OLICH: A reference library of DNA barcodes for Nordic lichens

Abstract

DNA barcodes are increasingly being used for species identification amongst the lichenised fungi. This paper presents a dataset aiming to provide an authoritative DNA barcode sequence library for a wide array of Nordic lichens.
We present 1324 DNA barcode sequences (nrITS) for 507 species in 175 genera and 25 orders. Thirty-eight species are new to GenBank and, for 25 additional species, ITS sequences are here presented for the first time. The dataset covers 20–21% of the Nordic lichenised species. Barcode gap analyses are given and discussed for the three genera Cladonia , Ramalina and Umbilicaria . The new combination Bryobilimbia fissuriseda (Poelt) Timdal, Marthinsen & Rui is proposed for Mycobilimbia fissuriseda and Nordic material of the species, currently referred to as Pseudocyphellaria crocata and Psoroma tenue ssp. boreale, are shown to belong in Pseudocyphellaria citrina and Psoroma cinnamomeum , respectively.

Full text

Biodiversity Data Journal 7: e36252
doi: 10.3897/BDJ.7.e36252
Data Paper
OLICH: A reference library of DNA barcodes for
Nordic lichens
Gunnhild Marthinsen , Siri Rui , Einar Timdal
‡ Natural History Museum, University of Oslo, Oslo, Norway
Corresponding author: Gunnhild Marthinsen (gunnhilm@nhm.uio.no)
Academic editor: Andreas Beck
Received: 16 May 2019 | Accepted: 31 Jul 2019 | Published: 19 Aug 2019
Citation: Marthinsen G, Rui S, Timdal E (2019) OLICH: A reference library of DNA barcodes for Nordic lichens.
Biodiversity Data Journal 7: e36252. https://doi.org/10.3897/BDJ.7.e36252
Abstract
Background
DNA barcodes are increasingly being used for species identification amongst the
lichenised fungi. This paper presents a dataset aiming to provide an authoritative DNA
barcode sequence library for a wide array of Nordic lichens.
New information
We present 1324 DNA barcode sequences (nrITS) for 507 species in 175 genera and 25
orders. Thirty-eight species are new to GenBank and, for 25 additional species, ITS
sequences are here presented for the first time. The dataset covers 20–21% of the Nordic
lichenised species. Barcode gap analyses are given and discussed for the three genera
Cladonia, Ramalina and Umbilicaria. The new combination Bryobilimbia fissuriseda (Poelt)
Timdal, Marthinsen & Rui is proposed for Mycobilimbia fissuriseda and Nordic material of
the species, currently referred to as Pseudocyphellaria crocata and Psoroma tenue ssp.
boreale, are shown to belong in Pseudocyphellaria citrina and Psoroma cinnamomeum,
respectively.
‡ ‡ ‡
© Marthinsen G et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are
credited.

Keywords
DNA barcoding, ITS, Lichenised fungi, Norway, Taxonomy
Introduction
Lichens are fungi living in mutualistic symbiosis with green algae and/or cyanobacteria.
They are highly polyphyletic and 20–30 independent lichenisation events may have
occurred during the evolution of higher Fungi (Schoch et al. 2009, Lücking et al. 2017a).
The lichenised fungi currently consists of more than 19,000 species, i.e. about 17% of all
known fungal species and 27% of all known ascomycete species (Lücking et al. 2017a).
They occur globally in all major terrestrial ecosystems, especially on trees, soil and rock.
Although a lichen thallus consists of a symbiotic association between a fungal exhabitant,
the mycobiont and one or more photosynthetic algal and/or cyanobacterial inhabitants, the
photobionts, the taxonomy and nomenclature follow that of the mycobiont. A lichen thallus
may house additional fungi, i.e. parasites, parasymbionts and endophytes (Hawksworth
1982, Richardson 1999, Spribille et al. 2016), making a lichen thallus a complex symbiotic
assemblage.
Identifying lichenised fungi has traditionally been based on morphological, anatomical and
chemical characters, but DNA-based identification is now increasingly being used. In 2012,
the internal transcribed spacer of nuclear ribosomal DNA (ITS) was recommended as the
primary fungal DNA barcode marker by the International Fungal Barcoding Consortium (Xu
2016), following studies on its suitability by for example Schoch et al. (2012). For
lichenised fungi, studies reporting successful use of the ITS marker for species
delimitations include Kelly et al. (2011), Zheng et al. (2011), Parnmen et al. (2013), Leavitt
et al. (2014), Magain and Serusiaux (2015) and Divakar et al. (2016). Pino-Bodas et al.
(2013), however, found ITS often failing for species delimitations in Cladonia and
investigated alternative markers.
In this paper, we present a DNA barcode dataset for Nordic lichens, including 1324
records, comprising 507 species in 175 genera and 25 different orders.
General description
Purpose: This dataset aims to provide an authoritative DNA barcode sequences library for
a wide array of Nordic lichens. The library should facilitate DNA-based identification of
species for both traditional molecular studies and DNA-metabarcoding studies and should
also be a resource for biosystematic research on Nordic lichens.
Additional information: Eighty sequences have been published previously, but are
included here as they were produced under the OLICH project, i.e. GenBank IDs
KT800002–07 (Westberg et al. 2015), KU687445–60 (McCune et al. 2016), KU873926–32
2Marthinsen G et al

(Timdal et al. 2016), KY682961 (Pykälä et al. 2018), 38 sequences within the series
MG838156–201 (Kistenich et al. 2018), MG968256–61 (Haugan and Timdal 2018) and
MK567159–64 (Haugan and Timdal 2019). In addition, 151 sequences (KY266831–981)
have been submitted to GenBank previously for an unpublished project.
Results: Thirty-eight species are new to GenBank: Arthonia phaeobaea, Arthopyrenia
salicis, Bactrospora corticola, Baeomyces carneus, Buellia arnoldii, B. dives, Candelariella
arctica, Cladonia cyanipes, Coccotrema citrinescens, Eopyrenula leucoplaca, Epilichen
glauconigellus, E. scabrosus, Gyalecta foveolaris, Lecanora atrosulphurea, L. leptacina,
Ochrolechia alaskana, O. alboflavescens, O. gowardii, O. mahluensis, Orphniospora
moriopsis, Phaeophyscia kairamoi, Pilophorus dovrensis, Pseudocyphellaria norvegica,
Pyrenula occidentalis, Ramalina cuspidata, Rhizocarpon badioatrum, R. bolanderi, R.
chioneum, R. jemtlandicum, R. leptolepis, R. rittokense, R. santessonii, R. subgeminatum,
Rinodina balanina, R. disjuncta, R. roboris, Stereocaulon capitellatum and S.
symphycheilum.
Twenty-five additional species are previously represented in GenBank, but not with ITS
sequences: Arthonia didyma, Bryobilimbia fissuriseda, Buellia epigaea, Callome
multipartita, Dendrographa latebrarum, Fuscidea mollis, Gyalecta flotowii, G. friesii, G.
geoica, Gyrographa gyrocarpa, Halecania alpivaga, Koerberiella wimmeriana, Opegrapha
vermicellifera, Pertusaria aspergilla, P. carneopallida, Phlyctis agelaea, Placynthiella
oligotropha, Psoronactis dilleniana, Pyrenula laevigata, Ramboldia elabens, Sagiolechia
protuberans, Stereocaulon grande, Thelotrema macrosporum, Tremolecia atrata and
Varicellaria lactea.
Project description
Title: The name OLICH refers to “contributions from the Oslo lichen herbarium”.
Personnel: Gunnhild Marthinsen (project coordinator), Siri Rui (project engineer), Einar
Timdal (taxonomic supervisor), all affiliated to the Natural History Museum, University of
Oslo.
Study area description: The Nordic countries, including Svalbard: Norway, mainland
(97.0% of the specimens), Sweden (2.0%), Finland (0.5%), Norway, Svalbard (0.5%),
Fig. 1.
Funding: This project is funded by the Norwegian Research Council (RCN 226134)
through the Norwegian Barcode of Life project (NorBOL). In addition, the Norwegian
Species Information Centre (Artsdatabanken) has funded four projects on lichens under
the Norwegian Taxonomy Initiative from which material has been collected and/or sampled
(project numbers 70184216, 70184227, 70184233, and 70184237).
OLICH: A reference library of DNA barcodes for Nordic lichens 3

Sampling methods
Sampling description: The material was partly freshly collected specimens from the field
and partly previously curated specimens in university herbaria. The distribution of
specimen age at seqencing is given in Fig. 2. The specimens were initially determined by
experts based on morphology and secondary chemistry and the identifications were re-
examined at the quality control after sequence acquisition. All collected material were, after
sampling, deposited in university herbaria and the entire material is housed in BG (2.1%),
H (0.1%), O (94.9%), SAV (0.2%), TRH (1.7%) and TROM (1.1%) (abbreviated according
to Index Herbariorum; http://sweetgum.nybg.org/science/ih).
Figure 1.
Distribution of the OLICH records
Figure 2.
Distribution of specimen age (years after collecting) at sequencing in the OLICH dataset.
4Marthinsen G et al

Thin-layer chromatography (TLC) was performed for identification of secondary
compounds on selected specimens in accordance with the methods of Culberson (1972),
as modified by Menlove (1974) and Culberson and Johnson (1982). Solvent system B' was
used for routine examination, A and C for additional confirmation.
Tissue of thallus or preferably apothecia (about 3 mm ) was assembled in 96 well plates
obtained from the Canadian Centre for DNA Barcoding (CCDB; http://ccdb.ca). One well
was left empty to serve as negative control. The freshly collected material was either
sampled in the field or at a field station. In the field, the specimen was first photographed,
then tissue was removed with a flame-decontaminated forceps and placed in the well and
finally the specimen was collected. Specimens requiring more controlled handling were
photographed and sampled either with forceps or scalpel under a binocular lens at a field
station. Wells containing fresh material were dried for several days with lids open in a
drying chamber containing silica gel. Herbarium material was sampled at the housing
institutions under binocular lenses.
Field data was stored on tablets and GPSs. Tissue sampling data and photography data
were transferred directly to standard BOLD spreadsheets, whereas the collection data
passed through herbarium databases for obtaining institution storing, voucher ID (= sample
ID), uniform nomenclature and taxonomical hierarchy and contact information. Material
sampled in herbaria used a similar pipeline, although simpler, as collection data was
already available.
The plates were sent to CCDB for DNA extraction (following Ivanova et al. 2011) and
sequencing of the ITS region (following Kuzmina and Ivanova 2011). Primers used were
ITS1-F or ITS5 (forward) and ITS4 (reverse) (Gardes and Bruns 1993, White et al. 1990).
Trace files and sequences were uploaded to BOLD (Ratnasingham and Hebert 2007). The
sequences were edited at CCDB, but occasionally inspected and re-edited at OLICH.
Quality control: All OLICH sequences were subjected to a local BLASTn search against a
database consisting of all lichen ITS sequences in GenBank (downloaded 26-06-2018) and
all unpublished lichen ITS sequences were produced at various projects at our own
institution, including the OLICH dataset. We inspected automatically generated distance-
based neighbour-joining trees for all genera represented in the OLICH dataset, i.e. we
merged all OLICH sequences with all GenBank ITS sequences from the same genus
before running ClustalW2 (http://www.clustal.org/clustal2) in batch mode (one run for each
genus) and saving the trees. For selected cases (Figs. 5–15), phylogenetic analyses were
performed through the following pipeline: selecting relevant additional sequences for
comparison from the GenBank download; aligning the dataset using the online version of
MAFFT (https://mafft.cbrc.jp/alignment/server); trimming ends and manually editing the
alignment using BioEdit 7.1.3.0 (Hall 1999); analysing the alignment by the maximum
likelihood method using RAxML v. 8.2.10 (Stamatakis 2014) by applying the “ML + rapid
bootstrap” method with 100 bootstrap repeats and the GTRGAMMA substitution model;
and editing the best-scoring tree using TreeGraph2 (Stöver and Müller 2010).
3
OLICH: A reference library of DNA barcodes for Nordic lichens 5

To further check that the sequences and species determinations were correct and also to
check whether species delimitation, based on traditional characters, coincides with ITS
species delimitation, we performed barcode gap analyses. The three genera in our dataset
with most sampled species were chosen for these analyses; Cladonia, Ramalina and
Umbilicaria. Sequences were aligned separately for each genus using the ClustalW
algorithm implemented in MEGA7 (Kumar et al. 2016), with additional manual editing.
Pairwise genetic distances between all specimens within each of the three genera were
calculated in MEGA7, using the K2P substitution model. The distances were summarised
in a bar plot, based on a histogram of all pairwise distances. For each species pair within
the genera, we plotted the maximum intraspecific distance against the minimum distance
to nearest neighbour-species.
Geographic coverage
Description: The Nordic countries, including Svalbard (Fig. 1).
Coordinates: 58.02°N and 78.94°N Latitude; 5.06°E and 31.08°E Longitude.
Taxonomic coverage
Description: The OLICH dataset comprises 1324 records of 507 species belonging to 25
orders, 8 classes and 2 phyla of fungi (Table 1). The Nordic lichen flora comprises some
2400–2500 lichenised species (2387 are listed in Santesson et al. (2004)), i.e. the dataset
comprises about 20–21% of the currently known Nordic species. Several taxonomic groups
are still poorly understood, however, making the production of authoritative DNA-barcodes
for many species premature due to uncertain taxonomy and/or lack of fresh material. The
dataset is hence biased towards the more well-known groups, mainly the macrolichens in
the orders Lecanorales (233 spp.), Peltigerales (62 spp.) and Caliciales (37 spp.) of the
class Lecanoromycetes (Ascomycota). The taxonomy and nomenclature follows Nordin et
al. (2019) unless commented on under Observations below.
Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Acarospora
glaucocarpa
Acarosporales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201295 OLICH3852-17 MK811746
Table 1.
List of specimens included in the OLICH dataset (sequence length >200bp, 1324 records, 507
species). Thin Layer Chromotagrophy (TLC, Chemistry-column) was only done in species
complexes where chemical data is relevant for the identification of the species or where there
are chemical strains within the species.
6Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Acarospora
molybdina
Acarosporales NO, Finnmark,
2014, E.
Timdal
O-L-195556 OLICH1514-14 KY266941
Acarospora
wahlenbergii
Acarosporales NO, Hedmark,
2013, E.
Timdal
O-L-184511 OLICH737-13 MK811773
Acarospora
wahlenbergii
Acarosporales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184363 OLICH742-13 MK812641
Acarospora
wahlenbergii
Acarosporales NO, Telemark,
2013, E.
Timdal
O-L-184419 OLICH870-13 MK811926
Acolium inquinans Caliciales NO, Nordland,
2012, J.T.
Klepsland
O-L-186246 OLICH2077-15 MK812481
Acolium inquinans Caliciales NO, Buskerud,
2012, T.H.
Hofton
O-L-194494 OLICH2080-15 MK811701
Acolium inquinans Caliciales NO, Aust-
Agder, 2014,
J.T. Klepsland
O-L-200169 OLICH3168-16 MK812233
Acolium inquinans Caliciales NO, Hedmark,
2014, R.
Haugan
O-L-207991 OLICH3827-17 MK811747
Alectoria nigricans Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195780 OLICH1652-14 MK812557
Alectoria nigricans Lecanorales NO, Finnmark,
2014, M.
Westberg
O-L-195858 OLICH1703-14 KY266967
Alectoria nigricans Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195886 OLICH1731-14 KY266922
Alectoria nigricans Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195950 OLICH1795-14 KY266831
OLICH: A reference library of DNA barcodes for Nordic lichens 7

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Alectoria nigricans Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196314 OLICH2244-15 MK812235
Alectoria nigricans Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200895 OLICH2548-16 MK812358
Alectoria
ochroleuca
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195746 OLICH1618-14 KY266936
Alectoria
ochroleuca
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195883 OLICH1728-14 KY266959
Alectoria
ochroleuca
Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195823 OLICH2195-15 MK811939
Alectoria
ochroleuca
Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200894 OLICH2547-16 MK812226
Alectoria
ochroleuca
Lecanorales NO, Oppland,
2012, K.A. Lye
O-L-206884 OLICH4129-17 MK812278
Alectoria
sarmentosa
Lecanorales NO, Nord-
Trøndelag,
2012, J.T.
Klepsland
O-L-186255 OLICH1441-14 MK812505
Alectoria
sarmentosa
Lecanorales NO, Sogn og
Fjordane,
2009, T.H.
Hofton
O-L-186870 OLICH1452-14 MK812223
Alectoria
sarmentosa
Lecanorales NO, Østfold,
2013, B.P.
Løfall & O.
Wergeland
Krog
O-L-187364 OLICH1460-14 MK812403
Alectoria
sarmentosa
Lecanorales NO, Sogn og
Fjordane,
2014, K.J.
Grimstad & O.
Olsen
O-L-193185 OLICH1478-14 MK812717
8Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Alectoria
sarmentosa
Lecanorales NO, Sør-
Trøndelag,
2014, S. Vatne
O-L-200758 OLICH3273-16 MK812509
Allantoparmelia
alpicola
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195748 OLICH1620-14 KY266874
Allantoparmelia
alpicola
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195850 OLICH1695-14 KY266926
Allantoparmelia
alpicola
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195899 OLICH1744-14 KY266946
Allantoparmelia
alpicola
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196324 OLICH2254-15 MK811991
Allantoparmelia
alpicola
Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201391 OLICH4173-17 MK811876
Anaptychia ciliaris Caliciales NO, Aust-
Agder, 2012,
J.T. Klepsland
O-L-186030 OLICH1419-14 MK812701
Anaptychia ciliaris Caliciales NO, Hedmark,
2011, T.H.
Hofton
O-L-194359 OLICH1483-14 MK812099
Anaptychia ciliaris Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195591 OLICH1532-14 MK812249
Anaptychia ciliaris Caliciales NO, Finnmark,
2014, G. Thor
O-L-195736 OLICH1608-14 KY266912
Anaptychia ciliaris Caliciales NO, Akershus,
2014, E.
Timdal
O-L-199982 OLICH1956-15 MK812295
Anaptychia ciliaris Caliciales NO, Oslo,
2014, E.
Timdal
O-L-198929 OLICH2482-16 MK812296
Anaptychia
runcinata
Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195767 OLICH1639-14 MK812680
OLICH: A reference library of DNA barcodes for Nordic lichens 9

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Anaptychia
runcinata
Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196019 OLICH1864-14 MK811742
Anaptychia
runcinata
Caliciales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208017 OLICH4189-17 MK811645
Arctocetraria
andrejevii
Lecanorales NO, Finnmark,
2012, H.
Henriksen
O-L-180951 OLICH869-13 MK811992 norrangiformic acid
(minor), rangiformic
acid (major)
Arctoparmelia
centrifuga
Lecanorales NO, Troms,
2014, R.
Haugan
O-L-198774 OLICH2399-16 MK812567
Arctoparmelia
centrifuga
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200879 OLICH2536-16 MK812513
Arctoparmelia
centrifuga
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184727 OLICH942-13 MK812611
Arctoparmelia
incurva
Lecanorales NO, Finnmark,
2014, T.
Tønsberg
O-L-195717 OLICH1589-14 KY266852
Arctoparmelia
incurva
Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195832 OLICH2204-15 MK811976
Arthonia didyma Arthoniales NO, Nord-
Trøndelag,
2013, J.T.
Klepsland
O-L-197934 OLICH4022-17 MK812348
Arthonia
phaeobaea
Arthoniales NO, Sør-
Trøndelag,
2013, J.T.
Klepsland
O-L-198129 OLICH3187-16 MK812215
Arthopyrenia
salicis
Pleosporales NO, Nord-
Trøndelag,
2016, J.T.
Klepsland
O-L-206752 OLICH3471-17 MK812299
10 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Arthopyrenia
salicis
Pleosporales NO, Møre og
Romsdal,
2016, J.T.
Klepsland
O-L-206774 OLICH3493-17 MK812520
Asahinea
chrysantha
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195754 OLICH1626-14 KY266872
Bacidia
bagliettoana
Lecanorales NO, Akershus,
2009, R.
Haugan
O-L-161668 OLICH282-13 MG838180
Bacidia
bagliettoana
Lecanorales NO, Troms,
2009, J.T.
Klepsland
O-L-164908 OLICH284-13 MG838178
Bacidia
subincompta
Lecanorales NO, Finnmark,
2011, E.
Timdal
O-L-170623 OLICH198-13 MG838157 no lichen
substances
Bacidia
subincompta
Lecanorales NO, Sogn og
Fjordane,
2013, J.T.
Klepsland
O-L-197862 OLICH2086-15 MG838186
Bacidia
subincompta
Lecanorales NO, Østfold,
2001, B.P.
Løfall
O-L-105331 OLICH310-13 MG838165
Bacidia
subincompta
Lecanorales NO, Oslo,
2014, J.T.
Klepsland
O-L-200148 OLICH3180-16 MG838175
Bacidia
subincompta
Lecanorales NO, Nordland,
2015, J.T.
Klepsland
O-L-206520 OLICH3420-17 MG838176
Bacidia vermifera Lecanorales NO, Troms,
2011, J.T.
Klepsland
O-L-177162 OLICH264-13 MG838160
Bacidia vermifera Lecanorales NO, Troms,
2009, J.T.
Klepsland
O-L-165007 OLICH299-13 MG838169
OLICH: A reference library of DNA barcodes for Nordic lichens 11

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bactrospora
corticola
Arthoniales NO, Nord-
Trøndelag,
2010, H.
Holien
TRH-
L-13774
OLICH1208-14 MK811961
Bactrospora
corticola
Arthoniales NO, Nord-
Trøndelag,
2009, H.
Holien
TRH-
L-12953
OLICH1209-14 MK811741
Bactrospora
corticola
Arthoniales NO, Nord-
Trøndelag,
2009, H.
Holien
TRH-
L-12974
OLICH1210-14 MK811643
Bactrospora
corticola
Arthoniales NO, Sør-
Trøndelag,
2010, G.
Gaarder
TRH-
L-651772
OLICH1211-14 MK811979
Bactrospora
corticola
Arthoniales NO, Sør-
Trøndelag,
2014, S. Vatne
O-L-200761 OLICH2438-16 MK812695
Baeomyces
carneus
Baeomycetales NO, Østfold,
2003, B. P.
Løfall
O-L-125189 OLICH4208-17 MK811898
Baeomyces
carneus
Baeomycetales NO, Østfold,
2010, J.T.
Klepsland
O-L-174982 OLICH4421-17 MK812113
Baeomyces
placophyllus
Baeomycetales NO, Telemark,
2014, E.
Timdal
O-L-196267 OLICH1942-15 MK812574
Baeomyces
placophyllus
Baeomycetales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196313 OLICH2243-15 MK812640
Baeomyces rufus Baeomycetales NO, Telemark,
2014, S. Rui &
E. Timdal
O-L-194160 OLICH1481-14 MK811923
Baeomyces rufus Baeomycetales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200015 OLICH1989-15 MK812462
12 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Baeomyces rufus Baeomycetales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201257 OLICH2765-16 MK812004
Baeomyces rufus Baeomycetales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201258 OLICH2766-16 MK812643
Baeomyces rufus Baeomycetales NO, Aust-
Agder, 2014,
J.T. Klepsland
O-L-200581 OLICH3271-16 MK812401
Bilimbia lobulata Lecanorales NO, Nordland,
2001, B.P.
Løfall
O-L-105772 OLICH469-13 MK812451
Bilimbia lobulata Lecanorales NO, Troms,
2009, J.T.
Klepsland
O-L-164992 OLICH470-13 MK812712
Bilimbia lobulata Lecanorales NO, Nordland,
2000, S. Rui &
E. Timdal
O-L-51771 OLICH471-13 MK812122
Bilimbia lobulata Lecanorales NO, Oppland,
1995, R.
Haugan & E.
Timdal
O-L-19936 OLICH630-13 MK812573
Bilimbia
sabuletorum
Lecanorales NO, Buskerud,
2008, J.T.
Klepsland
O-L-158388 OLICH206-13 MK812338
Bilimbia
sabuletorum
Lecanorales NO, Telemark,
2010, S. Rui &
E. Timdal
O-L-163841 OLICH207-13 MK812048 no lichen
substances
Bilimbia
sabuletorum
Lecanorales NO, Oslo,
2009, J.T.
Klepsland
O-L-164642 OLICH208-13 MK812294
Bilimbia
sabuletorum
Lecanorales NO, Aust-
Agder, 2010,
J.T. Klepsland
O-L-175077 OLICH209-13 MK812254
OLICH: A reference library of DNA barcodes for Nordic lichens 13

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bilimbia
sabuletorum
Lecanorales NO, Hedmark,
2004, R.
Haugan
O-L-131717 OLICH632-13 MK811845
Blastenia
ammiospila
Teloschistales NO, Finnmark,
2014, U. Arup
O-L-195889 OLICH1734-14 KY266873
Blastenia
crenularia
Teloschistales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196018 OLICH1863-14 MK811924
Blastenia
crenularia
Teloschistales NO, Nordland,
2014, J.T.
Klepsland
O-L-200301 OLICH3173-16 MK811921
Blastenia
crenularia
Teloschistales NO, Akershus,
2013, S. Rui &
E. Timdal
O-L-203051 OLICH3289-16 MK812322
Brianaria lutulata Lecanorales NO, Aust-
Agder, 2015,
J.T. Klepsland
O-L-206489 OLICH3390-17 MK812659
Brianaria
tuberculata
Lecanorales NO, Aust-
Agder, 2015,
J.T. Klepsland
O-L-206488 OLICH3389-17 MK811824
Brodoa atrofusca Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200891 OLICH2544-16 MK811833
Brodoa atrofusca Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184308 OLICH725-13 MK811829 atranorin (major),
physodic acid
(minor),
protocetraric acid
(major)
Brodoa
intestiniformis
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-190610 OLICH1474-14 MK811819
Brodoa
intestiniformis
Lecanorales NO,
Hordaland,
2014, E.
Timdal
O-L-195806 OLICH1678-14 MK811842
14 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Brodoa
intestiniformis
Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-195819 OLICH2191-15 MK812115
Brodoa
intestiniformis
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184709 OLICH928-13 MK812552
Brodoa oroarctica Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200939 OLICH2660-16 MK811789
Brodoa oroarctica Lecanorales NO, Hedmark,
2017, E.
Timdal
O-L-207794 OLICH4155-17 MK812012
Brodoa oroarctica Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184297 OLICH726-13 MK812600
Bryobilimbia
diapensiae
Lecideales NO, Finnmark,
2011, E.
Timdal
O-L-170679 OLICH1167-14 MK812130
Bryobilimbia
diapensiae
Lecideales NO, Nordland,
2012, R.
Haugan
O-L-182225 OLICH1168-14 MK811983
Bryobilimbia
diapensiae
Lecideales NO, Sør-
Trøndelag,
2004, R.
Haugan
O-L-142025 OLICH144-11 MK812681
Bryobilimbia
diapensiae
Lecideales NO, Troms,
2003, R.
Haugan
O-L-141675 OLICH145-11 MK812677
Bryobilimbia
diapensiae
Lecideales NO, Finnmark,
2014, S. Rui
O-L-195960 OLICH1805-14 KY266915
Bryobilimbia
fissuriseda
Lecideales NO, Oppland,
2005, E.
Timdal
O-L-148945 OLICH627-13 MK811695
Bryobilimbia
fissuriseda
Lecideales NO, Oppland,
2006, R.
Haugan
O-L-145176 OLICH628-13 MK812621
OLICH: A reference library of DNA barcodes for Nordic lichens 15

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bryobilimbia
fissuriseda
Lecideales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184390 OLICH647-13 MK811896
Bryobilimbia
hypnorum
Lecideales NO, Nordland,
2014, J.T.
Klepsland
O-L-200331 OLICH3196-16 MK812370
Bryobilimbia
hypnorum
Lecideales NO, Aust-
Agder, 2014,
J.T. Klepsland
O-L-200584 OLICH3198-16 MK811910
Bryocaulon
divergens
Lecanorales NO, Hedmark,
2013, R.
Haugan
O-L-190313 OLICH1472-14 MK812080
Bryocaulon
divergens
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195982 OLICH1827-14 KY266917
Bryocaulon
divergens
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196360 OLICH1919-15 MK812069
Bryocaulon
divergens
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196317 OLICH2247-15 MK811658
Bryoplaca
sinapisperma
Teloschistales NO, Troms,
2014, J.T.
Klepsland
O-L-200396 OLICH4055-17 MK812160
Bryoria bicolor Lecanorales NO, Buskerud,
2009, S.
Svantesson
O-L-157404 OLICH1369-14 MK811780
Bryoria bicolor Lecanorales NO, Rogaland,
2011, J.T.
Klepsland
O-L-176874 OLICH1396-14 MK812469
Bryoria bicolor Lecanorales NO, Hedmark,
2012, R.
Haugan
O-L-179764 OLICH1400-14 MK811677
Bryoria bicolor Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183745 OLICH1403-14 MK812128
16 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bryoria bicolor Lecanorales NO, Hedmark,
2012, R.
Haugan
O-L-183997 OLICH1404-14 MK812174
Bryoria capillaris Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-199996 OLICH1970-15 MK812045 alectorialic acid,
barbatolic acid
Bryoria capillaris Lecanorales SE, Lule
Lappmark,
2014, E.
Timdal
O-L-195685 OLICH2008-15 MK812229 alectorialic acid,
barbatolic acid
Bryoria capillaris Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-195816 OLICH2188-15 MK812443 alectorialic acid,
barbatolic acid
Bryoria capillaris Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184687 OLICH871-13 MK812425 alectorialic acid,
barbatolic acid,
fumarprotocetraric
acid
Bryoria fremontii Lecanorales FI, Perä-
Pohjanmaa,
2014, E.
Timdal
O-L-195488 OLICH1498-14 MK812707
Bryoria fremontii Lecanorales SE,
Härjedalen,
2014, E.
Timdal
O-L-195688 OLICH2011-15 MK812076
Bryoria fremontii Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-207993 OLICH3830-17 MK812563
Bryoria furcellata Lecanorales FI, Perä-
Pohjanmaa,
2014, E.
Timdal
O-L-195487 OLICH1497-14 MK812435
Bryoria furcellata Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195834 OLICH2206-15 MK811793
OLICH: A reference library of DNA barcodes for Nordic lichens 17

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bryoria furcellata Lecanorales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200527 OLICH2618-16 MK812653
Bryoria furcellata Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-207994 OLICH3831-17 MK812397
Bryoria implexa Lecanorales FI, Perä-
Pohjanmaa,
2014, E.
Timdal
O-L-195485 OLICH1495-14 MK812092 fumarprotocetraric
acid
Bryoria implexa Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195887 OLICH1732-14 KY266878 no lichen
substances
Bryoria implexa Lecanorales SE, Torne
Lappmark,
2014, E.
Timdal
O-L-195665 OLICH2004-15 MK812090 no lichen
substances
Bryoria implexa Lecanorales SE, Lule
Lappmark,
2014, E.
Timdal
O-L-195686 OLICH2009-15 MK812159 no lichen
substances
Bryoria implexa Lecanorales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197823 OLICH2087-15 MK812714 fumarprotocetraric
acid
Bryoria implexa Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196325 OLICH2255-15 MK812347 fumarprotocetraric
acid
Bryoria implexa Lecanorales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200539 OLICH2604-16 MK812375 fumarprotocetraric
acid
Bryoria implexa Lecanorales NO, Hedmark,
2017, E.
Timdal
O-L-207753 OLICH4156-17 MK811675 no lichen
substances
Bryoria implexa Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184688 OLICH872-13 MK812231 fumarprotocetraric
acid
18 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bryoria
nadvornikiana
Lecanorales NO, Sogn og
Fjordane,
2008, G.
Gaarder
O-L-155318 OLICH1367-14 MK812079
Bryoria
nadvornikiana
Lecanorales NO, Hedmark,
2012, R.
Haugan
O-L-183994 OLICH2172-15 MK811678
Bryoria
nadvornikiana
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-176834 OLICH2173-15 MK812465
Bryoria
nadvornikiana
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-198885 OLICH2397-16 MK812326
Bryoria
nadvornikiana
Lecanorales NO, Hedmark,
2017, E.
Timdal
O-L-207754 OLICH4157-17 MK812417 alectorialic acid
(major), barbatolic
acid (minor),
fumarprotocetraric
acid (minor)
Bryoria
nadvornikiana
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184689 OLICH873-13 MK812390
Bryoria nitidula Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195715 OLICH1587-14 KY266934 fumarprotocetraric
acid (major),
protocetraric acid
(minor)
Bryoria simplicior Lecanorales FI, Inarin
Lappi, 2014, E.
Timdal
O-L-195506 OLICH1505-14 MK812146
Bryoria simplicior Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195825 OLICH2197-15 MK812415
Bryoria simplicior Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200936 OLICH2562-16 MK811887
Bryoria smithii Lecanorales NO, Vest-
Agder, 2011,
J.T. Klepsland
O-L-177044 OLICH2165-15 MK811714
OLICH: A reference library of DNA barcodes for Nordic lichens 19

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Bryoria smithii Lecanorales NO, Rogaland,
2011, J.T.
Klepsland
O-L-176881 OLICH2166-15 MK812250
Bryoria tenuis Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196345 OLICH1905-15 MK812591
Buellia arnoldii Caliciales NO, Aust-
Agder, 2015,
H. Holien
TRH-
L-16501
OLICH3670-17 MK811634
Buellia badia Caliciales NO, Vestfold,
2015, E.
Timdal
O-L-198941 OLICH2498-16 MK812426
Buellia disciformis Caliciales NO, Troms,
2014, J.T.
Klepsland
O-L-200381 OLICH4052-17 MK812587
Buellia dives Caliciales NO, Nord-
Trøndelag,
2015, H.
Holien
TRH-
L-16419
OLICH3631-17 MK811893
Buellia epigaea Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195523 OLICH1507-14 KY266900
Buellia
erubescens
Caliciales NO, Nord-
Trøndelag,
2015, U.
Nordin
O-L-201275 OLICH3229-16 MK811955
Bunodophoron
melanocarpum
Lecanorales NO, Rogaland,
2011, H.
Fjeldstad
O-L-177411 OLICH1399-14 MK812059
Bunodophoron
melanocarpum
Lecanorales NO,
Hordaland,
2011, J.T.
Klepsland
O-L-177097 OLICH2161-15 MK812620
Bunodophoron
melanocarpum
Lecanorales NO, Sogn og
Fjordane,
2010, J.T.
Klepsland
O-L-175004 OLICH2163-15 MK811949
20 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Calicium
adspersum
Caliciales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200822 OLICH2410-16 MK812068
Calicium pinicola Caliciales NO, Oppland,
2015, R.
Haugan
O-L-204191 OLICH4087-17 MK811674
Calicium viride Caliciales NO, Hedmark,
2017, S. Rui &
E. Timdal
O-L-207800 OLICH4161-17 MK812371
Callome
multipartita
Peltigerales NO, Buskerud,
2013, S. Reiso
& T. Høitomt
O-L-194256 OLICH2054-15 MK811803
Calogaya
biatorina
Teloschistales NO, Sogn og
Fjordane,
2013, J.T.
Klepsland
O-L-197877 OLICH2459-16 MK812288
Calogaya
decipiens
Teloschistales NO, Oslo,
2014, R.
Haugan et al.
O-L-196061 OLICH1877-14 MK812143
Calogaya
decipiens
Teloschistales NO, Hedmark,
1998, R.
Haugan
O-L-55723 OLICH4389-17 MK812258
Calogaya
decipiens
Teloschistales NO, Akershus,
2010, R.
Haugan
O-L-167011 OLICH4390-17 MK811994
Caloplaca
chlorina
Teloschistales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-198079 OLICH4423-17 MK811786
Caloplaca
chlorina
Teloschistales NO, Telemark,
2014, J.T.
Klepsland
O-L-200720 OLICH4425-17 MK811988
Caloplaca
demissa
Teloschistales NO, Rogaland,
2010, J.T.
Klepsland
O-L-200085 OLICH3270-16 MK812156
OLICH: A reference library of DNA barcodes for Nordic lichens 21

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Caloplaca
demissa
Teloschistales NO, Sogn og
Fjordane,
2010, S. Vatne
O-L-169301 OLICH3953-17 MK812032
Caloplaca
monacensis
Teloschistales NO, Buskerud,
1998, H. Bratli
O-L-39356 OLICH4422-17 MK811632
Calvitimela
armeniaca
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195741 OLICH1613-14 KY266899 alectorialic acid
Calvitimela livida Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195842 OLICH1687-14 KY266947 atranorin, stictic
acid
Calvitimela
melaleuca
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195711 OLICH1583-14 KY266838 alectorialic acid,
atranorin
Calvitimela
melaleuca
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195915 OLICH1760-14 KY266897 alectorialic acid,
norstictic acid
Candelaria
concolor
Candelariales NO, Rogaland,
2015, T.
Tønsberg & E.
Nygaard
BG-
L-98168
OLICH3699-17 MK812350
Candelariella
arctica
Candelariales NO, Finnmark,
2014, J.T.
Klepsland
O-L-200509 OLICH3171-16 MK812015
Candelariella
aurella
Candelariales NO, Troms,
1998, R.
Armstrong
TROM_L
_50030
OLICH3532-17 MK812178
Candelariella
coralliza
Candelariales NO, Akershus,
2014, E.
Timdal
O-L-199992 OLICH1966-15 MK812334
Candelariella
coralliza
Candelariales NO, Buskerud,
2014, E.
Timdal
O-L-195833 OLICH2205-15 MK812024
Candelariella
coralliza
Candelariales NO, Aust-
Agder, 2004,
J.T. Klepsland
O-L-186001 OLICH2460-16 MK811884
22 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Candelariella
coralliza
Candelariales NO, Telemark,
2015, E.
Timdal & S.
Rui
O-L-200875 OLICH3836-17 MK812646
Candelariella
placodizans
Candelariales NO, Troms,
2003, J.W.
Bjerke
TROM_L
_521176
OLICH3583-17 MK812262
Candelariella
placodizans
Candelariales NO, Troms,
2009, J.T.
Klepsland
O-L-164896 OLICH3949-17 MK811687
Candelariella
placodizans
Candelariales NO, Nordland,
2000, E.
Timdal
O-L-51849 OLICH3950-17 MK811709
Candelariella
placodizans
Candelariales NO, Oppland,
2009, R.
Haugan
O-L-159822 OLICH3951-17 MK812369
Candelariella
placodizans
Candelariales NO, Telemark,
2008, J.T.
Klepsland
O-L-164600 OLICH3952-17 MK812523
Candelariella
vitellina
Candelariales NO, Nord-
Trøndelag,
1999, H. Bratli
O-L-41044 OLICH3941-17 MK811713
Candelariella
vitellina
Candelariales NO, Vest-
Agder, 2011,
H. Bratli
O-L-175901 OLICH3943-17 MK811697
Candelariella
vitellina
Candelariales NO, Oslo,
2010, R.
Haugan
O-L-166987 OLICH3948-17 MK812582
Cetraria aculeata Lecanorales NO, Finnmark,
2014, M.
Bendiksby
O-L-195862 OLICH1707-14 KY266883
Cetraria aculeata Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196315 OLICH2245-15 MK811967
Cetraria aculeata Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196032 OLICH3808-17 MK811881
OLICH: A reference library of DNA barcodes for Nordic lichens 23

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cetraria aculeata Lecanorales NO, Østfold,
2013, K.A. Lye
O-L-194757 OLICH4090-17 MK811694
Cetraria aculeata Lecanorales NO, Vest-
Agder, 2011,
H. Bratli
O-L-175907 OLICH4091-17 MK811732
Cetraria
ericetorum
Lecanorales NO, Hedmark,
2013, R.
Haugan
O-L-190276 OLICH1469-14 MK811667
Cetraria
ericetorum
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195750 OLICH1622-14 KY266868
Cetraria
ericetorum
Lecanorales NO, Buskerud,
2015, E.
Timdal & S.
Rui
O-L-200904 OLICH3839-17 MK811635
Cetraria
ericetorum
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184702 OLICH884-13 MK811886
Cetraria islandica Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-200003 OLICH1977-15 MK812195
Cetraria islandica Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-195818 OLICH2190-15 MK812355
Cetraria islandica Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196304 OLICH2234-15 MK812550
Cetraria islandica Lecanorales NO, Sør-
Trøndelag,
2017, E.
Timdal
O-L-208279 OLICH4164-17 MK812647
Cetraria islandica Lecanorales NO, Sør-
Trøndelag,
2017, E.
Timdal
O-L-208278 OLICH4165-17 MK812026
Cetraria islandica Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184703 OLICH885-13 MK812121
24 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cetraria muricata Lecanorales NO,
Hordaland,
2014, E.
Timdal
O-L-195799 OLICH1671-14 MK812154
Cetraria muricata Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-195815 OLICH2187-15 MK812716
Cetraria nigricans Lecanorales NO, Troms,
2012, J.T.
Klepsland
O-L-186197 OLICH1436-14 MK812225
Cetraria nigricans Lecanorales NO, Finnmark,
2011, E.
Timdal
O-L-170662 OLICH2159-15 MK812613
Cetraria odontella Lecanorales NO, Buskerud,
2012, T.H.
Hofton
O-L-194457 OLICH1485-14 MK812629
Cetraria sepincola Lecanorales NO, Hedmark,
2013, R.
Haugan
O-L-190020 OLICH1463-14 MK812418
Cetraria sepincola Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200881 OLICH2538-16 MK812133
Cetraria sepincola Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184705 OLICH886-13 MK811790
Cetrariella
commixta
Lecanorales NO, Finnmark,
2014, M.
Westberg
O-L-195926 OLICH1771-14 KY266843
Cetrariella delisei Lecanorales NO, Oppland,
2013, R.
Haugan
O-L-190044 OLICH1465-14 MK812542
Cetrariella delisei Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195559 OLICH1517-14 KY266903
Cetrariella delisei Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195763 OLICH1635-14 KY266858
OLICH: A reference library of DNA barcodes for Nordic lichens 25

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cetrariella delisei Lecanorales NO, Finnmark,
2014, M.
Westberg
O-L-195882 OLICH1727-14 KY266955
Cetrariella delisei Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195946 OLICH1791-14 KY266892
Cetrariella delisei Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200893 OLICH2546-16 MK811762
Cetrariella
fastigiata
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195560 OLICH1518-14 KY266875
Cetrariella
fastigiata
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195762 OLICH1634-14 KY266870
Cetrariella
fastigiata
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195985 OLICH1830-14 KY266890
Cetrariella
fastigiata
Lecanorales NO, Finnmark,
2011, E.
Timdal
O-L-170481 OLICH2154-15 MK811637
Cetrariella
fastigiata
Lecanorales NO, Troms,
2011, J.T.
Klepsland
O-L-177156 OLICH2155-15 MK812493
Cetrelia
cetrarioides
Lecanorales NO, Telemark,
2005, S. Reiso
O-L-141329 OLICH1354-14 MK811823 anziaic acid
(minor), atranorin
(major), perlatolic
acid (major), 4-O-
methylolivetoric
acid (minor)
Cetrelia
cetrarioides
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-176836 OLICH1395-14 MK812086 anziaic acid
(minor), atranorin
(major), perlatolic
acid (major), 4-O-
methylolivetoric
acid (minor)
26 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cetrelia
cetrarioides
Lecanorales NO, Sogn og
Fjordane,
2009, T.H.
Hofton
O-L-186891 OLICH1453-14 MK812180 anziaic acid
(minor), atranorin
(major), perlatolic
acid (major), 4-O-
methylolivetoric
acid (minor)
Cetrelia
cetrarioides
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200022 OLICH2470-16 MK812062 anziaic acid
(minor), atranorin
(major), perlatolic
acid (major), 4-O-
methylolivetoric
acid (minor)
Cetrelia
cetrarioides
Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201355 OLICH2740-16 MK812302 anziaic acid
(minor), atranorin
(major), perlatolic
acid (major), 4-O-
methylolivetoric
acid (minor)
Cetrelia
cetrarioides
Lecanorales NO, Buskerud,
2016, M.
Bendiksby & E.
Timdal
O-L-208029 OLICH4257-17 MK812008 anziaic acid
(minor), atranorin
(major), perlatolic
acid (major), 4-O-
methylolivetoric
acid (minor)
Cetrelia
monachorum
Lecanorales NO, Buskerud,
2012, T.H.
Hofton
O-L-194448 OLICH4095-17 MK811729 anziaic acid
(minor), atranorin
(major), imbricaric
acid (major), 4-O-
denmethylimbricaric
acid (minor)
Chaenotheca
chlorella
Coniocybales NO, Sør-
Trøndelag,
2014, S. Vatne
O-L-200765 OLICH2447-16 MK812013
Chaenotheca
ferruginea
Coniocybales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200838 OLICH2426-16 MK812287
OLICH: A reference library of DNA barcodes for Nordic lichens 27

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Chaenotheca
gracilenta
Coniocybales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201318 OLICH2790-16 MK812452
Chaenotheca
gracillima
Coniocybales NO, Sør-
Trøndelag,
2014, S. Vatne
O-L-200773 OLICH2446-16 MK812119
Cladonia
albonigra
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195659 OLICH1999-15 MK812147
Cladonia
amaurocraea
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195776 OLICH1648-14 MK811717
Cladonia
amaurocraea
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195877 OLICH1722-14 KY266961
Cladonia
amaurocraea
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196364 OLICH1923-15 MK812471
Cladonia
amaurocraea
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196316 OLICH2246-15 MK811885
Cladonia
arbuscula
Lecanorales NO, Finnmark,
2014, A.
Kosuthova
O-L-195866 OLICH1711-14 KY266933
Cladonia
arbuscula
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196358 OLICH1917-15 MK812466
Cladonia
arbuscula
Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-196268 OLICH1943-15 MK811810
Cladonia
arbuscula
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196301 OLICH2231-15 MK812036
28 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia
arbuscula
Lecanorales NO, Vestfold,
2015, E.
Timdal
O-L-198942 OLICH2497-16 MK811763
Cladonia
bellidiflora
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195740 OLICH1612-14 KY266902
Cladonia
bellidiflora
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195881 OLICH1726-14 KY266913
Cladonia
bellidiflora
Lecanorales NO, Finnmark,
2014, A.
Kosuthova
O-L-195906 OLICH1751-14 KY266845
Cladonia
bellidiflora
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195958 OLICH1803-14 KY266921
Cladonia
caespiticia
Lecanorales NO, Østfold,
2012, J.T.
Klepsland
O-L-186036 OLICH1420-14 MK811962
Cladonia
caespiticia
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200916 OLICH2576-16 MK812605
Cladonia
caespiticia
Lecanorales NO,
Hordaland,
2011, J.T.
Klepsland
O-L-183605 OLICH4081-17 MK811712
Cladonia cariosa Lecanorales NO, Buskerud,
2012, E.W.
Hanssen et al.
O-L-186370 OLICH1443-14 MK811826
Cladonia cariosa Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184343 OLICH743-13 MK812622
Cladonia carneola Lecanorales NO, Finnmark,
2014, A.
Kosuthova
O-L-195963 OLICH1808-14 KY266855
Cladonia cenotea Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196307 OLICH2237-15 MK811749
OLICH: A reference library of DNA barcodes for Nordic lichens 29

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia cenotea Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-198892 OLICH2393-16 MK812468
Cladonia cenotea Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-207986 OLICH3824-17 MK812541
Cladonia cenotea Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201300 OLICH3855-17 MK812267
Cladonia cenotea Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184714 OLICH932-13 MK812699
Cladonia
chlorophaea
Lecanorales NO, Buskerud,
2016, S. Rui &
E. Timdal
O-L-208034 OLICH4262-17 MK812242
Cladonia
coccifera
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195895 OLICH1740-14 KY266935
Cladonia
coniocraea
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196310 OLICH2240-15 MK812094
Cladonia
coniocraea
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200908 OLICH2569-16 MK812138
Cladonia cornuta Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196260 OLICH2267-15 MK812497
Cladonia cornuta Lecanorales NO, Hedmark,
2015, E.
Timdal
O-L-200857 OLICH2508-16 MK812414
Cladonia cornuta Lecanorales NO, Hedmark,
2016, E.
Timdal
O-L-201516 OLICH3979-17 MK811731
Cladonia cornuta Lecanorales NO, Telemark,
2017, E.
Timdal
O-L-208059 OLICH4167-17 MK812047
30 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia cornuta Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184715 OLICH933-13 MK812277
Cladonia crispata Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195880 OLICH1725-14 KY266865
Cladonia crispata Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195824 OLICH2196-15 MK811863
Cladonia crispata Lecanorales NO, Telemark,
2016, S. Rui &
E. Timdal
O-L-205005 OLICH3995-17 MK811728
Cladonia crispata Lecanorales NO, Vest-
Agder, 2016,
S. Rui & E.
Timdal
O-L-204984 OLICH3996-17 MK812514
Cladonia crispata Lecanorales NO, Telemark,
2017, E.
Timdal
O-L-208058 OLICH4275-17 MK811707
Cladonia
cyanipes
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195758 OLICH1630-14 KY266889
Cladonia
cyanipes
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195981 OLICH1826-14 KY266977
Cladonia
cyathomorpha
Lecanorales NO, Vest-
Agder, 2012,
J.T. Klepsland
O-L-186087 OLICH1430-14 MK811836
Cladonia
cyathomorpha
Lecanorales NO, Aust-
Agder, 2010,
J.T. Klepsland
O-L-175272 OLICH2147-15 MK811648
Cladonia
decorticata
Lecanorales SE, Torne
Lappmark,
2014, E.
Timdal
O-L-195679 OLICH2006-15 MK811738
OLICH: A reference library of DNA barcodes for Nordic lichens 31

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia
ecmocyna
Lecanorales NO,
Hordaland,
2014, E.
Timdal
O-L-195793 OLICH1665-14 MK812344
Cladonia
ecmocyna
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195956 OLICH1801-14 KY266972
Cladonia
ecmocyna
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200933 OLICH2559-16 MK812085
Cladonia fimbriata Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195743 OLICH1615-14 KY266953
Cladonia fimbriata Lecanorales NO, Vest-
Agder, 2011,
J.T. Klepsland
O-L-183619 OLICH2145-15 MK812488 fumarprotocetraric
acid
Cladonia fimbriata Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200909 OLICH2570-16 MK811629
Cladonia furcata Lecanorales NO, Buskerud,
2011, K.A. Lye
& T. Høitomt
O-L-194749 OLICH1492-14 MK811985
Cladonia furcata Lecanorales NO, Finnmark,
2014, T.
Tønsberg
O-L-195964 OLICH1809-14 KY266930
Cladonia furcata Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196298 OLICH2228-15 MK812624
Cladonia furcata Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196308 OLICH2238-15 MK811830
Cladonia furcata Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201273 OLICH3847-17 MK812251
Cladonia gracilis Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195764 OLICH1636-14 KY266906
32 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia gracilis Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196359 OLICH1918-15 MK812362
Cladonia gracilis Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196321 OLICH2251-15 MK811813
Cladonia gracilis Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196262 OLICH2269-15 MK811638
Cladonia gracilis Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196047 OLICH3802-17 MK812342
Cladonia grayi Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195757 OLICH1629-14 KY266978
Cladonia krogiana Lecanorales NO, Telemark,
2003, B.P.
Løfall & A.
Ognedal
O-L-127158 OLICH1337-14 MK811907 barbatic acid,
chlorovinetorin
Cladonia krogiana Lecanorales NO, Akershus,
2004, B.P.
Løfall
O-L-129021 OLICH1339-14 MK811663 barbatic acid,
chlorovinetorin
Cladonia krogiana Lecanorales NO, Akershus,
2012, J.T.
Klepsland
O-L-186068 OLICH2136-15 MK812057
Cladonia
macrophylla
Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-200002 OLICH1976-15 MK811782
Cladonia
macrophylla
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200021 OLICH2469-16 MK812683
Cladonia
macrophyllodes
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196379 OLICH1938-15 MK812614
OLICH: A reference library of DNA barcodes for Nordic lichens 33

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia maxima Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195829 OLICH2201-15 MK811693
Cladonia maxima Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184361 OLICH692-13 MK812518
Cladonia mitis Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184706 OLICH887-13 MK811894
Cladonia
parasitica
Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-207988 OLICH3825-17 MK811639
Cladonia
parasitica
Lecanorales NO, Hedmark,
2016, E.
Timdal
O-L-201525 OLICH3989-17 MK812470
Cladonia
phyllophora
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196375 OLICH1934-15 MK812644
Cladonia
phyllophora
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196320 OLICH2250-15 MK812551
Cladonia pleurota Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195959 OLICH1804-14 KY266968 usnic acid, zeorin
Cladonia
pulvinata
Lecanorales NO, Sør-
Trøndelag,
2015, H.
Holien
TRH-
L-16327
OLICH2920-16 MK811846 fumarprotocetraric
acid, psoromic acid
Cladonia pyxidata Lecanorales NO, Østfold,
2012, J.T.
Klepsland
O-L-186049 OLICH1425-14 MK812571 fumarprotocetraric
acid
Cladonia pyxidata Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195914 OLICH1759-14 KY266877 fumarprotocetraric
acid (major),
protocetraric acid
(minor)
34 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia pyxidata Lecanorales NO, Oslo,
2014, S. Rui &
E. Timdal
O-L-196073 OLICH1889-14 MK812676
Cladonia
rangiferina
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195755 OLICH1627-14 KY266884
Cladonia
rangiferina
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195774 OLICH1646-14 MK811970
Cladonia
rangiferina
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196302 OLICH2232-15 MK812460
Cladonia
rangiferina
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196037 OLICH3806-17 MK811708
Cladonia
rangiferina
Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201298 OLICH3854-17 MK812260
Cladonia
rangiformis
Lecanorales NO, Telemark,
2012, S. Reiso
O-L-194209 OLICH2062-15 MK812118 atranorin
Cladonia
rangiformis
Lecanorales NO, Rogaland,
2013, J.T.
Klepsland
O-L-198309 OLICH2351-15 MK811656
Cladonia rei Lecanorales NO, Telemark,
2014, S. Rui &
E. Timdal
O-L-194142 OLICH1479-14 MK811774
Cladonia rei Lecanorales NO, Oslo,
2014, J.T.
Klepsland
O-
L-200262b
OLICH4086-17 MK812389 fumarprotocetraric
acid, homosekikaic
acid
Cladonia
squamosa
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184733 OLICH1415-14 MK812303
Cladonia
squamosa
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195646 OLICH1558-14 MK811958
OLICH: A reference library of DNA barcodes for Nordic lichens 35

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia
squamosa
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195909 OLICH1754-14 KY266973
Cladonia
squamosa
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196309 OLICH2239-15 MK812360
Cladonia stellaris Lecanorales NO, Finnmark,
2014, A.
Kosuthova
O-L-195905 OLICH1750-14 KY266887
Cladonia stellaris Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196263 OLICH2270-15 MK812506
Cladonia stellaris Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184707 OLICH926-13 MK812280
Cladonia
straminea
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195976 OLICH1821-14 KY266880
Cladonia strepsilis Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-200000 OLICH1974-15 MK812544
Cladonia strepsilis Lecanorales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208340 OLICH4270-17 MK812494
Cladonia stygia Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-196269 OLICH1944-15 MK812684
Cladonia stygia Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196303 OLICH2233-15 MK812337
Cladonia stygia Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184708 OLICH927-13 MK812669
Cladonia
subcervicornis
Lecanorales NO, Rogaland,
2013, S. Vatne
O-L-194122 OLICH2072-15 MK812018
36 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia
subcervicornis
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196028 OLICH3794-17 MK812661
Cladonia
subcervicornis
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196053 OLICH3811-17 MK812155
Cladonia
subcervicornis
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-204985 OLICH3998-17 MK811825
Cladonia
subcervicornis
Lecanorales NO, Vest-
Agder, 2011,
J.T. Klepsland
O-L-176962 OLICH4113-17 MK812379
Cladonia
subfurcata
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195987 OLICH1832-14 KY266923
Cladonia
subfurcata
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196319 OLICH2249-15 MK812649
Cladonia
subfurcata
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200934 OLICH2560-16 MK812597
Cladonia subulata Lecanorales NO, Sør-
Trøndelag,
2013, R.
Haugan
O-L-190281 OLICH1471-14 MK812581
Cladonia subulata Lecanorales FI, Inarin
Lappi, 2014, E.
Timdal
O-L-195490 OLICH1500-14 MK811811
Cladonia subulata Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196299 OLICH2229-15 MK812279
Cladonia subulata Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200914 OLICH2574-16 MK811787
OLICH: A reference library of DNA barcodes for Nordic lichens 37

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Cladonia
sulphurina
Lecanorales NO, Finnmark,
2014, A.
Kosuthova
O-L-195911 OLICH1756-14 KY266846
Cladonia
symphycarpa
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195597 OLICH1535-14 MK811971 atranorin
Cladonia turgida Lecanorales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-199980 OLICH1954-15 MK811657
Cladonia turgida Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200932 OLICH2558-16 MK812027
Cladonia uncialis Lecanorales NO, Østfold,
2013, R.
Haugan
O-L-190182 OLICH1467-14 MK812589
Cladonia uncialis Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195939 OLICH1784-14 KY266974
Cladonia
verticillata
Lecanorales NO,
Hordaland,
2014, E.
Timdal
O-L-195797 OLICH1669-14 MK812461
Cladonia
verticillata
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196312 OLICH2242-15 MK811806
Cladonia
verticillata
Lecanorales NO, Sør-
Trøndelag,
2015, H.
Holien
TRH-
L-16326
OLICH2921-16 MK812385 fumarprotocetraric
acid
Cladonia zopfii Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196044 OLICH3804-17 MK811805
Coccotrema
citrinescens
Pertusariales NO, Nordland,
2015, T.
Tønsberg & A.
Botnen
BG-
L-99172
OLICH3741-17 MK811814
38 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Coccotrema
citrinescens
Pertusariales NO, Nordland,
2015, T.
Tønsberg
BG-
L-98897
OLICH3754-17 MK811895
Coccotrema
citrinescens
Pertusariales NO, Møre og
Romsdal,
2015, T.
Tønsberg & Z.
Palice
BG-
L-99069
OLICH3774-17 MK812329
Coccotrema
citrinescens
Pertusariales NO, Nordland,
2013, J.T.
Klepsland
O-L-198020 OLICH4212-17 MK812183
Coccotrema
citrinescens
Pertusariales NO, Sør-
Trøndelag,
2011, R.
Haugan
O-L-173864 OLICH4403-17 MK812186 stictic acid
Cornicularia
normoerica
Lecanorales NO, Aust-
Agder, 2014,
J.T. Klepsland
O-L-200754 OLICH2603-16 MK812042
Cornicularia
normoerica
Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201280 OLICH3850-17 MK811980
Cystocoleus
ebeneus
Capnodiales NO, Sør-
Trøndelag,
1998, R.
Haugan
O-L-55770 OLICH3942-17 MK811795
Dactylina
ramulosa
Lecanorales NO, Troms,
2012, J.T.
Klepsland
O-L-186199 OLICH1438-14 MK812632
Dendrographa
latebrarum
Arthoniales NO, Møre og
Romsdal,
2016, H.
Holien
TRH-
L-16972
OLICH3682-17 MK811843
Dermatocarpon
bachmannii
Verrucariales NO, Østfold,
2000, B. P.
Løfall et al.
O-L-78257 OLICH2297-15 MK812540
OLICH: A reference library of DNA barcodes for Nordic lichens 39

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Dermatocarpon
bachmannii
Verrucariales NO, Østfold,
2005, B. P.
Løfall
O-L-147654 OLICH2305-15 MK812444
Dermatocarpon
deminuens
Verrucariales NO, Østfold,
2000, B. P.
Løfall
O-L-78264 OLICH2298-15 MK812502
Dermatocarpon
deminuens
Verrucariales NO, Østfold,
1998, B. P.
Løfall
O-L-36303 OLICH2299-15 MK812476
Dermatocarpon
deminuens
Verrucariales NO, Østfold,
2000, B. P.
Løfall et al.
O-L-78258 OLICH2304-15 MK812075
Dermatocarpon
luridum
Verrucariales NO, Buskerud,
2009, T.H.
Hofton
O-L-187021 OLICH1455-14 MK812236
Dermatocarpon
meiophyllizium
Verrucariales NO, Østfold,
2006, B. P.
Løfall
O-L-147725 OLICH2296-15 MK812273
Dermatocarpon
meiophyllizium
Verrucariales NO, Hedmark,
2006, R.
Haugan
O-L-145251 OLICH2300-15 MK812522
Dermatocarpon
meiophyllizium
Verrucariales NO, Østfold,
2001, B. P.
Løfall
O-L-105979 OLICH2306-15 MK811820
Dermatocarpon
miniatum
Verrucariales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196009 OLICH1854-14 MK812245
Dibaeis
baeomyces
Pertusariales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200009 OLICH1983-15 MK812560
Dibaeis
baeomyces
Pertusariales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201296 OLICH2796-16 MK812219
40 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Diploschistes
scruposus
Ostropales NO, Finnmark,
2014, S.
Fernandez
O-L-195913 OLICH1758-14 KY266920
Diploschistes
scruposus
Ostropales NO, Aust-
Agder, 2013,
J.T. Klepsland
O-L-198319 OLICH2350-15 MK811671
Diplotomma
alboatrum
Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195577 OLICH1527-14 MK812220
Eopyrenula
leucoplaca
Vezdaeales NO, Telemark,
2016, J.T.
Klepsland
O-L-206771 OLICH3490-17 MK811948
Epilichen
glauconigellus
Rhizocarpales NO, Oppland,
2012, S. Rui &
E. Timdal
O-L-179907 OLICH1100-14 MK812566
Epilichen
glauconigellus
Rhizocarpales NO, Troms,
2003, R.
Haugan
O-L-141634 OLICH1101-14 MK811705
Epilichen
scabrosus
Rhizocarpales NO, Hedmark,
2012, S. Rui &
E. Timdal
O-L-179955 OLICH1102-14 MK812558
Epilichen
scabrosus
Rhizocarpales NO, Troms,
2009, J.T.
Klepsland
O-L-164936 OLICH832-13 MK812386
Evernia divaricata Lecanorales NO, Buskerud,
2011, T.H.
Hofton
O-L-194431 OLICH1484-14 MK812536
Evernia divaricata Lecanorales NO, Oppland,
2012, T.H.
Hofton
O-L-194526 OLICH2315-15 MK811891
Evernia
mesomorpha
Lecanorales NO, Hedmark,
2012, T.H.
Hofton
O-L-194511 OLICH1489-14 MK812555
Evernia
mesomorpha
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200008 OLICH1982-15 MK811689
OLICH: A reference library of DNA barcodes for Nordic lichens 41

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Evernia
mesomorpha
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-176825 OLICH2308-15 MK811679
Evernia
mesomorpha
Lecanorales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200526 OLICH2614-16 MK812464
Evernia prunastri Lecanorales NO, Oslo,
2014, S. Rui &
E. Timdal
O-L-196076 OLICH1892-14 MK811993
Evernia prunastri Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-199988 OLICH1962-15 MK812247
Evernia prunastri Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196292 OLICH2222-15 MK812058
Evernia prunastri Lecanorales NO, Hedmark,
2015, E.
Timdal
O-L-200856 OLICH2507-16 MK811838
Evernia prunastri Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200923 OLICH2531-16 MK811722
Flavocetraria
cucullata
Lecanorales NO, Østfold,
2013, B.P.
Løfall
O-L-187362 OLICH1459-14 MK812129
Flavocetraria
cucullata
Lecanorales NO, Finnmark,
2014, S. Rui
O-L-195752 OLICH1624-14 KY266924
Flavocetraria
cucullata
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195778 OLICH1650-14 MK811878
Flavocetraria
cucullata
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196338 OLICH2276-15 MK812660
Flavocetraria
cucullata
Lecanorales NO, Svalbard,
2015, S. Rui
O-L-203288 OLICH2707-16 MK812458
42 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Flavocetraria
cucullata
Lecanorales NO, Buskerud,
2015, E.
Timdal & S.
Rui
O-L-200903 OLICH3838-17 MK812490
Flavocetraria
cucullata
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184721 OLICH938-13 MK812603
Flavocetraria
nivalis
Lecanorales NO, Østfold,
2013, B.P.
Løfall
O-L-187361 OLICH1458-14 MK812252
Flavocetraria
nivalis
Lecanorales NO, Finnmark,
2014, S. Rui
O-L-195723 OLICH1595-14 KY266896
Flavocetraria
nivalis
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195777 OLICH1649-14 MK811972
Flavocetraria
nivalis
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196311 OLICH2241-15 MK811652
Flavocetraria
nivalis
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196339 OLICH2277-15 MK812419
Flavocetraria
nivalis
Lecanorales NO, Svalbard,
2015, S. Rui
O-L-203283 OLICH2702-16 MK811681
Flavocetraria
nivalis
Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-207989 OLICH3829-17 MK812064
Flavocetraria
nivalis
Lecanorales NO, Buskerud,
2015, E.
Timdal & S.
Rui
O-L-200902 OLICH3837-17 MK812356
Flavoparmelia
soredians
Lecanorales NO, Rogaland,
2013, E.
Nygaard
BG-
L-98015
OLICH2437-16 MK812200
Flavoplaca citrina Teloschistales NO, Nord-
Trøndelag,
2015, V.
Alstrup
O-L-201322 OLICH2845-16 MK811952
OLICH: A reference library of DNA barcodes for Nordic lichens 43

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Flavoplaca citrina Teloschistales NO, Sør-
Trøndelag,
2015, R.
Haugan
O-L-204037 OLICH3891-17 MK812194
Flavoplaca
flavocitrina
Teloschistales NO, Hedmark,
2015, R.
Haugan
O-L-204199 OLICH3887-17 MK811821
Flavoplaca marina Teloschistales NO, Rogaland,
2010, J.T.
Klepsland
O-L-200087 OLICH3206-16 MK811686
Flavoplaca marina Teloschistales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200616 OLICH3207-16 MK811757
Frutidella
caesioatra
Lecanorales NO, Troms,
2014, J.T.
Klepsland
O-L-200359 OLICH3257-16 MK812163
Frutidella
caesioatra
Lecanorales NO, Sogn og
Fjordane,
2010, E.
Timdal
O-L-163395 OLICH387-13 MK811625
Frutidella
caesioatra
Lecanorales NO, Nordland,
2010, E.
Timdal
O-L-163746 OLICH388-13 MK812446
Frutidella
caesioatra
Lecanorales NO, Nordland,
2010, E.
Timdal
O-L-163752 OLICH390-13 MK812346
Frutidella
caesioatra
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184399 OLICH657-13 MK812561
Fuscidea mollis Umbilicariales NO, Finnmark,
2014, M.
Westberg
O-L-195707 OLICH1579-14 KY266980 divaricatic acid
Fuscidea mollis Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195917 OLICH1762-14 KY266834 divaricatic acid
44 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Fuscopannaria
ahlneri
Peltigerales NO, Nordland,
2008, T.H.
Hofton
O-L-186679 OLICH2464-16 MK812382
Fuscopannaria
confusa
Peltigerales NO, Nordland,
2013, J.T.
Klepsland
O-L-198017 OLICH2465-16 MK812003
Fuscopannaria
ignobilis
Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201286 OLICH2675-16 MK811940
Fuscopannaria
ignobilis
Peltigerales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201281 OLICH2778-16 MK812023
Fuscopannaria
ignobilis
Peltigerales NO, Sør-
Trøndelag,
2015, H.
Holien
TRH-
L-16696
OLICH3629-17 MK812459
Fuscopannaria
ignobilis
Peltigerales NO, Sør-
Trøndelag,
2012, H.
Fjeldstad
O-L-203080 OLICH4062-17 MK812103
Fuscopannaria
mediterranea
Peltigerales NO, Sogn og
Fjordane,
2013, J.T.
Klepsland
O-L-197869 OLICH2096-15 MK812182
Fuscopannaria
mediterranea
Peltigerales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199530 OLICH3166-16 MK811995
Fuscopannaria
praetermissa
Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-196265 OLICH1940-15 MK812190
Fuscopannaria
praetermissa
Peltigerales NO, Nordland,
2013, J.T.
Klepsland
O-L-198031 OLICH2097-15 MK811734
OLICH: A reference library of DNA barcodes for Nordic lichens 45

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Fuscopannaria
praetermissa
Peltigerales NO, Finnmark,
2003, E.
Timdal
O-L-123194 OLICH3883-17 MK812710
Fuscopannaria
praetermissa
Peltigerales NO, Troms,
2009, J.T.
Klepsland
O-L-174932 OLICH3884-17 MK811837
Gyalecta flotowii Ostropales NO,
Hordaland,
2013, J.B.
Jordal
O-L-199472 OLICH3165-16 MK811791
Gyalecta flotowii Ostropales NO, Sogn og
Fjordane,
2013, H.
Fjeldstad
O-L-203030 OLICH4063-17 MK811730
Gyalecta
foveolaris
Ostropales NO, Finnmark,
2014, T.B.
Wheeler
O-L-195735 OLICH1607-14 KY266833
Gyalecta friesii Ostropales NO, Sør-
Trøndelag,
2014, S. Vatne
O-L-200781 OLICH2442-16 MK812083
Gyalecta geoica Ostropales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200544 OLICH3164-16 MK811956
Gyalecta jenensis Ostropales NO, Finnmark,
2014, M.
Westberg
O-L-195708 OLICH1580-14 KY266916
Gyalecta jenensis Ostropales NO, Finnmark,
2014, A.
Millanes
O-L-195860 OLICH1705-14 KY266964
Gyalecta jenensis Ostropales NO, Nordland,
2013, J.T.
Klepsland
O-L-198022 OLICH4034-17 MK811684
Gyalecta ulmi Ostropales NO, Sør-
Trøndelag,
2014, S. Vatne
O-L-200778 OLICH2443-16 MK812657
46 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Gyalolechia
bracteata
Teloschistales NO, Finnmark,
2011, E.
Timdal
O-L-170861 OLICH3215-16 MK811756
Gyalolechia
flavorubescens
Teloschistales NO, Telemark,
2014, E.
Timdal
O-L-195786 OLICH1658-14 MK811715
Gyrographa
gyrocarpa
Arthoniales NO, Nord-
Trøndelag,
2015, H.
Holien
TRH-
L-16879
OLICH3651-17 MK812164
Gyrographa
gyrocarpa
Arthoniales NO, Aust-
Agder, 2015,
J.T. Klepsland
O-L-204523 OLICH4234-17 MK812125
Haematomma
ochroleucum
Lecanorales NO,
Hordaland,
2015, T.
Tønsberg
BG-
L-99143
OLICH3704-17 MK812424
Haematomma
ochroleucum
Lecanorales NO, Nordland,
2015, T.
Tønsberg
BG-
L-99181
OLICH3749-17 MK812361
Haematomma
ochroleucum
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196048 OLICH3817-17 MK812222
Halecania
alpivaga
Leprocaulales NO, Finnmark,
2014, M.
Westberg
O-L-195847 OLICH1692-14 KY266832
Hertelidea
botryosa
Lecanorales NO, Hedmark,
2016, E.
Timdal
O-L-201530 OLICH3988-17 MK811999
Hertelidea
botryosa
Lecanorales NO, Sør-
Trøndelag,
2007, E.
Timdal
O-L-158480 OLICH415-13 MK812335
Heterodermia
speciosa
Caliciales NO, Buskerud,
2009, T.H.
Hofton
O-L-186983 OLICH1454-14 MK812396
OLICH: A reference library of DNA barcodes for Nordic lichens 47

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Heterodermia
speciosa
Caliciales NO, Oppland,
2011, J.T.
Klepsland
O-L-176833 OLICH2287-15 MK812055
Heterodermia
speciosa
Caliciales NO, Oppland,
2011, J.T.
Klepsland
O-L-176831 OLICH2288-15 MK812697
Heterodermia
speciosa
Caliciales NO, Oppland,
2011, J.T.
Klepsland
O-L-176827 OLICH2290-15 MK812504
Heterodermia
speciosa
Caliciales NO, Oppland,
2011, T.
Høitomt et al.
O-L-176666 OLICH2291-15 MK812537
Heterodermia
speciosa
Caliciales NO, Oppland,
2011, J.T.
Klepsland
O-L-176840 OLICH2292-15 MK812549
Heterodermia
speciosa
Caliciales NO, Oppland,
2011, J.T.
Klepsland
O-L-176824 OLICH2293-15 MK811755
Heterodermia
speciosa
Caliciales NO, Hedmark,
2011, R.
Haugan
O-L-173228 OLICH2294-15 MK812405
Hypocenomyce
scalaris
Umbilicariales SE, Dalarna,
2014, E.
Timdal
O-L-195692 OLICH2014-15 MK812269
Hypocenomyce
scalaris
Umbilicariales NO, Telemark,
2013, J.T.
Klepsland
O-L-198107 OLICH2352-15 MK811723
Hypocenomyce
scalaris
Umbilicariales NO, Telemark,
2013, J.T.
Klepsland
O-L-198111 OLICH2366-15 MK812170
Hypocenomyce
scalaris
Umbilicariales NO, Hedmark,
2016, E.
Timdal
O-L-201526 OLICH3987-17 MK811809
Hypocenomyce
scalaris
Umbilicariales SE, Jämtland,
2017, E.
Timdal
O-L-208182 OLICH4285-17 MK812211
48 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Hypogymnia
austerodes
Lecanorales NO, Hedmark,
2013, R.
Haugan
O-L-190094 OLICH1466-14 MK812650
Hypogymnia
austerodes
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196349 OLICH1908-15 MK812410
Hypogymnia
austerodes
Lecanorales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200537 OLICH2613-16 MK812531
Hypogymnia
bitteri
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184690 OLICH874-13 MK811649
Hypogymnia
farinacea
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196258 OLICH2265-15 MK811676
Hypogymnia
farinacea
Lecanorales NO, Hedmark,
2010, R.
Haugan
O-L-183979 OLICH2279-15 MK812081
Hypogymnia
farinacea
Lecanorales NO, Hedmark,
2010, R.
Haugan
O-L-183983 OLICH2280-15 MK812607
Hypogymnia
farinacea
Lecanorales NO, Hedmark,
2010, R.
Haugan
O-L-183984 OLICH2281-15 MK812314
Hypogymnia
farinacea
Lecanorales NO, Hedmark,
2011, R.
Haugan
O-L-174340 OLICH2283-15 MK812440
Hypogymnia
farinacea
Lecanorales NO, Hedmark,
2010, R.
Haugan
O-L-183985 OLICH2285-15 MK812305
Hypogymnia
farinacea
Lecanorales NO, Buskerud,
2013, J.T.
Klepsland
O-L-198103 OLICH2369-15 MK812704
OLICH: A reference library of DNA barcodes for Nordic lichens 49

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Hypogymnia
farinacea
Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201310 OLICH2785-16 MK811721
Hypogymnia
hultenii
Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201260 OLICH2767-16 MK811640
Hypogymnia
incurvoides
Lecanorales NO, Rogaland,
2012, J.T.
Klepsland
O-L-186120 OLICH1433-14 MK811919 atranorin,
physodalic acid,
physodic acid,
protocetraric acid
(minor)
Hypogymnia
incurvoides
Lecanorales NO, Rogaland,
2012, J.T.
Klepsland
O-L-186139 OLICH1435-14 MK812392 atranorin,
physodalic acid,
physodic acid,
protocetraric acid
(minor)
Hypogymnia
incurvoides
Lecanorales NO, Nord-
Trøndelag,
2014, J.T.
Klepsland
O-L-200733 OLICH2620-16 MK812655 atranorin,
physodalic acid,
physodic acid
Hypogymnia
incurvoides
Lecanorales NO,
Hordaland,
2014, J.T.
Klepsland
O-L-200683 OLICH3161-16 MK812638 atranorin,
physodalic acid,
physodic acid
Hypogymnia
physodes
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195888 OLICH1733-14 KY266854
Hypogymnia
physodes
Lecanorales NO, Finnmark,
2014, M.
Westberg
O-L-195921 OLICH1766-14 KY266919
Hypogymnia
physodes
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200883 OLICH2565-16 MK811818
50 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Hypogymnia
physodes
Lecanorales NO,
Hordaland,
2014, J.T.
Klepsland
O-L-200741 OLICH2621-16 MK812700 3-hydroxyphysodic
acid, atranorin,
physodalic acid,
physodic acid
Hypogymnia
physodes
Lecanorales NO, Nord-
Trøndelag,
2014, J.T.
Klepsland
O-L-200731 OLICH3158-16 MK812007 3-hydroxyphysodic
acid, atranorin,
physodalic acid,
physodic acid
Hypogymnia
physodes
Lecanorales NO, Nordland,
2014, J.T.
Klepsland
O-L-200736 OLICH3159-16 MK812416 3-hydroxyphysodic
acid, atranorin,
physodalic acid,
physodic acid
Hypogymnia
physodes
Lecanorales NO, Akershus,
2014, J.T.
Klepsland
O-L-200255 OLICH3160-16 MK812648
Hypogymnia
physodes
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184691 OLICH875-13 MK812139
Hypogymnia
tubulosa
Lecanorales NO, Telemark,
2014, S. Rui &
E. Timdal
O-L-194145 OLICH1480-14 MK811733
Hypogymnia
tubulosa
Lecanorales NO, Rogaland,
2015, T.
Tønsberg & E.
Nygaard
BG-
L-98167
OLICH3698-17 MK812120
Hypogymnia
tubulosa
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184692 OLICH876-13 MK811664
Hypogymnia
vittata
Lecanorales NO, Buskerud,
2008, G.
Gaarder
O-L-155319 OLICH1368-14 MK811984
Hypogymnia
vittata
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195648 OLICH1560-14 MK812063
Hypogymnia
vittata
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200917 OLICH2577-16 MK812228
OLICH: A reference library of DNA barcodes for Nordic lichens 51

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Hypogymnia
vittata
Lecanorales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200536 OLICH2612-16 MK812445 atranorin, physodic
acid, vittatolic acid
Hypogymnia
vittata
Lecanorales NO, Akershus,
2017, S. Rui &
E. Timdal
O-L-206863 OLICH4108-17 MK812679
Hypotrachyna
afrorevoluta
Lecanorales NO, Rogaland,
2011, J.T.
Klepsland
O-L-176869 OLICH2182-15 MK812595
Hypotrachyna
afrorevoluta
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-205001 OLICH3991-17 MK811750
Hypotrachyna
laevigata
Lecanorales NO, Rogaland,
2010, J.T.
Klepsland
O-L-175129 OLICH1391-14 MK812534
Hypotrachyna
laevigata
Lecanorales NO, Rogaland,
2011, J.T.
Klepsland
O-L-176890 OLICH1397-14 MK812711
Hypotrachyna
revoluta
Lecanorales NO, Rogaland,
2005, J.T.
Klepsland
O-L-141145 OLICH1352-14 MK812078
Hypotrachyna
revoluta
Lecanorales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197837 OLICH2099-15 MK812441
Hypotrachyna
revoluta
Lecanorales NO, Rogaland,
2011, T.
Høitomt
O-L-176684 OLICH2183-15 MK812141
Hypotrachyna
revoluta
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-205002 OLICH3990-17 MK812580
Hypotrachyna
sinuosa
Lecanorales NO, Rogaland,
2016, T.
Tønsberg
BG-
L-99501
OLICH3758-17 MK812199
Icmadophila
ericetorum
Pertusariales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184734 OLICH1416-14 MK812230
52 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Icmadophila
ericetorum
Pertusariales NO, Finnmark,
2014, E.
Timdal
O-L-195557 OLICH1515-14 KY266881
Icmadophila
ericetorum
Pertusariales NO, Finnmark,
2014, H.
Holien
O-L-195756 OLICH1628-14 KY266943
Icmadophila
ericetorum
Pertusariales NO, Finnmark,
2014, T.
Tønsberg
O-L-195874 OLICH1719-14 KY266876
Icmadophila
ericetorum
Pertusariales NO, Nord-
Trøndelag,
2015, H.
Holien
TRH-
L-16544
OLICH3627-17 MK811879
Imshaugia
aleurites
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-196257 OLICH2264-15 MK812672
Japewia
subaurifera
Lecanorales NO, Nord-
Trøndelag,
2012, T.
Tønsberg
BG-
L-94252
OLICH577-13 MK812331
Japewia
subaurifera
Lecanorales NO, Finnmark,
2012, T.
Tønsberg
BG-
L-94707
OLICH578-13 MK812698
Japewia
subaurifera
Lecanorales NO, Nordland,
2012, T.
Tønsberg
BG-
L-94659
OLICH596-13 MK812623
Japewia
tornoensis
Lecanorales NO, Buskerud,
2009, J.T.
Klepsland
O-L-165163 OLICH357-13 MK812546
Japewia
tornoensis
Lecanorales NO, Buskerud,
2011, J.T.
Klepsland
O-L-176857 OLICH358-13 MK812668
Koerberiella
wimmeriana
Lecideales NO,
Hordaland,
2010, E.
Timdal
O-L-163472 OLICH414-13 MK812168
OLICH: A reference library of DNA barcodes for Nordic lichens 53

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Lasallia pustulata Umbilicariales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-204990 OLICH4001-17 MK811849
Lecanactis
abietina
Arthoniales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200836 OLICH2424-16 MK812703
Lecanactis
abietina
Arthoniales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201261 OLICH2768-16 MK811953
Lecania aipospila Lecanorales NO, Finnmark,
2014, J.T.
Klepsland
O-L-200506 OLICH3254-16 MK812577
Lecania aipospila Lecanorales NO, Finnmark,
2003, E.
Timdal
O-L-123172 OLICH3962-17 MK811636
Lecania erysibe Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183713 OLICH3956-17 MK812191
Lecanora
atrosulphurea
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195558 OLICH1516-14 KY266931
Lecanora intricata Lecanorales NO, Finnmark,
2014, U. Arup
O-L-195841 OLICH1686-14 KY266891
Lecanora intricata Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195876 OLICH1721-14 KY266975
Lecanora
leptacina
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195896 OLICH1741-14 KY266950
Lecidea auriculata Lecideales NO, Finnmark,
2014, E.
Timdal
O-L-195916 OLICH1761-14 KY266956 no lichen
substances
54 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Lecidea
leucothallina
Lecideales NO, Oppland,
2013, T.
Tønsberg
BG-
L-96170
OLICH1065-14 MG968256 pannarin (thallus)
Lecidea
leucothallina
Lecideales NO, Nordland,
2010, E.
Timdal
O-L-163621 OLICH179-11 MG968259 pannarin
(apothecium), no
lichen substances
(thallus)
Lecidea
leucothallina
Lecideales NO, Nordland,
2010, E.
Timdal
O-L-163673 OLICH180-11 MG968258 pannarin (thallus)
Lecidea
leucothallina
Lecideales NO, Møre og
Romsdal,
2010, R.
Haugan
O-L-165815 OLICH181-11 MG968260 no lichen
substances
(thallus)
Lecidea
toensbergii
Lecideales NO, Oppland,
2013, T.
Tønsberg
BG-
L-95936
OLICH1010-13 MG968261 pannarin (thallus)
Lecidea
toensbergii
Lecideales NO, Møre og
Romsdal,
2010, R.
Haugan
O-L-165806 OLICH182-11 MG968257 pannarin (thallus)
Leprocaulon
quisquiliare
Leprocaulales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200884 OLICH2566-16 MK812636
Leprocaulon
quisquiliare
Leprocaulales NO, Akershus,
2015, S. Rui &
E. Timdal
O-L-200816 OLICH2587-16 MK811899
Leprocaulon
quisquiliare
Leprocaulales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208334 OLICH4265-17 MK812206
Leproplaca
cirrochroa
Teloschistales NO, Oppland,
2011, J.T.
Klepsland
O-L-183714 OLICH4395-17 MK812049
Leproplaca
cirrochroa
Teloschistales NO, Telemark,
2009, R.
Haugan & S.
Reiso
O-L-160423 OLICH4427-17 MK812423
OLICH: A reference library of DNA barcodes for Nordic lichens 55

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Leproplaca
cirrochroa
Teloschistales NO, Nord-
Trøndelag,
1998, R.
Haugan, R.
Hollas
O-L-44774 OLICH4428-17 MK812709
Leproplaca
obliterans
Teloschistales NO, Akershus,
2013, J.T.
Klepsland
O-L-197812 OLICH4426-17 MK812503
Leptogium
cochleatum
Peltigerales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199489 OLICH3149-16 MK812473
Leptogium
cochleatum
Peltigerales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199545 OLICH3150-16 MK811946
Leptogium
saturninum
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195592 OLICH1533-14 MK812662
Leptogium
saturninum
Peltigerales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199516 OLICH3250-16 MK812179
Leptogium
saturninum
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196046 OLICH3800-17 MK811641
Letharia vulpina Lecanorales NO, Hedmark,
2012, R.
Haugan
O-L-184032 OLICH1405-14 MK812266 unknown acid,
vulpinic acid
Letharia vulpina Lecanorales NO, Hedmark,
2012, R.
Haugan
O-L-184029 OLICH2179-15 MK812109
Letharia vulpina Lecanorales NO, Hedmark,
2011, B.
Skinderhaug
O-L-169053 OLICH2180-15 MK811718
56 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Letharia vulpina Lecanorales NO, Hedmark,
2010, R.
Haugan
O-L-165378 OLICH2181-15 MK811860
Letharia vulpina Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-198890 OLICH2395-16 MK811758
Lichenomphalia
alpina
Agaricales NO, Finnmark,
2014, H.
Holien
O-L-195732 OLICH1604-14 KY266895
Lichenomphalia
velutina
Agaricales NO, Oslo,
2014, E.
Timdal
O-L-200004 OLICH1978-15 MK811822
Lobaria
amplissima
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196021 OLICH1866-14 MK812432
Lobaria
amplissima
Peltigerales NO, Sør-
Trøndelag,
2010, J.T.
Klepsland
O-L-174993 OLICH2176-15 MK812486
Lobaria
amplissima
Peltigerales NO, Sør-
Trøndelag,
2013, J.T.
Klepsland
O-L-198062 OLICH2326-15 MK811832
Lobaria
amplissima
Peltigerales NO, Buskerud,
2011, T.H.
Hofton
O-L-194379 OLICH2327-15 MK811802
Lobaria
amplissima
Peltigerales NO, Sogn og
Fjordane,
2010, T.H.
Hofton
O-L-187118 OLICH2328-15 MK812689
Lobaria
amplissima
Peltigerales NO, Sør-
Trøndelag,
2011, J.T.
Klepsland
O-L-176912 OLICH2329-15 MK812498
OLICH: A reference library of DNA barcodes for Nordic lichens 57

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Lobaria
amplissima
Peltigerales NO, Nord-
Trøndelag,
2012, J.T.
Klepsland
O-L-186329 OLICH2330-15 MK811725
Lobaria
amplissima
Peltigerales NO, Nord-
Trøndelag,
2013, J.T.
Klepsland
O-L-198158 OLICH2387-16 MK812569
Lobaria
amplissima
Peltigerales NO, Nordland,
2003, A.
Granmo & L.
Molster
TROM_L
_46394
OLICH3586-17 MK811901
Lobaria linita Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195728 OLICH1600-14 KY266856
Lobaria linita Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195892 OLICH1737-14 KY266932
Lobaria
pulmonaria
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196024 OLICH1869-14 MK811957
Lobaria
pulmonaria
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196356 OLICH1915-15 MK811682
Lobaria
pulmonaria
Peltigerales NO, Østfold,
2013, O.
Wergeland
Krog
O-L-193007 OLICH2335-15 MK812447
Lobaria
pulmonaria
Peltigerales NO, Østfold,
2012, O.
Wergeland
Krog et al.
O-L-193009 OLICH2336-15 MK812330
Lobaria
pulmonaria
Peltigerales NO, Østfold,
2013, O.
Wergeland
Krog
O-L-193011 OLICH2337-15 MK812533
58 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Lobaria
pulmonaria
Peltigerales NO, Hedmark,
2013, R.
Haugan
O-L-190366 OLICH2338-15 MK812202
Lobaria
pulmonaria
Peltigerales NO, Buskerud,
2011, T.H.
Hofton & S.
Reiso
O-L-194285 OLICH2339-15 MK812005
Lobaria
pulmonaria
Peltigerales NO, Buskerud,
2012, T.H.
Hofton
O-L-194456 OLICH2340-15 MK812631
Lobaria
pulmonaria
Peltigerales NO, Vest-
Agder, 2011,
J.T. Klepsland
O-L-177093 OLICH2346-15 MK812031
Lobaria
pulmonaria
Peltigerales NO, Hedmark,
2012, R.
Haugan
O-L-183966 OLICH2347-15 MK812039
Lobaria
pulmonaria
Peltigerales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200863 OLICH2515-16 MK811954
Lobaria
pulmonaria
Peltigerales NO, Troms,
2005, A.
Granmo
TROM_L
_50934
OLICH3536-17 MK811932
Lobaria
scrobiculata
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196350 OLICH1909-15 MK812570
Lobaria
scrobiculata
Peltigerales NO, Østfold,
2013, B.P.
Løfall & O.
Wergeland
Krog
O-L-187365 OLICH2341-15 MK812073
Lobaria
scrobiculata
Peltigerales NO, Hedmark,
2013, R.
Haugan
O-L-190098 OLICH2342-15 MK812310
OLICH: A reference library of DNA barcodes for Nordic lichens 59

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Lobaria
scrobiculata
Peltigerales NO, Buskerud,
2012, T.H.
Hofton
O-L-194516 OLICH2343-15 MK812214
Lobaria
scrobiculata
Peltigerales NO, Rogaland,
2012, J.T.
Klepsland
O-L-186129 OLICH2344-15 MK811651
Lobaria
scrobiculata
Peltigerales NO, Nord-
Trøndelag,
2010, H. Bratli
O-L-168086 OLICH2345-15 MK812309
Lobaria
scrobiculata
Peltigerales NO, Troms,
2005, A.
Granmo
TROM_L
_50935
OLICH3537-17 MK812074
Lobaria
scrobiculata
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196051 OLICH3810-17 MK811855
Lobaria virens Peltigerales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201271 OLICH2774-16 MK811660
Lobaria virens Peltigerales NO, Nordland,
2014, J.T.
Klepsland
O-L-200346 OLICH3249-16 MK811736
Lobaria virens Peltigerales NO, Vestfold,
2003, T.H.
Hofton et al.
O-L-128546 OLICH4103-17 MK812046
Lobothallia
alphoplaca
Pertusariales NO, Finnmark,
2014, J.T.
Klepsland
O-L-200411 OLICH2658-16 MK812484
Lobothallia
melanaspis
Pertusariales SE, Torne
Lappmark,
2014, E.
Timdal
O-L-195680 OLICH2007-15 MK811796
Lobothallia
melanaspis
Pertusariales NO, Hedmark,
2014, R.
Haugan
O-L-198879 OLICH2390-16 MK811944
60 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Loxospora elatina Sarrameanales NO, Nord-
Trøndelag,
2014, J.T.
Klepsland
O-L-200270 OLICH2657-16 MK812411
Massalongia
carnosa
Peltigerales NO, Aust-
Agder, 2013,
J.T. Klepsland
O-L-198320 OLICH2371-15 MK812187
Massalongia
carnosa
Peltigerales NO, Buskerud,
2011, E.
Timdal
O-L-175425 OLICH3285-16 MK811716
Megalaria grossa Lecanorales NO, Akershus,
2010, J.T.
Klepsland
O-L-174947 OLICH1036-13 MK811839
Megalaria grossa Lecanorales NO, Sør-
Trøndelag,
2005, E.
Timdal
O-L-149362 OLICH1039-13 MK812151
Megalaria grossa Lecanorales NO, Nordland,
2014, J.T.
Klepsland
O-L-200308 OLICH2656-16 MK812492
Megalaria
pulverea
Lecanorales NO, Sogn og
Fjordane,
2009, J.T.
Klepsland
O-L-165033 OLICH1033-13 MK812535
Megalaria
pulverea
Lecanorales NO, Sør-
Trøndelag,
2009, R.
Haugan
O-L-167026 OLICH1034-13 MK812297
Megalaria
pulverea
Lecanorales NO, Nord-
Trøndelag,
2008, J.T.
Klepsland
O-L-158214 OLICH1035-13 MK812205
Megalaria
pulverea
Lecanorales NO,
Hordaland,
2011, J.T.
Klepsland
O-L-176983 OLICH1317-14 MK812565
OLICH: A reference library of DNA barcodes for Nordic lichens 61

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Megalaria
pulverea
Lecanorales NO, Nordland,
2013, J.T.
Klepsland
O-L-197986 OLICH2104-15 MK812321
Melanelia agnata Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196376 OLICH1935-15 MK812394
Melanelia
hepatizon
Lecanorales NO,
Hordaland,
2014, E.
Timdal
O-L-195807 OLICH1679-14 MK812512
Melanelia
hepatizon
Lecanorales NO, Finnmark,
2014, M.
Westberg
O-L-195864 OLICH1709-14 KY266879
Melanelia
hepatizon
Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201254 OLICH2762-16 MK812070
Melanelia
hepatizon
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184723 OLICH939-13 MK812188
Melanelia stygia Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184736 OLICH1418-14 MK812608
Melanelia stygia Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196377 OLICH1936-15 MK812312
Melanelixia
fuliginosa
Lecanorales NO, Vestfold,
2015, E.
Timdal
O-L-198943 OLICH2499-16 MK812485
Melanelixia
fuliginosa
Lecanorales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200866 OLICH2518-16 MK812619
62 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Melanelixia
fuliginosa
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196050 OLICH3818-17 MK812455
Melanelixia
glabratula
Lecanorales NO,
Hordaland,
2011, J.T.
Klepsland
O-L-176965 OLICH4122-17 MK812674
Melanelixia
subargentifera
Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-200810 OLICH2475-16 MK812517
Melanelixia
subargentifera
Lecanorales NO, Hedmark,
2015, E.
Timdal
O-L-200854 OLICH2505-16 MK812491
Melanelixia
subargentifera
Lecanorales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200867 OLICH2519-16 MK811964
Melanelixia
subargentifera
Lecanorales NO, Hedmark,
2014, R.
Haugan
O-L-207985 OLICH3820-17 MK812324
Melanelixia
subaurifera
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195997 OLICH1842-14 MK811631
Melanelixia
subaurifera
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-199993 OLICH1967-15 MK811942
Melanelixia
subaurifera
Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-198930 OLICH2483-16 MK812688
Melanohalea
exasperata
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196008 OLICH1853-14 MK812395
OLICH: A reference library of DNA barcodes for Nordic lichens 63

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Melanohalea
exasperata
Lecanorales NO, Telemark,
2015, E.
Timdal
O-L-208004 OLICH3878-17 MK812307
Melanohalea
exasperata
Lecanorales NO, Sogn og
Fjordane,
2004, A. Breili
O-L-151072 OLICH4123-17 MK812184
Melanohalea
infumata
Lecanorales NO, Troms,
2003, E.
Timdal
O-L-123135 OLICH4126-17 MK812651
Melanohalea
olivacea
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195933 OLICH1778-14 KY266962
Melanohalea
olivacea
Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195826 OLICH2198-15 MK812054
Melanohalea
olivacea
Lecanorales NO, Telemark,
2015, E.
Timdal
O-L-208005 OLICH3879-17 MK812340
Melanohalea
septentrionalis
Lecanorales NO, Akershus,
2017, S. Rui &
E. Timdal
O-L-206865 OLICH4110-17 MK812433
Menegazzia
subsimilis
Lecanorales NO, Rogaland,
2007, E.
Timdal
O-L-149092 OLICH864-13 MK812421
Menegazzia
subsimilis
Lecanorales NO, Rogaland,
2012, J.T.
Klepsland
O-L-186134 OLICH944-13 MK812020
Menegazzia
terebrata
Lecanorales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200861 OLICH2513-16 MK812093
Menegazzia
terebrata
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200924 OLICH2532-16 MK812635
64 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Menegazzia
terebrata
Lecanorales NO,
Hordaland,
2013, T.
Tønsberg
BG-
L-95692
OLICH858-13 MK812654
Menegazzia
terebrata
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-176828 OLICH861-13 MK812025
Menegazzia
terebrata
Lecanorales NO, Rogaland,
2012, J.T.
Klepsland
O-L-186132 OLICH949-13 MK812145
Micarea
botryoides
Lecanorales NO, Rogaland,
2013, J.I.
Johnsen
BG-
L-95780
OLICH1006-13 MK812289
Micarea denigrata Lecanorales NO, Oppland,
2011, R.
Haugan & A.M.
Haugan
O-L-174438 OLICH1013-13 MK811696
Micarea misella Lecanorales NO, Nord-
Trøndelag,
2013, H.
Holien
TRH-
L-15146
OLICH990-13 MK812399
Micarea paratropa Lecanorales NO, Sør-
Trøndelag,
2010, R.
Haugan
O-L-165711 OLICH101-11 MK812318 no lichen
substances
Montanelia
disjuncta
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195590 OLICH1531-14 MK811852
Montanelia
disjuncta
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196357 OLICH1916-15 MK811711
Montanelia
disjuncta
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-198675 OLICH2404-16 KY266910
Montanelia
panniformis
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195904 OLICH1749-14 KY266847
OLICH: A reference library of DNA barcodes for Nordic lichens 65

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Montanelia
panniformis
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200016 OLICH1990-15 MK812407
Montanelia
panniformis
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200878 OLICH2535-16 MK812634
Montanelia
panniformis
Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201338 OLICH2722-16 MK812263
Montanelia
sorediata
Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-195791 OLICH1663-14 MK811963
Montanelia
sorediata
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195658 OLICH1998-15 MK811965
Montanelia
sorediata
Lecanorales NO, Buskerud,
2016, M.
Stapnes Dahl
et al.
O-L-204941 OLICH4115-17 MK811977
Montanelia tominii Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201344 OLICH2728-16 MK812373
Mycobilimbia
berengeriana
Lecideales NO, Aust-
Agder, 2011,
J.T. Klepsland
O-L-177070 OLICH1113-14 MK812040
Mycobilimbia
berengeriana
Lecideales NO, Oppland,
2011, J.T.
Klepsland
O-L-176838 OLICH1114-14 MK812343
Mycobilimbia
berengeriana
Lecideales NO, Finnmark,
2011, E.
Timdal
O-L-170554 OLICH1117-14 MK811751 no lichen
substances
Mycobilimbia
berengeriana
Lecideales NO, Aust-
Agder, 2009,
J.T. Klepsland
O-L-167344 OLICH1170-14 MK811880
Mycobilimbia
berengeriana
Lecideales NO, Finnmark,
2014, E. Llop
Vallverdu
O-L-195696 OLICH1568-14 KY266871
66 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Mycobilimbia
berengeriana
Lecideales NO, Sør-
Trøndelag,
2004, R.
Haugan
O-L-142068 OLICH629-13 MK811889
Mycobilimbia
carneoalbida
Lecideales NO, Telemark,
2010, J.T.
Klepsland
O-L-175294 OLICH1109-14 MK812353
Mycobilimbia
carneoalbida
Lecideales NO, Aust-
Agder, 2005,
J.T. Klepsland
O-L-151121 OLICH1120-14 MK811646
Mycobilimbia
carneoalbida
Lecideales NO, Troms,
2009, J.T.
Klepsland
O-L-164904 OLICH1121-14 MK812149
Mycobilimbia
carneoalbida
Lecideales NO, Oppland,
2011, R.
Haugan
O-L-174298 OLICH1122-14 MK812290
Mycobilimbia
carneoalbida
Lecideales NO, Hedmark,
2011, R.
Haugan
O-L-174282 OLICH1123-14 MK812691
Mycobilimbia
carneoalbida
Lecideales NO, Telemark,
2007, E.
Timdal
O-L-149057 OLICH1124-14 MK812639
Mycobilimbia
epixanthoides
Lecideales NO, Hedmark,
2006, J.T.
Klepsland
O-L-164598 OLICH1105-14 MK812105
Mycobilimbia
epixanthoides
Lecideales NO, Aust-
Agder, 2006,
J.T. Klepsland
O-L-151124 OLICH1106-14 MK812388
Mycobilimbia
epixanthoides
Lecideales NO, Troms,
2007, J.T.
Klepsland
O-L-154903 OLICH1107-14 MK812645
Mycobilimbia
epixanthoides
Lecideales NO, Telemark,
2007, E.
Timdal
O-L-149055 OLICH1108-14 MK812402 no lichen
substances
Mycobilimbia
epixanthoides
Lecideales NO, Akershus,
2000, R.
Haugan
O-L-160303 OLICH1112-14 MK811772
OLICH: A reference library of DNA barcodes for Nordic lichens 67

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Mycobilimbia
pilularis
Lecideales NO, Vest-
Agder, 2009,
J.T. Klepsland
O-L-164694 OLICH1110-14 MK812275
Mycobilimbia
pilularis
Lecideales NO, Møre og
Romsdal,
2012, B.
Nordén & J.B.
Jordal
O-L-180158 OLICH1111-14 MK812666
Mycobilimbia
pilularis
Lecideales NO,
Hordaland,
2008, J.B.
Jordal
O-L-176020 OLICH1125-14 MK812525
Mycobilimbia
pilularis
Lecideales NO, Sogn og
Fjordane,
2009, A. Breili
O-L-159489 OLICH1126-14 MK812665
Mycobilimbia
tetramera
Lecideales NO, Finnmark,
2011, E.
Timdal
O-L-170621 OLICH1119-14 MK812422 no lichen
substances
Mycobilimbia
tetramera
Lecideales NO, Nordland,
2011, R.
Haugan
O-L-174132 OLICH1127-14 MK812216 no lichen
substances
Mycobilimbia
tetramera
Lecideales NO, Aust-
Agder, 2006,
J.T. Klepsland
O-L-183493 OLICH1128-14 MK811935
Mycobilimbia
tetramera
Lecideales NO, Finnmark,
2014, E. Llop
Vallverdu
O-L-195846 OLICH1691-14 MK812124
Mycobilimbia
tetramera
Lecideales NO, Akershus,
2014, J.T.
Klepsland
O-L-200258 OLICH3147-16 MK812192
Mycobilimbia
tetramera
Lecideales NO, Akershus,
2009, J.T.
Klepsland
O-L-164634 OLICH631-13 MK812158
Mycobilimbia
tetramera
Lecideales NO, Hedmark,
2008, R.
Haugan
O-L-160476 OLICH633-13 MK812208
68 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Mycoblastus
sanguinarius
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195768 OLICH1640-14 MK812482
Mycoblastus
sanguinarius
Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201263 OLICH2769-16 MK811859
Mycoblastus
sanguinarius
Lecanorales NO, Hedmark,
2010, E.
Timdal
O-L-163524 OLICH391-13 MK811665
Mycoblastus
sanguinarius
Lecanorales NO, Telemark,
2005, E.
Timdal
O-L-138597 OLICH410-13 MK812197
Mycoblastus
sanguinarius
Lecanorales NO, Oppland,
2006, A. Breili
O-L-150999 OLICH617-13 MK812135
Myriolecis
straminea
Lecanorales NO, Finnmark,
2014, M.
Westberg
O-L-195870 OLICH1715-14 KY266929
Myriospora
smaragdula
Acarosporales NO, Finnmark,
2014, E.
Timdal
O-L-195609 OLICH1543-14 MK811918
Nephroma
arcticum
Peltigerales NO, Troms,
2014, E.
Timdal
O-L-195654 OLICH1566-14 MK811882
Nephroma
arcticum
Peltigerales NO, Finnmark,
2014, S. Rui
O-L-195724 OLICH1596-14 KY266888
Nephroma
arcticum
Peltigerales NO, Finnmark,
2014, A.
Millanes
O-L-195873 OLICH1718-14 KY266963
Nephroma
arcticum
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196368 OLICH1927-15 MK812091
Nephroma
arcticum
Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-196272 OLICH1945-15 MK812336
OLICH: A reference library of DNA barcodes for Nordic lichens 69

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Nephroma
arcticum
Peltigerales NO, Nord-
Trøndelag,
2014, Z.
Fackovcova et
al.
SAV-36 OLICH3073-16 MK812204
Nephroma
arcticum
Peltigerales NO, Nord-
Trøndelag,
2014, Z.
Fackovcova et
al.
SAV-37 OLICH3074-16 MK812592
Nephroma
arcticum
Peltigerales NO, Finnmark,
2008, A.
Granmo & G.
Mathiassen
TROM_L
_46162
OLICH3570-17 MK811792
Nephroma
arcticum
Peltigerales NO, Hedmark,
2014, R.
Haugan
O-L-207992 OLICH3828-17 MK812028
Nephroma bellum Peltigerales NO, Nord-
Trøndelag,
2010, H. Bratli
O-L-168043 OLICH2027-15 MK811672 triterpenoids
Nephroma bellum Peltigerales NO, Sogn og
Fjordane,
2012, O. Roen
O-L-181644 OLICH2028-15 MK812308 triterpenoids
Nephroma bellum Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196348 OLICH2373-15 MK812483 triterpenoids
Nephroma
expallidum
Peltigerales NO, Finnmark,
2014, M.
Westberg
O-L-195701 OLICH1573-14 KY266905
Nephroma
expallidum
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195891 OLICH1736-14 KY266844
Nephroma
expallidum
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196346 OLICH1906-15 MK812408
70 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Nephroma
expallidum
Peltigerales NO, Finnmark,
2011, E.
Timdal
O-L-170548 OLICH2365-15 MK811945
Nephroma
expallidum
Peltigerales NO, Svalbard,
2015, S. Rui
O-L-203292 OLICH2712-16 MK812041
Nephroma
helveticum
Peltigerales NO, Buskerud,
2013, T.H.
Hofton
O-L-196491 OLICH2036-15 KT800007 triterpenoids
Nephroma
helveticum
Peltigerales NO, Oppland,
2011, J.T.
Klepsland
O-L-181601 OLICH2037-15 KT800006 triterpenoides (T4,
T1)
Nephroma
laevigatum
Peltigerales NO, Nord-
Trøndelag,
2010, H. Bratli
O-L-168088 OLICH2025-15 MK812687
Nephroma
laevigatum
Peltigerales NO, Sogn og
Fjordane,
2010, J.T.
Klepsland
O-L-175024 OLICH2038-15 MK812108 pigments,
triterpenoid
Nephroma
laevigatum
Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201288 OLICH2676-16 MK812261
Nephroma
laevigatum
Peltigerales NO, Troms,
2010, K.
Wannebo
Nilsen
TROM_L
_48759
OLICH3531-17 MK811868
Nephroma
laevigatum
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196055 OLICH3812-17 MK811969
Nephroma
laevigatum
Peltigerales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-204997 OLICH4002-17 MK812002
Nephroma parile Peltigerales NO, Finnmark,
2014, T.
Tønsberg
O-L-195875 OLICH1720-14 KY266927 triterpenoids
OLICH: A reference library of DNA barcodes for Nordic lichens 71

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Nephroma
resupinatum
Peltigerales NO, Hedmark,
2013, R.
Haugan
O-L-190528 OLICH2020-15 MK811650
Nephroma
resupinatum
Peltigerales NO, Sogn og
Fjordane,
2010, E.
Timdal
O-L-163401 OLICH2022-15 MK812367
Nephroma
resupinatum
Peltigerales NO, Troms,
2011, J.T.
Klepsland
O-L-177168 OLICH2023-15 MK812612
Nephroma
resupinatum
Peltigerales NO, Nord-
Trøndelag,
2011, J.T.
Klepsland
O-L-183598 OLICH2024-15 MK811912 no lichen
substances
Nephroma
resupinatum
Peltigerales NO, Troms,
2005, A.
Granmo
TROM_L
_50937
OLICH3538-17 MK812301
Nephroma
resupinatum
Peltigerales NO, Finnmark,
2008, A.
Granmo & G.
Mathiassen
TROM_L
_48235
OLICH3588-17 MK812553
Nephroma
tangeriense
Peltigerales NO,
Hordaland,
2012, J.T.
Klepsland
O-L-186150 OLICH2018-15 MK811986 pigments,
triterpenoides (T3,
T4, T6)
Nephroma
tangeriense
Peltigerales NO, Rogaland,
2010, J.T.
Klepsland
O-L-181602 OLICH2019-15 MK812084 pigments,
triterpenoides (T2,
T3, T6)
Nephroma
tangeriense
Peltigerales NO, Vest-
Agder, 2011,
J.T. Klepsland
O-L-183599 OLICH2039-15 MK812530 pigments,
triterpenoids
Nephroma
tangeriense
Peltigerales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197824 OLICH2105-15 MK812579 pigments
Nephroma
tangeriense
Peltigerales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197826 OLICH2106-15 MK812593
72 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Nevesia
sampaiana
Peltigerales NO, Sogn og
Fjordane,
2009, H.
Fjeldstad
O-L-203003 OLICH3277-16 MK812089
Normandina
pulchella
Verrucariales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-204993 OLICH4003-17 MK812131
Ochrolechia
alaskana
Pertusariales NO, Finnmark,
2008, A.
Granmo & G.
Mathiassen
TROM_L
_60552
OLICH3582-17 MK811874
Ochrolechia
alboflavescens
Pertusariales NO, Nord-
Trøndelag,
2015, F.
Jonsson
O-L-201276 OLICH3230-16 MK812244
Ochrolechia
androgyna
Pertusariales NO, Nord-
Trøndelag,
2015, H.
Holien
TRH-
L-16947
OLICH3684-17 MK812264
Ochrolechia
androgyna
Pertusariales NO, Oppland,
2015, R.
Haugan
O-L-203919 OLICH3885-17 MK811897 atranorin,
gyrophoric acid,
lecanoric acid,
unknown acid
Ochrolechia
androgyna
Pertusariales NO, Oppland,
2015, E.
Timdal
O-L-201397 OLICH4178-17 MK812682 gyrophoric acid
Ochrolechia
frigida
Pertusariales NO, Finnmark,
2014, A.
Launis
O-L-195945 OLICH1790-14 KY266952
Ochrolechia
gowardii
Pertusariales NO, Nord-
Trøndelag,
2014, J.T.
Klepsland
O-L-200283 OLICH3146-16 MK812101
Ochrolechia
gowardii
Pertusariales NO, Nord-
Trøndelag,
2012, J.T.
Klepsland
O-L-200100 OLICH3225-16 MK812175
OLICH: A reference library of DNA barcodes for Nordic lichens 73

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Ochrolechia
mahluensis
Pertusariales NO, Finnmark,
2014, H.
Holien
O-L-195912 OLICH1757-14 KY266861
Ochrolechia
microstictoides
Pertusariales NO, Aust-
Agder, 2014,
J.T. Klepsland
O-L-200753 OLICH3145-16 MK812077
Ochrolechia
parella
Pertusariales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196015 OLICH1860-14 MK811779
Ochrolechia
parella
Pertusariales NO, Nordland,
2014, J.T.
Klepsland
O-L-200300 OLICH3144-16 MK811835
Ochrolechia
subviridis
Pertusariales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200638 OLICH2655-16 MK812495 gyrophoric acid
Ochrolechia
szatalaensis
Pertusariales NO, Troms,
2011, J.T.
Klepsland
O-L-200097 OLICH2653-16 MK811817 murolic acid
(complex),
variolaric acid
Ochrolechia
szatalaensis
Pertusariales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199529 OLICH2654-16 MK812256
Ochrolechia
szatalaensis
Pertusariales NO, Nord-
Trøndelag,
2014, J.T.
Klepsland
O-L-200284 OLICH3143-16 MK811903
Ochrolechia
szatalaensis
Pertusariales NO, Finnmark,
2011, J.T.
Klepsland
O-L-200096 OLICH3227-16 MK811865 murolic acid
(complex),
variolaric acid
Ochrolechia
tartarea
Pertusariales NO, Finnmark,
2014, H.
Holien
O-L-195951 OLICH1796-14 KY266863
Ochrolechia
tartarea
Pertusariales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196041 OLICH3795-17 MK812341
74 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Ochrolechia
upsaliensis
Pertusariales NO, Finnmark,
2014, H.
Holien
O-L-195967 OLICH1812-14 KY266960
Ochrolechia
upsaliensis
Pertusariales NO, Oppland,
2013, J.T.
Klepsland
O-L-198354 OLICH2376-16 MK811642
Opegrapha
vermicellifera
Arthoniales NO,
Hordaland,
2016, J.T.
Klepsland
O-L-206757 OLICH3476-17 MK811737
Ophioparma
lapponica
Umbilicariales NO, Finnmark,
2014, T.B.
Wheeler
O-L-195838 OLICH1683-14 KY266901
Ophioparma
ventosa
Umbilicariales FI, Inarin
Lappi, 2014, E.
Timdal
O-L-195498 OLICH1503-14 MK812137
Ophioparma
ventosa
Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195543 OLICH1512-14 KY266864
Ophioparma
ventosa
Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195574 OLICH1525-14 MK812332
Ophioparma
ventosa
Umbilicariales NO, Oppland,
2015, E.
Timdal
O-L-201417 OLICH3307-16 MK811904
Ophioparma
ventosa
Umbilicariales NO, Troms,
2000, K. Lerfall
TROM_L
_565529
OLICH3550-17 MK811740
Ophioparma
ventosa
Umbilicariales NO, Hedmark,
2013, E.
Timdal
O-L-184520 OLICH701-13 MK811771
Orphniospora
moriopsis
Umbilicariales NO, Finnmark,
2014, H.
Holien
O-L-195716 OLICH1588-14 KY266869
Pannaria
conoplea
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196369 OLICH1928-15 MK811685
OLICH: A reference library of DNA barcodes for Nordic lichens 75

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Pannaria
conoplea
Peltigerales NO, Sogn og
Fjordane,
2012, T.H.
Hofton & T.
Høitomt
O-L-194474 OLICH2379-16 MK812100
Pannaria
conoplea
Peltigerales NO, Hedmark,
2011, R.
Haugan
O-L-173219 OLICH2380-16 MK812203
Pannaria
conoplea
Peltigerales NO, Hedmark,
2011, R.
Haugan
O-L-173231 OLICH2382-16 MK812521
Pannaria
conoplea
Peltigerales NO, Hedmark,
2011, R.
Haugan
O-L-173220 OLICH2383-16 MK811788
Pannaria
conoplea
Peltigerales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200927 OLICH2553-16 MK812604
Pannaria
conoplea
Peltigerales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201274 OLICH2776-16 MK812480
Pannaria hookeri Peltigerales NO, Nordland,
2013, J.T.
Klepsland
O-L-198018 OLICH3882-17 MK812291
Pannaria
rubiginosa
Peltigerales NO, Nordland,
2004, A.
Elvebakk
TROM_L
_1340509
OLICH3563-17 MK812061
Pannaria
rubiginosa
Peltigerales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201270 OLICH3846-17 MK812317
Parmelia ernstiae Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201279 OLICH2777-16 MK567160 atranorin,
lichesterinic acid,
lobaric acid,
protolichesterinic
acid, salazinic acid
76 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Parmelia ernstiae Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201277 OLICH3848-17 MK567162 atranorin,
lichesterinic acid,
lobaric acid,
protolichesterinic
acid, salazinic acid
Parmelia fraudans Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196374 OLICH1933-15 MK812511
Parmelia
omphalodes
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195947 OLICH1792-14 KY266894
Parmelia saxatilis Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195994 OLICH1839-14 MK567163 atranorin, lobaric
acid, salazinic acid
Parmelia saxatilis Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184724 OLICH940-13 MK567164 atranorin, lobaric
acid, salazinic acid
Parmelia serrana Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195995 OLICH1840-14 MK567161 atranorin,
lichesterinic acid,
lobaric acid,
protolichesterinic
acid, salazinic acid
Parmelia serrana Lecanorales NO, Vestfold,
2016, S. Rui &
E. Timdal
O-L-205007 OLICH4004-17 MK567159 atranorin,
lichesterinic acid,
lobaric acid,
protolichesterinic
acid, salazinic acid
Parmelia sulcata Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-195789 OLICH1661-14 MK811938
Parmelia sulcata Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196000 OLICH1845-14 MK812088
OLICH: A reference library of DNA barcodes for Nordic lichens 77

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Parmelia sulcata Lecanorales NO, Oslo,
2014, M.
Bendiksby et
al.
O-L-196057 OLICH1873-14 MK812065
Parmelia sulcata Lecanorales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-196331 OLICH2261-15 MK812499
Parmelia sulcata Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184694 OLICH878-13 MK812475
Parmeliella
parvula
Peltigerales NO, Nord-
Trøndelag,
2013, J.T.
Klepsland
O-L-197955 OLICH2107-15 MK812304
Parmeliella
triptophylla
Peltigerales NO, Telemark,
2015, E.
Timdal
O-L-207999 OLICH3873-17 MK812457
Parmelina
pastillifera
Lecanorales NO, Rogaland,
2013, J.T.
Klepsland
O-L-198298 OLICH2377-16 MK812633
Parmelina
pastillifera
Lecanorales NO,
Hordaland,
2006, H.
Fjeldstad & G.
Gaarder
O-L-202999 OLICH3231-16 MK812400
Parmelina
pastillifera
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-205003 OLICH4005-17 MK812564
Parmelina
pastillifera
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-204989 OLICH4006-17 MK812554
Parmelina
pastillifera
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-205000 OLICH4007-17 MK812365
Parmelina tiliacea Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-199984 OLICH1958-15 MK812323
78 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Parmelina tiliacea Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196287 OLICH2217-15 MK812693
Parmelina tiliacea Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-198924 OLICH2477-16 MK811628
Parmelina tiliacea Lecanorales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200873 OLICH2525-16 MK812259
Parmelina tiliacea Lecanorales NO, Akershus,
2015, S. Rui &
E. Timdal
O-L-200818 OLICH2589-16 MK812562
Parmelina tiliacea Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201334 OLICH2718-16 MK811862
Parmelina tiliacea Lecanorales NO, Oslo,
2016, S. Rui &
E. Timdal
O-L-203344 OLICH3314-16 MK812472
Parmelina tiliacea Lecanorales NO, Rogaland,
2015, T.
Tønsberg & E.
Nygaard
BG-
L-98117
OLICH3696-17 MK812477
Parmeliopsis
ambigua
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195902 OLICH1747-14 KY266965
Parmeliopsis
ambigua
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195934 OLICH1779-14 KY266851
Parmeliopsis
ambigua
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195990 OLICH1835-14 KY266850
Parmeliopsis
ambigua
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184695 OLICH879-13 MK811727
OLICH: A reference library of DNA barcodes for Nordic lichens 79

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Parmeliopsis
hyperopta
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195989 OLICH1834-14 KY266853
Parmeliopsis
hyperopta
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184696 OLICH880-13 MK812515
Parmotrema
crinitum
Lecanorales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197835 OLICH2109-15 MK812284
Parmotrema
crinitum
Lecanorales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197840 OLICH2110-15 MK811794
Parmotrema
perlatum
Lecanorales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200600 OLICH2651-16 MK812450
Parmotrema
perlatum
Lecanorales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200643 OLICH4358-17 MK812630
Pectenia
cyanoloma
Peltigerales NO, Nord-
Trøndelag,
2013, H.
Holien
O-L-193006 OLICH1477-14 MK811927
Pectenia
cyanoloma
Peltigerales NO, Sogn og
Fjordane,
2017, O. Olsen
O-L-207808 OLICH4320-17 MK812029
Peltigera
aphthosa
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195726 OLICH1598-14 KY266857
Peltigera
aphthosa
Peltigerales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200017 OLICH1991-15 MK811726
Peltigera
aphthosa
Peltigerales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184749 OLICH2048-15 MK812596
Peltigera
aphthosa
Peltigerales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200534 OLICH2611-16 MK811760
80 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera
aphthosa
Peltigerales NO, Telemark,
2015, E.
Timdal
O-L-208000 OLICH3874-17 MK811699
Peltigera
aphthosa
Peltigerales NO, Buskerud,
2016, M.
Bendiksby & E.
Timdal
O-L-208028 OLICH4256-17 MK811816
Peltigera
britannica
Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201329 OLICH2689-16 MK812626
Peltigera canina Peltigerales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200554 OLICH2610-16 MK811916
Peltigera canina Peltigerales NO, Østfold,
2012, K.A. Lye
O-L-206918 OLICH4134-17 MK812107
Peltigera canina Peltigerales NO, Østfold,
2012, K.A. Lye
O-L-206873 OLICH4135-17 MK811752
Peltigera collina Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196371 OLICH1930-15 MK812439
Peltigera collina Peltigerales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200865 OLICH2517-16 MK811975
Peltigera collina Peltigerales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200926 OLICH2540-16 MK812019
Peltigera collina Peltigerales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201268 OLICH3845-17 MK811743
Peltigera degenii Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-196276 OLICH1950-15 MK812238
OLICH: A reference library of DNA barcodes for Nordic lichens 81

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera degenii Peltigerales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-196335 OLICH2273-15 MK812333
Peltigera degenii Peltigerales NO, Buskerud,
2016, S. Rui &
E. Timdal
O-L-208031 OLICH4259-17 MK812618
Peltigera didactyla Peltigerales NO, Telemark,
2015, E.
Timdal
O-L-208003 OLICH3877-17 MK811844
Peltigera
elisabethae
Peltigerales NO, Buskerud,
2014, E.
Timdal
O-L-196071 OLICH1887-14 MK811869
Peltigera
extenuata
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196343 OLICH1903-15 MK811873
Peltigera
extenuata
Peltigerales NO, Oslo,
2014, E.
Timdal
O-L-200007 OLICH1981-15 MK811908
Peltigera
extenuata
Peltigerales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200018 OLICH1992-15 MK811654
Peltigera
horizontalis
Peltigerales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184750 OLICH2083-15 MK811925
Peltigera
horizontalis
Peltigerales NO, Hedmark,
2014, R.
Haugan
O-L-198894 OLICH2391-16 MK812678
Peltigera
horizontalis
Peltigerales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200862 OLICH2514-16 MK811673
Peltigera
horizontalis
Peltigerales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208021 OLICH4193-17 MK812656
82 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera
hymenina
Peltigerales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200019 OLICH1993-15 MK812274
Peltigera
hymenina
Peltigerales NO, Oslo,
2014, E.
Timdal
O-L-198933 OLICH2485-16 MK811966
Peltigera
hymenina
Peltigerales NO, Vestfold,
2015, E.
Timdal
O-L-198944 OLICH2495-16 MK812144
Peltigera
hymenina
Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201328 OLICH2688-16 MK811735
Peltigera
lepidophora
Peltigerales NO, Telemark,
2015, E.
Timdal & S.
Rui
O-L-200874 OLICH3835-17 MK812313
Peltigera
lepidophora
Peltigerales NO, Sør-
Trøndelag,
2013, E.
Timdal
O-L-184530 OLICH732-13 MK812625
Peltigera
leucophlebia
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196342 OLICH1902-15 MK812292
Peltigera
leucophlebia
Peltigerales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200020 OLICH1994-15 MK811981
Peltigera
leucophlebia
Peltigerales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200929 OLICH2555-16 MK811680
Peltigera
leucophlebia
Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201315 OLICH2684-16 MK812123
Peltigera
leucophlebia
Peltigerales NO, Svalbard,
2015, S. Rui
O-L-203280 OLICH2699-16 MK811857
OLICH: A reference library of DNA barcodes for Nordic lichens 83

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera
leucophlebia
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196030 OLICH3807-17 MK812241
Peltigera
leucophlebia
Peltigerales NO, Oppland,
2015, E.
Timdal
O-L-201400 OLICH4181-17 MK812102
Peltigera malacea Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195725 OLICH1597-14 KY266940
Peltigera malacea Peltigerales NO, Finnmark,
2014, R.
Haugan
O-L-195848 OLICH1693-14 KY266836
Peltigera malacea Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196362 OLICH1921-15 MK811739
Peltigera malacea Peltigerales NO, Oslo,
2014, E.
Timdal
O-L-200006 OLICH1980-15 MK812014
Peltigera malacea Peltigerales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184751 OLICH2085-15 MK811998
Peltigera malacea Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201331 OLICH2690-16 MK811990
Peltigera
membranacea
Peltigerales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195991 OLICH1836-14 MK811987
Peltigera
membranacea
Peltigerales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200024 OLICH2472-16 MK812374
Peltigera neckeri Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195753 OLICH1625-14 KY266937
84 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera neckeri Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195932 OLICH1777-14 MK811778
Peltigera neckeri Peltigerales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200910 OLICH2582-16 MK811968
Peltigera
neocanina
Peltigerales NO, Buskerud,
2014, R.
Haugan & E.
Timdal
O-L-196062 OLICH1878-14 MK812413
Peltigera
neopolydactyla
Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-196266 OLICH1941-15 MK812243
Peltigera
neopolydactyla
Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-196277 OLICH1946-15 MK812022
Peltigera
neopolydactyla
Peltigerales NO, Oslo,
2014, E.
Timdal
O-L-199994 OLICH1968-15 MK812548
Peltigera
neopolydactyla
Peltigerales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184753 OLICH2084-15 MK811653
Peltigera
neopolydactyla
Peltigerales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200025 OLICH2473-16 MK812286
Peltigera
neopolydactyla
Peltigerales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201332 OLICH2691-16 MK812172
Peltigera
polydactylon
Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-196264 OLICH1939-15 MK811815
Peltigera
polydactylon
Peltigerales NO, Telemark,
2014, E.
Timdal
O-L-195820 OLICH2192-15 MK812112
Peltigera
polydactylon
Peltigerales NO, Buskerud,
2016, S. Rui &
E. Timdal
O-L-206899 OLICH4136-17 MK812583
OLICH: A reference library of DNA barcodes for Nordic lichens 85

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera
praetextata
Peltigerales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184752 OLICH2044-15 MK811692
Peltigera
praetextata
Peltigerales NO, Oslo,
2014, E.
Timdal
O-L-195837 OLICH2209-15 MK811856
Peltigera
praetextata
Peltigerales NO, Akershus,
2009, T.
Starholm
O-L-201769 OLICH3880-17 MK812051
Peltigera
praetextata
Peltigerales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208022 OLICH4194-17 MK812578
Peltigera
retifoveata
Peltigerales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200533 OLICH2605-16 MK811996
Peltigera
rufescens
Peltigerales NO, Oslo,
2014, S. Rui &
E. Timdal
O-L-196072 OLICH1888-14 MK812663
Peltigera
rufescens
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196341 OLICH1901-15 MK811858
Peltigera
rufescens
Peltigerales NO, Akershus,
2014, E.
Timdal
O-L-199989 OLICH1963-15 MK812224
Peltigera
rufescens
Peltigerales NO, Oppland,
2015, E.
Timdal
O-L-201347 OLICH2733-16 MK812696
Peltigera
scabrosa
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195729 OLICH1601-14 KY266948
Peltigera
scabrosa
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195765 OLICH1637-14 KY266970
Peltigera
scabrosa
Peltigerales NO, Telemark,
2015, E.
Timdal
O-L-207997 OLICH3871-17 MK812044
86 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Peltigera
scabrosella
Peltigerales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201255 OLICH2763-16 MK812406
Peltigera
scabrosella
Peltigerales NO, Telemark,
2015, E.
Timdal
O-L-207998 OLICH3872-17 MK812463
Peltula euploca Lichinales NO, Oppland,
2013, J.T.
Klepsland
O-L-197859 OLICH2111-15 MK811928
Pertusaria alpina Pertusariales NO, Nordland,
2014, J.T.
Klepsland
O-L-200312 OLICH3141-16 MK811905
Pertusaria alpina Pertusariales NO, Nord-
Trøndelag,
2013, J.T.
Klepsland
O-L-197933 OLICH4038-17 MK811785
Pertusaria
aspergilla
Pertusariales NO, Sør-
Trøndelag,
2015, H.
Holien
TRH-
L-16775
OLICH3686-17 MK812282
Pertusaria
aspergilla
Pertusariales NO, Nordland,
2015, T.
Tønsberg
BG-
L-99158
OLICH3716-17 MK812384
Pertusaria
aspergilla
Pertusariales NO, Østfold,
2015, R.
Haugan
O-L-204101 OLICH3929-17 MK812496 fumarprotocetraric
acid
Pertusaria
carneopallida
Pertusariales NO, Nordland,
2013, J.T.
Klepsland
O-L-198006 OLICH4039-17 MK812218
Pertusaria
excludens
Pertusariales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-203909 OLICH3927-17 MK811670 norstictic acid
OLICH: A reference library of DNA barcodes for Nordic lichens 87

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Pertusaria
excludens
Pertusariales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-203523 OLICH3935-17 MK812675 norstictic acid
Pertusaria
flavicans
Pertusariales NO, Møre og
Romsdal,
2015, T.
Tønsberg
BG-
L-99071
OLICH3777-17 MK812409
Pertusaria flavida Pertusariales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200860 OLICH2512-16 MK811913
Pertusaria
leioplaca
Pertusariales NO, Troms,
2000, S. Werth
TROM_L
_565683
OLICH3551-17 MK811922
Pertusaria pertusa Pertusariales NO, Vestfold,
2015, E.
Timdal
O-L-198948 OLICH2502-16 MK812037
Phaeophyscia
endococcina
Caliciales NO, Finnmark,
2014, R.
Haugan
O-L-198738 OLICH2400-16 MK812217
Phaeophyscia
kairamoi
Caliciales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196355 OLICH1914-15 MK811724
Phaeophyscia
orbicularis
Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195734 OLICH1606-14 KY266882
Phaeophyscia
orbicularis
Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196004 OLICH1849-14 MK811854
Phaeophyscia
orbicularis
Caliciales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196291 OLICH2221-15 MK811807
Phaeophyscia
orbicularis
Caliciales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-196330 OLICH2259-15 MK812383
88 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Phaeophyscia
orbicularis
Caliciales NO, Hedmark,
2014, R.
Haugan
O-L-207984 OLICH3819-17 MK812363
Phaeophyscia
sciastra
Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195721 OLICH1593-14 KY266837
Phaeophyscia
sciastra
Caliciales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196352 OLICH1911-15 MK812372
Phlyctis agelaea Ostropales NO,
Hordaland,
2015, J.T.
Klepsland
O-L-206746 OLICH3465-17 MK812009
Phlyctis argena Ostropales NO, Nordland,
2015, T.
Tønsberg & A.
Botnen
BG-
L-98987
OLICH3760-17 MK811759
Phlyctis argena Ostropales NO, Nordland,
2015, T.
Tønsberg & A.
Botnen
BG-
L-99194
OLICH3761-17 MK811753
Phlyctis argena Ostropales NO, Buskerud,
2014, T.
Tønsberg
BG-
L-99532
OLICH3786-17 MK811872
Physcia
adscendens
Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196005 OLICH1850-14 MK811630
Physcia
adscendens
Caliciales NO, Vestfold,
2015, E.
Timdal
O-L-198947 OLICH2501-16 MK812201
Physcia aipolia Caliciales NO, Telemark,
2014, E.
Timdal
O-L-195783 OLICH1655-14 MK812601
OLICH: A reference library of DNA barcodes for Nordic lichens 89

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Physcia aipolia Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195992 OLICH1837-14 MK811770
Physcia aipolia Caliciales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200890 OLICH2528-16 MK811989
Physcia aipolia Caliciales NO, Oslo,
2014, J.T.
Klepsland
O-L-200142 OLICH2609-16 MK812087
Physcia aipolia Caliciales NO, Østfold,
2012, K.A. Lye
O-L-206860 OLICH4138-17 MK811719
Physcia alnophila Caliciales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201269 OLICH2773-16 MK812315
Physcia caesia Caliciales NO, Telemark,
2014, E.
Timdal
O-L-195792 OLICH1664-14 MK811883
Physcia caesia Caliciales NO, Oslo,
2014, M.
Bendiksby et
al.
O-L-196078 OLICH1894-14 MK812056
Physcia caesia Caliciales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-196333 OLICH2263-15 MK812325
Physcia caesia Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196035 OLICH3791-17 MK812524
Physcia dimidiata Caliciales NO, Buskerud,
2015, E.
Timdal
O-L-201418 OLICH3299-16 MK812556
Physcia dubia Caliciales NO, Oppland,
2013, J.T.
Klepsland
O-L-197852 OLICH2113-15 MK812017
90 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Physcia dubia Caliciales NO, Oppland,
2015, E.
Timdal
O-L-201373 OLICH2748-16 MK811700
Physcia
magnussonii
Caliciales NO, Sogn og
Fjordane,
2013, J.T.
Klepsland
O-L-197861 OLICH2114-15 MK811875
Physcia phaea Caliciales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196373 OLICH1932-15 MK811892
Physcia stellaris Caliciales NO, Troms,
2014, E.
Timdal
O-L-195769 OLICH1641-14 MK811861
Physcia stellaris Caliciales NO, Østfold,
2012, K.A. Lye
O-L-206903 OLICH4137-17 MK812381
Physcia tenella Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195561 OLICH1519-14 KY266867
Physcia tenella Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196007 OLICH1852-14 MK811920
Physcia tenella Caliciales NO, Oslo,
2014, M.
Bendiksby et
al.
O-L-196079 OLICH1895-14 MK811798
Physcia tenella Caliciales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196294 OLICH2224-15 MK811848
Physcia tenella Caliciales NO, Rogaland,
2013, J.T.
Klepsland
O-L-198307 OLICH2357-15 MK811950
Physcia tenella Caliciales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196034 OLICH3798-17 MK812588
OLICH: A reference library of DNA barcodes for Nordic lichens 91

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Physconia distorta Caliciales NO, Akershus,
2014, E.
Timdal
O-L-199983 OLICH1957-15 MK811745
Physconia distorta Caliciales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196290 OLICH2220-15 MK812071
Physconia
enteroxantha
Caliciales NO, Oslo,
2014, R.
Haugan et al.
O-L-196058 OLICH1874-14 MK811840
Physconia
enteroxantha
Caliciales NO, Akershus,
2014, E.
Timdal
O-L-199986 OLICH1960-15 MK811812
Physconia
enteroxantha
Caliciales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196293 OLICH2223-15 MK811936
Physconia
enteroxantha
Caliciales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-196328 OLICH2258-15 MK812142
Physconia
muscigena
Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195722 OLICH1594-14 KY266849
Physconia
muscigena
Caliciales NO, Finnmark,
2014, E.
Timdal
O-L-195920 OLICH1765-14 KY266925
Physconia
muscigena
Caliciales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196344 OLICH1904-15 MK811781
Pilophorus
dovrensis
Lecanorales NO, Troms,
2003, E.
Timdal
O-L-123083 OLICH211-13 MK812599
Placynthiella
oligotropha
Trapeliales NO, Oppland,
2012, E.
Timdal
O-L-182032 OLICH1176-14 MK811853
Placynthiella
oligotropha
Trapeliales NO, Oppland,
2011, M.
Bendiksby et
al.
O-L-175738 OLICH1177-14 MK812378
92 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Platismatia glauca Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195649 OLICH1561-14 MK812715
Platismatia glauca Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195993 OLICH1838-14 MK812351
Platismatia glauca Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200882 OLICH2564-16 MK812456
Platismatia glauca Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184697 OLICH881-13 MK812210
Platismatia
norvegica
Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201267 OLICH2772-16 MK812616
Platismatia
norvegica
Lecanorales NO, Akershus,
2017, S. Rui &
E. Timdal
O-L-206864 OLICH4109-17 MK812705
Pleopsidium
chlorophanum
Acarosporales NO, Troms,
2014, E.
Timdal
O-L-195643 OLICH1555-14 MK811850
Pleopsidium
chlorophanum
Acarosporales NO, Finnmark,
2014, E.
Timdal
O-L-195940 OLICH1785-14 KY266860
Pleopsidium
chlorophanum
Acarosporales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201252 OLICH2759-16 MK812117
Pleopsidium
chlorophanum
Acarosporales NO, Sør-
Trøndelag,
2017, E.
Timdal
O-L-208244 OLICH4322-17 MK812354
OLICH: A reference library of DNA barcodes for Nordic lichens 93

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Pleopsidium
chlorophanum
Acarosporales NO, Telemark,
2013, E.
Timdal
O-L-184414 OLICH695-13 MK812213
Pleopsidium
flavum
Acarosporales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200545 OLICH2624-16 MK811960
Pleopsidium
flavum
Acarosporales NO, Telemark,
2013, E.
Timdal
O-L-184415 OLICH696-13 MK812038
Pleopsidium
flavum
Acarosporales NO,
Hordaland,
2013, E.
Timdal
O-L-184439 OLICH697-13 MK811877
Pleurosticta
acetabulum
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-199985 OLICH1959-15 MK812572
Pleurosticta
acetabulum
Lecanorales NO, Vestfold,
2015, E.
Timdal
O-L-198935 OLICH2488-16 MK812152
Pleurosticta
acetabulum
Lecanorales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200869 OLICH2521-16 MK811775
Pleurosticta
acetabulum
Lecanorales NO, Rogaland,
2015, T.
Tønsberg & E.
Nygaard
BG-
L-98170
OLICH3700-17 MK811799
Pleurosticta
acetabulum
Lecanorales NO, Østfold,
2002, B. P.
Løfall
O-L-108808 OLICH4128-17 MK812479
Polycauliona
candelaria
Teloschistales NO, Akershus,
2014, E.
Timdal
O-L-199981 OLICH1955-15 MK811915
Polycauliona
candelaria
Teloschistales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196326 OLICH2256-15 MK812066
94 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Polycauliona
candelaria
Teloschistales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196340 OLICH2278-15 MK811959
Polychidium
muscicola
Peltigerales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201293 OLICH2795-16 MK812283
Protoblastenia
rupestris
Lecanorales NO, Finnmark,
2014, A.
Millanes
O-L-195859 OLICH1704-14 KY266981
Protomicarea
limosa
Lecanorales NO, Hedmark,
2012, R.
Haugan
O-L-182289 OLICH1084-14 MK811666
Protomicarea
limosa
Lecanorales NO, Hedmark,
2004, R.
Haugan
O-L-131761 OLICH1085-14 MK812532
Protomicarea
limosa
Lecanorales NO, Nord-
Trøndelag,
2009, J.T.
Klepsland
O-L-165102 OLICH1086-14 MK812181
Protomicarea
limosa
Lecanorales NO, Oppland,
2012, E.
Timdal
O-L-182031 OLICH1140-14 MK812033
Protomicarea
limosa
Lecanorales NO, Nordland,
2010, T.
Tønsberg
BG-
L-92492
OLICH601-13 MK812586
Protomicarea
limosa
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184281 OLICH821-13 MK812176
Protopannaria
pezizoides
Peltigerales NO, Finnmark,
2014, A.
Guillermo
O-L-195697 OLICH1569-14 KY266840
Protopannaria
pezizoides
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195890 OLICH1735-14 KY266909
OLICH: A reference library of DNA barcodes for Nordic lichens 95

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Protopannaria
pezizoides
Peltigerales NO, Finnmark,
2014, H.
Holien
O-L-195970 OLICH1815-14 KY266886
Protopannaria
pezizoides
Peltigerales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196347 OLICH1907-15 MK811767
Protopannaria
pezizoides
Peltigerales NO, Nord-
Trøndelag,
2013, J.T.
Klepsland
O-L-197950 OLICH2115-15 MK812467
Protopannaria
pezizoides
Peltigerales NO, Svalbard,
2015, S. Rui
O-L-203297 OLICH2706-16 MK811974
Protoparmelia
badia
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195568 OLICH1522-14 KY266939
Protoparmelia
badia
Lecanorales NO, Finnmark,
2014, A.
Launis
O-L-195839 OLICH1684-14 KY266945
Pseudephebe
minuscula
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196378 OLICH1937-15 MK811997
Pseudephebe
minuscula
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184305 OLICH736-13 MK812067
Pseudephebe
pubescens
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195619 OLICH1548-14 MK812357
Pseudephebe
pubescens
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195849 OLICH1694-14 KY266938
Pseudephebe
pubescens
Lecanorales NO, Finnmark,
2014, R.
Haugan
O-L-195986 OLICH1831-14 KY266914
96 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Pseudephebe
pubescens
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184396 OLICH694-13 MK812150
Pseudephebe
pubescens
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184306 OLICH735-13 MK811888
Pseudephebe
pubescens
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184725 OLICH941-13 MK812652
Pseudevernia
furfuracea
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-195999 OLICH1844-14 MK812319
Pseudevernia
furfuracea
Lecanorales NO, Hedmark,
2016, E.
Timdal
O-L-201522 OLICH3984-17 MK812161
Pseudevernia
furfuracea
Lecanorales NO, Aust-
Agder, 2016,
S. Rui & E.
Timdal
O-L-204977 OLICH4008-17 MK812508
Pseudevernia
furfuracea
Lecanorales NO, Telemark,
2016, S. Rui &
E. Timdal
O-L-205006 OLICH4009-17 MK811934
Pseudevernia
furfuracea
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184698 OLICH882-13 MK812628
Pseudocyphellaria
citrina
Peltigerales NO, Nord-
Trøndelag,
2013, J.T.
Klepsland
O-L-197920 OLICH2116-15 MK812391
Pseudocyphellaria
citrina
Peltigerales NO, Sør-
Trøndelag,
2013, J.T.
Klepsland
O-L-198050 OLICH2117-15 MK811890
OLICH: A reference library of DNA barcodes for Nordic lichens 97

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Pseudocyphellaria
citrina
Peltigerales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201266 OLICH2771-16 MK812527
Pseudocyphellaria
norvegica
Peltigerales NO,
Hordaland,
2006, H.
Fjeldstad
O-L-203000 OLICH3242-16 MK812692
Psilolechia
clavulifera
Lecanorales NO, Akershus,
2009, J.T.
Klepsland
O-L-164653 OLICH1081-14 MK811831
Psilolechia
leprosa
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195611 OLICH1544-14 MK812516
Psilolechia
leprosa
Lecanorales NO, Aust-
Agder, 2015,
J.T. Klepsland
O-L-204517 OLICH4229-17 MK811951
Psilolechia
leprosa
Lecanorales NO, Aust-
Agder, 2015,
J.T. Klepsland
O-L-204519 OLICH4251-17 MK812436
Psilolechia lucida Lecanorales SE, Närke,
2013, M.
Bendiksby et
al.
O-L-184487 OLICH665-13 MK811768
Psora decipiens Lecanorales NO, Oppland,
2013, J.T.
Klepsland
O-L-198348 OLICH2358-15 MK812030
Psora decipiens Lecanorales NO, Troms,
2014, J.T.
Klepsland
O-L-200362 OLICH2646-16 MK811804
Psora globifera Lecanorales NO, Nord-
Trøndelag,
2011, J.T.
Klepsland
O-L-177145 OLICH1319-14 KU873929 no lichen
substances
98 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Psora globifera Lecanorales NO, Sør-
Trøndelag,
2012, R.
Hjelmstad
O-L-184143 OLICH1320-14 KU873932 no lichen
substances
Psora globifera Lecanorales NO, Buskerud,
2014, J.T.
Klepsland
O-L-200597 OLICH2645-16 MK812110
Psora globifera Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183774 OLICH782-13 KU873928 no lichen
substances
Psora globifera Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184327 OLICH820-13 KU873930 no lichen
substances
Psora rubiformis Lecanorales NO, Oppland,
2012, H. Bratli
& R. Haugan
O-L-183466 OLICH1082-14 MK811688
Psora rubiformis Lecanorales NO, Nord-
Trøndelag,
2011, J.T.
Klepsland
O-L-177142 OLICH1321-14 MK812240
Psora rubiformis Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195525 OLICH1508-14 KY266957
Psora rubiformis Lecanorales NO,
Hordaland,
2013, E.
Timdal
O-L-184422 OLICH648-13 MK811706
Psora rubiformis Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184359 OLICH649-13 MK812000
Psora vallesiaca Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184392 OLICH650-13 KU873926 norstictic acid
OLICH: A reference library of DNA barcodes for Nordic lichens 99

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Psora vallesiaca Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183778 OLICH780-13 KU873927 norstictic acid
Psora vallesiaca Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183760 OLICH781-13 KU873931 norstictic acid
Psorinia
conglomerata
Lecanorales NO, Oppland,
2012, H. Bratli
& R. Haugan
O-L-182192 OLICH1061-14 MK812001 stictic acid
Psoroma
cinnamomeum
Peltigerales NO, Sør-
Trøndelag,
2013, E.
Timdal
O-L-184538 OLICH702-13 MK811978
Psoroma
hypnorum
Peltigerales NO, Finnmark,
2014, E.
Timdal
O-L-195857 OLICH1702-14 KY266971
Psoroma
hypnorum
Peltigerales NO, Sør-
Trøndelag,
2017, E.
Timdal
O-L-208202 OLICH4325-17 MK811800
Psoroma
hypnorum
Peltigerales NO, Sør-
Trøndelag,
2013, E.
Timdal
O-L-184537 OLICH703-13 MK811764
Psoronactis
dilleniana
Arthoniales NO, Nord-
Trøndelag,
2015, H.
Holien
TRH-
L-16932
OLICH3649-17 MK812257
Punctelia stictica Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201333 OLICH2698-16 MK812448
Punctelia
subrudecta
Lecanorales NO, Rogaland,
2015, T.
Tønsberg
BG-
L-98118
OLICH3697-17 MK812035
Punctelia
subrudecta
Lecanorales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200662 OLICH4106-17 MK812237
100 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Pycnothelia
papillaria
Lecanorales NO, Aust-
Agder, 2006,
J.T. Klepsland
O-L-151129 OLICH1364-14 MK811765 atranorin,
lichesterinic acid
Pycnothelia
papillaria
Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-200001 OLICH1975-15 MK812272
Pycnothelia
papillaria
Lecanorales NO, Nord-
Trøndelag,
2015, M.
Bendiksby et
al.
O-L-201299 OLICH2797-16 MK811947
Pyrenula
laevigata
Pyrenulales NO,
Hordaland,
2016, J.T.
Klepsland
O-L-206758 OLICH3477-17 MK812685
Pyrenula
laevigata
Pyrenulales NO, Møre og
Romsdal,
2016, J.T.
Klepsland
O-L-206773 OLICH3492-17 MK812185
Pyrenula
occidentalis
Pyrenulales NO, Møre og
Romsdal,
2016, J.T.
Klepsland
O-L-206777 OLICH3496-17 MK811633
Racodium
rupestre
Capnodiales NO, Rogaland,
2013, J.T.
Klepsland
O-L-198464 OLICH3945-17 MK812095
Ramalina capitata Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200012 OLICH1986-15 MK811917
Ramalina capitata Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-195831 OLICH2203-15 MK811626
Ramalina capitata Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200896 OLICH2549-16 MK812528
OLICH: A reference library of DNA barcodes for Nordic lichens 101

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Ramalina capitata Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184378 OLICH693-13 MK812398
Ramalina
cuspidata
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196016 OLICH1861-14 MK812006 usnic acid
Ramalina
cuspidata
Lecanorales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197829 OLICH2118-15 MK811797
Ramalina
farinacea
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196002 OLICH1847-14 MK812207 protocetraric acid,
usnic acid
Ramalina
farinacea
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-199987 OLICH1961-15 MK812169 protocetraric acid,
usnic acid
Ramalina
farinacea
Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196286 OLICH2216-15 MK812449 protocetraric acid ,
usnic acid
Ramalina
fastigiata
Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196289 OLICH2219-15 MK811906
Ramalina
fastigiata
Lecanorales NO, Vestfold,
2014, J.T.
Klepsland
O-L-200570 OLICH2644-16 MK811841
Ramalina
fastigiata
Lecanorales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208026 OLICH4197-17 MK812196
Ramalina fraxinea Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196288 OLICH2218-15 MK812590
Ramalina fraxinea Lecanorales NO, Vestfold,
2014, J.T.
Klepsland
O-L-200571 OLICH4105-17 MK812489
Ramalina fraxinea Lecanorales NO, Akershus,
2012, K.A. Lye
O-L-206861 OLICH4140-17 MK811900
102 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Ramalina fraxinea Lecanorales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208025 OLICH4196-17 MK812162
Ramalina
pollinaria
Lecanorales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196372 OLICH1931-15 MK812166
Ramalina
pollinaria
Lecanorales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200850 OLICH2516-16 MK812114
Ramalina
pollinaria
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200901 OLICH2552-16 MK811866
Ramalina
polymorpha
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195737 OLICH1609-14 KY266885
Ramalina
polymorpha
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195928 OLICH1773-14 MK811690
Ramalina
polymorpha
Lecanorales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-200011 OLICH1985-15 MK811661
Ramalina
polymorpha
Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200898 OLICH2551-16 MK812673
Ramalina
siliquosa
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196010 OLICH1855-14 MK811720 protocetraric acid
(minor), salazinic
acid (major), usnic
acid (trace)
Ramalina
subfarinacea
Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196284 OLICH2214-15 MK812104 norstictic acid
(major), usnic acid
(major)
Ramalina
subfarinacea
Lecanorales NO, Nordland,
2014, J.T.
Klepsland
O-L-200339 OLICH2643-16 MK812227 norstictic acid
(major), usnic acid
(major)
OLICH: A reference library of DNA barcodes for Nordic lichens 103

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Ramboldia
elabens
Lecanorales NO, Sør-
Trøndelag,
2007, E.
Timdal
O-L-158501 OLICH411-13 MK812637
Ramboldia
elabens
Lecanorales NO, Hedmark,
2008, R.
Haugan
O-L-160473 OLICH412-13 MK812132
Ramboldia
elabens
Lecanorales NO, Oppland,
2004, T.H.
Hofton
O-L-136707 OLICH413-13 MK812702
Ramboldia
elabens
Lecanorales NO, Nord-
Trøndelag,
2006, J.T.
Klepsland
O-L-183511 OLICH472-13 MK811668
Ramboldia
elabens
Lecanorales NO, Finnmark,
2011, E.
Timdal
O-L-170692 OLICH473-13 MK811870
Ramboldia
elabens
Lecanorales NO, Nordland,
2011, T.
Tønsberg
BG-
L-95299
OLICH580-13 MK812420
Rhizocarpon
badioatrum
Rhizocarpales NO, Oppland,
2010, E.
Timdal
O-L-163824 OLICH002-11 KU687450 no lichen
substances
Rhizocarpon
badioatrum
Rhizocarpales NO, Buskerud,
2010, E.
Timdal
O-L-163839 OLICH003-11 KU687453 no lichen
substances
Rhizocarpon
bolanderi
Rhizocarpales NO, Aust-
Agder, 2010,
E. Timdal
O-L-163373 OLICH023-11 KU687451 norstictic acid
(trace), stictic acid
(major)
Rhizocarpon
chioneum
Rhizocarpales NO, Finnmark,
2014, E.
Timdal
O-L-195535 OLICH1510-14 KY266951
Rhizocarpon
copelandii
Rhizocarpales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184329 OLICH766-13 KU687447
104 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Rhizocarpon
copelandii
Rhizocarpales NO, Hedmark,
2013, E.
Timdal
O-L-184521 OLICH790-13 KU687456 norstictic acid
Rhizocarpon
copelandii
Rhizocarpales NO, Oppland,
2012, S. Rui &
E. Timdal
O-L-180002 OLICH837-13 KU687455 norstictic acid
Rhizocarpon
geminatum
Rhizocarpales NO, Finnmark,
2014, E.
Timdal
O-L-195564 OLICH1520-14 KY266908
Rhizocarpon
jemtlandicum
Rhizocarpales NO, Telemark,
2010, E.
Timdal
O-L-163364 OLICH015-11 KU687445 norstictic acid
(trace), stictic acid
(major)
Rhizocarpon
jemtlandicum
Rhizocarpales NO, Nordland,
2010, E.
Timdal
O-L-163756 OLICH019-11 KU687446 norstictic acid
(minor), stictic acid
(major)
Rhizocarpon
jemtlandicum
Rhizocarpales NO, Finnmark,
2011, E.
Timdal
O-L-170452 OLICH803-13 KU687460 stictic acid
Rhizocarpon
leptolepis
Rhizocarpales FI, Inarin
Lappi, 2014, E.
Timdal
O-L-195500 OLICH1504-14 KU687449
Rhizocarpon
pycnocarpoides
Rhizocarpales NO, Nordland,
2012, R.
Haugan
O-L-183808 OLICH511-13 KT800004 no lichen
substances
Rhizocarpon
pycnocarpoides
Rhizocarpales NO, Nord-
Trøndelag,
2012, R.
Haugan
O-L-183810 OLICH512-13 KT800005 no lichen
substances
Rhizocarpon
pycnocarpoides
Rhizocarpales NO, Oppland,
2012, S. Rui &
E. Timdal
O-L-179903 OLICH513-13 KT800002
Rhizocarpon
pycnocarpoides
Rhizocarpales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184267 OLICH531-13 KT800003 no lichen
substances
Rhizocarpon
rittokense
Rhizocarpales NO, Finnmark,
2014, M.
Westberg
O-L-195695 OLICH1567-14 KU687458
OLICH: A reference library of DNA barcodes for Nordic lichens 105

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Rhizocarpon
rittokense
Rhizocarpales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184293 OLICH542-13 KU687459
Rhizocarpon
santessonii
Rhizocarpales NO, Oppland,
2011, R.
Haugan & E.
Timdal
O-L-169805 OLICH432-13 KU687454
Rhizocarpon
subgeminatum
Rhizocarpales NO, Hedmark,
2010, R.
Haugan
O-L-166501 OLICH033-11 KU687457 barbatic acid
Rhizocarpon
subgeminatum
Rhizocarpales NO, Sør-
Trøndelag,
2010, R.
Haugan
O-L-165477 OLICH034-11 KU687452 barbatic acid
Rhizocarpon
suomiense
Rhizocarpales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184330 OLICH769-13 KU687448
Rhizoplaca
melanophthalma
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195929 OLICH1774-14 MK812478 unknown
substances, usnic
acid
Rhizoplaca
melanophthalma
Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201366 OLICH2743-16 MK811669
Rinodina balanina Caliciales NO, Finnmark,
2014, T.B.
Wheeler
O-L-195705 OLICH1577-14 KY266842
Rinodina
disjuncta
Caliciales NO, Nord-
Trøndelag,
2014, H.
Holien
TRH-
L-15387
OLICH3675-17 MK812529
Rinodina
disjuncta
Caliciales NO, Nord-
Trøndelag,
2011, H.
Holien
TRH-
L-14116
OLICH3676-17 MK812438
106 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Rinodina
mniaraea
Caliciales NO, Svalbard,
2001, A.
Elvebakk
TROM_L
_565871
OLICH3552-17 MK812098
Rinodina roboris Caliciales NO, Rogaland,
2016, J.T.
Klepsland
O-L-206765 OLICH3484-17 MK811851
Romjularia lurida Lecideales NO, Akershus,
2010, E.
Timdal
O-L-169118 OLICH150-11 MK811769
Ropalospora
lugubris
Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195537 OLICH1511-14 KY266893
Ropalospora
lugubris
Umbilicariales NO, Nordland,
2015, T.
Tønsberg & A.
Botnen
BG-
L-98792
OLICH3732-17 MK812617
Ropalospora
lugubris
Umbilicariales NO, Finnmark,
2011, E.
Timdal
O-L-170519 OLICH3969-17 MK812431
Ropalospora
lugubris
Umbilicariales NO, Sør-
Trøndelag,
2011, R.
Haugan
O-L-168439 OLICH3970-17 MK812670
Ropalospora
lugubris
Umbilicariales NO, Sogn og
Fjordane,
2011, R.
Haugan
O-L-173455 OLICH3971-17 MK812610
Rusavskia
elegans
Teloschistales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196351 OLICH1910-15 MK812165
Rusavskia
elegans
Teloschistales NO, Oppland,
2013, J.T.
Klepsland
O-L-200120 OLICH2630-16 MK812212
Rusavskia
sorediata
Teloschistales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196353 OLICH1912-15 MK811744
OLICH: A reference library of DNA barcodes for Nordic lichens 107

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Sagiolechia
protuberans
Ostropales NO, Finnmark,
2014, M.
Westberg
O-L-195714 OLICH1586-14 KY266966
Sagiolechia
protuberans
Ostropales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201297 OLICH3853-17 MK811691
Sclerophora
peronella
Coniocybales SE,
Södermanland,
2015, S.
Kistenich & E.
Timdal
O-L-200846 OLICH2432-16 MK812193
Scoliciosporum
umbrinum
Lecanorales NO, Oslo,
2014, J.T.
Klepsland
O-L-200123 OLICH3322-16 MK812255
Scoliciosporum
umbrinum
Lecanorales NO, Akershus,
2012, E.
Timdal
O-L-177770 OLICH822-13 MK812271
Siphula ceratites Pertusariales NO, Finnmark,
2014, E.
Timdal
O-L-195919 OLICH1764-14 KY266949
Siphula ceratites Pertusariales NO, Nord-
Trøndelag,
2015, E.
Timdal et al.
O-L-201291 OLICH2679-16 MK812106
Siphula ceratites Pertusariales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196042 OLICH3790-17 MK812034
Solorina saccata Peltigerales NO, Buskerud,
2014, E.
Timdal
O-L-203070 OLICH3286-16 MK812538
Sphaerophorus
fragilis
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195650 OLICH1562-14 MK812306
108 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Sphaerophorus
fragilis
Lecanorales NO,
Hordaland,
2014, E.
Timdal
O-L-195800 OLICH1672-14 MK811867
Sphaerophorus
fragilis
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195977 OLICH1822-14 KY266958
Sphaerophorus
fragilis
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196043 OLICH3799-17 MK812393
Sphaerophorus
globosus
Lecanorales NO, Troms,
2014, E.
Timdal
O-L-195651 OLICH1563-14 MK811784 sphaerophorin
Sphaerophorus
globosus
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195855 OLICH1700-14 KY266841 sphaerophorin
Sphaerophorus
globosus
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195918 OLICH1763-14 KY266859 sphaerophorin,
squamatic acid
Sphaerophorus
globosus
Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195949 OLICH1794-14 KY266898
Sphaerophorus
globosus
Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-199995 OLICH1969-15 MK812694 sphaerophorin,
thamnolic acid,
unknowns
Sphaerophorus
globosus
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200918 OLICH2578-16 MK812575 sphaerophorin
Sphaerophorus
globosus
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196029 OLICH3793-17 MK811761 sphaerophorin,
squamatic acid
Sphaerophorus
globosus
Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201265 OLICH3844-17 MK812327 sphaerophorin,
thamnolic acid
OLICH: A reference library of DNA barcodes for Nordic lichens 109

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Staurolemma
omphalarioides
Peltigerales NO, Nordland,
2013, J.T.
Klepsland
O-L-197994 OLICH2467-16 MK811827
Staurolemma
omphalarioides
Peltigerales NO, Nordland,
2014, J.T.
Klepsland
O-L-200306 OLICH2640-16 MK811929
Stereocaulon
alpinum
Lecanorales NO, Telemark,
2006, E.
Timdal
O-L-149404 OLICH925-13 MK812253 atranorin, lobaric
acid
Stereocaulon
capitellatum
Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201401 OLICH4182-17 MK812320 anziaic acid,
atranorin, miriquidic
acid, perlatolic acid
Stereocaulon
coniophyllum
Lecanorales NO, Sogn og
Fjordane,
2010, S. Vatne
O-L-169267 OLICH713-13 MK811930 lobaric acid,
atranorin
Stereocaulon
cumulatum
Lecanorales NO, Finnmark,
2014, M.
Svensson
O-L-195924 OLICH1769-14 KY266904
Stereocaulon
cumulatum
Lecanorales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200899 OLICH2529-16 MK812171
Stereocaulon
cumulatum
Lecanorales NO, Hedmark,
2013, E.
Timdal
O-L-184515 OLICH674-13 MK812474
Stereocaulon
cumulatum
Lecanorales NO, Oppland,
2012, E.
Timdal
O-L-182037 OLICH711-13 MK812671
Stereocaulon
grande
Lecanorales NO, Troms,
2003, E.
Timdal
O-L-122991 OLICH917-13 MK812157 atranorin,
bourgeanic acid,
lobaric acid
Stereocaulon
pileatum
Lecanorales NO, Aust-
Agder, 2016,
S. Rui & E.
Timdal
O-L-204980 OLICH4012-17 MK812053
Stereocaulon
pileatum
Lecanorales NO, Østfold,
2002, B.P.
Løfall & A.
Often
O-L-113318 OLICH908-13 MK812510 atranorin, lobaric
acid
110 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Stereocaulon
pileatum
Lecanorales NO, Oslo,
2011, H. Bratli
& R. Haugan
O-L-174391 OLICH909-13 MK811864 atranorin, lobaric
acid
Stereocaulon
rivulorum
Lecanorales NO, Troms,
2003, E.
Timdal
O-L-127342 OLICH899-13 MK812658 anziaic acid,
atranorin, perlatolic
acid
Stereocaulon
rivulorum
Lecanorales NO, Finnmark,
2003, E.
Timdal
O-L-123189 OLICH901-13 MK811662 anziaic acid,
atranorin, perlatolic
acid
Stereocaulon
symphycheilum
Lecanorales NO, Oppland,
2012, H. Bratli
O-L-183927 OLICH706-13 MK811941 lobaric acid
Stereocaulon
symphycheilum
Lecanorales NO, Buskerud,
2012, S. Rui &
E. Timdal
O-L-184640 OLICH730-13 MK812328 atranorin, lobaric
acid
Stereocaulon
symphycheilum
Lecanorales NO, Sør-
Trøndelag,
2009, E.
Timdal
O-L-158545 OLICH895-13 MK812189 atranorin, lobaric
acid
Sticta fuliginoides Peltigerales NO, Rogaland,
2012, B.
Nordén & J.B.
Jordal
O-L-199414 OLICH2638-16 MK811703
Sticta sylvatica Peltigerales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200925 OLICH2539-16 MK811909
Tephromela atra Lecanorales NO, Finnmark,
2014, A.
Millanes
O-L-195861 OLICH1706-14 KY266944
Tephromela atra Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196054 OLICH3815-17 MK812606
Tephromela atra Lecanorales NO, Telemark,
2009, R.
Haugan & S.
Reiso
O-L-160434 OLICH399-13 MK812380
OLICH: A reference library of DNA barcodes for Nordic lichens 111

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Tephromela
grumosa
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-190787 OLICH2594-16 MK812127 atranorin,
lichesterinic acid
Tetramelas
chloroleucus
Caliciales NO, Sør-
Trøndelag,
2014, H.
Holien
TRH-
L-15560
OLICH3601-17 MK811644
Tetramelas
chloroleucus
Caliciales NO, Nordland,
2013, J.T.
Klepsland
O-L-198004 OLICH4032-17 MK812300
Thamnolia
vermicularis
Pertusariales NO, Finnmark,
2014, E.
Timdal
O-L-195710 OLICH1582-14 KY266979
Thamnolia
vermicularis
Pertusariales NO, Troms,
2014, E.
Timdal
O-L-195773 OLICH1645-14 MK812339
Thamnolia
vermicularis
Pertusariales NO, Finnmark,
2014, H.
Holien
O-L-195948 OLICH1793-14 KY266969
Thamnolia
vermicularis
Pertusariales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196363 OLICH1922-15 MK811783
Thamnolia
vermicularis
Pertusariales NO, Buskerud,
2015, E.
Timdal & S.
Rui
O-L-200905 OLICH3840-17 MK811627
Thelotrema
lepadinum
Ostropales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199496 OLICH3321-16 MK812434
Thelotrema
macrosporum
Ostropales NO,
Hordaland,
2013, B.
Nordén & J.B.
Jordal
O-L-199493 OLICH3320-16 MK812111
112 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Tholurna
dissimilis
Caliciales NO, Buskerud,
2014, E.
Timdal
O-L-195822 OLICH2194-15 MK812234
Tholurna
dissimilis
Caliciales NO, Nord-
Trøndelag,
2015, H.
Holien
TRH-
L-16038
OLICH2760-16 MK811683
Timdalia intricata Acarosporales NO, Telemark,
2013, E.
Timdal
O-L-194731 OLICH3244-16 MK812713
Timdalia intricata Acarosporales NO, Oppland,
2015, E.
Timdal
O-L-201379 OLICH4168-17 MK812246
Timdalia intricata Acarosporales NO, Hedmark,
2017, E.
Timdal
O-L-207764 OLICH4349-17 MK811937 psoromic acid
Timdalia intricata Acarosporales SE, Jämtland,
2017, E.
Timdal
O-L-208174 OLICH4350-17 MK811871 psoromic acid
Timdalia intricata Acarosporales SE,
Härjedalen,
2017, E.
Timdal
O-L-208190 OLICH4351-17 MK812454 psoromic acid
Toensbergia
leucococca
Rhizocarpales NO, Nordland,
2012, T.
Tønsberg
BG-
L-94650
OLICH592-13 MK812615
Toninia alutacea Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195931 OLICH1776-14 MG838194
Toninia alutacea Lecanorales NO, Buskerud,
2014, J.T.
Klepsland
O-L-200591 OLICH2636-16 MG838164
Toninia alutacea Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184350 OLICH551-13 MG838166
OLICH: A reference library of DNA barcodes for Nordic lichens 113

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Toninia alutacea Lecanorales NO, Troms,
2011, J.T.
Klepsland
O-L-177188 OLICH561-13 MG838197
Toninia alutacea Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183755 OLICH783-13 MG838156
Toninia aromatica Lecanorales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-L-200557 OLICH3317-16 MG838189
Toninia aromatica Lecanorales NO, Troms,
2009, J.T.
Klepsland
O-L-167368 OLICH560-13 MG838184
Toninia aromatica Lecanorales NO, Nordland,
2011, R.
Haugan
O-L-174042 OLICH614-13 MG838161
Toninia candida Lecanorales NO, Buskerud,
2014, E.
Timdal
O-L-196069 OLICH1885-14 MG838163
Toninia candida Lecanorales NO, Buskerud,
2011, K.A. Lye
& T. Høitomt
O-L-177883 OLICH557-13 MG838185
Toninia coelestina Lecanorales NO, Nordland,
2010, J.T.
Klepsland
O-L-175215 OLICH285-13 MG838190
Toninia
nordlandica
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195601 OLICH1537-14 MG838198
Toninia
nordlandica
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184351 OLICH552-13 MG838159
Toninia
opuntioides
Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195603 OLICH1539-14 MG838192
Toninia
opuntioides
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183630 OLICH559-13 MG838168 triterpenoid (minor)
114 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Toninia
opuntioides
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184387 OLICH635-13 MG838172 triterpenoid
(possible trace)
Toninia
opuntioides
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183757 OLICH784-13 MG838158
Toninia rosulata Lecanorales NO, Nord-
Trøndelag,
2011, J.T.
Klepsland
O-L-177141 OLICH1323-14 MG838195
Toninia sedifolia Lecanorales NO, Finnmark,
2014, E.
Timdal
O-L-195599 OLICH1536-14 MG838193
Toninia sedifolia Lecanorales NO, Aust-
Agder, 2010,
J.T. Klepsland
O-L-183585 OLICH558-13 MG838201 no lichen
substances
Toninia sedifolia Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184381 OLICH637-13 MG838196
Toninia sedifolia Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184332 OLICH817-13 MG838162
Toninia squalida Lecanorales NO, Oppland,
2011, R.
Haugan & E.
Timdal
O-L-169774 OLICH553-13 MG838183
Toninia squalida Lecanorales NO, Oppland,
2012, H. Bratli
& R. Haugan
O-L-182301 OLICH554-13 MG838182
Toninia squalida Lecanorales NO,
Hordaland,
2013, E.
Timdal
O-L-184426 OLICH638-13 MG838181
OLICH: A reference library of DNA barcodes for Nordic lichens 115

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Toninia squalida Lecanorales NO, Sør-
Trøndelag,
2013, E.
Timdal
O-L-184539 OLICH639-13 MG838200
Toninia taurica Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184333 OLICH546-13 MG838174
Toninia
verrucarioides
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184389 OLICH641-13 MG838167
Toninia
verrucarioides
Lecanorales NO, Oppland,
2011, J.T.
Klepsland
O-L-183717 OLICH850-13 MG838179
Trapelia coarctata Trapeliales NO, Oslo,
2014, J.T.
Klepsland
O-L-200174 OLICH3316-16 MK812584
Trapelia coarctata Trapeliales NO, Hedmark,
2012, R.
Haugan et al.
O-L-182063 OLICH562-13 MK812526
Trapelia coarctata Trapeliales NO, Telemark,
2012, E.
Timdal
O-L-179924 OLICH563-13 MK812177
Trapelia corticola Trapeliales NO, Nord-
Trøndelag,
2011, H.
Holien
TRH-
L-14083
OLICH993-13 MK811911
Trapelia
placodioides
Trapeliales NO, Nord-
Trøndelag,
2012, R.
Haugan
O-L-183857 OLICH1324-14 MK812664
Trapelia
placodioides
Trapeliales NO, Telemark,
1996, H. Bratli
& E. Timdal
O-L-20869 OLICH3976-17 MK812430
Trapelia
placodioides
Trapeliales NO, Hedmark,
2015, R.
Haugan
O-L-204154 OLICH4366-17 MK812690 gyrophoric acid,
lecanoric acid
116 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Trapelia
placodioides
Trapeliales NO, Oslo,
2014, J.T.
Klepsland
O-L-200245 OLICH4453-17 MK812050
Trapelia
placodioides
Trapeliales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184337 OLICH548-13 MK812359
Trapelia
placodioides
Trapeliales NO, Oppland,
2012, S. Rui &
E. Timdal
O-L-179918 OLICH569-13 MK812043
Trapeliopsis
flexuosa
Trapeliales NO, Nord-
Trøndelag,
2013, H.
Holien
TRH-
L-15151
OLICH994-13 MK811801
Trapeliopsis
granulosa
Trapeliales NO, Sør-
Trøndelag,
2007, E.
Timdal
O-L-158481 OLICH565-13 MK812609
Trapeliopsis
granulosa
Trapeliales NO, Oslo,
2011, R.
Haugan
O-L-177342 OLICH566-13 MK811943
Trapeliopsis
granulosa
Trapeliales NO, Troms,
2011, J.T.
Klepsland
O-L-177175 OLICH567-13 MK812708
Trapeliopsis
granulosa
Trapeliales NO, Finnmark,
2011, E.
Timdal
O-L-170698 OLICH568-13 MK812198
Trapeliopsis
granulosa
Trapeliales NO, Nord-
Trøndelag,
2013, H.
Holien
TRH-
L-15150
OLICH995-13 MK812377
Trapeliopsis
pseudogranulosa
Trapeliales NO, Telemark,
2015, E.
Timdal
O-L-208001 OLICH3875-17 MK812568
Trapeliopsis
pseudogranulosa
Trapeliales NO, Rogaland,
2011, J.T.
Klepsland
O-L-183692 OLICH852-13 MK812270
OLICH: A reference library of DNA barcodes for Nordic lichens 117

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Tremolecia atrata Hymeneliales NO, Sør-
Trøndelag,
2012, M.
Bendiksby et
al.
O-L-179677 OLICH4017-17 MK812298
Tremolecia atrata Hymeneliales NO, Finnmark,
2011, E.
Timdal
O-L-170578 OLICH606-13 MK811647
Tremolecia atrata Hymeneliales NO, Finnmark,
2011, E.
Timdal
O-L-170776 OLICH607-13 MK812097
Tuckermanopsis
chlorophylla
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196001 OLICH1846-14 MK811914
Tuckermanopsis
chlorophylla
Lecanorales NO, Akershus,
2014, S. Rui &
E. Timdal
O-L-196329 OLICH2260-15 MK812387
Umbilicaria arctica Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195898 OLICH1743-14 KY266911
Umbilicaria arctica Umbilicariales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200892 OLICH2545-16 MK812428
Umbilicaria arctica Umbilicariales NO, Aust-
Agder, 2016,
S. Rui & E.
Timdal
O-L-204983 OLICH4016-17 MK812268
Umbilicaria
cinereorufescens
Umbilicariales NO, Buskerud,
2014, E.
Timdal
O-L-195830 OLICH2202-15 MK811655
Umbilicaria
cinereorufescens
Umbilicariales NO, Oppland,
2015, E.
Timdal
O-L-201349 OLICH2734-16 MK812545
Umbilicaria
crustulosa
Umbilicariales NO,
Hordaland,
2014, E.
Timdal
O-L-195804 OLICH1676-14 MK811776
118 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Umbilicaria
crustulosa
Umbilicariales NO, Buskerud,
2015, S. Rui &
E. Timdal
O-L-200941 OLICH2662-16 MK812060
Umbilicaria
cylindrica
Umbilicariales NO,
Hordaland,
2014, E.
Timdal
O-L-195808 OLICH1680-14 MK811748
Umbilicaria
cylindrica
Umbilicariales NO, Finnmark,
2014, T.
Tønsberg
O-L-195872 OLICH1717-14 KY266907
Umbilicaria
cylindrica
Umbilicariales NO, Finnmark,
2014, H.
Holien
O-L-195957 OLICH1802-14 KY266835
Umbilicaria
cylindrica
Umbilicariales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196354 OLICH1913-15 MK812072
Umbilicaria
cylindrica
Umbilicariales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184711 OLICH929-13 MK812153
Umbilicaria
decussata
Umbilicariales NO, Sør-
Trøndelag,
2015, M.
Stapnes Dahl
et al.
O-L-201307 OLICH2692-16 MK812293
Umbilicaria
decussata
Umbilicariales NO, Oppland,
2015, E.
Timdal
O-L-201367 OLICH2744-16 MK812686
Umbilicaria
dendrophora
Umbilicariales NO, Sør-
Trøndelag,
2013, E.
Timdal
O-L-184529 OLICH731-13 MK812276
Umbilicaria
deusta
Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195766 OLICH1638-14 KY266862
Umbilicaria
deusta
Umbilicariales NO, Telemark,
2014, E.
Timdal
O-L-195790 OLICH1662-14 MK812011
OLICH: A reference library of DNA barcodes for Nordic lichens 119

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Umbilicaria
deusta
Umbilicariales NO, Vestfold,
2015, E.
Timdal
O-L-198940 OLICH2493-16 MK812096
Umbilicaria
deusta
Umbilicariales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184712 OLICH930-13 MK812412
Umbilicaria
havaasii
Umbilicariales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200930 OLICH2556-16 MK812376
Umbilicaria
hirsuta
Umbilicariales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196282 OLICH2212-15 MK812437
Umbilicaria
hirsuta
Umbilicariales NO, Vestfold,
2015, E.
Timdal
O-L-198937 OLICH2490-16 MK811754
Umbilicaria
hirsuta
Umbilicariales NO, Oppland,
2015, E.
Timdal
O-L-201350 OLICH2735-16 MK812136
Umbilicaria
hyperborea
Umbilicariales NO,
Hordaland,
2014, E.
Timdal
O-L-195803 OLICH1675-14 MK812311
Umbilicaria
hyperborea
Umbilicariales NO, Finnmark,
2014, R.
Haugan
O-L-195879 OLICH1724-14 KY266918
Umbilicaria
hyperborea
Umbilicariales NO, Finnmark,
2014, R.
Haugan
O-L-195925 OLICH1770-14 KY266976
Umbilicaria
hyperborea
Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195971 OLICH1816-14 KY266928
Umbilicaria
hyperborea
Umbilicariales NO, Troms,
2014, E.
Timdal
O-L-195656 OLICH1996-15 MK812016
Umbilicaria
hyperborea
Umbilicariales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184713 OLICH931-13 MK811931
120 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Umbilicaria lyngei Umbilicariales NO, Sør-
Trøndelag,
2000, B. P.
Løfall
O-L-78093 OLICH4074-17 MK812368 no lichen
substances
Umbilicaria lyngei Umbilicariales NO, Oppland,
2009, R.
Haugan
O-L-159920 OLICH4075-17 MK812559
Umbilicaria lyngei Umbilicariales NO, Sør-
Trøndelag,
2014, J.T.
Klepsland
O-
L-200558b
OLICH4096-17 MK811698
Umbilicaria
nylanderiana
Umbilicariales NO, Hedmark,
2015, M.
Stapnes Dahl
et al.
O-L-201305 OLICH2670-16 MK812167
Umbilicaria
nylanderiana
Umbilicariales NO, Hedmark,
2016, E.
Timdal
O-L-201508 OLICH3983-17 MK812209
Umbilicaria
nylanderiana
Umbilicariales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184729 OLICH950-13 MK812316
Umbilicaria
polyrrhiza
Umbilicariales NO, Oslo,
2014, E.
Timdal
O-L-199999 OLICH1973-15 MK812126
Umbilicaria
polyrrhiza
Umbilicariales NO, Østfold,
2013, R.
Braathen &
E.W. Hanssen
O-L-201768 OLICH2595-16 MK812352
Umbilicaria
polyrrhiza
Umbilicariales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208345 OLICH4384-17 MK812239
Umbilicaria
proboscidea
Umbilicariales NO, Finnmark,
2014, H.
Holien
O-L-195719 OLICH1591-14 KY266839
Umbilicaria
proboscidea
Umbilicariales NO, Troms,
2014, E.
Timdal
O-L-195772 OLICH1644-14 MK811704
OLICH: A reference library of DNA barcodes for Nordic lichens 121

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Umbilicaria
proboscidea
Umbilicariales NO, Finnmark,
2014, R.
Haugan
O-L-195923 OLICH1768-14 KY266848
Umbilicaria
proboscidea
Umbilicariales NO, Finnmark,
2014, A.
Millanes
O-L-195936 OLICH1781-14 KY266866
Umbilicaria
proboscidea
Umbilicariales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196361 OLICH1920-15 MK812366
Umbilicaria
proboscidea
Umbilicariales NO, Buskerud,
2014, S. Rui &
E. Timdal
O-L-196322 OLICH2252-15 MK812345
Umbilicaria rigida Umbilicariales SE, Torne
Lappmark,
2014, E.
Timdal
O-L-195668 OLICH2005-15 MK812082
Umbilicaria rigida Umbilicariales NO, Møre og
Romsdal,
2015, R.
Haugan
O-L-203950 OLICH4077-17 MK811777
Umbilicaria
spodochroa
Umbilicariales NO, Vestfold,
2015, E.
Timdal
O-L-198938 OLICH2491-16 MK812519
Umbilicaria
torrefacta
Umbilicariales NO, Troms,
2014, E.
Timdal
O-L-195634 OLICH1553-14 MK812116
Umbilicaria
torrefacta
Umbilicariales NO, Finnmark,
2014, E.
Timdal
O-L-195843 OLICH1688-14 KY266942
Umbilicaria
torrefacta
Umbilicariales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200900 OLICH2530-16 MK812602
Umbilicaria vellea Umbilicariales NO, Telemark,
2014, E.
Timdal
O-L-196275 OLICH1949-15 MK811710
122 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Umbilicaria
virginis
Umbilicariales NO, Møre og
Romsdal,
2015, R.
Haugan
O-L-203963 OLICH3162-16 MK812576
Usnea dasypoga Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184699 OLICH1411-14 MK812140
Usnea dasypoga Lecanorales NO, Telemark,
2014, E.
Timdal
O-L-196273 OLICH1947-15 MK812173
Usnea dasypoga Lecanorales NO, Oslo,
2014, E.
Timdal
O-L-200005 OLICH1979-15 MK812052
Usnea flammea Lecanorales NO, Vest-
Agder, 2013,
J.T. Klepsland
O-L-197827 OLICH2127-15 MK811847 constictic acid,
stictic acid, usnic
acid
Usnea
fragilescens
Lecanorales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200604 OLICH2634-16 MK812021
Usnea
subfloridana
Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184700 OLICH1412-14 MK811828
Usnea
subfloridana
Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200814 OLICH2585-16 MK811766
Usnea
subfloridana
Lecanorales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200728 OLICH2623-16 MK812453 thamnolic acid,
usnic acid
Usnea wasmuthii Lecanorales NO, Sør-
Trøndelag,
2013, J.T.
Klepsland
O-L-198061 OLICH2131-15 MK812232 barbatic acid,
salazinic acid, usnic
acid
Usnea wasmuthii Lecanorales NO, Rogaland,
2013, J.T.
Klepsland
O-L-197890 OLICH2132-15 MK812501 barbatic acid, usnic
acid
Varicellaria lactea Pertusariales NO, Oppland,
2015, R.
Haugan
O-L-204064 OLICH4380-17 MK812539 lecanoric acid,
variolaric acid
OLICH: A reference library of DNA barcodes for Nordic lichens 123

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Variospora
flavescens
Teloschistales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200626 OLICH4054-17 MK812667
Variospora
thallincola
Teloschistales NO, Rogaland,
2014, J.T.
Klepsland
O-L-200631 OLICH4056-17 MK812547
Verrucaria
tenebrosa
Verrucariales No, Nord-
Trøndelag,
2015, J.
Pykälä 48333
H-
WG2_2130
OLICH2807-16 KY682961
Vulpicida
juniperinus
Lecanorales NO, Oppland,
2013, M.
Bendiksby et
al.
O-L-184298 OLICH1406-14 MK812148
Vulpicida
juniperinus
Lecanorales NO, Nord-
Trøndelag,
2015, R.
Haugan
O-L-201311 OLICH3857-17 MK812404
Vulpicida
juniperinus
Lecanorales NO, Sør-
Trøndelag,
2017, E.
Timdal
O-L-208203 OLICH4356-17 MK812500
Vulpicida pinastri Lecanorales NO, Finnmark,
2014, H.
Holien
O-L-195944 OLICH1789-14 KY266954
Vulpicida pinastri Lecanorales NO, Telemark,
2015, S. Rui &
E. Timdal
O-L-200937 OLICH2563-16 MK811933
Vulpicida pinastri Lecanorales NO, Buskerud,
2013, S. Rui &
E. Timdal
O-L-184701 OLICH883-13 MK812364 pinastric acid,
vulpinic acid
Xanthomendoza
borealis
Teloschistales NO, Sør-
Trøndelag,
2014, E.
Timdal
O-L-196336 OLICH2274-15 MK812442
Xanthomendoza
borealis
Teloschistales NO, Oppland,
2013, J.T.
Klepsland
O-L-198333 OLICH2359-15 MK811659
124 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Xanthomendoza
fulva
Teloschistales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196295 OLICH2225-15 MK812265
Xanthomendoza
fulva
Teloschistales NO, Hedmark,
2006, B. P.
Løfall et al.
O-L-147714 OLICH4440-17 MK812487
Xanthomendoza
fulva
Teloschistales NO, Sogn og
Fjordane,
2012, B.
Nordén & J.B.
Jordal
O-L-199397 OLICH4442-17 MK811973
Xanthoparmelia
conspersa
Lecanorales NO, Telemark,
2014, S. Rui &
E. Timdal
O-L-196278 OLICH1951-15 MK812598
Xanthoparmelia
conspersa
Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196296 OLICH2226-15 MK812585
Xanthoparmelia
conspersa
Lecanorales NO, Rogaland,
2016, S. Rui &
E. Timdal
O-L-204999 OLICH3992-17 MK812248
Xanthoparmelia
loxodes
Lecanorales NO, Akershus,
2014, E.
Timdal
O-L-199990 OLICH1964-15 MK811808
Xanthoparmelia
loxodes
Lecanorales NO, Vestfold,
2014, S. Rui &
E. Timdal
O-L-196281 OLICH2211-15 MK811902
Xanthoparmelia
loxodes
Lecanorales NO, Aust-
Agder, 2013,
J.T. Klepsland
O-L-198317 OLICH2375-16 MK811982
Xanthoparmelia
loxodes
Lecanorales NO, Østfold,
2012, K.A. Lye
O-L-206876 OLICH4133-17 MK812427
Xanthoparmelia
loxodes
Lecanorales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208018 OLICH4190-17 MK812543
Xanthoparmelia
loxodes
Lecanorales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208343 OLICH4272-17 MK812627
OLICH: A reference library of DNA barcodes for Nordic lichens 125

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Xanthoparmelia
protomatrae
Lecanorales NO, Aust-
Agder, 2015,
H. Riddervold
O-L-201574 OLICH3243-16 MK812594
Xanthoparmelia
pulla
Lecanorales NO, Sør-
Trøndelag,
2014, R.
Haugan
O-L-196012 OLICH1857-14 MK812429
Xanthoparmelia
pulla
Lecanorales NO, Østfold,
2016, S. Rui &
E. Timdal
O-L-208019 OLICH4191-17 MK812706
Xanthoparmelia
pulla
Lecanorales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208344 OLICH4273-17 MK812285
Xanthoparmelia
stenophylla
Lecanorales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208338 OLICH4268-17 MK811834
Xanthoparmelia
tinctina
Lecanorales SE, Bohuslän,
2012, E.
Timdal
O-L-184585 OLICH3233-16 MK812507
Xanthoparmelia
tinctina
Lecanorales NO, Østfold,
2017, S. Rui &
E. Timdal
O-L-208333 OLICH4264-17 MK812642
Xanthoparmelia
verruculifera
Lecanorales NO, Oppland,
2015, E.
Timdal
O-L-201346 OLICH2731-16 MK812281
Xanthoria
parietina
Teloschistales NO, Telemark,
2014, E.
Timdal
O-L-195782 OLICH1654-14 MK812349
Xanthoria
parietina
Teloschistales NO, Oslo,
2014, M.
Bendiksby et
al.
O-L-196081 OLICH1897-14 MK812134
Xanthoria
parietina
Teloschistales NO, Østfold,
2012, K.A. Lye
O-L-206852 OLICH4143-17 MK811702
Xylographa
parallela
Trapeliales NO, Finnmark,
2011, E.
Timdal
O-L-170700 OLICH3294-16 MK812221 no lichen
substances
126 Marthinsen G et al

Species Order Voucher Sample ID Process ID GenBank ID Chemistry
Xylographa vitiligo Trapeliales NO, Nord-
Trøndelag,
2014, J.T.
Klepsland
O-L-200273 OLICH3315-16 MK812010
Temporal coverage
Notes: The sampled material was collected in the period 1995–2017, with 90% being after
2010 and with a peak of 532 specimens in 2014 (Fig. 3).
Usage rights
Use license: Open Data Commons Attribution License
Data resources
Data package title: DNA barcodes of Nordic lichens
Resource link: dx.doi.org/10.5883/DS-OLICH
Number of data sets: 1
Data set name: DS-OLICH
Figure 3.
Distribution of year of collection of the specimens in the OLICH dataset
OLICH: A reference library of DNA barcodes for Nordic lichens 127

Download URL: http://www.boldsystems.org/index.php/Public_SearchTerms?
query=DS-OLICH
Data format: dwc, xml, tsv, fasta, trace
Description: The OLICH dataset can be downloaded from the Public Data Portal of
BOLD (www.boldsystems.org) in different formats (data as dwc, xml or tsv, sequences
as fasta and trace files as trace). Alternatively, BOLD users can log in and access the
dataset via the Workbench platform of BOLD. All records are also searchable within
BOLD using the search function of the database.
Depending on future funding, the OLICH project will continue sequencing Nordic
lichens for the BOLD database, with the ultimate (although unrealistic) goal of complete
coverage. Hence, the dataset will be updated in the future with new sequences and
information. The version of the dataset at the time of writing the manuscript is included
as Suppl. materials 1, 2, 3, 4 in the form of two text files for record information as
downloaded from BOLD, one text file with the collecting and identification data in
Darwin Core Standard format (downloaded from BOLD and formatted by the authors)
and of a fasta file containing all sequences as downloaded from BOLD.
It should be noted that, as the BOLD database is not compliant with the Darwin Core
Standard format, the Darwin Core formatted file (dwc) that can be downloaded from
BOLD is not strictly Darwin Core formatted. For a proper Darwin Core formatted file,
see Suppl. material 3.
All records have images and geographical coordinates in BOLD.
Column labels below follow the labels downloaded in the tsv format. Columns with no
content in our dataset are left out in the list below.
Column label Column description
processid Unique identifier for the sample
sampleid Unique identifier for the voucher specimen, i.e. museum catalogue number in either lichen
herbarium or DNA bank at the Natural History Museum, University of Oslo
recordID Entry number in the database
catalognum Same as sampleid; Unique identifier for the voucher specimen, i.e. museum catalogue
number in either lichen herbarium or DNA bank at the Natural History Museum, University
of Oslo
institution_storing The full name of the institution where specimen is stored
phylum_taxID Taxonomic identifier of level Phylum
phylum_name Phylum name
class_taxID Taxonomic identifier of level Class
class_name Class name
128 Marthinsen G et al

order_taxID Taxonomic identifier of level Order
order_name Order name
family_taxID Taxonomic identifier of level Family
family_name Family name
genus_taxID Taxonomic identifier of level Genus
genus_name Genus name
species_taxID Taxonomic identifier of level Species
species_name Species name
identification_provided_by Name of primary individual repsonsible for providing taxonomic identification of the
specimen
identification_method The method(s) used to identify the specimen
identification_reference Authorship of species name
voucher_status Status of the specimen in an accessioning process
tissue_type A brief description of the type of tissue analysed
collectors Name(s) of the individual(s) responsible for collecting the sample in the field
habitat A description of the habitat
Notes Notes regarding species identification of the specimen or other note concerning the
specimen.
lat The geographical latitude (in decimal degrees) of the geographic centre of a location
long The geographical longitude (in decimal degrees) of the geographic centre of a location
coord_accuracy An estimation of the uncertainty of the coordinates given in the lat and long columns
(metres)
elev Elevation of sampling site. Measured in metres relative to sea level.
country Name of country in which the organism was collected
province_state Name of the province (in Norway "fylke") in which the organism was collected
region Name of municipality in which the organism was collected
exactsite Name of exact locality where the organism was collected
image_ids Entry number in the database
image_urls URL(s) from which the image(s) of the organism(s) can be accessed
media_descriptors Term depicting, in this case, side of organism photographed, i.e. dorsal
captions Short description of part of organism photographed
copyright_years The year of the licence declaration for the image
OLICH: A reference library of DNA barcodes for Nordic lichens 129

copyright_licences Type of licence associated with the image
copyright_institutions The primary licence holder's institutional affiliation
photographers The individual responsible for photographing
Additional information
Summary statistics and genetic distances
The dataset contains on average 2.6 records per species, ranging from one (184 species)
to 12 (one species) records per species. Average sequence length was 541 bp (including
sequences that did not include the whole ITS region due to unreadable trace files). The
shortest sequence in the dataset was an incomplete sequence, 229 bp long. The species
with the shortest complete ITS region was Haematomma ocroloeucum with 481 bp. The
longest ITS region was 856 bp from Leptogium cochleatum. However, the two sequences
of the latter species did not align with the others at the start of the sequence and it is
possible that introns in the SSU rDNA are included.
We calculated intra- and interspecific genetic distances for the three genera with most
sampled species and performed barcode gap analyses on those. Distances and key
statistics are found in Table 2.
Genus Ramalina Umbilicaria Cladonia
Number of species in genus in this dataset 9 19 47
Number of records 26 52 122
Average number of records per species 2.9 2.7 2.6
Average intraspecific distances 0–1.3% 0–2.3% 0–3.4%
Interspecific distances 0.4–8.2% 2.5–12.1% 0.2–15.6%
In Ramalina, there was an overlap between intra- and interspecific distances (Table 2, Fig.
4a). The overlap is caused by the one species pair with higher maximum intraspecific
distance than the distance to the nearest neighbour (Fig. 4b); i.e. R. farinacea had an
intraspecific distance of 0.9% and a distance of 0.4% to R. subfarinacea. If this species
pair is excluded from the distance-dataset, a barcode gap will occur at 2% (Fig. 4a).
Table 2.
Key statistics and genetic distances for the three genera with the most sampled species;
Ramalina, Umbilicaria and Cladonia.
130 Marthinsen G et al

a
b
c
d
e
f
Figure 4.
Genetic distances within the three lichen genera with most sampled species in the OLICH
dataset.
a: Frequency distribution of intraspecific distances (blue) and interspecific distances
(orange). A gap between intra- and interspecific distances illustrates the barcode gap in the
genus.
b: Scatterplots representing maximum intraspecific distances versus minimum distance to
closest neighbour for all species with more than one specimen per species, in each of the
three genera.
c: Frequency distribution of intraspecific distances (blue) and interspecific distances
(orange). A gap between intra- and interspecific distances illustrates the barcode gap in the
genus.
d: Scatterplots representing maximum intraspecific distances versus minimum distance to
closest neighbour for all species with more than one specimen per species, in each of the
three genera.
e: Frequency distribution of intraspecific distances (blue) and interspecific distances
(orange). A gap between intra- and interspecific distances illustrates the barcode gap in the
genus.
f: Scatterplots representing maximum intraspecific distances versus minimum distance to
closest neighbour for all species with more than one specimen per species, in each of the
three genera.
OLICH: A reference library of DNA barcodes for Nordic lichens 131

For Umbilicaria, there were no species pairs with higher maximum intraspecific distance
than the distance to the nearest neighbour, but one species had the same maximum
intraspecific distance as the distance to the nearest neighbour, namely U. proboscidea
whose closest neighbour was U. virginis (Fig. 4d). There is thus no barcode gap in our
Umbilicaria dataset, as the intra- and interspecific distances overlaps around 3% (Fig. 4c).
Whereas Ramalina and Umbilicaria, despite the above-mentioned issues, show a tendency
for displaying a barcode gap, Cladonia does not show the same pattern. There is no gap,
but rather overlapping, smooth curves of intra- and interspecific distances in the barplot
(Table 2, Fig. 4e). Most of the species pairs that disrupt the pattern are complexes where
species are known to be sometimes difficult to distinguish with traditional characters and
these are found below the line in Fig. 4f: C. coniocraea/gracilis, C. maxima/ecmocyna and
C. cenotea/subfurcata/crispata/squamosa.
ITS has previously been shown to give poor resolution amongst Cladonia species
(Fontaine et al. 2010, Kotelko and Piercey-Normore 2010, Pino-Bodas et al. 2013) and a
lack of barcode gap has been demonstrated (Kelly et al. 2011). This probably has to do
with their high morphological plasticity (Pino-Bodas et al. 2013) which may have led to
erroneous identifications/taxonomy, but there might also be evolutionary explanations,
such as incomplete lineage sorting (van Velzen et al. 2012).
Observations
From the phylogenetically and BLASTn based quality control, some specific taxonomic
observations were made that are listed below.
Alectoria sarmentosa (Ach.) Ach. ssp. vexillifera (Nyl.) D.Hawksw.: This subspecies is
currently accepted in the Nordic flora (Velmala and Myllys 2011, Nordin et al. 2019). It
differs morphologically from spp. sarmentosa in forming a more prostrate thallus with
broader and more flattened main branches and, ecologically, in growing on the ground or
on rocks mainly in coastal and alpine areas. Subspecies sarmentosa grows mainly on
trunks and branches of conifers in boreal forests. Our sequence of spp. vexillifera
(OLICH1478) is nested within our four sequences of spp. sarmentosa, however, which
does not support the recognition of the subspecies (Fig. 5).
Brodoa oroarctica (Krog) Goward : The species differs from B. atrofusca (Schaer.)
Goward in the shape of the thallus and lobes and in the lack of physodic acid in most of the
thallus except for in some lobe tips (Krog 1974). The former species is circumpolar arctic-
alpine, with southernmost occurrences in Europe in the southern Scandinavian mountains,
whereas the latter is a European species known mainly from the Alps and Pyrenées and
with scattered occurrences in Northern Europe. Our four barcode sequences of B.
oroarctica are however nested within the four sequences of B. atrofusca (Fig. 6), and the
complex seems to be in need of revision.
132 Marthinsen G et al

Bryobilimbia fissuriseda (Poelt) Timdal, Marthinsen & Rui, comb. nov.
Mycobank: MB 830330
Basionym: Lecidea fissuriseda Poelt, Mitteilungen der Botanischen Staatssammlung
München 4: 181, 1961. ≡ Mycobilimbia fissuriseda (Poelt) Poelt & Hafellner, Herzogia 8:
56, 1989. Type: Switzerland, Berner Oberland, Gipfelgrader Niesen, Nordhänge, alt. 2300
m, 1955, Poelt B55189 (M!, holotype).
Figure 5.
Maximum likelihood tree based on the OLICH sequences of Alectoria sarmentosa.
Numbers above branches and thick lines indicate bootstrap support ≥ 70%. Bold font
indicates focus taxon and grey font the outgroup. The two-letter ISO country codes are
included. This applies for all phylogram figures (Figs 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15).
Figure 6.
Maximum likelihood tree of Brodoa atrofusca and B. oroarctica based on all available ITS
sequences.
OLICH: A reference library of DNA barcodes for Nordic lichens 133

This rarely collected species was described from the Alps in the large, collective genus
Lecidea Ach. (Poelt 1961). It was later transferred to the segregate genus Mycobilimbia
Rehm by Poelt & Hafellner (in Hafellner 1989). Partly due to the presence of characteristic
blue, K+ green granules in the hymenium and the hypothecium, Poelt (1961) and Timdal
(1992) assumed it was closely related to M. hypnorum (Lib.) Kalb & Hafellner (syn. Lecidea
fusca (Schaer.) Th. Fr.). Fryday et al. (2014), based on a molecular study, excluded M.
hypnorum from Mycobilimbia and placed it, with four additional species, in the new genus
Bryobilimbia Fryday et al. Mycobilimbia fissuriseda was not included in that study, but our
phylogenetic tree, based on the DNA barcodes (Fig. 7), shows that it is, in fact, closely
related to B. diapensiae and B. hypnorum and it is hence here transferred to Bryobilimbia.
Figure 7.
Maximum likelihood tree of Bryobilimbia and Mycobilimbia based on all available ITS
sequences. The presumedly related genera Clauzadea and Romjularia are included and
Farnoldia and Lecidea are used as the outgroup.
134 Marthinsen G et al

Bryoria capillaris (Ach.) Brodo & D.Hawksw.: The species is still accepted in the Nordic
checklist (Nordin et al. 2019), despite several recent molecular phylogenies (Myllys et al.
2011, Velmala et al. 2014, Myllys et al. 2016, Boluda et al. 2019) showing it nested within
B. implexa (Hoffm.) Brodo & D. Hawksw. The OLICH dataset shows the same topography
(Fig. 8): B. capillaris nested within B. implexa. However, all our specimens of the B.
capillaris/implexa complex were identified by morphology prior to TLC analysis and the
correlation holds: all pale specimens (B. capillaris) contain alectorialic acid (plus
accessories) and barbatolic acid, whereas no dark specimens (B. implexa) do.
Fumarprotocetraric and protocetraric acids occur variably in both species. Either the DNA
barcode marker and the other markers used in recent phylogenetic studies do not
distinguish between two species which may be recognised by the correlation of thallus
colour and secondary chemistry or the two "species" are conspecific and their
morphological/chemical difference is caused by other factors. For example, the presence/
absence of an additional symbiont (basidiomycete yeast) were found likely to trigger the
production of vulpinic acid in the cortex of Bryoria fremontii/tortuosa by Spribille et al.
(2016).
Cetrelia cetrarioides (Duby) W.L. Culb. & C.F. Culb. and C. monachorum (Zahlbr.)
W.L. Culb. & C.F. Culb.: The three chemotypes of Cetrelia olivetorum (Nyl.) W.L. Culb. &
C.F. Culb. s. lat., currently recognised in the Nordic countries (Bjelland et al. 1997,
Obermayer and Mayrhofer 2007, Nordin et al. 2019), were recently shown to represent
three distinct species in the phylogenetic analysis by Mark et al. (2019). The diagnostic,
main compounds of those species are imbricaric acid in C. monachorum, olivetoric acid in
C. olivetorum s. str. and perlatolic acid in C. cetrarioides (Mark et al. 2019). The seven
specimens sequenced in the OLICH project fell into the C. cetrarioides and C.
monachorum clades (Fig. 9) and the TLC analyses of the sequenced specimens showed
the chemotypes were in accordance with the species identifications (Table 1).
Figure 8.
Maximum likelihood tree of Bryoria capillaris and B. implexa based on the OLICH
sequences.
OLICH: A reference library of DNA barcodes for Nordic lichens 135

Hypotrachyna afrorevoluta (Krog & Swinscow) Krog & Swinscow : The species is
morphologically and chemically quite similar to H. revoluta (Flörke) Hale and the OLICH
sequences (Fig. 10) confirm the presence of the two species in the Nordic countries. See
Elix and Thell (2011) for the subtile morphological differences between the species.
Figure 9.
Maximum likelihood tree of the OLICH sequences of Cetrelia, with selected ITS sequences
from Mark et al. (2019).
Figure 10.
Maximum likelihood tree of the OLICH sequences of Hypotrachyna afrorevolulta and H.
revoluta, with all available ITS sequences in GenBank.
136 Marthinsen G et al

Nephroma tangeriense (Maheu & A.Gillet) Zahlbr.: The species was recently reported
as new to the Nordic countries (Klepsland 2013) from two collections from Rogaland
county. The OLICH project has sequenced one of those (OLICH2019), as well as four
additional collections from Hordaland and Vest-Agder counties and confirms its identity and
distinction from N. laevigatum Ach. (Fig. 11).
Peltigera “ neocanina”: BLASTn search revealed the identity of OLICH1878, which was
originally identified as P. canina (L.) Willd., as P. “neocanina”. This is still an invalid name
despite the taxon, which must be regarded as a morphologically cryptic species, has been
known since the phylogenetic study of Peltigera by Miadlikowska and Lutzoni (2000).
Jüriado et al. (2017) showed its presence in Estonia and Magain et al. (2018) in Norway.
Pseudocyphellaria citrina (Gyeln.) Lücking, Moncada & S. Stenroos : The
Pseudocyphellaria species with yellow soralia, occurring in the Nordic countries, is
currently named P. crocata (L.) Vain. (Nordin et al. 2019). Lücking et al. (2017b) showed
that P. crocata was heterogeneous and split it into 13 species. European material was not
included in that study and it remained unknown which species occurs in our area. The
sequences obtained from three Norwegian specimens in the OLICH project all belong in
the P. citrina clade of Lücking et al. (2017b) (Fig. 12).
Psoroma cinnamomeum Malme : The specimen from which OLICH702 was generated
was originally identified as Psoroma tenue Henssen var. boreale Henssen, but BLASTn
search against GenBank indicated its identity as P. cinnamomeum. A phylogenetic
analysis, based on most of the ITS sequences of Psoroma in GenBank (including those
used by Park et al. 2018), supported this identification as OLICH702 and came out in a
strongly supported clade with the six other specimens of P. cinnamomeum, two of which
Figure 11.
Maximum likelihood tree based on the OLICH sequences of Nephroma laevigatum and N.
tangeriense, with selected ITS sequences from GenBank.
OLICH: A reference library of DNA barcodes for Nordic lichens 137

are from southernmost Chile and four from Svalbard (Fig. 13). The clade was the sister of
the strongly supported clade of the two specimens of P. tenue in GenBank, both from
Antarctica. Both P. cinnamomeum and P. tenue were described from Tierra del Fuego, var.
boreale from USA, Colorado (cf. Henssen and Renner 1981). OLICH702 contains
porphyrilic acid and two unknown compounds (apparently related to porphyrilic acid due to
the colour and fluorescence on the chromatograms) and hence fits the current concept of
P. tenue var. boreale (cf. Jørgensen 2007). Unless molecular studies show that there are
two morphologically very similar species in the Northern Hemisphere, both containing
porphyrilic acid, we believe P. cinnamomeum is the correct name for the taxon currently
known as P. tenue var. boreale.
Ramalina capitata (Ach.) Nyl. : The species has often been included in R. polymorpha
(Lilj.) Ach. in the past (e.g. Krog and James 1977), but is now usually accepted as distinct
(e.g. Nimis et al. 2018, Nordin et al. 2019). In our phylogeny (Fig. 14), Nordic and western
European material of the two species are mostly well separated, but the four sequences
from Turkey in GenBank are deviant and the complex seems to be in need of further study.
Ramalina subfarinacea (Cromb.) Nyl.: Our two sequences of this species, as well as the
two ITS sequences available in GenBank, are nested in the clade of Ramalina farinacea
(L.) Ach. in our phylogeny (Fig. 15). According to Krog and James (1977), there are subtle
morphological differences between the species, but apparently the major distinguishing
Figure 12.
Maximum likelihood tree of the Pseudocyphellaria crocata complex with the OLICH
sequences added to those of Lücking et al. (2017b).
138 Marthinsen G et al

feature is the ecology: mainly on coastal rock (R. subfarinacea) vs. mainly on bark of
deciduous, inland trees (R. farinacea). More study is needed in order to evaluate the
taxonomic status of R. subfarinacea.
Sticta fuliginoides Magain & Sérus. : BLASTn search of the sequence of an originally
identified Sticta fuliginosa (Hoffm.) Ach. (OLICH2638) showed 99.4–100% similarity with
the five sequences of S. fuliginoides and 90.0–90.2% similarity with the 9 sequences of S.
fuliginosa in the phylogeny of Magain and Serusiaux (2015). This confirms the presence of
S. fuliginoides in the Nordic countries, as recently reported by Ekman et al. (2019), based
on morphological characters.
Figure 13.
Maximum likelihood tree of Psoroma based on all available ITS sequences.
Figure 14.
Maximum likelihood tree of the Ramalina capitata/polymorpha complex, based on all
available ITS sequences.
OLICH: A reference library of DNA barcodes for Nordic lichens 139

Acknowledgements
We would like to thank the many contributors to this project who donated their collections
of lichens to the Natural History Museum, University of Oslo, for the purpose of sequencing
the DNA barcode marker, especially the major contributors Reidar Haugan and Jon T.
Klepsland. Håkon Holien and Tor Tønsberg gave access to material in the herbaria TRH
and BG, respectively. Mika Bendiksby assisted in several ways, for example, in the
arrangements of workshops and field meetings in Bergen, Oslo, Steinkjer and Varanger.
The project was funded by the Norwegian Research Council through the Norwegian
Barcode of Life project and by The Norwegian Species Information Centre through four
projects under the Norwegian Taxonomy Initiative.
Figure 15.
Maximum likelihood tree of the Ramalina farinacea/subfarinacea complex, based on all
available ITS sequences.
140 Marthinsen G et al

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Supplementary materials
Suppl. material 1: OLICH library - Processing details
Authors: Gunnhild Marthinsen, Siri Rui, Einar Timdal
Data type: Record information - processing details on ITS sequences
Brief description: The file includes information about all records in BOLD for the OLICH
library, i.e. the DS-OLICH dataset, at time of writing. It contains data on ITS sequences. The
data are as downloaded from BOLD without further processing.
Download file (492.24 kb)
Suppl. material 2: OLICH library - Specimen details
Authors: Gunnhild Marthinsen, Siri Rui, Einar Timdal
Data type: Record information - specimen data
Brief description: The file includes information about all records in BOLD for the OLICH
library, i.e. the DS-OLICH dataset, at time of writing. It contains collecting and identification
data. The data are as downloaded from BOLD, without further processing.
Download file (492.24 kb)
Suppl. material 3: OLICH library - Specimen details - Darwin Core Standard
Authors: Gunnhild Marthinsen, Siri Rui, Einar Timdal
Data type: Record information - specimen data in Darwin Core Standard format
Brief description: The file includes information about all records in BOLD for the OLICH
library, i.e. the DS-OLICH dataset, at time of writing. It contains collecting and identification
data. The data are downloaded from BOLD and further formatted to the DC standard by the
authors.
Download file (529.67 kb)
OLICH: A reference library of DNA barcodes for Nordic lichens 145

Suppl. material 4: OLICH library - DNA sequences
Authors: Gunnhild Marthinsen, Siri Rui, Einar Timdal
Data type: Genomic data, DNA sequences
Brief description: ITS sequences in fasta format. Each sequence is identified by the BOLD
ProcessID and species name, separated by pipe. The data are as downloaded from BOLD.
Download file (945.98 kb)
146 Marthinsen G et al