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VOLUMINA JURASSICA, 2017, XV: 181–186
DOI: 10.5604/01.3001.0010.7467
Developments with fixing a Tithonian/Berriasian (J/K) boundary
William A.P. WIMBLEDON1
Chairman, Berriasian Working Group (ISCS)
At the Vienna Cretaceous Symposium (August 2017), the Berriasian Working Group (International Subcommission on
Cretaceous Stratigraphy) gave an account of its concerted research activities since 2009 (Wimbledon et al., 2017) – that and
this text are abbreviated summaries of Tithonian–Berriasian studies carried out in preparation for the consideration and
choice of a GSSP. Past decisions have consistently stated that such a GSSP should be dened in Tethys. Of Panthalassa, only
a comparatively small area of Lower Cretaceous ocean oor survives (NW Pacic), and subduction effected similar losses in
the Arctic, and thus parts of tropical-subtropical Tethys constitute the largest geographical entity remaining from J/K times,
and the one with the richest, most widespread and most studied biota.
In the 1970s to 1990s, discussions in various working groups about a J/K boundary made no substantive progress
(Remane, 1991), and were suspended until more auspicious times came. By the early 2000s, J/K correlation had already
shifted away from a concentration on often endemic ammonites, which had repeatedly been recognised as a major obstacle
to correlation, even in western Tethys. In 2007, a new Berriasian Working Group (ISCS) was mooted and it initiated a new
phase of activity, concentrating on realities and rening Tithonian–Berriasian correlations, partly directed at xing a J/K
boundary. For this reason, the preoccupations of the earlier Cretaceous symposia and earlier working groups, dominated by
ammonite discussion, have been put aside and since 2009 work has refocussed on integrated high-resolution studies, includ-
ing ammonites, but more on the effective use of several microfossil groups (e.g. calpionellids, calcareous nannofossils, cal-
careous dinoagellates), which give greater precision, calibrated with magnetostratigraphy. The J/K interval is without any
marked chemostratigraphic event that can help us in xing a boundary, isotopic excursions are lacking (Price et al., 2016),
and thus correlative methods must be predominantly biostratigraphic and magnetostratigraphic. The WG has focussed on the
detailed documentation of numerous key proles, old and new, putative J/K levels, calibrating magnetostratigraphy with fos-
sil range data, and examining prospective primary marker levels formerly suggested: e.g. the bases of the Jacobi Subzone, the
calpionellid Alpina Subzone, of M18r, and the Grandis Subzone. Many localities, from California to Tibet and the Russian
Far East, have been documented and assessed, including classical localities in France, Crimea, etc. Of course, long-range
correlations to some remote boreal regions, with impoverished, endemic biotas (vis a vis Tethys), as well as the extensive
non-marine basins, remain approximate. Though much work has recently been put into the magnetostratigraphy of the Pur-
beck Formation in England.
Documentation of classical Berriasian areas has led to some signicant revisions: we have highlighted the absence
of ‘Berriasella’ jacobi in the lower part of the nominal Jacobi Subzone (Frau et al., 2016), and the predominance of Delphi-
nella (e.g. France and Ukraine): thus Strambergella jacobi (and the Jacobi Subzone) has been ruled out as a GSSP marker.
Continuing problems with separating the ammonite assemblages of the Jacobi and Grandis subzones of past authors have
meant that we have not pursued the Grandis Subzone’s base as a putative stage base (Fig. 1). Better resolution also has shown
that the base of the Alpina (calpionellid) Subzone and the Jacobi (ammonite) Subzone do not coincide, and the Elliptica Sub-
zone base is below that of the Occitanica Zone.
1 School of Earth Sciences, University of Bristol, U.K.; mishenka1@yahoo.co.uk.
182 William A.P. Wimbledon
Another boundary alternative put forward in the past, the base of magnetozone M18r, has not proved to be effective as
a J/K marker. It is widely identied, of course, but it cannot be calibrated with any widespread fossil marker: its proposal
(e.g. Arkadi’ev et al., 2017) has not been supported by any documentation of biotic events (calpionellid, ammonite or nan-
nofossil) at, or bracketing the base of M18r.
For the last twenty years, building on a considerable and growing body of literature. calpionellids have been seen by nu-
merous authors as the most useful fossil group that could provide a J/K marker. Further, the turnover from Crassicollaria and
large Calpionella to small orbicular Calpionella alpina (together with Crassicollaria parvula and Tintinopsella carpathica)
has been documented repeatedly as the most consistent and widespread marker in the middle of magnetic subzone M19n.2n,
as recognised by the clear consensus amongst specialists at the Warsaw J/K workshop (Wimbledon, 2013). Work then in
progress has since come to fruition and the Alpina level has been proven further east in Tethys, to Arabia and Iran, and west
into the Americas (North and South). In June 2016, a formal ballot of the Berriasian WG led to a decisive vote that selected
the Alpina Subzone base as the primary T/B boundary marker.
The FADs of calcareous nannofossil species are helpful indicators of the base of Alpina Subzone, as they bracket this
level. But their ranges have been modied signicantly due to recent research (Fig. 2), undermining the 1980s biozonation of
Bralower. However, records of the FAD of Hexalithus strictus [=H. geometricus] have a restricted vertical distribution, strad-
dling the base of the Alpina Subzone in 19n.2n. The FAD of Nannoconus steinmannii minor occurs in mid M19n.2n: imme-
diately above the Alpina Subzone base (Puerto Escaño) or just below it (Rio Argos). Cruciellipsis cuvillieri, N. wintereri and
N. globulus globulus occur a little lower, though at Puerto Escaño the FAD of the rst species, unusually, is low in M19n.2n.
The FAD of N. wintereri occurs just below the Alpina Subzone’s base (e.g. Le Chouet, Puerto Escaño). Nannoconus kampt-
neri minor has latterly been recorded as rst appearing in the upper half of M19n, e.g. at Theodosia, Puerto Escaño and Le
nodiger singularis rjasanensis
M17r
anguiformis
?
Jacobi Grandis
Western Tethys
M18r
M19r
prim.preplic. lamplughi runctoni kochi
Purbeck Limestone Formation
?
okensis taimyrensis chetae sibiricus kochi
Tithonian Berriasian
Colomi Alpina Ferasini
Elliptica
Crassicollaria Calpionella
α
β
Published (1965) colloquiumdecision on J/K boundary
Published (1975) colloquiumdecision on J/K boundary
β
α
Micr.
Sea
Substeueroceras
koeneni
damesi
?
?
?
M17r
M18r
M19r
?
?
Praetintinnop.
Chitinoidella
Interm.
Remanei
Andreaei
Occitanica Boiss.
napaensis tehamaensis knoxville.
Nordvik
Russian
Platform
?
fulgens subditus
M19n.2n M19n.2n
M18n M18n
N. steinmannii minor
N. kamptneri minor
N. wintereri
R. asper
M16r M16r
Cinder
Bed
Mammal
Bed
Corongoceras
alternans
*
?
?
?
Euxinus
N. steinmannii steinmannii
N. kamptneri minor
N. erbae
N. wintereri, H. strictus,
C. cuvillieri, N. globulus globulus
N. globulus minor
N. puer
L. carniolensis
C. octofenestratus
C. surirellus, R. asper
N. kamptneri kamptneri
Arroyo Las
Loncoche Loicas
W. i.
Argenticeras
noduliferum
S. koeneni
?
?
Spiticeras
damesi
Cr.
Ch.
A.n.
M20n.2nM20n.2n
N. steinmannii minor
North Dorset
Simplex
Calpio-
nellopsis
Spiticeras
Ch.
A.n.
Chitinoidella
Cr. Crassicollaria
Argentiniceras noduliferum Zone
W.i. Windhauseniceras internispinosum
Fig. 1. Correlations of Tethys with austral and boreal regions
183Developments with xing a Tithonian/Berriasian (J/K) boundary
Chouet. Even more striking, N. kamptneri kamptneri and N. steinmannii steinmannii, previously used as infallible biozonal
indicators in M17r, have been found widely in lower M18r and the upper half of M19n (Figs. 1, 2)
In recent times, the western Tethyan core area with denite and precise microfossil correlations has been considerably
expanded. In particular, there have been nds of Lower Berriasian nannofossil markers in N. Africa, Ukraine, Yemen, Iraq,
Tibet and the Andes, C. alpina records in the Middle East and the Andes, the unambiguous application of the “western
Tethyan” calpionellid scheme to Mexico (Lopez Martinez et al., 2013), Arabia (Celestino et al., 2016), northern Iraq (Wim-
bledon et al., 2016) and Iran (Benzaggagh et al., 2012), and magnetostratigraphic studies extended to California, the Andes
and N. Africa for the rst time. Denite calpionellid records in Australia, Papua and New Guinea etc. await fuller investiga-
tion, as do problematic nannofossils and radiometric dates (Liu et al., 2013) in southern Tibet.
Interesting recent unpublished results from Fiume Bosso and Puerto Escaño reveal the calpionellid Ferasini Subzone has
its base in M19n.2n, with, consequently, a more thinly developed Alpina Subzone. Which perhaps raises questions about the
accuracy of earlier results for the Ferasini Subzone at these and other sites, and discrimination between M19n.1r and M18r
in previous accounts.
Failed attempts at identifying calpionellids in the Andean sequences were related by Remane, but critical rst nds by
Fernandez Carmona and Riccardi (1998, 1999) have given impetus to a breakthrough, with identication of the Chitinoidella
Zone (Kietzmann et al., 2011) and the Crassicollaria Zone in the Corongoceras alternans biozone (Kietzmann, 2017).
Though putative nannofossil proxies for the base of the Alpina Subzone (thus circa mid M19n) were recorded at Las Loicas
(Vennari et al., 2014), close to the base of the Argentiniceras noduliferum Zone, no Calpionella was found. Palaeomagnetic
Nannoconus wintereri
Cruciellipsis cuvillieri
N. steinmannii minor
Cretarhabdus octofenestratus
N. kamptneri minor
N. steinmannii steinmannii
N. kamptneri kamptneri
Nannoconus globulus globulus
Cretarhabdus surirellus
Hexalithus strictus =H. geometricus[ ]
Rhagodisus asper
Nannoconus erbae
Nannoconus globulus minor
Umbria granulosa granulosa
Nannoconus puer
Umbria granulosa minor
N
annoconus infans
C. angustiforatus
142
143
144
145
146
147
19r
18r
17r
Alpina
Rem. Inter. Colomi
Chitin.
Para.
Dalmasi
Mircocanthum Andread. Jacobi
Grandis
Tintinopsella
Subalpina
/Privasensis
Ferasini Elliptica
18n
17n
19n.2n
20n.2n
TITHONIAN BERRIASIAN
Theodosia
Nutzhof
Puerto Escano
Rio Argos
Le Chouet
Arcevia
Foza
Sidi Khalif
Lokut
Kurovice
Font de
St Bertrand
Fig. 2. FADS of selected calcareous nannofossils in the Upper Tithonian to Lower Berriasian, calibrated with magnetozones and calpionellid biozones
(modified from Wimbledon, 2016, Wimbledon et al., 2017, Bakhmutov et al., in press)
Continuous vertical lines shows the distribution of FADS, in publications up to 2009 (after Casellato, 2010). Coloured spots represent records of FADS at lower
stratigraphic levels (After Speranza et al., 2005; Lukeneder et al., 2010; Casellato, 2010; Wimbledon et al., 2013; Hoedemaeker et al., 2016: Svobodova, Kostak,
2016; Grabowski et al., 2017; Švábenická et al., 2017; Bakhmutov et al., 2018, in press; Ebra et al., 2018, in press; Gardin, unpublished). NB: At Le Chouet,
magnetic subzone M19n.1r was not detected, so although N. steinmannii minor and N. kamptneri minor are shown in M19n.1n, their FADs may fall in M19n.2n.
The record of N. wintereri in M19r at Nutzhof is currently being re-examined
184 William A.P. Wimbledon
results from Las Loicas are still awaiting publication, but those from Arroyo Lonconche (Llanos et al., 2017) are intriguing,
as they put the Noduliferum Zone base not in magnetozone M19n, but higher, in M16r. At Las Loicas, Lopez Martinez et al.
(2017) have since identied an assemblage with predominant small globular Calpionella alpina (albeit co-occurring with
Crassicollaria brevis, C. remanei and C. massutiniana) and the Alpina Subzone at the base of the Noduliferum Zone (Fig. 1).
All this makes it an exciting time for Tethyan: Andean correlations.
Work must now focus on expanding efforts to identify proxies for the C. alpina level in the more problematic regions. For
instance, the application of belemnites in Siberia and the Pacic (Dzyuba, 2012). It seems that the comparatively little studied
Siberian boreal areas, though with disappointingly poor preservation of signicant nannofossil taxa (Zanin et al., 2012) and
a discontinuous record of endemic ammonites near the boundary (Schnabl et al., 2015) (Fig. 1), still have considerable cor-
relation potential with other fossil groups, and they need better (and much more) targeting of new localities for magnetostrati-
graphic study. In this context, Vishnevskaya (2017) records calcareous dinoagellates in Siberia (Stomiosphaerina proxima,
Colomisphaera tenuis, C. lapidosa, and C.? fortis) with radiolarians in the presumed Berriasian Bazhenovo Formation. This
demands more research, as, if conrmed, they would be of use as proxies for calpionellids.
Calcareous dinoagellate cysts are common fossils in the Tithonian to Berriasian of European and North African Tethys.
They appear to have much potential as accessories to the calpionellids, though there are rather few recent accounts and the
stratigraphic ranges of some species has yet to become static (e.g. Stomiosphaera moluccana and Colomisphaera pieniensis:
Wimbledon et al., 2013; and below). But of notably useful species, in the Balkans, the rst appearance of S. proxima has been
recorded in the upper Crassicollaria Zone, and said to have its FAD at the base of the Colomi Subzone, though it appears rst
in the Remanei Subzone in SE France. This restriction to the Crassicollaria Zone was emphasized by Rehakova (2000) (see
also Lukeneder et al., 2010: but note Lopez et al., 2013).
C. fortis only just pre-dates S. proxima (appearing pre-Crassicollaria Zone), and its range straddles the upper Crassicol-
laria and Calpionella zones, affording a wider stratigraphic bracket: this proves to be the case in some French sites (e.g. Tre
Maroua and St Bertrand). If such occurrences could be conrmed and consistently proved in Siberia (and perhaps the Russian
Platform), it would be a step forward in accuracy in trying to correlate with the Crassicollaria/Calpionella zonal boundary.
Vishnevskaya’s FAD of C.? fortis in Siberia appears to be very high (Analogus Zone) and that of S. proxima is shown lower
than one should expect (Exoticus Zone, ?M20n) as compared to Tethys, when the Alpina J/K boundary (M19n.2n) has been
correlated with the Siberian Taimyrensis Zone. However, the original assignment of magnetozones numbers in Siberia, at
Nordvik, may merit reconsideration. The reversed intervals, identied as M19n.1r, M18r and M17r, are of limited, and simi-
lar, thickness (Schnabl et al., 2015). With a fresh mind and no preconceptions, perhaps one might reconsider the original
magnetozone notation of Houša et al. and assign different numbers. That being as it may, more research on calcareous dino-
agellates is an absolute priority in boreal regions.
Most recently, Ivanova and Kietzmann (2017) record dinoagellate cysts in the Andes: Colomisphaera tenuis, C. fortis,
S. proxima (lower Noduliferum Zone) and S. wanneri (upper Noduliferum Zone). The S. wanneri biozone (Arroyo Lonco-
che) they characterise as Upper Berriasian (Noduliferum to Damesii zones). S. wanneri has been found higher in the
Berriasian in the Carpathians, but it has also been collected in the Lower Berriasian in southern Ukraine (Bakhmutov et al.,
2018, in press), and it seems that it appears similarly early in France (Font de St Bertrand – Elliptica Subzone: Tre Maroua
– Simplex Subzone), as does C. vogleri (Simplex Subzone). Kietzmann and Llanos (2017), discussing the results of Lopez
Martinez et al. (2017), would correlate S. wanneri and S. proxima with the European Berriasian, as above, and have Nanno-
conus steinmannii steinmannii in the Andes indicate correlation with M17r in Tethys: but see Fig. 2.
CONCLUSIONS
• No fossil species has anything like a global distribution in the Tithonian-Berriasian interval. Though Calpionella alpina
is the most widespread.
• The base of M18r does not coincide with any consistent biotic marker at sites recently studied for magnetostratigraphy,
and the same is true of the base of M19n.2n.
• The traditional Berriasella jacobi Subzone will play no part in any denition of the base of the Berriasian, nor can the
species Berriasella jacobi [=Strambergella jacobi].
• Calpionellids (calibrated with magnetostratigraphy) provide the most effective primary J/K boundary marker, the Cras-
sicollaria to Calpionella turnover, i.e. the Colomi/Alpina subzone boundary as shown in Figs 1, 2.
185Developments with xing a Tithonian/Berriasian (J/K) boundary
• In June 2016, the Berriasian Working Group voted, by a large majority (76%), to adopt the Crassicollaria/Calpionella
turnover and base of the Alpina Subzone as the primary marker for the base of the Berriasian Stage: the WG now focusses
on identication of the best candidate site for a GSSP.
• This level is bracketed by a number of nannofossil FADs (Fig 2).
• Correlative advances in Argentina are exciting, and amplication of the rst palaeomagnetic results and of recent C. al-
pina nds are anticipated.
• In austral regions, most notably in the Andes, some of the Tethyan nannofossil marker species have been identied (along-
side endemic ammonites), including species which are proxies for the base of the Alpina Subzone, as they have in Tibet.
• New nds of Tethyan calcareous dinoagellate cysts in the Andes indicate their usefulness and potential as secondary
markers.
• In Siberia, at one site, Nordvik, it is possible to approximate the horizon of Calpionella alpina (in mid M19n.2n): close
below the base of the Tehamaensis (belemnite) Zone and close above the ‘oating’ Taimyrensis (ammonite) zonal base.
Earlier nds of calcareous nannofossils in Siberia have yet to be followed up. Identications there of calcareous dinoag-
ellates have great potential, and it is to be hoped they will provide proxies for calpionellid datums, as in austral regions.
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