PNWMS Technical Report: 1998–2005 Green Peak Density Management Fungal Community Study: 2017 updated summary

Abstract

An unpublished report submitted in 2008 summarized the outcomes of a seven-year study evaluating the short-term response of epigeous ectomycorrhizal basidiomycetes to five different density treatments in a 65-year old Douglas-fir forest in the Oregon coast range. That report supplemented information provided in Norvell & Exeter’s 2004 publication, “Ectomycorrhizal epigeous basidiomycete diversity in Oregon’s coastal montane Pseudotsuga menziesii forests – preliminary observations.” This revision includes illustrations and analyses of the 2008 summary and provides new taxonomic reports, a complete bibliography, and a nomenclaturally refined species list.

Full text

PNW-MS Technical Report—
1998–2005 Green Peak Density Management
Fungal Community Study: 2017 updated summary
Hebeloma crustuliniforme complex, g2051109h1-7
Lactarius cf. chrysorheus g2051109h2-15
Inocybe kauffmanii, g2051109l2-5
Suillus caerulescens, g2051026u2-2
Hydnum cf. ‘oregonense’, g2041122h1-21

PNW-MS Technical Report: 1998–2005 Green Peak Density Management Fungal Community
Study: 2017 updated summary. Prepared by Lorelei L. Norvell. March 31, 2017. 28 pp.
ResearchGate link: www.researchgate.net/publication/315734877
AbstrAct— An unpublished report submitted in 2008 summarized the outcomes of a seven-year
study evaluating the short-term response of epigeous ectomycorrhizal basidiomycetes to ve
different density treatments in a 65-year old Douglas-r forest in the Oregon coast range. That
report supplemented information provided in Norvell & Exeter’s 2004 publication, “Ectomycorrhizal
epigeous basidiomycete diversity in Oregon’s coastal montane Pseudotsuga menziesii forests –
preliminary observations.” This technical report includes illustrations and analyses of the 2008
summary and provides new taxonomic reports, a complete bibliography, and a nomenclaturally
rened species list.

1998–2005 Green Peak Density Management Fungal Community Study:
2017 updated summary
Prepared by Lorelei L. Norvell, PhD
Pacic Northwest Mycology Service
Portland, Oregon
In October 1998, Pacic Northwest Mycology Service (PNWMS) mycologist Dr Lorelei Norvell
(Portland, Oregon) and USDI-BLM Mary’s Peak botanist Ron Exeter (Salem, Oregon) jointly initiated
a ve-year fungal community/density management study. Their goal was to assess the short-term
response of epigeous-fruiting ectomycorrhizal and other agged (“C-3”) basidiomycetes in a 65-
year old Douglas-r forest scheduled for ve different density treatments. For the next seven
years (of which six were funded), the research team surveyed and monitored fungi from the plots.
During the nal 2005 eld season, they gathered 480 fungal collections from the ten transects in
the Green Peak BLM Research Forest in Benton County that Norvell photographed, provisionally
identied, examined, and processed for curation in the PNWMS Mycological Herbarium, where
they remain for on-going reevaluation and examination. The nal 2005 eldwork brought to 4301
the total number of collections vouchered during the 7-year study.
Early interim reports were submitted for 1998 (on 6/2/00), 1999 (10/15/01), 2000 (1/8/02), 2001
(9/17/02), 2002 (9/13/04) and 2004 (1/8/07). (Work conducted during the unfunded 2003 period
was reported within the 2004 interim report and included in the ‘nal’ summary and database
submitted on June 30, 2008.) Norvell & Exeter’s 30-page book chapter, “Ectomycorrhizal epigeous
basidiomycete diversity in Oregon Coast Range Pseudotsuga menziesii forests: Preliminary
observations” (2004, in Fungi in Forest ecosystems) summarizing observations from the rst ve
project years served as the ofcial project publication. Although additional taxonomic research
is still needed for a nal analysis of the impact of timber thinning techniques upon the fungal
community, that paper has been well received by the mycological community and is cited in 40
research publications worldwide.

4 ... 1998–2005 Green Peak Density Management Fungal Community Study
1998–2005 Fungal Collections and Species Table: July 4, 2008 update
october 28, 1998 – november 9, 2005
Total sampling visits: 35 [Autumn: 26 Spring: 9]
Collections Total Targeted
Number of collections 4301 2980
Number microscopically veried to date 1134 [26%] 835 [28 %]
Number with macrophotos 2176 [51 %] 1373 [46 %]
Species Total Targeted
Number of species 524 (458 named) 264 (249 named)
Project species — annual totals
1998 174 118
1999** [harvest year: no fall patch cut] 131 70
2000 174 120
2001 186 121
2002 134 63
2003 autumn unfunded 115 54
2004 spring unfunded 283 146
2005 162 81
Number microscopically veried to date 302 [58 %] 190 [72 %]
Number with macrophotos 392 [75 %] 197 [75 %]
New species added annually 1998 base> 174 (157 named) 118 (109 named)
1999** [harvest year: no fall patch cut] 71 (66 named) 29 (28 named)
2000 63 (54 named) 39 (38 named)
2001 61 (55 named) 31 (28 named)
2002 31 (26 named) 5 (2 named)
2003 autumn unfunded 28 (24 named) 9 (9 named)
2004 spring unfunded 79 (55 named) 35 (31 named)
2005 17 (13 named) 4 (4 named)
Plot comparisons [1998–2005] Target collections
[% total targeted]
Target species
[% spp identied]
Control (uncut = U) 826 [28%] 159 [64%]
High retention (120 trees/acre = H) 763 [26%] 143 [57%]
Medium retention (80 trees/acre = M) 773 [26%] 147 [59%]
Low retention (40 trees/acre = L) 393 [13%] 102 [41%]
No retention (clear/patch cut = C) 221 [7%] 82 [33%]

2017 summary (Norvell) ... 5
1998–1999 v 2000–2005 target species Pre-treat targets Post-treat targets
Control (uncut = U)
98-05 targets = 159 75 [47%] 141 [89%]
High retention (120 trees/acre = H)
98-05 targets = 143 65 [45%] 125 [87%]
Medium retention (80 trees/acre = M)
98-05 targets = 147 60 [41%] 128 [87%]
Low retention (40 trees/acre = L)
98-05 targets = 102 65 [64%] 55 [54%]
No retention (patch cut = C)
98-05 targets = 82 (**= 18 spp subtracted;
found only at bases of uncut buffer trees)
59 [72%]
(no samples fall ’99)
39 [48%]
**21 [26%]
Data obtained from completion of an indifferent 2005 eld season did not substantially alter any
of the conclusions reported in the 2004 publication, although 4 new mycorrhizal targets (Lactarius
chrysorheus cf., L. subammeus, Russula pseudo-olivascens, R. mariae) and ve incidentals (Hygrophoropsis
aurantiaca, Mycena acicula, Mycena tenerrima, Polyporus badius, Psathyrella candolleana) were recorded
for the rst time. That book chapter, therefore, will remain the ofcial preliminary report along
with the project database accompanying this summary in the Salem BLM ofce. The above graph
and tables summarize the species and collections data contained in the database submitted in 2008;
they do not, however, reect the taxonomic and nomenclatural updates reported in 2017, and new
taxonomic discoveries are expected from re-evaluation of species concepts and further microscopical
work. Norvell will send further reports and updated databases as new information becomes available.
Observations regarding species obtained during the density management fungal project were also
published in Norvell & Redhead’s 2000 paper transferring the NWFP-agged species Pholiota albivelata
to the genus Stropharia . The 157-page monograph, RamaRia oF the PAciFic northwestern united stAtes
by Exeter, Norvell & Cázares (2006) quickly became a respected Ramaria resource cited in 29 research
publications worldwide. Norvell also presented the Exeter & Norvell Ramaria poster at the 2007 annual
Mycological Society of America meeting in Baton Rouge, LA. Both the book and poster cited observations
or published photos obtained during the Green Peak study. Finally, the 228-page monograph, Norvell
& Exeter’s Phaeocollybia in western north AmericA, published by the USDI-BLM Salem Ofce in 2009 (as
“2008”) also cited observations from the Green Peak Study. A complete bibliography of publications,
reports, and presentations incorporating data from the density management project along with a list of
publications citing these works is presented at the end of this summary.
2008 dAtAbAse Adjustments And keys to Fungi oF douglAs-Fir Forests: The Green Peak density management
database incorporates data on all 4301 project collections now curated in the Pacic Northwest
Mycology Service herbarium. Names of collections not yet microscopically conrmed were changed
in the 2008 database to link collection numbers with names of similar species to eliminate ‘singlet’
species; this provided more realistic tallies for computing plot totals for the above graph and tables, but
has not yet been updated to reect the recent nomenclatural changes shown in the species list below.

6 ... 1998–2005 Green Peak Density Management Fungal Community Study
The database will continue to evolve as collections are examined in species-by-species or genus-by-
genus lots, the most efcient way to resolve difcult species concepts and provide an accurate fungal
assessment of each treatment plot; it will also take time to consult with Cortinarius and Russula experts
regarding the taxonomy and nomenclature in those difcult genera. We hope that such systematic re-
evaluation of all collections and nomenclature will lead to reliable conclusions regarding the impact
of treatment type on the fungal community and the completion of keys to Cortinarius, Russula, Inocybe,
Galerina, Laccaria, Gomphidius, Lactarius, Suillus, Mycena and omphalinoid species in Douglas-r forests to
be added to the keys to Ramaria, Phaeocollybia, and chanterelles already completed.
PhotogrAPhic record: As of 2017, the DVD containing photos of ~200 identied target species and
~170 incidental species (a nal study goal) remains in current preparation. Unfortunately, a computer
failure in 2003 and the 2007 crash of an external hard drive was followed by a second massive computer
crash in 2009 has delayed preparation of a CD containing photos of all 2176 collections photographed.
Some digital les are irretrievably lost, and I still must scan negatives and transparencies for 1998–2001
collections. Once completed, all photos will be made available online or in the cloud and thus accessible
for taxonomic consultation and public use.
DiScuSSioN
PremAture hArvest comPromises Pre-treAtment bAseline dAtA: Norvell & Exeter (2004) noted that an
ecological study of the impact of different timber thinning strategies upon the associated fungal
communities must surmount many challenges to provide reliable data. We wish to emphasize some of
those concerns here. It is unfortunate that the timber harvest was carried out at the beginning of the
1999 fall season, not in 2000 as scheduled when we designed the study. At least two years of baseline
data are needed for reasoned statistical analysis. One year is insufcient, as much of that rst year
must be devoted to establishing and then ne-tuning efcient sampling protocols and identication
techniques.
The three partial thin plots had already been harvested prior to the rst 1999 sampling visit, the
timing of which was dictated by the onset of autumn rains and fungal fruiting. Even though the low-
retention, moderate-retention, and high-retention plots were already thinned, the fungi sampled that
fall must be counted as part of the pre-treatment totals. Unfortunately, fruitbodies were obscured in
the heavily thinned plots by branches and slash proving impossible to locate with slash several feet high
and virtually impenetrable along the low-retention transects, only somewhat more manageable on the
moderate-retention transects, yet easily managed within the high-retention plot, although there a deep
skid road cut deeply across both transects. [Interestingly, fruitbodies representing several uncommon
species proved particularly abundant along this groove in later years.]
Unfortunately, the upper control and clear (patch) cut plots had not undergone treatment on the
rst visits, so that there sampling surveys were conducted as usual. Also unfortunately, the team was
barred from monitoring the patch cut plot during the 2nd visit and both the control and cut plots during
the nal fall visit, thereby reducing the pre-treatment species tallies for those two important reference
sites.
Despite the sporadic sampling times and incomplete pre-treatment data, we decided to use what
data we do have by generating individual stand productivity ratios to compare pre-treatment and post-
treatment tallies separately with the cumulative tally (i.e., pre-treatment + post-treatment) for each
treatment stand.

2017 summary (Norvell) ... 7
other hArvest-relAted Problems: Retained trees were fairly evenly distributed in the high- and moderate-
retention plots. However, timber was removed so unevenly in the low-retention plot that the lower
transect resembled the patch cut transects (and similarly exposed to sun and rain) while the upper
transect retained so many trees that it resembled the high-retention stand. Likewise, numerous trees
marked for retention in the control plot were felled by ‘mistake’, exposing the west ends of both transects
to drying and sun. It is now obvious that we ought to have established all transects longitudinally along
an elevation gradient, although the tree retention pattern could not have been predicted.
Equally problematical was blow-down of several large trees above the control plot transects during
the winters of 2002 and 2003. Timber crews had weakened several large trees directly above the upper
control transect by using them to hoist timber from the adjacent patch cut up to the yarding area. As
they toppled during the winter storms, those trees in turn knocked over several others trees directly
onto the upper transect, destroying numerous prolic fungal microsites in the process. Such problems,
only to be expected during usual logging practices, nonetheless severely compromised our species
tallies and ability to analyze the study data accurately.
green PeAk densty mAnAgement FungAl study conclusions
As noted above, our nal statistical analysis of the impact of tree-thinning regimes upon the fungal
community awaits full microscopic (re)evaluation of most target collections. Nonetheless, even with
the problems in obtaining reliable baseline data, we feel condent that our summary graph and tables
above support the observation that over the short term, the fungal community is not threatened by
light to moderate thinning; there is little appreciable difference in the fungal species richness between
the control plot on the one hand and the high- and moderate-retention plots on the other.
It is difcult to rely, however, on the conclusions from the so-called light-retention plot, as the tree
retention varied so greatly between transects. Nonetheless the ‘average’ species tally summarized in
graph and table suggests that reduction of trees to 40/acre stresses ectomycorrhizal fungal fruiting
for the rst ve years after harvest. Only by 2005 had the exposed lower transect begun to generate
ectomycorrhizal fruitbodies.
The inability to monitor the patch cut plot during the height of the 1999 fall season obscures the
full impact of removal of all trees on fungal fruiting. It is nonetheless very clear that there is a strong
negative impact on fungal fruiting for ve years after treatment. Here we suggest that the smaller the
patch, the swifter the harvested area will reestablish itself, given the abundance of species that fruited
within a very small area at the base of the buffer trees even after the rst harvest year. Likewise,
timber workers who leave seedlings in place provide shelter for ectomycorrhizal fungi, as witness the
appearance of Phaeocollybia phaeogaleroides (recorded the rst time in 2004) near the bases of three
5–8 tall hemlocks in the middle of the patch cut.
With respect to the incidental collections, we should note that the comparatively larger number
of vouchers collected from the low-retention and patch cut transects can be attributed in part to
the scarcity of ectomycorrhizal targets on the same transects. As the collection boxes were relatively
empty, we lled them with incidentals instead.
imPortAnce oF the FungAl community density study For the Future
The collections obtained during our seven-year study provide an excellent overview of fungal
species richness in ‘working’ mature Douglas r and hemlock forests in the Oregon coast range. Seven
years of data will provide insights into which common Douglas r associates fruit annually and which
sporadically; they will also indicate which species are and are not more susceptible to a particular
harvesting technique. Using permanent transects to track host-fungus mycorrhizal interactions likewise
can provide valuable information on species-specic associations. As we recorded the location of

8 ... 1998–2005 Green Peak Density Management Fungal Community Study
the less frequent Tsuga heterophylla along the transect, we have been able to demonstrate support
for the hypothesis that both Phaeocollybia and Ramaria are hemlock-dependent genera. Acquisition of
1300 ‘incidental’ collections representing over 260 species of infrequently collected non-mycorrhizal
fungi will provide an excellent opportunity to research the ‘common’ saprophytic fungal community
dependent on humus and woody debris as well as provide a good basis for keys to Galerina and Mycena
species in mature Oregon coast range forests.
The fact that so many specimens representing both mycorrhizal and non-mycorrhizal species were
photographed and processed immediately after sampling makes the project collections a particularly
valuable resource for future taxonomic studies.
2017 uPDate
Although most of the unexamined collections still need microscopical verication, several have
contributed to taxonomic progress since 2008.
Taxonomic revision of Phaeocollybia is ongoing. Sequences sampled by Matt Gordon from the P. phaeo-
galeroides specimens we collected in 2004 and 2005 from the clear cut transects helped establish that
species as representing the most basal phylogenetic clade in the genus and helped draw attention to
at least three other cryptic taxa to which the name phaeogaleroides has been misapplied (Norvell & al.
2010 a, b). Gordon also generated a sequence from a phaeocollybia collected by Ron Exeter in 2009
from the high-retention transect that revealed an undescribed taxon closely related to P. ammiratii.
Matheny & al. (2013) published the new species Inocybe chondroderma Stuntz ex Matheny, Norvell &
Giles accompanied by Norvell’s photographs (p. 437) of two collections (obtained in 2002 and 2004)
from the Green Peak upper and lower high-retention transects illustrating the blue PDAB reaction
that readily distinguishes this surprisingly common PNW inocybe from a horde of look-alikes. The
phylogram based on ITS sequence analyses of specimens from British Columbia, Washington, Oregon,
and Japan (plus several other unrelated European inocybes) incorporates one sequence from an I .
chondroderma specimen we collected from the mid-retention upper transect in 2004. Nine collections
from the Green Peak control, high-retention, and mid-retention transects were cited in this paper.
Of perhaps greater signicance is recent progress made toward a world-wide revision of the genus
Hydnum. Shortly after attending the 2002 International Mycological Congress Finland foray with
Norvell, Slovenian Tine Grebenc requested samples of Hydnum umbilicatum from the PNW-MS fungal
collections; she returned two 2001 Green Peak collections (one made in May from the high-retention
upper transect and the second in November from the mid-retention lower transect). Grebenc (2009)
and several other scientists (Olariaga E al. 2012, Vizzini & al. 2013, Yanaga & al. 2015, Feng & al. 2016)
used ITS sequences from those specimens to represent H. umbilicatum in their worldwide phylogenies
of the genus.
However, in March 2016 Kare Liimatainen (a Hydnum specialist from Finland) contacted Norvell to
inform her that the Green Peak Hydnum ‘umbilicatum’ specimens represented instead a species new
to science. Norvell assumed that the spring collection would represent the new species with the
autumnal collection representing H. umbilicatum. This turned out not to be the case: Hydnum are highly
endemic, and Peck’s eastern North American type is thought unlikely to occur in the Pacic states.
ITS sequence analyses supported one species distributed in Oregon and California. (The specimen
referred to Hydnum ‘oregonense’ in the cover photos has yet to be molecularly conrmed.) The paper
by Niskanen & al. (submitted to Mycologia in 2017 and still in review) recognizes 36 Hydnum species
worldwide, of which 24 are named as species new to science.

2017 summary (Norvell) ... 9
References (updated 31 March 2017)
Published papers and abstracts citing project data
Exeter Ronald L. & Lorelei L. Norvell. 2008. Ramaria of the Pacic Northwestern United States. Abstract for 2007 MSA
poster presentation, in inoculum 59(2): 16.
Exeter Ronald L., Lorelei Norvell, Efren Cázares. 2006. rAmAriA oF the PAciFic northwestern united stAtes. USDI-BLM.
Salem District. 157 pp.
Grebenc T, Martín MP, Kraigher H. 2009. Ribosomal ITS diversity among the European species of the genus Hydnum
(Hydnaceae). AnAles del jArdín botánico de mAdrid 66S1:121–132.
Matheny, P. Brandon; Norvell, Lorelei L.,; Giles, Emily C.ikllk,. A common new species of Inocybe in the Pacic Northwest
with a diagnostic PDAB reaction. mycologiA 105(2): 436–446. doi: 10.3852/12-155
Niskanen, Tuula, Kare Liimatainen, Jorinde Nuytinck, Paul Kirk, Ibai Olariaga Ibarguren, Roberto Garibay Orijel,
Lorelei Norvell, Seppo Huhtinen, Ilkka Kytövuori, Juhani Ruotsalainen, Tuomo Niemelä, Joseph F, Ammirati &
Leho Tedersoo. 2017 [in edit.] Identifying and naming the currently known diversity of the genus Hydnum
(Cantharellales) with an emphasis on North American and European materials. mycologiA.
Norvell, Lorelei L. 2003. Ecology and management of commercially harvested edible chanterelles in western North
America. Abstract. 3rd internAtionAl workshoP on edible mycorrhizAl mushrooms, August 16-22, 2003.
University of Victoria, Victoria, British Columbia.
Norvell, Lorelei L. 2004. Phaeocollybia in western North America 4: Two new species with tibiiform cheilocystidia and
Section Versicolores reconsidered. mycotAxon 90: 241-260.
Norvell, Lorelei L. 2017. Pnw-ms technicAl rePort: Green Peak Density Management Fungal Community Study: 1998–
2005: 2017 updated summary. 28 pp. (this report).
Norvell, Lorelei L. & Ronald L. Exeter. 1999. 1161 -- The Oregon Douglas-r fungal community. xvi internAtionAl
botAnicAl congress: AbstrActs p. 486.
Norvell, Lorelei L. & Ronald L. Exeter. 1999. Oregon Douglas-r fungal communities. Abstract in northwest scientiFic
AssociAtion 1999 AnnuAl meeting bulletin: 45.
Norvell, Lorelei L. & Ronald L. Exeter. 2002a. 547 – The Douglas-r epigeous ectomycorrhizal basidiomycete
community in the western North American Northern Spotted Owl zone. imc7: book oF AbstrActs.
Oslo [www.uio.no/conferences/imc7]. pp. 166-67.
Norvell, Lorelei L. & Ronald L. Exeter. 2002b. The epigeous ectomycorrhizal basidiomycete Douglas-r
fungal community in “peace” and “war”. inoculum 53(3):47.
Norvell, Lorelei L. & Ronald L. Exeter. 2004. Ectomycorrhizal epigeous basidiomycete diversity in Oregon’s coastal
montane Pseudotsuga menziesii forests – preliminary observations. IN Fungi in Forest ecosystems: diversity,
ecology, And systemAtics, Cathy Cripps (ed.). Memoirs of the New York Botanical Garden 89: 159-189.
Norvell, Lorelei L. & Ronald L. Exeter. 2009 (“2008”). Phaeocollybia oF PAciFic northwest north AmericA.
USDI-BLM/OR/WA/GI-08/100-1792, Salem, Oregon 228 p.
Norvell, Lorelei L. & Scott A. Redhead. 2000a. Phaeocollybia in western North America 2:
The vernal P. pleurocystidiata sp. nov. and P. carmanahensis reconsidered. mycologiA 92: 984-991.
Norvell, Lorelei L. & Scott A. Redhead. 2000b. Stropharia albivelata and its basionym Pholiota albivelata.
mycotAxon 76: 315-320.
Norvell, Lorelei L., Ronald L. Exeter, Matthew Gordon, Scott A. Redhead. 2010a. Species concepts in a molecular age:
the Phaeocollybia waltz. Abstract in inoculum 61(4): 66.
Norvell, Lorelei L., Ronald L. Exeter, Matthew Gordon, Scott A. Redhead. 2010b.
Species concepts in a molecular age: the Phaeocollybia waltz. Abstract in imc9: the biology oF Fungi.
Oxford CD given to 1750 IMC9 delegates.
Pilz, David, Lorelei Norvell, Eric Danell & Randy Molina. 2003. ecology And mAnAgement oF commerciAlly hArvested
chAnterelle mushrooms. gen. tech. reP PNW-GTR-576. March 2003. Portland, OR: US Department of Agriculture,
Forest Service, Pacic Northwest Research Station. 89 pp.

10 ... 1998–2005 Green Peak Density Management Fungal Community Study
Ofcial reports on le in USDI-BLM, Salem District Ofce
(unpublished; available on request)
Norvell, Lorelei L. 2000 (June 1) – PNWMS / BLM Cooperative Oregon Douglas-r density management – fungal
community study: 1998-1999 Annual Report. Requisitioned by co-investigator & Mary’s Peak BLM Botanist Ronald L.
Exeter. 33 pp.
Norvell, Lorelei L. 2002 (January 18). Oregon’s Douglas-Fir ectomycorrhizal fungal communities: the Green Peak density
management study. Year 2 (1999-2000) & Year 3 (2000-2001). Requisitioned by co-investigator & Mary’s Peak BLM
Botanist Ronald L. Exeter. 73 pp.
Norvell, Lorelei L. 2002 (September 17). Oregon’s Douglas-Fir ectomycorrhizal fungal communities: the Green Peak
density management study: Year 3 (2001-2002). Requisitioned by co-investigator & Mary’s Peak BLM Botanist Ronald
L. Exeter. 25 pp.
Norvell, Lorelei L. 2004 (September 13). Oregon’s Douglas-Fir ectomycorrhizal fungal communities: the Green Peak
density management study: Year 4 (2002–2003). Requisitioned by co-investigator & Mary’s Peak BLM Botanist
Ronald L. Exeter. 49 pp.
Norvell, Lorelei L. 2007 (January 8). 2003-2004 annual report for USDI-BLM & PNW-MS 1998-2003 cost-share studies:
Oregon Douglas-r fungal communities—Green Peak density management study. Requisitioned by co-investigator
& Mary’s Peak BLM Botanist Ronald L. Exeter. 67 pp. (including 31-page reprint).
Norvell, Lorelei L. 2008 (July 4). Green Peak Density Management Fungal Community Study: 1998–2005 nal summary.
Requisitioned by co-investigator & Mary’s Peak BLM Botanist Ronald L. Exeter.18 pages. + MS Excel collections
database. (Photos are still incomplete due to 2003 and 2009 computer crashes; reclamation is ongoing).
Slide lectures & poster presentations
(all presented by Norvell; listed in reverse chronological order; many now accessible on ResearchGate)
2012. (July 11). Norvell, Lorelei L. Rooting Phaeocollybia: Lessons learned and a second chance. FESIN (North American
Mycoora Project) presentation (Yale), New Haven, CT.
2010 (August 1). Lorelei L. Norvell, Ronald L. Exeter, Matthew Gordon & Scott A. Redhead. Species concepts in a
molecular age: the Phaeocollybia waltz. Poster. International Mycological Congress 9. Edinburgh, Scotland.
2010 (June 28). Lorelei L. Norvell, Ronald L. Exeter, Matthew Gordon & Scott A. Redhead. Species concepts in a molecular
age: the Phaeocollybia waltz. Poster. Mycological Society of America. Lexington, Kentucky.
2008 (April 1). Lorelei L. Norvell: Kenning the seeds of time. 1-hour Power Point presentation. Vancouver Mycological
Society, Vancouver, BC. Canada.
2008 (April 3). Lorelei L. Norvell. To the root of a mushrooming problem: chanterelles, surveys & volunteers. 1-hour
Power Point presentation. South Vancouver Island Mycological Society, Victoria, BC. Canada.
2004 (July 19). Lorelei Norvell. Fungi & the Northwest Forest Plan: lessons learned about epigeous basidiomycetes in
surveyed & managed west slope forests. 20-minute Power Point presentation. University of North Carolina at
Asheville, North Carolina, USA.
2004 (June 12). Lorelei Norvell. Mushrooms! How to nd, know, and love them in their natural habitat or on the table.
1-hour Power Point presentation. Skamania Lodge Wild Mushroom Celebration, Skamania, Washington, USA.
2004 (June 1). Lorelei Norvell & Ron Exeter. Fungal response to thinning at Green Peak. USDI-BLM Density Management
Workshop. 25-minute Power Point presentation. Oregon State University, Corvallis, Oregon, USA.
2003 (August 17). Lorelei Norvell. Ecology and management of commercially harvested edible chanterelles in western
North America. 1-hour Power Point public lecture with poster. 3rd International Workshop on Edible Mycorrhizal
Mushrooms. University of Victoria, Victoria, British Columbia, Canada.
2003 (July 27-29). Lorelei Norvell, David Pilz, Eric Danell, Randy Molina. Ecology and management of commercially
harvested chanterelle mushrooms. Poster. Mycological Society of America, Asilomar Conference Center, Pacic
Grove, California, USA.

2017 summary (Norvell) ... 11
2003 (March 24). Lorelei Norvell. Fungal gems of Oregon’s Pseudotsuga Forests. 1 hour power point presentation. Oregon
Mycological Society. World Forestry Center, Portland, Oregon, USA.
2002 (August 12-14). Lorelei Norvell & Ron Exeter. The Douglas-r epigeous ectomycorrhizal basidiomycete community
in the western North American Northern Spotted Owl zone. Poster. VIIth International Mycological Congress,
Oslo, Norway.
2002 (June 24). Lorelei Norvell & Ron Exeter. Poster. The epigeous ectomycorrhizal basidiomycete Douglas-r fungal
community in “peace” and “war”. Annual Meeting of the Mycological Society of America. Corvallis, Oregon, USA.
2002 (April 10). Lorelei Norvell. The Northwest Forest Plan: Fungi, Herbaria, and the Perception of Rarity. Harvard
Herbaria Seminar Series: Farlow Herbarium, Cambridge, Massachusetts, USA.
2002 (April 8). Lorelei Norvell. Loggers, Litigation, and Lowlifes: Fungi of the Northwest Forest Plan. 2-hour lecture.
Boston Mycological Club, Farlow Herbarium, Cambridge, Massachusetts, USA.
2002 (April 7). Lorelei Norvell. Conversation with Lorelei – mushrooms, names, and projects. 4-hour workshop. Boston
Mycological Club. Farlow Herbarium seminar room: Cambridge, Massachusetts, USA.
2002 (January 28). Lorelei Norvell. Fungal community research in the Oregon forest ecosystem. 2-hour slide lecture, as
part of the “Communities, Ecosystems, and Landscapes in Conservation Theory and Practice” class, Portland State
University, Portland, Oregon, USA.
2002 (January 22). Lorelei Norvell. Field of names: rst you nd it… Los Angeles Mycological Society, Los Angeles County
Natural History Museum, Los Angeles, California, USA.
2000 (August 1). Lorelei Norvell. Practical mycology in the twenty-rst century. 15-minute slide lecture. Mycological
Society of America annual meeting. Burlington, Vermont, USA.
2000 (April 19). Lorelei Norvell. Taking the fungal pulse of America’s forests. 1-hour slide presentation. Fungus Federation
of Santa Cruz. Santa Cruz, California, USA.
2000 (April 18). Lorelei Norvell. Taking the fungal pulse of America’s forests. 1-hour slide presentation. Mycological
Society of San Francisco. San Francisco, California, USA.
1999 (October 23). Lorelei Norvell. So just what are they doing out there? Trophic strategies of Oregon mushrooms.
45-minute slide presentation, Oregon Mycological Society Fall Foray, Camp Cascade (Marion County), Oregon,
USA.
1999 (September 27). Lorelei Norvell. Taking the fungal pulse of America’s forests. 45-minute slide presentation. Oregon
Mycological Society, Portland, Oregon, USA.
1999 (August 1-7). Lorelei L. Norvell and Ronald L. Exeter. The Oregon Douglas-r community. Poster. XVI International
Botanical Congress, St. Louis, Missouri, USA.
1999 (May 7). Lorelei Norvell. The utility of transect sampling of mushrooms in Oregon Douglas-r forests and the Great
Smoky Mountain National Park. 40-minute slide presentation. Pacic Northwest Key Council Spring Workshop,
Rimrock, Washington, USA.
1999 (March 26). Lorelei L. Norvell & Ronald L. Exeter. Oregon Douglas-Fir fungal communities: The rst year. 15-minute
slide presentation: Northwest Scientic Association, Tacoma, Washington, USA.
1999 (February 17). Lorelei Norvell. The track of the mushroom: Fungal biodiversity inventories in Oregon, the Great
Smokies and the Redwoods. 45-minute slide presentation: Humboldt Bay Mycological Society, Arcata, California,
USA.

12 ... 1998–2005 Green Peak Density Management Fungal Community Study
Publications citing Green Peak Fungal Community Study publications
[publication/s cited]
Aguilar-Cruz, Yonatan; Villegas, Margarita. 2010. Espécies de Gomphales comestibiles en el Municipio de Villa del Carbón
Estado de México. revistA mexicAnA de micologíA. 31: 1–8. [Exeter & al. 2006]
Barnhart, Harley. 2004. Book review of Fungi in Forest ecosystems. mushroom, the journAl. 85(2): 24-26. [Norvell & Exeter
2004]
Barnhart, Harley. 2007. Book review of Exeter & al. Ramaria of the Pacic Northwestern United States. mcilvAineA.
[Exeter & al. 2006]
Barroetaveña, C., E. Cázares, M. Rajchenberg. 2006. Mycorrhizal fungi of Pseudotsuga menziesii, an introduced tree species
in Central Patagonia (Argentina). novA hedwigiA 83: 53–66. [Norvell & Exeter 2004]
Barroetaveña, C., E. Cázares & M. Rajchenberg. 2007. Ectomycorrhizal fungi associated with ponderosa pine and Douglas-
r: a comparison of species richness in native western North American forests and Patagonian plantations from
Argentina. mycorrhizA 17: 355–373. [Norvell & Exeter 2004]
Beeren, Christoph von; à l’Allemand, Soa Lizon; Hashim, Rosli; Wittke, Volker. 2014. Collective exploitation of a
temporally unpredictable food source: mushroom harvesting by the ant Euprenolepis procera. AnimAl behAviour
89: 71–77. [Norvell & Exeter 2004]
Beug, M. 2010. Pacic Northwest spring Ramaria species: Subgenera Laeticolora and Ramaria. mcilvAineA 19 (copyright by
PNW Key Council 2010). [Exeter & al. 2006]
Biodiversity Heritage Library. (accessed 31 March 2017): Full online text of Norvell & Exeter’s Phaeocollybia volume.
http://www.biodiversitylibrary.org/item/212150# [Norvell & Exeter 2008]
Cázares E, Guevara G, García J, Estrada A, Trappe JM. 2011. Tres nuevas especies de Ramaria de los bosques de encino del
centro de México. revistA mexicAnA de micologíA 33: 37–42. [Exeter & al. 2006]
Centko RM, Ramon-Garcia S, Taylor T, Patrick BO, Thompson CJ, Miao VP, Andersen RJ. 2012. Ramariolides A–D,
Antimycobacterial butenolids Isolated from the mushroom Ramaria cystidiophora. journAl oF nAturAl Products 75:
2178–2182. [Exeter & al. 2006]
Ceska O. A survey of macrofungi on observatory hill: fall 2009 and winter 2009/2010. e-FlorA bc: electronic AtlAs oF the
PlAnts oF british columbiA. 2010 Mar: 95p. [Exeter & al. 2006, Norvell & Exeter 2004]
Ceska O. 2012. A survey oF mAcroFungi on observAtory hill: sPring 2011 & winter 2011/2012. [Exeter & al. 2006]
Ceska, Oldriska. 2013. technicAl rePort: A survey of macrofungi on Observatory Hill: Spring 2012 and Winter 2012/2013.
96 p. www.researchgate.net/publication/277559543 [Exeter & al. 2006, Matheny & al. 2013, Norvell & Exeter 2004
]
Choi, Jae-Wook; Lee, Eun-Hwa; Eo, Ju-Kyeong; Koo, Chang-Duck; Eom, Ahn-Heum. 2014. Community changes of
ectomycorrhizal fungi by thinning in a forest of Korea. koreAn journAl oF mycology 42(2): 133–137.
http://dx.doi.org/10.4489/KJM.2014.42.2.133 [in Korean] [Norvell & Exeter 2004]
Cissel, John. 2005. Article posted by USDI-BLM as part of ‘thinning treAtment ProPosAl’. [Norvell & Exeter 2004]
Cissel, J.H.; Anderson, P.D.; Olson, Deanna; Puettmann, Klaus; Berryman, Shanti; Chan, Samuel; and Thompson, Charley.
2006, BLM Density Management and Riparian Buffer Study: Establishment Report and Study Plan, U.S. Geological
Survey, scientiFic investigAtions rePort 2006-5087, 151 p. [Exeter & al. 2006; Norvell & Exeter 2002a,b; 2004]
Cline, Erica. 2008. Book Review. inoculum (MSA Newsletter) 59(5): 26. [Exeter & al. 2006]
Coimbra VRM, Gibertoni TB, Wartchow F. Phaeocollybia nigripes (Agaricomycetes), a new species from Brazil. mycotAxon
120: 171–179. http://dx.doi.org/10.5248/120.171 [Norvell & Redhead 2002a]
Cortez VG, Silveira RMB da. 2007. A new species of Stropharia with hymenial acanthocytes. mycologiA 99: 135–138.
[Norvell & Redhead 2002b]
Cortez VG, Silveira RMB da. 2007. Species of Hypholoma (Fr.) P. Kumm. (Strophariaceae, Agaricales) in Rio Grande do Sul
State, Brazil. ActA bot. brAs. 21(3): ?? [Norvell & Redhead 2002b]
Cortez, Vagner G.; Silveira, Rosa Mara B. da. 2008. The agaric genus Stropharia (Strophariaceae, Agaricales) in Rio Grande do
Sul State, Brazil. FungAl diversity 32: 31–57. [Norvell & Redhead 2002b]

2017 summary (Norvell) ... 13
Daniels PP, Ripoll MAR, Christan J. 2012. Notas en Gomphales V: Pimeros registros de Ramaria cokeri RH Petersen, para
Macaronesia y Africa. cryPtogAmie mycologie 33.4: 481-488. [Exeter & al. 2006]
Daniels, PP; Pérez-Gorjón. 2009. Notas en Gomphales IV: Ramaria mediterranea, nuevas citas para la Peninsula Iberica. bol.
soc. micol. mAdrid 33: 75–79. [Exeter & al. 2006]
Desjardin, Dennis E.; Wood, Michael G.; Stevens, Frederick A. 2015. cAliForniA mushrooms: the comPrehensive identiFicAtion
guide. Timber Press, Portland OR. ISBN 978-1-60469-353-9. 560 p. [Exeter & al. 2006]
Esteve-Raventos F, Moreno G. 2014. Inocybe lanatopurpurea Esteve-Rav. & G. Moreno sp. nov. Fungal Planet 273 — 10
June 2014. 288–289 in Crous & al, Fungal Planet description sheets: 214–280. PersooniA 32: 184–306. http://dx.doi.
org/10.3767003158514X682395 [Matheny & al. 2013]
Fan YG, Bau T. 2014. Inocybe hainanensis, a new lilac-stiped species from tropical China. mycosystemA 33(5): 954–960.
http://dx.doi.org/10.13346/j.mycostema.140043 [Matheny & al. 2013]
Feng B, Wang X-H, Ratkowsky D, Gates G, Lee S.S, Grebenc T, Yang Z. L. 2016. Multilocus phylogenetic analyses reveal
unexpected abundant diversity and signicant disjunct distribution pattern of the hedgehog mushrooms (Hydnum
L.). scientiFic rePorts 6:25586. doi:10.1038/srep25586 [see Niskanen & al. 2017]
Gamiet S, Berch S, Roberts C, Winder R, MacKinnon A, Kroeger P. 2003. non-timber Forest Products oF british columbiA:
mushrooms. http://bcmushrooms.forrex.org/ntfp/. [Norvell & Exeter 2002a]
Gamiet, S., S.M. Berch, P. Kroeger, C. Roberts, R. Winder & A. MacKinnon. [July 4, 2008 downloaded]
http://bcmushrooms.forrex.org/ntfp/pages/cantherformo/cantharformo_syn.html [Norvell & Exeter 2002a]
Gates, Genevieve. 2009. coArse woody debris, mAcroFungAl AssemblAges, And sustAinAble Forest mAnAgement in A eucalyPtus
obliqua Forest oF southern tAsmAniA. PHD thesis, University of Tasmania. [Norvell & Exeter 2004]
Gates GM, Mohammed C, Ratkowsky DA, Wardlaw T, Davidson NJ. 2011. Diversity and ecology of epigeous
ectomycorrhizal macrofungal assemblages in a native wet eucalypt forest in Tasmania Australia. FungAl ecology
4(1): 56–67 http://dx.doi.org/10.1016/j.funeco.2010.07.005 [Norvell & Exeter 2004]
Gibson, Ian. Book Review. 2009. BEN (Botanical Electronic News) 414. Posted 23 September 2009:
http://www.ou.edu/cas/botany-micro/ben/ben414.html [Exeter & al. 2006]
Gonzalez-Avila, Patricia; Torres-Miranda, Andrés; Villegas-Rios, Margarita; Luna-Vega, Isolda. 2013. Species diversity and
ecological patterns of Phaeoclavulina species of Mexico with implications for conservation. north AmericAn Fungi 8:
1, February 2013. http://dx.doi.org/10.2509/naf2013.008.016 [Exeter & al. 2006, Norvell & Exeter 2004]
Gonzalez-Avila, Patricia; Villegas-Rios, Margarita; Estrada-Torres, Arturo. 2013. Especies del género Phaeoclavulina en
México. revistA mexicAnA de biodiversidAde: 84 (August supplement): S91–S110. http://dx.doi.org/10.7550/mb.31606
[Exeter & al. 2006, Norvell & Exeter 2004]
Gordon, Matthew; van Norman, Kelli. 2014. Molecular monitoring of protected fungi: mycelium persistence in soil after
timber harvest. FungAl ecology 9: 34–42. [Norvell & Exeter 2004]
Hughes, Karen; Petersen, Ronald H.; Justice, Jay. 2014. Two new species of Ramaria from Arkansas. mycokeys 8: 17–29.
http://dx.doi.org/10.3897/mycokeys.8.7356 [Exeter & al. 2006]
ISSSSP (Huff, ed.). 2009 (April). ISSSSP List Update news. interAgency sPeciAl stAtus And sensitive sPecies ProgrAm uPdAte.
Region 6 Forest Service & BLM Oregon/Washington.2(2): 1. https://www.fs.fed.us/r6/sfpnw/issssp/documents/
updates/update-2009-04.pdf [Norvell & Exeter 2009]
Karstedt F, Oliveira JJS de, Capelari M. A second record of Phaeocollybia megalospora var. tetraspora Singer, 1987 (Agaricales:
Hymenogastraceae) from Brazil. check list 9(6): 1600–1602. [Norvell & Redhead 2002a]
Knudson AG. 2012. The genus RamaRia in minnesotA. Master’s thesis: University of Minnesota. [Exeter & al. 2006]
Kranabetter JM, Friesen J, Gamiet S, Kroeger P. 2005. Ectomycorrhizal mushroom distribution by stand age in western
hemlock – lodgepole pine forests of northwestern British Columbia. cAnAdiAn journAl oF Forest reseArch 35(7):
1527–1539. [Norvell & Exeter 2004]
Kumar, S. Suresh; Jegadeesan, M. 2014. Studies of VAM fungi in and around Mucuna pruriens (L.) DC. — Anti-Parkinson
plant. electronic journAl oF biosciences 2(2): 99–106. http://ejbios.co.in [Norvell & Exeter 2004]
Luo, Hong; Li, Xuan; Li Guozhong; Pan, Yanbo; Zhang, Keqin. 2006. Acanthocytes of Stropharia rugosoannulata function as a
nematode-attacking device. APPlied And environmentAl microbiology. 72: 2982–2987. [Norvell & Redhead 2002b]

14 ... 1998–2005 Green Peak Density Management Fungal Community Study
Majka, Christopher G. 2007. The Ciidae (Cleoptera: tenebrionoidea) of the Maritime Provinces of Canada: new records,
distribution, zoogeography, and observations on bettle-fungi relationships in saproxylic environments. zootAxA
1654: 1–20. [Norvell & Exeter 2004]
Majka CG. 2007. The Erotylidae and Endomychidae (Coleoptera: Cucujoidea) of the Maritime Provinces of Canada: new
records, zoogeography, and observations on beetle-fungi relationships and forest. zootAxA 1546: 39–50.
[Norvell & Exeter 2004]
Majka CG, Langor DW. 2009. Clambidae (Coleoptera) of Atlantic Canada. j. AcAd. entomol. soc. 5: 32–40.
[Norvell & Exeter 2004]
Maneevum A, Dodgson J, Sanoamuang N. 2012. Phaeoclavulina and Ramaria (Gomphaceae, Gomphales) from Nam Nao
National Park, Thailand. TroPicAl nAturAl history 12.2 (2012): 147-164. [Exeter & al. 2006]
Miller Jr., Orson K. & Hope Miller. north AmericAn mushrooms: A Field guide to edible And inedible Fungi. The Globe Pequot
Press: Guilford (Ct). [Norvell & Redhead 2002a]
Mleczko P, Zubek S. 2011. Description of ectomycorrhiza and a new Central European locality of the rare hydnoid species
Sarcodon leucopus (Pers.) Maas Geest. et Nannf. (Thelephorales, Basidiomycota). novA hedwigiA 92: 257-272.
[Norvell & Exeter 2004]
Molina, Randy. 2008. Protecting rare, little known, old-growth forest-associated fungi in the Pacic Northwest USA: A case
study in fungal conservation. mycologicAl reseArch 112: 613-638.
[Exeter & al. 2006, Norvell & Exeter 2004, Norvell & Redhead 2000a,b]
Mycologues amateurs de l’Outaquais. 2007. Book review of Exeter & al. Ramaria of the Pacic Northwestern United
States. lA corne d’AbondAnce (Newsletter of the mycological society in Hull, Québec) 23(1—Mars): 7. (in French).
[Exeter & al. 2006]
Nasim, Ghazala; Ali, Muhammad; Shabbir, Asad. 2008. A study of genus Ramaria from Ayubia National Park, Pakistan.
mycoPAth 6: 43–46. [Exeter & al. 2006]
Oertel, Bernhard; Bandini, Ditte; Vauras, Jukka. 2014. Zwei aus Finnland beschriebene Risspilze in Deutschland
nachgewiesen: Inocybe urceolicystis Stangl & Vauras und Inocybe ericetorum Vauras & Kokkonen. zeitschriFt Für
mykologie 80: 43–79. [Matheny & al. 2013]
Olariaga I, Grebenc T, Salcedo I, Martín MP. 2012. Two new species of Hydnum with ovoid basidiospores: H. ovoideisporum
and H. vesterholtii. mycologiA 104(6):1443–1455. [see Niskanen & al. 2017]
Olson, Deanna H.; Van Norman, Kelli J.; Huff, Robert D. 2007. the utility oF strAtegic surveys For rAre And little-known
sPecies under the northwest Forest PlAn. Gen. Tech. Rep. PNW-GTR-708. Portland, OR: U.S. Department of
Agriculture, Forest Service, Pacic Northwest Research Station. 48 p.
[Norvell & Exeter 2002a,b; 2004; Norvell & Redhead 2002b]
ORNHIC. 2002. Oregon status factors: Phaeocollybia piceae. Oregon Natural Heritage Information Center.
http://inr.oregonstate.edu/sites/inr.oregonstate.edu/les/phaeocollybia_piceae_or.pdf [Norvell & Exeter 2004]
ORNHIC. 2003. California status factors. Oregon National Heritage Information Center.
oregonstate.edu/ornhic/survey/ pholiota_albivelata_ca.pdf [Norvell & Redhead 2000b]
ORNHIC. 2003. Heritage rank status factors. Oregon National Heritage Information Center.
oregonstate.edu/ornhic/survey/ pholiota_albivelata_global.pdf [Norvell & Redhead 2002b]
ORNHIC. 2003. Oregon status factors. Oregon National Heritage Information Center.
oregonstate.edu/ornhic/survey/ pholiota_albivelata_or.pdf [Norvell & Redhead 2002b]
ORNHIC. 2004. Oregon status factors for Phaeocollybia sipei. Oregon Natural Heritage Information Center via
oregonstate.edu/ornhic/ survey/phaeocollybia_sipei_or.pdf [Norvell & Exeter 2004]
ORNHIC. 2004. Washington status factors for Cortinarius cyanites.. Oregon Natural Heritage Information Center via
oregonstate.edu/ornhic/ survey/Cortinarius_cyanites_wa.pdf [Norvell & Exeter 2004]
ORNHIC. 2014. Fact sheet: Craterellus tubaeformis. (prepared by Helen Lau).
https://www.fs.fed.us/r6/sfpnw/issssp/.../sfs-fu-craterellus-tubaeformis-2014-02.docx
[Norvell & Exeter 2004, Pilz & al. 2003]

2017 summary (Norvell) ... 15
ORNHIC. 2014. Fact sheet: Ramaria amyloidea (prepared by Rick Dewey).
https://www.fs.fed.us/r6/sfpnw/issssp/.../sfs-fu-ramaria-amyloidea-2014-01.docx [Exeter & al. 2006]
ORNHIC. 2014. Fact sheet: Ramaria amyloidea (prepared by Jenifer Ferriel).
https://www.fs.fed.us/r6/sfpnw/issssp/.../sfs-fu--rubella-var-blanda-2014-02.docx [Exeter & al. 2006]
Outerbridge R, Trofymow JA, Lalumiere A. 2009. re-estAblishment oF ectomycorrhizAe From reFugiA bordering regenerAting
douglAs-Fir stAnds on vAncouver islAnd. Canadian Forest Service Pacic Forestry Centre information report
BC-X-418. 32 p. [Norvell & Exeter 2004]
Palfner, Götz; Casanova-Katny, M. Angélica; Read, David J. 2005. The mycorrhizal community in a forest chronosequence
of Sitka spruce [Picea sitchensis (Bong.) Carr.] in Northern England. mycorrhizA 15: 571–579.
[Norvell & Exeter 2002b, 2004]
Parladé, J, Martínez-Peña, F, Pera, J. 2017. Effects of forest management and climatic variables on the mycelium dynamics
and sporocarp production of the ectomycorrhizal fungus Boletus edulis. Forest ecology And mAnAgement 390
(2017): 73-79. [Norvell & Exeter 2004]
Petersen, Ronald H. 2007. Book review of Exeter & al. Ramaria of the Pacic Northwestern United States. mycotAxon
102: 439–440. [Exeter & al. 2006]
Ramirez-Lopez I, Villegas-Rios M, Cano-Santana S. 2012. Diversidad de Agaricomycetes clavarioides en la Estación de Biología
de Chamelam, Jalisco, México. revistA mexicAnA de biodiversidAd. http://www.scielo.org.mx/scielo.php?pid=S1870-
34532012000400017&script=sci_arttext&tlng=es [Exeter & al. 2006]
Ríos MV. Diversidad de Agaricomycetes clavarioides en la Estación de Biología de Chamela, Jalisco, México. revistA mexicAnA
de biodiversidAd. 83(4) (in document format posted on ResearchGate). [Exeter & al. 2006]
Scates, Kit & Tina Gospodnetich (1981) Ian Gibson rev. 2003. Trial eld key to the species of Pholiota in the Pacic
Northwest. PAciFic northwest key council. http://www.svims.ca/council/Pholio.rtf [Norvell & Redhead 2002b]
Siegel, Noah; Nguyen, Nhu H.; Vellinga, Else C. 2015. Pholiota olivaceophylla, a forgotten name for a common snowbank
fungus, and notes on Pholiota nubigena. mycotAxon 130: 517–532. [Norvell & Redhead 2002b]
Siegel, Noah & Christian Schwarz. 2016. mushrooms oF the redwood coAst—A comPrehensive guide to the Fungi oF
coAstAl northern cAliForniA. Ten speed press, Berkeley. ISBN 978-1-60774-817-5. 601 p.
[Exeter & al. 2006, Norvell & Exeter 2008, ]
Silva, Paula Santos da. 2008. EsPécies de stRoPhaRiaceae sing. & A.h. sm. (agaRicales, basidiomycota) nA FlorestA nAcionAl de sAo
FrAncisco de PAulA, rio grAnde do su, brAsil. Master’s thesis: Universidade Federal do Rio Grande do Sul. 111 pp.
[Norvell & Redhead 2002b]
Silva PS, Cortez VG, Silveira RMG. 2012. Synopsis of the Strophariaceae (Basidiomycota, Agaricales) from Floresta Nacional
de Sao Francisco de Paula, Río Grande do Sul State, Brazil. hoehneA 39, No. 3. [Norvell & Redhead 2002b]
Stone, Roshanna (District Ranger). 2008 (June). curtis timber sAle Project: environmentAl Assessment. Douglas County,
Oregon. 202 pp. [Exeter & al. 2006]
Thompson, Jennifer B. n.d. Allelopathic alkaloids of an invasive shrub and their effect on the growth of ectomycorrhizal
fungi. University of California, Santa Cruz. 24 p. [Norvell & Exeter 2004]
Trudell, Steve. 2005. Book Reviews. the mycoPhile May/June 2005: 14-15. [Norvell & Exeter 2004]
USDA-FS. 2013. Appendix II Additional Fungus species. 24 pp.
https://www.fs.fed.us/r6/sfpnw/issssp/.../ca-fu-appendix-ii-2013-08-15.docx [Exeter & al. 2006]
USDA-FS Gifford Pinchot Forest. 2009. environmentAl Assessment: Pinchot PArtners restorAtion PlAn (Cowlitz Villey
Ranger District, Lewis County, Washington). 110 p. [p. 3–9.] [Norvell & Exeter 2004]
USDA FS-PNW. 2009 (March) D-Bug hazard reduction timber sale project: drAFt environmentAl imPAct stAtement.
Diamond Lake Ranger District. 394 p. [Exeter & al.2006]
U.S. Fish & Wildlife Service. No date. willAPA nAtionAl wildliFe reFuge comPrehensive conservAtion PlAn. Volume 2. 561
pp. www.fws.gov/willapa/CCP/Willapa NWR CCP Volume 2 reduced [Norvell & Exeter 2004]
Vellinga, Else C. 2007. A mycological Florilegium: Fungal communities. mcilvAineA 17(1): 27–33.
[Norvell & Exeter 2004]

16 ... 1998–2005 Green Peak Density Management Fungal Community Study
Vizzini A, Picillo B, Ercole E, Voyron S, Contu M. 2013. Detecting the variability of Hydnum ovoideisporum
(Agaricomycetes, Cantharellales) on the basis of Italian collections, and H. magnorufescens sp. nov.
mycosPhere 4(1):32–44, doi 10.5943 /mycosphere/4/1/2 [see Niskanen & al. 2017]
Wei TZ, Fu SZ, Qu PP, Yao YJ. 2010. Phaeocollybia purpurea (Cortinariaceae), a new species from Wuyishan, China.
mycotAxon 113: 423–430. doi: 10.5248/113.423 [Norvell & Redhead 2002a]
Wook, Choi Jae; Hwa, Lee Eun; Lee, Joo Kyung. 2014. Clonal variation of extracellular mycorrhizal fungi
according to thinning treatment. mycobiology. 42(2):133-7. [in Korean] [Norvell & Exeter 2004]
Yanaga K, Sotome K, Ushijima S, Maekawa N. 2015. Hydnum species producing whitish basidiomata in Japan.
mycoscience 56:434-442. [see Niskanen & al. 2017]
York, Dana & Richard Helliwell. 2007. A habitat suitability model for six rare Pacic Northwest Fungi using
ecological niche factor analysis.(January – Draft (accessed 3-31-17):
www.fs.fed.us/r6/sfpnw/issssp/documents/inventories/inv-rpt-fu-ectomycorrhizal-ump-hab-suit-model-2007-01-04.pdf
[Exeter & al. 2006, Norvell & Exeter 2004]
Young, A. 2014. Australian coralloid fungi IV — Ramaria licola. muelleriA 33: 13–19. [Exeter & al. 2006]
Young AM. Fechner NA. 2009. Australian coralloid fungi II—a new species of Ramaria (Gomphales) from
Western Australia: Ramaria citrinocuspidata sp. n o v. AustrAlAsiAn mycology 28: 65–67. [Exeter & al. 2006]

2017 summary (Norvell) ... 17
Green Peak Density Management fungal species list:
2017 nomenclatural revisions
(subject to future periodic taxonomic and nomenclatural revisions)
The list of species submitted in 2008 has been revised to reect current taxonomic
concepts; red font denotes recent nomenclatural changes. I have reorganized the
targeted ectomycorrhizal basidiomycete families phylogenetically in accord with
several recent well supported ITS, RPB2, and LSU sequence analyses. Cortinariaceae (as
delimited in previous summaries) has been segregated into Cortinariaceae (Cortinarius) ,
Inocybaceae (Inocybe), and Hymenogastraceae (Hebeloma, Phaeocollybia). Among the
Agaricales, these three segregate families are listed rst below because together
they contain the largest number of species; the remainder are listed alphabetically
according to order and family. Species in the targeted non-ectomycorrhizal fungi and
in fungal ‘incidentals’ are listed in alphabetical, not phylogenetic, order. Species epithets
in bold indicate microscopically veried or otherwise ‘secure’ identications (in that
they adhere to a cohesive species concept). Nonetheless, this list remains preliminary.
u (control) = all trees/acre retained; h = 120 trees/acre retained; m = 80 trees/acre retained
L = 40 trees/acre retained; c (clear or patch cut) = 0 trees/acre retained
Abbreviations before § denote Pre-treAtment (1998–1999) transects
Abbreviations after § denote post-treatment (2000–2005) transects
— indicates no collections obtained pre- or post-treatment
green PeAk density mAnAgement ectomycorrhizAl bAsidiomycete tArget sPecies
Agaricales—Cortinariaceae [~75 species]
Cortinarius acutus chLmu§Cu hLmu
Cortinarius albovariegatus u§L
Cortinarius alboviolaceus —§mh
Cortinarius angelesianus c§hmu
Cortinarius anomalovelatus group [previously cited as ‘anomalus’] —§Lh
Cortinarius armeniacus —§m
Cortinarius balteatus Lm§—
Cortinarius biformis / ligniarius Peck cf§mh
Cortinarius brunneus [early default catch-all] group cLmhu§mhu
Cortinarius brunneus aff –[yellow veil, spring] cmu§—
Cortinarius brunneus var. glandicolor [f curtus] cu§hmu
Cortinarius caesiostramineus /caerulescens var. pallidipes u§mhu
Cortinarius cagei cf. [as bicolor] m§—
Cortinarius californicus cf. —§1u
Cortinarius calochrous var. coniferarum u§mu
Cortinarius camphoratus [cf. also traganus] g41013u1-5
Cortinarius cascadensis U2§H1
Cortinarius casimiri/decipiens/alnetorum/hemitrichus/stemmatus group —§Lmhu
[as brunneofulvus mh§u; as stemmatus cf. —§m]
Cortinarius cedriolens [aka parvannulatus] mhu§Lmu
Cortinarius clandestinus [coniferous; previous hardwood assoc. ‘cotoneus’ goes here] cu§cu hLu

18 ... 1998–2005 Green Peak Density Management Fungal Community Study
Cortinarius croceus group u2 [prev. C . ‘thiersii’ also goes here —§u]
Cortinarius damascenus u§m
Cortinarius diasemospermus [strong rose odor] ?§Lmhu
Cortinarius distans var. olympianus§m
Cortinarius dolabratus (cf. )§cu mhu
Cortinarius erubescens cf. §chu
Cortinarius fasciatus Lhmu§hmu
Cortinarius exipes ss Moser & var. abellus mu§mhu
Cortinarius gentilis cLu§hmu
Cortinarius glaucopus group u§Lmu
Cortinarius illuminus u§cu mhu
Cortinarius imbutus cf§mh
Cortinarius infractiavus [= infractus var. avus] cLmu§mu
Cortinarius juberinus m§—
Cortinarius laetus / ionosmus / duracinus§u
Cortinarius laniger group cLh§cu
Cortinarius kroegeri [as ‘limonius’] cf. g 981111c2-16
Cortinarius malicorius[check also L gentilis] —§1h [x] [refer earlier zakii m§mhu here]
Cortinarius miniatopus c§m
Cortinarius miser ss Moser CF. —§h
Cortinarius mucosus cf. —§u
Cortinarius multiformis group —§u (includes previous ‘allutus’ c§—)
Cortinarius obtusus [common; catch-all] cLmhu§cu Lmhu
Cortinarius olivaceofuscus 1u[x]§ [Check also chrysolitus/idahoensis/ humboldtensis/huronensis/ olivaceopictus/tubarius]
Cortinarius orichalceus —§hmu [in the luteicolor group; check also aureofulvus/ cedretorum/ cupreorufus/ elegantio-
occidentalis/ fulmineus/ vellingae/ xanthodryophilus]
Cortinarius pallidifolius cf. —§mu
Cortinarius papulosus (<—castaneicolor) —§h
Cortinarius smithii [PNW taxon previously referred to ‘phoeniceus va r. occidentalis’] 1h§—
Cortinarius pluvius —§Lh
Cortinarius purpurascens as ‘barlowensis’ —§h
Cortinarius renidens —§Lmhu
Cortinarius saginus [previously cited as ‘cliduchus’] —§m
Cortinarius scandens g 991124h2-8
Cortinarius sec Brunnei —§hu
Cortinarius sec Hydrocybe —§hu
Cortinarius sec Obtusi —§mu
Cortinarius subtortus cf. —§mu
Cortinarius superbus Lh§mh
Cortinarius traganus [the previous ‘var. odoratus’ is not retained; no such variety] cf. —§Lmu
Cortinarius umbilicatus —§Lu
Cortinarius umbrinolens (<— rigidus Moser) ch§hu
Cortinarius venetus L§—
Cortinarius vibratilis cLu§Lu
Cortinarius violaceorubens OR as yet undescr. sp. [previously cited as ‘cyanites’] u§mu
Cortinarius violaceus —§L1
Cortinarius “brunneopunctata” —§mh

2017 summary (Norvell) ... 19
Cortinarius “geranium”—§h
Cortinarius “sweet pea” u§—
Cortinarius “matchstick” u§—
Cortinarius “mystery carrot cort” #1 L§—
Cortinarius “mystery carrot cort” #2 —§u
Cortinarius zundet 1 —§cLmh
Cortinarius zundet 2 —§hu
Cortinarius zundet 3 cLmhu§cLmhu
Cortinarius zundet 4 m§hu
Agaricales—Inocybaceae [~59 species]
Inocybe agglutinata (early catchall) cLmhu§Lmh
Inocybe albodisca [in PNW ≠ grammata; see Matheny] h§mhu
Inocybe amethystina Lmh§mu
Inocybe assimilata [<-curvipes ss LN 2000] chu§mhu
Inocybe bresadolae aff u§—
Inocybe castanea Lh§Lmhu
Inocybe chondroderma [aka: posterula, submuricellata] cmhu§cLmhu
Inocybe cicatricata sensu Stuntz [typ. form under Quercus] —§u
Inocybe cincinnata [cin–ula, phaeocomis [var. major] cLmhu§cLmhu [Index Fungorum lists as syn: I. obscura c§—]
Inocybe cinnamomea h§—
Inocybe cookei cf. [chk also chondroderma, mixtilis, xanthomelas] cmhu§—
Inocybe curvipes [<– decipientoides; IF cites also boltonii, referred to subcarpta by Matheny] h§mhu
Inocybe decemgibbosa [previously cited as oblectabilis var. decemgibbosa; refer glabrodisca here? Matheny 2016)
—?§mh
Inocybe earleana CF. —§L
Inocybe eutheloides cf. 1h§—
Inocybe occulosa (+ var. crocifolia) m+?§cLmhu
Inocybe fuscidula aff u§L
Inocybe fuscodisca [one as ‘fulvodisca’] Lmu§—
Inocybe geophylla! cLmhu§Lmhu
Inocybe griseolilacina Lu§Lmhu
Inocybe griseoscabrosa —§u
Inocybe hirsuta v ar. maxima h§hu [IF refers to I. calamistrata; Matheny 2016 key does NOT]
Inocybe hotsoniana —§m
Inocybe inodora cf. —§m
Inocybe kauffmanii Lhu§Lmhu
Inocybe lacera [the universal shaggy catchall] cLmhu§Lmhu
Inocybe laetior u§—
Inocybe lanatodisca m§—
Inocybe langei / hirtella cf. —§L
Inocybe lanuginosa (I. ovatocystis Atk only) Lhu§c?Lmhu
Inocybe leptocystis cf. h§—
Inocybe leptophylla h§Lmh [IF & Matheny cite as syn casimiri]
Inocybe lilacina [<— geophylla var. lilacina; see Matheny 2016] cLmhu§cLmhu
Inocybe maragaritispora —§u
Inocybe minima cf. [‘fuzztop’] chu§?

20 ... 1998–2005 Green Peak Density Management Fungal Community Study
Inocybe mixtilis cLmhu§Lmhu
Inocybe napipes/cookei ? cLmhu§mhu
Inocybe nitidiuscula cLmhu§Lmhu
Inocybe obscurobadia h§—
Inocybe olympiana u§cu hu
Inocybe phaeoleuca [previously cited as splendens var. phaeoleuca] cf. (check also praecox) —§Lmhu
Inocybe praecox (spring: refer Mar–May ‘queletii’ here!) §u
Inocybe praetervisa m§cLmhu
Inocybe pruinosa L§—
Inocybe pudica [IF lists as syns whitei; Matheny retains pudica] cLmhu§cLmhu
Inocybe retipes / griseovelata —§hm
Inocybe rimosa [<—fastigiata] cLmh§mhu
Inocybe sindonia [not nal] —§hm
Inocybe sororia cLmhu§cLmhu
Inocybe subbrunnea cf. [previously cited as leiocephala/catalaunica; see Matheny key 2016)? —§cLmhu
Inocybe subcarpta sensu Matheny 2016 h+?§cLmhu
[previously cited as boltonii sensu Matheny 2000] Lh§mh
Inocybe subdestricta cLmhu§u
Inocybe subexilis —§mhu
Inocybe subochracea mhu§h
Inocybe umboninota Peck ?§mh
Inocybe xanthomelas cm§h
Inocybe “pyrenocystidiata” u§—
Inocybe zundet c§Lmhu
Inocybe zunk —§m
Agaricales—Hymenogastraceae [~14 species]
Hebeloma crustuliniforme complex [refer to incarnatulum?] cLmhu§cmhu; [refer here also H. longicaudum aff u§Lu ]
Hebeloma mesophaeum —§Lm
Hebeloma perplexum cf. —§u
Hebeloma praeolidum (fragrans also for now) mu§Lu
Hebeloma stenocystis cf. —§mhu
Hebeloma zunk —§u
Phaeocollybia ammiratii —§h
Phaeocollybia attenuata ch§h
Phaeocollybia dissiliens c§—
Phaeocollybia fallax c§h
Phaeocollybia phaeogaleroides —§c
Phaeocollybia pleurocystidiata c§—
Phaeocollybia sipei ch§h
Phaeocollybia spadicea c§h [+ syn. tibiikauffmanii ch§—]
Agaricales—Amanitaceae [~6 species]
Amanita citrina —§mhu
Amanita gemmata [refer to gemmata sp. 01 Amanitaceae.org 28 Mar 17] Lm§Lmhu
Amanita pachycolea —§hu
Amanita smithiana —§u
Amanita vaginata [Refer to Amanita sp-NW04 Amanitaceae.org 28 Mar 17; nsm] —§u
Limacella glioderma [probably unnamed sp.; cf. sp-L-CO01 or L-OR01 on Amanitaceae.org] —§u

2017 summary (Norvell) ... 21
Agaricales—Clavariaceae [1 species]
Clavulinopsis laeticolor cf. —§m
Agaricales—Hydnangiaceae [3 species]
Laccaria amethysteo-occidentalis cLmhu§mhu
Laccaria bicolor —§Lmhu
Laccaria laccata var. pallidifolia cLmhu§Lmhu
Agaricales—Hygrophoraceae [7 species]
Hygrophorus agathosmus m§u
Hygrophorus atramentosus —§u
Hygrophorus bakerensis mh§—
Hygrophorus camarophyllus —§u
Hygrophorus eburneus c-u§c-u
Hygrophorus avodiscus cf. —§u
Hygrophorus odoratus cf. —§h
Agaricales—Tricholomataceae [~10 species]
Tricholoma equestre [previously cited as avovirens] —§u
Tricholoma muricatum [previously cited as pessundatum] —§m
Tricholoma myomyces [PNW name for terreum complex; moseri also refers here] cLmhu§Lmu
Tricholoma pardinum —§Lm
Tricholoma portentosum mh§m
Tricholoma saponaceum —§mh
Tricholoma subsejunctum? [under sejunctum] mh§m
Tricholoma sulphureum cLmhu§cmu
Tricholoma vaccinum —§Lm
Tricholoma zundet u§mh
Boletales—Boletaceae [7 species]
Chalciporus piperatus c§m
Neoboletus luridiformis [previously cited in Boletus; also as B. erythropus] u§u
Phylloporus arenicola [PNW sp., misapplied rhodoxanthus goes here] cf. —§L
Suillus caerulescens u§mhu
Suillus lakei cLmhu§mhu
Xerocomellus diffractus [previously cited as Suillus zelleri] —§u
Xerocomus subtomentosus [previously cited in Boletus] h§h
Boletales—Gomphidiaceae [3 species]
Gomphidius glutinosus (incl. var. salmoneus) cLmhu§cuLmhu
Gomphidius subroseus Lhu§Lmhu
Gomphidius zundet (micro needed) u§mu
Cantharellales—Cantharellaceae [~4 species]
Cantharellus cascadensis cf. [all probably represent cafo] —§cLmh
Cantharellus formosus cLmhu§c(u)Lmhu
Cantharellus subalbidus —§u
Craterellus tubaeformis PNW [‘neotubaeformis’ nom. prov.] mu§Lmhu

22 ... 1998–2005 Green Peak Density Management Fungal Community Study
Cantharellales—Hydnaceae [~2 species]
Hydnum oregonense Norvell, Liimat. & Niskanen sp. nov. in ed. [ITS conrmed] —§mh;
probably most other H. umbilicatum collections (—§mhu) go here.
Hydnum repandum Lh§mhu
Gomphales—Clavariadelphaceae [1 species]
Clavariadelphus caespitosus —§u
Gomphales—Gomphaceae [~12 species]
Gomphus clavatus —§u
Ramaria abietina —§L
Ramaria acrisiccescens —§mu
Ramaria aurantiisiccescens —§c/u m
Ramaria formosa —§m
Ramaria gelatiniaurantia —§L
Ramaria leptoformosa —§c/u mu
Ramaria longispora u§cmhu
Ramaria rasilispora —§u
Ramaria rubella —§L
Ramaria stricta —§L
Ramaria zundet m§m
Russulales—Russulaceae [~47 species]
Lactarius chrysorheus cf. —§h
Lactarius fallax v concolor —§h
Lactarius kauffmanii m§—
Lactarius pseudomucidus —§Lmh
Lactarius rubidus [previously cited as fragilis var. rubidus] cLmu§Lmhu
Lactarius rubrilacteus cLmhu§c/uLmhu
Lactarius subammeus —§u
Lactarius subrufus cf. —§c/u
Lactarius zunk —§u
Russula abietina [tax work needed] chu§Lmhu
Russula adulterina aff. —§u
Russula aeruginea ss Thiers —§mu
Russula alutacea L§—
Russula atroviolacea cf. [?a pine associate] L§h
Russula badia cf. u§—
Russula bicolor m§mhu
Russula brevipes m§—
Russula brunnceoviolacea cf. —§u
Russula cessans ss Thiers —§Lhu
Russula consobrina —§u
Russula dissimulans h§mh
Russula foetens cf. [check also fragrantissima, laurocerasi, pectinata] u§u
Russula gracilis —§h
Russula graveolens —§mu
Russula grisea [re Roberts 2012: check instead R. medullata; grisea associates with beech]—§mh
Russula mariae cf. —§h
Russula murrillii chu§mhu
Russula olivacea m§h

2017 summary (Norvell) ... 23
Russula parazurea L§mu
Russula pectinata [check also cerolens, foetens] c§—
Russula placita —§Lu
Russula pseudo-olivascens cf. [IF cites for elaeodes, nom. dub.] —§u
Russula queletii [previously cited as lilacea] cf. —§mu
Russula raoultii cL§Lmh
Russula risagallina [IF cites for chameoleontina] cf. m§—
Russula robinsonae [‘puellaris’’rhodopoda’’viscida’] ?x§Lmhu
Russula rosea —§u
Russula sanguinea [previously cited also as rosacea ss Woo] Lmhu§m
Russula semirubra —§Lmh
Russula sphagnophila cf. —§h
Russula stuntzii cf. —§u
Russula turci cf. [check – no pine or spruce!!] cm§m
Russula vesca —§h
Russula veternosa cf. tax/nom probs —§hu
Russula xerampelina [w cfs, semi_grav_elaeos] cLmhu§Lmhu
Russula zundet (no spores) cLmhu§L mhu
Russula zunk m§Lmhu
Russulales—Thelephoraceae [~3 species]
Thelephora anthocephala —§mhu
Thelephora palmata [check also anthocephala] L§cLmhu
Thelephora terrestris —§hu
green PeAk density mAnAgement non-ectomycorrhizAl bAsidiomycete tArgets
[~8 species]
Asterophora parasitica —§h
Clavulina cristata h§cLmhu
Infundibulicybe trulliformis [previously cited as Clitocybe senilis, possibly strictly European, with the former Clitocybe
trulliformis (or an unnamed taxon) a more likely determination] Lu§cL mhu
Galerina atkinsoniana [Strategy 3] L§cLmu
Galerina vittiformis f tetraspora [now delisted] Lmhu§cLmhu
Gymnopilus punctifolius c§cu
Lichenomphalia umbellifera < Omphalina ericetorum —§cLm
Stropharia albivelata u§Lhu
green PeAk density mAnAgement incidentAl non-ectomycorrhizAl sPecies
[~250 species]
Agaricus diminutivus (‘semotus’ in PNW refers here; Kerrigan 2017) —§mhu
Agaricus incultorum [previously cited as ‘cupreobrunneus’] —§hu
Agaricus zunk§m
Agrocybe praecox —§Lh
Alloclavaria purpurea [previously cited in Clavaria] —§L
Armillaria nabsnona cf. —§§m
Armillaria ostoyae cf. m§cmu
Armillaria sinapina cf. Lh§Lu
Armillaria zundet —§hu
Arrhenia (as Omphalina) hohensis cf. L§—

24 ... 1998–2005 Green Peak Density Management Fungal Community Study
Arrhenia epichysium —§L
Atheniella adonis [cited previously in Mycena] cm§mhu
Atheniella aurantiidisca [cited previously in Mycena] mu§hu
Atheniella avoalba cf. [previously cited in Mycena] —§u
Auriscalpium vulgare —§u
Baeospora myosura cf. u§Lu
Bisporella citrina —§mu
Bogbodia uda [previously cited in ‘Stropharia’; should have been Hypholoma] cf. —§u
Calocera viscosa L§L
Cheimonophyllum candidissimum m§m
Chrysomphalina aurantiaca Lu§cLu
Ciboria rufofusca —§h
Clitocybe ‘subfragrans’ [??] —§hu
Clitocybe concava ss Bigelow 1985 —§L
Clitocybe fragrans —§cm
Clitocybe pinophila cf§cu
Clitocybe ramigena [99’s as ‘squamulosa va r. montana’] cL§Lmhu
Clitocybe subconcava —§c
Clitocybe subspadicea cf. —§h
Clitocybe zundet h§mhu
Clitocybe zunk —§c
Clitocybe zunk [Lyophyllum?] —§Lm
Clitopilus hobsonii cf. —§h
Coccomyces dentatus (orig as coronatus) cf. —§mhu
Connopus acervatus [previously cited in Gymnopus] —§cLu
Conocybe subovalis cf.—§hu
Conocybe tenera complex cu§cLmhu
Conocybe zundet —§u
Conocybe zunk —§L
Coprinopsis lagopides cf. —§u
Coprinopsis zundet —§u
Coprinus comatus —§u
Crepidotus mollis m§Lu
Crepidotus occidentalis L§m
Crepidotus zunk m§—
Cryptoporus volvatus —§u
Cystoderma amianthinum cm§cLmhu
Cystoderma zundet —§cmh
Cystodermella cinnabarina [previously cited in Cystoderma] [syn. terryi] h§Lm
Cystodermella granulosum [previously cited in Cystoderma] h§L
Cystolepiota [some as Lepiota] oregonensis/ petasiformis—§ch
Cystolepiota seminuda L§Lhu
Dacrymyces cf. zundet —§h
Deconica crobula [cited previously in Psilocybe] —§mhu
Deconica horizontalis [cited previously in Melanotus] Lm§Lm
Deconica inquilina [cited previously in Psilocybe] —§m
Deconica phyllogena (< rhombispora) [cited previously in Psilocybe] —§mh

2017 summary (Norvell) ... 25
Donadinia nigrella [as Plectania nannfeldtii] Lu§—
Entoloma zundet m§cu
Exidia zundet —§u
Flavoscypha cantharellua [cited previously as Otidea concinna] h§Lhu
Fomitopsis cajanderi / rosea —§c
Galerina autumnalis (in marginata complex) u§cLu
Galerina badipes mhu§cLhu
Galerina cedretorum cLmhu§cLmhu
Galerina fallax cL§—
Galerina lignatilis —§L
Galerina marginata L§cu
Galerina perplexa L§Lh
Galerina pubescentipes —§m
Galerina pumila var. subalpina (as cerina) cLm§cLhu
Galerina semilanceata —§cLmhu
Galerina sideroides —§ch
Galerina stylifera m§cLmhu
Galerina subochracea —§h
Galerina unicolor c§—
Galerina vittiformis f vittiformis L§Lh
Galerina zundet —§cLmhu
Gamundia leucophylla —§c
Geastrum saccatum —§Lm
Geopyxis carbonaria —§u
Gloeophyllum sepiarium —§c
Guepiniopsis alpina [all as Heterotextus] —§hu
Gymnopilus bellulus L§hL
Gymnopilus liquiritae —§u
Gymnopilus picreus / penetrans —§Lmu
Gymnopilus sapineus —§m
Gymnopilus zundet —§Lmhu
Gymnopus fuscopurpureus u§hm
Gymnopus perforans cf. [previously cited in Micromphale] u§u
Gymnopus zundet —§c
Gyromitra esculenta —§Lmhu
Gyromitra infula —§mhu
Helvella compressa —§u
Helvella lacunosa h§cLmhu
Helvella maculata L§u
Hemimycena ‘planopedes’ L§—
Hemimycena albicolor —§c
Hemimycena delectabilis u§—
Hemimycena lactea <– Mycena delicatella u§u
Hemimycena zundet m§cLhu
Hygrocybe conica —§cm
Hygrocybe zundet —§u
Hygrophoropsis aurantiaca —§c

26 ... 1998–2005 Green Peak Density Management Fungal Community Study
Hymenoscyphus or Cudoniella —§L
Hypholoma capnoides c§cLmhu
Hypholoma fasciculare —§cLmh
Hypholoma marginatum [previously cited as dispersum; synonymy not noted in IF) —§cLmhu
Hypholoma zundet —§c
Hypogymnia cf. zundet m§—
Icmadophila ericetorum h§—
Jahnoporus hirtus c§h
Kuehneromyces lignicola (<– vernalis) —§mh
Kuehneromyces mutabilis —§Lh
Lentaria pinicola cf. [IF erroneously follows Corner, placing this in Ramaria. See Giachini & al. 2010]—§L
Lepiota atrodisca —§ch
Lepiota castaneidisca —§h
Lepiota cristata [var. pallidior] —§cLhu
Lepiota ammeotincta —§cLmh
Lepiota magnispora —§chu
Lepiota zundet —§cLmu
Lepiota zunk —§L
Leptonia Lyophyllum rancidum? L§—
Leptonia trichomata —§L
Leptonia zundet —§L
Leptoporus mollis —§c
Leucoagaricus [as Lepiota] glabridiscus cf. {some ‘rubrotinctus’ collections go here] h§—
Leucoagaricus rubrotinctus h§cLmh
Lycoperdon perlatum —§u
Lycoperdon umbrinum ch§ch
Lyophyllum / Clitocybe C F. ‘obsoleta subditopoda pinophila’ L§cmhu
Lyophyllum cf. zundet Lh§chu
Lyophyllum cf. zunk —§mh
Lyophyllum cf. zunk sh / Rhodocybe carlottae —§mhu
Lyophyllum decastes —§u
Lyophyllum eustygium [cf. also helvelloides] u§—
Lyophyllum helvella cf. L§—
Macrotyphula juncea —§m
Marasmiellus candidus [previously cited in Tetrapyrgos; also as Marasmiellus albuscorticus] L§u
Marasmioid zunk —§u
Marasmius plicatulus c§—
Melanoleuca cognata cf§u
Melanoleuca zundet g01025m1-15
Mycena acicula cf. —§u
Mycena aurantiomarginata mu§h
Mycena epipterygia [var. lignicola micro but no pix] mh§cLhu
Mycena lopes cf. —§u
Mycena galericulata —§cu
Mycena galopus Lh§—
Mycena haematopus —§c
Mycena leptocephala hu§cLmhu
Mycena maculata —§hu
Mycena metata [w early ‘quinaultensis’ ‘tenax’] cLmu§chu

2017 summary (Norvell) ... 27
Mycena murina Lm§Lmhu
Mycena oregonensis —§Lu
Mycena pura [not referred to Prunulus] cLmhu§cLmhu
Mycena purpureofusca —§cu
Mycena robusta —§Lmh
Mycena rosella —§chu
Mycena rubromarginata L§—
Mycena rutilantiformis [not referred to Prunulus] —§L
Mycena sanguinolenta m§Lmu
Mycena stipata cf. cLmhu§Lu
Mycena tenerrima [previously cited as adscendens] —§h
Mycena zundet u§cLhu
mycenoid zundet —§cm
Mycetinus copelandii [previously cited in Marasmius; ‘scorodonius’ goes here] cL§Lmh
Mycetinus salalis [previously cited in Marasmius] cLh§Lmh
Neolentinus zundet m§—
Nidula niveotomentosa —§cLmh
Nolanea holoconiota (some as hirtipes) —§cLmhu
Nolanea proxima —§h
Nolanea verna cf. [as Entoloma] —§h
Nolanea zundet shy —§u
Nolanea zundet m§cmhu
Nolanea zunk m§—
Onnia tomentosa [previously cited in Inonotus] u§hu
Otidea onotica [pix also cf. ‘concinna’] cLmhu§Lmhu
Otidea tuomikoskii [as leporina] cLm§mu
Otidea zundet Lm§mu
Otidea zundet: cf. alutacea ch§m
Panellus stipticus —§mh
Paralepista inversa [previously cited in Lepista] —§cL
Peniophora ‘incrustans’ [?] cf. —§m
Peniophora incarnata cf. —§m
Pholiota astragalina m§u
Pholiota decorata mu§X
Pholiota ferrugineolutescens cf. —§u
Pholiota lenta —§hmu
Pholiota malicola Lm§cLmh
Pholiota subochracea cf. —§L
Pholiota terrestris —§u
Pholiota zundet [cf. also Hypholoma] —§Lmhu
Plectania melastoma mh§m
Pleurocybella [as Nothopanus] porrigens —§c
Pluteus cervinus —§m
Pluteus cyanopus [IF cites this as synonym of P. chrysophaeus] —§u
Polyporus badius cf. —§u
Polyporus leptocephalus [for elegans see Zhou et al. 2016 (as ‘Cerioporia’ in IF?)] / melanopus; cf. m§L
Postia caesia cf. [for Tyromyces, Oligoporus] —§mh
Postia fragilis [previously cited in Oligoporus] —§h
Psathyrella candolleana —§c

28 ... 1998–2005 Green Peak Density Management Fungal Community Study
Psathyrella brillosa u§—
Psathyrella longistriata —§cmhu
Psathyrella piluliformis [previously cited as hydrophila] cf. —§m
Psathyrella zundet —§cmu
Pseudohydnum gelatinosum —§Lmhu
Psilocybe pelliculosa Lhu§cLmhu
Psilocybe zundet —§cmh
Psilocybe-Tubaria zundet —§u
Resinomycena cf. zundet m§h
Resupinatus / Cheimonophyllum zundet —§m
Rhodocollybia butyracea —§L
Rhodocollybia maculata C F. —§m
Rhodocollybia oregonensis —§m
Rhodocollybia prolixa var. distorta cf. —§L
Rhodocybe caelata —§Lmhu
Rhodocybe trachyospora (as carlottae) —§mu
Rhodocybe zundet —§u
Rhodophana [previously cited in Rhodocybe] nitellina m§cmhu
Roridomyces / Resinomycena zundet —§h
Roridomyces roridus L§hu
Sclerotinia zundet —§m
Scytinotus longinquus [previously cited in Panellus] —§L
Stereum hirsutum —§u
Strobilurus albipilatus Lm§Lhu
Strobilurus trullisatus Lm§mh
Strobilurus zundet —§mh
Stropharia ambigua u§Lu
Stropharia hornemannii —§L
Tapinella atrotomentosa (2004 w Hypomyces) L§L
Tremella encephala —§cmh
Tremella mesenterica —§mh
Tremella zundet —§h
Trichaptum abietinum [as Hirschioporus] —§hu
Tricholomopsis rutilans —§L
Tubaria furfuracea C F. —§Lu
Tubaria minutalis C F. —§u
Tubaria zundet —§cu
Tuber gibbosum h§—
Urnula mexicana [previously cited in Sarcosoma] c§u
Xeromphalina campanella cL§Lmhu
Xeromphalina cirris /cornui cu§hu
Xeromphalina fulvipes u§cmhu
Xeromphalina parvibulbosa / cornui —§u
Xylaria hypoxylon pnw —§u
Xylaria zundet —§L
6 not yet assigned to order: 2000, 2001, 2003, 2004 (2), 2005