New floristic records in the Balkans: 28* Reports 1–8

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367
PHYTOLOGIA BALCANICA 21 (3): 367 – 399, Sofia, 2015
New floristic records in the Balkans: 28*
Compiled by Vladimir Vladimirov
1
, Feruzan Dane
2
& Kit Tan
3
1
Department of Plant and Fungal Diversity and Resources, Institute of Biodiversity and
Ecosystem Research, Bulgarian Academy of Sciences, Acad. Georgi Bonchev St., bl. 23,
1113 Sofia, Bulgaria, e-mail: vladimir_dv@abv.bg
2
Department of Biology,
Faculty of Science and Arts, University of Trakya, 22030
Edirne, Turkey, e-mail: feruzandane@gmail.com
3
Institute of Biology, University of Copenhagen, Øster Farimagsgade 2D, DK-1353
Copenhagen K, Denmark, e-mail: kitt@bio.ku.dk
Abstract: New chorological data are presented for 239 species and subspecies from Bulgaria (1-8, 93-101, 113-115,
117-121, 127-136), Greece (9, 10, 16-91, 102-112, 116, 122-126, 137-239) and Turkey-in-Europe (11-15, 92).
The taxa belong to the following families: Alliaceae (37, 79-81, 229), Apiaceae (109-111, 137-144), Araceae
(38), Asclepiadaceae (1), Asteraceae (2, 16-18, 51-59, 102-104, 145-153), Basellaceae (19), Boraginaceae (9, 93,
154-157, 239), Brassicaceae (20, 60, 92, 105, 158-165), Campanulaceae (166, 167), Caryophyllaceae (21-23,
61, 94, 95, 168-176), Chenopodiaceae (24, 62), Colchicaceae (230), Convolvulaceae (25), Crassulaceae (3, 26,
63, 177-179), Cucurbitaceae (122), Dipsacaceae (180-182), Eucommiaceae (117), Euphorbiaceae (183-185),
Fabaceae (11-13, 27-29, 64-70, 96, 116, 123, 186-189), Fagaceae (118), Geraniaceae (190), Globulariaceae
(191), Grossulariaceae (192), Hyacinthaceae (231), Hypericaceae (71, 72, 97, 193), Juncaceae (39-41, 82),
Lamiaceae (98, 194-204), Lauraceae (119), Liliaceae (42, 83, 232), Lythraceae (30), Molluginaceae (10),
Oleaceae (120), Onagraceae (106), Orchidaceae (113-115, 233-237), Orobanchaceae (205), Plantaginaceae (31,
206), Plumbaginaceae (207), Poaceae (43-50, 84-91, 126, 127-136, 238), Polygalaceae (208, 209), Polygonaceae
(32, 33, 73), Primulaceae (107), Ranunculaceae (14, 74, 112), Resedaceae (75), Rosaceae (4, 15, 34, 99, 100,
210), Rubiaceae (35, 76, 77, 101, 211-215), Rutaceae (5), Santalaceae (36, 216-218), Saxifragaceae (219),
Scrophulariaceae (220), Simaroubaceae (121), Solanaceae (6, 108, 124, 125), Tamaricaceae (221), Ulmaceae
(7), Urticaceae (8, 78), Veronicaceae (222-226) and Violaceae (227, 228).
New subspecies for science is: Centaurea athoa subsp. chelmea Kit Tan & al. (147).
New reports for countries are: Bulgaria – Eucommia ulmoides (117), Laurus nobilis (119); Greece –
Aeschynomene rudis (123), Euphorbia hyssopifolia (184).
The publication includes contributions by: I. Aneva & P. Zhelev (1-8), A. Asenov (9), P. Authier & Kit
Tan (10), M. Aybeke (11-13), M. Aybeke & C. Yarcı (14-15), B. Biel & Kit Tan (16-50), B. Biel & Kit Tan
(51-91), F. Dane & M. Aybeke (92), E. Filipova & K. Vassilev (93-101), K. Giannopoulos, Kit Tan & G. Vold
(102-108), K. Polymenakos & Kit Tan (109-112), A. Popatanasov (113-115), Kit Tan, G. Iszkuło, M. Dering
& E. Pers-Kamczyc (116), A. Tashev & N. Tashev (117-121), I. Tsialtas & Kit Tan (122-126), K. Vassilev &
H. Pedashenko (127-136), G. Zarkos, V. Christodoulou, Kit Tan & G. Vold (137-238), K.B. Simoglou, Ch.A.
Dordas & Th.V. Koutsos (239).
This is an ongoing report in the series dealing with the new chorological data on vascular plants in the Balkans.
For details on the presentation of information see Phytologia Balcanica, vol. 12(1), pp. 107-108 and vol. 12(2), p. 279.
*Reports for Bulgaria have been reviewed by V. Vladimirov, for Greece by Kit Tan, and for Turkey-in-Europe by F. Dane.

368 Vladimirov, V. & al • New floristic records in the Balkans: 28
Reports 1–8
Ina Aneva
1
& Petar Zhelev
2
1
Department of Plant and Fungal Diversity and Resources,
Institute of Biodiversity and Ecosystem Research,
Bulgarian Academy of Sciences, Acad. Georgi Bonchev St.,
bl. 23, 1113 Sofia, Bulgaria, e-mail: ina.aneva@abv.bg
2
Department of Dendrology, University of Forestry,
10 Kliment Ohridsky Blvd., 1797 Sofia, Bulgaria,
e-mail: zhelev@ltu.bg
Asclepiadaceae
1. Cynanchum acutum L.
Bu Tundzha Hilly Country: near a Quercus
pedunculiflora forest, about 2 km SE from Aitos,
42.65795°N, 27.30519°E, 04.08.2002, coll. P.
Zhelev (SOM 171501).
The find confirms distribution of the species in this
floristic region. Kuzmanov (1982) noted that the spe-
cies was reported for this floristic region near Sliven
by Velenovský (1898: 191).
Asteraceae
2. Carpesium cernuum L.
Bu Thracian Lowland: at the edge of a Quercus
pedunculiflora forest in the Krichim Park, KG96,
02.09.2013, coll. P. Zhelev (SOM 170826).
The species is new for the Thracian Plain. It could
be probably found in other lowland forests in the re-
gion.
Crassulaceae
3. Umbilicus erectus subsp. lassithiensis (Gand.) Stoj.
Bu Rhodopi Mts (Western): in the valley of river
Chepinska, in the vicinities of Marko Nikolov
railway station, KG66, 23.05.1993, coll. P. Zhelev
(SOM 154060).
The species has been known so far from the Rhodopi
Mts (Central, Eastern) (Assyov & Petrova 2012).
Rosaceae
4. Potentilla supina L.
Bu Valley of River Struma (Northern): on the gravel
edge of river Struma, close to Belo Pole village,
42.03685°N, 23.03785°E, 01.05.2014, coll. P.
Zhelev & I. Aneva (SOM 170830), 31.05.2014,
coll. P. Zhelev & I. Aneva (SOM 170831).
A new species for this floristic region.
Rutaceae
5. Haplophyllum suaveolens (DC.) G. Don.
Bu Mt Slavyanka: near Petrovo village, between the
road to Goleshevo village and river Petrovska,
41.4333°N, 23.5190°E, 12.07.2014, coll. I. Aneva
& P. Zhelev (SOM 170834).
A new species to this floristic region.
Solanaceae
6. Nicandra physalodes (L.) Gaertner
Bu Balkan Range (Eastern): at the edge of a mixed
oak forest in the Gorska Baraka locality,
42.9060°N, 27.7190°E, 01.08.2002, coll. P. Zhelev
(SOM 171500).
The closest localities of this adventive species are
along the Black Sea Coast.
Ulmaceae
7. Ulmus glabra Huds.
Bu Pirin Mts: along a river, about 12 km N from
Pirin village, near the road to Pirin chalet,
41.6108°N, 23.5279°E, 07.10.2012, coll. P. Zhelev
& I. Aneva (SOM 170829); close to Varbite
locality, along the road between Varbite and
Zagaza chalet, 41.7342°N, 23.2772°E, 14.10.2012,
coll. P. Zhelev & I. Aneva (SOM 170827).
These are the first records of this species in the Pirin Mts.
Urticaceae
8. Parietaria diffusa Mert. & W.D.J. Koch
Bu Valley of River Struma (Southern): on stony walls
in Melnik town, FL99, 07.10.2012, coll. I. Aneva
& P. Zhelev (SOM 170828).
First report of the species for this floristic region.
Report 9
Asen Asenov
Herbarium (SO), Faculty of Biology, St. Kliment Ohridski
Sofia University, 8 Dragan Tsankov Blvd., 1164 Sofia,
Bulgaria, e-mail: asenasenov71@yahoo.com
Boraginaceae
9. Echium russicum J.F. Gmel.
Gr Nomos & Eparchia Dramas: Mt Falakron, Aj.
Pnevma plateau, south of the EOS Katafigion,
1730–1750 m, subalpine pastures and rocky
outcrops, 41°19'N, 24°03'E, 01.07.1981,
Stamatiadou 22355 (ATH); south ridge of
Choros peak 1 km SW of plateau, 1746 m,
41°18'38"N, 24°02'54"E, 01.06.2015, coll. A.
Asenov (SO 107652).
Recently reported as new to the Greek flora (Simoglou
& al. 2015). In fact, the species was already collected

369Phytol. Balcan. 21(3) • Sofia • 2015
by Elli Stamatiadou in 1981. In 2015, a population
comprising ca. 30 flowering and 12 vegetative individ-
uals was noted in a small damp pothole. The plants
were growing together with Alchemilla xanthochlo-
ra, Anthyllis montana, Carex caryophyllea, Centaurea
tuberosa, Colchicum autumnale, Dactylorhiza sam-
bucina, Festuca rubra, Fragaria vesca, Galium verum,
Genista carinalis, Gentiana verna, Geranium sil-
vaticum, Hieracium cymosum, Luzula campestris,
Ranunculus montanus, Rhodax canus, Rosa pimpinel-
lifola, Plantago subulata, Serratula tinctoria, Sesleria
nitida, Taraxacum erythrospermum, Veronica chamae-
drys, etc.
Echium russicum occurs in Central and East
Europe, Caucasus and Anatolia, and is included in
Annex II of Directive 92/43/EEC.
Acknowledgements. I thank Kit Tan and Arne Strid for in-
formation on the earlier documentation of the species in Greece.
Report 10
Pierre Authier
1
& Kit Tan
2
1
Attaché Honoraire, Muséum National d’Histoire Naturelle,
Département Systématique et Évolution, CP 39, 57 rue
Cuvier, F-75231, Paris, France
2
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt @bio.ku.dk (corresponding author)
Molluginaceae
10. Glinus lotoides L. (Fig. 1)
Gr Nomos Ioanninon, Eparchia Dodonis: near
Kalpaki, ca. 500 m after Ioannina-Konitsa road
junction and road to Doliana, damp bare ground
near small pond, 400 m, 39°54'N, 20°36'E,
05.07.2013, Authier s.n. (herb. Authier); loc. ibid.,
466 m, 39°55'N, 20°36'E, 30.09.2015, fruiting,
Authier obs.; seasonally wet meadow near the
new road between Kalpaki and Albania, 397 m,
39°53'N, 20°35'E, 29.09.2015, Authier s.n. (herb.
Authier).
New for nomos, eparchia, Timfi area and phytogeo-
graphical region Northern Pindos (NPi). More than
a hundred plants in fruit and flower were noted. Salix
alba, Oenanthe fistulosa, Mentha pulegium, Alisma
lanceolatum, Xanthium spinosum, Trifolium fragifer-
um, Cirsium italicum, etc. occurred in the same local-
ities.
Reports 11–13
Mehmet Aybeke
Department of Biology, Faculty of Science,
University of Trakya, 22030 Edirne, Turkey,
e-mail: mehmetaybeke@yahoo.com
Fabaceae
11. Trifolium dubium Sibth.
Tu(E) A1(E) Çanakkale: Eceabat, in the environs
of Martyrdom Monument, 23 m, 40°11'03"N,
26°21'27"E, 11.05.1995, coll. M. Akpolat, det. M.
Aybeke (EDTU 2880).
— A1(E) Edirne: Keşan, Beykoy, in a pasture (04
B 03), 65 m, 41°16'N, 26°51'E, 05.05.2004, coll.
C. Kurt, A. Semerci & M. Aybeke, det. M. Aybeke
(TTAE 80);
New for A1(E) Edirne and Çanakkale in European
Turkey. According to Zohary (1970), this taxon was
reported for Tekirdağ.
12. Trifolium micranthum Viv.
Tu(E) A1(E) Edirne: Lalapasa, Kalkansöğüt,
in a pasture, 508 m, 41°56'26"N, 26°52'09"E,
17.07.2003, coll. C. Kurt, A. Semerci & M. Aybeke,
det. M. Aybeke (TTAE 1164).
New for A1(E) Edirne in European Turkey. According
to Zohary (1970), this taxon was reported for A2(E)
Istanbul.
13. Trifolium spumosum L.
Tu(E) A1(E) Edirne: Süloğlu, in Geçkinli pasture
(04 E 05), 183 m, 41°43'N, 26°51'E, 02.06.2004,
coll. C. Kurt, A. Semerci & M. Aybeke, det. M.
Aybeke (TTAE 1176).
Fig. 1. Glinus lotoides (photo P. Authier).

370 Vladimirov, V. & al • New floristic records in the Balkans: 28
New for A1(E) Edirne in European Turkey. According
to Zohary (1970), this taxon was reported for A1(E)
Kırklareli.
Reports 14–15
Mehmet Aybeke & Celal Yarcı
Department of Biology, Faculty of Science,
University of Trakya, 22030 Edirne, Turkey,
e-mail: mehmetaybeke@yahoo.com
Ranunculaceae
14. Ranunculus lanuginosus L.
Tu(E) A1(E) Kırklareli: Kırklareli, Demirköy, 1 km
along the road from Demirköy to Pınarhisar,
252 m, 41°49'30"N, 27°45'35"E, 03.03.1990, coll.
& det. C. Yarcı (EDTU 5055).
New for A1(E) Kırklareli in European Turkey. So far
the species has been known from A1(E) Edirne, A2(E)
Istanbul (Davis 1965).
Rosaceae
15. Filipendula vulgaris Moench
Tu(E) A1(E) Kırklareli: Demirköy, between
Demirköy and Pınarhisar, 1 km from Demirköy,
252 m, 41°49'30"N, 27°45'35"E, 02.07.1989, coll.
& det. C. Yarcı (EDTU 4161).
New for A1(E) Kırklareli in European Turkey.
According to Peşmen & Chamberlain (1972), this tax-
on was reported only for A2(E) Istanbul.
Reports 16–50
Burkhard Biel
1
& Kit Tan
2
1
Am Judengarten 3, D-97204 Höchberg, Germany
2
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku.dk (corresponding author)
This is the second report of new plant-records for
the island of Folegandros (phytogeographical region
Kiklades, Nomos Kikladon, Eparchia Milou) based on
three visits carried out in October 2004, March and
May 2005 respectively. The 35 records listed are new
to the island unless otherwise stated, and three species
were found to be new for the floristic region Kiklades
(Kik) as circumscribed in Flora Hellenica (Strid & Tan
1997), bringing the total number of new records for
this area to 34. Occurrence on the other Kikladean is-
lands is briefly summarized.
Asteraceae
16. Atractylis cancellata L.
Gr Folegandros: S of Ano Meria, abandoned vine
terraces above road, on schist, 100 m, 36°38'05"N,
24°51'24"E, 18.05.2005, Biel 05.130.
Also noted near Angali and Hora.
17. Crepis fraasii Sch. Bip. subsp. fraasii
Gr Folegandros: S-SW of Hora, pasture on limestone
slope, 320 m, 36°37'16"N, 24°54'57"E, 15.05.2005,
Biel 05.085.
18. Lactuca tuberosa Jacq.
Gr Folegandros: S-SW of Hora, rocky phrygana
limestone slope, 270 m, 36°37'24"N, 24°55'02"E,
15.05.2005, Biel 05.080.
Also noted near Karavostasis and Hora.
Basellaceae
19. Anredera cordifolia (Ten.) Steenis
Gr Folegandros: W of Hora, damp uncultivated
sandstone terraces, 180 m, 36°37'45"N,
24°55'05"E, 19.05.2005, Biel obs. (photo).
New for Kiklades; in Greece documented only from a
few islands (Karpathos, Kefalonia, Kerkira and Thasos).
Brassicaceae
20. Alyssum foliosum Bory & Chaub.
Gr Folegandros: SW of Karavostasis, grazed
phrygana near Livadi, on rocky limestone, 110 m,
36°36'16"N, 24°56'15"E, 07.03.2005, Biel 05.022.
Caryophyllaceae
21. Holosteum umbellatum L.
Gr Folegandros: E of Karavostasis, phrygana on
steep rocky schistose slope, 20 m, 36°37'02"N,
24°57'06"E, 07.03.2005, Biel 05.014.
22. Paronychia capitata (L.) Lam.
Gr Folegandros: W-NW of Karavostasis, stony
phrygana in valley, on limestone, 100 m,
36°37'10"N, 24°56'10"E, 07.03.2005, Biel 05.031.
New for Kiklades.
23. Silene pentelica Boiss.
Gr Folegandros: E of Karavostasis, phrygana on
steep rocky schistose slope, 20 m, 36°37'02"N,
24°57'06"E, 07.03.2005, Biel 05.013.
Chenopodiaceae
24. Atriplex prostrata DC.
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 15.05.2005, Biel 05.100.

371Phytol. Balcan. 21(3) • Sofia • 2015
Also noted S of Karavostasis.
Convolvulaceae
25. Cressa cretica L.
Gr Folegandros: S of Karavostasis, road margins in
west part of harbour, 5 m, 36°36'56"N, 24°56'52"E,
19.05.2005, Biel 05.157; S-SE of Karavostasis,
sandy and marshy ground at Livadi beach, 5 m,
36°36'30"N, 24°57'11"E, 19.05.2005, Biel 05.161.
Observed near Karavostasis in 2004.
Crassulaceae
26. Sedum rubens L.
Gr Folegandros: NE of Hora, concrete path
to monastery of Ag. Ioannis, phrygana on
limestone, 240 m, 36°37'50"N, 24°55'23"E,
17.05.2005, Biel obs. (photo).
Occurring in most of Kiklades.
Fabaceae
27. Medicago lupulina L.
Gr Folegandros: NE of Angali, phrygana on schistose
slope, 100 m, 36°38'07"N, 24°53'54"E, 08.03.2005,
Biel 05.034.
28. Melilotus siculus (Turra) B.D. Jacks.
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 19.05.2005, Biel 05.162.
29. Vicia lathyroides L.
Gr Folegandros: NE of Hora, concrete path to monastery
of Ag. Ioannis, phrygana on limestone, 240 m,
36°37'50"N, 24°55'23"E, 06.03.2005, Biel 05.060.
Lythraceae
30. Lythrum hyssopifolia L.
Gr Folegandros: N-NE of Angali, terraced olive slopes
in valley with seasonal stream, on schist, 30 m,
36°38'09"N, 24°53'47"E, 17.05.2005, Biel 05.133.
Also noted near Ano Meria.
Plantaginaceae
31. Plantago lanceolata L.
Gr Folegandros: W of Hora, wet uncultivated
sandstone terraces, 180 m, 36°37'45"N,
24°55'05"E, 19.05.2005, Biel 05.152a.
Polygonaceae
32. Rumex pulcher subsp. raulinii (Boiss.) Rech. f.
Gr Folegandros: W of Hora, wet uncultivated
sandstone terraces, 180 m, 36°37'45"N,
24°55'05"E, 19.05.2005, Biel 05.151.
33. Rumex scutatus L.
Gr Folegandros: W of Hora, wet uncultivated
sandstone terraces, 180 m, 36°37'45"N,
24°55'05"E, 19.05.2005, Biel 05.152.
Also noted near Ano Meria and Karavostasis. New for
Kiklades.
Rosaceae
34. Rosa agrestis Savi
Gr Folegandros: SW of Hora, uncultivated terraces
above road, 240 m, 36°37'35"N, 24°54'60"E,
15.05.2005, Biel obs. (photo).
Recorded from Andros.
Rubiaceae
35. Galium rotundifolium L.
Gr Folegandros: W of Hora, damp uncultivated
sandstone terraces, 180 m, 36°37'45"N,
24°55'05"E, 19.05.2005, Biel 05.152b.
Santalaceae
36. Thesium humile Vahl (Fig. 2)
Gr Folegandros: S-SE of Ano Meria, rocky phrygana
slope by path, on schist, 70 m, 36°38'07"N,
24°52'57"E, 18.05.2005, Biel 05.145.
Fig. 2. esium humile (photo B. Biel).

372 Vladimirov, V. & al • New floristic records in the Balkans: 28
Alliaceae
37. Allium dentiferum Webb & Berthel.
Gr Folegandros: SE of Angali, olive terraces along
dirt road, on limestone, 150 m, 36°37'40"N,
24°54'13"E, 17.05.2005, Biel 05.138.
Araceae
38. Arum italicum Mill.
Gr Folegandros: W of Hora, damp uncultivated
sandstone terraces, 180 m, 36°37'45"N,
24°55'05"E, 19.05.2005, Biel obs. (photo).
Also noted near Karavostasis.
Juncaceae
39. Juncus inflexus L.
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 19.05.2005, Biel 05.163.
40. Juncus maritimus Lam.
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 19.05.2005, Biel 05.165c.
41. Juncus subulatus Forssk.
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 09.03.2005, Biel 05.058.
Liliaceae
42. Gagea rigida Boiss. & Spruner (Fig. 3)
Gr Folegandros: SW of Karavostasis, grazed rocky
phrygana below Livadi, on marble/limestone,
110 m, 36°37'54"N, 24°56'27"E, 07.03.2005, Biel
05.021; S of Hora, phrygana in olive plantation,
on limestone, 250 m, 36°36'28"N, 24°55'28"E,
07.03.2005, Biel 05.061 (confirmed by J.-M.
Tison, October 2015).
Poaceae
43. Aegilops peregrina (Hack.) Maire & Weiller
Gr Folegandros: SW of Karavostasis, grazed rocky
phrygana at pass, on limestone, 260 m, 36°36'21"N,
24°56'01"E, 15.05.2005, Biel 05.097b; N of Ano
Meria, olive terraces, on schistose rock, 40 m,
36°39'45"N, 24°51'48"E, 16.05.2005, Biel 05.116; W
of Hora, limestone plateau with phrygana, 160 m,
36°37'43"N, 24°54'51"E, 17.05.2005, Biel 05.127; S
of Ano Meria, phrygana on rocky limestone slope
S of Trachilias hill, 100 m, 36°38'05"N, 24°51'56"E,
18.05.2005, Biel 05.140; S-SW of Hora, rocky
phrygana limestone slope near Ag. Nikolaos,
220 m, 36°36'51"N, 24°54'42"E, 20.05.2005, Biel
05.168.
44. Elytrigia atherica (Link) Kerguélen [syn.: Elymus
pycnanthus (Godr.) Melderis]
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 19.05.2005, Biel 05.165.
45. Elytrigia elongata (Host) Nevski
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 15.05.2005, Biel 05.099.
46. Hordeum marinum Huds.
Gr Folegandros: S-SE of Karavostasis, sandy and
marshy ground at Livadi beach, 5 m, 36°36'30"N,
24°57'11"E, 28.10.2004, Biel 04.007.
Also noted near Ano Meria.
47. Hordeum murinum subsp. glaucum (Steud.)
Tzvelev
Gr Folegandros: NW of Ano Meria, rocky phrygana
at road near Karkana, on schist, 210 m,
36°39'19"N, 24°51'29"E, 16.05.2005, Biel 05.119.
48. Phalaris aquatica L.
Gr Folegandros: S of Hora, clearing in olive
Fig. 3. Gagea rigida (photo B. Biel).

373Phytol. Balcan. 21(3) • Sofia • 2015
plantation with phrygana, on limestone, 250 m,
36°36'28"N, 24°55'28"E, 15.05.2005, Biel 05.096.
Also noted near Ano Meria.
49. Phalaris canariensis L.
Gr Folegandros: S of Karavostasis, grazed phrygana
in-between fields, 10 m, 36°36'22"N, 24°57'02"E,
19.05.2005, Biel 05.158.
50. Setaria adhaerens (Forssk.) Chiov.
Gr Folegandros: SE of Angali, grassy olive terraces
along dirt road, on limestone, 150 m, 36°37'40"N,
24°54'13"E, 17.05.2005, Biel 05.138a.
Cited vouchers are provisionally kept in the private
herbarium of B. Biel at Höchberg (herb. Biel). We
thank Prof. Hildemar Scholz
†
(Berlin) for kindly iden-
tifying our Poaceae specimens.
Reports 51– 91
Burkhard Biel
1
& Kit Tan
2
1
Am Judengarten 3, D-97204 Höchberg, Germany
2
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku.dk (corresponding author)
This is the second report of new plant-records for the
island of Sikinos (phytogeographical region Kiklades,
Nomos Kikladon, Eparchia Milou) based on three
visits carried out in October 2004, March and May
2005 respectively. All records listed are new to the is-
land unless otherwise stated. Occurrence on the other
Kikladean islands is briefly summarized.
Asteraceae
51. Centaurea laconica subsp. lineariloba (Halácsy
& Dörfl.) E. Gamal-Eldin & Wagenitz (Fig. 4)
Gr Sikinos: SW of Hora, uncultivated field at
hill saddle, on limestone, 400 m, 36°41'13"N,
25°06'28"E, 25.05.2005, Biel obs. (photo).
On several Kikladean islands.
52. Chondrilla juncea L.
Gr Sikinos: Hora-Apano, road margins and stone
steps in village, 280 m, 36°41'38"N, 25°07'01"E,
25.05.2005, Biel 05.189.
Almost throughout Kiklades.
53. Crepis setosa Haller f.
Gr Sikinos: E-NE of Hora, stony phrygana slopes
near chapel Prof. Ilias, 340 m, 36°42'26"N,
25°08'42"E, 24.05.2005, Biel 05.185.
Recorded from Andros.
54. Erigeron bonariensis L.
Gr Sikinos: Hora-Apano, road margins and stone
steps in village, 280 m, 36°41'38"N, 25°07'01"E,
25.05.2005, Biel 05.188.
On several Kikladean islands.
55. Filago eriocephala Guss.
Gr Sikinos: SW of Hora, terraced slopes near
chapel Ag. Pantes, on schist, 270 m, 36°41'24"N,
25°06'24"E, 22.05.2005, Biel 05.159.
Almost throughout Kiklades.
56. Lactuca acanthifolia (Willd.) Boiss.
Gr Sikinos: Hora-Apano, road embankments, stone
steps in village, 280 m, 36°41'38"N, 25°07'01"E,
25.05.2005, Biel obs. (photo).
On several Kikladean islands.
57. Senecio leucanthemifolius Poir.
Gr Sikinos: S of Hora, abandoned terraces along
old path below village, on schist, 220 m,
36°41'33"N, 25°07'10"E, 26.05.2005, Biel 05.001;
SW of Alopronia, phrygana slope near holiday
homes, on marble, 10 m, 36°40'32"N, 25°08'46"E,
28.05.2005, Biel 05.038.
Recorded from Amorgos, Naxos, Ofidoussa, Paros
and Sifnos. Also noted near Alopronia and Hora.
Fig. 4. Centaurea laconica subsp. lineariloba (photo B. Biel).

374 Vladimirov, V. & al • New floristic records in the Balkans: 28
58. Taraxacum minimum (Guss.) N. Terracc.
Gr Sikinos: S of Hora, stony terraces near threshing
place and chapel Ag. Stefanos, on schist, 340 m,
36°41'20"N, 25°07'05"E, 31.10.2005, Biel obs.
(photo).
Recorded from Amorgos, Folegandros, Kea, Kimolos,
Naxos and Paros.
59. Taraxacum hellenicum Dahlst.
Gr Sikinos: N-NE of Hora, steep phrygana slope
with spring near Vorino, on schist, 100 m,
36°42'02"N, 25°07'19"E, 28.05.2005, Biel 05.046.
On several Kikladean islands.
Brassicaceae
60. Sisymbrium irio L.
Gr Sikinos: W of Hora-Kastro, abandoned terraces,
road margins, on schist, 260 m, 36°41'24"N,
25°06'24"E, 24.05.2005, Biel 05.176.
Recorded from Amorgos, Denoussa, Kimolos,
Kithnos, Naxos, Siros and Thira.
Caryophyllaceae
61. Dianthus tripunctatus Sm. (Fig. 5)
Gr Sikinos: W-SW of Hora, stony phrygana and olive
terraces near manor Villa, on limestone, 270 m,
36°41'40"N, 25°06'50"E, 22.05.2005, Biel 05.146.
Recorded from Amorgos, Andros, Ios, Kea, Kithnos,
Serifos, Siros and Tinos.
Chenopodiaceae
62. Chenopodium opulifolium Schrad.
Gr Sikinos: W of Hora, steep terraced slope
with Euphorbia along steps, on schist, 130 m,
36°41'48"N, 25°06'54"E, 01.11.2004, Biel 04.030.
On several Kikladean islands.
Crassulaceae
63. Sedum caespitosum (Cav.) DC.
Gr Sikinos: SW of Alopronia, stony phrygana and
olive terraces along old path, on limestone, 100 m,
36°40'06"N, 25°07'47"E, 02.03.2005, Biel 05.079a.
Recorded from Anafi, Giaros, Giaros (Sourli), Milos,
Naxos and Paros.
Fabaceae
64. Ceratonia siliqua L.
Gr Sikinos: SW of Alopronia, stony phrygana and
olive terraces along old path, on limestone, 100 m,
36°40'06"N, 25°07'47"E, 02.03.2005, Biel 05.065.
On several Kikladean islands.
65. Lathyrus clymenum L.
Gr Sikinos: SW of Alopronia, phrygana slope near
holiday homes, on marble, 10 m, 36°40'32"N,
25°08'46"E, 28.05.2005, Biel 05.039.
On several Kikladean islands.
66. Medicago truncatula Gaertn.
Gr Sikinos: S-SW of Hora, stony phrygana with
Juniperus at path in Klosenas valley, on limestone,
130 m, 36°39'28"N, 25°05'53"E, 04.03.2005, Biel
05.105; W-NW of Hora-Kastro, abandoned
terraces with phrygana at steps, on schist, 170 m,
36°41'49"N, 25°06'58"E, 24.05.2005, Biel 05.178.
Almost throughout Kiklades.
67. Ononis pubescens L.
Gr Sikinos: SW of Alopronia, stony phrygana and
olive terraces at old path, on limestone, 100 m,
36°40'06"N, 25°07'47"E, 23.05.2005, Biel 05.174.
On several Kikladean islands. Also noted N and S of Hora.
68. Trifolium cherleri L.
Gr Sikinos: N-NW of Alopronia, grazed phrygana
slope near well, on limestone and schist, 140 m,
36°41'15"N, 25°08'23"E, 21.05.2005, Biel 05.140.
On several Kikladean islands.
69. Trifolium infamia-ponertii Greuter
Gr Sikinos: W-SW of Hora, grazed phrygana terraces
near chapel, on schist, 280 m, 36°41'33"N,
25°06'34"E, 22.05.2005, Biel 05.151.Fig. 5. Dianthus tripunctatus (photo B. Biel).

375Phytol. Balcan. 21(3) • Sofia • 2015
Common, almost throughout Kiklades.
70. Vicia lathyroides L.
Gr Sikinos: NE of Hora, stony phrygana with
Juniperus near monastery Zoodochou Pighi,
on limestone, 350 m, 36°41'55"N, 25°07'25"E,
03.03.2005, Biel 05.096.
On several Kikladean islands.
Hypericaceae
71. Hypericum perforatum L.
Gr Sikinos: SW of Hora, terraced phrygana in
valley near chapel Ag. Pantes, on schist, 270 m,
36°41'24"N, 25°06'24"E, 22.05.2005, Biel 05.160.
Recorded from Andros, Naxos, Serifos and Tinos.
72. Hypericum tetrapterum Fr.
Gr Sikinos: W-NW of Hora, stony phrygana with
Euphorbia and olive trees, on schist, 160 m,
36°41'48"N, 25°06'45"E, 05.03.2005, Biel 05.109.
Reported only from Andros.
Polygonaceae
73. Rumex pulcher subsp. raulinii (Boiss.) Rech. f.
Gr Sikinos: Hora-Kastro, ruderal places and
road margins in village, on limestone, 280 m,
36°41'47"N, 25°07'18"E, 21.05.2005, Biel 05.132.
On several Kikladean islands.
Ranunculaceae
74. Clematis cirrhosa L.
Gr Sikinos: SW of Hora, phrygana terraces near
channel, along old path, on schist, 300 m,
36°40'42"N, 25°05'51"E, 27.02.2005, Biel obs.
(photo).
On several Kikladean islands.
Resedaceae
75. Reseda alba L.
Gr Sikinos: S-SW of Hora, stony phrygana slope
near chapel Ag. Marina, on marble, 360 m,
36°39'37"N, 25°05'23"E, 23.05.2005, Biel 05.172.
On several Kikladean islands. Also noted near Hora.
Rubiaceae
76. Crucianella angustifolia L.
Gr Sikinos: NE of Hora, stony phrygana on ridge
near chapel Ag. Fotini, on limestone, 100 m,
36°42'37"N, 25°08'13"E, 01.11.2004, Biel 04.027.
On several Kikladean islands.
77. Valantia aprica (Sm.) Tausch
Gr Sikinos: Hora (Apano Chorio), ruderal places near
old threshing place, on limestone and schist, 310 m,
36°41'34"N, 25°06'58"E, 27.02.2005, Biel 05.019.
Reported only from Naxos.
Urticaceae
78. Urtica urens L.
Gr Sikinos: N of Alopronia, edge of road junction,
on limestone and schist, 30 m, 36°40'48"N,
25°08'34"E, 28.02.2005, Biel 05.044.
On several Kikladean islands.
Alliaceae
79. Allium ampeloprasum L.
Gr Sikinos: S-SE of Hora, rocky slope at stream,
phrygana on limestone, 60 m, 36°39'37"N,
25°05'23"E, 23.05.2005, Biel obs.
On several Kikladean islands. Also noted W and E of
Hora.
80. Allium roseum L.
Gr Sikinos: SW of Hora (Apano Chorio), terraces
and road margins, phrygana on limestone and
schist, 420 m, 36°41'20"N, 25°06'47"E, 22.05.2005,
Biel 05.164.
On several Kikladean islands.
81. Allium trifoliatum Cirillo (Fig. 6)
Gr Sikinos: SW of Hora, terraced phrygana in
valley near chapel Ag. Pantes, on schist, 270 m,
Fig. 6. Allium trifoliatum (photo B. Biel).

376 Vladimirov, V. & al • New floristic records in the Balkans: 28
36°41'24"N, 25°06'24"E, 22.05.2005, Biel obs.
(photo).
Recorded from Andros, Kimolos, Milos, Naxos, Paros,
Siros, Thira and Tinos.
Juncaceae
82. Juncus minutulus V. Krecz. & Gontsch.
Gr Sikinos: SW of Alopronia, stony phrygana in
dry stream bed by old path, on limestone, 50 m,
36°40'11"N, 25°07'56"E, 02.03.2005, Biel 05.083.
Recorded from Andros, Ios, Paros, Sifnos and Tinos.
Liliaceae
83. Ornithogalum arabicum L.
Gr Sikinos: SW of Hora (Apano Chorio), phrygana
terraces and road margins, on limestone and
schist, 420 m, 36°41'20"N, 25°06'47"E, 22.05.2005,
Biel 05.165.
Recorded from Anafi, Iraklia, Milos, Paros, Sifnos and
Siros.
Poaceae
84. Alopecurus myosuroides Huds.
Gr Sikinos: W-SW of Hora, stony phrygana and olive
terraces near manor Villa, on limestone, 270 m,
36°41'40"N, 25°06'50"E, 22.05.2005, Biel 05.150a.
On several Kikladean islands.
85. Bromus diandrus Roth
Gr Sikinos: E of Hora, road margins and olive
terraces below village, on schist, 60 m,
36°41'45"N, 25°07'33"E, 02.11.2004, Biel 04.033.
On several Kikladean islands. Also noted W of Hora.
86. Gastridium ventricosum (Gouan) Schinz & ell.
Gr Sikinos: E-SE of Hora, schistose roadside slope,
180 m, 36°41'26"N, 25°07'53"E, 21.05.2005, Biel
05.136; N-NW of Alopronia, grazed phrygana
slope near well by path, on limestone and schist,
140 m, 36°41'15"N, 25°08'23"E, 21.05.2005, Biel
05.142.
Almost throughout Kiklades.
87. Hordeum marinum Huds.
Gr Sikinos: Alopronia, beach and estuary, 2 m,
36°40'35"N, 25°08'43"E, 30.10.2004, Biel obs.
On several Kikladean islands.
88. Lolium perenne L.
Gr Sikinos: SW of Hora (Apano Chorio),
uncultivated field at saddle, on limestone, 400 m,
36°41'13"N, 25°06'28"E, 25.05.2005, Biel 05.192.
On several Kikladean islands. Also noted E of Hora.
89. Poa inrma Kunth
Gr Sikinos: SW of Alopronia, stony phrygana in dry
stream bed near old path, on limestone, 50 m,
36°40'11"N, 25°07'56"E, 02.03.2005, Biel 05.082.
On several Kikladean islands. Also noted N of
Alopronia and Hora.
90. Polypogon viridis (Gouan) Breistr.
Gr Sikinos: W-NW of Hora-Kastro, abandoned
terraces with well, on schist, 120 m, 36°41'50"N,
25°06'56"E, 24.05.2005, Biel 05.179.
On several Kikladean islands. Also in vicinity of Hora.
91. Rostraria smyrnacea (Trin.) H. Scholz
Gr Sikinos: W-SW of Hora, grazed phrygana slopes
near chapel, on schist, 280 m, 36°41'33"N,
25°06'34"E, 22.05.2005, Biel 05.153.
Only reported from Naxos and Paros.
Cited vouchers are provisionally kept in the private
herbarium of B. Biel at Höchberg (herb. Biel). We
thank Prof. Hildemar Scholz
†
(Berlin) for kindly iden-
tifying our Poaceae specimens.
Report 92
Feruzan Dane & Mehmet Aybeke
Department of Biology, Faculty of Science,
University of Trakya, 22030 Edirne, Turkey,
e-mail: mehmetaybeke@yahoo.com
Brassicaceae
92. Alyssum hirsutum M. Bieb.
Tu(E) A1(E) Edirne: Centre, 26 m, 41°40'33"N,
26°33'31"E, 10.04.2001, coll. F. D ane & G. Dalgıç,
det. G. Dalgıç (EDTU 8281) & 05.06.1988, coll. F.
Dane, det. N. Orcan (EDTU 2685).
This is a new species for European Turkey. According to
Dudley & al. (1965), this taxon was reported for A2(A)
Istanbul, but not from any region in European Turkey.
Reports 93–101
Eva Filipova
1
& Kiril Vassilev
2
1
Department of Ecology and Environmental Protection,
Faculty of Biology, St. Kliment Ohridski University of
Sofia, 8 Dragan Tzankov Blvd., 1164 Sofia, Bulgaria,
e-mail: eveto_filipova@abv.bg
2
Department of Plant and Fungal Diversity and Resources,
Institute of Biodiversity and Ecosystem Research,
Bulgarian Academy of Sciences, Acad. Georgi Bonchev St.,
bl. 23, 1113 Sofia, Bulgaria, e-mail: kiril5914@abv.bg

377Phytol. Balcan. 21(3) • Sofia • 2015
Boraginaceae
93. Alkanna tinctoria (L.) Tausch
Bu Rhodopi Mts (Western): in dry grasslands
between Velingrad town and Yundola village,
42.03811°N, 23.90109°E, 05.08.2015, coll. K.
Vassilev & E. Filipova (SOM 171515).
Found in xerothermic communities as a companion
species. According to Evstatieva (2015), it is an en-
dangered species and is known from seven floristic re-
gions. It is also protected by Annex 3 of the Biological
Diversity Act (2007). This locality is isolated from the
other populations in the country.
Caryophillaceae
94. Minuartia recurva (All.) Schinz & Thell.
Bu Balkan Range (Central): in dry rocky places
near Kazanlak town, LH62, 17.08.1941, coll.
A. Jurkovskyi (SOM 21133); in rocky grasslands
under peak Botev, 2100 m, LH33, 02.07.1972,
coll. P. P an o v (SOM 147542); on rocks SE of
Tazha chalet, under the Chin Koliba locality,
1580 m, LH33, 06.08.1956, coll. I. Bondev (SOM
157651); on rocks under Rai chalet, LH32, 1965,
coll. D. Peev & S. Kozhuharov (SOM 11519); on
rocks near Rai chalet, along the trail to peak
Botev, LH32, 17.08.2008, coll. D. Dimitrov (SOM
164514).
— Vitosha region: in dry grasslands near
Zlokuchene village, 42.38198°N, 23.56673°E,
GM09, 07.07.2015, coll. K. Vassilev & E. Filipova
(SOM 171516).
This is a new species for these floristic regions. So
far it has been known from nine floristic regions in
Bulgaria (Assyov & Petrova 2012). Its population near
Zlokuchene village is represented by 20–30 individu-
als and is developed on shallow soils and steep slopes.
95. Minuartia setacea (uill.) Hayek
Bu Balkan Range (Central): on rocks around peak
Kademliya, at 1600–2000 m, LH42, 04.06.1956,
coll. I. Bondev (SOM 177652).
— Pirin Mts (Northern): Bayuvi Dupki Reserve,
in the Golyama Bayuva Dupka circus in rocky
places around Parev Grob locality, 1880 m, FM94,
08.06.1976, coll. N. Andreev (SOM 135503).
This species is found in stony grasslands, from low-
lands up to high mountains. It is a companion species
in grasslands of classes Asplenietea trichomanes and
Festuco-Brometea. It is a new species for these floris-
tic regions.
Fabaceae
96. Genista januensis Viv.
Bu Rhodopi Mts (Eastern): in dry grasslands near
Chernichino village, MG00, 24.05.2015, coll. K.
Vassilev & E. Filipova (SOM 171510).
The species is widespread in Bulgaria, but has not
been reported for this floristic region before.
Hypericaceae
97. Hypericum linarioides Bosse
Bu Tundzha Hilly Country: in dry grasslands near
Dabravite village, 42.23706°N, 24.01347°E, KG57,
25.07.2015, coll. K. Vassilev & E. Filipova (SOM
171519).
This species in found locally in xerothermic grass-
lands. It is a new species for this floristic region.
Lamiaceae
98. Thymus striatus Vahl
Bu Vitosha region: in xerothermic grasslands near
Pasarel village, GN01, 42.54088°N, 23.51577°E,
06.07.2015, coll. K. Vassilev & E. Filipova (SOM
171522).
This is a new species for this floristic region found
in grasslands of the class Festuco-Brometea and alli-
ances Saturejion montanae and Festucion valesiacae.
Other common species in the area are: Festuca dal-
matica, Phleum phleoides, Koeleria nitidula, Astragalus
onobrychis, Dichantium ischaemum, Seseli pallasii,
Euphorbia barrelieri, Teucrium chamaedrys, etc.
Rosaceae
99. Potentilla pedata Willd.
Bu Rhodopi Mts (Western): in dry grasslands near
Dorkovo village, 42.04955°N, 24.01867°E, KG65,
05.08.2015, coll. K. Vassilev & E. Filipova (SOM
171512).
This is a widespread species in xerothermic grasslands
and is new for this floristic region.
100. Potentilla pilosa Willd.
Bu Rhodopi Mts (Central): in dry grasslands near
Yagodina village, 41.64240°N, 24.36369°E,
KG71, 07.08.2015, coll. K. Vassilev & E. Filipova
(SOM 171514); around St. Petka Monastery near
Asenovgrad town, LG24, 29.06.1966, coll. M.
Markova (SOM 127275).
This is a new species for this floristic region. Near
Yagodina, it is found in rocky grasslands around the
village and in dry grasslands of the classes Asplenieta
trichomanes and Festuco-Brometea. These com-

378 Vladimirov, V. & al • New floristic records in the Balkans: 28
munities host a great diversity of endemic and ra-
re species (Petrova & Vladimirov 2009, 2010),
such as Scabiosa rhodopensis, Micromeria dalmati-
ca, Achillea clypeolata, Sesleria latifolia, Trachelium
rumelianum, etc.
Rubiaceae
101. Asperula tenella Degen
Bu Vitosha region: in xerothermic grasslands near
Pasarel village, 42.54088°N, 23.51577°E, GN01,
06.07.2015, coll. K. Vassilev & E. Filipova (SOM
171520).
— Rhodopi Mts (Western): in dry grasslands near
Tsrancha village, 41.56428°N, 24.11204°E, KG50,
06.08.2015, coll. K. Vassilev & E. Filipova (SOM
171518).
The species is widespread in xerothermic grasslands
as a companion species, but this is its first record for
these floristic regions.
Reports 102–108
Konstantinos Giannopoulos
1
, Kit Tan
2
& Gert Vold
3
1
Dabaki 15, Pyrgos, Ilias 271 00, Greece
2
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku.dk (corresponding author)
3
Botanical Garden, State Natural History Museum, Øster
Farimagsgade 2C, DK-1353 Copenhagen K, Denmark
Continuing a series of new plant records based on fur-
ther floristic investigations in the prefecture of Ilia
in western Peloponnese. The records listed are new
for Eparchia Ilias or Olimbias, or for both eparchies
in Nomos Ilias. The floristic regions adopted follow
those circumscribed in Flora Hellenica (Strid & Tan
1997).
Asteraceae
102. Cirsium eriophorum (L.) Scop. (Fig. 7)
Gr Nomos & Eparchia Ilias: on road to Astras
plateau, Mt Lambia, 1200 m, 37°54'N, 21°47'E,
09.08.2015, Kit Tan, G. Vold & Giannopoulos s.n.
(herb. Giannopoulos).
New for nomos and eparchia. In Peloponnese, oc-
curring in nomi Achaias, Arkadias (Mt Likeo) and
Messinias (Mt Taigetos).
103. Cirsium hypopsilum Boiss. & Heldr.
Gr Nomos & Eparchia Ilias: on road to Astras
plateau, Mt Lambia, 1200 m, 37°54'N, 21°47'E,
09.08.2015, Kit Tan, G. Vold & Giannopoulos s.n.
(herb. Giannopoulos).
New for nomos and eparchia. On mountains of N, C
and S Peloponnese and Sterea Ellas (Mt Gerania).
104. Echinops sphaerocephalus subsp. albidus
(Boiss. & Spruner) Maire & Petitm. (Fig. 8)
Gr Nomos & Eparchia Ilias: on road to Astras
plateau, Mt Lambia, 1200 m, 37°54'N, 21°47'E,
Fig. 8. Echinops sphaerocephalus subsp. albidus (photo K. Gian-
nopoulos).
Fig. 7. Cirsium eriophorum (photo K. Giannopoulos).

379Phytol. Balcan. 21(3) • Sofia • 2015
09.08.2015, Kit Tan, G. Vold & Giannopoulos s.n.
(herb. Giannopoulos).
New for nomos and eparchia. In Peloponnese, in no-
mi Achaias, Arkadias and Korinthias.
Brassicaceae
105. Lepidium graminifolium L.
Gr Nomos Ilias, Eparchia Olimbias: outside
Andritsena, 780 m, 37°28'N, 21°54'E, 08.08.2015,
Kit Tan, G. Vold & Giannopoulos 31928 (herb.
Giannopoulos).
New for eparchia; also collected in nearby locality by
Willing (Willing 117981, B).
Onagraceae
106. Oenothera biennis L. (Fig. 9)
Gr Nomos & Eparchia Ilias: outside village
Lanzoi, 80 m, 37°42'N, 21°33'E, 09.08.2015,
Kit Tan, G. Vold & Giannopoulos 31935 (herb.
Giannopoulos).
New for Peloponnese; garden escape, naturalized at
outskirts of village. Native to N America and widely
naturalized in temperate regions.
Primulaceae
107. Cyclamen graecum Link (Fig. 10)
Gr Nomos Ilias, Eparchia Olimbias: outside village of
Taxiarches, 300 m, 37°27'N, 21°42'E, 29.11.2015,
Kit Tan, G. Vold & Giannopoulos obs. (photos);
Lepreo archaeological site, 425 m, 37°26'N, 21°43'E,
29.11.2015, Kit Tan, G. Vold & Giannopoulos obs.
(photos); Ancient Figalia, archaeological site,
510 m, 37°24'N, 21°50'E, 29.11.2015, Kit Tan, G.
Vold & Giannopoulos 32024 (herb. Giannopoulos);
Mt Minthi, 1210 m, 37°29'N, 21°46'E, 24.10.2015,
Giannopoulos & Vendras obs. (photos).
New for nomos and eparchia; widespread in N, C and
S Peloponnese and flowering at the same time as C.
hederifolium.
Solanaceae
108. Nicotiana glauca R.C. Graham
Gr Nomos & Eparchia Ilias: near village Lanzoi,
120 m, 37°42'N, 21°33'E, 09.08.2015, Kit
Tan, G. Vold & Giannopoulos 31934 (herb.
Giannopoulos).
New for Eparchia Ilias; also collected by Willing, SW
of Vardha (Willing 107069, B). Naturalized near vil-
lage and along roadsides. Native to S America.
Reports 109–112
Kostas Polymenakos
1
& Kit Tan
2
1
Psaron 67, Chalandri 152 32, Attikis, Greece
2
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku.dk (corresponding author)
Continuing a series of new plant records based on fur-
ther floristic investigations in Greece. The floristic re-
gions adopted follow those circumscribed in Flora
Hellenica (Strid & Tan 1997).
Apiaceae
109. Opopanax chironium (L.) W.D.J. Koch (Fig. 11)
Gr Nomos & Eparchia Fthiotidos: Mt Iti, SE of
peak Greveno, 1660–1665 m, 38°48'N, 22°17'E, Fig. 9. Oenothera biennis (photo K. Giannopoulos).
Fig. 10. Cyclamen graecum (photo K. Giannopoulos).

380 Vladimirov, V. & al • New floristic records in the Balkans: 28
27.08.2015, Polymenakos obs. (photos; confirmed
Kit Tan, October 2015).
New for Mt Iti, nomos and eparchia Fthiotidos. A few
plants noted on the serpentine slopes. In Greece often
photographed or collected in flowering state but rare-
ly with ripe fruits. O. hispidus common in southern
Greece has fruits with fewer commissural vittae.
110. Peucedanum aegopodioides (Boiss.) Vandas
(Fig. 12)
Gr Nomos & Eparchia Fthiotidos: by stream between
Ano and Kato Pavliani, 900 m, 38°44'N, 22°20'E,
26.08.2015, Polymenakos obs. (photos; confirmed
Kit Tan, October 2015).
A Balkan endemic, new for Mt Iti, nomos and epar-
chia Fthiotidos and phytogeographical region Sterea
Ellas. On mainland Greece occurring north of 39°N
in N & S Pindos, EC, NC and NE Greece. Numerous
plants found, distinct by the Aegopodium podagraria-
like leaves and fruits with broad wings.
111. Seseli parnassicum Boiss. & Heldr. (Fig. 13)
Gr Nomos Fokidos, Eparchia Parnassidos:
Pavliani to Pira, opening of Abies woodland,
1180‒1200 m, 38°42'N, 22°19'E, 26.08.2015,
Polymenakos obs. (photos; confirmed Kit Tan,
October 2015).
New for Mt Iti; in Sterea Ellas, recorded from Mts
Giona, Panetoliko, Parnassos and Timfristos. A sin-
gle plant observed.
Fig. 13. Seseli parnassicum (photo K. Polymenakos).
Fig. 12. Peucedanum aegopodioides (photo K. Polymenakos).
Fig. 11. Opopanax chironium (photo K. Polymenakos).
Ranunculaceae
112. Delphinium fissum Waldst. & Kit. subsp. fissum
Gr Nomos & Eparchia Fthiotidos: meadow south of
village Kastania, 1240‒1245 m, 38°50'N, 22°12'E,
21.07.2015, Polymenakos obs. (photos; confirmed
Kit Tan, October 2015).
New for Mt Iti; in Sterea Ellas, recorded from Mts
Chelidonia, Giona, Kaliakouda, Kitheronas, Parnitha,
Pateras, Timfristos, Xerovouni and Vardousia. Single
plant observed.
Reports 113–115
Andrey Popatanasov
Faculty of Biology, Paisij Hilendarski Plovdiv University,
Plovdiv, Bulgaria, e-mail: atanasov@abv.bg
Orchidaceae
113. Epipactis persica (Soó) Nannf. (Fig. 14)
Bu Rhodopi Mts (Central): at the shrub edge of
an old, open, mixed, mostly coniferous forest,
along a forest dirt road near Chereshkite village,
on a southwest-facing slope, with inclination
approx. 30 degrees, 1068 m, 41°29'40.044"N,
24°35'2.082"E, with flowers and fruits,
03.08.2015, A. Popatanasov obs.

381Phytol. Balcan. 21(3) • Sofia • 2015
Already reported for this floristic region from a near-
by locality (Petrova & Vassilev 2006). Only two plants
were found not far from each other above a forest dirt
road used for transportation of timber. Presumably,
the population was bigger, since several aerial stems
were found nearby that died in the flowering phase
because of excessive drought during this year’s vege-
tation season. They had similar morphology and size.
Other species from this family found in this forest
were Epipactis helleborine and Gymnadenia conopsea.
Although this taxon was reported for the first time
to the Bulgarian flora more than a quarter century ago
(Siering & Henning 1990), only one more location
in the country was reported ever since and only one
specimen has been deposited in the Bulgarian herbar-
ia (Petrova & Vassilev 2006).
Regardless of the rarity and peculiar vulner-
able biology of this species, it is not yet included in
the list of endangered plants in Bulgaria under the
Biodiversity Act, nor in Red List of Bulgarian vascular
plants (Petrova & Vladimirov 2009). This is its third
reported location which reconfirms the presence of
this taxon in the Bulgarian flora, providing further ar-
guments for getting an officially protected status from
the institutions in charge of these matters.
The location lies less than 200 meters away from
an area of active legal woodcutting activities which
highly endanger the survival of the population ei-
ther by direct damage or indirectly by anthropogen-
ic changes in the ecosystem.
114. Orchis laxiora Lam. (Fig. 15)
Bu Mt Pirin (Northern): in a waterlogged marshy
meadow, near a dirt road that connects to the
main road for Senokos village, few kilometres
downhill, on a west-facing slope, with inclination
from 1 to 10 degrees, 775 m, 41°50'47.16"N,
23°11'55.602"E, 18.05.2015, with flowers and
fruits, coll. A. Popatanasov (SO 107665).
The species is new for this floristic region and moun-
tain, according to Kitanov & Kitanov (1990) and
Assyov & Petrova (2012). The population occupies an
area of over 1 ha, adjacent to a water stream around
which a marshy meadow is formed, rich in mosses
and hygrophyte flora. The area probably extends fur-
Fig. 14. Epipactis persica (photo A. Popatanasov).
Fig. 15. Orchis laxiora (photo A. Popatanasov).

382 Vladimirov, V. & al • New floristic records in the Balkans: 28
ther downhill, but the land is mostly privately owned
and it is hard to make a correct estimate due to limited
access. The population is rather big for this rare and
endangered species, comprising over five hundred in-
dividuals. Over 50 % of the population occupies the
marshy territories near the stream. The rest of it occu-
pies the adjacent less moist areas with higher degree
of inclination.
A relatively large size of the population increases
the chance of occurrence of such genetically recessive
phenomena as hypochromia. Five individuals with
white flower pigmentation and 11 individuals with
pale-pink flower coloration were found randomly dis-
tributed among the individuals with the normal flow-
er coloration pattern.
Close to this population, seven spatially separat-
ed populations of O. elegans (Delforge 2006) – a close-
ly related to O. laxiflora species – were also found in
the appropriate moist to waterlogged habitats that are
characteristic for them. However, regardless of their
similar ecological requirements, after examination of
the populations no plants of the other closely related
species were found within the territory occupied by
them: the populations of O. elegans had no plants of
O. laxiflora among them and vice versa. Thus a geo-
graphical barrier of just few hundred meters is appar-
ently sufficient to ensure spatial populational isolation
for these closely related taxa.
Another species from this family found in this
meadow was Dactylorhiza saccifera.
Although the population of O. laxiflora was rath-
er big, it is quite vulnerable to anthropogenic damage
or even destruction, because of its closeness to the pri-
vate lands and the paths of the herds of local farmers.
115. Orchis pinetorum Boiss. & Kotschy
Bu Znepole region: Mt Ruy, in a mixed deciduous
forest not very far from the road to Erma River
Gorge, on a west-facing slope with inclination
from 10 to 30 degrees, 691 m, 42°51'28.2"N,
22°38'44.556"E, with flowers and fruits,
31.05.2015, coll. A. Popatanasov (SO 107666).
This species was recently reported for this floristic
region (Asenov 2015), however, it is a new record for
Mt Ruy (cf. Assyov & Petrova 2012; Asenov 2015).
In the location, over thirty plants were found on an
area of over one ha, in a mixed deciduous forest not
very far from the road from Tran town to Erma River
Gorge; most of the shoots were generative. In the
same forest, a closely related taxon was also found,
O. ovalis (Delforge 2006); however here it flowers
approximately two weeks earlier so the chances for
interspecies cross-pollination are significantly re-
duced. The forest had a rather poorly developed sec-
ondary floor but was relatively rich in fungus spe-
cies. In the autumn, numerous fruit bodies of over
ten Basidiomycota species were found. This is in
conformity with the observed richness of myco-de-
pendent forestry orchids found on such a relatively
small area: nine orchid species.
Other species from this family found in this local-
ity were: Cephalantera damasonium, Dactylorhiza sac-
cifera, Epipactis helleborine, Orchis ovalis, O. purpurea,
Listera ovata, Platanthera chlorantha.
Report 116
Kit Tan
1
, Grzegorz Iszkuło
2
, Monika Dering
2
&
Emilia Pers-Kamczyc
2
1
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku dk (corresponding author)
2
Institute of Dendrology, Polish Academy of Sciences,
Parkowa 5, 62-035 Kórnik, Poland
Fabacaeae
116. Laburnum alpinum (Mill.) Bercht. & J. Presl
(Fig. 16)
Gr Nomos & Eparchia Karditsis: N of the village
Petrilo, steep eroded schistose slopes and ravine,
in openings of Fagus-Abies cephalonica forest,
20–30 trees and large shrubs 2–5 m tall, 1300 m,
Fig. 16. Laburnum alpinum habitat near Plastira Lake (photo G.
Iszkuło).

383Phytol. Balcan. 21(3) • Sofia • 2015
39°20'N, 21°37'E, 14.06.1997, Strid, Kit Tan &
Vold 43982 (ATH!G!LD! herb. Strid!); ca. 40 km
SW of Karditsa (near Plastira Lake), 1100 m,
39°12'N, 21°42'E, 11.06.2015, Iszkuło, Dering &
Pers-Kamczyc (KOR 51232!).
New for nomos and eparchia Karditsis in S Pindos.
The species is rare in Greece, first reported from no-
mos and eparchia Trikalon in S Pindos by Browicz &
Zieliński (1989: 85) and subsequently by others in the
same area, e.g., Sfikas & Sfikas (1995: 2). In the check-
list of Dimopoulos & al. (2013) Laburnum alpinum
was erroneously reported from northeastern Greece;
this was based on a collection by Pavlides (1976:
59) from nomos & eparchia Chalkidiki: Sithonia
Peninsula, Tripotamos; the record however, is of L.
anagyroides Medik. In Bulgaria the latter species is
planted in parks and gardens and along roads as an or-
namental where it often escapes, becoming fully natu-
ralized in both natural and disturbed habitats (Petrova
& al. 2013).
Reports 117–121
Alexander Tashev
1
& Nikolay Tashev
2
1
Department of Dendrology, Faculty of Forestry, University
of Forestry, 10 Kl. Ochridski Blvd., Sofia 1756, Bulgaria,
e-mail: atashev@mail.bg
2
University of Forestry, 10 Kl. Ochridski Blvd., Sofia 1756,
Bulgaria, e-mail: wanderernat@yahoo.com
Eucommiaceae
117. Eucommia ulmoides Oliv.
Bu Black Sea Coast (Southern): between
Chernomorets and Sozopol towns, along the
coastline on the territory of Gradina Camping
Site, on a sand substrate, among shrubbery
and Pinus nigra culture, along with Fraxinus
americana, etc., in close proximity to the sea
coast, on an east-facing slope with inclination
of 2°, 4 m, NG59, 42°24'51.2"N, 27°39'04.1"E,
20.09.2015, coll. A. Tashev & N. Tashev (SOM
171741, 171742; SO 107681).
This is a new adventive species for the Bulgarian flora,
originating from West and Central China. It is culti-
vated as a decorative tree in some regions of Bulgaria.
Fagaceae
118. Quercus rubra L.
Bu Black Sea Coast (Southern): between
Chernomorets and Sozopol towns, along the
coastline on the territory of Gradina Camping
Site, in a thinned-out spot among 30-year culture
of Pinus nigra, single individuals were found or
small groups of about 3–4 individuals reaching
3 m in height, the underbrush was 5–7 years old,
the terrain was level, with sandy soil cover, 2 m,
NG59, 42°24'24.7"N, 27°39'38.5"E, 26.08.2015,
coll. A. Tashev & N. Tashev (SOM 171755;
171756; 171757).
Along with Quercus rubra, there was usually found an
undergrowth of Q. frainetto, Q. cerris, Q. polycarpa, Q.
pubescens, Fraxinus oxycarpa, Prunus cerasifera, and
Pinus nigra. Among the identified shrubs were: Cornus
sanguinea, Vitis sylvestris, Ligustrum vulgare, Prunus
spinosa, Cionura erecta, Rosa canina, Rubus sp., and
of the grasses – Calamagrostis epigeios, Althaea can-
nabina, Agrimonia eupatoria, Hypericum perforatum,
Plantago lanceolata, Cychorium inthybus, etc.
This is a new find of this adventive North
American species in the Bulgarian flora, so far report-
ed from Vitosha region and Tundzha Hilly Country
(Tashev & al. 2013; Vladimirov 2013).
Lauraceae
119. Laurus nobilis L.
Bu Black Sea Coast (Southern): between
Chernomorets and Sozopol towns, along the
coastline on the territory of Gradina Camping
Site, several scores of individuals were found
under the canopy of Robinia pseudoacacia on a
sandy substrate, with northeastern exposition
and an incline of 4°, 25 m, NG59, 42°26'13.0"N,
27°38'41.9"E, 08.08.2015, coll. A. Tashev &
N. Tashev (SOM 171750); with northeastern
exposition and an incline of 2°, 23 m, NG59,
42°26'09.9"N, 27°38'43.6"E, 08.08.2015, coll. A.
Tashev & N. Tashev (SOM 171751); with eastern
exposition and an incline of 3°, 30 m, NG59,
42°26'18.1"N, 27°38'42.5"E, 08.08.2015, coll. A.
Tashev & N. Tashev (SOM 171752); with eastern
exposition and an incline of 4°, 26 m, NG59,
42°26'19.7"N, 27°38'42.4"E, 08.08.2015, coll. A.
Tashev & N. Tashev (SOM 171753).
Several scores of individuals reaching 50–60 cm in
height were randomly scattered under the canopy of
about 40-year old culture of Robinia pseudoacacia,
along with Prunus cerasifera, P. avium, Acer campes-
tre, A. pseudoplatanus, A. negundo, Ailanthus altissi-
ma, Ulmus minor, Fraxinus oxycarpa, Quercus cer-

384 Vladimirov, V. & al • New floristic records in the Balkans: 28
ris, Juglans regia, and Ficus carica. The identified
shrubs were: Rubus caesius, Hedera helix, Clematis
vitalba, Smilax exelsa, and Euonimus europaeus.
Myrrhoides nodosa dominated among the grasses,
but also occurred Tamus communis, Galium aparine,
Brachypodium pinnatum, Geum urbanum, Sambucus
ebulus, Buglossoides purpurocoerulea, Arum macula-
tum, Stellaria media, Allium sp., etc.
This is the first report of spontaneous develop-
ment of this Mediterranean species in Bulgaria, its
seeds probably being carried out by birds from the
yards of the nearby towns, where it is cultivated as a
decorative species. The spontaneous development of
this warm-loving species suggests an actual warmup
of the climate in the last years, which permits its sur-
vival during the cold winter periods of the year.
Oleaceae
120. Fraxinus americana L.
Bu Black Sea Coast (Southern): between
Chernomorets and Sozopol towns, along the
coastline on the territory of Gradina Camping
Site, on a snady substrate, among shrubs and a
culture of Pinus nigra, in close proximity to the
coastline, on an east-facing slope with inclination
of 2°, 4 m, NG59, 42°24'51.2"N, 27°39'04.1"E,
20.09.2015, coll. A. Tashev & N. Tashev (SOM
171765; SO 107680).
This is a new locality of this adventive species for
Bulgaria, so far reported from the Danube Plain and
Sofia region. The population was numerous in the area
between Chernomorets and Sozopol (Cheshmedzhiev
2011: 282; Assyov & Petrova 2012: 197).
Simaroubaceae
121. Ailanthus altissima (Mill.) Swingle
Bu Valley of River Mesta: in the lands of Godeshevo
village, along the motorway to Gotse Delchev
town, in a lay-by with a water fountain, in
the upper part of a slope with west-northwest
exposition and inclination of 10°, 792 m, KF59,
41°29'16.8"N, 24°03'00.1"E, 06.09.2015, coll. A.
Tashev (SOM 171747; SO 107685); in the lands
of Slashten village, along the motorway to Gotse
Delchev town, along with Robinia pseudoacacia,
Ulmus minor and Prunus spinosa on a south-
southwest facing slope with inclination of
30°, 677 m, KF59, 41°30'13.7"N, 24°00'33.7"E,
06.09.2015, coll. A. Tashev (SOM 171745, 171746;
SO 107684).
This find reconfirms distribution of this species in this
floristic region (Assyov & Petrova 2012: 51; Petrova &
al. 2013: 41, 43).
Reports 122–126
Ioannis Tsialtas
1
& Kit Tan
2
1
Aristotle University of Thessaloniki, Faculty of Agriculture,
Lab. of Agronomy, 541 24 Thessaloniki, Greece
2
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku.dk (corresponding author)
Cucurbitaceae
122. Sicyos angulatus L.
Gr Nomos & Eparchia Kavalas: Krinides, in a maize
field at Tenagi Filippon, 45 m, 40°58'Ν, 24°15'Ε,
09.10.2015, Tsialtas obs. (photos; det. Kit Tan,
October 2015).
This was first reported for Greece (in nomos Evrou)
by Giannopolitis (2003). It is now noted in moderate
numbers along the drainage channels in the Tenagi
Filippon region. According to local growers S. angu-
latus, originating from N America, is one of the most
harmful weeds of corn (Zea mays) along with other
climbing species of Ipomoea. It evades the action of
chemical herbicides due to the high organic content of
the soils and also by its late emergence.
Fabaceae
123. Aeschynomene rudis Benth. (Fig. 17)
Gr Nomos & Eparchia Serron: rice field at Nea
Tyroloi overtaken by Aeschynomene plants 2 m
tall, ca. 30 m, 41°06'N, 23°22'E, 04.09.2015,
Tsialtas obs.; rice field and margins, near raised
banks of Belitsa river, ca. 35 m, 41°06'N, 23°21'E,
04.09.2015, Tsialtas obs.; at edge of rice fields to
the right of the Vamvakia to Anagennisi road, ca.
32 m, 41°05'N, 23°20'E, 11.09.2015, Tsialtas obs.
(photos; det. Kit Tan, September 2015).
First report for Greece. Aeschynomene indica was re-
ported by us from the same area (Tsialtas & Tan 2014)
but we were not aware of the existence of the rather sim-
ilar A. rudis which is native to Mexico and S America.
Both species are distinguished locally by their vernac-
ular names ‘little-tree’ and ‘lentil-tree’, and are well-es-
tablished weeds in rice fields together with its rotation
crop maize. According to growers, both Aeschynomene
species were inadvertently introduced some 10 to 15
years ago, possibly as a contaminant of crop seed. The

385Phytol. Balcan. 21(3) • Sofia • 2015
species are easily controlled using chemical herbicides
but without, or with unsuccessful, herbicide applica-
tion, Aeschynomene takes over and dominates instead
of the cultivated crop. The species grow intermixed in
wet places at the edge of rice fields and raised banks
of drainage channels together with Sorghum halepense,
Cynodon dactylon, Echinochloa crus-galli, Persicaria
lapathifolia and Phragmites australis. However,
Aeschynomene rudis seems more tolerant of powdery
mildew infection whereas A. indica usually succumbs.
Solanaceae
124. Datura stramonium f. tatula (L.) Geerinck &
Walravens
Gr Nomos Lesvou, Eparchia Limnou: Tsimandria to
Diapori, ca. 7 m, 39°51'N, 25°10'E, 24.08.2015,
Tsialtas obs. (photos; det. Kit Tan, September
2015).
New for nomos, eparchia and island of Limnos, not
reported by Baliousis (2014). Numerous flowering
plants in field planted with Vigna unguiculata (black-
eyed bean), causing a serious weed problem togeth-
er with Amaranthus retroflexus, Portulaca oleracea,
Sorghum halepense and Xanthium spinosum. Less
abundant weeds are Amaranthus albus, Convolvulus
arvensis and Heliotropium europaeum.
This purple-flowered variant has been observed in
many localities in north, south and central Greece, of-
ten co-existing with the more com-
mon white-flowered form. In field
notes it is usually presented as D.
stramonium without formal recog-
nition of the variant. The two forms
differ only by a single pair of genes
responsible for the flower colour.
125. Solanum luteum Mill.
Gr Nomos & Eparchia Kavalas:
Krinides, in a field sown to Glycine
max (soyabean), ca. 65 m, 41°00'Ν,
24°19'Ε, 09.10.2015, Tsialtas obs.
(photos; det. Kit Tan, October 2015).
New for eparchia. Several plants
were found together with the
more abundant Solanum nigrum,
Abutilon theophrasti, Sorghum ha-
lepense, Xanthium strumarium and
Cheno podium album.
Poaceae
126. Panicum capillare L. (Fig. 18)
Gr Nomos & Eparchia Kozanis: the former drained
Lake Sari Giol, in a field sown to Phaseolus
communis (dwarf dry bean), 655 m, 40°23'N,
21°51'E, 11.08.2015, Tsialtas obs. (photos; det. Kit
Tan, August 2015).
New for nomos and eparchia. A few plants were not-
ed in the bean field with a rich weed flora dominat-
ed by Sorghum halepense, Abutilon theophrasti and
Xanthium strumarium. According to local growers,
the species, originating from N America, is common
in shady places and on the banks of drainage chan-
nels.
Fig. 17. Aeschynomene rudis (photo I. Tsialtas).
Fig. 18. Panicum capillare (photo I. Tsialtas).

386 Vladimirov, V. & al • New floristic records in the Balkans: 28
Reports 127–136
Kiril Vassilev & Hristo Pedashenko
Department of Plant and Fungal Diversity and Resources,
Institute of Biodiversity and Ecosystem Research,
Bulgarian Academy of Sciences, Acad. Georgi Bonchev
St., bl. 23, 1113 Sofia, Bulgaria, e-mail: kiril5914@abv.bg,
hristo_pedashenko@yahoo.com
Poaceae
127. Agrostis stolonifera L.
Bu Rhodopi Mts (Central): in grasslands between
Shiroka Laka village and Smolyan town, LG01,
41.61911°N, 24.66191°E, 09.08.2015, coll. K.
Vassilev (SOM 171566).
This is a new species for this floristic region found
in mesophytic grasslands of the class Molinio-
Arrhenatherateae. It grows together with Festuca prat-
ensis, F. r ubra, Cynosurus cristatus, Achillea mile-
folium, Trifolium pratense, T. repens, T. campestre,
Lathyrus pratensis, Dactylis glomerata, etc.
128. Anthoxanthum aristatum Boiss.
Bu Rhodopi Mts (Central): in dry grasslands near
Zmeitsa village, KG71, 41.65034°N, 24.22931°E,
07.08.2015, coll. K. Vassilev & H. Pedashenko
(SOM 171563).
This is a Euro-Asiatic species distributed in the south-
ern parts of Bulgaria, which has not been reported so
far for this floristic region. Its distribution in the coun-
try was probably wider and potential floristic regions
of its possible finding are Mt Sredna Gora (Eastern),
Mt Slavyanka, Pirin Mts (Southern), and the Valley of
River Mesta.
129. Bromus moesiacus Velen.
Bu Valley of River Struma (Southern): Kresna Gorge,
under Oshtava village, FM82, 31.07.2007, coll. D.
Dimitrov (SOM 163781).
— Rila Mts: Rila Monastery Forest Reserve, in
xero-mesophytic grasslands, FM97, 42.14096°N,
23.34269°E, 14.06.2015, coll. K. Vassilev (SOM
171568, 178570).
This is a new species for these floristic regions. It is a
Bulgarian endemic known from eight floristic regions
in the country, according to Assyov & Petrova (2012).
On the territory of the Rila Monastery Forest Reserve,
it appears in grasslands in the beech belt as subdom-
inant species, together with Festuca callieri, Koeleria
nitidula and Stipa balcanica. This vegetation is close
to the class Festuco-Brometea, but a future analysis will
show their syntaxonomical position.
130. Elymus elongatus (Host) Runemark
Bu Balkan Range (Western): Mt Mala Planina, in
xerothermic grasslands near Gradets village,
FN75, 42.89149°N, 23.18901°E, 03.08.2015, coll.
K. Vassilev & H. Pedashenko (SOM 171573).
The species is locally distributed in an area covering
only several square meters. It is a new species for this
floristic region.
131. Festuca callieri (St-Yves) Markgr.-Dann.
Bu Rhodopi Mts (Central): in dry grasslands near
Trigrad village, KG80, 41.60907°N, 24.38181°E,
08.08.2015, coll. K. Vassilev & H. Pedashenko
(SOM 171546).
— Mt Sredna Gora (Western): in dry grasslands
between Strelcha and Koprivshtitsa towns, KH81,
42.58847°N, 24.37037°E, 13.08.2015, coll. K.
Vassilev & H. Pedashenko (SOM 171548).
This is a new species for these floristic regions. It is
known from six floristic regions in the Bulgaria, ac-
cording to Assyov & Petrova (2012). Its communities
are locally distributed in rocky grasslands but their
syntaxonomical position is still unclear. They are rich
in different xerothermic species such as Trifolium ar-
vense, Thymus callieri, Rhodax canus, Plantago subu-
lata, etc.
132. Festuca rupicola Heu.
Bu Rhodopi Mts (Western): in dry grasslands
between Velingrad and Sarnitsa towns, KG53,
41.97276°N, 23.95003°E, 05.08.2015, coll. K.
Vassilev & H. Pedashenko (SOM 171562).
A new species for this floristic region. It is a wide-
spread grass in the lowlands and mountainous regions
of Bulgaria. In the study region, it forms communities
together with Chrysopogon gryllus, Asperula depres-
sa, A. purpurea, Koeleria nitidula, Scabiosa triniifolia,
Teucrium chamaedrys, Phleum phleoides and should
be referred to class Festuco-Brometea.
133. Lolium multiorum Lam.
Bu Valley of River Struma (Southern): Kresna Gorge,
above Peyo Yavorov Railway Station, 31.07.2007,
coll. D. Dimitrov (SOM 165837).
— Rhodopi Mts (Western): in dry grasslands
between Dorkovo and Kostandovo villages,
KG65 42.03406°N, 24.12135°E, 05.08.2015, coll.
K. Vassilev & H. Pedashenko (SOM 171565).
This species has not been reported for these floristic
regions before. It is widely distributed in pastures as a
companion species.

387Phytol. Balcan. 21(3) • Sofia • 2015
134. Poa molinerii Balb.
Bu Balkan Range (Central): in grasslands on NE
slope of peak Malka Ambaritsa, 1890 m, LH13,
15.07.1965, coll. H. Kochev (SOM 113043).
— Mt Slavyanka: Mt Slavyanka, GL18, 31.07.2008,
coll. D. Dimitrov (SOM 169484); in calcareous
grasslands around peak Gotshev, 2212 m, GL18,
26.08.2005, coll. D. Dimitrov (SOM 162402).
— Rhodopi Mts (Central): in dry grasslands between
Gyovren and Trigrad villages, KG81, 41.61423°N,
24.38059°E, 08.08.2015, coll. K. Vassilev & H.
Pedashenko (SOM 171552); on calcareous rocks
in Durdaga locality near Trigrad village, KG80,
02.08.2011, coll. D. Dimitrov (SOM 167664); S of
peak Chaeva Chuka above Mugla village, KG91,
03.08.2011, coll. D. Dimitrov (SOM 167665)
This species is distributed in the mountainous areas of
Bulgaria in rocky places. It is a new species for these
floristic regions.
135. Poa timoleontis Boiss.
Bu Vitosha region: in grasslands near Dolni
Okol village, GN00, 42.48759°N, 23.50128°E,
07.07.2015, coll. K. Vassilev & H. Pedashenko
(SOM 171575); Mt Lyulin, on rocks under
the peak below Tabakov's property, FN72,
19.05.1935, coll. B. Achtaroff (SOM 5559).
— Rhodopi Mts (Western): in rocky grasslands near
Peshtera town, 450 m, KG75, 17.06.1926, coll. B.
Davidoff (SOM 5767).
— Thracian Lowland: between Sladun and Varnik
villages, MG53, 14.05.2011, coll. D. Dimitrov
(SOM 168084); on rocks near Plovdiv town,
LG16, 06.1984, coll. V. Stribrny (SOM 5759); in
grasslands near Hisarya town, Plovdiv district,
LH10, 06.05.1927, coll. K. Urumov (SOM 5756)
— Tundzha Hilly Country: in grasslands near Aitos
town, NH22, 09.05.1915, coll. I. Mrkvička & B.
Davidoff (SOM 5804, 5806).
According to Assyov & Petrova (2012), the species is
known from seven floristic regions.
136. Vulpia muralis (Kunth) Nees
Bu Vitosha region: Mt Lozenska, in meadows of
Vsred Rakita locality near Dolni Lozen village,
GN01, 01.07.1954, coll. I. Ganchev (SOM 40295);
Mt Lozenska, in dry meadows of Rusama locality
near Dolni Pasarel village, GN01, 26.06.1953,
coll. I. Ganchev (SOM 40296).
— Rhodopi Mts (Central): in dry grasslands near
Zmeitsa village, KG71, 41.65034°N, 24.22931°E,
07.08.2015, coll. K. Vassilev & H. Pedashenko
(SOM 171564).
It is a new species for these floristic regions.
Reports 137–238
George Zarkos
1
, Vasilis Christodoulou
2
, Kit Tan
3
&
Gert Vold
4
1
Kolokotroni 37A, Kiato, 202 00, Korinthias, Greece
2
Apellou sidestreet, Kiato, 202 00, Korinthias, Greece
3
Institute of Biology, University of Copenhagen, Øster
Farimagsgade 2D, DK-1353 Copenhagen K, Denmark,
e-mail: kitt@bio.ku.dk (corresponding author)
4
Botanical Garden, State Natural History Museum, Øster
Farimagsgade 2C, DK-1353 Copenhagen K, Denmark
The following are new plant records based on floristic
investigations in the prefecture of Korinthias in north
central Peloponnese. The list also includes 88 new re-
cords for Ntourntouvana (Dourndouvana), a moun-
tain range bordering Nomos Achaias and Nomos Ko-
rinthias. This natural border lies between Mt Chelmos
and Mt Saitas in nomos Achaias/Korinthias, Epar-
chia Kalavriton/Korinthias and is considered a south-
ern extension of Chelmos. The range is distinguished
by its steep NW-facing slopes; there are two summits,
Ntourntouvana (2078 m) and Triadafilia (2109 m).
Apiaceae
137. Bupleurum glumaceum Sm.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31960.
138. Carum graecum Boiss. & Heldr.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1600 m, 37°55'N,
22°14'E, 09.07.2015, Zarkos obs.; NE slopes of
Ntourntouvana, 5 km from Panorama along
forest road to the summit, 1320–1350 m, 37°55'N,
22°15'E, 12.08.2015, Kit Tan & G. Vold 31974 (in
fruit).
139. Carum meioides (Griseb.) Halácsy
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits Ntourntouvana and Triadafilia, 1956 m,
37°54'N, 22°14'E, 09.07.2015, Zarkos obs.

388 Vladimirov, V. & al • New floristic records in the Balkans: 28
140. Laserpitium pseudomeum Orph., Heldr. &
Sartori
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, 1956 m, 37°54'N, 22°14'E, 09.07.2015,
Zarkos obs.
Endemic to Greece (Peloponnese and Sterea Ellas).
141. Physospermum cornubiense (L.) DC.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31970.
142. Sanicula europaea L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies
cephalonica forest on N slopes, 1317 m, 37°55'N,
22°15'E, 24.05.2015, Zarkos & Christodoulou obs.
143. Selinum silaifolium (Jacq.) Beck
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31969.
144. Seseli aroanicum Hartvig
Gr Nomos & Eparchia Korinthias: Mt Killini,
crevices of vertical limestone cliff, 1216 m,
37°58'N, 22°21'E, 27.08.2015, Zarkos obs. (several
photos: det. Kit Tan, August 2015); 1615 m,
37°53'N, 22°21'E, 30.08.2015, Zarkos obs.; 1375–
1402 m, 37°54'N, 22°21'E, 30.08.2015, Zarkos s.n.
New for Mt Killini, nomos and eparchia Korinthias;
previously recorded from Mts Chelmos (locus classi-
cus), Erimanthos and Kallifoni. Endemic to Greece
(Peloponnese). A thousand plants were noted but on-
ly two were flowering. On Mt Erimanthos, the plants
collected (Strid, Kit Tan & Vold 44800) are taller and
with longer ultimate leaf segments than in the holo-
type from Chelmos; this tall habit was also noted in
the plants from Killini.
Asteraceae
145. Achillea umbellata Sm.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies
cephalonica forest on NW slopes, 1692 m,
37°55'N, 22°14'E, 09.07.2015, Zarkos obs.
Endemic to Greece (Peloponnese, Sterea Ellas and Evvia).
146. Bidens tripartitus L.
Gr Nomos & Eparchia Korinthias: Xylokastro,
coastal pine forest, 7 m, 38°04'N, 22°39'E,
23.09.2015, Zarkos obs.
New for nomos and eparchia. In Pinus halepensis and
Pinus pinea woodland.
147. Centaurea athoa subsp. chelmea Kit Tan,
Zarkos, V. Christodoulou & G. Vold, subsp. nov.
(Fig. 19)
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana,
stony ground along forest road in open Abies
cephalonica woodland, 1125 m, 37°55'N, 22°16'E,
24.05.2015, Zarkos & Christodoulou obs.; loc.
ibid., 12.08.2015, Kit Tan & G. Vold 31936
(holotype C; isotype ATH).
Endemic to Greece (Peloponnese). Differing from C.
athoa DC. subsp. athoa (Mt Athos in NE Greece, W &
S Anatolia) and C. athoa subsp. parnonia (Halácsy) E.
Gamal-Eldin & Wagenitz (Mts Parnon and Taigetos
in S Peloponnese) by its basal leaves with 6–11 pairs
of narrowly linear-lanceolate and less densely arach-
noid lateral segments (see Fig. 19). The achenes are
5–6 mm long, in C. athoa subsp. athoa they were ob-
served to be 4.5–5 mm. Centaurea athoa subsp. par-
nonia has shorter and very densely arachnoid-lanate
basal leaves.
148. Centaurea grisebachii (Nyman) Heldr.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31946.
Fig. 19. Centaurea athoa subsp. chelmea, basal leaves and capitu-
lum (photos G. Zarkos).

389Phytol. Balcan. 21(3) • Sofia • 2015
149. Centaurea zuccariniana DC.
Gr Nomos & Eparchia Korinthias: Mt Killini,
1470 m, 37°58'N, 22°22'E, 06.07.2015, Zarkos obs.
New for Mt Killini. In Korinthias, recorded from
Stymfalia (Zarkos & al. 2011) and near Sarandapicho.
150. Echinops ritro subsp. ruthenicus (M. Bieb.)
Nyman
Gr Nomos & Eparchia Korinthias: Mougosto forest,
835 m, 37°59'N, 22°36'E, 13.08.2015, Zarkos s.n.
(det. Kit Tan, September 2015).
New for nomos and eparchia Korinthias. The plants
were 20–40 cm tall. Rarely documented at subspecies
rank, this is apparently the third confirmed record
from the Peloponnese. Echinops ritro subsp. ritro oc-
curs on mainland Greece.
151. Echinops ritro subsp. sartorianus (Boiss. &
Heldr.) Kožuharov
Gr Nomos & Eparchia Korinthias: Mt Killini, along
road to the peak Simio,1590 m, 37°56'N, 22°25'E,
12.08.2015, Zarkos s.n. (det. Kit Tan, September
2015).
New for Mt Killini. Collected in Korinthias from
the monastery of Ag. Giorgos, Feneos by Maire &
Petitmengin in August 1906, more than a century ago
(Maire & Petitmengin 1908: 120). The plants are tall-
er than in E. r. subsp. ruthenicus being 80–100 cm in
height, with pale blue capitula, unbranched or spar-
ingly branched stems and less divided leaves. Endemic
to Greece (Peloponnese and Sterea Ellas).
152. Lactuca muralis (L.) Gaertn.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31971.
153. Tanacetum balsamita L. (Fig. 20)
Gr Nomos & Eparchia Korinthias: village of
Sarandapicho,1227m, 38°01'N, 22°22'E,
16.08.2015, Zarkos s.n.
New for the Peloponnese, and second record from
Greece. It was first documented by Elli Stamatiadou
who found it in open Fagus forest near the village of
Hania on Mt Pilion (East Central Greece). Naturalized.
Boraginaceae
154. Buglossoides incrassata (Guss.) I.M. Johnst.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between summits,
rocky limestone slopes and scree, 2030 m, 37°54'N,
22°14'E, 24.05.2015, Zarkos & Christodoulou obs.
155. Myosotis refracta Boiss. subsp. refracta
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1995 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
156. Myosotis sylvatica subsp. cyanea (Hayek)
Vestergren
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1995 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
157. Onosma erecta subsp. malickyi Teppner
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31964.
Endemic to Greece (Peloponnese).
Brassicaceae
158. Aethionema saxatile (L.) R. Br.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, along
path to summit, in Abies cephalonica and
Juniperus communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.; NE slopes of Ntourntouvana,
5 km from Panorama along forest road to
the summit, 1320–1350 m, 37°55'N, 22°15'E,
12.08.2015, Kit Tan & G. Vold obs.
Fig. 20. Tanacetum balsamita (photo G. Zarkos).

390 Vladimirov, V. & al • New floristic records in the Balkans: 28
159. Arabis alpina L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, path
to summit, in Abies cephalonica and Juniperus
communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
160. Aubrieta deltoidea (L.) DC.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, path
to summit, in Abies cephalonica and Juniperus
communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
161. Draba lacaitae Boiss.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1951 m, 37°54'N, 22°14'E, 09.07.2015, Zarkos obs.
162. Draba laconica Stevanović & Kit Tan
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1620 m, 37°55'N,
22°14'E, 24.05.2015, Zarkos & Christodoulou obs.
Endemic to Greece (Peloponnese).
163. Draba lasiocarpa Rochel subsp. lasiocarpa
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, path
to summit, in Abies cephalonica and Juniperus
communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
164. Malcolmia graeca subsp. bicolor (Boiss. &
Heldr.) Stork
Gr Nomos Achaias/Korinthias, Eparchia
Kalavriton/Korinthias: Ntourntouvana, summit
of Ntourntouvana, rocky limestone slopes,
2051 m, 37°54'N, 22°15'E, 24.05.2015, Zarkos &
Christodoulou obs.
Endemic to Greece.
165. laspi graecum Jord.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, path
to summit, in Abies cephalonica and Juniperus
communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.; NE slopes of Ntourntouvana,
5 km from Panorama along forest road to
the summit, 1320–1350 m, 37°55'N, 22°15'E,
12.08.2015, Kit Tan & G. Vold obs.
Endemic to Greece (Peloponnese and Sterea Ellas).
Campanulaceae
166. Campanula aizoon subsp. aizoides (Greuter)
Fedorov
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between summits,
rocky limestone slopes and scree, 1995 m, 37°54'N,
22°14'E, 24.05.2015, Zarkos & Christodoulou obs.
Endemic to Greece (Peloponnese).
167. Campanula spatulata subsp. spruneriana
(Hampe) Hayek
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1956 m, 37°54'N, 22°14'E, 09.07.2015, Zarkos obs.
Caryophyllaceae
168. Arenaria cretica Spreng.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1790 m, 37°54'N,
22°14'E, 09.07.2015, Zarkos obs.
169. Bolanthus graecus (Schreb.) Barkoudah
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: summit of Ntourntouvana, limestone
scree, 2051 m, 37°54'N, 22°15'E, 24.07.2015,
Zarkos s.n. (det. Kit Tan, September 2015).
New for Peloponnese, endemic to Greece (Sterea Ellas,
Kiklades, E Aegean and W Aegean). Petals white with
a transverse purple stripe.
170. Cerastium candidissimum Correns
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest
on steep NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
Endemic to Greece.
171. Herniaria hirsuta L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31945.
172. Minuartia verna (L.) Hiern
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between

391Phytol. Balcan. 21(3) • Sofia • 2015
summits, rocky limestone slopes and scree,
2080 m, 37°54'N, 22°14'E, 09.07.2015, Zarkos obs.
173. Silene caesia Sm.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, path
to summit, in Abies cephalonica and Juniperus
communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
174. Silene multicaulis Guss. subsp. multicaulis
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest
on steep NW slopes, 1544 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
175. Silene parnassica Boiss. & Spruner
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest
on steep NW slopes, 1834 m, 37°54'N, 22°14'E,
09.07.2015, Zarkos obs.
176. Silene radicosa Boiss. & Heldr.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1978 m, 37°54'N, 22°15'E, 24.07.2015, Zarkos obs.
Crassulaceae
177. Sedum dasyphyllum L. subsp. dasyphyllum
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1790 m, 37°54'N,
22°14'E, 09.07.2015, Zarkos obs.
178. Sedum hispanicum L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest
on steep NW slopes, 1682 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
179. Sedum magellense Ten. subsp. magellense
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1790 m, 37°54'N,
22°14'E, 09.07.2015, Zarkos obs.
Dipsacaceae
180. Cephalaria flava subsp. setulifera (Boiss. &
Heldr.) Kokkini
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31965.
Endemic to Greece (Peloponnese, Sterea Ellas and Evvia).
181. Pterocephalus perennis Coult. subsp. perennis
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between summits,
rocky limestone slopes and scree, 1984 m, 37°54'N,
22°15'E, 24.07.2015, Zarkos obs.; NE slopes of
Ntourntouvana, 5 km from Panorama along
forest road to the summit, 1320–1350 m, 37°55'N,
22°15'E, 12.08.2015, Kit Tan & G. Vold 31961.
Endemic to Greece.
182. Scabiosa ochroleuca L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31963.
Euphorbiaceae
183. Euphorbia herniariifolia Willd.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between summits,
rocky limestone slopes and scree, 2030 m, 37°54'N,
22°14'E, 24.05.2015, Zarkos & Christodoulou obs.
184. Euphorbia hyssopifolia L. (Fig. 21)
Gr Nomos & Eparchia Korinthias: Xylokastro,
Fig. 21. Euphorbia hyssopifolia (photo G. Zarkos).

392 Vladimirov, V. & al • New floristic records in the Balkans: 28
coastal pine forest, 7 m, 38°04'N, 22°39'E,
23.09.2015, Zarkos obs. (photos; confirmed Kit
Tan, October 2015).
New for Greece. Under Pinus halepensis and P pinea
together with Asparagus aphyllus, Bidens tripartitus,
Euphorbia peplis, Tamarix smyrnensis and Rhamnus
alaternus.
185. Euphorbia peplis L.
Gr Nomos & Eparchia Korinthias: Xylokastro,
coastal pine forest, 7 m, 38°04'N, 22°39'E,
23.09.2015, Zarkos obs.
New for nomos and eparchia. In Pinus halepensis and
Pinus pinea woodland.
Fabaceae
186. Astragalus depressus L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
2030 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
187. Ononis pusilla L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1600 m, 37°55'N,
22°14'E, 09.07.2015, Zarkos obs.
188. Tripodion graecum (Boiss.) Lassen (Fig. 22)
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
Occurring only in Peloponnese and S Anatolia.
189. Vicia cracca L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes,
forest road in Abies cephalonica woodland,
1317 m, 37°55'N, 22°15'E, 24.05.2015, Zarkos &
Christodoulou obs.
Geraniaceae
190. Geranium rotundifolium L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31975.
Globulariaceae
191. Globularia stygia Boiss. (Fig. 23)
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
2030 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Endemic to Greece (Peloponnese).
Grossulariaceae
192. Ribes uva-crispa subsp. austro-europaeum
(Bornm.) Bech.
Gr Nomos Achaias/Korinthias, Eparchia
Kalavriton/Korinthias: Ntourntouvana, steep
Fig. 22. Tripodion graecum (photo G. Zarkos). Fig. 23. Globularia stygia (photo G. Zarkos).

393Phytol. Balcan. 21(3) • Sofia • 2015
NW slopes, limestone rock and scree, 1600 m,
37°55'N, 22°14'E, 09.07.2015, Zarkos obs.;
Ntourntouvana, NE slopes, along forest road
in open Abies cephalonica woodland, 1120–
1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit Tan &
G. Vold obs.
Hypericaceae
193. Hypericum perforatum L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31975.
Lamiaceae
194. Acinos alpinus subsp. meridionalis (Nyman)
P.W. Ball
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
195. Marrubium velutinum subsp. cylleneum (Boiss.
& Heldr.) Nyman
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
Endemic to Greece (Peloponnese, Sterea Ellas and S
Pindos).
196. Melittis melissophyllum L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31968.
197. Micromeria juliana (L.) Rchb.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31972.
198. Origanum vulgare subsp. hirtum (Link) A.
Terracc.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31941.
199. Phlomis samia L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31948.
200. Scutellaria rupestris Boiss. & Heldr. subsp.
rupestris
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1980 m, 37°54'N, 22°15'E, 24.07.2015, Zarkos
obs.; Ntourntouvana, NE slopes, along forest
road in open Abies cephalonica woodland, 1120–
1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit Tan &
G. Vold 31947.
Endemic to Greece (Peloponnese).
201. Sideritis clandestina subsp. peloponnesiaca
(Boiss. & Heldr.) Baden
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, steep NW slopes,
limestone rock and scree, 1600 m, 37°55'N,
22°14'E, 09.07.2015, Zarkos obs.; NE slopes of
Ntourntouvana, 5 km from Panorama along
forest road to the summit, 1320–1350 m, 37°55'N,
22°15'E, 12.08.2015, Kit Tan & G. Vold 31966.
Endemic to Greece (Peloponnese).
202. Teucrium montanum L. subsp. montanum
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between summits,
rocky limestone slopes and scree, 1980 m, 37°54'N,
22°15'E, 24.07.2015, Zarkos obs.; NE slopes of
Ntourntouvana, 5 km from Panorama along
forest road to the summit, 1320–1350 m, 37°55'N,
22°15'E, 12.08.2015, Kit Tan & G. Vold 31962.
203. ymus leucotrichus Halácsy subsp.
leucotrichus
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
204. ymus longicaulis subsp. chaubardii (Rchb. f.)
Jalas
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.

394 Vladimirov, V. & al • New floristic records in the Balkans: 28
Orobanchaceae
205. Odontites linkii Boiss.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31949.
Endemic to Greece.
Plantaginaceae
206. Plantago holosteum Scop.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31944.
Plumbaginaceae
207. Armeria canescens (Host) Boiss.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
2030 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Polygalaceae
208. Polygala alpestris Rchb.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1956 m, 37°54'N, 22°14'E, 09.07.2015, Zarkos obs.
209. Polygala rausiana U. Raabe & al. (Fig. 24)
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, E slopes, edge of
forest path, 1135 m, 37°54'N, 22°16'E, 14.07.2015,
Zarkos obs.
Endemic to Greece (Peloponnese), southernmost oc-
currence of species.
Rosaceae
210. Potentilla speciosa Willd.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: summit of Ntourntouvana, limestone
scree, 2051 m, 37°54'N, 22°15'E, 24.05.2015,
Zarkos & Christodoulou obs.
Rubiaceae
211. Asperula laevigata L.
Gr Nomos & Eparchia Korinthias: edge of Lake Doxa,
885 m, 37°55'N, 22°17'E, 19.06.2015, Zarkos obs.
(photos); loc. ibid., Kit Tan & G. Vold obs.
New for nomos and eparchia Korinthias.
212. Asperula oetaea (Boiss.) Halácsy
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1835 m, 37°54'N, 22°14'E,
09.07.2015, Zarkos obs.; ridge between summits,
rocky limestone slopes and scree, 1956 m,
37°54'N, 22°14'E, 09.07.2015, Zarkos obs.
Endemic to Greece (Peloponnese, Sterea Ellas and S
Pindos).
213. Galium rotundifolium L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1545 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
214. Galium thymifolium Boiss. & Heldr.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1956 m, 37°54'N, 22°14'E, 09.07.2015, Zarkos obs.
Fig. 24. Polygala rausiana (photo G. Zarkos).

395Phytol. Balcan. 21(3) • Sofia • 2015
Endemic to Greece (Peloponnese, Sterea Ellas and
Evvia).
215. Galium verticillatum Danthoine
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1896 m, 37°54'N, 22°14'E, 24.07.2015, Zarkos obs.
Santalaceae
216. Viscum album subsp. abietis (Wiesb.) K. Malý
Gr Nomos & Eparchia Korinthias: east of
Sarandapicho village, 1403 m, 38°01'N, 22°23'E,
03.11.2015, Zarkos obs. (photos; confirmed Kit
Tan, November 2015); loc. ibid., 01.12.2015, Kit
Tan & G. Vold s.n.
From Korinthias, there are a few reports from the foot-
hills of Mt Killini. Observed parasitizing Abies cepha-
lonica on both sides of the road.
217. Viscum album L. subsp. album (Fig. 25)
Gr Nomos & Eparchia Korinthias: east of
Sarandapicho village, 1369 m, 38°01'N,
22°23'E, 03.11.2015, Zarkos obs. (photos; host
confirmed by Kit Tan, November 2015); loc. ibid.,
01.12.2015, Kit Tan & G. Vold 32028
New for nomos and eparchia. Parasitic on the inva-
sive Robinia pseudoacacia, which is a new host report
for Greece.
218. Viscum album subsp. austriacum (Wiesb.) Vollm.
Gr Nomos & Eparchia Korinthias: along road east of
Sarandapicho village, 1403 m, 38°01'N, 22°23'E,
03.11.2015, Zarkos obs. (photos; confirmed Kit
Tan, November 2015); loc. ibid., 01.12.2015, Kit
Tan & G. Vold 32026.
New for nomos and eparchia. On Pinus nigra, also
on both sides of the road. Thus in the same area in
Korinthias, all three subspecies of Viscum album were
found. It is possible Maire & Petitmengin (1908: 101)
had already collected Viscum album subsp. austria-
cum from Korinthias (from Flabouritsa, Mt Killini)
more than a century ago, describing the plant as “sur
les branches du Pinus laricio [P. n i g r a ]”. Because only
V. album subsp. abietis had been later noted in Abies
cephalonica forest at Flabouritsa, this report had not
received the attention it deserved or perhaps it was
considered as referring to V. album subsp. abietis al-
though the host was Pinus nigra.
Saxifragaceae
219. Saxifraga scardica Griseb.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1995 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Scrophulariaceae
220. Verbascum epixanthinum Boiss. & Heldr.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
Endemic to Greece.
Tamaricaceae
221. Tamarix smyrnensis Bunge
Gr Nomos & Eparchia Korinthias: Xylokastro,
coastal pine forest, 7 m, 38°04'N, 22°39'E,
23.09.2015, Zarkos obs.
New for nomos and eparchia. Introduced planting in
Pinus halepensis and P. p i n e a woodland, together with
Rhamnus alaternus.
Veronicaceae
222. Digitalis laevigata subsp. graeca (Ivanina)
Werner
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31967.
223. Veronica hederifolia L.
Gr Nomos Achaias/Korinthias, Eparchia
Kalavriton/Korinthias: Ntourntouvana,
Fig. 25. Viscum album subsp. album on Robinia pseudoacacia
(photo G. Zarkos).

396 Vladimirov, V. & al • New floristic records in the Balkans: 28
summit of Triadafilia, stony limestone scree,
2085 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Widespread in N and C Peloponnese.
224. Veronica praecox All.
Gr Nomos Achaias/Korinthias, Eparchia
Kalavriton/Korinthias: Ntourntouvana,
summit of Triadafilia, stony limestone scree,
2085 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
225. Veronica thymifolia Sm.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
1995 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Endemic to Greece (Peloponnese, Sterea Ellas and
Crete).
226. Veronica verna L.
Gr Nomos Achaias/Korinthias, Eparchia
Kalavriton/Korinthias: Ntourntouvana,
summit of Triadafilia, stony limestone scree,
2085 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Violaceae
227. Viola graeca (W. Becker) Halácsy
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, ridge between
summits, rocky limestone slopes and scree,
2030 m, 37°54'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Endemic to Greece (Peloponnese and Sterea Ellas).
228. Viola mercurii Halácsy
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, along
path to the summit, in Abies cephalonica and
Juniperus communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Endemic to Greece (Peloponnese).
Alliaceae
229. Allium guttatum subsp. tenorei (Parl.) Soldano
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in open Abies cephalonica woodland,
1120–1125 m, 37°55'N, 22°16'E, 12.08.2015, Kit
Tan & G. Vold 31940.
Colchicaceae
230. Colchicum atticum Spruner ex Tomm. (Fig. 26)
Gr Nomos & Eparchia Korinthias: pine forest
at Palaeo Kalamaki, 19 m, 37°53'N, 23°00'E,
18.11.2015, Zarkos obs.; loc. ibid., 01.12.2015,
Kit Tan, G. Vold & Zarkos obs.; loc. ibid., 28 m,
18.11.2015, Zarkos obs.; loc. ibid., 01.12.2015,
Kit Tan, G. Vold & Zarkos obs.; Mt Onia, stony
phrygana, 214 m, 37°52'N, 22°59'E, 26.11.2015,
Zarkos obs.; loc. ibid., 01.12.2015, Kit Tan, G. Vold
& Zarkos 32030 (living plants also collected).
New for the Peloponnese. Recorded from Sterea Ellas,
North Central, North East, North, East and West
Aegean islands.
Hyacinthaceae
231. Muscari botryoides (L.) Mill.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, along
path to the summit, in Abies cephalonica and
Juniperus communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Liliaceae
232. Fritillaria graeca Boiss. & Spruner
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: summit of Ntourntouvana, rocky and
stony limestone scree, 2051 m, 37°54'N, 22°15'E,
24.05.2015, Zarkos & Christodoulou obs.
Orchidaceae
233. Cephalanthera longifolia (L.) Fritsch
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, open Abies forest on
Fig. 26. Colchicum atticum (photo G. Zarkos).

397Phytol. Balcan. 21(3) • Sofia • 2015
steep rocky NW slopes, 1692 m, 37°55'N, 22°14'E,
09.07.2015, Zarkos obs.
234. Dactylorhiza sambucina (L.) Soó
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, along
path to the summit, in Abies cephalonica and
Juniperus communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
235. Epipactis helleborine (L.) Crantz
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, NE slopes, along
forest road in Abies cephalonica woodland,
1317 m, 37°55'N, 22°15'E, 24.05.2015, Zarkos &
Christodoulou obs.
236. Orchis pallens L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, along
path to summit, in Abies cephalonica and
Juniperus communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
237. Orchis spitzelii W.D.J. Koch subsp. spitzelii
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: Ntourntouvana, W slopes, along
path to summit, in Abies cephalonica and
Juniperus communis subsp. hemisphaerica forest,
1620 m, 37°55'N, 22°14'E, 24.05.2015, Zarkos &
Christodoulou obs.
Poaceae
238. Melica ciliata L.
Gr Nomos Achaias/Korinthias, Eparchia Kalavriton/
Korinthias: NE slopes of Ntourntouvana, 5 km
from Panorama along forest road to the summit,
1320–1350 m, 37°55'N, 22°15'E, 12.08.2015, Kit
Tan & G. Vold 31973.
Report 239
Konstantinos B. Simoglou
1
, Christos A. Dordas
2
&
Theodoros V. Koutsos
3
1
Rural Economy and Veterinary Directorate of Drama, e-mail:
kbsimoglou@fast-mail.org (corresponding author)
2
Laboratory of Agronomy, School of Agriculture, Aristotle
University of Thessaloniki
3
Hellenic Agricultural Organisation “DEMETER”
Thessaloniki
Fig. 28. Echium russicum owers (photo K.B. Simoglou).
Fig. 27. Echium russicum in full ower, visited by butteries
(photo K.B. Simoglou).
Boraginaceae
239. Echium russicum J.F. Gmel. (Figs. 27, 28)
Gr Nomos & Eparchia Dramas: Mt Falakron, Aj.
Pavlos, SE-facing slope, subalpine pastures
and rocky outcrops, 1450‒1600 m, 41°15'50"N,
23°53'13"E, 13.07.2012, Simoglou, Dordas &
Koutsos s.n. (photos).
This is the second record of Echium russicum from
Greece, the first being a collection by Elli Stamatiadou

398 Vladimirov, V. & al • New floristic records in the Balkans: 28
in 1981 (Stamatiadou 22355, ATH). Comments on
ecology, habitat and distribution are provided in the
following paragraphs. We found a small population
of 100 plants for the first time in 2012 on a SE-facing
slope of Mt Falakron at an altitude of 1450‒1600 m.
They were in full flower from mid-June to mid-July.
The tubular corollas were a bright coral red and the
ripe fruits black. We realized this was Echium russi-
cum which to our knowledge, does not seem to have
been documented from Greece and reported our dis-
covery in a Greek publication which we sent to Kit Tan
(Simoglou & al. 2015).
In the two localities of ca. 1.5 ha, rather shallow
soil overlies the limestone slope and at the edges of the
slope, apparently deeper soil supports a more dense
Fagus forest cover. The vegetation in this small area
includes Festuca rubra, Valeriana officinalis, Thymus
spp., Sideritis scardica, Centaurea salonitana, Stachys
officinalis, Rhinanthus rumelicus, Delphinium fis-
sum, Gladiolus palustris, Anthyllis aurea, Trifolium al-
pestre, Teucrium montanum, Senecio macedonicus,
Coronilla varia, Jurinea mollis, Asyneuma limonifoli-
um, Campanula persicifolia, Filipendula vulgaris and
Rosa pimpinellifolia, etc. In the following years of
2013, 2014 and 2015 we realized that Echium russicum
is well-established on Mt Falakron, with steady num-
bers of ca. 100 plants.
According to Taravati & al. (2013) Echium rus-
sicum has high concentrations of anthocyanins
(65 mg/g), the level of which depends on the intensi-
ty of the solar radiation in its habitat, and lower con-
centration of tannins (26 mg g). We noted that the
plants were very attractive to various species of bees
and butterflies; it is considered a honey-plant (Tashev
& Pancheva 2011), producing sufficient quantities of
nectar throughout the flowering period, with high
(58 %) sugar content (Chwil & Wesyszko-Chmielewska
2007). It also attracts the beetle Phytoecia coerulescens
(Cerambycidae) which is a known pest of the genus
(Kirk & Wapshere 1979.).
The seeds which contain a high concentration of
omega-3 and omega-6 fatty acids, make it a valuable
plant for cultivation and production of useful fatty ac-
ids (Guil-Guerrero 2003). The plant is at risk in the wild
probably due to over-collecting in the countries where
it grows as it is considered a medicinal plant with heal-
ing properties (Altundag & Ozturk 2011). It has much
ornamental value, being known horticulturally as Red-
flowered Viper’s Bugloss. Echium russicum is thus listed
as endangered in the European Red List of Angiosperm
Plants (Bilz & al. 2011). It is also included in Annexes II
and IV of the Habitats Directive 92/43/EU (as amended
by the Directive 2013/17/EU) which covers European
species requiring strict protection and Special Areas of
Conservation. The less easily accessible localities on Mt
Falakron would thus help to protect the plant and in-
crease its survival possibilities. The populations have
been monitored every year since the summer of 2012
and the plants were found to remain in good stand.
Acknowledgements. We wish to thank Dr Kit Tan for her kind
help in preparing our text.
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