The lichen biota of the Drawieński National Park (NW Poland, Western Pomerania)

Abstract

The whole known lichen biota of the Drawieński National Park is presented. In total 290 species (262 lichenized, 25 lichenicolous and 3 lichen-related, saprotrophic fungi) are listed. Trichonectria anisospora and Milospium lacoizquetae are reported as new to Poland. Lecanora stenotropa and Phaeophyscia pusilloides are reported for the first time from Polish lowlands. The most lichenologically interesting and richest habitat complexes are the river valleys with their beech slope forests, their alluvial forests and their fast running rivers. Further habitats of high nature conservation value are roadside trees and pine forests, which inhabit a rich lichen biota as well.

Full text

Folia Cryptog. Estonica, Fasc. 49: 59–71 (2012)
INTRODUCTION
First lichen records from the Drawieński Na-
tional Park originated from Karl Schulz-Korth
(Berlin) who visited the present territory and its
surroundings together with Johannes Hillmann
(Berlin) and the local teacher E. Putzler from
Choszczno (Arnswalde) in 1928. They explored
the western part of the national park walking
from the small town Drawno (Neuweddel) via
Barnimie (Fürstenau) and Moczele (Marzelle)
to Stare Osieczno (Hochzeit). Schulz-Korth pre-
sented his collections partly in his dissertation
on the lichen ora of the Mark Brandenburg
(Schulz-Korth 1931). Shortly after, the botanist
Adolf Straus (Berlin) collected lichens mainly
in the surrounding of Głusko (Steinbusch),
Sitnica (Marienthal) and Rogoźnica (Räumde),
and the teacher Aloys Mallach (Lubstorf) close
to Miradz (Grüneberg). Their data were pub-
lished later by Hillmann together with his own
results (Hillmann 1932–1933, 1936, 1942).
Finally, all results were compiled by Hillmann
and Grumman (1957) in the lichen ora of the
Mark Brandenburg.
Despite of the activities in the rst half of the
20th century the lichen ora of the Drawieński
National Park was poorly known until the work
of Lipnicki (1993). He investigated the southern
and western part of the park between Płociczna
river and Ostrowieckie lake, and reported 206
taxa, including several lichens new for Poland.
Inspired by the results of Lipnicki (1993) and
the nature value of the national park we decided
to conduct more studies on lichen biota of this
area. In this paper we present a compiled list of
lichenized, lichenicolous and allied fungi of the
Drawieński National Park, including historical
literature records and the results of our recent
surveys.
e lichen biota of the Drawieński National Park
(NW Poland, Western Pomerania)
Ulf Schiefelbein1, Paweł Czarnota2, Holger üs3 & Martin Kukwa4
1Blücherstr. 71, D-18055 Rostock, Germany.
E-mail: ulf.schiefelbein@gmx.de
2Department of Agroecology and Landscape Architecture, University of Rzeszów,
Ćwiklińskiej 2, PL-35-601 Rzeszów, Poland.
E-mail: pawczarnota@poczta.onet.pl
3Natural History Museum, Cromwell Road, London, SW7 5BD.
E-mail: h.thues@nhm.ac.uk
4Department of Plant Taxonomy and Nature Conservation, University of Gdańsk,
Al. Legionów 9, PL-80-441 Gdańsk, Poland.
E-mail: dokmak@ug.edu.pl
Abstract: e whole known lichen biota of the Drawieński National Park is presented. In total 290 species (262 lichenized,
25 lichenicolous and 3 lichen-related, saprotrophic fungi) are listed. Trichonectria anisospora and Milospium lacoizquetae are
reported as new to Poland. Lecanora stenotropa and Phaeophyscia pusilloides are reported for the rst time from Polish lowlands.
e most lichenologically interesting and richest habitat complexes are the river valleys with their beech slope forests, their
alluvial forests and their fast running rivers. Further habitats of high nature conservation value are roadside trees and pine
forests, which inhabit a rich lichen biota as well.
Kokkuvõte: Drawieński rahvuspargi (Lääne-Pomeraania, Loode-Poola) samblike elustik
Tutvustatakse Drawieński rahvuspargi kõiki teadaolevaid samblikke, nimekiri sisaldab 290 liiki, neist 262 on lihheniseerunud
seened, 25 on lihhenikoolsed seened ja 3 on samblikele lähedased saprotroofsed seened. Trichonectria anisospora ja Milospium
lacoizquetae on Poolale uued; Lecanora stenotropa ja Phaeophyscia pusilloides on esmasleiud Poola tasandikualadele. Lihheno-
loogiliselt kõige huvipakkuvamad ja liigirikkamad on jõgede orgude nõlvadel kasvavad pöögimetsad ja alluviaalsed metsad.
Ka teedeäärsed puud ning männimetsad, kus leidub rikkalikult samblikke, on kõrge looduskaitselise väärtusega kasvukohad.

60 Folia Cryptog. Estonica
STUDY AREA
The Drawieński National Park was established
in 1990 and is located in north-western Poland,
on the border of the Greater Poland, Lubusz
and West Pomeranian Voivodeships (Polish:
Wielkopolskie, Lubuskie i Zachodniopomorskie)
(Fig. 1). The park covers 113.42 km² and is a
part of Puszcza Drawska Forest, lying on the
outwash plain with deep crevices of the Drawa
and Płociczna rivers, and by a long string of
gully lakes.
The northern (between Zatom and Drawno) and
central part (Radęcin Reserve) of the study area
belong to the ground moraine and are covered
mainly by boulder-clay. The rusty soil is the
most widespread soil type in the area (c. 87%),
but also brown-earth soils and semihydrogenic
or hydrogenic soils occur there (Biały et al.
1997).
The climate of the Drawieński National Park is
characterized by the low amplitude of annual
temperature differences, late spring, long, warm
autumn, mild, not very snowy winter, mostly
western and north-western winds and long peri-
ods of springtime dryness. The average monthly
temperatures range from -2 ºC in January to
17 ºC in July and August (www.dpn.pl).
96.14 km² of the national park are covered by
forests of which 80% are pure or mixed pine
plantations (Pawlaczyk 1997); those stands
are now under the reconstruction in order to
develop natural forests. Present-day natural for-
est communities are acidophilic or mesotrophic
beech forests, alder swamp forests and ash-alder
alluvial forests, but those are rare and grown
almost exclusively in the valleys of the rivers and
in the nature reserve Radęcin. The oligo- and
mesotrophic bogs are developed in former small
POLAND
Fig.1. Location of the Drawieński National Park in Poland and collecting sites (asterisks) in the
study area.

61
lakes, in areas of intensive underground water
leakages and in river valleys, and cover 3% of
the total area. Also small mid-forest meadows
are located in the river valleys, which are regu-
larly mowed to protect the oristic and faunistic
species richness. Settlements within the park
are mostly abandoned, only some outposts and
remains of churches, churchyards and farm-
yards still exist.
MATERIALS AND METHODS
The list is based on observations and collections
made during eld trips in 2004 (P. Czarnota, lo-
calities nos 16–19) and 2010 (M. Kukwa and U.
Schiefelbein, localities nos 1–15) (Fig. 1), and on
published data. Specimens are deposited in the
herbaria UGDA, GPN and the private herbarium
of the rst author, with some duplicates in BM.
Lichen substances were analyzed by standard
technique of thin-layer chromatography in sol-
vent systems A, B, C and G (Orange et al. 2001).
The nomenclature follows mostly Fałtynowicz
(2003), Blanco et al. (2004), Santesson et al.
(2004), Nordin et al. (2010), Arup & Sandler
Berlin (2011), Kukwa (2011), Spribille et al.
(2011), and Diederich et al. (2012).
Following symbols and abbreviations are used
in the species list.: # – lichenicolous fungus, + –
non-lichenized saprobic fungus, Ac – Acer spp.,
Acpl – Acer platanoides, Acps – Acer pseudopla-
tanus, Alg – Alnus glutinosa, Ali – Alnus incana,
Be – Betula spp., br – brick, Cab – Carpinus
betulus, co – concrete, Coa – Corylus avellana,
de – detritus, Fas – Fagus syvatica, Fre – Fraxi-
nus excelsior, gr – granite, Lad – Larix decidua,
li – lichenicolous, loc. – locality, Ma – Malus
spp., mos – mosses, mor – mortar, Paa – Padus
avium, Pia – Picea abies, Pis – Pinus sylvestris,
Po – Populus spp., Pot – Populus tremula, Qu –
Quercus spp., Rop – Robinia pseudoacacia, sa
– sandstone, Sa – Salix spp., San – Sambucus
nigra, so – soil, Soa – Sorbus aucuparia, Ti – Tilia
spp., tu – turf, Ul – Ulmus spp., Vam – Vaccinium
myrtillus, wo – wood. Categories of threats ac-
cording to Cieśliński et al. (2006) are included
in square brackets. CR means critically endan-
gered, EN – endangered, VU – vulnerable and
NT – near threatened. The excluded species are
listed separately, after the main list of species.
List of localities
1. N of Głusko village, 53°03'10"N, 15°56'25"E,
roadside trees; 5 May 2010.
2. C. 1 km N of Głusko village, E of Drawa river,
53°03'10"N, 15°56'22–25"E, pine and mixed
forests, forest edge and row of ash trees; 5
May 2010.
3. Valley of Drawa river, c. 1.2 km N of Głusko,
53°03'14"N, 15°56'11"E, riverside, alluvial
and deciduous slope forests; 5 May 2010.
4. ‘Tragankowe Urwisko’ range, c. 2 km NE of
Zatom, 53°09'12"N, 15°51'49–54"E, steep
slopes with deciduous forests and some
open areas; 6 May 2010.
5. C. 2 km S of Sitnica by the road to Głusko
village, by the crossroad to Jelenie village,
53°05'22–23"N, 15°55'51"E, pine forests,
oak forests and old roadside oaks; 6 May
2010.
6. Former village Dąbrowa, c. 3 km N of Głuskie
and c. 2 km E of Moczele village, 53°05'45"N,
15°55'36"E, mesotrophic grassland and in
glades, old cemetery; 7 May 2010.
7. Road from Głusko to Jelenie village, c. 1
km N of the crossroad Głuskie – Drawno,
53°05'34"N, 15°56'05"E, c. 30 years old pine
forest; 7 May 2010.
8. Former settlement Pustelnia, river valley
around the bridge Gromnik, 53°07'23"N,
15°59'20"E, riverside, alluvial and decidu-
ous slope forests; 7 May 2010.
9. C. 1.5 km S of Pustelnia settlement,
53°07'05"N, 15°59'13"E, transition bog
woodland surrounded by old oaks and other
trees; 7 May 2010.
10. Ostrowite village, 53°04'05"N, 15°58'30"E,
ruins of the church, churchyard, roadside
trees and open area; 7 May 2010.
11. Stara Węgornia scenic spot, valley of
Płociczna river, c. 2 km NNE of Ostrowite
village, 53°04'42–46"N, 15°59'31–41"E, deep
river valley with alluvial and deciduous slope
forests; 8 May 2010.
12. Dębowy peninsula in Ostrowieckie lake,
53°05'14"N, 15°58'20"E, old lime trees and
old pine forest; 8 May 2010.
13. Bridge on Drawa river, c. 2 km S of Głusko
village, boundary stones along the road,
53°01'58"N, 15°56'43"E; 8 May 2010.
14. Bridge on Drawa river, c. 5 km N of Głusko
village, 53°04'57"N, 15°54'45"E, bridge and
surrounding slopes; 8 May 2010.

62 Folia Cryptog. Estonica
15. Vicinity of ‘Wydrzy Głaz’ nature monument,
c. 3 km N of Moczele village, 53°06'29"N,
15°54'17"E, erratic block in the river, alluvial
and deciduous slope forests; 8 May 2010.
16. Forest section no. 307, W of Ostrowieckie
lake, 53°04'30–32"N, 15°56'47–55"E, pine
forest; 18 May 2004.
17. ‘R adęcin’ nature reserve, 53°05'56"N,
15°53'40"E, Pomeranian beech forest; 18
May 2004.
18. Forest section no. 248, 53°05'47"N,
15°53'39"E, Pomeranian beech forest; 18
May 2004.
19. Border of Drawieński National Park near
‘Radęcin’ nature reserve, 53°05'38"N,
15º53'52"E, Pomeranian beech forest; 18
May 2004.
RESULTS AND DISCUSSION
Prior to our studies the lichen biota of the
Drawieński National Park included 188 lichen-
forming, one lichenicolous (Microcalicium
disseminatum) and one saprotrophic allied
(Leptorhaphis atomaria) species. Schulz-Korth
(1931) and Hillmann (1932–1933, 1936, 1942)
recorded only 30 species, but Lipnicki (1993)
identied 178 species accepted in this paper
(some names are considered to be synonymous
at present, and some taxa are excluded as
probably misidentied; see the list). During our
trips 228 species (201 lichenized, 25 licheni-
colous and two saprotrophic fungi) have been
recorded. Consequently, the lichen biota of the
Drawieński National Park consists of 290 taxa
(262 lichenized, 25 lichenicolous and 3 sapro-
trophic taxa). Two lichenicolous fungi, Trichonec-
tria anisospora and Milospium lacoizquetae, are
reported as new to Poland. Lecanora stenotropa
and Phaeophyscia pusilloides are reported for
the rst time from Polish lowlands. Several other
lichen species are rare in Poland; many of them
are commented in the list.
Four species (Bacidia circumspecta, Chrysothrix
candelaris, Lecanographa amylacea, Usnea or-
ida) are considered to be critically endangered
in Poland, further 25 species are endangered
to extinction and 30 species are thought to be
vulnerable; 16 taxa are in the category ‘near
threatened’ (Cieśliński et al. 2006). The occur-
rence of Usnea orida and Lecanora albella, L.
intumescens, Lobaria pulmonaria and Usnea
suboridana as well as Stereocaulon conden-
satum, Usnea dasypoga and U. hirta was not
conrmed nor by Lipnicki (1993) neither by
us; those taxa are most probably extinct there.
Also Caloplaca lobulata and C. pyracea have not
been recently conrmed, but at least the later
one can be found.
From the lichenological point of view the conser-
vation value of the Drawieński National Park is
largely determined by the deep valleys with their
fast running river, old, well-preserved beech
forests on steep slopes and alluvial forests. The
lichen biota of Fagus sylvatica is the richest and
70 species were found on its bark; also Carpinus
betulus is a host for many lichen taxa (42 spe-
cies). According to Wirth et al. (2009) nine of
them (Bacidia circumspecta, B. rosella, Biatora
eforescens, Lecanographa amylacea, Lobaria
pulmonaria, Microcalicium disseminatum, My-
cobilimbia pilularis, Peltigera horizontalis and
Phaeophyscia pusilloides) have a strong afnity
to old-growth forests, and additional twelve are
considered to be indicators of old-growth forests
in Polish lowland (Czyżewska & Cieśliński 2003).
Further important lichen habitats are the old
roadside trees, e.g., on the old maple trees in
Ostrowite village and its surrounding 44 species
occur, among other many threatened species
(e.g. Anaptychia ciliaris, Physconia distorta, Ph.
perisidiosa, Ramalina fastigiata) and other note-
worthy lichens (e.g. Phaeophyscia pusilloides).
Pine forests were usually strongly managed be-
fore the establishment of the national park, but
they are rich in terricolous lichens, especially in
Cladonia species.
SPECIES LIST
AbsconditellA lignicolA Vězda & Pišút – loc. 9:
wo.
AcAros porA fuscAt A (Ach.) Th. Fr. – Lipnicki
(1993); loc. 6, 13: gr.
AcArosporA moenium (Vain.) Räsänen [syn. As-
picilia moenium (Vain.) G. Thor & Timdal] –
Lipnicki (1991b, 1993, as Aspicilia excavata
G. Thor & Timdal): mor.
AlyxoriA vAriA (Pers.) Ertz & Tehler – Lipnicki
[1993, as Opegrapha diaphora (Ach.) Ach.,
O. lichenoides Pers., O. lichenoides var. chlo-
rina (Pers.) Redgr. and O. pulicaris (Hoffm.)
Schrad.]; loc. 4, 6, 8, 10, 15: Acpl, Acps,
Cab, Fas, Pot, Rop, wo. [NT]

63
AmAndineA punctAtA (Hoffm.) Coppins & Scheid. –
Lipnicki [1993, as Buellia punctata (Hoffm.)
A. Massal.]; loc. 2, 4–6, 10: Acpl, Acps, gr,
Fas, Pot, Qu, Rop, Sa, wo.
AnAptychiA ciliAris (L.) Körb. – Lipnicki (1993);
loc. 10: Acpl. [EN]
Anisomeridium poly pori (Ellis & Everh.) M.E.
Barr. – loc. 2, 4, 6, 8, 15: Cab, Fas, Fre, Pot,
Rop, San, Ul.
ArthoniA At rA (Pers.) A. Schneid. – Lipnicki
(1993, also as Opegrapha atra Pers. and as
Opegrapha atra var. arthonioidea Leight.);
loc. 4, 6: Ac, Alg, Coa, Fas, Fre, Pot.
ArthoniA didymA Körb. – Lipnicki (1991a, 1993,
as A. aspersella Leight.): Soa. [EN]
ArthoniA rAdiAtA (Pers.) Ach. – Lipnicki (1993);
loc. 4, 11, 15: Cab, Coa, Fas.
ArthoniA spAdiceA Leight. – Lipnicki [1993, as A.
spadicea var. subspadicea (Nyl.) Redinger];
loc. 2–4, 8, 11, 15: Alg, Cab, Fas, Fre.
Arthothelium ruAnum (A. Massal.) Körb. – Lip-
nicki (1993); loc. 11: Cab. [NT]
#ArthrorhAphis AeruginosA R. Sant. & Tønsberg
– loc. 3, 9, 11, 16: li (on Cladonia coniocraea
and C. digitata).
AspiciliA cinereA (L.) Körb. – Lipnicki (1993): gr.
#AtheliA ArAchnoideA (Berk.) Jülich – loc. 8: li
(on Physcia sp. and Xanthoria parietina).
bAcidiA circumspectA (Norrl. & Nyl.) Malme – loc.
11: Fas. [CR]
bAcidiA rosellA (Pers.) De Not. – Lipnicki (1993):
Fas. [EN]
bAcidiA rubellA (Hoffm.) A. Massal. – Hillmann &
Grummann [1957, as B. luteola (Ach.) Mudd
var. pachythallina Vain.], Lipnicki (1993);
loc. 10: Acpl. [VU]
bAcidiA subincomptA (Nyl.) Arnold – loc. 5: Qu.
[EN]
bAcidinA AdAstrA (Sparrius & Aptroot) M. Hauck
& V. Wirth – loc. 2, 6: Fre, San.
bAcidinA cAligAns (Nyl.) Llop & Hladún – loc. 2,
6: Fre, Rop.
bAcidinA delicAtA (Larbal. & Leight.) V. Wirth &
Vězda – loc. 6: San.
bAcidinA neosquAmulosA (Aptroot & Herk) S. Ek-
man – loc. 2, 11: Cab, Fre.
bAcidinA sulphurellA (Samp.) M. Hauck & V.
Wirth – loc. 2, 8, 11: Ali, Cab, Fre. Only
recently distinguished species, previously
included in B. arnoldiana (Körb.) V. Wirth
& Vězda (Brand et al. 2009); so far reported
from two localities in Poland (Brand et al.
2009), but most probably much more com-
mon.
bAeomyces rufus (Huds.) Rebent. – Lipnicki
(1993); loc. 4: so.
biAtorA efflorescens (Hedl.) Räsänen – Lipnicki
(1993): Qu. [VU]
biAtorA globulosA (Flörke) Fr. – Lipnicki [1993,
as Bacidia globulosa (Flörke) Hafellner & V.
Wirth]; loc. 5, 6, 10: Acpl, Acps. Qu. [VU]
biAtorA ponticA Printzen & Tønsberg – loc. 11:
Cab. Recently found in Poland (Kukwa et
al. 2012).
bilimbiA sAbuletorum (Schreb.) Arnold – loc. 8,
10: co, mor, mos. Most probably also the
record of Mycobilimbia tetramera (De Not.)
Vitik. et al. [see Lipnicki 1993, as M. fusca
(A. Massal.) Hafellner & V. Wirth] belongs
to this species.
#briAncoppinsiA cytosporA (Vouaux) Diederich,
Ertz, Lawrey & van den Boom [syn. Phoma
cytospora (Vouaux) D. Hawksw.] – loc. 5: li
(on Hypogymnia physodes).
bryoriA fuscescens (Gyeln.) Brodo & D. Hawksw.
– Lipnicki (1993): Be. [VU]
buelliA AethAleA (Ach.) Th. Fr. – Lipnicki (1993):
gr.
buelliA griseovirens (Turner & Borrer ex Sm.)
Almb. – Lipnicki (1993); loc. 2–6, 8, 11, 15:
Ali, Cab, Fas, Fre, Pot, Qu, Rop.
cAlicium Adspersum Pers. – Lipnicki (1993); loc.
17: Qu. [EN]
cAlicium glAucellum Ach. – loc. 11: Alg. [VU]
cAlicium sAlicinum Pers. – Lipnicki (1993); loc.
4, 5: Acpl, Fas (wo), Qu. [VU]
cAlicium viride Pers. – Lipnicki (1993); loc. 5, 10:
Acpl, Acps, Alg, Fas, Qu. [VU]
cAloplAcA citrinA (Hoffm.) Th. Fr. – Lipnicki
(1993); loc. 2, 14: br, co.
cAloplAcA decipiens (Arnold) Blomb. & Forssell
– Lipnicki (1993): co.
cAloplAcA flAvocitrinA (Nyl.) H. Olivier – loc.
8: co.
cAloplAcA lobulAtA (Flörke) Hellb. – Hillmann
[1932–1933, as Xanthoria lobulata (Flörke)
B. de Lesd.], Hillmann & Grummann (1957,
as X. lobulata): San.
cAloplAcA oAsis (A. Massal.) Szatala – loc. 10, 14:
co. The occurrence of this species in Poland
has only been recently conrmed by Wilk
(2011), with two localities reported from Po-
merania. The species is most probably more
common than the morphologically similar

64 Folia Cryptog. Estonica
Caloplaca holocarpa; records of the latter
reported by Lipnicki (1993) most probably
belong here as well.
cAloplAcA pusillA (A. Massal.) Zahlbr. – loc.
10: co, mor. Caloplaca pusilla has previ-
ously been led under the name C. saxicola
(Hoffm.) Nordin (Wilk 2011), but probably
it is more common in the lowland than the
latter taxon. The records of C. saxicola by
Lipnicki (1993) may belong here.
cAloplAcA pyrAceA (Ach.) Th. Fr. – Hillmann
(1942), Hillmann & Grummann (1957): Pot.
cAndelAriellA AurellA (Hoffm.) Zahlbr. – Lipnicki
(1993); loc. 10: co, mor.
cAndelAriellA corAllizA (Nyl.) H. Magn. – Lipnicki
(1993); loc. 15: co, gr.
cAndelAriellA efflorescens R.C. Harris & W.R.
Buck – loc. 6, 12: Qu, San, Ul.
cAndelAriellA vitellinA (Hoffm.) Müll. Arg. – Lip-
nicki (1993); loc. 6, 13: co, gr.
cAndelAriellA xAnthostigmA (Pers. ex Ach.) Let-
tau – Hillmann (1932–1933), Lipnicki (1993);
loc. 5, 6, 10: Acpl, Acps, Pot, Qu.
cAtillAriA nigroclAvAtA (Nyl.) Schuler – loc. 6, 8:
Ma, Pot, San.
cetrAriA AculeAtA (Schreb.) Fr. – Lipnicki [1993,
as Coelocaulon aculeatum (Schreb.) Link.]:
so.
cetrAriA sepincolA (Ehrh.) Ach. – loc. 9: Lad. [EN]
chAenothecA brAchypodA (Ach.) Tibell – Lipnicki
(1993); loc. 6: Fas, Rop. [EN]
chAenothecA chrysocephAlA (Turner ex Ach.) Th.
Fr. – Lipnicki (1993); loc. 3, 5, 9–12: Acpl,
Acps, Alg, Qu, Ti, wo.
chAenothecA ferrugineA (Turner ex Sm.) Mig. –
Hillmann [1932–1933, as Chaenotheca mel-
anophaea (Ach.) Zwack.], Lipnicki (1993),
Tobolewski & Kupczyk (1974); loc. 2–6, 9,
10, 12: Acpl, Alg, Be, Ma, Pis, Pot, Qu, Ti, wo.
chAenothecA furfurAceA (L.) Tibell – Lipnicki
(1993); loc. 11, 15, 16: Alg, Coa, Fas, Qu,
wo. [NT]
chAenothecA stemoneA (Ach.) Müll. Arg. – Lip-
nicki (1993); loc. 11: Alg, Qu. [EN]
chAenothecA trichiAlis (Ach.) Th. Fr. – Lipnicki
(1993); loc. 3, 5, 6, 9: Acpl, Alg, Fas, Qu,
Rop. [NT]
#chAenothecopsis sAvonicA (Räsänen) Tibell – loc.
15: li (on Chaenotheca furfuracea).
#chA en ot hec op si s pusillA (Flörke) A.F.W.
Schmidt – loc. 17: li (associated also with
algae).
chrysothrix cAndelAris (L.) J.R. Laundon – Lip-
nicki (1993): Qu, Ti. [CR]
circinAriA cAesiocinereA (Nyl. ex Malbr.) A. Nor-
din, S. Savić & Tibell – loc. 15: gr.
circinAriA cAlcAreA (L.) A. Nordin, S. Savić &
Tibell – Lipnicki [1993, as Aspicilia calcarea
(L.) Mudd]: mor.
circinAriA contortA (Hoffm.) A. Nordin, S. Savić
& Tibell – Lipnicki [1993, as Aspicilia con-
torta (Hoffm.) Kremp.]; loc. 14: co.
clAdoniA ArbusculA (Wallr.) Flot. subsp. ber-
ingiana Ahti – Lipnicki [1993, as Cladina
arbuscula (Wallr.) Hale & W.L. Culb.]; loc.
4, 12: so.
clAdoniA ArbusculA subsp. mitis (Sandst.) Ruoss
– Lipnicki [1993, as Cladina mitis (Sandst.)
Hustich]; loc. 4, 12: so.
clAdoniA cAespiticiA (Pers.) Flörke – loc. 4: so.
[EN]
clAd oniA ceno teA (Ach.) Schaer. – Lipnicki
(1993); loc. 5, 7, 9, 12: Pis, so, wo.
clAdoniA cervicornis (Ach.) Flot. subsp. verticil-
lAtA (Hoffm.) Ahti – Lipnicki (1993): so.
clAdoniA chlorophAeA (Flörke) Speng. – Lipnicki
(1993); loc. 4: so, wo.
clAdoniA cil iAtA Stirt. – Lipnicki [1993, as
Cladina ciliata (Stirt.) Trass]; loc. 4, 7: so.
clAdoniA coniocrAeA (Flörke) Spreng. (syn. C.
ochrochlora Flörke) – Hillmann (1932–1933,
as C. ochrochlora), Hillmann & Grummann
(1957, as C. ochrochlora), Lipnicki (1993,
also as C. ochrochlora); loc. 2–7, 9–12, 15:
Alg, Be, Cab, Fas, Fre, Qu, so, mos, wo.
clAdoniA cornutA (L.) Hoffm. – Lipnicki (1993);
loc. 7, 12: so.
clAdoniA crispAtA (Ach.) Flot. – Lipnicki (1993):
so.
clAdoniA deformis (L.) Hoffm. – Lipnicki (1993);
loc. 4, 7: so, wo.
clAdoniA digitAtA (L.) Hoffm. – Lipnicki (1993);
loc. 2–5, 7–9, 11, 12: Alg, Pis, so, wo.
clAdoniA fimbriAtA (L.) Fr. – Lipnicki (1993); loc.
4, 6–10, 12: Ali, Be, Pot, Rop, so, wo.
clAdoniA foliAceA (Huds.) Willd. – Lipnicki
(1993): so.
clAdoniA furcA tA (Huds.) Schrad. – Lipnicki
(1993); loc. 4, 7: so.
clAdoniA glAucA Flörke – Lipnicki (1993); loc. 7,
9: Be, so, wo.
clAdoniA grAcilis (L.) Willd. – Lipnicki (1993);
loc. 7: so.
clAdoniA grAyi G. Merr. ex Sandst. – loc. 9: wo.

65
clAdoniA incrAssAtA Flörke – loc. 9: wo. [EN]
clAdoniA mAcilentA subsp. floerkeAnA (Fr.)
V. Wirth – Lipnicki [1993, as Cladonia
oerkeana (Fr.) Flörke]; loc. 5: so, wo.
clAdoniA mAcilentA Hoffm. subsp. mAcilentA –
Lipnicki [1993, as Cladonia bacillaris (Ach.)
Nyl. and C. macilenta Hoffm.]; loc. 3–5, 9,
12: Be, Pis, so, wo.
clAdoniA merochlorophAeA Asahina – loc. 5, 7,
12: mos, so.
clAdoniA phyllophorA Ehrh. ex Hoffm. – Lipnicki
(1993); loc. 4, 7: so.
clAdoniA pleurotA (Flörke) Schaer. – Lipnicki
(1993); loc. 4: so.
clAdoniA portentosA (Dufour) Coem. – Lipnicki
[1993, as Cladina portentosa (Dufour) Foll-
mann]: so.
clAdoniA rAmulosA (With.) J.R. Laundon – loc.
4: so.
clAdoniA rAngiferinA (L.) Weber ex F. H. Wigg. –
Lipnicki [1993, as Cladina rangiferina (L.)
Nyl.]; loc. 7: so.
clAdoniA rAngiformis Hoffm. – Lipnicki (1993):
so.
clAdon iA squAmosA (Scop.) Hoffm. – Lipnicki
(1993); loc. 6: Be, Rop, so, wo.
clAdoniA subulAtA (L.) Weber ex F.H. Wigg. – Lip-
nicki (1993): so, wo.
clAdoniA unciAlis (L.) Weber ex F.H. Wigg. – Lip-
nicki (1993); loc. 4: so.
#clypeococcum hypocenomycis D. Hawksw. –
loc. 2, 4–6, 9, 12, 16: li (on Hypocenomyce
scalaris).
coenogonium pineti (Schrad.) Lücking & Lumb-
sch – Lipnicki [1993, as Dimerella pineti
(Schrad. ex Ach.) Vězda]; loc. 2–4, 9, 11, 15:
Alg, Fas, Pia, Pis, Qu, Ti, Vam.
dibAeis bAeomyces (L. f.) Rambold & Hertel –
Lipnicki (1993, as Baeomyces roseus Pers.):
so. [NT]
diplotommA cf. AlboAtrum (Hoffm.) Flot. – loc.
10: gr.
#epiclAdoniA sAndstedei (Zopf) D. Hawksw. – loc.
9: li (on Cladonia coniocraea)
#epiclAdoniA stenosporA (Harm.) D. Hawksw. –
loc. 9: li (on Cladonia coniocraea).
everniA prunAstri (L.) Ach. – Lipnicki (1993); loc.
5, 6, 8–10, 12: Acpl, Ali, Be, Qu, Rop, Ti. [NT]
fellhAnerA subtilis (Vězda) Diederich & Sérus.
– loc. 18: Pic.
flAvopArmeliA cApe rAtA (L.) Hale – Lipnicki
(1993): Qu. [EN]
fuscideA ArboricolA Coppins & Tønsberg – loc.
15: Fas.
fuscideA pusillA Tønsberg – loc. 9: Be.
grAphis scriptA s.l. – Lipnicki (1993); loc. 3, 4,
11: Cab, Fas. [NT]
hAlecAniA viridescens Coppins & P. James – loc.
2, 6, 8: Fre, Ma, San.
hypocenomyce AnthrAcophilA (Nyl.) P. James &
Gotth. Schneid. – loc. 5: Pis.
hypocenomyce cArAdocensis (Leight. ex Nyl.) P.
James & Gotth. Schneid. – Lipnicki (1993);
loc. 16: Pis, wo.
hypocenomyce scAlAris (Ach. ex Lilj.) M. Choisy –
Lipnicki (1993); loc. 2–7, 9, 11, 12, 16: Alg,
Fre, Pis, Qu, Ti, wo.
hypogymniA fArinAceA Zopf – Hillmann [1932–
1933, as Hypogymnia bitteriana (Zahlbr.)
Räsänen], Hillmann (1936, as Parmelia
bitteriana Zahlbr.), Hillmann & Grummann
(1957, as P. bitteriana); loc. 4: Fas. [VU]
hyp ogy mn iA p hys ode s (L.) Nyl. – Hillmann
[1932–1933, as Parmelia physodes (L.) Ach.],
Lipnicki (1993); loc. 2, 3, 5–7, 9–12, 15, 17:
Acpl, Acps, Alg, Be, Cab, Fas, Fre, Ma, Pia,
Pis, Pot, Qu, Rop, so, Soa, Ti, wo.
hypogymniA tubulosA (Schaer.) Hav. – Lipnicki
(1993); loc. 2, 5, 10, 12, 15, 17: Acpl, Acps,
Fas, Fre, Qu, Ti, wo. [NT]
imshAugiA Aleurites (Ach.) S.L.F. Mey. – Lipnicki
(1993); loc. 12: Pis, wo.
lecAniA cyrtellA (Ach.) Th. Fr. – Lipnicki (1993);
loc. 4–6, 8: Fas, Qu, San, wo.
lecAniA nAegelii (Hepp) Diederich & van den
Boom – loc. 2, 4, 5, 8: Fas, Fre, Qu, San.
lecAniA sylvestris (Arnold) Arnold – Lipnicki
(1993): co.
lecAnogrAph A AmylAceA (Ehrh. ex Pers.) Egea
& Torrente – Hillmann [1932–1933, as Le-
canactis amylacea (Ehrh. ex Pers.) Arnold],
Hillmann & Grummann (1957, as L. amyla-
cea), Lipnicki (1993, as L. amylacea): Ac,
Fas, Qu, wo. [CR]
lec Ano rA Alb ell A (Pers.) Ach. – Hillmann
[1932–1933, as L. pallida (Schreb.) Ra-
benh.], Hillmann & Grummann (1957, as
L. pallida). [EN]
lecAnorA Albescens (Hoffm.) Branth & Rostr. –
Lipnicki (1993); loc. 10: br, co.
lecAnorA AllophAnA Nyl. – Hillmann (1932–1933),
Lipnicki (1993); loc. 6: Acpl, Alg.
lecAnorA ArgentAtA (Ach.) Malme – Hillmann
(1932–1933, as L. subfusca (L.) Ach. var. ex-

66 Folia Cryptog. Estonica
crescens Hillmann), Hillmann & Grummann
(1957, as L. subfuscata), Lipnicki (1993); loc.
3, 4, 11, 12, 15: Cab, Fas, Ti.
lecAnorA cArpineA (L.) Vain. – Lipnicki (1993);
loc. 2, 4, 6, 11: Acpl, Cab, Fas, Fre.
lecAnorA chlArotherA Nyl. – Lipnicki (1993); loc.
6: 10: Acpl, Fas, Pot.
lecAnorA compAllens Herk & Aptroot – loc. 5:
Acpl.
lecAnorA conizAeoides Nyl. ex Cromb. – Lipnicki
[1993, also as L. conizaea (Ach.) Nyl.]; loc.
4–7, 9–12: Be, Pis, wo.
lecAnorA crenulAtA (Dicks.) Hook. – Lipnicki
(1993): co.
lecAnorA dispersA (Pers.) Röhl. – Lipnicki (1993);
loc. 10: co.
lecAnorA expAllens Ach. – Lipnicki (1993); loc.
2–6, 8, 10–12, 17: Acpl, Acps, Alg, Ali, Fas,
Fre, Ma, Pot, Qu, Rop, San, Ti.
lecA norA glAbr AtA (Ach.) Malme – Lipnicki
(1993): Cab, Fas.
lecAnorA hAgenii (Ach.) Ach. – Lipnicki [1993,
also as L. umbrina (Ehrh.) Röhl.]; loc. 10:
Acpl, co.
lecAnorA horizA (Ach.) Lindsay – loc. 3: Cab.
lecAnorA intumescens (Rebent.) Rabenh. – Hill-
mann (1932–1933), Hillmann & Grummann
(1957): Fas. [EN]
lecAnorA persimilis Th. Fr. – loc. 2, 4, 6, 10, 12:
co, Fas, Fre, Pot, Qu, Ti. [DD]
lecAnorA polytropA (Ehrh. ex Hoffm.) Rabenh.
– Lipnicki (1993): gr.
lecAnorA pulicAris (Pers.) Ach. – Lipnicki (1993);
loc. 3, 4, 6, 15: Cab, Fas, Fre, Pot.
lecAnorA rugosellA Zahlbr. – Lipnicki (1993):
Alg, Fas.
lecAnorA sAlignA (Schrad.) Zahlbr. – Lipnicki
(1993); loc. 6, 8, 12: Ali, wo, bark of decidu-
ous trees.
lecA norA sAxi colA (Pollich) Ach. – Lipnicki
(1993); loc. 6: gr.
lecAnorA semipAllidA H. Magn. – loc. 10: co.
lecAnorA stenotropA Nyl. – loc. 13: gr. The spe-
cies has been only recently reported from
Poland (Flakus 2007). This is the rst Polish
lowland locality of the species.
lecAnorA subrugosA Nyl. – loc. 2, 6: Fre, Ma.
lecAnorA symmictA (Ach.) Ach. – Lipnicki (1993);
loc. 2: Fre, wo.
lecAnorA vAriA (Hoffm.) Ach. – Lipnicki (1993):
Bark of deciduous trees, wo.
lecideA fuscoAtrA (L.) Ach. – Lipnicki (1993);
loc. 13: gr.
lecideA nylAnderi (Anzi) Th. Fr. – loc. 4, 8, 12:
Ali, Pis.
lecidellA elAeochromA (Ach.) M. Choisy – Lip-
nicki [1993, also as L. euphorea (Flörke)
Hertel]; loc. 2–4, 6, 8, 13: Acpl, Alg, Cab,
Fas, Fre, Pot, San, Ti.
lecidellA flAvosorediAtA (Vězda) Hertel & Leuck-
ert – loc. 10: Acpl.
lecidellA stigmAteA (Ach.) Hertel & Leuckert –
Lipnicki 1993, also as Lecidella cinnamomea
Helb.); loc. 6, 10: co, mor.
leimonis errAticA (Körb.) R.C. Harris & Lendemer
– loc. 16, 19: gr. [NT]
leprAriA elobAtA Tønsberg – loc. 8, 9, 11, 15:
Alg, Ali, Cab, Soa.
leprAriA incAnA (L.) Ach. – Lipnicki (1993); loc.
2–12, 15: Acpl, Alg, Ali, Be, Cab, Fas, Fre,
Ma, Pis, Qu, Rop, San, Ti, wo.
leprAriA jAckii Tønsberg – loc. 2, 7, 12: Pis.
leprAriA lobificAns Nyl. – loc. 2–4, 6, 8, 10–12,
15: Cab, Fas, Fre, mor, Qu, Rop, San, Ti.
leprAriA rigidulA (de Lesd.) Tønsberg – loc. 2,
4–6, 8–11, 15: Acpl, Alg, Ali, Cab, Fas Fre,
Pot, Qu, San, Soa.
leprAriA vouAuxii (Hue) R.C. Harris – loc. 2, 5, 6,
10, 15: Acpl, Acps, co, Fas, Fre, Ma, Pis, Rop.
+leptorhAphis AtomAriA (Ach.) Szatala – Lipnicki
[1993, as Arthopyrenia punctiformis (Pers.)
A. Massal. var. atomaria (Ach.) Anzi]: Cab.
#lichenoconium erodens M.S. Christ. & D.
Hawksw. – loc. 2, 10, 11: li (on Hypogymnia
physodes, Lecanora conizeoides and Parme-
lia sulcata).
#lichenoconium lecAnorAe (Jaap) D. Hawksw. –
loc. 7, 10: li (on Lecanora conizeoides and
Pleurosticta acetabulum).
#lichenoconium pyxidAtAe (Oudem) Petrak & H.
Sydow – loc. 6, 12: li (on Cladonia mbriata
and C. cornuta).
#lichenoconium xAnthoriAe M.S. Christ. – loc. 1:
li (on Xanthoria parietina).
#lichenodiplis lecAnorAe (Vouaux) Dyko & D.
Hawksw. – loc. 6: li (on Lecanora saligna).
#lichenodiplis pertusAriicolA (Nyl.) Diederich –
loc. 3: li (on Pertusaria leioplaca).
lichenomphAliA umbelliferA (L.) Redhead, Lut-
zoni, Moncalvo & Vilgalys – loc. 9: mos, tu,
wo. [NT]
lobAriA pulmonAriA (L.) Hoffm. – Hillmann (1932–
1933), Schulz-Korth (1931). [EN]
melAnelixiA glAbrAtulA (Lamy) Sandler & Arup
– Lipnicki [1993, as Melanelia fuliginosa
(Fr. ex Duby) Essl.]; loc. 2–6, 8–12, 15, 17:

67
Acpl, Acps, Alg, Ali, Be, Cab, Fas, Fre, Pot,
Qu, Rop, Ul.
melAnelixiA subArgentiferA (Nyl.) O. Blanco et
al. – Hillmann [1932–1933, as Parmelia
verruculifera Nyl.), Hillmann & Grummann
(1957, as P. subargentifera Nyl. f. conspur-
cata (Schaer.) Grummann), Lipnicki [1993,
as Melanelia subargentifera (Nyl.) Essl.]; loc.
10: Ac, Acpl. [VU]
melAnelixiA subAuriferA (Nyl.) O. Blanco et al.
– Lipnicki [1993, as Melanelia subaurifera
(Nyl.] Essl.): Fas.
melAnohAleA elegAntulA (Zahlbr.) O. Blanco et
al. – Lipnicki [1993, as Melanelia incolorata
(Parr.) Essl.]; loc. 10: Acps, on bark of de-
ciduous trees. [VU]
melAnohAleA exAsperAtulA (Nyl.) O. Blanco et al.
– Lipnicki [1993, as Melanelia exasperatula
(Nyl.) Essl.]; loc. 9: Ac, Be, Cab.
micAreA byssAceA (Th. Fr.) Czarnota, Guzow-
Krzemińska & Coppins – loc. 11: Alg.
micAreA denigrAtA (Fr.) Hedl. – Hillmann [1932–
1933, as Catillaria synothea (Ach.) Beltr.],
Hillmann & Grummann [1957, as C. deni-
grata (Fr.) Hedl.], Lipnicki (1993); loc. 7, 8,
12, 16, 18, 19: Ali, Pis, wo.
micAreA melAenA (Nyl.) Hedl. – loc. 8, 11: wo. [NT]
micAreA micrococcA (Körb.) Gams ex Coppins –
loc. 16: Be, Pis.
micAreA misellA (Nyl.) Hedl. – loc. 4, 6, 10, 12: wo.
micAreA nitschkeAnA (J. Lahm ex Rabenh.) Harm.
– loc. 16, 18: Pic, Pis.
micAreA pr AsinA Fr. – Lipnicki (1993); loc. 2,
11, 17: Alg, Cab, Qu, so, Ul, wo. Records
by Lipnicki (1993) may also belong to other
species of the complex, M. byssacea and M.
micrococca.
#microcAlicium disseminAtum (Ach.) Vain. – Lip-
nicki (1993); loc. 11, 17: li (on Chaenotheca
stemonea).
#milospium lAcoizquetAe Etayo & Diederich – loc.
9: li (on Cladonia digitata). The species is
new to Poland.
#monodictys epileprAriA Kukwa & Diederich –
loc. 2, 3, 8, 9, 11, 15: li (on Lepraria elobata,
L. incana and L. rigidula).
mycobilimbiA epixAnthoides (Nyl.) Vitik., Ahti,
Kuusinen, Lommi & T. Ulvinen – loc. 15: Fas.
mycobilimbiA pilulAris (Körb.) Hafellner & Türk
– Lipnicki [1993, as Biatora pilularis (Körb.)
Hepp]: Fas. [VU]
ochrolechiA bAhusiensis H. Magn. – loc. 2, 4:
Acps, Fas.
ochrolechiA subviridis (Høeg) Erichsen – Lip-
nicki (1993): Cab. [VU]
ochrolechiA turneri (Sm.) Hasselrot – loc. 5,
10: Acpl.
opegrAphA niveoAtrA (Borrer) Laundon – loc. 15:
Fas. [VU]
opegrAphA vermicelliferA (Kunze) J.R. Laundon
– Lipnicki (1993): Cab. [EN]
pArmeliA sAxAtilis (L.) Ach. – Lipnicki (1993); loc.
3–5, 9, 12: Alg, Be, Fas, Fre, Qu, Soa, Ti, Ul.
pArmeliA sulcAtA Taylor – Hillmann (1932–1933),
Hillmann & Grummann (1957, also as P.
sulcata f. pruinosa (Harm.) Maas Geest.),
Lipnicki (1993); loc. 2–6, 8–12: Acpl, Acps,
Alg, Ali, Be, Cab, Fas, Fre, Ma, Pot, Qu, Rop,
San, Ti, Ul.
pArmelinA til iAce A (Hoffm.) Hale – Lipnicki
(1993): Qu. [VU]
pArmeliopsis AmbiguA (Wulfen) Nyl. – Lipnicki
(1993); loc. 2–4, 7, 8, 11, 12: Alg, Cab, Be,
Fas, Pis, Qu, wo.
pArmeliopsis hyperoptA (Ach.) Arnold – loc. 9:
Be. [VU]
peltigerA cAninA (L.) Willd. – Lipnicki (1993):
so. [VU]
peltigerA horizontAlis (Huds.) Baumg. – Lipnicki
(1993): so. [EN]
peltigerA prAetextAtA (Flörke ex Sommerf.) Vain.
– Hillmann (1932–1933, also as P. subcanina
Gyeln.), Hillmann & Grummann (1957, also
as P. subcanina), Lipnicki (1993), Schulz-
Korth (1931, as P. subcanina); loc. 4, 11,
14: Qu, so, wo. [VU]
peltigerA rufescens (Weiss) Humb. – Lipnicki
(1993): so.
pertusAriA Albescens (Huds.) M. Choisy & Wer-
ner – Hillmann [1932–1933, as P. orbiculata
(Schreb.) Zahlbr.], Lipnicki (1993); loc. 10,
12: Acpl, Acps, Ti.
pertusAriA AmArA (Ach.) Nyl. – Lipnicki (1993);
loc. 2–6, 8, 10–12, 15: Acpl, Acps, Alg, Cab,
Fas, Fre, Qu, Ti, wo.
pertusAriA coccodes (Ach.) Nyl. – Hillmann
& Grummann [1957, as P. coccodes var.
phymatodes (Ach.) Almb.], Lipnicki (1993),
Tobolewski (1988); loc. 3–5, 10, 12: Acpl,
Acps, Cab, Fas, Ti. [NT]
pertusAriA coro nAtA (Ach.) Th. Fr. – Lipnicki
[1993, as P. coronata var. isidiifera (Erich-
sen) Almb.]: Cab. [VU]
per tusAriA flAvid A (DC.) J.R. Laundon – To-
bolewski (1988); loc. 4, 12, 15: Fas, Ti. [EN]

68 Folia Cryptog. Estonica
per tusAriA hemisphAericA (Flörke) Erichsen –
Hillmann & Grummann (1957), Lipnicki
(1993); loc. 12, 15: Fas, Ti, bark of decidu-
ous trees. [VU]
pertusAriA leioplAcA DC. – Hillmann (1932–
1933), Hillmann & Grummann (1957),
Lipnicki (1993); loc. 2–4, 11, 15: Cab, Fas,
Fre. [NT]
per tusAriA pe rtusA (Weigel) Tuck. – Lipnicki
(1993); loc. 4, 10, 15: Acpl, Fas, wo. [VU]
phAeophysciA nigricAns (Flörke) Moberg – Hill-
mann & Grummann [1957, as Physcia
nigricans (Flörke) Stizenb.], Lipnicki (1993);
loc. 6, 10: co, Pot, Qu.
phAeophysciA orbiculAris (Neck.) Moberg – Lip-
nicki (1993); loc. 6, 8, 10, 12: Acpl, co, Pot,
San, Ul.
phAeophysciA pusilloides (Zahlbr.) Essl. – loc. 10:
Acpl. This is very rarely reported species in
Poland (see Fałtynowicz 2003); it is the rst
record from Polish lowlands. [DD]
#phAeospo robulus sp. – loc. 4: li (on Buellia
griseovirens). It is perhaps an undescribed
species; material needs more study.
phlyctis AgelAeA (Ach.) Flot. – Lipnicki (1993):
Cab, Fas, Pot. [EN]
phlyctis ArgenA (Spreng.) Flot. – Lipnicki (1993);
loc. 2, 4, 5, 8, 10–12, 15: Acpl, Acps, Ali,
Cab, Fas, Fre, Qu, San, Ti, Ul, wo.
physciA Adscendens (Th. Fr.) H. Olivier – Lipnicki
(1993); loc. 1, 6, 8, 12: Paa, Pot, San, Ul.
physciA cAesiA (Hoffm.) Fürnr. – Lipnicki (1993);
loc. 6, 10: co.
physciA dubiA (Hoffm.) Lettau – Lipnicki (1993):
Ac.
physciA stellAris (L.) Nyl. – loc. 8: San.
physciA tenellA (Scop.) DC. – Lipnicki (1993);
loc. 4, 6, 8, 9, 12, 15, 17: Be, Cab, Fas, gr,
Ma, Paa, Rop, San, Ul.
physconiA distortA (With.) J.R. Laundon – Lip-
nicki (1993); loc. 5, 10: Ac, Acpl. [EN]
physconiA enteroxAnthA (Nyl.) Poelt – Lipnicki
(1993); loc. 6, 10: Acpl, San.
physconiA griseA (Lam.) Poelt – Lipnicki (1993);
loc. 6, 10: Ac, Acpl, co, Pot.
physconiA perisidiosA (Erichsen) Moberg – Lip-
nicki (1993); loc. 5, 10: Acpl, Acps, co. [EN]
piccoliA ochrophorA (Nyl.) Hafellner – loc. 6: Qu.
plAcynthiellA dAsAeA (Stirt.) Tønsberg – loc. 3,
4, 6, 7, 9, 11: Be, co, Pic, wo.
plAcynthiellA icmAleA (Ach.) Coppins & P. James
– Lipnicki (1993); loc. 8, 9, 12: Ali, Be, so, wo.
plAcynthiellA oligotrophA (J.R. Laundon) Cop-
pins & P. James – Lipnicki (1993); loc. 12:
de, so, wo.
plAcynthiellA uliginosA (Schrad.) Coppins & P.
James – Lipnicki (1993); loc. 12: de, so, wo.
plAtismAtiA glAucA (L.) W.L. Culb. & C.F. Culb. –
Lipnicki (1993); loc. 3–5, 8, 9, 12, 15: Acpl,
Ali, Fas, Qu, Ti, wo.
pleurostictA AcetAbulum (Neck.) Elix & Lumbsch
– Lipnicki [1993, as Melanelia acetabulum
(Neck.) Essl.]; loc. 10: Acpl, Acps. [EN]
polysporinA simplex (Davies) Vězda – Lipnicki
(1993): gr.
#polysporinA subfuscescens (Nyl.) K. Knudsen &
Kocourk. – loc. 13: li (on Acarospora fuscata).
porinA AeneA (Wallr.) Zahlbr. – Lipnicki (1993);
loc. 2–4, 6, 10, 11, 15: Acpl, Acps, Cab, Cor,
Fas, Fre.
porinA chloroticA (Ach.) Müll. Arg. – Lipnicki
(1993); loc. 10: br, gr.
porpidiA crustulAtA (Ach.) Hertel & Knoph – Lip-
nicki (1993): gr.
porpidiA soredizodes (Lamy ex Nyl.) J.R. Laun-
don – Lipnicki (1993); loc. 6: gr.
porpidiA tuberculosA (Sm.) Hertel & Knoph –
Lipnicki (1993); loc. 10: gr.
protopArmeliA oleAginA (Harm.) Coppins – loc.
4: Fas.
pseudeverniA furfurAceA (L.) Zopf – Hillmann
[1932–1933, as Parmelia furfuracea (L.)
Ach. var. ceratea (Ach.) Th. Fr.], Hillmann &
Grummann (1957, as P. f. var. soreumatica
Wallr. and P. f. var. ceratea), Lipnicki (1993);
loc. 4, 5, 12, 15: Acpl, Be, Fas, Pis, Qu, wo.
pycnorA sorophorA (Vain.) Hafellner – loc. 12:
Pis, wo.
pyrenulA nitidA (Weigel) Ach. – Lipnicki (1993);
loc. 3, 4, 15: Cab, Fas, Ti. [VU]
rAmAlinA fArinAceA (L.) Ach. – Lipnicki (1993);
loc. 5, 10: Acpl, Acps, Po, Ti. [VU]
rAmAlinA fAstigiAtA (Pers.) Ach. – Lipnicki (1993);
loc. 10: Ac, Acps, Po. [EN]
rAmAlinA frAxineA (L.) Ach. – loc. 10: Ac, Acpl,
Fas. [EN]
rAmAlinA pollinAriA (Westr.) Ach. – Lipnicki
(1993): Ac, Po. [VU]
rhizocArpon distinctum Th. Fr. – Hillmann [1932–
1933, as R. ambiguum (Schaer.) Zahlbr.],
Hillmann & Grumman (1957), Lipnicki
(1993); loc. 13: gr.
rhizocArpon reductum Th. Fr. – Lipnicki (1993,
as R. obscuratum auct.); loc. 6: co, gr.

69
rinodinA oleAe Bagl. – Lipnicki (1993, as Rino-
dina gennarii Bagl.): br.
ropAlosporA viridis (Tønsberg) Tønsberg – loc. 3,
4, 8, 11, 15, 17: Ali, Cab, Coa, Fas.
sArcogyne regulAris Körb. – Lipnicki (1993): co.
+sAreA resinAe (Fr.) Kuntze – loc. 6: Pic.
scoliciosporum chlorococcum (Graewe ex
Stenh.) Vězda – Lipnicki (1993); loc. 4, 5:
Fas, Qu, wo.
scoliciosporum sArothAmni (Vain.) Vězda – loc.
5, 12: Qu, Ti.
scoliciosporum umbrinum (Ach.) Arnold – Lipnicki
[1993, also as Micarea intrusa (Th. Fr.) Cop-
pins & Kilias]; loc. 13: gr.
+stenocybe pullAtulA (Ach.) Stein – loc. 8: Ali.
stereocAulon condensAtum Hoffm. – Hillmann
(1932–1933): so. [VU]
stereocAulon tomentosum Fr. – Lipnicki (1993):
so. [EN]
strAngosporA moriformis (Ach.) Stein – Lipnicki
(1993): wo.
#syzygosporA physciAceArum Diederich – loc. 6, 8,
9: li (on Physcia adscendens and Ph. tenella).
#tAeniolellA punctAtA M.S. Christ. & D. Hawksw.
– loc. 11: li (on Graphis scripta).
thelocArpon intermediellum Nyl. – loc. 12: wo.
[VU]
thelocArpon lAureri (Flot.) Nyl. – Lipnicki (1993):
gr.
#thelocArpon lichenicolA (Fuckel) Poelt & Hafell-
ner – loc. 11: wo.
trApeliA coArctAtA (Turner ex Sm.) M. Choisy –
Lipnicki (1993): gr.
trApeliA glebulosA (Sm.) J.R. Laundon – Lip-
nicki [1993, as Trapelia involuta (Taylor)
Hertel]: br.
trApeliopsis flexuosA (Fr.) Coppins & P. James
– Lipnicki (1993); loc. 3, 4, 6, 9, 11, 12, 17:
Be, Pic, Pis, Qu, wo.
trApeliopsis glAucolepideA (Nyl.) Gotth. Schneid.
– loc. 2, 17: tu, wo. [DD]
trApeliopsis grAnulosA (Hoffm.) Lumbsch – Lip-
nicki (1993); loc. 4–6, 9, 11, 12: Be, de, Pis,
so, wo.
trApeliopsis pse udogr Anul osA Coppins & P.
James – loc. 4, 11: so, wo.
#tremellA hypogymniAe Diederich & M.S. Christ.
– loc. 5, 15: li (on Hypogymnia physodes).
#tremellA lichenicolA Diederich – loc. 11, 15: li
(on Violella fucata).
#trichonectriA AnisosporA (Lowen) van den Boom
& Diederich – loc. 9: li (on Hypogymnia
physodes). The species is new to Poland.
tuckermAnopsis chlorophyllA (Willd.) Hale – Lip-
nicki [1993, as Cetraria chlorophylla (Willd.)
Vain.]; loc. 5, 6, 8, 10, 12, 17: Acps, Ali, Cab,
Fas, Ma, Pis, Qu. [VU]
usneA dAsypogA (Ach.) Nyl. – Hillmann & Grum-
mann [1957, as U. dasypoga var. tuberculata
(Motyka) Räsänen]: Fas. [VU]
usneA floridA (L.) Weber ex F.H. Wigg. – Hill-
mann (1932–1933), Hillmann & Grummann
(1957), Schulz-Korth (1931): Fas, Pis. [CR]
usneA hirtA (L.) Weber ex F.H. Wigg. – Hillmann
(1932–1933), Hillmann & Grummann
(1957): Pis. [VU]
usneA subfloridAnA Stirt. – Hillmann & Grum-
mann [1957, as Usnea comosa (Ach.) Röhl.
var. similis (Motyka) Erichsen], Tobolewski
(1988): Fas. [EN]
verrucAriA dolosA Hepp. – loc. 8: gr.
verrucAriA hydrelA Ach. – loc. 3, 8, 15: gr. [VU]
verrucAriA murAlis Ach. – Lipnicki (1993, also
as V. rupestris Schrad. and V. conuens A.
Massal.); loc. 6: co.
verrucAriA nigrescens Pers. – Lipnicki (1993);
loc. 6, 10: br, co.
verrucAriA prAetermissA (Trevis.) Anzi – loc. 3,
15: gr. [NT]
verrucAriA sublobulAtA Eitner ex Servít – loc.
3: gr, sa. [DD]
violellA fucAtA (Stirt.) T. Sprib. [syn. Mycoblas-
tus fucatus (Stirt.) Zahlbr.] – loc. 3, 4, 8, 11,
15: Cab, Fas, wo.
vulpicidA pinAstri (Scop.) J.-E. Mattsson & M.J.
Lai – Lipnicki [1993, as Cetraria pinastri
(Scop.) Ach.]: wo. [NT]
xAnthomendozA ulophyllodes (Räsänen) Søcht-
ing, Kärnefelt & S. Y. Kondr. – loc. 10: Acpl.
[VU]
xAnthopArmeliA conspersA (Ehrh. ex Ach.) Hale
– Lipnicki (1993): gr.
xAnthopArmeliA loxodes (Nyl.) O. Blanco et al. –
Lipnicki [1993, as Neofuscelia loxodes (Nyl.)
Essl.]: co.
xAnthopArmeliA cf. pullA (Ach.) O. Blanco et al.
– loc. 15: gr. [NT]
xAnt horiA c Andel AriA (L.) Th. Fr. – Lipnicki
(1993): bark of deciduous roadside trees.
xAnthoriA elegAns (Link) Th. Fr. – loc. 10: co.
xAnthoriA pArietinA (L.) Th. Fr. – Lipnicki (1993);
loc. 1, 4, 6, 8: co, Fas, Paa, San.
xAnthoriA polycArpA (Hoffm.) Rieber – loc. 12: Ti.

70 Folia Cryptog. Estonica
Excluded species
Arthonia exilis (Flörke) Anzi – Lipnicki (1993):
Cab. Taxonomical classication could not
be claried.
Cladonia bellidiora (Ach.) Schaer. – Lipnicki
(1993): so. Determination considered to be
doubtful.
Cladonia pyxidata (L.) Fr. – Lipnicki (1993): so.
The records most probably belong to other
morphologically similar species.
Coenogonium cf. luteum (Dicks.) Kalb & Lück-
ing – Lipnicki [1993, as Dimerella cf. lutea
(Dicks.) Trevis]: Cab. Determination needs
re-examination.
Haematomma ochroleucum (Neck.) J.R. Laundon
var. porphyrium (Pers.) J.R. Laundon – Lip-
nicki (1993): Fas. Determination considered
to be doubtful; most Polish records of this
lichen belong to Lecanora thysanophora R.C.
Harris (Kukwa 2005).
Lecanora populicola (DC.) Duby – Lipnicki
(1993): Cab. Determination needs re-
examination; the species usually grows on
Populus tremula.
Lepraria membranacea (Dicks.) Vain. – Hillmann
[1932–1933, as Crocynia membranacea
(Dicks.) Zahlbr.], Lipnicki [1993, Leproloma
membranaceum (Dicks.) Vain.]: co, bark of
deciduous trees. Determinations considered
to be doubtful; this is a montane species in
Poland (Kukwa 2006).
Lepraria neglecta (Nyl.) Lettau – Lipnicki (1993):
co. Determination needs re-examination;
it is mostly a montane species, which was
often incorrectly reported from lowland
(Kukwa 2006).
Ochrolechia androgyna (Hoffm.) Arnold – Lip-
nicki (1993): Be, Qu. Determination con-
sidered to be doubtful; the records belong
most probably to O. bahusiensis, which
was recently proved to be distinct from O.
androgyna (see Jabłońska & Kukwa 2007).
Opegrapha subparallela Müll. Arg. – Lipnicki
(1993): wo. Taxonomical classication could
not be claried.
Pertusaria lactea (L.) Arnold – Lipnicki (1993):
co. Determination considered to be doubtful;
it is a montane species in Poland.
Pertusaria maculata Erichs. – Lipnicki (1993):
Coa. This name has been applied in Poland
to many different species; the determination
needs re-examination.
Verrucaria athiobola Ach. – Lipnicki (1993, as V.
laevata Ach.): gr. Name of unclear identity.
Verrucaria elaeomelaena (A. Massal.) Arnold –
Lipnicki (1993): gr. Determination needs
re-examination.
ACKNOWLEDGMENTS
We thank Dorota Janicka and Agata Suchta
from the Regional Direction for Protection of the
Environment in Szczecin (Regionalna Dyrekcja
Ochrony Środowiska w Szczecinie) for mak-
ing possible the investigations. Furthermore,
we are grateful to Elżbieta Hołubczat from the
Drawieński National Park for the permission for
lichen investigations and arranging the visits.
We thank also Beata Krzewicka (Kraków) for
the identication of Verrucaria sublobulata and
Piret Lõhmus (Tartu) for determining Calicium
glaucellum.
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72 Folia Cryptog. Estonica