), Paleocene. Camulos quad., Ventura Co. [CASG 66532.]
Holotype SU 8003 [=CASG 66532.01] (of Venericardia (Leuroactis) schencki Verástegui, 1953).

Holotype UCMP 15626, paratype (of Venericardia () smileyi (Vokes, 1939)) UCMP 15627.

Venericardia () vokesi Verástegui, 1953) SU 8016 [=CASG 72185].
Venericardia () argentea
Paleocene. Near junction of Panoche and Silver Creeks, Panoche quarangle, Fresno Co. SU 2073 [=CASG 66535].
Holotype SU 7995 [=CASG 66535.02]. Remarks: Junior synonym of Glyptoactis (Claibornicardia) keenae (Verá-
stegui, 1953),  Moore (1992: p. E27). [Carditidae].
Venericardia ()-
gine Fm., Eocene. Vicinity of Oil City, Fresno Co. Hypotype CASG 1828. Remarks: Venericardia () hornii
 Stewart, 1930,  Squires (1984: p. 46). [Carditidae].
-
lotype UCMP 31450. Remarks: Venericardia () Stewart, 1930,  Squires (1984: p. 46).
    
[=CASG 72167]. Remarks: Venericardia () Stewart, 1930,  Squires (1984: p. 46).
Venericardia ()




La Jolla quad., San Diego Co. UCMP 3990. Holotype (of Venericardia () hertleini Verástegui, 1953) UCMP
30415.
Venericardia ()
176 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Fm., Eocene. Grapevine Canyon, Kern Co. CASG 245. Holotype CASG 685 (not CASG 686 as in pl. caption) [=CASG

Venericardia ()
Eocene. Vicinity of Vickers Hot Springs, Ventura Co. CASG 829. Holotype (of Venericardia () durhami
Verástegui, 1953) SU 8005 [=CASG 66528.01]. [Carditidae].
Venericardia () Verástegui, 1953. Moore 1992. USGS
16–18. Santa Susana Fm. (of Verástegui 1953), Paleocene. McCray Oil Wells, Oil Canyon, Camulos quad., Ventura
Co. SU CA-33 [CASG 66527]. Holotype SU 8004 [=CASG 66527.01)] [Carditidae].
Venericardia ()-
sana Fm., Paleocene/Eocene. Santa Susana quad., Ventura Co. UCMP 7009. Hypotype UCMP 37431. [Carditidae].
Venericardia () carlosensis (Vokes, 1939). [Megacardita]. Verástegui 1953. Pal. Amer. 3(25):36–37, pl. 13,
Remarks: Junior synonym
of Venericardia (Venericor) vallecitosensis (Vokes, 1939),  Moore (1992: p. E25). [Carditidae].

Venericardia () clarki clarki
2. Coldwater Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23724. Hypotype LACMIP 11660.
[Carditidae].

Venericardia () clarki popenoei
      
Holotype (photocopy of Verástegui 1953,  Moore (1992: pl. 9 caption), UCLA (not UCMP as in pl. caption)
15689 [=LACMIP 10845]. [Carditidae].
Venericardia () diabloensis

61967.01]. Remarks: Venericardia () agaronia diabloensis,  Moore (1992: p. E20). [Carditidae].
Venericardia (Glyptoactis) domenginica-
gine Fm., Eocene. Domengine Ranch, Coalinga quad., Fresno Co. UCMP A1219. Holotype UCMP 15611. Remarks:
In the genus/subgenus Glyptoactis (Claibornicardia),  Moore (1992: p. E27). [Carditidae].
Venericardia () durhami
Vicinity of Vickers Hot Springs, Ventura Co. CASG 829 [=CASG 66528]. Holotype CASG 8005 [=CASG 66528.01].
Remarks: Junior synonym of Venericardia () Turner, 1938,  Moore (1992: p. E20).
[Carditidae].
Venericardia () gabbi-
cene. Grapevine Canyon, Kern Co. Holotype CASG 685 (not CASG 686 as in pl. caption). Remarks: Venericardia
() Verástegui, 1953,  Moore (1992: p. E22). [Carditidae].
Venericardia () hertleini Verástegui, 1953 (n
Fm.” [=possibly Ardath Shale], Eocene. Near junction of San Clemente and Rose Canyons, San Diego Co. UCMP
3990. Holotype UCMP 30415. Remarks: Junior synonym of Venericardia () Stewart, 1930,
 Moore (1992: p. E21). [Carditidae].
Venericardia () hornii (Gabb, 1864). [Cardita
Tejon Fm., Eocene. Tejon, Kern Co. Lectotype (of Stewart 1930: p. 167) ANSP. 4558.


GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 177
Venericardia () hornii (Gabb, 1864). [Cardita
Canyon Fm.” [=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH 1667. Hypotype SDSNH 4145].
[Carditidae].
Venericardia () hornii (Gabb, 1864). [Cardita-
cene. Vicinity of Alamo Creek, Dry Canyon, and the Cuyama River Valley, Reyes Peak quad. (1943), Ventura Co.
UCR 4745. Hypotype UCR 4745/1. [Carditidae].
Venericardia hornii (Gabb, 1864). [Cardita
Live Oak Canyon, Kern Co. Lectotype (of Stewart 1930: p. 167) ANSP 4558. [Carditidae].
Venericardia hornii (Gabb, 1864) var. [Cardita

Hypotype CASG 5982. [Carditidae].
Venericardia () hornii (Gabb) cf. subsp. 
Fm., Eocene. Near Thorn Meadows and Thorn Pt., Lion Canyon quad. (1943), Ventura Co. UCR 4684. Hypotype UCR
4684/6. Remarks: Venericardia () Stewart, 1930,  Moore (1992: p. E21). [Carditidae].
Venericardia ()
Eocene. Bus-Trough Canyon area, Calabasas quad., Ventura Co. UCLA 7071 [=LACMIP 27071]. Hypotype UCLA
59383 [=LACMIP 10438]. Remarks: Venericardia () Stewart, 1930,  Moore (1992: p.
E21). [Carditidae].

59378 [=LACMIP 10439].
Venericardia ()    
Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypo-
types LACMIP 6573, 6574. Remarks: Venericardia () Stewart, 1930,  Moore (1992:
p. E21). [Carditidae].
Venericardia () Stewart, 1930. Squires 1997b. Simi Valley Hist. Soc. vol., p-
 Fm., Eocene. Ventura Co. CSUN374 [=LACMIP 7242). Hypotype LACMIP
6573]. Remarks: Venericardia () Stewart, 1930,  Moore (1992: p. E21). [Carditidae].
Venericardia () hornii lutmani 
Fm., Eocene. Hot Springs Creek, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4662. Hypotype UCR 4662/40.
Remarks: Venericardia () Turner, 1938,  Moore (1992: p; E20). [Carditidae].
Venericardia () hornii lutmani
Eocene. Hot Springs Canyon, Topatopa Mtns. quad., Ventura Co. UCLA 4244 [=LACMIP 24244]. Hypotype UCLA
59381 [=LACMIP 10437]. Remarks: Venericardia () Turner, 1938,  Moore (1992: p.
E21). [Carditidae].
Venericardia () hornii lutmani
Cobblestone Mtn. quad. (1958), Los Angeles Co. CSUN 824 [=LACMIP 16402]. Hypotype LACMIP 7524]. Remarks:
Venericardia () Turner, 1938,  Moore (1992: p. E20). [Carditidae].
Venericardia () hornii susanaensis       
Santa Susana Fm., Paleocene. North of Las Llajas Canyon, Santa Susana quad., Ventura Co. UCMP 7000. Hypotype
UCMP 37431. Remarks: Venericardia () Verástegui, 1953,  Squires (1999b: p. 25).
[Carditidae].
Venericardia ionensis
Remarks: Venericardia (Paci-
) agaronia joaquinensis (Vokes, 1939),  Moore (1992: p. E23). [Carditidae].
Venericardia (Leuroactis) joaquinensis (Vokes, 1939). [Megacardita]. Verástegui 1953. Pal. Amer. 3(25):60–61, pl.

178 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Holotype UCMP 15616. Remarks: Venericardia () agaronia joaquinensis (Vokes, 1939),  Moore (1992:
p. E23). [Carditidae].

72176].

Venericardia (Glyptoactis) keenae      
Paleocene. Near junction of Panoche and Silver creeks, Panoche quad., Fresno Co. SU 2072 (not SU 2073 as pub-
lished) [=CASG 66535]. Holotype CASG 7993 [=CASG. 66535.01]. Remarks: In the genus/subgenus Glyptoactis
(Claibornicardia),  Moore (1992: p. E27). [Carditidae].
Venericardia (Glyptoactis) keenae
Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717].
Hypotype SU 7993 (not SU 7992 as in pl. caption) [=CASG 66535]. Remarks: In the genus/subgenus Glyptoactis
(Claibornicardia), Moore (1992: p. E27). [Carditidae].
Venericardia () lisa Verástegui, 1953. Weaver and Kleinpell 1963. UCPG 43:200, 
Gaviota Fm., Eocene. Near Cañada San Onofre, Gaviota quad., Santa Barbara Co. UCMP. B7026. Hypotype SU 9281
[=CASG 70258]. Remarks: Junior synonym of Venericardia () clarki popenoei Verástegui, 1953 (in part),
 Moore (1992: p. E24). [Carditidae].
        
UCMP B6955. Hypotype SU 9280 [=CASG 70272]. Remarks: Venericardia () hornii (Gabb, 1864), 
Moore (1992: p. E23). [Carditidae].
Venericardia (Glyptoactis) marksi-
cene. Live Oak Canyon, Kern Co. SU 183 [=CASG 66536]. Holotype SU 8021 [= CASG 66536.01]. Remarks: In the
genus/subgenus Glyptoactis (Claibornicardia),  Moore (1992: p. E27). [Carditidae].
Venericardia (Glyptoactis) marksi-
ter” Sandstone, Eocene. Vicinity of Brush Peak on Camino Cielo, San Rafael Mtn. quad., Santa Barbara Co. UCMP
B6946. Hypotype SU 9279 [=CASG 70265]. Remarks: In the genus/subgenus Glyptoactis (Claibornicardia), 
Moore (1992: p. E27). [Carditidae].
Venericardia (Glyptoactis) mcmastersi
Fm., Eocene. San Clemente Canyon, San Diego Co. [CASG 66534]. Holotype SU 8011 [=CASG 66534.01]. Remarks:
Junior synonym of Glyptoactis (Claibornicardia) sandiegoensis (M.A. Hanna, 1927),  Moore (1992: p. E26).
[Carditidae].
Venericardia () mulleri-
cene. Near junction of Panoche and Silver creeks, Panoche quad., Fresno Co. SU 2073 [=CASG 66535]. Holotype
SU 7994 [=CASG 72179, listed as voucher in CASG online database]. [Carditidae].
Venericardia mulleri
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype
UCMP 14207. [Carditidae].
Venericardia () mulleri
[=Santa Susana Fm.] ("Martinez" Stage), Paleocene. Garapito Creek, Santa Monica Mtns., Los Angeles Co. LACMIP
16403. Hypotype UCLA 59137 [=LACMIP 10426]. [Carditidae].
Venericardia () mulleri
Paleocene. Near junction of Panoche and Silver creeks, Panoche quad., Fresno Co. SU 2073. Holotype SU 7994
[=CASG 72179, as voucher in CASG online database]. [Carditidae].
Venericardia () nelsoni-
    
Camulos quad., Ventura Co. UCMP 3765. Holotype UCMP 32804. [Carditidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 179
Venericardia () nelsoni Verástegui, 1953
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6825. Hypotype UCR 6838/100 (not UCR 6825/1 as in pl.
caption). [Carditidae].
Venericardia () nelsoni
Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCMP 3765. Holotype (of Venericardia () transversaria
Verástegui, 1953) UCMP 32804a determined here as missing. [Carditidae].
Venericardia () nelsoni
Fm. (of Zinsmeister, 1983a: p. 62), Paleocene. Simi Hills, Camulos quad., Ventura Co. UCMP 3765. Holotype UCMP
32804 determined here as missing. [Carditidae].
Venericardia (Leuroactis) popenoei-
ter” Sandstone, Eocene. On east side of main branch of Sespe Gorge, Ventura Co. UCLA L581 [=LACMIP 20581].
Holotype UCLA 15689 [=LACMIP 10845]. Remarks: Venericardia () clarki popenoei Verástegui, 1953,
 Moore (1992: p. E24). [Carditidae].
Venericardia cf.  (M.A. Hanna, 1927). [Cardita
Sandstone, Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS 14492. Hypotype USNM 498737. Remarks: In
the genus/subgenus Glyptoactis (Claibornicardia),  Moore (1992: p. E26). [Carditidae].
Venericardia (Leuroactis) schencki
Fm., Eocene. Simi Hills area, Camulos quad., Ventura Co. CASG 66532. Holotype SU 8003[=CASG 66532.01]. Re-
marks: Junior synonym of Venericardia () aragonia joaquinensis (Vokes, 1939),  Moore (1992: p.
E23). [Carditidae].
Venericardia (Venericor) simiana
       
quad., Ventura Co. SU 2697 [=CASG 66533]. Holotype SU 8001 [=CASG 66533.01], paratype SU 8002 [=CASG
72180]. Remarks: Junior synonym of Venericardia (Venericor) venturensis Waring, 1914,  Moore (1992: p.
E25). [Carditidae].
Venericardia (Leuroactis) smileyi (Vokes, 1939). [Megacardita
4, 5. Avenal Sandstone, Eocene. Coalinga quad., Fresno Co. UCMP 672. Holotype UCMP 15626. Remarks: Junior
synonym of Venericardia () aragonia joaquinensis (Vokes, 1939),  Moore (1992: p. E23). [Carditidae].
Venericardia () susanaensis
Fm., Eocene. From McCray Oil Wells, Oil Canyon, Camulos quad., Ventura Co. SU Ca-33 [=CASG 66527]. Holotype
SU 8004 [=CASG 66527.01]. Remarks: Venericardia () Verástegui, 1953,  Moore
(1992: p. E19). [Carditidae].
Venericardia () taliaferroi
Fm., Paleocene. Vicinity of Nacimiento River, Adelaida quad., San Luis Obispo Co. CASG 66526]. Holotype SU 7996
[=CASG 66526.01], paratype SU 7997 [=CASG 72181]. [Carditidae].
Venericardia () taliaferroi Verástegui, 1953. Saul 1986. SEPM 
uppermost Cretaceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo
Co. UCLA 6525 [=LACMIP 26525]. Hypotype LACMIP 7558. [Carditidae].
Venericardia () taliaferroi
Fm. (of Verástegui 1953:433), Cretaceous/Paleocene. Vicinity of Nacimiento River, Adelaida quad., San Luis
Obispo Co. CASG 66526]. Holotype SU 7996 [=CASG 66526.01], paratype (hypotype in pl. caption) CASG 7997
[=CASG 72181]. [Carditidae].
Venericardia () transversaria

quad., Ventura Co. UCMP 3765. Holotype UCMP 32804a determined here as missing. Remarks: Junior synonym
of Venericardia () nelsoni Verástegui, 1953,  Moore (1992: p. E19). [Carditidae].
Venericardia () vallecitosensis (Vokes, 1939). [Megacardita]. Verástegui 1953. Pal. Amer. 3(25):35–36, pl. 12,
180 PALEOBIOS, VOLUME 38, FEBRUARY 2021

UCMP 15614, hypotype [topotype] UCMP 30177. [Carditidae].
Venericardia (Venericor) vallecitosensis (Vokes, 1939). [Megacardita]. Moore 1992. USGS PP 1228-E:E25–E26, pl.
Venericardia
() hornii carlosensis Verástegui, 1953) UCMP 15619. [Carditidae].

UCMP 15614, paratype UCMP 15615.
Venericardia (Venericor) venturensis

Valley, Ventura Co. Hypotype SU 7998 [=CASG 72183]. [Carditidae].

[=CASG 72183].

Venericardia (Venicor)* venturensis
Susana Fm., Paleocene. Meier Canyon, Simi Valley, Ventura Co. UCR 4572]. Hypotype UCR 4572/1. Remarks:
Error for Venericor. [Carditidae].
Venericardia (Venericor venturensis

[=LACMIP 22689]. Hypotype UCLA 59362 [=LACMIP 10425]. [Carditidae].
Venericardia (Venericor) venturensis Waring, 1917. Moore 1992. USGS PP 1228-E:E25, pl. 
14. Santa Susana Fm. (of Zinsmeister, 1983a: p. 62), Paleocene. Las Virgenes Canyon area, Simi Hills, Camulos
quad., Ventura Co. SU 2697. Holotype CASG 159 [=CASG 61667.08, as syntype in CASG online database], hypo-
types [paratypes,  Keen and Bentson 1944: p. 120] CASG 318, 319 [=CASG 72182–72184, unable to match
specimens to numbers], holotype SU/CASG 8001 [=CASG 66533], paratype SU/CASG 8002 [=CASG 72180] (both
of Venericardia (Venericardia) simiana Verástegui, 1953). [Carditidae].
Venericardia (Leuroactis) vokesi
Eocene. Vicinity of Big Tar Canyon, Cholame quad., Kings Co. UCMP A819. Holotype UCMP 15618. Remarks: Junior
synonym of Venericardia () aragonia joaquinensis (Vokes, 1939),  Moore (1992: p. E23). [Carditidae].

Venericardia
Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8758 [= SBMNH 60181]. Remarks: Determined
here as Venericardia (Venericor). [Carditidae].
  

Venerupis laciniata hannibali (Howe, 1922). [Paphia
Etchegoin Fm. (Upper Pseudocardium Zone), Pliocene. La Cima, North Dome, Kettleman Hills, Kings Co. USGS
12414. Hypotype USNM 495787. Remarks: Protothaca staleyi (Gabb) hannibali (Howe),  Adegoke (1969: p.
146). Possibly in the genus Leukoma,  P. Valentich-Scott (personal communication, 2019). [Veneridae].
Ventricolaria welchensis
Hercules, Contra Costa Co. SU 3255 [=CASG 100138]. Holotype SU 8460 [=CASG 69989]. [Veneridae].

Paratype SU 8461 [=CASG 69967].
Venus (Chione) securis

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 181
[=CASG 71594]. Remarks: In the genus Securella,  Parker (1949: p. 593). [Veneridae].
Chione securis (Shumard)). [Jacalitos
Fm.], Miocene. Coalinga District, Fresno Co. USGS. 4763. Hypotype USNM 165599].
Venus   
(1943), Alameda Co. SU 3252. Hypotype SU 8464 (not SU 8446 as in pl. caption) [=CASG 69988]. [Veneridae].
Verticordia (Trigonulina) ornata (d'Orbigny, 1853). [Trigonulina]. Hertlein and Grant 1972. SDSNH Mem. 2(2B):344,
  Diego Co., San Ysidro quad. (1943), San
Diego Co. LACMIP 305. Hypotype LACMIP 4884. Remarks: Trigonulina  Jung, 1996,  Coan et al. (2000:
p. 563). [Verticordiidae].
Vertipecten alexclarki-
stone Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotypes USNM
46523, 646525. [Pectinidae].

646524.
   
USNM 646528.
     
Hypotype USNM 646529 [holotype, 


Moore, 1984: p. B62; 


USNM 646539 as in pl. caption; [syntype in USNM print out]). Remarks:
(as Vertipecten n. sp.).
Vertipecten alexclarki-
blor Fm. (Wygal Sandstone Member), Oligocene. Central Temblor Range, Las Yeguas Ranch quad., Kern Co. USGS

Vertipecten alexclarki-
gal Sandstone Member), Oligocene. Las Yeguas Ranch quad., Kern Co. USGS M3281. Lectotype (of Moore 1984:


Vertipecten alexclarki
Member), Oligocene. Media Agua Creek area, Las Yeguas Ranch quad., Kern Co. USGS M3281. Lectotype (of Moore,
1984:B62) USNM 646531. [Pectinidae].
 area, Carneros Rocks quad., Kern Co. CASG 27625. Hypotype CASG
60862.

quad., Kern Co. USGS M3985. Hypotype USNM 646538 (not USNM 646539 as in pl. caption).

Ranch quad., Kern Co. USGS. M3772. Hypotype USNM 646529.
182 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Vertipecten bowersi (Arnold, 1906). [Pecten
Fm., Miocene. Santa Inez [sic] Canyon, Santa Ynez quad., Santa Barbara Co. Holotype UCMP 12075. [Pectinidae].
Vertipecten bowersi (Arnold, 1906). [Pecten


Salisbury Canyon quad., San Luis Obispo Co. USGS M3436. Hypotype USNM 335951.
Vertipecten diabloensis (Clark in G D. Hanna, 1924). [Pecten

paratype UCMP 11139. [Pectinidae].
Vertipecten fucanus (Dall, 1898). [Pecten
North of Poso [=Ocoya Creek], Knob Hill quad., Kern Co. CASG 1452. Hypotype CASG 6098. [Pectinidae].
Vertipecten kernensis (Hertlein, 1925a). [Pecten
(Monterey Fm. in pl. caption), Miocene. Near Pyramid Hill, Caliente quad., Kern Co. SU 150 [=CASG 61886]. Ho-
lotype SU 128 [=CASG 61886.01]. [Pectinidae].
Vertipecten kernensis (Hertlein, 1925a). [Pecten
Miocene. Pyramid Hil areal, Rio Bravo Ranch quad., Kern Co. USGS M1590. Hypotype USNM. 335002. [Pectinidae].

quad., Kern Co. USGS M5211. Hypotype (of Vertipecten perrini
4), determined here as missing. Remarks: Hypotype never received from author, according to label in USNM
collection.

462. Hypotypes LACMIP 5873, 5874.

Kern Co. UCMP B1673. Hypotype (of Vertipecten nevadanus
36554.
, Junipero Serra quad., Monterey
Co. CASG 31524. Hypotype CASG 60986.

SU 150. Holotype SU 128 [=CASG 61886.01]

quad., Contra Costa Co. UCMP 1311. Holotype (of Pecten (Lyropecten) gabbi Clark, 1918 and of Pecten diabloensis
Clark in G D. Hanna, 1924) UCMP 11138.
Vertipecten nevadanus (Conrad, 1855b). [Pecten
Poso Creek, north of Coffee Canyon, Pine Mtn. quad., Kern Co. UCMP B1673. Hypotype UCMP 36554. Remarks:
Junior synonym of Vertipecten kernensis (Hertlein, 1925a),Moore (1984: p. B64). [Pectinidae].
Vertipecten nevadanus (Conrad, 1855b). [Pecten
Fm. (Carneros Sand Member) (of Stinemeyer et al. 1959:9, 13), Miocene. Temblor Range, Kern Co. USGS 4941.
Hypotype USNM 646793a (not USNM 646793 as in pl. caption). Remarks: Vertipecten bowersi (Conrad, 1855b),
 Smith (1991a: p. 81), [=Vertipecten kernensis (Hertlein, 1925a),Smith (1991a: p. 81). [Pectinidae].
Vertipecten nevadanus (Conrad, 1855b). [Pecten
Vertipecten perrini). Jewett Sand, Miocene. Pyramid
Hill area, Rio Bravo Ranch quad., Kern Co. USGS M5211. Hypotype USNM 647085. Remarks: Label in USNM col-

Vertipecten kernensis (Hertlein, 1925a),Moore (1984: p. B64). [Pectinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 183

Vertipecten perrini (Arnold, 1906). [Pecten-
mid Hill Sand Member), Miocene. Kern River area, Kern Co. USGS M5211. Hypotype USNM 647085. Remarks:

Vertipecten kernensis (Hertlein, 1925a),  Smith (1991a: p. 81). [Pectinidae].
Vertipecten perrini (Arnold, 1906). [Pecten
5. Vaqueros Fm., Oligocene. Between Morro and Torro creeks, Morro Bay North quad., San Luis Obispo Co. UCLA
6204 [=LACMIP 26204]. Holotype SU 13 [=CASG 61824.01]. [Pectinidae].
Vertipecten perrini (Arnold, 1906). [Pecten
Oligocene/Miocene. Between Morro and Toro Creeks, San Luis Obispo Co. Holotype SU 13 [=CASG 61824.01].
[Pectinidae].
Vertipecten perrini (Arnold, 1906). [Pecten
Fm., Oligocene/Miocene. Between Morro and Toro Creeks, Santa Lucia Range, Cayucos quad., San Luis Obispo
Co. Type area rediscovered,  Hall and Prior (1975), to be on Morro Bay North quad., UCLA 6204,  Smith
(1991a: p. 84). Holotype SU 13 [=CASG 61824.01]. [Pectinidae].
Vertipecten cf.  (Conrad, 1849a). [Pecten      
Fm., Miocene. Reef Ridge, near Roundtop, Reef Ridge quad., Fresno Co. USGS 14393. Hypotype USNM 498742.
Remarks: Patinopecten propatulus (Conrad, 1849a),  Moore (1984: p. B79). However, also included in the
synonymy of Vertipecten fucanus (Dall, 1898),  Moore (1984: p. B63). [Pectinidae].
Vertipecten yneziana (Arnold, 1907). [Pecten
Eocene/Oligocene. Nojoqui Creek, Los Olivos quad., Santa Barbara Co. UCMP B6985. Hypotype (of Pecten (Ver-
tipecten) yneziana yneziana
Vertipecten yneziana yneziana (Arnold, 1907). [Pecten
Fm., Eocene. Lompoc, Lompoc quad., Santa Barbara Co. USGS 4507. Holotype USNM 165313a, paratype USNM
165313. [Pectinidae].
Vertipecten yneziana (Arnold, 1907). [Pecten-
blee 1950:29–30), Eocene. Nojoqui Creek, Santa Ynez Mtns., Los Olivos quad., Santa Barbara Co. UCMP B6985
[=SU 2908a]. Hypotype (of Pecten (Vertipecten) yneziana yneziana Arnold of Weaver and Kleinpell 1963, pl. 31,


Santa Barbara Co. UCMP B6955. Hypotype (of Pecten (Vertipecten) yneziana yneziana Arnold of Weaver and

  
M4507. Holotype USNM 165313a, paratype USNM 165313.
Vertipecten yneziana subyneziana (Weaver and Kleinpell, 1963). [Pecten]. Moore 1984. USGS PP 1228-B:B61, pl.

Mtn. quad., Santa Barbara Co. UCMP B6940. Holotype SU/CASG 9265. [Pectinidae].
Vertipecten-
ber), Oligocene. South side of Stone Corral, Las Yeguas Ranch quad., Temblor Range, Kern Co. USGS M3985 (not
M3984 as in pl. caption). Hypotype USNM 646538. Remarks: Vertipecten alexclarki n. sp.,  Addicott (1973b:
p. 26). [Pectinidae].
Remarks: Ver-
tipecten alexclarki n. sp.,  Addicott (1973b: p. 26).
Vesicomya (Vesicomya) aff  ( ) tschudi
Wagonwheel Fm., Eocene. Wagonwheel Mtn. Emigrant Hill quad. (1953, PR 1973), Kern Co. CSUN 1580 [=LACMIP
16886]. Hypotypes LACMIP 12360–12362. [Vesicomyidae].
184 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Seco, 
Vesicomyidae indeterminate. Powell et al. 2019b. PaleoBios
excavation, Berkeley Hills, Oakland Hills, Alameda Co. UCMP IP16025. Hypotype UCMP 123686. [Vesicomyidae].

IP16027. Hypotype UCMP 123690.

IP16022. Hypotype UCMP 123682.
Volsella cf.  (Conrad, 1837). [Modiola-
quin Fm. (Pecten zone), Pliocene. Arroyo Pequeño and Arroyo Finito area, North Dome, Kettleman Hills, Kings
Co. USGS 12343. Hypotype USNM 495778. Remarks: In the genus Modiolus,  Coan and Valentich-Scott (2012:
p. 146). [Mytilidae].
Volsella (Modiolus) cf. gabbi (Clark, 1915). [Modiolus
Concord quad. (1915), Contra Costa Co. UCMP A2569. Hypotype UCMP 30671. Remarks: In the genus/subgenus
Brachidontes (Aeidimytilus),  Moore (1983: p. A69). [Mytilidae].
Volsella cf.  (Linneaus, 1758). [Modiolus
Etchegoin, Pliocene. Vicinity of El Vejon, South Dome, Kettleman Hills, Kings Co. USGS 12652. Hypotypes USNM
495364, 495365. Remarks: In the genus Modiolus,  Moore (1983: p. A77). [Mytilidae].
Volsella cf.  (Conrad, 1837). [Modiolus-
quin Fm. (Pecten zone), Pliocene. Near Arroyo Vadoso and Arroyo Largo, North Dome, Kettleman Hills, Kings Co.
USGS 12331. Hypotype USNM 495740. Remarks: In the genus Modiolus,  Moore (1983: p. A76). [Mytilidae].
Neverita
USGS 12493. Hypotype USNM 495330.

495328.
Volsella recta (Conrad, 1837). [Modiolus
Modiolus rectus Conrad)). [Etchegoin Fm.], Pliocene. Coalinga
District, Fresno Co. USGS 4698. Hypotype USNM 165535]. Remarks: In the genus Modiolus,  Moore (1983:
p. A76). [Mytilidae].
Volsella

Joaquin Fm. (Neverita Zone), Pliocene. North Dome, Kettleman Hills, Kings Co. USGS 12499. Hypotype USNM
(slab with Neverita reclusiana (=Glossaulax reclusiana), Olivella cf.  (=Calianax alectrona) and Anadara
trilineata) determined here as missing. [Mytilidae].
Vulsellaclarki-
ber), Paleocene/Eocene. Near where Urruttia Canyon enters Salt Creek, Coalinga quad., Fresno Co. UCMP 1817.
Holotype UCMP 15743. [Malleidae].
Yoldia (Kalayoldia carnarosensis
Fm. in text), Miocene. Carneros Creek, Napa Co. Holotype UCMP 11539. [Yoldiidae].
Yoldia (Kalayoldia) cf.  () carnarosensis
Miocene. Oil Center quad. (1954), Kern Co. USGS. M1596. Hypotype USNM 646526. [Yoldiidae].
Yoldia cooperi supramontereyensis
Near Pacheco, Contra Costa Co. UCMP A4929. Hypotype UCMP 34123. Remarks: Probably junior synonym of
Yoldia (Kalayoldia) cooperi Gabb, 1865,  Coan et al. (2000: p. 113). [Yoldiidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 185
Yoldia gala Woodring in
(diatomaceous strata), Miocene. Gato Ridge, Santa Barbara Co. USGS 15023. Hypotype USNM 560027. Remarks:
Provisionally in the genus Yoldia Moore (1983: p. A28). [Yoldiidae].



Yoldiagala Woodring in
6. Sisquoc Fm. (Todos Santos Claystone Member), Miocene/Pliocene. Solomon Hills, Vicinity of Los Alamos, Santa
Barbara Co. USGS 15020. Holotype USNM 560024. [Yoldiidae].
Yoldia (Kalayoldia gesteri-
cene. East of Lower Lake village, near Herndon Creek, Lake Co. UCMP 790. Holotype UCMP 11685. [Yoldiidae].
Yoldia gesteri
Simi Hills, Ventura Co. UCR 6684. Hypotype UCR 6684/17. Remarks: Provisionally in the subgenus Kalayoldia
 Moore (1983: p. A26). [Yoldiidae].
Yoldia impressa-
ton quad.], Santa Cruz Co. SCCM 6. Hypotype SCCM 21006]. Remarks: Junior synonym of Litorhadia astoriana
(Henderson, 1920),  Moore (1983: p. A23). [Nuculanidae].
Yoldia (Cnesterium) seminuda Dall, 1871. Moore 1983. USGS PP 1228
Co. UCMP 7100. Holotype (of Yoldia (Cnesterium) imleri Waterfall, 1929) UCMP 31420. [Yoldiidae].


Yoldia (Kalayoldia) submontereyensis
Fm. (=Unnamed Miocene of Dibblee 1966), Oligocene/Miocene. On San Francisquito Creek (below dam), near
   
165460]. Remarks: The hypotype of Turritella ineziana Conrad, 1857 of Arnold, 1908 also carries the number
USNM 165459 on specimen and label, but is apparently corrected on USNM list in specimen tray as USNM 165460.
Yoldia
(Kalayoldia) submontereyensis is USNM.165460. [Yoldiidae].
Yoldia cf.  Arnold, 1908. Powell et al. 2019b. PaleoBios
Caldecott Tunnel excavation, Berkeley Hills, Oakland, Alameda Co. UCMP IP 13003. Hypotypes UCMP 218771,
218804. [Yoldiidae].
Yoldia supramontereyensis
(Hames Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M2044. Hypotype
USNM 254354. Remarks: In the subgenus Kalayoldia,  Moore (1983: p. A25). [Yoldiidae].
Yoldia supramontereyensis

San Luis Obispo Co. USGS M2044. Hypotype USNM 254354. Remarks: In the subgenus Kalayoldia,  Moore
(1983: p. A25). [Yoldiidae].
Yoldia (Kalayoldia) supramontereyensis 

Santa Cruz quad., Santa Clara Co. Holotype CASG 5362 [=CASG 69778] . [Yoldiidae].
Yoldia cf.  Arnold, 1908. Powell et al. 2019b. PaleoBios 
Caldecott Tunnel excavation, Berkeley Hills, Oakland, Alameda Co. UCMP IP 13003. Hypotype UCMP 218714.
[Yoldiidae].
186 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Yoldia (Yoldia temblorensis
Mtn. Silt, Miocene. North bank of Kern River, Caliente quad., Kern Co. CASG 68. Holotype CASG 106. [Yoldiidae].
Yoldia tenuissima
Jaro Creek area, Lompoc quad., Santa Barbara Co. UCMP B7017. Hypotype SU 9248 [=CASG 70244]. Remarks:
In the subgenus Kalayoldia,  Addicott (1973b: pp. 22, 23); Moore 1983: p. A24. [Yoldiidae].
Yoldia (Kalayoldia) tenuissima Clark, 1918. 2, 3, 5. Temblor Fm.
(Wygal Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotypes USNM 646513–
646515. [Yoldiidae].

at base of bluff below where fossils were collected at USGS 6623, Kern Co.] USGS 6622. Hypotype USNM 646516.
Yoldia (Kalayoldia) tenuissima-

Yoldia thraciaeformis
Member), Pliocene. Arroyo Cañada Verde area, Half Moon Bay quad., San Mateo Co. Locality F685-3M. Hypotype
determined here as missing. Remarks: In the genus Megayoldia,  Coan et al. (2000: p. 114). [Yoldiidae].
Yoldia (Orthoyoldia-
cene. Monocline Ridge, Fresno Co. UCMP D6528]. Hypotype UCMP 37159. [Yoldiidae].
Yoldia (Orthoyoldia
Monocline Ridge quad., Fresno Co. UCMP D6528. Hypotype UCMP 37159. [Yoldiidae].
Yoldia
[=Wilson Grove Fm.], Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotype CASG, determined here as
missing. [Yoldiidae].
Yoldia

YoldiaPaleoBios
Hills, Oakland, Alameda Co. UCMP IP 13003. Hypotype UCMP 218702. [Yoldiidae].
Zirfaea dentata



[possible holotype,  Kennedy 1974: p. 31].
Zirfaea gabbi Tryon, 1863, var. Woodring et al. 1940 [1941]. [Zirphaea
(Macoma zone), Pliocene. Vicinity of El Chichon, North Dome, Kettleman Hills, Kings Co. USGS 12421. Hypotype
USNM 495837. Remarks: Zirfaea pilsbryi Lowe, 1931,  Kennedy (1974: p. 31). [Pholadidae].
Zirfaea gabbi femii San
Joaquin Fm., Pliocene. Reef Ridge quadangle, Kings Co. UCMP.B6538. Holotype UCMP 36795, paratypes UCMP
36796–36798. Remarks: Junior synonym of Zirfaea pilsbryi Lowe, 1931,  Kennedy (1974: p. 31). [Pholadidae].
Zirfaeaquad.,
Santa Clara Co. USGS M1715. Hypotype USNM 650968. Remarks: Zirfaea pilsbryi Lowe, 1931,  Kennedy
(1974: p. 31). [Pholadidae].
GASTROPODA
Acanthina perrini-
anton, La Costa Valley quad. (1943), Alameda Co. SU 2739. Hypotypes SU 8477, 8478 [=CASG 68839, 68840].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 187
[Muricidae].
Acanthina
Miocene/Pliocene. Northeast of San Lucas, Monterey Co. USGS M903. Hypotype USNM 254350 determined here
as missing. [Muricidae].
Acanthinucella emersoni (Hertlein and Allison, 1959). [Acanthina
22. Pico Fm., Pliocene. Southwest of Valencia, Oat Mtn. quad. (1952, PR 1969), Los Angeles Co. LACMIP 15719.
Hypotype LACMIP 13351. [Muricidae].
Acmaea Eschscholtz in Rathke, 1833. Vedder and Moore 1976. AAPG Misc. Pub. 24:124, 130, pl.

M6501. Hypotype USNM 235586. [Acmaeidae].
Acrilla (Ferminoscala) durhami-
sana Fm., Paleocene. Meier Canyon, Calabasas quad., Simi Hills, Ventura Co. UCR 6670. Holotype UCR 6670/240
determined here as missing. Paratype UCR 6670/241. Remarks: Ferminoscala =Amaea,  WoRMS (2020),
thus making the name Amaea durhami (Zinsmeister, 1983b) a secondary homonym of Amaea durhami (Keen,
-
dition, Zinsmeister’s species is missing its morphologically diagnostic aperture. His species might not even be
an epitoniid. [Undetermined].
Acrilla procerus     
Paleocene. Runkle Canyon, Calabasas quad., Simi Hills, Ventura Co. UCR 4579. Holotype UCR 4579/100 (not UCR.
4579/10 as in text) paratype UCR 4579/101. [Epitoniidae].
Acrilla-
tura Co. UCR 4579. Hypotype UCR 4579/101 (not UCLA 4579/4 as in pl. caption). Remarks: Described as Acrilla
procerus n. sp.,  Zinsmeister (1983b: p. 1291). [Epitoniidae].
Acrilla (FerminoscalaSusana Fm., Paleocene.
Simi Hills, Ventura Co. UCR 6670. Hypotype UCR 6670/12 [same number also listed for Cancellaria crassa War-
ing, 1917 of Zinsmeister 1983a]. Remarks: Described as Acrilla (Ferminoscala) durhami n. sp.,  Zinsmeister
(1983b: p. 1291). [Epitoniidae].
Acteocina culcitellaBulla (Akera
Newhall area, Los Angeles Co. LACMIP 7760. Hypotype LACMIP 14416. [Tornatinidae].
Acteon boulderana

7529 [=CASG 72019]. Remarks: Acteon boulderanus Etherington, 1931,  Addicott (1970: p. 137). [Acteonidae].
Acteon cf. 
Castaic Canyon Los Angeles Co. CIT 2093 [=LACMIP 11853[. Hypotype LACMIP 4215. Remarks: Acteon boul-
deranus Etherington, 1931,  Addicott (1970: p. 137). [Acteonidae].
Acteon boulderanus
Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650271. [Acteonidae].

Tivela bed Rio Bravo Ranch quad. (1954), Rio Bravo Ranch quad., Kern Co. USGS M1600. Hypotype
USNM 650273.
Acteon quercus
Near Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4722. Hypotype UCR 4722/14. [Acteonidae].
Acteon (Rictaxis) weaveri
Rio Bravo Ranch quad. (1954), Kern Co. USGS M2601. Holotype USNM 650282. Remarks: Determined here as
in the genus Rictaxis. [Acteonidae].
188 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Acteon
Los Angeles Co. LACMIP 17850. Hypotype LACMIP 13435. [Acteonidae].
Admete clatskaniensis
Domengine Ranch quad., Fresno Co. UCMP D1078. Hypotype UCMP 36899. [Cancellariide].
Admete gracilior (Carpenter in Gabb 1869a) of Arnold. [Cancellaria]. Woodring and Bramlette 1950 [1951]. USGS PP

Hypotype USNM 560088. Remarks: Gabb’s description of this species was quoted from Carpenter (in lit.) and
he is clearly not the author , Groves and Squires (2018: p. 118). [Cancellariide].
Agaronia mathewsonii (Gabb, 1864). [Olivella 
Eocene. Reef Ridge, southeast of Big Tar Canyon, Reef Ridge quad., Kings Co. USGS 14495. Hypotype USNM
498702. [Olividae].
Agaronia Fowkes 1982. Coalinga Guidebook, p. 22, unnumbered 
Kings Co. Remarks: Determinded here as Agaronia mathewsonii (Gabb, 1864). [Olividae].
Agaronia-
ertine Rock, Imperial Co. (not San Diego Co. as in locality description). USGS M9779 [=USGS M9778]. Hypotype
USNM 486906. [Olividae].
Agasoma sinuatum (Gabb, 1866). [Clavella
Koilopleura sinuata
Agasoma sinuata)). [Briones Fm.], Miocene. San Pablo, Walnut Creek, Contra Costa
Co. UCMP 1354. Hypotype [UCMP 11900]. [Buccinidae].
Agasoma sinuatum (Gabb, 1866). [Clavella
Fm.], Miocene. Cuyama Valley, Santa Barbara Co. USGS 13021. Hypotype USNM 562011. [Buccinidae].
    
Barbara Co. UCLA 1718 [=LACMIP 21718]. Hypotype UCLA 27800 [=LACMIP 10977].
Akera maga
Co. LACMIP 26624. Hypotype LACMIP 12746. [Akeridae].
Aletes aquamigerus*          
(Tahana Member), Pliocene. Vicinity of Pomponio Creek, San Gregorio quad., San Mateo Co. locality F467-7M.
Hypotype determined here as missing. Remarks: Error for squamigerus. In the genus Thylacodes,  Golding
et al. (2014: pp. 9, 14). [Vermetidae].
Alia tuberosa (Carpenter, 1864b). [Amycla
area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14399. Remarks: In the genus Mitrella,  deMaintenon
(2019: p. 176). [Columbellidae].
Amaea (Scalina) sp., cf. A. (S.) edwilsoni
Pliocene. Newhall area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14393. [Epitoniidae].
Amaurellina caleocia-
gine Fm., Eocene. Parson's Peak area, north of Coalinga, Coalinga quad., Fresno Co. UCMP 672. Holotype UCMP
33781, paratype UCMP 15862. [Naticidae].

UCMP 15864 (both of Amaurellina garzaensis Vokes, 1939).
Amaurellina caleocia
Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 219 [=LACMIP 16100]. Hypotype LACMIP 7470.
[Naticidae].
Amaurellina caleocia Vokes, 1939. Squires 1988a. Northern
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 189
Lockwood Valley area, Ventura Co. CSUN 979 [=LACMIP 16081]. Hypotype LACMIP 7707. [Naticidae].
Amaurellina caleocia
Valley, Sawmill Mtn. quad., Ventura Co. CSUN 979 [=LACMIP 16081]. Hypotype LACMIP 7707. [Naticidae].
Amaurellina caleocia 

662]. Hypotype LACMIP 11466. [Naticidae].
Amaurellina caleocia
Mtn. quad. (1952, PR 1969), southwestern Santa Susana Mtns., Ventura Co. LACMIP 7418. Hypotype LACMIP
12735. [Naticidae].
Amaurellina (Euspirocrommium clarki
Sandstone, Eocene. Reef Ridge, near Big Tar Canyon, Reef Ridge quad., Kings Co. USGS 14495. Hypotype USNM
498704. Remarks: In the genus Pachycrommium,  Marincovich (1977: p. 238). [Naticidae].
Amaurellina clarki
Somersville, Contra Costa Co. CASG 261. Hypotype CASG 6725]. Remarks: In the genus Pachycrommium, 
Marincovich (1977: p. 238). [Naticidae].
Amaurellina clarki

UCMP 31385]. Remarks: In the genus Pachycrommium,  Marincovich (1977: p. 238). [Naticidae].
Amaurellina (Euspirocrommium) clarki
15. Gaviota Fm., Eocene. Stratigraphically just above Sacate-Gaviota contact in narrow gorge of branch of Cañada
del Corral, Los Olivos quad., Santa Barbara Co. UCMP B6982. Hypotype SU 9202 [=CASG 70129]. Remarks: In
the genus Pachycrommium,  Marincovich (1977: p. 238). [Naticidae].
Amaurellina lajollaensis 
Eocene. Reef Ridge, Vicinity of Canoas Canyon, Dark Hole quad., Fresno Co. USGS 14503. Hypotype USNM 498703.
Remarks: Junior synonym of Tejonia moragai (Stewart, 1926 [1927],  Marincovich (1977: p. 233). [Naticidae].
Amaurellina moragai
Shale, Eocene. Nojoqui Creek area, Santa Barbara Co. UCMP B6932. Hypotype SU 9203 [=CASG 70132]. Remarks:
In the genus Tejonia,  Marincovich (1977: p. 233). [Naticidae].
Amaurellina moragai lajollaensis

of Rose Canyon, San Diego Co. UCMP 3976. Holotype UCMP 31401]. [Naticidae]. Remarks: Junior synonym of
Tejonia moragai (Stewart, 1926 [1927]),  Marincovich (1977: p. 233). [Naticidae].
Amaurellina moragai lajollaensis Stewart, 1926 [1927]. Rose
Canyon Fm.” [=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. [SDSNH 2363. Hypotype SDSNH 4182].
Remarks: Junior synonym of Tejonia moragai (Stewart, 1926 [1927]),  Marincovich (1977: p. 233). [Naticidae].
Amaurellina

Amauropsis martinezensis
Fm., Paleocene. Near Stewartville, Contra Costa Co. UCMP 154]. Holotype UCMP 11702. [Naticidae].
-
potype UCR. 6814/500.
Remarks: Described as
Amauropsis meirensis n. sp.,  Zinsmeister (1983b: p. 1292). Holotype UCR 6825/100.

190 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Amauropsis martinezensis
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6814. Hypotype UCR 6814/500 (not UCR. 6814/1 as in pl.
caption). [Naticidae].
Amauropsis martinezensis-
sana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6814. Hypotype UCR 6814/500 (not UCR 6814/100 as in pl.
caption). [Naticidae].
Amauropsis meierensis        
Paleocene. Calabasas quad., Simi Hills, Ventura Co. UCR 6825. Holotype UCR 6825/100 (not UCR 6825/4 as in
text). [Naticidae].

Amauropsis
Ventura Co. UCR 6825. Hypotype UCR 6825/100 (not UCR 6825/4 as in pl. caption). Remarks: Described as
Amauropsis meierensis n. sp.,  Zinsmeister (1983b: p. 1292). [Naticidae].
  
Remarks: Described as Amauropsis meierensis n. sp.,  Zinsmeister )1983b: p. 1292).
Amphissa reticulata
Twelvemile Creek area, San Mateo Co. UCMP D3690. Hypotype USNM 251808. [Columbellidae].
Amphissa versicolor Dall, 1871 var. G D. Hanna and Hertlein
Pliocene. Northeast of Schumann [sic], Santa Barbara Co. CASG. 568. Hypotype CASG 5728. [Columbellidae].
Amphissa versicolor
locality). Line drawing, source specimen unknown. [Columbellidae].
Amphissa wahrhaftigi
-
types UCMP 36863, 36864. [Columbellidae].
Amphissa
17918. Hypotype LACMIP 14400. [Columbellidae].
Ampullella hewitti
Domengine Fm., Eocene. Vicinity of Carneros Creek, Kern Co. CASG 32388A [=CASG 61884.01]. Holotype CASG

Remarks: In the genus Euspirocrommium,  Marincovich (1977:p.
237). [Naticidae].
Ampullella hewitti
Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4668. Hypotype UCR 4668/21. Remarks: In the
genus Euspirocrommium,  Marincovich (1977: p. 237). [Naticidae].
Ampullela schencki          

4865/7. Remarks: Junior synonym of Crommium pinyonensis (Dickerson, 1914),  Marincovich (1977: p.
227). [Naticidae].
Ampullospira susanaensis
Susana Fm.], Paleocene. South of Simi Valley, Ventura Co. [CASG 391]. Holotype CASG 985. [Naticidae].
Ampullina
Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8720 [=SBMNH 60194].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 191
Ameranella kewi (Dickerson, 1915). [Nyctilochus
Fm., Eocene. Live Oak Canyon, near Tejon, Kern Co. UCMP 7200. Hypotype (topotype) UCMP 51052 (published
as UCMP 7200). [Ranellidae].
Anachis watsonae-
cene. Caliente quad., Kern Co. SU. 2121 [=CASG 71992]. Holotype SU 7530 [=CASG 71992.00]. Remarks: In the
subgenus  Addicott (1970: p. 86). [Columbellidae].
Anachis (Costoanachis) watsonae Keen, 1943. Addicott, 1970. USGS PP 
Miocene. North of Kern River, Kern Co. USGS 6065. Hypotype USNM. 650139. [Columbellidae].

Anachis (Costoanachis
Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, west side of George F. Canyon, Palos Verdes
Hills, Los Angeles Co. USGS 13127. Hypotype USNM 497073. [Columbellidae].
Ancilla (Oliverato) californica

   Remarks: In the genus Cryptochorda
(family Volutidae),  Vokes (1939: p.140). [Volutidae].
Ancilla (Spirancilla) gabbi 
Susana quad. (1951, PR 1969), Ventura Co. CSUN 489 [=LACMIP 16103]. Hypotype LACMIP 6544. [Ancillariidae].
Ancilla gabbi
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080. Hypotype LACMIP 7483. [Ancillariidae].
Ancillarina
Susana quad. (1951, PR 1969), Ventura Co. CSUN 961 [=LACMIP 17191]. Hypotype LACMIP 12686. [Olividae].
Ancistrolepis? carolineae
Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Holotype UCLA 59401
       
[Buccinidae].
Anomalosipho
area, Los Angeles Co. CIT 1671 [=LACMIP 2556]. [Hypotypes LACMIP 4201, 4202]. [Buccinidae].
Antillophos posunculensis (Anderson and Martin, 1914). [Amphissa]. Addicott 1970. USGS PP 642:96–98, pl. 11,
       
[Columbellidae].


M1604. Hypotype USNM 650159.


Antillophos posunculensis (Anderson and Martin, 1914). [Amphissa]. Addicott et al. 1978. USGS OFR 78-446:58, 60,
. Monterey Fm. (Saltos Shale Member), Miocene. Indian Creek, La Panza Range, San Luis Obispo Co.
USGS M3814. Hypotype USNM 254359. [Columbellidae].
Antillophos posunculensis (Anderson and Martin, 1914). [Amphissa]. Addicott et al. 1980. SEPM Guidebook 6:18, 20,

Indian Creek, La Panza Range, San Luis Obispo Co. USGS M3814. Hypotype USNM 254359. [Columbellidae].
192 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Antillophos cf. posunculensis (Anderson and Martin, 1914). [Amphissa

[Columbellidae].
Antillophos woodringi
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650165; hypotypes USNM 650164,
650166, 650167. [Columbellidae].
Antillophos woodringi
(Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4312. Hypotype SDSNH 83242. [Columbellidae].
Antillophos-
ber), Oligocene. The Indians, Junipero Serra quad., Santa Lucia Range, Monterey Co. SU 1756. Hypotype CASG
60370. [Columbellidae].
Antiplanes perversa (Gabb, 1865). [Pleurotoma
Pliocene. Ferndale quad., Humboldt Co. UCMP B7950. Hypotype UCMP 37799. [Pseudomelatomidae].
Antiplanes voyi (Gabb, 1866). [Pleurotoma    
Fm., Pliocene. Vicinity of Fleener Creek, Ferndale quad., Humboldt Co. UCMP B7939. Hypotype UCMP 37800.
[Pseudomelatomidae].
Antiplanes
canyons, Los Angeles Co. CIT 1849 [=LACMIP 2557]. Hypotype LACMIP 4203. [Pseudomelatomidae].
Apiotoma andersoni (Dickerson, 1914a). [Surcula (Surculites)]. Zinsmeister 1983a. SEPM Guidebook 35:70, pl. 4,

here as missing. [Cochlespiridae].
Apiotoma californiana
quad. (1958), Los Angeles Co. CSUN 360 [=LACMIP 16109]. Holotype LACMIP 7494. [Cochlespiridae].
Apiotoma californiana
Sawmill Mtn. quad. (1943), Ventura Co. CSUN 983 [=LACMIP 16110]. Hypotype LACMIP 7718. [Cochlespiridae].
Architectonia* llajasensis. Llajas Fm., Eocene. Las Llajas Can-
yon area, Santa Susana Mtns., Ventura Co. LACMIP 461B [=LACMIP 16812]. Holotype LACMIP 1140. Remarks:
Error for Architectonica. [Architectonicidae].
Architectonica (Stellaxis) cognata-
Architectonica cognata)). Tejon Fm., Eocene. Martinez, Contra
Costa Co. MCZ 1043. Hypotype MCZ 27811 [=MCZ 112952]. [Architectonicidae].
Architectonica (Stellaxis) cognata   
Eocene. Reef Ridge, southeast of Big Tar Canyon, Reef Ridge quad., Kings, Co. USGS 14495. Hypotype USNM
498705. [Architectonicidae].
Architectonica (Stellaxis) cognata
of San Guillermo Mtn., San Guillermo quad. (1943), Ventura Co. UCR 4750. Hypotype UCR 4750/20. [Architec-
tonicidae].
Architectonica (Stellaxis) cognata
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP
6518 (not LACMIP 6516 as in pl. caption). [Architectonicidae].
Architectonica (Stellaxis) cognata
Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=LACMIP 16114]. Hypotype LACMIP 7455.
[Architectonicidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 193
Architectonica (Stellaxis) cognata     
Eocene. Northern Lockwood Valley area, Ventura Co. CSUN 983 [=LACMIP 16110]. Hypotype LACMIP 7698.
[Architectonicidae].
Architectonica (Stellaxis) cognata         
Lockwood Valley, Sawmill Mtn. quad. (1943). Ventura Co. CSUN 983 [=LACMIP 16110]. Hypotype LACMIP 7698.
[Architectonicidae].
Architectonica (Architectonica) hornii

UCR 4741. Hypotype UCR 4741/60. [Architectonicidae].
Architectonica hornii-
cene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH
20775. [Architectonicidae].
Architectonica (Architectonica) hornii
Matilija Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23688. Hypotype LACMIP 11629. [Archi-
tectonicidae].
Architectonica (Architectonica) llajasensis Fm.,
Eocene. Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP
16115]. Hypotype LACMIP 6517 (not LACMIP 6515 as in pl. caption). [Architectonicidae].
Architectonica (Architectonica) llajasensis
Santa Susana Fm., Eocene. North side Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP
26609. Hypotype LACMIP 12689. [Architectonicidae].
Architectonica nobilis
Fm., Miocene/Pliocene. Alverson Canyon, Imperial Co. SDSNH 351. Hypotype SDSNH 23148. [Architectonicidae].
Architectonica nobilis-
cene. Goat Canyon, just north of the U.S./Mexico border, San Diego Co. LACMIP 305. Hypotype LACMIP 14850].
[Architectonicidae].
Architectonica (Solariaxis) ullreyana
Thorn Meadows area, San Guillermo quad. (1943), Ventura Co. UCR 4690. Hypotype UCR 4690/9. [Architec-
tonicidae].
Architectonica 
Remarks: Determined here as Architectonica (Stellaxis) cognata Gabb, 1864.
[Architectonicidae].
Architectonica

D8157. UCMP unnumbered hypotype lost,  C. Throckmorton (personal communication, 2002). [Architec-
tonicidae].
Arene mcleani  
Sawmill Mtn. quad. (1943), Ventura Co. CSUN 983 [=LACMIP 16110]. Holotype LACMIP 7691, paratypes LACMIP
7692, 7693. [Arenidae].
Arene mcleani

Arene (Arene) stephensoni
[=Latrania Fm.], Pliocene. Whitewater River area, San Jacinto quad., Riverside Co. UCMP A1415] [=UCR 7477].


194 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Argobuccinum (Fusitriton) arnoldi-
cene. Eel River area, near Scotia and Rio Dell, Humboldt Co. UCMP A4539. Hypotype UCMP 37829. Remarks:
Beringinus” [sic] [=Beringius]  Smith (1970: p. 520). [Ranellidae].
Argobuccinum (Fusitriton) coosense Dall, 1909. Faustman 1964. UCPG 41(2):-
cene. Scotia quad., Humboldt Co. UCMP B7643. Hypotype UCMP 37827. Remarks: Junior synonym of Fusitriton
oregonense (, 1846),  Smith (1970: p. 520). [Ranellidae].
Asperiscala hemphilli (Dall, 1878a). [ScalariaScalaria hemphilli). San
Diego Fm., Pliocene. Balboa Park, San Diego Co. Holotype USNM 7991. [Epitoniidae].
Asperiscala sp., cf.  (De Boury, 1912). [Scala
Pliocene. Newhall area, Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14392. [Epitoniidae].
Assorted marine invertebrate fossils. Rugh 2001. Quarterly Dose (IDEC Pharmaceuticals Corp.) 21(3):8, unnum-

San Diego Co. SDSNH 4572. Hypotype SDSNH. 158088]. Remarks: Turritella sp. cf.  Conrad, 1855, 
S. Rugh (personal communication, 2019).
Astraea (Pomaulax) biangulataPachypoma  
7–9. Upper Olcese Sand/lower Round Mtn. Silt, Miocene. Temblor locality 1, Caliente quad. (presumably from the
Round Mtn. Silt near Kern River), Kern Co. UCMP 2713. Hypotypes UCMP 11333, 12147, 12148. [Turbinidae].

Astraea biangulataPachypoma 
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2859. Hypotype USNM
254371. [Turbinidae].
Astraea biangulataPachypoma-

Luis Obispo Co. USGS M2859. Hypotype USNM 254371. [Turbinidae].
Astraea cf. (Pomaulax) gradata
Castaic Fm., Miocene. Southeast of Elderberry Canyon, Los Angeles Co. CIT 2077 [=LACMIP 16117]. [Hypotype
LACMIP 4162]. Remarks: Determined here as in the genus Megastraea. [Turbinidae].
-
potype LACMIP 4163].
Astraea (Pomaulax) gradata
LACMIP 16118. Hypotype UCLA 49523 [=LACMIP 10622]. Remarks: Determined here as in the genus Megas-
traea. [Turbinidae].
Astraea (Pomaulax) gradata Grant and Gale, 1931. Vedder and Moore 1976. AAPG misc. Pub. 24:123, 130, pl. 2, 
9 (peel). Unnamed strata, Pliocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS
M6501. Hypotype USNM 235587. [Turbinidae]. Remarks: Determined here as in the genus Megastraea.
Astraea
Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS M6501. Hypotype USNM 235588.
Remarks: Determined here as in the genus Megastraea. [Turbinidae].
Astraea (Pomaulax  
quad. (1914), Kern Co. UCMP B1616. Hypotype UCMP 11064. Remarks: In the genus Pomaulax (J.H. McLean,
personal communication, 2011). [Turbinidae].
Athleta roddai
(1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Holotype LACMIP 7488, paratype LACMIP 7489. [Volutidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 195
Atys
Rock, Imperial Co. (not San Diego Co. as in locality description). USGS M9779 [=USGS M9778]. Hypotype USNM
486900. [Haminoeidae].
Austrotrophon kernensis (Anderson, 1905). [Trophon 
Fm., Miocene. Garza Creek, Reef Ridge quad., Kings Co. UCMP D1061. Hypotype UCMP 36844. [Muricidae].
Balcis conchita

[Eulimidae].
....., Caliente quad., Kern Co. LACMIP 16140 (ex
....., SDSNH 5422 (ex
Balcis conchita
Rio Bravo Ranch quad. (1954), Kern Co. USGS M1612. Hypotypes USNM 650274, 650275. [Eulimidae].
   
quad. (1954), Kern Co. USGS M1597. Holotype USNM 650309, paratypes USNM 650307, 650308. [Eulimidae].
Balcis cf.  (Bartsch, 1917b). [Melanella
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650303. Eulimidae].
Balcis petrolia
Kern Co. USGS 6623. Hypotype USNM 650286. [Eulimidae].
          
Holotype UCMP. 37112.
Barbarofusus arnoldi (Cossmann, 1903). [Fusus 

1219. Hypotypes current whereabouts unknown. [Fasciolariidae].
Barbarofusus barbarensis (Trask, 1855). [Fusus
Newhall area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14401. [Fasciolariidae].
Bathybembix-
cene/Oligocene. San Lorenzo River, Big Basin quad., Santa Cruz Co. SCCM37. Hypotype SCCM 21002. [Eucyclidae]

Simi Valley, Ventura Co. LACMIP 22371. Holotype LACMIP 12744. [Borsoniidae].
Batillaria californica
USGS M3091. Holotype USNM 305208, paratypes USNM 305209–305211. [Batillariidae].
Batillaria ocoyana (Anderson and Martin, 1914). [Drillia
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650069. [Batillariidae].

       
649170, determined here as missing, 650070.
Batillaria ocoyana (Anderson and Martin, 1914). [Drillia-
cene. Orange Co. Hypotype possibly at SDSNH (current whereabouts unknown). Remarks: Ex Grouard collection.
[Batillariidae].
Batillaria ocoyana (Anderson and Martin, 1914). [Drillia  
Miocene (Cold Creek Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22635 [=UCLA
196 PALEOBIOS, VOLUME 38, FEBRUARY 2021
2635]. Hypotype LACMIP 13075 [=UCLA 57334]. [Batillariidae].
Benoistia cantonensis 
Peak quad. (1958), Los Angeles Co. CSUN 802 [=LACMIP 16144]. Holotype LACMIP 7458. [Littorinidae].
Benoistea* umpquaensis Turner, 1938. Givens and Kennedy 1976. Jour. Paleo. 50(5 
17. Llajas Fm., Eocene. Simi Valley, Ventura Co. UCMP 7005. Holotype (of Tectarius ligniticus Vokes 1939) UCMP
15838. Remarks: Error for Benoistia. [Littorinidae].
Between Vista and San Marcos, San Diego Co. UCR 4865.
Hypotypes UCR 4865/40–4865/42.
Benoistia umpquaensis
quad. (1951, PR 1967) Ventura Co. CSUN 374 [=LACMIP 22312, =LACMIP 7242]. Hypotype UCLA 59274 [=LAC-
MIP 10550]. [Littorinidae].
Beringinus* arnoldi (Martin, 1914). [Argobuccinum
Fm., Pliocene. West of the branching of the Little South Fork of the Elk River, Fortuna quad., Humboldt Co. UCMP
A6855. Hypotype UCMP 10585. Remarks: Error for Beringius. [Buccinidae].

Beringius stantoni (Arnold, 1908). [Chrysodomus
Capitola beach area, Santa Cruz Co. Hypotype determined here as missing. [Buccinidae].
Bittium arnoldi
Elizabeth Lake Canyon, Los Angeles Co. CIT 279 [=LACMIP 7777]. Hypotype LACMIP 4169]. [Cerithiidae].
Bittium asperum (Gabb, 1866). [Turbonilla
Fm. (Pseudocardium zone), Pliocene. Arroyo Conchoso area, North Dome, Kettleman Hills, Kings Co. USGS 12524.
Hypotype USNM 495797. Remarks: In the genus Lirobittium,  Squires (2012: p. 77). [Cerithiidae].
Bittium asperum (Gabb, 1866). [Turbonilla-
ing, source specimen unknown. [Cerithiidae].
Bittium (Lirobittium) asperum (Gabb, 1866). [Turbonilla
Fm., Pliocene. Coalinga quad., Fresno Co. UCMP B6546. Hypotype UCMP 36812. Remarks: In the genus Lirobit-
tium,  Squires (2012: p. 77). [Cerithiidae].
Bittium casmaliense    
Mudstone, Pliocene. Casmalia Hills, near Shuman, Santa Barbara Co. USGS. 14876. Hypotype USNM 560043.
[Cerithiidae].

Bittium casmaliense-
cho Rico Fm., Miocene/Pliocene. Pancho Rico Creek area, San Ardo quad., Monterey Co. USGS M1929. Hypotype
USNM 649108. [Cerithiidae].

Co. USGS M1967. Hypotype USNM 649109.
Bittium casmaliense Bartsch, 1911. Addicott 1969. Pr.CAS 37(3):
Palo Alto quad., Santa Clara Co. USGS M1715. Hypotypes USNM. 650950, 650951. [Cerithiidae].
Bittium casmaliense arnoldi
Careaga Fm. (Cebada Member), Pliocene. Graciosa Ridge near Orcutt, Santa Barbara Co. USGS 14648. Hypotype
USNM 560066. [Cerithiidae]. Remarks: The age of this member is Plio/Pleistocene, 
Bittium dumblei (Dickerson, 1916). [Cerithium
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 197
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7456. [Cerithiidae].
Bittium dumbleiCerithium
Eocene. Whitaker Peak area, Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7456. [Cerithiidae].
Bittium dumblei (Dickerson, 1916). [Cerithium
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 979 [=LACMIP 16081]. Hypotype LACMIP 7699.
[Cerithiidae].
Bittium (Semibittium?) frankeli
Fm., Pliocene. Ferndale quad., Humboldt Co. UCMP B7952. Holotype UCMP 37830, paratypes UCMP 37831,
37832. [Cerithiidae].
Bittium preussi (M.A. Hanna, 1927). [Cerithiopsis

4865/50–4865/53. [Cerithiidae].
Bittium cf.  (M.A. Hanna, 1927). [Cerithiopsis-

LACMIP 11460. [Cerithiidae].
Bittium topangensis (Arnold, 1907). [Cerithium
Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1600. Hypotype USNM 650071. [Cerithiidae].


Bittium topangensis (Arnold, 1907). [Cerithium
Canyon, Santa Monica Mtns., Los Angeles Co. UCLA 2632 (=LACMIP 22632]. Hypotype UCLA 49517 [=LACMIP
10616]. Remarks: Ex Grouard collection. [Cerithiidae].
Bittium topangensis (Arnold, 1907) [Cerithium
Creek Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. [LACMIP 22632 (=UCLA 2632). Hypotype
LACMIP 13076 [=UCLA 57337]. [Cerithiidae].
Bittium cf. B Dall and Bartsch, 1910. Cummings et al. 1962:202, ph
Member), Pliocene. Vicinity of Purisima Creek, Half Moon Bay quad., San Mateo Co. Locality F701M. Hypotype
determined here as missing. [Cerithiidae].
Bittium
area, San Mateo Co. UCMP D5920. Hypotypes USNM 251809, 251810. [Cerithiidae].
Boetica hertleini
most likely is Castaic Fm. (Miocene). Mint Canyon quad., Los Angeles Co. LACMIP 291. Holotype LACMIP 1145,

Bonellitia paucivaricata (Gabb, 1864). [Tritonium
Sandstone, Eocene. Nojoqui Creek area, Lompoc quad., Santa Barbara Co. UCMP B6920. Hypotype SU 9235
[=CASG 70138]. [Cancellariidae].
Bonellitia (Admetula) paucivaricata (Gabb, 1864). [Tritonium

LACMIP 7717. [Cancellariidae].
Boreotrophon durhami
Near Fleener Creek, Ferndale quad., Humboldt Co. UCMP B7939 (not UCMP B7949 as in text and pl. caption for
     Remarks: The age of this member is
Pliocene/Pleistocene, 
198 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Boreotrophon multicostatus Trophon]. Woodring and Bramlette 1950 [1951].

560081. [Muricidae].
Boreotrophon stuarti var. cierboensis Grant and Allen in
Cierbo Fm., Miocene. Caliente Range, San Luis Obispo Co. UCLA 1856 [=LACMIP 21856]. Holotype UCLA 8935
[=LACMIP 8962]. [Muricidae].
Brachysphingus gabbi
Martinez Fm., Eocene [Paleocene]. Contra Costa Co. CASG 27174. Hypotype CASG 5700. Remarks: Brachysphingus
sinuatus Gabb, 1869b,  Squires (1997a: p. 855). [Buccinidae].
Brachysphingus gabbi
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 4573. Hypotype UCR 4573/26. Remarks: Brachysphingus
gibbosus Nelson, 1925,  Squires (1997a: p. 851). [Buccinidae].
Brachysphingus gibbosus Nelson, 1925. Nuttall and Cooper 1973. Bull    
Martinez
G12834/1. [Buccinidae].
Brachysphingus gibbosus
Fm., Paleocene. Meier canyon, south side Simi Valley, Simi Hills, Calabasas quad. (1952, PR 1967), Ventura Co.
LACMIP 22330. Hypotypes LACMIP 7924, 7925. [Buccinidae].

Holotype UCMP 30526, paratype UCMP 30527.

[=LACMIP 17086]. Hypotype LACMIP 7926.

  
1974), Los Angeles Co. LACMIP 21581. Hypotype LACMIP 7928.
        
Nacimiento, Lime Mtn. quad. (1948), San Luis Obispo Co. LACMIP 26526. Hypotype LACMIP 7563.
        
Hypotype LACMIP 7929.
Brachysphingus mammilatus
Paleocene. Near Benicia Arsenal, Solano Co. UCMP A4785. [Hypotype UCMP 32298]. [Buccinidae].
Brachysphingus mammilatus
Hot Springs Creek, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4662. Hypotype UCR 4662/5. [Buccinidae].
Brachysphingus mammilatus
Meganos Fm., Paleocene/Eocene. Deer Valley area, Antioch South quad. (1980), Contra Costa Co. LACMIP 7173.
Hypotypes LACMIP 7934, 7935. [Buccinidae].


Co. LACMIP 26456. Hypotype LACMIP 7937.

UCMP 3157. Holotype UCMP 31234.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 199
 Co. UCMP 3577 (not UCMP 3159 as in pl. caption). Para-
type UCMP 31235.

PR 1971), Fresno Co. LACMIP 7044. Hypotypes LACMIP 7938, 7939.
Brachysphingus mammilatus
Fm., Eocene. Bulldozer-generated exposure now under houses, Santa Susana quad. (1951, PR 1969), Ventura Co.
CSUN 958 [=LACMIP 40958]. Hypotype LACMIP 12685. [Buccinidae].
Brachysphingus sinuatus       

[1927]:392, ANSP 4258), [Paralectotypes (6) ANSP 79503]. [Buccinidae].
Brachysphingus sinuatus
Fm., Paleocene. Simi Hills, Ventura Co. UCR 6670. Hypotype UCR 6670/29. [Buccinidae].
Brachysphingus sinuatus-
cene. East of Lower Lake, Copsey Creek area, Lower Lake quad. (1975), Lake Co. UCMP A1523 [=LACMIP 7047].
Hypotypes UCMP 154001, 154002. [Buccinidae].

(=LACMIP 7051). Hypotypes LACMIP 7930–7932.
 
Costa Co. Holotype (of Buccinum liratum

22701. Hypotype LACMIP 7933.

Costa Co. Lectotype (of Stewart, 1926 [1927), p. 392), ANSP 4258, paralectotype ANSP 79503.
Brachysphingus-
ermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA 6525 [=LACMIP
26526]. Hypotype LACMIP 7563. Remarks: Brachysphingus gibbosus Nelson, 1925,  Squires (1997a: p. 851).
[Buccinidae].
Brachysphingus
Contra Costa Co. UCMP A1620. Hypotype UCMP number unknown. Remarks: Brachysphingus sinuatus Gabb,
1869b,  Squires (1997a: p. 855). [Buccinidae].
Bruclarkia barkeriana (Cooper, 1894). [Agasoma]. G D. Hanna and Hertlein 
19. Temblor Fm., Miocene. Hills just north of Kern River, Kern Co. CASG 65. Hypotype CASG 5741. [Buccinidae].
Bruclarkia barkeriana (Cooper, 1894). [Agasoma

Bruclarkia barkeriana (Cooper, 1894). [Agasoma
Mtn. Silt, Miocene. Caliente quad. (1914), Kern Co. UCMP B1638. Hypotype UCMP 32774. [Buccinidae].





200 PALEOBIOS, VOLUME 38, FEBRUARY 2021
-
potype USNM 650144.
Bruclarkia barkeriana (Cooper, 1894). [Agasoma-
terey Fm. (Saltos Shale Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M5026.
Hypotype USNM 254362. [Buccinidae].
Bruclarkia barkeriana (Cooper, 1894). [Agasoma-

La Panza Range, San Luis Obispo Co. USGS M5026. Hypotype USNM 254362. [Buccinidae].
Bruclarkia barkeriana forma santacruzana (Arnold, 1908). [Agasoma

Hypotype UCMP 36556. Remarks: Junior synonym of Bruclarkia barkeriana (Cooper, 1894),  Addicott (1970:
p. 88). [Buccinidae].
Bruclarkia barkeriana forma santacruzana (Arnold, 1908). [Agasoma]. Addicott 1970. USGS PP 642:88–89, pl. 10,

pl. caption). Hypotype UCMP 32806, 36556. Remarks: Junior synonym of Bruclarkia barkeriana (Cooper, 1894),
 Addicott (1970: p. 88). [Buccinidae].

Bruclarkia barkeriana forma santacruzana (Arnold, 1908). [Agasoma]. Addicott 1973b. USGS PP 791:7, 21, 22, pl.
Center quad. (1950), Kern Co.
UCMP. B1598. Hypotype UCMP 36556. Remarks: Junior synonym of Bruclarkia barkeriana (Cooper, 1894), 
Addicott (1970: p. 88). [Buccinidae].
Bruclarkia columbiana (Anderson and Martin, 1914) [Agasoma]. Addicott 1972b. SEPM Symposium vol., p. 6, pl. 1,

Kern Co. UCR 1106. Hypotype UCR 1106/1 (not UCR 1106 as in pl. caption). Remarks: Junior synonym of Bru-
clarkia gravida (Gabb, 1866),  Vendetti (2009: pp. 40, 44).[Buccinidae].
Bruclarkia columbiana (Anderson and Martin, 1914). [Agasoma
23. Temblor Fm. (Cymric Shale Member), Oligocene. Kern Co. UCR 1106. Hypotype UCR 1106/1 (not UCR 1106
as in pl. caption). Remarks: Junior synonym of Bruclarkia gravida (Gabb, 1866),  Vendetti (2009: pp. 40, 44).
[Buccinidae].
Bruclarkia columbiana (Anderson and Martin, 1914). [Agasoma]. Phillips et al. 1974. SEPM Guidebook, p. 123, pl. 3,

Remarks: Junior synonym of Bruclarkia gravida (Gabb, 1866),  Vendetti (2009: pp. 40, 44). [Buccinidae].
Bruclarkia columbiana (Anderson and Martin 1914). [Agasoma-
hagen Fm., Oligocene. Monocline RidgeMonocline Ridge quad., Fresno Co. UCMP D4272]. Hypotype UCMP 37169.
Remarks: Junior synonym of Bruclarkia gravida (Gabb, 1866),  Vendetti (2009: pp. 40, 44). [Buccinidae].
Bruclarkia gravida (Gabb, 1866). [Clavella


Bruclarkia gravida (Gabb, 1866). [Clavella-
Clavella gravida)). San Ramon Fm., Miocene. Martinez, Contra Costa Co.

Bruclarkia gravida (Gabb, 1866). [Clavella
Near Muir Station, Contra Costa Co. UCMP A4661. Hypotype UCMP. 32426]. [Buccinidae].
Bruclarkia gravida (Gabb, 1866). [Clavella
Fm., Miocene. Martinez, Contra Costa Co. Lectotype (of Stewart 1926 [1927]:398) ANSP 4345, hypotype ANSP.
4345a (invalid number,  K. Estes-Smargianni, personal communication, 2019). [Buccinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 201
Bruclarkia oregonensis (Conrad, 1848). [Fusus
Near junction of Norris Creek with Crow Creek, Hayward quad., Alameda Co. UCMP A4565. Hypotypes UCMP
34162, 34163. [Buccinidae].
Bruclarkia oregonensis (Conrad, 1848). [Fusus]. Hall 1958. UCPG 34(1 hy-
potype SU 8479). Oursan Sandstone, Miocene. Northwest of Pleasanton, Dublin quad. (1943), Alameda Co. SU
3229. Hypotype SU 8479 [=CASG 69956]. [Buccinidae].
Bruclarkia oregonensis (Conrad, 1848). [Fusus
Remarks: Bruclarkia sp. aff.
Bruclarkia barkeriana (Cooper, 1894), 

650145.
Bruclarkia oregonensis (Conrad, 1848). [Fusus-
blor Fm. (Buttonbed Sand Member) (of Stinemeyer et al. 1959:8, 13), Miocene. Shale Pt. quad., Temblor Range,
Kern Co. USGS M3986. Hypotypes USNM 646804, 646805. [Buccinidae].
Bruclarkia oregonensis (Conrad, 1848). [Fusus
San Juan Creek area, Orange Co. [Hypotype at CC]. [Buccinidae].
Bruclarkia oregonensis yaquinana (Anderson and Martin, 1914). [Agasoma
15, 16. Temblor Fm., Miocene. Reef Ridge quad., Kings Co. UCMP D1056. Hypotype UCMP 36871. [Buccinidae].
Bruclarkia seattlensisPhacoi-
des Sand member) (of Stinemeyer et al. 1959:12–13), Miocene. Las Yeguas Ranch quad., Temblor Range, Kern
Co. USGS M3280. Hypotype USNM 646616. Remarks: Junior synonym of Bruclarkia acuminata (Anderson and
Martin, 1914),  Vendetti (2009: pp. 9, 40). [Buccinidae].

Bruclarkia seattlensis-
stone Member), Miocene. Zemorra Creek area, Carneros Rocks quad., Kern Co. USGS M3979. Hypotype USNM
646613. Remarks: Junior synonym of Bruclarkia acuminata (Anderson and Martin, 1914),  Vendetti (2009:
pp. 9, 40). [Buccinidae].

quad., Kern Co. USGS M3772. Hypotypes USNM 646614, 646615. [Buccinidae].

USGS M3280. Hypotype USNM 646616.
Bruclarkia seattlensis
(lower sandstone member), Oligocene. The Indians, Junipero Serra quad., Santa Lucia Range, Monterey Co. SU
1756. Hypotypes CASG 60365, 60372 determined here as missing. Remarks: Junior synonym of Bruclarkia
acuminata (Anderson and Martin, 1914),  Vendetti (2009: pp. 9, 40). [Buccinidae].
, 15. Approximately 1 km southeast of Indian Ranger Station, Monterey Co. USGS M6557. Hypotypes
USNM 264140, 264142, determined here as missing.
Bruclarkia yaquinana (Anderson and Martin, 1914). [Agasoma 
8, 12, 13. Basal Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotypes USNM
650149, 650151. Remarks: Junior synonym of Bruclarkia acuminata (Anderson and Martin, 1914),  Vendetti
(2009: pp. 9, 40). [Buccinidae].
Bruclarkia
Co. UCMP A4929. Hypotype UCMP 34164. Remarks: Bruclarkia oregonensis (Conrad, 1848),  Addicott (1970:
p. 90). [Buccinidae].
202 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Bruclarkia          
Miocene]. Santa Monica Mtns., Los Angeles Co. Hypotype SAMO 3204. Remarks: Determined here as Bruclarkia
barkeriana (Cooper, 1894). [Buccinidae].
Indeterminate Bruclarkia Powell et al. 2019b. PaleoBios
excavation, Berkeley Hills, Oakland, Alameda Co. UCMP IP13004. Hypotype UCMP 412181. [Muricidae].

Pliocene. Fish Creek/Vallecito Creek, San Diego Co. [ABDSP 5081.2. Hypotype ABDSP I 11151]. Remarks: Pos-
sibly in the genus Solenosteira. [Buccinidae].


 C. Throckmorton (personal communication, 2002). [Buccinidae].
Bulla cantuaensis
Silt, Miocene. Caliente Quad., Kern Co. SU 2121. Hypotype SU/CASG. 9941 determined here as missing. [Bullidae].
Bulla cantuaensis       
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2859. Hypotype USNM
254372. [Bullidae].
Bulla cantuaensis

Obispo Co. USGS M2859. Hypotype USNM 254372. [Bullidae].
Bulla sp. Redman et al. 2007. Palaios-
yon, Imperial Co. Hypotype probably at SDSU, current whereabouts unknown. [Bullidae]
Bullia (Buccinanops clarki Wagner and Schilling, 1923. Allmon 1990. BAP 99(335):
Fm., Oligocene. San Emigdio Grant, Mt. Pinos quad., Kern Co. UCMP 3182. Holotype UCMP 11430. [Nassariidae].
Bullina
quad. (1954), Kern Co. USGS M1612. Hypotype USNM 650336. [Aplustridae].
Calicantharus cf.  (Clark and Arnold, 1923). [Searlesia
Temblor Fm. (Wygal Sandstone Member), Miocene. Carneros Creek area, Carneros Rocks quad., Kern Co. USGS
M4471. Hypotype USNM 646622. Remarks: In the genus Lirabuccinum,  Vermeij (1991: p. 269). [Buccinidae].
Calicantharus dalli (Clark, 1918). [Searlesia
Sandstone Member), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3772. Hypotype USNM
646628. Remarks: In the genus Lirabuccinum
Calicantharus cf.  (Carpenter, 1866). [Pisania
8 (cast). Pancho Rico Fm., Miocene/Pliocene. Base of bluff on south side of Pancho Rico Creek, San Ardo quad.,
Monterey Co. USGS M1939. Hypotype USNM 649106. Remarks: In the genus Pusio,  Vermeij (2001: p. 296).
[Buccinidae].
Calicantharus fortis (Carpenter, 1866 [not 1864 as cited]). [Pisania
Pico Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14398. Remarks: In the
genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Calicantharus fortis (Carpenter, 1866 [not 1864 as cited]). [Pisania
3. Reworked shells, Pliocene. Shells found in a modern-day shrimp trap off San Clemente Is., California Channel
Ids. CASG 72573. Hypotype CASG 72573. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Calicantharus fortis var. cf. angulata (Arnold, 1907). [Pisania]. Woodring and Bramlette 1950 [1951]. USGS PP 222:75,

Hypotype CIT 4453 [=LACMIP 4194]. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 203

Calicantharus cf.  (Arnold, 1907). [Pisania]. Durham and Addicott 1965. USGS PP 524-A:A13, pl.

M980. Hypotype USNM 649102. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Calicantharus humerosus (Gabb, 1869a). [Neptunea
Elsmere Canyon, Los Angeles Co. UCLA 242 [=LACMIP 20242]. Hypotype UCLA 49524 [=LACMIP 10623]. Re-
marks: In the genus Pusio,  Vermeij (2001: p. 296). Ex Grouard collection. [Buccinidae].
Calicantharus humerosus (Gabb, 1869a). [Neptunea     
Pliocene. Orange, Co. SDSNH 4080]. Remarks: SDSNH hypotype determined here as missing. In the genus Pusio,
 Vermeij (2001: p. 296).
Calicantharus humerosus (Gabb, 1869a). [Neptunea]. Squires 2012. NHMLAC CS
Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14397. Remarks: In the genus Pusio,  Vermeij
(2001: p. 296). [Buccinidae].
Calicantharus kernensis (Anderson and Martin, 1914). [Chrysodomus]. Addicott 1970. USGS PP 642:92–93, pl. 7,
      
Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].

Calicantharus kettlemanensis (Arnold, 1909 [1910]). [Thais]. Durham and Addicott 1965. USGS PP 524-A:A13, pl. 5,

M1929. Hypotypes USNM 649100, 649101. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Calicantharus cf. kettlemanensis (Arnold, 1909 [1910]). [Thais
Round Mtn. Silt, Miocene. North Bank of Kern River, Kern Co. USGS 6063. Hypotype USNM 650162. Remarks: In
the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Calicantharus kettlemanensis (Arnold 1909, [1910]). [Thais
3. Panorama Hills Fm. (of Dibblee 1968), Pliocene. Near Panorama Pt. Panorama Hills quad., Temblor Range, Kern
Co. USGS 12876. Hypotype USNM 646813. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].

Calicantharus portolaensis (Arnold, 1908). [Fusus]. Woodring and Bramlette 1950 [1951]. USGS PP 222:75, pl. 14,
   
14771. Hypotype USNM 560080. Remarks: In the genus Lirabuccinum,  Vermeij (1991: p. 269). Remarks:
Remarks: The age of this member is Pliocene/Pleistocene, 
Calicantharus rancherianus-
ceneRio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Holotype USNM 650156, hypotypes USNM 650154,
650155. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Calicantharus woodfordi
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650161, paratype USNM 650146.
Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Californiconus californicus
Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14404. [Conidae].
Califostoma rotundivaricosum     
Diego Fm., Pliocene. El Rancho Del Rey, San Diego Co. SDSNH 4411. Holotype SDSNH.137443, paratypes SD-
SNH.137444, 137445. [Muricidae].
SDSNH 137446, 137448.
204 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Califostoma turris (Nomland, 1916a). [Purpura-
chegoin Fm., Pliocene. West of Coalinga, Fresno Co. UCMP 2110. Holotype UCMP 12049. [Muricidae].


Callianax baetica (Carpenter, 1864b). [Oliva
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14403. Remarks: Junior synonym of Calianax alectona
(Duclos, 1835),  Powell et al. (2020: p. 5). [Olividae].
Calliostoma carsoni
Rio Bravo Ranch quad., Kern Co. UCLA 3456 [=LACMIP 23456]. Holotype UCLA 45763 [=LACMIP 8429]. [Cal-
liostomatidae].
Calliostoma coalingense
Joaquin Fm. (Acila Zone), Pliocene. Arroyo Vadoso, North Dome, Kettleman Hills, Kings Co. USGS 12330. Hypotype
USNM 495710. [Calliostomatidae].
Pecten Zone), Pliocene. Arroyo Torcido area, North Dome, Kettleman Hills,
Kings Co. USGS 12334. Hypotype USNM 495768.
Calliostoma coalingense
cast). Pancho Rico Fm., Miocene/Pliocene. Indian Valley, Valleton quad., Monterey Co. USGS M1930. Hypotype
USNM 649104. [Calliostomatidae].

USNM 649105.
Calliostoma coalingense catoteron Woodring in Woodring and Bramlette, 1950 [1951] (n. var.). USGS PP 222:69,

Holotype USNM 560061. [Calliostomatidae].
Calliostoma coalingense privum Stewart in
17. Etchegoin Fm., Pliocene, Coalinga quad., Fresno Co. UCMP B6546. Hypotype UCMP 36801. [Calliostomatidae].
Calliostoma gemmulatum
Unnamed strata, Pliocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Is. USGS M6501.
Hypotype USNM 235583. [Calliostomatidae].
Calliostoma cf.  Carpenter, 1864b. Woodring et al. 1940 [1941
San Joaquin Fm. (Pecten Zone), Pliocene. La Lomica, North Dome, Kettleman Hills, Kings Co. USGS 12365a [=USGS
12768). Hypotypes USNM 495774, 495775. [Calliostomatidae].
Calliostoma kerri
Fm. (Pecten zone), Pliocene. Vicinity of Arroyo Pequeño and Arroyo Finito, North Dome, Kettleman Hills, Kings
Co. USGS 12343. Hypotype USNM 495773. [Calliostomatidae].
Calliostoma olssoni
Pliocene. Vicinity of Whitewater River, San Jacinto quad., Riverside Co. UCMP A1407. Hypotype UCMP 16086.
[Calliostomatidae].

cited).
Calliostoma platinum       
found in a modern-day shrimp trap off San Clemente Is., California Channel Ids. CASG 72574. Hypotype CASG
72574. Remarks: In the genus Akoya,  Tuskes and McGowan-Tuskes (2019: pp. 21–23). [Calliostomatidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 205
Calliostoma sp., aff. 
Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14377. [Calliostomatidae].
Calliostoma sp., cf. splendens
Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14376. [Calliostomatidae]. Remarks: Synonym
of Calliostoma supragranosum,  Tuskes and McGowan-Tuskes (2019: p. 20). [Calliostomatidae].
Calliostoma
CASG 37468. Hypotype CASG 12930. [Calliostomatidae].
Calliostoma
(1954), Kern Co. USGS M1591. Hypotype USNM 650335. [Calliostomatidae].
Calliovarica eocensis

quad., Fresno Co. UCMP 1817. Holotype UCMP 15884]. [Chilodontidae].
Calorebama dilleri kirbyi (Clark, 1938). [Pseudoliva
Eocene. Grapevine Canyon, Grapevine quad., Kern Co. UCLA 2340 [=LACMIP 22340]. Hypotype LACMIP 7866 [ex
UCLA 43896]. Remarks: Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p. 1) for com-
ments about Calorebama and Sulcobuccinum. [Pseudolividae].

38211.
Calorebama dilleri lineata (Gabb, 1864). [Pseudoliva]. Squires 1989a. Jour. 
of Martinez, Contra Costa Co. Lectotype (of Stewart 1926 [1927]:400) ANSP. 4200 (not
paralectotypes (9) ANSP 79424]. Remarks: Sulcobuccinum linea-
tum,  Vermeij (1998: p. 81); see Squires (2001: p. 10) for comments about Calorebama and Sulcobuccinum.
[Pseudolividae].

Calorebama dilleri lineata (Gabb, 1864). [Pseudoliva
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype
LACMIP 11470. Remarks: Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p. 10) for com-
ments about Calorebama and Sulcobuccinum. [Pseudolividae].
Calorebama inornata (Dickerson, 1915). [Pseudoliva-
jon Fm., Eocene. Grapevine Canyon, Grapevine quad., Kern Co. UCMP A971. Hypotype UCMP 38212. Remarks:
Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p. 10) for comments about Calorebama
and Sulcobuccinum. [Pseudolividae].
Calorebama inornata (Dickerson, 1915). [Pseudoliva
Matilija Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 24260. Hypotype LACMIP 11628. Remarks:
Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p. 10) for comments about Calorebama
and Sulcobuccinum. [Pseudolividae].
Calorebama inornata (Dickerson, 1915). [Pseudoliva-
cene. North side of Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 7445. Hypotype LACMIP
12737. Remarks: Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p. 10) for comments
about Calorebama and Sulcobuccinum. [Pseudolividae].
Calorebama volutaeformis (Gabb, 1864). [Pseudoliva
Fm., Eocene. Grapevine Canyon area, Kern Co. Lectotype (of Stewart 1926[1927]:400) ANSP 4201 (not CASG
Remarks: Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p.
10) for comments about Calorebama and Sulcobuccinum. [Pseudolividae].
Calorebama volutaeformis (Gabb, 1864). [Pseudoliva
Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23710. Hypotype LACMIP 11655. [Pseudolividae].
206 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Remarks: Sulcobuccinum lineatum,  Vermeij (1998: p. 81); see Squires (2001: p. 10) for comments about
Calorebama and Sulcobuccinum. [Pseudolividae].
Calyptraea coreyi
Ranch quad. (1954), Kern Co. USGS M1591. Holotype USNM 650085. [Calyptraeidae].
 Jewett Sand, Miocene. Caliente quad. (1914), Kern Co. UCMP B1665. Paratype UCMP 15452.
Calyptraea coreyi
Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP22634 [=UCLA 2634). Hypotype LACMIP 13073
[=UCLA 57622]. [Calyptraeidae].
Calyptraea cf. costellata (Conrad, 1857d). [Trochita
Canyon, Los Angeles Co. UCLA 2634 [=LACMIP 22634. Hypotype UCLA 49576 [=LACMIP 10615]. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita  
-
type (of Stewart, 1926 [1927]: p. 340, of Galerus excentricus
(12) ANSP 79473]. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita
Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS 14494. Hypotype USNM 498706. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita -
ferentiated Sacate/Gaviota fms., Eocene. Nojoqui Creek area, Lompoc quad., Santa Barbara Co. UCMP B6954.
Hypotype SU 9195 [=CASG 70140]. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita
Fm. (Wygal Sandstone Member), Miocene. Media Agua Creek area, Las Yeguas Ranch quad., Kern Co. USGS M3280.
Hypotype USNM 646607. [Calyptraeidae].

Calyptraea diegoana (Conrad, 1855b). [Trochita
Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hy-
potype SDSNH 20776. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita     
Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115].
Hypotype LACMIP 6519. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita-
ker Peak quad. (1958), Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype LACMIP 7461. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 983 [=LACMIP 16110]. Hypotype LACMIP 7700.
[Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita]. Squires et al. 1988. SEPM 
Eocene. El Paso Mtns., Inyokern SE quad. (1972), Kern Co. USGS M9067. Hypotype USNM 424891 determined
here as missing. Remarks: Hypotype never received from authors,  USNM label. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype
LACMIP 11462. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita
Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 24260. Hypotype LACMIP 11627. [Calyptraeidae].

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 207
11648.
Calyptraea diegoana (Conrad, 1855b). [Trochita  
Eocene. North side of Simi Valley, bulldozer-generated exposure now under houses, Santa Susana quad. (1951,
PR 1969), Ventura Co. CSUN 958 [=LACMIP 17190]. Hypotype LACMIP 12682. [Calyptraeidae].
Calyptraea diegoana (Conrad, 1855b). [Trochita
Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13419. [Calyptraeidae].
Calyptraea diegoana
Grapevine Canyon, Tejon quad., Kern Co. LACMIP 22340. Hypotype LACMIP 13458. [Calyptraeidae].
Calyptraea fastigiata
Member), Pliocene. Lobitos Creek area, Half Moon Bay quad., San Mateo Co. locality F692M. Hypotype determined
here as missing. [Calyptraeidae].
Calyptraea fastigiata
Santa Cruz Co. CASG 66087. Hypotype CASG 66087.01. [Calyptraeidae].
Calyptraea cf.  (Gabb, 1866). [Trochita
Fm. (Upper Mya Zone), Pliocene. Las Alturas, Middle Dome, Kettleman Hills, Kings Co. USGS 12685. Hypotype
USNM 495291. Remarks:  (Gabb,1866),  Addicott (1970: p. 60). [Calyptraeidae].
 (Gabb, 1866). [Trochita-
vington, La Costa Valley quad. (1943), Alameda Co. SU 3252. Hypotype SU 8468 [=CASG. 69957]. [Calyptraeidae].
 (Gabb, 1866). [Trochita        
Miocene. North of Kern River, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1604. Hypotype USNM 650086.
[Calyptraeidae].

. UCMP B1638 (not UCMP B1683 as in pl. caption). Hypotype
UCMP 15453.
Calyptraea cf.  (Gabb, 1869a). [Trochita
Joaquin Fm. (Acila Zone), Pliocene. North side of Arroyo Hondo, North Dome, , Kettleman Hills, Kings Co. USGS
12340. Hypotype USNM 495714. Remarks: Calyptraea inornata (Gabb, 1869a),  Adegoke (1969: p. 164).
[Calyptraeidae].
Pecten Zone), Pliocene. West of El Portillo, Middle Dome, Kettleman Hills, Kings
Co, USGS 12385. Hypotype USNM 495763. Remarks: May be a worn specimen of  cf.  (Gabb 1866), 
Calyptraea inornata (Gabb 1869a), Adegoke (1969: p. 164).
Neverita Zone), Pliocene. Arroyo Torcido and Arroyo Hondo area, North Dome,
Kettleman Hills, Kings Co. USGS 12513. Hypotype USNM 495324. Remarks: Calyptraea inornata (Gabb, 1869a),
 Adegoke (1969: p. 164).
Calyptraea cf. martini
Contra Costa Co. UCMP A2569. Hypotype UCMP 30672. [Calyptraeidae].
Calyptraea
Remarks: Determined here as Calyptraea diegoana (Conrad, 1855b). [Calyptraeidae].
Calyptraea-
stone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312.
Hypotype SDSNH 85244 (error) [corrected as SDSNH 136498 in errata, p. 50]. [Calyptraeidae].
Calyptraea (Trochita
Co. LACMIP 17918. Hypotype LACMIP 14384. [Calyptraeidae].
208 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Campanile dilloni (G D. Hanna and Hertlein, 1949). [Campanilopa]. Squires 1991c. SEPM Guidebook 68:220, pl. 1,
  
16155]. Hypotype LACMIP 7165. [Campanilidae].
Campanile dilloni (G D. Hanna and Hertlein, 1949). [Campanilopa
7. Santa Susana Fm., Paleocene/Eocene. Bus Canyon, Thousand Oaks quad. (1950, PR 1967), Ventura Co. CSUN
1565 [=LACMIP 16505]. Hypotype LACMIP 12235. [Campanilidae].

16199]. Hypotype LACMIP 12236.

San Marcos Pass quad. (1959, PR 1988), Barbara Co. CSUN. 1566 [= LACMIP 16507]. Hypotype LACMIP 12237.
     
16506]. Hypotype LACMIP 12238.
Campanile dilloni (G D. Hanna and Hertlein, 1949). [Campanilopa
Santa Susana Fm., Eocene. Bus Canyon, Thousand Oaks quad. (1950, PR1967), Ventura Co. CSUN 1565 [=LACMIP
16505]. Hypotype LACMIP 12335. [Campanilidae].
Campanile greenellum
Paleocene. Trailer Canyon, Santa Monica Mtns., Topanga quad. (1952, PR 1967), Los Angeles Co. LACMIP 24433.
Hypotype LACMIP 12232. [Campanilidae].
      
Hypotype LACMIP 12233.

Angeles Co. LACMIP 24716. Hypotype LACMIP 12234.
Campanile greenellum
German Rancho Fm., Paleocene. Gualala, Mendocino Co. Wentworth W730 [=UCMP W730]. Hypotype UCMP
39915. [Campanilidae].
Campanile            
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 837 [=LACMIP 16156]. Hypotypes LACMIP 7459, 7460.
[Campanilidae].
Campanilopa dilloni
Domengine Fm., Eocene Media Agua Creek, Kern Co. CASG 32388A [=CASG 30677]. Holotype CASG 9425 [=CASG
Remarks:
In the genus Campanile,  Squires (1993a: p. 327). [Campanilidae].
Campanilopa dilloni
Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4668. Hypotype UCR 4668/101. Remarks:
In the genus Campanile,  Squires (1993a: p. 327). [Campanilidae].
Campanilopa dilloni
2.1. Maniobra Fm., Eocene. Northeast Orocopia Mtns., Canyon Spring SW quad. (1963), Riverside Co. CSUN 668
[=LACMIP 16155]. Hypotype LACMIP 7165. Remarks: In the genus Campanile,  Squires (1993a: p. 327).
[Campanilidae].
Cancellaria altispira
Angeles Co. LACMIP 17934. Hypotype LACMIP 14410. [Cancellariidae].
Cancellaria arnoldi-
stone, Pliocene. Santa Barbara Co. USGS 4473. Hypotype USNM 560049. [Cancellariidae].
  
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 209
14648. Hypotype USNM 560085.

Cancellaria arnoldi
Palo Alto quad., Santa Clara Co. USGS M1715. Hypotypes USNM 650955–650957. [Cancellariidae].
Cancellaria arnoldi
Santa Cruz Co. Hypotype determined here as missing. [Cancellariidae].
Cancellaria arnoldi
Cruz Co. Hypotype determined here as missing. [Cancellariidae].
Cancellaria barkeri (Anderson in G D. Hanna, 1924). [Fossarus
Fm., Miocene. Garza Creek, Reef Ridge quad., Kings Co. UCMP D1061. Hypotype UCMP 36894. [Cancellariidae].
Cancellaria (Narona) birchi
Mtn. Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Hypotypes USNM 650215, 650216.
[Cancellariidae].

Cancellaria (Euclia) circumspinosa
Miocene. Kern River, Kern Co. USGS 3886. Hypotype USNM 650186. [Cancellariidae].
    
Holotype USNM 650187.
-
potype UCMP 33480.

Cancellaria (Euclia) condoni
Mtn. Silt, Miocene. Caliente quad. (1914), Kern Co. UCMP B1618. Hypotype UCMP 33019. [Cancellariidae].

650182.

UCMP 33349.

Cancellaria cf.  Anderson, 1905. Woodring et peel). Monterey
Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, Palos Verdes Hills, Los Angeles
Co. USGS 13127. Hypotype USNM 497076. Remarks: Cancellaria (Euclia) oregonensis Conrad, 1849a,  Addicott
(1970: p. 111). [Cancellariidae].
Cancellaria condoni
quad., Fresno Co. UCMP D1085. Hypotype UCMP 36889. [Cancellariidae].
Cancellaria crassa-
cene. Simi Hills, Ventura Co. UCR. 6670. Hypotype UCR. 6670/12 [same number as Acrilla (Ferminoscala) sp. [=A.
() durhami Zinsmeister, 1983b] determined here as missing. Remarks: Cancellaria simiana G D. Hanna, 1924:
p. 160, new name for Cancellaria crassa Waring, 1917, preocupied. [Cancellariidae].
Cancellaria dalli (Anderson and Martin, 1914). [Fossarus
Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650202. Remarks:
Cancellaria barkeri Anderson in G D. Hanna, 1924, new name for Cancellaria dalli (Anderson and Martin, 1914),
210 PALEOBIOS, VOLUME 38, FEBRUARY 2021
preoccupied. [Cancellariidae].
-
potype USNM 650203.


Cancellaria dalliana            
quad. (1950), Kern Co. UCMP B1598. Hypotype UCMP 36558. Remarks: In the subgenus Euclia,  Addicott
(1970: p. 108). [Cancellariidae].
Cancellaria (Euclia) dalliana
Upper Olcese Sand, Miocene. Oil Center quad. (1950), Kern Co. UCMP B1598. Hypotypes UCMP 33483, 36558,
33476. [Cancellariidae].


Cancellaria dalliana-
potype possibly at SDSNH, current whereabouts unknown. Remarks: Ex Grouard collection. [Cancellariidae].
Cancellaria galei
Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650212, paratype USNM 650213. [Cancel-
lariidae].

Co. USGS M1590. Holotype USNM 650214.
Cancellaria hamlini. 96. Pico Fm., Pliocene. Newhall area,
Los Angeles Co. LACMIP. 17919. Hypotype LACMIP 14413. [Cancellariidae].
Cancellaria hemphilli
Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. Alex Clark locality. Hypotypes (2) USNM 560086.
[Cancellariidae].
Cancellaria hemphilli
Ranch quad., Fresno Co. UCMP B6529. Hypotype UCMP 36896. [Cancellariidae].
Cancellaria hemphilli 
Angeles Co. LACMIP 7757. Hypotype LACMIP 14411. [Cancellariidae].
Cancellaria irelaniana
Narona)). Capay Fm., Eocene. Marysville Buttes, Sutter Co. CSMB 13738. Holotype CASG 10.
[Cancellariidae].
Cancellaria (Euclia) joaquinensis
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1601. Hypotype USNM 650197. [Cancellariidae].


UCMP 33738.

Cancellaria keenae
River (presumably from the same stratigraphic interval and geographic area as USGS localities M1608, M1612,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 211
and M1613, which are on the south side of Kern River, Rio Bravo Ranch quad.), Kern Co. UCLA AC-2-34 [=LACMIP
16157]. Holotype UCLA 45771 [=LACMIP 8430]. [Cancellariidae].


Cancellaria (Crawfordina) kernensis    
Round Mtn., Miocene. Kern River, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1608. Holotype USNM 650207.
[Cancellariidae].
  
database) CASG 12936 [=CASG 28790].
Cancellaria (Pyruclia) lickana Anderson and Martin, 1914.
Upper Olcese Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1599. Hypotype USNM 650198.
[Cancellariidae].


Cancellaria lipara Woodring in  
14. Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. Alex Clark locality. Holotype USNM.
560082. [Cancellariidae].
Cancellaria nevadensis-
cene. Garza Creek, Reef Ridge quad., Kings Co. UCMP D1056. Hypotype UCMP 36890. [Cancellariidae].
Cancellaria (Euclia nevadensis
Lower Round Mtn. Silt, Miocene. Kern River, Kern Co. USGS 6641. Hypotype USNM 650200. [Cancellariidae].


Cancellaria obesa-
trania Fm., Miocene/Pliocene. Coyote Mtn., Imperial Co. SDSNH 2895. Hypotype SDSNH 12142. [Cancellariidae].
Cancellaria (Euclia) ocoyana
Miocene. Woody quad. (1952), Kern Co. USGS M1593. Hypotype USNM 650189. [Cancellariidae].

UCMP 33484.

33485.

Cancellaria (Euclia) oregonensis
Miocene. Kern River, Kern Co. CASG 198. Neotype (of Keen and Bentson 1944: p. 138, rejected by Addicott 1970:
   Cancellaria dalliana Anderson,
1905) CASG 198. [Cancellariidae].

Cancellaria condoni Anderson” 1905, CASG 65.01.
Cancellaria (Euclia) oregonensis
Sand, Miocene. Caliente quad. (1914), Kern Co. UCMP B1641. Hypotype UCMP 33577. [Cancellariidae].
212 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Cancellaria cf.  Clark. Briones Fm., Miocene. South of
Pleasanton, Niles quad. (1943), Alameda Co. SU 3232. Hypotype SU 8480 [=CASG 69958]. Remarks: Cancellaria
pabloensis Clark, 1915,  Adegoke (1969: p. 190). [Cancellariidae].
Cancellaria (Euclia)
Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1600. Neotype (of Addicott 1970:114) USNM.
650205. [Cancellariidae].



Cancellaria posunculensis  
63-5. Temblor Fm., Miocene. Near Kern River, Kern Co. CASG 65. Hypotype CASG 5725. [Cancellariidae].
Cancellaria posunculensis
Miocene. Crow Canyon Hayward quad., Alameda Co. UCMP A4565. Hypotype UCMP 34168. Remarks: Possibly
Antilliophos posunculensis (Anderson and Martin, 1914),  Addicott (1970: p. 116). [Cancellariidae].
Cancellaria (Coptostoma) posunculensis Anderson and Martin, 1914. Addicott 1970. USGS PP 642:116–117, pl. 15,

33811. [Cancellariidae].


Cancellaria posunculensis
Monterey Fm. (Saltos Shale Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS
M2868. Hypotype USNM 254367. [Cancellariidae].
Cancellaria posunculensis

area, La Panza Range, San Luis Obispo Co. USGS M2868. Hypotype USNM 254367. [Cancellariidae].
Cancellaria rapa
(Pecten Zone), Pliocene. Vicinity of Arroyo Vadoso and Arroyo Largo, North Dome, Kettleman Hills, Kings Co.
USGS 12331. Hypotype USNM 495755. [Cancellariidae].
Cancellaria rapa perrini Carson, 1926. Woodring and Bramlette 1950 [1951]. USGS PP 222: 77, pl
Careaga Fm. (Graciosa Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14618. Hypotypes
USNM 560123, 560124. [Cancellariidae].

Cancellaria (Barkeria) sanjosei
Upper Round Mtn. Silt, Miocene. Cottonwood Creek area, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1605.
Hypotypes USNM 650210, 650211. [Cancellariidae].

Cancellaria (Euclia) simplex
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotypes USNM 650191, 650192. [Cancellariidae].


GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 213

Cancellaria (Euclia) simplex
19, 27. Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. Upper Olcese, Miocene. USGS M1602. Hypotypes
USNM 650193, 650194. [Cancellariidae].

Hypotype USNM 650195.
Cancellaria (Euclia) simplex
Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1602. Hypotype USNM 650196. [Cancellariidae].
Cancellaria tritonidea 
Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. USGS 14769. Hypotype USNM 560083.
Remarks: Cancellaria tritonidea Gabb, 1866,  Adegoke (1969: p. 191). [Cancellariidae].
Cancellaria tritonidea
Miocene/Pliocene. King City quad., Monterey Co. UCMP B4927 (not UCMP B4926 as in pl. caption). Hypotype
UCMP 36470. [Cancellariidae].
Cancellaria tritonideaPolinices reclusianus).
-
potypes current whereabouts unknown. [Cancellariidae].
Cancellaria tritonidea   . Purisima Fm., Miocene/ Pliocene. Capitola,
Santa Cruz Co. Hypotype CASG 66147.01. [Cancellariidae].
Cancellaria tritonidea
Angeles Co. LACMIP 7752. Hypotype LACMIP 14412. [Cancellariidae].
Cancellaria tritonidea altispira       

Angeles Co. LACMIP 1219. Hypotypes (2) current whereabouts unknown. [Cancellariidae].
Cancellaria
Clara quad., Santa Clara Co. USGS M1715. Hypotype USNM 650960. [Cancellariidae].
Cancellaria        

lost,  C. Throckmorton (personal communication, 2002). [Cancellariidae].
Cantharus fortis (Carpenter, 1866). [Pisania
Remarks: In the genus
Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Cantharus fortis (Carpenter, 1866). [Pisania      
source specimen unknown. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Cantharus aff. 
Shells found in a modern-day shrimp trap off San Clemente Is., California Channel Ids. CASG 72573. Hypotype
CASG 72575.
Cantharus resembling Gemophos ringens (Reeve, 1846). [Buccinum
2-2. Deguynos Fm. (Jackson Creek Member), Pliocene. Fish Creek/Vallecito Creek, San Diego Co. ABDSP 5070.1.
Hypotype ABDSP I 11115]. [Buccinidae].
Cancellaria [in text] with Polinices
reclusianus
1219. Hypotype current whereabouts unknown. Remarks: Determined here as Boreotrophon sp. [Muricidae].
214 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Cassis subtuberosa-
rial Fm. [=Latrania Fm.], Miocene/Pliocene. Alverson Canyon, Coyote Mtn., Imperial Co. Holotype UCMP 31275.
[Cassidae].
Cassis subtuberosa*-
drade Member), Miocene. Alverson Canyon, Imperial Co. ABDSP 5008.2. Hypotype ABDSP I 9608]. Remarks:
Error for subtuberosus. [Cassidae].
Cassis (Coalingodea) tuberculata (Gabb, 1864). [Morio]. Abbott 1968:59–60, pl. 34. Domengine Fm., Eocene. Par-
son's Peak, Coalinga, 
[UCMP 11783] (middle Remarks: Galeodea tuberculiformis G.D. Hanna, 1924,  (Squires
2019: p. 11). [Cassidae].
Ceratostoma delorae

SU 3245. Holotype SU 8473 [=CASG 69945]. [Muricidae].
Ceratostoma delorae-
cene. East of Irvington, Alameda Co. SU 3245 [=CASG 69945]. Holotype SU 8473 [=CASG 69945.00]. [Muricidae].
Ceratostoma foliatum (Gmelin, 1791). [Purpura
Pancho Rico Fm., Miocene/Pliocene. Long Valley, Monterey Co. Hypotype SU 9754 [=CASG 70322]. [Muricidae].
Ceratostoma nannum (Nomland, 1917b). [Pupura
Margarita Fm., Miocene. North of Coalinga, Domengine Ranch quad., Kings Co. UCMP 2279]. Holotype UCMP
11315. [Muricidae].
Ceratostoma perangulatum (Nomland, 1916b). [Murex
Jacalitos Fm., Pliocene [Miocene]. West of Coalinga, Kings Co. UCMP 2649. Holotype UCMP 12057. [Muricidae].
Ceratostoma (Dall, 1909). [Purpura
Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650120. [Muricidae].
Cerithiopsis llajasensis
area, Santa Susana quad. (section cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115]. Holotype
LACMIP 6515, paratype LACMIP 6516 (not LACMIP 6517 and 6518 as in pl. caption). [Cerithiopsidae].
Cerithiopsis aff.  
-
thiopsidae].
Cerithium cliffensis  
quad. (1943), Ventura Co. UCR 4703. Hypotype UCR 4703/30. [Cerithiidae].
Cerithium vaderensis (Dickerson, 1915). [Melania
Fm., Eocene. Cañada del Corral, Los Olivos quad., Santa Barbara Co. UCMP B6951. Hypotype SU 9193 [=CASG
70144]. [Cerithiidae].
         
Carneros Rocks quad., Kern Co. USGS M3578. Hypotype USNM 646603. [Cerithiidae].
Charonia. Miocene/Pliocene, Imperial Fm. [=Latrania Fm.]. Fossil Canyon
[=Alverson Canyon], Coyote Mtn.], Imperial Co. LACMIP 17740. Hypotype LACMIP 8188]. [Ranellidae].
Chedevillia saltonensis  
Fm., Eocene. Northeastern Orocopia Mtns., Canyon Spring NW quad. (1963), Riverside Co. CSUN 662 [=UCLA

10565]. [Strombidae].
Chedevillia saltonensis
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 215
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 830 [=LACMIP 16166]. Hypotype LACMIP 7464. [Strombidae].
Chedevillia saltonensis

28987 [=LACMIP 10564]. [Strombidae].
Chedevillia sp. cf. 
Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3779 [=LACMIP 23779]. Hypotype UCLA 28987 [=
LACMIP 10564]. Remarks: Described as Chedevillia saltonensis n. sp.,  Squires and Advocate (1986: p. 855).
[Strombidae].
Chlorostoma gallina form 
Newhall area, Los Angeles Co. LACMIP 7753. Hypotype LACMIP 14378. [Tegulidae]. Remarks: Junior synonym
of Tegula gallina,  Alf (2019: p. 73).
Chlorostoma
Monterey Fm., Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312. Hy-
potype SDSNH 93707. Remarks: Tegula sp. [Tegulidae].
Chrysallida rotundomontana

paratype SU 7959 [=CASG. 7959]. Remarks: In the genus/subgenus Odostomia (Besla),  Addicott (1970: p.
143). [Pyramidellidae].
....., SDSNH5422 (ex
Cidarina cidaris (A. Adams inMargarites]. Woodring and Bramlette 1950 [1951]. USGS
-
ity. Hypotype USNM. 560062. [Eucyclidae].
Cirsotrema
(1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242 =LACMIP 22312]. Hypotype UCLA. 59275 [=LACMIP
10551]. [Epitoniidae].
Clathrodrillia mercedensis (Martin, 1914). [Drillia
Merced Fm. [=Purisima Fm.], Pliocene. Two localities listed: Palo Alto and Stanford quads., Santa Clara Co. CASG
28617. Hypotype CASG 5716. [Drilliidae].
Clathrodrillia mercedensis (Martin, 1914). [Drillia]. 
Line drawing, source specimen unknown. [Drilliidae].
Clavatula labiata (Gab, 1881). [Clavatula
Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, Palos Verdes Hills,
Los Angeles Co. USGS 13127. Hypotype USNM 497080. Remarks: Turricula piercei (Arnold, 1909 [1910]), 
Addicott (1970: p. 130). [Clavatulidae].
Clavilithes sp. cf.  (Dickerson, 1913). [Clavella
7. Maniobra Fm., Eocene. Shaver Summit area, Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3779
[=LACMIP 23779]. Hypotype UCLA 28989 [=LACMIP 8839]. Remarks: Clavilithes tabulatus (Dickerson, 1913),
Givens (1974: p. 85);  Squires (1987: p. 41). [Fasciolariidae].
Clavilithes tabulatus (Dickerson, 1913). [Clavella
Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4661. Hypotype UCR 4661/5. [Fasciolariidae].

Clavilithes tabulatus (Dickerson, 1913). [Clavella  
Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115].
Hypotype LAMCIP 6538. [Fasciolariidae].
216 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Clavilithes tabulatus (Dickerson, 1913). [Clavella
Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotypes LACMIP 7478, 7479. [Fasciolariidae].
Clavilithes tabulatus (Dickerson, 1913). [Clavella
Eocene. Whitaker Peak area, Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype LACMIP 7478. [Fasciolari-
idae].
Clavilithes tabulatus (Dickerson, 1913). [Clavella-
cene. Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN. 984 [=LACMIP 16178]. Hypotype LACMIP
7714. [Fasciolariidae].
Clavilithes tabulatus (Dickerson, 1913). [Clavella
-
MIP 11469. [Fasciolariidae].
Clavilithes-

Remarks: Clavilithes n. sp. Clark and Vokes 1936,  Squires (1984: p. 31); Clavilithes tabulatus (Dickerson,
1913),  Squires (1987: p. 41). [Fasciolariidae].
Clavilithes n. sp. Squires 1983a Llajas Canyon, Santa Susana
quad. (1951, PR 1969), Ventura Co. CSUN 445 [=LACMIP 16176]. Hypotype UCLA 59194 [=LACMIP 10547].
Remarks: Clavilithes n. sp. B,  Squires 1984: p. 31. [Fasciolariidae].
-
type UCLA 59195 [=LACMIP 10548]. Remarks: Clavilithes n. sp. B,  Squires (1984: p. 31).
Clavilithes
and Trough canyons, Thousand Oaks quad. (1950, PR 1967), Ventura Co. CSUN 704 [=LACMIP 16177]. Hypotype
LACMIP 6539. Remarks: Clavilithes tabulatus (Dickerson, 1913),  Squires (1987: p. 41). [Fasciolariidae].
Clavilithes         
Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN. 444 [=LACMIP 16203]. Hypotype UCLA 59195
[=LACMIP 10548]. [Fasciolariidae].
Clavocerithium (Clavocerithium) santanum (Loel and Corey, 1932). [Terebra]. Squires 1993b. Veliger 36(4):401–403,
    
Orange Co. CSUN 1185 [=LACMIP 16508]. Hypotypes LACMIP 12102, 12104–12105. [Cerithiidae].

Orange Co. UCMP 6128. Holotype UCMP 31608.

1988), Orange Co. LACMIP 7700. Hypotypes LACMIP 12103, 12106, 12107.


Coalingodea tuberculiformis (G D. Hanna, 1924). [Galeodea
Domengine Fm., Eocene. North of Coalinga, Fresno, Co. UCMP. 672. Hypotypes UCMP 11783, 15807. Remarks:
In the genus Galeodea,  Squires (2019: p. 9). [Cassidae].
Coalingodea tuberculiformis (G D. Hanna, 1924). [Galeodea
Eocene. San Guillermo quad. (1943), Ventura Co. UCR 4752. Hypotype UCR 4752/1. Remarks: In the genus
Galeodea,  Squires (2019: p. 9). [Cassidae].
?Cochlespiropsis jenkinsi-
yon, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 22312. Holotype LACMIP 12741. [Paladmetidae].

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 217
CSUN 371 [=LACMIP 16654]. Paratype LACMIP 12742.
Cockerella* conradiana (Gabb, 1866). [Clathurella
Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14406. Remarks: Error for Crockerella. [Clath-
urellidae].
Colubraria (Colubraria) n. sp. Smith 1975b. SEPM Symposium. Near
junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype SU
10231 [=CASG 61666.21]. [Fasciolariidae].
Columbarium   
Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype SU 10224 [=CASG
61666.19]. [Turbinellidae].
Colus recurvus (Gabb, 1866). [Neptunea
Carson Creek area, Ft. Ross quad., Sonoma Co. UCMP B2155. Hypotype UCMP 14890. [Buccinidae].
Colwellia antiquata (Gabb, 1864). [Nassa          
Domengine Fm., Eocene. Big Tar Canyon area, Cholame quad., Kings Co. UCMP A819. Hypotype UCMP 15939.
[Nassariidae].
Colwellia antiquata (Gabb, 1864). [Nassa
Big Tar Canyon, Cholame quad.], Kings Co. UCMP A819. Hypotype UCMP 15939. [Nassariidae].

Colwellia  

Conomitra aff.  (Weaver, 1912). [Mitra-
cene. West of Reyes Peak, Reyes Peak quad. (1943), Ventura Co. UCR 4706. Hypotype UCR 4706/26. [Mitridae].
Conus bramkampi
Pliocene. Carrizo Mtn., Imperial Co. UCMP A1269. Holotype UCMP 34199. Remarks: In the genus Lithoconus,
Tucker and Tenorio (2009: p. 107). [Conidae].
Conus bramkampi-
nia Fm.], Pliocene. Carrizo Mtn., Imperial Co. UCMP A1269. Holotype UCMP 34199 (not UCMP 34198 as in text).
Remarks: In the genus Lithoconus, Tucker and Tenorio (2009: p. 107). [Conidae].
Conus bramkampi G D. Hanna and Strong, 1949. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert, p. 56,

Remarks: In the genus Lithoconus, Tucker and Tenorio (2009: p. 107). [Conidae].
Conus caleocius
quad. (1952, PR 1969), Los Angeles Co. CSUN. 498 [=LACMIP 16189]. Hypotype LACMIP. 6558. Remarks: In the
genus Dalliconus, Tucker and Tenorio (2009: p. 141). [Conidae].
Conus caleocius Vokes,         Eocene. Canton Canyon,
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 828 [=LACMIP 16141]. Hypotype LACMIP 7498. Remarks:
In the genus Dalliconus, Tucker and Tenorio (2009: p. 141). [Conidae].
Conus caleocius
Ventura Co. CSUN 498 [=LACMIP 16189]. Hypotype LACMIP 6558. Remarks: In the genus Dalliconus, Tucker
and Tenorio (2009: p. 141). [Conidae].
Conus caleocius    
Sawmill Mtn. quad. (1943), Ventura Co. CSUN 979 [=LACMIP 16081]. Hypotype LACMIP 7721. Remarks: In the
genus Dalliconus, Tucker and Tenorio (2009: p. 141). [Conidae].
218 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Conus californicus

1219]. Hypotypes current whereabouts unknown. Remarks: In the genus Californiconus, Tucker and Tenorio
(2009: pp. 155, 156). [Conidae].
Conus durhami
Pliocene. Carrizo Mtn., Imperial Co. UCMP A1269. Holotype UCMP 34200. Remarks: In the genus Cylinder, 
Tucker and Tenorio (2009: p. 68). [Conidae].
Conus durhami
Fm.], Pliocene. Carrizo Mtn., Imperial Co. UCMP A1269. Holotype UCMP 34200. Remarks: In the genus Cylinder,
Tucker and Tenorio (2009: p. 68). [Conidae].
Conus durhami
3.11B. Latrania Fm., Miocene/Pliocene. Alverson Canyon, Imperial Co. SDSNH 5282. Hypotype SDSNH 103738.
Remarks: In the genus Cylinder, Tucker and Tenorio (2009: p. 68). [Conidae].
Conus (Lithoconus) hayesi
Silt, Miocene. Near junction of Kern River and Cottonwood Creek, Kern Co. USGS 6611. Hypotype USNM 650220.
Remarks: In the genus Pyruconus, Tucker and Tenorio (2009: p. 116). [Conidae].
Conus (Lithoconus) hayesi-
Remarks: In
the genus Pyruconus, Tucker and Tenorio (2009: p. 116). [Conidae].
Conus hornii
Canyon quad. (1943), Ventura Co. UCR 4731. Hypotype UCR 4731/14. Remarks: In the genus Profundiconus,
Tucker and Tenorio (2009: p. 150). [Conidae].
Conus hornii piruensis
Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Holotype LACMIP
7500. Remarks: In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus hornii umpquaensisTurner, 1938. Stewart 1946. USGS PP 205-C:91, pl. 
Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS 14497a. Hypotype USNM 498707. Remarks: In the genus
Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus hornii umpquaensis
Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6559. Remarks: In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus hornii umpquaensis
Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype LACMIP 11476.
Remarks: In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus owenianus
63-10. Temblor Fm., Miocene. Kern River area, Kern Co. CASG 65. Hypotype CASG 5708. Remarks: In the genus
Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus owenianus
Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, Palos Verdes Hills, Los Ange-
les Co. USGS 13127. Hypotypes USNM. 497077, 497078. Remarks: In the genus Profundiconus, Tucker and
Tenorio (2009: p. 150). [Conidae].
Conus (Chelyconus) owenianus
Mtn. Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1612. Hypotypes USNM 650221, 650224.
Remarks: In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 219


caption). Hypotypes USNM 650223, 650225.

Hypotype UCMP 36934.
Conus owenianus
Shale Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M2862. Hypotype USNM.
254360. Remarks: In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus owenianus Anderson, 1905. Addicott et al. 1980. SEPM Guidebook 6:18, 20, pl  
Addicott et al., 1978, pl Creek area, La Panza
Range, San Luis Obispo Co. USGS M2862 (not USGS M2869 as in pl. caption). Hypotype USNM 254360. Remarks:
In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus (Chelyconus) owenianus-
Remarks: In the genus
Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus owenianus
Fm., Miocene. Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP2906]. Hypotype LACMIP 8158.
Remarks: In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus cf. owenianus
Monterey Fm., Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to LACMIP.
Remarks: In the genus Profundiconus,  Tucker and Tenorio (2009: p. 150). [Conidae].
Conus remondi
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 219 [=LACMIP 16100]. Hypotype LACMIP 7499. Remarks:
In the genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus remondi         
Whitaker Peak area, Los Angeles Co. CSUN 219 [=LACMIP 16100]. Hypotype LACMIP 7499. Remarks: In the
genus Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus remondi
Mtn. quad. (1943), Ventura Co. CSUN 984 [=LACMIP 16178]. Hypotype LACMIP 7722. Remarks: In the genus
Profundiconus, Tucker and Tenorio (2009: p. 150). [Conidae].
Conus schencki    
Cañada de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP B7013. Holotype SU 9237 [=CASG 70145], para-
type SU 9238 (not SU 9500 as in text) [=CASG 70146]. Remarks: In the genus Profundiconus, Tucker and
Tenorio (2009: p. 150). [Conidae].
Conus spurius
[=Latrania Fm.], Miocene. Coyote Mtns., Imperial Co. LACMIP 9802. Hypotype LACMIP 8274]. Remarks: In the
genus Spuriconus, Tucker and Tenorio (2009: pp. 121, 122). [Conidae].
Conus (Conrad, 1855b). [Volutatithes*
Fm., Eocene. West of Reyes Peak, Reyes Peak quad. (1943), Ventura Co. UCR 4706. Hypotype UCR 4706/141.
Remarks: Error for Volutilithes
Conus
(1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13433. [Conidae].
Cophocara stantoni Stewart, 1926 [1927]. Smith 1975b. SEPM Symposium 
[=LagunaHypotype SU 10240 [=CASG 61616.01]
determined here as missing. Remarks: Perissitys stewarti (Zinsmeister, 1983),  Popenoe and Saul (1987: p.
220 PALEOBIOS, VOLUME 38, FEBRUARY 2021
20). [Perissityidae].

Remarks: Perissitys stewarti (Zinsmeister, 1983),  Popenoe and Saul (1987: p. 20).
Cophocara stewarti
Meier Canyon, Calabasas quad., Ventura Co. UCR 6670. Holotype UCR 6670/900 (not UCR 6670/9 as in text),
paratype UCR 6670/901 [=Pyropsis fantozzi n. sp.,  Saul (1988b: p. 884)]. Remarks: Not a Pyropsis, 
Squires (2011: p. 1204). [Perissityidae].
Corsania (Januncia) janus
Paleocene. Upper part of Well Canyon, Santa Cruz Is. A quad. (1943), Santa Cruz Is., California Channel Ids. LAC-
MIP 23348 (not LACMIP 26720 as in pl. caption). Hypotype LACMIP 12907. [Neritidae].
Corsania (Januncia) rhoga
Near Lower Lake, Lower Lake quad. (1975), Lake Co. LACMIP 7047. Holotype LACMIP 7889. [Neritidae].
Corsania (Januncia) rhoga
Susana Fm., Paleocene. Trailer Canyon, Topanga quad. (1952, PR 1981), Santa Monica Mtns., Los Angeles Co.
LACMIP 10508. Hypotype LACMIP 12906. [Neritidae].
Corsania (Januncia) susana
Eocene. Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 969 [=LACMIP 16849]. Holotype
LACMIP 7890. [Neritidae].

LACMIP 7891.

paratype LACMIP 6441.
Corsania (Januncia) susana
Fm., Eocene. Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 969 (=LACMIP 16894). Ho-
lotype LACMIP 7890. [Neritidae].
Crassispira olcesensis
Kern River, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1608. Holotype USNM 650264. [Pseudomelatomidae].
Crassispira zizyphus (Berry, 1940). [Clavus 

Angeles Co. LACMIP 1219. Hypotypes (7) current whereabouts unknown. [Pseudomelatomidae].
Crassispira sp. Squires 2012. NHMLAC CS 520:77, 
17918. Hypotype LACMIP 14408. [Pseudomelatomidae].
Crawfordina fugleri (Arnold, 1907). [Cancellaria]. Woodring and Bramlette 1950 [1951]. USGS PP 222:77, pl. 16,

560087. [Cancellariidae].
Crenilabrum venturaensis (Nelson, 1925). [Acteon
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6684. Hypotype UCR 6684/27 determined here as missing.
[Acteonidae].
Crepidula aculeata (Gmelin, 1791). [Patella     
Newhall area, Los Angeles Co. LACMIP 17918. Hypotypes LACMIP 14385, 14386. Remarks: Two specimens
vertically stacked. [Calyptraeidae].
Crepidula bractea    
Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650088. [Calyptraeidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 221

CASG 2064 [=CASG 65]. Paratype CASG 12931 [=CASG 2064.01].

Crepidula inornata
Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963). Hypotype LACMIP
12453. [Calyptraeidae].
Crepidula nummaria
Fm., Miocene/Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M1929. Hypotype USNM 649095.
[Calyptraeidae].
Crepidula cf.   
(Siphonalia Zone), Pliocene. Los Jinetes, North Dome, Kettleman Hills, Kings Co. USGS 12393. Hypotype USNM
495814. [Calyptraeidae].
Crepidula onyx
possibly at SDSNH, current whereabouts unknown. Remarks: Ex Grouard collection. [Calyptraeidae].
Crepidula onyx
Fm., Pliocene [=Wilson Grove Fm.], Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotype CASG, determined
here as missing. [Calyptraeidae].
Crepidula (Spirocrypta) pileum (Gabb, 1864). [Crypta (Spirocrypta)
Crepidula pileum)). Tejon Fm., Eocene. Tejon, [Fresno
Co.]. Lectotype (of Stewart 1926 [1927]: p. 342) ANSP. 4221]. Remarks: In the genus Spirocrypta,  Saul and
Squires (2008a: p. 131). [Calyptraeidae].
Crepidula pileum (Gabb, 1864). [Crypta (Spirocrypta)]. Weaver and Kleinpell 1963. 
Middle Gaviota Fm., Eocene. Near town of Las Cruces, Los Olivos quad., Santa Barbara Co. UCMP B7008. Hypotype
SU 9196 [=CASG. 70157]. Remarks: In the genus Spirocrypta,  Saul and Squires (2008a: p. 131). [Calyptraeidae].

Co. UCMP.B6955. Hypotype SU 9197 [=CASG 70159].
Crepidula (Spirocrypta) pileum (Gabb, 1864). [Crypta (Spirocrypta)]. Squires 1994b. SEPM Guidebook 74:54, pl. 2,

Remarks: In the genus Spirocrypta,  Saul and Squires (2008a: p. 131). [Calyptraeidae].
Crepidula princeps
Pliocene. Santa Barbara Co. Hypotype CASG 5937 (originally CSMB. 14639). Remarks: Supplemental description

genus Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps
Fm. (diatomaceous strata), Miocene Purisima Hills, Santa Barbara Co. USGS 14969. Hypotype USNM 560021.
Remarks:
illustrated in Conrad, 1857b. In the genus Grandicrepidula,  Squires 2012: p. 77. [Calyptraeidae].


Hypotype USNM 560045.
Crepidula princeps
specimen unknown. Remarks:  
Grandicrepidula,  Squires, (2012: p. 77). [Calyptraeidae].
Crepidula princeps          
222 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pliocene. La Honda quad., San Mateo Co. 711. Hypotype determined here as missing. Remarks: Supplemental
Grandicrepidula,  Squires (2012: p. 77). [Ca-
lyptraeidae].
Crepidula princeps
San Diego, San Diego Co. SDSNH 2383. Hypotype SDSNH 4123]. Remarks:
illustrated in Conrad, 1857b. In the genus Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula cf. 
Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650969. Remarks: Supplemental description
Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps
Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotypes USNM 650092, 650094. Remarks: Supple-
Grandicrepidula,  Squires (2012: p.
77). [Calyptraeidae].
19, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1590. Hypotype USNM 650093.
Crepidula princeps

Angeles Co. LACMIP 1219]. Hypotypes (3) current whereabouts unknown. Remarks: Supplemental description
Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps         
Capitola area, Santa Cruz Co. SCCM1. Hypotypes SCCM 9603.00, 9603.02]. Remarks: Supplemental description
Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula aff. -
tocene. Twelvemile Creek area, San Mateo Co. UCMP D3364. Hypotype USNM 251805. Remarks: Supplemental
Grandicrepidula,  Squires (2012: p. 77). [Ca-
lyptraeidae].
Crepidula princeps
Fm.], Pliocene. Northern Simi Valley, Ventura Co. CSUN 725 [LACMIP 29227]. Hypotype UCLA 59412 [=LACMIP
8325]. Remarks:Grandicrepidula,
 Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps  
Co. [SDSNH 4080]. SDSNH hypotype determined here as missing. Remarks:
illustrated in Conrad, 1857b. In the genus Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps       
Escalona Gulch, Santa Cruz Co. CASG66095. Hypotype CASG 66095.02. Remarks: Supplemental description
Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps
Fm., Pliocene. Los Angeles, Los Angeles Co. LACMIP 466. Hypotype LACMIP 6619 (not LACMIP 6722 as published).
Remarks: Grandicrepidula, 
Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps

29227]. Hypotype UCLA. 59412 [=LACMIP 8325]. Remarks:
Conrad, 1857b. In the genus Grandicrepidula,  Squires (2012: p. 77). [Calyptraeidae].
Crepidula princeps-
Remarks: Supplemental description
Grandicrepidula,  Squires (2012: p. 77). [Calyptraeaidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 223
Crepidula rostralis (Conrad, 1865). [Crypta  
Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1600. Hypotype USNM 650089. [Calyptraeidae].
....., pl. 4, Bravo Ranch quad. (1954) Kern Co. USGS M1608.
Hypotype USNM 650090.
-
potype UCMP 15458.

Crepidula rostralis (Conrad, 1865). [Crypta
Fm. (Buttonbed Sand Member) (of Stinemeyer et al. 1959), Miocene. Shale Pt. quad., Temblor Range, Kern Co.
USGS M3986. Hypotype USNM 646798. [Calyptraeidae].
Crepidula cf. Phacoides
Sand member) (of Stinemeyer et al. 1959: pp. 12, 13), Miocene. Las Yeguas Ranch quad., Temblor Range, Kern
Co. USGS M3280 [M3281 in Addicott 1973]. Hypotype USGS 646606. [Calyptraeidae].
Crepidula cf. -
ber), Miocene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM 646604.
[Calyptraeidae].
     
USNM 646605.

Hypotype USNM 646606.
Crepidula      
  
Hypotype LACMIP 7462. [Calyptraeidae].
Crepidula         
Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 305 [=LACMIP 16197]. Hypotype UCLA 58212 [=LACMIP
10572]. [Calyptraeidae].
Crepidula
Fm.], Miocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype SBNMH 60193]. Remarks:
Determined here as Crepidula praerupta Conrad, 1849a. [Calyptraeidae].
Crommium andersoni (Dickerson, 1914c). [Amauropsis
Hot Springs Creek, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4659. Hypotype UCR 4659/5. [Naticidae].
Crommium andersoni (Dickerson, 1914c). [Amauropsis]. Givens and Kennedy 1976. Jour. Paleo. 50(5):963, 969, pl.
-
types UCR 4865/80, 4865/81. [Naticidae].
Crommium sp. cf. 
Eocene. Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963]. Hypotype
LACMIP 12454. [Naticidae].
Crommium pinyonensis (Dickerson, 1914c). [Natica]. Marincovich -
tinez Fm. [=San Francisquito Fm.], Paleocene. Pinyon Ridge, Rock Creek quad., Los Angeles Co. UCMP 2245].
Holotype UCMP 11761. [Naticidae].

Holotype (of Ampullella schencki Vokes, 1939) UCMP 15856. Remarks: Ampullella schencki Vokes, 1939, 
Squires (1987: p. 35).
224 PALEOBIOS, VOLUME 38, FEBRUARY 2021

UCMP A9717. Hypotypes UCMP 14398, 14399, 32707.
Remarks: Ampullella
schencki Vokes, 1939,  Squires (1987: p. 35).
Crommium pinyonensis (Dickerson, 1914b). [Natica
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 802 [=LACMIP 16144]. Hypotype LACMIP 7467. [Naticidae].
Crucibulum (Dispotaea) papulum
Miocene. Kern River, Kern Co. USGS 6623. Holotype USNM 650098, paratype USNM. 650097. [Calyptraeidae].

UCMP 15457.
Crucibulum cf.  (Wood, 1828) [Patella
Round Mtn. Silt, Miocene. Cottonwood Creek, Caliente quad. (1914), Kern Co. UCMP B1588. Hypotype UCMP
15463. [Calyptraeidae].
Crucibulum cf. scutellatum (Wood, 1828). [Patella
(Jackson Fork Member), Pliocene. Fish Creek/Vallecito Creek, San Diego Co. ABDSP 5070.4]. Hypotype ABDSP I
11189]. [Calyptraeidae].
Crucibulum-
stone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312.
Hypotype SDSNH 93719 [in errata, p. 50]. [Calyptraeidae].
Cryptochorda (Cryptochorda) californica (Cooper, 1894). [Ancilla
Fm., Eocene. Thousand Oaks quad. (1950, PR 1967), Ventura Co. CSUN 703 [=LACMIP 16199]. Hypotype LACMIP
6549. [Volutidae].
Cryptochorda (Cryptochorda) californica (Cooper, 1894). [Ancilla
Juncal Fm., Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 361 [=LACMIP 16200]. Hypotype LACMIP
7487. [Volutidae].
Cryptochorda (Lyrianella) lyrata Nelson, 1925. Wenz 1943-
son, 1925, pl., Paleocene]. Simi Hills, Camulos quad.,
Ventura Co. UCMP 3808. Holotype UCMP 30543. [Volutidae].
Cryptochorda-
ermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA 6525 [=LACMIP
26525]. Hypotype LACMIP 7565. [Volutidae].
Cryptoconus cooperi (Dickerson, 1916). [Drillia    
Whitaker Peak area, Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype LACMIP 7497. Remarks: Crypto-
conus injucundus G D. Hanna, 1924: p. 164, new name for Cryptoconus cooperi Dickerson, 1916, preoccupied.
[Cryptoconidae].
Cryptoconus cooperi (Dickerson, 1916). [Drillia
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 975 [=LACMIP 16201]. Hypotype LACMIP 7720.
Remarks: Cryptoconus injucundus G D. Hanna, 1924: p. 164, new name for Cryptoconus cooperi (Dickerson,
1916), preoccupied. [Cryptoconidae].
Cryptoconus cooperi (Dickerson, 1916). [Drillia-
mere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13432. Remarks:
Cryptoconus injucundus G D. Hanna, 1924: p. 164, new name for Cryptoconus cooperi (Dickerson, 1916), preoc-
cupied. [Cryptoconidae].
Cryptonatica aleutica (Dall, 1919a). [Natica
Foxen Mudstone, Pliocene. Santa Barbara Co. USGS 4473. Hypotype USNM 560046. Remarks: Junior synonym
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 225
of Natica (Cryptonatica) clausa Broderip and G.B. Sowerby I, 1829,  Marincovich (1977: p. 411). [Naticidae].
Cryptonatica aleutica (Dall, 1919a). [Natica      
Pliocene. Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650954. Remarks: Junior synonym of
Natica (Cryptonatica) clausa Broderip and G.B. Sowerby I, 1829,  Marincovich (1977: p. 411). [Naticidae].

Cryptonatica clausa (Broderip and Sowerby I, 1829). [Natica]
Pliocene. Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14390. [Naticidae].
Cylichna? loismartinae       
         
[=CASG. 7966]. [Cylichnidae].
....., LACMIP 16140 (ex
....., SDSNH5422 (ex
Cylichna? loismartinae
Silt, Miocene. Caliente quad. (1914), Kern Co. UCMP B1637. Hypotypes UCMP 37104–37106. [Cylichnidae].
Cylichna temblorensis Keen,       13,14. Lowermost Round Mtn.
Silt, Miocene. Caliente quad., Kern Co. 
7960–7962]. [Cylichnidae].
....., LACMIP16140 (ex
....., [SDSNH 5422 (ex
Cylichna temblorensis      
Miocene. Caliente quad. (1914), Kern Co. UCMP B1637. Hypotypes UCMP 37107, 37108. [Cylichnidae].

[Cylichnidae].
     
USNM 650284. [Cylichnidae].
Cylichnina costata (Gabb, 1864). [Cylichna
Fm., Paleocene. Simi Hills, Ventura Co. UCR 6871. Hypotype UCR 6871/9. Remarks: In the genus/subgenus
Scaphander (Mirascapha),  Squires (1984: p. 40, 1987: p. 53). [Cylichnidae].
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella  
11. Avenal Sandstone, Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS 14497a. Hypotype USNM 498712.
[Cylichnidae].
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella
Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP 16115]. Hypotype LACMIP 6562.
[Cylichnidae].
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella
Fm., Eocene. Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP
16115]. Hypotype LACMIP 6562 (not LACMIP 6561 as in pl. caption). [Cylichnidae].
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella
-
MIP 7501. [Cylichnidae].
Cylichnina tantilla (Anderson and G D.Hanna, 1925). [Cylichnella
226 PALEOBIOS, VOLUME 38, FEBRUARY 2021

LACMIP 7723. [Cylichnidae].
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella]. Squires 1991c. SEPM Guidebook 68:222, pl. 2,

Hypotype LACMIP 11477. [Cylichnidae].
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella
Susana Fm., Eocene. Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 964 [=LACMIP 21551]. Hypotype
LACMIP 12691. [Cylichnidae].
 12692.
Cylichnina tantilla (Anderson and G D. Hanna, 1925). [Cylichnella
Fm., Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP
13434. [Cylichnidae].
Cymatium (Reticutriton) elsmerense (English, 1914a). [Gyrineum
Diego Fm.], Pliocene. [Hills south of Tijuana River], San Diego Co. LACMIP 305. [Hypotypes LACMIP 12743–12745].
[Ranellidae].
Cymatium sp., cf. C. (Reticutriton) elsmerense (English, 1914a). [Gyrineum
75. Pico Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14394. [Ranellidae].
Cymatium (Septa) janetae
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 444 [=LACMIP 16203]. Holotype UCLA
59191 [=LACMIP 10543]. [Ranellidae].
-
type UCLA 59192 [= LACMIP 10544].
-
type UCLA 59193 [=LACMIP 10545]. Remarks: Sassia bilineata (Dickerson, 1916),  Squires (1987: p. 40).
Cymatium (Septa) janetae-
sana quad. (1951, PR 1969) Ventura Co. CSUN 444 [=LACMIP 16203]. Holotype UCLA 59191 [=LACMIP 10543].
[Ranellidae].
Cymatium (Septa) janetae

59191 [=LACMIP 10543]. [Ranellidae].
Cymatium
just north of the Kern River, Kern Co. CASG 2064 [=CASG 65]. Hypotype CASG. 12932. [Ranellidae].
Cypraea castacensis
Remarks: In the
genus Eocypraea,  Groves (1992: p. 106). [Ovulidae].
      
UCMP A1282. Hypotype UCMP 15815].
Cypraea castacensis Stewart, 1926 [1927]. Squires 1984. NHMLAC CS 350:23–24
Llajas Canyon, Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP. 6523. Remarks: In the genus Eocyp-
raea,  Groves (1992: p. 106). [Ovulidae].
Cypraea fresnoënsis
Northwest of Coalinga, Fresno Co. CASG 61713. Holotype CASG. 50 [=CASG 61713.01]. Remarks: In the genus
Bernaya,  Groves (1992: p. 106). [Cypraeidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 227
Cypraea fresnoënsis Cypraea fresnoensis in pl.
caption). Avenal Sandstone, Eocene. Northwest of Coalinga, Fresno Co. CASG 61713. Holotype CASG 50 [=CASG
61713.01]. Remarks: In the genus Bernaya,  Groves (1992: p. 106). [Cypraeidae].
Cypraea kemperae
Eocene [=Paleocene]. South of Simi Valley, Ventura Co. UCMP 3764. Syntype [=paratype in UCMP online database]
UCMP 30541. Remarks: In the genus Propustularia,  Groves (1992: p. 106). [Cypraeidae].
Cypraea kemperae
Fm.], Eocene [=Paleocene]. South of Simi Valley, Ventura Co. CASG 391. Holotype CASG 987 [=CASG 391.01].
Remarks: In the genus Propustularia,  Groves (1992: p. 106). [Cypraeidae].
Cypraea mathewsonii       
Contra Costa Co. Holotype ANSP 4217. Remarks: In the genus Sulcocypraea,  Squires and Groves (1993: p.
83). [Ovulidae].
Cypraea mathewsonii
Contra Costa Co. Holotype ANSP 4217]. Remarks: In the genus Sulcocypraea,  Squires and Groves (1993: p.
83). [Ovulidae].
Cypraea novasumma (Nelson, 1925). [Ovula
Susana Fm.], Eocene [=Paleocene]. South side of Simi Valley, Ventura Co. UCMP. 3776. Holotype UCMP 30449.
Remarks: In the genus Eocypraea,  Groves (1992: p. 106). [Ovulidae].
Cypraea novasumma (Nelson, 1925). [Ovula
Susana Fm.], Eocene [=Paleocene]. South side of Simi Valley, Ventura Co. UCMP 3776. Holotype UCMP 30449.
Remarks: In the genus Eocypraea,  Groves (1992: p. 106). [Ovulidae].
Cypraea simiensis
Fm.], Eocene [=Paleocene]. South side of Simi Valley, Ventura Co.. UCMP 3818. Holotype UCMP 30498. Remarks:
In the genus Propustularia,  Groves (1992: p. 106). [Cypraeidae].
Cypraea simiensis
Fm.], Eocene [=Paleocene]. South of Simi Valley, Ventura Co. UCMP 3818. Holotype UCMP 30498. Remarks: In
the genus Propustularia,  Groves (1992: p. 106). [Cypraeidae].
Cypraea spadicea Swainson, 1823a. Shimer and Shrock 1944. Index Foss
Angeles Co. SDSNH 217a. Hypotype
SDSNH. 270. Remarks: In the genus Neobernaya,  Groves, (2019: pp. 131–136). [Cypraeidae].
Cypraea
Rd. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=CASG. 61666]. Hypotype
SU 10236 [=CASG 61666.04]. [Ovulidae].

UCMP 14208.
Cypraea
Sawmill Mtn. quad. (1943), Ventura Co. CSUN 984 [=LACMIP 16178]. Hypotype LACMIP 7704. [Ovulidae].
Calyptraea sp. in caption,

site, Laguna Hills, Orange Co., El Torro quad. SDSNH 4520. Hypotype SDSNH 98688. [Cypraeidae].
DemondiaPaleoBios
Hills, Oakland, Alameda Co. UCMP IP13003. Hypotypes UCMP 218821, 218822. [Nassariidae].
Dentimitra clementensis (M.A. Hanna, 1927). [Mitra

UCMP 5051. Holotype UCMP 30991. [Mitridae].
228 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Dentimitra murietta (Anderson and G D. Hanna, 1925). [Mitra]. Cernohorsky 1976. Indo-Pac. Moll. 3(17):510, pl.

Creek, Kern Co. CASG 245]. Holotype CASG 834, paratype CASG 835. [Mitridae].
Dentrimitra simplicissima (Cooper, 1894). [Mitra      


-
ceous/earliest Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad. (1948, PR 1979), San Luis Obispo
Co. CASG 61598. Holotype CASG 61598.01 (ex CASG 30031). [Buccinidae].
Deussenia Cretaceous/lowermost
Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. CASG 61598. Hypotype CASG
61598.01. Remarks:  n. sp.,  Squires and Saul (2000: p. 123). [Buccinidae].
Diodora arnoldi
Pliocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS M6501. Hypotype USNM
235585. [Fissurellidae].
Diodora arnoldi-
SNH 4080]. Hypotype SDSNH 75910. [Fissurellidae].
Diodora cf.  (Eschscholtz in Rathke, 1833). [Fissurella

Hypotype UCMP 37680 (not UCMP 30680 as in text). [Fissurellidae].
Diadora* aspera (Eschscholtz in Rathke, 1833). [Fissurella]. Bedrossian 1974. Calif. Geol. 27(8):177, unnumbered

Co. Hypotype CASG determined here as missing. Remarks: Error for Diodora. [Fissurellidae].
Diodora cayenensis (Lamarck, 1822). [Fissurella
16–21. Imperial Fm. [=Latrania Fm.], Pliocene. Whitewater River, San Jacinto quad., Riverside Co. UCMP A1407.
Hypotypes UCMP 16080–16083. [Fissurellidae].
Diodora aff.  (Weaver and Palmer, 1922). [Fissuridea]. Squires and Deméré 1991. SEPM Guidebook

Diodora (Sowerby I, 1835). [Fissurella]. Durham and Addicott 1965.USGS PP 524-A:A12, pl.

M977. Hypotype USNM 649103. [Fissurellidae].
Diodora
Kern Co. UCMP B1605. Hypotype UCMP 11031. [Fissurellidae].
Diodora (Diodora
basal part of the Jewett Sand, Woody quad. (1952), Kern Co. UCMP B1674. Hypotype UCMP 10064. [Fissurellidae].
Diodora
Palms, San Gorgonio Pass, Riverside Co. UCR 4777. Hypotype UCR 4777/10. [Fissurellidae].
Diodora
Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13413. [Fissurellidae].
Diodora-
cality now covered by gate-guarded housing tract, Santa Monica Mtns.,
Topanga quad. (1952, PR 1981), Los Angeles Co. LACMIP 16888. Hypotypes LACMIP 7940, 7941. [Fissurellidae].
Domenginella claytonensis (Gabb, 1864). [Turris 
      
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 229
15776]. [Clathurellidae].
Domenginella claytonensis (Gabb, 1864). [Turris

15775, 15776]. [Clathurellidae].
Domenginella claytonensis (Gabb, 1864). [Turris
Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115].
Hypotype LACMIP 6556. Remarks: Trypanotoma stocki (Dickerson, 1916),  Squires (1988c: p. 16). [Clath-
urellidae].
Domenginella aff.  (Gabb, 1864). [Turris-

LACMIP 11475. [Clathurellidae].
Drillia columbiana
Garza Creek, Reef Ridge quad., Kings Co. UCMP D1056. Hypotype UCMP 36900. Remarks: In the genus Ptycho-
syrinx,  Hickman (1976a: p. 94). [Drilliidae].
Drillia graciosana
Careaga Fm. (Graciosa Member), Pliocene. Graciosa Ridge, Santa Barbara Co. USGS 14613. Hypotypes USNM
560130–560133. 62. Remarks: In the genus Ophiodermella,  Addicott (1969: p. 62). [Drilliidae].

Echinophoria dalliGaleodea-
ota Fm., Eocene. Cañada del Corral, Gaviota quad., Santa Barbara Co. UCMP B6968. Hypotype SU 9215 [=CASG
70162]. [Cassidae].
Echinophoria trituberculata (Weaver, 1912). [Morio
Fm., Eocene. Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP B6920. Hypotype SU 9216 [=CASG 70163].
Remarks:
Echinophoria trituberculata (Weaver, 1912). [Morio]. Squires 2019. PaleoBios
Grapevine Canyon, Tehachapi Mtns., Kern Co. LACMIP 22340. Hypotype 14836. [Cassidae].
Ectinochilus (Cowlitzia) canalifer (Gabb, 1864). [Rostellaria (Rimella)]. Weaver and Kleinpell 1963. UCPG 43:189,

9207 [=CASG 70164]. Remarks: Junior synonym of Rimella supraplicata (Gabb, 1864),  Squires (2013: p.
835). [Strombidae].
Ectinochilus (Cowlitzia) canalifer (Gabb, 1864). [Rostellaria (Rimella)]. Givens 1974. UCPG 109:72, pl. 7, 
Fm., Eocene. Near Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4714. Hypotype UCR 4714/30.
Remarks: Junior synonym of Rimella supraplicata (Gabb, 1864),  Squires (2013: p. 835). [Strombidae].
Ectinochilus (Cowlitzia) canalifer supraplicatusNeptunea]. Weaver and Kleinpell 1963. UCPG 43:189,
   -
potype SU 9208 [=CASG 70165]. Remarks: Junior synonym of Rimella supraplicata (Gabb, 1864),  Squires
(2013: p. 835). [Strombidae].
Ectinochilus (Vaderos) cf. ( ) elongata (Weaver, 1912). [Rimella]. Squires 1991c. SEPM Guidebook 68:220, pl.

662). Hypotype LACMIP 11463. [Strombidae]. Remarks: Junior synonym of Rimella macilenta White, 1889, 
Squires (2013: p. 833). [Strombidae].
Ectinochilus (Macilentos) macilentus (White, 1889). [Rimella-
Rimella macilenta)). Chico Fm., Cretaceous [=Domengine Fm., Eocene].
New Idria, Fresco [sic] Co. USGS 212. Holotype USNM 20114. Remarks: In the genus Rimella,  Squires (2013:
p. 831). [Strombidae].
230 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Ectinochilus macilentum* (White, 1889). [Rimella

212. Holotype USNM. 20114 (25 syntypes as per online database), CASG paratype determined here as missing.
Remarks: Error for  In the genus Rimella,  Squires (2013: p. 831). [Strombidae].
Ectinochilus macilentus (White, 1889). [Rimella-
stone, Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS.14500. Hypotypes USNM 498713, 498716. Remarks:
In the genus Rimella,  Squires (2013: p. 831). [Strombidae].

Ectinochilus (Macilentos) macilentus (White, 1889). [Rimella
Eocene. Thorn Meadows area, Lion Canyon quad. (1943), Ventura Co. UCR 4680. Hypotype UCR 4680/17. Re-
marks: In the genus Rimella,  Squires (2013: p. 831). [Strombidae].

[Strombidae].
Ectinochilus (Macilentos) macilentus (White, 1889). [Rimella-
jas Fm., Eocene. Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP 16115]. Hypotype LACMIP 6521.
Remarks: In the genus Rimella,  Squires (2013: p. 831). [Strombidae].
Ectinochilus (Macilentos) macilentus (White, 1889). [Rimella]. Squires 1984. NHMLAC CS
Fm., Eocene. Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LAC-
MIP 16115]. Hypotype LACMIP 6521. Remarks: In the genus Rimella,  Squires (2013: p. 831). [Strombidae].
Ectinochilus (Macilentos) macilentus (White 1889). [Rimella
Eocene. Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype
LACMIP 7463. Remarks: In the genus Rimella,  Squires (2013: p. 831). [Strombidae].
Ectinochilus (Macilentos) macilentus (White, 1889). [Rimella

LACMIP 7702. Remarks: In the genus Rimella,  Squires (2013: p. 831). [Strombidae].
Ectinochilus (Macilentos) macilentus (White, 1889). [Rimella
Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13420.
Remarks: In the genus Rimella,  Squires (2013: p. 831). [Strombidae].
Ectinochilus (Cowlitzia) supraplicatus (Gabb, 1864). [Neptunea
Eocene. San Guillermo quad. (1943), Ventura Co. UCR 4753. Hypotype UCR 4753/131. Remarks: In the genus
Rimella,  Squires (2013: p. 831). [Strombidae].
Elaeocyma
7752. Hypotype LACMIP 14407. [Drilliidae].
Eocernina hannibali (Dickerson, 1914c). [Natica
Las [sic] Llajas Fm., Eocene. Aliso Canyon of Devil Creek, Santa Susana quad., Los Angeles Co. CASG 393. Hypotype
CASG 5962. [Naticidae].
Eocernina hannibali (Dickerson, 1914c). [Natica
Fm., Eocene. Santa Susana quad., Ventura Co. UCMP.7004. Hypotypes UCMP 14401, 14402. [Naticidae].


Eocernina hannibali (Dickerson, 1914c). [Natica
Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP 7242. Hypotype LACMIP 6524. [Naticidae].
Eocernina hannibali (Dickerson, 1914c). [Natica
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 231
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP
6524. [Naticidae].
Eocernina hannibali (Dickerson, 1914c). [Natica]. Squires 1987. NHMLAC CS 388:36, 
Eocene. Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype
LACMIP 7468. [Naticidae].
Eocernina hannibali (Dickerson, 1914c). [Natica
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 984 [=LACMIP 16178]. Hypotype LACMIP 7705.
[Naticidae].
Eocernina hannibali (Dickerson, 1914c). [Natica
-
MIP 11464. [Naticidae].
Eocernina hannibali (Dickerson, 1914c). [Natica

LACMIP 6524. [Naticidae].
Eocernina hanniballi (Dickerson, 1914c). [Natica-
man Rancho Fm., Eocene. South of Gualala, Mendocino Co. Wentworth W241 [=UCMP W241]. Hypotype UCMP
39916. [Naticidae].
Eocernina hanniballi (Dickerson, 1914c). [Natica-
mere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13422. [Naticidae].
Eocernina hespericosta
Simi Hills, Thousand Oaks quad., Ventura Co. UCR 7320. Holotype UCR 7320/4. [Naticidae].
Eocernina
Hills, Ventura Co. UCR 7320. Hypotype UCR 7320/4. Remarks: Eocernina hespericosta n. sp.,  Zinsmeister
(1983b: p. 1293). [Naticidae].
Eocithara mutica californiensis (Vokes, 1937b). [Harpa

Holotype UCMP 30438. [Harpidae].
Eocithara mutica californiensis (Vokes, 1939). [Harpa
Devil Canyon, Santa Susana quad. (sec. cor. is in the Oat Mtn. quad.), Los Angeles Co. CSUN 373 [=LACMIP 16211].
Hypotype LACMIP 6548. [Harpidae].
Eocithara clarki Vokes, 1937b. [Harpa
-
lotype UCMP 15792. [Harpidae].
Eocypraea? maniobraensis
Fm., Eocene. Orocopia Mtns., Canyon Spring NW quad. (1963), Riverside Co. CSUN 662 [=UCLA 3779) [=LACMIP
23779]. Holotype UCLA 48431 [=LACMIP 10566]. Remarks: In the genus Eocypraea,  Groves (1992: p. 106).
[Ovulidae].
Eocypraea maniobraensis Squires and Advocate, 1986. Squires 1991c. SEPM Guidebook   
. (1987), Riverside Co. LACMIP 23779 [=CSUN 662]. Holotype UCLA 48431
[=LACMIP 10566]. Remarks: In the genus Eocypraea,  Groves (1992: p. 106). [Ovulidae].
Eocypraea () sp., cf. Eocypraea () castacensis (Stewart, 1926 [1927]). [Cypraea]. Squires 2008. NHMLAC CS

Hypotype LACMIP 13421. Remarks: In the genus Eocypraea,  Groves (1992: p. 106). [Ovulidae].
Eocypraea (Eocypraea) sp. cf. () (Lamarck, 1802). [Cypraea

232 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Eocypraea novasumma (Nelson 1925). [Ovula
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6871. Hypotype UCR 6871/13 (not UCR 3776/1 as in pl. cap-
tion). [Ovulidae].
Eocypraea (Eocypraea) takeosusukii-
leocene Herndon Creek near Lower Lake, Lake Co. LACMIP 7045. Holotype LACMIP 13645. [Ovulidae].
Eopleurotoma whitakerpeakensis
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 827 [=LACMIP 16153]. Holotype LACMIP 7496. [Turridae].
Eosurcula aff. Eosurcula inconstans (Cooper, 1894). [Surcula 
Llajas Fm., Eocene Simi Valley, Santa Susana quad. (1903), Ventura Co. LAMCIP 7434. Hypotype LACMIP 12743.
[Turridae].
Epitonium (Gyroscala) barkerianum
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650077. [Epitoniidae].
Epitonium (Cirsotrema) clallamensis
Miocene. Caliente quad., Kern Co. UCMP B1665. Hypotype UCMP 36553. [Epitoniidae].
Epitonium (Cirsotrema) clallamense
Jewett Sand, Miocene. Caliente quad. (1914), Kern Co. UCMP B1665. Hypotypes UCMP 15428, 36553 (not UCMP
36533 as in pl. caption). [Epitoniidae].


Epitonium (Cirsostrema) clallamensis

determined here as missing. [Epitoniidae].
Epitonium (Asperiscala) hemphilli (Dall, 1878a). [Scalaria]. Stadum 1984a. NHFOC 
Fm., Pliocene. Orange Co. SDSNH 4080. Hypotype SDSNH 75912]. [Epitoniidae].
Epitonium (Cirsotrema) posoense 
Lower Round Mtn. Silt, Miocene. Kern River, Kern Co. UCLA AC-14-34 [=LACMIP 26326]. Hypotype UCLA 45765
[=LACMIP 8431]. [Epitoniidae].
Epitonium (Nitidiscala) tedfordi
Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650075. [Epitoniidae].

Epitonium (Nitidiscala) tinctum
5. Unnamed strata, Pliocene. Near Lemon Tank Reservoir, San Clemente Is., California Channel Ids. USGS M6501.
Hypotype USNM 235584. [Epitoniidae].
Erato cf.  G. B. Sowerby I, 1832 [not Gray as in pl. caption]. Woodring and Bramlette 1950 [1951].

Co. USGS 14648. Hypotype USNM 560074. Remarks: Determined here to be Hespererato columbella (Menke,
1847). [Eratoidae].
Eulima gabbiana (Anderson and Martin, 1914). [Eulimella
Mtn. Silt (possibly uppermost Olcese Sand), Miocene. Kern River, Caliente quad., Kern Co. SU 2641. Hypotype SU
7543 [=CASG 72000]. Remarks: In the genus Melanella,  Hanna (1924: p. 170). [Eulimidae].
Eulima gabbiana (Anderson and Martin, 1914). [Eulimella 
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 233
Round Mtn. Silt, Miocene. Caliente quad. (1914), Kern Co. UCMP B1637. Hypotype UCMP 37103. Remarks: In
the genus Melanella,  Hanna (1924: p. 170). [Eulimidae].

Euspira clementensis (M.A. Hanna, 1927). [Natica
Thorn Meadows area, Lion Canyon quad. (1943), Ventura Co. UCR 4688. Hypotype UCR 4688/25. [Naticidae].
Euspira nuciformis (Gabb, 1864). [Lunatia-
cene. Reef Ridge, Big Tar Canyon, Reef Ridge quad., Kings Co. USGS 14495. Hypotype USNM 498717. [Naticidae].
Euspira nuciformis (Gabb, 1864). [Lunatia
Mtn. Lodge, Reyes Peak quad. (1943), Ventura Co. UCR 4671. Hypotype UCR 4671/2. Remarks: In the genus/
subgenus Polinices (Euspira),  Marincovich (1977: p. 281);  Squires (1987: p. 37). [Naticidae].
Euspira

Euspirocrommium hewitti (G D. Hanna and Hertlein, 1949). [Ampullella]. Marincovich 1977. BAP 70 (294):237–238,
Eocene. Media Agua Creek, Kern Co. CASG 61884. Holotype CASG 9430, hypotype CASG
32388. [Naticidae].

quad. (1943), Ventura Co. UCR 4668. Hypotype UCR. 4668/21.
Euthriofusus (Whitneyella) sinuatus (Gabb, 1864). [Fasciolaria   
Whitneyella sinuata)). Tejon Fm., Eocene. Tejon, Kern Co.
Lectotype (of Stewart 1926 [1927]: p. 402) UCMP 31404]. Remarks: Determined here as in the genus Whit-
neyella. [Fasciolariide].
Exilia fausta
Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4726. Hypotype UCR 4726/19. [Turbinellidae].
Exilia llajasensis
Susana quad. (1951, PR 1969), Ventura Co. CSUN 539 [=LACMIP 16231]. Hypotype LACMIP 6557. [Turbinellidae].
Falsifusus cf.  (Merriam and Turner, 1937). [Fusinus-
cal Fm., Eocene. Hot Springs Creek, Topatopa Mtn. quad. (1943), Ventura Co. UCR 4659. Hypotype UCR 4659/9.
[Fasciolariidae].
 
(Saddleback Valley Limestone member). Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4312. Hypotype SDSNH 93739. [Fasciolariidae].
Favartia       
Canyon quad., Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype LACMIP 11468. [Muricidae].
Ferminoscala durhami

[=CASG 7963]. Remarks: In the genus Scalina,  Addicott (1970: p. 56). Ferminoscala and Scalina =Amaea, 
WoRMS (2020), thus making Amaea durhami (Keen, 1954) the senior secondary homonym of Acrilla durhami

genus. In addition, Zinsmeister’s specimen is missing its morphologically diagnostic aperture. His species might
not even be an epitoniid. [Undetermined].
Ferminoscala whitei-

SU 7983 [=CASG 7983]. Remarks: In the genus Scalina,  Addicott (1970: p. 57). [Epitoniidae].
....., SDSNH 5422 (ex
234 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Ficopsis cooperiana Stewart, 1926 [1927]. Moore 1968, SDSNH Occas. Pap. 15:26, pl. 
[=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH4169. Hypotype SDSNH 4147. Remarks:
In the genus ,  Squires (2014: p. 8). [Ficidae].
Ficopsis cooperiana
Meadows Lion Canyon quad. (1943), Ventura Co. UCR 4679. Hypotype UCR 4679/111. Remarks: In the genus
,  Squires (2014: p. 8). [Ficidae].

Ficopsis cooperiana 
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype UCLA
59277 [=LACMIP 10553]. Remarks: In the genus ,  Squires (2014: p. 8). [Ficidae].
Ficopsis crescentensis
Eocene. Reef Ridge, Reef Ridge quad., Kings Co. USGS 14494. Hypotype USNM 498718. Remarks: Ficopsis re-
mondii crescentensis Weaver and Palmer 1922,  Squires (1984: p. 30). In the genus Ficus (as Ficus remondii
(Gabb)),  Squires (2014: p. 11). [Ficidae].
Ficopsis hornii (Gabb, 1864). [Fusus (Hemifusus-
entiated Sacate/Gaviota fms., Eocene. Lompoc quad., Santa Barbara Co. UCMP B6963. Hypotype SU 9218 [=CASG
70166]. [Ficidae].
Ficopsis hornii (Gabb, 1864). [Fusus (Hemifusus

4741. Hypotype UCR. 4741/201. [Ficidae].
Ficopsis hornii (Gabb, 1864). [Fusus (Hemifusus           
Shale (Circle B Sandstone member), Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23687. Hypotype LACMIP
11642. [Ficidae].

11654.
Ficopsis remondi (Gabb, 1864). [Fusus (Hemifusus

missing]. [Ficidae].
          
Contra Costa Co. Lectotype (of Stewart 1926 [1927]: p. 376) ANSP 4184].
Ficopsis remondi (Gabb, 1864). [Fusus (Hemifusus
Fusus (Hemifusus) remondi)). Tejon Fm., Eocene. [Ft. Tejon,

[Ficidae].
Ficopsis remondi (Gabb, 1864). [Fusus (Hemifusus)]. Weaver and Kleinpell 1963. UCPG 43:191, pl
Fm., Eocene. Nojoqui Creek area, Lompoc quad., Santa Barbara Co. UCMP B6920. Hypotype SU 9220 [=CASG
70167]. [Ficidae].
Ficopsis remondi (Gabb, 1864). [Fusus (Hemifusus-
sion Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962.
Hypotype SDSNH 20723. [Ficidae].
Ficus remondii (Gabb, 1864). [Fusus (Hemifusus
Eocene. Rose Canyon, San Diego Co.. LACMIP 7185. Hypotype LACMIP 14483. [Ficidae].
         Ficopsis angulatus
Weaver, 1905) UCMP 11887.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 235

        
14484, 14486.

14485.


14488.
Ficopsis remondi crescentensis
Shale, Eocene. Rose Canyon, San Diego Co. SDSNH2363. Hypotype SDSNH 4162]. [Ficidae].
Ficopsis remondi crescentensis-
cene. Park Canyon area, San Guillermo quad. (1943), Ventura Co. UCR 4703. Hypotype UCR 4703/13. [Ficidae].
Ficopsis remondi crescentensis-
cene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969) Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype
LACMIP 6535. [Ficidae].
Ficopsis remondi crescentensis
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7477. [Ficidae].
Ficopsis remondi crescentensis
Eocene. Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 983 [=LACMIP 16110]. Hypotype LAC-
MIP 7713. [Ficidae].
Ficopsis remondi crescentensis          .

Hypotype LACMIP 11467. [Ficidae].
Ficopsis remondi crescentensis

Hypotype LACMIP 6535. [Ficidae].
Ficopsis remondi crescentensis
Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13426. [Ficidae].
Ficus gesteri
Eocene. Near Cañada de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP B7020. Hypotype SU 9221 [=CASG
70168]. [Ficidae].
Ficus gesteri        
Eocene. San Emigdio Canyon, San Emigdio Mtns., Kern Co. UCMP 3182. Holotype UCMP 11425, paratype UCMP
11426. [Ficidae].

Ficus (Trophosycon) kerniana (Cooper, 1894). [Agasoma?
Basal Jewett Sand, Miocene. Vicinity of Pyramid Hill, Caliente quad., Kern Co. UCMP B1660. Hypotype UCMP
15543. [Ficidae].



236 PALEOBIOS, VOLUME 38, FEBRUARY 2021
650116.



Ficus (Trophosycon) kerniana (Cooper, 1894). [Agasoma
  Remarks:
Determined here as in the genus Trophosycon. [Ficidae].
Ficus mamillata
Lion Canyon quad. (1943), Ventura Co. UCR 4715. Hypotype UCR 4715/9. [Ficidae].
Ficus mamillata Gabb, 1864. Squires 2014. NHMLAC CS 522:13  
Kern Co. Holotype ANSP 4230. [Ficidae].
   
LACMIP 14489.
Ficus mamillatus
Creek area, Lompoc quad., Santa Barbara Co. UCMP B6920. Hypotype SU 9222 [=CASG 70169]. [Ficidae].
Ficus modesta (Conrad, 1848). [Pyrula-
east of Irvington, La Costa Valley quad. (1943), Alameda Co. SU 3248. Hypotype SU/CASG 8470. [Ficidae].
Ficus cf.  (Conrad, 1848). [Pyrula
(Phacoides Sand member) (of Stinemeyer et al. 1959: pp. 12, 13), Miocene. Carneros Rocks quad., Temblor Range,
Kern Co. USGS M3578. Hypotype USNM 646625. [Ficidae].
Ficus cf.  (Conrad, 1848). [Pyrula
      
M3578. Hypotype USNM 646625. [Ficidae].

Ficus (Trophosycon) ocoyana (Conrad, 1855b). [Sycotopus     
Fiscus]. Castaic Fm., Miocene. Elizabeth Lake Canyon area, Los Angeles Co. CIT 279 [=LACMIP 7777]. Hypotype
LACMIP 4170. [Ficidae].
Ficus (Trophosycon) ocoyanus (Conrad, 1855b). [Syctopus
Ficus (Trophosycon) ocoyana)). Temblor Fm., Miocene. Poso Creek,
Kern Co. Hypotype (topotype) CAS 648 [=CASG 71566]. [Ficidae].
Ficus (Trophosycon) ocoyanus (Conrad, 1855b). [Sycotopus
Sobrante Fm., Miocene. Near junction of Norris Creek with Crow Creek, Hayward quarangle, Alameda Co. UCMP
A4565. Hypotypes UCMP 34159–34161. Remarks: Ficus (Trophosycon) kerniana (Cooper, 1894), Addicott
(1970: p. 74). [Ficidae].
-
potype LACMIP 4171].
Ficus
area, Santa Cruz Is., California Channel Ids. Hypotype SBMNH 60197. Remarks: Determined here as Scaphander
cf. (Mirascapha) costatus Gabb, 1864. [Scaphandridae].
Fissurella rixfordi Hertlein, 1928a. Stadum and Finger 2016. NHMLAC CS 524:38, 40, 
(Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 237
quad. SDSNH 4312. Hypotypes SDSNH 136497, 140187. [Fissurellidae].
Fissurella? stantoni Powell and Geiger, 2019 (n. sp.) PaleoBios
Santa Monica Mtns., Los Angeles Co. LACMIP 31511. Holotype LACMIP 14847. [Fissurellidae].
Forreria bartoni (Arnold, 1909 [1910]). [Trophon
Monterey Fm. (Saddleback Valley Limestone member), Miocene. Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4521. Hypotype SDSNH 83243. [Muricidae].
Forreria belcheri (Hinds, 1844–1845). [Chorus      
Pecten coalingaensis Zone, Pliocene. North Dome of Kettleman Hills, Kings Co. CASG 2364.
Hypotype CASG 5723. [Muricidae].
Forreria belcheri (Hinds, 1844–1845). [Chorus

[=USGS M1341). Hypotypes UCMP 15177, 15178. [Muricidae].
Forreria belcheri (Hinds, 1844–1845). [Chorus
Hills Fm. (of Dibblee 1968), Pliocene. Panorama Hills quad., Temblor Range, Kern Co. USGS M1059. Hypotype
USNM 646818. [Muricidae].
Forreria cancellaroides
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650114. [Muricidae].

15544.
Forreria carisaensis (Anderson, 1905). [Chorus
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2382. Hypotype USNM
254379. [Muricidae].

Forreria carisaensis (Anderson, 1905). [Chorus-

San Luis Obispo Co. USGS M2382. Hypotype USNM 254379. [Muricidae].

Co. USGS 4930. Hypotype USNM 254383.
Forreria cf. carisaensis (Anderson, 1905). [Chorus

Forreria carisaensis mirandaensis (Grant and Eaton in Eaton et al, 1941). [Trophon]. Adegoke 1969. UCPG 80:172,
Ranch quad., Fresno Co. UCMP B6532. Hypotype UCMP
36834. [Muricidae].
Forreria cf.  (Arnold, 1909 [1910]). [Trophon]. Durham and Addicott 1965. USGS PP 524-A:A12, pl.
     
Hypotype UCMP 36472. [Muricidae].
Forreria emersoni
Bravo Ranch quad. (1954), Kern Co. USGS M1591. Holotype USNM 650118; hypotypes USNM 650117, 650119.
[Muricidae].
Forreria magister (Nomland, 1916b). [Trophon-
chegoin Fm. (Patinopecten Zone), Pliocene. Vicinity of Arroyo Toreido, North Dome, Kettleman Hills, Kings Co.
USGS 12423. Hypotype USNM 495848. [Muricidae].

238 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Forreria magister (Nomland, 1916b). [Trophon
drawing, source specimen unknown. [Muricidae].
Forreria
area, La Panza Range, San Luis Obispo Co. USGS 4930. Hypotype USNM 254376. [Muricidae].
Forreria
8). Santa Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS 4930. Hypotype
USNM 254376. [Muricidae].
Indeterminate Forreria Powell et al. 2019b. PaleoBios-
vation, Berkeley Hills, Oakland, Alameda Co. UCMP IP13003. Hypotype UCMP 218695. [Buccinidae].
Fulgurofusus darraghi-
leocene. Vicinity of Meier and Runkle canyons, Calabasas quad., Simi Hills, Ventura Co. UCR 4573. Holotype UCR
4573/900. [Turbinellidae].

Fulgurofusus waringi
Meier Canyon, Calabasas quad., Simi Hills, Ventura Co. UCR 6671. Holotype UCR 6671/12. [Turbinellidae].
Fulgurofusus
Ventura Co. UCR 4573. Hypotype UCR 4573/900 (not UCR 4573/9 as in pl. caption). Remarks: Fulgurofusus
darraghi n. sp.,  Zinsmeister (1983b: p. 1296). [Turbinellidae].
Remarks: Fulgurofusus darraghi n. sp.,
 Zinsmeister (1983b: p. 1296).
Fulgurofusus n. sp. B. Zinsmeister 1983a. SEPM Guidebook Simi
Hills, Ventura Co. UCR 6671. Hypotype UCR 6671/12 (not 5571/12 as in pl. caption). Remarks: Fulgurofusus
waringi n. sp.,  Zinsmeister (1983b: p. 1297). [Turbinellidae].
Fulgoraria (Psephaea) -
vincial Stage” [=Santa Susana Fm.], Paleocene. Vicinity of Runkle and Meier canyons, Simi Hills, Calabasas quad.
(1967), Ventura Co. UCR 6602. Holotype UCR. 6602/1, paratypes UCR 6602/2, 6602/3. [Volutidae].


Fulgoraria zinsmeisteri
Fm., Paleocene. Simi Hills, Ventura Co. UCR 6602. Hypotype UCR 6602/1. [Volutidae].
Fusinus (Barbarofusus) arnoldi (Cossmann, 1903). [Fusus    
Fm., Pliocene. Orange Co. SDSNH 4080. Hypotype SDSNH 75914. [Fasciolariidae].
Fusinus teglandae-
ows, Lion Canyon quad. (1943), Ventura Co. UCR 4684. Hypotype UCR 4684/5. [Fasciolariidae].

4700/4.
Fusinus teglandae
Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242. Hypotype LAMCIP 6540. [Fascio-
lariidae].
Fusinus aff. 
Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6541. [Fasciolariidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 239
Fusinus waringi
Simi Hills, Ventura Co. UCR 6670. Hypotype UCR 6670/35 determined here as missing. [Fasciolariidae].
Fusitriton oregonensisTriton
Pliocene. Newport Beach, Orange Co. LACMIP 466. Hypotype LACMIP 5050 (not LACM 41 as in pl. caption).
[Ranellidae].
Fusitriton oregonensis1846). [Triton5, 3. Reworked shells,
Pliocene. Shells found in a modern-day shrimp trap off San Clemente Is., California Channel Ids. CASG 72573.
Hypotype CASG 72576. [Ranellidae].
Fusitriton scotiaensis (Martin, 1914). [Argobuccinum
Fm., Pliocene. Ferndale quad., Humboldt Co. UCMPA4556. Hypotypes UCMP 10577, 10578. [Ranellidae].
Fusiturricula (Crenaturricula) crenatospira (Cooper, 1894). [Surcula
   
UCMP 12166. [Drilliidae].
Fusiturricula (Crenaturricula) crenatospira (Cooper, 1894). [Surcula 
9h. Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP
7242 = LACMIP 22312]. Hypotype UCLA 59278 [=LACMIP 10558]. [Drilliidae].
Fusiturricula (Crenaturricula) crenatospira (Cooper, 1894). [Surcula 

16080]. Hypotype LACMIP 7492. [Drilliidae].
Fusiturricula (Crenaturricula) crenatospira domenginica   
Llajas Fm., Eocene. Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371
[=LACMIP 16115]. Hypotype LACMIP 6554. [Drilliidae].
Fusus (Levifusus  ()” calabasasensis
strata, uppermost Cretaceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis
Obispo Co. CASG 61599. [Hypotype CASG 61599.01]. [Fasciolariidae].
Fusus
Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8831 [=SBMNH 60198]. Remarks:
Determined here as indet. buccinid sp. [Buccinidae].
Galeodea californica
Ventura Co. LACMIP 7242. Hypotype LACMIP. 6530. [Cassidae].
….., Fig. 4C. Llajas Fm., Las Llajas Canyon, Simi Valley, Ventura Co. LACMIP 22312. Hypotype LACMIP 14834.
Galeodea (Caliagaleodea) californica
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP
6530. Remarks: Galeodea californica Clark, 1942,  Squires (2019: p. 9). [Cassidae].
Galeodea (Caliagaleodea) californica
Eocene. Simi Valley, Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 6530. Remarks: Galeodea cali-
fornica Clark, 1942,  Squires (2019: p. 9). [Cassidae].
Galeodea (Caliagaleodea) californica Clark, 1942. Squires 1988c. NHMLAC CS
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 987 [=LACMIP 16175]. Hypotype LACMIP 7710.
Remarks: Galeodea californica Clark, 1942,  Squires (2019: p. 9). [Cassidae].
Galeodea cf. 
Fm., Eocene. Orocopia Mtns., Canyon Spring NW quad. (1963), Riverside Co. CSUN 662 [=UCLA 3779) =LACMIP
23779]. Hypotype UCLA. 28988 [=LACMIP. 8836]. [Cassidae].
Galeodea louella (Squires and Advocate, 1986). [Phalium (Semicassis)]. Squires 2019. Paleobios 36:8–9, Fig. N.
240 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Maniobra Fm., Eocene. Orocopia Mtns., Riverside Co. LACMIP 16335. Paratype LACMIP 7167. See Phalium (Semi-
cassis) louella Squires and Advocate (1986). [Cassidae].

Galeodea cf. 
        
[Cassidae].
Galeodea (Gomphopages) meganosensis       
Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7474.
Remarks: Galeodea meganosensis,  Squires (2019: p. 4). [Cassidae].
Galeodea (Gomphopages) meganosensis
Fm., Eocene. Whitaker Peak area, Los Angeles Co. CSUN 845 [=LACMIP 16137]. Hypotype LACMIP 7474. Remarks:
Galeodea meganosensis,  Squires (2019: p. 4). [Cassidae].
Galeodea meganosensis Vokes, 1939. Squires 2019. PaleoBios
Peak, Los Angeles Co. LACMIP 40827. Hypotypes LACMIP 14829, 14830. [Cassidae].
Galeodea aff.   (Deshayes, 1835). [Cassidaria  

Hypotype LACMIP 7711. Remarks: Junior synonym of Galeodea meganosensis Vokes,  Squires (2019: p. 4).
[Cassidae].
Galeodea schencki
Gaviota fms., Eocene.Nojoqui Creek, west of hill 1477, Lompoc quad., Santa Barbara Co. UCMP B6893. Holotype
SU. 9210 [=CASG 70173], paratype SU. 9211 [=CASG 701740]. Remarks:Galeodeaschencki is a nomen dubium,
 Squires (2019: p. 14). [Cassidae].
Galeodea (Gomphopages) susanae        
Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115].
Hypotype LACMIP 6531. Remarks: Junior synonym of Galeodea sutterensis Dickerson, 1916,  Squires (2019:
p. 7). [Cassidae].
Galeodea (Mambrina) susanae. 24 (as Galeodea (Mamabrina)
in pl. caption). Juncal Fm., Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848.
Hypotype LACMIP 13425. Remarks: Junior synonym of Galeodea sutterensis Dickerson, 1916,  Squires (2019:
p. 7). [Cassidae].
Galeodea sutterensis Dickerson 1916. Squires 2019. PaleoBios
Simi Valley, Los Angeles Co. LACMIP 40371. Hypotypes LACMIP 14831,14832. [Cassidae].

14833.
Galeodea (Gomphopages) sutterensis
Eocene. Marysville Buttes, Marysville quad., Sutter Co. UCMP 1853. Holotype UCMP 11782. Remarks: Galeodea
sutterensis,  Squires (2019: p. 7). [Cassidae].
Galeodea sutterensis

Galeodea sp. cf. 
Fm., Eocene. Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3779 [=LACMIP 23779]. Hypotype UCLA
28988 [=LACMIP 8836]. Remarks: Galeodea cf.  Wrigley, 1934,  Squires and Advocate (1986: p.
857). [Cassidae].
Galeodea (Gomphopages) sutterensis
Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4658. Hypotype UCR 4658/5. Remarks:
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 241
Galeodea sutterensis,  Squires (2019: p. 7). [Cassidae].
Galeodea tuberculiformis G D. Hanna, 1924. Squires 2019. PaleoBios
north side Simi Valley, Los Angeles Co. LACMIP 16115. Hypotype LACMIP 6532. [Cassidae].

Galeodea 
del Capitan and Las Yegas Canyon, Gaviota quad., Santa Barbara Co. UCMP B6983. Hypotype SU 9214 [=CASG
70170] [Cassidae].

Hypotype SU 9213 [=CASG 70171].

SU 9212 [=CASG 70172].
Galerus
Galerus excentricus Gabb 1864)). Domengine
Fm. [as Tejon Fm.], Eocene. West of Coalinga, Fresno Co. Hypotype USNM 165643]. Remarks: Determined here
as Calyptraea diegoana (Conrad, 1855b). [Calyptraeidae].
Gastropod. Adegoke 1967a. Veliger 9(3):300, pl. 40,
Joaquin Rocks quad., Fresno Co. UCMP A9731. Hypotype UCMP 12158 (Massive bryozoan colony completely
encrusting a gastropod shell). Remarks: Determined here as possibly a naticid (family Naticidae).
Gemmula sp., aff. -
sana Fm., Eocene. Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 969 (=LACMIP 16894).
Hypotype LACMIP 12687. [Turridae].
Gemmula
21). Tesla Fm., Eocene. Midway quad., Alameda Co. UCMP. D8151. UCMP unnumbered hypotype lost,  C.
Throckmorton (personal communication, 2002). [Turridae].
Gibbula (Tumulus?) baileyi
Miocene. Kern River, Kern Co. USGS 6065. Holotype USNM 650048, paratype USNM 650047, hypotype USNM
650049. [Trochidae].
Gilbertina popenoei-
basas quad., Simi Hills, Ventura Co. UCR 6806. Holotype UCR 6806/4. [Ringiculidae].
Gilbertina
Ventura Co. UCR 6806. Hypotype UCR. 6806/4. Remarks: Described as Gilbertina popenoei n. sp.,  Zinsmeister
(1983b: p. 1301). [Ringiculidae].

Gisortia clarki
Co. UCMP 4052. Holotype UCMP 14844. [Cypraeidae].
Gisortia clarki
Co. UCMP 4052. Holotype UCMP 14844. [Cypraeidae].
Gisortia sp. cf. -
leocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717 =
CASG 61666]. Hypotype SU 10235 [=CASG 61666.06]. Remarks: Determined here as Sphaerocypraea martini
(Dickerson, 1914a). [Ovulidae].
Gisortia clarkiGisortia-
tion). Juncal Fm., Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 848 [=LACMIP 16243]. Hypotype
242 PALEOBIOS, VOLUME 38, FEBRUARY 2021
LACMIP 7466. [Cypraeidae].
Gisortia (Megalocypraea) clarki
Las Llajas Canyon, Santa Susana Mtns., Ventura Co. UCMP 4052. Holotype UCMP 14844. [Cypraeidae].
Globularia (Eocernina) hannibali (Dickerson, 1914c). [Natica]. Stewart 1946. USGS 205-C:
Sandstone, Eocene. Reef Ridge, just southeast of Big Tar Canyon, Reef Ridge quad., Kings Co. USGS 14495. Hypo-
type USNM 498719. Remarks: In the genus Eocernina,  Marincovich (1977: p 229). [Naticidae].
Globularia (Eocernina) hanniballi (Dickerson, 1914c). [Natica
Eocene. Park Canyon, San Guillermo quad. (1943), Ventura Co. UCR 4699. Hypotype UCR 4699/18. Remarks:
In the genus Eocernina,  Marincovich (1977: p. 229). [Naticidae].
      
UCR 4658/11.
Globularia

Glyphostoma carinata
Oil Center quad. (1950), Kern Co. UCMP B1598. Hypotype UCMP 37097. [Clathurellidae].

Goniobasis kettlemanensis-
quin Fm. (Pecten and Trachycardium Zones), Pliocene. Arroyo Pino, North Dome, 1500, Kettleman Hills, Kings Co.
USGS 12360. Hypotype USNM 495759. Remarks: In the genus Juga,  Taylor (1966: p. 119). [Pleuroceridae].

Hypotypes USNM 495760, 495767.
Goniobasis kettlemanensis   
Pliocene. Reef Ridge quad., Kings Co. UCMP B6536. Hypotype UCMP 12209. Remarks: In the genus Juga, 
Taylor (1966: p. 119). [Pleuroceridae].
Goniobasis olequaensis (Arnold and Hannibal in Hannibal, 1912). [Ambloxus]. Squires and Deméré 1991. SEPM
Goiobasas olequahensis). Friars Fm., Eocene. San Diego Co. SDSNH 3278. Hypotype
SDSNH 28379. [Pleuroceridae].
Grandicrepidula princeps (Conrad, 1855b). [Crepidula
Newhall area, Los Angeles Co. LACMIP 17921. Hypotype LACMIP 14387. Remarks: Supplemental description

Gyrineum judithi-
cene. Calabasas quad., Simi Hills, Ventura Co. UCR 6893. Holotype UCR. 6893/1 (not UCR 6899/100 as in text
and pl. caption), paratype UCR 6893/5 (not UCR 6899/101 as in pl. caption). Remarks: Species name emended
to judithae
Gyrineum judithae Zinsmeister, 1983b. Beu 1988. Saito Ho-on Simi Conglom-
erate, Paleocene. Simi Hills, Ventura Co. UCR 6893]. Plastoholotype [cast from UCR 6893/1, not UCR 6899/100
as cited in pl. caption], NZGS WM 11835. Remarks: Species name emended to judithae
1–3). [Ranellidae].
Gyrineum lewisii
of Boulder Creek, adjacent to the Wiggins Ranch, Humboldt Co. UCMP A4233. Hypotype UCMP 34175]. Remarks:
Junior synonym of Mediargo mediocris (Dall, 1909),  Smith (1970: p. 504). [Ranellidae].
Gyrineum mediocre lewisii

locality. Hypotypes (3) USNM 560075. Remarks: Junior synonym of Mediargo mediocris (Dall, 1909),  Smith
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 243
(1970: p. 504). [Ranellidae].
Gyrineum
Ventura Co. UCR 6893. Hypotype UCR 6893/1 (not UCR.7311/9 as cited in pl. caption). Remarks: Described as
Gyrineum judithi n. sp.,  Zinsmeister 1983b: p. 1294; species name emended to judithae by Beu (1988: pl. 4,

Gyrodes robustus
Susana Fm.], Paleocene. Simi Hills, Ventura Co. SU 2697 [=CAS 61667]. Lectotype (of Marincovich, 1977:244)
SU.147 [=CAS 61667.09], paralectotype SU 148 [=CAS 61667.10]. [Naticidae].

Gyrodes robustus-
leocene. Simi Hills, Ventura Co. UCR 6899. Hypotype UCR 6899/2. [Naticidae].
Gyrodes Gyrodes) robustus
Fm., Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCLA 2582 [=LACMIP 22582]. Hypotype UCLA 59730
[=LACMIP 10041]. [Naticidae].
Haliotis assimilis
Drive quarry, Balboa Park, San Diego, San Diego Co. LACMIP 107. Hypotype LACMIP 14848 determined here as
missing]. [Haliotidae].
Haliotis elsmerensis

CASG 286]. Remarks: Determined here as Haliotis koticki Hertlein, 1937. [Haliotidae].
Haliotis lasia

Holotype USNM 371767, paratype USNM 371768]. [Haliotidae].
Haliotis palaea
Monica Mtns., Los Angeles Co. CIT 52 [=LACMIP 7830]. Holotype LACMIP 4156. [Haliotidae].
Haliotis (Padollus) pourtalesii Dall, 1881. Schremp 1981. Jour. Paleo
Fm.], Pliocene. San Gorgonio Pass, Riverside Co. UCMP A3197. Hypotype UCMP 16074. [Haliotidae].
Haliotis pourtalesii
 
UCMP 16074. [Haliotidae].
Haliiotis rufescens

Pliocene. Gillibrand Quarry, Santa Susana Mtns., Ventura Co. LACMIP 29227. Hypotype LACMIP 12660. [Haliotidae].
Haliotis walallensis
-
cene. Hills south of the Tijuana River, San Diego Co. LACMIP 305c [=LACMIP 16817]. Hypotype LACMIP 12658.
[Haliotidae].
Haliotis

Arroyo Santa Rosa, near Moorpark, Ventura Co. LACMIP 16896. Hypotype LACMIP 12659. [Haliotidae].
Haliotis 
17921. Hypotype LACMIP 14375. [Haliotidae].
Haminoea virescens (Sowerby I, 1833). [Bulla     
Olcese Sand, Miocene. Kern River area, Kern Co. CASG 2147. Hypotype CASG 12943. [Haminoeidae].
244 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Hastula gnomon-

[=CASG 7984]. [Terebridae].
....., LACMIP 16140 (exex UCLA 46034).
....., SDSNH 5422 (ex
Hastula gnomon
River, Kern Co. USGS 6641. Hypotype USNM 650240. [Terebridae].


Hemitoma (Montfortia) cantonensis
Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=LACMIP 16114]. Holotype LACMIP
7440. Remarks: Determined here as in the genus Montfortia. [Fissurellidae].
Hemitoma (Montfortia) emarginata (Blainville, 1825). [Emarginula]. Schremp 1981. Jour. Paleo. 55(5):1125–1127,
pl. 1,  Jacinto quad., Riverside Co. UCMP A1415.
Hypotype UCMP 16075 (not 16074 as in pl. caption). Remarks: Determined here as in the genus Montfortia.
[Fissurellidae].

Heteroterma(?) acrita
Meier Canyon, Santa Susana quad., Simi Hills, Ventura Co. UCLA 2307 [=LACMIP 22307]. Holotype LACMIP 7598.
[Perissityidae].

61666. Paratype CASG 61666.03.

22330]. Paratype LACMIP 7600.
Heteroterma gabbi
7. Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6899. Hypotype UCR 6899/2 (not UCR. 6899/8 as
in pl. caption). [Perissityidae].
Heteroterma gabbi -
leocene. Calabasas quad., Simi Hills, Ventura Co. UCR 6899. Hypotypes UCR 6899/2 (not UCR 6899/8 as in pl.
caption). Remarks:Heteroterma acrita n. sp.,  Saul (1988a: p. 14). [Perissityidae].
Heteroterma striata
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073. Hypotype CASG 10245 [=CASG.
61666.11]. Remarks: Pyropsis judithae n. sp.,  Saul (1988b: p. 886). Not a Pyropsis, Squires (2011: p.
1204). [Perissityidae].

UCMP 14209. Remarks: Pyropsis judithae n. sp.,  Saul (1988b: p. 14).
Heteroterma striata-
cene. Simi Hills, Ventura Co. UCLA 3117 [=LACMIP 23117]. Hypotype UCLA 59294 [=LACMIP 10979]. Remarks:
Pyropsis judithae n. sp.,  Saul (1988b: p. 14). Not a Pyropsis, Squires (2011: p. 1204). [Perissityidae].
Heteroterma striata
Simi Hills, Ventura Co. UCLA 3117 [=LACMIP 23117]. Hypotype UCLA 59294 [=LACMIP 10979]. Remarks: Py-
ropsis judithae n. sp.,  Saul (1998b: p. 14). Not a Pyropsis, Squires (2011: p. 1204). [Perissityidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 245
 
Remarks: Pyropsis judithae n. sp.,  Saul (1988b: p. 14).
Heteroterma trochoidea Gabb, 1869b. Zinsmeister 1983b. Jour
1969, pl. Lectotype (of Stewart 1926 [1927]: p.
423) ANSP 4237. [Perissityidae].
Heteroterma trochoidea-
cene. Meier Canyon, Santa Susana quad., Ventura Co. UCLA 2330 [=LACMIP 22330]. Hypotype LACMIP 7596.
[Perissityidae].


Heteroterma
Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA 6525 [=LACMIP 26525].
Hypotype LACMIP 7562. [Perissityidae].
Hexaplex whitneyi (Gabb, 1864). [Tritonium
Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4715. Hypotype UCR 4715/3. [Muricidae].
Hipponix pristinus      
Thousand Oaks quad., Simi Hills, Ventura Co. UCR 6887. Holotype UCR 6887/200, paratype UCR 6887/201.
[Hipponicidae].
Hipponix    
Hills, Ventura Co. UCR 6887. Hypotype UCR 6887/200 (not UCR 6887/2 as in pl. caption). Remarks: Described
as Hipponix pristinus n. sp,  Zinsmeister (1983b: p. 1292). [Hipponicidae].
Homalopoma (Leptothyropsis) maiquetiana (Weisbord, 1962). [Arene]. Schremp 1981. Jour. Paleo. 55(5):1133, pl.

A1415. Hypotype UCMP 16088 (not UCMP 16087 as in pl. caption). [Colloniidae].
Homalopoma cf.  (Dall, 1871). [Leptothyra]. Woodring and Bramlette 1950 [1951]. USGS PP 222:70,
 
Hypotype CIT 4557 [=LACMIP 4166]. [Colloniidae].
Homalopoma umpquaensis
Lockwood Valley quad. (1943), Ventura Co. UCR 4676. Hypotype UCR 4676/6. [Colloniidae].
Homalopoma umpquaensis domenginensis-

7694. [Colloniidae].
Homalopoma umpquaensis domenginensis       

LACMIP 7694. [Colloniidae].
Homalopoma wattsi (Dickerson, 1916). [Monodonta]. Squires 2008. NHMLAC CS
Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13416.
[Colloniidae].
Homalopoma aff.  (Dickerson, 1916). [Monodonta 
Maniobra Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082].
Hypotype LACMIP 11453. [Colloniidae].
Homoploma* paucicostatumLeptothyra
Pliocene. Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14379. Remarks: Error for Homol-
apoma. [Colloniidae].
246 PALEOBIOS, VOLUME 38, FEBRUARY 2021
-

hypotype lost,  C. Throckmorton (personal communication, 2002).
Iselica fenestrata (Carpenter, 1864b) var. [Isapis]. Woodring and Bramlette 1950 [1951]. USGS PP 222:70, pl. 13,
    
Hypotype CIT 4558 [=LACMIP 4214], determined here as missing. [Amathinidae].
Janiopsis sanmarcosensis-
mar Fm., Eocene. Near Vista and San Marcos, San Diego Co. UCR. 4865. Holotype UCR 4865/11, paratype UCR
4865/110. [Buccinidae].
Jaton cf.  (Dall, 1899a). [Pteronotus
9, 11. Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. CIT. 869 [LACMIP 6108]. Hypotype
CIT. 4555 [=LACMIP 4198]. [Muricidae].
Jaton cf.  (Hinds, 1844). [Murex
Fm. (Pecten Zone), Pliocene. Vicinity of Arroyo Finito, North Dome, Kettleman Hills, Kings Co. USGS 12343. Hy-
potype USNM 495756. Remarks: In the genus Pteropurpura,  McLean (1978: p. 42). [Muricidae].
Jaton festivus (Hinds, 1844) n. var. [Murex
Fm. (Pecten Zone), Pliocene. La Lomica, North Dome, Kettleman Hills, Kings Co. USGS 12365 [=USGS 12767].
Hypotype USNM 495762. Remarks: In the genus Pteropurpura,  McLean (1978: p. 42). [Muricidae].
Kelletia cf.  (Forbes, 1852). [Fusus      
Pancho Rico Fm., Miocene/Pliocene. Sarah Canyon area, Wunpost quad., Monterey Co. USGS M1934. Hypotype
USNM 649096. Remarks: Kelletia kellettii (Forbes, 1852),  Addicott (1970: p. 91) [Buccinidae].
Kelletia kettlemanensis (Arnold, 1909 [1910]). [Thais
65-16. Etchegoin Fm. (Upper Mulinia Zone), Pliocene. Kettleman Hills, Kings Co. CASG 27072. Hypotype CASG
5902. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Kelletia lorata Addicott, 1970 (n. sp.). USGS    Miocene. Oil
Center quad. (1950), Kern Co. UCMP B1598. Holotype UCMP 32834. [Buccinidae].
Kelletia posoensis (Anderson and Martin, 1914). [Siphonalia
Basal Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650152.
[Buccinidae].
Kelletia cf.  (Anderson and Martin, 1914). [Siphonalia
(cast). Lower Jewett Sand, Miocene. Northwest of Pyramid Hill, ‘Vaqueros’, Kern Co. UCLA WR163 [=LACMIP
16252]. Hypotype UCLA 45770 [=LACMIP 8435]. [Buccinidae].
Kelletia vladimiri Kanakoff, 1954 (n. sp.). BSCAS 53(2):114–117, pls. 29–31. Originally referred to as Pico Fm.
(Pliocene) but most likely is Castaic Fm. (Miocene). Mint Canyon quad., Los Angeles Co. LACMIP 291. Holotype
LACMIP 1097, paratypes LACMIP 1098, 1099. [Buccinidae].

Kelletia    
Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM 646620. [Buccinidae].

Kleinacteon moodyi (Dickerson, 1916). [Acteon 
   
UCMP 3117]. [Acteonidae].
Knefastia cf.  (Kiener, 1840, ex Valenciennes MS). [Pleurotoma]. Woodring et al. 1946. USGS PP 207:27,

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 247
Hills, Los Angeles. USGS 13127. Hypotype USNM 497079. [Pseudomelatomidae].
Knefastia garcesana
Miocene. Kern River area., Rio Bravo Ranch quad. (1954), Kern Co. USGS M1607. Holotype USNM 650255. [Pseu-
domelatomidae].
Lacuna carpenteri
Rio Bravo Ranch quad. (1954), Kern Co. USGS M1600. Hypotype USNM 650068. [Littorinidae].

Lacunaria striata (Gabb, 1869b). [Ampullina
11). Martinez Fm., Paleocene. Martinez, Contra Costa Co. Lectotype (of Stewart 1926 [1927]:339) ANSP 4241,

Fresno Co. SU 422. Hypotype CASG 10275
[=CASG 72659].
Sinum
dickersoni Waring, 1917) SU 163 [=CASG 69891].
Laevityphis (Laevityphis) antiquus (Gabb, 1864). [Typhis      
Eocene. Devil Canyon, Santa Susana quad. (sec. cor. is in Oat Mtn.quad.), Los Angeles Co. CSUN 371 [=LACMIP
16115]. Hypotype LACMIP. 6536. [Muricidae].
Lapparia eomagna        
Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 22312. Hypotype LACMIP 12740. [Volutidae].
Latirus” cf.  (Dickerson, 1914a). [Tritonium
11, 12. Lodo Fm., Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073
[=UCMP A9717]. Hypotype SU 10239 [=CASG 61666.18]. Remarks: Error for buwaldi. [Fasciolariidae].
Leiorhinus? reedi     
Kettleman Plain quad., Kings Co. USGS 14482a. Holotype USNM 498723. [Strombidae].
Liotia carinata Carpenter, 1857c. [Liotia-
cene. Humphreys quad. (1932, RP 1946), Los Angeles Co. CIT 1663 [=LACMIP 2555. Hypotype LACMIP 4158,
determined here as missing. Remarks: Arene lurida (Dall, 1913),  J.H. McLean (personal communication,
2005). [Liotiidae].
Lirabuccinum dalli (Clark, 1918). [Searlesia 
10a, 10b. San Ramon Fm., Miocene. San Pablo Creek, Contra Costa Co. UCMP 2754. UCMP holotype 11215 (not
UCMP 12115 as in Keen and Bentson 1944: p. 192), paratype 11214. [Buccinidae].
Lirobittium asperum (Gabb, 1866). [Turbonilla-
whall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14382. [Cerithiidae].
Lirofusus ashleyi Arnold, 1908 (n. sp.). Proc. USNM 34(1617):372. San Lorenzo Fm., Miocene. San Lorenzo River,
ex CIT 3710]. [Fasciolariidae].
Littorina mariana
(Upper Mya Zone), Pliocene. Middle Dome, Las Alturas, Kettleman Hills, Kings Co. USGS 12697. Hypotype USNM
495285. [Littorinidae]. Remarks: Junior synonym of Littorina petricola Arnold, 1908,  Reid (1996: p. 92).

 Littorina Zone), Pliocene. North Dome area, El Piso on west side of La Cima,
Kettleman Hills, Kings Co. USGS 12391. Hypotypes USNM 495790, 495791.
Littorina mariana Arnold, 1909 [1910]. G D. Hanna
248 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Fm., Pliocene. Kettleman Hills, Kings Co. CASG 26811. Hypotype CASG 5722. Remarks: Junior synonym of Litto-
rina petricola Arnold, 1908,  Reid (1996: p. 92). [Littorinidae].
Littorina mariana
source specimen unknown. [Littorinidae]. Remarks: Junior synonym of Littorina petricola Arnold, 1908, 
Reid (1996: p. 92).
Littorina cf. 
Fm. (Graciosa Member), Pliocene. Graciosa Ridge near Orcutt, Santa Barbara Co. USGS 14619. Hypotype USNM.
560118. [Littorinidae].
Littorina petricola
Alto quad., Santa Clara Co. USGS M1715. Hypotypes USNM 650947–650949. [Littorinidae].
Littorina (Littorina) petricola
Kettleman Hills, Kings Co. USNM 4718. Holotype (of Littorina mariana Arnold, 1909 [1910]) USNM 165481.
Remarks: Arnold, 1909 [1910] originally listed this locality as Etchegoin Fm.
Littorina mari-
ana var. alta Arnold, 1909 [1910]. USNM 4730. Remarks: Arnold, 1909 [1910] originally listed this locality as
Etchegoin Fm.

Hypotype LACMIP 20697.1 and LACM 14876.
Littorina Zone, La Cima quad., Kings Co. UCMP D6369. Hypotypes UCMP
110605–110609.

Littorina (Littorina) remondii
Kirker’s Pass, Contra Costa Co. Lectotype (of Keen and Bentson, 1944: p. 167) ANSP 4334, paralectotypes (2)
ANSP 77549.

Littorina pittsbur-
gensis Clark, 1915) UCMP 11649.
Littorina pittsburgensis Clark, 1915) UCMP
11620. Remarks: Listed as Neroly Fm. in UCMP on-line database.
Littorina Fowkes 1982. Coalinga Guidebook, p. 33,  
2 (both as Littorina mariana var. alta n.var.)). San
Joaquin Fm. [as Etchegoin Fm.], Pliocene. Central Kettleman Hills, Kings Co. Hypotype USNM 165487]. Remarks:
Determined here as Littorina mariana Arnold, 1909 [1910]. [Littorinidae].
Lora turricula (Montagu, 1803). [Murex
Ferndale quad., Humboldt Co. UCMP B7950. Hypotype UCMP 37797. Remarks: Determined here as in the genus
Propebela. [Mangeliidae].
Lottia asmi (Middendorff, 1848a). [Acmaea    

LACMIP 8406. [Lottidae].
Lottia digitalis (Eschscholtz in Rathke, 1833). [Acmaea
Beach Fm., Pliocene/Pleistocene. Vicinity of Moonstone Beach Trinidad quad., Humboldt Co. UCMP B7348. Hy-
potype determined here as missing. [Lottidae].
Loxotrema turritum
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 249
Canyon, San Diego Co. SDSNH2314. Hypotype SDSNH 4181]. [Melanopsidae].
Loxotrema turritum
Reyes Peak quad. (1943), Ventura Co. UCR 4747. Hypotype UCR 4747/401. [Melanopsidae].
Loxotrema turritum
      
4865/202. [Melanopsidae].
Loxotrema turritum      
Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 671 [=LACMIP 16076]. Hypotype LACMIP 11459.
Remarks: Loxotrema turritum Gabb, 1868,  Squires (1998: p. 299). [Melanopsidae].
Loxotrema turritum
Fork Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963]. Hypotype LACMIP
7162. [Melanopsidae].

Loxotrema turritum
fork of Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963]. Hypotype
LACMIP 12452. [Melanopsidae].

Loxotremafrom Arnold 1909 [1910], pl. 4,
Loxotrema turritum Gabb, 1868)). Do-
mengine Fm., Eocene [as Tejon Fm.]. West of Coalinga, sec. 26, T20S, R14E, Fresno Co. Hypotype USNM 165649].
Remarks: Loxotrema turritum Gabb, 1868,  Squires (1999c: p. 299). [Melanopsidae].
Lucunaria* striata (Gabb, 1869). [Ampullina-
sana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6805. Hypotype UCR 6805/5. Remarks: Error for Lacunaria.
[Naticidae].
Lunatia cf.  (Gould, 1847). [Natica
Fm. (Pecten Zone), Pliocene. Vicinity of Arroyo Vadoso and Arroyo Largo, North Dome, Kettleman Hills, Kings Co.
USGS 12331. Hypotype USNM 495761. Remarks: In the genus Euspira,  McLean (2007: p. 737). [Naticidae].
Siphonalia Zone), Pliocene. Los Jinetes, North Dome, Kettleman Hills, Kings Co. USGS
12393. Hypotype USNM 495813.
Lunatia cf.  (Gould, 1847). [Natica
Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. USGS 14769. Hypotype USNM 560071.
Remarks: In the genus Euspira,  McLean (2007: p. 737). [Naticidae].
Lunatia
Remarks:
and Pliocene localities, the exact locality of Lunatia of Fowkes 1982 is unknown. [Naticidae].
Glycymeris Zone”), Miocene/Pliocene. Southwest of Coalinga,

Lyria andersoni Waring, 1917 (n. sp.). Pr.CAS 7(4):97. Llajas Fm., Eocene. McCray Wells, Ventura Co. LACMIP 16272.
ex CIT].
Lyria andersoni
Aliso Creek, Los Angeles Co. CASG 364. Hypotype CASG 5712. [Volutidae].
Lyria andersoni        
Mtn. quad. (1952, PR 1969), Los Angeles Co. CSUN 498 [=LACMIP 16189]. Hypotype LACMIP 6550. [Volutidae].
250 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Lyria andersoni-
ker Peak quad. (1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype LACMIP 7490. [Volutidae].
Lyria andersoni
Ventura Co. CSUN 498 [=LACMIP 16189]. Hypotype LACMIP 6550. [Volutidae].
Lyria andersoni
Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LACMIP 7716. [Volutidae].
Lyria andersoni Waring, 1917. Squires., Eocene. Maniobra
Valley, Red Canyon quad. (1986), Riverside Co. CSUN 675 [=LACMIP 16151]. Hypotype LACMIP 11471. [Volutidae].
Lyria andersoni
Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13429. [Volutidae].
Lyria hannibali-
cene. Simi Hills, Ventura Co. UCLA 2330 [=LACMIP 22330]. [Hypotype UCLA 59580 [=LACMIP 10981). [Volutidae].
Lyriscapha* lajollaensis (M.A. Hanna, 1927). [Pejonia
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 672 [=LACMIP 16274]. Hypotype
LACMIP 11472. Remarks: Error for Lyrischapa. [Volutidae].
Lyrischapa lajollaensis (M.A. Hanna, 1927). [Pejonia
Shale, Eocene Torrey Pines State Park, Del Mar quad. (1967), San Diego Co. UCMP 5062 [=UCR 4922]. Holotype
UCMP 30911 (missing  Givens 1979: p. 124), neotype (of Givens 1979: p. 124) UCMP 14634. Remarks: Ho-


UCR 7719/4.


Lyrischapa lajollaensis (M.A. Hanna, 1927). [Pejonia
Devil Canyon, Santa Susana quad. (sec. cor. is in the Oat Mtn. quad.), Ventura Co. CSUN 373 [=LACMIP 16211].
Hypotype LACMIP 6551. [Volutidae].
Lyrischapa lajollaensis (M.A. Hanna, 1927). [Pejonia 
Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype LACMIP 7491.
[Volutidae].
Lyrischapa lajollaensis (M.A. Hanna, 1927). [Pejonia
Fm., Eocene. Simi Valley, Ventura Co. CSUN 373 [=LACMIP 16211]. Hypotype LACMIP 6551. [Volutidae].
Macrarene diegensis

LACMIP 1300–1656]. [Liotiidae].

Macron aethiops (Reeve, 1847). [Buccinum]. Addicott 1970 Sand,
Miocene. Woody quad., Kern Co. SU 26854. Hypotype SU 9940, determined here as missing. [Buccinidae].
-
gest the material is from the lower part of the Olcese Sand exposed on the south side of Poso Creek, Woody quad.,
Kern Co. UCLA MB1400 [=LACMIP 16278]. Hypotype UCLA 45769 [=LACMIP 8436].
Malea ringens (Swainson, 1822). [Cassis
[=Latrania Fm.], Miocene/Pliocene. Coyote Mtn., Imperial Co. Line drawing, source specimen unknown. [Cassidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 251
Malea ringens (Swainson, 1822). [Cassis
3.2, 3.8B. Latrania Fm., Miocene/Pliocene. Alverson Canyon, Imperial Co. SDSNH 345. Hypotype SDSNH 12234.
[Cassidae].

Mangelia (Notocytharella?) hartensis
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650265. [Mangeliidae].
Mangelia (Agathotoma) howei
Round Mtn. Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M2480. Hypotype USNM 650266.
[Mangeliidae].
Mangelia (Cacodaphnella kernensis (Anderson and Martin, 1914). [Mangilia]. Keen 1943. Tr. SDSNH 10(2):47, 48,

72005, listed as Manella (Cacodaphnellakernensis and as a bivalve in the CASG online database. [Mangeliidae].
Mangelia (Notocytharella) kernensis (Anderson and Martin, 1914). [Mangilia]. Addicott 1970. USGS PP 642:135–136,

USNM 650267–650270. [Mangeliidae].

Margarites semiornata

6876/800c (not UCR 6873/81 (as in text) or UCR 6873/800 (as in plate caption), paratype UCR 6876/801 (not
UCR 6873/801 as in pl. caption). Remarks: An apparent inadvertent illustration of two specimens labeled as the

two specimens as syntypes. A measurement of 14 mm noted in the text is meaningless as personal measurements
made by author LTG yielded larger sizes for both specimens. [Margaritidae].
Margarites n. sp. Zinsmeister 1983a. SEPM Paleocene. Simi
Hills, Ventura Co. UCR 6876. Hypotypes UCR 6876/800a, 6876/800c (not UCR 6876/7 as cited in pl. caption).
Remarks: Described as Margarites semiornata n. sp.,  Zinsmeister (1983b: p. 1289). [Margaritidae].
Marginella cf. -
cene. Humphreys quad. (1932, RP 1946), Los Angeles Co. CIT 2069 [=LACMIP 2558]. Hypotype [=LACMIP 4197].
[Marginellidae].
Mayena kewi (Dickerson, 1915). [Nyctilochus     
Eocene. Grapevine Canyon, Kern Co. UCMP 458. Holotype UCMP 11052. Remarks: In the genus Ameranella, 
Beu (1988: p. 82). [Ranellidae].
Mayena” (Nyctilochus) kewi (Dickerson, 1915). [Nyctilochus  Mayena
kewi). Tejon Fm., Eocene. Grapevine Canyon, Kern Co. LACMIP 22340. Line drawings, source specimen unknown.
Remarks: In the genus Ameranella,  Beu (1988: p. 82). [Ranellidae].
Mediargo dilleri (Anderson and Martin, 1914). [Argobuccinum-
per Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1602. Hypotype USNM 650109. Remarks:
In the genus Fusitriton,  Smith (1970: p. 502). [Ranellidae].



Hypotype CASG 12942.
Mediargo mathewsoni (Gabb, 1866). [Ranella
Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650108. [Ranellidae].
252 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Mediargo mathewsonii (Gabb, 1866). [Ranella
 Smith (1970: p.
510). [Ranellidae].

Bursa
trampasensis Clark, 1915) UCMP 11611.
     
Holotype (of Tritonium newsomi Arnold, 1908) USNM 165436 (not USNM 16543) as in pl. caption].
Mediargo mediocris (Dall, 1909). [Gyrineum]. Terry 1968. Veliger
(reworked. [LACMIP 16441]. Hypotype LAC-
MIP 1190 (not LACMIP 2155.96 as cited). [Ranellidae].

Gyrineum
lewisii Carson, 1926) SU 138 [=CASG 61893.06].

Mediargo mediocris (Dall, 1909). [Gyrineum
Pliocene. From a well in San Diego, San Diego Co. Hypotype CASG 11689. [Ranellidae].

[=CASG 61893.01], paratype SU 138 [=CASG 61893.06] (both of Gyrineum lewisii Carson, 1926).
ex SU 31625, [=CASG 70385].
Megasurcula carpenteriana (Gabb, 1865). [Cryptoconus
e. San Diego Fm., Pliocene. Reynard Wy., San Diego, San Diego Co. SDSNH429. Hypotype SDSNH 4169]. [Pseu-
domelatomidae].
Megasurcula carpenterianaCryptoconus]. Woodring and Bramlette 1950 [1951]. USGS

Co. USGS 14618. Hypotype USNM 560128. [Pseudomelatomidae].
Megasurcula carpenteriana tryoniana (Gabb, 1866) strongly shouldered form. [Pseudotoma]. Woodring and Bram-

Hypotype USNM 560050. [Pseudomelatomidae].
Megasurcula carpenteriana tryoniana (Gabb, 1866). [Pseudotoma]. Weaver 1971. West (LA Times), April 18, 1971:11,
th and Flower
streets, downtown Los Angeles, Los Angeles Co. LACMIP1219]. Hypotypes (3) current whereabouts unknown.
[Pseudomelatomidae].
Megasurcula condonana (Anderson and Martin, 1914). [Bathytoma
Basal Jewett Sand, Miocene. Pyramid Hill, Kern Co. CASG 37468. Hypotype CASG 12940. [Pseudomelatomidae].

Megasurcula howei G D. Hanna and Hertlein, 1938. Miocene.
Float collection, Oil Center quad. (1950), Kern Co. UCMP B1597. Hypotype UCMP 36560. [Pseudomelatomidae].
Megasurcula howei
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650254. [Pseudomelatomidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 253


Megasurcula keepi (Arnold, 1907). [Pleurotoma

[Pseudomelatomidae].


Megasurcula keepi (Arnold, 1907). [Pleurotoma-

Megasurcula keepi (Arnold, 1907). [Pleurotoma
Monterey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San
Juan Capistrano quad. SDSNH 4312. Hypotype SDSNH 83240. [Pseudomelatomidae].
Megasurcula remondi (Gabb, 1866). [Metula
Pliocene. Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650978. [Pseudomelatomidae].
Megasurcula remondi (Gabb, 1866). [Metula
Fm., Pliocene. Santa Cruz Co. SCCM hypotype determined here as missing. [Pseudomelatomidae].
Megasurcula remondi (Gabb, 1866). [Metula
Capitola, Santa Cruz Co. Hypotype CASG 66125.01 determined here as missing. [Pseudomelatomidae].
Megasurcula tryoniana (Gabb, 1866). [Pleurotoma
Pliocene. Coalinga quad., Fresno Co. UCMP B6546. Hypotype UCMP 36909. [Pseudomelatomidae].
Megasurcula wynoocheensis (Weaver, 1912). [Turris 
Round Mtn. Silt, Miocene. Kern River, Caliente quad. (1914), Kern Co. UCMP B1618. Hypotype UCMP 37094.
[Pseudomelatomidae].
 Mtn. Silt, Miocene. Temblor locality 1, Caliente quad., Kern Co. UCMP
2713. Hypotype UCMP 15906.
Megasurcula      
Pliocene [=Wilson Grove Fm.], Miocene/Pliocene. Santa Rosa Valley, Sonoma Co. Hypotype CASG determined
here as missing. [Pseudomelatomidae].
Megathura
Kern Co. CASG 245. Hypotype CASG 760. [Fissurellidae].
Megistostoma gabbianum (Stoliczka, 1868). [Bullaea
Canyon Fm.” [=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH2367. Hypotype SDSNH 4165.
Remarks: Megistostoma striata Gabb, 1864, see Groves and Squires 2018: p. 103 for the complicated nomencla-
tural history of this species. [Philinidae].
Megistostoma gabbianum (Stoliczka, 1868). [Bullaea
Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 458 [=LACMIP 16280]. Hypotype LACMIP
6553. Remarks: Megistostoma striata Gabb, 1864, see Groves and Squires 2018: p. 103 for the complicated no-
menclatural history of this species. [Philinidae].
Melanatria markleyensis (Clark, 1938). [Thiara     
Sandstone, Eocene. Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1445 [=LACMIP 24259]. Hypotypes LAC-
MIP 12450, 12451. [Thiaridae].
Mesalia clarki (Dickerson, 1914a). [Turritella
254 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Fm., Paleocene. Simi Hills, Ventura Co. UCR 4570. Hypotype UCR 4570/14. Remarks: Mesalia martinezensis (Gabb,
1869b),  Squires and Saul (2007: p. 8). [Turritellidae].
Mesalia clarki (Dickerson, 1914a). [Turritella 
Susana Fm., Paleocene. Trailer canyon, Topanga quad. (1952, PR 1981), Santa Monica Mtns., Los Angeles Co.
LACMIP 10508. Hypotypes LACMIP 13405–13411. [Turritellidae].
Mesalia martinezensis (Gabb, 1869b). [Turritella


Mesalia martinezensis (Gabb, 1869b). [Turritella

Mesalia martinezensis (Gabb, 1869b). [Turritella
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6668. Hypotype UCR 6668/3. [Turritellidae].
Mesalia martinezensis (Gabb, 1869b). [Turritella]. Saul 1983b. SEPM Guidebook 35:74, pl. 1
Virgenes Fm., Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCLA 3111 [=LACMIP 23111]. Hypotype UCLA
59360 [=LACMIP 10424]. [Turritellidae].
Mesalia martinezensis (Gabb, 1869b). [Turritella

Mesalia martinezensis (Gabb, 1869b). [Turritella    
Santa Susana Fm., Paleocene. Temescal Canyon, Topanga quad. (1952, PR 1967), Santa Monica Mtns., Los Angeles
Co. LACMIP 26897. Hypotypes LACMIP. 13399, 13403. [Turritellidae].
-
potype LACMIP 13400.


Hypotype LACMIP 13402.

LACMIP 13404.
Metacerithium packardi
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU. 2073 [=UCMP A9717]. Hypotype
UCMP 14196.
Miopleiona oregonensis Dall, 1907 (not Arnold as in caption) Cummings et al. 1962. CDMG Bull. 181:202, ph. 16,

F701M. Hypotype determined here as missing. Remarks: Determined here as in the genus/subgenus Musashia
(Nipponomelon). [Volutidae].
Mirachelus imperialis-
trania Fm.], Pliocene. Whitewater River, San Jacinto quad., Riverside Co. UCMP A1415. Holotype UCMP 14659,



Mitra (Atrimitra) andersoni
Silt, Miocene. North side of Kern River, Kern Co. USGS 6641. Holotype USNM 650183. [Mitridae].
    
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 255
Hypotype UCMP 33479.
Mitra (Mitra) andersoni
11, 13). Lower Round Mtn. Silt, Miocene Kern River Kern Co. USGS 6641. Holotype USNM 650183]. [Mitridae].
Mitra simplicissima
Santa Susana quad. (1951, PR 1969), Ventura Co. LACMIP 20586. Hypotypes LACMIP 12738, 12739. [Mitridae].
Mitra (Fusimitra) uvasana 

711. Hypotype CASG 358. [Mitridae].
Mitra 
area, Santa Cruz Is., California Channel Ids. Hypotype UCSB. [Mitridae].
Mitrella (Columbellopsis) alta        
sand, MiocenOil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650135, paratype USNM 650134.
[Columbellidae].
Mitrella (Mitrella) anchuela 
          
[=CASG 7965]. [Columbellidae].
....., LACMIP 16140 (ex
....., SDSNH 5422 (ex
Mitrella anchuela
Ranch quad. (1954), Kern Co. USGS M2601. Hypotype USNM 650136. [Columbellidae].



Mitrella cf.  (Gould, 1850). [Columbella
San Joaquin Fm. (Acila Zone), Pliocene. Arroyo Pino, North Dome, Kettleman Hills, Kings Co. USGS 12363. Hypo-
type USNM 495712. Remarks: Mitrella richthofeni (Gabb, 1866),  Adegoke (1969: p. 179). [Columbellidae].
Mitrella gausapata (Gould, 1850). [Columbella   
Joaquin Fm. (Neverita Zone), Pliocene. Arroyo Mellado, North Dome, Kettleman Hills, Kings Co. USGS 12519.
Hypotype USNM 495323. [Columbellidae].
Mitrella gausapata (Gould, 1850). [Columbella
7. Careaga Fm. (Cebada Member), Pliocene. Casmalia Hills, northeast of Shuman, Santa Barbara Co. USGS 14608.
Hypotype USNM 560089. [Columbellidae].
Mitrella gausapata (Gould, 1850). [Columbella]. Adegoke 1969.    
Fm., Pliocene. Near Skunk Hollow, Domengine Ranch quad., Fresno Co. UCMP B6548. Hypotype UCMP 36856.
[Columbellidae].
Mitrella gausapata (Gould, 1850). [Columbella
Capitola, Santa Cruz Co. Hypotype SCCM determined here as missing. [Columbellidae].
Mitrella gouldii (Carpenter in Gould and Carpenter, 1857). [Nitidella]. Cummings et al. 1962. CDMG Bull. 181:202,

Locality F701M. Hypotype determined here as missing. Remarks: Alia carinata (Hinds, 1844),, deMaintenon

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Mitrella cf.  (Carpenter in Gould and Carpenter, 1857). [Nitidella]. Addicott 1969. Pr.CAS 37(3):62. 64,

USNM 650959. [Columbellidae].
Mitrella gouldii (Carpenter in Gould and Carpenter, 1857). [Nitidella]. Perry 1977. Foss. Santa Cruz Co., p. 23, bot-

Remarks: Alia carinata (Hinds, 1844),
Mitrella (Bastrophia) llajasensis     
Santa Susana quad. (1969 photo revised), south side Simi Valley, Ventura Co. LACMIP 41604. Holotype LACMIP
14514. [Columbellidae].
Mitrella richthofeni (Gabb, 1866). [Columbella-
cene. Coalinga quad., Fresno Co. UCMP B6546. Hypotype UCMP 36858. [Columbellidae].
Mitrella tuberosa 
Careaga Fm. (Cebada Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14648. Hypotypes
(2) USNM 560090. [Columbellidae].
Molopophorus sp., cf. aequicostatus
Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13427. Remarks:
Molopophorus,”  Allmon (1990: p. 69). [Nassariidae].
Molopophorus anglonana (Anderson, 1905). [Bullia      
64-14. Temblor Fm., Miocene. Near Kern River, Kern Co. CASG 65. Hypotype CASG 5724. Remarks: In the genus
Molopophorus,”  Allmon (1990: pp. 64–65). [Nassariidae].
Molopophorus anglonana (Anderson, 1905). [Bullia
Molopophorus angloanus). Temblor Fm., Miocene. Kern River, Kern Co. USGS 6613. Hypotype USNM 563136.
Remarks:Molopophorus,”  Allmon (1990: pp. 64–65). [Nassariidae].
Molopophorus anglonanus (Anderson, 1905). [Bullia]. Addicott 1965b. USGS PP 525-C:
Miocene. Oil Center quad. (1950), Kern Co. UCMP B1598. Hypotype UCMP 36562. Remarks:Mo-
lopophorus,”  Allmon (1990: pp. 64–65). [Nassariidae].
Molopophorus anglonanus (Anderson, 1905). [Bullia
Olcese Sand, Miocene. Oil Center quad. (1950), Kern Co. UCMP B1598. Hypotypes UCMP 32851, 36562. [Nas-
sariidae]. Remarks:Molopophorus,”  Allmon (1990: pp. 64–65).

Molopophorus anglonanus (Anderson, 1905). [Bullia  
3. Temblor Fm., Miocene. Kern River, Kern Co. USGS 6613. Hypotype USNM 563136 (of Moore, 1963 [1964]).
Remarks:Molopophorus,”  Allmon (1990: pp. 64–65). [Nassariidae].
Molopophorus anglonanus (Anderson, 1905). [Bullia      

[Nassariidae].
Remarks: Erroneously listed as being from California, 

Molopophorus antiquatus (Gabb, 1864). [Nassa
Creek, San Guillermo quad. (1943), Ventura Co. UCR 4707. Hypotype UCR 4707/301. Remarks: In the genus
Colwellia,Nuttall and Cooper (1973: p. 209); Allmon (1990: p. 70). [Nassariidae].
Molopophorus biplicatus (Gabb, 1866). [Cuma      

1926 [1927]:389); ANSP 4340, paralectotype ANSP 79478. Remarks:Molopophorus,”  Allmon
(1990: p. 66). [Nassariidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 257
Molopophorus biplicatus (Gabb, 1866). [Cuma-
cene. Probably south of Martinez, Contra Costa Co. Hypotype MCZ 27846. [Nassariidae].
Molopophorus californicus
        
D8156. UCMP unnumbered hypotype lost, C. Throckmorton (personal communication, 2002). Remarks: In
Molopophorus,”  Allmon (1990: p. 68). [Nassariidae].
Molopophorus cretaceus (Gabb, 1864). [Nassa

1926 [1927]: p. 391) ANSP. 4197, 13 paralectotypes ANSP. 79432]. Remarks: In the genus Colwellia,  Nuttall
and Cooper (1973: p. 209); Allmon (1990: p. 70). [Nassariidae].
Molopophorus cretaceus (Gabb, 1864). [Nassa]. Squires 1984. NHMLAC CS 
Canyon, Santa Susana quad., Los Angeles Co. CSUN 467 [=LACMIP 16079]. Hypotype LACMIP. 6537. Remarks:
In the genus Colwellia,  Nuttall and Cooper (1973: p. 209); Colwellia,”  Allmon (1990: p. 70).
[Nassariidae].
Molopophorus dalli
Oligocene. San Emigdio region, Kern Co. CASG 165. Holotype CASG 168, paratype CASG 169. [Nassariidae].
Molopophorus striatus (Gabb, 1869b). [Bullia (Molopophorus-

[1927]:389) ANSP 4249, paralectotypes (3) ANSP 79505. Remarks:Molopophorus,”  Allmon
(1990: p. 64). [Nassariidae].
Molopophorus striatus (Gabb, 1869b). [Bullia (Molopophorus)]. Shimer and Shrock 1944. Index Foss. N.A., p. 517,

      Remarks: In the genus
Molopophorus,”  Allmon (1990: p. 64). [Nassariidae].
Molopophorus striatus (Gabb, 1869b). [Bullia (Molopophorus)]. Nuttall and Cooper 1973. Bull. BM(NH) 23(3):209,
-
lectotypes (3) ANSP 79505. Remarks:Molopophorus,”  Allmon (1990: p. 64). [Nassariidae].

(hypotype cited as UCMP 7182, which is actually the UCMP locality number).
Molopophorus tejonensis
Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4722. Hypotype UCR 4722/401. Remarks: In the genus
Colwellia,  Allmon (1990: pp. 69–70). [Nassariidae].
Moniliopsis electilis
Caliente quad., Kern Co. SU 2121. Holotype SU 7540 [=CASG 72007]. [Turridae].
Monodonta (Incisilabium?) piruensis        
Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 833 [=LACMIP 16281]. Holotype LACMIP 7441,

Morula (Morunella) granti
Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650160. [Muricidae].
Morula granti      
Heights, Santa Ana Mtns., Orange Co. Hypotype possibly at SDSNH, current whereabouts unknown. Remarks:
Ex Grouard collection. [Muricidae].
Muracypraea Arnold, 1998. The Festivus 30(8):90–92, nd through 5th specimens), middle row (1st,
4th and 5th specimens), and bottom row (all specimens)]. Latrania Fm., Miocene. Fossil Canyon, Imperial Co.
Hypotypes (12) SDSNH [ex SDSU] unnumbered (far left specimen in middle row missing, far right specimen
bottom row missing). [Cypraeidae].
258 PALEOBIOS, VOLUME 38, FEBRUARY 2021
nd and 3rd specimens from left). LACMIP 16811. Hypotypes LACMIP 11535, 14869.
Muracypraeast specimen), 3, 5 (p. 92). Latrania
Fm., Miocene. Fossil Canyon, Imperial Co. LACMIP 16811. [Hypotype LACMIP 11535]. [Cypraeidae].

Muracypraea
Alverson Canyon, Imperial Co. SDSNH 345. Hypotype SDSNH 88085. [Cypraeidae].

Murex (Ocenebra) dalli           
Pleasanton, La Costa Valley quad. (1943), Alameda Co. SU 2739. Hypotype SU 8471 [=CASG 69941]. Remarks:
Determined here as possibly in the genus Paciocinebrina. [Muricidae].

69942].
Murex rodeoensis
Ranch quad., Kings Co. UCMP D1103. Hypotype UCMP 36840. [Muricidae].
Murex whitneyi (Gabb, 1864). [Tritonium
Eocene. Nojoqui Creek area, Lompoc quad., Santa Barbara Co. UCMP B6920. Hypotype SU 9230 [=CASG 70190].
Remarks: In the genus Hexaplex Givens (1974: p. :83). [Muricidae].
Muricopsis vistaensis
Fm. [=Santiago Fm.], Eocene. Near Vista and San Marcos, San Diego Co. UCR 4865. Holotype UCR 4865/9, para-
types UCR 4865/90, 4865/91. [Muricidae].
Nassa coalingensis
Fm. (Pecten Zone), Pliocene. Arroyo Degollado, North Dome, Kettleman Hills, Kings Co. USGS 12359. Hypotype
USNM 495757. Remarks: In the genus/subgenus Nassarius (Caesia),  Addicott (1965a: p. B9). [Nassariidae].
Nassa mendica
Careaga Fm. (Cebada Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14648. Hypotypes
(2) USNM 560078. Remarks: In the genus Hima,  McLean (2007: p. 743). [Nassariidae].
Nassa moraniana Martin, 1914. Woodring et al. 1940 [1941]. 
Pliocene. North Dome, locality uncertain, Kettleman Hills, Kings Co.
Hypotypes USNM 495834, 495835. Remarks: Nassarius grammatus (Dall, 1917),  Addicott (1965a: p. B8).
[Nassariidae].
Nassa moraniana
Careaga Fm. (Cebada Member), Pliocene. Solomon Hills, Santa Barbara Co. USGS 14677. Hypotype USNM 560076.
Remarks: Nassarius gramatus (Dall, 1917),  Addicott (1965a: p. B6). [Nassariidae].

560120.
Nassa waldorfensis
Fm. (diatomaceous strata), Miocene. Casmalia Hills, near Shuman, Santa Barbara Co. USGS 14878. Hypotype
USNM 560022. Remarks: Junior synonym of Nassarius californianus (Conrad, 1857a),  Addicott (1965a: p.
B3). [Nassariidae].

USNM 560048.
       
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 259
14608. Hypotype USNM 560077.

14618. Hypotypes (2) USNM 560121.
“Nassa waldorfensis
Fm. (Acila Zone), Pliocene. Arroyo Estrecho, North Dome, Kettleman Hills, Kings Co. USGS 12364. Hypotype USNM
495854. Remarks: Junior synonym of Nassarius (Demondia) californianus (Conrad, 1856),  Addicott (1965a:
p. B3);  Adegoke (1969: p. 187). [Nassariidae].
Nassa
Member), Miocene/Pliocene. Foxen Canyon, Santa Barbara Co. USGS 14997. Hypotype USNM 560014. Remarks:
Probably referable to Nassarius (Demondia) grammatus (Dall, 1917),  Addicott (1965a: p. B6). [Nassariidae].
Nassarius (Catilon antiselli (Anderson and Martin, 1914). [Nassa
13, 14. Monterey (Saltos Shale Member) of Hill et al. (1958: pp. 2989–2990), Miocene. Wells Ranch quad., San
Luis Obispo Co. USGS M1923. Hypotype USNM 648580. [Nassariidae].
Nassarius (Catilon) arnoldi (Anderson, 1905). [Nassa  
Round Mtn. Silt of Diepenbrock (1933: pp. 14–16), Miocene. Rio Bravo Ranch quad., Kern Co. USGS M1604. Hy-
potype USNM 648575. [Nassariidae].
1933: pp. 14–16), Miocene. Oil Center quad., Kern Co. USGS M1597
[=UCMP B1598]. Hypotype (topotype) USNM 648576, neotype (of Addicott (1965a: p. B12) USNM 648577.
Nassarius arnoldi (Anderson, 1905). [Nassa
Miocene. Vicinity of Kern River, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1604. Hypotype USNM 649169.
Remarks: In the subgenus Catilon,  Addicott (1970: p. 98). [Nassariidae].
Nassarius (Catilon) arnoldi (Anderson, 1905). [Nassa
Round Mtn. Silt, Miocene. Vicinity of Kern River, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1604. Hypotype
USNM 648575. [Nassariidae].
-
potype USNM 650170.


650172, neotype (of Addicott (1965a: p. B12) USNM 648577.

Nassarius arnoldi (Anderson, 1905). [Nassa
(Buttonbed Sand Member) (of Stinemeyer et al. 1959: pp. 8, 13), Miocene. Shale Pt. quad., Temblor Range, Kern
Co. USGS M3986. Hypotype USNM 646795. [Nassariidae].
Nassarius sp cf. arnoldi
San Juan Capistrano, Orange Co. Monterey Fm., Miocene. Hypotype at CC. [Nassariidae].
Nassarius californianus (Conrad, 1857a). [Schizopyga
65-8, 65-9. Etchegoin Fm., Pliocene. Tulare Co. CASG 1397. Hypotypes CASG 5720, 5721. Remarks: In the sub-
genus Demondia,  Addicott (1965a: p. B3). [Nassariidae].
Nassarius californianus (Conrad, 1857a). [Schizopyga
Line drawing, source specimen unknown. Remarks: In the subgenus Demondia,  Addicott (1965a: p. B3).
[Nassariidae].
Nassarius (Demondia) californianus (Conrad, 1857a). [Schizopyga]. Addicott 1965a. USGS PP 503-B:B3–B5, pl. 1,

260 PALEOBIOS, VOLUME 38, FEBRUARY 2021
UCMP 7033. Hypotype UCMP 30809. [Nassariidae].

648548–648550, 648591, neotype (of Addicott 1965a: p. B4) USNM 648596.

 Addicott (1965a: p. B4).
    
14608. Hypotypes USNM. 649001, 649002.

Nassarius californianus (Conrad, 1857a). [Schizopyga   
Remarks: In
the subgenus Demondia,  Addicott (1965a: p. B3). [Nassariidae].
Remarks: Addicott (1965c:
p. 52).

Nassarius aff.  (Conrad, 1857a). [Schizopyga]. Durham and Addicott 1965. USGS PP 524-A:A13, pl.

USNM 649112. Remarks: In the subgenus Demondia,  Addicott (1965a: p. B3). [Nassariidae].
Nassarius (Demondia) californianus (Conrad, 1857a). [Schizopyga

Nassarius californianus (Conrad, 1857a). [Schizopyga
Pliocene. Capitola, Santa Cruz Co. CASG66180. Hypotype CASG 66180.03. Remarks: In the subgenus Demondia,
 Addicott (1965a: p. B4). [Nassariidae].
Nassarius (Demondia) californianus (Conrad, 1857a). [Schizopyga
Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14402. [Nassariidae].
Nassarius (Caesia) coalingensis (Arnold, 1909 [1910]). [Nassa  

Kings Co. USGS 4758. Holotype USNM 165508. [Nassariidae].
Pecten Zone and Tra-
chycardium Zone), Pliocene. Arroyo Degollado, North Dome, Kettleman Hills, Kings Co. USGS 12359. Hypotype
USNM 495757.
Nassarius (Caesia) coalingensis (Arnold, 1909 [1910]). [Nassa]. Durham and Addicott 1965. USGS PP 524-A:A13, pl.

Co. USGS M979. Hypotype USNM 649117. [Nassariidae].
Nassarius (Caesia) grammatus (Dall, 1917). [Alectrion
    
[Nassariidae].
Nassarius grammatus (Dall, 1917). [Alectrion
Pliocene. Santa Clara Co. USGS M1715. Hypotypes USNM 648568, 648569, 648592. Remarks: In the subgenus
Caesia,  Addicott (1965a: p. B6). [Nassariidae].
Nassarius (Caesia) grammatus (Dall, 1917). [Alectrion
Diego Fm., Pliocene. Reynard Wy., San Diego, San Diego Co. SDSNH 2383. Hypotype SDSNH 4170]. Remarks: In
the subgenus Caesia,  Addicott (1965a: p. B6). [Nassariidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 261
Nassarius (Caesia) grammatus (Dall, 1917). [Alectrion
      
648570. [Nassariidae].
Nassarius grammatus (Dall, 1917). [Alectrion
Helicotaphrichnus commensalis
n. sp. [spionid tube] in Nassarius grammatus =LACMIP 9398]. Remarks: In the subgenus Caesia,  Addicott
(1965a: p. B6). [Nassariidae].
Nassarius grammatus (Dall, 1917). [Alectrion      
Pliocene. Capitola, Santa Cruz Co. SCCM 1. Hypotype SCCM 9605.10. Remarks: In the subgenus Caesia,  Ad-
dicott (1965a: p. B6). [Nassariidae].
Nassarius grammatus (Dall, 1917). [Alectrion
Capitola, Santa Cruz Co. CASG. 66178. Hypotype CASG 66178.01. Remarks: In the subgenus Caesia,  Addicott
(1965a: p. B6). [Nassariidae].
Nassarius (Caesia) grammatus addicotti      -
chegoin Fm., PlioceneCoalinga quad., Fresno Co. UCMP B6541. Holotype UCMP 36877, paratype UCMP 36878.
[Nassariidae].
Nassarius (Caesia) grammatusAlectrion
Rio Del Fm. of Ogle (1953: pp. 31–33), Pliocene. Rio Del Fm., Scotia quad. (1951), Humboldt Co. UCMP B7642.
Hypotype UCMP 15074. [Nassariidae].
Nassariius (Catilon) hamlini (Arnold, 1907). [Nassa

648583, 648584. [Nassariidae].
Nassarius (Phrontis) harrellensis
Silt, Miocene. Kern River, Kern Co. USGS 3886. Holotype USNM 650173. [Nassariidae].
Nassarius harrelensis-
tonbed Sand member) of Stinemeyer et al. (1959: pp. 8, 13), Miocene. Shale Pt. quad., Temblor Range, Kern Co.
USGS M3986 (not USNM 3986 as in pl. caption). Hypotype USNM 646794. [Nassariidae].
Nassarius hildegardae Kanakoff, 1956 (n. sp.). BSCAS 55(2):112–113, pl. 31. Originally referred to as Pico Fm.
(Pliocene) but most likely is Castaic Fm. (Miocene). Mint Canyon quad., Los Angeles Co. LACMIP 291. Holotype
Remarks: In the subgenus Catilon,  Addicott (1965a:
p. B16). [Nassariidae].


Nassarius (Catilon) hildegardae
referred to as Pico Fm. (Pliocene) but most likely is Castaic Fm. (Miocene). Mint Canyon quad.. Los Angeles Co.
LACMIP 291. Hypotypes (topotypes) LACMIP 1131, 1132. [Nassariidae].
Nassarius (Catilon) iniquus (Stewart in Woodring et al., 1940 [1941]). [Nassa]. Addicott 1965a. USGS PP 503-B:B16,
Merriamaster coalingensis Zone” in La Cima quad., Kings Co. UCMP A3166.
Hypotype UCMP 15079. [Nassariidae].
Nassarius moranianus (Martin, 1914). [Nassa
Pliocene. Humboldt Co. UCMP A4233. Hypotype UCMP34173. Remarks: In the subgenus Caesia,  Addicott
(1965a: p. B8). [Nassariidae].
Nassarius moranianus (Martin, 1914). [Nassa-
cene. Bolinas, Marin Co. CSMB 12688. Hypotype CASG 8481. Remarks: In the subgenus Caesia,  Addicott
(1965a: p. B8). [Nassariidae].
262 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Nassarius (Caesia) moranianus (Martin, 1914). [Nassa
Merced Fm., Pliocene. Bolinas, Marin Co. UCMP 1549. Holotype UCMP 12338, hypotypes (topotypes) UCMP
15069–15071. [Nassariidae].
Nassarius moranianus (Martin, 1914). [Nassa
Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D3688. Hypotype USNM 251807. Remarks: In the
subgenus Caesia,  Addicott (1965a: p. B8). [Nassariidae].
Nassariusocoyanus (Anderson and Martin, 1914). [Nassa]. Addicott 1965a. USGS PP 503-B:B16–B17,

UCMP B1637. Hypotypes UCMP 15080–15082. [Nassariidae].
Nassarius (Phrontis) ocoyanus (Anderson and Martin, 1914). [Nassa]. Addicott 1970. USGS PP 642: 100–101,
  
15081,15082. [Nassariidae].
Nassarius (Catilon) pabloensis (Clark, 1915). [Nassa
(casts). Santa Margarita Fm., Miocene. Arvin quad., Kern Co. USGS M1619. Hypotypes USNM 648585–648588.
[Nassariidae].
Nassarius cf.  (Clark, 1915). [Nassa
Santa Margarita Fm., Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M3568. Hypotype
USNM 254382. Remarks: In the subgenus Catilon,  Addicott (1965a: p. B13). [Nassariidae].
Nassarius cf.  (Clark, 1915). Addicott et al. 1980. SEM Guidebook 6:19, 33, 
Addicott et al. 1978, area, La Panza Range, San Luis
Obispo Co. USGS M3568.Hypotype USNM 254382. Remarks: In the subgenus Catilon,  Addicott (1965a: p.
B13). [Nassariidae].
Nassarius cf.  (Hinds, 1844). [Nassa
(Pomponio Member), Pliocene. Purisima Creek, Half Moon Bay quad., San Mateo Co. locality F701M. Hypotype
determined here as missing. Remarks: Error for perpinguis. Nassarius (Demondia) californianus,  Addicott
(1965a: p. B3). [Nassariidae].
Nassarius (Phrontis?) posoensis
Olcese Sand, Miocene. Woody quad. (1952), Kern Co. USGS.M1593. Holotype USNM 650168. [Nassariidae].
Nassarius (Catilon?) salinasensis-
cho Rico Fm., Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M977. Holotype USNM 648597,
hypotype USNM 648598. [Nassariidae].
Nassarius salinasensis
Rico Fm., Miocene/Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M977. Hypotype USNM
648598. Remarks: In the subgenus Catilon Addicott (1965a: p. B15). [Nassariidae].
Nassarius (Catilon) smooti-
brock (1933: pp. 14–16), Miocene. Oil Center quad., Kern Co. USGS M1597. Holotype USNM 648578 (as USNM
648575 in pl. caption), hypotype (topotype) USNM 648579. [Nassariidae].
Nassarius (Phrontis) smooti
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 648578, hypotypes USNM 648579, see
also Addicott 1965b, 650174. [Nassariidae].
Nassarius (Phrontis) cf. ( ) smooti
Silt, Miocene. North side of Kern River, Kern River, Kern Co. USGS 6609. Hypotype USNM 650169. [Nassariidae].
Nassarius cf. (Catilon) smooti
Orange Co. SDSNH 4080. Hypotype SDSNH 75913. [Nassariidae].
Nassarius stocki
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 263
(Pliocene) but most likely is Castaic Fm. (Miocene). Mint Canyon quad.. Los Angeles Co. LACMIP 291. Holotype
Remarks: In the subgenus
Catilon,  Addicott (1965a: p. B14). [Nassariidae].
ex SU 8663).

Nassarius (Catilon) stocki Kanakoff, 1956. Addicott 1965a. USGS PP 503-B:B14, pl. 3
as Pico Fm. (Pliocene) but most likely is Castaic Fm. (Miocene). Mint Canyon quad, Los Angeles Co. CASG 36827
[=LACMIP 291). Hypotypes (topotypes) CASG 12599,12600. [Nassariidae].
Nassarius whitneyi (Trask, 1922). [Nassa
Muir station, Contra Costa Co. UCMP A4924. Plastotype UCMP 34167. Remarks: In the subgenus Caesia, 
Addicott (1965a: p. B9). [Nassariidae].
Nassarius (Caesia) whitneyi (Trask, 1922). [Nassa       
Briones Fm., Miocene. South of Pacheco, Concord quad., Contra Costa Co. UCMP A4920. Hypotype UCMP 15075.
[Nassariidae].

Costa Co. UCMP A4925. Hypotype UCMP 15076.
     
Briones quad. (1949), Contra Costa Co. UCMP B4749. Hypotypes UCMP 15077, 15078.
Nassarius
[=Wilson Grove Fm., Miocene/Pliocene]. Sonoma Co. Hypotype CASG determined here as missing. [Nassariidae].
Nassarius
Miocene/Pliocene]. Santa Rosa Valley, Sonoma Co. Hypotype CASG determined here as missing. [Nassariidae].
Natica (Cryptonatica) clausa Broderip and G.B. Sowerby I, 1829. Marincovich 1977. BAP 70(294):410–418, pl.
Natica convexa
Nomland 1916a) UCMP 12064. Remarks: In the genus Cryptonatica,  J.H. McLean (personal communication,
1999). [Naticidae].

Natica clausa
Capitola slump block, Santa Cruz Co. CASG 66177. Hypotype CASG 66177.01. Remarks: In the genus Cryptonatica,
J.H. McLean (personal communication, 1999). [Naticidae].
Natica (Naticarius) domengenica
Domengine Fm., Eocene. Parson's Peak, Coalinga quad., Fresno Co. UCMP 672. Holotype UCMP 15843. [Naticidae].
Natica (Tectonatica) gabbi 
Miocene. Near town of Walnut Creek, Contra Costa Co. UCMP 1311 . Holotype UCMP 11252. [Naticidae].
Natica (Euspira) galianoi Dall, 1909. [Polinices (Euspira
Miocene. Near junction of Norris Creek with Crow Creek, Hayward quarangle, Alameda Co. UCMP A4565. Hy-
potype UCMP 34157. [Naticidae].
Natica pabloensis Clark, 1915. Hall
1948, in the Tesla quad., mapped as Oursan Sandstone by Huey (1948 : pp. 38, 40, pl. 1), here considered Briones
Fm.”), Miocene. Vicinity of Livermore, Tesla quad., Alameda Co. SU 2740. Hypotype SU 8469 [=CASG 69944].
[Naticidae].
Natica (Naticarius posuncula
Upper Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650096. [Naticidae].
264 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Natica (Naticarius) posuncula
Temblor Fm., Miocene. Kern River, Kern Co. CASG65. Holotype CASG. 7084. [Naticidae].

Natica (Naticarius) posuncula
13. Temblor Fm., Miocene. Kern River, Kern Co. CASG 65. Holotype CASG 7084. [Naticidae].

Natica (Carinacca) rosensis 
Canyon, San Guillermo quad. (1943), Ventura Co. UCR 4703. Hypotype UCR 4703/7. [Naticidae].
Natica (Carinacca) rosensis  
Jolla Group,” Eocene. Near La Jolla, San Diego Co. UCMP 3990]. Holotype UCMP 30296, paratype UCMP 30927.
[Naticidae].

[=LACMIP 25730]. Hypotype UCLA 38778 [=LACMIP 9740].
Natica rosensis-
cene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH
20726. [Naticidae].
Natica (Tectonatica) russa
Fm., Pliocene. Vicinity of French Gulch, Fortuna quad., Humboldt Co. UCMP B7908 (not UCMP B7891 as in plate
caption). Hypotype UCMP 37836. Remarks: Junior synonym of Natica (Cryptonatica) clausa Broderip and G.B.
Sowerby I, 1829,  Marincovich (1977: p. 410). [Naticidae].
-
type UCMP 37835.
Natica (Natica) teglandae G D. Hanna and Hertlein, 1938. Addicott 1970. USGS 
Mtn. Silt, Miocene. Kern River, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1608. Hypotype USNM 650095.
Remarks: In the subgenus  Marincovich (1977: p. 388). [Naticidae].

Natica (Naticarius) teglandae G.D. Hanna and Hertlein, 1938. Marincovich 1977. BAP 70(294): 388–390, pl. 39,


Natica (Natica) uvasana-
cene. West of Cuarta Canyon, Pt. Conception quad., Santa Barbara Co. UCMP B6992. Hypotype SU 9198 [=CASG
70195]. Remarks: In the subgenus Naticarius,  Marincovich (1977: p. 390). [Naticidae].
Natica (Natica) uvasana
Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4715. Hypotype UCR 4715/4. Remarks: In the subgenus
Naticarius,  Marincovich (1977: p. 390). [Naticidae].

Natica (Naticarius) uvasana
Near Ft. Tejon, Kern Co. Lectotype (of Stewart 1926 [1927]: p. 322) ANSP 4233, paralectotypes (2) ANSP 79479,
 E. Benamy (personal communication, 1996)]. [Naticidae].
Natica (Naticarius) uvasana-
cene. Near Ft. Tejon, Kern Co. Lectotype (of Stewart 1926 [1927]: p. 322) ANSP 4233, paralectotypes (2) ANSP
4233 [=ANSP 79479]. [Naticidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 265

Natica (Naticarius) aff. () uvasana
Devil Canyon, Santa Susana quad. (sec. corner is in Oat Mtn. quad.), Los Angeles Co. CSUN 371 [=LACMIP 16115].
Hypotype LACMIP 6529. [Naticidae].
Natica (Naticarius) uvasana
Northern Lockwood Valley area, Ventura Co. CSUN 987 [=LACMIP. 16175]. Hypotype LACMIP 7709. [Naticidae].
Natica (Naticarius) uvasana
Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 987 [=LACMIP 16175]. Hypotype LACMIP 7709. [Naticidae].
Natica
Hypotype USNM 650100 (not USNM 950100 as in pl. caption). Remarks:
from this species [Natica (Natica) teglandae] or possibly ,”  Addicott (1970: p. 66). [Naticidae].
Natica (Natica) n. sp. Addicott 1973b. USGS PP 791:7, 19, pl. 8
Miocene Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotypes USNM 646609, 646610.
[Naticidae].
Natica
(1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13424. [Naticidae].

Ridge quad. (1955), Fresno Co. UCMP D6538. Hypotype UCMP 37168. [Naticidae].

(Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4312. Hypotype SDSNH 140186. [Naticidae].

Iverson Pt., Saunders Reef quad., Mendocino Co. UCMP D7149. Hypotype UCMP 14437. [Naticidae].


C. Throckmorton (personal communication, 2002). [Naticidae].
Indeterminate Naticidae Powell et al. 2019b. PaleoBios    
excavation, Berkeley Hills, Oakland, Alameda Co. UCMP IP13005. Hypotype UCMP 218737. [Naticide].
       

47313 [=LACMIP 9811]. [Cancellariidae].


Nekewis io (Gabb, 1864). [?Fasciolaria
of Alamo Creek, Dry Canyon, and Cuyama River valley, Reyes Peak quad. (1943), Ventura Co. UCR 4741. Hypotype
UCR 4741/11. [Perissityidae].
Nekewis ioFasciolaria
Canyon area, Kern Co. SU30116, [=CASG 245]. SU hypotype.10207 [=CASG 72820. [Perissityidae].
Nekewis simiensis
Meier Canyon, Calabasas quad., Simi Hills, Ventura Co. UCR 6805/1. Holotype UCR 6805/100. [Perissityidae].
Neptunea andersoni (Martin, 1914). [Chrysodomus. Wildcat Series, of
Lawson (1894: pp. 255–263), Pliocene. Eel River near Scotia, Humboldt Co. UCMP 1878. Holotype UCMP 12336.
266 PALEOBIOS, VOLUME 38, FEBRUARY 2021
[Buccinidae].
Neptunea (Sulcosipho) lawsoni (Martin, 1914). [Chrysodomus

Neptunea (Neptunea) lyrata altispira  
Fortuna quad. (1942), Humboldt Co. UCMP B7879. Hypotype UCMP 14532. [Buccinidae].
Neptunea (Neptunea) pribiloffensis (Dall, 1919a). [Chrysodomus 
Upper Rio Dell Fm., Pliocene. Fleener Creek, Ferndale quad., Humboldt Co. UCMP B7944. Hypotype UCMP 37807.
[Buccinidae].
Neptunea (Colus) cf. () recurva-
ced” Fm. [=Purisima Fm.], Pliocene. Vicinity of San Vicente Creek, San Mateo Co. UCMP B4793. Hypotype UCMP
37661. [Buccinidae].
Neptunea (Sulcosipho) scotiaensis (Martin, 1914). [Chrysodomus
Dell Fm., Pliocene. Eel River area, Scotia quad., Humboldt Co. UCMP B7644. Hypotype UCMP 37812. [Buccinidae].
Neptunea (Neptunea) smirnia (Dall, 1919b). [Chrysodomus
Fm., Pliocene. French Gulch area, Fortuna quad., Humboldt Co. UCMP B7891. Hypotype UCMP 37806. [Buccinidae].
Neptunea (Golikovia) smirnia (Dall, 1919b). [Chrysodomus
Pliocene. Deep Canyon, Dana Pt. quad. (1949), Orange Co. USGS M2106. Hypotype USNM 250505. [Buccinidae].
Neptunea stantoni (Arnold, 1908). [Chrysodomus
Fm. (Pomponio Member), Pliocene. Purisima Creek area, Half Moon Bay quad., San Mateo Co. 645-1M. Hypotype
determined here as missing. [Buccinidae].
Neptunea (Sulcosipho stocki (Howe, 1922). [Buccinum
Fm., Pliocene. Eel River, in vicinity of Scotia and Rio Dell, Humboldt Co. UCMP A4539. Hypotype UCMP 37810.
[Buccinidae].
Neptunea (Sulcosipho) tabulata (Baird, 1863). [Chrysodomus
Fm., Pliocene. Nanning Creek Scotia quad., Humboldt Co. UCMP B7642. Hypotype UCMP 37808. [Buccinidae].
Neptunea cf. Ntabulata (Baird, 1863). [Chrysodomus]. Addicott 1969. Pr.CAS 37(3):62, 64, 67, pl. 1, . 17. Merced
FmPliocene. Palo Alto quad., Santa Clara Co. USGS M1715. Hypotype USNM 650958. [Buccinidae].
Neptunea tabulata (Baird, 1863). [Chrysodomus]. Weaver 1971. West (LA Times), 18 April, 1971:11, unnumbered
th and Flower streets, downtown Los
Angeles, Los Angeles Co. LACMIP 1219. Hypotypes (2) current whereabouts unknown. [Buccinidae].
Neptunea (Sulcosipho) tabulata (Baird, 1863). [Chrysodomus]. Nelson 1978. Veliger 21(2):206, . 13Fernando
Fm.,” Pliocene. Upper Newport Bay, Newport Beach quad., Orange Co. USGS M2753. Hypotype USNM. 250507.
[Buccinidae].
Neptunea tabulata (Baird, 1863). [Chrysodomus]. Powell 2013. Festivus 45(3):20, pl. 1, . 8. Reworked shells,
Pliocene. Shells found in a modern-day shrimp trap off San Clemente Is., California Channel Ids. CASG 72573.
Hypotype CASG.72577. [Buccinidae].
Nerita beali
Fm., Miocene. Lower Cañada Garanon, Santa Rosa Is., California Channel Ids. UCSB SR67-55 [=LACMIP 16427].
Hypotype UCSB M9 [=LACMIP 6318]. Remarks: Determined here as Glossaulax reclusiana (Deshayes, 1839).
[Naticidae].
Nerita cf.  (Amphinerita) eorex 
Canyon, Thousand Oaks quad. (1950, PR 1967), Ventura Co. CSUN 700 [=LACMIP.16289]. Hypotype LACMIP 6508.
Remarks: Velates perversus (Gmelin, 1791),  Saul and Squires (1997: pp. 135, 136). [Neritidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 267
Nerita (Theliostyla?) joaquinensis
Miocene. Piso Creek, Kern River area, Kern Co. USGS M1640. Paratype USNM 650050. [Neritidae].

Nerita (Theliostyla?) kennedyi
Eocene. Vista area, San Marcos quad. (1968, PR 1983), San Diego Co. SDSNH 4105. Holotype SDSNH 67066.
[Neritidae].
Nerita quadrangulata 
Gaviota fms., Eocene. Gaviota Pass area, Los Olivos quad., Santa Barbara Co. UCMP B6962 [=66550] Holotype SU
9175 [=CASG 66550.01]. Remarks: Junior synonym of Nerita (Theliostyla) triangulata Gabb, 1869b,  Squires
(1992: p. 325). [Neritidae].
Nerita scabricosta-
trania Fm., Miocene/Pliocene. Alverson Canyon [=Fossil Canyon), Imperial Co. SDSNH 3135]. Hypotype SDSNH
53865. [Neritidae].
Nerita triangulata
Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH1667. Hypotype SDSNH 4160. Remarks: In the sub-
genus Theliostyla,  Squires (1992: p. 325). [Neritidae].
Nerita (Theliostyla) triangulata Matilija Fm., Eocene. Beartrap
Creek area, Reyes Peak quad. (1943), Ventura Co. UCR 4747. Hypotype UCR. 4747/3. [Neritidae].
Nerita (Theliostyla) triangulata
      
4865/100, 4865/101. [Neritidae].
Nerita (Theliostyla) triangulata
Cobblestone Mtn. quad. (1958), Ventura Co. CSUN 809 [=LACMIP 16291]. Hypotype LACMIP 7443. [Neritidae].
Nerita (Theliostyla) triangulata     
Southwest of San Marcos, San Marcos quad. (1968, PR 198]), San Diego Co. SDSNH 3522. Hypotypes SDSNH


Holotype CASG 9175 [=CASG 66550.01], paratype CASG 9175a [ CASG 66550.02] (both of Nerita quadrangulata
Weaver and Kleinpell, 1963).
Nerita (Theliostyla) triangulata-
stone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 23720. Hypotype LACMIP 11646. [Neritidae].
Nerita
Co. CIT 2072 [=LACMIP 16190]. [Hypotype LACMIP 4159]. [Neritidae].
Nerita Fowkes 1982. Coalinga Guidebook, p. 21 [as Nerita triangulata
Nerita triangulata
Gabb 1869)). Domengine Fm. [as Tejon Fm.], Eocene. West of San Joaquin Coal mine, west of Coalinga, Fresno Co.
Hypotype USNM 165700. Remarks: Determined here as Nerita triangulata Gabb, 1869. [Neritidae].
Nerita
UCMP A1297. Hypotype UCMP 30490. [Neritidae].
Neverita (Glossaulax) alta
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotypes USNM 650104, 650105. Remarks: Junior
synonym of Neverita (Glossaulax) reclusiana (Deshayes, 1839),  Marincovich (1977: p. 317); valid species in
the genus  McLean (2007: p. 737). [Naticidae].
Neverita (Glossaulax) andersoni (Clark, 1918). [Natica
268 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Remarks: In the genus Glos-
saulax,  Majima (1989: p. 58). [Naticidae].
Neverita (Glossaulax) andersoni (Clark, 1918). [Natica 22. Upper
UCMP B1593. Hypotype UCMP 36557. Remarks: In
the genus Glossaulax,  Majima (1989: p. 58). [Naticidae].


Neverita (Glossaulax) andersoni (Clark, 1918). [Natica
Ramon Fm., Miocene. Walnut Creek, Contra Costa Co. UCMP 1131. Holotype UCMP 11212. Remarks: In the genus
Glossaulax,  Majima (1989: p. 58). [Naticidae].
Natica (Nev-
erita) pabloensis Clark 1915) UCMP 11592.


14421.
Neverita andersoni (Clark, 1918). [Natica
(lower sandstone member), Oligocene. The Indians, Junipero Serra quad., Santa Lucia Range, Monterey Co. SU
1756. Hypotype CASG 60367 determined here as missing. Remarks: In the genus Glossaulax  Majima (1989:
p. 58). [Naticidae].
Neverita (Glossaulax) globosa
       
Hypotypes UCR 4865/60–4865/65. Remarks: In the subgenus Neverita,  Squires (1999c: p. 17). [Naticidae].
Neverita (Neverita) globosa

27859 [=MCZ 108518]. [Naticidae].
      Neverita
globosa reefensis Vokes, 1939) UCMP 15855.
Neverita (Neverita) globosa
of Bus and Trough canyons, Thousand Oaks quad. (1950, PR 1967), Ventura Co. CSUN 704 [=LACMIP 16177].
Hypotype LACMIP 6527. [Naticidae].
Neverita (Neverita) globosa
Peak quad. (1958), Los Angeles Co. CSUN 827 [=LACMIP 16153]. Hypotype LACMIP 7472. [Naticidae].
Neverita (Neverita) globosa
Eocene. Matilija Creek, Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1450 [=LACMIP 16963). Hypotype
LACMIP 12455. [Naticidae].
Neverita (Glossaulax) jamesae Moore, 1963 . Round Mtn. Silt,
Miocene. Temblor locality 1, Caliente quad., Kern Co. UCMP 2713. Hypotype UCMP 15498. Remarks: In the genus
Glossaulax,  Majima (1989: p. 58). [Naticidae].
Neverita (Glossaulax) jamesae
Remarks:
In the genus Glossaulax,  Majima (1989: p. 58). [Naticidae].
Neverita (Neverita) kirkensis (Clark, 1915). [Natica
Pablo Group, Miocene. Southeast of Oyster Pt., Contra Costa Co. UCMP 467. Holotype UCMP 11591. [Naticidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 269
Natica (Neverita) arnoldi
Clark, 1915) UCMP 11594.
Nevertia reclusiana (Deshayes, 1839). [Natica
20. San Joaquin Fm. (Trachycardium Zone), Pliocene. Arroyo Conchoso, North Dome, Kettleman Hills, Kings Co.
USGS 12375. Hypotypes USNM. 495764, 495771, 495772. Remarks:Neverita
(Glossaulax) andersoni (Clark, 1918)],  Addicott (1970: p. 67). In the genus Glossaulax,  McLean (2007:
p. 737). [Naticidae].

Neverita Zone), Pliocene. North Dome,
Kettleman Hills, Kings Co. USGS 12499. Hypotype USNM (slab with Olivella cf.  [=Callianax pedroana],
Anadara trilineata, and Volsella fragments), determined here as missing. [Naticidae].


   
495321.

Neverita reclusiana (Deshayes, 1839). [Natica
Careaga Fm. (Graciosa Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14622. Hypotype
USNM 560119. Remarks: In the genus Glossaulax,  McLean (2007: p. 737). [Naticidae].
Neverita (Glossaulax) reclusiana (Deshayes, 1839). [Natica]. Marincovich 1977. BAP 70 (294):317–328, pl. 29,
  
Neverita secta Gabb, missing, 
K. Estes-Smargianni (personal communication, 2019). Remarks: In the genus Glossaulax,  McLean (2007:
p. 737). [Naticidae].

Solano Co. UCMP A1289. Holotype (of Neverita secta hemisecta Clark, 1938) UCMP 12482.
     
UCMP 14420.
Neverita (Glossaulax) reclusiana (Deshayes, 1839). [Natica
Fm., Pliocene. Orange Co. SDSNH4080. SDSNH hypotype determined here as missing]. Remarks: In the genus
Glossaulax,  McLean (2007: p. 737). [Naticidae].
Neverita (Glossaulax) reclusiana (Deshayes, 1839). [Natica
Miocene (Cold Creek Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP22645 [=UCLA
2645). Hypotype LACMIP 13084 [=UCLA 57698]. Remarks: In the genus Glossaulax,  McLean (2007: p. 737).
[Naticidae].
Neverita reclusiana (Deshayes, 1839). [Natica
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14389. Remarks: In the genus Glossaulax,  McLean
(2007: p. 737). [Naticidae].
Neverita (Glossaulax) cf. () reclusiana (Deshayes, 1839). [Natica

Remarks: In the genus Glossaulax,  McLean (2007: p. 737). [Naticidae].
Neverita reclusiana forma thomsonae-
    
Monterey Co. SU 1756. Hypotype CASG 60368 determined here as missing. Remarks: In the genus Glossaulax,
 McLean (2007: p. 737). [Naticidae].
270 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Neverita reclusianus (Deshayes, 1839). [Natica]. Shapiro 1998. AAPG Guidebook MP 45:129, pl. 2 (pl. 1 in caption),

[=SBMNH 60195]. Remarks:Ficopsis
cassid sp. [Naticidae, Ficidae, Cassidae].
Neverita thomsonaePhacoi-
des Sand member) (of Stinemeyer et al.1959: pp. 12, 13), Miocene. Carneros Rocks quad., Temblor Range, Kern
Co. USGS M3978. Hypotype USNM 646617. Remarks: Determined here as in the genus Glossaulax. [Naticidae].
Neverita (Glossaulax) thomsonae

Remarks: Determined
here as in the genus Glossaulax, . [Naticidae].
NeveritaNeverita Zone”), Pliocene.

Neverita (Glossaulax
Kern Co. USGS 6622. Hypotype USNM 650107. Remarks:
specimen as Neverita (Glossaulax) andersoni (Clark, 1918); Polinices (Euspira) galianoi Dall, 1909,  Marinco-
vich (1977: p. 265). [Naticidae].
Remarks: Junior synonym of
Polinices (Euspira) galianoi Dall, 1909, Marincovich (1977: p. 265).
Niso antiselli
Miocene. Kern River, Kern Co. USGS 6065. Hypotype USNM 650293. [Eulimidae].


Niso cottonwoodensis
Kern Co. USGS M1640. Holotype USNM 650291. [Eulimidae].

Nucella analoga (Forbes, 1852). [Purpura
Pliocene. Seven Mile Beach, San Mateo Co. Hypotype CASG determined here as missing. [Muricidae].
Nucella canaliculata (Duclos, 1832). [Purpura
Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D5765. Hypotype USNM 251806. [Muricidae].
Nucella collomi (Carson, 1926). [Thais-
cene/Pliocene. Northeast of San Lucas, Monterey Co. USGS M903. Hypotype USNM 254388. [Muricidae].
Nucella etchegoinensis (Arnold, 1909 [1910]). [Thais
San Joaquin Fm. (Cascajo Conglomerate Member), Pliocene. El Arco, South Dome, Kettleman Hills, Kings Co. USGS
12796 [=USGS 12654). Hypotype USNM 495352. [Muricidae].


495354.
Patinopecten Zone), Pliocene. Vicinity of Arroyo Torcido, North Dome, Kettleman
Hills, Kings Co. USGS 12422. Hypotypes USNM 495844, 495845.
Nucella n. sp. aff.  (Arnold, 1909 [1910]). [Thais]. Powell et al. 2019a USGS SIR 2019-5021:3 (as
Nucella n. sp. cf. 
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 271
Co., CASG 54135, Hypotype CASG. 784117. [Muricidae].
Nucella megastoma     
Seven Mile Beach, San Mateo Co. CASG. 69251. Holotype CASG 69251. [Muricidae].
ex CASG.
54164. Paratype CASG 69252.
Nucella packi (Clark) var. talea
Ridge, near Beltran, Dark Hole quad., Fresno Co. USGS 14390. Holotype USNM 498749. Remarks: Junior synonym
of Nucella tokudai (Yokoyama, 1932),  Amano et al. (1993: p. 240). [Muricidae].
Nucella rankini Grant and Eaton in
Caliente Range, San Luis Obispo Co. UCLA 1644 [=LACMIP 21644]. Holotype UCLA 8936 [=LACMIP 8963]. [Mu-
ricidae].
Ocenebra cierboensis (Grant and Eaton in Eaton et al., 1941). [Boreotrophon]. Addicott 1978. USGS OFR 78-446:88–89,
        
USNM 254346. Remarks: Determined here as possibly in the genus Paciocinebrina. [Muricidae].
Ocenebra clarki
Kern Co. UCLA AC 8-34 [=UCLA 6361 and LACMIP. 26361]. Holotype UCLA 45765 [LACMIP 8432]. Remarks:
Determined here as possibly in the genus Paciocinebrina. [Muricidae].

Ocenebra coryphaena (Woodring in Woodring et al., 1946). [Tritonalia]. Weaver 1971. West (LA Times), 18 April,

downtown Los Angeles, Los Angeles Co. LACMIP 1219. Hypotypes (3) whereabouts unknown. [Muricidae].
Ocenebra gabbiana (Anderson, 1905). [Trophon
Miocene. Kern Co. USGS 6624. Hypotype USNM 650113. Remarks: Determined here as possibly in the genus
Trophon. [Muricidae].
Ocenebra gabbiana (Anderson, 1905). [Trophon
Fm. (Buttonbed Sand Member) (of Stinemeyer et al. 1959: pp. 8, 13), Miocene. Shale Pt. quad., Temblor Range,
Kern Co. USGS M3986. Hypotype USNM 646806. Remarks: Determined here as possibly in the genus Trophon.
[Muricidae].
Ocenebra interfossa
Pliocene. Santa Clara Co. CASG 28617. Hypotype CASG 13108. CASG hypotype determined here as missing. Re-
marks: In the genus Paciocinebrina,  Houart et al. (2019: p. 209). [Muricidae].
Ocenebra milicentana (Loel and Corey, 1932). [Purpura
Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650130. Remarks:
Determined here as possibly in the genus Trophon. [Muricidae].
Ocenebra topangensis
Fm., Miocene. South of Calabasas, Los Angeles Co. CASG 567. Hypotype CASG 5719. Remarks: Determined here
as possibly in the genus Paciocinebrina. [Muricidae].
Ocenebra topangensis
Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650122. Remarks: Determined here as pos-
sibly in the genus Paciocinebrina. [Muricidae].
Ocenebra topangensis-
type possibly at SDSNH, current whereabouts unknown. Remarks: Ex Grouard collection. Remarks: Determined
here as possibly in the genus Paciocinebrina. [Muricidae].
Ocenebra topangensis      
272 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22644 [=UCLA 2644]. Hypotype LAC-
MIP 13080 [=UCLA 57361]. Remarks: Determined here as possibly in the genus Paciocinebrina. [Muricidae].
Ocenebra trophonoides (Anderson and Martin, 1914). [Thais
Upper Olcese Sand, Miocene. Caliente quad. (1914), Kern Co. UCMP B1621. Hypotype UCMP 15992. Remarks:
Determined here as possibly in the genus Paciocinebrina. [Muricidae].

Ocenebra wilkesana (Anderson, 1905). [Fusus
Miocene. Caliente quad. (1914), Kern Co. UCMP B1599. Hypotype UCMP. 30122. Remarks: Determined here as
possibly in the genus Paciocinebrina. [Muricidae].

Ocenebra cf.  (Anderson, 1905). [Fusus-
ett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650121. Remarks:
Determined here as in the genus Paciocinebrina. [Muricidae].
Ocenebra
CASG 37468. Hypotype CASG 12934 (This is the number on specimen, however label and pl. caption read CASG.
12933 as does online database). Remarks: Determined here as in the genus Paciocinebrina. [Muricidae].
Ocenebra n. sp. Addicott 1978. Fm., Miocene/Pliocene. North-
east of San Lucas, Monterey Co. USGS M903. Hypotype USNM 254347. Remarks: Equivalent to Ocenebra n. sp.
A of Durham and Addicott (1965: p. A12). Remarks: Determined here as possibly in the genus Paciocinebrina.
[Muricidae].
Ocinebrina atropurpurea (Carpenter, 1865). [Ocenebra
Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14395. Remarks: In the genus Paciocinebrina,
 Houart et al. (2019: p. 209). [Muricidae].
Ocinebrina sp., aff.  (Oldroyd, 1920). [Tritonalia
Pliocene. Newhall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14396. Remarks: In the genus Pacio-
cinebrina,  Houart et al. (2019: p. 209). [Muricidae].
Odostomia (Evalea) andersoni
Silt (possibly uppermost Olcese Sand), Miocene. Kern River Caliente quad., Kern Co. SU 2641. Hypotypes SU 7541,
7541a [=CASG 72009]. [Pyramidellidae].
Odostomia (Evalea) andersoni
Sand, Miocene. Oil Center quad., Kern Co. USGS M1597. Hypotypes USNM 650304, 650305. [Pyramidellidae].


USNM 650306.
Odostomia (Evalea) aff. () donilla
Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650319. [Pyramidellidae].
Odostomia (Menestho) repenningi
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Paratype USNM 650313, holotype USNM 650314.
[Pyramidellidae].
Odostomia (Besla) rotundomontana (Keen, 1943). [Chrysallida  
Lower Round Mtn. Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Hypotype USNM. 650318.
[Pyramidellidae].

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 273

Odostomia (Chrysallida?) sequoiana    
8, 9, 19. Upper Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650302,
paratypes USNM 650300, 650301, Hypotype USNM 650285. [Pyramidellidae].
Odostomia (Chrysallida
Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650328. [Pyramidellidae].
Odostomia (Evalea      
quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650317.
Odostomia (Evalea
Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650316. [Pyramidellidae].
Olequahia domenginica (Vokes, 1939). [Ranella-
wood Valley quad. (1943), Ventura Co. UCR 4686. Hypotype UCR 4686/8. [Bursidae].

[Bursidae].
Olequahia domenginica (Vokes, 1939). [Ranella
Llajas Canyon, 1950 ft., Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype
LACMIP 6533. [Bursidae].
Olequahia domenginica (Vokes, 1939). [Ranella
Eocene. Simi Valley, Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 6533. [Bursidae].
Olequahia domenginica (Vokes, 1939). [Ranella
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LACMIP 7712.
[Bursidae].
Olequahia domengenica (Vokes, 1939). [Ranella
Eocene. Runckle [sic] Ranch, Simi Valley, Ventura Co. Line drawings, source specimen unknown. [Bursidae].
Olequahia hornii (Gabb 1864). [Tritonium
Eocene. Cañada de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP B6922. Hypotype SU 9223 [=CASG.
70205]. [Bursidae].
Olequahia lorenzana (Wagner and Schilling, 1923). [Strepsidura]. Weaver and Kleinpell 1963. UCPG 43:192, pl. 26,

-
potype SU 9226 [=CASG 70199].
    
UCMP B6985B. Hypotype SU 9225 [=CASG 70201].
Olequahia washingtoniana (Weaver, 1912). [Cassidaria]. 
from Dickerson Nyctilochus washingtoniana). Tejon Fm., Eocene. Tejon quad., Kern Co. CASG
245. Holotype CASG 311 [=CASG 7611]. Remarks: Surcula ioformis Anderson and G D. Hanna, 1925,  Keen
and Bentson (1944: p. 197). [Bursidae].
Oliva californica            
quad. (1914), Kern Co. UCMP B1600. Hypotype UCMP 36559. [Olividae].
Oliva (Oliva) californica
Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS. M1613. Hypotype USNM 650217. [Olividae].

274 PALEOBIOS, VOLUME 38, FEBRUARY 2021
UCMP 36936 (not UCMP 15507 as in pl. caption).
    
650218.

Oliva spicata (Röding, 1798). [Porphyria
West of Haskell Canyon, Los Angeles Co. CIT 230 [=LACMIP 10271]. Hypotype LACMIP 4199]. [Olividae].
Oliva spicata (Röding, 1798). [Porphyria
[=Latrania Fm.], Miocene/Pliocene. Coyote Mtn., Imperial Co. Line drawing, source specimen unknown. [Olividae].
Oliva spicata (Röding 1798). [Porphyria
Latrania Fm., Miocene/Pliocene. Painted Gorge, Imperial Co. SDSNH 378. Hypotype SDSNH 12332. [Olividae].
Oliva (Oliva
(1906), Kern Co. UCMP B1587. Hypotype UCMP 33871. [Olividae].
Olivella cf. 
Fm. (Cebada Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14648. Hypotype USNM
560093. Remarks: Junior synonym of Callianax alectona (Duclos, 1835),  Powell et al. (2020: p. 5). [Olividae].
Olivella biplicata (Sowerby I, 1825). [Oliva
[=Wilson Grove Fm.], Pliocene. Vicinity of Russian River, Sonoma Co. CASG 546. Hypotype CASG 5732. Remarks:
In the genus Callianax, McLean (2007: p. 746.) [Olividae].
Olivella biplicata (Sowerby I, 1825), slender var. [Oliva]. Woodring and Bramlette 1950 [1951]. USGS PP 222:78, pl.

Remarks: In the genus Callianax, McLean (2007: p. 746). [Olividae].

Olivella biplicata (Sowerby I, 1825). [Oliva       
drawing, source specimen unknown. Remarks: In the genus Callianax, McLean (2007: p. 746). [Olivlidae].
Olivella biplicata (Sowerby I, 1825). [Oliva        
Santa Clara quad., Santa Clara Co. USGS M1715. Hypotype USNM 650977. Remarks: In the genus Callianax,
McLean (2007: p. 746). [Olividae].
Olivella biplicata (Sowerby I, 1825). [Oliva   
-
type CASG determined here as missing. Remarks: In the genus Callianax, McLean (2007: p. 746). [Olividae].
Olivella ischnon        

SU 7973 [=CASG 7973]. Remarks: Determined here as in the genus Callianax. [Olividae].
....., SDSNH 5422 (ex
Olivella (Olivella) ischnon-
cene. Oil Center quad. (1950), Kern Co. UCMP.B1598. Hypotype UCMP 36935. Remarks: Determined here as in
the genus Callianax. [Olividae].

Olivella aff. 
Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype SDSNH
20774. [Olividae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 275
Olivella mathewsonii
Susana quad., Los Angeles Co. CSUN 467 [=LACMIP 16079]. Hypotype LACMIP 6545. [Olividae].
Olivella mathewsonii
Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP 7484.
[Olividae].
Olivella mathewsonii
Sawmill Mtn. quad. (1943), Ventura Co. CSUN 987 [=LACMIP 16175]. Hypotype LACMIP 7715. [Olividae].
Olivella mathewsonii
Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13428. [Olividae].
Olivella cf.  (Conrad, 1856b). [Strephona]. Woodring et al. 1940 [
     
     Fm. (Neverita Zone), Pliocene. Butte east of Lower
Road, North Dome, Kettleman Hills, Kings Co. USGS 12499. Hypotype USNM (slab with Neverita reclusiana (=
Glossaulax reclusiana), Anadara trilineata, and Volsella fragments), determined here as missing. Remarks: In the
genus Callianax,  Powell et al. (2020: p. 22). [Olividae].
      
USNM 495325, 495326.
Olivella pedroana (Conrad, 1855b). [Strephona
Capitola, Santa Cruz Co. CASG 66179. Hypotype CASG 66179.01. Remarks: In the genus Callianax,  Powell
et al. (2020: p. 22). [Olividae].
Olivella-
potype USNM 650226. Remarks: Determined here as in the genus Callianax. [Olividae].
Olivella     Neverita Zone”), Pliocene.
Remarks: Determined here as in the genus Callianax. [Olividae].
Olivia* sp. Redman et al., 2007. Palaios
Imperial Co. Hypotype probably at SDSU, current whereabouts unknown. Remarks: Error for Oliva. [Olividae].
Olivula marysvillensis-

Holotype UCMP 11770. [Olivellidae].
Oniscidia plectata (Waring, 1917). [Ficus 
Sandstone, Paleocene. South side Simi Valley, Ventura Co. CASG 61667. Holotype CASG 61667.05. [Harpidae].
-
ley, Ventura Co., LACMIP 41691. Hypotype LACMIP 14748.
Opalia anomala
Diego Co. Hypotype CASG 11861. Remarks: Junior synonym of Opalia varicostata Stearns, 1875,  DuShane
(1979: p. 128). [Epitoniidae].
Opalia varicostataPatinopec-
ten Zone), Pliocene. North Dome, Kettleman Hills, Kings Co. USGS 12428. Hypotype USNM 495846. [Epitoniidae].
Opalia varicostata
Pliocene. Casmalia Hills, near Waldorf, Santa Barbara Co. USGS 14879. Hypotype USNM 560047. [Epitoniidae].
Opalia varicostata Stearns, 1875. Moore et al. 
specimen unknown. [Epitoniidae].
Opalia varicostata
276 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Beach, San Diego Co. SDSNH 2452. Hypotype SDSNH 4124]. [Epitoniidae].
Opalia varicostata  
Beach, San Diego Co. Hypotype CASG 11860. [Epitoniidae].

214040a.
-
potype LACMIP 6287.
 
Hypotypes LACMIP 6288–6291.
Opalia varicostata
Las Llajas Canyon, Santa Susana Mtns., Ventura Co. CSUN 1161 [=LACMIP 12600. Hypotype LACMIP 11615].
[Epitoniidae].
Opalia varicostata  
Goat Canyon, near U.S./Mexico border, San Diego Co. LACMIP 305. Hypotype LACMIP 14853 determined here
as missing. [Epitoniidae].
Opalia varicostata
in a modern-day shrimp trap off San Clemente Is., California Channel Ids. CASG 72573. Hypotype CASG.72578.
[Epitoniidae].
Opalia (Rugatiscala) williamsoni (Anderson and Martin, 1914). [Epitonium]. Addicott 1970. USGS PP 642:25, 56, pl. 3,


15448.
Opalia williamsoni (Anderson and Martin, 1914). [Epitonium]. Addicott 1972b. SEPM Symposium vol., p. 12, pl. 3,
   
Temblor Range, Kern Co. USGS M3986. Hypotype USNM 646800. [Epitoniidae].
Opalia (Opalia) wroblewskyi (Mörch, 1874). [Scala
Unnamed strata, Pliocene. San Clemente Is. Central quad. (1950), San Clemente Is., California Channel Ids. UCLA
6322 [=LACMIP 26322]. Hypotype UCLA 38798 [=LACMIP. 10834]. [Epitoniidae].
Opalia        
hypotype determined here as missing. [Epitoniidae].
Ophiodermella graciosana (Arnold, 1907). [Drillia]. Addicott 1969. Pr.CAS 37(3):62, 68–69, pl
., Santa Clara Co. USGS M1715. Hypotype USNM 650974. [Borsoniidae].
Ophiodermella graciosana (Arnold, 1907). [Drillia-
cene/Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D3693. Hypotype USNM 251811. [Borsoniidae].
Ophiodermella graciosana (Arnold, 1907). [Drillia-
cene. Capitola, Santa Cruz Co. CASG hypotype 66145.01 determined here as missing. [Borsoniidae].
Ophiodermella incisa (Carpenter, 1865). [Drillia-
cene/Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP B4811. Hypotype USNM 251813. [Borsoniidae].
Ophiodermella inermis (Reeve, 1943b). [Pleurotoma
Newhall area, Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14405. [Borsoniidae].
Ophiodermella mercedensis (Martin, 1914). [Drillia-
cene/Pleistocene. Twelvemile Creek area, San Mateo Co. UCMP D3364. Hypotype USNM 251812. [Borsoniidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 277
Ophiodermella temblorensis (Anderson and Martin, 1914). [Drillia]. Addicott 1970. USGS PP 642:134–135, pl. 17,

B1638 as in pl. 17 caption). Hypotype UCMP 36937. [Borsoniidae].

650259–650262.


Ornopsis
Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA 6525 [=LACMIP 26525].
Hypotype LACMIP 7564. [Fasciolariidae].
Otostoma aethes   
Cretaceous/lowermost Paleocene. Dip Creek, at the narrows, south shore of Lake Nacimiento, San Luis Obispo
Co. UCMP B2368. Holotype UCMP 398607. [Neritide]
Otostoma bisculptata (M.A. Hanna, 1927). [Philne*
or Scripps Fm., Eocene. Rose Canyon, San Diego Co. CASG 826. Holotype CASG 2189 [=CASG 826.01]. Remarks:
Error for Philine. [Neritide].

Pachychilus (Loxotrema) turritum (Gabb, 1868). [Loxotrema]. Wenz 1939. Handbh. Palaoz. 
from Stewart, 1926 [1927], pl. 26, Loxotrema turritum)). Domengine Fm., Eocene. West of Griswold's,
Fresno Co.. Lectotype (of Stewart, 1926 [1927]: p. 347, ANSP 4228, paralectotype ANSP 79510]. Remarks: In
the genus Loxotrema,  Squires (1999c: p. 16). [Melanopsidae].
Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina
Eocene. San Guillermo quad. (1943), Ventura Co. UCR 4699. Hypotype UCR 4699/101. Remarks: In the genus
Pachycrommium,  Squires (1987: p. 36). [Naticidae].

Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina
4, 5. Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana quad., Ventura Co. UCMP 7004. Holotype UCMP 31385,
paratype UCMP 31386. [Naticidae].
 Amau-
rellina multiangulata Vokes 1939) UCMP 15866.
 
UCMP 14410.
Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina    
Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP16115. Hypotype LACMIP 6526.
[Naticidae].
Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina
Fm., Eocene Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6526. [Naticidae].
Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina
Eocene. Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=LACMIP 16114]. Hypotype
LACMIP 7469. [Naticidae].
Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina

278 PALEOBIOS, VOLUME 38, FEBRUARY 2021
LACMIP 7706. [Naticidae].
Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina
Maniobra Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 672 [=LACMIP 16274].
Hypotype LACMIP 11465. [Naticidae].
Pachycrommium clarki (Stewart, 1926 [1927]) (not 1917 as in pl caption). [Amaurellina]. Squires 1994b. SEPM

Hypotype LACMIP 11651. [Naticidae].

Pachycrommium clarki (Stewart, 1926 [1927]). [Amaurellina35, 36. Santa
Susana Fm., Eocene. Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 967 [=LACMIP 12648].
Hypotype LACMIP 12683. [Naticidae].
Pachycromium clarki (Stewart, 1926 [1927]). [Amaurellina
Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13423.
[Naticidae].
Pachycrommium   
Canyon, Topatopa Mts. quad. (1943), Ventura Co. UCR 4668. Hypotype UCR 4668/11. [Naticidae].
Paraseraphs erraticus (Cooper, 1894). [Tornatina 
  Terebellum californicum Vokes 1939)
UCMP 15820 [=NMB H16551]. [Seraphsidae].
Terebellum cali-
fornicum Vokes, 1939) UCMP 15822 [=NMB H16553].
Terebellum
californicum Vokes, 1939, UCMP 15821[=NMB H16553].

66186.01 and plastotype NMB 16550].
Paraseraphs erraticus (Cooper, 1894). [Tornatina
Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6522. [Seraphsidae].
Paraseraphs erraticus (Cooper, 1894). [Tornatina
Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype LACMIP
7465. [Seraphsidae].
Paraseraphs erraticus (Cooper, 1894). [Tornatina
Eocene. Whitaker Peak area, Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype LACMIP 7456. [Seraphsidae].
Paraseraphs erraticus (Cooper, 1894). [Tornatina
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LACMIP 7703.
[Seraphsidae].
Parasyrinx boundeyi (Waring, 1917). [Bathytoma  
Fm., Paleocene Runkle Canyon, Simi Hills, Calabasas quad. (1952, PR 1967), Ventura Co.. CASG 61919 (not CASG
61901 as published). Holotype CASG 61919.01 (not CASG 61901.01 as in text). [Turridae].
Para-
syrinx hickmani Zinsmeister, 1983b) UCR. 4572/100.

Holotype (of Parasyrinx hickmani Zinsmeister, 1983b) UCR 6668/36.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 279
Parasyrinx hickmani 
Meier Canyon, Calabasas quad., Simi Hills, Ventura Co. UCR 6668. Holotype UCR 6668/36. Remarks: Although
hickmanae,” it is left here as is, for
taxonomic stability, as should subsequent authors, in accordance with ICZN Article 32.5.1 (ICZN, 1999). Junior
synonym of Parasyrinx boundeyi (Waring, 1917),  Squires (2000: p. 282). [Turridae].

Parasyrinx
Ventura Co. UCR 6668. Hypotype UCR 6668/36. Remarks: Parasyrinx boundeyi (Waring, 1917),  Squires
(2000:p. 282). [Turridae].
Patelloida tejonensisAcmaea
3.2. Tejon Fm., Eocene. Liveoak Canyon, Kern Co. Lectotype (of Stewart 1926 [1927]: p. 312) ANSP 4229 .

10851]. Hypotype UCMP 38520.
        
[=UCMP 10852]. Hypotypes UCMP 38521, 38522.
Patelloida triquetrus
Pastoria Creek quad. (1974), Kern Co. CSUN 1203 [=UCMP 10852]. Holotype UCMP 38523, paratypes UCMP

Patelloida vokesi (Hickman, 1980). [Acmaea
6.8. Tejon Fm., Eocene. Edmonston Pumping Plant area, Pastoria Creek quad. (1974), Kern Co. CSUN 1203 [=UCMP
10852]. Hypotypes UCMP 38525, 38526.
Patelloida
Plant area, Pastoria Creek quad. (1974), Kern Co. CSUN.1203 [=UCMP 10852]. Hypotypes UCMP 38527, 38528.
   
452886].
Pegocomptus howardi (Dickerson, 1914b). [Molophorus   
Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6679. Hypotype UCR 6679/110. Remarks: In the genus
Sulcobuccinum,  Vermeij (1998: p. 81). [Pseudolividae].

Penion titan (Waring, 1917). [Trachytriton Fm.,
Paleocene. Simi Hills, Ventura Co. UCLA 2688 [=LACMIP 22688]. Hypotype UCLA 59253 [=LACMIP 10983]. Re-
marks: In the genus Saxituberosa Squires and Saul (2006: p. 73). [Fasciolariidae].

Penion titon* (Waring, 1917). [Tachytriton
Fm., Paleocene. Simi Hills, Ventura Co. UCLA 2701 [=LACMIP 22701]. Hypotype UCLA 59254 (not UCLA 59353
as cited in pl. caption) [=LACMIP 10982]. Remarks: Error for titan. In the genus Saxituberosa [family Fasciolari-
 Squires and Saul (2006: p. 73). [Fasciolariidae].

cited in pl. caption) [=LACMIP 10983].
Perissitys stewarti (Zinsmeister, 1983). [Cophocara
Seca Fm., Paleocene. Ortigalita Creek, Charleston School quad., Merced Co. UCMP. A3262. Hypotype UCMP. 37996.
[Perissityidae].

280 PALEOBIOS, VOLUME 38, FEBRUARY 2021
61617]. Hypotype CASG 61617.01.

[=CASG 61619]. Hypotype CASG 61619.01.
 
61616.01, 61616.02.
Perissolax blakei (Conrad, 1855b). [Busycon-

4189 (missing,  K. Estes-Smargianni, personal communication, 2019). Remarks: In the genus Pseudoperis-
solax and family Muricidae, Squires (2015: pp. 578, 581). [Muricidae].
Perissolax blakei (Conrad, 1855b). [Busycon
Remarks: In the genus Pseu-
doperissolax and family Muricidae,  Squires (2015: pp. 578, 581). [Muricdae].
Persististrombus obliteratus (G D. Hanna, 1926). [Strombus
cast) (as Strombus obliteratus). Imperial Fm. [=Latrania Fm.], Pliocene. Near Travertine Rock, Imperial Co. (not
San Diego Co. as in locality description). USGS M9779 [=USGS M9778]. Hypotype USNM 486909. [Strombidae].
Perse corrugatum
       
UCMP 11233]. [Fasciolariidae].
Perse martinez (Gabb, 1864). [Fusus]. Givens 1974. UCPG 109:85–86, pl. 10
above the mouth of Wagon Rd. Canyon, San Guillermo quad. (1943), Ventura Co. UCR 4707. Hypotype UCR
4707/201. [Fasciolariidae].
Perse sinuata (Gabb, 1864). [Fasciolaria
Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4719. Hypotype UCR 4719/2. [Fasciolariidae].
Phalium (Semicassis) louella     
Maniobra Fm., Eocene. Orocopia Mtns., Canyon Spring SW quad. (1963), Riverside Co. CSUN 665 [=LACMIP
Remarks: In the genus  Squires
(2019: p. 8). [Cassidae].
Phalium (Semicassis) louella      
Maniobra Fm., Eocene. Red Canyon quad. (1986), Riverside Co. CSUN 665 [=LACMIP 16335]. Holotype LACMIP
7166. Remarks: In the genus Galeodea,  Squires (2019: p. 8). [Cassidae].
Phalium (Semicassis) tuberculiformis (G D. Hanna, 1924). [Morio
Fm., Eocene. Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP
6532. Remarks: In the genus Galeodea,  Squires (2019: p. 8). [Cassidae].
Phalium (Semicassis) tuberculiformis (G D. Hanna, 1924). [Morio

Hypotype LACMIP 7475. Remarks: In the genus Galeodea,  Squires (2019: p. 8). [Cassidae].
Phalium (Semicassis) tuberculiformis (G D. Hanna, 1924). [Morio
Susana Fm., Eocene. Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 967 [=LACMIP 12648.
Hypotype LACMIP 12684. [Cassidae]. Remarks: In the genus Galeodea,  Squires (2019: p. 8). [Cassidae].
Philine (Megistostoma) gabbianum (Stoliczka, 1868). [Bullaea
Megistostoma gabbianum
Costa Co.. Holotype ANSP 4216. Remarks: Megistostoma striata Gabb, 1864, see Groves and Squires (2018: p.
103) for the complicated nomenclatural history of this species. [Philinidae].
Phos dumbleanus Anderson in
Monterey Fm. (Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, west side of George
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 281
F. Canyon, Palos Verdes Hills, Los Angeles Co. USGS 13127. Hypotypes USNM 497074, 497075. [Buccinidae].

Arroyo area, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 548 [= LACMIP 15450. Hypotype LACMIP
12734. [Strombidae].
Pleurofusia fresnoensis (Arnold, 1909 [1910]). [Pleurotoma-
cene. West of Reyes Peak, Reyes Peak quad. (1943), Ventura Co. UCR 4706. Hypotype UCR 4706/231. [Drilliidae].
Pleurofusia sp. aff.  (Arnold, 1909 [1910]). [Pleurotoma]. Smith 1975b. SEPM Symposium vol., p. 376,
9. Lodo Fm., Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU
2073 [=UCMP A9717]. Hypotype UCMP 14199. [Drilliidae].
Pleurofusia fresnoensis (Arnold, 1909 [1910]). [Pleurotoma
Eocene Devil Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6553.
[Drilliidae].
Pleurofusia fresnoensis (Arnold, 1909 [1910]). [Pleurotoma
Eocene. Canton Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype
LACMIP 7495. [Drilliidae].
Pleurofusia fresnoensis (Arnold, 1909 [1910). [Pleurotoma
-
potype LACMIP 11473. [Drilliidae].
Pleurofusia fresnoensis (Arnold, 1909 [1910]). [Pleurotoma
Eocene. Elsmere Canyon, San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13431.
[Drilliidae].
Pleuroploca princeps (Sowerby I, 1825). [Fasciolaria]. Deméré and Rugh 2006. Foss. Treas. Anza-Borrego Desert,

Remarks: In the genus Triplofusus,  Snyder et al. (2012: p. 43). [Fasciolariidae].

Pleurotoma” cf. 
Joaquin Fm. (Acila Zone), Pliocene. Arroyo Hondo, North Dome, Kettleman Hills, Kings Co. USGS 12340. Hypotype
USNM 495711. [Turridae].
Pleurotomaria vacavillensis-
known Fm., Upper Cretaceous or lower Eocene. Near Vacaville, Solano Co. UCMP A6294. Holotype UCMP 14237.
In the genus Leptomaria,  Harasewych et al. (2009: p. 758). [Pleurotomariidae]
Pleurotomaria (Entemnotrochus
Putah Creek, Yolo Co. UCMP A3990. Hypotype UCMP 14236. [Pleurotomariidae]
Plicifusus (Latifusus) indomitus
(of Ogle 1953:24–26), Pliocene. Eel River, Scotia and Rio Dell, Humboldt Co. [CASG 61763]. Holotype CASG 13651
[=CASG 61763.01]. [Buccinidae].
Polinices (Euspira clementensis (M.A. Hanna, 1927). [Natica]. Marincovich 1977. BAP 70(294):290–291, pl. 27,

31003. [Naticidae].
Polinices (Euspira) diabloensis (Clark, 1915). [Natica]. Marincovich 1977. 
Pablo Fm., Miocene. Walnut Creek, Contra Costa Co. UCMP 1947]. Holotype UCMP 11595. [Naticidae].
Polinices (Euspira) galianoi-
cene. Fresno Co. UCMP 2679. Holotype (of Natica orbicularis Nomland 1916b) UCMP 12058. [Naticidae].
282 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Polinices (Polinices) gesteri (Dickerson, 1916). [Natica
Lodo Fm., Paleocene. Salt Creek area, Coalinga quad., Fresno Co. UCMP. 1817. Holotype UCMP 11829. [Naticidae].

14411.
Polinices hornii (Gabb, 1864). [Lunatia
Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4720. Hypotype UCR 4720/2. [Naticidae].
Polinices (Polinices) hornii (Gabb, 1864). [Lunatia
Tejon Fm., Eocene. Alizos Creek [=Live Oak Canyon], near Ft. Tejon, Kern Co. Lectotype (of Stewart, 1926 [1927]:
p. 324) ANSP 4214, paralectotypes ANSP. 79445. [Naticidae].

UCR 7529/1.
Polinices hornii (Gabb, 1864). [Lunatia
Fm., Paleocene. Simi Hills, Ventura Co. UCR 6670. Hypotype UCR 6670/6. [Naticidae].
Polinices (Neverita) inezana (Conrad, 1857d). [Natica  
Gaviota Fm., Eocene. West of Cuarta Canyon, Pt. Conception quad., Santa Barbara Co. UCMP B6992. Hypotype
SU 9199 [=CASG 70207]. [Naticidae].
Polinices lewisii (Gould, 1847). [Natica       
Santa Cruz Co. Hypotype determined here as missing,. Remarks: In the genus Euspira,  McLean (2007: p.
737). [Naticidae]
Polinices lewisii (Gould, 1847). [Natica
Santa Cruz Co. CASG. 66130. Hypotype CASG 66130.01. Remarks: In the genus Euspira,  McLean (2007: p.
737). [Naticidae].
Polinices (Euspira) lincolnensis (Weaver, 1916). [Natica 
Round Mtn. Silt, Miocene. Kern Co. Hypotype USNM 339910 determined here as missing. [Naticidae].
Polinices (Euspira) nuciformis (Gabb, 1864). [Lunatia6. Tejon
Fm., Eocene. Alizos Creek [=Live Oak Canyon], near Ft. Tejon, Kern Co. Lectotype (of Stewart 1926 [1927]: p. 323)

Polinices (Euspira) nuciformis (Gabb, 1864). [Lunatia
Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype UCLA
59276 [=LACMIP 10552]. [Naticidae].
Polinices (Euspira) nuciformis      Lunatia]. Squires 1987. NHMLAC CS
-
potype LACMIP 7471. [Naticidae].
Polinices (Euspira) nuciformis (Gabb, 1864). [Lunatia-
water Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 20581. Hypotype LACMIP 11653. [Naticidae].
Polinices pinyonensis Dickerson, 1914b. [Natica
Lodo Fm., Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. UCMP A9717. Hy-
potype UCMP 14210. Remarks: Crommium pinyonensis (Dickerson, 1914),  Squires (1987: p. 35). [Naticidae].

10243 [=CASG 61666.12]. Remarks: Crommium pinyonensis (Dickerson, 1914),  Marincovich (1977: p. 227).
Polinices (Euspira) ramonensis (Clark, 1918). [Natica (Euspira)]. Marincovich 1977. BAP 70 (294):269–270, pl. 26,

UCMP 11257. [Naticidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 283
Polinices reclusianus (Deshayes, 1839). [Natica Cancel-
laria tritonideaBoreotrophon sp. (not Cancellaria)] and middle row [right 6

Co. LACMIP 1219]. Hypotypes current whereabouts unknown. Remarks: In the genus Glossaulax,  McLean
(2007: p. 737). [Naticidae].
Polinices (Neverita) reclusiana var. andersoni Clark, 1918. ). [Natica (Neverita)]. Lutz 1951. UCPG 28(13):391, pl.
     Remarks: Neverita
(Glossaulax) andersoni (Clark, 1918),  Addicott (1970: p. 67);  Marincovich (1977: p. 328). In the genus
Glossaulax,  Majima (1989: p. 58). [Naticidae].
Polinices (Neverita) secta (Gabb, 1864). [Neverita
Fm., Eocene El Jaro Creek Lompoc quad., Santa Barbara Co. UCMP B7016. Hypotype SU 9200 [=CASG 70245].
Remarks: Junior synonym of Neverita (Glossaulax) reclusiana (Deshayes, 1839), Marincovich (1977: p. 318).
Determined here as in the genus Glossaulax. [Naticidae].
Polinices (Euspira) simiensis 
[=Santa Susana Fm.], Paleocene. Simi Hills, Calabasas quad., Ventura Co. [UCMP 3776]. Holotype UCMP 30710.
[Naticidae].
Polinices simiensis Nelson, 1925. Zinsmeister 1983a. SEPM Guidebook 35:69, pl. 3, -
ocene. Simi Hills, Ventura Co. UCR 4571. Hypotype UCR 4571/7. [Naticidae].
Polinices susanaensis
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype
SU/CASG 10242. [Naticidae].
Polinices (Polinices) susanaensis
Fm. [=Santa Susana Fm.], Paleocene. Simi Hills, Camulos quad., Ventura Co. UCMP 4009. Holotype UCMP 30711.
[Naticidae].

UCMPA9717]. Hypotypes UCMP 14413, 14414.

Polinices victoriana  
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotypes USNM 650101–650103. Remarks:
Polinices (Euspira) lincolnensis (Weaver, 1916),  Marincovich (1977: p. 286). [Naticidae].
Polinices
Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM 646619. [Naticidae].
Polinices (Euspira
lowermost Paleocene. Dip Creek, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo Co. UCLA 6526 [=LACMIP
26526]. Hypotype LACMIP 7561. [Naticidae].
Polinices-
son Grove Fm.], Mio-Pliocene. Sonoma Co. CASG hypotype determined here as missing. [Naticidae].
Polinices         

hypotype determined here as missing. [Naticidae].
cf. Polinices
F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to LACMIP [Naticidae].
Polinices
and bryozoans). Pancho Rico Fm., Pliocene. Priest Valley quad., San Benito Co. UCMP A3293. Hypotype UCMP
39687. [Naticidae].
284 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Polystira englishi
Miocene. This locality occupies an area in the hills just north of the Kern River, Kern Co. CASG 2064 [=CASG 65).
Holotype CASG 12937 [=CASG 2064.02), paratype CASG 12938 [=CASG 2064.03). [Turridae].
. Silt, Kern Co. Temblor locality 4, Caliente quad., Kern Co. UCMP 2716. Hypotype
UCMP 36939.
Polytropa funkeanaAnadara bed
Reef Ridge quad., Kings Co. UCMP D1205. Holotype UCMP. 36847. [Muricidae].

Pomaulax gradata
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14380. Remarks: Determined here as in the genus Mega-
straea [Turbinidae].
Pomaulax gradata-
cene. Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14381. Remarks: Determined here as in
the genus Megastraea. [Turbinidae].
Popenoeum maritimus-
quito Fm., Paleocene. Warm Springs Mtn., Fish Canyon, Warm Springs Mtn. quad. (1958), Los Angeles Co. LACMIP
21579 [exRemarks: In the genus Sulcobuccinum,  Vermeij
(1998: pp. 81, 82). [Pseudolividae].
....., Redrock Mtn. quad. (1936), Los Angeles Co. LACMIP 21579A [ex
....., North of Warm Springs Mtn., Warm Springs Mtn. quad. (1958), Los Angeles Co. LACMIP. 21587 [ex UCLA 1587].

Potamides (Potamidopsis) californica
Eocene. Reyes Peak quad. (1943), in the vicinity of Beartrap Creek, Ventura Co. LACMIP 7226. Holotype LACMIP
11300, paratypes LACMIP. 11301, 11302. [Potamididae]. Remarks: In the genus Potamidopsis,  Reid et al.
(2008: table 4 on p. 691).
Potamides (Potamidopsis) californica       
Sandstone, Eocene. Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1453 [= LACMIP 16962]. Hypotype LACMIP
12440, 12441. [Potamididae]. Remarks: In the genus Potamidopsis,  Reid et al. (2008: table 4 on p. 691).
Potamides carbonicola
Cerithidea carbonicola

[ex
Potamides (Potamides carbonicola Cooper, 1894. Givens and Kennedy 1976. Jour. Paleo. 50 (5):960, 963–964, pl. 1,

UCR 4865/30–4865/34. [Potamididae].
Potamides davisiana Cooper, 1894. Coan 1981 [1982]. Univ. 
1894,  13739. Holotype CASG 13 [ex CSMB
13739]. Remarks: Surculites mathewsonii davisiana Cooper, 1894,  Vokes (1939: p. 123). [Potamididae].
Potamides sespeensis        
Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 401 [=LACMIP 16342]. Hypotype
UCLA 58223 [=LACMIP 10571]. [Potamididae].
Potamides 
Potamides carbonicola Cooper, 1894)).
Domengine Fm. [as Tejon Fm.], Eocene. West of Coalinga, Fresno Co. Hypotype USNM 165651. Remarks: Deter-
mined here as Potamides carbonicola Cooper, 1894. [Potamididae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 285


Co. UCMP D8152. UCMP unnumbered hypotypes lost,  C. Throckmorton (personal communication, 2002).
[Potamididae].
 (Gabb, 1869b). [Uroscya
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCLA2330 [=LACMIP 23330)]. Hypotype UCLA 59252 [=LACMIP
10978]. Remarks: Error for . [Ficidae].
 (Gabb, 1869b). [Uroscya-


 (Gabb, 1869b). [Urosyca
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCLA 2330 [=LACMIP 22330]. Hypotype UCLA 59252 [=LACMIP
10978]. [Ficidae].
 (Stewart, 1926 [1927]). [Fusus (Hemifusus
Shale, Eocene. San Diego Co. UCMP 2226. Holotype UCMP 11691. [Ficidae].


LACMIP 14477.
-
MIP 14478.

14479.
 (Gabb, 1864). [Fusus (Hemifusus. 20–22. Tejon Fm.,


Priscofusus geniculus (Conrad, 1849a). [Fusus-
Fusus hecoxi n. sp.)). San Lorenzo Shale, Oligocene. Boulder Creek, Santa Cruz Co. (not
Santa Barbara Co. as in text). USGS 100. Holotype USNM 165446. Remarks: The original spelling of this species
is Fusus hecoxi, and it was named for Miss Laura J. F. Hecox. Keen and Bentson (1944: p. 162) noted, however,
that it should be amended to hecoxae. Both names are correct.[Fasciolariidae].
Priscofusus geniculus (Conrad, 1849a). [Fusus     
Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotypes USNM 650178, 650179.
[Fasciolariidae].
Priscofusus cf. geniculus (Conrad, 1849a). [Fusus

Priscofusus geniculus (Conrad, 1849a) axial ribbed form [Fusus
26, 28. Basal Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotypes USNM
650180, 650181. [Fasciolariidae].
Priscofusus geniculus (Conrad, 1849a) [Fusus-
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4312. Hypotype SDSNH 83245. [Fasciolariidae].
Priscofusus hecoxae (Arnold, 1908). [Fusus
Oligocene. San Lorenzo River, Santa Cruz Co. USGS 100. Holotype USNM 165446. [Fasciolariidae].
286 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Priscofusus medialis (Conrad, 1849a). [Cerithium
Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotypes USNM 650175, 650177.
[Fasciolariidae].



ft., Kern Co. UCR 1306. Hypotype UCR 1306/1 (not UCR.1306 as in pl. caption).
Priscofusus sp. Addicott 1979 sandstone member),
Oligocene. Near Indian Ranger Station, Santa Lucia Range, Monterey Co. USGS M6557. Hypotype USNM 264141.
[Fasciolariidae].
ProgabbiaPecten zone), Pliocene.
North Dome, Kettleman Hills, Kings Co. USGS 12335. Hypotype USNM 495758. [Cancellariidae].
Propustularia kempera* (Nelson, 1925). [Cypraea
Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 4573. Hypotype UCR 4573/20. Remarks: Error for
kemperae. [Cypraeidae].
Propustularia simiensis (Nelson, 1925). [Cypraea  
Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 4573. Hypotype UCR 4573/21. [Cypraeidae].
Protobusycon judithae
Creek area, Tumey Hills quad., Tumey Hills, Fresno Co. UCLA 6456 [=LACMIP 26456]. Holotype LACMIP 7621
(not LACMIP 7612 as in text), paratype LACMIP 7622. [Melongenidae].

-
leocene. Right Angle Canyon, Panoche Hills, Chounet Ranch quad., Fresno Co. Hypotype current whereabouts
unknown. Remarks: Aporrhaid sp.,  L.R. Saul (personal communication, 2005). [Aporrhaidae].
Proximitra cretacea (Gabb, 1864). [Mitra-
yon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6546. [Volutomitridae].
Proximitra cretacea (Gabb, 1864). [Mitra      
Eocene. Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype
LACMIP 7485. [Volutomitridae].
Psephaea (Miopleiona) cf.   (M) (Conrad, 1849a). [Rostellaria]. Addicott 1970. USGS PP 642:105, pl. 13,

UCMP 3229. Hypotype UCMP 12136. [Volutidae].

2-1, depth 4657 ft., Kern Co. UCR 1305. Hypotype UCR 1305/1 (not UCR 1305 as in pl. caption). [Volutidae].
Psephaea oregonensis (Dall, 1907). [Miopleiona]. Woodring and Bramlette 1950 [1951]. USGS PP 222:78, pl. 14,
 
locality. Hypotypes USNM 560091, 560092. Remarks: Determined here as in the genus/subgenus Musashia
(Nipponomelon). [Volutidae].
Co. CIT 869 [=LACMIP 6108]. CIT 869 [=LACMIP 6108]. Hypotype CIT
4554 [=LACMIP 4213].
Psephaea (Miopleiona) weaveri (Tegland, 1933). [Miopleiona
17. Basal Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotype USNM 650184.
[Volutidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 287
Psephaea (Miopleiona) cf.   () weaveri (Tegland, 1933). [Miopleiona]. Addicott 1970. USGS PP 642:104–105, pl.

650185. [Volutidae].
Pseudoliva dilleri
Whitaker Peak quad., Los Angeles Co. CSUN 359 [=LACMIP 16101]. Hypotype LACMIP 7481. Remarks: Calore-
bama dilleri lineata (Gabb, 1864),  Squires (1989a: p. 42); in the genus Sulcobuccinum,  Vermeij (1998: p.
8). See Squires (2001: p. 10) for comments about Calorebama versus Sulcobuccinum. [Pseudolividae].
Pseudoliva n. gen. howardi (Dickerson, 1914b). [Molopophorus  

6679/11 as in pl. caption). Remarks: In the genus Pegocomptus,  Zinsmeister (1983b: p. 1296); in the genus
Sulcobuccinum,  Vermeij (1998: p. 81). [Pseudolividae].
Pseudoliva inornata
Guillermo quad. (1943), Ventura Co. UCR 4707. Hypotype UCR 4707/31. Remarks: In the genus Calorebama,
 Squires (1989a: p. 43); in the genus Sulcobuccinum,  Vermeij (1998: p. 81). See Squires (2001: p. 10) for
comments about Calorebama versus Sulcobuccinum. [Pseudolividae].
Pseudoliva lineata
Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6542. Remarks: In the genus
Calorebama,  Squires (1989a: p. 41); in the genus Sulcobuccinum,  Vermeij (1998: p. 81). See Squires
(2001: p. 10) for comments about Calorebama versus Sulcobuccinum. [Pseudolividae].
Pseudoliva volutaeformis
Peak, Reyes Peak quad. (1943.), Ventura Co. UCR 4706. Hypotype UCR 4706/151. Remarks: Calorebama dilleri
kirbyi (Clark, 1938),  Squires (1989a:p. 44); in the genus Sulcobuccinum,  Vermeij (1998: p. 81). See Squires
(2001: p. 10) for comments about Calorebama versus Sulcobuccinum. [Pseudolividae].
Pseudoliva
and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=CASG 61666 = UCMP. A9717]. Hypotype SU 10299
[=CASG 61666.02]. Remarks: Brachysphingus mammilatus Clark and Woodford, 1927,  Squires (1997a: p. 856).

UCMP 14202. Remarks: Brachysphingus mammilatus Clark and Woodford 1927,  Squires (1997a: p. 856).
Pseudomelatoma  (Martin, 1914). [Drillia]. Faustman 1964. UCPG 41(2):127, pl.  
Dell Fm., Pliocene. Fleener Creek, Ferndale quad., Humboldt Co. UCMP B7939. Hypotype UCMP 37801. [Pseu-
domelatomidae].
Pseudoperissolax blakei (Conrad, 1855b) (not Gabb as in pl. caption). [Busycon]. G D. Hanna and Hertlein 1943.

Remarks: Family Muricidae,  Squires (2015: p. 578). [Muricidae].
Pseudoperissolax blakei (Conrad, 1855b). [Busycon-
cene. [Ft. Tejon], Kern Co. UCR 1807. Hypotype UCR 1807/15. Remarks: Family Muricidae,  Squires (2015:
p. 578). [Muricidae].
Pseudoperissolax blakei (Conrad, 1855b). [Busycon
Eocene. Grapevine Canyon, Kern Co. UCLA 2340 [=LACMIP 22340]. Hypotypes LACMIP 7647, 7648. Remarks:
Family Muricidae,  Squires (2015: p. 578). [Muricidae].
Pseudoperissolax blakei (Conrad, 1855b). [Busycon 
Remarks:
Family Muricidae,  Squires (2015: p. 578). [Muricidae].
-
MIP 22340. Hypotypes LACMIP.7647, 14498, 14500–14508.

288 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pseudoperissolax blakei praeblakei
Fm.” [=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH 1667. Hypotype SDSNH 4148]. Re-
marks: Junior synonym of  and in family Muricidae,  Squires (2015: p. 578). [Muricidae].
Pseudoperissolax blakei praeblakei
Thorn Meadows, Lockwood Valley quad. (1943), Ventura Co. UCR 4682. Hypotype UCR 4682/7. Remarks: Junior
synonym of  and in family Muricidae,  Squires (2015: p. 578). [Muricidae].
        
4685/121.
Pseudoperissolax blakei subsp. praeblakei 
Lodo Fm., Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU.2073 [=UCMP
A9717]. Hypotype SU 10238 [=CASG 61666.22]. Remarks: Junior synonym of  and in family Muricidae,
 Squires (2015: pp. 578, 583). [Muricidae].
Pseudoperissolax blakei praeblakei
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP
6547 Remarks: Junior synonym of  and in family Muricidae,  Squires (2015: pp. 578, 583). [Muricidae].
Pseudoperissolax blakei praeblakei
Piru Creek, Cobblestone Mtn. quad. (1958), Ventura Co. CSUN 808 [=LACMIP 16344]. Hypotype LACMIP 7486.
Remarks: Junior synonym of  and in family Muricidae,  Squires (2015: pp. 578, 583). [Muricidae].
Pseudoperissolax merriami
Sobrante Ridge, Contra Costa Co. UCMP 3055. Paratype UCMP 11286. Remarks: Aforia clallamensis (Weaver,
1916) (family Turridae),  Javidpour (1973: p. 201). [Turridae].
Pseudoperissolax tricarinatus (Weaver, 1905). [Perissolax  
5. Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 4569. Hypotype UCR 4569/26, determined here as
missing. Remarks: Family Muricidae,  Squires (2015: p. 578). [Muricidae].
Pseudoperissolax tricarinatus (Weaver, 1905). [Perissolax
Santa Susana Fm., Paleocene. Santa Ynez Canyon, Los Angeles Co. LACMIP 16888. Hypotype LACMIP 14492.
Remarks: Family Muricidae,  Squires (2015: p. 578). [Muricidae].
-
potypes LACMIP 14493, 14494, 11496, 11497.

UCMP 11882.

LACMIP 14495.
Pseudovertagus (Pseudoaluco 
quad. (1958), Los Angeles Co. CSUN 360 [=LACMIP 16109]. Hypotype LACMIP 7457. [Cerithiidae].
  
quad., Mendocino Co. UCMP D7149. Hypotype 14435. [Limacinidae].
Pteropurpura
Fm., Pliocene. Arco Towers excavation, 6th and Flower streets, downtown Los Angeles, Los Angeles Co. [LACMIP
1219]. Hypotype current whereabouts unknown. [Muricidae].
Pterynotus (Pterynotus) cf. Pterynotus (Pterynotus) washingtonicus Squires and Goedert, 1995. Squires 2001. NHM-

LACMIP 16654. Hypotype LACMIP 12736. [Muricidae].
Puncturella cooperi aphales Woodring in Woodring and Bramlette, 1950 [1951] (n. var.). USGS PP 222:69, pl. 12,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 289

Holotype USNM 560060. [Fissurellidae].
Puncturella cucullata (Gould, 1846). [Rimula]. Woodring and Bramlette 1950 [1951]. USGS 
5, 6. Foxen Mudstone, Pliocene. Santa Barbara Co. USGS 4473. Hypotype USNM 560042. [Fissurellidae].
Purpura milicentana      
Vaqueros Fm., Miocene. North coast Santa Rosa Is., California Channel Ids. UCSB SR67-79 [=LACMIP 16432].
Hypotype UCSB M21 [=LACMIP 6310]. [Muricidae].
Purpura turris
West of Coalinga, Fresno Co. UCMP 2110. Holotype UCMP 12049. Remarks: In the genus Califostoma,  Bean
and Vermeij (2016: p. 2). [Muricidae].
Purpura-
rial. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8815 [=SBMNH 60196]. Remarks:
Determined here as indet. muricid sp. [Muricidae].
Pyramidella (Longchaeus) cooperi 
Lower Round Mtn. Silt, Miocene. Caliente quad. (1914), Kern Co. UCMP B1638. Hypotype UCMP 37109. [Pyra-
midellidae].

Pyramidella (Syrnola) ochsneri (Anderson and Martin, 1914). [Eulimella]. Addicott, 1970. USGS PP 642:142, pl. 21,

[Pyramidellidae].


Pyramidella preblei     
Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP
7503. [Pyramidellidae].
Pyrgulifera (Pyrgulifera) lajollaensis (M.A. Hanna, 1927). [Trichotropis]. Squires 1999c. NHMLAC CS 480:13–14,

Hypotype LACMIP 12449. [Thiaridae].
-
cene. Meier Canyon, Simi Hills, Calabasas quad., Ventura Co. UCR 6670. Paratype UCR 6670/901. Remarks: Not
a Pyropsis, Squires (2011: p. 1204). [Turbinellidae].



Las Virgenes Fm., Paleocene. Las Virgenes Canyon, Simi Hills, Calabasas quad., Ventura Co.
UCLA 3111 [=LACMIP 23111]. Paratype LACMIP 7616.

Pyropsis gabbi (Stanton, 1896). [Heteroterma    

Remarks: Not a Pyropsis, Squires (2011: p. 1204). [Turbinellidae].

UCLA 2307 [=LACMIP 22307]. Hypotype LACMIP 7599.
290 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Pyropsis striata (Stanton, 1896). [Heteroterma
Paleocene. Pacheco syncline area, Contra Costa Co. LACMIP 10488 (ex UW 2080). Hypotypes LACMIP 7618, 7619.
Remarks: Not a Pyropsis, Squires (2011: p. 1204). [Turbinellidae].
Pyruconus cf. -
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4312. Hypotype SDSNH 140190. [Conidae].
Pyruconus cf. -
terey Fm. (Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan
Capistrano quad. SDSNH 4312. Hypotype SDSNH 83241. [Conidae].
Ranella californica (Gabb, 1866). [Tritonium
Gyrineum kewi (Dickerson,
1915)) UCMP 30878. [Ranellidae].

(lectotype, Smith (1970: p. 522)) ANSP 4205.


Ranella katherineae
Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. UCLA 2312 [=CSUN 374, =LACMIP 7242]. Holotype
UCLA 45969 [=LACMIP 10546]. [Ranellidae].
Ranella katherineae
Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Holotype UCLA 45969 [=LACMIP
10546]. [Ranellidae].
Ranella louella
junction of Panoche Creek and Silver Creek, Tumey Hills, Fresno Co. UCLA 6456 [=LACMIP 26456]. Holotype
LACMIP 7689. [Ranellidae].
Ranella pilsbryi Stewart, 1926 [1927]. Givens Near Pine Mtn.
Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4721. Hypotype UCR 4721/112. [Ranellidae].
Ranella pilsbryi
Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6534. [Ranellidae].
Rapana vaquerosensis (Arnold, 1907). [Purpura
2. Vaqueros Fm., Miocene. Sespe Creek area, Topatopa Mtns., Ventura Co. CSUN 384 [=LACMIP 16351. Hypotype
UCLA 58218 [=LACMIP 10575]. [Muricidae].
     
UCLA 58219 [=LACMIP 10576].
     
UCLA 58221 [=LACMIP 10577].

UCLA 58200,58201 [=LACMIP 10573,10574].
Rapana vaquerosensis imperialis Hertlein and Jordan, 1927. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):176,
ex CSMB 12322)
determined here as missing. [Muricidae].
Rapana vaquerosensis (Arnold, 1907) imperialis Hertlein and Jordan, 1927. Avila and Weaver 1969. AAPG/SEPM

UCSB. SR64-12 [=LACMIP 16433]. Hypotype UCSB M22 [=LACMIP 6312]. [Muricidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 291
Rapana vaquerosensis (Arnold, 1907) imperialis
Vaqueros Fm., Miocene. Santa Ana Mtns., Orange Co. UCMP 6128 [=LACMIP 16443]. Hypotype LACMIP10612.
[Muricidae].
Rapana vaquerosensis imperialis Hertlein and Jordan, 1927. Squires and Filewicz (eds.) 1983. SEPM Guidebook

12720]. [Muricidae].
Rapana vaquerosensis imperialis

Field Area, north side Simi Valley, Ventura Co. LACMIP 17097. Hypotype LACMIP 1272]. [Muricidae].
Ravinella lipmanorum
Susana Fm., Paleocene. Temporary locality now covered by landscaping and sprinklers, north side of Simi Valley,
Ventura Co. LACMIP 41691. Holotype LACMIP 14724, paratypes LACMIP 14725–14728. [Acteonidae].
Retipirula crassitesta (Gabb, 1869b). [Turbinella]. G D. Hanna and Hertlein 1943. CDM Bull. 
62-24. Martinez Fm., Eocene [Paleocene]. Santa Susana Mtns., Los Angeles Co. CASG. 396. Hypotypes CASG 5915,
5915A. Remarks: In the genus/subgenus Volutocorbis (Retipirula),  Zinsmeister (1977: p. 178). [Volutidae].
Retipirula crassitesta (Gabb, 1869b). [Turbinella
Santa Susana Fm., Paleocene. Santa Susana quad., NW slope of Simi Hills, Ventira Co. LACMIP 22307. Hypotype
LACMIP 13221. [Volutidae].
Retipirula pinguis   
Fm., Paleocene. Calabasas quad., Ventura Co. LACMIP 22701. Holotype LACMIP 13222, paratypes LACMIP 13223,
13224. [Volutidae].

Rictaxis painei grandior
area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14415. Remarks: Junior synonym of Rictaxis painei (Dall
1903c),  Valdés (2019: p. 246). [Acteonidae].
Rimella macilenta White, 1889. [Rimella
Eocene. Devil Canyon, Santa Susana Mtns., Los Angeles Co., LACMIP 40371 [=CSUN 371]. Hypotypes LACMIP
14445, 14453. [Strombidae].

     
Hypotype LACMIP 14456.

14459.

14458.

14459.

Rimella supraplicata Neptunea     
Fm. (Metralla Sandstone Member), Eocene. Tehachapi Mtns., Kern Co, most likely LACMIP 7155. Lectotype (of
Stewart 1926 [1927]: p. 366) ANSP 4231, of Ectinochilus (Cowlitzia) canalifer (Gabb, 1864), paralectotypes (8)
ANSP 79467. Remarks:

292 PALEOBIOS, VOLUME 38, FEBRUARY 2021
771 as in plate caption). Hypotype LACMIP 14462.

LACMIP 14466.
Diego Co., UCMP 5062. Holotype (of Cowlitzia
problematica M.A. Hanna, 1927) UCMP 31144.

Rimula cancellata
Pliocene. Whitewater River, San Jacinto quad., Riverside Co. UCMP A1407. Holotype UCMP 16076, paratype UCMP





Ringicula (Ringicula) pinguis (Gabb, 1864). [Cinulia
Fm., Eocene. South side of Simi Valley, Calabasas quad. (1952, PR 1967), Ventura Co. CSUN 1344 [=LACMIP 17192].
Hypotype LACMIP 12690. [Ringiculidae].
Sassia bilineata (Dickerson, 1916). [Fasciolaria
Alamo Creek, San Guillermo quad. (1943), Ventura Co. UCR 4707. Hypotype UCR 4707/23. [Ranellide].
Sassia bilineata (Dickerson, 1916). [Fasciolaria    
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 802 [=LACMIP 16144]. Hypotype LACMIP 7476. [Ranellidae].
Saxituberosa fons
Paleocene (Turritella peninsularis qualeyi zone of Saul, 1983:30–33). Towards Warm Springs Summit, Los Angeles
Co. LACMIP 21581 (not LACMIP 1588 as in text). Holotype LACMIP 13354. [Fasciolariidae].
-
MIP 13355.

21588 (not LACMIP 1588 as in pl. caption). Paratype LACMIP 13356.
Saxituberosa titan (Waring, 1917). [Trachytriton
Susana Fm., Paleocene. Simi Hills, Calabasas quad. (1952, PR 1967), Ventura Co. LACMIP 22688. Hypotype LAC-
MIP 10983 [ex UCLA 59253]. [Fasciolariidae].
ex UCLA 59254].
Scalina durhami (Keen, 1943). [Ferminoscala]. Addicott 1970. USGS PP 642:56, pl. 3
Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Hypotype USNM 650082. [Epitoniidae].
....., pl. 3, . 24. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1611. Hypotype USNM 650083.
Scalina whitei (Keen, 1943). [Ferminoscala
Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Hypotypes USNM 650078, 650079. [Epitoniidae].


GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 293
Scaphander (Mirascapha) costatus (Gabb, 1864). [Cylichna
Scaphander costatus
Co. Lectotype (of Stewart 1926 [1927]: p. 437) ANSP 4338, paralectotypes (8) ANSP 79477]. [Cylichnidae].
Scaphander (Mirascapha) costatus (Gabb, 1864). [Cylichna    

4163]. [Cylichnidae].
Scaphander (Mirascapha) costatus (Gabb, 1864). [Cylichna
Eocene. Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 493 [=LACMIP 16198]. Hypotype LACMIP 6561
(not LACMIP 6562 as in pl. caption). [Cylichnidae].
Scaphander (Mirascapha) costatus (Gabb, 1864). [Cylichna

[=LACMIP 16100]. Hypotype LACMIP 7502. [Cylichnidae].
Scaphander jugularis (Conrad, 1855b). [Bulla

UCMP 15179. [Cylichnidae].
Scaphander jugularis (Conrad, 1855b) [Bulla
Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650278. [Cylichnidae].


Scaphander jugularis (Conrad, 1855b). [Bulla
Fm. (Saltos Shale Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M5026. Hypo-
type USNM 254357. [Cylichnidae].
Scaphander jugularis (Conrad, 1855b). [Bulla]. Addicott et al. 1980. SEPM Guidebook 6:18,20, pl. 
from Addicott et al., 1978, pl. Creek area, La Panza
Range, San Luis Obispo Co. USGS M5026. Hypotype USNM 254357. [Cylichnidae].
Scaphander cf. stewarti
Monocline Ridge, Monocline Ridge quad. (1955), Fresno Co. UCMP D6532]. Hypotype UCMP 37170. [Cylichnidae].
Scelidotoma aldersoni Powell and Geiger, 2019 (n. sp.). PaleoBios
Old Topanga [Canyon] Rd., Los Angeles Co. LACMIP 30503. Holotype LACMIP 14846, paratype SMBNH 634129.
[Fissurellidae].
Scobinella claytonensis (Gabb, 1864). [Turris
-
totype (of Stewart 1926 [1927]: p. 417) ANSP 4190, paralectotype ANSP 4190a (=ANSP 79429]. [Clathurellidae].
Searlesia portolaensis (Arnold, 1908). [Fusus-
cene. Coalinga quad., Fresno Co. UCMP B6545. Hypotype UCMP 36873. Remarks: In the genus Lirabuccinum,
 Vermeij (1991: p. 269). [Buccinidae].
Searlesia portolaensis (Arnold, 1908). [Fusus
Capitola, Soquel quad. Santa Cruz Co. SCCM 40. Hypotype SCCM 21015. Remarks: In the genus Lirabuccinum,
 Vermeij (1991: p. 269). [Buccinidae].
?Serpulorbis llajasensis
Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. UCLA 2313 [=LACMIP 22313]. Holotype LACMIP 8091,
paratype LACMIP. 8092. [Vermetidae].
Serratocerithium usanium
[=Santa Susana Fm.], Paleocene. Ridge east of Santa Ynez Canyon, Santa Monica Mtns., Los Angeles Co. SU 2691.
294 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Holotype SU 7756 [=CASG 72063], paratype SU 7757 (not SU 7758 as in pl. caption) [=CASG 72064]. [Cerithiidae]
Sinum obliquum (Gabb, 1864). [Naticina
[=possibly Ardath Shale], Eocene. Rose Canyon, San Diego Co. SDSNH 1667. Hypotype SDSNH 4159]. [Naticidae].
Sinum obliquum (Gabb, 1864). [Naticina  
  
ANSP 79450. [Naticidae].
Sinum coryliforme Anderson and G D.Hanna,
1915) CASG 851.
Sinum obliquum (Gabb, 1864). [Naticina
Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6528. [Naticidae].
Sinum obliquum (Gabb, 1864). [Naticina.
Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP
7473. [Naticidae].
Sinum obliquum (Gabb, 1864). [Naticina
Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 983 [=LAMCIP 16110]. Hypotype LACMIP 7708. [Naticidae].
Sinum perrini (Arnold, 1907). [Sigaretus
Head of Topanga Canyon, south of Calabasas, Camulos quad., Los Angeles Co. USGS 4402. Holotype USNM 164979.
Sinum scopulosum (Conrad, 1849a). [Sigaretus
5. Careaga Fm. (Cebada Member), Pliocene. Fugler Pt., Santa Barbara Co. Alex Clark locality. Hypotype USNM
560072. [Naticidae].
Sinum scopulosum (Conrad, 1849a). [Sigaretus
Pliocene. Reynard Wy., San Diego, San Diego Co. SDSNH 2383. Hypotype SDSNH 4172]. [Naticidae].
Sinum scopulosum (Conrad, 1849a). [Sigaretus 
Miocene. Kern River, Kern Co. USGS 6622. Hypotype USNM 650110. [Naticidae].


650112.
Sinum cf.  (Conrad, 1849a). [Sigaretus-
blor Fm. (Wygal Sandstone Member), Miocene. Carneros Rocks quad., Kern Co. USGS M3578. Hypotypes USNM
646623, 646624. [Naticidae].
Sinum scopulosum (Conrad, 1849a). [Sigaretus
Pliocene. San Fernando, Los Angeles Co. Lectotype (of Stewart 1926 [1927]:328) ANSP 4326 (of Sinum planico-
stum

(of Sinum (Sigaretus) trigenarium Trask, 1922. UCMP 12386.
Sinum scopulosum (Conrad, 1849a). [Sigaretus-
whall area, Los Angeles Co. LACMIP 7757. Hypotype LACMIP 14391. [Naticidae].
Sinum cf.  (Conrad, 1849a). [Sigaretus
Monterey Fm. (Saddleback Valley Limestone member). Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San
Juan Capistrano quad. SDSNH 4312. Hypotype SDSNH 93721. [Naticidae].
Siphonalia (Eosiphonalia) andersoni Wagner and Schilling. Pleito Fm.,
Oligocene. North of Devil's Kitchen, San Emigdio Mtns., Kern Co. UCMP 3200. Holotype UCMP 11431. [Buccinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 295
Siphonalia (Nassicola) bicarinata crooki-
kley Fm., Eocene. North of Vacaville, Solano Co. UCMP A1297. Holotype UCMP 30899, paratype UCMP 12462,
hypotype UCMP 30427. [Buccinidae].
Siphonalia (Nassicola) bicarinata wheatlandensis

UCMP 30421, paratype UCMP.11271, hypotypes UCMP 30421, 30422, 30424, 30425, 30433. [Buccinidae].
Siphonalia (Eosiphonalia) californica (Arnold, 1908). [Strepsidura    

Santa Cruz Co. UCMP 108. Holotype USNM 165450. [Buccinidae].
Siphonalia cf.  (Gabb, 1869a). [Neptunea
Joaquin Fm. (Acila Zone), Pliocene. Arroyo Hondo, North Dome, Kettleman Hills, Kings Co. USGS 12337. Hypotype
USNM 495716. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Siphonalia kettlemanensis (Arnold, 1909 [1910]). [Thais]. Woodring et al. 1940 [1941]. USGS PP 195:87, pl. 31,
Siphonalia Zone), Pliocene. La Aleta, North Dome, Kettleman Hills, Kings Co. USGS 12712.
Hypotype USNM 495807. Remarks: In the genus Pusio,  Vermeij (2001: p. 296). [Buccinidae].

Siphonalia kettlemanensis (Arnold, 1909 [1910]). [Thais-
chegoin Fm., Pliocene. Coalinga quad., Fresno Co. UCMP B6544. Hypotype UCMP 36866. Remarks: In the genus
Pusio,  Vermeij (2001: p. 296). [Buccinidae].
Siphonalia (Kelletia) merriami
Fm., Oligocene. San Emigdio Canyon, Kern Co. UCMP 3177. Holotype UCMP 11427. [Buccinidae].

Siphonalia merriami
Fm., Eocene. Near Gaviota Pass, Santa Ynez Mtns., Lompoc quad., Santa Barbara Co. UCMP B6964. Hypotype SU/
CASG 9503 determined here as missing. Remarks: Label with CASG. 9229 [=Siphonalia sp.] in text, p. 192, credits

        
B6980. Hypotype SU 9228 [=CASG 70231].
Siphonalia (Kelletia) posoënsis Anderson and Martin, 1914. Ruth 1942. UCPG 26(3):
from Anderson and Martin Obispo Co. CASG 126. Holotype
CASG 174. Remarks: In the genus Kelletia,  Addicott (1970: p. 92). [Buccinidae].
Siphonalia sopenahensis (Weaver, 1912). [Hemifusus
Northeast of Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4719. Hypotype UCR 4719/16. [Buc-
cinidae].
Siphonalia (Nassicola) sutterensis
Eocene. West Butte, Marysville Buttes, Sutter Co. UCMP 1853. Holotype UCMP 11764. [Buccinidae].
Siphonalia sutterensis
West of South Butte, Marysville Buttes quad., Sutter Co. CASG 248. Hypotype CASG. 5710. [Buccinidae].
Siphonalia (Nassicola) tularensis-

Holotype CASG 854. [Buccinidae].
Siphonalia (Nassicola) thunani (Dickerson 1916). [Nyctilochus
(Siphonalia sutterensis Zone), Eocene. North of Oroville, Butte Co. UCMP 2225. Holotype UCMP 12262. [Buccinidae].
296 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Siphonalia vistaensis
   
paratype UCR 4865/121. [Buccinidae].
Siphonalia
Remarks: In the genus Pusio,  Vermeij (2001: p. 296).
[Buccinidae].
Siphonalia-
cene. Media Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3280. Hypotype USNM. 646618. [Buccinidae].
Solariella crenulata (Gabb, 1864). [Margaritella
Thorn Meadows, Lockwood Valley quad. (1943), Ventura Co. UCR 4681. Hypotype UCR 4681/201. [Solariellidae].
Solariella walkeri
San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Holotype LACMIP 13414, paratype LACMIP 13415.
[Solariellidae].
Solariella  
Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14648. Hypotype USNM 560064. [So-
lariellidae].
Solenosteira sp. Deméré and Rugh 2006, 3.9. Deguynos Fm.
(Camels Head Member), Pliocene. Fish Creek/Vallecito Creek, San Diego Co. ABDSP 2553. Hypotypes ABDSP I
326, I 327. Remarks: Fig. 3.3 =Cymia sp.,  S. Rugh (personal Calicantharus
sp.,  L. Gilbert (personnel communication, 2019). [Buccinidae].
   -
leocene. Santa Ynez Canyon area, Santa Susana Mtns., Topanga quad. (1952, PR 1981), Los Angeles Co. LACMIP
16869. Holotype LACMIP 12873. [Strombidae].
Spirocrypta pileum (Gabb, 1864). [Crypta (Spirocrypta
Tejon Fm., Metralla Sandstone Member, Eocene. Live Oak Canyon, Pastoria Creek quad. (1958, PR 1974), Kern
Co. LACMIP 22386. Hypotype LACMIP 13395. [Calyptraeidae].
Sthenorytis stearnsii (Dall, 1892). [Scala (StenorytisScala in pl. cap-

 (Gabb, 1864). [Whitneya
)). Tejon Fm., Eocene. Tejon quad., Kern Co. CASG 245. [=CASG
66189]. Hypotype CASG 332 [=CASG 66189.02]. [Fasciolariidae].
 (Gabb, 1864). [Whitneya-


 (Gabb, 1864). [Whitneya-
ated Sacate/Gaviota fms., Eocene. Nojoqui CreekLompoc quad., Santa Barbara Co. UCMP B6933. Hypotypes SU
9233, 9234 [=CASG 70250, 70251]. [Fasciolariidae].
 (Gabb, 1864). [Whitneya  
Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4721. Hypotype UCR 4721/131. [Fasciolariidae].
 (Gabb, 1864). [Whitneya
Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH 2962. Hypotype
SDSNH 20773. [Fasciolariidae].
 (Gabb,1864). [Whitneya
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 452 [=LACMIP 16364]. Hypotype LACMIP.
6543. [Fasciolariidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 297
 (Gabb, 1864). [Whitneya-
ker Peak quad. (1958), Los Angeles Co. CSUN 360 [=LACMIP 16109]. Hypotype LAMCIP 7482. [Fasciolariidae].
 (Gabb, 1864). [Whitneya
Simi Valley, Ventura Co. CSUN 452 [=LACMIP 16364]. Hypotype LACMIP 6543. [Fasciolariidae].
 (Gabb, 1864). [Whitneya. 1.7–1.10. Tejon Fm.,
Eocene. Grapevine Creek, Kern Co. CASG 245. Hypotypes CASG 66189.01, 66189.02, 66190.01. [Fasciolariidae].
Streptochetus californiana-
ker Peak quad. (1958), Los Angeles Co. CSUN 845 [=LACMIP 16137]. Holotype LACMIP 7480. [Fasciolariidae].
Strictispira (Crassispira) zizyphus        
Fm., Pliocene. Goat Canyon, near U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotype LACMIP 14855.
[Strictispiridae].
Strioterebrum martini (English, 1914a). [Terebra]. Woodring and Bramlette 1950 [1951]. USGS PP 222:80, pl. 16,
      
560094. [Terebridae].
Strombina    
Ranch quad. (1954), Kern Co. USGS M1605. Hypotype USNM. 650142. [Columbellidae].
Strombus galeatus       
Latrania Fm., Miocene/Pliocene. Coyote Mtn., Imperial Co. SDSNH 2895. Hypotype SDSNH 12148. Remarks: In
the genus Lobatus,  Liverani (2013: p. 26). [Strombidae].
Strombus galeatus
Pliocene. Oasis quad., Imperial Co. (not San Diego Co. as in locality description). UCMP A1708. Hypotype UCMP
29878. Remarks: In the genus Lobatus,  Liverani (2013: p. 26). [Strombidae].
Strombus cf. 
(Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, Palos Verdes Hills, Los Angeles Co.
USGS 13127. Hypotypes USNM 497071, 497072. [Strombidae].
Strombus obliteratus           
[=Latrania Fm.], Miocene. Alverson Canyon area, southern Coyote Mtns., Imperial Co. UCR. 7267. Hypotype UCR
7267/9. Remarks: In the genus Persististrombus,  Kronenberg and Lee (2007: p. 257). [Strombidae].
Strombus obliteratus
B. Latrania Fm., Miocene/Pliocene. [Coyote Mtn., Imperial Co. SDSNH 2895]. Hypotype SDSNH 12155. Remarks:
In the genus Persististrombus,  Kronenberg and Lee (2007: p. 257). [Strombidae].
Strombus
Imperial Co. Hypotype probably at SDSU, current whereabouts unknown.
Sulcoretusa? israelskyi Addicott, 1970 (n. sp.). USGS PP 642:140–141, 
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650290, paratypes USNM 650287,
650288, hypotype USNM 650289. [Acteonidae].
Surcula crenatospira        
ex CSMB
ex CSMB 13733]. Remarks: In the genus/subgenus Fusiturricula (Crenaturricula),
 Vokes (1939: p. 115). [Clavatulidae].
Surculites mathewsonii (Gabb, 1864). [Fusus          
Thorn Meadows, Lion Canyon quad. (1943), Ventura Co. UCR 4679. Hypotype UCR 4679/4. [Fasciolariidae].
-
type UCR 4682/2. [Fasciolariidae].
298 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Surculites mathewsonii (Gabb, 1864). [Fusus
Paleocene. Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717].
Hypotype SU 10237 [=CASG 61666.13]. Remarks: In the genus Surculites,  Squires (1987: p. 48.) [Fascio-
lariidae].
Surculites mathewsonii (Gabb, 1864). [Fusus
Jolla Fm.,” Eocene. San Clemente Canyon., San Diego Co. SU 30628, [=CASG 72184]. SU hypotype 10206 [=CASG
72814]. [Fasciolariidae].
Surculites cf.  (Gabb, 1864). [Fusus
Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6670. Hypotype UCR 6670/14. [Fasciolariidae].
Surculites mathewsonii (Gabb, 1864). [Fusus
Canyon, Santa Susana quad., Los Angeles Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6555. [Fasciolariidae].
Surculites mathewsonii (Gabb, 1864). [Fusus
Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=LACMIP 16114]. Hypotype LACMIP 7493.
[Fasciolariidae].
Surculites mathewsonii (Gabb, 1864). [Fusus
Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 671 [=LACMIP 16076]. Hypotype LACMIP
11474. [Fasciolariidae].
Surculites sinuatus (Gabb, 1864). [Conus   
Surcula (Surculites) sinuata)). Tejon Fm., Eocene. Ft. Tejon
 
 Keen and Bentson, (1944: p. 197). [Fasciolariidae].
Symmetrocapulus
Co. CSUN 1201 [=LACMIP 12022]. Hypotype LACMIP 8048. [Symmetrocapulidae].
Syrnola scandix-
cene. Kern Co. SU 2121. Holotype SU 7544 and paratypes SU 7544a, b [=CASG 72011]. Remarks: Junior synonym
of Pyramidella (Syrnola) ochsneri (Anderson and Martin 1914),  Addicott (1970: p. 142). [Pyramidellidae].
Taranis strongi (Arnold, 1903). [Mangilia
Pliocene. Fleener Creek area, Ferndale quad., Humboldt Co. UCMP B7939. Hypotype UCMP 37802. [Borsoniidae].
Tegula (Chlorostoma) dalli
Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22643 [=UCLA 2643]. Hypotype LACMIP
13077 [=UCLA 57650]. [Tegulidae].
Tegula (Omphalius) dalli arnoldi
Miocene. Caliente quad. (1914), Kern Co. UCMP B1599. Paratype UCMP 11062. [Tegulidae].


Tegula hemphilli
Beach area, San Diego Co. UCMP 12661. Holotype UCMP 31208. Remarks: Pliocene age based on specimen pres-
ervation, which is distinctly different from Pleistocene preservation at this locality,  G.L. Kennedy (personal
communication, 2005). [Tegulidae].
Tegula (Omphalius) laevis
Rio Bravo Ranch quad. (1954), Kern Co. USGS M1598. Paratype USNM 650042. [Tegulidae].

Tegula varistriata
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 299
Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2878. Hypotype USNM 254374. [Tegulidae].
Tegula varistriata-

Co. USGS M2878. Hypotype USNM 254374. [Tegulidae].
Tegula
Kern Co. UCR 1235. Hypotype UCR 1235/1 (not UCR 1235 as in pl. caption). [Tegulidae].

Teinostoma (Teinostoma?) lens Keen, 1943 (n. sp.). Tr. 
Silt, Miocene. Kern Co. SU locality 2121. Holotype SU 7545 [=CASG 72012],  paratype SU 7964 [=CASG
7964]. Remarks: In the genus/subgenus Vitrinella (Vitrinellopsis),  Addicott (1970: p. 46). [Vitrinellidae].
Tejonia lajollaensis (Stewart, 1926 [1927]). [Amaurellina]. G D. Hanna and Hertlein 1943. CDM Bull. 118(2):172,
sic] Llajas Fm., Eocene. In Aliso Canyon of Devil Creek, Santa Susana quad., Los Angeles Co. CASG
393. Hypotype CASG 5666. Remarks: Tejonia moragai (Stewart, 1926 [1927]),  Marincovich (1977: p. 233).
[Naticidae].
Tejonia lajollaensis (Stewart, 1926 [1927]). [Amaurellina
Park Canyon, San Guillermo quad. (1943), Ventura Co. UCR 4703. Hypotype UCR 4703/151. Remarks: Tejonia
moragai (Stewart, 1926 [1927]),  Marincovich (1977: p. 233). [Naticidae].
Tejonia moragai (Stewart, 1926 [1927]). [Natica
Peak area, Reyes Peak quad. (1943), Ventura Co. UCR 4706. Hypotype UCR 4706/101. [Naticidae].
Tejonia moragai (Stewart, 1926 [1927]). [Amaurellina   
Tejon Fm., Eocene. Live Oak Canyon, Kern Co. UCMP 7200. Holotype UCMP 31387. [Naticidae].
Amaurellina
moragai lajollaensis Stewart, 1926 [1927]) UCMP 31401.


       
UCMP 14408.
Tejonia moragai (Stewart, 1926 [1927]). [Natica
Canyon, Santa Susana quad., Los Angeles Co. CSUN 467 [=LACMIP 16079]. Hypotype LACMIP 6525. [Naticidae].
Tenagodus californiensis
Canyon, Simi Hills, Thousand Oaks quad. (1950, PR 1967), Ventura Co. CSUN 1290 [=LACMIP 16370]. Holotype
LACMIP 8086. [Siliquariidae].

(1952, PR 1967), Los Angeles Co. UCLA 7108 [=CSUN 354, =LACMIP 27108]. Paratypes LACMIP 8087, 8088.
Terebellum (Terebellum) erraticum (Cooper, 1894). [Tornatina]. Weaver and Kleinpell 1963. UCPG 43:189, pl. 25,
de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP B6922. Hypotype
SU 9209 [=CASG 70246]. Remarks: In the genus Paraseraphs (family Seraphsidae),  Squires (1984: p. 23;
1987: p. 33). [Seraphsidae].
Terebra (Fusoterebra?) adelaidana
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650232. [Terebridae].
Terebra californica
Susana quad., Los Angeles Co. CSUN 467 [=LACMIP 16079]. Hypotype LACMIP 6560. [Terebridae].
300 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Terebra (Terebra) cooperi    
Upper Olcese Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotypes USNM 650235–650238.
[Terebridae].

Terebra cooperi
Heights, Santa Ana Mtns., Orange Co. Hypotype possibly at SDSNH, current whereabouts unknown. Remarks:
Ex Grouard collection. [Terebridae].
Terebra gausapata-
cene. Coyote Mtn., Imperial Co. ABDSP 5155.1. [Hypotype ABDSP I 12525. [Terebridae].
Terebra (Strioterebrum) martini
Pliocene. Reynard Wy., San Diego, San Diego Co. SDSNH2383. Hypotype SDSNH 4143]. [Terebridae].
Terebra (Strioterebrum) martini-
whall area, Los Angeles Co. LACMIP 17918. Hypotype LACMIP 14409. [Terebridae].
Terebra (Strioterebrum) stirtoni
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650228, 650229. [Terebridae].

Terebra wattsiana
 Keen and Bentson (1944: p.
200). [Terebridae].
Terebra (Strioterebrum
Oil Center quad., Kern Co. USGS M1597. Hypotypes USNM. 650230, 650231. [Terebridae].
Terebra (Terebra) n. sp. Addicott Upper Olcese Sand, Miocene. Oil
Center quad. (1954), Kern Co. USGS M1597. Hypotypes USNM 650233, 650234. [Terebridae].
Terebralia susana
Quarry Canyon, Santa Monica Mtns., Topanga quad. (1952, PR 1981), Los Angeles Co. LACMIP 26814. Holotype
LACMIP 7942. [Potamididae]. Remarks:Terebralia as used by Reid et al.
(2008),  Reid et al. (2008: p. 692), but they gave no indication as to what genus it is.
Thais (Thaisella) blakei
Sand, Miocene. Locality occupies an area nearly half a mi. square in hills just north of the Kern River, Kern Co.
CASG 65. Hypotype CASG 12934. [Muricidae].
Thais collomi-
cene/Pliocene. Priest Valley quad., Monterey Co. UCMP A4946. Hypotype UCMP 36471. Remarks: In the genus
Nucella,  Addicott (1978: p. 88). [Muricidae].
Thais (Thaisella) edmondi
Oil Center quad. (1954), Kern Co. USGS M1601. Hypotype USNM 650141. [Muricidae].

UCLA 45767 [=LACMIP 8434].
Thais etchegoinensis   
Rico Fm., Miocene/Pliocene. Vicinity of San Lucas, Monterey Co. SU C1001. Hypotype SU 9755 [=CASG 70325].
[Muricidae].
Thais imperialis (Dall, 1909). [Chrysodomus
of Irvington, La Costa Valley quad. (1943), Alameda Co. SU 3252. Hypotype SU 8475 [=CASG 69927]. [Muricidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 301
Thais (Nucella) lamellosa (Gmelin, 1791). [Buccinum

Hypotype UCMP 37669. Remarks: In the genus Nucella,  Abbott (1974: p. 182. [Muricidae].
Thais lima (Gmelin, 1791). [Murex
Pleasanton, La Costa Valley quad. (1943), Alameda Co. SU 2739. Hypotype SU 8474 [=CASG 69981]. Remarks:
Nucella tokudai (Yokoyama, 1932),  Amano et al. (1993: p. 240). [Muricidae].
Thais (Nucella) cf. () lima (Gmelin, 1791 [not Martyn, 1788 as cited]). [Murex]. Glen, 1959. UCPG 36(2):182, pl.

UCMP B4794. Hypotype UCMP 37670. Remarks: In the genus Nucella,  Abbott (1974: p. 182). [Muricidae].
Thais (Nucella) cf.   () lima (Gmelin, 1791). [Murex]. Addicott. 10. 11.
 Co. USGS M1715. Hypotype USNM 650986. Remarks: In
the genus Nucella,  Abbott (1974: p. 182). [Muricidae].
Thais packi     
quad. (1954), Kern Co. USGS M1591. Hypotype USNM 649167. Remarks: Nucella tokudai (Yokoyama, 1932),
 Amano et al. (1993: p. 240). [Muricidae].
Thais (Nucella) packi
Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Hypotypes USNM 649167, 650131, 650132. Remarks:
Nucella tokudai (Yokoyama, 1932),  Amano et al. (1993: p. 240). [Muricidae].

Thais (Nucella) packi
[San Lorenzo Series], Oligocene. Northeast of Muir Station, Contra Costa Co. UCMP 1309. Holotype UCMP 11246.
Remarks: In the genus Nucella,  Stewart (1946: p. 102). [Muricidae].
Thais (Nucella) trancosanaThais
(Nucella) transcosana-
types USNM 650942–650946, 650976. Remarks: Determined here as in the genus Nucella. [Muricidae].
Thais
Ridge, Contra Costa Co. UCMP A4931. Hypotype UCMP 34165. Remarks: Thais (Nucella) packi Clark, 1918, 
Addicott 1970:84; Nucella tokudai (Yokoyama, 1932),  Amano et al. (1993: p. 240). [Muricidae].
Remarks: Thais (Nucella)
packi Clark 1918,  Addicott (1970: p. 84); Nucella tokudai (Yokoyama, 1932),  Amano et al. (1993: p. 240).
Thais cf. 
La Costa Valley quad. (1943), Alameda Co. SU 3248. Hypotype SU 8476 [=CASG 69982]. Remarks: Thais (Nucella)
packi Clark 1918,  Addicott (1970: p. 84); determined here as in the genus Nucella. [Muricidae].
Thesbia ocoyana (Anderson and Martin, 1914). [Drillia
Miocene. Oil Center quad. (1954), Kern Co. USGS M1602. Hypotype USNM 649170. Remarks: In the genus Bat-
tillaria,  Addicott (1970: p. 53). [Battilariidae].
Tornatella pinguis (Gabb, 1864). [Cinulia
Fm., Paleocene. Simi Hills, Ventura Co. UCR 6884. Hypotype UCR 6684/10 UCR determined here as missing.
Remarks: In the genus Ringicula,  Squires (1999b: p. 22). [Ringiculidae].
Trichotropis (?) durhami Weaver and Kleinpell, 1963 (n. sp.). UCPG 43:188–189
Eocene. Near Las Cruces, Los Olivos quad., Santa Barbara Co. UCMP B6999. Paratype SU 9206 [=CASG 70241].
[Capulidae].

9204 [=CASG 70201].
302 PALEOBIOS, VOLUME 38, FEBRUARY 2021

Paratype SU 9205 [=CASG 70240].
Trichotropis lajollaensis
Creek area, Reyes Peak quad. (1943), Ventura Co. UCR 4747. Hypotype UCR 4747/501. Remarks: In the genus
Pyrgulifera,  Squires (1999c: p. 13). [Capulidae].
Trichotropis tricarinata
Bravo Ranch quad. (1954), Kern Co. USGS M1590. Holotype USNM 650091. [Capulidae].
Trichotropis
(1954), Fresno Co. USGS M1590. Hypotype USNM 650084. [Capulidae].
Trichotropis 
Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8715 [=SBMNH 60192]. Remarks: Determined
here as turrid sp. [Turridae].
Tritonalia cf.   (Middendorff, 1848b). [Tritonium]. Woodring et al. 1940 [1941]. USGS PP 195:88, pl. 15,
Trachycardium Zone), Pliocene. La Porteria, Middle Dome, Kettleman Hills, Kings Co.
USGS 14126. Hypotype USNM 495765. Remarks: In the genus Paciocinebrina,  Houart et al. (2019: p. 209).
[Muricidae].
Tritonalia lurida (Middendorff, 1848b). [Tritonium
Fm., Pliocene. Reef Ridge quad., Kings Co. UCMP D1201. Hypotype UCMP 36857. Remarks: In the genus Pacio-
cinebrina,  Houart et al. (2019: p. 209). [Muricidae].
Tritonalia topangensis (Arnold, 1907). [Ocinebra
drawing, source specimen unknown. Remarks: Determined here as in the genus Ocenebra. [Muricidae].
Tritonidea kreyenhageni
Tritonium
californicum Gabb, 1869b)). Domengine Fm. (as Tejon Fm.), Eocene. North of Coalinga, Fresno Co. Hypotype
USNM 165644]. [Buccinidae].
Triumphis
Hill], Sand, Poso Creek.” Matrix and other specimens from this locality suggest the material is from the lower
part of the Olcese Sand exposed on the south side of Poso Creek, Woody quad., Kern Co. UCLA MB1400 [=LACMIP
16278]. Hypotype UCLA 45768 [=LACMIP. 8437]. [Buccinidae].
Trivia cf.  (Gray in G.B. Sowerby I, 1832) [not Gray, 1832, as cited]. . [Cypraea]. Woodring and Bramlette
1950 [Cebada Member), Pliocene. Graciosa Ridge, near
Orcutt, Santa Barbara Co. USGS 14648. Hypotype USNM 560073. Remarks: Determined here to be Pusula cali-
forniana (Gray, 1827). [Triviidae].
Trochita cf.  (Clark, 1915) [Calyptraea


North of Kern River, Kern Co. CASG 65. Hypotype CASG. 5731. Remarks: Determined here as in the genus Ca-
lyptraea. [Calyptraeidae].
Trochita radians (Lamarck, 1816), small var. [Trochus]. Woodring and Bramlette 1950 [1951]. USGS PP 222:72, pl.

560070. Remarks: Junior synonym of Trochita trochiformis (Born, 1778),  Pastorino and Urteaga, 2012: pp.
68–69. [Calyptraeidae].
Trochita cf. 
sandstone member), Oligocene. The Indians, Junipero Serra quad., Santa Lucia Range, Monterey Co. SU 1756.
Hypotype CASG 60366 determined here as missing. Remarks: Junior synonym of Trochita trochiformis (Born,
1778),  Pastorino and Urteaga, 2012: pp. 68– 69. [Calyptraeidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 303
Trochita cf.  (Born, 1778). [Patella-
cene. Humphreys quad. (1932, PR 1946), Los Angeles Co. CIT 1663 [=LACMIP 2555]. Hypotype LACMIP 4168.
[Calyptraeidae].
Trochita

Trophon cf. bartoni

Trophon carisaensis var. mirandaensis Grant and Eaton in
Neroly Fm., Miocene. Caliente Range, San Luis Obispo Co. UCLA 1777A [=LACMIP. 21777]. Holotype UCLA 8939
[=LACMIP 8966]. [Muricidae].
Trophon clarki var. cuyamanus Grant and Eaton in
Fm., Miocene. Caliente Range, San Luis Obispo Co. UCLA 1725 [=LACMIP 21725]. Holotype UCLA 8938 [=LACMIP
8965]. [Muricidae].
Trophon dickersoni-
gine Ranch quad., Fresno Co. UCMP D1090. Hypotype UCMP 36841. [Muricidae].
Trophon dickersoni nomlandi 
Fm., Miocene.,Domengine Ranch quad., Fresno Co. UCMP D702. Holotype UCMP 36842. [Muricidae].
Trophon gillulyi Grant and Eaton in Eaton et al., 1941 (n. Miocene.
Caliente Range, San Luis Obispo Co. UCLA 1849 [=LACMIP 21849]. Holotype UCLA 8937 [=LACMIP 8964]. [Mu-
ricidae].
Trophon kernensis
Miocene. San Luis Obispo Co. CASG 126. Hypotype CASG. 5718. Remarks: In the subgenus Austrotrophon, 
Addicott (1970: p. 81). [Muricidae].
Trophon kernensis-
ente quad. (1914), Kern Co. UCMP B1600. Hypotype UCMP 36561. Remarks: In the subgenus Austrotrophon,
 Addicott (1970: p. 81). [Muricidae].
Trophon (Austrotrophon) kernensis
Olcese Sand, Miocene. Rio Bravo Ranch quad. (1954 Kern Co. USGS M1599. Hypotype USNM 650124. [Muricidae].



B1619. Hypotype UCMP 30823.

Trophon kernensis 
(Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4312. Hypotype SDSNH 93835. Remarks: In the subgenus Austrotrophon,  Addicott (1970: p.
81). [Muricidae].
Trophon (Austrotrophon) kernensis medialis
Jewett Sand, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1591. Holotype USGS 650127, hypotype
USNM 650128. [Muricidae].
Trophon

304 PALEOBIOS, VOLUME 38, FEBRUARY 2021


Trophosycon kernianum (Cooper, 1894). [Agasoma
21. Temblor Fm., Miocene. Kern River, Kern Co. CASG 64. Hypotype CASG 5717. [Ficidae].
Trophosycon kernianum (Cooper, 1894). [Agasoma     
Monterey Fm. (Saltos Shale Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS
M2862. Hypotype USNM 254356. [Ficidae].

Hypotype USNM 254380.
Trophosycon kernianum (Cooper, 1894). [Agasoma   
), Miocene. Indian Creek
area, La Panza Range, San Luis Obispo Co. USGS M2862. Hypotype USNM 254356. [Ficidae].

La Panza Range, San Luis Obispo Co. USGS M3568. Hypotype USNM 254380.
Trophosycon ocoyanum (Conrad, 1855b). [Sycotypus
-
potype SU 648, [=CASG 71566]. Remarks: Ficus (Trophosycon) kerniana (Cooper, 1894),  Addicott (1970: p.
74). [Ficidae].
-
terey Co. Hypotype SU 645 [=CASG 71568]. Remarks: Ficus (Trophosycon) kerniana (Cooper, 1894),  Addicott
(1970: p. 74). [Ficidae].
Trophosycon-

as SDSNH 4520, p. 50). Hypotype SDSNH 98691. [Ficidae].
   
Hypotype SDSNH 83239.
Trophosycon sp. Powell et al. 2019b. PaleoBios-
tion, Berkeley Hills, Oakland, Alameda Co. UCMP IP13004. Hypotype UCMP 411945. [Ficidae].
Trypanotoma stocki (Dickerson, 1916). [Turris
Eocene. Simi Valley, Ventura Co. CSUN 371 [=LACMIP 16115]. Hypotype LACMIP 6556. [Turridae].
Trypanotoma stocki (Dickerson, 1916). [Turris
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 975 [=LACMIP 16201]. Hypotype LACMIP 7719.
[Turridae].
Tudicla (Pseudoperissolax) blakei (Conrad, 1855b). [Busycon-
Pseudoperissolax blakei)). Tejon Fm., Eocene. Grapevine
Creek, Kern Co. [CASG 711]. [Hypotype CASG 847]. Remarks: In the genus Pseudoperissolax and family Muricidae,
 Squires (2015b: p. 583). [Muricidae]
Turbo (Senectus) mounti-
trania Fm.], Pliocene. Whitewater area, San Gorgonio Pass, Riverside Co. UCR 5032. Holotype UCR 5032/28.
[Turbinidae].

....., Super Creek, Riverside Co. UCR 7477. Hypotypes UCR 7477/7–7477/9.

GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 305



Turbonilla (Pyrgiscus) bravoensis
Round Mtn. Silt, MioceneCaliente quad., Kern Co. SU 2121. Holotype SU 7546 and paratypes SU. 7546a, b [=CASG

....., SDSNH 5422 (ex
Turbonilla (Pyrgiscus) bravoensis
Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS. M1612. Hypotype USNM 650320. [Pyramidellidae].

Turbonilla (Ptycheulimella) edisonensis
Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Holotype USNM 650299. [Pyramidellidae].
Turbonilla (Tragula) greenhornensis
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650326, paratype USNM 650327.
[Pyramidellidae].
Turbonilla () hannai
Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650324, hypotype USNM 650323.
[Pyramidellidae].

Turbonilla (Pyrgiscus) hormigacuesta
Sand, MioceneOil Center quad. (1954), Kern Co. USGS M1597. Holotype USNM 650322, paratype USNM 650321.
[Pyramidellidae].
Turbonilla (Pyrgolampros) mariposa
Mtn. Silt, Miocene. Caliente quad., Kern Co. SU 2121. Holotype SU 7547 and paratype SU 7547a [=CASG 72015],

....., SDSNH 5422 (ex
Turbonilla (Pyrgolampros) mariposa Keen, 1943. Addicott 1970. USGS PP
Sand, Miocene. Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650310. [Pyramidellidae].

Hypotypes USNM 650332–650334.
Turbonilla cf. 
Oil Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650312. [Pyramidellidae].
Turbonilla (Chemnitzia
Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650325. [Pyramidellidae].
Turbonilla (Chemnitzia   
Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650311. [Pyramidellidae].
Turbonilla (Pyrgiscus
Center quad. (1954), Kern Co. USGS M1597. Hypotype USNM 650315. [Pyramidellidae].
Turbonilla 
17918. Hypotype LACMIP 14414. [Pyramidellidae].
306 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turcica imperialis brevis Stewart in Woodring et al., 1940 [1941]. Woodring and Bramlette 1950 [1951]. USGS PP
-
ity Hypotype USNM 560063. [Eucyclidae].

 Arnold, 1908 (n. sp.). Pr.USNM 34:373. San Lorenzo, Oligocene. San Lorenzo River, Santa

Turricula sp., aff.  (Nelson, 1925). [Turris
Fm., Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 961 [=LACMIP 17191].
Hypotype LACMIP 12688. [Clavatulidae].
Turricula buwaldana (Anderson and Martin, 1914). [Drillia

650258. [Clavatulidae].



Turricula (Anderson and Martin, 1914). [Drillia]. Addicott 1970. USGS PP 642:130, 131, pl. 18,
Superior Oil Co., 4949–5957 ft. depth,
Kern Co. CASG 28909. Hypotype CASG 12941. [Clavatulidae].
Turricula cohni (Dickerson, 1915). [Surcula  
junction of Alamo Creek, Dry Canyon, and Cuyama River valley, Reyes Peak quad. (1943), Ventura Co. UCR 4741.
Hypotype UCR 4741/251. [Clavatulidae].
Turricula (Crenaturricula) crenatospira (Cooper, 1894). [Surcula  
Fusiturricula (Crenaturricula) crenatospira)). Arroyo Hondo Fm.,
Eocene. Marysville Buttes, Sutter Co. UCMP 1853. Hypotype (topotype) UCMP 12166. [Clavatulidae].
Turricula ochsneri (Anderson and Martin, 1914). [Drillia 
Lower Round Mtn. Silt, Miocene. Rio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Hypotypes USNM
650242–650244. [Clavatulidae].


Turricula piercei (Arnold, 1909 [1910]). [Pleurotoma (Bathytoma
Upper Olcese Sand, Miocene. Caliente quad. (1914), Kern Co. UCMP B1586. Hypotype UCMP 37091. [Clavatulidae].



Turricula piercei (Arnold, 1909 [1910]). [Pleurotoma (Bathytoma)]. Addicott 1972b. SEPM Symposium vol., p. 12,

Temblor Range, Kern Co. USGS M3986. Hypotype USNM 646797. [Clavatulidae].
Turricula piercei (Arnold, 1909 [1910]). [Pleurotoma (Bathytoma)]. Addicott et al. 1978. USGS OFR 78-446:58, 60,

USGS M2862. Hypotype USNM 254353. [Clavatulidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 307

Turricula piercei (Arnold, 1909 [1910]). [Pleurotoma (Bathytoma)]. Addicott et al. 1980. SEPM Guidebook 6: 18,

Creek area, La Panza Range, San Luis Obispo Co. USGS M2862 (not USGS M2861 as in pl. caption). Hypotype
USNM. 254353. [Clavatulidae].
Turricula piercei babykingensis Adegoke, 1969. (n. subsp.). UCPG 80:193, pl. 13-
cene. Vicinity of Baby King
paratype UCMP 36905. [Clavatulidae].
Turricula praeattenuata (Gabb, 1869b). [Surcula
Lockwood Valley quad. (1943), Ventura Co. UCR 4676. Hypotype UCR 4676/4.

Turricula waringi        
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717). Hypotype
UCMP 14200. [Clavatulidae].
Turricula? waringi-
leocene. Simi Hills, Ventura Co. UCR 4569. Hypotype UCR 4569/27 determined here as missing. [Clavatulidae].
Turricula wilsoni (Anderson and Martin 1914). [Drillia
Miocene. Anadara bed, Reef Ridge quad., Kings Co. UCMP D1064. Hypotype UCMP 36902. [Clavatulidae].
Turricula wilsoni (Anderson and Martin, 1914). [Drillia]. Addicott et al. 1978. USGS OFR 78-446:58 (as Turricula
piercei
Luis Obispo Co. USGS M2869. Hypotype USNM 254361. [Clavatulidae].
Turricula wilsoni (Anderson and Martin, 1914). [Drillia]. Addicott et al. 1980. SEPM Guidebook 6:18 (as Turricula
piercei
(Saltos Shale Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS. M2869. Hypotype
USNM 254361. [Clavatulidae].
Turricula
and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073. Hypotype SU 10227 [=CASG 61666.23]. [Clavatulidae].

UCMP 14198. [Clavatulidae].
Turricula 
(1995), Los Angeles Co. LACMIP 17850. Hypotype LACMIP 13430. [Clavatulidae].


lost, 
Turritella andersoni Dickerson, 1916. G D. Hanna and Hertlein 
Domengine Fm., Eocene. Salt Creek, Fresno Co. CASG 316. Hypotypes CASG 5621, 5622, 7737. [Turritellidae].
Turritella andersoni
Creek, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4659. Hypotypes UCR 4659/201–4659/203. [Turritellidae].

Turritella andersoni
area, Santa Susana quad. (1951, PR 1960), Ventura Co. CSUN 469d [=LACMIP 16389]. Hypotype LACMIP 6510.
[Turritellidae].
308 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turritella andersoni  
Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 362 [=LACMIP 16273]. Hypotype LACMIP 7448.
[Turritellidae].

Turritella andersoni       
Northern Lockwood Valley area, Ventura Co. CSUN 979 [= LACMIP 16081]. Hypotype LACMIP 7695. [Turritellidae].
Turritella andersoni
junction of Urruttia Canyon and Salt Creek, Coalinga quad. (1902) [=Joaquin Rocks quad. (1969)], Fresno Co.
UCMP 1817. Neotype (of Squires 1988b: p. 205) UCMP 38193. [Turritellidae].
Turritella andersoni-
ley, Sawmill Mtn. quad. (1943) Ventura Co. CSUN 979 [=LACMIP 16081]. Hypotype LACMIP 7695. [Turritellidae].
Turritella andersoni
Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 676 [=LACMIP 16192]. Hypotype LACMIP 11455.
[Turritellidae].
Turritella andersoni
San Fernando quad. (1995), Los Angeles Co. LACMIP 17848. Hypotype LACMIP 13417. [Turritellidae].
Turritella andersoni
Seep Canyon” [=Chivo Canyon], Santa Susana quad., Ventura Co. LACMIP 16388. Hypotypes UCLA 59364–59367
[=LACMIP 10429–10432]. Remarks: Turritella andersoni susanae Merriam, 1941,  Squires (1999b: p. 16).
[Turritellidae].
Turritella andersoni Dickerson n. var. Crowell and Susuki 1959. GSA   -
cene. Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA. 3780 [=LACMIP 23780]. Hypotype UCLA. 28983
[=LACMIP 8838]. [Turritellidae].
Turritella andersoni lawsoni   
Fm., Eocene. Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3780 [=LACMIP 23780]. Hypotype UCLA
28984 [=LACMIP 8844]. [Turritellidae].
Turritella andersoni lawsoni
Rd. Canyon, San Guillermo quad. (1943), Ventura Co. UCR 4701. Hypotype UCR 4701/11. [Turritellidae].


4694. Hypotype UCR 4694/10.
Turritella andersoni lawsoni       
Eocene. Simi Arroyo, Santa Susana quad., Ventura Co. UCLA 5837 [=LACMIP 25837]. Hypotypes UCLA 59379,
59397 [=LACMIP 10440, 10441]. [Turritellidae].
Turritella andersoni lawsoni
Las Llajas Canyon, Santa Susana Mtns., Ventura Co. [LACMIP 7242]. Hypotype LACMIP 6511. [Turritellidae].
Turritella andersoni lawsoni
Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotypes LACMIP
6511, 6512. [Turritellidae].
Turritella andersoni lawsoni 
 
LACMIP 6511. [Turritellidae].
Turritella andersoni lawsoni Dickerson, 1916. Elder et al. 1998. SEPM Guidebook 84:160, 162 (as Turritella sp. cf.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 309

W200 [=UCMP W200]. Hypotype UCMP 39917. [Turritellidae].
Turritella andersoni lawsoni forma secondaria
9, 11 [in part]. Maniobra Fm., Eocene. Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3780 [=LACMIP
23780]. Hypotypes UCLA 28981, 28982 [=LACMIP 8841, 8842]. Remarks: Figure 11 is Turritella andersoni
lawsoni Dickerson, 1916,  Squires (1984: p. 18). [Turritellidae].
Turritella andersoni lawsoni forma secondaria  
Llajas Fm., Eocene. Simi Arroyo, Santa Susana quad., Ventura Co. UCLA 5837 [=LACMIP 25837]. Hypotype UCLA
59380 [=LACMIP 10442]. Remarks: Turritella andersoni lawsoni Dickerson, 1916, Squires (1984: p. 18).
[Turritellidae].
Turritella andersoni susanae Merriam, 1941. Saul 1983b. SEPM
Las Llajas Canyon, Santa Susana quad., Ventura Co. UCMP A993. Hypotype UCMP 15295. Remarks: Turritella
andersoni Dickerson, 1916,  Squires (1987: p. 24). [Turritellidae].
Turritella andersoni susanae
Chivo Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 969 [=LACMIP 16894]. Hypotype LACMIP
12677. [Turritellidae].
Turritella applinae
drainage, Cañada de Santa Ana, Lompoc quad., Santa Barbara Co. SU 2105. Hypotype SU 7413 [=CASG 72021].
[Turritellidae].
Turritella buwaldana
Reef Ridge, Reef Ridge quad., Kings Co. USGS 14495. Hypotype USNM 498725. [Turritellidae].
Turritella buwaldana-
cene. Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3779 [=LACMIP 23779]. Hypotype UCLA 28985
[=LACMIP 8840]. [Turritellidae].
Turritella buwaldana    

Turritella buwaldana
area, San Guillermo quad. (1943), Ventura Co. UCR 4705. Hypotype UCR 4705/201. [Turritellidae].
Turritella buwaldana   

Turritella buwaldana
Eocene. Devil Canyon, Santa Susana quad., Los Angeles Co. UCLA 2777 [=LACMIP 22777]. Hypotype UCLA 59374
[=LACMIP 10443]. [Turritellidae].

UCLA 586 [=LACMIP 20586]. Hypotypes UCLA 59375, 59376 [=LACMIP 10444, 10445].
Turritella buwaldana
Canyon, Santa Susana Mtns., Ventura Co. LACMIP 16211. Hypotype LACMIP 6513. [Turritellidae].
Turritella buwaldana
450 ft. Santa Susana quad., Los Angeles Co. CSUN 373 [=LACMIP 16211]. Hypotype LACMIP 6513. [Turritellidae].
Turritella buwaldana. 24. Juncal Fm., Eocene. Sharps Canyon,
Whitaker Peak quad. (1958), Los Angeles Co. CSUN 237 [=LACMIP 16080]. Hypotype LACMIP 7452. [Turritellidae].
Turritella buwaldana 
Northern Lockwood Valley area, Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LACMIP 7696. [Turritellidae].
310 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turritella buwaldana-

unnumbered hypotype lost, C. Throckmorton (personal communication, 2002). [Turritellidae].
Turritella buwaldana-
wood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 981 [=LACMIP 16142]. Hypotype LACMIP 7696.
[Turritellidae].
Turritella buwaldana
Eocene. El Paso Mtns., Inyokern SE quad. (1972), Kern Co. USGS M9176. Hypotype USNM 424889. [Turritellidae].
Turritella buwaldana
Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 674 [=LACMIP 16082]. Hypotype LACMIP 11457.
[Turritellidae].
Turritella buwaldana
Santa Susana quad. (1959, PR 1961), Ventura Co. LACMIP 26615. Hypotype LACMIP 10434 [=UCLA hypotype
59372]. [Turritellidae].

17186]. LACMIP hypotype 12678.
Turritella buwldana
San Fernando quad. (1995), Los Angeles Co. LACMIP. 17848. Hypotype LACMIP 13418. [Turritellidae].
Turritella buwaldana subsp. coosensis?

D8151. UCMP unnumbered hypotype lost, C. Throckmorton (personal communication, 2002). [Turritellidae].
Turritella buwaldana crooki
Eocene. Hot Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4656. Hypotypes UCR 4656/201,
4656/202. [Turritellidae].
Turritella buwaldana crooki
Fm., Eocene. Chivo Canyon, Santa Susana quad., Ventura Co. UCLA 6615 [=LACMIP 26615]. Hypotypes UCLA
59372, 59373 [=LACMIP 10434, 10435]. Remarks: Junior synonym of Turritella buwaldana Dickerson, 1916,
 Squires (1999b: p. 16). [Turritellidae].
Turritella buwaldana subuvasana Nelson, 1925. Saul 1983b. SEPM
Fm., Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCLA 3172 [=LACMIP 23172]. Hypotype UCLA 59361
[=LACMIP 10427]. Remarks: Junior synonym of Turritella buwaldana Dickerson, 1916,  Squires (1999b: p.
16). [Turritellidae].
Turritella carisaensis*
Fm., Miocene. [East side Newsome Canyon, Cuyama Valley, San Luis Obispo Co.] UCLA 1832 [=LACMIP 21832].
Hypotypes UCLA 8961, 8962 [=LACMIP 8947, 8948]. Remarks: Error for carrisaensis. [Turritellidae].
Turritella carrisaensis
Margarita Fm., Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2834. Hypotype USNM
254381. [Turritellidae].
Turritella carrisaensisTur-
ritella carisaensis
area, La Panza Range, San Luis Obispo Co. USGS M2834. Hypotype USNM 254381. [Turritellidae].
Turritella carisaensis* var. padronesensis Grant and Eaton in Eaton et al., 1941 (n. var.). AAPG Bull. 25(2):221, pl.


UCLA 7532C [=LACMIP 8946]. Remarks: Error for carrisaensis. [Turritellidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 311
Turritella cooperi   
source specimen unknown. [Turritellidae].
Turritella cooperi

Turritella cooperi        
Rico Fm., Miocene/Pliocene. Pancho Rico Creek, San Ardo quad., Monterey Co. USGS M1929. Hypotype USNM
649107. [Turritellidae].
Turritella cooperi
junction of Reynier and Sand canyons, Los Angeles Co. CIT 1849 [=LACMIP 2557]. Hypotypes LACMIP 4176–4180].
[Turritellidae].
Turritella cooperi  

Angeles Co. LACMIP1219]. Hypotypes (3) current whereabouts unknown. [Turritellidae].
Turritella cooperi      
Fm. (of Dibblee 1968), Pliocene. Panorama Hills quad., Temblor Range, Kern Co. USGS M1059. Hypotype USNM
646819. [Turritellidae].
Turritella cooperi
Hypotype determined here as missing. [Turritellidae].
Turritella cooperi Carpenter, 1864b. Squires and White 1983. SEPM   
[=Pico Fm.], Pliocene. Northern Simi Valley, Ventura Co. CSUN 725 [=LACMIP 29227]. Hypotype UCLA 59411
[=LACMIP 8324]. [Turritellidae].
Turritella cooperi
SDSNH 4080. Hypotype SDSNH 75911]. [Turritellidae].
Turritella cooperi
        
[=LACMIP 29227). Hypotype UCLA 59411 [=LACMIP 8324)]. [Turritellidae].
Turritella cooperi
Los Angeles Co. LACMIP 17917. Hypotype LACMIP 14383. [Turritellidae].
Turritella cooperi forma nova
Rico Fm., Miocene/Pliocene. Salinas River, Hames Valley quad., Monterey Co. USGS. M1676. Hypotype USNM
649098. [Turritellidae].
Turritella cooperi Carpenter 1864b, strongly sculptured var. Woodring and Bramlette, 1950 [1951]. USGS PP 222:71,
        
USNM 560068. [Turritellidae].
-
ritellidae].
Turritella coreyi-

UCMP B6940. Hypotypes SU 9178, 9179 [=CASG 70270, 70271]. [Turritellidae].

Turritella cornuatus-
cene. Calabasas quad., Simi Hills, Ventura Co. UCR 6825. Holotype UCR 6825/100 (not UCR 6825/1 (as in text)
or UCR 6835/100 (as in pl. caption)), paratype UCR 6825/101. [Turritellidae].
312 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turritella aff. 
Near junction of Reynier and Sand canyons, Los Angeles Co. CIT 1849 [=LACMIP. 2557]. Hypotypes LACMIP
4181, 4182. [Turritellidae].
Turritella gonostoma hemphilli Applin in Merriam, 1941. Woodring and Bramlette 1950 [1951]. USGS PP 222:71,
   
560069. [Turritellidae].
Turritella gonostoma hemphilli Applin in Merriam, 1941. Durham and Addicott 1965. USGS PP 524-A:A12, pl. 
18, 19 (casts). Pancho Rico Fm., Miocene/Pliocene. Wunpost quad., Monterey Co. USGS M982. Hypotypes USNM
649115, 649116. [Turritellidae].
Turritella hannai
Simi Hills, Ventura Co. UCR 4573. Hypotype UCR 4573/13 (not UCR 4571/22 as in pl. caption). [Turritellidae].
Turritella imperialis
Fm.], Pliocene. Imperial Co. UCMP. 12107. Hypotypes UCMP 32817 (not UCMP 32187 as in pl. caption), 32819.
[Turritellidae].
Turritella imperialis
Member) [=Latrania Fm.], Miocene. Base of Coyote Mtn. Carrizo Mtn. quad., Imperial Co. UCMP 738. Hypotype
USNM 418205. [Turritellidae].
Turritella imperialis          
Miocene/Pliocene. Base of Coyote Mtn., Carrizo Mtn. quad., Imperial Co. UCMP 738. Hypotype USNM 418205.
[Turritellidae].
Turritella imperialis     
Deguynos Fm., Pliocene. Carrizo Badlands, Imperial Co. Hypotypes in outcrop were not collected. [Turritellidae].

Turritella inezana
Mtn., San Luis Obispo Co. UCLA 485 (not UCLA 574 as in pl. caption) [= LACMIP 20485]. Hypotype UCLA. 8844
(not UCLA 8840 as in pl. caption) [=LACMIP 8937]. [Turritellidae].

Hypotype UCLA 8840 (not UCLA 8844 as in pl. caption) [=LACMIP 8936].
Turritella inezana
Fm., Miocene. Cajon Pass, Hesperia quad., San Bernardino Co. CIT 568 [=LACMIP 7673]. Hypotypes CIT 3896–3898
[=LACMIP 4183–4185]. [Turritellidae].

Turritella inezana
Miocene. San Augustine Canyon, Santa Rosa Is., California Channel Ids. CASG 1153. Hypotype CASG 5543. [Tur-
ritellidae].
Turritella inezana

4514. Hypotype USNM 165321]. [Turritellidae].
Turritella inezana Conrad, 1857d. Avila and Weaver 1969. AAPG/SEPM Guidebook, p
Miocene. Santa Rosa Is., California Channel Ids. UCSB SR64-14 [=LACMIP 16436]. Hypotype UCSB M25 [=LACMIP
6313]. [Turritellidae].
Turritella inezana
Sand Member) (of Stinemeyer et al. 1959: pp. 9, 13), Miocene. Temblor Range, Kern Co. USGS 4941. Hypotype
USNM 165748. [Turritellidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 313

Turritella inezana var. altacorona-
ros Fm., Miocene. Ernest's Canyon, Caliente Mtns., San Luis Obispo Co. UCLA 535 [=LACMIP 20535]. Hypotypes
UCLA 8946, 8947 [=LACMIP 8938, 8939]. [Turritellidae].
Turritella inezana var. bicarinata
Fm., Miocene. Simi Valley, Ventura Co. LACMIP 16391. Hypotype UCLA 1704 [=LACMIP 8931]. [Turritellidae].
    
LACMIP 8932]. [Turritellidae].
Turritella inezana var. hoffmanni -
cene. Cuyama Valley, San Luis Obispo Co. UCLA 548 [=LACMIP 20548]. Hypotype UCLA 3926 [=LACMIP 8930].
[Turritellidae].
Turritella inezana forma hoffmanni
Fm., Miocene. South of Pt. Arena, Mendocino Co. USGS M2638. Hypotype USNM 649594. [Turritellidae].

Turritella (Torcula) inezana pervulgata    


Turritella inezana var. santana-
ros” Fm.. Miocene. Santa Ana Mtns., Orange Co. UCMP 6128 [=LACMIP 16392]. Paratype UCLA 2302 [=LACMIP
8935]. [Turritellidae].
Turritella inezana santana
Trabuco, Santa Ana Mtns., Orange Co. LACMIP 16392. Hypotype LACMIP 8935. [Turritellidae].
Turritella inezana var. sespeensis-
cene. Piru Creek, Ventura Co. LACMIP 16394. Hypotype UCLA 3925 [=LACMIP 8934]. [Turritellidae].
 
[=LACMIP 8933].
Turritella infragranulata Gabb, 1864. Smith 1975b. SEPM Symposium vol., p
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype
SU 10225 [=CASG 61666.14]. [Turritellidae].
Turritella infragranulata
Fm.], Paleocene. Quarry Canyon, Topanga quad., Santa Monica Mtns., Los Angeles Co. UCLA 6574 [=LACMIP
26574]. Hypotype UCLA 57963 [=LACMIP 10319]. [Turritellidae].

6572 [=LACMIP 26572]. Hypotypes UCLA 58930, 58931, 58940, 58941 [=LACMIP 10322, 10323, 10324,10325].

[=LACMIP 26571]. Hypotypes UCLA. 58928, 58929, 59033 [=LACMIP 10320, 10321, 10326].

Hypotype UCLA 39736 [=LACMIP 10318].

Turritella infragranulata
Susana Fm.], Paleocene. Garapito Creek, Santa Monica Mtns., Los Angeles Co. UCLA 6572 [=LACMIP 26572].
Hypotype UCLA 58931 [=LACMIP 10323]. [Turritellidae].
314 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turritella infragranulata Gabb, 1864. Specimen illustrated on the front cover of the present paper. Santa Susana Fm.,
Garapito Creek, Santa Monica Mtns., Los Angeles Co. LACMIP 40354. Hypotype LACMIP 14781. [Turritellidae].
Turritella infragranulata pachecoensis
Canyon Fm. [=Santa Susana Fm.], Paleocene. Quarry Canyon, Topanga quad., Santa Monica Mtns., Los Angeles Co.
UCLA 6574 [=LACMIP 26574]. Hypotype UCLA 58921 [=LACMIP 10317]. [Turritellidae].

Simi Hills, Ventura Co. UCLA 3121 [=LACMIP 23121]. Hypotype UCLA 58915 [=LACMIP 10313].

Canoga Park quad., Los Angeles Co. UCLA 6583 [=LACMIP 26583]. Hypotype UCLA 58920 [=LACMIP 10316].

quad. (1929), Simi Hills, Ventura Co. UCLA 3163 [=LACMIP 23163]. Hypotype UCLA 59039 [=LACMIP 10314].

Channel Ids. UCLA 3348 [=LACMIP 23348]. Hypotype UCLA 59038 [=LACMIP 10315].
Turritella infragranulata pachecoensis Stanton, 1896. Saul 1983b. SEPM Guidebook 35:74
Fm. [=Santa Susana Fm.], Paleocene. Vicinity of Encino Reservoir, Santa Monica Mtns., Los Angeles Co. UCLA 6583
[=LACMIP 26583]. Hypotype UCLA 58920 [=LACMIP 10316]. [Turritellidae].
Turritella infragranulata pachecoensis Stanton, 1896. Elder et al. 1998. SEPM Guidebook 84:160, 166 (as Turritella
pachecoensis
W730 [=UCMP W730]. Hypotype UCMP 39914. [Turritellidae].
Turritella infragranulata pachecoensis Stanon, 1896. Koch et al. 2004. NPS Tech. Rept. 04/01:8 (as ),
  
Monica Mtns., Los Angeles Co. LACMIP 26853. Hypotype LACMIP 10316. [Turritellidae].
Turritella infragranulata pachecoensis

[Turritellidae].
Turritella kernensis
Oil Center quad. (1954), Kern Co. USGS M1597. Hypotypes USNM 650058, 650060, 650061. [Turritellidae].


Turritella lawsonisic] Llajas
Fm., Eocene. West of Aliso Canyon, Los Angeles Co. CASG 549. Hypotype CASG 5916. Remarks: Turritella ander-
soni lawsoni Dickerson, 1916, Squires (1984: p. 18). [Turritellidae].
Turritella lawsoni
Reef Ridge, Kettleman Plain quad., Kings Co. USGS 14482. Hypotypes USNM 498728, 498730. Remarks: Tur-
ritella andersoni lawsoni Dickerson, 1916, Squires (1984: p. 18). [Turritellidae].

Turritella lawsoni
    Remarks: Turritella andersoni lawsoni Dickerson, 1916, Squires
(1984: p. 18). [Turritellidae].
Turritella lorenzana
Emigdio Fm.), Oligocene. Pleito Creek, Mt. Pinos quad., Kern Co. UCMP 3217. Holotype UCMP 11424. Remarks:
Turritella variata lorenzana,  Merriam (1941: p. 99). [Turritellidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 315
Turritella meganosensis Clark and Woodford, 1927. Saul 1983b
Fm., Paleocene [Eocene]. Las Llajas Canyon, Santa Susana quad., Ventura Co. UCMP 7000. Hypotype UCMP. 37430.
[Turritellidae].
Turritella meganosensis
Eocene. Las Llajas Canyon, Santa Susana quad., Ventura Co. UCMP 7000. Hypotype UCMP 37430. [Turritellidae].
Turritella meganosensis protumescens
Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana quad., Ventura Co. UCLA 6616 [=LACMIP 26616]. Hypotype
UCLA 59371 [=LACMIP 10433]. [Turritellidae].
Turritella meganosensis protumescens
Llajas Fm., Eocene. Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP 16395]. Hypotype LACMIP 6509.
[Turritellidae].
Turritella meganosensis protumescens
Fm., Eocene. Las Llajas Canyon, Santa Susana quad. (1951, PR 1969) Ventura Co. CSUN 454 [=LACMIP 16395].
Hypotype LACMIP 6509. [Turritellidae].
Turritella meganosensis protumescens-
cal Fm., Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 846 [=LACMIP 16396]. Hypotype LACMIP
7450. [Turritellidae].
Turritella meganosensis protumescens Merriam and Turner, 1937. Squires 1991c. SEPM Guidebook 68:220, pl. 1,
  
16076]. Hypotype LACMIP 11456. [Turritellidae].
Turritella merriami
Eocene. Buttes Well 4 at a depth of about 2800 ft., north of Sutter City, Sutter Co. CASG 29138. Hypotype CASG.
5709. [Turritellidae].
Turritella merriami      
Canyon, , Whitaker Peak quad. (1958), Los Angeles Co. CSUN 358 [=LACMIP 16114]. Hypotype LACMIP 7451.
[Turritellidae].
Turritella merriami
      
D8157. UCMP unnumbered hypotypes lost, C. Throckmorton (personal communication, 2002). [Turritellidae].
Turritella moodyi
Caliente quad. (1914), Kern Co. UCMP B1637. Hypotype UCMP 12150. [Turritellidae].


Turritella ocoyana Conrad, 1855b. Eaton et Fm., Miocene. Cuyama
Valley, San Luis Obispo Co. UCLA 540 [=LACMIP 20540]. Hypotype UCLA 8950 (not UCLA. 8850 as in pl. caption)
[=LACMIP 8940]. [Turritellidae].

UCLA 8951 (not UCLA 8851 as in pl. caption) [=LACMIP 11303].

8852 as in pl. caption) [=LACMIP 11304].
Turritella ocoyana  
Miocene. Vicinity of Cottonwood, Kern Co. CASG 66. Hypotype CASG 5963. [Turritellidae].
Turritella ocoyana
316 PALEOBIOS, VOLUME 38, FEBRUARY 2021
-
potype USNM 165593]. [Turritellidae].
Turritella ocoyana
Reef Ridge quad., Kings Co. USGS 14402. Hypotype USNM 498748. [Turritellidae].
Turritella ocoyana
(Altamira Shale Member), Miocene. George F. Canyon-Miraleste Canyon area, George F. Canyon, Palos Verdes
Hills, Los Angeles Co. USGS 13127. Hypotypes USNM 497069, 497070. [Turritellidae].
Turritella ocoyana

Turritella ocoyana-
wood Creek area, Caliente quad. (1914), Kern Co. UCMP B1588. Hypotype UCMP 36563. [Turritellidae].
Turritella ocoyana
Oil Center quad. (1950), Kern Co. UCMP. B1598. Hypotype UCMP 12550. [Turritellidae].



M1689 as in pl. caption). Hypotype (topotype) USNM 650064.
Turritella ocoyanaTemblor Fm. (But-
tonbed Sand Member) (of Stinemeyer et al. 1959:8, 13), Miocene Shale Pt. quad., Temblor Range, Kern Co. USGS
M3986. Hypotype USNM 646808. [Turritellidae].
Turritella ocoyana
possibly at SDSNH, current whereabouts unknown. Remarks: Ex Grouard collection. [Turritellidae].
Turritella ocoyana
Shale Member), Miocene. Indian Creek area, La Panza Range, San Luis Obispo Co. USGS M3814. Hypotype USNM
254366. [Turritellidae].
Turritella ocoyana        

Range, San Luis Obispo Co. USGS M3814. Hypotype USNM 254366. [Turritellidae].
Turritella ocoyana-

Northeast of Oil City, Fresno Co. Hypotype USNM 165593]. [Turritellidae].
Turritella ocoyana
Reef,” Saddleback Valley, Orange Co. SDSNH 4312. Hypotype SDSNH 83244. [Turritellidae].
Turritella ocoyana-
ber). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22635 [=UCLA 2635]. Hypotype LACMIP
13081 [=UCLA 57741]. [Turritellidae].
Turritella ocoyana
Fm., Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotypes UCSB 6516, 8771 [=SBMNH
60188, 60189]. Remarks: Determined here as Turritella sp. [Turritellidae].
Turritella ocoyanaEntobiaCau-
lostrepis trace fossil). Round Mtn. Silt (not Temblor Fm. as in pl. caption), Miocene. Cottonwood Creek, Kern Co.
LACMIP 7907. Hypotypes LACMIP 12425, 12427. [Turritellidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 317
Turritella ocoyana
(Saddleback Valley Limestone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano
quad. SDSNH 4312. Hypotype SDSNH 140188. [Turritellidae].
Turritella ocoyana 
Orange, northern Santa Ana Mtns., Orange Co. Hypotype possibly at SDSNH, current whereabouts unknown.
Remarks: Ex Grouard collection. [Turritellidae].
Turritella ocoyana var. bösei Hertlein and Jordan, 1927. Eaton et al. 1941. 
Fm., Miocene. Ernest's Canyon, Cuyama Valley, San Luis Obispo Co. UCLA 537 [=LACMIP. 20537]. Hypotype UCLA.
8955 (not UCLA 8855 as in pl. caption) [=LACMIP 8941]. Remarks: Junior synonym of Turritella ocoyana Conrad,
1855b,  Addicott (1970: p. 50). [Turritellidae].

Co. UCLA 535 [=LACMIP 20535]. Hypotype UCLA 8956 (not UCLA 8856 as in pl. caption) [=LACMIP 8942].

UCLA 8957 (not UCLA 8857 as in pl. caption) [=LACMIP 8943].
Turritella ocoyana bösei
22. Temblor Fm., Miocene. San Juan River area, San Luis Obispo Co. CASG 146. Hypotype CASG 5702. Remarks:
Turritella ocoyana forma topangensis Merriam, 1941,  Addicott (1970: p. 50). [Turritellidae].
Remarks: Junior synonym of Turritella
ocoyana Conrad, 1855b,  Addicott (1970: p. 50).
Turritella ocoyana (Conrad) bosei Hertlein and Jordan, 1927. Avila and Weaver 1969. AAPG/SEPM Guidebook, p. 59,

16433]. Hypotype UCSB M26 [=LACMIP 6314]. Remarks: Junior synonym of Turritella ocoyana Conrad, 1855b
 Addicott 1970: p. 50. [Turritellidae].
Turritella ocoyana forma topangensis
Mtn. Silt, Miocene. Turritella bed west of Cottonwood Creek, Rio Bravo Ranch quad. (1954), Kern Co. USGS M1605.
Hypotype USNM 650066. [Turritellidae].


Turritella ocoyana topangensis
Co. Hypotype possibly at SDSNH, current whereabouts unknown. Remarks: Ex Grouard collection. [Turritellidae].
Turritella pachecoensis
Fm., Eocene [Paleocene]. Santa Cruz Is., California Channel Ids. CASG 28645. Hypotype CASG 5670. Remarks:
Turritella infragranulata pachecoensis,  Saul (1983a: p. 95). [Turritellidae].
Turritella (Haustator) pacheoensis*

4189. Holotype USNM 157843-22, paratype USNM 157843-21]. Remarks: Error for pachecoensis. [Turritellidae].
Turritella pachecoensis Stanton, 1896. Doerner 1969. AAPG/SEPM Guidebook      
Paleocene. Well Canyon, Santa Cruz Is., California Channel Ids. UCSB DWW-1 [=LACMIP 16437]. Hypotypes
UCSB M30, M31 [=LACMIP 6292, 6293]. Remarks: Turritella infragranulata pachecoensis,  Saul (1983a: p.
95). [Turritellidae].
Turritella pachecoensis
Angeles Co. UCLA2687 [=LACMIP 22687]. Hypotype LACMIP 10308] determined here as missing. [Turritellidae].
Turritella pachecoensis
Near junction of Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073. Hypotype SU 10244 [=CASG
318 PALEOBIOS, VOLUME 38, FEBRUARY 2021
1666.17]. Remarks: Turritella infragranulata pachecoensis,  Saul (1983a: p. 95). [Turritellidae].

UCMP 14197. Remarks: Turritella infragranulata pachecoensis,  Saul (1983a: p. 95).
Turritella pachecoensis


Ridge north of McLure Valley, Fresno Co. Hypotype USNM 165636. [Turritellidae].
Turritella pachecoensis
Paleocene. Simi Hills, Ventura Co. UCR 6833. Hypotype UCR 6833/1 determined here as missing. [Turritellidae].
]. Simi Hills, Ventura Co. UCR 6821. Hypotype UCR 6821/1 determined here as
missing.. Remarks: Error for pachecoensis.
Turritella pachecoensis Stanton, 1896. Boles and Weigand 1997. San Diego Assoc. Geologists Ann. Field Trip, p. 19,

Remarks: Turritella infragranulata pachecoensis,  Saul (1983a: p. 95). [Turritellidae].
Turritella pachecoensis Stanton, 1896. Boles and Weigand 2001. GSA and Pac. Sect-AAPG Cordilleran Sect. Guidebook,

Channel Ids. Hypotype current whereabouts unknown. Remarks: Turritella infragranulata pachecoensis, 
Saul (1983a: p. 95). [Turritellidae].
Turritella pachecoensis      
Pozo Fm., Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotypes SBMNH 60190, 60200].
Remarks: Turritella infragranulata pachecoensis,  Saul (1983a: p. 95). [Turritellidae].
Turritella pachewen's*

Remarks: Turritella infragranulata pachecoensis,  Saul (1983a: p. 95). Error for
pachecoensis. [Turritellidae].
Turritella (Torcula(  padronesensis Eaton et al., 1941. Addicott 1970. USGS PP 642: 49
Olcese, Miocene. Caliente quad. (1914), Kern Co. UCMP B1616. Hypotype UCMP 12549. Remarks: Determined
here as Turritella carriasaensis padronesensis. [Turritellidae].
Turritella peninsularis
  
3777. Holotype UCMP 15315 (not UCMP 15314 as in pl. caption), paratype UCMP 15314 (both of 
renodata Merriam, 1941). [Turritellidae].

Hypotype UCLA 58900 [=LACMIP 10311].

quad., San Gabriel Mtns., Los Angeles Co. UCLA. 2333 [=LACMIP 22333]. Hypotype UCLA. 58907 [=LACMIP 10312].
   
Canyon, Calabasas quad., Simi Hills, Ventura Co. UCLA 3133 [=LACMIP 23133]. Hypotype UCLA 58903 [=LACMIP
10309].
         
49512 [=LACMIP 10308].

Hypotype UCLA 59037 [=LACMIP 10310].
Turritella peninsularis
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 319
San Francisquito Fm., Paleocene. Pinyon Ridge, Valyermo quad., Los Angeles Co. UCLA 2333 [=LACMIP 22333].
Hypotype UCLA 58907 [=LACMIP 10312]. [Turritellidae].
-
MIP 22687]. Hypotype UCLA 49512 [=LACMIP 10308].
Turritella peninsularis Anderson and G.D. Hanna, 1935. Boles and Weigand in Gordon et al. in Dunne and Cooper

California Channel Ids. Hypotypes, which are mostly cross-sections of weathered uncollected specimens, are
determined here as Turritella infragranulata pachecoensis. [Turritellidae].
Turritella peninsularis adelaidana
strata, uppermost Cretaceous/lowermost Paleocene. Machesna Mtn., La Panza quad., San Luis Obispo Co. USGS
M3086. Hypotypes USNM 307685, 307686. [Turritellidae].
Turritella peninsularis adelaidana. 44. Unnamed strata, up-
permost Cretaceous/lowermost Paleocene. Dip Creek area, Lake Nacimiento, Lime Mtn. quad., San Luis Obispo
Co. UCLA 6526 [=LACMIP 26526]. Hypotype UCLA 58849 [=LACMIP 10281]. [Turritellidae].

Hypotype UCLA 58855 [=LACMIP. 10285].

Hypotype UCLA 58846 [=LACMIP 10279].
Turritella peninsularis quaylei-
cisquito Fm., Paleocene. Warm Springs Mtn., Warm Springs Mtn. quad., Los Angeles Co. UCLA 1595 [=LACMIP

11164–11167]. [Turritellidae].
-
        
10330–10336].
     
21579]. Paratypes UCLA 58881–58884 [=LACMIP 10300–10303].
-
types UCLA 58876 [=LACMIP 10296, 10297, 10337, 10338].

UCLA 58879 [=LACMIP 10299, 10339, 10340].
Turritella peninsularis quaylei intergrades to   Anderson and G D. Hanna, 1935. Saul 1983a. UCPG

quad., Simi Hills, Ventura Co. UCLA 3111 [=LACMIP. 23111]. Hypotypes UCLA 58887, 58888 [=LACMIP 10305,
10306]. [Turritellidae].

21579]. Hypotype UCLA 58878 [=LACMIP 10298].

Hypotype UCLA 58885 [=LACMIP 10304].
Turritella peninsularis quaylei
Fm., Paleocene. Simi Hills, Calabasas quad., Ventura Co. UCLA 3111 [=LACMIP 23111]. Hypotype UCLA 58888
[=LACMIP 10306]. [Turritellidae].
Turritella reversa
Paleocene. Poison Oak Canyon, Santa Susana quad., Ventura Co. UCLA 6547 [=LACMIP 26547]. Hypotype UCLA
320 PALEOBIOS, VOLUME 38, FEBRUARY 2021
58932 [=LACMIP 10307]. [Turritellidae].
Turritella reversa-
cene. Simi Hills, Ventura Co. UCR 4569. Hypotype UCR 4569/28. [Turritellidae].
Turritella reversa Waring, 1917. Saul 1983b. SEPM Guidebook 35:74,     
Paleocene. Poison Oak Canyon, Santa Susana quad., Ventura Co. UCLA 6547 [=LACMIP 26547]. Hypotype UCLA
58932 [=LACMIP 10307]. [Turritellidae].
Turritella sargeanti
Tejon Fm., Eocene. Live Oak Creek Kern Co. CASG 244. Hypotype CASG 5903. [Turritellidae].
Turritella schencki delaguerrae  

B6939. Holotype SU 9180 [=CASG 70259], paratype SU 9181 [=CASG 70260]. [Turritellidae].
Turritella scrippsensis
de Santa Anita, Santa Barbara Co. SU 2090. Hypotype SU 7415 [=CASG 72022]. [Turritellidae].
Turritella scrippsensis
Cañada de Santa Anita, Lompoc quad., Santa Barbara Co. UCMP. B6922. Hypotype SU/CASG 7415. [Turritellidae].
Turritella scrippsensis
Meadows, San Guillermo quad. (1943), Ventura Co. UCR 4690. Hypotype UCR 4690/10. [Turritellidae].
Turritella stocki
junction of Panoche and Silver Creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP. A9717]. Hypotype SU
10226 [=CASG 61666.20]. [Turritellidae].
Turritella susanaensis Nelson, 1925 nomen dubium
Paleocene/Eocene. South side of Simi Valley, Calabasas quad. (1952, PR 1967), Ventura Co. CSUN 1343 [=LACMIP
17186]. Hypotype LACMIP 12681. [Turritellidae].
Turritella tembloris*
Miocene. Dry Canyon, near Calabasas, Los Angeles Co. CASG 567. Paratype CASG 2984. Remarks: Error for tem-
blorensis. [Turritellidae].
Turritella temblorensis
Canyon, Los Angeles Co. UCMP B7853. Hypotype UCMP 36493. [Turritellidae].
Turritella temblorensis
Fm., Miocene. Santa Rosa Is., California Channel Ids. UCSB SR64-6 [=LACMIP 16438]. Hypotypes UCSB M27, M28
[=LACMIP 6315, 6316]. [Turritellidae].
Turritella temblorensis-
potype possibly at SDSNH, current whereabouts unknown. Remarks: Ex Grouard collection. [Turritellidae].
Turritella temblorensis Wiedey, 1928. Addicott et al. 1978. USGS OFR  
(Saltos Shale Member), Miocene. Shell Creek area, La Panza Range, San Luis Obispo Co. USGS M2869. Hypotype
USNM 254363. [Turritellidae].
Turritella temblorensis
      
Range, San Luis Obispo Co. USGS. M2869. Hypotype USNM 254363. [Turritellidae].
Turritella temblorensis
Member). Topanga Canyon, Santa Monica Mtns., Los Angeles Co. LACMIP 22638 [=UCLA 2638]. Hypotype LACMIP
13079 [=UCLA 57325)]. [Turritellidae].
Turritella temblorensis Wiedey var. tritschi Hertlein, 1928a. Avila and Weaver 1969. AAPG/SEPM Guidebook, p. 59,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 321
     
UCSB SR67-55 [=LACMIP 16427]. Hypotype UCSB M29 [=LACMIP 6317]. [Turritellidae].
Turritella uvasana

USNM 165653. [Turritellidae].
Turritella uvasana
Reef Ridge, Reef Ridge quad., Kings Co. USGS 14494. Hypotypes USNM 498726, 498727. [Turritellidae].
Turritella uvasana 
Eocene. Orocopia Mtns., Canyon Spring quad., Riverside Co. UCLA 3779 (not UCLA 2779 as in pl. caption) [=LAC-
MIP 23779]. Hypotype UCLA 28986 [=LACMIP 8843]. Remarks: Turritella uvasana infera Merriam, 1941, 
Givens (1974: p. 65). [Turritellidae].
Turritella uvasana

Turritella uvasana
Tequipis Canyon west of San Marcos Pass, Santa Barbara Co. Dibblee locality 29. Hypotype SU 9182 [=CASG
70263]. [Turritellidae].
Turritella uvasana
Lodge, 900 ft. Lion Canyon quad. (1943), Ventura Co. UCR 4722. Hypotype UCR 4722/210. [Turritellidae].
Turritella uvasana-

Turritella uvasana uvasana
Sandstone, Eocene. Upper Sespe Creek, Ventura Co. LACMIP 24260. Hypotype LACMIP 11626. [Turritellidae].
....., pl. 2, 
11647. [Turritellidae].
Turritella uvasana
Fm., Eocene. Santa Cruz Is., California Channel Ids. Line drawing, source specimen unknown. [Turritellidae].
Turritella uvasana
Fm., Paleocene. Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8773 [=SBMNH 60191].
Remarks: Determined here as Turritella infragranulata pachecoensis. [Turritellidae].

Mt. Diablo St. Park, Contra Costa Co. UCMP A4311. Hypotype UCMP 30279]. [Turritellidae].
Turritella uvasana applinae
Fm.” [=possibly Ardath Shale], Eocene. San Clemente Canyon, San Diego Co. SDSNH2395. Hypotype SDSNH 4149].
[Turritellidae].
Turritella uvasana applinae
of Piru Creek and Alamo Creek watersheds, San Guillermo quad. (1943), Ventura Co. UCR 4694. Hypotype UCR
4694/21. [Turritellidae].


Turritella uvasana applinae-
cene. Las Llajas Canyon, Santa Susana quad., Ventura Co. UCLA 2312 [=LACMIP 22312]. Hypotype UCLA 59377
[=LACMIP 10446]. [Turritellidae].
322 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turritella uvasana applinae
Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP7242. Hypotype LACMIP 6514. [Turritellidae].
Turritella uvasana applinae 
Llajas Canyon, Santa Susana quad. (1951, PR 1969) Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP
6514. [Turritellidae].
Turritella uvasana applinae
Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 219 [=LACMIP 16100]. Hypotype LACMIP 7454.
[Turritellidae].
Turritella uvasana hendoni
Eocene. Lockwood Valley area, Ventura Co. CSUN 987 [=LACMIP. 16175]. Hypotype LACMIP 7697. [Turritellidae].
Turritella uvasana hendoni       
Lockwood Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 987 [=LACMIP 16175]. Hypotype LACMIP 7697.
[Turritellidae].
Turritella uvasana hendoni

11458. [Turritellidae].
Turritella uvasana infera
Creek, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4662. Hypotype UCR.4662/201. [Turritellidae].


Turritella uvasana infera-
leocene. Las Llajas Canyon, Santa Susana quad., Ventura Co. UCMP 7000. Hypotype UCMP 37429. [Turritellidae].

Hypotype UCLA 59359 [=LACMIP 10436].
Turritella uvasana infera
Canyon, Santa Susana quad., Ventura Co. UCLA 6616 [=LACMIP 26616]. Hypotype UCLA 59359 [=LACMIP 10436].
[Turritellidae].
Turritella uvasana infera-
ker Peak quad. (1958), Los Angeles Co. CSUN 805 [=LACMIP 16397]. Hypotype LACMIP 7453. [Turritellidae].
Turritella uvasana infera  
North side Simi Valley, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 967 (=LACMIP 12648). Hypotype
LACMIP 12679. [Turritellidae].

Hypotype LACMIP 12680.
Turritella uvasana cf. 
Cozy Dell Fm., Eocene. North of Camino Cielo, San Rafael Mtn. quad., Santa Barbara Co. UCMP B6927. Hypotype
SU 9183 [=CASG 70264]. [Turritellidae].
Turritella uvasana neopleura        
West of Reyes Peak, Reyes Peak quad. (1943), Ventura Co. UCR 4706. Hypotypes UCR 4706/201, 4706/204.
[Turritellidae].


GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 323
Turritella uvasana sargeanti
Eocene. Near Pine Mtn. Lodge, Lion Canyon quad. (1943), Ventura Co. UCR 4726. Hypotypes UCR. 4726/101–
4726/103. [Turritellidae].
            
4732/301, 4732/302.
Turritella uvasana sargeanti
8. Mission Valley Fm., Eocene. Scripps Ranch, southeast of Miramar Reservoir, San Diego, San Diego Co. SDSNH
2962. Hypotype SDSNH 20777. [Turritellidae].
Turritella vanvlecki
Fm. (Siphonalia Zone), Pliocene. North Dome, ridge east of La Maseta, Kettleman Hills, Kings Co. USGS 12464.
Hypotype USNM 495809. [Turritellidae].
Turritella vanvlecki
Rico Fm., Miocene/Pliocene. North side of Walker Canyon, Wunpost quad., Monterey Co. USGS. M1935. Hypotype
USNM 649110. [Turritellidae].

Turritella vanvlecki  Entobia endobiont), 8d (with
spionid borings). Pancho Rico Fm., Miocene/Pliocene. Lynch Canyon, San Ardo quad., Monterey Co. LACMIP
22587. Hypotypes LACMIP 12426, 12428. [Turritellidae].
Turritella variata

CASG 5665 determined here as missing. [Turritellidae].
Turritella variata
Lompoc quad., Santa Barbara Co. UCMP A948. Hypotypes UCMP 33905, 33906. [Turritellidae].
Turritella variata
Wheatland Fm., Eocene. Dry Slough [=Dry Creek], near Wheatland, Smartsville quad., Yuba Co. UCMP A1889.
Hypotypes UCMP 11012, 11024, 11058. [Turritellidae].
Turritella variata  -
cene. Las Varas Canyon, Goleta quad., Santa Barbara Co. UCMP B6966. Hypotypes SU 9184, 9185 [=CASG 70261,
70262]. [Turritellidae].
sic] Mtns., Santa Barbara Co. Holotype USNM 13346.
Turritella variata juliana Merriam, 1941. Weaver and Kleinpel 1963. UCPG 43:185, pl
Near junction of Yridsis Creek with El Jaro Creek Lompoc quad., Santa Barbara Co. UCMP B7007A. Hypotype SU
9186 [=CASG 70274]. [Turritellidae].
Turritella variata Conrad cf. var. juliana
Nojoqui Creek, Lompoc quad., Santa Barbara Co. UCMP A948. Hypotype UCMP 33911. [Turritellidae].
Turritella variata lompocensis
Eocene. Near junction of Yridsis Creek with El Jaro Creek, Lompoc quad., Santa Barbara Co. UCMP B7007E. Hy-
potype SU 9187 [=CASG 70229]. [Turritellidae].
Turritella variata lorenzana
Alegria Fm., Oligocene. Vicinity of Arroyo el Bulito, Lompoc quad., Santa Barbara Co. UCMP B7032. Hypotype SU
9188 [=CASG 70266]. [Turritellidae].

Barbara Co. UCMP B7017. Hypotype SU 9189 [=CASG 70267].
324 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Turritella variata sanmarcosensis        
            
Santa Barbara Co. UCMP B6939. Paratypes SU 9191, 9192 [=CASG 70257.10, 70257.20]. [Turritellidae].
 
UCMP B6940. Holotype SU 9190 [=CASG 70257].
Turritella cf. -
ter” Sandstone, Eocene. Santa Barbara Co. Dibblee locality 238. Hypotype SU 9194 (not SU 9501 as in text and
pl. caption) [=CASG 70256]. [Turritellidae].
Turritella variata wheatlandensis
9. Wheatland Fm., Eocene. Northwest of Wheatland, Smartsville quad., Yuba Co. UCMP A1889]. Holotype UCMP
11059. [Turritellidae].
Turritella wheatlandensis
Eocene. Northwest of Wheatland, Smartsville quad., Yuba Co. UCMP A1889. Holotype UCMP 11059. Remarks:
Determined here as Turritella variata wheatlandensis. [Turritellidae].
Turritella

Turritella
Ventura Co. UCR 6825. Hypotype UCR 6825/100 (not UCR 6825/1 as cited in pl. caption). Remarks: Described
as Turritella cornuatus,  Zinsmeister (1983b: p. 1289). [Turritellidae].
Turritella n. sp. Saul 1983b. SEPM Guidebook 35:74, pl. 1, 
Ventura Co. CIT 531 [=LACMIP. 7153]. Hypotype UCLA 59363 [=LACMIP 10428]. Remarks: Determined here as
Turritella buwaldana Dickerson 1916. [Turritellidae].
Turritella
Valley, Orange Co. Hypotype determined here as missing. [Turritellidae].
Turritella
Sandstone, Paleocene. Just west of Burro Flats, Simi Hills, Ventura Co. Hypotype current whereabouts unknown.
Turritella-
ton quad., Santa Cruz Co. SCCM. 22. Hypotype SCCM. 21000]. Remarks: Turritella infragranulata pachecoensis
Stanton 1896,  L.R. Saul (personal communication, 1998). [Turritellidae].
Turritella
Inyokern SE quad. (1972), Kern Co. USGS M9067. Hypotype USNM 424890. [Turritellidae].
Turritella-
stone member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312.
Hypotype SDSNH 85244 (error) [83244 in errata, p. 50]. [Turritellidae].
Turritella    
Santa Barbara Co. SU. 1484. Hypotype SU 7414 [=CASG 72020]. Remarks: Turritella variata lompocensis Arnold,
1907,  Weaver and Kleinpell (1963: p. 185). [Turritellidae].
Turritella-
tura Co. SU. 2379. Hypotype SU/CASG 7238 determined here as missing. [Turritellidae].
Turris
Anita, Lompoc quad., Santa Barbara Co. UCMP B7013. Hypotypes SU 9232 (not SU 9504 as in text), 9236 [=CASG
70252, 70253]. [Turridae].
Tympanotonus (Tympanotonus) californicus
Sandstone, Eocene. Matilija quad. (1952, PR 1967), Ventura Co. CSUN 1453 [=LACMIP 16962). Holotype LACMIP
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 325
12442, paratypes LACMIP 12443–12448. Remarks: In the genus Certhideopsis,  Reid et al. (2008: table 4 on
p. 691).
Typhis (Talityphis) lampada

7949 [=CASG 72017, 7949]. [Muricidae].
Typhis (Talityphis) lampada
Silt, MioceneRio Bravo Ranch quad. (1954), Kern Co. USGS M1613. Hypotypes USNM 650129, 650130 detemined
here as missing. [Muricidae].
....., pl. 9, 
Typhis
Panoche and Silver creeks, Tumey Hills quad., Fresno Co. SU 2073 [=UCMP A9717]. Hypotype UCMP 14203.
[Muricidae].
Umpquaia oregonensis         
        
4865/102–4865/107. [Buccinidae].

11–14). Mobil Oil Co. (General Petroleum Corp.) 3891 ft. depth, Kern Co. UCR 1199. Hypotype UCR 1199/1 (not
UCR 1199 as in pl. caption). Remarks: Determined here as possible nassariid. [Nassariidae].


Los Angeles Co. Hypotype to be donated to LACMIP. [Naticidae].
-
per). Monterey Fm., Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to
LACMIP. [Olividae].

Fm., Miocene. George F. Canyon, Palos Verdes Hills, Los Angeles Co. Hypotype to be donated to LACMIP.
        

determined here as missing. Remarks: Determined here as possibly a buccinid.
Urosyca caudata
area, Contra Costa Co.]. Lectotype (of Stewart 1926 [1927]:379) ANSP 4208, [paralectotypes (4) ANSP. 79507].
Remarks: Name reassigned back to the original name Uroscya caudata Gabb from (Gabb),
 Squires (2014: p. 5. [Ficidae].


Hypotype LACMIP 14773.

-
types 14474, 14475.
Urosyca robusta Weaver, 1905. Squires, 2014a, Contra
Costa Co. UCMP 1888. Hypotype UCMP 11865. [Ficidae].

326 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Uzita arnoldi (Anderson, 1905). [Nassa     
[=Lower Round Mtn. Silt], Miocene. Kern River, Kern Co. USGS 6623. Hypotype USNM 563158. Remarks: In the
genus/subgenus Nassarius (Catilon),  Addicott (1965a: p. B12). [Nassariidae].
Vasum (Vasum) pufferi
Carrizo Mtn. [=Coyote Mtn.], Imperial Co. UCMP A1269. Holotype UCMP 15089. [Turbinellidae].
ex CASG
2447].
Vasum pufferi
Fm., Miocene/Pliocene. Coyote Mtn., Imperial Co. SDSNH 252. Hypotype SDSNH 23112. [Turbinellidae].
Velates californicus Vokes, 1935a. Squires and Filewicz 1983 (eds.). SEPM Guidebook 35:80 [follows Saul 1983b],

3173 [=LACMIP 23173]. Hypotype UCLA 58275 [=LACMIP 8881]. [Neritidae].
Velates californicus
Santa Susana Fm., Paleocene. Southwest of Runkle Canyon, Calabasas quad., Simi Hills, Ventura Co. UCLA 3173
[=LACMIP 23173]. Hypotypes UCLA 58274, 58275 [=LACMIP 8880, 8881]. [Neritidae].
Velates perversus (Gmelin, 1791). [Nerita
Springs Canyon, Topatopa Mtns. quad. (1943), Ventura Co. UCR 4668. Hypotypes UCR 4668/33, 4668/34. [Ne-
ritidae].
Velates perversus (Gmelin, 1791). [Nerita
Hills, Ventura Co. UCMP 7193 [=CSUN 702). Hypotype UCMP. 37435. [Neritidae].
Velates perversus (Gmelin, 1791). [Nerita
Bus Canyon Rd., Thousand Oaks quad., Simi Hills, Ventura Co. UCLA 7069 [=LACMIP 27069]. Hypotype UCLA
59278 [=LACMIP 8886]. [Neritidae].

[=LACMIP 24244]. Hypotypes UCLA. 58376, 59579 [=LACMIP 8884, 8887].
Velates perversus (Gmelin, 1791). [Nerita]. Squires 1987  
Eocene. Whitaker Peak quad. (1958), Los Angeles Co. CSUN 850 [=LACMIP 16400. Hypotypes LACMIP 7444,
7446, 7447. [Neritidae].

Velates perversus (Gmelin, 1791). [Nerita         
Fm., Eocene. Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 665 [=LACMIP 16335]. Hypotype
LACMIP 11454. [Neritidae].
Velates perversus (Gmelin, 1791). [Nerita
Paleocene/Eocene. Calabasas quad. (1952, PR 1967), Ventura Co. LACMIP 7124. Hypotype LACMIP 12676. [Ne-
ritidae].

member), Miocene. Fossil Reef Park, Laguna Hills, Orange Co., San Juan Capistrano quad. SDSNH 4312 [4512 in
errata is erroneous]. Hypotype SDSNH 142016. [Vermetidae].
Vitrinella (Vitrinellops) lens (Keen, 1943). [Teinostoma
Mtn. Silt, MioceneRio Bravo Ranch quad. (1954), Kern Co. USGS M1612. Hypotype USNM 650052. [Vitrinellidae].

Vitrinella stearnsi
Fm. (Cebada Member), Pliocene. Graciosa Ridge, near Orcutt, Santa Barbara Co. USGS 14648. Hypotype USNM
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 327
560065. [Vitrinellidae].
Volutilithes orocopiaensis
Fm., Eocene. Orocopia Mtns., Canyon Spring NW quad. (1963), Riverside Co. CSUN 662 [=UCLA 3779, =LACMIP

Volutilithes orocopiaensis-

LACMIP 7168. [Volutidae].
Volutilithes torreyensis
Pines St. Park, San Diego Co. UCR 4920. Holotype UCR 4920/1. [Volutidae].
Volutocorbis (Retipirula) crassitesta (Gabb, 1869b). [Turbinella   
Retipirula crassitesta)). Martinez Fm., Paleocene. Marti-
nez, Contra Costa Co. Lectotype (of Stewart, 1926 [1927]: p. 406) ANSP 4332 missing,  K. Estes-Smargiassi
(personal communication, 2019), paralectotypes (3) ANSP 79506]. Remarks: In the genus Retipirula,  Saul
and Squires (2008b: p. 234). [Volutidae].
Volutocorbis (Retipirula) crassitesta (Gabb, 1869b) [Turbinella]. Zinsmeister 1977. Jour. 
Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6846. Hypotype UCR.6846/12. [Volutidae]. Remarks:
In the genus Retipirula,  Saul and Squires (2008b: p. 234). [Volutidae].



Volutocorbis (Retipirula) crassitesta (Gabb, 1869b). [Turbinella
15. Santa Susana Fm., Paleocene. Simi Hills, Ventura Co. UCR 6863. Hypotype UCR 6863/151 (not UCR 6803/1 as
in pl. caption). [Volutidae]. Remarks: In the genus Retipirula,  Saul and Squires (2008b: p. 234). [Volutidae].
Volutocorbis virginiae       
Paleocene. Vicinity of Meier and Runkle canyons, Calabasas quad. (1952 [PR 1967), Simi Hills, Ventura Co. UCR

[Volutidae].
Volutocorbis virginea*
Fm., Paleocene. Simi Hills, Ventura Co. UCR 6887. Hypotype UCR. 6887/1. Remarks: Error for virginiae. [Volutidae].
Volutocristata lajollaensis (M.A. Hanna, 1927). [Plejona
62-29. Llajas Fm., Eocene. Aliso Creek, Los Angeles Co. CASG 364. Hypotype CASG 5714. Remarks: In the genus
Lyrischapa, Squires (1984: p. 36; 1987: p 47). [Volutidae].
Volvulella gluma


....., SDSNH5422 (ex
Volvulella gluma
quad. (1950), Kern Co. UCMP B1598. Hypotype UCMP 37102 [Rhizoridae].

Volvulella joaquinensis
Caliente quad. (1914), Kern Co. UCMP B1637. Holotype UCMP 37101. [Rhizoridae].

328 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Oak, Kern Co. CASG 244. Hypotype CASG 5713. Remarks: In the genus Strepsidura,  Wenz (1943: p. 1270).
[Fasciolariidae].
Whitneyella sinuata (
Creek, Lompoc quad., Santa Barbara Co. UCMP B6985. Hypotype SU 9231 [=CASG 70273, as Whitneyella sinuata
aff.  in CASG online database]. [Fasciolariidae].
Whitneyella         

lost, C. Throckmorton (personal communication, 2002). [Fasciolariidae].
Xenophora simiensis
Paleocene. Simi Hills, Ventura Co. UCR 6871. Hypotype UCR 6871/20. [Xenophoridae].
Xenophora stocki
Lion Canyon quad. (1943.), Ventura Co. UCR 4680. Hypotype UCR 4680/7. [Xenophoridae].
Xenophora stocki
Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN [=LACMIP 7242]. Hypotype LACMIP 6520. [Xenophoridae].
Xenophora stocki   
Valley, Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 6520. [Xenophoridae].
Xenophora stocki-
ley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 979 [=LAMCIP 16081]. Hypotype LACMIP 7701. [Xenophoridae].
Xenophora stocki-
cene. Upper Sespe Creek, Ventura Co. LACMIP 23735. Hypotype LACMIP 11650. [Xenophoridae].
Xenophora zitteli Weaver, 1905 (not Dickerson as cited in pl. caption). Zinsmeister 1983a. SEPM Guidebook 35:69, pl.

Xenoturris antiselli (Anderson and Martin, 1914). [Drillia 
Olcese Sand, Miocene. Caliente quad., Kern Co. SU 29501. Hypotype SU 9942 [=CASG 73129]. [Turridae].

650257.
Zonaria (Zonaria) emmakingae, 3. Topanga Canyon Fm. (Cold Creek
Member), Miocene. Santa Monica Mtns., Los Angeles Co. LACMIP 5136. Holotype LACMIP 12277. [Reprinted
in Bull. SCPA 26(9-10): pp. 81, 82, 1994]. Remarks: In the genus Pseudozonaria,  Lorenz, (2017: p. 399).
[Cypraeidae].
Zonaria (Neobernaya) spadicea (Swainson, 1823a). [Cypraea
Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 7752. Hypotype LACMIP 14388. Remarks: In the genus
Neobernaya
Zonaria spadicea fernandoensis (Arnold, 1907). [Cypraea]. Cate 1969. Veliger 12(1):116, pl
Fm.” [=Pico Fm.], Pliocene. Elsmere Canyon, near Newhall, Los Angeles Co. Holotype USNM 164961. Remarks:
Junior synonym of Neobernaya spadicea (Swainson 1823a),  Groves (2019: p. 133). [Cypraeidae].
CLASS POLYPLACOPHORA 
Amicula solivaga 
Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Holotype LACMIP
13846. Paratypes LACMIP 13842–13845, 13847–13850, 13852–13856. [Mopaliidae].

LACMIP 16817 [=LACMIP 305C]. Paratype LACMIP 13851.
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 329
Callistochiton palmulatus 
Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes


Co. LACMIP 16817 [=LACMIP 305C]. Hypotypes LACMIP 13756, 13758–13763.
Callistochiton sphaerae
Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Holotype LACMIP


16817 [=LACMIP 305C]. Paratype LACMIP 13854.
Dendrochiton         
Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP
14285–14291. [Mopaliidae].
Leptochiton nexus
Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP
13739–13746. [Leptochitonidae].
Leptochiton rugatus (Pilsbry, 1892) species complex. [Lepidopleurus]. Vendrasco et al. 2012. NHMLAC CS 520:20–21,
Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP
305. Hypotypes LACMIP 13730–13738. [Leptochitonidae].

Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Holotype LACMIP
13810, paratypes LACMIP 13801–13809, 13811.13812, 13814–13816. [Ischnochitonidae].

LACMIP 16817 [=LACMIP 305C]. Paratype LACMIP 13813.
Lepidozona mertensii (Middendorff, 1847). [Chiton
9.14, 9.15. San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP
305. Hypotypes LACMIP 13780, 13781, 13785, 13786. [Ischnochitonidae].

Co. LACMIP 16817 [=LACMIP 305C]. Hypotypes LACMIP 13777–13779, 13782–13784.
Lepidozona pectinulata (Carpenter in Pilsbry, 1892). [Ischnochiton
10.1. 10.12, 10.14–10.24. San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego
Co. LACMIP 305. Hypotypes LACMIP 13787–13792, 13974–13798. [Ischnochitonidae].
    
Diego Co. LACMIP 16817 [=LACMIP 305C]. Hypotype LACMIP 13793, 13799, 13800.
Lepidozona sp. cf.  (Carpenter in Pilsbry, 1892). [Ischnochiton]. Vendrasco et al. 2012. NHMLAC CS 520:30,
33, 11.6–11.10. San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co.
LACMIP 305. Hypotypes LACMIP 13804–13806. [Ischnochitonidae].
Lepidozona sp. cf. 
Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP
13801–13803. [Ischnochitonidae].
Mopalia sinuata
Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP
13894, 13895. [Mopaliidae].
Mopalia sp. cf. 
330 PALEOBIOS, VOLUME 38, FEBRUARY 2021
San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypo-
types LACMIP 13858–13872. [Mopaliidae].
  
LACMIP 16817 [=LACMIP 305C]. Hypotype LACMIP 13857.
Mopalia sp. 15.27. San Diego Fm., Pliocene. Goat Can-
yon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotype LACMIP 13873. [Mopaliidae].
Nuttallina
Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP 13892.
[Lepidochitonidae].
Oldroydia percassa (Dall, 1894). [Lepidopleurus        
Diego Fm., Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP

Placiphorella sp. cf. 
Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LAC-
MIP 13879–13883. [Mopaliidae].
Placiphorella velata         
Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP
13874–13878. [Mopaliidae].
Stenoplax circumsenta
Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes LACMIP
13817–13820. [Ischnochitonidae].
 
LACMIP 16817 [=LACMIP 305C]. Hypotype LACMIP 13819, 13820.
Stenoplax fallax (Carpenter in Pilsbry, 1892). [Ischnochiton
12.14, 12.17–12.21. San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co.


LACMIP 16817 [=LACMIP 305C]. Hypotype LACMIP 13827.
Stenoplax sp. cf. 
13.19. San Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305.
Hypotypes LACMIP 13832–13838, 13840, 13841. [Ischnochitonidae].

LACMIP 16817 [=LACMIP 305C]. Hypotype LACMIP 13819.
Tonicella sp. cf.   Clark, 1999. Vendrasco et al. 2012. NHMLAC CS 
Diego Fm., Pliocene. Goat Canyon, just north of the U.S./Mexico boundary, San Diego Co. LACMIP 305. Hypotypes
LACMIP 13884–13891. [Mopaliidae].
CLASS SCAPHOPODA 
Cadulus (Platyschides?) addicotti-

[Gadilinidae].
Cadulus (Platyschides) gabbi

paralectotype ANSP 79583. [Gadilinidae].
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 331
Cadulus (Cadulus) mcfarlandae-
caville Shale”], Eocene. Ualatis Creek, southwest of Dunns Peak, Vaca Valley, Solano Co. UCMP A3365. Holotype

Coccodentalium emersoni
Valley, Sawmill Mtn. quad. (1943), Ventura Co. CSUN 979 [=LACMIP . 16081]. Holotype LACMIP 7690. [Dentaliidae].
Dentalium (Laevidentalium)-
cene. Las Llajas Canyon, Santa Susana Mtns., Ventura Co. LACMIP 16211. Hypotype LACMIP 6507. [Dentaliidae].
Dentalium (Laevidentalium)
Canyon, Santa Susana quad., Los Angeles Co. CSUN 373 [=LACMIP. 16211]. Hypotype LACMIP 6507. [Dentaliidae].
Dentalium cf. 
of Pleasanton, Dublin quad. (1943), Alameda Co. SU 3229. Hypotype SU 8481 [=CASG 69951]. [Dentaliidae].
Dentalium (Antalis) cooperi -

Remarks: 

certain species, it is entirely probable that the type of  came from there.” [Dentaliidae].
Dentalium laneensis
Fm. (Phacoides Sand member) (of Stinemeyer et al. 1959:12, 13), Miocene. Las Yeguas Ranch quad., Temblor
Range, Kern Co. USGS M3772. Hypotype USNM 646611. Remarks: -
ceived from author,” and is presumed missing. Fig. 9 is missing from pl. 1 as noted in ‘errata’ sheet provided by
the author. [Dentaliidae].
Dentalium laneensis

Agua Creek, Las Yeguas Ranch quad., Kern Co. USGS M3772. Hypotype USNM 646611. Remarks: Label in USNM

Dentalium (Fissidentalium?) mcganna Powell, et al., 2019b (n. sp.). PaleoBios 36:15–17
Fm., Miocene. Caldecott Tunnel excavation, Berkeley Hills, Oakland, Alameda Co. UCMP IP13003. Holotype UCMP
251012, paratypes 218713, 218799. Remarks: Although the species name should have been correctly formed
macgannae,” it is left here as is, for taxonomic stability, as should subsequent authors, in
accordance with ICZN Article 32.5.1 (ICZN, 1999). [Dentaliidae].

UCMP 14436.
Dentalium neohexagonum Sharp and Pilsbry in 
101. Pico Fm., Pliocene. Newhall area, Los Angeles Co. LACMIP 7752. Hypotypes LACMIP 14417, 14418. [Den-
taliidae].
Dentalium stentor
Las Llajas Canyon, Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 444 [=LACMIP 16203]. Hypotype
LACMIP 6506. [Dentaliidae].
Dentalium stentor
Eocene. Sharps Canyon, Whitaker Peak quad. (1958), Los Angeles Co. CSUN 219 [=LACMIP 16100]. Hypotype
LACMIP 7439. [Dentaliidae].
Dentalium stentor
Dell Shale (Circle B Sandstone Member), Eocene. Upper Sespe Creek, Ventura Co. CSUN 531 [=LACMIP 16668].
Hypotype LACMIP 11641. [Dentaliidae].
Dentalium stentor

332 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Hypotype LACMIP. 6506. [Dentaliidae].
Dentalium stramineum
Maniobra Valley, Red Canyon quad. (1986), Riverside Co. CSUN 676 [=LACMIP 16192]. Hypotype LACMIP 11451.
[Dentaliidae].
Dentalium
Anita, Lompoc quad., Santa Barbara Co. UCMP B7013. Hypotype SU 9241 [=CASG 70160]. [Dentaliidae].
Dentalium
Angeles Co. CIT 1670 [=LACMIP 7751]. Hypotype LACMIP 4216. [Dentaliidae].
Dentalium    
Monocline Ridge quad. (1955), Fresno Co. UCMP D6531]. Hypotype UCMP 37160. [Dentaliidae].
Dentalium sp. Phillips et al. 1976.    Fm., Oligocene. Vicinity of
Iverson Pt., Saunders Reef quad., Mendocino Co. UCMP D7149. Hypotype UCMP 14436. Remarks: Described as
Dentalium (Fissidentaliummcganna by Powell et al. 2019b. [Dentaliidae].
CLASS CEPHALOPODA 
Aturia angustata (Conrad, 1849a). [Nautilus-
cene. Along Deer Creek, Santa Cruz Co. Hypotype SU 5379 [=CASG 69748], determined here as missing. [Aturidae].

Hypotype SU 612 [=CASG 71609].
Aturia angustata (Conrad, 1849a) [Nautilus

Aturia angustata (Conrad, 1849a) [Nautilus
  
LACMIP 13064. [Aturidae].
Aturia dickersoni
Fm., Paleocene. Mt. Diablo area, Contra Costa Co. UCMP 150. Holotype UCMP 11840. [Aturidae].
Aturia kerniana-
cene. Liveoak Canyon, Kern Co. CASG 61661. Hypotype SU 7917 [=CASG 72106]. [Aturidae].

CASG 5222.
Aturia myrlae
Member),” Eocene. Tecolote Valley, La Jolla quad., San Diego Co. UCMP 3989. Holotype UCMP 31089. [Aturidae].

7004]. Hypotype UCMP 31894.

SU 7891 [=CASG 72107].
Aturia myrlae
Santa Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 7242]. Hypotype LACMIP 6595. [Aturidae].
Aturia myrlae

6595]. [Aturidae].
Aturia myrlae M.A. Hanna, 1927. Specimen illustrated on front cover of the present paper. Llajas Fm., Eocene,
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 333
north side of Simi Valley, Ventura Co.. LACMIP 40374. Hypotype LACMIP 14872. Remarks: A juvenile specimen.
[Aturidae].
Aturia sp. [of . Agasoma gravidum beds, Contra Costa Co.
UCMP 1173]. Hypotype (of
AturiaAturia sp. in pl. caption). Capay Fm.
(Vacaville Shale Member), Eocene. Putah Creek, Yolo Co. UCMP A3990. Hypotype UCMP 30285. [Aturidae].
Aturia cf.  Hanna in pl. caption). Arroyo Hondo Fm., Eocene. Near where Urruttia
Canyon enters Salt Creek, north of Coalinga, Fresno Co. UCMP 1817. Hypotype UCMP 15877.
Aturoidea mathewsoni (Gabb, 1864). [Aturia

lost,  Miller (1947: p. 71). [Hercoglossidae].

Eutrephoceras hallidayi (Waring, 1914). [Nautilus          
[=Santa Susana Fm.], Paleocene. Simi Hills, Calabasas quad., Ventura Co. CASG 69612. Holotype SU 560 =[CASG
69725]. [Nautilidae].
Eutrephoceras hannai
Shale Member),” Eocene. Rose Canyon area, La Jolla quad., San Diego Co. UCMP IP2720. Holotype UCMP 32296.
[Nautilidae].

Co. CASG 61655. Hypotypes SU 7896. 7897 [=CASG 72111].
Eutrephoceras marksi
area, Tejon quad., Kern Co. CASG 72114. Holotype SU 7899 [=CASG 72114]. [Nautilidae].
Eutrephoceras stephensoni (Dickerson, 1914a). [Nautilus
Fm., Paleocene. Muir Station area, Concord quad., Contra Costa Co. UCMP 243. Holotype UCMP 11841]. [Nautilidae].

UCMP 15983.

Eutrephoceras aff.  (Dickerson, 1914a). [Nautilus]. Miller and Downs 1950. Jour. Paleo. 24(1):3, pl. 1,

Eutrephoceras
Yolo Co. UCMP A3073]. Hypotype UCMP.11412. [Nautilidae].
Eutrephoceras sp. Shapiro
Cañada Posa area, Santa Cruz Is., California Channel Ids. Hypotype UCSB 8704 [=SBMNH 60177]. [Nautilidae].
Hercoglossa merriami
Near Martinez, Contra Costa Co. Hypotype ANSP 16122, missing,  K. Estes-Smargiassi (personal communica-
tion, 2019). [Hercoglossidae].

3763. Hypotype UCMP 15984.

UCMP 11842.
Kummelonautilus  
334 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Susana quad. (1951, PR 1969), Ventura Co. CSUN 374 [=LACMIP 22312]. Hypotype LACMIP 12759. [Nautilidae].

   
13056. [Argonautidae].



LACMIP 13060.

Co. LACMIP 17558. Paratype LACMIP 13061.
-
MIP 12021. Paratype LACMIP 13063.

LACMIP 17598. Paratype LACMIP 13057.

Hills quad., Santa Monica Mtns., Los Angeles Co. LACMIP 23350. Paratype LACMIP 13062.
Rhyncolites
Diego, San Diego Co. Hypotype KU 115,355 (not KU 155,355 as published,  R. Kaesler (personnal ommunica-
tion, 1998)), missing,  R. Kaesler (personal communication, 1998). [family uncertain].
Sepiamorph sepiid Squires 1991e. BSCAS 90(2):
Fm.], Eocene. Rose Canyon, San Diego Co. LACMIP 16359]. Hypotype LACMIP 8407. [Sepiidae].

Susana quad. (1951, PR 1969), Ventura Co. CSUN 493 [=LACMIP 16198]. Hypotype UCLA 59197 [=LACMIP
12885]. [Spirulidae].

1969), Ventura Co. CSUN 493 [=LACMIP 16198]. Hypotype UCLA 59197 [=LACMIP 12885]. [Spirulidae].
Woodringia simiensis (Vokes, 1937a). [Hercoglossa . Martinez
Fm. [=Santa Susana Fm.], Paleocene. Simi Valley, Santa Susana quad., Ventura Co. UCMP 7016. Holotype UCMP
15985. [Hercoglossidae].
ACKNOWLEDGEMENTS
The completion of this check list was facilitated greatly
by the generous cooperation and hospitality of many re-
searchers, curators, collection managers, and colleagues.
We express our sincere thanks to you all. In alphabetic
order by institution and/or company we thank Linda Gil-
bert and George T. Jefferson (ABDSP), Elana Benamy, Paul
Callomon, Katy Estes-Smargiassi, Ned Gillmore, and Gary
Rosenberg (-
lege, CA), George L. Kennedy (Brian F. Smith & Associates
and SDSU), the late Jean DeMouthe, Christine (Chrissy)
Garcia, Lou Marincovich, Jr., Peter U. Rodda, and Peter
D. Roopnarine (CASG), Eugene V. Coan, Amanda Grimes,
and Elizabeth Kools (CASIZ), Brian J. Swanson (California
Geological Survey, Los Angeles), Ángel Valdés (California
Polytechnic State University, Pomona, CA), the late Alan
Horowitz (IU), Lori Wear (Kern County Museum, Bakers-
. Alderson, Harry F. Filkorn, Austin J.W.
Hendy, the late Philip G. Owen, LouElla R. Saul, Mary S.
Stecheson, and Lindsay Walker (LACMIP), the late James
H. McLean and Jann E. Vendetti (LACM Malacology), Lau-
rie C. Anderson (LSU), the late Fred Collier (MCZ), the late
Rodney Watkins (MPM), Cam Wallis (NHFOC), Warren
D. Allmon and Wendy Taylor (PRI), Daniel L. Geiger and
Paul Valentich-Scott (SBMNH), Frank Perry (SCCM),
Tom Deméré, Kesler Randall, N. Scott Rugh, and Patricia
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 335
Don Vito (SDSNH), Susan Kidwell (UC), Mark Goodwin,
David Haasl, Carole S. Hickman, Pat Holroyd, David R.
Lindberg, Dave Smith, and Karen Wetmore-Grycewicz
(UCMP), Marylin Kooser (UCR), the late Robert Norris,
Russell S. Shapiro (CSU), and Bruce Tiffney (UCSB), the
late Warren C. Blow, Charyl Ito, Jann Thompson, and
Thomas R. Waller (USNM) all of whom made collections,
locality information, and libraries available to us. The
late Warren O. Addicott (Ashland, OR), the late Yvonne
Z. Albi (Playa del Rey, CA), the late Richard R. Alexander
(Rider University, Lawrenceville, NJ), the late Ray Givens
(Nicholls State University, Thibodaux, LA), the late Ellen J.
Moore (Corvalis, OR), Daniel R. Muhs (USGS, Denver, CO),
Charles L. Powell, II (USGS, Menlo Park, CA), Judy Terry
Smith (SU and USNM), Carol J. Stadum (Carlsbad, CA),
Jerry Treiman (CDMG), and Sally E. Walker (University
of Georgia) provided reprints of published papers and/
or personal insights. George L. Kennedy (La Mesa, CA)
and Patrick I. LaFollette (Cathedral City, CA) kindly re-
viewed the Pholadidae and Pyramidellidae text entries
respectively and made helpful corrections and sugges-
tions. Eugene V. Coan (CASIZ) provided photocopies of
annotated pages from Myra Keen's personal copy of the
1944 checklist. We especially thank LouElla R. Saul for
sharing her unequaled knowledge of the former UCLA
invertebrate-fossil collection. Dr. Bentson H. McFarland
(Seattle, WA) is thanked for supplying us with heretofore
unpublished biographic information and the image of
his
Herbert, Richard Hulser, and the late Don McNamee
(LACM, Research Library) and Jean Crampon, Melinda
Hayes, and Suzanne Henderson (USC research libraries)
all aided in acquisition of rare and obscure references.
Janet Squires   
problems encountered during the preparation of the
    sincerely
thank his wonderful wife Cathy L. Groves (LACM Echi-
noderms Department) for accompanying him [while on

ABDSP, CASG, SBMNH, UCSB, and USNM. James L. Goed-
ert (Gig Harbor, WA), Austin J.W. Hendy (LACMIP), and
Charles L. Powell, II (USGS ret.) reviewed the manuscript
and provided meaningful suggestions and improvements.
Special thanks to PaleoBios Senior editor Diane M. Erwin
who did a superb job throughout the review and editing
process. Her expertise and patience are greatly appreci-
ated. We sincerely thank you Diane.
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in 
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     
coast of North America: An annotated bibliography, 1797–
1969. 
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paleontology of the lower part of the type Temblor Formation,
California.     791:iv +



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condoni Dall, a unique Pliocene and Pleistocene bivalve from
The Veliger
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of the Salinas Basin during the latest part of the Miocene,
with notes on macrofossils from near San Lucas, California.
   in W.O. Addicott (ed.). Neogene
336 PALEOBIOS, VOLUME 38, FEBRUARY 2021
biostratigraphy of selected areas in the California Coast Ranges.
 78-446.
Addicott, W.O. 1979. Oligocene molluscan biostratigraphy of the
Indians, Santa Lucia Range, central California. Pp. 45–49, pl.
1 in S.A. Graham (ed.). Tertiary and Quaternary geology of
the Salinas Valley and Santa Lucia Range, Monterey County,
California. 
and Mineralogists Guidebook 14.
-
Geological Society of America
Special Paper
Addicott, W.O., and J.S.Galehouse.1973. Pliocene marine fossils in
the Paso Robles Formation, California. Journal of Research of

Addicott, W.O., R.Z. Poore, J.A. Barron, H.D. Gower, and K. McDou-
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Creek area, northern La Panza Range, California. Pp. 49–81,
  in W.O. Addicott (ed.). Neogene biostra-
tigraphy of selected areas in the California Coast ranges. 
Geological Survey Open File Report 78.
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area, northern La Panza Range, San Luis Obispo County, Cali-
in G.H. Blake (ed.). Neo-
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Paleontologists and Mineralogists Guidebook 6.
Chaceia
(Pholadidae: Martesiinae) from the late Miocene of California.
The Veliger 9(2):233–235, pl. 21.
Adegoke, O.S. 1967a. Bryozoan-mollusk relationships. The Veliger
9(3):298–300, pl. 40.
Adegoke, O.S. 1967b. California late Miocene records of Swifto-
pecten Hertlein, 1935 (Pelecypoda: Pectinidae). The Veliger
9(3):337–339, pl. 47.
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ical Survey Professional Paper 1228-F:” 107 non-consecutive
pages, pls. 1–14 [currently only available via electronic for-
mat of Chintimini Press, Corvalis OR: http://members.peak.
org/~chintimp/Tertiary.pelecypods.htm ].
Moore, E.J. 2003b. Tertiary marine pelecypods of California and
Baja California: Tellinidae  -
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pages, pls. 1–11 [currently only available via electronic for-
mat of Chintimini Press, Corvalis OR: http://members.peak.
org/~chintimp/Tertiary.pelecypods.htm ]. Note: These im-
portant on-line references by Ellen J. Moore remain available
as per our updated submission date of January 2021. In 2003
Ellen announced the
Papers 1228-F and 1228-G” and that they were last updated on
July 1, 2003. The senior author corrected many of the errors
that they contained and strongly urged Ellen to make these
corrections. The corrected versions were subsequently posted
but many errors remain.
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Natural History Museum of Los Angeles County Contributions
in Science 

in M.V. Filewicz
and R.L. Squires (eds.). Paleogene stratigraphy, west coast of
North America. -
gists and Mineralogists Guidebook 58.
Squires, R.L. 1988b. Rediscovery of the type locality of Turritella
andersoni and its geologic age implications for west coast
in M.V. Filewicz and R.L.
Squires (eds.). Paleogene stratigraphy, west coast of North
America.    
and Mineralogists Guidebook 58.
Squires, R.L. 1988c. Eocene macropaleontology of northern
Lockwood Valley, Ventura County, California. Natural His-
tory Museum of Los Angeles County Contributions in Science

Squires, R.L. 1989a. A new pseudolivine gastropod genus from
the lower Tertiary of North America. Journal of Paleontology

Squires, R.L. 1989b. New stratigraphic and geographic occur-
rences of Isognomon (Mollusca: Bivalvia) from the Eocene of
California and Oregon. Transactions of the San Diego Society of
Natural History 
Squires, R.L. 1989c. First Tertiary occurrence of a rare patelliform
gastropod (Archaeogastropoda: Symmetrocapulidae), Eocene
Tejon Formation, Tehachapi Mountains, California. The Veliger

Squires, R.L. 1990a. New Paleogene siliquariid and vermetid gas-

The Veliger
Squires, R.L. 1990b. First occurrence of the Tethyan bivalve Naya-
dina (Exputens) in Mexico, and a review of all species of this
North American subgenus. The Veliger
Squires, R.L. 1990c. New Paleogene Fimbria (Mollusca: Bivalvia)
Journal
of Paleontology
Squires, R.L. 1991a. New early Eocene species of Arca s.s. (Mollus-
ca: Bivalvia) from southern California. The Veliger 34(1):67–72,

Squires, R.L. 1991b. New morphologic and stratigraphic data on
Calyptogena (Calyptogena) gibbera Crickmay, 1929 (Bivalvia:
Vesicomyidae) from the Pliocene and Pleistocene of southern
California. The Veliger
Squires, R.L. 1991c. Molluscan paleontology of the lower Eocene
Maniobra Formation, Orocopia Mountains, southern California.
in P.L. Abbott and J.A. May (eds.).
Eocene geologic history San Diego region. 
of Economic Paleontologists and Mineralogists Guidebook 68.
Squires, R.L. 1991d. A new middle Eocene potamidid gastropod
from brackish-marine deposits, southern California. The Veliger

Squires, R.L. 1991e. A new sepiamorph sepiid (Mollusca: Cepha-
lopoda) from the Eocene of California. Bulletin of the Southern
California Academy of Sciences
Squires, R.L. 1992. New morphologic and geographic data on the
neritid gastropod Nerita (Theliostyla) triangulata Gabb, 1869,
       The
Veliger
Squires, R.L. 1993a. New reports of the large gastropod Campanile
     
America. The Veliger
Squires, R.L. 1993b. The gastropod Terebra santana Loel and
Corey, 1932, from the lower Miocene Vaqueros Formation,
southern California, belongs in the cerithiid genus Clavoceri-
thium s.s. The Veliger 
Squires, R.L. 1994a. New morphologic information on the bivalve
Isognomon (Isognomon) panzana (Loel and Corey, 1932) from
the lower Miocene Vaqueros Formation, southern California.
The Veliger
Squires, R.L. 1994b. Macropaleontology of Eocene marine rocks,
upper Sespe Crek area, Ventura County, southern California.
in A.E. Fritsche (ed.). Sedimentol-
ogy and paleontology of Eocene rocks in the Sespe Creek area,
Ventura County, California. 
Paleontologists and Mineralogists Guidebook 74.
354 PALEOBIOS, VOLUME 38, FEBRUARY 2021
Squires, R.L. 1995. The gastropod Philene bisculpta Hanna, 1927,
from the Eocene of southern California, belongs in the neritid
genus Otostoma d'Archiac, 1859. The Veliger 38(2):176–180,

Squires, R.L. 1997a. Taxonomy and distribution of the buccinid
gastropod Brachysphingus from uppermost Cretaceous and
     
America. Journal of Paleontology

in P. Havens (ed.). Simi Valley: A journey
through time. 
Squires, R.L. 1998. New information on morphology, stratigraphy,
and paleoclimate inplications of the Eocene brackish-marine
gastropod Loxotrema turritum Gabb, 1868, from the west coast
of the United States. The Veliger
Squires, R.L. 1999a. William More Gabb: First paleontologist of the
Geological Survey of California. California Geology 52(4):11–14,

Squires, R.L. 1999b. Upper Paleocene to lower Eocene ("Me-
ganos Stage") marine megafossils in the uppermost Santa
Susana Formation, Simi Valley, southern California. Natural
History Museum of Los Angeles County Contributions in Science

Squires, R.L. 1999c. Middle Eocene brackish-marine mollusks
from the Matilija Sandstone at Matilija Hot Springs, Ventura
County, southern California. Natural History Museum of Los
Angeles County Contributions in Science 
Squires, R.L. 2000. New information on a poorly known late Pa-
leocene turrid gastropod from southern California and vicinity.
The Veliger
Squires, R.L. 2001. Additions to the Eocene megafossil fauna of
the Llajas Formation, Simi Valley, southern California. Natural
History Museum of Los Angeles County Contributions in Science

Squires, R.L. 2003. Turnovers in marine gastropod faunas dur-
ing the Eocene-Oligocene transition, west coast of the United
in D.R. Prothero, L.C. Ivany,
and E.A. Nesbitt, E.A. (eds.). From greenhouse to icehouse.
The marine Eocene-Oligocene transition. Columbia University
Press, New York.
Squires, R.L. 2008. Eocene megapaleontology, stratigraphy, and
depositional environments, Elsmere Canyon, Los Angeles
County, southern California. Natural History Museum of Los
Angeles County Contributions in Science 
    
Paleocene glycymeridid bivalves. Journal of Paleontology

Pyropsis
(Neogastropoda: Pyropsidae) and paleobiogeography of the
genus. Journal of Paleontology
Squires, R.L. 2012. Late Pliocene megafossils of the Pico Formation,
Newhall area, Los Angeles County, southern California. Natural
History Museum of Los Angeles County Contributions in Science

Squires, R.L. 2013. West coast North America record of the Paleo-
cene marine stromboid gastropod Rimella and paleobiogeog-
raphy of the genus. Journal of Paleontology 87(5):826–841,

Urosyca,
, and Ficus from coastal-western North America:
Phylogenetic and global paleobiogeographic implications.
Natural History Museum of Los Angeles County Contributions
in Science 
Squires, R.L. 2014b. Cretaceous and Paleogene Pteria bivalves from
The Veliger 51(4):216–236,

Squires, R.L. 2015a. First report of the Eocene gastropod Mitrella
(Bastrophia) (Neogastropoda: Columbellidae) from the north-
The Nautilus

     
genus Pseudoperissolax (Neogastropoda: Muricidae: Muri-
cinae) and its paleobiogeography. Journal of Paleontology

Squires, R.L. 2018a. Paleocene and Eocene oysters from the west
coast of the United States: Revision of named species and
recognition of new species. Natural History Museum of Los
Angeles County Contributions in Science 
Squires, R.L. 2018b. First report of Oniscidia Mörch, 1852 (Neo-

paleobiogeographic review of the genus. Journal of Paleontol-
ogy
Squires, R.L. 2018c. Catalogue of the taxonomic updates of

and gastropods named from 1874 to 1966. The Nautilus
132(3–4):91–100.
Squires, R.L. 2018d. Late Cretaceous endemic shallow-marine
gastropod genera of the northeast : Biodiversity and
faunal changes. PaleoBios 35:1–16, . 1–5.
Squires, R.L. 2019. Eocene warm-water cassid gastropods from
coastal southwestern North America taxonomic revisions,
paleobiogeographic aspects, and faunal-turnover implications.
PaleoBios 
Squires, R.L., and D.M. Advocate. 1986. New early Eocene mollusks
from the Orocopia Mountains, southern California. Journal of
Paleontology
Squires, R.L., B.F. Cox, and C.L. Powell, II. 1988. Late Paleocene
or early Eocene mollusks from the uppermost part of the
in M.V.
Filewicz and R.L. Squires (eds.).Paleogene stratigraphy, west
coast of North America.    
Paleontologists and Mineralogists Guidebook 58.
Squires, R.L., and T.A. Deméré. 1991. A middle Eocene marine
molluscan assemblage from the usually nonmarine Friars

in P.L. Abbott and J.A. May (eds.). Eocene geologic history San
Diego region. 
and Mineralogists Guidebook 68.
Squires, R.L., and M.V. Filewicz (eds.). 1983. Cenozoic geology of
the Simi Valley area, southern California.  Section Society
of Economic Paleontologists and Mineralogists Guidebook 35.
Squires R.L., and A.E. Fritsche. 1978. Miocene macrofauna along
    
pls. 1–4 in A.E. Fritsche (ed.). Depositional environments of
Tertiary rocks along Sespe Creek, Ventura County, California.
-
alogists Guidebook 13.

bivalves from suspect cold seeps, Wagonwheel Mountain, cen-
tral California. Journal of Paleontology
GROVES & SQUIRES—CHECKLIST OF CALIFORNIA MOLLUSCA SINCE KEEN AND BENTSON (1944) 355
Squires, R.L., and L.T. Groves. 1993. First report of the ovulid
gastropod Sulcocypraea mathewsonii (Gabb, 1869) from the
Eocene of Washington and Oregon and an additional report
from California. The Veliger
Squires, R.L., L.T. Groves, and J.T. Smith. 2006. New information
on molluscan paleontology and depositional environments of
the upper Pliocene Pico Formation, Valencia area, Los Angeles
County, southern California. Natural History Museum of Los
Angeles County Contributions in Science 
Squires, R.L., and G.L. Kennedy. 1998. Additions to the late Paleo-
cene molluscan fauna from the Santa Monica Mountains, Los
Angeles County, southern California. The Veliger 41(2):157–

Squires, R.L., and P. Lipman. 2018. First occurrence of the Paleo-
cene gastropod Ravniella Rosenkrantz, 1970 (Heterobranchia:
Acteonidae) in North America. The Nautilus 132(2):51–57,

Squires, R.L., and L.R. Saul. 1993. A new species of Otostoma
(Gastropoda: Neritidae) from near the Cretaceous/Tertiary
boundary at Dip Creek, Lake Nacimiento, California. The Veliger

Squires, R.L., and L.R. Saul. 1997. Review of the bivalve genus Pli-
catula from Cretaceous and lower Cenozoic strata of California
and Baja California. Journal of Paleontology 71(2):287–298,

Squires, R.L., and L.R. Saul. 1998. New upper Paleocene species
of the bivalve Plicatula from southern California. Journal of
Paleontology
Squires, R.L., and L.R. Saul. 2000. The buccinid gastropod Deus-
senia from Upper Cretaceous strata of California. The Veliger

Squires, R.L., and L.R. Saul. 2001. A new genus of aporrhaid gas-
tropod from upper Paleocene rocks in southern California. The
Veliger
Squires, R.L., and L.R. Saul. 2002. New information on Late Cre-
taceous, Paleocene, and Eocene neritid gastropods from the
   The Veliger 45(3):177–192,

Squires, R.L., and L.R. Saul. 2006. New buccinoid gastropods from
uppermost Cretaceous and Paleocene strata of California and
Baja California, Mexico. The Nautilus
Squires, R.L., and L.R. Saul. 2007. Paleocene pareorine turritellid
The Nau-
tilus
Squires, R.L., and D.R. White. 1983. Common megafossils of the
lower Saugus Formation, northern Simi Valley area, Califor-
nia. Pp. 221–224, pl. 1 in R.L. Squires and M.V. Filewicz (eds.).
Cenozoic geology of the Simi Valley area, southern California.
-
alogists Guidebook 35.
Squires, R.L., W.J. Zinsmeister, and L.M. Paredes-Mejia 1989.
Popenoeum, a new pseudolovine genus: Widespread and
Journal of Paleontology

Stadum, C.J. 1973. A student guide to Orange County fossils.
Chapman College Press, Orange, California. 64 pp., numerous

Stadum, C.J. 1984a. The fossils of the Niguel Formation of south-
in B. Butler,
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County. Memoirs of the Natural History Foundation of Orange
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Stadum, C.J. 1984b. Saddleback Valley's fossil reef –A window to
in B. Butler, J. Gant, and
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Memoirs of the Natural History Foundation of Orange County 1.
Stadum, C.J., and K.L. Finger. 2016. Paleontology and stratigraphy
of the Miocene Saddleback Valley Limestone, Orange County,
southern California. Natural History Museum of Los Angeles
County Contributions in Science
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ogy
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
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
of Scaphopoda (Mollusca). Zoosystema 23(3):433–460.
     in
R.C. Moore (ed.).  
N, Mollucsa 6(3).  

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
Stinemeyer, E.S., R.S. Beck, R.A. Ortalda, E.K. Espenscheid, J.D. Bain-

[California].  15 pp.
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in L.A. York (ed.).
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360 PALEOBIOS, VOLUME 38, FEBRUARY 2021
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... Thus, it would appear that both Patinopecten healeyi and Euvola bella do range into the Pleistocene but are likely not younger than about 1.75 Ma. The most recent summary of other localities hosting E. bella are all in Pliocene formations (Groves and Squires, 2021). Mytilus coalingensis is known only from the Miocene and Pliocene, based on the compilation by Groves and Squires (2021). ...
... The most recent summary of other localities hosting E. bella are all in Pliocene formations (Groves and Squires, 2021). Mytilus coalingensis is known only from the Miocene and Pliocene, based on the compilation by Groves and Squires (2021). ...
Article
Glacial isostatic adjustment (GIA) models hypothesize that along coastal California, last interglacial (LIG, broadly from ~130 to ~115 ka) sea level could have been as high as +12 m to +14 m, relative to present, substantially higher than the commonly estimated elevation of +6 m. Areas with low uplift rates can test whether such models are valid. Marine terraces on Santa Cruz Island have previously been reported to occur at low (<10 m) elevations, but ages of many such localities are not known. Using lidar imagery as a base, marine terraces on Santa Cruz Island were newly mapped, elevations were measured, fossils were collected for U-series dating (corals), strontium isotope compositions and amino acid geochronology (mollusks), and paleozoogeography (all taxa). Sr isotope compositions of mollusks from the highest of three marine terraces give ages of ~2.5 Ma to 1.9 Ma, along with Pliocene ages, from shells interpreted to be reworked. U-series ages of corals from the western part of the island indicate that low-elevation terraces north of the Santa Cruz Island fault correlate to the LIG. Where corals are lacking, amino acid ratios and faunal aspects support terrace correlation to the LIG high stand of sea. Elevations of most terrace localities north of the east-west trending Santa Cruz Island fault, in both the western and eastern parts of the island, range from 5.75 m to 8 m above sea level, well below the modeled paleo-sea-level range. Subsidence is ruled out as a mechanism for explaining the lower-than-modeled elevations, because higher-elevation terraces are present along much of the Santa Cruz Island coast north of the fault, indicating long-term tectonic uplift. The low elevations of the LIG terrace fragments are, however, consistent with a low rate of uplift derived from the higher, ~2.5-1.9 Ma terrace. A number of other localities on the Pacific Coast, also dated to the LIG, have marine terrace elevations below the modeled level. GIA models may have overestimated last interglacial sea level by a substantial amount and need to be revised if used for forecasts for future sea-level rise.
... 12577) and San Miguel Island (see below). Elsewhere on the southern California mainland, it has been reported in marine deposits as old as Pliocene (Groves, 1991;Squires, 2012;Groves and Squires, 2021). With one exception on San Nicolas Island, which is interpreted to be a single specimen reworked from a higher terrace (Muhs et al., 2012), this species has not been reported from marine terrace deposits as young as 120 ka (MIS 5e). ...
... Names for seventeen species and eight subspecies have been attributed to the genus Nucella in the California Neogene (Grant and Gale 1931, Keen and Bentson 1943, Groves and Squires 2021. These species and their stratigraphic range are Nucella canaliculata (syn. ...
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Nucella demouthae n. sp. is a new muricid gastropod species described from the late Miocene part of the Wilson Grove Formation in Sonoma County, central California. It is distinguished from other fossil and modern California Nucella by its small size and faint to moderately strong radial sculpture, and its thickened and recurved outer lip with two denticles within the interior of the aperture. To date, this fossil rocky shore predatory gastropod has been found only at Bloomfield Quarry, north of the town of Bloomfield in Sonoma County, in the lowest part of the Wilson Grove Formation, which has been dated between 9.3 and 6.2 Ma. Thus, this species is part of the malacofauna of the late Miocene “Jacalitos” California provincial molluscan stage.
... Small and well-preserved fossil mollusks occur in remarkable numbers in some of the late Miocene Super Creek exposures of the "Imperial'' Formation north of Palm Springs, Riverside County, California. Among these are three members of the family Caecidae which are the first fossils record of the family from the Tertiary of California (Keen andBentson 1944, Groves andSquires 2021), although a few authors have recorded modern species in Pleistocene deposits along coastal California (Woodring et al. 1946, Valentine, 1959, 1989. ...
... Small and well-preserved fossil mollusks occur in remarkable numbers in some of the late Miocene Super Creek exposures of the "Imperial'' Formation north of Palm Springs, Riverside County, California. Among these are three members of the family Caecidae which are the first fossils record of the family from the Tertiary of California (Keen andBentson 1944, Groves andSquires 2021), although a few authors have recorded modern species in Pleistocene deposits along coastal California (Woodring et al. 1946, Valentine, 1959, 1989. ...
... Neritidae are generally rare in the Eocene of the U.S., only four to date from the Eocene of the Gulf Coastal Plain, Neritina unidentata Aldrich, 1911 from the early Eocene Hatchetigbee Formation of Alabama, Theodoxus domicilium Garvie, 1996, from the middle Eocene Reklaw Formation of Texas, Velates floridanus Richards, 1946, from the late Eocene, Inglis Formation of Florida, and Pseudodostia ocalana Vermeij and Portell, 2013, from the late Eocene Ocala Limestone of Florida. Many more have been described from the Eocene of the Pacific Coast (Groves and Squires, 2021). The species described herein, with the exception of C. traceyi new species, are all known from only from a single specimen, indicating that they were living at some distance from the localities at which they were found, and were transported there by ocean currents. ...
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ABSTRACT Nine new species of neritoidean gastropods are described from the fossil record of the Gulf Coastal Plain of the United States, eight of which are named and one is left in open nomenclature. Otostoma cattoi new species, Otostoma sp., and Neridomus mccallae new species, are from the Cretaceous (Albian) Edwards Formation of Texas. Neridomus minutissimus new species is from the Cretaceous (Albian) Walnut Formation of Texas. The genus Neridomus has not thus far been reported from the Cretaceous of the USA. Clithon cooncreekense new species is from the Cretaceous (Maastrichtian) Coon Creek type locality in Tennessee and is the first neritid reported from that site. Pseudodostia lisbonensis new species is from the Eocene (Bartonian) Lisbon Formation of Alabama. From the Eocene (Lutetian) Cook Mountain Formation of Texas, Clithon duomillias new species is from the Landrum Member, Clithon lisae new species is from the Wheelock Member, and Clithon traceyi new species is from the Stone City Member. These are the first reported neritids from the Eocene (Claibornian Stage) of the Gulf Coastal Plain.
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In areas of low uplift rate on the Pacific Coast of North America, reoccupation of emergent marine terraces by later high sea‐stands has been hypothesised to explain the existence of thermally anomalous fauna (mixtures of warm and cool species) of last interglacial age. If uplift rates have been low for much of the Quaternary, it follows that higher (older) terraces should also show evidence of reoccupation. Strontium isotope analyses of fossils from a high‐elevation marine terrace on Anacapa Island, California, yield ages ranging from ~2.4–2.3 Ma to ~1.4–1.5 Ma. These results indicate that terrace reoccupation and fossil mixing on Anacapa Island could have taken place over several interglacial periods in the early Pleistocene. Terrace reoccupation over this time period is likely a function of both a low uplift rate and the timing of orbital forcing of glacial–interglacial cycles. Climate change in the early Pleistocene was modulated by the 41 ka obliquity cycle, and glacial–interglacial cycles were much shorter than later in the Pleistocene. Nearby San Miguel Island also has evidence of terrace reoccupation, with Sr isotope ages of shells from several high‐elevation terraces ranging from ~1.21–1.25 Ma to ~0.43–0.50 Ma. However, the frequency of terrace reoccupation was lower than on Anacapa Island. The uplift rate of San Miguel Island is higher than that of Anacapa Island and terraces formed when glacial–interglacial cycles were longer. The frequency of marine terrace reoccupation is controlled by the rate of tectonic uplift and the timing of orbital forcing of sea level change during glacial–interglacial cycles.
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Thesis (Ph. D.)--University of California, Berkeley, 1979. Includes bibliographical references (leaves 717-775). Microfiche. s
Article
The Castaic formation comprises the upper Miocene marine sediments deposited in the Soledad Basin, northwestern Los Angeles County, California. The Soledad Basin is topographically the eastern extension of the Ventura Basin. Geologically, the two basins of deposition are separated by the San Gabriel fault along which an uncertain amount of lateral movement has taken place. Therefore, detailed correlation of the sediments of the Castaic formation with contemporary sediments of the Ventura Basin is not possible. The age of the Castaic formation is Mohnian and Delmontian. The sediments were deposited in part along an open coast and in part within a broad, open embayment. Late Miocene movement on the San Gabriel fault, relatively up on the southwest, defined the western margin of the basin of deposition. A land mass on the west side of the fault northwest of Castaic, created the embayment of the northwest part of the basin. It is likely that a submerged extension of this land southeast of Castaic formed a sill along the western margin of the rest of the basin and restricted the circulation of bottom water. The formation is about 7000 feet thick. It consists mainly of interbedded sandstone and mudstone. The two rock types occur in units 25 to 250 feet thick. Coarse-grained basal sediments were deposited at the margin of the basin of deposition. Megafossils are largely confined to the basal sediments. The nature of the basal sediments deposited at the margin of the transgressing late Miocene sea is related to the provenance and the relief along shore. Conglomeratic basal sediments were deposited along the northeast margin of the basin along a steep shore composed of the well-indurated Martinez formation. These sediments were derived locally from the conglomerate and sandstone beds of the Martinez formation. Sandy and thin conglomeratic basal sediments were deposited along the southeast margin of the basin. Near-shore relief was low. The sediments were derived largely from the finer-grained, less consolidated sediments of the Mint Canyon formation. The very coarse-grained, poorly-sorted Violin breccia was derived from a nearby source of considerable relief that was northwest of the embayment. The fauna of the Castaic formation is largely molluscan. Other phyla are represented by one species of brachiopod, two species of echinoderm, a balanid, a bryozoan, foraminifers and fish remains--both bones and scales. Approximately 100 species are present. Probably new species of Nerita, Colus (Anomalosipho), Marginella, and Arene occur in the formation. Other forms in the Castaic formation previously unreported or rarely reported from upper Miocene sediments of California are Anadara (Anadara) trilineata trilineata, Glycymeris cf. G. branneri, Chlamys hodgei, Sondylus sp., Eucrassatella cf. E. subgibbosa, Pseudochama sp., Corbula luteola, Periploma cf. P. discus, Tegula gallina, Calyptraea (Trochita) sp., Polinices (P.) uber, Oliva spicata. The fauna is comparable to that found at present at the northern limit of the recent Panamic molluscan province. All the near-shore assemblages in the formation are essentially contemporaneous. Faunal variations are indicative of environmental variations which existed along the margin of the basin. In particular, the geographic distribution of the fauna can be related to differences in water depth and substrate, and to environmental differences associated with bay versus open coast habitat. Analysis of the fauna suggests that: oxygen content of the water was probably less than normal at the bottom of the basin; the water was of normal marine salinity; most of the megafauna and associated sediments were deposited in water less than 25 fathoms deep; the maximum depth of the sill which probably existed along the western margin of the basin was about 100 fathoms; the marine climate was apparently uniform geographically within the basin of deposition and was much like that found at present off the southwest coast of Baja California. The average maximum annual surface temperature was about 22°C. The average minimum annual surface temperature was about 19°C. The Castaic formation was deposited at the northern limit of the late Miocene Panamic molluscan province. Bathymetric distribution as well as geographic distribution of a fauna must be considered in any paleozoogeographic study in order to determine paleoclimate.
Article
Thesis (M.A. in Paleontology)--University of California, Berkeley, Jan. 1950. "Literature cited": leaves 59-60. Microfilm. s
Article
Thesis (M.S.)--University of California, Davis.
Article
Thesis (M.S.)--University of California, Davis. Includes bibliographical references (p. 46-51).
Article
Thesis (M.A. in Paleontology)--University of California, Berkeley, Sept. 1957. Includes bibliographical references (leaves 104-112).
Article
Thesis (Ph. D.)--University of California, Los Angeles, 1969. Includes bibliographical references (leaves 328-344). Microform of typescript. -- Ann Arbor, Mich. : University Microfilms, 1970. -- 1 microfilm reel ; 35 mm.