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Sciarid flies from Dominican Amber (Diptera, Sciaridae)

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Das Ergebnis der nachfolgenden Untersuchung ist die Neubeschreibung von 20 fossilen Arten aus Dominikanischen Bernstein und einer neuen Gattung mit fossilen und einer rezenten Art aus Afrika. Die Arten können acht Gattungen zugeordnet werden: Trichosia (Subgenus Mouffetina [1]); Leptosciarella (Subgenus Leptosciarella [1]), Cratyna (Subgenus Cratyna [1]), Cratyna unplaziert [2], Archicratyna gen. nov. [4], Chaetosciara [1], Bradysiopsis [1], Bradysia [3], Corynoptera [3] und Epidapus [4]. - Keine Art aus dem Dominikanischen Bernstein ist identisch mit Arten aus dem Baltischen bzw. dem Sächsischen (Bitterfeld) Bernstein, gehören aber zu gemeinsamen Gattungen wie Trichosia, Leptosciarella, Cratyna, Corynoptera and Epidapus. Alle Arten vom Dominikanischen Bernstein gehören zu rezenten Gattungen, während viele Arten vom Baltischen und Sächsischen Bernstein ausgestorbenen Gattungen bzw. Untergattungen repräsentieren. Außerdem ist die Merkmalsausprägung moderner als bei den Arten aus Europäischem Bernstein. Erstaunlich ist weiterhin, dass die fossile Sciaridenfauna aus dem Dominikanischen Bernstein mehr Ähnlichkeiten mit der rezenten Fauna der Holarktischen Region als mit der rezenten Fauna von Mittel- und Südamerika aufweist.StichwörterSciaridae, Dominican Amber.Nomenklatorische Handlungenarcana Mohrig & Röschmann, 2005 (Archicratyna), spec. n.subarcana Mohrig & Röschmann, 2005 (Archicratyna), spec. n.trichoarcana Mohrig & Röschmann, 2005 (Archicratyna), spec. n.zimbabweensis Mohrig, 2005 (Archicratyna), spec. n.antiqua Mohrig & Röschmann, 2005 (Bradysia), spec. n.dimidiata Mohrig & Röschmann, 2005 (Bradysia), spec. n.exoleta Mohrig & Röschmann, 2005 (Bradysia), spec. n.repentina Mohrig & Röschmann, 2005 (Bradysiopsis), spec. n.obsoleta Mohrig & Röschmann, 2005 (Chaetosciara), spec. n.iocosa Mohrig & Röschmann, 2005 (Corynoptera), spec. n.prisca Mohrig & Röschmann, 2005 (Corynoptera), spec. n.pristina Röschmann & Mohrig, 2005 (Corynoptera), spec. n.interposita Mohrig & Röschmann, 2005 (Cratyna), spec. n.villosoantennata Mohrig & Röschmann, 2005 (Cratyna), spec. n.tempestiva Mohrig & Röschmann, 2005 (Cratyna (Cratyna)), spec. n.adstrictosetus Mohrig & Röschmann, 2005 (Epidapus), spec. n.longisetus Mohrig & Röschmann, 2005 (Epidapus), spec. n.macrospinatus Mohrig & Röschmann, 2005 (Epidapus), spec. n.microspinus Mohrig & Röschmann, 2005 (Epidapus), spec. n.manifesta Mohrig & Röschmann, 2005 (Leptosciarella (Leptosciarella)), spec. n.Archicratyna Mohrig, 2005 (Sciaridae), gen. n.nova Mohrig & Röschmann, 2005 (Trichosia (Moufettina)), spec. n.
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
55
  
55  
Sciarid flies from Dominican Amber
(Diptera, Sciaridae)


Summary
 
Trichosia
MouffetinaLeptosciarellaLeptosciarellaCratynaCratynaCratyna
ArchicratynaChaetosciaraBradysiopsisBradysiaCorynoptera
Epidapus
   
           Trichosia, Leptosciarella,
Cratyna, CorynopteraEpidapus

   


Zusammenfassung

   
     Trichosia  Mouffetina  Leptosciarella 
Leptosciarella  Cratyna  Cratyna  Cratyna   Archicratyna   
ChaetosciaraBradysiopsisBradysiaCorynopteraEpidapus
              
  Trichosia, Leptosciarella,
Cratyna, Corynoptera  Epidapus        

     



Key words

DOI: 10.21248/contrib.entomol.55.2.319-361
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 























       
   Trichosia,
Leptosciarella, Cratyna, Epidapus, Corynoptera  Lycoriella   Succinosciara






    


CorynopteraTrichosia/LeptosciarellaLycoriella
EpidapusCratyna

   Hymenaea      
       


   

DOI: 10.21248/contrib.entomol.55.2.319-361
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55






               


 

      














 
    


             
       

  



Order Insecta
Family Sciaridae BILLBERG, 1820
Genus Trichosia WINNERTZ, 1867
Type species:
Trichosia splendens
DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Trichosia 

Trichosia
meunieriTrichosia
Palaeotrichosia
Archaeosciara
  Mouffetina     
T. filispina

Subgenus Mouffetina FREY, 1942
Type species:
Plastosciara pulchricornis
Trichosia (Mouffetina) nova sp. n. †

Description:








Thorax:



   












DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Abdomen:
  




Material:










 




Remarks:
TrichosiaMouffetina


TrichosiaMpulchricor-
nisT.Mfilispina



Genus Leptosciarella TUOMIKOSKI, 1960
Type species:
Sciara elegans    Sciara scutellata  

Leptosciarella
Leptosciarella




Protosciarella 
DOI: 10.21248/contrib.entomol.55.2.319-361
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 
   Hirtipennia  Trichosiopsis
Leptospina 


 Leptosciarella 



Protosciarella
Leptosciarella

Subgenus Leptosciarella TUOMIKOSKI, 1960 s. str.
Type species:
Sciara elegansSciara scutellata

Leptosciarella (Leptosciarella) manifesta sp. n. †

Description:
Head.


   
  



Thorax:












DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Abdomen:
   




Material:






Remarks:
Leptosciarella



Leptosciarella nudivenosa

Genus Cratyna WINNERTZ, 1867
Type species:
Cratyna atra
Cratyna atraPseudosciara pictiventris
 Plastosciara        
Cratyna
 Plastosciara  Termitosciara
PeyerimhoffiaSpathobdella
Spathobdella
C. nobilis
Cratyna archaica

      Cratyna        
C. atra PeyerimhoffiaC. vagabunda

Cratyna
  



Cratyna
DOI: 10.21248/contrib.entomol.55.2.319-361
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 


            
Plastosciara keilbachi
Subgenus Cratyna WINNERTZ s. str.
Type species:
Cratyna atra
Cratyna (Cratyna) tempestiva sp. n. †

Description:
Head.






Thorax:
     












Abdomen:







DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Material:

 

 



Remarks:
Cratyna

 
Cr. subalpinaCr. curaosi
Cr. fumoalata      



Unplaced species of Cratyna
Cratyna villosoantennata sp. n. †

Description:
Head.     
       




Thorax:





 
 

   



DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Abdomen:
   
   

 

Material:

 

Remarks:



 Cratyna

Cr. interposita
Culiginosa

Cratyna interposita sp. n. †

Description:
Head.
      
 
  


Thorax:






         
      




DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Abdomen:




Material:


 



Remarks:



 Cratyna
Genus Archicratyna MOHRIG gen. n.
Type species:
Archicratyna zimbabweensis
Diagnosis:
Head.
    

     
     

       






Remarks:


Cratyna
DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Archicratyna zimbabweensis MOHRIG sp. n.

Description:
Head.




Thorax:
 

              


 



Abdomen:
    




Material:



Remarks:
 





DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Archicratyna arcana sp. n. †

Description:
Head.
    
   

 

Thorax:

       





  





Abdomen:





Material:






Remarks:




DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Archicratyna trichoarcana

Archicratyna trichoarcana sp. n. †

Description:
Head.  
          




Thorax:

  
       
             









Abdomen:





Material:



Remarks:
Archicratyna arcanaArchicratynasubar-
cana
DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Archicratyna subarcana MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.

    



Thorax:
   










Abdomen:

  

   

Material:




Remarks:
Archicratyna arcana

Chaetosciara FREY, 1942

Type species:
Sciara estlandica
DOI: 10.21248/contrib.entomol.55.2.319-361
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 




Chaetosciara gilva 



Chaetosciara obsoleta MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.


   




Thorax:



  









Abdomen:
   
   



Material:

DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
 



Remarks:
Chaetosciara


Chaetosciara gilva
Ch. estlandica

Genus Bradysiopsis TUOMIKOSKI, 1960

Type species:Lycoriella (Bradysiopsis) leucotricha      

      
 

 
     

Bradysiopsis repentina MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.






Thorax:

      
          



DOI: 10.21248/contrib.entomol.55.2.319-361
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 



      


 


Abdomen:





Material:



 

Remarks:
Bradysiopsis




 
Bradysiopsis vittata  

Genus Bradysia WINNERTZ, 1867

Type species:
Bradysia angustipennis
Bradysia
      



Bradysia

DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Bradysia.



Bradysia antiqua MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.     
       




Thorax:
     












Abdomen:
   





Material:

 

 

DOI: 10.21248/contrib.entomol.55.2.319-361
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 

  

Remarks:
Bradysia




Bradysia exoleta MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.

    


Thorax:
    









Abdomen:






Material:



DOI: 10.21248/contrib.entomol.55.2.319-361
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
55





Remarks:
Bradysia


Bradysia


Bradysia dimidiata MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.     
       




Thorax:


          










Abdomen:

         


DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Material:



Remarks:
Bradysia 



Bradysia
Genus Corynoptera WINNERTZ, 1867

Type species:
Corynoptera perpusilla
CorynopteraBradysia
Bradysia, 
       


      Corynoptera       


  Corynoptera,  Trichosia,
LeptosciarellaEpidapus

Corynoptera





  





Corynoptera subtilis
DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Corynoptera pristina RÖSCHMANN & MOHRIG n. sp. †

Description:
Head.     
      

    

  

Thorax:
   











Abdomen:
  



Material:

 

Remarks:
Corynoptera


   
C. perpusillaC.sphenoptera
       


DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Corynoptera prisca MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.     
       




Thorax:



  
        

 




Abdomen:





Material:

 


 
            








DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Remarks:
Corynoptera

C. pristinasubtilis
            


Corynoptera iocosa MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.
      




Thorax:
 

            








Abdomen:




              

Material:

         


DOI: 10.21248/contrib.entomol.55.2.319-361
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 



Remarks:
Corynoptera


C. blanda

Genus Epidapus Haliday, 1851

Type species: Chionea venaticaTipula atomaria
E. atomarius
E. gracilisE. microthorax

 Epidapus




   Epidapus            



Epidapus





Epidapus
Epidapus microspinus MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.     
       
             
 

DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Thorax:












Abdomen:





Material:





Remarks:
 Epidapus 
Epidapus

Epidapus
Epidapus macrospinatus MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.


Thorax:


DOI: 10.21248/contrib.entomol.55.2.319-361
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 





Abdomen:

          



Material:




Remarks:
  Epidapus
Epidapus

Epidapus adstrictosetus MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.
       
 

Thorax:





 

 

  


DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Abdomen:





Material:



 
E. longisetus
Remarks:


Pseudoaptanogyna
E. ignava

Epidapus longisetus MOHRIG & RÖSCHMANN n. sp. †

Description:
Head.
       
    

Thorax:












DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Abdomen:

          



Material:


E. adstrictosetus


Cpristina
Remarks:
EpidapusEmi-
crothorax

 


  
        



   Trichosia,
Leptosciarella, Cratyna, Corynoptera Epidapus
Chaetosciara, Archicratyna, Bradysiopsis
Bradysia












DOI: 10.21248/contrib.entomol.55.2.319-361
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
55














Trichosia,
Leptosciarella, Corynoptera

Pseudosciara, Rhynchosciara, OdontosciaraZygomma, Pseudolycoriella
Mouffettina
Leptosciarella

Chaetosciara
Bradysiopsis 
Corynoptera
Bradysia

CratynatempestivaArchicratyna
 Cratyna       
Cratyna tempestiva

 Archicratyna
       
             
      






   



      
DOI: 10.21248/contrib.entomol.55.2.319-361
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 


       

    

        


       
     












Literature



22





 62


  2
2


5 

6

28 
DOI: 10.21248/contrib.entomol.55.2.319-361
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55
Sciara
SchwenckfeldinaAerumnosa  .Cratyna,PhytosciaraChaetosciara 6

            
Scythropochroa Cratyna  Pictosciara   Pseudozygomma   Epidapus
Macrotarsus Pnyxia CorynopteraKeilbachiaScatopsciara undPelliciplanta


7

41 



71 
    
Epidapus


          
44

1 

42 
            

69 

10 

36
12 

  
86
            
21 
Author’ s address:





DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Fig. 1:Trichosia Mouffetina novaa)b)c)
d)e)
Fig. 2:LeptosciarellaLeptosciarellamanifesta a)b)c)
d) e)f)
DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Fig. 3:CratynaCratynatempestivaa)b)
c)d)e)f)g)h)
DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Fig. 4:Cratyna villosoantennataa)b)c)d)

Fig. 5:Cratyna interpositaa)b)c)d)
e) 
DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Fig. 6:Archicratyna zimbabwae
a) b)
c)d)
e)
Fig. 7:Archicratyna ar-
cana a)
   b)
 c)
   
  
d)e)
f )
g)

DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Fig. 8:Archicratyna trichoarcana
 a)  
b)
 c) 
d)
   
   e)  f)  

Fig. 9:Archicratyna subarcana
a)b)
c)d)
e)

DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Fig. 10:Chaetosciara obsoletaa)b)c)
d)e)
DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Fig. 11:Bradysiopsis repentinaa)b)c)d)
e)
Fig. 12:Bradysia antiquaa)b)c)
d)e)
DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Fig. 13:Bradysia exoleta a)b)c)
d)
Fig. 14:Bradysia dimidiata a)b)c)d)
e)
DOI: 10.21248/contrib.entomol.55.2.319-361
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 
Fig. 18:Epidapus microspinusa)b)c)d)
e)f)
Fig. 15:Corynoptera pristina a)b)c)d)
e)f)
Fig. 16:Corynoptera priscaa)
b)c)

Fig. 17:Corynoptera iocosa
a)b)
c)
DOI: 10.21248/contrib.entomol.55.2.319-361
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
55
Fig. 20:Epidapus adstrictosetusa) 
b)c)
d)
Fig. 19:Epidapus macrospinatusa)b)
c)d)
Fig. 21:Epidapus longisetusa)
b)c)d)
e)
DOI: 10.21248/contrib.entomol.55.2.319-361
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... It is estimated that at least one-half of these species names are in need of verification by type study. Taxonomic Hippa et al. 1998;Menzel 2017;Menzel & Smith 2007Menzel et al. 2013;Mohrig & Röschmann 2005;Rudzinski 1997aRudzinski -e, 1999Rudzinski , 2000Rudzinski , 2003Rudzinski & Baumjohann 2012;Steffan 1967;. Afrotropical sciarid material examined and referred to in publications in the years 1838-2013 originates from Cameroon, Côte d'Ivoire, Democratic Republic of Congo, Equatorial Guinea, Ethiopia, Ghana, Guinea, Kenya, Madagascar, Seychelles, Sierra Leone, South Africa, Tanzania, Togo, Tristan da Cunha Is., Uganda, United Arab Emirates and Zimbabwe. ...
... Archicratyna Mohrig. A genus of four species, three of which were described from Dominican amber (Miocene) (Mohrig & Röschmann 2005). The extant type species, A. zimbabween sis Mohrig, 2005, was described from Matopus National Park, Zimbabwe. ...
... A genus of four species, three of which were described from Dominican amber (Miocene) (Mohrig & Röschmann 2005). The extant type species, A. zimbabween sis Mohrig, 2005, was described from Matopus National Park, Zimbabwe. The biology and immature stages remain unknown. ...
Chapter
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Considering taxonomic decisions and other results (published descriptions, synonymies and records) into account, there are currently 101 species of Sciaridae known to occur in the Afrotropics. Of these, 78 species in 23 genera remain to be properly classified and the remaining 23 species have not yet been studied and are therefore, regarded as Sciaridae incertae sedis for the time being. Kirk-Spriggs & Stuckenberg (2009) estimated the Afrotropical fauna to contain 2,000+ species of Sciaridae.
... At present, 41 fossil species from European ambers (Baltic amber and Bitterfeld or Saxonian amber, perhaps of the same age) have been identified (Mohrig & Röschmann 1994a,b, Röschmann & Mohrig 1995a). Recent studies of Dominican amber, which is considerably younger than the Baltic amber and of Miocene origin with estimated age between 15 and 20 million years (IturraldeVinnent 2001), detected 20 new fossil species (Mohrig & Röschmann 2005 ). All the Dominican amber species possess more modern features than those from the European ambers and show more similarities to the recent fauna of the Holarctic region than to the recent fauna of Central or South America. ...
... For microscopic observation the amber piece was cut down to a thickness of about 3 mm, polished with fine leather . The specimen was studied and drawn under cover slips on a slide in a drop of paraffin oil (Mohrig & Röschmann 2005). For conservation the piece of amber was embedded in Canada balsam on a microscope slide. ...
... The sciarid amber fauna from the Dominican Republic shows a dominance of species with connections to the recent Holarctic fauna (Mohrig et al. 2004 ). None of the species found in Dominican amber represented a genus typical for the recent fauna of Central or South America (Mohrig & Röschmann 2005). Like the species from Dominican amber, the new fossil species from Mexican amber shows in all details modern morphological structures, too. ...
Article
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Schwenckfeldina archoica sp. nov. is the first sciarid fly described from Mexican amber, approximately 20 million years old. The species shows in all details modern morphological structures and is very closely related to recent representatives of the Schwenckfeldina forcipigera species group, presently known from at least ten species distributed in Central America.
... Using the 161 morphological characters proposed for cladistic analysis by Menzel and Mohrig (2000), the fossil material (Röschmann and Mohrig, 1995) has been referred to three groups (Leptosciarella + Trichosia) + ((Cratyna + Epidapus) + (Lycoriella + Corynoptera)) (Menzel and Mohrig, 2000). Interestingly, the generic relationships of the Baltic amber fossils [35-50 MYA, Eocene] were identical to our molecular phylogenetic results, except for the Megalosphyinae, of which fossil species have not been discovered in Europe (Mohrig and Röschmann, 2005). Although the genus Bradysia is presently the most species-rich group, only three species have been found in Dominican amber [20-23 MYA, Oligocene] (Mohrig and Röschmann, 2005). ...
... Interestingly, the generic relationships of the Baltic amber fossils [35-50 MYA, Eocene] were identical to our molecular phylogenetic results, except for the Megalosphyinae, of which fossil species have not been discovered in Europe (Mohrig and Röschmann, 2005). Although the genus Bradysia is presently the most species-rich group, only three species have been found in Dominican amber [20-23 MYA, Oligocene] (Mohrig and Röschmann, 2005). The strict comb-like row of bristles on the apex of the fore tibia (synapomorphy of Megalosphyinae) might have appeared in subfamily Megalosphyinae after divergence from the Pseudolycoriella group (Mohrig and Röschmann, 2005). ...
... Although the genus Bradysia is presently the most species-rich group, only three species have been found in Dominican amber [20-23 MYA, Oligocene] (Mohrig and Röschmann, 2005). The strict comb-like row of bristles on the apex of the fore tibia (synapomorphy of Megalosphyinae) might have appeared in subfamily Megalosphyinae after divergence from the Pseudolycoriella group (Mohrig and Röschmann, 2005). ...
... Sciaridae is composed of more than 2,000 species worldwide (Menzel & Mohrig, 2000 ) and more than 600 species in Europe (Heller & Menzel, 2010). The bodies and wing membranes of adult sciarids have a distinctly dark colouration (Mohrig & Menzel, 2009 ). The wings are fragile , with a markedly " Y " shaped vein in each wing and the antennae are distinctly long and segmented (Cloyd & Eur. ...
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Herbivory on liverworts is rarely reported. We studied the effects of feeding by larvae of the sciarid fly Scatopsciara cunicularius on the growth of the thalloid liverwort Marchantia polymorpha at two different constant temperatures, 12°C and 22°C. Larvae reared at the lower temperature fed slower and over a longer period of time, which resulted in more damage and a greater reduction in the growth of the liverwort than that caused by those reared at the higher temperature. The reduction in growth of the liverwort was positively density-dependent in terms of number of larvae at both temperatures. These results indicate that the larvae of S. cunicularius are likely to be an effective means of controlling M. polymorpha, which is a common weed in plant nurseries and greenhouse cultures.
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The genus Bradysiopsis Tuomikoski is new for the sciarid fauna of North America. It includes 6 species. Four of them are new for science. These are Bradysiopsis postvittigera sp. n., Br. praevittata sp. n., Br. praevittigera sp. n. and Br. Subvittigera sp. n. Two species, Br. vittata (Meigen, 1830) and Br. vittigera (Zetterstedt, 1851), are distributed in the Holarctic but new for North America. In this paper species are described, illustrated by figures and a key for classification as well as distribution data are given.
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This revision covers 40 species of the genera Cratyna Winnertz, 1867 [4 species], Euricrium Enderlein, 1911 [8 species], Metangela Rübsaamen, 1894 [4 species], Pseudosciara Schiner, 1866 [21 species] and Sciara Meigen, 1803 [3 species]. The species are redescribed and illustrated. Sixty-three species are newly combined and four names identified as synonyms: Plastosciara barretoi Lane, 1960 is a new synonym of Cratyna boracensis (Lane, 1960); Plastosciara downsi Lane, 1960 is a new synonym of Cratyna cucarisi (Lane, 1960); Euricrium fulgescens Enderlein is a new synonym of Euricrium ruebsaameni (Enderlein); Pseudosc. aitkeni Lane, 1959 is a new synonym of Pseudosc. forceps (Pettey, 1918). In addition, 3 names are raised from synonymy [Metangela spinata Lane, 1955 restit., Pseudosc. ribeiroi (Lane, 1955) restit., Trichomegalosphys Enderlein, 1911 restit.], and 2 preoccupied names were replaced: Pseudos. willistoni Menzel nom. nov. for Sciara concinna Williston, 1896 [preocc.] and Pseudos. helleri Menzel nom. nov. for Sciara debilis Williston, 1896 [preocc.]. It was further found necessary to designate a lectotype for Pseudosc. willistoni Menzel nom. nov. and neotypes for Pseudosc. muricata (Lengersdorf, 1926), Pseudosciara thoracica (Lengersdorf, 1930) and Metangela bicolorata (Lengersdorf, 1930). Unknown males of two species are described and illustrated [Euricrium varians (Lane), Pseudosc. humeropicta (Lengersdorf)]. The genus group names Zygomma Enderlein, 1911 [preocc], Mapiria Edwards, 1934 and Muhabbetiola Koçak, 2009 are junior synonyms of Euricrium Enderlein, 1911. The genus Pseudosciara Schiner was split into two subgenera Pseudosciara s. str. and Pseudosciarella subgen. nov. After this revision the genera Apelmocreagris Enderlein, 1911, Trichomegalosphys Enderlein, 1911 and Trichosia Winnertz, 1867 are not represented within the Neotropical region and must be deleted from the catalog of Neotropical sciarids. The genus Pseudozygoneura Steffan, 1969 is also a synonym of Cratyna Winnertz. The 41 described species of Pseudozygoneura revised by Hippa et al. (1998) are therefore new combined in the subgenus Cratyna s. str. [in German]
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Corynoptera Winnertz, s. str., is defined. In the Holarctic region it includes 123 species. The species are diagnosed and keyed, and distribution data are given. The following 50 new species are described: Corynoptera adustula (Nepal), C. aequispina (Canada: Quebec), C. alneti (Europe, Canada: British Columbia), C. andalusica (Spain, Greece), C. angustior (W Europe), C. anodon (Japan), C. badia (Italy), C. caesula (Spain), C. collicola (Japan), C. condyloma (Japan), C. controversa (Russia: Primorsk region), C. curvapex (Japan), C. decepta (Russia: Primorsk region), C. digemina (Japan), C. dioon(Nepal), C. distenta (Japan), C. exerta (Russia: Primorsk region), C. fimbriata (Japan), C. flava (Germany, Sweden), C. gemellata (Czech Republic), C. iberica (Spain), C. inclinata (Morocco), C. latibula (Spain), C. lobata (Japan), C. micula(Japan), C. minax (Japan), C. ninae (Finland), C. pacifica (USA: California), C. paracantha (Japan), C. patula (W Europe, Sri Lanka), C. phili (Canada: Ontario, Quebec), C. plusiochaeta (Finland, Sweden, USA: Alaska), C. primoriensis (Russia: Primorsk region), C. redunca (Canada: Quebec), C. romana (Italy), C. serotina (Spain), C. sinedens (Japan), C. sphaerula (Japan), C. spiciceps (Morocco), C. spicigera (Nepal), C. stellaris (Austria, Germany), C. subclinochaeta (Germany, Greece, Russia: Krasnodar region), C. tarda (Russia: Altay region), C. trichistylis (Canada: British Columbia), C. truncatula (Spain), C. tumidula (Finland, Sweden), C. umbrata (Greece, Spain), C. uncinula (Canada, Russia: Yamal Peninsula), C. undulosa (Italy, Portugal, Spain), and C. vulcani (Spain: Canary Islands). The status of C. consumpta (Freeman, 1987), C. defecta (Frey, 1948), C. dubitata Tuomikoski, 1960 and C. arboris Fritz, 1982, is restored as valid species, whereas the following species are regarded as junior synonyms: C. bisulca Mohrig & Mamaev, 1987 = C. bipartita Mohrig & Krivosheina, 1985, C. fritzi Mohrig & Rulik, 2001 = C. triacantha Tuomikoski, 1960, C. praepiniphila Mohrig & Dimitrova, 1992= C. alticola (Kieffer, 1919), C. simonae Rudzinski, 1992 = C. macricula Mohrig & Krivosheina, 1986 and C. spungisi Mohrig & Krivosheina, 1985 = C. irmgardis (Lengersdorf, 1930). Lectotypes are designated for C. dubitata Tuomikoski, 1960, C. gymnops Tuomikoski, 1960, C. inexspectata Tuomikoski, 1960, C. levis Tuomikoski, 1960, C. saccataTuomikoski, C. sphenoptera Tuomikoski, 1960, C. tetrachaeta Tuomikoski, 1960, and Bradysia pachycerca Frey, 1948.
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Examination of 278 pieces of fossil resins from the lower Miocene Saxonian amber (opencast mining Goitsche/Bitterfeld/Germany) yielded to a total of 60 Sciarids in 29 pieces of amber. They belong to 9 species from 3 genera. The speciesEpidapus Primarius, Trichosia (Palaeotrichosia) voelsgeni andTrichosia (Palaeotrichosia) kedingi are new. The other species are known also from Baltic amber.
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Based on ecofaunistical indices (dominance, similarity indices:Jaccard-, Sørensen-, Renkonen-Index) Sciaridae and Ceratopogonidae of Baltic amber and Saxonian amber from Bitterfeld are compared. Despite of the age difference of the amber-bearing deposits of 13–18 Mio. years the included sciarids have a species identity of 34.2% (Jaccard-Index) and a dominance identity of 49.5% (Renkonen-Index). Because of the less carefully studied material from Saxonian amber the ceratopogonids have a species identity of only 9.5 % (Jaccard-Index) (Szadziewski 1993); remarkable is the dominance identity of 37 % (Renkonen-Index). Compared with the data from recent faunas the high degree of identity is the result of a pronounced similarity of the nematoceran faunas of Baltic and Saxonian amber. The ecofaunistic comparison of the sciarids and ceratopogonids provides evidence for two geographically separated amber producing forests. They are more closely related with respect to their age than indicated by the deposits.
Geologia de la region amberifera oriental de la Republica Dominicana
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Die Trauermücken (Diptera: Sciaridae) von Papua-Neuguinea. Teil I -Gattung Sciara, Schwenckfeldina, Aerumnosa gen. nov., Cratyna, Phytosciara und Chaetosciara
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