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e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 1
The wild bees (Hymenoptera, Apoidea)
of the island of Cyprus
Androulla I. Varnava1, Stuart P.M. Roberts2, Denis Michez3, John S. Ascher4,
eodora Petanidou5, Stavroula Dimitriou5, Jelle Devalez5, Marilena Pittara1,
Menelaos C. Stavrinides1
1 Department of Agricultural Sciences, Biotechnology and Food Science, Cyprus University of Technology, Arch.
Kyprianos 30, Limassol, 3036, Cyprus 2 CAER, School of Agriculture, Policy and Development, e University
of Reading, Reading, UK 3 Research Institute of Bioscience, Laboratory of Zoology, University of Mons, Place
du parc 23, 7000 Mons, Belgium 4 Department of Biological Sciences, National University of Singapore, 14
Science Drive 4, Singapore 117543, Singapore 5 Laboratory of Biogeography & Ecology, Department of Geo-
graphy, University of the Aegean, 81100 Mytilene, Greece
Corresponding author: Androulla I. Varnava (androulla.varnava@cut.ac.cy);
Menelaos C. Stavrinides (m.stavrinides@cut.ac.cy)
Academic editor: Michael S. Engel|Received 18 July 2019|Accepted 25 November 2019|Published 6 April 2020
http://zoobank.org/596BC426-C55A-40F5-9475-0934D8A19095
Citation: Varnava AI, Roberts SPM, Michez D, Ascher JS, Petanidou T, Dimitriou S, Devalez J, Pittara M, Stavrinides
MC (2020) e wild bees (Hymenoptera, Apoidea) of the island of Cyprus. ZooKeys 924: 1–114. https://doi.
org/10.3897/zookeys.924.38328
Abstract
Cyprus, the third largest island in the Mediterranean, constitutes a biodiversity hotspot with high rates
of plant endemism. e wild bees of the island were studied extensively by the native George Mavro-
moustakis, a world-renowned bee taxonomist, who collected extensively on the island from 1916 to 1957
and summarised his results in a series of eight Cyprus-specic papers published from 1949 [“1948”] to
1957. e current work represents the rst modern checklist of the wild bees of Cyprus, based on a
compilation of previous publications, museum specimens and authors’ recent collections. Overall, 369
veried wild bee species have been recorded on the island, with eleven species reported from Cyprus for
the rst time. e island hosts all six of the globally widespread bee families, with Apidae represented
by 110 species, Megachilidae with 91, Andrenidae with 76, Halictidae with 72, Colletidae with 19, and
Melittidae with1. Twenty-one of the recorded bee species are endemic (i.e., 5.7% endemism rate) and
ZooKeys 924: 1–114 (2020)
doi: 10.3897/zookeys.924.38328
http://zookeys.pensoft.net
Copyright Androulla I. Varnava et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
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RESEARCH ARTICLE
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
2
Cyprus ranks third after Lesvos and Sicily in known bee species richness among the Mediterranean is-
lands. Previously unpublished records from various locations on Cyprus for 156 previously reported bee
species are also provided in the study. e current work provides a baseline for future studies of wild bee
diversity on the island of Cyprus and neighbouring regions.
Keywords
Bee species richness, biodiversity conservation, pollination, wild bees
Introduction
Cyprus, the third largest island in the Mediterranean, is situated in the eastern part of
the basin, in the active tectonic zone between the African and the Eurasian plates. e
island is a global biodiversity hotspot (Myers et al. 2000) characterised by high rates
of plant endemism (Christodes 2017). Cyprus is an ideal place for the study of wild
bees, because there is a wealth of past information on the diversity of species, a result of
the considerable work of George Mavromoustakis, a world-renowned bee taxonomist
native to the island.
e rst list of the bee fauna of Cyprus was published by Pittioni (1950) who
reported the results of a 1939 general scientic expedition on the island by Harald
Lindberg, one of the most distinguished botanists in Finland in the 20th century, and
his two sons, Hakan and Par Harald. e most detailed records on the bee fauna of the
island were compiled by Mavromoustakis in a series of eight papers that summarised
his Cyprus records which included 237 currently valid species (Mavromoustakis 1949
[“1948”], 1951, 1952, 1953, 1954, 1955, 1957a, 1958 [“1957”]). e rst paper
in the series is commonly cited according to its imprint date of 1948, but its date of
publication as established for nomenclatural purposes by Evenhuis (2003) is 14 Janu-
ary 1949. Some other Mavromoustakis papers also have ICZN dates postdating their
imprint dates as notes. Mavromoustakis’ data included in his eight Cyprus-specic
papers are currently available in an accessible online database (Varnava and Stavrinides
2015), which includes data on the location, month and plant species on which bees
were collected. e wild bee species had been reported by Mavromoustakis to visit 177
dierent species of plants, of which 13 are endemic (Varnava and Stavrinides 2015).
Georghiou (1977) listed all the bees reported on the island by Pittioni (1950), as well
as some species in the collections of the Department of Agriculture, virtually all of
which were collected and identied by Mavromoustakis.
Recent work on selected genera of wild bees of Cyprus was carried out by Ebmer
(2014). Ascher and Pickering’s (2018) online compilation lists 335 species of wild bees
on the island, while Kuhlmann et al. (2015) report 305 species. e aim of the current
work is to provide the rst comprehensive checklist of the bees of Cyprus, including all
past information on authors’ reports for each species, enriched with lists of new records
and species collected by the authors of the current study.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 3
Materials and methods
Study area
e geological history of Cyprus comprises a series of complex geological processes
that started approximately 90 million years ago (mya), when the subduction of the
African plate beneath the Eurasian created new oceanic crust; this was later cut o and
uplifted to create the Troodos range, in the centre of the island, with a peak at 1,951
m (Wagstae 2016). e uplifting of limestone depositions created the Pendadaktylos
range in the north of the country, with a maximum elevation of 1,024 m. e current
form of the island took shape approximately two mya.
Pollen evidence suggests that early Holocene Cyprus was covered by dense stands
of typical Mediterranean trees and shrubs, such as Ceratonia siliqua, Quercus coccifera,
Quercus infectoria, Laurus nobilis, Olea europaea, Cupressus sempervirens and species in
the genus Juniperus (Delipetrou et al. 2008). Noteworthy extinct species of animals
that lived on the island during the Pleistocene until around 11,000 years BP include
the Cyprus dwarf elephant (Elephas cypriotes) and Cyprus pygmy hippopotamus (Pha-
nourios minutus), which probably arrived on the island by swimming from the main-
land (Nicolaou et al. 2016).
e current land surface area of Cyprus is 9,251 km2, of which 42 % is covered
with forest/shrubland, and 23 % is devoted to agriculture (Vogiatzakis et al. 2016).
ere are 52 dierent habitat types listed in the EU Habitats Directive (92/43/EC),
with ve of them being exclusive to the island. e island is very rich in plant diversity
with more than 1,900 plant species, of which 130 are endemic (Christodes 2017).
Most of the endemic species of plants occur on the Troodos and Pentadaktylos moun-
tain ranges.
ere are 29 species of terrestrial mammals recorded on Cyprus, with bats rep-
resented by 19 species (Nicolaou et al. 2016). ere are nine species of snakes, 11
species of lizards (one endemic species) (Sparrow and Baier 2016a), three species
of native turtles (Sparrow and Baier 2016a), and three species of amphibians (one
endemic) (Sparrow and Baier 2016b). Among the 412 bird species that have been
recorded on Cyprus, 57 are resident breeders with two endemic species that breed
nowhere else: the Cyprus wheatear (Oenanthe cypriaca) and the Cyprus warbler (Syl-
via melanothorax), with an additional of four endemic subspecies (Charalambidou
et al. 2016).
Insects in Cyprus represent a highly diverse group with more than 5,000 re-
corded species according to Fauna Europaea (2012), with the number of species per
order varying in more recent accounts. Coleoptera is the most species-rich order,
followed by Hymenoptera and Lepidoptera. Well-studied groups include Odonata
with 37 species (Sparrow et al. 2016), Orthoptera with 71 species of which 12 are
endemic (Siedle et al. 2016), and butteries with 49 species of which three are en-
demic (John 2016).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
4
Methodology
e updated checklist was based on published records of wild bees and species present
in authors’ personal collections and in those of numerous correspondents. e earliest
published descriptions based on Cyprus specimens date to 1870 (Dours, 1870) and
1910 (Cockerell, 1910), however these include specimens collected earlier, as it was
very common for ornithologists or sometimes palaeontologists working for European
museums to collect dierent taxa during their expeditions and provide them to experts
for identication.
We used as a starting point for this survey a compilation of species occurrence
records available for the island by J.S Ascher, accessible online through the website
Discover Life (Ascher and Pickering 2018) some of which had associated specimens re-
cords captured through the project on the collaborative databasing of North American
bee collections within a global informatics network (Ascher 2016). Each species in this
initial list was evaluated by the experts, and species were retained in the checklist only
if a primary paper based on examined specimens conrmed the presence of the species
on the island. In addition, we retained in the checklist species for which a reference to
a specic museum specimen was available.
Furthermore, a review of the literature known to the authors to cover Cyprus wild
bee species was made to identify additional species recorded from the island. In gen-
eral, we avoided redundant citation of distributional listings in global or regional com-
pilations (e.g., Ascher and Pickering 2018, Kuhlmann et al. 2015) or other secondary
sources, such as Gusenleitner and Schwarz (2002) and Müller (2018). However, for
some species for which no records for Cyprus existed in primary sources and which the
authors of the study considered valid for Cyprus, we cited secondary sources.
Published works covering the wild bees of Cyprus as known to the authors were
listed in chronological order under each species referenced. Georghiou (1977) was
used as a reference only for species that were collected/examined in the Mavromousta-
kis collection during the Georghiou survey, but not for species whose presence in Cy-
prus was based on other published works (notably Pittioni, 1950).
Experts’ samplings were carried out at dierent sites on the island (Fig. 1) by hand-
netting and pan trapping. Sampling locations were concentrated on the southern part
of the island, where the Republic of Cyprus exercises eective control (see Fig. 2A for
an example of a sampling location). Hand-netting focused on collecting specimens
during their visits to owering plants (e.g., Fig 2B-D). For each species from the per-
sonal collection of authors we provide information on the collector (leg.), expert who
identied the specimen (det.), locality, collection day, number of individuals of each
sex and the collection specic number (museum collections/authors collections) when
available. A list of museum acronyms is available in Suppl. material 1, Table S1.
For species for which Cyprus is the type locality, we provide details on the type
locality, status, repository, and collection event information (i.e., collector and date of
collection), when available, extracted from a compilation of type data for world bees
(J. S. Ascher, unpublished). For taxa described from Cyprus, we note the name under
which the specimen was described from the island, to highlight the potentially inter-
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 5
Figure 1. Map of Cyprus indicating locations of type specimens (for both valid and synonymised taxa)
and unpublished records. See Supplem. material 1, Table S3, S4 for the geographic coordinates. e new
reports of bees are shown with black dots and type specimen locations with grey dots.
esting variation within the taxon. In addition, collection localities are provided for all
specimens from the Mavromoustakis collection. Virtually all GPS coordinates for type
specimens and specimens from the Mavromoustakis collection represent the centre
of the administrative region of the village / city from where samples were collected,
as no GPS data were available at the time of collection. For the unpublished records,
the horizontal distance (30 m) from the given decimal latitude and decimal longitude
describe the smallest circle containing the whole of the location.
e global distribution of each species is reported based on IUCN (2019) with
additional data from Ascher and Pickering (2018) in cases where IUCN data did not
list all continents from which a species was reported. Ascher and Pickering (2018)
data were used to include additional continents in the distribution but not to amend
the distribution data at country level for continents reported in IUCN (2019). We
recognise that species distribution is dynamic, and therefore there might be dierences
between the actual distribution and the one described in the current study. Countries
are grouped into geographic regions based on a United Nations categorisation (Unit-
ed Nations 2017 - ST/ESA/SER.A/408 – see Suppl. material 1, Table S2). Specic
countries are listed when the species is present in less than four countries within the
geographic region. e nomenclature in the present work follows Nieto et al. (2014)
for species names, Michener (2007) for most supra-specic taxa, and Dorchin et al.
(2018) for the tribe Eucerini.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
6
Results
e present checklist for the Cyprus wild bee fauna includes a total of 369 species with
conrmed records, 21 of which are endemic to Cyprus. Eleven species are reported
for the rst time from the island: Andrena (Taeniandrena) leucopsis Warncke, 1967, A.
(Chlorandrena) gordia Warncke, 1975, Anthophora (Pyganthophora) dalmatica Pérez,
1902, Colletes creticus Noskiewicz, 1936, Eucera (Hetereucera) aequata Vachal, 1907, E.
(Eucera) palaestinae Friese, 1922, E. (Eucera) sulamita Vachal, 1907, Megachile (Eutri-
charaea) inexspectata Rebmann, 1968, yreus picaron Lieftinck, 1968, Sphecodes ephip-
pius (Linnaeus, 1767) and S. pseudofasciatus Blüthgen, 1925.
Additionally, unpublished records for 156 species of bees previously reported on
the island are provided. e 369 wild bee species are classied into six families as
follows: Apidae 110, Megachilidae 91, Andrenidae 76, Halictidae 72, Colletidae 19,
and Melittidae 1 species. Most of the 21 endemic species belong to the Andrenidae
and Apidae families. We also provide a list of 29 additional species (see relevant sec-
Figure 2. A Landscape with diverse bee fauna in Cyprus (Yermasoyia Dam area) B Melitturga syriaca
foraging on Trifolium clypeatum C Eucera dimidiata foraging on Anchusa undulata subsp. hybrida D Meg-
achile cypricola foraging on Onobrychis venosa. Pictures provided by Nicolas J. Vereecken (A, B, C ) and
Androulla Varnava (D).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 7
tion of Supplementary Material) for which there is high uncertainty for their presence
on the island, even though they had been reported as present in Cyprus in the past.
A total of 101 species or subspecies were described from Cyprus (80 of which by
G. Mavromoustakis) from 44 distinct locations (Fig. 1). Currently, 46 are valid at the
species, and 18 at the subspecies level (Suppl. material 1, Table S3). Most type loca-
tions are concentrated on the southern part of the island, in the District of Limassol,
where Mavromoustakis resided and recent collections were carried out. Most of the
type insects from Cyprus (52) are located at the Department of Agriculture collection
in Nicosia (Cyprus Department of Agriculture, 1989).
Updated checklist of the bees of Cyprus
MELITTIDAE
DASYPODAINI
Genus Dasypoda LATREILLE, 1802
1 species.
Dasypoda (Megadasypoda) suripes (Christ, 1791)
References. Michez et al. (2004).
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Albania, It-
aly), Northern Europe (Lithuania), Eastern Europe, Western Asia (Turkey, Armenia).
Notes. D. suripes has been considered by some authors to be a nomen dubium and
therefore the species has been cited in some sources as D. aurata Rudow, 1881.
ANDRENIDAE
ANDRENINI
Genus Andrena FABRICIUS, 1775
73 species.
Andrena (Aciandrena) aciculata Morawitz, 1886
References. Scheuchl and Willner (2016).
Material examined. Limassol District: Polemidia, 34.71178°N, 33.004775°E,
8.III.2017, (1♀); Sovereign Base Area, Akrotiri, Bishop's Pool, 34.597305°N,
32.984521°E, 10.III.2017, (1♀); Sovereign Base Area, Akrotiri, 34.600657°N,
32.971419°E, 10.III.2017, (2♀); Anogyra to Pachna Road km 4, 34.764269°N,
32.757736°E, 5.V.2015, (1♀), all records S.P.M Roberts leg., B. Tomozei det.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
8
Distribution. Cyprus, Western Europe (Austria), Southern Europe (North Mac-
edonia, Greece), Eastern Europe, Western Asia (Turkey), Southern Asia (Iran).
Andrena (Suandrena) aegypticola Friese, 1922
References. Mavromoustakis (1954); Warncke (1967).
Mavromoustakis localities. Famagusta, Larnaca.
Distribution. Cyprus, Northern Africa (Libya), Western Asia.
Notes. Described from Cyprus as Andrena larnacensis Mavromoustakis 1954, from
Famagusta: 35.125°N, 33.941667°E, 15.II.1951, G.A. Mavromoustakis leg., G.A.
Mavromoustakis det. ♀, (DAAN).
Andrena (Aenandrena) aeneiventris Morawitz, 1872
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1957a, b); Georghiou (1977); Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol, Polemedia Hills, Zakaki, Asomatos, Fas-
souri, Yermasoyia river, Salamis, Near Pano Kivides, Kaloiri Hills near Yermasoyia
River, Morphou.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa, Central Asia, Western Asia.
Andrena (Melandrena) albopunctata (Rossi, 1792)
References. Gusenleitner and Schwarz (2002).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa, Western Asia, Central Asia, Southern Asia (Iran, Afghanistan,
Pakistan).
Andrena (Micrandrena) alfkenelloides Warncke, 1965
References. Gusenleitner and Schwarz (2002).
Material examined. Limassol District: 1 km E of Pissouri, 34.677579°N,
32.722066°E, 27.IV.2015, (3♀, 1♂), collecting pollen on Brassicaceae; 0.5 km E of
Vasa, 34.831384°N, 32.79744°E, 29.IV.2015, (1♀), visiting Apiaceae; Anogyra to
Avdimou Road km 2, 34.723986°N 32.736892°E, 3.V.2015, (21♀), collecting pollen
and visiting Apiaceae; 0.7 km N of Anogyra, 34.745537°N, 32.73385°E, 3.V.2015,
(14♀, 1♂), collecting pollen and visiting Apiaceae; Anogyra to Pachna Road km 4,
34.764269°N, 32.757736°E, 5.V.2015, (3♀), collecting pollen at Apiaceae, all records
S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia (Turkey).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 9
Andrena (Chlorandrena) astica Warncke, 1967
References. Gusenleitner and Schwarz (2002); Schwenninger (2015).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey).
Andrena (Euandrena) bicolor Fabricius, 1775
References. Warncke (1965, 1975); Scheuchl & Willner (2016).
Material examined. Limassol District: Chionistra, 34.9317°N, 32.8664°E,
14.V.2012, 16.V.2012, S. Dimitriou leg., E. Scheuchl det. (1♂, 1♀), pan trap (UAEG).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Alge-
ria, Tunisia), Western Asia (Turkey, Israel), Central Asia (Kazakhstan), Southern Asia
(Iran), Eastern Asia (China).
Notes. Described from Cyprus as Andrena bicolor ssp. apricaria Warncke 1975,
from Limassol: 34.66839°N, 33.03252°E, II.1939, K. Warncke leg./det. ♀, (OLML).
Andrena (Cryptandrena) brumanensis Friese, 1899
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952).
Material examined. Limassol District: 0.7 km N of Anogyra, 34.745537°N,
32.73385°E, 3.V.2015, (1♀), collecting pollen on Apiaceae; Yermasoyia Dam,
34.755799°N, 33.096194°E, 7.III.2017, all records S.P.M. Roberts leg., B. Tomozei
det. (10♀).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Hungary, Romania, Slovakia), Western Asia (Turkey).
Andrena (Truncandrena) caneae Strand, 1915
References. Pittioni (1950); Mavromoustakis (1953, 1957a); Warncke (1967).
Mavromoustakis localities. Limassol, Amathus, Yerasa, Polemedia, Famagusta.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey).
Notes. Described from Cyprus as Andrena mavromoustakisi Pittioni, 1950, from
Geroskipou, E of Paphos: 34.76666°N, 32.46666°E, 8.V.1946, G.A. Mavromoustakis
leg., B. Pittioni det., (MZHF).
Andrena (Micrandrena) cervina Warncke, 1975
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 10.IV.1967,
G.A. Mavromoustakis leg., K. Warncke det. ♀, (OLML).
References. Warncke (1975).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
10
Material examined. Limassol District: 2 km N of Anogyra, 34.748126°N,
32.732248°E, 1.V.2015, (1♀); Famagusta District: Cape Greco, 34.963264°N,
34.066211°E, 15.III.2017, (3♂), all records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus. ENDEMIC.
Andrena (Aenandrena) chaetogastra Pittioni, 1950
Type locality–country. Cyprus, Mt. Troodos, Chionistra: 34.9364°N, 32.8636°E,
17.VI.1939, H. Lindberg leg., B. Pittioni det. ♀, (MZHF).
References. Pittioni (1950).
Distribution. Cyprus, Western Asia (Israel).
Andrena (Chlorandrena) cinereophila Warncke, 1965
References. Warncke (1965).
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Eastern Eu-
rope (Romania, Bulgaria), Western Asia (Israel), Southern Asia (Afghanistan).
Andrena (Brachyandrena) colletiformis Morawitz, 1874
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952, 1957a); Warncke (1967).
Mavromoustakis localities. Limassol, Cherkes, Yermasoyia River, Amathus, Sa-
lamis, Larnaca.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa (Morocco, Tunisia, Algeria), Western Asia (Turkey, Israel), Cen-
tral Asia (Turkmenistan, Kazakhstan), Southern Asia (Iran).
Notes. Described from Cyprus as Andrena colletiformis insulana Pittioni, 1950,
from Kouklia: 34.6978°N, 32.592°E, 26.VI.1939, H. Lindberg leg., B. Pittioni det.,
(MZHF).
Andrena (Simandrena) combinata (Christ, 1791)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1954, 1958 ["1957"]);
Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol, Cherkes, Yermasoyia River, Amathus, Ye-
rasa, Lania, Near Paramytha, Northern Mountains Kantara, Kellaki, Morphou (West
Mesaoria plains), Larnaca.
Material examined. Limassol District: Makria Kontarka, 34.9095°N, 32.8971°E,
31.V.2012, 2.VI.2012 (1♀), pan trap (UAEG), all records S. Dimitriou leg., E.
Scheuchl det.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Algeria), Western
Asia, Central Asia, Southern Asia (Iran), Eastern Asia (China).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 11
Andrena (Chlorandrena) crepidis Schwenninger, 2015
Type locality–country. Cyprus, Limassol District, 4 km SW Kaminaria: 34.904°N,
32.761°E, 24.III.2013, H.R. Schwenninger leg., H.R. Schwenninger det. ♂, (SMNS).
References. Schwenninger (2015).
Material examined. Limassol District: Kaminaria, 34.93°N, 32.78°E, 22.III.2013,
(1♂); Kaminaria, 34.93°N, 32.78°E, 24.III.2013, (1♂); 4 km SW Kaminaria,
24.III.2013, (1♂); 5 km SW Kaminaria, 24.III.2013, (1♀, 1♂); Kaminaria, 34.93°N,
32.78°E, 2.IV.2009 (2♂); Amiantos, 34.918°N, 32.9472°E, 5-7.IV.2012 (1♂), pan
trap (UAEG); Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 5-7.IV.2012, (3♂),
pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 25-27.IV.2012, (1♀), pan trap
(UAEG), all Kaminaria records H.R. Schwenninger leg., H.R. Schwenninger det., all
Amiantos records S. Dimitriou leg., E. Scheuchl det., all Kakopetria records S. Dimi-
triou leg., E. Scheuchl and J. Devalez det.
Distribution. Cyprus. ENDEMIC.
Andrena (Notandrena) curvana Warncke, 1965
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (Austria), Southern Europe, Eastern Europe.
Andrena (Cordandrena) cypria Pittioni, 1950
Type locality–country. Cyprus, Mt. Troodos, Chionistra: 34.9364°N, 32.8636°E,
17.VI.1939, H. Lindberg leg., B. Pittioni det. ♀, (MZHF).
References. Pittioni (1950); Mavromoustakis (1953).
Mavromoustakis localities. Mt. Troodos.
Distribution. Cyprus, Western Asia, Southern Asia (Iran).
Andrena (Plastandrena) cypricola Mavromoustakis, 1952
Type locality–country. Cyprus, Kato Amiandos, 762–914m: 34.9059°N, 32.9431°E,
4.IV.1946, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Pittioni (1950); Mavromoustakis (1952, 1953, 1954); Warncke
(1967).
Mavromoustakis localities. Pera Pedi, Yermasoyia River, Trimiklini, Yerasa, Saet-
tas, Ayia Phyla, Amyrou Monastery-near Apsiou, Amathus, Rotsou spring-near Par-
amytha, Kato Amiandos, Mt. Troodos.
Material examined. Nicosia District: Linou, 35.0755°N, 32.9164°E, 5-7.IV.2012,
(1♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 5-7.IV.2012, (1♀), pan
trap (UAEG), all records S. Dimitriou leg., E. Scheuchl det.
Distribution. Cyprus. ENDEMIC.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
12
Andrena (Micrandrena) dargia Warncke, 1965
References. Gusenleitner and Schwarz (2002).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey).
Andrena (Holandrena) decipiens Schenck, 1861
References. Mavromoustakis (1949 [“1948”]); Warncke (1965); Scheuchl and Will-
ner (2016).
Mavromoustakis localities. Limassol, Fassouri, Amathus, Akrounda, Nicosia.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa, Western Asia (Turkey), Southern Asia (Iran).
Andrena (Melandrena) elmaria Gusenleitner, 1998
References. Gusenleitner (1998).
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (1♂); Sovereign Base Area Akrotiri, Bishop's Pool,
34.597305°N, 32.984521°E, 10.III.2017, (1♀, 1♂); Famagusta District: Cape Greco,
34.963264°N, 34.066211°E, 15.III.2017, (1♀), all records S.P.M. Roberts leg., B.
Tomozei det.
Distribution. Cyprus, Western Asia, Southern Asia (Iran).
Andrena (Parandrenella) gurata Morawitz, 1866
References. Gusenleitner and Schwarz (2002).
Distribution. Cyprus, Southern Europe, Eastern Europe, Western Asia (Turkey,
Georgia, Azerbaijan), Southern Asia (Iran).
Andrena avilabris Schenck, 1874
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (France, Switzerland, Germany), Southern
Europe (Spain), Eastern Europe (Ukraine), Western Asia.
Andrena (Zonandrena) avipes Panzer, 1799
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1953, 1954, 1957a); Georghiou (1977); Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Zakaki, Fas-
souri, Trimiklini, Eftagonia, Pissouri, Yerasa, Lania, Platania Forest Station, Fasoula,
Northern Mountains Kantara, Larnaca, sand dunes near Amathus.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 13
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 5-7.
IV.2012, (3♂, 2♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 14-
16.V.2012, (2♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 31.V.2012,
2.VI.2012, (1♀), pan trap (UAEG); Limassol District: Amiantos, 34.918°N, 32.9472°E,
5-7.IV.2012, (12♂, 3♀), pan trap (UAEG); Amiantos, 34.918°N, 32.9472°E, 14-
16.V.2012, (1♂, 1♀), pan trap (UAEG); Amiantos, 34.918°N, 32.9472°E, 31.V.2012,
2.VI.2012, (1♀), pan trap (UAEG); Makria Kontarka, 34.9095°N, 32.8971°E, 14-
16.V.2012, (31♂, 3♀), pan trap (UAEG); Makria Kontarka, 34.9095°N, 32.8971°E,
31.V.2012, 2.VI.2012 (4♂, 17♀), pan trap (UAEG); Almirolivado, 34.9333°N,
32.9004°E, 14-16.V.2012, (58♂, 13♀), pan trap (UAEG); Almirolivado, 34.9333°N,
32.9004°E, 31.V.2012, 2.VI.2012, (1♂, 12♀), pan trap (UAEG); Troodos, Chionistra,
34.9317°N, 32.8664°E, 14-16.V.2012, (77♂, 1♀), pan trap (UAEG); Troodos, Chion-
istra, 34.9317°N, 32.8664°E, 31.V.2012, 2.VI.2012, (8♂, 40♀), pan trap (UAEG);
Anogyra to Pachna Road km 4, 34.764269°N, 32.757736°E, 5.V.2015, (1♀); Ye-
rmasoyia Dam, 34.755799°N, 33.096194°E, 7.III.2017, (1♀, 1♂); Paphos District:
Nr Arminou Reservoir, 34.883435°N, 32.750988°E, 29.IV.2015, (3♀), collecting
pollen on Brassicaceae; 2.7 km SW of Acheleia, Potamos tis Ezouzas, 34.729004°N,
32.457544°E, 30.IV.2015, (1♀, 1♂), all Kakopetria, Amiantos, Makria Kontarka,
Almirolivado and Troodos-Chionistra records S. Dimitriou leg., J. Devalez det., all Ano-
gyra, Yermasoyia Dam and Paphos district records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (India, Nepal), Eastern Asia (China).
Andrena (Holandrena) forsterella Osytshnjuk, 1978
References. Schönitzer et al (1995); Gusenleitner and Schwarz (2002).
Distribution. Cyprus, Southern Europe, Eastern Europe (Bulgaria), Western Asia,
Southern Asia (Iran).
Notes. In older literature often not separated from A. variabilis.
Andrena (Melanapis) fuscosa Erichson, 1835
References. Cockerell (1910); Mavromoustakis (1951, 1952, 1954); Warncke (1967);
Scheuchl and Willner (2016).
Mavromoustakis localities. Episkopi, Zakaki, Cherkes, Fassouri, Famagusta, Larnaca.
Material examined. Famagusta District: Cape Greco, 34.963264°N, 34.066211°E,
15.III.2017, S.P.M. Roberts leg., B. Tomozei det. (7♂).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia Southern Asia (India).
Notes. Described from Cyprus as Andrena cyprica Cockerell 1910, from Nico-
sia: 35.166667°N, 33.366667°E, 17.III, C. Glaszner leg., T.D.A. Cockerell det. ♂,
(NHMUK).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
14
Andrena (Ptilandrena) glidia Warncke, 1965
References. Warncke (1974); Gusenleitner and Schwarz (2002).
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 5-7.
IV.2012, (1♀), pan trap (UAEG); Limassol District: Almirolivado, 34.9333°N,
32.9004°E, 14-16.V.2012, (2♀), pan trap (UAEG); Troodos, Chionistra, 34.9317°N,
32.8664°E, 14-16.V.2012, (3♀), pan trap (UAEG); Troodos, Chionistra, 34.9317°N,
32.8664°E, 31.V.2012, 2.VI.2012, (3♀), pan trap (UAEG), all records S. Dimitriou
leg., E. Scheuchl det.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Israel).
Andrena (Chlorandrena) gordia Warncke, 1975
Material examined. Paphos District: Nr Arminou Reservoir, 34.883435°N,
32.750988°E, 29.IV.2015, (1♂), S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Western Asia (Turkey).
Andrena (Melandrena) grandilabris Pérez, 1903
References. Warncke (1969, 1974).
Distribution. Cyprus, Western Asia.
Andrena (Chrysandrena) hesperia Smith, 1853
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1957a); Scheuchl
and Willner (2016).
Mavromoustakis localities. Limassol, Akrotiri Forest, Fassouri, Amathus, Yer-
masoyia River, Yerasa, Omodos, Trimiklini, Mesayitonia-Fasoula, Amyrou Monastery
(near Apsiou), Nicosia, Larnaca, Famagusta.
Material examined. Limassol District: Amiantos, 34.918°N, 32.9472°E, 25-
27.IV.2012, (1♀), pan trap (UAEG); Sovereign Base Area, Akrotiri, 34.600657°N,
32.971419°E, 17.III.2017, (2♀), collecting pollen on Crepis sancta (Asteraceae); Sov-
ereign Base Area, Avdimou Bay Clis, 34.656698°N, 32.773339°E, 13.III.2017, (1♀),
all Amiantos records S. Dimitriou leg., E. Scheuchl det., all Akrotiri and Avdimou Bay
records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey, Israel), Central Asia.
Andrena (Margandrena) hyacinthina Mavromoustakis, 1958
Type locality–country. Cyprus, Yermasoyia River plain: 34.7182°N, 33.08788°E,
3.III.1952, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 15
References. Pittioni (1950); Mavromoustakis (1958); Warncke (1967).
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, S.P.M. Roberts leg., B. Tomozei det. (1♀).
Distribution. Cyprus, Western Asia (Turkey, Israel, Lebanon).
Andrena (Ptilandrena) kornosica Mavromoustakis, 1954
Type locality–country. Cyprus, Mt. Kornos (Northern Mountains), 762m:
35.1379°N, 33.1379°E, 3.III.1936, G.A. Mavromoustakis leg., G.A. Mavromoustakis
det. ♀, (NHMUK).
References. Pittioni (1950); Mavromoustakis (1954, 1957a); Warncke (1967).
Mavromoustakis localities. Mt. Kornos, Lania, Near Trooditissa.
Distribution. Cyprus. ENDEMIC.
Andrena (Aciandrena) lamiana Warncke, 1965
References. Warncke (1965).
Material examined. Limassol District: Anogyra, 34.741952°N, 32.734845°E,
3.V.2015, S.P.M. Roberts leg., B. Tomozei det. (1♀), collecting pollen on Brassicaceae.
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Eastern Eu-
rope (Bulgaria), Western Asia (Turkey, Syria), Southern Asia (Iran).
Andrena (Simandrena) lepida Schenck, 1861
References. Warncke (1965); Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey, Israel, Syria), Southern Asia (Iran).
Andrena (Taeniandrena) leucopsis Warncke, 1967
Material examined. Limassol District: Almirolivado, 34.9333°N, 32.9004°E, 14-
16.V.2012, S. Dimitriou leg., E. Scheuchl det. (2♀), pan trap (UAEG).
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Eastern Eu-
rope (Romania, Bulgaria), Western Asia (Turkey, Lebanon), Southern Asia (Iran).
Andrena (Poecilandrena) limassolica Mavromoustakis, 1949
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 19.I.1936, G.A.
Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1949 [“1948”], 1952); Warncke (1967).
Mavromoustakis localities. Limassol, Polemedia Hills.
Distribution. Cyprus, Western Asia (Jordan).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
16
Andrena (Melandrena) limata Smith, 1853
References. Cockerell (1910, 1914); Mavromoustakis (1953, 1954, 1957a); Warncke
(1967); Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol, Zakaki, Pera Pedi.
Material examined. Limassol District: Sovereign Base Area Akrotiri, Bishop's
Pool, 34.597305°N, 32.984521°E, 10.III.2017, (2♀, 1♂); Sovereign Base Area, Ak-
rotiri, 34.600657°N, 32.971419°E, 10.III.2017, (1♀), collecting pollen on Asterace-
ae, all records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa, Western Asia, Central Asia, Southern Asia (Iran).
Notes. Described from Cyprus as Andrena batesiae Cockerell, 1910, from Nicosia:
35.166667°N, 33.366667°E, D.M.A. Bate leg., T.D.A. Cockerell det. ♀, (NHMUK).
Andrena (Micrandrena) lindbergella Pittioni, 1950
Type locality–country. Cyprus, Mt. Troodos, Chionistra: 34.9364°N, 32.8636°E,
17.VI.1939, H. Lindberg leg., B. Pittioni det. ♀, (MZHF).
References. Pittioni (1950); Warncke (1967).
Distribution. Cyprus, Western Asia (Israel, Lebanon).
Andrena (Truncandrena) medeninensis Pérez, 1895
References. Warncke (1967, 1974).
Distribution. Cyprus, Southern Europe (Spain, Greece), Northern Africa, West-
ern Europe (Turkey).
Notes. Records pertain to subspecies usura Warncke, 1967.
Andrena (Simandrena) mehelyi Alfken, 1936
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (Austria), Southern Europe, Eastern Eu-
rope, Western Asia (Turkey), Southern Asia (Iran).
Andrena (Chrysandrena) merula Warncke, 1969
Type locality–country. Cyprus, Ayia Eirini: 34.9816°N, 32.9712°E, 20.IV.1939,
G.A. Mavromoustakis leg., K. Warncke det. ♂, (OLML).
References. Warncke (1969).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Israel),
Southern Asia (Afghanistan).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 17
Andrena (Cryptandrena) monacha Warncke, 1965
References. Warncke (1965).
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, S.P.M. Roberts leg., B. Tomozei det. (1♂).
Distribution. Cyprus, Southern Europe (Greece), Western Europe (Turkey, Leba-
non, Syria).
Andrena (Melandrena) morio Brullé, 1832
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951 1953, 1954, 1957a); Warncke (1967); Georghiou (1977); Scheuchl and Will-
ner (2016).
Mavromoustakis localities. Limassol, Pera Pedi, Trimiklini, Lania, Moni, Po-
lemedia, Zakaki, Cherkes, Fasoula, Amathus, Potamitissa, Famagusta, Ayia Varvara
(Stavrovouni).
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E,
31.V.2012, 2.VI.2012, (1♀), pan trap (UAEG); Polemidia, 34.71178°N, 33.004775°E,
8.III.2017, (1♀); Famagusta District: Cape Greco, 34.963264°N, 34.066211°E,
15.III.2017, (7♂), all Kakopetria records S. Dimitriou leg., J. Devalez det., all Po-
lemidia and Cape Greco records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe, Northern Africa, Western Asia, Central Asia.
Notes. Described from Cyprus as Andrena morio athalassae Pittioni, 1950, from
Limassol: 34.66839°N, 33.03252°E, 4-6.VI.1939, H. Lindberg leg., B. Pittioni det.,
(MZHF).
Andrena (Ulandrena) neocypriaca Mavromoustakis, 1956
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 16.IV.1949,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1956, 1957b); Warncke (1967).
Mavromoustakis localities. Near Limassol, Yermasoyia Hills.
Material examined. Limassol District: 2 km N of Anogyra, 34.748126°N,
32.732248°E, 1.V.2015, S.P.M. Roberts leg., B. Tomozei det. (1♀).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey).
Andrena (Melandrena) nigroaenea (Kirby, 1802)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1954, 1956); Georghiou
(1977); Scheuchl and Willner (2016).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
18
Mavromoustakis localities. Limassol, Polemedia Hills, Zakaki, Mesayitonia, Ap-
siou, Yermasoyia River, Amathus, Pissouri, Potamitissa, Trimiklini, Lania, Kitromili
near Polemedia, Famagusta, Near Kyperounta, Larnaca.
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 5-7.
IV.2012, (1♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 14-16.V.2012,
(8♀), pan trap (UAEG); Limassol District: Amiantos, 34.918°N, 32.9472°E, 14-
16.V.2012, (3♀), pan trap (UAEG); Almirolivado, 34.9333°N, 32.9004°E, 14-
16.V.2012, (3♀), pan trap (UAEG); Makria Kontarka, 34.9095°N, 32.8971°E, 14-
16.V.2012, (5♀), pan trap (UAEG); Troodos, Chionistra, 34.9317°N, 32.8664°E, 14-
16.V.2012, (3♀), pan trap (UAEG); Almirolivado, 34.9333°N, 32.9004°E, 31.V.2012,
2.VI.2012, (5♀), pan trap (UAEG), all records S. Dimitriou leg., J. Devalez det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Southern Asia (Afghanistan, Iran).
Andrena (Parandrenella) nisoria Warncke, 1969
References. Warncke (1974).
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, S.P.M. Roberts leg., B. Tomozei det. (1♂).
Distribution. Cyprus, Western Asia, Southern Asia (Iran).
Andrena (Plastandrena) oligotricha Mavromoustakis, 1952
Type locality–country. Cyprus, Pera Pedi, 609m: 34.859444°N, 32.876111°E,
22.V.1929, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1949 [“1948”], 1952, 1953, 1954, 1957a);
Warncke (1967).
Mavromoustakis localities. Limassol, Cherkes, Zakaki, Pera Pedi, Amathus, La-
nia, Kellaki, Ayia Varvara, Saettas, Potamitissa, Kato Amiantos, Ayios Kostantinos,
Yerasa, Apsiou, Fasoula, Ayia Phyla, Amathus, Kykkos, Yermasoyia River.
Material examined. Paphos District: Nr Arminou Reservoir, 34.883435°N,
32.750988°E, 29.IV.2015, (2♀, 1♂), collecting pollen on Brassicaceae; Limassol Dis-
trict: 2 km N of Anogyra, 34.748126°N, 32.732248°E, 1.V.2015, (1♀), collecting
pollen on Brassicaceae, all records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Western Europe, Southern Europe, Northern Africa (Mo-
rocco, Algeria, Tunisia).
Notes. ere has been considerable confusion over the true identity of this taxon.
E. Scheuchl (2017, in litt.) regards Andrena oligotricha Mavromoustakis as a good spe-
cies and that it replaces Andrena bimaculata in Cyprus. All previous records attributed
to A. bimaculata are here treated as A. oligotricha.
Andrena (Chlorandrena) orientana Warncke, 1965
References. Schwenninger (2015).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 19
Material examined. Limassol District: 2 km N of Anogyra, 34.748126°N,
32.732248°E, 1.V.2015, (1♀), collecting pollen on Asteraceae; Sovereign Base Area,
Akrotiri, 34.600657°N, 32.971419°E, 10.III.2017, (5♀), collecting pollen on Aster-
aceae, all records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Eastern Eu-
rope (Bulgaria, Ukraine), Western Asia (Turkey, Israel).
Andrena (Taeniandrena) ovatula (Kirby, 1802)
References. Mavromoustakis (1949 [“1948”], 1957b); Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol, Amathus, Yermasoyia River.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, S.P.M. Roberts leg., B. Tomozei det. (1♀, 1♂).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia (Is-
rael), Central Asia, Southern Asia (Iran, Afghanistan).
Andrena (Chlorandrena) panurgimorpha Mavromoustakis, 1957
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 16.IV.1949,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1957b, 1958).
Mavromoustakis localities. Yermasoyia Hills.
Material examined. Limassol District: Anogyra to Pachna Road km 4,
34.764269°N, 32.757736°E, 5.V.2015, S.P.M. Roberts leg., B. Tomozei det. (1♀, 2♂).
Distribution. Cyprus, Southern Europe (Italy, Greece), Eastern Europe (Ukraine),
Western Asia (Turkey, Israel), Southern Asia (Iran).
Andrena (Truncandrena) pareklisiae Mavromoustakis, 1957
Type locality–country. Cyprus, Lania: 34.82444°N, 32.920833°E, 11.IV.1953, G.A.
Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1957b); Warncke (1967).
Distribution. Cyprus. ENDEMIC.
Andrena (Ulandrena) polemediana Mavromoustakis, 1956
Type locality–country. Cyprus, Ayia Eirini (near Paramytha): 34.9816°N, 32.9712°E,
12.IV.1939, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1956, 1957b).
Mavromoustakis localities. Near Limassol.
Material examined. Limassol District: Anogyra, 34.741952°N, 32.734845°E,
3.V.2015, S.P.M. Roberts leg., B. Tomozei det. (1♀), visiting Asteraceae.
Distribution. Cyprus. ENDEMIC.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
20
Andrena (Melandrena) pyropygia Kriechbaumer, 1873
References. Mavromoustakis (1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Episkopi, Pano
Kivides, Eleousa Monastery, Karpasian Peninsula, Apostolos Varnavas (near Famagusta).
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Ukraine),
Western Asia (Turkey, Israel), Southern Asia (Iran).
Andrena (Poliandrena) pyrozonata Friese, 1921
References. Mavromoustakis (1958); Warncke (1967).
Distribution. Cyprus, Western Asia (Turkey).
Notes. Described from Cyprus as Andrena perapedica Mavromoustakis, 1958,
from Pera Pedi, 609 m: 34.859444°N, 32.876111°E, G.A. Mavromoustakis leg., G.A.
Mavromoustakis det. ♀, (DAAN).
Andrena (Truncandrena) paramythensis Mavromoustakis, 1957
References. Mavromoustakis (1957a); Warncke (1967).
Mavromoustakis localities. Near Paramytha, Yerasa, Apsiou, Fasoula, Lania, Pera
Pedi, Kellaki, Trimiklini.
Distribution. Cyprus, Western Asia (Turkey).
Notes. e subspecies described from Cyprus is Andrena rufomaculata paramythen-
sis Mavromoustakis, 1957, from Paramytha vicinity (Vrisi tou Rotsou): 34.757°N,
32.972°E, 11.III.1944, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀,
(DAAN).
Andrena (Troandrena) saettana Warncke, 1975
Type locality–country. Cyprus, Saittas: 34.8708333°N, 32.9166667°E, 2.V.1961,
G.A. Mavromoustakis leg., K. Warncke det. ♀, (OLML).
References. Warncke (1975).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Jordan).
Andrena (Margandrena) sibthorpi Mavromoustakis, 1952
Type locality–country. Cyprus, Polemedia Hills: 34.7134°N, 32.9812°E, 9.XII.1949,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1952); Warncke (1967).
Mavromoustakis localities. Polemedia Hills.
Distribution. Cyprus. ENDEMIC.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 21
Andrena (Taeniandrena) similis Smith, 1849
References. Warncke (1975); Scheuchl and Willner (2016).
Material examined. Paphos District: Nr Arminou Reservoir, 34.883435°N,
32.750988°E, 29.IV.2015, (1♀); Limassol District: 0.5 km E of Vasa, 34.831384°N,
32.79744°E, 29.IV.2015, (1♀), visiting Asteraceae; Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (1♀), all records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Widespread in Europe, Western Asia, Central Asia, South-
ern Asia (Iran, Afghanistan), Eastern Asia (China).
Notes. Described from Cyprus as Andrena ocreata ssp. cyprisina Warncke 1975,
from Limassol: 34.66839°N, 33.03252°E, 8.III.1959, G.A. Mavromoustakis leg., K.
Warncke det. ♀, (KW).
Andrena (Micrandrena) spreta Pérez, 1895
References. Warncke (1974, 1975).
Distribution. Cyprus, Western Europe, Southern Europe, Northern Africa, West-
ern Asia, Southern Asia (Iran, Afghanistan).
Notes. Cyprus records pertain to subspecies scirpacea Warncke, 1975 (type local-
ity: Turkey).
Andrena (Aciandrena) tenuiformis Pittioni, 1950
Type locality–country. Cyprus, Mt. Troodos, 1200–1952 m: 34.9364°N, 32.8636°E,
16-22.VI.1939, H. Lindberg leg., B. Pittioni det. ♀, (MZHF).
References. Pittioni (1950); Warncke (1967).
Distribution. Cyprus, Western Asia (Turkey, Israel).
Andrena (Simandrena) thomsonii Ducke, 1898
References. Gusenleitner and Schwarz (2002).
Distribution. Cyprus, Western Europe (France), Southern Europe, Western Asia
(Turkey), Southern Asia (Iran).
Andrena (Melandrena) thoracica (Fabricius, 1775)
References. Mavromoustakis (1949 [“1948”], 1953, 1954, 1957a, b); Georghiou
(1977); Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol, Zakaki, Episkopi, Yermasoyia River, Famagusta.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Algeria,
Tunisia), Western Asia, Central Asia, Southern Asia (Iran, Afghanistan), Eastern Asia
(China, Korea).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
22
Notes. Described from Cyprus as Andrena thoracica ssp. kotschyi Mavromoustakis
1953, from Limassol: 34.66839°N, 33.03252°E, 12.V.1930, G.A. Mavromoustakis
leg./det. ♀, (DAAN).
Andrena (Cordandrena) torda Warncke, 1965
References. Warncke (1965).
Material examined. Famagusta District: Cape Greco, 34.963264°N, 34.066211°E,
15.III.2017, S.P.M. Roberts leg., B. Tomozei det. (1♀).
Distribution. Cyprus, Southern Europe (Greece), Western Asia.
Andrena (Simandrena) transitoria Morawitz, 1871
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1954, 1957a).
Mavromoustakis localities. Limassol, Cherkes, Episkopi, Yermasoyia River, Ayia
Phyla, Apsiou, Amathus, Pissouri, Potamitissa, Kato Amiandos, Fasoula, Yerasa, Moni,
Trimiklini, Kellaki, Phinikaria River, Platania Forest.
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Eastern
Europe, Western Asia, Central Asia (Kyrgyzstan, Tajikistan, Turkmenistan), Southern
Asia (Afghanistan).
Andrena (Troandrena) troodica Warncke, 1975
References. Pittioni (1950).
Distribution. Cyprus, Western Asia (Turkey).
Andrena (Truncandrena) truncatilabris Morawitz, 1877
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1957a, b).
Mavromoustakis localities. Limassol, Cherkes, Apsiou, Fassouri, Yermasoy-
ia River, Amathus, Pissouri, Yerasa, Near Paramytha, Trimiklini, Morphou, Near
Famagusta.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (1♂, 2♀); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017,
(1♂, 1♀); Sovereign Base Area, Akrotiri, Bishop's Pool, 34.597305°N, 32.984521°E,
10.III.2017, (1♀); Paphos District: Nr Arminou Reservoir, 34.883435°N,
32.750988°E, 29.IV.2015, (1♂, 2♀), collecting pollen on Sinapis (Brassicaceae), all
records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa (Algeria, Libya), Western Asia (Israel, Lebanon, Syria), Central
Asia (Kazakhstan, Turkmenistan), Southern Asia (Iran).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 23
Andrena (Notandrena) ungeri Mavromoustakis, 1952
Type locality–country. Cyprus, Zakaki: 34.663951°N, 32.999785°E, 18.I.1949
[“1948”], G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♂, (DAAN).
References. Pittioni (1950); Mavromoustakis (1952).
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Eastern Eu-
rope (Hungary, Romania, Bulgaria), Northern Africa (Morocco), Western Asia (Turkey).
Andrena (Notandrena) urdula Warncke, 1965
References. Gusenleitner and Schwarz (2002).
Distribution. Cyprus, Southern Europe (Spain - only in the west, Greece), East-
ern Europe (Romania, Bulgaria), Northern Africa (South Morocco), Western Asia.
Andrena (Holandrena) variabilis Smith, 1853
References. Mavromoustakis (1949 [“1948”], 1951).
Mavromoustakis localities. Limassol, Cherkes, Yermasoyia River, Near Zakaki,
Near Enkomi of Famagusta.
Material examined. Limassol District: Almirolivado, 34.9333°N, 32.9004°E,
31.V.2012, 2.VI.2012, S. Dimitriou leg., E. Scheuchl det. (1♂), pan trap (UAEG);
Anogyra to Pachna Road km 4, 34.764269°N, 32.757736°E, 5.V.2015, S.P.M. Rob-
erts leg., B. Tomozei det. (1♀).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Algeria,
Tunisia), Western Asia, Central Asia (Kazakhstan, west Turkmenistan), Southern Asia
(Iran).
Andrena (Cryptandrena) ventricosa Dours, 1873
References. Mavromoustakis (1949 [“1948”], 1951, 1957a); Warncke (1975).
Mavromoustakis localities. Cherkes, Amathus, Pissouri, Pyrgos, Ayia Phyla,
Paramytha.
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe, Northern Africa (Morocco, Tunisia), Western Asia, Central Asia.
Notes. Described from Cyprus as Andrena ventricosa ridibundus Warncke 1975,
from Cherkes: 34.65°N, 32.975°E, 11.III.1951, G.A. Mavromoustakis leg., K.
Warncke det. ♀, (KW).
Andrena (Ptilandrena) vetula Lepeletier, 1841
References. Mavromoustakis (1949 [“1948”], 1951, 1954, 1958 ["1957"]); Georghiou
(1977).
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24
Mavromoustakis localities. Limassol, Fassouri, Yermasoyia River, Pera Pedi, Am-
athus, Near Famagusta.
Material examined. Limassol District: Anogyra, 34.73663°N, 32.732715°E,
26.IV.2015, (1♂, 1♀), collecting pollen on Sinapis (Brassicaceae); Anogyra,
34.741952°N, 32.734845°E, 3.V.2015, (3♀), collecting pollen on Sinapis (Brassi-
caceae); Anogyra to Avdimou Road km 2, 34.723986°N, 32.736892°E, 3.V.2015,
(1♀), collecting pollen on Sinapis (Brassicaceae); Anogyra to Pachna Road km 4,
34.764269°N, 32.757736°E, 5.V.2015, (1♀), collecting pollen on Brassicaceae; Sov-
ereign Base Area, Paramali Bay, 34.661805°N, 32.804261°E, 10.III.2017, (1♂), visit-
ing Sinapis alba (Brassicaceae); Paphos District: Nr Arminou Reservoir, 34.883435°N,
32.750988°E, 29.IV.2015, (1♂, 5♀), collecting pollen on Sinapis (Brassicaceae);
Famagusta District: Cape Greco, 34.963264°N, 34.066211°E, 15.III.2017, (2♂), all
records S.P.M. Roberts leg., B. Tomozei det.
Distribution. Cyprus, Western Europe (France), Southern Europe (Portugal,
Spain, Italy), Northern Africa, Western Asia, Central Asia (Turkmenistan), Southern
Asia (Iran).
Andrena (Holandrena) wilhelmi Schuberth, 1995
References. Schuberth (1995).
Distribution. Cyprus, Southern Europe (North Macedonia, Italy, Greece), East-
ern Europe (Ukraine), Western Asia (Turkey, Israel).
e only known Cyprus records are listed as Paratypes in Schuberth (1995): Li-
massol, III.1932, 1m J. Schuberth det.; Salamis (north of Famagusta), 23.III.1971,
K.M. Guichard leg., J. Schuberth det.
Andrena (Taeniandrena) wilkella (Kirby, 1802)
References. Warncke (1965); Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Western Asia, Central Asia, South-
ern Asia (India), Eastern Asia (China).
MELITTURGINI
Genus Melitturga LATREILLE, 1809
1 species.
Melitturga (Melitturga) syriaca Friese, 1899
References. Mavromoustakis (1949 [“1948”], 1951, 1957a).
Mavromoustakis localities. Near Limassol, Mesayitonia, Near Palodia.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 25
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (5♀); Paphos District: N of Elia Bridge, 34.900977°N,
32.776759°E, 29.IV.2015, (3♂), all records S.P.M. Roberts leg./det.
Distribution. Cyprus, Eastern Europe (Bulgaria), Western Asia.
PANURGINI
Genus Panurginus NYLANDER, 1848
2 species.
Panurginus lactipennis Friese, 1897
References. Pittioni (1950); Mavromoustakis (1952).
Mavromoustakis localities. Mt. Troodos.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Romania),
Western Asia (Turkey).
Panurginus turcomanicus Popov, 1936
References. Warncke (1972b).
Distribution. Cyprus, Eastern Europe (Ukraine), Western Asia (Turkey, Israel,
Azerbaijan), Central Asia.
HALICTIDAE
HALICTINI
Genus Halictus LATREILLE, 1804
14 species.
Halictus (Halictus) asperulus Pérez, 1895
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1953, 1957a); Ebmer (2014).
Mavromoustakis localities. Pera Pedi, Mt. Troodos, Platania Forest Station,
Ayia Varvara (Stavrovouni), Eftagonia, Trimiklini, Pyrga, Xerokolimbi Stream near
Trooditissa, Platres.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (1♀).
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe (Spain,
Italy, Albania), Eastern Europe, Western Asia (Israel), Southern Asia (Iran).
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26
Halictus (Halictus) brunnescens (Eversmann, 1852)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1957a,b); Ebmer (2014).
Mavromoustakis localities. Limassol, Polemedia Hills, Akrotiri village, Cherkes,
Episkopi, Yermasoyia River, Fassouri, Near Zakaki, Ayios Athanasios, Amathus, Eft-
agonia, Trimiklini, Mt. Troodos Kannoures Springs, Younaros of Zakaki, Pyrga (Larn-
aca), Near Ayios eodoros (Pitsillia), Larnaca.
Material examined. Nicosia District: Linou, 35.0755°N, 32.9164°E, 5-7.
IV.2012, (1♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 5-7.IV.2012,
(3♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 25-27.IV.2012, (1♀),
pan trap (UAEG); Limassol District: Amiantos, 34.918°N, 32.9472°E, 31.V.2012,
2.VI.2012, (2♀), pan trap (UAEG); Almirolivado, 34.9333°N, 32.9004°E, 16-18.
IX.2011, (1♀), pan trap (UAEG); SBA, 8 km S Limassol, Akrotiri (near Airbase),
34.60°N, 32.97°E, 20.VI.2013, (4♂); Paphos District: 15 km SE Paphos, Kouklia,
34.72°N, 32.55°E, 20.VI.2013, (10♂); 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (1♀, 5♂), all Linou, Kakopetria, Amiantos and Almirolivado
records S. Dimitriou leg., J. Devalez det., all Akrotiri and Paphos district records C.
Schmid-Egger leg., A.W. Ebmer det.
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Spain, Greece),
Eastern Europe, Northern Africa, Western Asia (Turkey), Central Asia, Southern Asia.
Halictus (Seladonia) cephalicus Morawitz, 1874
References. Mavromoustakis (1949 [“1948”]); Mavromoustakis (1951, 1952, 1957a);
Ebmer (2014).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Akrotiri Bay,
Moni, Yermasoyia River, Kathikas, Amathus, Near Eftagonia.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (20♀, 1♂); 20 km N Paphos, Kathikas, 34.90°N, 32.42°E,
20.VI.20153, (3♀); 6 km NE Polis, beach, 35.06°N, 32.46°E, 20.VI.2013, (2♀); 6 km
W Polis, botanical garden, 35.03°N, 32.37°E, 20.VI.2013, (1♀); Polis, 35.053539°N,
32.351197°E, 30.X.2016, (9♂), visiting Dittrichia viscosa (Asteraceae); 8 km N
Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (5♀, 1♂); Limas-
sol District: Troodos, Mt. Olympos, 34.93°N, 32.86°E, 20.VI.2013, (1♀); 1 km E of
Pissouri, 34.677579°N, 32.722066°E, 27.IV.2015, (1♀), visiting Brassicaceae; Ano-
gyra to Pachna Road km 4, 34.764269°N, 32.757736°E, 5.V.2015, (2♀), collecting
pollen on Brassicaceae; Sovereign Base Area, Akrotiri, 34.628771°N, 32.941031°E,
29.X.2016, (1♂, 1♀), visiting Dittrichia viscosa (Asteraceae), all Kouklia, Kathikas,
NE Polis, Polis botanical garden, Mavrokolympos Reservoir and Troodos, Mt. Olym-
pos records C. Schmid-Egger leg., A.W. Ebmer det., all Polis, Pissouri, Anogyra, and
Akrotiri records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Southern Europe, Eastern Europe, Western Asia, Southern
Asia (Iran).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 27
Halictus (Mucoreohalictus) cypricus Blüthgen, 1937
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 7.III.1931,
G.A. Mavromoustakis leg., P.A.V. Blüthgen det. ♀, (MFNB).
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”]); Pittioni (1950);
Mavromoustakis (1951, 1952); Georghiou (1977); Ebmer (2014).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Zakaki, Episko-
pi, Pera Pedi, Akrotiri Bay, Pissouri, Near Zakaki, Famagusta.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (1♂).
Distribution. Cyprus, Western Asia (Turkey, Israel), Southern Asia (Iran).
Halictus (Halictus) fatsensis Blüthgen, 1936
References. Ebmer (2014).
Distribution. Cyprus, Northern Africa (Egypt), Western Asia (Turkey, Israel, Syria).
Halictus (Halictus) graecus Blüthgen, 1933
References. Pesenko (2005).
Distribution. Cyprus, Southern Europe (Greece, Croatia), Eastern Europe (Bul-
garia, Ukraine), Western Asia (Turkey).
Halictus (Halictus) nicosiae Blüthgen, 1923
Type locality–country. Cyprus, Nicosia: 35.166667°N, 33.366667°E, Staudinger
leg., P. A.V. Blüthgen det.
References. Blüthgen (1923); Mavromoustakis (1949 [“1948”]); Pittioni (1950);
Mavromoustakis (1951, 1952, 1953, 1954); Ebmer (2014).
Mavromoustakis localities. Polemedia Hills, Apsiou, Pera Pedi, Near Pano Ki-
vides, Near Amathus, Trimiklini, Saettas, Platres, Fasoula, Yerasa, Lania, Livadin of
Cedars (Paphos Forest), Platania Forest Station, Karpasian Peninsula, Eleousa Monas-
tery, Xerokolimbi Stream near Trooditissa, Kykkou Monastery.
Ebmer localities: Platres, Mochi, Lefkara, Coral Bay, Agios Nikolaos, Troodos-
Pano Platres road, Moni Trooditissa, Caledonia Waterfall, Tripylos Cedar Valley, south
of Mt. Olympus,Kannaviou, Arminou to Filousa, Agios Nikolaos to Mandria, Pano
Panagia, Moni Trooditissa,north of Mt. Olympus, South of Kakopetria, East of Plata-
nia, 3 km north east Troodos, north of Platres.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N, 32.55°E,
20.VI.2013, (2♀); 20 km N Paphos, Kathikas, 34.90°N, 32.42°E, 20.VI.2013, (1♀);
8 km N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (2♀);
Limassol District: Troodos, Mt. Olympos, 34.93°N, 32.86°E, 20.VI.2013, C. (2♀); 2
km N of Anogyra, 34.748126°N, 32.732248°E, 1.V.2015, (1♀); Anogyra to Avdimou
Road km 2, 34.723986°N, 32.736892°E, 3.V.2015, (1♀), visiting Asteraceae; 0.7 km
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
28
N of Anogyra, 34.745537°N, 32.73385°E, 3.V.2015, (1♀); Anogyra to Pachna Road
km 4, 34.764269°N, 32.757736°E, 5.V.215, (1♀), all Kouklia, Kathikas, Mavroko-
lympos Reservoir and Troodos Mt. Olympos records C. Schmid-Egger leg., A.W. Eb-
mer det., all Anogyra records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus. ENDEMIC.
Halictus (Seladonia) phryganicus (Pauly & Devalez, 2015)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1957a); Georghiou (1977); Ebmer (2014); Pauly (2015).
Mavromoustakis localities. Limassol, Polemedia Hills, Mt. Troodos Kannoures
springs, Erimi, Chiicoudia marshes, Amathus, Pissouri, Yermasoyia River/Hills,
Moni, Pyrga, Near Mesayitonia, Alassa.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (5♀); 6 km NE Polis, beach, 35.06°N, 32.46°E, 20.VI.2013,
(4♀); 6 km W Polis, botanical garden, 35.03°N, 32.37°E, 20.VI.2013, (1♀); 8
km N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (1♀,
1♂); Limassol District: SBA, 8 km S Limassol, Akrotiri (near Airbase), 34.60°N,
32.97°E, 20.VI.2013, (1♀, 1♂); Sovereign Base Area, Episkopi, Kensington Clis,
34.670772°N, 32.846923°E, 4.V.2015, (2♀), all Paphos district and 8 km S Limas-
sol Akrotiri records C. Schmid-Egger leg., A. Pauly det., all Episkopi records S.P.M.
Roberts leg., A. Pauly det.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia (Turkey, Israel), Central Asia, Southern Asia (Iran).
Notes. All records of Halictus smaragdulus Vachal, 1895 from Cyprus are now at-
tributable to this recently described species.
Halictus (Mucoreohalictus) pollinosus Sichel, 1860
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”], 1951, 1954, 1957a);
Ebmer (2014).
Mavromoustakis localities. Limassol, Pera Pedi, hills near Trimiklini, Amiandos,
Moni, Mt. Troodos Kannoures Springs.
Material examined. Limassol District: Amiantos, 34.918°N, 32.9472°E,
31.V.2012, 2.VI.2012, (2♀), pan trap (UAEG); Anogyra to Pachna Road km 4,
34.764269°N, 32.757736°E, 5.V.2015, (2♀), visiting Asteraceae; Paphos District: 2.7
km SW of Acheleia, Potamos tis Ezouzas, 34.729004°N, 32.457544°E, 30.IV.2015,
(1♀), collecting pollen at Chrysanthemum sp. (Asteraceae), all Amiantos records S.
Dimitriou leg., J. Devalez and A. Pauly det., all Anogyra and Paphos district records
S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Western Europe (France, Germany, Austria), Southern Eu-
rope, Eastern Europe, Northern Africa, Western Asia (Turkey, Israel, Jordan), Central
Asia, Southern Asia (Iran, Afghanistan, Pakistan).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 29
Notes. e subspecies from Cyprus is Halictus pollinosus limissicus Blüthgen, 1937,
from Limassol: 34.66839°N, 33.03252°E, 2.VI.1928, G.A. Mavromoustakis leg.,
P.A.V. Blüthgen det. ♂, (MFNB).
Halictus (Vestitohalictus) pulvereus Morawitz, 1874
References. Ebmer (2014).
Material examined. Paphos District: 20 km N Paphos, Kathikas, 34.90°N,
32.42°E, 20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (2♀).
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Ukraine),
Western Asia (Turkey), Central Asia (Turkmenistan, Uzbekistan), Southern Asia (Iran,
Afghanistan), Eastern Asia.
Halictus (Halictus) quadricinctus (Fabricius, 1776)
References. Georghiou (1977).
Material examined. Limassol District: 1 km E of Pissouri, 34.677579°N, 32.722066°E,
27.IV.2015, (1♀), visiting Asteraceae; 2 km N of Anogyra, 34.748126°N, 32.732248°E,
1.V.2015, (1♀), visiting Asteraceae, all records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Europe (Denmark, Finland), Western Asia, Central Asia, Southern Asia (Iran, Af-
ghanistan), Eastern Asia (China, Mongolia).
Halictus (Halictus) resurgens Nurse, 1903
References. Blüthgen (1923); Mavromoustakis (1949 [“1948”], 1951, 1952, 1953,
1954, 1957a, b); Georghiou (1977); Ebmer (2014).
Mavromoustakis localities. Limassol, Mesayitonia, Chiicoudia marshes, Moni,
Yermasoyia River, Amathus, Lania, Pera Pedi, Famagusta, Nicosia, Livadin of Cedars
(Paphos Forest), Near Zakaki, Near Amathus, Near Ayios eodoros (Pitsillias).
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (2♀); 8 km N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E,
20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (2♀, 2♂); Limassol District: Ano-
gyra, 34.73663°N, 32.732715°E, 26.IV.2015, (4♀), visiting Chrysanthemum (Asterace-
ae); Sovereign Base Area, Avdimou Bay Clis, 34.656698°N, 32.773339°E, 27.IV.2015,
(1♀); Sovereign Base Area, Akrotiri, 34.628771°N, 32.941031°E, 29.X.2016, (3♂, 1♀),
visiting Dittrichia viscosa (Asteraceae); 2 km N of Anogyra, 34.748126°N, 32.732248°E,
1.V.2015, (1♀); 0.7 km N of Anogyra, 34.745537°N, 32.73385°E, 3.V.2015, (1♀); Lar-
naca District: Zygi, 34.731233°N, 33.343487°E, 28.X.2016, (1♂), visiting Dittrichia
viscosa (Asteraceae), all Paphos district records C. Schmid-Egger leg., A.W. Ebmer det., all
Limassol and Larnaca district records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Southern Europe, Eastern Europe, Northern Africa (Egypt),
Western Asia (Turkey), Central Asia, Southern Asia, Eastern Asia (China).
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30
Halictus (Halictus) subsenilis Blüthgen, 1955
References. Ebmer (1975, 2014).
Distribution. Cyprus, Western Asia (Israel).
Halictus (Halictus) tetrazonianellus Strand, 1909
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1953); Ebmer (2014).
Mavromoustakis localities. Limassol, Cherkes, Moni, Yermasoyia River, Near
Paramytha, Near Zakaki, Near Ayios eodoros (Pitsilia), Larnaca.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (13♀); 20 km N Paphos, Kathikas, 34.90°N, 32.42°E,
20.VI.2013, (2♀); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E, 20.VI.2013,
(4♀); Limassol District: 1 km E of Pissouri, 34.677579°N, 32.722066°E, 27.IV.2015,
(1♀), all Paphos district records C. Schmid-Egger leg., A.W. Ebmer det., all Limassol
district records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Ukraine,
Moldova, Russian Federation), Western Asia (Turkey), Central Asia (Turkmenistan),
Southern Asia (Iran).
Genus Lasioglossum CURTIS, 1833
37 species.
Lasioglossum (Lasioglossum) aegyptiellum (Strand, 1909)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1957a); Ebmer (2014).
Mavromoustakis localities. Limassol, Cherkes, Chiicoudia marshes, Fassouri,
Amathus, Moni, Yermasoyia Hills, Near Mesayitonia, Larnaca.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (1♀).
Distribution. Cyprus, Southern Europe (Croatia, Greece), Eastern Europe (Bul-
garia), Northern Africa (Libya, Egypt), Western Asia, Central Asia (Turkmenistan),
Southern Asia (Iran), Eastern Asia (China).
Lasioglossum (Dialictus) akroundicum (Blüthgen, 1937)
Type locality–country. Cyprus, Akrounda: 34.768889°N, 33.079444°E, 24.IV.1931,
G.A. Mavromoustakis leg., P.A.V. Blüthgen det. ♀, (MFNB).
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”]); Pittioni (1950);
Mavromoustakis (1951, 1952); Ebmer (2014).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 31
Mavromoustakis localities. Pyrgos, Near Paramytha, Akrounda, Near Eftagonia,
Mt. Troodos, Listovounos.
Material examined. Limassol District: Troodos, Chionistra, 34.9317°N,
32.8664°E, 31.V.2012, 2.VI.2012, S. Dimitriou leg., A. Ebmer det. (1♀), pan trap
(UAEG); Troodos, Mt. Olympos, 34.93°N, 32.86°E, 20.VI.2013, C. Schmid-Egger
leg., A.W. Ebmer det. (3♀).
Distribution. Cyprus. ENDEMIC.
Lasioglossum (Sphecodogastra) anellum (Vachal, 1905)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1957a); Ebmer (2014).
Mavromoustakis localities. Limassol, Pera Pedi, Mt. Troodos, Amiantos, Ayia
Varvara (Stavrovouni).
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (1♀); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E,
20.VI.2013, (1♀); 6 km NE Polis, beach, 35.06°N, 32.46°E, 20.VI.2013, (1♀); 8 km
N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (1♀, 2♂); Nr
Arminou Reservoir, 34.883435°N, 32.750988°E, 29.IV.2015, (1♀); Asprokremmos
Dam, 34.720825°N, 32.551994°E, 30.IV.2015, (1♀), collecting pollen at Sisymbrium
(Brassicaceae); Limassol District: 1 km E of Pissouri, 34.677579°N, 32.722066°E,
27.IV.2015, (1♀), collecting pollen at Brassicaceae; Pissouri 2 km S, 34.654385°N,
32.717924°E, 30.x.2016, (2♂), visiting Ceratonia siliqua (Fabaceae); Larnaca District:
Zygi, 34.731233°N, 33.343487°E, 28.X.2016, (1♂), visiting Ceratonia siliqua (Fa-
baceae); Zygi, 34.746277°N, 33.384472°E, 28.X.2016, (1♂), visiting Ceratonia sili-
qua (Fabaceae), all Kouklia, Lara beach, Polis and Mavrokolympos Reservoir records
C. Schmid-Egger leg., A.W. Ebmer det., all Arminou Reservoir, Asprokremmos Dam,
Limassol and Larnaca district records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Southern Europe, Eastern Europe (Bulgaria), Western Asia,
Southern Asia (Iran).
Lasioglossum (Lasioglossum) aphrodite Ebmer, 2014
Type locality–country. Cyprus, Mt. Troodos, S of Troodos, Caledonia Waterfall,
1400–1500 m: 34.9045°N, 32.86849°E, 10.VII.1987, A.W. Ebmer leg., A.W. Ebmer
det. ♂, (AWE).
References. Ebmer (2014).
Material examined. Limassol District: Troodos, Chionistra, 34.9317°N,
32.8664°E, 14-16.V.2012, (1♂, 2♀), pan trap (UAEG); Troodos, Chionistra,
34.9317°N, 32.8664°E, 31.V.2012, 2.VI.2012, (7♂, 34♀), pan trap (UAEG), all re-
cords S. Dimitriou leg., A. Ebmer det.
Distribution. Cyprus. ENDEMIC.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
32
Lasioglossum (Hemihalictus) clypeiferellum (Strand, 1909)
References. Mavromoustakis (1949 [“1948”]); Ebmer (2014).
Mavromoustakis localities. Limassol.
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Northern
Africa (Egypt), Western Asia (Israel), Central Asia, Southern Asia (Iran, Afghanistan),
Eastern Asia (Mongolia).
Lasioglossum (Hemihalictus) convexiusculum (Schenck, 1853)
References. Mavromoustakis (1949 [“1948”]); Ebmer (2014).
Mavromoustakis localities. Mt. Troodos.
Material examined. Limassol District: Troodos, Mt. Olympos, 34.93°N, 32.86°E,
20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (6♀).
Distribution. Cyprus, Western Europe, Southern Europe (Spain, Italy, Greece),
Eastern Europe (Czech Republic, Poland, Ukraine), Northern Europe (Lithuania),
Western Asia (Turkey), Central Asia (Tajikistan), Southern Asia (Iran).
Lasioglossum (Sphecodogastra) damascenum (Pérez, 1910)
References. Mavromoustakis (1954, 1957a); Ebmer (2014).
Mavromoustakis localities. Pera Pedi, Yerasa.
Material examined. Limassol District: Sovereign Base Area, Episkopi, Kensington
Clis, 34.670772°N, 32.846923°E, 4.V.2015, S.P.M. Roberts leg., A. Pauly det. (1♂).
Distribution. Cyprus, Southern Europe, Eastern Europe (Hungary, Romania,
Bulgaria), Western Asia (Turkey, Israel, Syria).
Lasioglossum (Hemihalictus) dolichocephalum (Blüthgen, 1923)
References. Mavromoustakis (1949 [“1948”]); Ebmer (2014).
Mavromoustakis localities. Ayios Athanasios.
Distribution. Cyprus, Southern Europe (North Macedonia, Croatia, Greece),
Western Asia (Turkey, Israel, Lebanon).
Lasioglossum (Hemihalictus) elegans (Lepeletier, 1841)
References. Mavromoustakis (1949 [“1948”], 1951); Ebmer (2014).
Mavromoustakis localities. Apostolos Varnavas (near Famagusta), Ayios Athana-
sios, Evdhimou River.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe (Hungary, Romania), Northern Africa (Algeria), Western Asia
(Turkey, Israel, Georgia), Central Asia, Southern Asia (Iran).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 33
Lasioglossum (Sphecodogastra) epipygiale (Blüthgen, 1924)
References. Ebmer (2014).
Distribution. Cyprus, Western Asia (Turkey), Central Asia, Southern Asia.
Lasioglossum (Hemihalictus) erraticum (Blüthgen, 1931)
References. Mavromoustakis (1949 [“1948”]); Ebmer (2014).
Mavromoustakis localities. Ayios Athanasios.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Armenia).
Lasioglossum (Hemihalictus) griseolum (Morawitz, 1872)
References. Mavromoustakis (1949 [“1948”], 1957a); Pittioni (1950); Ebmer (2014).
Mavromoustakis localities. Limassol, Cherkes.
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (3♀); 6 km NE Polis, beach, 35.06°N, 32.46°E, 20.VI.2013,
(1♀), all records C. Schmid-Egger leg., A.W. Ebmer det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe (Bul-
garia), Northern Africa (Morocco, Tunisia), Western Asia, Central Asia, Southern Asia
(Afghanistan).
Lasioglossum (Sphecodogastra) imbecillum Ebmer, 1974
References. Ebmer (2014).
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia (Turkey, Jordan).
Lasioglossum (Lasioglossum) kotschyi Ebmer, 1981
Type locality–country. Described from Cyprus as Halictus eurasicus torquillus Warncke,
1982, from Mt. Troodos: 34.9045°N, 32.86849°E, 7.VII.1935, A.W. Ebmer det. ♂.
References. Ebmer (1981); Warncke (1982); Pittioni (1950); Ebmer (2014).
Material examined. Limassol District: Troodos, Chionistra, 34.9317°N,
32.8664°E, 14-16.V.2012, (14♀), pan trap (UAEG); Troodos, Chionistra, 34.9317°N,
32.8664°E, 31.V.2012, 2.VI.2012, (1♀), pan trap (UAEG), all records S. Dimitriou
leg., A. Ebmer det.
Distribution. Cyprus. ENDEMIC.
Lasioglossum (Hemihalictus) laevidorsum (Blüthgen, 1923)
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”], 1951); Warncke
(1982); Ebmer (2014).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
34
Mavromoustakis localities. Mt. Troodos Kannoures springs, Pera Pedi, Between
Mozaras Station and Kato Aminados, Mt. Troodos Pasha Livadin.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern Eu-
rope, Northern Africa (Egypt), Western Asia (Turkey, Israel), Southern Asia (Iran, India).
Notes. e subspecies described from Cyprus is Lasioglossum laevidorsum troodi-
cum (Blüthgen, 1937), from Mt. Troodos: 34.9234°N, 32.8833°E, 19.VI.1935 G.A.
Mavromoustakis leg., P.A.V. Blüthgen det. ♀, (KW).
Lasioglossum (Sphecodogastra) laticeps (Schenck, 1870)
References. Pittioni (1950); Mavromoustakis (1957b); Ebmer (2014).
Mavromoustakis localities. Limassol, Apsiou, Moni River, Near Amathus, Ka-
loiri Hill, Morphou, Trimiklini, Kellaki, Monagroulli, Near Mesayitonia, Polemedia.
Distribution. Cyprus, Widespread in Europe, Western Asia (Turkey, Israel),
Southern Asia (Iran).
Lasioglossum (Leuchalictus) leucozonium (Schrank, 1781)
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”], 1951, 1952, 1953);
Ebmer (2014).
Mavromoustakis localities. Limassol, Polemedia Hills, Yermasoyia River, Ama-
thus, Trimiklini, Near Famagusta, Saettas, Kilani.
Material examined. Nicosia District: Linou, 35.0755°N, 32.9164°E, 5-7.IV.2012,
(7♀), pan trap (UAEG); Linou, 35.0755°N, 32.9164°E, 25-27.IV.2012, (6♀), pan
trap (UAEG); Linou, 35.0755°N, 32.9164°E, 14-16.V.2012, (1♀), pan trap (UAEG);
Kakopetria, 34.992°N, 32.9082°E, 5-7.IV.2012, (1♀), pan trap (UAEG); Kakope-
tria, 34.992°N, 32.9082°E, 14-16.V.2012, (1♀), pan trap (UAEG); Limassol District:
Amiantos, 34.918°N, 32.9472°E, 5-7.IV.2012, (2♀), pan trap (UAEG), all records S.
Dimitriou leg., J. Devalez det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran), Eastern Asia (Mongolia, China).
Lasioglossum (Hemihalictus) limbellum (Morawitz, 1876)
References. Ebmer (2014).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa (Morocco, Algeria, Tunisia), Western Asia (Turkey, Israel), Central Asia,
Southern Asia (Iran, Afghanistan), Eastern Asia (China).
Lasioglossum (Sphecodogastra) lineare (Schenck, 1870)
References. Mavromoustakis (1949 [“1948”], 1952, 1954); Ebmer (2014).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 35
Mavromoustakis localities. Limassol, Akrotiri Forest, Polemedia Hills, Ayios
Athanasios, Yermasoyia River, Pera Pedi.
Material examined. Limassol District: Troodos, Chionistra, 34.9317°N,
32.8664°E, 14-16.V.2012, S. Dimitriou leg., A. Ebmer det. (1♂), pan trap (UAEG);
Paphos District: 2.7 km SW of Acheleia, Potamos tis Ezousas, 34.729004°N,
32.457544°E, 30.IV.2015, S.P.M. Roberts leg., A. Pauly det. (1♀).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, West-
ern Asia, Central Asia (Turkmenistan, Tajikistan, Uzbekistan), Southern Asia (Iran).
Lasioglossum (Hemihalictus) lucidulum (Schenck, 1861)
References. Ebmer (2014).
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (1♀).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey, Israel, Georgia), Central Asia, Southern Asia (Iran, Afghanistan, Pakistan),
Eastern Asia.
Lasioglossum (Sphecodogastra) malachurum (Kirby, 1802)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1953, 1954, 1957a, b); Georghiou (1977); Ebmer (2014).
Mavromoustakis localities. Limassol, Polemedia Hills, Yermasoyia, Cherkes,
Episkopi, Chiicoudia marshes, Famagusta.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (1♂); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E,
20.VI.2013, (6♀); 2.7 km SW of Acheleia, Potamos tis Ezouzas, 34.729004°N,
32.457544°E, 30.IV.2015, (1♀); Famagusta District: Rizokarpaso, 5 km E, 35.63°N,
34.50°E, 10.IV.2007, (1♂); Limassol District: Sovereign Base Area, Akrotiri, Bishop's
Pool, 34.597305°N, 32.984521°E, 28.IV.2015, (1♀), all Kouklia, Lara Beach, and
Rizokarpaso records C. Schmid-Egger leg., A.W. Ebmer det., all Potamos tis Ezousas
and Akrotiri records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Turkmenistan), Southern Asia (Iran).
Lasioglossum (Dialictus) mandibulare (Morawitz, 1866)
References. Mavromoustakis (1949 [“1948”], 1952, 1957a); Ebmer (2014).
Mavromoustakis localities. Limassol, Cherkes, Pernera coast of Paralimni, Chi-
icoudia marshes, Famagusta.
Distribution. Cyprus, Western Europe (Switzerland), Southern Europe (Spain,
Bosnia and Herzegovina, Greece), Eastern Europe (Romania, Bulgaria, Russian Fed-
eration), Western Asia.
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36
Lasioglossum (Evylaeus) marginatum (Brullé, 1832)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1954); Ebmer (2014).
Mavromoustakis localities. Polemedia Hills, Ayia Phyla, Pera Pedi, Amathus, Tri-
miklini, Kellaki, Lania, Platania Forest Station, Amyrou Monastery, Ayia Irini Station.
Material examined. Kyrenia District: 15 km E Kyrenia, 'Turtle Beach',
35.334413°N, 33.494187°E, 10.IV.2007, (15♀); Limassol District: Anogyra,
34.73663°N, 32.732715°E, 26.IV.2015, (1♀); Sovereign Base Area, Avdimou Bay Clis,
34.656698°N, 32.773339°E, 27.IV.2015, (1♀); 2 km N of Anogyra, 34.748126°N,
32.732248°E, 1.V.2015, (4♀); 0.7 km N of Anogyra, 34.745537°N, 32.73385°E,
3.V.2015, (1♀), visiting Cistus creticus (Cistaceae); Anogyra to Pachna Road km 4,
34.764269°N, 32.757736°E, 5.V.2015, (1♀), collecting pollen on Brassicaceae; Paphos
District: Nr Arminou Reservoir, 34.883435°N, 32.750988°E, 29.IV.2015, (5♀); 2.7
km SW of Acheleia, Potamos tis Ezouzas, 34.729004°N, 32.457544°E, 30.IV.2015,
(1♀), all Kyrenia district records C. Schmid-Egger leg., A.W. Ebmer det., all Limassol
and Paphos district records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Western Asia, Southern Europe (Spain, Greece), Eastern
Europe (Czech Republic, Hungary, Russian Federation), Western Asia (Israel, Arme-
nia), Central Asia, Southern Asia.
Lasioglossum (Hemihalictus) medinai (Vachal, 1895)
References. Mavromoustakis (1949 [“1948”], 1954); Ebmer (2014) [all as L. villosu-
lum (Kirby).
Integrative taxonomy (Pauly et al. 2019) resuscitated two cryptic species in this
complex including L. medinai, the form veried to occur on Cyprus, whereas Lasioglos-
sum villosulum villosulum sensu stricto has been conrmed from Greece including Crete
and from Israel but not from Cyprus.
References. Mavromoustakis (1949 [“1948”], 1954); Ebmer (2014).
Distribution. Western Europe (Austria), Southern Europe (Spain, France, Italy,
Greece including Crete), Eastern Europe (Romania, Russia (Volgograd)), Northern
Africa (Algeria), Western Asia.
Lasioglossum (Hemihalictus) mesosclerum (Pérez, 1903)
References. Ebmer (2014).
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (1♀); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E,
20.VI.2013, (1♀); 8 km N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E,
20.VI.2013, (1♂), all records C. Schmid-Egger leg., A.W. Ebmer det.
Distribution. Cyprus, Western Asia (France, Austria), Southern Europe, Eastern
Europe, Northern Africa (Morocco, Libya, Egypt), Western Asia, Central Asia, South-
ern Asia (Iran, Afghanistan).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 37
Lasioglossum (Hemihalictus) minutissimum (Kirby, 1802)
References. Ebmer (2014).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey), Southern Asia (Pakistan).
Lasioglossum (Hemihalictus) nitidiusculum (Kirby, 1802)
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Algeria,
Tunisia), Western Asia (Turkey), Central Asia (Turkmenistan, Kazakhstan, Tajikistan),
Southern Asia (Iran).
Lasioglossum (Sphecodogastra) obscuratum (Morawitz, 1876)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1954, 1958 ["1957"]).
Mavromoustakis localities. Polemedia Hills, Yermasoyia River, Episkopi, Ama-
thus, Fasoula, Yerasa, Mt. Troodos Kannoures Springs, Mt. Troodos open slopes, Ami-
antos, Near Paramytha, Lania, Kykkou Monastery, Famagusta.
Material examined. Limassol District: Troodos, Chionistra, 34.9317°N,
32.8664°E, 14-16.V.2012, (1♂, 1♀), pan trap (UAEG); Troodos, Chionistra,
34.9317°N, 32.8664°E, 31.V.2012, 2.VI.2012, (2♂, 4♀), pan trap (UAEG); Paphos
District: 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E, 20.VI.2013, (1♀), all
Limassol district records S. Dimitriou leg., J. Devalez det., all Paphos district records
C. Schmid-Egger leg., A.W. Ebmer det.
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Croatia,
Greece), Eastern Europe, Western Asia, Central Asia, Southern Asia (Iran, Afghani-
stan).
Lasioglossum (Lasioglossum) pallens (Brullé, 1832)
References. Warncke (1982); Ebmer (2014).
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 25-27.
IV.2012, S. Dimitriou leg., A. Ebmer det. (1♀), pan trap (UAEG); Limassol District:
2 km N of Anogyra, 34.748126°N, 32.732248°E, 1.V.2015, S.P.M. Roberts leg., A.
Pauly det. (2♀).
Distribution. Cyprus, Western Europe, Southern Europe (Spain, Italy, Greece),
Northern Europe (Lithuania), Eastern Europe (Czech Republic, Hungary), North-
ern Africa (Morocco), Western Asia (Turkey), Southern Asia (Iran), Eastern Asia
(Mongolia).
Notes. e subspecies described from Cyprus is Lasioglossum pallens kanta-
rae (Warncke, 1982), from Kantara Castle: 35.4064°N, 33.9233°E, 7.IV.1953, K.
Warncke det. ♂, (KW).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
38
Lasioglossum (Sphecodogastra) pauxillum (Schenck, 1853)
References. Pittioni (1950); Ebmer (2014).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco to Tuni-
sia), Western Asia, Central Asia, Southern Asia (Iran).
Lasioglossum (Dialictus) podolicum (Noskiewicz, 1925)
References. Ebmer (2014).
Distribution. Cyprus, Western Europe (France, Switzerland), Southern Europe,
Eastern Europe (Hungary, Ukraine), Western Asia (Turkey), Southern Asia (Iran).
Lasioglossum (Hemihalictus) puncticolle (Morawitz, 1872)
References. Ebmer (2014).
Distribution. Cyprus, Western Europe, Southern Europe, Northern Europe
(United Kingdom), Eastern Europe, Northern Africa, Western Asia (Turkey, Israel,
Georgia), Southern Asia (Iran).
Lasioglossum (Hemihalictus) pygmaeum (Schenck, 1853)
References. Ebmer (2014).
Material examined. Kyrenia District: 15 km E Kyrenia, 'Turtle Beach',
35.334413°N, 33.494187°E, 10.IV.2007, (1♀); Limassol District: 2 km N of Ano-
gyra, 34.748126°N, 32.732248°E, 1.V.2015, (1♀), collecting pollen at Cistus cre-
ticus (Cistaceae); Sovereign Base Area, Episkopi Kensington Clis, 34.670772°N,
32.846923°E, 4.V.2015, (3♂); Anogyra to Pachna Road km 4, 34.764269°N,
32.757736°E, 5.V.2015, (1♀), all Kyrenia district records C. Schmid-Egger leg., A.W.
Ebmer det., all Limassol district records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa (Morocco to Tunisia), Western Asia, Central Asia (Turkmenistan, Kazakh-
stan, Tajikistan), Southern Asia.
Lasioglossum (Hemihalictus) transitorium (Schenck, 1870)
References. Erlandsson (1979).
Material examined. Kyrenia District: 15 km E Kyrenia, 'Turtle Beach',
35.334413°N, 33.494187°E, 10.IV.2007, (1♀); Limassol District: Anogyra to Avdi-
mou Road km 2, 34.723986°N, 32.736892°E, 3.V.2015, (1♂, 1♀), visiting Apiace-
ae; Sovereign Base Area, Episkopi, Kensington Clis, 34.670772°N, 32.846923°E,
4.V.2015, (2♀), all Kyrenia district records C. Schmid-Egger leg., A.W. Ebmer det.,
all Limassol district records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 39
Lasioglossum (Sphecodogastra) tricinctum (Schenck, 1874)
References. Ebmer (2014).
Material examined. Famagusta District: 15 km E Rizokarpaso, 'Golden Sands',
35.64°N, 34.55°E, 9.VIII.2001, (1♀); Paphos District: 20 km N Paphos, Kathikas,
34.90°N, 32.42°E, 20.VI.2013, (1♂); 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, C. Schmid-Egger leg., A.W. Ebmer det. (1♀); 8 km N Paphos,
Mavrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (1♀, 1♂), all records C.
Schmid-Egger leg., A.W. Ebmer det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe (Hun-
gary, Poland, Ukraine), Western Asia (Turkey, Azerbaijan), Southern Asia (Iran).
Notes. e subspecies described from Cyprus is Lasioglossum tricinctum lonicerae
Ebmer, 2014, from Mt. Troodos: 34.9234°N, 32.8833°E, 11.VII.1987, A.W. Ebmer
leg., A.W. Ebmer det. ♂, (AWE).
Lasioglossum (Hemihalictus) truncaticolle (Morawitz, 1877)
References. Ebmer (2014).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa (Algeria), Western Asia (Israel), Central Asia (Kazakhstan),
Southern Asia (Iran).
Lasioglossum (Leuchalictus) zonulum (Smith, 1848)
References. Ebmer (2014).
Distribution. Cyprus, Widespread in Europe, Western Asia (Turkey, Azerbaijan),
Southern Asia (Iran), Eastern Asia (China).
Genus Sphecodes LATREILLE, 1804
14 species.
Sphecodes albilabris (Fabricius, 1793)
References. Pittioni (1950); Mavromoustakis (1951, 1952, 1953, 1957a) (Warncke,
1992).
Mavromoustakis localities. Cherkes, Moni River, Near Limassol, Nicosia, Poleme-
dia Hills, Salipes marshes near Akrotiri, Trimiklini, Yermasoyia River, Zakaki marshes.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran, India).
Notes. Includes specimens labelled S. fumipennis by Mavromoustakis, which are
referable to this species. According to Warncke (1992) Cyprus specimens refer to S.
a. rubripes Spinola, 1838, variously treated by recent authors as a valid subspecies or
as a species.
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40
Sphecodes alternatus Smith, 1853
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952, 1954).
Mavromoustakis localities. Asomatos, Pera Pedi, Akrounda.
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia (Iran), Eastern
Asia (China).
Notes. Described from Cyprus as Sphecodes alternatus lindbergi Pittioni, 1950,
from Geroskipou, E of Paphos: 34.76666°N, 32.46666°E, 20.VII.1939, H. Lindberg
leg., B. Pittioni det., (MZHF).
Sphecodes croaticus Meyer, 1922
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Limassol, Episkopi, Yermasoyia River.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa (Morocco), Western Asia (Turkey), Central Asia (Turkmenistan).
Notes. e subspecies described from Cyprus is Sphecodes croaticus cypricus Blüth-
gen, 1937, from Limassol: 34.66839°N, 33.03252°E, 7.III.1931, G.A. Mavromousta-
kis leg., P.A.V. Blüthgen det. ♀, (MFNB).
Sphecodes ephippius (Linnaeus, 1767)
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N, 33.096194°E,
7.III.2017, Bee Course students leg., M. Schwarz det. (1♀).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey), Eastern Asia.
Sphecodes gibbus (Linnaeus, 1758)
References. Blüthgen (1937); Mavromoustakis (1949 [“1948”]); Pittioni (1950); Ma-
vromoustakis (1952, 1954, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Episkopi, Pera Pedi, Akrounda, Moni Riv-
er, Younarka (near Zakaki), Near Asomatos.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia, Eastern Asia (China, Mongolia).
Notes. Described from Cyprus as Sphecodes pergibbus Blüthgen, 1937, from [Li-
massol; Episcopi]: 34.675134°N, 32.882598°E, IV-VI, [G.A. Mavromoustakis] leg.,
P.A.V. Blüthgen, det., (MFNB).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 41
Sphecodes longuloides Blüthgen, 1923
References. Warncke (1992).
Distribution. Cyprus, Southern Europe (Spain, Italy, Portugal), Northern Africa
(Morocco, Algeria, Tunisia).
Sphecodes longulus Hagens, 1882
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Tunisia, Libya),
Western Asia (Turkey, Syria, Armenia), Central Asia, Southern Asia (Iran), Eastern
Asia (China, Japan).
Sphecodes monilicornis (Kirby, 1802)
References. Mavromoustakis (1949 [“1948”], 1952, 1957a); Pittioni (1950).
Mavromoustakis localities. Limassol, Fassouri, Mt. Troodos, Cherkes, Moni
River, Pyrga (Larnaca).
Material examined. Limassol District: 1 km E of Pissouri, 34.677579°N,
32.722066°E, 27.IV.2015, S.P.M. Roberts leg./det. (2♀).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Turkmenistan, Uzbekistan, Tajikistan), Southern Asia (Iran, Pakistan,
India), Eastern Asia.
Notes. Described from Cyprus as Sphecodes quadratus cephalotiformes Pittioni,
1950, from Mt. Troodos, Chionistra: 34.9364°N, 32.8636°E, 17.VI.1939, H. Lind-
berg leg., B. Pittioni det., (MZHF). Sphecodes monilicornis is recognised sensu lato fol-
lowing Nieto et al. (2014), including forms that have been considered distinct such as
Sphecodes cephalotes.
Sphecodes pellucidus Smith, 1845
References. Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Episkopi, Pera Pedi, Ayios Kostantinos of Pitsilia.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Turkmenistan, Kyrgyzstan), Southern Asia (Iran), Eastern Asia (China,
Mongolia).
Sphecodes pinguiculus Pérez, 1903
References. Warncke (1992).
Distribution. Cyprus, Western Europe (France), Southern Europe (Spain, Italy,
Greece), Eastern Europe, Northern Africa, Western Asia (Turkey), Eastern Asia (Mongolia).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
42
Sphecodes pseudofasciatus Blüthgen, 1925
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N, 33.096194°E,
7.III.2017, Bee Course students leg., M. Schwarz det. (2♀).
Distribution. Cyprus, Western Europe, Southern Europe (Spain, Italy. Portugal),
Eastern Europe, Western Asia (Turkey).
Sphecodes puncticeps omson, 1870
References. Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Cherkes, Yermasoyia River.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Egypt), Western
Asia (Israel), Central Asia (Uzbekistan, Tajikistan), Southern Asia (Iran), Eastern Asia
(Mongolia).
Sphecodes ruventris (Panzer, 1798)
References. Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Episkopi, Pera Pedi.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Alge-
ria), Western Asia, Central Asia (Uzbekistan, Kazakhstan, Tajikistan), Southern Asia
(Iran).
Sphecodes schenckii Hagens, 1882
References. Warncke (1992).
Distribution. Cyprus, Western Asia, Southern Europe, Eastern Europe, Northern
Africa, Western Asia (Turkey), Southern Asia (Iran).
NOMIINI
Genus Pseudapis KIRBY, 1900
4 species.
Pseudapis (Nomiapis) bispinosa (Brullé, 1832)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1957a); Ebmer (2014).
Mavromoustakis localities. Limassol, Akrotiri Forest, Cherkes, Zakaki, Near Ak-
rotiri, Moni, Pyrgos, Larnaca, Eftagonia.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa, Western Asia, Central Asia, Southern Asia, Eastern Asia (China).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 43
Pseudapis (Nomiapis) diversipes (Latreille, 1806)
References. Mavromoustakis (1948); Pittioni (1950); Mavromoustakis (1951, 1952,
1957a); Ebmer (2014).
Mavromoustakis localities. Limassol, Cherkes, Akrotiri Bay, Yermasoyia River,
Moni, Tsada, Erimi, Pera Pedi, Chiicoudia marshes, Near Akhyritou, Kathikas, Near
Enkomi of Famagusta, Larnaca, Ayia Varvara Stavrovouni.
Material examined. Famagusta District: Achna Dam, 35.05519°N, 33.814011°E,
28.X.2016, (1♀), visiting Dittrichia viscosa (Asteraceae); Paphos District: Polis,
35.053539°N, 32.351197°E, 30.X.2016, (2♂, 1♀), visiting Dittrichia viscosa (Aster-
aceae), all records S.P.M. Roberts leg., A. Pauly det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, Cen-
tral Asia.
Pseudapis (Nomiapis) equestris (Gerstäcker, 1872)
References. Ebmer (2014).
Distribution. Cyprus, Southern Europe (Greece), Western Asia, Central Asia
(Turkestan), Southern Asia (Iran).
Pseudapis (Nomiapis) valga (Gerstäcker, 1872)
References. Ebmer (2014).
Distribution. Cyprus, Southern Europe, (North Macedonia, Spain, Greece), East-
ern Europe (Czech Republic), Western Asia, Central Asia (Turkmenistan, Kazakhstan,
Tajikistan), Southern Asia (Iran, Pakistan).
NOMIOIDINI
Genus Ceylalictus STRAND, 1913
1 species.
Ceylalictus (Ceylalictus) variegatus (Olivier, 1789)
References. Cockerell (1931); Mavromoustakis (1949 [“1948”]); Pittioni (1950); Ma-
vromoustakis (1951, 1952); Georghiou (1977); Ebmer (2014).
Mavromoustakis localities. Limassol, Cherkes, Zakaki, Akrotiri Bay and Forest,
Yermasoyia River, Moni River, Pernera coast of Paralimni, Akrotiri Bay Bogazi shore.
Material examined. Paphos District: 2.7 km SW of Acheleia, Potamos tis Ezou-
zas, 34.729004°N, 32.457544°E, 30.IV.2015, (7♀), collecting pollen on Tama-
rix (Tamaricaceae); Polis, 35.037305°N, 32.391073°E, 30.X.2016, (1♂), visiting
Ceratonia siliqua (Fabaceae); Larnaca District: Zygi, 34.731233°N, 33.343487°E,
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
44
28.X.2016, (1♂), visiting Ceratonia siliqua (Fabaceae), all records S.P.M. Roberts
leg., A. Pauly det.
Distribution. Cyprus, Widespread in Europe, Western Africa, Northern Africa,
Western Asia, Central Asia, Southern Asia, Eastern Asia.
Genus Nomioides SCHENCK, 1867
1 species.
Nomioides (Nomioides) minutissimus (Rossi, 1790)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952); Ebmer (2014).
Mavromoustakis localities. Limassol, Akrotiri Bay, Yermasoyia River.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey, Israel, Georgia), Central Asia, Southern Asia, Eastern Asia (China, Mongolia).
ROPHITINI
Genus Dufourea LEPELETIER, 1841
1 species.
Dufourea (Cyprirorophites) cypria Mavromoustakis, 1952
Type locality–country. Cyprus, Famagusta: 35.125°N, 33.941667°E, 24.IV.1949,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♂, (DAAN).
References. Mavromoustakis (1952); Ebmer (2014).
Mavromoustakis localities. Famagusta.
Distribution. Cyprus, Western Asia (Turkey).
COLLETIDAE
COLLETINI
Genus Colletes LATREILLE, 1802
7 species.
Colletes brevigena Noskiewicz, 1936
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1957a, b).
Mavromoustakis localities. Limassol, Polemedia Hills, Akrotiri Forest, Kantara
Mountains, Chiicoudia marshes, Platania Forest, Yermasoyia River, Kellaki, Ama-
thus, Kellaki, Zakaki, Stavrovouni Monastery, Monagroulli, Paramali.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 45
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E,
31.V.2012, 2.VI.2012, (1♀), pan trap (UAEG); Makria Kontarka, 34.9095°N,
32.8971°E, 31.V.2012, 2.VI.2012, (1♀), pan trap (UAEG); Pedoulas, 34°57.711"N,
32°49.733"E, 14.X.2017, (1♂), visiting Hedera helix; Between Pedoulas-Kalopanay-
iwtis, 34°58.308"N, 32°49.775"E, 14.X.2017, (1♂), visiting Hedera helix; Limassol
District: Troodos, Chionistra, 34.9317°N 32.8664°E, 31.V.2012, 2.VI.2012, (1♀), pan
trap (UAEG); Troodos Natural Bath, 34 52.892°N, 32 52.618°E, 27.IV.2016, (1♀), vis-
iting Cistus creticus; Yermasoyia Dam, 34.745849°N, 33.083579°E, 28.X.2016, (1♂),
visiting Dittrichia viscosa (Asteraceae); Yermasoyia Dam, 34.745849°N, 33.083579°E,
28.X.2016, (1♂), visiting Dittrichia viscosa; Sovereign Base Area, Paramali, 34.676011°N
32.794947°E, 30.X.2016, (3♂), visiting Dittrichia viscosa (Asteraceae); 0.7 km N of
Anogyra, 34.743911°N, 32.732439°E, 26.X.2016, (41♂, 3♀), visiting Ceratonia si-
liqua (Fabaceae); Anogyra, 34.729369°N, 32.738368°E, 11.III.2016, (1♂), visiting
Ceratonia siliqua (Fabaceae); Anogyra, 34.742646°N, 32.730431°E, 11.III.2016,
(2♂), visiting Ceratonia siliqua (Fabaceae); Anogyra, 34.742646°N, 32.730431°E,
22.X.2016, (1♀), visiting Ceratonia siliqua (Fabaceae); Anogyra, 34.729369°N,
32.738368°E, 26.X.2016, (1♂), visiting Ceratonia siliqua (Fabaceae); Anogyra,
34.729369°N, 32.738368°E, 11.XII.2016, (1♀), visiting Ceratonia siliqua (Fabaceae);
Anogyra, 34.729369°N, 32.738368°E, 11.III.2017, , (1♀), visiting Ceratonia siliqua
(Fabaceae); Akrotiri, 34.600657°N, 32.971419°E, 11.VI.2016, (3♀, 1♂); Paramali,
34.676011°N, 32.794947°E, 30.X.2016, (1♂), visiting Ceratonia siliqua (Fabaceae);
Akrotiri, 34.600657°N, 32.971419°E, 30.X.2017, (1♀, 1♂), visiting Dittrichia viscosa;
Sovereign Base Area, Akrotiri, 34.600657 32.971419, 29.X.2016, (3♂, 4♀), visiting
Dittrichia viscosa (Asteraceae); Larnaca District: Zygi, 34.731233°N, 33.343487°E,
28.X.2016, (5♂, 1♀), visiting Ceratonia siliqua (Fabaceae) and Dittrichia viscosa
(Asteraceae); Zygi, 34.746277°N, 33.384472°E, 28.X.2016, (11♂, 1♀), visiting Cera-
tonia siliqua (Fabaceae); Zygi, 34.731233°N, 33.343487°E, 11.IV.2016, (1♂), visiting
Ceratonia siliqua (Fabaceae); Zygi, 34.746277°N, 33.384472°E, 11.IV.2016, (1♀),
visiting Ceratonia siliqua (Fabaceae); Zygi, 34.77458°N, 33.448023°E, 11.IV.2016,
(1♀), visiting Ceratonia siliqua (Fabaceae); Zygi, 34.731233°N, 33.343487°E,
23.X.2016, (1♀), visiting Ceratonia siliqua (Fabaceae); Avdellero, 35 00.598°N 33
35.041°E, 11.IV.2017, (1♂), visiting Ceratonia siliqua (Fabaceae); Famagusta Dis-
trict: Achna Dam, 35.05519°N, 33.814011°E, 28.X.2016, (1♂), visiting Dittrichia
viscosa (Asteraceae); Paphos District: Polis, 35.053539°N, 32.351197°E, 30.X.2016,
(10♂), (2♀), visiting Ceratonia siliqua (Fabaceae) and Dittrichia viscosa (Asteraceae);
Polis, 35.036538°N, 32.373117°E, 30.X.2016, (3♂), visiting Ceratonia siliqua (Fa-
baceae); Polis, 35.037305°N, 32.391073°E, 30.X.2016, (3♂, 1♀), visiting Ceratonia
siliqua (Fabaceae); Polis, 35.032203°N, 32.413732°E, 30.X.2016, det. (5♂, 5♀), vis-
iting Ceratonia siliqua (Fabaceae); Mesogi, 34.810747°N, 32.450907°E, 30.X.2016,
(7♂, 1♀), visiting Ceratonia siliqua (Fabaceae); Polis, 35.053539°N, 32.351197°E,
11.II.2016, (2♂), visiting Ceratonia siliqua (Fabaceae) and Dittrichia viscosa (Aster-
aceae); Polis, 35.037305°N, 32.391073°E, 11.II.2016, (1♀); Polis, 35.032203°N,
32.413732°E, 11.II.2016, (1♀); Polis, 35.036538°N, 32.373117°E, 21.X.2016,
(1♂); Polis, 35.037305°N, 32.391073°E, 21.X.2016, (1♀); Polis, 35.037305°N,
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
46
32.391073°E, 19.XI.2016, (1♀); Polis, 35.037305°N, 32.391073°E, 11.XII.2016,
(2♀); Polis, 35.036538°N, 32.373117°E, 11.II.2017, (2♀); Polis, 35.053539°N,
32.351197°E, 11.II.2017, (2♂); Polis, 35.053539°N, 32.351197°E, 11.XII.2017,
(2♀); Polis, 35.037305°N, 32.391073°E, 11.XII.2017, (1♀), all Polis records visiting
Ceratonia siliqua (Fabaceae), all Kakopetria, Makria Kontarka and Troodos Chionistra
records S. Dimitriou leg., J. Devalez det., all Pedoulas, Between Pedoulas-Kalopanay-
iotis, Troodos Natural Bath, Yermasoyia Dam, Anogyra, Paramali, Akrotiri, Zygi and
Polis records A. Varnava leg., M. Kuhlmann det., all Yermasoyia Dam, Sovereign Base
Area, Paramali, 0.7 km N of Anogyra, Sovereign Base Area, Akrotiri, Zygi, Achna
Dam, Polis, Mesogi records S.P.M. Roberts leg., M. Kuhlmann det.
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe (Hungary, Bulgaria, Ukraine), Western Asia (Turkey).
Colletes creticus Noskiewicz, 1936
Material examined. Limassol District: Sovereign Base Area, Episkopi, Kensington
Clis, 34.670772°N, 32.846923°E, 4.V.2015, (1♂); Pissouri 2 km S, 34.654385°N,
32.717924°E, 30.X.2016, (3♂), visiting Dittrichia viscosa (Asteraceae) and Ceratonia
siliqua (Fabaceae), all records S.P.M. Roberts leg., M. Kuhlmann det.
Distribution. Cyprus, Southern Europe (Greece).
Colletes cyprius Noskiewicz, 1936
Type locality–country. Cyprus, Kykkos mountains, 1500m: 34.984°N, 32.741°E,
V-VI.1922/1923 or 1930, G.A. Mavromoustakis leg., J. Noskiewicz det.
References. Noskiewicz (1936); Mavromoustakis (1949 [“1948”], 1951, 1952,
1957a).
Mavromoustakis localities. Limassol, Mesayitonia, Cherkes, Yermasoyia Hills,
Asomatos, Alassa River, Pera Pedi, Akrotiri Bay, Stroumbi, Livadin of Cedar (Paphos
Forest), Potamitissa, Akrounda, Moni, Kellaki, Larnaca, Kykkou Monastery, Ayia
Phyla, Mt. Troodos, Near Mesayitonia, Bogazi, Kathikas, Eftagonia, Xerokolimbi near
Trooditissa.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N, 32.55°E,
20.VI.2013, (1♂); 8 km N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E,
20.VI.2013, (1♂); Polis, 35.053539°N, 32.351197°E, 30.X.2016, (1♂), visiting Cera-
tonia siliqua (Fabaceae); Polis, 35.036538°N, 32.373117°E, 11.II.2017, (1♂), visiting
Ceratonia siliqua (Fabaceae); Limassol District: Anogyra, 34.729369°N, 32.738368°E,
11.V.2015, (1♀), visiting Ceratonia siliqua (Fabaceae); Akrotiri, 34.600657°N,
32.971419°E, 5.VIII.2016, (1♀); Anogyra, 34.729369°N, 32.738368°E, 26.X.2016
(1♀), visiting Ceratonia siliqua (Fabaceae); Anogyra, 34.729369°N, 32.738368°E,
11.III.2017, (1♂), visiting Ceratonia siliqua (Fabaceae); Anogyra, 34.729369°N,
32.738368°E, 18.X.2017, (2♀), visiting Ceratonia siliqua (Fabaceae), all 15 km SE
Paphos, Kouklia and Mavrokolympos Reservoir records, C. Schmid-Egger leg., M.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 47
Kuhlmann det., all Polis and Anogyra records A. Varnava leg., M. Kuhlmann det., all
Anogyra (11.V.2015) records S. Louca leg., M. Kuhlmann det.
Distribution. Cyprus. ENDEMIC.
Colletes hylaeiformis Eversmann, 1852
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (France, Germany, Austria), Southern Eu-
rope, Eastern Europe, Western Asia (Turkey, Georgia, Azerbaijan), Central Asia.
Colletes maidli Noskiewicz, 1936
References. Mavromoustakis (1949 [“1948”], 1952); Georghiou (1977).
Mavromoustakis localities. Asomatos, Yermasoyia River, Evdhimou River, Famagusta.
Material examined. Limassol District: Akrotiri, 34.583676°N, 32.949306°E,
20.VI.2016, (1♀); Akrotiri, 34.352103°N, 32.562132°E, 20.VI.2016, (1♂), all re-
cords A. Varnava leg., M. Kuhlmann det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Western Asia, Central Asia.
Colletes similis Schenck, 1853
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953).
Mavromoustakis localities. Limassol, Akrotiri Bay, Asomatos, Moni, Yerasa, Kato
Platres, Pera Pedi, Monagroulli, Platania Forest Station, Mt. Troodos.
Material examined. Limassol District: Sovereign Base Area, Akrotiri, 34.600657°N,
32.971419°E, 29.X.2016, (1♀), visiting Dittrichia viscosa (Asteraceae); Paphos District:
Polis, 35.053539°N, 32.351197°E, 30.X.2016, (5♀), visiting Dittrichia viscosa (Aster-
aceae), all records S.P.M. Roberts leg., M. Kuhlmann det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran), Eastern Asia (China).
Colletes squamulosus Noskiewicz, 1936
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Larnaca.
Distribution. Cyprus, Western Asia, Central Asia (Turkmenistan).
HYLAEINI
Genus Hylaeus FABRICIUS, 1793
12 species.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
48
Hylaeus (Paraprosopis) clypearis (Schenck, 1853)
References. Erlandsson (1986).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa, Western Asia (Turkey, Georgia).
Hylaeus (Abrupta) cornutus Curtis, 1831
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Limassol, Zakaki, Asomatos, Pera Pedi, Larnaca.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia (Is-
rael), Central Asia (Turkmenistan), Southern Asia (Iran).
Hylaeus (Spatulariella) cypricola (Warncke, 1972)
References. Pittioni (1950); Mavromoustakis (1957a).
Mavromoustakis localities. Cherkes.
Material examined. Limassol District: Anogyra, 34.741952°N, 32.734845°E,
3.V.2015, (2♂), visiting Sinapis (Brassicaceae); Sovereign Base Area, Episkopi Kensing-
ton Clis, 34.670772°N, 32.846923°E, 4.V.2015, (5♂, 3♀); Larnaca District: Zygi,
34.731233°N, 33.343487°E, 28.X.2016, (1♀), visiting Ceratonia siliqua (Fabaceae),
all records S.P.M. Roberts leg., G. Holmström and H. Dathe det.
Distribution. Cyprus, Western Asia (Turkey), Northern Africa (Egypt).
Notes. Described from Cyprus as Spatulariella (Brachyspatulariella) dimidiatus Pit-
tioni, 1950, from Limassol: 34.66839°N, 33.03252°E, G.A. Mavromoustakis leg., K.
Warncke det., (MZHF).
Hylaeus (Prosopis) gibbus Saunders, 1850
References. Alfken (1912).
Material examined. Limassol District: Sovereign Base Area, Akrotiri,
34.628771°N, 32.941031°E, 29.X.2016, (1♂, 2♀), visiting Dittrichia viscosa (Aster-
aceae); Paphos District: 2.7 km SW of Acheleia, Potamos tis Ezouzas, 34.729004°N,
32.457544°E, 30.IV.2015, (1♂, 2♀), all records S.P.M. Roberts leg., G. Holmström
and H. Dathe det.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Egypt),
Western Asia Central Asia (Kazakhstan), Southern Asia (Iran).
Notes. Although the Cyprus specimens listed below were determined by H. Dathe,
he himself believes that DNA analysis could reveal that they might be specimens of H.
incongruus Forster, 1871, H. stigmorhinus (Pérez, 1895) or the dark form of H. pictus
(Smith, 1853) (with black scutellum) (Dathe, 2018 in litt.). It is clear that a revision of
this group is required to elucidate the nature of this species group.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 49
Hylaeus (Dentigera) imparilis Förster, 1871
References. Alfken (1928); Pittioni (1950).
Material examined. Limassol District: 1 km E of Pissouri, 34.677579°N,
32.722066°E, 27.IV.2015, (2♂, 1♀), visiting Brassicaceae; Anogyra to Pachna Road km
4, 34.764269°N, 32.757736°E, 5.V.2015, (1♂); 0.7 km N of Anogyra, 34.743911°N,
32.732439°E, 26.X.2016, (1♀), visiting Ceratonia siliqua (Fabaceae); 1 km S of Ano-
gyra, 34.727647°N, 32.73462°E, 26.X.2016, (1♀), visiting Ceratonia siliqua (Fabace-
ae); Paphos District: Nr Arminou Reservoir, 34.883435°N, 32.750988°E, 29.IV.2015,
(1♂); Asprokremmos Dam, 34.720825°N, 32.551994°E, 30.IV.2015, (1♂, 2♀),
visiting Sisymbrium (Brassicaceae); 2.7 km SW of Acheleia, Potamos tis Ezousas
34.729004°N, 32.457544°E, 30.IV.2015, (1♂); Larnaca District: Zygi, 34.731233°N,
33.343487°E, 28.X.2016, (6♀), visiting Dittrichia viscosa (Asteraceae) and Ceratonia
siliqua (Fabaceae), all records S.P.M. Roberts leg., G. Holmström and H. Dathe det.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa (Morocco, Tunisia), Western Asia, Southern
Asia (Iran).
Notes. Described from Cyprus as Prosopis brevicornis var. cypria Alfken, 1928,
from [Yermasoyia; Episcopi]: 34.72231°N, 33.08497°E, 34.67513°N, 32.88260°E,
G.A. Mavromoustakis leg., J.D. Alfken det.
Hylaeus (Paraprosopis) lineolatus (Schenck, 1861)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1957a).
Mavromoustakis localities. Finikaria River, Mesapotamos, Near Zakaki, Near
Mesayitonia.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa (Morocco, Algeria), Western Asia (Israel, Lebanon, Azerbaijan), Southern
Asia (Iran).
Notes. Described from Cyprus as Prosopis lineolata rudis Pittioni, 1950, from
Kykkos: 34.984°N, 32.741°E, 15-17.VII.1939, H. Lindberg leg., B. Pittioni det.,
(MZHF).
Hylaeus (Prosopis) meridionalis Förster, 1871
References. Alfken (1928); Mavromoustakis (1949 [“1948”], 1951, 1957a).
Mavromoustakis localities. Akrotiri Forest, Cherkes, Zakaki, Asomatos, Fassouri,
Gape Gata, Larnaca, Pernera coast of Paralimni.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa (Morocco), Western Asia, Central Asia (Kyr-
gyzstan), Southern Asia (Iran).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
50
Notes. Described from Cyprus as Prosopis maculiscutum Alfken, 1928, from
Cherkes: 34.65°N, 32.975°E, 7 or 24.VIII.1924, G.A. Mavromoustakis leg., J.D.
Alfken det.
Hylaeus (Prosopis) pictus (Smith, 1853)
References. Mavromoustakis (1951, 1952).
Mavromoustakis localities. Near Limassol, Cherkes, Chiicoudia marshes, Pera
Pedi, Near Akrotiri, Yermasoyia River, Potamitissa, Mt. Troodos Kannoures springs,
Near Cape Gata.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Hungary, Bulgaria), Northern Africa, Western Asia.
Hylaeus (Dentigera) rubicola Saunders, 1850
References. Alfken (1928); Mavromoustakis (1949 [“1948”]); Pittioni (1950).
Mavromoustakis localities. Yermasoyia.
Distribution. Cyprus, Western Europe (France), Southern Europe (Spain, Italy,
Greece), Eastern Europe (Bulgaria), Northern Africa (Egypt), Western Asia.
Notes. Described from Cyprus as Prosopis rectanguliceps Alfken, 1928, from Ye-
rmasoyia: 34.7182°N, 33.08788°E, 1-20.IX.1924, G.A. Mavromoustakis leg., J.D.
Alfken det.
Hylaeus (Lambdopsis) scutellatus (Spinola, 1838)
References. Alfken (1928); Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mav-
romoustakis (1951, 1957a).
Mavromoustakis localities. Limassol, Cherkes, Zakaki, Moni, Pera Pedi, Stroum-
bi, Tsada.
Material examined. Paphos District: Asprokremmos Dam, 34.720825°N,
32.551994°E, 30.IV.2015, S.P.M. Roberts leg., G. Holmström and H. Dathe det.
(1♂), visiting Sisymbrium (Brassicaceae).
Distribution. Cyprus, Southern Europe, Northern Africa (Egypt), Western Asia
(Palestine to Turkey), Southern Asia (Iran).
Hylaeus (Paraprosopis) soror (Pérez, 1903)
References. Pittioni (1950).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria, Russian Federation), Northern Africa, Southern Asia (Iran).
Hylaeus (Paraprosopis) taeniolatus Förster, 1871
References. Erlandsson (1986).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 51
Material examined. Limassol District: Anogyra, 34.73663°N, 32.732715°E,
26.IV.2015, (1♂), visiting Brassicaceae; 1 km E of Pissouri, 34.677579°N,
32.722066°E, 27.IV.2015, (4♀, 1♂), visiting Brassicaceae; 2 km N of Ano-
gyra, 34.748126°N, 32.732248°E, 1.V.2015, (1♀); Anogyra to Avdimou Road
km 2, 34.723986°N, 32.736892°E, 3.V.2015, (1♂), visiting Apiaceae; Anogyra,
34.741952°N, 32.734845°E, 3.V.2015, (1♀, 3♂), visiting Brassicaceae; 0.7 km N
of Anogyra, 34.745537°N, 32.73385°E, 3.V.2015, (8♂); 0.7 km N of Anogyra,
34.743911°N, 32.732439°E, 26.X.2016, (1♀), visiting Fabaceae; Yermasoyia Dam,
34.755799°N, 33.096194°E, 7.III.2017, (1♂); Paphos District: Nr Arminou Res-
ervoir, 34.883435°N, 32.750988°E, 29.IV.2015, (1♀, 1♂); Asprokremmos Dam,
34.720825°N, 32.551994°E, 30.IV.2015, (1♀, 1♂), visiting Brassicaceae; 2.7 km
SW of Acheleia, Potamos tis Ezouzas, 34.729004°N, 32.457544°E, 30.IV.2015, (2♂);
Polis, 35.053539°N, 32.351197°E, 30.X.2016, (1♀, 1♂), visiting Fabaceae; Larnaca
District: Zygi, 34.731233°N, 33.343487°E, 28.X.2016, (1♀, 4♂), visiting Asteraceae
and Fabaceae, all records S.P.M. Roberts leg., G. Holmström and H. Dathe det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe (Bul-
garia), Northern Africa, Western Asia.
MEGACHILIDAE
LITHURGINI
Genus Lithurgus BERTHOLD, 1827
2 species.
Lithurgus (Lithurgus) chrysurus Fonscolombe, 1834
References. Mavromoustakis (1949 [“1948”], 1952, 1954, 1957a).
Mavromoustakis localities. Near Limassol, Cherkes, Pera Pedi, Mandria, Near
Kalliana.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa (Morocco), Western Asia, Central Asia (Ta-
jikistan), Southern Asia (Iran), Eastern Asia (China).
Lithurgus (Lithurgus) tibialis Morawitz, 1875
References. Mavromoustakis (1949 [“1948”], 1957a).
Mavromoustakis localities. Cherkes, Zakaki, Asomatos, Episkopi, Pera Pedi, Fas-
souri, Near Famagusta, Symboulas.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N, 32.55°E,
20.VI.2013, (1♂); 20 km N Paphos, Kathikas, 34.90°N, 32.42°E, 20.VI.2013, (2♂),
all records C. Schmid-Egger leg., C. Schmid-Egger det.
Distribution. Cyprus, Southern Europe, Northern Africa, Western Asia, Central
Asia (Turkmenistan), Southern Asia (Iran, Pakistan).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
52
OSMIINI
Genus Chelostoma LATREILLE, 1809
3 species.
Chelostoma (Chelostoma) comosum Müller, 2012
Type locality–country. Cyprus, Troodos, Kato Platres S of Foini: 34.8775°N,
32.8386°E, 1.V.2011, C. Sedivy, A. Müller leg., A. Müller det. ♂, (ETHZ).
References. Müller (2012).
Material examined. Larnaca District: 2 km S of Ora, 450m, 34.85°N, 33.20°E,
3.IV.2009, H.R. Schwenninger leg., A. Müller det. (1♀); Limassol District: Anogyra,
34.73663°N, 32.732715°E, 26.IV.2015, M. Jenner leg., G. Le Go det. (1♂); Fama-
gusta District: 5 km E of Rizokarpaso, 35.62°N, 34.46°E, C. Schmid-Egger leg., A.
Müller det. (1♂).
Distribution. Cyprus, Western Asia.
Chelostoma (Chelostoma) diodon Schletterer, 1889
References. Mavromoustakis (1951, 1952, 1957a).
Mavromoustakis localities. Yerasa, Platania Forest Station.
Material examined. Limassol District: Anogyra to Pachna Road km 4,
34.764269°N, 32.757736°E, 5.V.2015, S.P.M. Roberts leg., A. Müller det. (1♀), visit-
ing Asteraceae.
Distribution. Cyprus, Southern Europe (Greece), Western Asia, Southern Asia
(Iran).
Notes. e subspecies described from Cyprus is Chelostoma diodon cypriacum (Ma-
vromoustakis, 1951), from Yerasa: 34.8025°N, 32.998056°E, 22.III.1945, G.A. Mav-
romoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
Chelostoma (Chelostoma) lucens (Benoist, 1928)
References. Mavromoustakis (1951, 1952, 1957a).
Mavromoustakis localities. Apsiou, Yerasa.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia.
Genus Heriades SPINOLA, 1808
4 species.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 53
Heriades (Rhopaloheriades) clavicornis Morawitz, 1875
References. Ungricht, Müller and Dorn (2008).
Distribution. Cyprus, Southern Europe (Greece), Western Asia, Central Asia (Ta-
jikistan), Southern Asia (Iran).
Heriades (Michenerella) punctulifera Schletterer, 1889
References. Mavromoustakis (1951).
Mavromoustakis localities. Mt. Troodos.
Distribution. Cyprus, Southern Europe, Eastern Europe (Bulgaria), Western Asia.
Notes. e subspecies described from Cyprus is Heriades punctulifera troodica
(Mavromoustakis, 1951), from Mt. Troodos, open slopes, 1676.4m: 34.9234°N,
32.8833°E, 18.VII.1949 [“1948”], G.A. Mavromoustakis leg. G.A. Mavromoustakis
det. ♀, (DAAN).
Heriades (Heriades) rubicola Pérez, 1890
References. Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Near Limassol, Akrotiri Bay, Cherkes, Chiicoudia
marshes, Erimi, Alassa River, Evdhimou River.
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (4♀); Polis, 35.053539°N, 32.351197°E, 30.X.2016, (2♂), vis-
iting Dittrichia viscosa (Asteraceae); Polis, 35.053539°N, 32.351197°E, 30.X.2016,
(6♂), visiting Dittrichia viscosa; Limassol District: Sovereign Base Area, Akrotiri,
34.628771°N, 32.941031°E, 29.X.2016, (1♀), visiting Dittrichia viscosa (Asterace-
ae); Cherkes, 34.653067°N, 32.974233°E, 22.V.2016, (1♂); Cherkes, 34.65096°N,
32.99091°E, 26.VIII.2016, (1♀); Cherkes, 34.65096°N, 32.99091°E, 12.VIII.2016,
(1♀); Cherkes, 34.6432°N 32.9952°E, 6.XI.2016, (1♀); Akrotiri, 34.628817°N,
32.940667°E, 3.VII.2016, (2♂); Akrotiri, 34.628817°N, 32.940667°E, 26.VIII.2016,
(1♀); Akrotiri, 34.600657°N, 32.971419°E, 11.IX.2016, (1♂); Akrotiri,
34.628817°N, 32.940667°E, 8.X.2016, (1♀); Akrotiri, 34.628817°N, 32.940667°E,
6.XI.2016, (1♀); Akrotiri, 34.628817°N, 32.940667°E, 21.IV.2017, (1♀); Akrotiri,
34.628817°N, 32.940667°E, 3.VII.2017, (1♀, 1♂), all 20 km NNW Paphos and Lara
Beach (20.VI.2013) records, C. Schmid-Egger leg., A. Müller det., all Polis (30.X.2016,
2♂ ) and Sovereign Base Area, Akrotiri records S.P.M. Roberts leg., S.P.M. Roberts and
G. Le Go det., all Polis (30.X.2016, 6♂), all Cherkes and Akrotiri records A. Varnava
leg., A. Müller det.
Distribution. Cyprus, Western Europe (France, Switzerland), Southern Europe,
Eastern Europe (Bulgaria, Slovakia), Northern Africa, Western Asia, Central Asia
(Turkmenistan, Kazakhstan, Kyrgyzstan), Southern Asia.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
54
Heriades (Heriades) truncorum (Linnaeus, 1758)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952).
Mavromoustakis localities. Near Limassol, Chiicoudia marshes, Erimi, Near
Platania Forest Station, Rizokarpaso.
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (1♀); Polis, 35.053539°N, 32.351197°E, 30.X.2016, (1♀),
visiting Dittrichia viscosa (Asteraceae); Limassol District: Cherkes, 34.653067°N,
32.974233°E, 22.V.2016, (2♀, 1♂); Cherkes, 34.659801°N, 32.99051°E, 22.V.2016,
(1♀); Cherkes, 34.659801°N, 32.99051°E, 5.VI.2016, (2♀); Akrotiri, 34.600657°N,
32.971419°E, 22.V.2016, (1♂); Akrotiri, 34.628817°N, 32.940667°E, 13.X.2017,
(1♀); Platres, 34.886528°N, 32.862465°E, 14.X.2017, (1♀), visiting Hedera helix,
all 20 km NNW Paphos, Lara Beach records C. Schmid-Egger leg., A. Müller det.,
all Polis records S.P.M. Roberts leg., S.P.M. Roberts and G. Le Go det., all Limassol
district records A. Varnava leg., A. Müller det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia.
Genus Hoplitis KLUG, 1807
9 species.
Hoplitis (Alcidamea) acuticornis (Dufour & Perris, 1840)
References. Mavromoustakis (1939b, 1949 [“1948”]).
Mavromoustakis localities. Sphalangiotissa Monastery.
Material examined. Limassol District: Sovereign Base Area, Akrotiri, Bishop's
Pool, 34.597305°N, 32.984521°E, 28.IV.2015, M. Jenner leg., G. Le Go det. (1♀);
Polemidia, 34.71178°N, 33.004775°E, 8.III.2017, Bee Course students leg., A. Mül-
ler det. (2♂, 1♀).
Distribution. Cyprus, Widespread in Europe (except Great Britain and northern
Europe), Northern Africa, Western Asia, Central Asia, Southern Asia (Iran).
Hoplitis (Anthocopa) anipuncta (Alfken, 1935)
References. Müller (2018).
Distribution. Cyprus, Western Asia.
Hoplitis (Hoplitis) annulata (Latreille, 1811)
References. Mavromoustakis (1939b, 1949 [“1948”], 1951).
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (2♀); 8 km N Paphos, Mavrokolympos Reservoir, 34.85°N,
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 55
32.40°E, 20.VI.2013, (1♀); Asprokremmos Dam, 34.720825°N, 32.551994°E,
30.IV.2015, (1♂, 1♀), visiting Echium angustifolium (Boraginaceae); Limassol Dis-
trict: Sovereign Base Area, 8 km S Limassol, Akrotiri (near Airbase), 34.60°N, 32.97°E,
20.VI.2013, (2♀); Sovereign Base Area, Akrotiri, Bishop's Pool, 34.597305°N,
32.984521°E, 28.IV.2015, (1♂, 4♀), visiting Echium angustifolium (Boraginace-
ae); Cherkes, 34.65096°N, 32.99091°E, 9.IV.2016, (1♀); Akrotiri, 34.600657°N,
32.971419°E, 19.IV.2016, (1♂); Akrotiri, 34.600657°N, 32.971419°E, 22.V.2016,
(2♂); Akrotiri, 34.601506°N, 32.986197°E, 22.V.2016, (2♂); Akrotiri, 34.600657°N,
32.971419°E, 5.V.2017, (1♀, 1♂); Akrotiri, 34.600657°N, 32.971419°E, 24.V.2017,
(1♀); Akrotiri, 34.600657°N, 32.971419°E, 15.VI.2017, (1♀); Akrotiri, 34.600657°N,
32.971419°E, 26.IV.2018, (1♀); Akrotiri, 34.601506°N, 32.986197°E, 26.IV.2018,
(1♂) Yermasoyia Dam, 34.755799°N, 33.096194°E, 7.III.2017, (1♂), all 20 km
NNW Paphos, Lara Beach, Mavrokolympos Reservoir and Sovereign Base Area, 8 km
S Limassol, Akrotiri (near Airbase) records C. Schmid-Egger leg., A. Müller det., all
Asprokremmos Dam and Sovereign Base Area, Akrotiri, Bishop's Pool records S.P.M.
Roberts leg., A. Müller det., all Yermasoyia Dam records S.P.M. Roberts leg., G. Le
Go det., all Cherkes and Akrotiri records A. Varnava leg., A. Müller det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria), Northern Africa, Western Asia.
Hoplitis (Anthocopa) cypriaca (Mavromoustakis, 1938)
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 21.IV.1938,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1938, 1949 [“1948”], 1952, 1953, 1954).
Mavromoustakis localities. Limassol, Evdhimou River, Near Enkomi of Fama-
gusta, Xylophagou, Arakapas, Deryneia.
Material examined. Limassol District: Akrotiri, 34.583676°N, 32.949306°E,
27.IV.2018, A. Varnava leg, A. Müller det. (1♀).
Distribution. Cyprus, Western Asia (Israel, Jordan, Syria).
Hoplitis (Anthocopa) fasciculata (Alfken, 1934)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis (1952).
Mavromoustakis localities. Cherkes, Yermasoyia River, Younaros of Zakaki,
Deryneia.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (4♀); Limassol District: Akrotiri, 34.583676°N, 32.949306°E,
24.V.2017, (1♂); Akrotiri, 34.583676°N, 32.949306°E, 26.IV.2018, (1♀), all Paphos
district records C. Schmid-Egger leg., A. Müller det., all Limassol district records A.
Varnava leg., A. Müller det.
Distribution. Cyprus, Southern Europe (Italy, Greece), Western Asia, Southern
Asia (Iran).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
56
Notes. Described from Cyprus as Osmia idalia Mavromoustakis, 1949 [“1948"],
from Yermasoyia River: 34.7182°N, 33.08788°E, 5.VI.1938, G.A. Mavromoustakis
leg., G.A. Mavromoustakis det. ♀, (DAAN).
Hoplitis (Hoplitis) holmboei (Mavromoustakis, 1949)
Type locality–country. Cyprus, Yermasoyia River: 34.7182°N, 33.08788°E,
20.III.1937, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1949 [“1948”], 1952, 1957a).
Mavromoustakis localities. Polemedia Hills, Yermasoyia River, Near Amathus.
Material examined. Limassol District: Almirolivado, 34.9333°N, 32.9004°E, 14-
16.V.2012, S. Dimitriou leg., A. Müller det. (1♂), pan trap (UAEG).
Distribution. Cyprus, Southern Europe (Greece).
Hoplitis (Alcidamea) limassolica (Mavromoustakis, 1937)
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 18.III.1935,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1937b, 1938, 1949 [“1948”], 1951, 1952, 1953,
1957a).
Mavromoustakis localities. Polemedia Hills, Yerasa, Kellaki, Trimiklini, Fama-
gusta, Ayia Varvara (Stavrovouni).
Material examined. Limassol District: Polemidia, 34.71178°N, 33.004775°E,
8.III.2017, Bee Course students leg., A. Müller det. (1♂).
Distribution. Cyprus, Northern Africa, Western Asia.
Hoplitis (Pentadentosmia) pomarina (Warncke, 1991)
References. Müller (2018).
Material examined. C. Schmid-Egger leg., A. Müller det., 2013.
Distribution. Cyprus, Southern Europe (North Macedonia, Greece), Western
Asia.
Hoplitis (Anthocopa) yermasoyiae (Mavromoustakis, 1938)
References. Mavromoustakis (1938, 1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Polemedia Hills, Potamitissa, Yerasa, Kellaki, Near
Eftagonia, Ayia Irini Station, Eleousa Monastery (Karpasian Peninsula).
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 5-7.
IV.2012, (6♂, 2♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 25-27.
IV.2012, (14♂, 10♀), pan trap (UAEG); Limassol District: 1 km E of Pissouri,
34.677579°N, 32.722066°E, 27.IV.2015, (1♀); 0.7 km N of Anogyra, 34.745537°N,
32.73385°E, 3.V.2015, (1♀); Anogyra to Pachna Road km 4, 34.764269°N,
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 57
32.757736°E, 5.V.2015, (2♀), visiting Asteraceae; Akrotiri, 34.583676°N,
32.949306°E, 23.IV.2016, (1♀); Akrotiri, 34.600657°N, 32.971419°E, 23.IV.2016,
(1♀); Yermasoyia Dam, 34.755799°N, 33.096194°E, 7.III.2017, (1♂); Akrotiri,
34.601506°N, 32.986197°E, 7.IV.2017, (1♀); Akrotiri, 34.601506°N, 32.986197°E,
21.IV.2017, (1♂); Akrotiri, 34.601506°N, 32.986197°E, 14.IV.2018, (1♀, 1♂);
Akrotiri, 34.601506°N, 32.986197°E, 26.IV.2018, (2♀); Paphos District: N of Elia
Bridge, 34.900977°N, 32.776759°E, 29.IV.2015, (1♂), all Nicosia district records S.
Dimitriou leg., A. Müller det., all 1 km E of Pissouri, 0.7 km N of Anogyra, Anogyra
to Pachna Road km 4 and Paphos district records S.P.M. Roberts leg., A. Müller det.,
all Akrotiri records A. Varnava leg., A. Müller det., all Yermasoyia Dam records Bee
Course students leg., A. Müller det.
Distribution. Cyprus, Southern Europe (North Macedonia, Albania, Greece),
Western Asia.
Notes. e subspecies described from Cyprus is Hoplitis yermasoyiae yermasoyiae,
from Yermasoyia: 34.7182°N, 33.08788°E, 21.III.1937, G.A. Mavromoustakis leg./
det. ♂, (DAAN).
Genus Osmia PANZER, 1806
25 species.
Osmia (Pyrosmia) amathusica Mavromoustakis, 1937
Type locality–country. Cyprus, Amathus ruins: 34.7125°N, 33.1419°E, 26.III.1935,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1937b, 1949 [“1948”], 1957a).
Mavromoustakis localities. Polemedia Hills, Choirokitia.
Material examined. Limassol District: Polemidia, 34.71178°N, 33.004775°E,
8.III.2017, Bee Course students leg., A. Müller det. (1♂).
Distribution. Cyprus, Southern Europe (Italy, Greece), Western Asia.
Osmia (Erythrosmia) andrenoides Spinola, 1808
References. Mavromoustakis (1939b, 1949 [“1948”], 1951).
Mavromoustakis localities. Limassol, Polemedia Hills.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, West-
ern Asia.
Osmia (Osmia) bicornis (Linnaeus, 1758)
References. Mavromoustakis (1939b, 1949 [“1948”], 1951, 1952, 1953).
Mavromoustakis localities. Apsiou, Yerasa, Trimiklini, Pera Pedi, Mt. Troodos
open slopes, Listovounos Kakopetria.
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58
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 25-27.
IV.2012, S. Dimitriou leg., J. Devalez and A. Müller det. (1♂), pan trap (UAEG).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Turkmenistan, Kazakhstan, Kyrgyzstan), Southern Asia (Iran).
Osmia (Metallinella) brevicornis (Fabricius, 1798)
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran, Afghanistan).
Notes. Records pertain to ssp. leucogastra Morawitz, 1875.
Osmia (Helicosmia) caerulescens (Linnaeus, 1758)
References. Mavromoustakis (1939b, 1949 [“1948”]); Pittioni (1950); Mavromousta-
kis (1951, 1952, 1953, 1957a).
Mavromoustakis localities. Episkopi Forest, Potamitissa, Trimiklini, Platres, Ay-
ios Ioannis tou Agrou, Asbestotos spring near Amathus, Platania Forest Station, Mt.
Troodos open slopes, Livadin of Cedars (Paphos Forest).
Material examined. Limassol District: Amiantos, 34.918°N, 32.9472°E, 25-27.
IV.2012, (1♂), pan trap (UAEG); Troodos, Chionistra, 34.9317°N, 32.8664°E, 14-
16.V.2012, (1♂), pan trap (UAEG); Yermasoyia Dam, 34.755799°N, 33.096194°E,
7.III.2017, (1♂); Sovereign Base Area, Akrotiri, Bishop's Pool, 34.597305°N,
32.984521°E, 10.III.2017, (2♂); Sovereign Base Area, Episkopi, Kensington Clis,
34.670772°N, 32.846923°E, 4.V.2015, (1♀); Paphos District: 8 km N Paphos, Ma-
vrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (1♀); Famagusta District:
Cape Greco, 34.963264°N, 34.066211°E, 15.III.2017, (1♂), all Amiantos records S.
Dimitriou leg., J. Devalez and A. Müller det., all Troodos, Chionistra records S. Dimi-
triou leg., A. Müller det., all Yermasoyia Dam, Sovereign Base Area, Akrotiri, Bishop's
Pool and Famagusta district records S.P.M. Roberts leg., G. Le Go det., all Sovereign
Base Area, Episkopi, Kensington Clis records S.P.M. Roberts leg., A. Müller det., all
Paphos district records C. Schmid-Egger leg., A. Müller det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (India), Eastern Asia, New Zealand.
Osmia (Pyrosmia) cephalotes Morawitz, 1870
References. Mavromoustakis (1951, 1952, 1953).
Mavromoustakis localities. Nicosia, Apsiou, Yerasa, Trimiklini, Platania Forest
Station, Kykkou Monastery, Prodromos, Rizokarpaso.
Material examined. Limassol District: Anogyra, 34.73663°N, 32.732715°E,
26.IV.2015, M. Jenner leg., G. Le Go det. (1♀).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 59
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa, Western Asia, Central Asia (Turkmenistan, Uzbekistan), South-
ern Asia (Iran).
Osmia (Osmia) cornuta (Latreille, 1805)
References. Mavromoustakis (1938, 1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Apsiou, Potamitissa, Kitromili near Polemedia.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa, Western Asia, Central Asia (Turkmenistan, Kazakhstan), Southern Asia
(Iran).
Notes. e subspecies described from Cyprus is Osmia cornuta neoregaena Mav-
romoustakis, 1938, from Saittas, 762m: 34.8708333°N, 32.9166667°E, 10.III.1937,
G.A. Mavromoustakis leg./det. ♀, (DAAN).
Osmia (Pyrosmia) cyanoxantha Pérez, 1879
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Near Limassol, Mesayitonia, Yermasoyia River, Ama-
thus, Akrounda.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria), Northern Africa, Western Asia, Southern Asia (Iran).
Osmia (Helicosmia) dimidiata Morawitz, 1870
References. Mavromoustakis (1949 [“1948”], 1952, 1957a).
Mavromoustakis localities. Limassol, Mesayitonia, Apsiou, Famagusta, Near En-
komi of Famagusta, Deryneia, Near Trimiklini.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria, Ukraine, Russian Federation), Northern Africa, Western Asia, Central
Asia (Kyrgyzstan), Southern Asia (Iran).
Osmia (Helicosmia) dives Mocsáry, 1877
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Limassol, Episkopi, Fassouri, Yermasoyia River, Ama-
thus, Mesayitonia.
Material examined. Limassol District: Amiantos, 34.918°N, 32.9472°E, 25-27.
IV.2012, (9♂), pan trap (UAEG); 1 km E of Pissouri, 34.677579°N, 32.722066°E,
27.IV.2015, (1♀), collecting pollen at Echium angustifolium (Boraginaceae); Cherkes,
34.641933°N, 32.963433°E, 26.IV.2018, (1♂); Akrotiri, 34.601506°N, 32.986197°E,
19.IV.2016, (1♀); Akrotiri, 34.628817°N, 32.940667°E, 30.III.2018, (1♀); Akrotiri,
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60
34.583676°N, 32.949306°E, 14.IV.2018, (2♀); Akrotiri, 34.583676°N, 32.949306°E,
27.IV.2018, (1♀); Paphos District: 1.75 km N of Ag. Georgios, 34.911736°N,
32.327703°E, 30.IV.2015, (1♀), visiting Asteraceae, all Amiantos records S. Dimi-
triou leg., J. Devalez and A. Müller det., all 1 km E of Pissouri and Paphos district
records S.P.M. Roberts leg., A. Müller det., all Cherkes and Akrotiri records A. Varnava
leg., A. Müller det.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Hungary, Bul-
garia), Northern Africa, Western Asia, Central Asia, Southern Asia (Iran).
Osmia (Erythrosmia) erythrogastra Ferton, 1905
References. Müller (2018).
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, Bee Course students leg., A. Müller det. (1♂).
Distribution. Cyprus, Western Europe (France), Southern Europe, Western Asia.
Osmia (Pyrosmia) ferruginea Latreille, 1811
References. Mavromoustakis (1939b, 1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Cape Gata, Larnaca, Morphou, Near Deryneia.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (3♀); Yermasoyia Dam, 34.755799°N, 33.096194°E,
7.III.2017, (1♂); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017, (2♀); Polemidia,
34.71178°N, 33.004775°E, 8.III.2017, (2♂, 10♀); Sovereign Base Area, Avdimou Bay
Clis, 34.656698°N, 32.773339°E, 10.III.2017, (3♀), visiting Onobrychis (Fabaceae);
Akrotiri, 34.601506°N, 32.986197°E, 25.III.2017, (1♂); Akrotiri, 34.583676°N,
32.949306°E, 25.III.2017, (1♂); Akrotiri, 34.600657°N, 32.971419°E, 1.III.2018,
(1♂); Akrotiri, 34.5886°N, 32.9389°E, 1.III.2018, (1♀); Akrotiri, 34.628817°N,
32.940667°E, 1.III.2018, (1♂); Larnaca District: Koshi, 34.96663°N, 33.544391°E,
9.III.2018, (1♀, 1♂), visiting Onobrychis venosa, Yermasoyia Dam (7.III.2017, 3♀),
Polemidia (8.III.2017, 2♀) and Sovereign Base Area, Avdimou Bay Clis records
S.P.M. Roberts leg., G. Le Go det., Yermasoyia Dam (7.III.2017, 1♂) and Polemidia
(8.III.2017, 2♂, 10♀) records Bee Course students leg., A. Müller det., all Akrotiri
and Larnaca district records A. Varnava leg., A. Müller det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Northern Af-
rica, Western Asia.
Osmia (Pyrosmia) hellados van der Zanden, 1984
References. Müller (2018).
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (1♂); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017,
(5♂, 2♀) all records Bee Course students leg., A. Müller det.
Distribution. Cyprus, Southern Europe, Eastern Europe (Bulgaria), Western Asia.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 61
Osmia (Helicosmia) latreillei (Spinola, 1806)
References. Mavromoustakis (1939b, 1951, 1952, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Cherkes, Amathus, sand dunes of Ayios
Memnon (2 miles from Famagusta).
Material examined. Limassol District: Anogyra, 34.73663°N, 32.732715°E,
26.IV.2015, (1♀); 0.7 km N of Anogyra, 34.745537°N, 32.73385°E, 3.V.2015, (1♀);
Polemidia, 34.71178°N, 33.004775°E, 8.III.2017, (6♂, 2♀); Cherkes, 34.643188°N,
32.995281°E, 25.III.2017, (1♂); Cherkes, 34.643188°N, 32.995281°E, 7.IV.2017,
(1♀); Cherkes, 34.653067°N, 32.974233°E, 1.III.2018, (1♂); Cherkes, 34.659801°N,
32.99051°E, 16.III.2018, (1♀); Cherkes, 34.65096°N, 32.99091°E, 16.III.2018,
(1♀); Cherkes, 34.65096°N, 32.99091°E, 30.III.2018, (1♀); Cherkes, 34.653067°N,
32.974233°E, 30.III.2018, (1♀); Paphos District: Aspro Pools, 34.720825°N,
32.551994°E, 30.IV.2015, (1♀), all Anogyra and Paphos district records M. Jenner
leg., G. Le Go det., all Polemidia records Bee Course students leg., A. Müller det., all
Cherkes records A. Varnava leg., A. Müller det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Northern Af-
rica, Western Asia, Southern Asia (Iran).
Osmia (Hoplosmia) ligurica Morawitz, 1868
References. Mavromoustakis (1939b, 1949 [“1948”], 1951, 1952, 1953).
Mavromoustakis localities. Evdhimou River, Potamitissa, Yerasa, Eftagonia, Tri-
miklini, Arakapas, sand dunes of Ayios Memnon.
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe, Northern Africa, Western Asia, Central Asia (Turkmenistan), Southern
Asia (Iran).
Osmia (Helicosmia) melanogaster Spinola, 1808
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe, Northern Africa, Western Asia, Central Asia (Turkmenistan, Uzbekistan),
Southern Asia (Iran), Eastern Asia (China).
Osmia (Pyrosmia) nana Morawitz, 1874
References. Mavromoustakis (1951).
Mavromoustakis localities. Cape Apostolos Andreas.
Distribution. Cyprus, Southern Europe (Italy, Croatia, Greece), Eastern Europe
(Bulgaria, Russian Federation), Western Asia, Central Asia (Turkmenistan), Southern
Asia (Iran).
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62
Osmia (Helicosmia) niveata (Fabricius, 1804)
References. Mavromoustakis (1949 [“1948”]); Georghiou (1977).
Mavromoustakis localities. Episkopi, Potamitissa.
Material examined. Limassol District: Amiantos, 34.918°N, 32.9472°E,
25.iv.2012, 27.iv.2012, (2♂), pan trap (UAEG); Sovereign Base Area, Akrotiri, Bish-
op's Pool, 34.597305°N, 32.984521°E, 28.IV.2015, (6♀), collecting pollen on Aster-
aceae (Cardueae); Anogyra, 34.741952°N, 32.734845°E, 28.IV.2015, (1♂); Cherkes,
34.65096°N, 32.99091°E, 17.IV.2016, (1♀); Cherkes, 34.65096°N, 32.99091°E,
16.III.2018, (1♀); Cherkes, 34.65096°N, 32.99091°E, 14.IV.2018, (1♀); Cherkes,
34.65096°N, 32.99091°E, 26.IV.2018, (2♀); Akrotiri, 34.601506°N, 32.986197°E,
16.III.2018, (1♂), all Amiantos records S. Dimitriou leg., J. Devalez and A. Müller
det., all Sovereign Base Area, Akrotiri, Bishop's Pool records S.P.M. Roberts leg., A.
Müller det., all Anogyra records M. Jenner leg., G. Le Go det., all Cherkes and Ak-
rotiri records A. Varnava leg., A. Müller det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Southern Asia (Iran), Eastern Asia (China).
Osmia (Pyrosmia) saxicola Ducke, 1899
References. Mavromoustakis (1937b, 1939b, 1949 [“1948”], 1951, 1952, 1953).
Mavromoustakis localities. Near Pano Kivides, Platania Forest Station, Yerasa
Hills, hills near Paramytha.
Distribution. Cyprus, Western Europe (France), Southern Europe, Western Asia,
Central Asia (Tajikistan), Southern Asia (Iran).
Notes. Described from Cyprus as Osmia cypricola Mavromoustakis, 1937, from
Pera Pedi: 34.859444°N, 32.876111°E, 20.V.1929, G.A. Mavromoustakis leg./det.
♀, (DAAN).
Osmia (Hoplosmia) scutellaris Morawitz, 1868
References. Mavromoustakis (1939b, 1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Polemedia Hills, Akrotiri Forest, Cape Apostolos An-
dreas, Evdhimou River, Potamitissa, Yerasa.
Material examined. Limassol District: Cherkes, 34.65096°N, 32.99091°E,
9.IV.2016, (3♂); Akrotiri, 34.600657°N, 32.971419°E, 19.IV.2016, (1♀); Akrotiri,
34.600657°N, 32.971419°E, 8.V.2016, (1♀); Akrotiri, 34.600657°N, 32.971419°E,
5.V.2017, (1♂); Cherkes, 34.659801°N, 32.99051°E, 26.IV.2018, (1♀), all records A.
Varnava leg., A. Müller det.
Distribution. Cyprus, Western Europe (France, Switzerland), Southern Europe,
Eastern Europe, Northern Africa, Western Asia, Southern Asia (Iran).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 63
Osmia (Helicosmia) signata Erichson, 1835
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Georghiou (1977).
Mavromoustakis localities. Limassol, Fassouri, Randidi Forest.
Material examined. Paphos District: 20 km N Paphos, Kathikas, 34.90°N, 32.42°E,
20.VI.2013, (1♀); Limassol District: Cherkes, 34.65096°N, 32.99091°E, 14.IV.2018,
(2♀); Cherkes, 34.65096°N, 32.99091°E, 26.IV.2018, (1♀); Akrotiri, 34.583676°N,
32.949306°E, 10.III.2017, (1♂); Akrotiri, 34.583676°N, 32.949306°E, 26.IV.2018,
(3♀); Akrotiri, 34.583676°N, 32.949306°E, 28.IV.2018, (2♀), all Paphos district re-
cords C. Schmid-Egger leg., A. Müller det., all Limassol district records A. Varnava
leg., A. Müller det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Ukraine), Northern Africa, Western Asia, Central Asia (Turkmenistan), South-
ern Asia (Iran), Eastern Asia (China).
Osmia (Pyrosmia) submicans Morawitz, 1870
References. Mavromoustakis (1939b, 1949 [“1948”], 1951, 1957a).
Mavromoustakis localities. Akrotiri forest, Near Platania Forest Station,
Potamitissa, Cape Apostolos Andreas, Morphou, Choirokitia, Deryneia.
Material examined. Limassol District: Sovereign Base Area, Akrotiri, Bishop's Pool,
34.597305°N, 32.984521°E, 28.IV.2015, (1♀); Anogyra, 34.741952°N, 32.734845°E,
28.IV.2015, (1♀); Anogyra, 34.741952°N, 32.734845°E, 3.V.2015, (1♀); Ano-
gyra, 34.724979°N, 32.737225°E, 1.IV.2016, (1♂); Troodos, Kannoures Spring,
34.940989°N, 32.872738°E, 21.VIII.2016, (1♂); Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (1♀); Yermasoyia Dam, 34.755799°N, 33.096194°E,
7.III.2017, (1♂); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017, (1♀); Polemidia,
34.71178°N, 33.004775°E, 8.III.2017, (7♀); Akrotiri, 34.601506°N, 32.986197°E,
26.IV.2018, (1♀), all Sovereign Base Area, Akrotiri, Bishop's Pool records S.P.M. Rob-
erts leg., A. Müller det., Anogyra records M. Jenner leg., G. Le Go det., Anogyra
(1.IV.2016, 1♂), Troodos, Kannoures Spring and Akrotiri records A. Varnava leg., A.
Müller det., Yermasoyia Dam (7.III.2017, 1♀) and Polemidia (8.III.2017, 1♀) records
S.P.M. Roberts leg., G. Le Go det., Yermasoyia Dam (7.III.2017, 1♂) and Polemidia
(8.III.2017, 7♀) records Bee Course students leg., A. Müller det.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa, Western Asia, Central Asia (Kazakhstan).
Osmia (Allosmia) sybarita Smith, 1853
References. Mavromoustakis (1939b, 1949 [“1948”]).
Mavromoustakis localities. Polemedia Hills, Near Paramytha, Randidi Forest,
Near Deryneia.
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64
Material examined. Limassol District: Filitos, 34.800043°N, 33.000918°E,
1.IV.2016, (1♀); Akrotiri, 34.600657°N, 32.971419°E, 23.IV.2016, (1♀); Akrotiri,
34.583676°N, 32.949306°E, 16.III.2018, (1♂); Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (1♂); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017,
(1♂, 3♀), all Filitos and Akrotiri records A. Varnava leg., A. Müller det., all Yermasoyia
Dam and Polemidia records Bee Course students leg., A. Müller det.
Distribution. Cyprus, Southern Europe (Albania, Greece), Eastern Europe (Bul-
garia), Western Asia.
Osmia (Pyrosmia) teunisseni van der Zanden, 1981
References. Ungricht, Müller and Dorn (2008).
Distribution. Cyprus, Southern Europe (Italy, Croatia, Greece), Western Asia.
Osmia (Pyrosmia) viridana Morawitz, 1874
References. Mavromoustakis (1939b, 1949 [“1948”], 1951, 1952, 1953, 1957a).
Mavromoustakis localities. Polemedia Hills, Trimiklini, Near Paramytha, Near
Pano Kivides, Near Cape Gata, Morphou, Panagia ton Katharadon Monastery (Kyrenia).
Material examined. Limassol District: 0.7 km N of Anogyra, 34.745537°N,
32.73385°E, 3.V.2015, (1♀), visiting Trifolium (Fabaceae); Anogyra, 34.741952°N,
32.734845°E, 3.V.2015, (2♀); Anogyra to Pachna Road km 4, 34.764269°N,
32.757736°E, 5.V.2015, (2♀); Filitos, 34.800043°N, 33.000918°E, 1.IV.2016, (1♀);
Polemidia, 34.71178°N, 33.004775°E, 8.III.2017, (2♂); Polemidia, 34.71178°N,
33.004775°E, 8.III.2017, (5♂, 1♀), all 0.7 km N of Anogyra and Anogyra to Pachna
Road km 4 records S.P.M. Roberts leg., A. Müller det., all Anogyra records M. Jen-
ner leg., G. Le Go det., all Filitos records A. Varnava leg., A. Müller det., Polemidia
(8.III.2017, 2♂) records S.P.M. Roberts leg., G. Le Go det., Polemidia (8.III.2017,
5♂, 1♀) records Bee Course students leg., A. Müller det.
Distribution. Cyprus, Western Europe (France, Italy, Germany), Southern Eu-
rope, Eastern Europe (Bulgaria, Ukraine, Russian Federation), Northern Africa, West-
ern Asia, Central Asia, Southern Asia (Iran).
Notes. e subspecies described from Cyprus is Osmia viridana nicosiana Mavro-
moustakis, 1939, from Ayia Fyla: 34.71947°N, 33.01962°E, 22.II.1937, G.A. Mavro-
moustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
Genus Protosmia DUCKE, 1900
3 species.
Protosmia (Protosmia) glutinosa (Giraud, 1871)
References. Mavromoustakis (1951).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 65
Mavromoustakis localities. Mt. Troodos.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria), Northern Africa, Western Asia (Turkey).
Protosmia (Protosmia) monstrosa (Pérez, 1895)
References. Mavromoustakis (1939b).
Distribution. Cyprus, Southern Europe (Greece), Northern Africa, Western Asia.
Protosmia (Protosmia) paradoxa (Friese, 1899)
References. Mavromoustakis (1939b, 1949 [“1948”]); Pittioni (1950).
Mavromoustakis localities. Cape Apostolos Andreas, Eleousa Monastery (Karpasia).
Material examined. Limassol District: 1 km E of Pissouri, 34.677579°N,
32.722066°E, 27.IV.2015, (1♂); Sovereign Base Area, Akrotiri, Bishop's Pool,
34.597305°N, 32.984521°E, 28.IV.2015, (1♀), collecting pollen on Echium angus-
tifolium (Boraginaceae); 2 km N of Anogyra, 34.748126°N, 32.732248°E, 1.V.2015,
(1♂), visiting Asteraceae; Anogyra to Pachna Road km 4, 34.764269°N, 32.757736°E,
5.V.2015, (1♂, 1♀), visiting Asteraceae; Filitos, 34.800043°N, 33.000918°E,
1.IV.2016, (2♂); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017, (1♂), all 1 km
E of Pissouri, Sovereign Base Area, Akrotiri Bishop's Pool, 2 km N of Anogyra and
Anogyra to Pachna Road km 4 records S.P.M. Roberts leg., A. Müller det., all Filitos
records A. Varnava leg., A. Müller det., all Polemidia records S.P.M. Roberts leg., G.
Le Go det.
Distribution. Cyprus, Southern Europe (Greece), Western Asia.
ANTHIDIINI
Genus Anthidiellum COCKERELL, 1904
2 Species.
Anthidiellum (Anthidiellum) breviusculum (Mavromoustakis, 1949)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1957a).
Mavromoustakis localities. Mt. Troodos, Livadin of Cedars (Paphos Forest),
Ayios Ilarion, Xerokolimbi (near Trooditissa), Platania Forest, Mt. Troodos Chionistra.
Distribution. Cyprus, Southern Europe (Spain, Greece), Eastern Europe (Bul-
garia, Romania), Northern Africa (Tunisia), Western Asia (Turkey, Lebanon, Israel),
Southern Asia (Iran).
Notes. e subspecies described from Cyprus is Anthidiellum breviusculum troodi-
cum Mavromoustakis, 1949 [“1948”], from Mt. Troodos, 1524 m: 34.908°N, 32.866°E,
14.VI.1935, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
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66
Anthidiellum (Anthidiellum) strigatum (Panzer, 1805)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a).
Mavromoustakis localities. Limassol, Cherkes, Erimi, Alassa River, Chiicoudia
marshes, Livadin of Cedars (Paphos Forest), Ayia Irini (Kyrenia), Moni River, Finika-
ria, Symboulas, Paramali, Zakaki, Asomatos, Salipes marshes near Akrotiri.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran), Eastern Asia.
Genus Anthidium FABRICIUS, 1804
5 species.
Anthidium (Anthidium) cingulatum Latreille, 1809
References. Mavromoustakis (1949 [“1948”], 1952, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Akrotiri Forest and Bay,
Asomatos, Yermasoyia River, Eftagonia, Lania, Cape Gata, Trimiklini, Platres, Salipes
marshes, Curium (near Episkopi), Episkopi, Moni, Evdhimou River, Near Platania
Forest Station, Livadin of Cedars (Paphos Forest), Mt. Troodos, Symboulas Forest,
Ayia Irini Station, Kykkou Monastery, Famagusta.
Material examined. Limassol District: Sovereign Base Area, Episkopi, Kensington
Clis, 34.670772°N, 32.846923°E, 4.V.2015, S.P.M. Roberts leg., J. Praz det. (1♀),
visiting Lamiaceae.
Distribution. Cyprus, Western Europe (France, Austria, Germany), Southern Eu-
rope, Eastern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia
(Iran), Eastern Asia (China).
Anthidium (Anthidium) diadema Latreille, 1809
References. Mavromoustakis (1949 [“1948”], 1951, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Mesayitonia, Ayios Atha-
nasios, Yermasoyia River, Younarka (near Zakaki), Curium (near Episkopi), Near
Akhyritou, Pernera coast of Paralimni.
Material examined. Limassol District: Sovereign Base Area, Episkopi, Kensington
Clis, 34.670772°N, 32.846923°E, 4.V.2015, S.P.M. Roberts leg., J. Praz det. (1♀),
visiting Asteraceae.
Distribution. Cyprus, Western Europe (France, Belgium), Southern Europe,
Eastern Europe (Bulgaria), Northern Africa (Morocco, Algeria,), Central Asia, South-
ern Asia (Iran), Eastern Asia (China).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 67
Anthidium (Anthidium) orentinum (Fabricius, 1775)
References. Cockerell (1910); Mavromoustakis (1949 [“1948”], 1957).
Mavromoustakis localities. Ayios Ioannis (Agros), Odou, Stavros (Paphos For-
est), Krios River, Zakaki.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa, Western Asia, Central Asia, Southern Asia (Iran), Eastern Asia (China).
Notes. Described from Cyprus as Anthidium orentinum cypriacum Mavromousta-
kis, 1949 ["1949 [“1948”]"], from Agios Ioannis, 34.86777°N, 32.69027°E, VII.1930,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
Anthidium (Anthidium) loti Perris, 1852
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Episkopi, Chii-
coudia marshes, Pera Pedi, Fassouri, Moni River, Zakaki, Episkopi Forest.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Hungary, Bulgaria, Romania), Western Asia (Turkey), Central Asia, Southern
Asia (Iran).
Anthidium (Proanthidium) undulatum Dours, 1873
References. Mavromoustakis (1939a, 1949 [“1948”], 1951, 1952, 1957a).
Mavromoustakis localities. Limassol, Yermasoyia River, Eftagonia, Mesayitonia,
Polemedia Hills, Episkopi, Erimi, Ayia Varvara (Stavrovouni), Famagusta, Glypha
(near Akanthou), Mt. Troodos.
Distribution. Cyprus, Western Europe (France), Southern Europe (Croatia, North
Macedonia, Greece), Northern Africa (Algeria), Western Asia, Southern Asia (Iran).
Notes. e subspecies described from Cyprus is Anthidium undulatum holozonium
(Mavromoustakis, 1939), from Limassol: 34.66839°N, 33.03252°E, G.A. Mavro-
moustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
Genus Icteranthidium MICHENER, 1948
2 species.
Icteranthidium ferrugineum (Fabricius, 1787)
References. Mavromoustakis (1949 [“1948”], 1951).
Mavromoustakis localities. Akrotiri Bay, Famagusta, Ayios Memnon, Yermasoyia
River, Near Episkopi.
Distribution. Cyprus, Southern Europe (Spain), Northern Africa (Morocco, Tu-
nisia to Egypt), Western Africa (Mauritania), Western Asia, Central Asia (Turkmeni-
stan, Kazakhstan), Southern Asia (Iran), Eastern Asia (China).
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68
Icteranthidium grohmanni (Spinola, 1838)
References. Pittioni (1950); Mavromoustakis (1957a).
Mavromoustakis localities. Mt. Troodos, Yermasoyia River.
Distribution. Cyprus, Southern Europe, Eastern Europe (Bulgaria, Ukraine),
Northern Africa (from Morocco to Egypt), Western Asia, Central Asia (Kyrgyzstan),
Southern Asia (Iran), Eastern Asia (China).
Genus Rhodanthidium ISENSEE, 1927
1 species.
Rhodanthidium (Rhodanthidium) septemdentatum (Latreille, 1809)
References. Mavromoustakis (1949 [“1948”], 1951).
Mavromoustakis localities. Polemedia Hills, Northern Mountains Kantara, Near
Amathus, Episkopi Forest, Symboulas Forest.
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, West-
ern Asia.
Genus Eoanthidium POPOV, 1950
1 species.
Eoanthidium (Eoanthidium) insulare (Morawitz, 1874)
References. Mavromoustakis (1937a, 1949 [“1948”], 1951, 1952, 1953, 1957a).
Mavromoustakis localities. Polemedia Hills, Moni River, Evdhimou River, Mt.
Troodos Kannoures springs, Mt. Troodos open slopes, Livadin of Cedars (Paphos For-
est), Platania Forest Station, Ayios Ilarion (Northern Mountain), Xerokolimbi (near
Trooditissa), Pasha Livadin (Mt. Troodos), Eagle’s Bath (Mt. Troodos).
Material examined. Limassol District: Almirolivado, 34.9333°N, 32.9004°E, 16-
18.IX.2011, S. Dimitriou leg., J. Devalez det. (1♀), pan trap (UAEG).
Distribution. Cyprus, Southern Europe (Croatia, Greece), Western Asia (Turkey),
Central Asia (Tajikistan).
Notes. Described from Cyprus as Dianthidium insulare var. lemesium Mavro-
moustakis, 1937, from Limassol: 34.66839°N, 33.03252°E, 26.VI.1931, G.A. Mav-
romoustakis leg./det. ♀, (DAAN).
Genus Stelis PANZER, 1806
2 species.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 69
Stelis (Stelis) murina Perez, 1884
References. Warncke (1992); Kasparek (2015).
Distribution. Cyprus, Southern Europe and Northern Africa.
Notes. Recorded as a subspecies of S. phaeoptera (Kirby, 1802) by Warncke (1992).
Stelis phaeoptera sensu stricto may occur on Cyprus as depicted on the map in Kasparek
(2015), but the area of overlap, hosts, and identication criteria of these forms remain
uncertain as noted in text of the revision and as conrmed by its author (M. Kasparek
in litt.). Pending further verication, we treat S. phaeoptera sensu stricto as unveried in
the supplementary text.
Stelis (Protostelis) signata (Latreille, 1809)
References. Popov (1944); Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Alassa River, Li-
vadin of Cedars (Paphos Forest), Near Akrotiri (village), Yermasoyia River.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Algeria,
Tunisia), Central Asia (Kazakhstan), Southern Asia (Iran).
DIOXYINI
Genus Stelis CAMERON, 1901
1 species.
Stelis tridentata (Nylander, 1848)
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Polemedia Hills (Limassol).
Distribution. Cyprus, Widespread in Europe, Western Asia, Central Asia (Kyr-
gyzstan), Eastern Asia (China).
Notes. Described from Cyprus as Dioxoides tridentata limassolica Mavromousta-
kis, 1949 [“1948”]"], from Polemedia Hills, 4 miles NW of Limassol: 34.7134°N,
32.9812°E, 20.V.1939, G.A. Mavromoustakis leg./det. ♀, (DAAN).
Genus Dioxys LEPELETIER & SERVILLE, 1825
2 species.
Dioxys cinctus (Jurine, 1807)
References. Popov (1944); Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Polemedia Hills, Ayia Phyla, Sphalangiotissa Monastery.
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70
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern Eu-
rope, Eastern Europe, Northern Africa, Western Asia (Israel), Central Asia (Uzbekistan).
Dioxys pumilus Gerstäcker, 1869
References. Popov (1944); Mavromoustakis (1949 [“1948”], 1957a).
Mavromoustakis localities. Limassol, Episkopi.
Distribution. Cyprus, Southern Europe (Spain, Italy, Greece), Northern Africa
(Morocco, Algeria, Tunisia), Western Asia (Turkey, Israel, Syria), Southern Asia (Iran).
Notes. Described from Cyprus as Dioxys cypriaca Popov, 1944, from Limassol:
34.66839°N, 33.03252°E, V.1935; IV.1936, G.A. Mavromoustakis leg., V.V. Popov
det. (ZISP).
MEGACHILINI
Genus Coelioxys LATREILLE, 1809
12 species.
Coelioxys (Allocoelioxys) acanthopyga Alfken, 1940
References. Alfken (1940); Mavromoustakis (1952).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Israel).
Notes. Described from Cyprus as Coelioxys carinulata Alfken, 1940, from Cherkes:
34.65°N, 32.975°E, 29.VI.1939, G.A. Mavromoustakis leg., J.D. Alfken det. ♀,
(MFNB).
Coelioxys (Allocoelioxys) acanthura (Illiger, 1806)
References. Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Limassol, Cherkes, Evdhimou River, Near Akrotiri.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Hungary, Ukraine), Northern Africa, Western Asia (Turkey).
Coelioxys (Allocoelioxys) afer Lepeletier, 1841
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a).
Material examined. Paphos District: 20 km N Paphos, Kathikas, 34.90°N,
32.42°E, 20.VI.2013, (2♀); Limassol District: Troodos, Mt. Olympos, 34.93°N,
32.86°E, 20.VI.2013, (2♀), all records C. Schmid-Egger leg., C. Schmid-Egger det.
Distribution. Cyprus, Widespread in Europe, Northern Africa (from Morocco to
Tunisia), Western Asia (Turkey, Oman), Central Asia, Southern Asia (Iran).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 71
Coelioxys (Allocoelioxys) argenteus Lepeletier, 1841
References. Mavromoustakis (1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Limassol, Yermasoyia River, Cherkes, Polemedia
Hills, Asomatos, Episkopi, Chiicoudia marshes, Fassouri, Ayia Irini (Kyrenia), Near
Akrotiri, Moni, Paramali, Near Enkomi of Famagusta.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Romania), Northern Africa, Western Asia (Turkey), Central Asia (Turkmeni-
stan), Southern Asia (Iran), Eastern Asia (China).
Coelioxys (Allocoelioxys) brevis Eversmann, 1852
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Limassol, Cherkes, Near Enkomi of Famagusta.
Distribution. Cyprus, Western Europe (France, Germany, Austria), Southern Eu-
rope, Eastern Europe, Northern Africa, Western Asia (Turkey), Central Asia, Southern
Asia (Iran), Eastern Asia (China).
Coelioxys (Melissoctonia) conoideus (Illiger, 1806)
References. Mavromoustakis (1949 [“1948”], 1952, 1957a).
Mavromoustakis localities. Limassol, Cherkes, Potamitissa, Odou, Erimi,
Trooditissa Monastery.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey), Southern Asia (Iran).
Coelioxys (Liothyrapis) decipiens Spinola, 1838
Remarks. Reported by Mavromoustakis under its junior synonym Paracoelioxys de-
cipiens.
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952, 1957).
Mavromoustakis localities. Limassol, Erimi, Cherkes, Famagusta.
Distribution. Cyprus, Southern Europe (Greece), Northern Africa, Western Asia
(Turkey), Central Asia, Southern Asia (Iran, India), Eastern Asia (China).
Coelioxys (Allocoelioxys) echinatus Förster, 1853
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa (Morocco, Algeria, Egypt), Western Asia (Turkey, Israel, Iraq), Southern
Asia (Iran, Afghanistan).
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72
Coelioxys (Allocoelioxys) elegantulus Alfken, 1934
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Chiicoudia
marshes, Yermasoyia River.
Distribution. Cyprus, Southern Europe (Greece), Northern Africa (Algeria,
Egypt), Western Asia (Turkey, Israel, Palestine).
Coelioxys (Allocoelioxys) haemorrhoa Förster, 1853
References. Alfken (1940); Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mav-
romoustakis (1952).
Mavromoustakis localities. Limassol, Cherkes, Asomatos, Chiicoudia marshes.
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Spain,
Greece), Eastern Europe (Romania), Northern Africa, Western Asia (Turkey, Israel),
Central Asia, Southern Asia (Iran, Pakistan, India), Eastern Asia (China).
Coelioxys (Paracoelioxys) inermis Kirby, 1802
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco, Al-
geria), Western Asia (Turkey), Central Asia (Uzbekistan, Kyrgyzstan), Southern Asia
(Iran), Eastern Asia (China).
Coelioxys (Allocoelioxys) polycentris Förster, 1853
References. Pittioni (1950).
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, Eastern
Europe, Southern Asia (Iran), Southern Asia (Iran, Pakistan), Eastern Asia (Mongolia).
Genus Megachile LATREILLE, 1802
17 species.
Megachile (Creightonella) albisecta (Klug, 1817)
References. Cockerell (1931); Mavromoustakis (1939b, 1949 [“1948”]); Pittioni
(1950); Mavromoustakis (1951, 1954, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Erimi, Evdhi-
mou River, Pera Pedi, Pissouri, Ayia Irini (Kyrenia), Near Akrotiri village, Yermasoyia
River, Moni River, Pernera coast of Paralimni, Younarka (near Zakaki), Near Ayios
eodoros (Pitsillia).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 73
Material examined. Paphos District: 20 km N Paphos, Kathikas, 34.90°N,
32.42°E, 20.VI.2013, C. Schmid-Egger leg., C. Praz det. (1♀, 1♂).
Distribution. Cyprus, Western Europe (France, Switzerland), Southern Europe,
Eastern Europe, Northern Africa, Western Asia, Central Asia, Eastern Asia (China).
Notes. e subspecies described from Cyprus is Megachile albisecta cyprica Cock-
erell, 1931, from Limassol: 34.66839°N, 33.03252°E, 15-17.VII.1928, G.A. Mavro-
moustakis leg., T.D.A. Cockerell det. ♀.
Megachile (Eutricharaea) anatolica Rebmann, 1968
References. Soltani et al. (2017).
Distribution. Cyprus, Southern Europe (Italy, Greece), Western Asia, Southern
Asia (Iran).
Megachile (Eutricharaea) apicalis Spinola, 1808
References. Mavromoustakis (1939b, 1949 [“1948”]); Pittioni (1950); Mavromousta-
kis (1951, 1952, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Near Akrotiri,
Paramali, Davlos coast, Tsada, Chiicoudia marshes, Livadin of Cedars (Paphos For-
est), Krios River (Kaledonia Falls), Glypha (near Akanthou), Younarka (near Zakaki),
Near Mesayitonia, Ayios Memnon near Famagusta, Younaros of Zakaki, Akrotiri (vil-
lage), Ayios Ilarion, Mt. Pentadactylos, Garillis River, Pachna.
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (1♀, 2♂); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E,
20.VI.2013, (1♀); 6 km W Polis, botanical garden, 35.03°N, 32.37°E, 20.VI.2013,
(1♀); Limassol District: 8 km S Limassol, Akrotiri (near Airbase), 34.60°N, 32.97°E,
20.VI.2013, C. (1♀, 1♂), all records Schmid-Egger leg., C. Praz det.
Distribution. Cyprus, Widespread in Europe, Western Asia, Central Asia (Uz-
bekistan), Southern Asia (Iran, Pakistan), Eastern Asia (China).
Megachile (Megachile) centuncularis (Linnaeus, 1758)
References. Mavromoustakis (1953, 1954, 1957a, b).
Mavromoustakis localities. Limassol, Cherkes, Famagusta, Platania Forest Sta-
tion, Platres, Pera Pedi, Kilani, Odou, Yermasoyia River, Chiicoudia marshes, Eagle’s
Bath (Mt. Troodos).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Kazakhstan), Southern Asia (Iran, India).
Notes. Described from Cyprus as Megachile centuncularis nesiotica Mavromousta-
kis, 1953, from Limassol: 34.66839°N, 33.03252°E, VIII.1936, G.A. Mavromousta-
kis leg., G.A. Mavromoustakis det. ♀, (DAAN).
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74
Megachile (Chalicodoma) cypricola Mavromoustakis, 1938
Type locality–country. Cyprus, Ayia Phyla: 34.7198°N, 33.0195°E, 23.III.1938,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1938, 1949 [“1948”], 1951, 1952, 1957a).
Mavromoustakis localities. Apsiou, Near Pano Kivides, Yerasa Hills, Choirokitia,
Ayia Napa.
Material examined. Limassol District: Sovereign Base Area, Avdimou Bay
Clis, 34.656698°N, 32.773339°E, 27.IV.2015, (1♀); Sovereign Base Area, Avdi-
mou Bay Clis, 34.656698°N, 32.773339°E, 27.IV.2015, (1♀); 3.5 km SE of Moni,
34.706873°N, 33.211916°E, 14.III.2017, (2♂, 1♀), visiting Onobrychis venosa (Fa-
baceae); 3.5 km SE of Moni, 34.706873°N, 33.211916°E, 14.III.2017, (2♀, 1♂),
visiting Onobrychis venosa (Fabaceae); 0.5 km W of Agios Georgios, 34.706786°N,
33.229364°E, 14.III.2017, (1♀); 0.5 km W of Agios Georgios, 34.706786°N,
33.229364°E, 14.III.2017, (2♀); Agios Georgios, 34.706786°N, 33.229364°E,
27.II.2018, (1♀, 3♂), visiting Onobrychis venosa (Fabaceae); Avdimou Bay Clis,
34.656698°N, 32.773339°E, 17.III.2018, (5♀), visiting Onobrychis venosa (Fabace-
ae); Paramali, 34.66198333°N, 32.80439444°E, 17.III.2018, (3♀), visiting Ono-
brychis venosa (Fabaceae); Larnaca District: 2.1 km S of Choirokoitia, 34.777521°N,
33.33622°E, 14.III.2017, (1♂); Choirokitia, 34.777521°N, 33.33622°E, 4.III.2018,
(1♀), Sovereign Base Area, Avdimou Bay Clis records M. Jenner leg., C. Praz det. and
S.P.M. Roberts leg., C. Praz det., 3.5 km SE of Moni (14.III.2017, 2♂, 1♀), 0.5 km
W of Agios Georgios (14.III.2017, 1♀) and 2.1 km S of Choirokoitia records S.P.M.
Roberts leg., S.P.M. Roberts det., 3.5 km SE of Moni (14.III.2017, 2♀, 1♂) and 0.5
km W of Agios Georgios (14.III.2017, 2♀) records A. Varnava leg., S.P.M. Roberts
det., Agios Georgios, Avdimou Bay Clis, Paramali, and Choirokitia (4.III.2018) re-
cords A. Varnava leg., A. Varnava det.
Distribution. Cyprus. ENDEMIC.
Megachile (Pseudomegachile) ericetorum (Lepeletier, 1841)
References. Georghiou (1977).
Material examined. Limassol District: Yermasoyia Dam, 34.745849°N,
33.083579°E, 28.X.2016, S.P.M. Roberts leg., G. Le Go det. (1♀), visiting Dittrichia
viscosa (Asteraceae).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Europe (Finland), Northern Africa, Western Asia, Southern Asia (Iran), Eastern
Asia (China).
Megachile (Pseudomegachile) farinosa Smith, 1853
References. Dorchin and Praz (2018).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 75
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (1♂); 8 km N Paphos, Mavrokolympos Reservoir, 34.85°N,
32.40°E, 20.VI.2013, (3♂), all records C. Schmid-Egger leg., C. Praz det.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Syria,
Iraq), Southern Asia (Iran).
Megachile (Eutricharaea) inexspectata Rebmann, 1968
Material examined. Paphos District: 6 km NE Polis, beach 35.06°N, 32.46°E,
20.VI.2013, C. Schmid-Egger leg., C. Praz det. (2♂).
Distribution. Cyprus, Northern Africa (Morocco, Algeria, Tunisia), Western Asia.
Megachile (Eutricharea) leachella Curtis, 1828
References. Georghiou (1977).
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (4♀, 3♂); 20 km N Paphos, Kathikas, 34.90°N, 32.42°E,
20.VI.2013, (1♂); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E, (12♀,
7♂); 6 km W Polis, botanical garden, 35.03°N, 32.37°E, 20.VI.2013, (1♀); 8 km N
Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E, 20.VI.2013, (2♀, 1♂); Polis,
35.053539°N, 32.351197°E, 30.X.2016, (1♂), visiting Dittrichia viscosa (Asteraceae);
Limassol District: Troodos, Mt. Olympos, 34.93°N, 32.86°E, (1♂); Yermasoyia Dam,
34.745849°N, 33.083579°E, 28.X.2016, (1♂, 4♀), visiting Dittrichia viscosa (Aster-
aceae); Sovereign Base Area, Akrotiri, 34.600657°N, 32.971419°E, 29.X.2016, (1♀),
visiting Dittrichia viscosa (Asteraceae); Sovereign Base Area, Paramali, 34.676011°N,
32.794947°E, 30.X.2016, (1♂), visiting Dittrichia viscosa (Asteraceae); Larnaca Dis-
trict: Zygi, 34.731233°N, 33.343487°E, 28.X.2016, (1♀), visiting Dittrichia viscosa
(Asteraceae); Famagusta District: Achna Dam, 35.05519°N, 33.814011°E, 28.X.2016,
(2♂, 1♀), visiting Dittrichia viscosa (Asteraceae), all 15 km SE Paphos Kouklia, 20 km
N Paphos Kathikas, 20 km NNW Paphos Lara Beach, 6 km W Polis botanical gar-
den, 8 km N Paphos Mavrokolympos Reservoir and Troodos Mt. Olympos records
C. Schmid-Egger leg., C. Praz det., Polis, Yermasoyia Dam, Sovereign Base Area Ak-
rotiri, Sovereign Base Area Paramali, Zygi, and Achna Dam records S.P.M. Roberts
leg., S.P.M. Roberts and G. Le Go det.
Distribution. Cyprus, Widespread in Europe, Central Asia.
Megachile (Eutricharaea) marginata Smith, 1853
References. Mavromoustakis (1939b).
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, C. Schmid-Egger leg., C. Praz det. (2♀, 2♂).
Distribution. Cyprus, Southern Europe, Northern Africa (Tunisia), Western Asia
(Turkey, Iraq), Central Asia (Kyrgyzstan, Tajikistan), Southern Asia (Afghanistan, Pakistan).
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Megachile (Megachile) melanopyga Costa, 1863
References. Mavromoustakis (1957a); Georghiou (1977).
Mavromoustakis localities. Platania Forest Station, Odou, Yermasoyia River,
Near Zakaki.
Distribution. Cyprus, Western Europe, Southern Europe (Spain, Italy, Slovenia),
Eastern Europe, Western Asia, Eastern Asia (China).
Notes. Described from Cyprus as Megachile melanopyga zakakica Mavromoustakis,
1957, from Platania Forest Station, Mt. Troodos: 34.9473°N, 32.9284°E, 5.IX.1951,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀.
Megachile (Eurymella) patellimana Spinola, 1838
References. Pittioni (1950); Mavromoustakis (1951, 1952).
Mavromoustakis localities. Famagusta, Near Akrotiri, Moni River, Akrotiri (vil-
lage), Ayios Memnon.
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, (10♀, 7♂); 6 km W Polis, botanical garden, 35.03°N, 32.37°E,
20.VI.2013, (1♂); 8 km N Paphos, Mavrokolympos Reservoir, 34.85°N, 32.40°E,
20.VI.2013, (1♂), all records C. Schmid-Egger leg., C. Praz det.
Distribution. Cyprus, Western Africa (Niger, Nigeria), Southern Africa (Namibia,
Botswana), Northern Africa (Egypt, Sudan), Eastern Africa (Mozambique), Western
Asia (Israel, Saudi Arabia, United Arab Emirates), Southern Asia (Pakistan).
Megachile (Eutricharea) pilidens Alfken, 1890
References. Cockerell (1931).
Material examined. Limassol District: Amiantos, 34.918°N, 32.9472°E, 15-17.
IX.2011, S. Dimitriou leg., J. Devalez det. (1♀), pan trap (UAEG).
Distribution. Cyprus, Northern Africa (Morocco), Western Europe, Southern
Europe, Eastern Europe, Central Asia.
Megachile (Eutricharaea) posti Mavromoustakis, 1952
Type locality–country. Cyprus, Polemedia Hills: 34.699444°N, 32.996944°E,
5.VII.1949, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1952).
Mavromoustakis localities. Polemedia Hills.
Distribution. Cyprus. ENDEMIC.
Notes. e status of this species has long been in question but Praz (2017) is of the
opinion that M. posti is a good species and distinct from M. basilaris Morawitz, 1875,
with which it has been synonymised in the past.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 77
Megachile (Chalicodoma) roeweri (Alfken, 1928)
References. Mavromoustakis (1939b, 1949 [“1948”], 1952, 1954).
Mavromoustakis localities. Limassol, Pera Pedi, Akrotiri (village).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey).
Notes. e subspecies described from Cyprus is Megachile roeweri akrotirica Ma-
vromoustakis, 1939, from Akrotiri Forest: 34.5843°N, 32.9676°E, 23.V.1938, G.A.
Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
Megachile (Eutricharaea) rotundata (Fabricius, 1787)
References. Mavromoustakis (1939b, 1952).
Mavromoustakis localities. Chiicoudia marshes, Erimi, Salipes marshes near
Akrotiri.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey, Georgia, Azerbaijan), Central Asia, Southern Asia (Iran, Pakistan, India),
Eastern Asia (China, Mongolia), Australia, New Zealand.
Megachile (Paracella) troodica Mavromoustakis, 1953
References. Mavromoustakis (1953); Zanden (1992).
Mavromoustakis localities. Mt. Troodos, Chionistra, Xerokolimbi.
Distribution. Cyprus. ENDEMIC.
Notes. Described from Cyprus as Megachile (Eutricharaea) mavromoustakisi van
der Zanden, 1992, from Mt. Troodos, 1676m (open slopes in the forest of Pinus nigra
pallasiana Lamb.): 34.9234°N, 32.8833°E, 7.VII.1936, G.A. Mavromoustakis leg.,
G.A. Mavromoustakis det. ♀ (DAAN).
APIDAE
ANTHOPHORINI
Genus Amegilla FRIESE, 1897
4 species.
Amegilla (Zebramegilla) albigena (Lepeletier, 1841)
References. Mavromoustakis (1949 [“1948”], 1951).
Mavromoustakis localities. Limassol, Polemedia Hills, Ayios Athanasios, Tsada,
Mt. Troodos, Chionistra, Kato Platres, Ayios Ioannis Agrou, Famagusta, Stroumbi,
Kathikas.
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 15-
17.IX.2011, (1♂), pan trap (UAEG); Limassol District: Almirolivado, 34.9333°N,
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
78
32.9004°E, 16-18.IX.2011, (4♀, 3♂), pan trap (UAEG), all records S. Dimitriou leg.,
J. Devalez det.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia
(Iran, Pakistan), Eastern Asia (China).
Amegilla (Micramegilla) glauca (Alfken, 1926)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1957a).
Mavromoustakis localities. Chiicoudia marshes, Akrotiri Bay and Forest,
Cherkes, Famagusta, Paramali.
Distribution. Cyprus, Northern Africa (Egypt).
Amegilla (Amegilla) quadrifasciata (de Villers, 1789)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Pissouri, Akrotiri Bay, Fasouri, Polemedia,
Cherkes, Asomatos, Chiicoudia marshes, Moni, Eftagonia, Evdhimou River, Platania
Forest Station, Famagusta, Mt. Troodos, Chionistra, Kannoures springs, Kathikas, Li-
vadin of Cedars (Paphos Forest), Akanthou, Ayia Irini (Kyrenia district).
Material examined. Limassol District: Almirolivado, 34.9333°N, 32.9004°E, 16-
18.IX.2011, S. Dimitriou leg., J. Devalez det. (1♀), pan trap (UAEG).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, South-Eastern Asia (Myanmar), Eastern Asia.
Amegilla (Zebramegilla) salviae (Morawitz, 1876)
References. Scheuchl and Willner (2016).
Mavromoustakis localities. Limassol.
Distribution. Cyprus, Western Europe (Switzerland, Austria), Southern Europe,
Eastern Europe, Northern Africa, Western Asia (Turkey, Azerbaijan), Central Asia,
Southern Asia (Iran), Eastern Asia (China).
Genus Anthophora LATREILLE, 1803
8 species.
Anthophora (Anthophora) canescens Brullé, 1832
References. Lepeletier (1841); Dours (1869); Costa (1883); Mavromoustakis (1949
[“1948”], 1951, 1952, 1953, 1954, 1957a); Georghiou (1977).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 79
Mavromoustakis localities. Limassol, Polemedia, hills, Episkopi, Ayios Athana-
sios, Apsiou, Pera Pedi, Akrotiri Bay, Fassouri, Yermasoyia River, Pissouri, Fasoula,
Yerasa, Pyrgos, Near Paramytha, Moni, Kellaki, Zakaki, Saettas, Morphou, Amathus,
Kyrenia, Famagusta, Nicosia.
Material examined. Nicosia District: 35.1688°N, 33.367°E, 13.III.2006, A Grace
leg., J. Devalez det. (2♂), hand net (UAEG); Paphos District: 34.7626°N, 32.4108°E,
25.II.2000, 8.III.2000, H. Wolf leg., J. Devalez det. (2♀), hand net (UAEG).
Distribution. Cyprus, Southern Europe, Western Europe (France, Switzerland),
Northern Africa, Western Asia.
Notes. is is the taxon sometimes referred to as Anthophora subterranea Germar,
1826.
Anthophora (Pyganthophora) dalmatica Pérez, 1902
Material examined. Limassol District: Almirolivado, 34.9333°N, 32.9004°E,
14.V.2012, 16.V.2012, S. Dimitriou leg., J. Devalez det. (1♀), pan trap, (UAEG);
Nicosia District: 35.1688°N, 33.367°E, 13.III.2006, A. Grace leg., A. Grace det. (2♀),
insect net, (UAEG).
Distribution. Cyprus, Southern Europe.
Notes. e true status of this taxon is not certain and Rasmont (in litt.) suggests
that it could be a ssp. of A. atroalba Lepeletier, 1841. Whatever the status, the taxon is
newly recognised in Cyprus.
Anthophora (Paramegilla) harmalae Morawitz, 1877
References. Mavromoustakis (1949 [“1948”], 1951).
Mavromoustakis localities. Limassol, Tsada, Krios River near Kilani, Mt. Troodos
Kannoures springs.
Distribution. Cyprus, Eastern Europe (Russian Federation), Northern Africa,
Western Asia, Central Asia, Southern Asia (Iran).
Anthophora (Anthophora) plumipes (Pallas, 1772)
References. Mavromoustakis (1949 [“1948”], 1957); Georghiou (1977).
Mavromoustakis localities. Limassol, Yermasoyia River, Cherkes, Fassouri, Tri-
miklini, Episkopi, Kilani, Amathus, Fasoula, Yerasa, Fassouri, Pera Pedi, Potamitissa,
Pyrgos, Near Paramytha, Saettas, Kellaki, Ayios Kostantinos Pitsilia, Kyrenia, Ayios
Ilarion, Nicosia.
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 5-7.
IV.2012, (1♀), pan trap (UAEG); Makria Kontarka, 34.9095°N, 32.8971°E, 14-
16.V.2012, (1♀), pan trap (UAEG), all records S. Dimitriou leg., J. Devalez det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Kazakhstan), Southern Asia (Iran, Afghanistan).
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80
Notes. Described from Cyprus as Anthophora acervorum cypriaca Mavromoustakis,
1957, from Amathus ruins: 34.7125°N, 33.1419°E, 27.II.1935, G.A. Mavromousta-
kis leg./det. ♂.
Anthophora (Lophanthophora) robusta (Klug, 1845)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Episkopi, Ayios Athana-
sios, Pera Pedi, Kato Platres, Mt. Troodos Chionistra, Curium (near Episkopi), Trimik-
lini, Ayios Athanasios, Mt. Troodos Pasha Livadin.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria, Romania, Ukraine), Northern Africa (Morocco, Algeria), Western
Asia, Central Asia (Turkmenistan, Kazakhstan).
Anthophora (Pyganthophora) rogenhoferi Morawitz, 1871
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1954, 1957a);
Georghiou (1977).
Mavromoustakis localities. Limassol, Mesayitonia, Akrotiri Forest, Episkopi, Ap-
siou, Kantara Mountains, Kaloiri Hills, Kellaki, Pera Pedi, Fassouri, Amathus, Fasoula,
Yerasa, Pyrgos, Larnaca, Nicosia, Karpasian Peninsula, Eleousa Monastery, Morphou,
Ayios Ilarion, Randidi Forest, Rizokarpaso.
Distribution. Cyprus, Southern Europe (Italy, Greece), Western Asia, Central
Asia, Southern Asia (Iran).
Anthophora (Pyganthophora) rubricrus Dours, 1870
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1954, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Mesayitonia, Akrotiri
Forest/Bay, Apsiou, Pera Pedi, Fassouri, Yermasoyia River, Amathus, Fasoula, Yerasa,
Pyrgos, Near Paramytha, Akrounda, Kellaki, Lania, Pareklisia, Trimiklini, Morphou,
Amathus, Kyrenia, Famagusta.
Material examined. Nicosia District: 35.1688°N, 33.367°E, 13.III.2006, A.
Grace leg., J. Devalez det. (5♂, 8♀), insect net (UAEG); Paphos District: 34.7626°N,
32.4108°E, 25.II.2000, 8.III.2000, H. Wolf leg., J. Devalez det. (4♀), insect net
(UAEG).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Israel).
Anthophora (Lophanthophora) rutilans Dours, 1870
Type locality–country. Cyprus, Nicosia: 35.166667°N, 33.366667°E, Sichel leg.,
Dours collection, J.A. Dours det. ♂ (MNHN).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 81
References. Dours (1870); Mavromoustakis (1949 [“1948”], 1951, 1952, 1953,
1954, 1957a).
Mavromoustakis localities. Limassol, Polemedia, Kitromili near Polemedia,
near Ayios Athanasios, Cherkes, Akrotiri Forest, Mesayitonia, Episkopi, Fassouri, Yer-
masoyia River, Pera Pedi, Amathus, Potamitissa, Fassoula, Yerasa, Kellaki, Moni, Near
Paramytha, Amyrou Monastery (near Apsiou), Ayios Ilarion, Near Famagusta, Mor-
phou, Mesayitonia-Fasoula.
Material examined. Paphos District: 34.7626°N, 32.4108°E, 25.II.2000,
8.III.2000, H. Wolf leg., J. Devalez det. (1♂), insect net (UAEG).
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Russian Fed-
eration), Western Asia.
AMMOBATINI
Genus Ammobates LATREILLE, 1809
3 species.
Ammobatoides abdominalis (Eversmann, 1852)
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (Austria), Southern Europe, Eastern Eu-
rope, Western Asia (Turkey, Lebanon), Central Asia (Kazakhstan), Southern Asia
(Iran), Eastern Asia (China).
Ammobates (Ammobates) biastoides Friese, 1895
References. Mavromoustakis (1954).
Mavromoustakis localities. Cherkes.
Distribution. Cyprus, Northern Africa (Algeria), Western Asia (Turkey, Israel).
Notes. e subspecies described from Cyprus is Ammobates biastoides globosus Ma-
vromoustakis, 1954, from Cherkes: 34.65°N, 32.975°E, 16.VI.1939, G.A. Mavro-
moustakis leg., G.A. Mavomoustakis det. ♀ (DAAN).
Ammobates (Ammobates) mavromoustakisi Popov, 1944
References. Popov (1944); Mavromoustakis (1949 [“1948”], 1951, 1954).
Mavromoustakis localities. Limassol, Cherkes, Zakaki, Ayios Athanasios, Faran-
gas near Famagusta, Paramali, Yermasoyia.
Distribution. Cyprus, Western Asia (Turkey, Palestine, Israel), Central Asia (Uz-
bekistan, Tajikistan), Southern Asia (Iran).
Notes. e subspecies described from Cyprus is Ammobates mavromoustakisi ma-
vromoustakisi, from Limassol, Ayios Athanasios, Cherkes: 34.72075°N, 33.05327°E,
34.65635°N, 32.98748°E, G.A. Mavromoustakis leg., V.V. Popov. det. (ZISP).
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82
Genus Chiasmognathus ENGEL, 2006
1 species.
Chiasmognathus orientanus (Warncke, 1983)
References. Warncke (1983).
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria, Ro-
mania), Western Asia (Turkey, Palestine, Israel).
Notes. Described from Cyprus as Pasites (Parammobatodes) orientanus cyprius
Warncke, 1983, from Yermasoyia River: 34.7182°N, 33.08788°E, 27.VII.1967, G.A.
Mavromoustakis leg., K. Warncke det. ♀, (KW).
All records of Ch. orientanus from Cyprus have been assigned cyprius.
Genus Parammobatodes POPOV, 1931
1 species.
Parammobatodes minutus (Mocsáry, 1878)
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Greece), East-
ern Europe, Western Asia.
Genus Pasites JURINE, 1807
1 species.
Pasites maculatus Jurine, 1807
References. Cockerell (1910); Popov (1944); Mavromoustakis (1949 [“1948”], 1951,
1952, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Cherkes, Moni River, Fa-
rangas near Famagusta, Ayia Napa.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia
(Iran, Afghanistan, Pakistan), Eastern Asia (China, Mongolia).
ANCYLAINI
Genus Ancyla LEPELETIER, 1841
1 species.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 83
Ancyla holtzi Friese, 1902
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Limassol, Cherkes, Zakaki, Asomatos, Near Enkomi
of Famagusta.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia (Turkey, Iraq), Southern Asia (Iran).
Genus Tarsalia MORAWITZ, 1895
2 species.
Tarsalia ancyliformis Popov, 1935
References. Pittioni (1950); Mavromoustakis (1952, 1953).
Mavromoustakis localities. Limassol, Cherkes, Yermasoyia River, Moni, Salipes
marshes near Akrotiri, Yermasoyia Hills, Near Asomatos, Near Trachoni.
Distribution. Cyprus, Southern Europe (Italy), Western Asia (Turkey), Central
Asia (Turkmenistan, Uzbekistan, Tajikistan), Southern Asia (Iran).
Notes. Described from Cyprus as Tarsalia ancyliformis mediterranea Pittioni, 1950,
from Geroskipou: 34.7559°N, 32.4516°E, 20.VII.1939, H. Lindberg leg., B. Pittioni det.
Includes T. mediterranea which is now regarded as a ssp. of Tarsalia ancyliformis
(MZHF).
Tarsalia hirtipes Morawitz, 1894
References. Mavromoustakis (1952, 1953, 1957a).
Mavromoustakis localities. Limassol, Cherkes.
Distribution. Cyprus, Western Asia (Turkey), Central Asia (Turkmenistan, Uz-
bekistan), Southern Asia (Iran).
Notes. Described from Cyprus as Tarsalia hirtipes cypriaca Mavromoustakis, 1952,
from Cherkes: 34.65°N, 32.975°E, 3.VIII.1933, G.A. Mavromoustakis leg./det. ♀,
(DAAN). Includes T. cypriaca which is now regarded as a ssp. of Tarsalia hirtipes.
APINI
Genus Apis LINNAEUS, 1758
1 species.
Apis (Apis) mellifera Linnaeus, 1758
References. Pittioni (1950); Georghiou (1977).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
84
Distribution. Cyprus, Widespread in Europe, Africa, Western Asia, Central Asia,
Southern Asia, Australia, New Zealand.
Notes. e subspecies described as Apis mellifera cypria Pollmann, 1879, from
unknown location, det. A. Pollmann. Beekeeping is widely practised throughout the
island.
BOMBINI
Genus Bombus LATREILLE, 1802
2 species.
Bombus (Sibiricobombus) niveatus Kriechbaumer, 1870
References. Rasmont and Iserbyt (2014); Rasmont et al. (2015).
Distribution. Cyprus, Southern Europe (Greece, Albania), Eastern Europe (Bulgar-
ia, Ukraine), Western Asia (Lebanon, Syria), Southern Asia (Iran), Eastern Asia (China).
Notes. Lack of recent records for Cyprus indicates the need for a conservation as-
sessment of its status on the island.
Bombus (Bombus) terrestris (Linnaeus, 1758)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Georghiou (1977).
Material examined. Nicosia District: Linou, 35.0755°N, 32.9164°E, 14-
16.V.2012, (1♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 15-17.
IX.2011, (2♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 5-7.IV.2012,
(2♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 14-16.V.2012, (2♀), pan
trap (UAEG); Limassol District: Amiantos, 34.918°N, 32.9472°E, 14-16.V.2012,
(1♀), pan trap (UAEG); Amiantos, 34.918°N, 32.9472°E, 31.V.2012, 2.VI.2012,
(1♂, 3♀), pan trap (UAEG); Almirolivado, 34.9333°N, 32.9004°E, 16-18.IX.2011,
(2♀), pan trap (UAEG); Almirolivado, 34.9333°N, 32.9004°E, 14-16.V.2012, (29♀),
pan trap (UAEG); Almirolivado, 34.9333°N, 32.9004°E, 31.V.2012, 2.VI.2012, (6♀),
pan trap (UAEG); Makria Kontarka, 34.9095°N, 32.8971°E, 16-18.IX.2011, (3♀),
pan trap (UAEG); Makria Kontarka, 34.9095°N, 32.8971°E, 14-16.V.2012, (15♀)
(UAEG); Makria Kontarka, 34.9095°N, 32.8971°E, 31.V.2012, 2.VI.2012, (2♂,
1♀), pan trap (UAEG); Troodos, Chionistra, 34.9317°N, 32.8664°E, 14-16.V.2012,
(3♂, 18♀), pan trap (UAEG); Troodos, Chionistra, 34.9317°N, 32.8664°E, 16-18.
IX.2011, (1♂), pan trap (UAEG); Troodos, Chionistra, 34.9317°N, 32.8664°E,
31.V.2012, 2.VI.2012, (12♀), pan trap (UAEG); Troodos, Mt. Olympos, 34.93°N,
32.86°E, 20.VI.2013, (6♀); Sovereign Base Area, Avdimou Bay Clis, 34.656698°N,
32.773339°E, 27.IV.2015; Sovereign Base Area, Episkopi, Kensington Clis,
34.670772°N, 32.846923°E, 4.V.2015, all Linou, Kakopetria, Amiantos, Almiroli-
vado, Makria Kontarka and Troodos Chionistra records S. Dimitriou leg., J. Devalez
det., Troodos Mt. Olympos records C. Schmid-Egger leg., C. Schmid-Egger det., Sov-
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 85
ereign Base Area, Avdimou Bay Clis and Sovereign Base Area, Episkopi, Kensington
Clis records S.P.M. Roberts leg., S.P.M. Roberts det.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Central Asia (Ka-
zakhstan, Kyrgyzstan, Uzbekistan), Southern Asia (Iran, Afghanistan), Eastern China
(China, Mongolia), Australia, New Zealand.
CERATININI
Genus Ceratina LATREILLE, 1802
8 species.
Ceratina (Neoceratina) bispinosa Handlirsch, 1889
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1952, 1957a).
Mavromoustakis localities. Limassol, Cherkes, Eftagonia, Zakaki, Ayios Atha-
nasios.
Distribution. Cyprus, Southern Europe (Croatia, Greece), Eastern Europe (Ro-
mania), Western Asia.
Ceratina (Euceratina) chrysomalla Gerstäcker, 1869
References. Mavromoustakis (1949 [“1948”]); Mavromoustakis (1951, 1952, 1954,
1957a,b).
Mavromoustakis localities. Limassol, Cherkes, Apsiou, Pera Pedi, Moni, Yer-
masoyia River/Hills (Kaloiri), Yerasa, Lania, Ayia Varvara (Stavrovouni), Platres, Pyrga.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Romania,
Ukraine, Bulgaria), Western Asia, Southern Asia (Iran).
Ceratina (Euceratina) cypriaca Mavromoustakis, 1949
Type locality–country. Cyprus, Pera Pedi: 609 m, 34.8580°N, 32.8730°E, 27.V.1929,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1953, 1957a).
Distribution. Cyprus. ENDEMIC.
Ceratina (Euceratina) dallatorreana Friese, 1896
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Polemedia Hills, Apsiou, Livadin of Cedars (Paphos
Forest), Mt. Troodos, Trimiklini, Eftagonia, Pera Pedi, Kykkou Monastery, Stavro-
vouni, Mesapotamos, Xerokolimbi (near Trooditissa), Potamitissa.
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86
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa (Morocco, Algeria, Tunisia), Western Asia, Central Asia (Turk-
menistan, Kyrgyzstan, Uzbekistan), Southern Asia (Iran).
Ceratina (Euceratina) mandibularis Friese, 1896
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952, 1953, 1954, 1958 ["1957"]).
Mavromoustakis localities. Cherkes, Chiicoudia marshes, Yermasoyia River, Ye-
rasa, Trimiklini, Erimi, Kathikas.
Material examined. Paphos District: 2.7 km SW of Acheleia, Potamos tis Ezou-
zas, 34.729004°N, 32.457544°E, 30.IV.2015, S.P.M. Roberts leg./det. (3♂).
Distribution. Cyprus, Northern Africa (Egypt), Western Asia (Turkey).
Ceratina (Euceratina) moricei Friese, 1899
References. Mavromoustakis (1949 [“1948”], 1952, 1957a).
Mavromoustakis localities. Limassol, Cherkes, Asomatos, Yermasoyia River, Pyr-
gos, Trimiklini, Yermasoyia Hills.
Distribution. Cyprus, Western Asia (Turkey), Southern Asia (Iran).
Ceratina (Dalyatina) parvula Smith, 1854
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951).
Mavromoustakis localities. Limassol, Yermasoyia River.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria), Northern Africa, Western Asia, Central Asia (Turkmenistan).
Ceratina (Neoceratina) schwarzi Kocourek, 1998
References. Terzo (1998).
Distribution. Cyprus, Southern Europe, Eastern Europe (Romania, Bulgaria),
Western Asia, Southern Asia (Iran).
EPEOLINI
Genus Epeolus LATREILLE, 1802
2 species.
Epeolus bischo (Mavromoustakis, 1954)
References. Bogusch and Hadrava (2018).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 87
Mavromoustakis localities. Mavromoustakis specimens in the Snow Entomolog-
ical Museum Collection, Kansas, USA: Akrotiri Bay, 34.62°N, 33.00°E, 12.VII.1943;
Zakaki, 23.VI.1949; Salamis, 18.VI.1957; Akrotiri Bay, 34.62°N, 33.00°E,
20.VII.1933; Akrotiri Bay, 34.62°N, 33.00°E, 1.VIII.1933; Akrotiri Bay, 34.62°N,
33.00°E, 12.VII.1943, all records G.A. Mavromoustakis leg./det.
Distribution. Cyprus, Western Asia.
Epeolus transitorius Eversmann, 1852
References. Bogusch and Hadrava (2018).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Russian Federation), Northern Africa (Morocco), Western Asia, Central Asia
(Turkmenistan, Kazakhstan, Uzbekistan), Southern Asia (Iran).
EUCERINI
Genus Eucera SCOPOLI, 1770
21 species.
Eucera (Hetereucera) aequata Vachal, 1907
Material examined. Limassol District: Akrotiri, 34.600657°N, 32.971419°E,
8.V.2016, (1♀); Akrotiri, 34.600657°N, 32.971419°E, 7.IV.2017, (2♂); Akroti-
ri, 34.600657°N, 32.971419°E, 21.IV.2017, (1♀, 2♂); Akrotiri, 34.600657°N,
32.971419°E, 5.V.2017, (1♀); Akrotiri, 34.600657°N, 32.971419°E, 30.III.2018,
(2♂); Akrotiri, 34.600657°N, 32.971419°E, 28.IV.2019 (1♀), all records A. Varnava
leg., A. Dorchin det.
Distribution. Cyprus, Western Asia (Turkey, Israel, Syria).
Eucera (Tetralonia) alticincta (Lepeletier, 1841)
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Odou, Krios River near Kilani.
Distribution. Cyprus, Widespread in Europe except in the North, Northern Af-
rica, Western Asia (Turkey).
Eucera (Hetereucera) bidentata Pérez, 1887
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Mesayitonia, Fassouri, Yermasoyia River,
Nicosia, Famagusta, Amathus, Fasoula, Morphou, Rizokarpaso, Ayia Irini station
(near Limassol).
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88
Material examined. Kyrenia District: 15 km E Kyrenia, 'Turtle Beach',
35.334413°N, 33.494187°E, 10.IV.2013, (1♀); Famagusta District: Protaras, Cape
Greco, 34.963264°N, 34.066211°E, 15.III.2017, (1♀); Achna Dam, 35.05519°N,
33.814011°E, 19.II.2018, (1♂); Limassol District: Cherkes, 34.653067°N,
32.974233°E, 10.III.2017, (1♂); Cherkes, 34.641933°N, 32.963433°E, 25.III.2017,
(1♂); Cherkes, 34.653067°N, 32.974233°E, 25.III.2017, (1♂); Cherkes,
34.653067°N, 32.974233°E, 14.II.2018, (2♂); Cherkes, 34.653067°N, 32.974233°E,
1.III.2018, (2♂); Cherkes, 34.650960°N, 32.990910°E, 1.III.2018, (1♂); Cherkes,
34.641933°N, 32.963433°E, 1.III.2018, (1♂); Cherkes, 34.650960°N, 32.990910°E,
30.III.2018, (1♀); Akrotiri, 34.601506°N, 32.986197°E, 25.III.2017, (1♂); Ak-
rotiri, 34.628817°N, 32.940667°E, 25.III.2017, (1♂); Akrotiri, 34.601506°N,
32.986197°E, 1.III.2018, (1♂), all Kyrenia district records C. Schmid-Egger leg., S.
Risch det., all Famagusta and Limassol district records A. Varnava leg., A. Dorchin det.
Distribution. Cyprus, Southern Europe (Italy, Greece), Western Asia (Turkey, Is-
rael, Syria).
Eucera (Hetereucera) caerulescens Friese, 1899
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a).
Mavromoustakis localities. Limassol, Polemedia Hills, Mesayitonia, Cherkes,
Apsiou, Cape Apostolos Andreas, Yermasoyia River, Yerasa, Pyrgos, Akrounda, Parekli-
sia, Randidi Forest.
Material examined. Kyrenia District: 5 km E of Kyrenia, "Turtle Beach",
35.334413°N, 33.494187°E, 10.IV.2007; Famagusta District: 15 km E of Rizokar-
paso, "Golden Sands", 35.64°N, 34.55°E, 10.IV.2007, 1 (♀) (AMNH), all records C.
Schmid-Egger leg., S. Risch det.
Distribution. Cyprus, Western Asia (Turkey, Israel), Southern Asia (Iran).
Eucera (Eucera) cypria Alfken, 1933
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 13-27.II.1927,
G.A. Mavromoustakis leg., J.D. Alfken det.
References. Alfken (1933); Mavromoustakis (1949 [“1948”], 1951, 1953, 1954,
1957a); Tkalců (1984).
Mavromoustakis localities. Limassol, Polemedia Hills, Mesayitonia, Pera Pedi,
Fassouri, Yermasoyia River, Famagusta, Amathus, Yerasa, Larnaca, Near Paramytha,
Mesayitonia-Fasoula, Moni, Lania, Trimiklini.
Material examined. Limassol District: Anogyra, 34.724979°N, 32.737225°E,
23.II.2016, (1♂); Cherkes, 34.650960°N, 32.990910°E, 1.III.2018, (1♂), all records
A. Varnava leg., A. Dorchin det.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Israel), Southern
Asia (Iran).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 89
Eucera (Eucera) dalmatica Lepeletier, 1841
References. Mavromoustakis (1949 [“1948”], 1952, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Akrotiri Forest, Near Enkomi of Famagus-
ta, Younarka, Pyrga, Xylophagou, Ormideia, Near Deryneia, Ayios Memnon (Fama-
gusta).
Material examined. Limassol District: Akrotiri, 34.628817°N, 32.940667°E,
24.V.2017, A. Varnava leg., A. Dorchin det. (1♂).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa (Morocco), Western Asia, Southern Asia (Iran).
Eucera (Eucera) dimidiata Brullé, 1832
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1953, 1954, 1957a);
Georghiou (1977).
Mavromoustakis localities. Limassol, Polemedia Hills, Episkopi, Fassouri, Yer-
masoyia River, Mesayitonia, Fasoula, Pera Pedi, Amathus, Trimiklini, Kyrenia, Mor-
phou, Famagusta, Sandy shore of Salamis (near Famagusta), Larnaca.
Material examined. Nicosia District: city centre, 35.1688°N, 33.367°E,
13.III.2006, (5♂, 16♀), insect net (UAEG); Limassol District: Anogyra, 34.724979°N,
32.737225°E, 23.II.2016, (1♀, 3♂); Ayios Dimitrianos, 8.III.2016, (1♂); Aswma-
tos, 8.III.2016, (1♂); Pissouri, 34.65093056°N, 32.72539722°E, 5.II.2017, (1♂);
Cherkes, 34.641933°N, 32.963433°E, 23.II.2017, (1♂); Cherkes, 34.650960°N,
32.990910°E, 25.III.2017, (2♀); Cherkes, 34.641933°N, 32.963433°E, 25.III.2017,
(1♀); Cherkes, 34.659801°N, 32.990510°E, 25.III.2017, (3♀); Cherkes,
34.641933°N, 32.963433°E, 21.IV.2017, (2♀); Cherkes, 34.641933°N, 32.963433°E,
1.II.2018, (1♂); Cherkes, 34.653067°N, 32.974233°E, 14.II.2018, (1♂); Cherkes,
34.641933°N, 32.963433°E, 1.III.2018, (1♀); Cherkes, 34.659801°N, 32.990510°E,
1.III.2018, (1♀); Cherkes, 34.659801°N, 32.990510°E, 16.III.2018, (2♀); Akrotiri,
34.588600°N, 32.938900°E, 1.III.2018, (1♀); Akrotiri, 34.628817°N, 32.940667°E,
1.III.2018, (1♂); Larnaca District: Choirokitia, 34.777521°N 33.33622°E,
27.II.2018, (1♀); Famagusta District: Deryneia, 23.III.2016, (1♀); Protaras, Cape
Greco, 34.963264°N, 34.066211°E, 5.ii.2018, (1♂); Achna Dam, 35.05519°N,
33.814011°E, 19.II.2018, (1♀), Nicosia district records A. Grace leg., S. Risch det.,
all Limassol, Larnaca, and Famagusta district records A. Varnava leg., A. Dorchin det.
Distribution. Cyprus, Southern Europe (Greece), Northern Africa (Algeria, Tunisia,
Egypt), Western Asia (Turkey, Israel, Saudi Arabia), Southern Asia (Iran, Afghanistan).
Eucera (Hetereucera) furfurea Vachal, 1907
References. Alfken (1933); Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Limassol, Akrotiri Forest, Pissouri, Nicosia.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Israel).
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90
Eucera (Hetereucera) gaullei Vachal, 1907
References. Alfken (1933); Mavromoustakis (1949 [“1948”], 1951); Georghiou (1977).
Mavromoustakis localities. Limassol, Akrotiri Forest, Episkopi, Yermasoyia Riv-
er, Kato Platres, Potamitissa, Kyrenia, Sphalangiotissa Monastery, Northern Moun-
tains Kantara, Near Salamis, Ayios Ilarion.
Material examined. Nicosia District: city centre, 35.1688°N, 33.367°E,
13.III.2006, (4♂), insect net (UAEG); Limassol District: Cherkes, 34.641933°N,
32.963433°E, 8.V.2016, (1♀); Akrotiri, 34.601506°N, 32.986197°E, 21.IV.2017,
(1♀); Akrotiri, 34.600657°N, 32.971419°E, 26.IV.2018, (1♀), all Nicosia district
records A. Grace leg., S. Risch det., all Limassol district records A. Varnava leg., A.
Dorchin det.
Distribution. Cyprus, Western Asia.
Eucera (Tetralonia) glauca (Fabricius, 1775)
References. Pittioni (1950); Mavromoustakis (1952, 1957a); Tkalců (1979).
Mavromoustakis localities. Pyrga (Larnaca).
Mavromoustakis specimens in the Snow Entomological Museum Collection, Kan-
sas, USA: Asomatos, 34.64°N, 32.96°E, 26.VI.1951, G.A. Mavromoustakis leg./det.,
(1♀) and 3 unspecied specimens.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Iraq),
Southern Asia (Iran).
Eucera (Tetralonia) inulae (Tkalců, 1979)
References. Tkalců (1979).
Mavromoustakis localities. Mavromoustakis specimens in the Snow Entomo-
logical Museum Collection, Kansas, USA: Limassol, Kilani, 14.VII.1937, G.A. Mav-
romoustakis leg., D.B. Baker det., 3 specimens.
Material examined. Limassol District: Akrotiri; 34.600657°N, 32.971419°E,
29.X.2016, S.P.M. Roberts leg./det., foraging on Dittrichia viscosa (Asteraceae), (1♂, 1♀).
Distribution. Cyprus, Southern Europe, Eastern Europe (Ukraine, Bulgaria, Rus-
sian Federation), Southern Asia (Iran).
Eucera (Synhalonia) mavromoustakisi (Tkalců, 1984)
Type locality–country. Cyprus, Lania: 34.82444°N, 32.92083°E, 2.VI.1964, G.A.
Mavromoustakis leg., B. Tkalců det. ♂ (FSAG).
References. Tkalců (1984).
Distribution. Cyprus. ENDEMIC.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 91
Eucera (Tetralonia) malvae (Rossi, 1790)
References. Mavromoustakis (1949 [“1948”]).
Mavromoustakis localities. Polemedia Hills, Mesayitonia, Akrotiri Forest, Sym-
boulas Chiik (near Limassol), Near Platania Forest Station, Lania, Ayia Varvara (Stav-
rovouni), Near Ayios Athanasios, hills near Trimiklini.
Distribution. Cyprus, Widespread across Europe, Western Asia (Turkey).
Eucera (Eucera) palaestinae Friese, 1922
Material examined. Limassol District: Akrotiri, 34.601506°N, 32.986197°E,
21.IV.2017, (1♂); Akrotiri, 34.601506°N, 32.986197°E, 1.III.2018, (3♀); Akrotiri,
34.600657°N, 32.971419°E, 1.III.2018, (2♀), all Limassol district records A. Varnava
leg., A. Dorchin det.
Distribution. Cyprus, Western Asia, Southern Asia (Iran).
Eucera (Eucera) proxima Morawitz, 1875
References. Mavromoustakis (1949 [“1948”], 1951, 1952).
Material examined. Limassol District: Troodos, Chionistra, 34.9317°N,
32.8664°E, 31.V.2012, 2.VI.2012, (1♂), pan trap, (UAEG); Akrotiri, 34.628817°N,
32.940667°E, 25.III.2017, (1♀); Akrotiri, 34.601506°N, 32.986197°E, 7.IV.2017,
(1♀); Akrotiri, 34.601506°N, 32.986197°E, 21.IV.2017, (1♀); Akrotiri, 34.628817°N,
32.940667°E, 21.IV.2017, (1♀); Akrotiri, 34.600657°N, 32.971419°E, 1.III.2018,
(1♂), all Troodos, Chionistra records S. Dimitriou leg., S. Risch det., all Akrotiri re-
cords A. Varnava leg., A. Dorchin det.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Western Asia (Turkey, Syria), Central Asia (Turkmenistan, Uzbekistan, Tajik-
istan), Southern Asia (Iran).
Notes. Also recorded as E. graeca Radoszkowski, 1876 e.g., Rasmont et al. 2017.
Precedence is given to proxima following Nieto et al. (2014; see methodology).
Eucera (Eucera) seminuda Brullé, 1832
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe (North
Macedonia, Italy, Greece), Eastern Europe, Northern Africa (Tunisia), Western Asia
(Turkey, Armenia).
Eucera (Eucera) sinufascia Dorchin, 2018
References. Tkalců (1979).
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92
Material examined. Limassol District: Cherkes, 34.641933°N, 32.963433°E,
10.III.2017, A. Varnava leg., A. Dorchin det. (1♂).
Distribution. Cyprus, Southern Europe (Greece (Aegean islands)), Western Asia.
Notes. Synonymising Tetraloniella under Eucera, the name became a junior homo-
nym of Eucera (Tetralonia) penicillata (Friese, 1905) and Eucera sinufascia Dorchin
2018 was proposed as a replacement Dorchin et al. (2018).
Eucera (Eucera) sulamita Vachal, 1907
Material examined. Northern Cyprus, 10 km N of Famagusta, Salamis, 35.187°N,
33.899°E, 4.X.2007, C. Schmid-Egger leg., S. Risch det. (1♂) (AMNH).
Distribution. Cyprus, Western Asia (Israel).
Eucera (Hetereucera) syriaca Dalla Torre, 1896
References. Mavromoustakis (1949 [“1948”], 1951, 1957a); Georghiou (1977).
Mavromoustakis localities. Limassol, Polemedia Hills, Mesayitonia, Ayia Phyla,
Nicosia, Sphalangiotissa Monastery, Fassouri, Famagusta, Amathus, Near Enkomi of
Famagusta.
Material examined. Famagusta District: 5 km E of Rizokarpaso, 35.63°N,
34.50°E, 10.IV.2007,; Limassol District: Akrotiri, 34.583676°N, 32.949306°E,
14.IV.2018, (1♀); Akrotiri, 34.583676°N, 32.949306°E, 28.IV.2018, (1♀), all Fama-
gusta district records C. Schmid-Egger leg., S. Risch det., all Limassol district records
A. Varnava leg., A. Dorchin det.
Distribution. Cyprus, Western Asia, Southern Asia (Iran).
Eucera (Synhalonia) tricincta Erichson, 1835
References. Tkalců (1984).
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Spain, Italy,
Greece), Eastern Europe, Northern Africa, Western Asia.
Eucera (Synhalonia) zeta Dalla Torre, 1896
References. Mavromoustakis (1952).
Mavromoustakis localities. Pera Pedi, Potamitissa, Moni, Kellaki, Saettas, Kitro-
mili, Mt. Kornos.
Distribution. Cyprus, Southern Europe (Greece), Western Asia, Central Asia
(Turkmenistan), Southern Asia (Iran).
Notes. Described from Cyprus as Eucera vernalis sintenisi Mavromoustakis, 1952
from, Potamitissa: 34.9075°N, 32.989444°E, 16.III.1947, G.A. Mavromoustakis leg.,
G.A. Mavromoustakis det. ♀, (DAAN).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 93
MELECTINI
Genus Melecta LATREILLE, 1802
7 species.
Melecta (Melecta) albifrons (Forster, 1771)
References. Mavromoustakis (1949 [“1948”], 1951); Lieftinck (1980).
Mavromoustakis localities. Nicosia, Mt. Troodos, Yermasoyia River, Amathus,
Potamitissa, Yerasa, Near Amathus, Prodromos, Dragontospillios cave.
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Southern Asia (Iran).
Melecta (Melecta) duodecimmaculata (Rossi, 1790)
References. Mavromoustakis (1949 [“1948”]); Georghiou (1977); Lieftinck (1980).
Mavromoustakis localities. Polemedia Hills, Mesayitonia, Limassol, Ayia Phyla,
Fassouri, Yermasoyia River, Amathus, Yerasa.
Distribution. Cyprus, Western Europe (France), Southern Europe, Northern Af-
rica, Western Asia, Central Asia, Southern Asia (Iran).
Melecta (Melecta) italica Radoszkowski, 1876
References. Lieftinck (1980).
Distribution. Cyprus, Western Europe (France), Southern Europe, Northern Af-
rica, Western Asia, Eastern Asia.
Melecta (Melecta) leucorhyncha Gribodo, 1894
References. Lieftinck (1980).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Ukraine), Northern Africa, Western Asia, Eastern Asia.
Melecta (Melecta) luctuosa (Scopoli, 1770)
References. Scheuchl and Willner (2016).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran, Pakistan).
Melecta (Melecta) mundula Lieftinck, 1983
References. Lieftinck (1983); Schwarz (1999).
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94
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Russian Fed-
eration), Western Asia (Turkey, Israel, Jordan), Central Asia, Southern Asia (Iran).
Notes. Described from Cyprus as Melecta megaera Lieftinck, 1980, from Poleme-
dia: 34.699444°N, 32.996944°E, 9.III.1950, G.A. Mavromoustakis leg., M.A. Lieft-
inck det. ♂, (MFNB).
Melecta (Melecta) tuberculata Lieftinck, 1980
References. Lieftinck (1980).
Distribution. Cyprus, Western Europe (France), Southern Europe (Portugal,
Spain, Greece), Eastern Europe (Bulgaria, Russian Federation), Western Asia (Turkey,
Israel, Lebanon), Eastern Asia.
Genus yreus PANZER, 1806
5 species.
yreus anis (Morawitz, 1874)
References. Pittioni (1950); Lieftinck (1968); Mavromoustakis (1951, 1952).
Mavromoustakis localities. Limassol, Polemedia Hills, Akrotiri Bay, Near Ak-
rotiri, Younaros of Zakaki.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa, Western Asia, Central Asia, Southern Asia (Iran, Pakistan).
yreus elegans (Morawitz, 1877)
References. Lieftinck (1968).
Distribution. Cyprus, Southern Europe, Northern Africa, Western Asia, Central
Asia (Turkmenistan, Kyrgyzstan), Southern Asia (Iran, Pakistan).
yreus histrionicus (Illiger, 1806)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1957a); Lieftinck (1968);
Georghiou (1977).
Mavromoustakis localities. Limassol, Polemedia Hills, Mesayitonia, Cherkes,
Akrotiri Bay, Near Akrotiri, Near Enkomi of Famagusta, Yermasoyia River, Akrounda,
Moni River, Mt. Troodos Chionistra, Pernera coast of Paralimni, Younarka (near Zaka-
ki), Krios River near Kilani, near Fassouri.
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, C. Schmid-Egger leg., M. Schwarz det. (1♀).
Distribution. Cyprus, Western Europe (France, Austria), Southern Europe, East-
ern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia (Iran), Eastern
Asia (China).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 95
yreus picaron Lieftinck, 1968
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, C. Schmid-Egger leg., M. Schwarz det. (1♂).
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope (Bulgaria, Romania).
yreus ramosus (Lepeletier, 1841)
References. Mavromoustakis (1949 [“1948”], 1951, 1952, 1954); Lieftinck (1968);
Georghiou (1977).
Mavromoustakis localities. Limassol, Cherkes, Episkopi, Ayios Athanasios, Pera
Pedi, Akrotiri Bay, Moni, Near Akrotiri, Yermasoyia River, Eftagonia, Xerokolimbi
Stream near Trooditissa, Platres, Pera Pedi.
Material examined. Paphos District: 20 km NNW Paphos, Lara Beach, 34.94°N,
32.31°E, 20.VI.2013, C. Schmid-Egger leg., M. Schwarz det. (1♂).
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia,
Eastern Asia (China).
NOMADINI
Genus Nomada SCOPOLI, 1770
39 species.
Nomada babiyi Schwarz & Standfuss, 2007
References. Smit (2018).
Distribution. Cyprus, Southern Europe (Croatia, Greece), Eastern Europe
(Ukraine, Bulgaria), Western Asia (Turkey, Syria).
Nomada bifasciata Olivier, 1812
References. Smit (2018).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia
(Turkey).
Nomada caspia Morawitz, 1894
References. Smit (2018).
Distribution. Cyprus, Southern Europe (Croatia, Greece), Western Asia (Turkey,
Israel, Lebanon), Central Asia (Turkmenistan).
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96
Nomada cherkesiana Mavromoustakis, 1955
Type locality–country. Cyprus, Cherkes: 34.65°N, 32.975°E, 30.III.1950, G.A. Ma-
vromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1955, 1957a); Smit (2018).
Mavromoustakis localities. Cherkes, Yermasoyia River, Yerasa.
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Turkey, Israel).
Nomada connis Schmiedeknecht, 1882
References. Smit (2018).
Distribution. Cyprus, Western Europe (Switzerland), Southern Europe, Eastern
Europe (Bulgaria), Western Asia, Southern Asia (Iran).
Nomada cypria Mavromoustakis, 1952
Type locality–country. Cyprus, Cherkes: 34.65°N, 32.975°E, 15.III.1950, G.A. Ma-
vromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Schwarz (1999); Mavromoustakis (1952); Smit (2018).
Distribution. Cyprus, Southern Europe (Greece), Western Asia (Israel).
Nomada cypricola Mavromoustakis, 1955
Type locality–country. Cyprus, Zakaki: 34.6563°N 33.0029°E, 17.VI.1949, G.A.
Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1957a); Smit (2018).
Mavromoustakis localities. Limassol, Polemedia Hills, Ayia Irini (near Palodkia),
Sphalangiotissa Monastery (near Limassol).
Distribution. Cyprus. ENDEMIC.
Nomada erythrocephala Morawitz, 1871
References. Mavromoustakis (1949 [“1948”]); Smit (2018).
Mavromoustakis localities. Limassol, Polemedia Hills, Episkopi Forest, Ayia Irini
Station (near Limassol).
Distribution. Cyprus, Western Europe (France), Southern Europe (Spain,
Greece), Eastern Europe, Western Asia (Turkey).
Nomada licornis Schwarz & Smit, 2018
References. Smit (2018).
Remarks. Smit paratype localities: Limassol, Vavla, Akrotiri, Kapedhes.
Distribution. Cyprus, Southern Europe (Italy, Greece), Western Asia (Turkey, Jor-
dan, Syria).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 97
Nomada avinervis Brullé, 1832
References. Mavromoustakis (1955, 1957a); Smit (2018).
Mavromoustakis localities. Limassol, Pera Pedi, Lania, Kellaki, Fasoula, Trimik-
lini, Episkopi, Yermasoyia River.
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia (Turkey, Israel).
Nomada avoguttata (Kirby, 1802)
References. Mavromoustakis (1949 [“1948”]); Smit (2018).
Mavromoustakis localities. Limassol.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, Bee Course students leg., M. Schwarz det. (4♀).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran), Eastern Asia (China).
Nomada fucata Panzer, 1798
References. Mavromoustakis (1949 [“1948”]); Georghiou (1977); Smit (2018).
Mavromoustakis localities. Limassol, Yermasoyia River, Amathus, Potamitissa.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, Bee Course students leg., M. Schwarz det. (1♀).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia (Kazakhstan, Kyrgyzstan), Southern Asia (Pakistan).
Nomada fulvicornis Fabricius, 1793
References. Pittioni (1950); Smit (2018).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia, Southern Asia (Iran, Pakistan, India), Eastern Asia (China).
Nomada furva Panzer, 1798
References. Smit (2018).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe, North-
ern Africa (Algeria), Western Asia (Turkey, Georgia), Central Asia.
Nomada gageae Schwarz & Smit, 2018
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 25.I.1949
[“1948”], G.A. Mavromoustakis leg., J. Smit det. ♀, (MSAA). Paratypes: Cyprus,
Limassol, 28.I.1949 [“1948”], G.A. Mavromoustakis leg., J. Smit/M. Schwarz
det. 2♂.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
98
References. Smit (2018).
Distribution. Cyprus. ENDEMIC.
Nomada goodeniana (Kirby, 1802)
References. Smit (2018).
Distribution. Cyprus, Widespread in Europe, Western Asia, Central Asia, South-
ern Asia (Iran).
Nomada immaculata Morawitz, 1874
References. Mavromoustakis (1949 [“1948”]); Smit (2018).
Mavromoustakis localities. Polemedia Hills, Episkopi, Yerasa.
Distribution. Cyprus, Southern Europe (Greece, North Macedonia), Eastern
Europe (Hungary, Russia), Northern Africa, Western Asia (Armenia), Southern Asia
(Afghanistan, Pakistan).
Nomada incisa Schmiedeknecht, 1882
References. Smit (2018).
Distribution. Cyprus, Southern Europe, Eastern Europe, Western Asia (Turkey).
Nomada integra Brullé, 1832
References. Smit (2018).
Distribution. Cyprus, Widespread in Europe, Northern Africa, Western Asia,
Central Asia.
Nomada kohli Schmiedeknecht, 1882
References. Smit (2018).
Distribution. Cyprus, Western Europe (France, Germany, Austria), Southern Eu-
rope, Eastern Europe, Northern Africa, Western Asia (Turkey, Israel).
Nomada kornosica Mavromoustakis, 1958
Type locality–country. Cyprus, Mt. Kornos (Northern Mountains): 35.1379°N,
33.1379°E, 23.III.1953, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀.
References. Mavromoustakis (1958); Smit (2018).
Distribution. Cyprus, Southern Europe (Greece).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 99
Nomada limassolica Mavromoustakis, 1955
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 13.III.1953,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♂, (DAAN).
References. Mavomoustakis (1955, 1957); Smit (2018).
Distribution. Cyprus, Southern Europe (Greece), Eastern Europe (Bulgaria),
Western Asia (Turkey, Israel).
Nomada lucidula Schwarz, 1967
References. Smit (2018).
Distribution. Cyprus, Southern Europe (Albania, Greece), Eastern Europe (Bul-
garia), Western Asia (Turkey, Israel).
Nomada mutica Morawitz, 1872
References. Smit (2018).
Distribution. Cyprus, Western Europe, Southern Europe, Eastern Europe (Hun-
gary, Romania), Western Asia.
Nomada nesiotica Mavromoustakis, 1958
Type locality–country. Cyprus, Fasoula [8 km N of Limassol]: 34.761667°N, 33.026944°E,
9.III.1951, G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1958 ["1957"]); Smit (2018).
Mavromoustakis localities. Limassol, Fasoula, Apsiou, Yerasa, Amathus, Yer-
masoyia River, Pera Pedi, Lania, Trimiklini, Kellaki.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, Bee Course students leg., M. Schwarz det. (8♀).
Distribution. Cyprus, Southern Europe (Greece).
Nomada numida Lepeletier, 1841
References. Mavromoustakis (1949 [“1948”]); Smit (2018).
Mavromoustakis localities. Ayios Athanasios, Yermasoyia River, Near Paramytha,
Ayia Varvara (Stavrovouni), Pernera coast of Paralimni.
Distribution. Cyprus, Western Europe (France), Southern Europe, Eastern Eu-
rope, Northern Africa, Western Asia (Turkey, Israel, Iraq).
Nomada pallispinosa Schwarz, 1967
References. Smit (2018).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
100
Material examined. Limassol District: Polemidia, 34.71178°N, 33.004775°E,
8.III.2017, Bee Course students leg., M. Schwarz det. (2♀).
Distribution. Cyprus, Southern Europe, Eastern Europe (Bulgaria, Russian Fed-
eration), Western Asia (Turkey, Israel).
Nomada pleurosticta Herrich-Schäer, 1839
References. Smit (2018).
Distribution. Cyprus, Western Europe, Southern Europe (Spain, Greece), Eastern
Europe (Czech Republic, Hungary, Slovakia), Northern Africa (Tunisia), Western Asia
(Turkey), Southern Asia (Iran).
Nomada polemediana Mavromoustakis, 1957
Type locality–country. Cyprus, Limassol: 34.66839°N, 33.03252°E, 17.IV.1954,
G.A. Mavromoustakis leg., G.A. Mavromoustakis det. ♀, (DAAN).
References. Mavromoustakis (1957b; Warncke (1967); Smit (2018).
Distribution. Cyprus. ENDEMIC.
Nomada propinqua Schmiedeknecht, 1882
References. Mavromoustakis (1952); Smit (2018).
Mavromoustakis localities. Apsiou, hills near Paramytha, Mt. Kornos.
Distribution. Cyprus, Southern Europe, Western Asia (Israel).
Nomada pyrgosica Schwarz & Smit, 2018
Type locality–country. Cyprus, Famagusta, Rizokarpaso: 35.59881°N, 34.2772°E,
29.III.2012, Schwenninger leg., J. Smit det. ♀, (MSAA).
References. Smit (2018).
Distribution. Cyprus, Western Asia (Turkey).
Nomada stigma Fabricius, 1804
References. Mavromoustakis (1955, 1957b); Smit (2018).
Distribution. Cyprus, Widespread in Europe, Northern Africa (Algeria), Western
Asia (Turkey), Central Asia (Kazakhstan).
Notes. e subspecies described from Cyprus is Nomada stigma cypricola Mav-
romoustakis, 1955, from Zakaki: 34.6563°N, 33.0029°E, 17.VI.1949, G.A. Mavro-
moustakis leg. G.A. Mavromoustakis det. ♀, (DAAN).
Nomada striata Fabricius, 1793
References. Mavromoustakis (1951, 1957a); Smit (2018).
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 101
Mavromoustakis localities. Chiicoudia marshes (near Limassol), Pera Pedi,
Amathus.
Distribution. Cyprus, Widespread in Europe, Northern Africa (Morocco), West-
ern Asia, Central Asia (Kazakhstan).
Nomada succincta Panzer, 1798
References. Scheuchl and Willner (2016); Smit (2018).
Distribution. Widespread in Europe, Northern Africa, Western Asia, Central Asia
(Turkmenistan, Kazakhstan).
Nomada teunisseni Schwarz & Smit, 2018
Type locality–country. Cyprus, Akanthou: 35.3741°N, 33.7558°E, 12.III.1981, H.
Teunissen leg., J. Smit det. ♂ (MSAA). Paratypes: Cyprus, Mt Kornos, 9.III.1981, H.
Teunissen leg., Smit/ Schwarz det. 1 ♂; Cyprus, Kantara, 2000m, 19.III.1971, K.M.
Guichard leg., Smit/ Schwarz det. 1 ♀.
References. Smit (2018).
Distribution. Cyprus. ENDEMIC.
Nomada thersites Schmiedeknecht, 1882
References. Smit (2018).
Distribution. Cyprus, Southern Europe (Spain, Italy, Greece), Eastern Europe
(Hungary, Ukraine, Russian Federation), Western Asia (Turkey, Israel), Central Asia
(Kazakhstan, Kyrgyzstan), Southern Asia (Iran).
Nomada tridentirostris Dours, 1873
References. Smit (2018).
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe (Poland, Hungary, Bulgaria), Northern Africa, Western Asia
(Turkey, Israel), Southern Asia (Iran).
Nomada trispinosa Schmiedeknecht, 1882
References. Mavromoustakis (1952, 1953); Smit (2018).
Mavromoustakis localities. Limassol, Cherkes, Apsiou, Trimiklini.
Material examined. Limassol District: Yermasoyia Dam, 34.755799°N,
33.096194°E, 7.III.2017, (6♀); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017,
(3♀), all records Bee Course students leg., M. Schwarz det.
Distribution. Cyprus, Western Europe (Austria), Southern Europe (Greece, Slo-
venia), Eastern Europe, Western Asia (Israel, Jordan, Azerbaijan), Eastern Asia (China).
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
102
Nomada yermasoyiae Schwarz, Smit & Gusenleitner, 2018
References. Schwarz, Smit and Gusenleitner (2018).
Type localities. Cyprus, Cherkes: 23.III.1950, ♀, G.A. Mavromoustakis leg., in
coll. M. Schwarz. Allotype: Cyprus, Cherkes, 23.III.1950, ♂, G.A. Mavromousta-
kis leg., in coll. M. Schwarz. Paratypes: Cyprus: Limassol, 28.III.1959, G.A. Mavro-
moustakis leg., 1 ♀, 2 ♂; Cyprus, Amathus, 7.III.1966, 1 ♂, G.A. Mavromoustakis leg.
Distribution. Cyprus, Western Asia (Israel).
XYLOCOPINI
Genus Xylocopa LATREILLE, 1802
4 species.
Xylocopa (Copoxyla) iris (Christ, 1791)
References. Pittioni (1950); Mavromoustakis (1949 [“1948”], 1952).
Mavromoustakis localities. Moni River, Livadin of Cedars (Paphos Forest).
Material examined. Paphos District: 15 km SE Paphos, Kouklia, 34.72°N,
32.55°E, 20.VI.2013, (1♂); 20 km N Paphos, Kathikas, 34.90°N, 32.42°E, 20.VI.2013,
(1♀, 1♂); 20 km NNW Paphos, Lara Beach, 34.94°N, 32.31°E, 20.VI.2013, (1♀,
1♂); 6 km NE Polis, beach, 35.06°N, 32.46°E, 20.VI.2013, (1♂); N of Elia Bridge,
34.900977°N, 32.776759°E, 29.IV.2015, (1♂); Limassol District: 8 km E Limas-
sol, on road, 34.67°N, 32.85°E, 20.VI.2013, (1♀); Anogyra to Avdimou Road km 2,
34.723986°N, 32.736892°E, 3.V.2015, (1♀); 0.7 km N of Anogyra, 34.745537°N,
32.73385°E, 3.V.2015, (1♀), all 15 km SE Paphos Kouklia, 20 km N Paphos Kathi-
kas, 20 km NNW Paphos Lara Beach, 6 km NE Polis, beach and 8 km E Limassol,
on road records C. Schmid-Egger leg./det., all N of Elia Bridge, Anogyra to Avdimou
Road km 2 and 0.7 km N of Anogyra records S.P.M. Roberts leg./ det.
Distribution. Cyprus, Western Europe (France, Switzerland, Austria), Southern
Europe, Eastern Europe, Northern Africa, Western Asia, Central Asia, Southern Asia
(Iran, Afghanistan).
Xylocopa (Proxylocopa) olivieri Lepeletier, 1841
References. Pittioni (1950); Mavromoustakis (1949 [“1948”], 1951, 1952).
Mavromoustakis localities. Limassol, Akrotiri Forest, Evdhimou River, Pera Pedi,
Platres, Near Kilani, Near Cape Akamas, Near Trimiklini.
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 15-17.
IX.2011, (1♀), pan trap (UAEG); Kakopetria, 34.992°N, 32.9082°E, 14-16.V.2012,
(1♂, 1♀), pan trap (UAEG), all records S. Dimitriou leg., J. Devalez det.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 103
Distribution. Cyprus, Southern Europe (Albania, North Macedonia, Greece),
Eastern Europe (Russian Federation), Northern Africa (Egypt), Western Asia (Israel),
Central Asia (Turkmenistan, Kyrgyzstan), Southern Asia (Iran).
Xylocopa (Koptortosoma) pubescens Spinola, 1838
References. Rasmont et al. (2017).
Material examined. Nicosia District: city centre, 35.1688°N, 33.367°E,
13.III.2006, (1♂), insect net (UAEG); Agios Sozomenos, 35.06687°N, 33.43580°E,
4.III.2019, (1♀) (from photograph); Paphos District: 15 km SE Paphos, Kouk-
lia, 34.72°N, 32.55°E, 20.VI.2013, (1♀); Paphos, Venus Beach Hotel, 34.78°N,
32.40°E, 7.VIII.2016, (1♀) (from photograph); Larnaca District: Prosfygikos Sy-
noikismos EAC, 34.983633°N, 33.745922°E, 2.IV.2016, (1♀); Prosfygikos Sy-
noikismos EAC, 34.983633°N, 33.745922°E, VII.2017, (1♀); Prosfygikos Syn-
oikismos EAC, 34.983633°N 33.745922°E, 2.III.2018, (1♀); Limassol District:
Akrotiri, 34.583602°N 32.949536°E, 17.IV.2016, (1♀); Akrotiri, 34.583602°N,
32.949536°E, 8.V.2016 (1♀); Polemidia, 34.71178°N, 33.004775°E, 8.III.2017,
(1♀); Akrotiri, 34.600427°N 32.971111°E, 14.IV.2018, (1♀), Nicosia city centre re-
cords A. Grace leg., J. Devalez and A. Pauly det., Agios Sozomenos records E. Tzirkalli
leg., S.P.M. Roberts det., 15 km SE Paphos, Kouklia records C. Schmid-Egger leg., C.
Schmid-Egger det., Paphos, Venus Beach Hotel records S. Bagshaw leg., S.P.M. Rob-
erts det., Prosfygikos Synoikismos EAC (2.IV.2016) A. Varnava leg., S.P.M. Roberts
det., Prosfygikos Synoikismos EAC (VII.2017, 2.III.2018) records A. Varnava leg., A.
Varnava det., Akrotiri (records 17.IV.2016, 8.V.2016) A. Varnava leg., S.P.M. Roberts
det., all Polemidia records S.P.M. Roberts leg./det., Akrotiri records (14.IV.2018) A.
Varnava leg./det.
Distribution. Cyprus, Southern Europe (Greece), Widespread in Africa, Western
Asia, South-eastern Asia (Myanmar), Central Asia, Southern Asia.
Xylocopa (Xylocopa) violacea (Linnaeus, 1758)
References. Mavromoustakis (1949 [“1948”]); Pittioni (1950); Mavromoustakis
(1951, 1952).
Mavromoustakis localities. Limassol, Polemedia Hills, Akrotiri Bay, Yerasa,
Kitromili near Polemedia, Chiicoudia marshes, Bogazi shore.
Material examined. Nicosia District: Kakopetria, 34.992°N, 32.9082°E, 25-27.
IV.2012, (1♀), pan trap (UAEG); Limassol District: Amiantos, 34.918°N, 32.9472°E,
31.V.2012, 2.VI.2012, (1♀), pan trap (UAEG); Paphos District: 20 km NNW Paphos,
Lara Beach, 34.94°N, 32.31°E, 20.VI.2013, (4♀), all Nicosia district and Amiantos
records S. Dimitriou leg., J. Devalez det., all Paphos district records C. Schmid-Egger
leg., C. Schmid-Egger det.
Distribution. Cyprus, Widespread in Europe.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
104
Discussion
e rst modern checklist of the wild bees of Cyprus provides information on the spe-
cies reported on the island based on previous publications and new collections. e list
contains 369 species of wild bees that were accepted/veried by the authors to occur on
the island. e Mediterranean is well known for its high bee diversity, a result of the
high oral diversity and optimal weather conditions (Michener 2007, Nieto et al. 2014).
Cyprus has greater species richness of bees than larger islands, such as New Guinea,
Honshu, Great Britain, and Borneo (Ascher and Pickering 2018). e bee species rich-
ness of the island compares well to that of other Mediterranean islands (Fig. 3), with
only Lesvos (at least 600 species: Nielsen et al. 2011, Petanidou et al. unpublished
data) and Sicily (562 species: [cf. 575 species according to J. S. Ascher, unpublished,
based in part on Scheuchl and Willner 2016]) having greater richness than Cyprus.
Crete, Sardinia, and Corsica are reported to host 351, 296, and 263 bee species, re-
spectively (Ascher unpublished, cf. Scheuchl and Willner 2016; Ascher and Pickering
2018), while Mallorca 175 (Baldock 2014), Malta 108 (Balzan et al. 2016, 2017),
Ibiza and Formentera 77 (Baldock 2014), Madeira and Porto Santo 19 (Fellendorf
et al. 1999) and Menorca 13 species (Baldock 2014). A recent study by Kaloveloni
et al. (2018) reported the following bee species diversity for islands in the Aegean
Archipelago: Ana (59), Chios (208), Folegandros (46), Ikaria (102), Ios (78), Karpa-
thos (105), Kea (114), Kos (142), Kythnos (59), Limnos (178), Milos (38), Mykonos
(38), Paros (60), Samothraki (128), Santorini (67), Serifos (68), Syros (68), assos
(169), and Tinos (95). However, dierences in the sampling eort between islands
may bias bee diversity estimates. Sampling for wild bees on the island of Cyprus has
focused traditionally on the southern part of the island (Fig. 1). Collections of samples
reported for the rst time in the current study also concentrated on the southern part
of the island (Fig. 1), where the Republic of Cyprus exercises full control, because of
diculties in reaching and collecting samples from the northern part of Cyprus. e
under-sampling in the north of the island, both historical and present, probably leads
to an underestimation of the bee species richness on the island.
e bee fauna of Cyprus shows similarity to that of southern Europe and Middle
East (Levant), while the similarity with North Africa appears weaker. Cyprus meets the
denition of an oceanic island as it emerged from the sea, but the Messinian Salinity
Crisis that led to the virtual drying up of the Mediterranean around 6 mya (Nicolaou et
al. 2016) probably facilitated the arrival of many species of bees, as has been proposed
for other species, including mammals (Nicolaou et al. 2016). e more recent drop of
the sea level during the last glacial maxima of the Pleistocene (25,000 to 18,000 years
ago) that exposed underwater seamounts could also have aided bee dispersal from Asia
Minor (Anatolia) in a stepping stone model, even though bees would need to y over
40 km over water to reach the dierent islets (Nicolaou et al. 2016). A detailed analysis
of the biogeographical anities of the species present on the island will improve our
understanding of historical dispersal events.
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 105
ere are 2,051 bee species in Europe (Rasmont et al. 2017), as listed in the Eu-
ropean Red List of Bees, with 400 species endemic to Europe (20.4 %) (Nieto et al.
2014). Most endemic species occur in southern Europe with a higher diversity in the
Mediterranean, predominantly in the Iberian, Italian, and Balkan Peninsulas. Cyprus
with 21 endemic species ranks second after Crete with 52 endemics followed by Sicily
(10), Corsica (5) and Sardinia (1). e wild bee endemism rate for Cyprus (5.7 %)
is similar to that for butteries (6.1 %), with three endemics out of 49 species (John
2016), but much lower than that for Orthoptera (16.9 %, Siedle et al. 2016). e
endemic species are mainly medium- to small-sized bees and mostly ground nesters.
ey have been reported to forage on many plant species, some of which are endemic
to the island, including Onobrychis venosa (Fig. 2D), Teucrium cyprium, and Nepeta
troodi (Edwards et al. 2016).
Approximately 9 % of assessed European bee species are classied as threatened
[0.4 % – Critically Endangered, 2.4 % – Endangered, 1.2 % – Vulnerable, 5.2 % –
Near reatened, (Nieto et al. 2014)]. In Cyprus, the endemic Megachile cypricola is
listed as Critically Endangered. Field work conducted in support of the current study
has shown that the species is still present in Cyprus, but the population size and trend
still need to be evaluated to review its status. As the specimens were collected only on
the endemic plant Onobrychis venosa, a rst mitigation measure could be the conser-
vation of strong populations of the host plant. Moreover, Ammobatoides abdominalis,
Bombus niveatus and Parammobatodes minutus, listed as Endangered in Europe (Nieto
et al. 2014), require re-evaluation of their conservation status, as they have not been
collected from the island in recent years.
0
0.5
1
1.5
2
2.5
3
012345
log (species no)
Island area [log(km2)]
Cyprus
Figure 3. Diversity of bee fauna in 30 Mediterranean islands as a function of island area (logarithm trans-
formed data). See text for the islands and their untransformed bee species numbers. Cyprus represented
by the open circle.
Androulla I. Varnava et al. / ZooKeys 924: 1–114 (2020)
106
Climate change, degradation and fragmentation of natural habitats, urbanisa-
tion, monoculture farming and frequent use of pesticides aect the diversity of bees
throughout the world (Potts et al. 2010). Cyprus is no exception to the rule, as natural
areas are converted into development projects to meet the increasing needs of the
tourism and housing industry. e intensive use of pesticides in farm elds is another
factor that potentially impacts wild bee populations (Michener 2007), and anecdotal
evidence suggests that honeybee poisonings are frequent on the island. e value of
the 32 insect-pollinated crops grown on Cyprus exceeds €37 million euros annually
(Agricultural Statistics 2015), underlining the importance of taking measures to con-
serve both honeybee and wild bee populations. While the role of wild bees in pollinat-
ing agricultural crops has never been studied before on Cyprus, it is very likely that
they support and enhance agricultural production, especially in areas where honeybee
populations are low.
Conserving wild bees requires the establishment of a monitoring program to assess
the most signicant pressures on their populations and to identify eective conserva-
tion practices. Climate change and/or other anthropogenic pressures including agri-
cultural intensication and conversion of natural areas to urban fabric are prominent
factors that need to be studied, but without more recent data on population trends it is
almost impossible to design eective conservation practices. Many of the original loca-
tions where Mavromoustakis sampled for bees have been or are being converted into
urban areas. Future studies need to document the impact of urban development on bee
conservation and the potential value of anthropogenic habitats for wild bee conserva-
tion. e current work provides a baseline for future studies of wild bee diversity on
the island of Cyprus and elsewhere.
Conclusions
Mediterranean islands are well known for their bee diversity, a result of their isolation,
high oral diversity and optimal nesting conditions (Nieto et al. 2014, Kaloveloni et
al. 2018). Cyprus hosts 369 bee species, with 5.7 % endemic to the island. e island
hosts species of conservation concern in Europe, such as the Critically Endangered
Megachile cypricola and the Endangered Ammobatoides abdominalis, Bombus niveatus,
and Parammobatodes minutus. Conserving wild bees requires the establishment of a
monitoring program to assess their population trends, the most signicant threats on
their populations and to identify eective conservation practices. e current study is
the rst step towards conserving wild bees on the island of Cyprus.
Acknowledgements
We thank omas Wood (UK) (Andrena), Gerard Le Go (FR), Andreas Müller (CH)
and Christophe Praz (CH) (Megachilidae), Achik Dorchin (IL) (Eucera), Erwin Scheuchl
e wild bees (Hymenoptera, Apoidea) of the island of Cyprus 107
(DE), Bogdan Tomozei (RO) (Andrena), Michael Kuhlmann (DE) (Colletes), Alain Pau-
ly (BE) (Halictidae), Pierre Rasmont (BE) (Anthophorini), Göran Holmström (SE)
and Holger Dathe (DE) (Hylaeus), and Jan Smit (NL) (Nomada), for the identication,
comments, and suggestions on the species of the catalogue; Nicolas J. Vereecken for the
photos; and we acknowledge the support of the COST Action FA 1307 Sustainable
Pollination in Europe (SUPER-B) and the projects STEP – Status and Trends in Euro-
pean Pollinators (FP7) and ALARM – Assessing Large-scale environmental Risks with
tested Methods (FP6). We want to acknowledge Christian Schmid-Egger (DE), Martin
Jenner, Sophie Bagshaw, Nick Owens (UK), Elli Tzirkalli (CY), Stavroulla Louka (CY),
and Yiannis Varnava (CY) for their contribution in providing data and Ong Xin Rui for
her recommendations. We should also thank the Department of Agriculture, Ministry
of Agriculture, Rural Development and Environment of Cyprus people for access to the
Mavromoustakis collection and the Snow Entomological Museum Collection (Kansas,
USA) who provided online digitised data of specimens collected by Mavromoustakis
under a Creative Commons Attribution (CC-BY) 4.0 License.
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Supplementary material 1
Species list
Authors: Androulla I. Varnava, Stuart P.M. Roberts, Denis Michez, John S. Ascher,
eodora Petanidou, Stavroula Dimitriou, Jelle Devalez, Marilena Pittara, Menelaos
C. Stavrinides
Data type: species data
Explanation note: e list below is of species reported from Cyprus, but which are of
uncertain occurrence on the island or have been excluded recently.
Copyright notice: is dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). e Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Link: https://doi.org/10.3897/zookeys.924.38328.suppl1
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