![Pachydissus Newman, 1838 and Dymasius J. Thomson, 1864, habitus, dorsal view, and pronotum. 74-P. patricius Holzschuh, 1991 (after Holzschuh [1991], photograph by Luboš Dembický); 75-P. borneoensis sp. n.; 76-P. pullus Holzschuh, 2017; 77-P. murzini sp. n.; 78-P. birmanicus Gardner, 1926 (photograph by Sudhir Singh); 79-P. argentatus Pic, 1923 (photograph by Gérard L. Tavakilian); 80D. querceus Holzschuh, 2015 (after Holzschuh [2015], photograph by Luboš Dembický). 74-75, 77-80-holotypes; 74, 76, 78-79-females; 75, 77, 80-males. Рис. 74-80. Pachydissus Newman, 1838 и Dymasius J. Thomson, 1864, общий вид сверху и переднеспинка. 74-P. patricius Holzschuh, 1991 (по [Holzschuh, 1991], фотография Л. Дембицкого); 75-P. borneoensis sp. n.; 76-P. pullus Holzschuh, 2017; 77-P. murzini sp. n.; 78-P. birmanicus Gardner, 1926 (фотография С. Сингха); 79-P. argentatus Pic, 1923 (фотография Ж. Тавакиляна); 80-D. querceus Holzschuh, 2015 (по [Holzschuh, 2015], фотография Л. Дембицкого). 74-75, 77-80-голотипы; 74, 76, 78-79-самки; 75, 77, 80-самцы.](https://www.researchgate.net/publication/330475550/figure/fig1/AS:716418584678406@1547818909895/80-Pachydissus-Newman-1838-and-Dymasius-J-Thomson-1864-habitus-dorsal-view-and_Q320.jpg)
![Dymasius J. Thomson, 1864, habitus, dorsal view. 137-D. macilentus (Pascoe, 1859); 138-139-D. strigosus J. Thomson, 1864; 140-D. tatianae sp. n.; 141-D. indigus Holzschuh, 2008; 142-D. nodifer Holzschuh, 2005. 137, 140-142-holotypes; 137-138, 140, 142-males; 139, 141-females; 141-142-after Holzschuh [2005, 2008], photographs by Luboš Dembický. Рис. 137-142. Dymasius J. Thomson, 1864, общий вид сверху. 137-D. macilentus (Pascoe, 1859); 138-139-D. strigosus J. Thomson, 1864; 140-D. tatianae sp. n.; 141-D. indigus Holzschuh, 2008; 142-D. nodifer Holzschuh, 2005. 137, 140-142-голотипы; 137-138, 140, 142-самцы; 139, 141-самки; 141-142-по [Holzschuh, 2005, 2008], фотографии Л. Дембицкого.](https://www.researchgate.net/publication/330475550/figure/fig3/AS:716418588880904@1547818910333/142-Dymasius-J-Thomson-1864-habitus-dorsal-view-137-D-macilentus-Pascoe-1859_Q320.jpg)
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CAUCASIAN ENTOMOLOGICAL BULLETIN
РОССИЙСКАЯ АКАДЕМИЯ НАУК
Южный научный центр
RUSSIAN ACADEMY OF SCIENCES
Southern Scientific Centre
Том 14. Вып. 2
Vol. 14. No. 2
Ростов-на-Дону
2018
Кавказский энтомол. бюллетень 14(2): 197–246 © CAUCASIAN ENTOMOLOGICAL BULL. 2018
DOI: 10.23885/181433262018142-197246
e longicorn beetle tribe Cerambycini Latreille, 1802
(Coleoptera: Cerambycidae: Cerambycinae) in the fauna of Asia.
4. New or little-known taxa, mainly from Indochina and Borneo,
with reviews or annotated checklists of species of some genera
Жуки-дровосеки трибы Cerambycini Latreille, 1802
(Coleoptera: Cerambycidae: Cerambycinae) фауны Азии.
4. Новые и малоизвестные таксоны,
преимущественно из Индокитая и Борнео,
с обзорами или аннотированными списками видов некоторых родов
© A.I. Miroshnikov1, 2
© А.И. Мирошников1, 2
1Russian Entomological Society, Krasnodar, Russia. E-mail: miroshnikov-ai@yandex.ru
2Sochi National Park, Moskovskaya str., 21, Sochi, Krasnodar Region 354002 Russia
1Русское энтомологическое общество, Краснодар, Россия
2Сочинский национальный парк, ул. Московская, 21, Сочи, Краснодарский край 354002 Россия
Key words: Coleoptera, Cerambycidae, Cerambycini, reviews of genera, annotated checklists of species, new species, new
combinations, new synonymy, Southeastern Asia.
Ключевые слова: Coleoptera, Cerambycidae, Cerambycini, обзоры родов, аннотированные списки видов, новые
виды, новые комбинации, новая синонимия, Юго-Восточная Азия.
Abstract. Full generic statuses of Plavichydissus Pic,
1946, stat. rest., Laomargites Pic, 1923, stat. rest. and
Lamellocerambyx Pic, 1923,stat.rest. are restored. Reviews
of these genera, as well as keys to species of the former two
are given. Annotated checklists of the Asian species of the
genera Pachydissus Newman, 1838 and Margites Gahan,
1891, as well as of all species of Diorthus Gahan, 1891 are
presented. e following new species are described and
new specific combinations established: Plavichydissus
grossepunctatus (Gressitt et Rondon, 1970), comb. n.,
P. irinae sp. n. (Vietnam), P. aggregatus (Holzschuh,
1999), comb. n., P. sulcicollis (Gahan, 1893), comb. n.,
P. myanmarensissp. n. (Myanmar), P. makarovisp. n.
(ailand), P. nataliaesp. n. (Vietnam), P. decipiens
(Holzschuh, 1989), comb. n., P. penangensissp. n.
(Western Malaysia), P.sodalis (Holzschuh, 1999),comb.n.,
P. dembickyisp. n. (Western Malaysia), Pachydissus
murzinisp. n. (Yunnan, China), P. borneoensissp. n.
(Eastern Malaysia), Laomargites fedorenkoisp. n.
(Vietnam), Dymasius tatianaesp. n. (Eastern Malaysia),
D.solodovnikovisp.n. (ailand), D.barclayisp.n. (Western
Malaysia), Zatrephus jaklisp.n. (Java, Indonesia), Diorthus
kabakovisp. n. (Afghanistan), Tapinolachnus uniformis
(Pic, 1933), comb. n., T. xyliae (Fisher, 1940), comb. n.
e following specific combinations are restored:
Plavichydissus semiplicatus (Pic, 1926), comb. rest.,
P.rufipennis (Pic, 1923),comb.rest., Laomargites singularis
Pic, 1923, comb. rest. and Lamellocerambyx laosensis
Pic, 1923, comb. rest. e synonymization of the genus
Diorthus with the genus Tapinolachnus J.omson, 1865
is confirmed as being wrong. e following new synonymy
is established: Tapinolachnus = Mimoderolus (Aeolesthes
subgen.) Pic, 1933, syn. n. (non syn. pro Derolus Gahan,
1891). Dymasius strigosus J. omson, 1864, sp. rest. is
resurrected from the synonymy with Dymasius macilentus
(Pascoe, 1859). e genus Derolydnus Hüdepohl, 1989 is
reported from Indochina for the first time. New records of
a number of species from other genera are given as well,
thus one way or another extending their known distribution
areas, sometimes very significantly so. e lectotypes of
Margites modicus Gahan, 1906, Diorthus sericeus Gardner,
1939 and Tapinolachnus xyliae (Fisher, 1940), comb. n.
are designated. Abundant pictures of the species studied,
including numerous type specimens, are provided.
Резюме. Восстановлены родовые статусы
Plavichydissus Pic, 1946, stat. rest., Laomargites Pic,
1923,stat. rest. и Lamellocerambyx Pic, 1923, stat.rest.
Даны обзоры этих родов и предложены таблицы
для определения видов двух первых из них.
Представлены аннотированные списки азиатских
видов родов Pachydissus Newman, 1838 и Margites
Gahan, 1891, а также всех видов рода Diorthus
Gahan, 1891. Описаны следующие новые виды и
установлены новые комбинации видовых названий:
Plavichydissus grossepunctatus (Gressitt et Rondon,
1970),comb.n., P.irinaesp. n. (Вьетнам), P. aggregatus
(Holzschuh, 1999), comb. n., P. sulcicollis (Gahan,
1893), comb. n., P. myanmarensissp. n. (Мьянма),
P. makarovisp. n. (Таиланд), P. nataliaesp. n.
(Вьетнам), P. decipiens (Holzschuh, 1989), comb. n.,
P. penangensissp. n. (Западная Малайзия), P. sodalis
(Holzschuh, 1999), comb. n., P. dembickyisp. n.
(Западная Малайзия), Pachydissus murzinisp. n.
(Юньнань, Китай), P. borneoensissp. n. (Восточная
Малайзия), Laomargites fedorenkoisp. n. (Вьетнам),
Dymasius tatianaesp. n. (Восточная Малайзия),
D. solodovnikovisp. n. (Таиланд), D. barclayisp. n.
(Западная Малайзия), Zatrephus jaklisp. n. (Ява,
Индонезия), Diorthus kabakovisp. n. (Афганистан),
Tapinolachnus uniformis (Pic, 1933), comb. n., T. xyliae
(Fisher, 1940), comb. n. Восстановлены комбинации
следующих видовых названий: Plavichydissus
semiplicatus (Pic, 1926), comb. rest., P. rufipennis
(Pic, 1923), comb. rest., Laomargites singularis Pic,
1923, comb. rest. и Lamellocerambyx laosensis Pic,
1923, comb. rest. Подтверждена ошибочность
синонимизации рода Diorthus с родом Tapinolachnus
J. omson, 1865. Установлена следующая новая
синонимия: Tapinolachnus= Mimoderolus (Aeolesthes
subgen.) Pic, 1933, syn. n. (non syn. pro Derolus Gahan,
1891). Восстановлен из синонимов Dymasius strigosus
J. omson, 1864, sp. rest., non syn. pro Dymasius
macilentus (Pascoe, 1859). Род Derolydnus Hüdepohl, 1989
впервые приведен для Индокитая. Отмечены также
новые находки целого ряда видов из других родов,
расширяющие их ареалы. Обозначены лектотипы
Margites modicus Gahan, 1906, Diorthus sericeus Gardner,
1939 и Tapinolachnus xyliae (Fisher, 1940), comb. n.
Представлено большое количество иллюстраций
исследуемых видов, в том числе многих типовых
экземпляров.
Introduction
In the initial publication of this series [Miroshnikov,
2017], some preliminary remarks concerning the
taxonomically confused genera (or their representatives)
Pachydissus Newman, 1838, Margites Gahan, 1891 and
Plavichydissus Pic, 1946 were made. e present paper
provides a review of the latter genus, with the restoration
of its generic status, and annotated checklists of the Asian
species are given for both former genera. Primary generic
statuses are also substantiated here for Laomargites Pic,
1923 and Lamellocerambyx Pic, 1923 (with their reviews
presented as well), considered by some researchers as
subgenera of the genera Margites and Diorthus Gahan,
1891, respectively. e fallacy of the synonymization of
the latter genus with the genus Tapinolachnus J.omson,
1865 is confirmed, and annotated checklists of Diorthus
and Tapinolachnus species are given together with some
synonymies.
Besides this, 14new species of the genera Plavichydissus
(6species), Pachydissus (2species), Laomargites (1species),
Dymasius J. omson, 1864 (3species), Zatrephus Pascoe,
1857 (1 species) and Diorthus (1 species) are described
below. New data on the distribution of many other species
from various genera are given, to some extent expanding
their distribution areas, as well as other new information
is presented. e previously expressed deep doubt
[Miroshnikov, 2017] concerning the synonymy Dymasius
macilentus = Dymasius strigosus which has been in use
until recently is substantiated, the species status of the
latter taxon being resurrected.
e material treated in this work belongs to the
following institutional and private collections:
BM – Bishop Museum (Honolulu, USA);
BMNH – Natural History Museum (London, United
Kingdom);
IRSN – Institut Royal de Sciences naturelles de
Belgique (Bruxelles, Belgium);
MNHN – Muséum national d’Histoire naturelle
(Paris, France);
NFIC – National Forest Insect Collection, Forest
Research Institute (Dehradun, India);
NHMD – Natural History Museum of Denmark,
University of Copenhagen (Copenhagen, Denmark);
NHRS – Swedish Museum of Natural History
(Stockholm, Sweden);
ZMMU – Zoological Museum of the Moscow State
University (Moscow, Russia);
ZIN – Zoological Institute of the Russian Academy of
Sciences (StPetersburg, Russia);
ZMUK – Zoologisches Museum der Universität (Kiel,
Germany);
cAM – collection of Alexandr Miroshnikov
(Krasnodar, Russia);
cCH – collection of Carolus Holzschuh (Villach,
Austria);
cLD – collection of Luboš Dembický (Brno, Czech
Republic);
cSM – collection of Sergey Murzin (Moscow, Russia).
Tribe Cerambycini Latreille, 1802
Genus Plavichydissus Pic, 1946, stat. rest.
Plavichydissus Pic, 1946: 107; Gressitt, Rondon, 1970: 71
(Pachydissus subgen.); Miroshnikov, 2017: 223 (preliminary
remarks).
Type species: Pachydissus semiplicatus Pic, 1926.
Diagnosis. is genus, which some researchers
consider as a subgenus of the genus Pachydissus or species
of which have been described in the genus Margites, differs
clearly at least from all Asian representatives of both genera
(see Remarks below) in the distinctive sculpture of the
pronotum, the pattern of the dorsal setation, the somewhat
peculiar sculpture of the elytra, as well as in some other
traits indicated below.
When detailing the structure of
Plavichydissusstat. rest., the following features must be
noted as being characteristic of this genus: head short,
with more or less well developed antennal tubercles;
eyes large, strongly convex or considerably less strongly
developed, moderately convex; male antennae in most
species much longer than body, in some representatives
only very clearly or slightly reaching beyond the apex of
elytra; antennomere 1 without cicatrix; antennomere 2
distinctly or very clearly longitudinal (while in Pachydissus,
antennomere 2 distinctly or very clearly transverse, only
sometimes barely longitudinal or subequal in length and
198 A.I. Miroshnikov
width); male antennomeres 3 and 4 or 3–5 one way or
another broadened towards or near apex, but cannot be
inflated in apical part as in males of almost all species of
Margites; apical external angle of at least antennomeres3–5
in male and female more or less rounded, not drawn
laterad, that of following antennomeres, except for last
one, sometimes obtuse-angled or sharpened, only weakly
or moderately protruding (whereas in some representatives
of Pachydissus, apical external angle of at least
antennomeres3–5 in males more or less strongly sharpened
and strongly drawn laterad, that of several following
antennomeres, except for last one, strongly sharpened and
clearly or strongly drawn laterad, thereby apical external
angle of antennomeres 3 and 4 in females more or less
right, clearly drawn laterad, at least of antennomeres5–7
or 5–8 strongly or very strongly sharpened and more or
less strongly drawn laterad); pronotum with a sharp or
very sharp constriction before base and near apex (while
in Margites, constriction before base of pronotum usually
less sharp), with deep or very deep longitudinal grooves,
resulting in a median, wide or very wide, sometimes very
strong, high elevation flanked either by coarse or very
coarse longitudinal folds (ribs) or such folds, combined
with coarse or very coarse, irregular, sinuous folds, as in
Color plate 4: 27–30, Figs 43–58 (vs neither Pachydissus
nor Margites with a pronotum sculpture similar to the
above, Color plate6:76–77, Figs102–105), as a rule, with
numerous, very long, erect setae (sometimes these setae
partly obliterated and appearing less numerous) and, in
addition, median elevation in some species with dense
or at least numerous, recumbent, light setae forming a
characteristic horseshoe-shaped pattern, as in Figs 43–47
(whereas in Pachydissus and Margites, pronotum only with
individual, long or very long, erect setae, without forming a
pattern of dense, recumbent, light setae similar to the above);
elytra moderately elongated, sometimes more strongly
elongated, without distinct longitudinal ribs, with both a
rough or coarse, sometimes very large, sparse, irregular
and very small, dense, double, very contrastingly differing
puncturation to some degree resembling Imbrius Pascoe,
1866 (whereas in Pachydissus, elytra usually with small,
more or less uniform, dense puncturation, only sometimes,
in addition, with sparse, more or less large, but weakly
expressed punctures generally not forming such a sculpture
as in Plavichydissusstat.rest.; in Margites, elytra with this
or that puncturation, but in general also clearly or at least
somewhat different from that of Plavichydissusstat.rest.);
apical external angle of elytra more or less uniformly
rounded, not clearly expressed, only sometimes obtuse-
angled and well-expressed, apical sutural angle more or
less right, sometimes with only a small, weakly-expressed
denticle (while in Pachydissus, apical external angle of
elytra obtuse-angled or almost right, clearly or sharply
expressed, apical sutural angle in most species drawn into
a more or less long tooth, in some species with a small, but
well-expressed denticle); elytra with a recumbent, more
or less dense, light setation, in one way or another hiding
the puncturation, all along with suberect, this or that way
protruding, but always clearly or sharply prominent setae
and, in addition, with (or sometimes without) long or very
long, more or less numerous or at least separate, erect setae,
as in Figs31–42; thereby recumbent setation of elytra, one
way or another revealing their large puncturation, in most
cases forming a characteristic speckled general surface
(while in Pachydissus and Margites, elytra only with a
recumbent light setation, as in Figs98–101, thereby in the
former usually this or that way irregular, often patterned
and to a varying degree iridescent, but not speckled, as in
Color plate5: 69–73; sometimes only at the very base of
elytra with individual, erect, moderately long, gentle setae;
elytra of Margites not looking speckled either); prosternum
with a heterogeneous, partly rough or moderately coarse
sculpture, with an unclear or distinctly (but not too
sharply) expressed transverse groove in apical part in front
of middle; prosternal process with a weakly expressed,
sometimes very clear tubercle at apex or, conversely, without
such; mesosternal process without tubercle dorsally; legs
moderately long; at least profemora, especially on ventral
side, with a rough, very dense and confluent, partly rugose
puncturation or with an even coarser sculpture; meso-
and metafemora usually with a less coarse sculpture,
but sometimes with a sculpture more or less similar to
that of profemora, especially on mesofemora (while in
Pachydissus, femora with a small, dense or very dense,
partly or predominantly rugose puncturation, usually only
somewhat sharper on profemora, which sometimes, in
addition, with transverse, more or less gentle wrinkles);
tibiae with a very clear or less distinct, sometimes partly
or predominantly poorly expressed, but nonetheless
visible carina along each side (while in Pachydissus, tibiae
without carina); metatarsomere 1 noticeably or clearly
shorter than metatarsomeres2 and3 combined (whereas
in Pachydissus, metatarsomere 1 longer than or subequal
to metatarsomeres 2 and 3 combined, only sometimes
barely shorter than both); body length 10.6–28.3 mm,
thereby in the vast majority of species up to 20mm (while
in Pachydissus, body length 18.7–34 mm, thereby in most
species not less than 23 mm).
By the combination of the above features,
Plavichydissusstat.rest. differs not only from Pachydissus
and Margites, but also from all other similar genera of the
tribe.
Remarks. Taking into account the features of the
distribution of the genus Plavichydissus stat. rest. (see
below), it seemed to me expedient to show in detail its
differences only from the Asian representatives of the
genera Pachydissus and Margites. However, a part of the
differences discussed above, at least in the sculpture of the
pronotum and elytra, the elytral setation and some other
details of the structure, also belong to the species of both
latter genera, distributed outside of Asia (including the
type species of the genus Pachydissus, P.sericus Newman,
1838, and other Australian congeners). In addition, without
a diagnostic re-evaluation of the genus Pachydissus
as a whole, the necessity of which I noted recently
[Miroshnikov, 2017], no more extensive diagnosis of the
genus Plavichydissusstat.rest. is presently warranted.
Composition. e genus includes 13species, six of
which are described as new.
Distribution. Southern Asia (continental part),
Indochina, including Malay Peninsula; very likely also
southern China.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 199
Plavichydissus semiplicatus (Pic, 1926), comb. rest.
(Color plate1: 1, 4; Figs 204, 205)
Pachydissus semiplicatus Pic, 1926a: 23 (“Tonkin”). Type
locality: Northern Vietnam, Hoa Binh Province (according to the
original description and the labels of the syntypes). Plavilstshikov,
1931: 84.
Plavichydissus semiplicatus: Pic, 1946: 107, 108; Miroshnikov,
2017: 223 (preliminary combination).
Pachydissus (Plavichydissus) semiplicatus: Gressitt, Rondon,
1970: 71.
Material. 1♂, syntype (MNHN) (photograph; Color plate 1:1),
“Tonkin, Hoa Binh”, “semiplicatus n. sp.”, “Type”, “Plavichydissus n. g.”,
“Museum Paris, Coll. M. Pic”, “Holotype” (incorrect label) (Fig. 204); 1♀,
syntype (MNHN) (photograph; Color plate 1: 4), Vietnam, “Hoa Binh”,
“Type”, “Museum Paris, Coll. M.Pic”, “Allotype” (incorrect label) (Fig.205).
Morphological notes. Body length of male and
female syntypes 24.2 or 23.1mm, respectively (Dr.Gérard
L. Tavakilian, personal communication).
Distribution. Vietnam.
Plavichydissus grossepunctatus
(Gressitt et Rondon, 1970), comb. n.
(Color plate1: 2, 5; Figs 31, 43, 44, 59, 206, 207)
Pachydissus (Plavichydissus) grossepunctatus Gressitt et
Rondon, 1970: 71. Type locality: Laos, Borikhane Province,
Pakkading (according to the original description and the label of
the holotype). Hua, 1984: 80.
Plavichydissus grossepunctatus: Miroshnikov, 2017: 223
(preliminary combination).
Material. 1♂, holotype (BM) (Color plate 1: 2), “Laos: Borikhane
Prov., Pakkading, 18.III.1965” (sic, should read “18.III.1963”), “Pakkading,
18.3.[19]63” (handwritten), “J.A.Rondon Collection Bishop Mus.”, “Holotype
Pachydissus grossepunctatus Gressitt & Rondon”, “8293” (Fig. 206); 1♀,
paratype (BM) (Color plate 1: 5), “Laos: Borikhane Prov., Pakkading”,
“Pakkading, 26.5.[19]63” (handwritten), “J.A. Rondon Collection Bishop
Mus.”, “Allotype Pachydissus grossepunctatus Gressitt et Rondon”, “8293”
(Fig. 207); 1♀, paratype (BM), “Laos: Sedone Province, Pakse”, “Pakse,
31.3.[19]65”, (handwritten), “J.A .Rondon Collection Bishop Mus.”, “Paratype
Pachydissus grossepunctatus Gressitt et Rondon”.
Morphological notes. Body length 18.3–21.8 mm,
humeral width 4.45–5.6mm, thereby the holotype is the
largest, while the “allotype” is the smallest.
Distribution. Laos.
Plavichydissus irinae Miroshnikov, sp. n.
(Color plate2: 7; Figs 32, 45, 60)
Material. Holotype, ♂ (cAM) (Color plate2: 7): Vietnam, Gia Lai
Province, ~55km ENE of Pleiku, 14°17ʹ45ʺN/ 108°26ʹ57ʺE, Kon Ka Kinh
National park, 600m, at light, 8–20.05.2017 (leg. D.Fedorenko).
Diagnosis. Based on male characters, this
new species seems to be especially similar to
P.grossepunctatuscomb.n., but differs clearly by the elytra
being shorter and more strongly narrowed towards apex,
as in Color plate2: 7, the coloration of their integument
and their dense recumbent setation; the generally darker
coloration; the shorter erect setae and the predominantly
smaller and less sharp puncturation of the elytra (discarding
very small puncturation), as in Color plate 2: 7, Fig. 32;
the more strongly elongated several apical antennomeres,
especially the last one, as in Color plate 2: 7; the sparser,
recumbent, light setation and the much more obliterated
sculpture of the median elevation of the pronotum, as in
Fig.45; the generally sharper sculpture of the submentum;
the distinctly broader process of the prosternum, the well-
expressed tubercle near its apex; the coarser sculpture
of the profemora ventrally; the clearly larger body sizes.
Plavichydissus irinae sp. n. can also be compared to
P.semiplicatuscomb.rest., but differs very clearly at least
by the same features of the elytral and antennal structure
as P.grossepunctatuscomb.n., only an even more strong
difference in the puncturation of the elytra, as well as by
the somewhat larger body sizes (cf.Color plate1: 1,2, 4, 5,
Figs31, 43,44).
Description. Male. Body length 28.3 mm, humeral width
7.7 mm. Eyes, almost entirely dorsum, metasternum and
visible sternites, mostly mesosternum and mandibles black (in
P. grossepunctatus comb. n. and P. semiplicatus comb. rest., at
least elytra reddish brown); epipleura brownish red; head ventrally,
apical one-third of prosternum, prosternal process and partly
mesosternum brown-red; mostly antennae and legs combines
black-brown and dark brown tones, partly with red tint.
Head with a distinct median groove between upper lobes
of eyes; antennal tubercles moderately developed; eyes relatively
small, moderately convex; submentum with a heterogeneous,
predominantly rough and coarse sculpture; antennae much longer
than body, nearly reaching the apex of elytra by antennomere7;
length ratio of antennomeres1–11, 31:10: 42: 35: 48: 54: 59: 62:
66: 66 : 107; antennomere 1 with a heterogeneous, partly rough
sculpture; antennomere2 clearly longitudinal.
Pronotum barely transverse, 1.05times as wide as long; base
1.08times as wide as apex; with a much sharper constriction near
apex than in front of base; broadened somewhat angularly at the
middle; on disc with a very wide, barely convex, median elevation,
sparsely and more or less roughly punctured mainly near lateral
margins and apex; lateral to elevation with a sharply expressed
longitudinal fragment of sculpture formed by very sinuous coarse,
partly very short, transverse and partly strongly shiny folds; lateral
to this fragment with separate, longitudinal, coarse folds.
Scutellum triangular, with an unclear sculpture.
Elytra very distinctly narrowed towards apex, 2.4times as long
as humeral width (in male holotype of P.grossepunctatuscomb.n.
and male syntype of P.semiplicatuscomb.rest. 2.6 or 2.58times,
respectively); with both a more or less rough sparse and very small
dense puncturation; apical external angle rounded, sutural angle
nearly right.
Prosternum with heterogeneous, rough, predominantly
transverse folds in apical part; prosternal process rather wide
between coxae, with a well-expressed apical tubercle; mesosternal
process between coxae clearly wider than prosternal process,
without tubercle dorsally; metasternum and sternites with a small,
clear, dense puncturation; metasternum with a well-expressed
median groove; last (visible) sternite truncate at apex; last (visible)
tergite widely rounded apically.
Legs moderately long; profemora ventrally, predominantly in
basal part with a very coarse sculpture; all tibiae with a distinct
carina along each side; metatarsomere 1 noticeably shorter than
metatarsomeres2 and3 combined.
Recumbent setation of dorsum, prosternum, partly
mesosternum, antennae and legs golden-yellow and yellow, those
of remaining parts yellowish and yellowish grey (recumbent
setation of elytra silver-grey in P. grossepunctatus comb. n.
and P. semiplicatuscomb. rest.); recumbent moderately dense
setae on median elevation of pronotum forming a characteristic
horseshoe-shaped pattern, as in Fig.45; head, pronotum and elytra
with moderately long, erect, sparse, but numerous, light setae;
elytra, in addition, with numerous, suberect, short, light setae;
antennae with long light setae predominantly on both inner and
ventral sides, more numerous on basal antennomeres.
Etymology. I am pleased to dedicate this new species
to Irina, my elder daughter.
Distribution. Vietnam.
200 A.I. Miroshnikov
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) Color plate 1
Figs 1–6. Plavichydissus Pic, 1946, stat. rest., habitus, dorsal view.
1, 4 – P. semiplicatus (Pic, 1926),comb.rest. (photographs by Gérard L. Tavakilian); 2, 5 – P. grossepunctatus (Gressitt et Rondon),comb.n.; 3,6–
P.aggregatus (Holzschuh, 1999),comb. n. (6– after Holzschuh[1999], photograph by Luboš Dembický). 1, 4– syntypes; 2, 6 – holotypes; 5 – paratype;
1–2–males;3–6– females.
Рис. 1–6. Plavichydissus Pic, 1946, stat. rest., общий вид сверху.
1, 4 – P. semiplicatus (Pic, 1926), comb. rest. (фотографии Ж. Тавакиляна); 2, 5 – P. grossepunctatus (Gressitt et Rondon), comb. n.; 3, 6 – P. aggregatus
(Holzschuh, 1999), comb. n. (6 – по [Holzschuh, 1999], фотография Л.Дембицкого). 1, 4– синтипы; 2, 6– голотипы; 5– паратип; 1–2–самцы;3–6– самки.
Color plate 2 e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae)
Figs 7–12. Plavichydissus Pic, 1946, stat. rest., habitus, dorsal view.
7 – P. irinae sp.n.; 8–9 – P. sulcicollis (Gahan, 1893), comb. n.; 10–11– P. myanmarensis sp.n.; 12 – P. nataliae sp.n. 7–8, 10, 12– holotypes; 11–
paratype; 7–male;8–12– females.
Рис. 7–12. Plavichydissus Pic, 1946, stat. rest., общий вид сверху.
7 – P. irinae sp.n.; 8–9 – P. sulcicollis (Gahan, 1893), comb. n.; 10–11– P. myanmarensis sp.n.; 12 – P. nataliae sp.n. 7–8, 10, 12– голотипы; 11–
паратип; 7–самец;8–12– самки.
Plavichydissus aggregatus (Holzschuh, 1999), comb. n.
(Color plate1: 3, 6; Figs 33, 46, 47, 61)
Margites aggregatus Holzschuh, 1999: 21. Type locality:
Southern Vietnam, 40km NW of An Khe, Buon Luoi, 14°10ʹN/
108°30ʹE, 620–750m (according to the original description). Nga
et al., 2014: 435.
Plavichydissus aggregatus: Miroshnikov, 2017: 223
(preliminary combination).
Material. 1♀, holotype (cCH) (photograph; Color plate1:6).
Morphological notes. Body length 19.4 mm
[Holzschuh, 1999].
e ZIN collection contains a female (Color plate1:3)
very similar to the holotype. In addition, it was collected
in the type locality of P. aggregatus (Vietnam, Gia Lai
Province, Buon Luoi, 29.04.1995, leg. Gorochov). I have
preliminarly attributed the female in question to this
species, albeit it differs from the holotype by the somewhat
peculiar sculpture of the pronotal disc, the sparser and
less strongly developed, recumbent, light setation at its
median elevation and near apex in the middle, as in Fig.47
(cf.Fig.46), as well as by the somewhat smaller size of the
largest sparse punctures and the sparser, recumbent, light
setation of the elytra, which generaly more weakly masks
their puncturation. erefore it seems to me appropriate to
give a description of this female.
Body length 17.4 mm, humeral width 4.3 mm. Coloration
of integument mainly red-brown; eyes, partly mandibles and
pronotal disc black.
Head with longitudinal folds between upper lobes of eyes;
antennal tubercles well-developed; eyes medium-sized, mo derately
convex; submentum with a heterogeneous, predominantly rough
and coarse sculpture; antennae reaching beyond apex of elytra by
last antennomere; length ratio of antennomeres1–11, 32:10: 39:
28: 38: 39: 41: 36: 34: 30: 32; antennomere1 w ith a heterogeneous,
partly rough sculpture; antennomere2 clearly longitudinal.
Pronotum subequal in length and width; base 1.14times as
wide as apex; with a sharp constriction both in front of base and
near apex; on disc with a wide, barely convex, median elevation,
very sparsely and roughly punctured mainly near lateral margins
and apex; lateral to elevation with coarse longitudinal folds,
thereby the nearest of them somewhat sinuous, in basal part
branching into two folds with a very narrow gap between them (in
holotype, fold nearest to median elevation, also branching into two
folds, but with a wider gap between them).
Scutellum triangular, with an unclear sculpture.
Elytra predominantly nearly parallel-sided starting from
base, 2.62times as long as humeral width; with both a more or less
rough sparse and very small dense puncturation; apical external
angle rounded, sutural angle obtuse.
Prosternum in apical part with rough transverse folds;
prosternal process moderately wide between coxae, with a well-
expressed apical tubercle; mesosternal process between coxae
clearly wider than prosternal process, without tubercle dorsally;
metasternum and sternites with a small, clear, dense puncturation;
metasternum with a clear median groove; both last (visible)
sternite and tergite widely rounded apically.
Legs moderately long; profemora ventrally mostly with a
coarse sculpture; all tibiae with a poorly visible carina along each
side; metatarsomere 1 noticeably shorter than metatarsomeres2
and3 combined.
Recumbent setation of dorsum (except for anterior part of
head), prosternum, partly mesosternum, antennae and legs golden-
yellow and yellow, those of remaining parts mainly yellowish
and greyish; recumbent setae on median elevation of pronotum
forming a characteristic horseshoe-shaped pattern, as in Fig. 47;
head, pronotum and elytra with long and very long, erect, sparse,
but numerous (except for head), light setae; elytra, in addition,
with numerous, suberect, short, light setae; most antennomeres
with long, sparse, light setae predominantly on both inner and
ventral sides.
Distribution. Vietnam.
Plavichydissus sulcicollis (Gahan, 1893), comb. n.
(Color plate2: 8, 9; Figs 36, 37, 49, 50, 56, 62, 208)
Margites sulcicollis Gahan, 1893: 378. Type locality: Burma
(now Myanmar), Paungdé (according to the original description
and the label of the holotype). Gahan, 1906: 138; Aurivillius, 1912:
59; Plavilstshikov, 1931: 89; Holzschuh, 1999: 21.
Plavichydissus sulcicollis: Miroshnikov, 2017: 223
(preliminary combination).
Material. 1♀, holotype, by monotypy (BMNH) (Color plate 2: 8),
“Burma, 91–100”, “Paungdé”, “Margites sulcicollis Gahan, Type”, “Type”
(Fig.208); 1♀ (BMNH) (Color plate2:9), Myanmar, “Paungdé”, “Pachydissus
(Margites) sulcicollis Gahan”, “Andrewes Bequest, B.M. 1922–221”.
Morphological notes. Body length 13.7–14.7 mm,
humeral width 3.7–4mm, thereby holotype smallest.
Pronotum barely transverse, 1.01–1.05times as wide
as long; base 1.23–1.24times as wide as apex; with a very
sharp constriction both in front of base and near apex;
broadened angularly at the middle, as in Figs49,50.
Distribution. Myanmar; has also been recorded from
India [Plavilstshikov, 1931].
Plavichydissus myanmarensis Miroshnikov, sp. n.
(Color plate2: 10, 11; Figs 34, 35, 51, 52, 63)
Margites sulcicollis (non Gahan, 1893): Gahan, 1906: 138
(partim, “Burma, North Chin Hills”).
Material. Holotype, ♀ (BMNH) (Color plate2:10): “Burma. N[orth].
Chin Hills. 95–28”. Paratype: 1♀ (BMNH) (Color plate 2: 11), Myanmar,
“Myittha, F.W.T. Bodeker” (upperside), “Mawlaik, 6.4.[19]32” (underside),
“Margites sulcicollis Gah., D.J. Atkinson det. 1948”, “Pres. by Com. Inst. Ent.
B.M. 1948–165”.
Diagnosis. Based on female characters, this new
species is very similar to P.sulcicolliscomb.n., but differs
by the somewhat peculiar sculpture of the pronotal disc,
including a median elevation being distinctly narrower near
the apex, and by the presence of at least two longitudinal
folds, both rather rough and different in length, between
the elevation and the nearest, very coarse, longitudinal fold
on either of its sides, as in Figs51,52; the slightly narrower
scutellum; the lighter coloration of the elytra, antennae and,
partly, legs, as in Color plate2: 10,11, Figs34,35 (cf.Color
plate2: 8,9, Figs36,37, 49,50,56).
Description. Female. Body length 13.2–14.6mm, humeral
width 3.25–3.85 mm, thereby holotype largest. Eyes, head
dorsally, at least partly, pronotum mainly on median elevation
and longitudinal folds black; elytra brownish red; remaining parts
mainly red-brown, partly darkened.
Head with longitudinal folds between upper lobes of eyes;
antennal tubercles moderately developed; eyes very large, strongly
convex, lower lobes close together; submentum subequal in length
and width near middle, but not transverse, with a heterogeneous,
predominantly rough sculpture; antennae clearly or distinctly not
reaching the apex of elytra; length ratio of antennomeres 1–11,
26: 8 : 24 : 17 : 25: 27: 29: 27: 26: 25 : 30 (holotype taken as
an example); antennomere 1 with a dense rough puncturation;
antennomere2 very clearly longitudinal.
Pronotum barely transverse, 1.02–1.04times as wide as long;
base 1.16–1.24times as wide as apex; with a very sharp constriction
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 201
both in front of base and near apex; broadened angularly at the
middle; on disc with a wide, moderately convex, median elevation,
with both heterogeneous, partly rough puncturation and a clear or
at least noticeable longitudinal impression in basal part; lateral to
median elevation with very coarse folds, thereby between elevation
and nearest very coarse longitudinal fold at least with one long and
one shorter fold, both longitudinal, rather coarse, but significantly
lower than a very coarse longitudinal fold (see also Key to species
below).
Scutellum triangular, with an unclear sculpture.
Elytra predominantly nearly parallel-sided starting from
base, 2.61–2.65times as long as humeral width; with both a rough
sparse and very small dense puncturation; apical external angle
broadly rounded, sutural angle nearly right or narrowly rounded.
Prosternum in apical part with heterogeneous, rough,
predominantly transverse folds; prosternal process moderately
wide between coxae, without clear apical tubercle; mesosternal
process between coxae significantly wider than prosternal process,
without tubercle dorsally; metasternum and sternites with a small,
clear, dense puncturation; metasternum with a well-expressed
median groove; both last (visible) sternite and tergite widely
rounded apically.
Legs moderately long; profemora ventrally with a clearly
rough sculpture; all tibiae with a distinct carina along each side;
metatarsomere 1 noticeably shorter than metatarsomeres2 and3
combined.
Recumbent setation, except for elytra, mainly greyish,
partly silver-grey, that of elytra golden-yellow and yellow; head,
pronotum and elytra with rather long, erect, sparse, but numerous,
light setae; elytra, in addition, with numerous, suberect, short,
light setae; antennae, legs and venter with sparse, more or less
long, light setae.
Etymology. e name of the new species is derived
from Myanmar which it inhabits.
Distribution. Myanmar.
Plavichydissus rufipennis (Pic, 1923), comb. rest.
(Color plate3: 13, 15,16, 18, 20, 22;
Figs 38,39, 53,54, 57,58, 209)
Pachydissus rufipennis Pic, 1923a: 8. Type locality: “Laos, Ban
Saloueun” (according to the original description and the label of
the holotype). Plavilstshikov, 1931: 84.
Plavichydissus rufipennis: Pic, 1946: 107, 108 (Laos);
Miroshnikov, 2017: 223 (preliminary combination).
Margites (Margites) rufipennis: Gressitt, Rondon, 1970: 78
(Laos).
Margites rufipennis: Hua, 1984: 60; Holzschuh, 1999: 21.
Material. 1♀, holotype, by monotypy (MNHN) (photograph;
Color plate 3: 15), “Laos, B[an]. Saloueun, le 9.III.1920, R. Vitalis de
Salvaza”, “Pachydissus rufipennis n. sp.”, “Type”, “Museum Paris, Coll.
M. Pic”, “Holotype” (Fig. 209); 1♀ (BM), “Laos: Wapikhamthong Prov.,
Khong Sedone, 31.III.1965, “Khongsedone, 31.3.[19]65” (handwritten),
“J.A. Rondon Collection Bishop Mus.”, “Margites (s. str.) rufipennis (Pic),
J.L. Gressitt det.”; 1♂ (BM) (Color plate 3: 13), “Laos: Borikhane Prov.,
Pakkading, 15.IV.1966, “Pakkading, 15.4.[19]66” (handwritten), “J.A.Rondon
Collection Bishop Mus.”, “Margites (s.str.) rufipennis (Pic), J.L.Gressittdet.”;
1♀ (BM) (Color plate3:16), “Laos: Borikhane Prov., Pakkading, 17.III.1965,
“Pakkading, 17.3.[19]65” (handwritten), “J.A. Rondon Collection Bishop
M u s.”, “ Margites(s.str.) rufipennis (Pic), J.L.Gressittdet.”.
Morphological notes. Body length 9–13 mm
[Gressitt, Rondon, 1970]; the holotype is 15 mm long
(Dr. Gérard L. Tavakilian, personal communication); in
the specimens I have studied the body length was 12.5–
13.7mm, the humeral width between 3.15–3.5mm.
Pronotum subequal in length and width; base 1.14–
1.22 times as wide as apex; with a sharp constriction
both in front of base and near apex; broadened angularly
at the middle, as in Figs53, 54; with a strong, high, wide,
median elevation on disc, as in Figs 53, 54, 57, 58, with a
heterogeneous, mainly rough puncturation dorsally; lateral
to elevation with very coarse longitudinal folds.
Distribution. Laos.
Plavichydissus makarovi Miroshnikov, sp. n.
(Color plate 3: 14, 17, 19, 21; Figs 40, 55, 64)
Material. Holotype, ♂ (NHMD) (Color plate 3: 14): “ailand,
River Kwae, Erawan [National Park], 13.II.1994, [leg.]Mahuaka”, “Margites
rufipennis (Pic), Ole Mehl det. 2005”.
Diagnosis. is new species is very similar to
P. rufipennis comb. rest., but differs clearly by the less
strongly protruding, suberect, short setae and the presence
of only a small number of evidently shorter and significantly
more inclined erect setae on the elytra, as in Fig. 40; the
longitudinal pronotum which is less angularly broadened
at the middle, as in Fig. 55; the somewhat shorter male
antennae, as in Color plate3:14; the darker coloration of the
elytra and antennae, as in Color plate3:14; the more strongly
elongated parameres, as in Color plate3:19, the narrower
penis, including the apical part, as in Color plate3:17, the
darker coloration of the tegmen, penis and tergite8, as in
Color plate3: 17,19,21 (cf.Color plate3: 13,15, 16, 18, 20,
22, Figs38, 39, 53,54). Plavichydissus makarovisp.n. can
also be compared to the next new species, the differences
from which are given in its diagnosis.
Description. Male. Body length 10.6 mm, humeral width
2.65 mm. Head dorsally, eyes, almost entirely pronotun and
antennomere1 black; elytra dark reddish brown (elytra red-brown
in P. rufipennis comb. rest.); remaining parts mainly combines
dark brown and black-brown tones, partly with a reddish tint.
Head with longitudinal folds between upper lobes of eyes;
antennal tubercles well-developed; eyes large, strongly convex;
submentum with a heterogeneous, rough, partly coarse sculpture;
antennae reaching beyond apex of elytra by apex of penultimate
antennomere (male antennae of P. rufipennis comb. rest.
reach the apex of elytra by antennomere 9); length ratio of
antennomeres1–11, 21:6: 20: 14: 21: 24: 25: 25: 25: 24:30;
antennomere1 with a partly rough puncturation; antennomere2
very clearly longitudinal.
Pronotum barely longitudinal, 1.04 times as long as wide
(inP. rufipenniscomb.rest., pronotum both in male and female
subequal in length and width; see above); base 1.16times as wide as
apex; with a sharp constriction both in front of base and near apex;
broadened somewhat angularly at the middle; on disc with a strong,
high, wide, median elevation (like in P. rufipennis comb. rest.),
with a heterogeneous, mainly rough puncturation dorsally;
lateral to elevation with very coarse longitudinal folds (like in
P.rufipenniscomb.rest.).
Scutellum triangular, sharpened apically, with an unclear
sculpture.
Elytra barely narrowed towards apex, 2.57 times as long as
humeral width; with both a rough (but not too deep) sparse and
very small dense puncturation; apical external angle rounded,
sutural angle with a poorly expressed obtuse denticle.
Prosternum in apical part with transverse wrinkles;
prosternal process without clear apical tubercle; mesosternal
process between coxae noticeably wider than prosternal process,
without tubercle dorsally; metasternum and sternites with a small,
clear, dense puncturation; metasternum with a well-expressed
median groove; last (visible) sternite widely truncate at apex; last
(visible) tergite rounded apically.
Legs moderately long; profemora ventrally with a clearly
rough sculpture; all tibiae with a distinct carina along each side;
202 A.I. Miroshnikov
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) Color plate 3
Figs 13–22. Plavichydissus Pic, 1946, stat. rest., habitus, dorsal view, and male genitalia.
13, 15–16, 18, 20, 22 – P. rufipennis (Pic, 1923), comb. rest. (15 – photograph by Gérard L. Tavakilian); 14, 17, 19, 21 – P.makarovi sp.n. 14–15–
holotypes; 13–14 –males;15–16– females; 17–18– apical part of penis, ventral view; 19–20–apical part of tegmen, ventral view; 21–22–apical part of
tergite 8, dorsal view.
Рис. 13–22. Plavichydissus Pic, 1946, stat. rest., общий вид сверху и гениталии самца.
13, 15–16, 18, 20, 22 – P. rufipennis (Pic, 1923), comb. rest. (15 – фотография Ж . Тавакиляна); 14, 17, 19, 21 – P. makarovi sp.n. 14–15– голотипы;
13–14–самцы;15–16– самки; 17–18– вершинная часть пениса снизу; 19–20–вершинная часть тегмена снизу; 21–22–вершинная часть 8-го тергита сверху.
Color plate 4 e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae)
Figs 23–30. Plavichydissus Pic, 1946, stat. rest., habitus, dorsal view, and pronotum, holotypes.
23, 27 – P. decipiens (Holzschuh, 1989), comb. n. (after Holzschuh [1989], photographs by Luboš Dembický); 24, 28– P. penangensis sp.n.; 25, 29 – P.
sodalis (Holzschuh, 1999), comb. n. (after Holzschuh [1999], photographs by Luboš Dembický); 26, 30– P. dembickyi sp.n. 23–25–males;26– female.
Рис. 23–30. Plavichydissus Pic, 1946, stat. rest., общий вид сверху и переднеспинка, голотипы.
23, 27 – P. decipiens (Holzschuh, 1989), comb. n. (по [Holzschuh, 1989], фотографии Л. Дембицкого); 24, 28– P. penangensis sp.n.; 25, 29 – P. sodalis
(Holzschuh, 1999), comb. n. (по [Holzschuh, 1999], фотографии Л. Дембицкого); 26, 30– P. dembickyi sp.n. 23–25–самцы;26– самка.
metatarsomere1 noticeably shorter than metatarsomeres2 and3
combined.
Recumbent setation, except for elytra, mainly greyish, partly
yellowish greyish, that of elytra golden-yellow; pronotum with
more or less long, erect, sparse, but numerous, light setae; elytra
with separate, long, more or less inclined, reddish setae and, in
addition, with numerous, suberect, short, reddish setae; head,
antennae, legs and venter with sparse, more or less long, light setae.
Etymology. I am pleased to dedicate this new
species to my colleague and friend, Dr.KirillV. Makarov
(Moscow Pedagogical State University, Russia), a master
of microphotography who rendered his invaluable help in
taking the pictures presented in this work.
Distribution. ailand.
Plavichydissus nataliae Miroshnikov, sp. n.
(Color plate2: 12; Figs 41, 48, 65)
Material. Holotype, ♀ (cAM) (Color plate 2: 12): Vietnam, Gia Lai
Province, ~55km ENE of Pleiku, 14°17ʹ45ʺN / 108°26ʹ57ʹE, Kon Ka Kinh
National Park, 600m, at light, 8–20.05.2017 (leg.D.Fedorenko).
Diagnosis. is new species seems to be especially
similar to P.rufipenniscomb.rest. and P.makarovisp.n.,
but differs clearly from both by the peculiar shape of the
pronotum, as in Fig.48; the characteristic sculpture of its
disc, as in Fig. 48, including the less strongly developed,
much lower, median elevation, as in Fig. 41; the more
strongly elongated elytra, as in Color plate 2: 12. Besides
this, P. nataliae sp.n. differs from the former species by
the less strongly protruding, suberect, short setae and the
absence of very long, numerous, erect setae on the elytra,
as in Fig. 41 (somewhat similar to P. makarovisp. n.),
the longer antennae of the female, as in Color
plate2:12,Fig.41, while from the latter species by the wider
prosternal process between the coxae, the predominantly
shorter, recumbent, light setae on the prosternum and
probably the longer antennae of the female (the female of
P.makarovisp. n. is not yet known, but the antennae of
the male of this species are even shorter than in the male
of P.rufipennis comb.rest., see above) (cf.Color plate3:
13–16, Figs38–40, 53–55, 57–58).
Description. Female. Body length 15.5 mm, humeral
width 3.6 mm. Head dorsally, eyes, pronotum, almost entirely
antennomere 1 and femora, prosternum in basal part, meso-
and metasterna, mostly sternites black; elytra reddish brown;
remaining parts mainly combines dark brown and black-brown
tones, partly with a reddish tint.
Head with longitudinal folds between upper lobes of eyes;
antennal tubercles well-developed; eyes large, strongly convex;
submentum with a heterogeneous, rough, partly coarse sculpture;
antennae reaching beyond apex of elytra by last antennomere;
length ratio of antennomeres1–11, 29:9 : 30 : 21: 32: 35: 37 :
36: 34: 31:39; antennomere1 with a rough dense puncturation;
antennomere2 strongly longitudinal.
Pronotum distinctly longitudinal, 1.08 times as long as
wide; base 1.2 times as wide as apex; with a sharp constriction
both in front of base and near apex; broadened angularly at the
middle; on disc with a rather wide, moderately developed, median
elevation, with heterogeneous, partly rough, irregular folds
and heterogeneous sparse punctures dorsally; lateral to median
elevation with very coarse longitudinal folds.
Scutellum triangular, sharpened apically, with a very small
poorly expressed puncturation.
Elytra predominantly nearly parallel-sided starting from base,
2.75times as long as humeral width (in P.rufipennis comb.rest.
and P. makarovisp. n. 2.47–2.58 or 2.57 times, respectively);
with both a rough (but not too deep) sparse and very small dense
puncturation; apical external angle obtuse, well-expressed, sutural
angle with a poorly developed, but distinct, obtuse denticle.
Prosternum in apical part with transverse wrinkles;
prosternal process moderately wide, without clear apical tubercle;
mesosternal process between coxae noticeably wider than
prosternal process, without tubercle dorsally; metasternum and
sternites with a small, clear, dense puncturation; metasternum
with a well-expressed median groove; both last (visible) sternite
and tergite widely rounded apically.
Legs moderately long; profemora ventrally with a clearly
rough sculpture; all tibiae with a distinct carina along each side;
metatarsomere 1 clearly shorter than metatarsomeres 2 and 3
combined.
Recumbent setation mainly greyish, including that of elytra;
pronotum with more or less long, erect, sparse, but numerous,
light setae; elytra with separate, more or less long, but strongly
inclined, reddish setae and, in addition, with numerous, suberect,
short, reddish setae; head, antennae, legs and venter with sparse,
more or less long, light setae.
Etymology. I am pleased to dedicate this new species
to Natalia, my younger daughter.
Distribution. Vietnam.
Plavichydissus decipiens (Holzschuh, 1989), comb. n.
(Color plate4: 23, 27)
Margites decipiens Holzschuh, 1989: 393. Type locality:
Western Bhutan, Chimakothi (south of imphu) (according to
the original description).
Margites (Margites) decipiens: Catalogue..., 2010: 161.
Plavichydissus decipiens: Miroshnikov, 2017: 223 (preliminary
combination).
Material. 1♂, holotype (cCH) (photograph; Color plate3:23).
Morphological notes. Body length 11.4 mm
[Holzschuh, 1989].
Distribution. Bhutan.
Plavichydissus penangensis Miroshnikov, sp. n.
(Color plate4: 24, 28; Fig. 66)
Material. Holotype, ♂ (BMNH) (Color plate4:24): Western Malaysia,
“Penang”, ”Bowring , 63–47*”.
Diagnosis. Based on male characters, this new species
seems to be especially similar to P.decipienscomb.n., but
differs clearly by the peculiar sculpture of the pronotum,
including an obviously broader median elevation, as
in Color plate 4: 28, the partly smaller puncturation of
the elytra (discarding very small punctures), the weakly
expressed groove between the upper lobes of the eyes
which is completely invisible on the vertex, and the
generally darker coloration, as in Color plate4:24 (cf.Color
plate4:23,27).
Description. Male. Body length 11.8 mm, humeral width
3.05mm. Head dorsally, eyes, antennomere1, mostly pronotum
and partly scutellum black; elytra brownish red; remaining parts
mainly dark reddish brown, partly red-brown and black-brown tones.
Head with longitudinal folds between upper lobes of
eyes; antennal tubercles well-developed; eyes large, strongly
convex; submentum with a heterogeneous, rough, partly coarse
sculpture; antennae reaching beyond apex of elytra obviously
by last antennomere; length ratio of antennomeres1–11, 23:6:
23: 15 : 22 : 26 : 27: 27 : 27 : 26 : (last antennomere missing);
antennomere1 with a dense rough puncturation; antennomere2
very distinctly longitudinal.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 203
Figs 31–35. Plavichydissus Pic, 1946, stat. rest., habitus, lateral view.
31 – P. semiplicatus (Pic, 1926), comb. rest.; 32– P. irinae sp.n.; 33 – P. aggregatus (Holzschuh, 1999), comb. n.; 34–35– P. myanmarensis sp. n.
31–32, 34 – holotypes; 35– paratype; 31–32–males;33–35– females.
Рис. 31–35. Plavichydissus Pic, 1946, stat. rest., общий вид сбоку.
31 – P. semiplicatus (Pic, 1926), comb. rest.; 32– P. irinae sp.n.; 33 – P. aggregatus (Holzschuh, 1999), comb. n.; 34–35– P. myanmarensis sp. n.
31–32, 34 – голотипы; 35– паратип; 31–32–самцы;33–35– самки.
204 A.I. Miroshnikov
Figs 36–40. Plavichydissus Pic, 1946, stat. rest., habitus, lateral view.
36–37 – P. sulcicollis (Gahan, 1893), comb. n.; 38–39– P. rufipennis (Pic, 1923), comb. rest.; 40 – P. makarovi sp. n. 36, 40 – holotypes; 38,
40–males;36–37, 39– females.
Рис. 36–40. Plavichydissus Pic, 1946, stat. rest., общий вид сбоку.
36–37 – P. sulcicollis (Gahan, 1893), comb. n.; 38–39– P. rufipennis (Pic, 1923), comb. rest.; 40 – P. makarovi sp. n. 36, 40 – голотипы; 38,
40–самцы;36–37, 39– самки.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 205
Figs 41–48. Plavichydissus Pic, 1946, stat. rest., habitus, lateral view, and pronotum.
41, 48 – P. nataliae sp.n.; 42– P. dembickyi sp.n.; 43–44 – P. grossepunctatus (Gressitt et Rondon, 1970), comb. n.; 45 – P. irinae sp.n.; 46–47 – P.
aggregatus (Holzschuh, 1999), comb. n. (46 – after Holzschuh [1999], photograph by Luboš Dembický). 41–43, 45–46, 48 – holotypes; 44 – paratype; 41–42,
44, 46–48 –females;43, 45– males.
Рис. 41–48. Plavichydissus Pic, 1946, stat. rest., общий вид сбоку и переднеспинка.
41 – P. nataliae sp. n.; 42– P. dembickyi sp.n.; 43–44 – P. grossepunctatus (Gressitt et Rondon, 1970), comb. n.; 45 – P. irinae sp.n.; 46–47 – P.
aggregatus (Holzschuh, 1999), comb. n. (46 – по [Holzschuh, 1999], фотография Л. Дембицкого). 41–43, 45–46, 48 – голотипы; 44 – паратип; 41–42, 44,
46–48 –самки;43, 45– самцы.
206 A.I. Miroshnikov
Figs 49–58. Plavichydissus Pic, 1946, stat. rest., pronotum, dorsal and frontal views.
49–50, 56 – P. sulcicollis (Gahan, 1893), comb. n.; 51–52– P. myanmarensis sp.n.; 53–54, 57–58 – P. rufipennis (Pic, 1923), comb. rest.; 55 – P.
makarovi sp.n. 49, 51, 55 – holotypes; 52 – paratype; 49–53, 56, 58 –females;54–55, 57 –males.
Рис. 49–58. Plavichydissus Pic, 1946, stat. rest., переднеспинка сверху и спереди.
49–50, 56 – P. sulcicollis (Gahan, 1893), comb. n.; 51–52– P. myanmarensis sp.n.; 53–54, 57–58 – P. rufipennis (Pic, 1923), comb. rest.; 55 – P.
makarovi sp.n. 49, 51, 55 – голотипы; 52 – паратип; 49–53, 56, 58 –самки;54–55, 57 –самцы.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 207
Figs 59–67. Plavichydissus Pic, 1946, stat. rest., head, ventral view.
59 – P. grossepunctatus (Gressitt et Rondon, 1970), comb. n.; 60– P. irinae sp.n.; 61 – P. aggregatus (Holzschuh, 1999), comb. n.; 62 – P. sulcicollis
(Gahan, 1893), comb. n.; 63 – P. myanmarensis sp.n.; 64 – P. makarovi sp.n.; 65 – P. nataliae sp.n.; 66 – P. penangensis sp.n.; 67 – P. dembickyi sp.n.
59–60, 62–67 – holotypes; 59–60, 64, 66 –males;61–63, 65, 67 – females.
Рис. 59–67. Plavichydissus Pic, 1946, stat. rest., голова снизу.
59 – P. grossepunctatus (Gressitt et Rondon, 1970), comb. n.; 60– P. irinae sp.n.; 61 – P. aggregatus (Holzschuh, 1999), comb. n.; 62 – P. sulcicollis
(Gahan, 1893), comb. n.; 63 – P. myanmarensis sp.n.; 64 – P. makarovi sp.n.; 65 – P. nataliae sp.n.; 66 – P. penangensis sp.n.; 67 – P. dembickyi sp.n.
59–60, 62–67 – голотипы; 59–60, 64, 66 –самцы;61–63, 65, 67 – самки.
208 A.I. Miroshnikov
Pronotum barely longitudinal, 1.03 times as long as wide;
base 1.16times as wide as apex; with a sharp constriction both in
front of base and near apex; broadened angularly at the middle;
on disc with a wide, moderately developed, median elevation, with
rough irregular folds dorsally; lateral to elevation with very coarse,
longitudinal, partly sinuous folds, thereby fold nearest to elevation,
in apical part branching into two folds.
Scutellum triangular, with an unclear sculpture.
Elytra distinctly narrowed towards apex, 2.66 times as long
as humeral width; with both a rough sparse and very small dense
puncturation; apical external angle rounded, sutural angle nearly
right.
Prosternum in apical part with a heterogeneous sculpture,
partly transverse wrinkles; prosternal process moderately wide,
without distinct apical tubercle; mesosternal process between
coxae clearly wider than prosternal process, without tubercle
dorsally; metasternum and sternites with a small, clear, dense
puncturation; metasternum with a well-expressed median groove;
last (visible) sternite truncate at apex; last (visible) tergite rounded
apically.
Legs moderately long (posterior legs of holotype missing);
profemora ventrally with a rough dense puncturation; tibiae with a
distinct carina along each side.
Recumbent setation mainly greyish, including that of elytra;
pronotum with more or less long, erect, sparse, but numerous,
light setae; elytra at least with suberect, short, yellowish setae
(holotype with a very strongly obliterated setation of elytra); head,
antennae, legs and venter with sparse, more or less long, light setae
(partly abraded).
Etymology. e name of the new species is derived
from Penang Island, off the northwestern coast of Malay
Peninsula, the terra typica.
Distribution. Western Malaysia.
Plavichydissus sodalis (Holzschuh, 1999), comb. n.
(Color plate4: 25, 29)
Margites sodalis Holzschuh, 1999: 21. Type locality: Western
Malaysia, Pahang, Tioman Island, KajangMt., Wslope (according
to the original description).
Plavichydissus sodalis: Miroshnikov, 2017: 223 (preliminary
combination).
Material. 1♂, holotype (cCH) (photograph; Color plate4:25).
Morphological notes. Body length 13.9 mm
[Holzschuh, 1999].
Distribution. Western Malaysia.
Plavichydissus dembickyi Miroshnikov, sp. n.
(Color plate4: 26, 30; Fig. 67)
Material. Holotype, ♀ (cLD) (Color plate4:26): WesternMalaysia,
Perak, Banjaram Bintang, Bukit Berapit (Talping), 22–23.02.1997
(leg.I.Jeniš).
Diagnosis. is new species seems to be especially
similar to P.sodaliscomb.n., but differs by the somewhat
peculiar sculpture of the pronotum, as in Color plate4:30;
the seemingly sparser, recumbent, light setation of the
elytra and, as a consequence, the more strongly expressed
punctures; the absence of a distinctly red or reddish colour
in the coloration of the suberect short setae of the elytra;
and the generally darker coloration, as in Color plate4:26
(cf.Color plate4:25,29).
Description. Female. Body length 16 mm, humeral width
3.8 mm. Head dorsally, eyes, almost entirely pronotum black;
elytra dark reddish brown; remaining parts mainly combines red-
brown and dark brown tones with a red tint.
Head with longitudinal folds between upper lobes of eyes;
antennal tubercles very clearly expressed; eyes relatively well
developed, moderately convex; submentum with a heterogeneous,
rough, partly coarse sculpture; antennae freely reaching
beyond apex of elytra by last antennomere; length ratio of
antennomeres1–11, 29:9: 33: 22: 33: 36: 36: 35: 34: 32:43;
antennomere1 with a dense rough puncturation; antennomere2
very distinctly longitudinal.
Pronotum barely longitudinal, 1.05 times as long as wide;
base 1.17 times as wide as apex; with a sharp constriction both
in front of base and near apex; broadened somewhat angularly
at the middle; on disc with wide, moderately developed, median
elevation, with heterogeneous, transverse, partly rough folds;
lateral to elevation with irregular very coarse folds, in general
forming relatively wide longitudinal fragment of sculpture, on
either side of which with separate, coarse, longitudinal folds.
Scutellum triangular, with an unclear sculpture.
Elytra predominantly nearly parallel-sided starting from
base, 2.59times as long as humeral width; with both a rough, sharp,
sparse and very small dense puncturation; apical external angle
rounded, sutural angle with a poorly developed obtuse denticle.
Prosternum in apical part with well-expressed transverse
folds; prosternal process with a poorly noticeable apical tubercle;
mesosternal process between coxae very clearly wider than
prosternal process, without tubercle dorsally; metasternum and
sternites with a small, clear, dense puncturation; metasternum with
a distinct median groove; last (visible) sternite widely rounded at
apex; last (visible) tergite rounded apically.
Legs moderately long; profemora ventrally with a rough
sculpture; all tibiae with a distinct carina along each side;
metatarsomere 1 clearly shorter than metatarsomeres 2 and 3
combined.
Recumbent setation, except for elytra and scutellum, greyish
and greyish yellowish, of elytra and scutellum olive; head, pronotum
and elytra with more or less long, erect, partly inclined, sparse, but
numerous, light setae; elytra, in eddition, with numerous, sub erect,
short, yellowish setae; antennae, legs and venter with sparse, more
or less long, light setae.
Etymology. I am pleased to dedicate this new species
to my colleague and friend, Mr.Luboš Dembický (Brno,
Czech Republic), who constantly provides a very important
assistance to my research.
Distribution. Western Malaysia.
Key to species of Plavichydissus stat. rest.
1. Dense or at least abundant, recumbent, light setae on
median elevation of pronotum forming a characteristic
horseshoe-shaped pattern, as in Color plate 1: 1, 4,
Figs43–47; if this pattern poorly expressed due to a
small number of setae (obviously partly obliterated),
then elytra partly with very large sparse punctures
(discarding very small puncturation), as in Color
plate1:1,4; sculpture of median elevation of pronotum
usually mostly strongly obliterated, as in Figs 43–47 ...
............................................................................................... 2
– Median elevation of pronotum at most with sparse
recumbent setae forming no pattern, with a coarse or
rough sculpture at least partly in the form of irregular
folds, punctures or their combination, as in Color
plate4:27–30, Figs48, 49–55 ........................................ 5
2. Elytra with a dense, recumbent, grey or silver-grey
setation, as in Color plate1:1,2, 4,5,Fig.31 .............. 3
– Elytra with a dense, recumbent, cream or yellowish
cream setation, as in Color plate1:3, 6, Color plate2:7,
Figs32, 33 ........................................................................... 4
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 209
3.Elytra predominantly with a very large puncturation, as
in Color plate1:1,4; median elevation of pronotum
with pale, greyish yellowish or greyish, more or less
numerous, recumbent setae, as in Color plate1:1,4 ...
................................................ P. semiplicatus comb. rest.
– Elytra with a significantly smaller puncturation, as in
Color plate 1: 2, 5; median elevation of pronotum
with bright, golden-yellow or yellow, dense or at least
numerous, recumbent setae, as in Figs43,44 ................
.............................................. P. grossepunctatus comb. n.
4. Head dorsally, pronotum (Figs46, 47) and elytra with
a combination of dark red-brown and red-brown
tones; at least two longitudinal folds of pronotal disc
adjacent to and flanked by a median elevation only
partly sinuous, as in Figs46,47; puncturation of elytra
appearing sharper, as in Color plate 1: 3, 6; body
smaller, up to 19.4mm in length ....................................
........................................................ P. aggregatus comb. n.
–Head dorsally, pronotum (Fig.45) and elytra black; two
longitudinal folds of pronotal disc adjacent to and
flanked by a median elevation completely sinuous, as
in Fig.45; puncturation of elytra appearing weaker, as
in Color plate2:7; body larger, 28.3mm in length .......
....................................................................... P. irinae sp. n.
5. Pronotum with a strong, very high, wide, median
elevation, as in Figs57,58 ............................................... 6
– Pronotum with a less strongly developed, much lower
and usually narrower, median elevation, as in Fig.56 ....
............................................................................................... 7
6. Elytra darker, with clearly less strongly protruding,
suberect, short setae and, in addition, with a small
number of moderately long and strongly inclined
setae, as in Fig.40; pronotum longitudinal, at least in
the male less angularly broadened at the middle, as in
Fig.55; male antennae shorter, as in Color plate3:14;
male genitalia darker, as in Color plate3: 17, 19, 21,
parameres more strongly elongated, as in Color
plate3:19, penis narrower, as in Color plate3:17 ........
................................................................. P. makarovi sp. n.
– Elytra lighter, with clearly more strongly protruding,
suberect, short setae and, in addition, with
numerous, very long, erect setae, as in Figs 38–39;
pronotum subequal in length and width, more
angularly broadened at the middle, as in Figs53,54;
male antennae longer, as in Color plate 3: 13; male
genitalia significantly lighter, as in Color plate3: 18,
20,22, parameres less strongly elongated, as in Color
plate3:20, penis wider, as in Color plate3:18 ................
.................................................... P. rufipennis comb. rest.
7. At least 2–3 coarse or very coarse longitudinal folds
of pronotum, adjacent to and flanked by a median
elevation, more or less narrow, clearly separated from
each other, without distinct folds connecting them
(sometimes connected only at the very apex and/or at
the very base), weakly sinuous, as in Figs48–52, only
sometimes one of the folds about basal one-third can
be somewhat wider than in the remaining part ........... 8
– Sculpture of pronotum adjacent to a median elevation
on either of its sides formed by coarse or very coarse,
irregular, partly transverse folds or by two coarse,
longitudinal, sinuous folds (sometimes these in basal
parts fused into one irregularly intertwined fold) partly
connected by irregular folds, as in Color plate4:27–30
............................................................................................. 10
8. Pronotum barely transverse, 1.01–1.05 times as wide
as long, as in Figs 49–52; lower lobes of eyes close
together, thereby submentum subequal in length
and width near middle, as in Figs 62, 63; elytra less
strongly elongated, 2.46–2.65times as long as humeral
width, as in Color plate2: 8–11, with more strongly
protruding suberect setae and long or very long erect
setae all along elytra, as in Figs34–37 ........................... 9
–Pronotum distinctly longitudinal, 1.08 times as long as
wide, peculiar in shape, as in Fig. 48; lower lobes of
eyes relatively widely spaced, submentum very clearly
transverse, as in Fig.65; elytra more strongly elongated,
2.75 times as long as humeral width, as in Color
plate 2: 12, with less strongly protruding suberect
setae and separate, relatively long, erect setae only at
base of elytra, as in Fig.41 ................... P.nataliae sp. n.
9.Pronotum between median elevation and nearest, very
coarse, longitudinal fold on either side of elevation
with a rather wide and deep groove supplied with
only weakly expressed, individual, longitudinal, short
tubercles at bottom, as in Figs 49, 50; elytra darker,
as in Color plate2:8, 9; scutellum wider, as in Color
plate2:8,9 ..................................... P. sulcicollis comb. n.
– Pronotum between median elevation and nearest very
coarse longitudinal fold on either side of elevation
with a rather wide and deep groove showing one long
and one shorter fold, both longitudinal, rather coarse,
but significantly lower than a very coarse longitudinal
fold, as in Figs 51, 52; elytra lighter, as in Color
plate2:10,11; scutellum narrower, as in Color plate2:
10,11 ............................................ P.myanmarensis sp. n.
10. Elytra with a red or brownish red coloration of
integument and a pale greyish coloration of a
recumbent setation, as in Color plate4: 23, 24; male
with much shorter antennae (with many antennomeres
less strongly elongated), reaching beyond apex of
elytra by only last antennomere, as in Color plate4:23
............................................................................................. 11
–Elytra with a dark brown coloration of integument and
a bright olive coloration of a recumbent setation, as
in Color plate 4: 25, 26; male (if known) with much
longer antennae (with many antennomeres more
strongly elongated), reaching beyond apex of elytra by
antennomere 9, as in Color plate4:25 ........................ 12
11. Median elevation of pronotum clearly wider, with a
somewhat coarser sculpture, as in Fig. 28; elytra,
antennae and legs darker, as in Color plate 4: 24;
puncturation of elytra mostly smaller (discarding very
small puncturation), as in Color plate 4: 24; groove
between upper lobes of eyes weakly expressed, as in
Color plate4:24. Penang, WMalaysia ............................
........................................................... P. penangensis sp. n.
–Median elevation of pronotum clearly narrower, with a
somewhat less coarse sculpture, as in Color plate4:27;
elytra, antennae and legs lighter, as in Color plate4:23;
puncturation of elytra mostly larger (discarding very
small puncturation), as in Color plate 4: 23; groove
between upper lobes of eyes very well-expressed, as in
Color plate4:23. Bhutan ............. P. decipiens comb. n.
210 A.I. Miroshnikov
12. Pronotum on either side of median elevation with
clearly more strongly developed irregular folds
forming a generally much wider longitudinal
fragment of sculpture, as in Fig.30; median elevation
of pronotum itself wider in middle part, as in Color
plate4:30; recumbent light setation somewhat sparser
and, as a consequence, elytral punctures more sharply
expressed, as in Color plate4:26; coloration at least
of antennae and, partly, legs clearly darker, as in Color
plate4:26 ............................................ P. dembickyi sp. n.
– Pronotum on either side of median elevation with
clearly less strongly developed irregular folds forming
a generally much narrower longitudinal fragment of
sculpture, as in Color plate4:29; median elevation of
pronotum itself narrower in middle part, as in Color
plate4:29; recumbent light setation somewhat denser
and, as a consequence, elytral punctures less sharply
expressed, as in Color plate4:25; coloration at least
of antennae and, partly, legs clearly lighter, as in Color
plate4:25 ............................................ P. sodalis comb. n.
Genus Pachydissus Newman, 1838
Pachydissus Newman, 1838: 494; omson, 1864: 231;
Lacordaire, 1868: 265; Gemminger in Gemminger, Harold, 1872:
2804; Gahan, 1891: 24; Reitter, 1894: 356; Gahan, 1906: 133;
Aurivillius, 1912: 56; Plavilstshikov, 1931: 83; Gressitt, 1951: 141;
Gressitt, Rondon, 1970: 71; Adlbauer, 2002: 158; Catalogue…,
2010: 162; Ślipiński, Escalona, 2016: 223; Kariyanna et al., 2017: 34;
Miroshnikov, 2017: 220.
Type species: Pachydissus sericus Newman, 1838, by
monotypy.
Pachydissus parvicollis Gahan, 1891
(Color plate5:68; Fig. 210)
Pachydissus parvicollis Gahan, 1891: 29. Type locality:
Northern India (according to the original description and the
label of the syntype male). Gahan, 1906: 134; Aurivillius, 1912:
57; Plavilstshikov, 1931: 84; Hayashi, 1981: 7; Weigel, 2006: 498;
Catalogue..., 2010: 162; Kariyanna et al., 2017: 34; Miroshnikov,
2017: 221, fig.398.
Material. 1♂, syntype (BMNH) (Color plate 5: 68), “N. India”
(upperside), “Col. [illegible further on]” (underside), “60–15 E.I.C.”,
“Pachydissus parvicollis Gahan, Type”, “Type”, “Syntype” (Fig.210); 1♂, 1♀
(BMNH), NorthernIndia.
Morphological notes. Body length 30–32 mm,
humeral width 8–8.5mm [Gahan, 1906].
Distribution. Northern India; has also been recorded
from Nepal [Hayashi, 1981; Weigel, 2006].
Pachydissus schmutzenhoferi Holzschuh, 1990
(Color plate5: 70)
Pachydissus schmutzenhoferi Holzschuh, 1990: 185. Typ e
locality: Western Bhutan, Paro Distr., Gedu, 2000m (according to
the original description). Catalogue..., 2010: 162; Kariyanna et al.,
2017: 34; Miroshnikov, 2017: 221, fig.399.
Material. 1♂, holotype (cCH) (photograph; Color plate 5: 70); 1♀
(BMNH), “India”, “Pascoe Coll. 93–60”, “Pachydissus schmutzenhoferi
Holzschuh, 1990 ♀ det.A.Miroshnikov 2018”.
Morphological notes. Body length 19.8–29 mm
[Holzschuh, 1990]; the female I have studied has a body
length of 24.7mm and a humeral width of 6.2mm.
Distribution. Bhutan, northern India.
Pachydissus obsolescens Holzschuh, 2017
(Color plate5:69)
Pachydissus obsolescens Holzschuh, 2017: 66. Type locality:
Myanmar, Kachin State, ree River Junction (one chaung
sone), 26°23ʹ12ʺN/ 98°41ʹ04ʺE, 2044 m (according to the original
description).
Material. 1♂, holotype (cCH) (photograph; Color plate5:69).
Morphological notes. Body length 27–29 mm
[Holzschuh, 2017].
Distribution. Myanmar.
Pachydissus pullus Holzschuh, 2017
(Color plate5:71, 72; Color plate6: 76)
Pachydissus pullus Holzschuh, 2017: 67. Type locality:
“ailandN, Chiang MaiN, Doi Pha Hom Pok, 20°05ʹN, 99°15ʹE
(H = 2044 m)” (according to the original description) (see
Remarks).
Material. 1♀, holotype (cCH) (photograph; Color plate 5: 71); 1♀
(NHMD) (Color plate5:72), ailand, Chiang Mai Prov., Ban San Pakia,
1700 m, 25.04–7.05.1996 (leg.S.Bilý), “Pachydissus pullus Holzschuh, 2017
♀ det.A.Miroshnikov 2018”.
Morphological notes. Body length 26–32 mm
[Holzschuh, 2017]; the female I have studied has a body
length of 34mm and a humeral width of 9mm.
Remarks. e coordinates and altitude of the type
locality of this species as given in the original description
[Holzschuh, 2017] strongly mismatch. At least one if not
both of these parameters is wrong. It is highly suspicious
that the altitude accurate to one meter (2044m) matches
the altitude of the type locality of the previous species
[Holzschuh, 2017: 66]. In this connection, the type locality
of P.pullus requires clarification.
Distribution. ailand.
Pachydissus murzini Miroshnikov, sp. n.
(Color plate5:73; Color plate6: 77)
Material. Holotype, ♂ (cSM) (Color plate 5: 73): China, Yunnan
Province, 54km E of Tengchong, 2150m, 4–9.11.2004 (leg.S.Murzin).
Diagnosis. is new species seems to be especially
similar to P.pullus, but differs by the somewhat peculiar
sculpture of the pronotum, as in Color plate 6: 77;
the recumbent light setation of the elytra forming a
comparatively less strongly expressed, mottled, iridescent
pattern, as in Color plate 5: 73; and the scutellum more
strongly rounded on the sides. Pachydissus murzinisp. n.
can also be compared to P.obsolescens, P.schmutzenhoferi
and P.parvicollis, but differs from the former by the clearly
longer and seemingly more slender male antennae with
many antennomeres, including antennomere3, being more
strongly elongated, as in Color plate5: 73, and the more
slender legs, as in Color plate5:73, while from latter two
species at least by the darker, mainly black and brown-black
coloration, the somewhat peculiar shape and sculpture of
the pronotum, the more strongly protruding or sharper
apical external angle of at least several antennomeres
starting with the 3rd (cf. Color plate 5: 68–72, Color
plate6:76).
Description. Male. Body length 29.5 mm, humeral width
7 mm. Coloration of integument mainly black, only partly
mesosternum, both first and second (visible) sternites and mostly
epipleura reddish brown.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 211
Head with a deep median groove between upper lobes of
eyes and partly on vertex; antennal tubercles very well-developed;
eyes moderately convex; submentum with a heterogeneous,
rough, partly coarse sculpture; antennae much longer than body,
reaching beyond apex of elytra by antennomere7; length ratio of
antennomeres1–11, 32:5: 54: 35: 60: 56: 54: 49: 46: 40:58;
antennomere1 with a small dense puncturation; antennomere2
very clearly transverse; apical external angle of antennomeres3–10
one way or another sharpened, thereby on antennomeres 3–5,
especially on4th, strongly drawn towards external side.
Pronotum barely transverse, 1.02times as wide as long; base
1.17times as wide as apex; with a sharper constriction near apex
than in front of base; angularly broadened at the middle; with very
coarse, partly sinuous, irregular, mostly transverse folds, finely and
sparsely punctured.
Scutellum widely rounded apically, with an unclear sculpture.
Elytra predominantly barely narrowed towards apex starting
from base, 2.8times as long as humeral width; with a very small,
clear, dense puncturation; apical external angle with a short, but
well-expressed, obtuse tooth, sutural angle drawn into a long sharp
tooth.
Prosternum predominantly with coarse, partly sinuous,
transverse folds; prosternal process truncate apically and
dorsally, sharply protruding in this place; mesosternal process
between coxae distinctly wider than prosternal process, without
tubercle dorsally; metasternum and sternites with a small dense
puncturation, but less clear on sternites; metasternum with a sharp
median groove; last (visible) sternite truncate at apex; last (visible)
tergite with a poorly developed emargination apically.
Legs long and slender; metatarsomere 1 clearly longer than
metatarsomeres2 and3 combined.
Recumbent setation of dorsum mainly golden-yellow bright
while of venter, antennae and legs predominantly paler, mainly
yellowish and greyish yellowish tones; elytral setation irregular,
patterned and iridescent; more or less long, erect, light setae
mainly developed on pronotum and head.
Etymology. I am pleased to dedicate this new species
to my colleague and friend, Dr.SergeyV. Murzin (Moscow,
Russia), who collected the holotype and, over the many
years, supports my entomological research.
Distribution. China (Yunnan).
Pachydissus patricius Holzschuh, 1991
(Color plate6:74)
Pachydissus patricius Holzschuh, 1991: 36. Type locality:
ailand, NE Bangkok, Saraburi (according to the original
description). Miroshnikov, 2017: 222, fig.404.
Material. 1♀, holotype (cCH) (photograph; Color plate6:74).
Morphological notes. Body length 28.2 mm
[Holzschuh, 1991].
Distribution. ailand.
Pachydissus borneoensis Miroshnikov, sp. n.
(Color plate6:75)
Material. Holotype, ♂ (NHMD) (Color plate 6: 75): E Malaysia,
Sabah, Crocker Range, 03.2003 (local collector).
Diagnosis. is new species is similar to P.patricius,
but differs by the peculiar sculpture of the pronotum, as
in Color plate 6: 75 (see also Remarks below), the shape
of the elytral apex, as in Color plate6:75; the absence of
a clear groove between the upper lobes and on the vertex,
the more or less significant presence of black colour in the
coloration of several basal antennomeres, some features
of the coloration of the recumbent setation (cf. Color
plate6:74).
Description. Male. Body length 26.6 mm, humeral width
6.2mm. Mostly dorsum and tarsi and partly pro- and mesosterna
dark brown; eyes, partly mandibles, antennomeres 1 and 3–5,
almost entirely antennomere 2 black; femora, tibiae, epipleura,
partly antennae brownish red; remaining parts red-brown.
Head without distinct groove between upper lob es of eyes and
on vertex; antennal tubercles moderately developed; eyes relatively
weakly convex; submentum with a heterogeneous puncturation,
small and dense predominantly in middle part, vs. rough, partly
confluent near lateral margins; antennae much longer than body,
reaching beyond apex of elytra by antennomere7; length ratio of
antennomeres1–11, 29:11: 49: 23: 63: 56: 56: 52: 56: 59:100;
antennomere1 with a very coarse sculpture forming, in addition
to everything else, in middle part dorsally a strong longitudinal
rib, the latter occupying more than half of antennomere length
starting from base, as well as with a small dense puncturation;
antennomere 2 barely longitudinal; apical external angle of
antennomeres3–10 rounded or obtuse, not drawn towards laterad;
antennomeres3 and4 partly with a clear longitudinal impression
both dorsally and ventrally.
Pronotum distinctly longitudinal, 1.07times as long as wide;
base 1.09times as wide as apex; with a well-expresed constriction
both in front of base and near apex; with coarse and very coarse,
partly sinuous, transverse folds, these being very finely, irregularly,
in places densely punctured.
Scutellum rounded apically, with an unclear sculpture.
Elytra in basal one-third nearly parallel-sided, but then very
clearly narrowed towards apex, 2.6times as long as humeral width;
with a small, very clear, dense puncturation; apical external angle
with a well-expressed denticle, sutural angle drawn into a relatively
short, but very clear, sharp tooth.
Prosternum in apical one-third with somewhat rough
transverse folds, in middle part with very coarse transverse
folds; prosternal process truncate apically and dorsally, sharply
protruding in this place; mesosternal process between coxae
significantly wider than prosternal process, with a small tubercle
dorsally; metasternum and sternites with a small, clear, dense
puncturation; metasternum with a well-expressed median groove;
last (visible) sternite widely truncate at apex; last (visible) tergite
rounded apically.
Legs moderately long; femora relatively robust;
metatarsomere 1 barely shorter than metatarsomeres 2 and 3
combined.
Recumbent setation mainly greyish, partly yellowish or with
a yellowish tint (in P. patricius, recumbent setation at least of
dorsum and antennae dorsally seemingly without yellowish setae),
elytral setation irregular (in holotype largely abraded); more or less
long, erect, light setae mostly developed on pronotum and head.
Remarks. Although the differences between the new
species and P.patricius are based only on single specimens
of opposite sex, nonetheless these taxa are most likely to also
be distinguished by the shape of the pronotum. At least such
very clear differences in the shape of the pronotum as those
observed between the male of P.borneoensissp.n. and the
female of P.patricius are not encountered between the male
and female of the same taxon in some other Asian species
of the genus (e.g. P. parvicollis and P. schmutzenhoferi).
Besides this, unlike the male of P. borneoensissp. n. and
the female of P. patricius, the pronotum in the males of
P. obsolescens and P. murzinisp. n., is, on the contrary,
somewhat more strongly broadened on the sides than in
the female of P. pullus.
Etymology. A separate genus, Falsopachydissus
Miroshnikov, 2017, has recently been established for
the sole previously known representative of the genus in
Borneo, Pachydissus foveiscapus Holzschuh, 2011. In this
212 A.I. Miroshnikov
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) Color plate 5
Figs 68–73. Pachydissus Newman, 1838, habitus, dorsal view.
68 – P. parvicollis Gahan, 1891; 69 – P. obsolescens Holzschuh, 2017; 70 – P. schmutzenhoferi Holzschuh, 1990; 71–72 – P. pullus Holzschuh, 2017;
73 – P. murzini sp.n. 68– syntype; 69–71, 73 – holotypes; 68–70, 73 –males;71–72 – females; 69–71 – after Holzschuh [1990, 2017], photographs by Luboš
Dembický.
Рис. 68–73. Pachydissus Newman, 1838, общий вид сверху.
68 – P. parvicollis Gahan, 1891; 69 – P. obsolescens Holzschuh, 2017; 70 – P. schmutzenhoferi Holzschuh, 1990; 71–72 –P. pullus Holzschuh, 2017;
73 – P. murzini sp.n. 68– синтип; 69–71, 73 – голотипы; 68–70, 73 –самцы;71–72 – самки; 69–71 – по [Holzschuh, 1990, 2017], фотографии Л. Дембицкого.
Color plate 6 e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae)
Figs 74–80. Pachydissus Newman, 1838 and Dymasius J. omson, 1864, habitus, dorsal view, and pronotum.
74 – P. patricius Holzschuh, 1991 (after Holzschuh [1991], photograph by Luboš Dembický); 75 – P. borneoensis sp. n.; 76 – P. pullus Holzschuh, 2017;
77 –P. murzini sp.n.; 78 – P. birmanicus Gardner, 1926 (photograph by Sudhir Singh); 79 – P. argentatus Pic, 1923 (photograph by Gérard L. Tavakilian); 80 –
D. querceus Holzschuh, 2015 (after Holzschuh [2015], photograph by Luboš Dembický). 74–75, 77–80 – holotypes; 74, 76, 78–79 – females; 75, 77, 80–males.
Рис. 74–80. Pachydissus Newman, 1838 и Dymasius J. omson, 1864, общий вид сверху и переднеспинка.
74 – P. patricius Holzschuh, 1991 (по [Holzschuh, 1991], фотография Л. Дембицкого); 75 – P. borneoensis sp. n.; 76 – P. pullus Holzschuh, 2017;
77– P. murzini sp.n.; 78 – P. birmanicus Gardner, 1926 (фотография С. Сингха); 79 – P. argentatus Pic, 1923 (фотография Ж. Тавакиляна); 80 – D.
querceus Holzschuh, 2015 (по [Holzschuh, 2015], фотография Л.Дембицкого). 74–75, 77–80 – голотипы; 74, 76, 78–79 – самки; 75, 77, 80 –самцы.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 213
connection, P. borneoensissp. n. is currently the first
member of the genus Pachydissus to be found in Borneo,
and its epithet is intentionally formed on the basis of the
name of the locality it supports.
Distribution. Eastern Malaysia.
Pachydissus argentatus Pic, 1923
(Color plate6:79; Fig. 211)
Pachydissus argentatus Pic, 1923b: 8. Type locality: China,
“ibet, Vrianatang” (according to the original description and
the label of the holotype). Plavilstshikov, 1931: 84; Pic, 1946: 107,
108; Gressitt, 1951: 141; Hua, 2002: 222; Wang, Hua, 2009: 180,
Catalogue..., 2010: 162.
Material. 1♀, holotype, by monotypy (MNHN) (photograph; Color
plate 6: 79), “ibet, Vrianatang”, “Pachydissus argentatus n. sp.”, “Type”,
“Museum Paris, Coll. M.Pic”, “Holotype” (Fig.211).
Morphological notes. Body length of holotype
18.75 mm (Dr. Gérard L. Tavakilian, personal
communication).
Remarks. e IRSN collection contains a female I
have studied (“ibet, Coll. Le Moult”) which is very similar
to the holotype. Its body length is 19.2 mm, the humeral
width is 5.2mm, and the antennae are clearly longer than
the body, reaching beyond the apex of the elytra by the
penultimate antennomere.
Distribution. China (Xizang).
Pachydissus thibetanus Pic, 1946
Pachydissus thibetanus Pic, 1946: 108 (“ibet”). Type
locality: China, Xizang Province (according to the original
description). Gressitt, 1951: 632; Hua, 2002: 222; Wang, Hua, 2009:
180; Catalogue..., 2010: 162.
Morphological notes. Body length 23 mm [Pic, 1946].
Remarks. A recent attempt to relocate the type
specimen of this species in the Muséum national d’Histoire
naturelle, Paris, kindly undertaken by Dr. Gérard L.
Tavakilian upon my request, was unsuccessful. Instead,
Pic’s collection contains a label written by André Villiers,
where he noted to have never seen the P.thibetanus type.
Distribution. China (Xizang).
Pachydissus birmanicus Gardner, 1926
(Color plate6:78; Fig. 212)
Pachydissus birmanicus Gardner, 1926: 199. Type locality:
Burma (now Myanmar), Bondaung, S of Toungoo (according to
the original description and the label of the holotype).
Material. 1♀, holotype, by monotypy (NFIC) (photograph; Color
plate 6: 78), “For. Zool. Coll. / Bondaung, S. Toungoo, 18.5.1918. C.F.C.
Beeson” (upperside), “Found under bark of Xylia dolabriformis” (underside),
“Pachydissus birmanicussp.n. J.C.M.Gardner, Type”, “Type”, “332” (Fig.212).
Morphological notes. Body length 34.4mm, humeral
width 10mm (Dr.Sudhir Singh, personal communication).
Distribution. Myanmar.
Pachydissus elegans Nonfried, 1895
Pachydissus elegans Nonfried, 1895: 307. Type locality:
Indonesia, W Sumatra, Siboga (according to the original
description).
Remarks. is species, like several other taxa describe d
by Nonfried [Miroshnikov, 2017: 204], are known to me
only through the original description [Nonfried, 1895],
even though I have made repeated attempts to relocate its
type specimen(s) in a number of European museums and
some other institutions.
At the same time, based on its description, the
species in question most likely does not belong to the
genus Pachydissus. At least none of the Asian species of
Pachydissus has a pronotum being strongly sharpened on
each side, a relatively large body (48 mm in length) and
some other features as in P.elegans [Nonfried, 1895: 307–
308]. erefore, the present taxon is only conditionally
considered here within the genus Pachydissus.
Distribution. Indonesia (Sumatra).
“Pachydissus langsonius Fairmaire, 1895”
Remarks. Vitali et al. [2017] have recently shown
that the holotype of this species, kept in the Muséum
national d’Histoire naturelle, Paris, they revised, does not
differ significantly from Trirachys holosericeus (Fabricius,
1787) and in some morphological features, in particular,
the structure of the antennae, does not correspond to the
original description. ey regard this holotype to be false
and propose Pachydissus langsonius to provisionally be
considered as incertae sedis.
At the same time, on the basis of some specimens
kept at the Institut Royal de Sciences naturelles de
Belgique, Bruxelles, identified by Fairmaire as “Pachydissus
langsonius” and referred to by Vitali et al. [2017], as well
as considering the original description [Fairmaire, 1895:
176–177], the possibility that the holotype of P.langsonius
is false [Vitali et al., 2017], and the combination “Aeolesthes
langsonius” proposed by Aurivillius [1912:47], the species
in question is likely to belong either to the genus Trirachys
Hope, 1843 (if not a synonym of T. holosericeus) or to
the genus Aeolesthes Gahan, 1890, but anyway not to
Pachydissus.
“Dymasius querceus Holzschuh, 2015”
(Color plate6:80)
Remarks. In my opinion, the generic attribution of
D. querceus requires clarification, bearing in mind that
this species most likely belongs to the genus Pachydissus
[Miroshnikov, 2017].
Genus Margites Gahan, 1891
Margites Gahan, 1891: 26 (Pachydissus subgen., “section”);
Gahan, 1906: 137; Aurivillius, 1912: 59; Winkler, 1929: 1142;
Plavilstshikov, 1931: 88; 1940: 113, 641; Gressitt, 1951: 143;
Kojima, Hayashi, 1969: 48; Gressitt, Rondon, 1970: 77; Lee, 1982:
28; Kusama, Takakuwa, 1984: 255; Adlbauer, 2006: 63; Catalogue...,
2010: 161; Heffern, 2013: 10; Nga et al., 2014: 435; Kariyanna et al.,
2017: 31; Miroshnikov, 2017: 223.
Type species: Cerambyx egenus Pascoe, 1858, by
subsequent designation [Gahan, 1906].
Margites egenus (Pascoe, 1858)
(Figs 81, 98, 102, 106, 112, 215)
Cerambyx egenus Pascoe, 1858: 236 (“China Borealis”). Type
locality: Northern China (according to the original description and
the label of the holotype).
214 A.I. Miroshnikov
Pachydissus (Margites) egenus: Gahan, 1891: 26.
Margites egenus: Aurivillius, 1912: 59; Winkler, 1929: 1142;
Plavilstshikov, 1931: 89; Gressitt, 1951: 144 (“China: N. China;
Szechuan (Chengtu); Kwangtung (Lien)”); Wang, Hua, 2009: 174.
Margites (Margites) egenus: Catalogue..., 2010: 161 (China:
“Northern Territory, Sichuan and Guandong provinces”).
Material. 1♀, holotype, by monotypy (BMNH) (Fig. 81), “NChina”,
“Cerambyx egenus Pasc[oe]. Type”, “Type”, “Pascoe Coll. 93–60”, “Margites
egenus Pasc. NChina” (Fig.215).
Morphological notes. According to the original
description [Pascoe, 1858], the body length of the holotype
is “9 lines”, i.e. about 19 mm, while based on my own
measurements, the body length is 16.1 mm, the humeral
width is 4.2 mm. According to Plavilstshikov [1931], the
body length of the beetles of this species is 12–18mm.
Distribution. China.
Margites fulvidus (Pascoe, 1858)
(Figs 82, 83, 99, 103, 107, 216)
Cerambyx fulvidus Pascoe, 1858: 236 (“China Borealis”). Type
locality: Northern China (according to the original description and
the label of the holotype).
Pachydissus (?) fulvidus: Bates, 1873: 152.
Pachydissus (Margites) fulvidus: Gahan, 1891: 26.
Margites fulvidus: Aurivillius, 1912: 59; Winkler, 1929: 1142;
Plavilstshikov, 1931: 90; Gressitt, 1951: 144; Ohbayashi, 1964: 38;
Kojima, Hayashi, 1969: 48, pl.15, fig.5; Lee, 1982: 28, pl.4, fig.60;
Hua, 2002: 214; Chou, 2004: 140; Hua et al., 2009: 41, fig. 484
(possibly wrong determination), 172; Wang, Hua, 2009: 174.
Margites (Margites) fulvidus: Kusama, Takakuwa, 1984: 255,
pl.27, figs185, 185a; Catalogue..., 2010: 161.
Material. 1♀, holotype, by monotypy (BMNH) (Fig.83), “N. China”,
“Cerambyx fulvidus Pasc[oe]. Type”, “Type”, “Pascoe Coll. 93–60”, “Margites
fulvidus Pasc. N. China” (Fig. 216); 1♂ (ZMMU), “China, Ningpo [now
Ningbo], Coll. J.Clermont”, “Margites fulvidus Psc., N.Plavilstshikovdet.”;
2♀ (cSM), China, Shaanxi Prov., Haozhenzi, 1350–2000m, 27.05–8.06.1999
(leg.S.Murzin), “Margites fulvidus (Pasc.), S.Murzindet. 1999”; 1♂ (cSM),
China, Shaanxi Prov., Taibaishan Nat. Forest Park, 1350 m, 10.06.1999
(leg.M. Murzin), “Margites fulvidus (Pascoe, 1858) ♂ det.A.Miroshnikov
2018”; 1♂ (NHMD) (Fig.82), Taiwan, DatunMt., 22.06.1997 (leg.J.Chen),
“Margites fulvidus (Pascoe), Ole Mehl det.”; 1♂, 2♀ (NHMD), Japan, Amami
Ooshima Isl., Yuwan, 27.06.1978 (leg. N. Yamamoe), “Margites fulvidus
(Pascoe), Ole Mehl det.”.
Morphological notes. Body length of holotype
17.2mm, humeral width 4.25mm; in the specimens I have
studied (inaddition to the holotype) 14.8–16mm and 3.8–
4mm, respectively. According to Plavilstshikov [1931], the
body length is 12–18mm.
Distribution. China (including Taiwan), Korea, Japan.
Margites exiguus (Gahan, 1894)
Pachydissus (Margites) exiguus Gahan, 1894: 10. Ty pe
locality: Burma (now Myanmar), Mandalay (according to the
original description).
Margites exiguus: Gahan, 1906: 137 (Burma: “Pegu:
arawaddy; Mandalay”); Plavilstshikov, 1931: 89; Roonwal, 1954:
71, 81 (larvae in “Anogeissus acuminata and Aporosa villosula”).
Remarks. is species is known to me only from the
original description, as well as from the Gahanʼs monograph
[1906]. Dr.MaxwellV.L. Barclay kindly provided me with
the type specimens of all Asian Margites species kept under
his care at BMNH for study, but the type of M. exiguus
was absent among them. ere is no photograph in the
collection of Mr.Luboš Dembický (Brno, Czech Republic)
either, a person who kindly provided me with his pictures of
the types of all Margites species available to him, including
the types he photographed in BMNH.
One male (with a body length of 19mm) (Fig.92)
from BMNH that I have revised has the following
labels: “Siam. 1930 W.R.S. Ladell”, “Bangkok, March
1930, at light”, “9.1784”, “Margites exiguus Gah., fr[om].
description. D.J.Atkinsondet. 1948”. However, it shows
no clear differences from the Laotian specimens
identified by J.L.Gressitt as Margites grisescens Pic, 1937
(see below).
Morphological notes. Body length 11–16 mm
[Gahan, 1894, 1906].
Distribution. Myanmar; has also been recorded from
India [Plavilstshikov, 1931].
Margites modicus Gahan, 1906
(Figs 84, 100, 104, 108, 113, 217)
Margites modicus Gahan, 1906: 138. Type locality: India,
Nilgiri Hills (according to the original description and the label of
the lectotype). Aurivillius, 1912: 59; Weigel, 2006: 498; Kariyanna
et al., 2017: 31.
Margites (Margites) modicus: Catalogue..., 2010: 161.
Material. 1♂, lectotype, here designated (BMNH) (Fig.84), “India,
Nylghirri” [= Nilgiri], “Margites modicus Gahan, Type”, “Type”, “Fry Coll.
1905.100”, “26306” (Fig.217), “Lectotypus ♂ Margites modicus Gahan, 1906,
A.Miroshnikovdes., 2018”.
Morphological notes. Body length 13–17 mm,
humeral width 3.75–5 mm [Gahan, 1906], thereby the
lectotype is 14.3mm and 3.8mm, respectively.
Remarks. Based on the original publication [Gahan,
1906], this species was described from no less than three
specimens of both sexes. As noted above, Dr.MaxwellV.L.
Barclay kindly provided me with the type specimens of
all Asian Margites species kept under his care at BMNH
for study, but there is only a single type specimen of
M.modicus among them.
Distribution. India (Tamil Nadu, Maharashtra, Uttar
Pradesh); has also been recorded from Nepal [Weigel,
2006].
Margites auratonotatus Pic, 1923
(Figs 85, 86, 213, 214)
Margites auratonotatus Pic, 1923a: 7 (“Chine”). Type locality:
China, Xujiahui (according to the original description and the
labels of the syntypes). Winkler, 1929: 1142; Plavilstshikov, 1931:
89 (as a species with the dubious differences from M.egenus and
M.exiguus); Gressitt, 1951: 143 (M.egenus?= M.auratonotatus);
Hua, 2002: 214; Hua et al., 2009: 41 (fig. 482), 172; Wang, Hua,
2009: 174.
Margites (Margites) auratonotatus: Catalogue..., 2010: 161.
Material. 1♂, syntype (MNHN) (photograph; Fig. 85), “Zi-ka-wei
[= Xujiahui], 10.5.[19]23”, “Margites auratonotatus Pic”, “Type”, “Muséum
Paris, Coll. E. Licent”, “Holotype” (incorrect label) (Fig. 213); 1♀, syntype
(MNHN) (photograph; Fig. 86), “Zi-ka-wei [= Xujiahui], 11.5.[19]22”,
“Margites auratonotatus Picn.sp.”, “Type”, “Muséum Paris, Coll. E.Licent”,
“P. 33”, “Paratype” (incorrect label) (Fig. 214); 1♀ (photograph), China,
“Chekiang, Chusan [modern transliteration: Zhejiang, Zhoushan],
Musée Heude”, “16–6–[19]31, O. Piel, coll.”, “Margites auratonotatus Pic”
(photograph).
Morphological notes. B ody length of male and female
syntypes 16.4 or 16.5 mm, respectively (Dr. Gérard L.
Tavakilian, personal communication).
Distribution. China (including Taiwan).
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 215
Figs 81–86. Margites Gahan, 1891, habitus, dorsal view.
81 – M. egenus (Pascoe, 1858); 82–83 – M. fulvidus (Pascoe, 1858); 84 – M. modicus Gahan, 1906; 85–86 –M. auratonotatus Pic, 1923 (photographs
by Gérard L. Tavakilian). 81, 83 – holotypes; 84 – lectotype; 85–86 – syntypes; 81, 83, 86 – females; 82, 84–85 –males.
Рис. 81–86. Margites Gahan, 1891, общий вид сверху.
81 – M. egenus (Pascoe, 1858); 82–83 – M. fulvidus (Pascoe, 1858); 84 – M. modicus Gahan, 1906; 85–86 –M. auratonotatus Pic, 1923 (фотографии
Ж. Тавакиляна). 81, 83 – голотипы; 84 – лектотип; 85–86 – синтипы; 81, 83, 86 – самки; 82, 84–85 –самцы.
216 A.I. Miroshnikov
Figs 87–92. Margites Gahan, 1891, habitus, dorsal view.
87–88 – M. luteopubens Pic, 1926 (88 – photograph by Gérard L. Tavakilian); 89 – M. lajoyei Pic, 1926 (photograph by Gérard L. Tavakilian); 90–92 –
M. grisescens Pic, 1937 (90–91 – from Laos, 92 – from ailand). 88–89 – holotypes; 87, 89, 91–92 – males; 88, 90 –females.
Рис. 87–92. Margites Gahan, 1891, общий вид сверху.
87–88 – M. luteopubens Pic, 1926 (88 – фотография Ж. Тавакиляна); 89 – M. lajoyei Pic, 1926 (фотография Ж. Тавакиляна); 90–92 – M. grisescens
Pic, 1937 (90–91 – из Лаоса, 92 – из Таиланда). 88–89 – голотипы; 87, 89, 91–92 – самцы; 88, 90 –самки.
Figs 93–97. Margites Gahan, 1891, habitus, dorsal view.
93–94 – M. mucidus Holzschuh, 1995; 95 – M. pumilus Holzschuh, 1999; 96 – M. minutulus Holzschuh, 2008; 97 – M. alutaceus Holzschuh, 2006. 93,
95–97 – holotypes; 93, 95–96 – males; 94, 97 –females; 93, 95–97 – after Holzschuh [1995, 1999, 2006, 2008], photographs by Luboš Dembický.
Рис. 93–97. Margites Gahan, 1891, общий вид сверху.
93–94 – M. mucidus Holzschuh, 1995; 95 – M. pumilus Holzschuh, 1999; 96 – M. minutulus Holzschuh, 2008; 97 – M. alutaceus Holzschuh, 2006. 93,
95–97 – голотипы; 93, 95–96 – самцы; 94, 97 –самки; 93, 95–97 – по [Holzschuh, 1995, 1999, 2006, 2008], фотографии Л. Дембицкого.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 217
Figs 98–101. Margites Gahan, 1891, habitus, lateral view.
98 – M. egenus (Pascoe, 1858); 99 – M. fulvidus (Pascoe, 1858); 100 – M. modicus Gahan, 1906; 101 – M. grisescens Pic, 1937 (from ailand). 98–99 –
holotypes; 100 – lectotype; 98–99 – females; 100–101 –males.
Рис. 98–101. Margites Gahan, 1891, общий вид сбоку.
98 – M. egenus (Pascoe, 1858); 99 – M. fulvidus (Pascoe, 1858); 100 – M. modicus Gahan, 1906; 101 – M. grisescens Pic, 1937 (из Таиланда). 98–99 –
голотипы; 100 – лектотип; 98–99 – самки; 100–101 –самцы.
218 A.I. Miroshnikov
Margites luteopubens Pic, 1926
(Figs 87, 88, 218)
Margites luteopubens Pic, 1926a: 23. Type locality: China,
Yunnan (according to the original description and the label of
the holotype). Winkler, 1929: 1142; Plavilstshikov, 1931: 90 (syn.
pro Margites fulvidus; wrong synonymy; see also below); Gressitt,
1951: 144; Hua, 1984: 60; 2002: 214; Hua et al., 2009: 41, fig.483
(possibly wrong determination), 172; Wang, Hua, 2009: 174;
Weigel et al., 2013: 72, 161, pl. 6, fig.f; Nga et al., 2014: 435.
Margites (Margites) luteopubens: Gressitt, Rondon, 1970: 77;
Catalogue…, 2010: 161.
Material. 1♀, holotype, by monotypy (MNHN) (photograph;
Fig. 88), China, “Yunnan”, “Margites luteopubens n. sp.”, “Type”,
“21”, “Museum Paris, Coll. M. Pic”, “Holotype” (Fig. 218);
1♂ (ZIN) (Fig. 87), Vietnam, Hanoi City, 5.04.1962, at light
(leg.O.N.Kabakov), “Margites luteopubens Pic?, Kabakov det. 1985”,
“Margites luteopubens Pic, 1926 ♂ det. A.Miroshnikov 2018”; 2♂
(ZIN), NVietnam, Tam Dao, 5.06.1995 (leg. Gorochov), “Margites
luteopubens Pic, 1926 ♂ det. A. Miroshnikov 2018”; 1♂ (cLD),
NVietnam, 70km NW of Hanoi, Tam Dao, 21°27ʹN/ 105°39ʹE,
900–1200 m, 9–19.05.1998 (leg. L. Dembický, P. Pacholátko),
“Margites luteopubens Pic, 1926 ♂ det. A.Miroshnikov 2018”; 1♂
(ZIN), Vietnam, Lao Cai Prov., Sa Pa Distr., Fan Si PanMt., 22°20ʹN/
103°46ʹE, 1900–2500m, 20.04–9.05.1999 (leg.N.Orlov), “Margites
luteopubens Pic, 1926 ♂ det. A.Miroshnikov 2018”; 1♀ (NHMD),
NVietnam, Tam Dao Nat. Park, 21°16ʹ16ʺN/ 105°23ʹ17ʺE, 900m,
4–5.05.2005 (leg. A. Kun), “Margites luteopubens Pic, 1926 ♀
det. A. Miroshnikov 2018”; 1♀ (NHMD), Laos, Hua Phan Prov.,
Ban Saleui, Phou Pan Mt., ~20°12ʹN / 104°01ʹE, 1500–1900 m,
23.04–16.05.2008 (leg.C.Holzschuh), “Margites luteopubens Pic,
det.Holzschuh 2009”.
Morphological notes. Body length 12–20 mm
[Gressitt, Rondon, 1970], thereby the holotype is 17mm
long (Dr.GérardL. Tavakilian, personal communication);
in the specimens I have studied body length 13.1–
17mm, humeral width of 3.3–4.6mm. e indication of
“39mm” [Weigel et al., 2013: 161] is without any doubt
a misprint.
Margites luteopubens is morphologically very similar
to M. fulvidus, thereby both species showing significant
individual variability, which to some extent complicates
their diagnostics. In my opinion, the differences between
these species require a detailed elaboration using a more
extensive material.
Distribution. China (Yunnan), Laos, Vietnam.
Margites lajoyei Pic, 1926
(Figs 89, 219)
Margites lajoyei Pic, 1926b: 76. Type locality: “Cochinchine,
Cap Sain-Jacques” (now Vũng Tàu, southern Vietnam) (according
to the original description and the label of the holotype).
Plavilstshikov, 1931: 90 (syn. pro Margites fulvidus; wrong
synonymy).
Margites (Margites) lajoyei: Catalogue..., 2010: 161.
Material. 1♂, holotype, by monotypy (MNHN) (photograph; Fig.89),
“Cochinchine, Cap St.Jacques”, “Margites lajoyein. sp.”, “Type”, “exLajoye”,
“Museum Paris, Coll. M.Pic”, “Holotype” (Fig.219).
Morphological notes. Body length of holotype
12.7 mm (Dr. Gérard L. Tavakilian, personal
communication).
Distribution. Southern Vietnam; in my opinion, the
record in Yunnan Province, China [Catalogue..., 2010]
requires confirmation and possibly concerns another
species.
Margites grisescens Pic, 1937
(Figs 90–92, 101, 105, 109, 114)
Margites grisescens Pic, 1937: 7. Type locality: northern
Vietnam, “Annam” (according to the original description). Hua,
1984: 60; Nga et al., 2014: 435.
Margites (Margites) grisescens: Gressitt, Rondon, 1970: 78.
Material. 2♂ (BM), “Laos: Borikhane Prov., Paksane”, [?27.III.1962],
“J.A. Rondon Collection Bishop M u s . ”, “Margites (s. str.) grisescens
Pic, J.L. Gressitt det.”; 1♂ (BM) (Fig. 91) (body length 23 mm!), “Laos:
Khammouane Prov., Phon Tiou, 25.2.1964”, “J.A.Rondon Collection Bishop
M u s .”, “Margites(s. str.) grisescens Pic, J.L.Gressittdet.”; 1♀ (BM), “Laos:
Wapikhamthong Prov., Khong Sedone, 31.3.1965”, “J.A.Rondon Collection
Bishop M u s .”, “Margites(s.str.) grisescens Pic, J.L.Gressittdet.”; 1♀ (BM),
“Laos: Ban Van Heue, 20km E of Phou-kow-kuei [=Phou Khao Khouei],
15–31.V.1965”, “J.A. Rondon Collection Bishop M u s .”, “Margites (s. str.)
grisescens Pic, J.L . Gressitt det.”; 1♂, 1♀ (BM), “Laos: Vientiane
Prov., Vientiane, 24.II.1966”, “J.A . Rondon Collection Bishop M u s . ”,
“Margites(s.str.) grisescens Pic, J.L.Gressittdet.”; 1♀ (BM) (Fig.90), “Laos:
Sedone Prov., Pakse, 30.IV.1967”, “J.A. Rondon Collection Bishop M u s . ”,
“Margites(s.str.) grisescens Pic, J.L.Gressittdet.”; 1♀ (cLD), NWailand,
Mae Hong Son Prov., Soppong, 19°27ʹN/ 98°20ʹE, 1200m, 26–28.05.2000
(leg. P. Spáčil), “Margites grisescens Pic, 1937 ♀ det. A. Miroshnikov
2018”, “Compared to the specimens identified by J.L.Gressitt”; 1♂ (cAM),
Nailand, Chiang Rai Prov., Doi Chang env., 640–750m, 19°46ʹ01ʺN/
99°28ʹ11ʺE – 9°47ʹ44ʺN / 99°27ʹ06ʺE, 11–15.05.2013 (leg. I. Melnik),
“Margites grisescens Pic, 1937 ♂ det.A. Miroshnikov 2018”, “Compared
to the specimens identified by J.L .Gressitt”; 1♂, 1♀ (IRSN), Cambodia,
Kampong Speu, Chambok, 11°21ʹ25ʺN / 104°7ʹ9ʺE, 4–8.05.2015, light
trap (leg. J.Constant, V.Sougnez), “Margites grisescens Pic, 1937 [♂ or
♀, respectively] det.A.Miroshnikov 2018”, “Compared to the specimens
identified by J.L.Gressitt”.
Remarks. e type of this species is known to me only
from the original description. Arecent attempt to relocate
it in the Muséum national d’Histoire naturelle, Paris, kindly
undertaken by Dr.GérardL. Tavakilian upon my request,
was unsuccessful, like in the case of the type of Pachydissus
thibetanus (see above). Iwas informed that Pic’s collection
also contained a label, where André Villiers noted to have
never seen the type of M.grisescens.
Margites grisescens is here understood as being
represenred by the specimens identified by J.L. Gressitt
that I have examined.
Morphological notes. Body length 11–15 mm [Pic,
1937; Gressitt, Rondon, 1970]; in the specimens that I have
studied the body length was 12.3–23 mm, the humeral
width between 3.3–5.8mm.
Distribution. Vietnam, Laos; based on the material
studied, M.grisescens is being recorded here from ailand
and Cambodia for the first time.
Margites mucidus Holzschuh, 1995
(Figs 93, 94)
Margites mucidus Holzschuh, 1995: 17. Type locality:
Northern ailand, Chiang Mai, Doi Suthep Mt., 1100 m
(according to the original description).
Material. 1♂, holotype (cCH) (photograph; Fig. 93); 1♀ (cAM)
(Fig. 94), Laos, Xaignabouri City, 16–18.04.2005 (unknown collector),
“Margites mucidus Holzschuh, 1995 ♀ det.A.Miroshnikov 2018”; 1♀ (cLD),
“Laos”, “Margites mucidus Holzschuh, 1995 ♀ det.A.Miroshnikov 2018”.
Morphological notes. Body length 15.5–19.3 mm
[Holzschuh, 1995]; in the specimens I have studied the
body length was 18.8–19.4mm, the humeral width between
4.6–4.8mm.
Distribution. ailand; based on the material studied,
M.mucidus is being recorded here from Laos for the first
time.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 219
Margites pumilus Holzschuh, 1999
(Fig. 95)
Margites pumilus Holzschuh, 1999: 20. Type locality:
Indonesia, Sumatra, Kebun Sei Kopas, 2°49ʹN / 99°18ʹE, 200 m
(according to the original description). Heffern, 2013: 10.
Material. 1♂, holotype (cCH) (photograph; Fig.95).
Morphological notes. Body length 9.2 mm
[Holzschuh, 1999].
Distribution. Indonesia (Sumatra); has also been
recorded from Borneo [Heffern, 2013].
Margites alutaceus Holzschuh, 2006
(Fig. 97)
Margites alutaceus Holzschuh, 2006: 221. Type locality:
Malaysia, Sabah, Sipitang, Mendolong (according to the original
description). Heffern, 2013: 10.
Material. 1♀, holotype (cCH) (photograph; Fig. 97).
Morphological notes. Body length 26 mm
[Holzschuh, 2006].
Distribution. Eastern Malaysia.
Margites minutulus Holzschuh, 2008
(Fig. 96)
Margites minutulus Holzschuh, 2007: 200 (nom. nudum).
Margites minutulus Holzschuh, 2008: 239. Type locality:
India, Karnataka, 20km SE Sagar (14°03ʹ49ʺN/ 75°05ʹ23ʺE), 600m
(according to the original description). Kariyanna et al., 2017: 31.
Material. 1♂, holotype (cCH) (photograph; Fig.96).
Morphological notes. Body length 8.2–9.2 mm
[Holzschuh, 2008].
Distribution. India.
Genus Laomargites Pic, 1923, stat. rest.
Laomargites Pic, 1923a: 8; Gressitt, Rondon, 1970: 78
(Margites subgen.).
Type species: Laomargites singularis Pic, 1923, by
monotypy.
Diagnosis. is genus considered by some researchers
as a subgenus of the genus Margites differs clearly from it
in the structure of the eyes, the sculpture in the area of the
base of the antennae, the pronotal sculpture, the structure
of the femora and tibiae, as well as by some other traits
indicated below.
When detailing the structure of Laomargitesstat.rest.,
the following features must be noted as being characteristic
of this genus: eyes, albeit strongly convex, but in general
significantly less strongly developed compared to Margites,
with both upper and lower lobes clearly more narrow, as
in Figs 110, 111, 115, 116 (cf. Figs 106–109, 112–114),
with a greater distance between both upper and lower
lobes; submentum strongly transverse (vssubmentum only
moderately transverse in Margites); bases of antennae with
a very strong, sharply protruding bordure embracing the
antennal cavities over most of their perimeter and forming
a wide and deep depression between inner margins of
antennal bases, as in Figs 110, 111 (vsbases of antennae
usual in structure, with neither a very strong bordure
nor a deep depression between their inner margins in
Margites, as in Figs 106–109); antennomere 2 both in
male and female distinctly or very clearly longitudinal,
in male somewhat inflated (vs antennomere 2 distinctly
transverse or subequal in length and width, in male
sometimes barely or slightly longitudinal, but not inflated
in Margites); antennomeres 3 and 4 in male distinctly
inflated; pronotum (Figs 110, 111) in apical part with a
very well-expressed, peculiar, sculptural formation in the
form of a scutum (somewhat reminding of Imbrius Pascoe,
1866, but larger), sharply bound from behind by a ledge
and along margin framed by a bordure, mainly with coarse,
mostly longitudinal wrinkles, folds and a clear, more or
less sharp puncturation; behind this formation with very
coarse, longitudinal and obliquely longitudinal, more or
less long, partly sinuous folds in places connected with
each other by transverse folds, in general forming a large
fragment of discal sculpture extending almost to base of
pronotum; lateral to this fragment with a coarse and very
coarse, mostly longitudinal, cellular sculpture (vsno similar
sculpture of pronotum is observed in Margites – Figs81–
97, 102–105); elytra with a well-developed recumbent
setation, but weakly hiding their puncturation, and, in
addition, sometimes with very clearly expressed, suberect,
short setae (vs elytra sometimes with a dense recumbent
setation, strongly hiding their puncturation, but without
clearly expressed, suberect, short setae in Margites); femora
with a gentle rugose sculpture and a small, more or less
dense puncturation (vs at least profemora, especially on
ventral side, with a rough or moderately coarse, dense and
confluent, rugose puncturation; meso- and metafemora
usually with a less coarse sculpture, but sometimes with
a sculpture more or less similar to that of profemora,
especially on mesofemora, in Margites); tibiae without
carina (vstibiae with a very clear or less distinct, sometimes
partly or predominantly poorly expressed carina, this
nonetheless being present along each side in Margites).
Composition. e genus includes two species, one of
which is described as new.
Distribution. Indochina (Laos, Vietnam, ailand).
Laomargites singularis Pic, 1923, comb. rest.
(Figs 110, 115, 117,118, 120,121, 123,
124, 126–129, 131, 133, 135, 220)
Laomargites singularis Pic, 1923a: 8. Type locality: Laos,
[Ban] Paklung (according to the original description and the label
of the holotype).
Margites (Laomargites) singularis: Gressitt, Rondon, 1970: 78.
Margites singularis: Hua, 1984: 60.
Material. 1♂, holotype, by monotypy (MNHN) (Fig. 128), “Laos,
Paklung, le 8.III.1920, R.Vitalis de Salvaza”, “Laomargitesn. g. singularisn.sp.”,
“Type”, “Museum Paris, Coll. M.Pic”, “3000”, “Holotype” (Fig.220); 2♂ (BM),
“Laos: Khammouane Prov., Phon Tiou, 25.2.1964”, “J.A.Rondon Collection
Bishop Mus.”, “Margites (Laomargites) singularis (Pic), J.L. Gressitt det.”;
1♂ (BMNH) (Fig.127), “Laos: Khammouane Prov., Phon Tiou, 17.III.1965”,
“J.A. Rondon Collection Bishop Mus.”, “Margites (Laomargites) singularis
(Pic), J.L. Gressitt det.”; 1♀, 1♀ (Fig. 129) (BM), “Laos: Vientiane Prov.,
Nongtevada, 15.II.1965, 17.III.1965”, “J.A. Rondon Collection Bishop Mus.”,
“Margites (Laomargites) singularis (Pic), J.L. Gressitt det.”; 1♂, 1♀ (BM),
“Laos: Vientiane Prov., Ban Van Eue, 16.III.1966”, “J.A.Rondon Collection
Bishop Mus.”, “Margites (Laomargites) singularis (Pic), J.L.Gressittdet.”; 2♂
(BM), “Laos: Vientiane Prov., Phou Kou Khouei, 19.III.1966, 15.IV.1966”,
“J.A. Rondon Collection Bishop Mus.”, “Margites (Laomargites) singularis
(Pic), J.L. Gressitt det.”; 1♂ (BMNH) (Fig. 126), “Laos: Vientiane Prov.,
Nongtevada, 2.II.1967”, “J.A. Rondon Collection Bishop Mus.”, “Margites
(Laomargites) singularis (Pic), J.L.Gressittdet.”; 1♀ (BM), “Laos: Sayaboury
Prov., Sayaboury, 25.III.1966”, “J.A. Rondon Collection Bishop Mus.”,
220 A.I. Miroshnikov
Figs 102–111. Margites Gahan, 1891 and Laomargites Pic, 1923, stat. rest., head, dorsal view, and pronotum.
102, 106 – M. egenus (Pascoe, 1858); 103, 107 – M. fulvidus (Pascoe, 1858); 104, 108 – M. modicus Gahan, 1906; 105, 109 – M. grisescens Pic, 1937
(from ailand); 110 –L. singularis Pic, 1923, comb. rest.; 111 –L. fedorenkoi sp. n. 102–103, 106–107, 111 – holotypes; 104, 108 – lectotype; 102–103,
106–107– females; 104–105, 108–111 –males.
Рис. 102–111. Margites Gahan, 1891 и Laomargites Pic, 1923, stat . rest., голова сверху и переднеспинка.
102, 106 – M. egenus (Pascoe, 1858); 103, 107 – M. fulvidus (Pascoe, 1858); 104, 108 – M. modicus Gahan, 1906; 105, 109 – M. grisescens Pic, 1937
(из Таиланда); 110 –L. singularis Pic, 1923, comb. rest.; 111 – L. fedorenkoi sp. n. 102–103, 106–107, 111 – голотипы; 104, 108 – лектотип; 102–103,
106–107– самки; 104–105, 108–111 –самцы.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 221
Figs 112–125. Margites Gahan, 1891 and Laomargites Pic, 1923, stat. rest.
112 – M. egenus (Pascoe, 1858); 113 – M. modicus Gahan, 1906; 114 – M. grisescens Pic, 1937 (from ailand); 115, 117–118, 120–121,
123–124–L.singularis Pic, 1923, comb. rest.; 116, 119, 122, 125 –L. fedorenkoi sp. n. 112, 116, 119, 122, 125 – holotypes; 113 – lectotype; 112 – female;
113–125–males; 112–116 – head, ventral view; 117–119 – apex of elytra, dorsal view (at an angle of about 45 degrees); 120–122 – apical part of elytra , lateral
view; 123–125– apical part of elytra, dorsal view.
Рис. 112–125. Margites Gahan, 1891 и Laomargites Pic, 1923, stat . rest.
112 – M. egenus (Pascoe, 1858); 113 – M. modicus Gahan, 1906; 114 – M. grisescens Pic, 1937 (из Таиланда); 115, 117–118, 120–121,
123–124–L.sing ularis Pic, 1923, comb. rest.; 116, 119, 122, 125 –L. fedorenkoi sp. n. 112, 116, 119, 122, 125 – голотипы; 113 – лектотип; 112 – самка;
113–125 –самцы; 112–116 – голова снизу; 117–119 – вершина надкрылий сверху (под углом примерно 45 градусов); 120–122 – вершинная часть
надкрылий сбоку; 123–125 – вершинная часть надкрылий сверху.
222 A.I. Miroshnikov
Figs 126–136. Laomargites Pic, 1923, stat. rest., habitus, dorsal view, and male genitalia.
126–129, 131, 133, 135 –L. singularis Pic, 1923, comb. rest.; 130, 132, 134, 136 –L. fedorenkoi sp. n. 128, 130 – holotypes; 126–128, 130 – males;
129 – female; 131–132 – apical part of penis, ventral view; 133–134 – apical part of tegmen, ventral view; 135–136 – apical part of tergite 8, dorsal view.
Рис. 126–136. Laomargites Pic, 1923, stat. rest., общий вид сверху и гениталии самца.
126–129, 131, 133, 135 –L. singularis Pic, 1923, comb. rest.; 130, 132, 134, 136 –L. fedorenkoi sp. n. 128, 130 – голотипы; 126–128, 130 – самцы;
129 – самка; 131–132 – вершинная часть пениса снизу; 133–134 – вершинная часть тегмена снизу; 135–136 – вершинная часть 8-го тергита сверху.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 223
“Margites (Laomargites) singularis (Pic), J.L.Gressittdet.”; 1♀ (BM), “Laos:
Tonpheng, 15.V.1966”, “J.A. Rondon Collection Bishop Mus.”, “Margites
(Laomargites) singularis (Pic), J.L.Gressittdet.”; 1♂ (IRSN), “Laos, Takek,
Coll. Le Moult”, “Laomargites singularis Pic, 1923 ♂ det. A. Miroshnikov
2018”.
Morphological notes. Body length 11–20 mm
[Gressitt, Rondon, 1970]; in the specimens I have studied
the body length was 13.1–22.2 mm, the humeral width
between 3.45–5.7 mm, thereby the holotype is 17.8 mm
and 4.6mm, respectively.
Distribution. Laos, ailand.
Laomargites fedorenkoi Miroshnikov, sp. n.
(Figs 111, 116, 119, 122, 125, 130, 132, 134, 136)
Material. Holotype, ♂ (cAM) (Fig. 130): Vietnam, Kon Tum Prov.,
Kon Plong Distr., Dak Khe River, 14°43ʹ20ʺN / 108°18ʹ58ʺE, 1030 m,
8–23.04.2015, at light (leg.D.Fedorenko).
Diagnosis. is new species is very similar to
L.singulariscomb.rest., but differs clearly by the presence
of numerous, very well-expressed, suberect, short setae on
the elytra, as in Figs119, 122, 125, and in the conformation
of the male genitalia (Figs132, 134, 136), in particular, the
widely spaced parameres, as in Fig. 134 (cf.Figs117,118,
120,121, 123,124, 131, 133,135).
Description. Male. Body length 12 mm, humeral width
3.1mm. Coloration of integument mainly red-brown, thereby head
dorsally and mostly pronotum darkest; eyes, partly mandibles and
most of pronotal coarse and rough folds black.
In general, structure of head, including areas of antennal
bases, as in L. singularis comb. rest. (see Diagnosis of genus
above); antennae longer than body, about reaching the apex of
elytra by antennomere 8; length ratio of antennomeres 1–11,
29:11: 26: 21: 29: 31: 34: 31: 31: 27:36; antennomere1 with
a heterogeneous, dense, partly rough puncturation, noticeably
impressed in basal part dorsally; antennomere 2 very clearly
longitudinal; antennomeres2–4 inflated in apical part.
Pronotum subequal in length and width; at base barely
wider than at apex; with a sharp constriction both in front of base
and near apex; in general, structure of pronotum, including its
sculpture, similar to L. singularis comb.rest. (see Diagnosis of
genus above).
Scutellum triangular, with a small, partly quite clear
puncturation.
Elytra predominantly nearly parallel-sided starting from
base, 2.5times as long as humeral width; with both a moderately
rough, more or less regular and very small puncturation; apical
external angle widely rounded, sutural angle narrowly rounded.
Prosternum in apical part with well-expressed transverse
folds; prosternal process without apical tubercle; mesosternal
process between coxae 2.7 times as wide as prosternal process,
without tubercle dorsally; metasternum and sternites with a small,
clear, dense puncturation; metasternum with a well-expressed
median groove; last (visible) sternite truncate at apex; last (visible)
tergite widely rounded apically.
Legs moderately long; tibiae without carina along each side;
metatarsomere 1 noticeably shorter than metatarsomeres2 and3
combined.
Recumbent setation, except for elytra, mainly greyish, partly
with yellowish tint, that of elytra yellowish golden, weakly masking
their puncturation; elytra, in addition, with numerous, well-
expressed, suberect, short, yellowish golden setae; more or less
long, erect, light setae mainly developed on pronotum and head.
Etymology. I am pleased to dedicate this new species
to my colleague and friend, Dr. Dmitry N. Fedorenko
(Institute for Problems of Ecology and Evolution, Russian
Academy of Sciences, Moscow, Russia), who has collected
in Vietnam a rich and very valuable material on Coleoptera,
including cerambycids.
Distribution. Vietnam.
Key to species of Laomargites stat. rest.
1. Elytra, in addition to recumbent setation, with sparse,
barely protruding, short setae very poorly visible
against general background, as in Figs 117, 118,
120, 121, 123, 124; parameres close together, as in
Fig.133 ...................................... L. singularis comb. rest.
– Elytra, in addition to recumbent setation, with numerous,
strongly protruding, short setae very well visible
against general background, as in Figs119, 122, 125;
parameres widely spaced, as in Fig.134 ..........................
............................................................... L. fedorenkoi sp. n.
Genus Dymasius J. omson, 1864
Dymasius J. omson, 1864: 234; Lacordaire, 1868: 261;
Gemminger in Gemminger, Harold, 1872: 2803; Gahan, 1891: 22;
1906: 139; Aurivillius, 1912: 60; Plavilstshikov, 1931: 92; Gressitt,
1951: 144; Kojima, Hayashi, 1969: 48; Gressitt, Rondon, 1970: 78;
Kusama, Takakuwa, 1984: 284; Catalogue..., 2010: 160; Heffern,
2013: 9; Kariyanna et al., 2017: 29; Miroshnikov, 2017: 199.
Type species: Dymasius strigosus J. omson, 1864, by
monotypy.
Dymasius strigosus J. omson, 1864, sp. rest.
(Figs 138,139, 161, 163, 165)
Dymasius strigosus J. omson, 1864: 234. Type locality:
“India” (according to the original description and the label of the
holotype). Lacordaire, 1868: 262; Gemminger in Gemminger,
Harold, 1872: 2803; Gahan, 1891: 22; Gahan, 1906: 139 (syn. pro
D. macilentus; wrong synonymy); Aurivillius, 1912: 60 (syn. pro
D.macilentus; wrong synonymy); Gressitt, Rondon, 1970: 78 (syn.
pro D. macilentus; wrong synonymy); Kusama, Takakuwa, 1984:
254 (syn. pro D.macilentus; wrong synonymy); Catalogue..., 2010:
160 (syn. pro D. macilentus; wrong synonymy); Heffern, 2013:
9 (syn. pro D. macilentus; wrong synonymy); Kariyanna et al.,
2017: 29 (syn. pro D.macilentus; wrong synonymy); Miroshnikov,
2017: 199 (the synonymy D. macilentus= D. strigosus requires
unequivocal evidence).
Material. 1♂, holotype, by monotypy (MNHN) (photographs); 1♂
(BMNH), “Ceylon”, “Bowr. Chevr. 63-47*”, “Dymasius strigosus oms.
♂, comp. with type”; 1♂ (BMNH), “Ceylon”, “Bowr. Chevr. 63-47*”; 1♂
(Fig. 138), 3♀ (BMNH), “Ceylon. G. Lewis. 1910–320”, “Dikoya. 3,800–
4,200ft., 6.XII.[18]81–16.I.[18]82”; 1♀ (Fig.139), 1♂, 2♀ (BMNH), “Ceylon”,
“Fry Coll. 1905.100”, “Ex Mus. Parry”; 1♀ (BMNH), “Ceylon”, “Pascoe Coll.
93–60”; 1♀ (BMNH), “Ceylon” (upperside), “62/84”(underside); 1♂, 1♀
(BMNH), “Ceylon”.
Comparative material. Dymasius macilentus (Pascoe, 1859): 1♂,
holotype, by monotypy (BMNH) (Fig.137), “Ceylon” (upperside), “59.106”
(underside), “Cerambyx macilentus Pascoe, Type”, ”Type”.
Remarks. Since based on a comparison of the holotype
male of D.strigosus and the holotype male of D.macilentus
(Pascoe, 1859) some significant morphological differences
were recently revealed between them, I concluded that
the synonymy D. macilentus= D. strigosus [Gahan,
1906; Aurivillius, 1912; Gressitt, Rondon, 1970; Kusama,
Takakuwa, 1984; Catalogue..., 2010 and others] required
undeniable evidence [Miroshnikov, 2017]. I thereby was
able to revise the holotype of D.strigosus, kept in MNHN,
based only on high-quality photographs.
Now, however, I have been able to examine in detail
5males and 9females of D.strigosus (kept in BMNH) kindly
provided by Dr.MaxwellV.L. Barclay. e results of this
study confirmed significant differences I showed previously
to be observed between D. macilentus and D. strigosus
in the structure of the male antennae and of the elytral
224 A.I. Miroshnikov
Figs 137–142. Dymasius J. omson, 1864, habitus, dorsal view.
137 – D. macilentus (Pascoe, 1859); 138–139 – D. strigosus J. omson, 1864; 140 – D. tatianae sp.n.; 141 –D. indigus Holzschuh, 2008; 142 – D.
nodifer Holzschuh, 2005. 137, 140–142 – holotypes; 137–138, 140, 142 – males; 139, 141 –females; 141–142 – after Holzschuh [2005, 2008], photographs
by Luboš Dembický.
Рис. 137–142. Dymasius J. omson, 1864, общий вид сверху.
137 – D. macilentus (Pascoe, 1859); 138–139 – D. strigosus J. omson, 1864; 140 – D. tatianae sp.n.; 141 –D. indigus Holzschuh, 2008; 142 – D. nodifer
Holzschuh, 2005. 137, 140–142 – голотипы; 137–138, 140, 142 – самцы; 139, 141 – самки; 141–142 – по [Holzschuh, 2005, 2008], фотографии Л.Дембицкого.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 225
Figs 143–148. Dymasius J. omson, 1864, habitus, dorsal view.
143 – D. nodifer Holzschuh, 2005 (from ailand); 144–147 – D. simplex Gressitt et Rondon, 1970 (146–147 – from ailand); 148 –D. prominor
Gressitt et Rondon, 1970. 144, 148 – holotypes; 145 – paratype; 143, 145, 147–148 – females; 144, 146 –males.
Рис. 143–148. Dymasius J. omson, 1864, общий вид сверху.
143 – D. nodifer Holzschuh, 2005 (from ailand); 144–147 – D. simplex Gressitt et Rondon, 1970 (146–147 – из Таиланда); 148 – D. prominor
Gressitt et Rondon, 1970. 144, 148 – голотипы; 145 – паратип; 143, 145, 147–148 – самки; 144, 146 –самцы.
226 A.I. Miroshnikov
apex. us, in the male of D. strigosus, antennomere 3 is
1.62–1.68 or 1.45–1.6 times as long as antennomere 1
and antennomere4, respectively, antennomere5 is 1.54–
1.6times as long as antennomere4, while in the holotype
male of D. macilentus, antennomere 3 is only 1.36 or
1.32times as long as antennomere1 and antennomere4,
respectively, antennomere 5 is only 1.32 times as long
as antennomere 4. e male antennae of D. strigosus
(Fig. 138), including the holotype, are stable in being
significantly longer than those of the holotype male of
D.macilentus (Fig.137), in each of these species the shape
of the inflated apical part of antennomeres3–5 is thereby
somewhat peculiar. e sculpture of male antennomere1
of D.strigosus is more or less variable, but always, at least
partly, clearly coarser than that in the holotype male of
D.macilentus. Both in the male and female of D.strigosus,
a tooth at the apical external angle of the elytra is somewhat
variable in length and sometimes considerably drawn
towards the external side, but is always clearly shorter than
that in the holotype male of D.macilentus.
Besides this, all examined specimens of D. strigosus
are characterized by a combination of dark red-brown
and red-brown coloration of the integument (except for
the eyes), including the elytra, antennae and legs, while in
the holotype of D.macilentus, the dorsum, antennae and
mostly legs black, only the pronotum is in places with weak
dark reddish brown tint. e recumbent light setation of the
prosternum in D.strigosus is sparser than in D.macilentus.
ere are also some clear differences in the structure of the
male genitalia of these species (Figs161–166).
As a result, Dymasius strigosus J. omson,
1864,sp.rest., non syn. pro Dymasius macilentus (Pascoe,
1859).
According to the original description and label
[Miroshnikov, 2017: 231, fig. 452], the holotype of
D.strigosus comes from India. As all non-type specimens
of this species I have studied derive from Sri Lanka, it
seems very likely that the holotype had also been caught in
Sri Lanka, not India.
In the specimens of D. strigosus I have studied body
length 25.3–34.8mm, humeral width 5.8–8.6mm.
Dymasius tatianae Miroshnikov, sp. n.
(Fig. 140)
Dymasius indigus (non Holzschuh, 2008): Miroshnikov,
2017: 204, figs241–243.
Material. Holotype, ♂ (NHMD) (Fig. 140): EMalaysia, Sabah, Trus
MadiMt., 03.2004 (local collector), “Dymasius mandibularis, Ole Mehl det.
2014”, “Dymasius indigus Holzschuh, 2008 ♂ det.A.Miroshnikov 2017”.
Diagnosis. is new species is very similar to
D. indigus Holzschuh, 2008, but differs clearly by the
generally darker coloration of the integument; the
coloration of the recumbent setation at least of the dorsum,
antennae and legs as in Fig. 140; the somewhat peculiar
sculpture and pattern of the recumbent light setation of the
pronotum, as in Fig.140; the longer median groove of the
head dorsally, well-developed not only between the eyes,
but also partly on the vertex; the significantly more strongly
developed recumbent light setation of the head behind the
upper lobes of the eyes, as in Fig. 140; and possibly the
larger body, at least so on the average (cf.Fig. 141).
Description. Male. Body length 30.1 mm, humeral width
7.1 mm. Head dorsally, eyes, pronotum, partly mandibles, basal
antennomeres and tarsi black; remaining parts combines red-
brown and dark reddish brown tones, thereby elytra dark reddish
brown.
Head with a very deep median groove between upper
lobes of eyes, partly, and on vertex; antennal tubercles very well-
developed; eyes moderately convex; submentum mainly with a
more or less small puncturation; antennae much longer than body,
almost reaching the apex of elytra by antennomere6; length ratio
of antennomeres1–11, 34 : 11 : 54 : 34 : 72 : 71 : 68 : 64 : 68 :
69: 122; antennomere1 devoid of a cicatrix (apical carina), with
a heterogeneous, partly very coarse sculpture, predominantly on
inner side with coarse transverse folds; antennomere 2 clearly
longitudinal; antennomeres 5–11 very strongly elongated,
especially so last one.
Pronotum barely longitudinal, 1.03times as long as wide;
with a sharper constriction near apex than in front of base; with
coarse and very coarse, partly sinuous, mainly transverse folds.
Scutellum triangular, with an unclear sculpture.
Elytra clearly narrowed towards apex, 2.7 times as long
as humeral width; with a very small dense puncturation; apical
external angle obtuse, well-expressed, sutural angle with a short,
but very clear, sharp tooth.
Prosternum in apical one-third with somewhat rough
transverse folds, in middle part with coarse, partly sinuous,
transverse folds; prosternal process with a very clear apical
tubercle; mesosternal process between coxae noticeably wider than
prosternal process, with a deep median impression; metasternum
and sternites with a very small, but clear, dense puncturation;
metasternum with a well-expressed median groove; both last
(visible) sternite and tergite truncate apically.
Legs long; femora not claviform, without longitudinal carina;
metatarsomere 1 slightly longer than metatarsomeres 2 and 3
combined.
Recumbent setation yellowish, partly with a golden shine,
including on elytra (in D. indigus, recumbent setation yellowish
brown, shiny– “Behaarung anliegend, gelblichbraun glänzend...”
[Holzschuh, 2008: 177]); pronotal setation forming a peculiar
pattern, as in Fig.140, thereby, compared to D.indigus, folds on
disc generally to a lesser degree masked by dense setae and more
strongly visible; elytral setation irregular; basal antennomeres with
numerous, erect, light setae in the form of a sparse gentle brush
(somewhat resembling members of the genus Elydnus Pascoe,
1868); more or less long, erect, light setae mainly developed on
pronotum and head.
Genitalia– see Miroshnikov [2017: 197, figs241–243].
Remarks. Based on the very clear morphological
similarity of the holotype of this new species to the holotype
of D.indigus (based on its picture) and on the origin of both
taxa from one and the same locality, initially I attributed
the former to D.indigus [Miroshnikov, 2017: 197 (figs 241–
243), 204]. However, the results of a more detailed study of
this male show that it should be considered as a separate
species.
Etymology. I am pleased to dedicate this new species
to my wife, TatianaP. Miroshnikova, who, over many years,
selflessly supports my entomological research and provides
an invaluable editing assistance in preparing very numerous
photographs and various other scientific materials.
Distribution. Eastern Malaysia.
Dymasius nodifer Holzschuh, 2005
(Figs 142, 143)
Dymasius nodifer Holzschuh, 2005: 11. Type locality:
Malaysia, Sabah, Trus Madi Mt. (according to the original
description).
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 227
Material. 1♂, holotype (cCH) (photograph; Fig. 142); 1♂ (cAM
ex NHMD), E Malaysia, Sabah, Trus Madi Mt., 03.2003 (local collector),
“Dymasius nodifer Holz., Ole Mehl det. 2007”; 1♀ (NHMD), E Malaysia,
Sabah, Crocker Range, 04.2005 (local collector), “Dymasius nodifer Holz.,
Ole Mehl det. 2006”; 1♀ (cAM ex NHMD), E Malaysia, Sabah, Crocker
Range, 03.2005 (local collector), “Dymasius nodifer Holz., Ole Mehl det.
2007”; 1♂ (NHMD), E Malaysia, Sabah, Trus Madi Mt., 04.2013 (local
collector), “Dymasius nodifer Holz., Mehldet. 2014”; 1♀ (BMNH) (Fig.143),
“Peninsular Siam, Nakon Sri Tamarat, Khao Ram, 750ft., February24th, 1922,
at light, H.M.Pendlebury”, “1927.428”, “318”, “Dymasius nodifer Holzschuh,
2005, ♀, det.A.Miroshnikov 2017”.
Distribution. Until now, this species has only been
known from Borneo [Holzschuh, 2005]. Based on the
material studied, D. nodifer is being recorded here from
ailand, as from Indochina in general, for the first time.
Dymasius simplex Gressitt et Rondon, 1970
(Figs 144–147, 221, 222)
Dymasius (Elydnus) simplex Gressitt et Rondon, 1970: 81.
Type locality: Laos, Borikhane Province, Pakkading (according to
the original description and the label of the holotype).
Dymasius simplex: Miroshnikov, 2017: 183.
Material. 1♂, holotype (BM) (Fig. 144), “Laos: Borikhane Prov.,
Pakkading, 6.IV.1963”, “Pakkading, 6.4.[19]63” (handwritten), “J.A. Rondon
Collection Bishop Mus.”, “Holotype D ymasius (Elydnus) simplex Gressitt
& Rondon”, “8300” (Fig.221); 1♀, paratype (BM) (Fig.145), “Muong Wapi,
25.IV.[19]67”, “Allotype Dymasius (Elydnus) simplex Gressitt et Rondon”,
“8300” (Fig.222); 5♂, 1♀ (cAM) (Figs146,147), Nailand, Lamphun, Mae
a, 20.04.2011 (local collector), “Dymasius simplex Gressitt et Rondon,
1970 [♂or♀, respectively] det.A.Miroshnikov 2018”.
Morphological notes. e body length of the
holotype and female paratype (allotype) is 10.3 or 10.5mm,
the humeral width is 2.45 or 2.6mm, respectively; in the
specimens from ailand 9.2–11.6 mm and 2.1–2.7 mm,
respectively.
Distribution. Until now, this species has only been
known from Laos [Gressitt, Rondon, 1970]. Based on the
material studied, D. simplex is being recorded here from
ailand for the first time.
Dymasius prominor Gressitt et Rondon, 1970
(Figs 148–150, 223)
Dymasius (Microdymasius) prominor Gressitt et Rondon,
1970: 82. Type locality: Laos, Vientiane Province, a Ngone
(according to the original description and the label of the
holotype).
Material. 1♀, holotype (BM) (Fig.148), “Laos: Vientiane Prov., a
Ngone”, “Vientiane, a Ngone, 29.4.[19]63” (handwritten), “J.A. Rondon
Collection Bishop Mus.”, “Holotype D ymasius (Microdymasius) prominor
Gressitt & Rondon”, “8301” (Fig. 223); 5♂, 4♀ (cAM) (Figs 149, 150),
N ailand, Lamphun, Mae a, 20.04.2011 (local collector), “Dymasius
prominor Gressitt et Rondon, 1970 [♂or♀, respectively] det.A.Miroshnikov
2018”.
Morphological notes. is species was described
from a single female which I have revised, its body length
being 10.2mm and humeral width 2mm.
Male (Fig.149). Closely resembling the female. Body
length 8.7–10.5 mm, humeral width 1.9–2.1 mm (in the
females from ailand I have studied, 9.4–11 and 1.9–2.15,
respectively). In comparison with the female, antennae
barely longer, as in Fig.149 (cf.Figs148,150).
Distribution. Until now, this species has only been
known from Laos [Gressitt, Rondon, 1970]. Based on the
material studied, D.prominor is being recorded here from
ailand for the first time.
Dymasius parvus Gressitt et Rondon, 1970
(Figs 151–153, 224)
Dymasius (Microdymasius) parvus Gressitt et Rondon,
1970: 82. Type locality: Laos, Wapikhamthong Province, Khong
Sédone (according to the original description and the label of the
holotype).
Material. 1♂, holotype (BM) (Fig.151), “Laos: Wapikhamthong Prov.,
Khong Sédone, 18.IV.1965”, “Khongsédone, 18.4.[19]65” (handwritten),
“J.A.Rondon Collection Bishop Mus.”, “Holotype Dymasius (Microdymasius)
parvus Gressitt & Rondon” (Fig. 224); 5♂, 2♀ (cAM) (Figs 152, 153),
N ailand, Lamphun, Mae a, 20.04.2011 (local collector), “Dymasius
parvus Gressitt et Rondon, 1970 [♂ or ♀, respectively] det.A.Miroshnikov
2018”.
Morphological notes. is species was described
from two males; the body length of the holotype is 10.3mm,
the humeral width is 2mm.
Female (Fig.153). Closely resembling the male. Body
length 11.1–12.5mm, humeral width 2.2–2.5mm (in the
males from ailand I have studied, 8.8–11.7 and 1.8–2.4,
respectively). In comparison with the male, antennae
slightly shorter, body clearly more robust, as in Fig. 153
(cf.Figs 151,152).
Distribution. Until now, this species has only been
known from Laos [Gressitt, Rondon, 1970]. Based on the
material studied, D. parvus is being recorded here from
ailand for the first time.
Dymasius niger Gressitt et Rondon, 1970
(Figs 155, 167, 225)
Dymasius (Microdymasius) niger Gressitt et Rondon, 1970:
83. Type locality: Laos, Vientiane Province, Ban Van Eue (according
to the original description and the label of the holotype).
Material. 1♀, holotype (non ♂; see Remarks below) (BM) (Fig.155),
“Laos: Vientiane Prov., Ban Van Eue, 15.IV.1966”, “B an Van Eue, 15.4.[19]66”
(handwritten), “J.A.Rondon Collection Bishop Mus.”, “Holotype D ymasius
(Microdymasius) niger Gressitt & Rondon”, “8304” (Fig.225).
Remarks. In the original description of this species
[Gressitt, Rondon, 1970], the holotype was indicated to be
a male with a body length of 13mm and a humeral width
of 2.9mm. Aphotograph was presented in fig.16d (p.84)
showing a male (implying the holotype). In the description,
in addition to the male, there was also a female with a body
length of 11mm and a humeral width of 1.9mm. However,
there is only information pertaining to the holotype
male, the sole kept in BM, contained below the text of
the description: “Holotype ♂ (Bishop 8304)...”. Without
doubt, the reference to the female, i.e. one more specimen
in addition to the holotype, in the original description is
erroneous.
At the same time, the holotype I have studied is actually
a female with a body length of 13 mm and a humeral width
of 3 mm. It is this specimen that is shown in fig. 16d (p.84)
in the original description (I have properly remounted the
holotype). Ialso received a photograph of the holotype from
Dr.Nobuo Ohbayashi, which corresponds to the picture
in the original description, but the image was horizontally
mirrored.
Dymasius solodovnikovi Miroshnikov, sp. n.
(Fig. 156, 168)
Material. Holotype, ♀ (cAM) (Fig. 156): N ailand, Lamphun,
Mae a, 20.04.2011 (local collector). Paratypes: 1♀ (cSM), NW Laos,
228 A.I. Miroshnikov
Figs 149–154. Dymasius J. omson, 1864 and Zatrephus Pascoe, 1857, habitus, dorsal view.
149–150 – D. prominor Gressitt et Rondon, 1970 (from ailand); 151–153 – D. parvus Gressitt et Rondon, 1970 (152–153 – from ailand);
154–Z.jakli sp.n. 151, 154 – holotypes; 149, 151–152 – males; 150, 153–154 –females.
Рис. 149–154. Dymasius J. omson, 1864 и Zatrephus Pascoe, 1857, общий вид сверху.
149–150 – D. prominor Gressitt et Rondon, 1970 (из Таиланда); 151–153 – D. par vus Gressitt et Rondon, 1970 (152–153 – из Таиланда);
154–Z.jaklisp.n. 151, 154 – голотипы; 149, 151–152 – самцы; 150, 153–154 –самки.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 229
Figs 155–160. Dymasius J. omson, 1864, habitus, dorsal view, females.
155 – D. niger Gressitt et Rondon, 1970; 156 – D. solodovnikovi sp. n.; 157 – D. barclayi sp. n.; 158 – D. makarovi Miroshnikov, 2017; 159–
160–D.cuneatulus Holzschuh, 2005. 155–157 – holotypes; 158 – paratype.
Рис. 155–160. Dymasius J. omson, 1864, общий вид сверху, самки.
155 – D. niger Gressitt et Rondon, 1970; 156 – D. solodovnikovi sp. n.; 157 – D. barclayi sp. n.; 158 – D.makarovi Miroshnikov, 2017; 159–
160–D.cuneatulus Holzschuh, 2005. 155–157 – голотипы; 158 – паратип.
230 A.I. Miroshnikov
Luang Namtha Prov., Muang Sing env., 21°08ʹ51ʺN/ 101°10ʹ13ʺE, 750 m,
23.03–5.04.2010 (leg. S. Murzin); 1♀ (cSM), NW Laos, Luang Nam a
Prov., 65km NW of Luang Namtha, Nam a NPA, 1050m, 8–15.04.2010
(leg.S.Murzin).
Diagnosis. Based on female characters, this new
species is very similar to D. niger, but differs by the less
strongly elongated antennomere 3, the length ratio of
antennomere1 to3, the peculiar, less strongly developed,
recumbent, light setation and the somewhat peculiar
sculpture of the pronotum, as in Fig. 168, and the less
strongly developed, recumbent, light setation of the head
dorsally, as in Fig.168 (cf.Fig.167).
Description. Female. Body length 11.9–15.4mm, humeral
width 2.9–3.55 mm, thereby holotype largest. Coloration of
integument predominantly combines red-brown and dark reddish
brown tones; head dorsally almost entirely or partly, eyes, partly
mandibles and mostly pronotum black.
Head without median groove between upper lobes of
eyes; antennal tubercles well-expressed; eyes moderately
convex; submentum with rough, partly coarse transverse folds;
antennae longer than body, about reaching the apex of elytra by
antennomere9 or freely reaching beyond it by this antennomere;
length ratio of antennomeres1–11 (holotype taken as an example),
25 : 6 : 33 : 26 : 37 : 41 : 40 : 36 : 34 : 34 : 37; antennomere 1
devoid of a cicatrix (apical carina), with a heterogeneous, partly
rough sculpture; antennomere 2 subequal in length and width;
antennomere 3, 1.22–1.32 times as long as antennomere 1 (in
holotype of D.niger, 1.47times).
Pronotum barely longitudinal, 1.04–1.05times as long as wide
or subequal in length and width (inD.niger, pronotum 1.11times
as long as wide); base 1.06–1.25times as wide as apex; with a sharp
constriction both in front of base and near apex; on disc almost
flat, with a heterogeneous, rough, cellular sculpture, obliterated
both in front of base and near apex, partly with confluent cells, as
well as with a wide, relatively short, shiny, median area in basal part
behind the middle.
Scutellum triangular, with a poorly expressed puncturation.
Elytra predominantly nearly parallel-sided starting from
base, 2.5–2.6times as long as humeral width; with a small, dense,
in places confluent puncturation; apical external angle widely
rounded, sutural angle obtuse.
Prosternum mostly with irregular, mainly short, partly
transverse, more or less rough folds; prosternal process without
apical tubercle; mesosternal process between coxae more than
twice as wide as prosternal one, without tubercle dorsally;
metasternum and sternites with a small dense puncturation;
metasternum with a distinct median groove; last (visible) sternite
widely rounded at apex; last (visible) tergite truncate apically.
Legs relatively short; femora without longitudinal carina;
metatarsomere1 noticeably shorter than metatarsomeres2 and3
combined.
Recumbent setation mainly greyish, relatively uniform on
elytra and predominantly spotty on pronotum, as in Fig.168; more
or less long, erect, light setae mostly developed on pronotum and
head.
Etymology. I am pleased to dedicate this new species
to my colleague and friend, Dr.Alexey Yu. Solodovnikov,
curator of the Coleoptera collection of the Natural History
Museum of Denmark (University of Copenhagen), who
constantly provides his great and versatile help to my
research.
Distribution. ailand, Laos.
Dymasius barclayi Miroshnikov, sp. n.
(Figs 157, 169)
Material. Holotype, ♀ (BMNH) (Fig.157): Western Malaysia , “Perak:”,
“Doherty”, “Fry Coll. 1905.100”, “35548”.
Diagnosis. Based on female characters, this new
species seems to be especially similar to D. cuneatulus
Holzschuh, 2005 and D.makarovi Miroshnikov, 2017, but
differs clearly from both by the obviously longer antennae,
as in Fig.157, the more strongly flattened antennomere1,
the more strongly elongated at least antennomere 5, as
in Fig. 157, the length ratio of antennomere 3 to 5, the
almost entirely red-brown coloration of the integument,
the posteriorly more sharply bounded tubercle of the
mesosternal process. Besides this, D.barclayisp.n. differs
from the former species by the coloration of the dorsal
setation which is more similar to that of D.makarovi, while
from the latter species by the shape and sculpture of the
pronotum (Fig. 169) which are more similar to those of
D.cuneatulus (cf.Figs158–160, 170–172).
Description. Female. Body length 23 mm, humeral width
5.8 mm. Coloration of integument mainly red-brown; eyes and
partly mandibles black; folds of pronotum partly blackish (in
males and females of D.cuneatulus and D.makarovi, at least head
dorsally, pronotum, antennae, legs, partly venter black, thereby
elytra of females of these species black-brown while in female of
D.cuneatulus sometimes black or black-brown as well).
Head with a coarse median fold partly between bases of
antennae and partly between eyes, with a short median groove on
vertex just behind eyes; antennal tubercles moderately developed;
submentum with individual transverse folds and a heterogeneous,
clear, partly rough, more or less dense puncturation; antennae
longer than body, freely reaching beyond apex of elytra by
antennomere9 (inD.cuneatulus and D.makarovi, female antennae
reaching beyond apex of elytra only by penultimate antennomere);
length ratio of antennomeres1–11, 24:8 : 32 : 20: 32: 37: 36 :
33: 30: 26:29; antennomere1 devoid of a cicatrix (apical carina),
relatively strongly flattened, with somewhat heterogeneous, more
or less small, dense puncturation and coarse surface dorsally;
antennomere 2 distinctly longitudinal; antennomere 3 subequal
in length to 5th (in females of D. cuneatulus and D. makarovi,
antennomere3, 1.28–1.32times as long as antennomere5).
Pronotum barely transverse, 1.04times as wide as long; base
1.22times as wide as apex; with a sharper constriction near apex
than in front of base; on disc weakly convex, with coarse and very
coarse, transverse, partly fused folds, as in Fig.169.
Scutellum rounded apically, with a very small distinct
puncturation.
Elytra moderately narrowed towards apex, 2.7times as long
as humeral width; with both a rough, more or less regular and small
dense puncturation; apical external angle very well-expressed,
subrectangular, sutural angle with a very short denticle.
Prosternum with a very well-developed transverse groove in
apical part, with coarse, irregular, more or less short folds behind
it; prosternal process clearly broadened towards apex dorsally,
with a clear apical tubercle; mesosternal process with a strong
tubercle dorsally, between coxae clearly wider than prosternal
process; mesosternum partly, metasternum and sternites with
a clear, small, dense puncturation; metasternum and sternites,
in addition, with individual rough punctures; metasternum with
a sharply expressed median groove; last (visible) sternite barely
rounded, almost truncate at apex; last (visible) tergite with a very
poorly noticeable emargination apically.
Legs long; femora not claviform, without longitudinal carina;
metatarsomere 1 slightly shorter than metatarsomeres 2 and 3
combined.
Recumbent dense setation greyish, including on pronotum
and elytra.
Etymology. I am pleased to dedicate this new species
to my colleague, Dr.MaxwellV.L. Barclay, the curator of the
collection of Coleoptera at the Natural History Museum,
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 231
London, United Kingdom, who, over a number of years,
has kindly provided his great assistance to my study of the
museum material.
Distribution. Western Malaysia.
Dymasius makarovi Miroshnikov, 2017
(Figs 158, 170)
Dymasius makarovi Miroshnikov, 2017: 199. Type locality:
Western Malaysia, Pahang, Cameron Highlands, Tanah Rata
(according to the original description and the label of the holotype).
Material. 1♂, holotype (cAM),W Malaysia, Pahang, Cameron
Highlands, Tanah Rata, 04.2015 (local collector); 1♂, 1♀ (Fig. 158),
paratypes, (cAM), same label as holotype; 1♂ (BMNH), W Malaysia,
Perak, “Larut Hills [= Maxwell Hill], 3300–4300 ft., S.S. Flower. 99–248.”
(upperside), “Ap. 1898” (underside), “Dymasius makarovi Miroshnikov,
2017 ♂ det.A.Miroshnikov 2018”.
Distribution. Until now, this species has only been
known from the type locality. e record quoted here
indicates a wider distribution of D. makarovi in Western
Malaysia.
Dymasius maculatus Gressitt et Rondon, 1970
Dymasius (Dymasius) maculatus Gressitt et Rondon, 1970:
80. Type locality: Laos, NofVientiane, Phou Khao Khoay, 1040m
(according to the original description).
Material. 1♂, holotype (BM) (photograph); 1♀, paratype (allotype)
(BM) (photograph); 1♂ (BMNH), “Siam, Renong”, “Doherty”, “Fry Coll.
1905.100”, “62447”, “Dymasius maculatus Gressitt et Rondon, 1970 ♂
det.A.Miroshnikov 2018 [preliminary determination]”.
Remarks. e male I have studied is attributed to
this species only preliminarily, since it was compared to
the holotype male and the allotype female of D. maculatus
based only on their photographs.
Distribution. e species in question has hitherto
been known only from Laos [Gressitt, Rondon, 1970].
Based on the new material, this species, albeit preliminarily,
is being recorded from ailand for the first time.
Genus Zatrephus Pascoe, 1857
Zatrephus Pascoe, 1857: 94; omson, 1864: 235; Pascoe,
1869: 523; Lacordaire, 1868: 267; Gemminger in Gemminger,
Harold, 1872: 2805; Aurivillius, 1912: 62; Gressitt, Rondon, 1970:
88; Catalogue..., 2010: 162; Heffern, 2013: 12; Miroshnikov, 2017:
208.
Type species: Zatrephus pannosus Pascoe, 1857, by
subsequent designation [Gressitt, Rondon, 1970].
Remarks. A review of this genus was published
recently, in which seven species were considered, including
one new [Miroshnikov, 2017]. Two females from Java were
also discussed in that paper, suggesting they were very
likely to belong to a new form. In preparing this review,
I have postponed its description in the hope to find a
corresponding male in any collections. However, until
now this has not happened. Only one of the females from
Java is currently available to me, on the basis of which the
following new species is described.
Zatrephus jakli Miroshnikov, sp. n.
(Fig. 154)
Zatrephus sp.: Miroshnikov, 2017: 208, figs266, 286 (Java).
Material. Holotype, ♀ (cLD) (Fig. 154): Indonesia, E Java, Meru
Betiri National Park, Sukamade, 8°15ʹS / 113°30ʹE, 0–200 m, 02–03.1996
(leg.S.Jákl), “Zatrephus pannosus Pasc. det.L.Dembický 2000”.
Diagnosis. Based on female characters, this new
species seems to be especially similar to Z. pannosus
Pascoe, 1857, but differs clearly by the structure of the
pronotum, in particular, the more obliterated peculiar
sculpture and the more strongly developed, recumbent,
light setation, thereby forming no contrasting, prominent,
lateral spot on each side near the apex; the absence of a
hairless shiny spot in the apical quarter of each elytron; the
less strongly elongated last (visible) sternite; the recumbent
light setation of the scutellum more widely separated by a
median bare strip; the somewhat more spotty recumbent
setation of the elytra, metasternum, sternites, femora,
and tibiae (in contrast to the vast majority of specimens
of Z.pannosus Ihave studied); the less strongly developed
recumbent setation of the submentum restricted mainly
to its middle third; and the smaller body size. Zatrephus
jakli sp. n. differs from the Javan Z. javanicus Fischer,
1936 by almost all features making it distinguished from
Z.pannosus, at least so from the male, because the female
Z.javanicus still remains unknown to me (cf.Miroshnikov
[2017: 201 (figs 263–265), 205 (figs275–277)]).
Description. Female. Body length 22.1mm, humeral width
6mm. Eyes, almost entirely head dorsally and pronotum, partly
mandibles black; remaining parts combines dark red-brown and
red-brown tones.
Head with a very deep median groove between upper lobes
of eyes, partly, and on vertex; antennal tubercles poorly developed;
genae moderately short; eyes weakly convex; gula with gentle
transverse wrinkles; neck predominantly with rough transverse
folds; antennae short, barely extending beyond middle of elytra;
length ratio of antennomeres1–11, 19:5: 13: 11: 12: 14: 16: 15:
14: 13:16; antennomere1 with a moderately rough very dense
puncturation; antennomere2 clearly transverse; antennomeres3–5
inflated, as in Fig.154; antennomeres6–10 moderately serrate.
Pronotum distinctly transverse, 1.08times as wide as long;
base 1.15times as wide as apex; with a sharper constriction near
apex than in front of base; on disc almost flat, predominantly with
a rough sculpture clearly obliterated in middle area and there
with neither transverse nor longitudinal folds sharply expressed
(inZ.pannosus and Z.javanicus, pronotum in middle area with
as very coarse, sharply expressed, differently oriented folds as in
adjacent areas).
Scutellum triangular, shortly truncate at the very base.
Elytra predominantly nearly parallel-sided starting from
base, 2.45times as long as humeral width; lateral to scutellum with
a very clear tubercle at the very base; with a heterogeneous, more
or less small, partly very small puncturation, behind the middle
partly with larger punctures; apical external angle obtuse, sutural
angle drawn into a clear, but small tooth, thereby both angles more
or less strongly masked under a dense setation.
Prosternum with a well-expressed transverse groove in front
of middle, with a transverse, moderately wide, roughly sculptured
elevation before it; prosternal process with a strong apical tubercle;
mesosternal process without tubercle dorsally, between coxae
significantly wider than prosternal process; mesosternum partly,
metasternum and sternites with a small dense puncturation;
metasternum with a weakly expressed median groove; last (visible)
sternite unclear rounded, almost truncate at apex, last (visible)
tergite with a poorly developed emargination apically.
Legs relatively short; metatarsomere 1 very clearly shorter
than metatarsomeres2 and3 combined.
Recumbent dense setation predominantly clearly spotted,
especially so on elytra and venter, combines red/reddish and
232 A.I. Miroshnikov
Figs 161–172. Dymasius J. omson, 1864.
161, 163, 165 – D. strigosus J. omson, 1864; 162, 164, 166 – D. macilentus (Pascoe, 1859); 167 – D. niger Gressitt et Rondon, 1970; 168 –
D.solodovnikovisp.n.; 169 – D. barclayi sp.n.; 170 – D. makarovi Miroshnikov, 2017, paratype; 171–172 – D. cuneatulus Holzschuh, 2005. 162, 164,
166–169 – holotypes; 161–162 – apical part of tegmen, ventral view; 163–164 – apical part of penis, ventral view; 165–166 – apical part of tergite 8, dorsal
view; 167–168 – head, dorsal view, and pronotum; 169–172 – pronotum.
Рис. 161–172. Dymasius J. omson, 1864.
161, 163, 165 – D. strigosus J. omson, 1864; 162,164, 166 – D. macilentus (Pascoe, 1859); 167 – D. niger Gressitt et Rondon, 1970; 168 –
D.solodovnikovisp.n.; 169 – D. barclayi sp.n.; 170 – D. makarovi Miroshnikov, 2017, паратип; 171–172 – D. cuneatulus Holzschuh, 2005. 162, 164,
166–169– голотипы; 161–162 – вершинная часть тегмена снизу; 163–164 – вершинная часть пениса снизу; 165–166 – вершинная часть 8-го тергита
сверху; 167–168 – голова сверху и переднеспинка; 169–172 – переднеспинка.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 233
white/whitish tones; setation in apical part of elytra partly rarefied
or missing, as a result forming a relatively wide and dark fascia; red
setae prevailing or strongly dominating mainly on head dorsally
and basal antennomeres, whereas white setae prevailing at least
on elytra, as in Fig.154; more or less long, erect, thin setae mainly
developed on pronotum and head.
Etymology. I am pleased to dedicate this new species
to Mr.Stanislav Jákl (Praha, Czech Republic), who collected
the holotype of this new species, as well as many other
little-known or rare Oriental cerambycids.
Distribution. Indonesia (Java).
Genus Diorthus Gahan, 1891
Diorthus Gahan, 1891: 27 (Pachydissus subgen., “section”);
Gahan, 1906: 132; Plavilstshikov, 1931: 81; Gressitt, Rondon, 1970:
70; Adlbauer, 2006: 62; Catalogue..., 2010: 160; Nga et al., 2014:
433; Kariyanna et al., 2017: 30; Miroshnikov, 2017: 223.
Diorthrus (misspelling): Aurivillius, 1912: 56.
Tapinolachnus auct. (non J. omson, 1865): Özdikmen,
Turgut, 2009: 302 (part.).
Type species: Hammaticherus simplex White,
1853= Cerambyx cinereus Fabricius, 1793, by subsequent
designation [Gahan, 1906].
Remarks. is genus, without any explanation, was
synonymized by some modern authors [Özdikmen, Turgut,
2009] with the genus Tapinolachnus J. omson, 1865.
However, other researchers [Weigel et al., 2013; e first
Web-site..., 2018] have found this in no way justified. Ialso
consider this synonymy to be completely erroneous.
Diorthus differs clearly from Tapinolachnus by the
structure of the antennae, including the presence of a
cicatrix on antennomere 1, the length ratio of the basal
antennomeres, the sculpture of antennomeres 3–5 and
the inflated male antennomeres3 and 4; the structure of
the elytra, in particular, a sharp, at least mostly clearly
larger puncturation and a very distinctly coarser, dense,
recumbent setation, and some other features.
Diorthus cinereus (Fabricius, 1793)
(Figs 173, 174, 184, 186, 189, 192, 227)
Cerambyx cinereus Fabricius, 1793: 265. Type locality: India,
“Tranquebariae” (according to the original description). Fabricius,
1801: 281.
Diorthus cinereus: Aurivillius, 1912: 56; Plavilstshikov,
1931:81; Hua, 1984: 36 (“Diorthrus”, misspelling); Adlbauer, 2006:
62; Makihara et al., 2008: 100; Catalogue..., 2010: 160; Nga et al.,
2014: 433; Kariyanna et al., 2017: 30; Kariyanna et al., 2018: 165.
Diorthus (Diorthus) cinereus: Gressitt, Rondon, 1970: 71.
Cerambyx holosericeus Olivier, 1795: 14 (No. 67), pl. 17,
fig.127 (Indes orientales).
Hammaticherus simplex White, 1853: 130 (W.Africa).
Pachydissus (Diorthus) simplex: Gahan, 1891: 31; 1894: 10.
Diorthus simplex: Gahan, 1906: 133.
Cerambyx vernicosus Pascoe, 1859: 19 (Ceylon).
Pachydissus inclemens J. omson, 1865b: 576 (India);
omson, 1878: 7.
Neocerambyx sordidus Pascoe, 1888: 491 (Laos).
Material. 1♂, type specimen (?holotype by monotypy) (ZMUK)
(photograph); 1♀, holotype (by monotypy) of Diorthus simplex
(White, 1853) (BMNH) (Fig. 173), “W. Afr[ica].” (upperside), “48/88”
(underside), “Pachydissus (Hammaticherus) simplex White, T y p e ”, “ T y p e ”,
“Hammaticherusn. sp. near H.sericeus”, “Hammaticherus simplex n.sp.,
W. Africa” (Fig. 227); 1♀ (NHMD), Sri Lanka, Southern Province,
Hambantota, 25–29.12.1994, ex larva from Tamarindus indica, 10.1995
(leg. O. Mehl), “Diorthus cinereus (Fab.), Ole Mehl det.”; 1♂ (NHMD),
Sri Lanka, Southern Province, Hambantota, 26–30.06.2003, ex larva
from Prosopis juliflora, 17.04.2004 (leg. O. Mehl), “Diorthus cinereus
(Fab.), Ole Mehldet.”; 1♂, 1♀ (NHMD), Sri Lanka, Western Province,
Henarathgoda Bot.Gar., 18.07.2003, 07.2004 (leg. O. Mehl), “Diorthus
cinereus (Fab.), Ole Mehl det. 2005”; 1♀ (cAM), Mautitius, Triolet
env., 2016 (leg .N. Kholiushkina), “Diorthus cinereus (Fabricius, 1793)
♀ det. A.Miroshnikov 2017”; 1♀ (BMNH), “Madras, India”, “G.Briant
Coll. 1919–147”; 1♂, 1♀ (cSM), SE India, Madras env., Manapakkam,
20.04.1997 (leg S.Saluk), “Diorthus cinereus (Fabricius, 1793) [♂ or ♀,
respectively] det. A. Miroshnikov 2018”; 1♂ (BMNH), “arrawaddy,
Burma”, “Pachydissus simplex White”; 1♂ (BMNH), Myanmar, “Paungdé”,
“Pachydissus simplex White”; 1♂ (ZMMU), “Siam mer., Sala-pa-[illegible
further on], Staudinger”, “Diorthus cinereus(F.), N.Plavilstshikovdet.”; 1♀
(NHMD), NELaos, Hua Phan Prov., Ban Saleui, Phou Pan Mt., 20°12ʹN/
104°01ʹE, 27.06.2013 (leg . C. Holzschuh), “Diorthus cinereus (Fab.),
Ole Mehl det. 2014”; 1♂ (cAM), Vietnam, Cao Bang Prov., Phja-Den
env., 950m, 30.04–5.05.2012 (leg.Lingafelter, Jendek, Pham), “Diorthus
cinereus (Fabricius, 1793) ♂ det. A. Miroshnikov 2017”; 1♂ (BMNH),
“Hamaticherus sericeus mihi h. in ins. J a v a ”, “Bowr. Ch e vr. 63–47*”,
“♂”; 1♂ (BMNH), “ J a v a ”, “Fry Coll. 1905–100”, “e x Mus . Laferte”; 1♂
(BMNH), “Senegal”, “Fry Coll. 1905–100”, “exM us . Laferte”; 1♀ (BMNH),
“Hamaticherus sericeus Dej., J a v a ”, “1047”, a red rectangle 9 × 6mm
(without inscription).
Morphological notes. Body length 14–32 mm
[Gressitt, Rondon, 1970]; in the specimens I have studied
the body length was 21.3–30 mm, the humeral width
between 6.2–8.5 mm (holotype of Diorthus simplex:
23.3mm and 6.9mm, respectively).
Remarks. A picture of the type male specimen of
Cerambyx cinereus Fabricius, 1793 is available on the
website of NHMD [http://www.daim.snm.ku.dk/search-in-
types].
Distribution. is species is very widely distributed
and covers Africa (at least from Mauritania south to
Tanzania; Mauritius), Southwest Asia (United Arab
Emirates, Yemen, southern Iran), South Asia, Indochina,
Indonesia (Java).
Diorthus pellitulus Holzschuh, 1984
(Fig. 175)
Diorthus pellitulus Holzschuh, 1984: 144. Type locality:
Nepal, Monari, Mitte Mai (according to the original description).
Weigel, 2006: 498.
Material. 1♀, holotype (cCH) (photograph; Fig.175).
Morphological notes. Body length 19 mm
[Holzschuh, 1984].
Distribution. Nepal.
Diorthus intricarius Holzschuh, 1984
(Fig. 176)
Diorthus intricarius Holzschuh, 1984: 145. Type locality:
Pakistan, Swat, Madyan, “71°90ʹL/ 35°70ʹB”, 1400m (according to
the original description).
Material. 1♀, holotype (cCH) (photograph; Fig.176).
Morphological notes. Body length 22 mm
[Holzschuh, 1984].
Distribution. Pakistan.
Diorthus kabakovi Miroshnikov, sp. n.
(Figs 177, 188, 191)
Diorthus sp.: Miroshnikov, 2017: 185, fig.147 (Afghanistan).
Material. Holotype, ♂ (ZIN) (Fig. 177): Afghanistan, Nurestan
[= Nuristan], S Čapa Dara [= Capa Dara], 1800 m, 14.06.1971
(leg.O.N.Kabakov), “Derolus”.
234 A.I. Miroshnikov
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 235
Diagnosis. is new species seems to be especially
similar to D. intricarius, but differs clearly by the darker
general coloration; the more obliterated sculpture of
the pronotum that more strongly masks its recumbent
light setation, as in Fig.177; the somewhat more strongly
elongated scutellum; the seemingly longer head behind
the eyes and, accordingly, the longer temples, as in
Fig. 177. Diorthus kabakovi sp. n. can also be compared
to D.pellitulus, but differs by some features like from the
previous species, including the darker general coloration,
the shape of the scutellum, the longer head behind the eyes,
as well as by the somewhat different shape of the pronotum
(albeit compared in the holotypes belonging to the opposite
sex), the pattern of its recumbent setation and seemingly
certain features of its sculpture (cf.Figs175,176).
Description. Male. Body length 26.5 mm, humeral width
7.7mm. Coloration of integument mainly dark red-brown, apical
antennomeres lightest; eyes and mandibles black.
Head with a well-expressed median groove between upper
lobes of eyes; antennal tubercles moderately developed; temples
rather long, almost twice as long as genae; antennae nearly reaching
the apex of elytra; length ratio of antennomeres1–11, 29:6: 26:
22: 24: 26: 27: 26: 25: 24:41 (length ratio of antennomeres4 and5
given taking into account their peculiarly distinguished bases);
antennomere1 with a sharp expressed cicatrix (apical carina), with
a heterogeneous, rough, irregular sculpture and, in addition, with
small dense punctures; antennomere 2 very clearly transverse;
basal part of antennomeres 3–5, predominantly dorsally, with a
coarse and rough puncturation; bases of antennomeres 4 and 5
with a wide fragment of a scabrous dull surface, sharply different
from adjacent parts of shiny surface of these antennomeres,
and, in addition, clearly delimited from this surface by a sharp
constriction; last antennomere with a well-expressed appendage.
Pronotum very clearly transverse, 1.22times as wide as long;
base 1.17times as wide as apex; with a well-expressed constriction
both in front of base and near apex; on disc weakly convex, only
with rough irregular folds and, in addition, with a clear, naked,
strongly shiny, median area in apical one-quarter.
Scutellum triangular, sharpened apically, with a very small,
but clear, very dense puncturation.
Elytra predominantly nearly parallel-sided starting from
base, 2.25 times as long as humeral width; with a somewhat
heterogeneous, more or less small, but sharp, very dense
puncturation obliterated towards apex; apical external angle
obtuse, sutural angle with a small denticle.
Prosternum in apical part with rough transverse folds;
prosternal process moderately broad; mesosternal process between
coxae about 1.3 times as wide as prosternal process, without
tubercle dorsally; metasternum and sternites with a small dense
puncturation; metasternum with a sharply expressed median
groove; last (visible) sternite with a clear impression, truncate at
apex; last (visible) tergite with a well-exspressed emargination
apically.
Legs moderately long; all femora with a clear carina along each
side; metatarsomere1 noticeably shorter than metatarsomeres2
and3 combined.
Recumbent dense setation of dorsum, except for scutellum,
consisting predominantly of grey and less numerous reddish setae,
those of remaining parts mainly grey, those of metasternum,
sternites, femora and tibiae speckled; more or less long, erect, light
setae mainly developed on pronotum and head.
Etymology. is new species is dedicated to the
memory of Oleg Nikolaevich Kabakov (1928–2009), a
famous Russian entomologist and an excellent collector of
beetles.
Distribution. Afghanistan.
Diorthus sericeus Gardner, 1939
(Figs 178, 179, 230, 231)
Diorthus sericeus Gardner, 1939: 2. Type locality: “South
Mangalore, 400m, Madras, India” (according to the label of the
lectotype). Kariyanna et al., 2017: 30.
Material. 1♂, lectotype, here designated (BMNH) (Fig.178), India,
“S.Mangalore, 400[m], Madras, J.C.M. Gardner, 30.IV.1931” (upperside),
“No. 86.M.” (underside), “Ex Pterocarpus marsupium”, “R.R.D. 119,
B.C.R. 126, Cage 779”, “Diorthus sericeus J.C.M. Gardner sp. n., Type”,
“Type”, “Brit. Mus. 1939–414”, “NHMUK 011220588” (Fig.231), “Lectotypus
♂ Diorthus sericeus Gardner, 1939, A. Miroshnikov des., 2018”; 1♀,
paralectotype (BMNH) (Fig. 179), India, “Sappal, 1700 [m], Palghat,
Madras, J.C.M. Gardner, 2.V.1931” (upperside), “No. M.74” (underside),
“Ex Acacia sp.”, “R.R.D. 119, B.C.R. 154, Cage 768”, “Diorthus sericeus
J.C.M.Gardnersp.n., Allotype”, “Type”, “Brit. Mus. 1939–414”, “NHMUK
011220589” (Fig.230), “Paralectotypus ♀ Diorthus sericeus Gardner, 1939,
A.Miroshnikovdes., 2018”.
Morphological notes. Body length 17–20 mm
[Gardner, 1939], thereby the body length of the lectotype
and female paralectotype (both in BMNH) is 19.9 or
20.5mm, the humeral width is 5.9 or 6.05mm, respectively.
Remarks. In the original description, Gardner [1939:
2] noted the following: “Two males and three females
reared from Pterocarpus marsupium and Acacia sp.,
Palghat, Madras (J.C .M.Gardner). Type (male) and allotype
(female) in British Museum; paratypes in Forest Research
Institute ”. However, another locality is marked on the label
of the type, namely, “S[outh]. Mangalore, Madras”. In this
regard, it seems to me necessary to designate the lectotype
and to clarify the type locality.
is species is morphologically not a quite
characteristic representative of the genus. At least it does
not have a longitudinal carina on the femora, as well as,
unlike other species, the structural features of the bases of
male antennomeres4 and5 (described below, see Diagnosis
of the genus Lamellocerambyx) are poorly expressed.
Distribution. Southern India.
Diorthus vagus (Gahan, 1891)
(Figs 181, 182, 228, 229)
Pachydissus (Diorthus) vagus Gahan, 1891: 32. Type locality:
“Senegal?” (according to the original description and the label of
the holotype). Aurivillius, 1912: 56.
Diorthus vagus: Adlbauer, 2006: 62.
Material. 1♂, holotype, by monotypy (BMNH) (Fig. 181),
“N.sp.Senegal?”, “Bowr. Chevr. 63–47*”, “Pachydissus vagus Gahan ♂, Type”,
“Type” (Fig.228); 1♀ (BMNH) (Fig.182), “Nova Holland” (wrong locality),
“Fry Coll. 1905.100”, “Ex Mus. Parry”, “Pachydissus vagus Gahan♀”, “43005”
(Fig.229).
Morphological notes. Body length 24.5–30.5 mm,
humeral width 7.1–9.6mm, thereby holotype smallest.
Remarks. Gahan’s comments [1891: 32] to the original
description must be mentioned here: “is species has a
strong resemblance and an evident affinity to P. simplex
(White), and its habitat might have thrown some light
upon the distribution of the latter. Unfortunately, however,
of the two specimens one (the male type) is ticketed
‘Senegal ?,’ the other (afemale, in Mr.Fryʼs collection) is
ticketed ‘Nov. Holland.’ e latter locality can scarcely be
correct”. In addition, as regards the female, Adlbauer [2006:
62–63] noted that it “...kam 1871 in die Fry Collection
(ex Mus. Parry, Australien) und dann ins BMNH. Das
Determinationsetikett „Pachydissus vagus ♀ Gahan“ wurde
offenbar später hinzugefügt. (S.Shute, in litteris)”.
Distribution. ? Senegal.
236 A.I. Miroshnikov
Figs 173–177. Diorthus Gahan, 1891, habitus, dorsal view.
173–174 – D. cinereus (Fabricius, 1793) (173 – holotype of Diorthus simplex (White, 1853)); 175 – D. pellitulus Holzschuh, 1984; 176 –D. intricarius
Holzschuh, 1984; 177 –D. kabakovi sp. n. 173, 175–177 – holotypes; 173, 175 – 176 – females; 174, 177 –males.
Рис. 173–177. Diorthus Gahan, 1891, общий вид сверху.
173–174 – D. cinereus (Fabricius, 1793) (173 – holotype of Diorthus simplex (White, 1853)); 175 – D. pellitulus Holzschuh, 1984; 176 –D. intricarius
Holzschuh, 1984; 177 –D. kabakovi sp. n. 173, 175–177 – голотипы; 173, 175 – 176 – самки; 174, 177 –самцы.
Figs 178–183. Diorthus Gahan, 1891 and Lamellocerambyx Pic, 1923, stat. rest., habitus, dorsal view.
178–179 – D. sericeus Gardner, 1939; 181–182 – D. vagus (Gahan, 1891); 180, 183 – L. laosensis Pic, 1923, comb. rest. 178 – lectotype; 179 –
paralectotype; 181, 183 – holotypes; 178, 180–181 – males; 179, 182–183 – females.
Рис. 178–183. Diorthus Gahan, 1891 и Lamellocerambyx Pic, 1923, stat. rest., общий вид сверху.
178–179 – D. sericeus Gardner, 1939; 181–182 – D. vagus (Gahan, 1891); 180, 183–L. laosensis Pic, 1923, comb. rest. 178 – лектотип; 179 –
паралектотип; 181, 183 – голотипы; 178, 180–181 – самцы; 179, 182–183 – самки.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 237
Figs 184–193. Diorthus Gahan, 1891 and Lamellocerambyx Pic, 1923, stat. rest., males.
184, 186, 189, 192 – D. cinereus (Fabricius, 1793); 185, 187, 190, 193 – L. laosensis Pic, 1923, comb. rest.; 188, 191 –D. kabakovi sp. n., holotype. 184–
185 – head, dorsal view; 186–187 – right eye; 188–190 – right antennomeres 3–4 and basal part of antennomere 5; 191–193 – base of right antennomere 4.
Рис. 184–193. Diorthus Gahan, 1891 и Lamellocerambyx Pic, 1923, stat. rest., самцы.
184, 186, 189, 192 – D. cinereus (Fabricius, 1793); 185, 187, 190, 193 – L. laosensis Pic, 1923, comb. rest.; 188, 191 – D. kabakovi sp. n., голотип.
184–185 – голова сверху; 186–187 – правый глаз; 188–190 – правые 3–4-й членики усиков и основная часть 5-го членика; 191–193 – основание
правого 4-го членика усиков.
238 A.I. Miroshnikov
Diorthus sp.
Remarks. e two males from southern Iran referred
to as Diorthus cinereus [Longhorn beetles..., http://www.
cerambyx.uochb.cz/] actually belong to another, probably
still undescribed species. I have studied quite extensive
material of D.cinereus from various regions and none of
the males has such short antennae and many antennomeres
so strongly shortened as in both southern Iranian males.
e male antennae of D. cinereus are much longer than
the body, reaching beyond the apex of the elytra usually
by antennomere7, while many antennomeres are strongly
elongated, as in Fig.174.
Genus Lamellocerambyx Pic, 1923, stat. rest.
Lamellocerambyx Pic, 1923a: 8; Gressitt, Rondon, 1970: 71
(Diorthus subgen.); Weigel et al., 2013: 52 (Diorthus subgen.).
Type species: Lamellocerambyx laosensis Pic, 1923, by
monotypy.
Diagnosis. is genus which some researchers
consider as a subgenus of the genus Diorthus differs
clearly from it by the structure of the eyes; the pattern of
elytral setation; the structure of the antennae, including
the sculpture of male antennomeres4 and 5 or 3–5; the
somewhat more slender body (at least from almost all
representatives of Diorthus); as well as by some other traits
indicated below.
When detailing the structure of
Lamellocerambyx stat. rest., the following features must
be noted as being characteristic of this genus: eyes almost
completely divided into two lobes, both connected to each
other by a relatively long and very narrow bridge entirely
devoid of ocelli, as in Fig. 187, upper lobe thereby being
disposed obliquely vertically, as in Fig. 185 (whereas in
Diorthus, albeit eyes also almost completely divided into
two lobes, a connecting bridge very short in narrowest
place, uniformly widening in both directions from this
place and often showing here one or more ocelli, as in
Fig.186, upper lobe thereby being disposed clearly more
horizontally, at an angle of about 45degrees, as in Fig.184);
male antennae more than 2 times longer than body
(whereas in Diorthus, male antennae if long, then only less
than 2times longer than body, sometimes relatively short,
only about reaching the apex of elytra or insignificantly
surpassing it); antennomere1, like in Diorthus, with a more
or less coarse sculpture, but with a clearly more obliquely
disposed cicatrix , as in Fig.185 (cf.Fig.184); antennomere2
subequal in length and width, but not transverse, as in
Fig. 185 (whereas in Diorthus, antennomere 2 distinctly
or very clearly transverse – Fig. 184); in male, bases
of antennomeres 4 and 5 usual in structure, at least
dorsally sculpture rather similar to adjacent parts of these
segments and, in addition, not separated from them by
any constriction, as in Figs 190, 193, antennomeres 3–5
without coarse sculpture (while in known males of almost
all species of Diorthus, bases of antennomeres4 and5 with
a more or less wide fragment of a scabrous dull surface,
sharply different from adjacent parts of shiny surface of
these antennomeres, and, in addition, usually or at least
often delimited from this surface by a distinct or sharp
constriction, as in Figs188,189, 191,192; antennomeres3–5
of male sometimes with a heterogeneous, partly or mostly
rough and coarse sculpture); pronotum with coarse,
mostly transverse folds, can only be with a median,
longitudinal, more or less short, narrow fold (while in a
number of Diorthus species, pronotum with less coarse
and mostly or predominantly irregular folds); elytra with
a recumbent setation, appearing velvety and forming, at
least partly, distinct longitudinal stripes, as in Figs180, 183
(while in Diorthus, setation of elytra neither forming clear
longitudinal stripes nor appearing velvety, as in Figs173–
177, 178,179, 181,182); legs moderately long; at least meso-
and metafemora each without carina, only profemora
ventrally sometimes with a more or less noticeable, gentle
carina (while in almost all species of Diorthus, femora
usually with a clear, often sharp, sometimes less distinct
carina along each side).
Composition. e genus includes a single species.
Distribution. Indochina and southern China.
Lamellocerambyx laosensis Pic, 1923, comb. rest.
(Figs 180, 183, 185, 187, 190, 193, 226)
Lamellocerambyx laosensis Pic, 1923a: 8. Type locality: Laos,
“Nam Mia” (according to the original description and the label of
the holotype).
Diorthus (Lamellocerambyx) laosensis: Gressitt, Rondon,
1970: 71; Weigel et al., 2013: 52 (Laos; China, Yunnan).
Diorthus laosensis: Weigel et al., 2013: 72, 161, pl.6, figsc, d.
Material. 1♀, holotype, by monotypy (MNHN) (photograph;
Fig. 183), “Laos, Nam Mia, le 17.IV.1918, R. Vitalis de Salvaza”,
“Lamellocerambyx n. g. laosensis n. sp.”, “Type”, “Museum Paris, Coll.
M. Pic”, “Holotype” (Fig. 226); 1♂ (cAM), Laos, Xaignabouri City,
16–18.04.2005 (unknown collector), “Lamellocerambyx laosensis Pic,
1923 ♂ det. A. Miroshnikov 2018”; 1♂ (Fig. 180), 1♀ (cSM), NW Laos,
Luang Namtha Prov., Muang Sing env., 21°08ʹ51ʺN/ 101°10ʹ13ʺE, 750 m,
26.03–5.04.2010 (leg. S. Murzin), “Lamellocerambyx laosensis Pic, 1923
[♂ or ♀, respectively] det. A. Miroshnikov 2018”; 1♂, 1♀ (cSM), same
locality, 1–10.04.2011 (leg. S. Murzin), “Lamellocerambyx laosensis Pic,
1923 [♂ or ♀, respectively] det. A. Miroshnikov 2018”; 1♂ (cLD), “Laos”,
“Lamellocerambyx laosensis Pic, 1923 ♂ det. A. Miroshnikov 2018”; 1♂
(NHMD), NELaos, Hua Phan Prov., Ban Saleui, Phou PanMt., 20°12ʹN /
104°01ʹE, 27.06.2013 (leg. C. Holzschuh), “Diorthus (Lamellocerambyx)
laosensis (Pic, 1923), Ole Mehldet. 2014”.
Morphological notes. Body length 16–24 mm
[Gressitt, Rondon, 1970], thereby the holotype is 18 mm
(Dr. Gérard L. Tavakilian, personal communication); in
the specimens I have studied the body length was 21.5–
26.2mm, the humeral width between 5.7–7mm.
Distribution. Laos, ailand and southern China.
Genus Tapinolachnus J. omson, 1865
Homalolachnus J. omson, 1864: 232 (nom. praeocc., non
LaFerté-Sénectère, 1851, Carabidae); Gemminger in Gemminger,
Harold, 1872: 2804.
Tapinolachnus J. omson, 1865a: 445; Aurivillius, 1912: 61.
Mimoderolus (Aeolesthes subgen.) Pic, 1933: 11, syn. n. (non
syn. pro Derolus Gahan, 1891: Vitali et al., 2017).
Pachydissus auct. (non Newman, 1838): Fisher, 1940: 202 (part.).
Derolus auct. (non Gahan, 1891): Gressitt, Rondon, 1970: 74
(part.); Vitali et al., 2017: 59 (part.).
Type species: Homalolachnus lacordairei J. omson,
1864.
Composition. e genus includes a single species.
Distribution. Oriental realm.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 239
Figs 194–200. Tapinolachnus J. omson, 1865, habitus, dorsal view, and pronotum.
194, 197 – T. lacordairei (J. omson, 1864), syntypes (photographs by Azadeh Taghavian); 195–196, 198–200 – T.?lacordairei ( J. omson, 1864)
(195, 200 – lectotype of Tapinolachnus xyliae (Fisher, 1940), comb. n.). 194–196, 199–200 – males; 197–198 – females.
Рис. 194–200. Tapinolachnus J. omson, 1865, общий вид сверху и переднеспинка.
194, 197 – T. lacordairei (J. omson, 1864), синтипы (фотографии А. Тагвьян); 195–196, 198–200 – T.?lacordairei (J. omson, 1864) (195, 200 –
лектотип Tapinolachnus xyliae (Fisher, 1940), comb. n.). 194–196, 199–200 – самцы; 197–198 – самки.
240 A.I. Miroshnikov
Tapinolachnus lacordairei (J. omson, 1864)
(Figs 194, 197, 232, 233)
Homalolachnus lacordairei J. omson, 1864: 232. Type
locality: “Malasia” (according to the original description and the labels
of the syntypes). Gemminger in Gemminger, Harold, 1872: 2804.
Tapinolachnus [lacordairei]: omson, 1865a: 445.
Tapinolachnus lacordairei: Lacordaire, 1868: 265 (“Malaisie”);
omson, 1878: 7; Aurivillius, 1912: 61 (“Malay. Archipel”) .
Aeolesthes (Mimoderolus) uniformis Pic, 1933: 11 (indicated
here as a synonym of T.lacordairei only preliminarily); Vitali et al.,
2017: 59 (asDerolus).
Pachydissus xyliae Fisher, 1940: 202 (indicated here as a
synonym of T. lacordairei only preliminarily); Gressitt, Rondon,
1970: 74 (as Derolus); Vitali et al., 2017: 59 (syn. pro Derolus
uniformis).
Material. 1♂, syntype (MNHN) (photographs; Fig. 194),
“Tapinolachnus oms. S.C. 445. Homalolachnus oms. S.C. 232.
nom.pr.”, “lacordairei oms. 232. Type Malas.”, “./Type”, “Tapinolachnus
lacordairei”, “Museum Paris, Coll. J.omson 1952” (Fig.232); 1♀, syntype
(MNHN) (photographs; Fig.197), “Museum Paris, Coll. J.omson 1952”,
“Paratype” (Fig.233).
Note. On the first two labels of the above syntypes that
are shared by both the male and female, the numerals 232
and 445 denote the page numbers of omson’s original
descriptions [1864, 1865a]. e modern label reading
“Paratype” attached to the female syntype is incorrect.
Body length and humeral width of male and female
syntypes 29.3 or 27.7mm and 7.6 or 7.5mm, respectively
(Mrs.Azadeh Taghavian, personal communication).
Additional material. e following specimens I have studied are
provisionally attributed to Tapinolachnus lacordairei: 1♂ (IRSN), “Malacca”,
“2636”; 2♂ (IRSN), “Tonkin, de Lang-Son Province, an-Moi”; 1♀ (BMNH)
(Fig. 198), “Bangkok, 11.3.[19]30, Ariant” (handwritten), “Siam. 1930.
W.R.S.Ladell”, “Press. by Com. Inst. Ent. B.M. 1948–165”, “Tapinolachnus
lacordairei oms., D.J.Atkinsondet. 1948”; 1♂ (BMNH) (Figs196,199),
Vietnam, 8°43ʹN / 106°36ʹE, “Poulo Condor”, “Sharp Coll. 1905–313.”,
“269”; 1♀ (BMNH), Western Malaysia, “Penang (Lamb.) Pascoe Coll.”,
“Neocerambyx”; 1♀ (BMNH), “Java.”, “Bowring, 63–47*”, “7.”, “Tapinolachnus
lacordairei oms.” (upperside), “from description”(underside); 1♀
(BMNH), Lesser Sunda Islands, Indonesia, “Sumbava”, “Sharp Coll. 1905–
313.”; 1♂ (BMNH), Lesser Sunda Islands, Indonesia, “Flores.” “Fry Coll.
1905.100.”, “56980”, “Tapinolachnus lacordairei oms.”; 1♂, lectotype of
Aeolesthes (Mimoderolus) uniformis Pic, 1933 (MNHN) (photographs),
Figs 201–203. Derolydnus Hüdepohl, 1989 and Derolus Gahan, 1891, habitus, dorsal view, males.
201 – Derolydnus bisulcatus (Aurivillius, 1914) (photograph by Luboš Dembický); 202 – Derolus glauciapicalis Gressitt et Rondon, 1970 (from
ailand); 203 –D. argentesignatus Gressitt et Rondon, 1970 (from ailand).
Рис. 201–203. Derolydnus Hüdepohl, 1989 и Derolus Gahan, 1891, общий вид сверху, самцы.
201 – Derolydnus bisulcatus (Aurivillius, 1914) (фотография Л. Дембицкого); 202 – Derolus glauciapicalis Gressitt et Rondon, 1970 (из Таиланда);
203 –D. argentesignatus Gressitt et Rondon, 1970 (из Таиланда).
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 241
“Hoo-Binh, Tonkin”, “Aeolesthes sg. Mimoderolus uniformisn.sp.”, “Type”,
“Museum Paris, Coll. M. Pic”, “Holotype” (incorrect label); 2♂ (cAM),
Laos, Wapikhamthong Province, “Khong Sédone, 31.03.[19]65, 18.04.
[19]65”; 1♂, lectotype of Pachydissus xyliae Fisher, 1940, here designated
(BMNH) (Figs 195, 200), body length 32.3 mm, humeral width 9.5 mm,
Myanmar, “Dawebauk Res., Ataran, R.Hla Ogh Coll. 18.X.1937”, “exXylia
dolabriformis”, “R .R.S. 1073, B.C.R. 712”, “Cage 105, D.S.R. 382”, “Pachydissus
xyliae Fisher”, “Type”, “Brit. Mus. 1946–[?78]”, “I.R. 3080”, “330” (Fig. 234),
“Lectotypus ♂ Pachydissus xyliae Fisher, 1940, A.Miroshnikovdes., 2018”.
Remarks. When studying the above specimens,
I could not find any clear morphological differences
between them. On this basis, I suppose that Tapinolachnus
lacordairei (J.omson, 1864)= Tapinolachnus uniformis
(Pic, 1933), comb. n.= Tapinolachnus xyliae (Fisher,
1940), comb. n. However, given that so far I have been
able to revise the syntypes of the former two taxa from
photographs alone, this synonymy is established here only
provisionally. In addition, it is noteworthy that Vitali et al.
[2017] have recently synonymized T.uniformis comb. n.
and T.xyliaecomb.n., although the type specimens of the
latter species is not mentioned in the material they studied.
It seems also important that both a male and a female
with body lengths of 30–32 mm are indicated in the
original description of Aeolesthes (Mimoderolus) uniformis
[Pic, 1933: 11], i.e., Pic had in mind at least two specimens
which must be considered as syntypes. At the same time,
a male kept at MNHN, besides Pic’s designation “Type”,
carries a modern label “Holotype”. It is in this quality (i.e.
the holotype) that Vitali et al. [2017] referred to that type
specimen. Ido not know yet if the female mentioned in
Pic’s original description is still kept at MNHN or any
other collection, nor is it clear if he somehow designated it.
Nevertheless, taking into account the above, the male type
cannot be considered as the holotype (by monotypy), but is
to be designated as the lectotype.
Notes on the type locality. In the original
description, omson [1864: 232] referred to this species
as coming from “Malasia”, this also being noted (only in
an abbreviated form) on the label of one of the syntypes
(male) I have examined. In the same monograph, omson
described many other species from “Malasia”. In later
publications by various authors, including modern ones,
the distributions of the above taxa of omson are given
in different ways. So far some of them have been recorded
only from the continental part of Southeast Asia (mainly
Indochina) or, in addition, from the mainland South Asia
and/or southern China. Some other species are known only
in Borneo or, in addition, in Sumatra and/or other islands
of the region, whereas some further taxa are characterized
by wider distributions, being found in continental and/
or insular parts of these areas. As regards the above work
by omson [1864], it is also noteworthy that, in addition
to some of his new species, he referred to “Malasia” some
species that Pascoe [1857] had described from “Borneo”
and/or “Malacca”. At the same time, omson allotted
many of Pascoe’s species the same locality, i.e. “Borneo” or
“Malacca”.
However, whereas an insular distribution pattern has
since been confirmed for Utopia castelnaudii J. omson,
1864, which has hitherto been known only from Borneo
and Sumatra [Heffern, 2013]; plus the material from
various museums and private collections I have studied),
some records of Mythodes plumosa J.omson, 1864, on
the contrary, indicate that up to now this species has been
found only in Western Malaysia and Singapore.
Considering all above, it is impossible to find out,
even presumably, a more specific area of origin of the type
specimens of T.lacordairei than the one indicated in the
original description of the species.
Distribution. e distribution area of this species
is probably extensive. According to preliminary data, it
covers at least Indochina, including Malay Peninsula, as
well as Java and the Lesser Sunda Islands.
Genus Derolydnus Hüdepohl, 1989
Derolydnus Hüdepohl, 1989: 51; Heffern, 2013: 9.
Type species: Elydnus bisulcatus Aurivillius, 1914.
Composition. e genus includes a single species.
Distribution. Oriental realm.
Derolydnus bisulcatus (Aurivillius, 1914)
(Fig. 201)
Elydnus bisulcatus Aurivillius, 1914: 269, taf.1, fig.2 (“Borneo:
Lawas”). Type locality: Malaysia, Sarawak, Lawas (according to the
original description).
Derolydnus bisulcatus: Hüdepohl, 1989: 52; Heffern, 2013: 9.
Material. 1♂ (according to the original description), holotype, by
monotypy (NHRS) (photograph; Fig.201); 1♀ (NHMD), Burma, Tenasserim,
03.1996 (local collector), “Derolydnus bisulcatus Aur., O.Mehldet. 2014”;
1♀ (NHMD), same locality, 04.1996 (local collector), “Derolydnus bisulcatus
Aur., O.Mehl det. 2014”; a large series of males and females from Borneo
and Sumatra (NHMD; cAM).
Distribution. Until now, this species has only
been known from Borneo and Sumatra [Aurivillius,
1914; Hüdepohl, 1989]. Based on the material studied,
D. bisulcatus is being recorded here from Myanmar, as
from Indochina in general, for the first time. Iam also aware
of individual records from central Vietnam, according to
some data to be verified.
Genus Derolus Gahan, 1891
Derolus Gahan, 1891: 26 (Pachydissus subgen.); Gahan, 1906:
135; Aurivillius, 1912: 58; Winkler, 1929: 1142; Plavilstshikov, 1931:
85; 1940: 111, 640; Gressitt, 1951: 141; Gressitt, Rondon, 1970: 72;
Catalogue..., 2010: 159; Heffern, 2013: 9; Nga et al., 2014: 432;
Kariyanna et al., 2017: 28; Vitali et al., 2017: 59; Miroshnikov, 2017: 223.
Capnocerambyx Reitter, 1894: 356 (type species:
“C.mauritanicus Luc[as].” (sic)).
Type species: Hammaticherus mauritanicus Buquet,
1840, by subsequent designation [Gahan, 1906].
Remarks. e largest genus of the tribe in terms of the
number of species, comprising almost 70species. Adlbauer
[2009] reviewed its African representatives. e Asian
group of species needs a detailed revision and a diagnostic
re-evaluation of the genus as a whole.
Below are some new records of two little-known
species, both somewhat expanding their distribution areas.
Derolus glauciapicalis Gressitt et Rondon, 1970
(Fig. 202)
Derolus glauciapicalis Gressitt et Rondon, 1970: 75. Typ e
locality: Laos, Sayaboury, 170 m (according to the original
description).
242 A.I. Miroshnikov
Figs 204–234. Cerambycini Latreille, 1802, labels of type and other specimens.
Рис. 204–234. Cerambycini Latreille, 1802, этикетки типовых и других экземпляров.
204–205 – Plavichydissus semiplicatus (Pic, 1926), comb. rest.; 206–207 – P. grossepunctatus (Gressitt et Rondon), comb.n.; 208 – P. sulcicollis
(Gahan, 1893), comb. n.; 209 – P. rufipennis (Pic, 1923), comb. rest.; 210 – Pachydissus parvicollis Gahan, 1891; 211 – P. argentatus Pic, 1923; 212 –
P.birmanicus Gardner, 1926; 213–214 – Margites auratonotatus Pic, 1923; 215 – M. egenus (Pascoe, 1858); 216 – M. fulvidus (Pascoe, 1858); 217 – M.modicus
Gahan, 1906; 218 – M. luteopubens Pic, 1926; 219 – M. lajoyei Pic, 1926; 220 – Laomargites singularis Pic, 1923, comb. rest.; 221–222 – Dymasius simplex
Gressitt et Rondon, 1970; 223 – D. prominor Gressitt et Rondon, 1970; 224 – D. parvus Gressitt et Rondon, 1970; 225 – D. niger Gressitt et Rondon, 1970;
226– Lamellocerambyx laosensis Pic, 1923, comb. rest.; 227 – Diorthus cinereus (Fabricius, 1793) (holotype of Diorthus simplex (White, 1853)); 228–229 –
D.vagus (Gahan, 1891); 230–231 – D. sericeus Gardner, 1939; 232–233 – Tapinolachnus lacordairei (J. omson, 1864); 234 – T. ?lacordairei (J. omson,
1864) (lectotype of Tapinolachnus xyliae (Fisher, 1940), comb. n.).
204–205, 210, 213–214, 232–233 – syntypes; 206, 208–209, 211–212, 215–216, 218–221, 223–228 – holotypes; 217, 231 – lectotypes; 207, 222 –
paratypes; 230 – paralectotype. 204–205, 209, 211, 213–214, 218–219 – photographs by Gérard L. Tavakilian; 212 – photograph by Sudhir Singh; 232–233–
photographs by Azadeh Taghavian.
204–205, 210, 213–214, 232–233 – синтипы; 206, 208–209, 211–212, 215–216, 218–221, 223–228 – голотипы; 217, 231– лектотипы; 207,
222– паратипы; 230 – паралектотип. 204 –205, 209, 211, 213–214, 218–219 – фотографии Ж.Тавакиляна; 212 – фотография С. Сингха; 232–233 –
фотографии А. Тагвьян.
e longicorn beetle tribe Cerambycini Latreille, 1802 (Coleoptera: Cerambycidae: Cerambycinae) 243
Material. 1♂, holotype (BM) (photograph); 1♂ (cAM) (Fig. 202),
NWailand, Lamphun, Mae a, 20.04.2011 (local collector).
Distribution. Until now, this species has only been
known from Laos [Gressitt, Rondon, 1970]. Based on the
material studied, D. glauciapicalis is being recorded here
from ailand for the first time.
Derolus argentesignatus Gressitt et Rondon, 1970
(Fig. 203)
Derolus argentesignatus Gressitt et Rondon, 1970: 76. Type
locality: Laos, Vientiane Province, Nong Tevada, 170m (according
to the original description).
Material. 1♂, holotype (BM) (photograph); 1♂ (cAM) (Fig. 203),
NW ailand, Mae Hong Son Prov., Pai env., ~600m, road on Mae Yen
waterfall, 19°21ʹ42ʺN / 98°27ʹ46ʺE – 19°22ʹ01ʺN / 98°30ʹ29ʺE, 27.04–
9.05.2013 (leg. I. Melnik); 1♀ (cAM), N ailand, Chiang Rai Prov., Doi
Chang env., 640–750m, 19°46ʹ01ʺN/ 99°28ʹ11ʺE– 9°47ʹ44ʺN/ 99°27ʹ06ʺE,
11–15.05.2013 (leg.I.Melnik).
Distribution. Until now, this species has only been
known from Laos [Gressitt, Rondon, 1970]. Based on the
material studied, D.argentesignatus is being recorded here
from ailand for the first time.
Errata
Since several of my previous publications
[Miroshnikov, 2016, 2017, 2018; Miroshnikov, Tichý, 2018]
contain some misprints, their corrections are listed below.
In these works, when comparing the lengths of
individual segments of the posterior tarsus, mainly in
the species descriptions, instead of the correct terms and
connotations “metatarsomere 1” and “metatarsomeres 2
and3 combined”, “tarsomere 1” and “tarsomeres 2 and 3
combined” are mistakenly indicated.
e corrections that should also be made to
Miroshnikov [2017] see in Table 1.
Besides this, in preparing the layout of the manuscript
of Miroshnikov [2017] and checking the spelling of often
repeated names of its sections, due to a software failure,
the spelling of the Distribution section in a number of
cases turned out to be erroneous, namely, “Disribution”.
Unfortunately, this incorrect spelling remained unnoticed
when the layout of the corresponding journal volume was
delivered to the printing house.
Acknowledgements
I am very grateful to Svetlana V. Andreeva (ZIN),
Maxwell V.L. Barclay and Michael F. Geiser (BMNH),
JamesH. Boone (BM), ierry Deuve, Azadeh Taghavian
and GérardL. Tavakilian (MNHN), Alain Drumont (IRSN),
AlexeyA. Gusakov (ZMMU), AlexeyYu. Solodovnikov and
Sree Gayathree Selvantharan (NHMD) for the opportunity
to study the museum material, to Luboš Dembický (Brno,
Czech Republic) and SergeyV. Murzin (Moscow, Russia),
who have provided various specimens from their private
collections. I would like to express my sincere thanks to
AlexeyYu. Solodovnikov for his kind permission to retain
some material in my personal collection, to Dmitry N.
Fedorenko (Institute for Problems of Ecology and Evolution,
Moscow, Russia), who was funded by the Russia-Vietnam
Tropical Center, for the valuable material he rendered
to me for study. I am deeply indebted to Alexandr G.
Kirejtshuk (ZIN), again to SergeyV. Murzin and AlexeyYu.
Solodovnikov who helped a lot in my prompt receipt of
the material for study, to Sudhir Singh (NFIC), HemantV.
Ghate (Department of Zoology, Modern College of Arts,
Science and Commerce, Shivajinagar, Pune, India), Karl
Adlbauer (Graz, Austria), Nobuo Ohbayashi (Kamimiyada,
Miura City, Japan), again to Alexey Yu. Solodovnikov,
Azadeh Taghavian and GérardL. Tavakilian for the helpful
provision of various pictures and/or valuable information,
to omas Pape (NHMD), who advised me on certain
nomenclatural issues, to the reviewers for their helpful
comments. I give special thanks to Kirill V. Makarov
(Moscow Pedagogical State University, Moscow, Russia) for
having rendered his great help in the preparation of almost
all photographs, and to Luboš Dembický who generously
shared the pictures of the holotypes of many species of the
tribe Cerambycini. Last but not least, Iam most grateful to
my wife Tatiana who helped a lot in the preparation of the
illustrations for publication.
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Received / Поступила: 6.11.2018
Accepted / Принята: 15.12.2018
246 A.I. Miroshnikov
