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Fauna Ryukyuana ISSN 2187-6657
http://w3.u-ryukyu.ac.jp/naruse/lab/Fauna_Ryukyuana.html
1
Taxonomic notes on two species of xanthid crabs of the genera
Hepatoporus Serène, 1984 and Gaillardiellus Guinot, 1976 from the Ryukyu Islands
Masahiro Marumura1, 3 & Masatsune Takeda2
13262 Shinjo-cho, Tanabe, Wakayama, 646-0011 Japan
2Department of Zoology, National Museum of Nature and Science,
4–1–1 Amakubo, Tsukuba, Ibaraki, 305-0005 Japan
3Corresponding author (mmarumura@ares.eonet.ne.jp)
Abstract. This report deals with two crab species of
the family Xanthidae, Hepatoporus sp. and
Gaillardiellus bathus Davie, 1997, with some
taxonomic comments and photographs. A female
specimen of the genus Hepatoporus from the
Kerama Group in the Ryukyu Islands is similar to H.
pumex Mendoza & Ng, 2008 reported from the
Bohol Sea, the Philippines, and also to H. orientalis
(Sakai, 1935) known from Japan and the Philippines.
The female specimen was compared in detail with
the original description of H. pumex and with some
specimens of H. orientalis. We, however, did not
identify it to species as we could not assess the range
of intraspecific variation or male characters.
Gaillardiellus bathus, previously known from New
Caledonia and the Kermadec Islands in the South
Pacific, is recorded from Okinoerabu-jima Island in
the Ryukyu Islands, the Kii Peninsula and the
Ogasawara Islands. Comparative notes on G. bathus
and allied G. rueppelli (Krauss, 1843) are provided.
Introduction
Continuous discoveries of undescribed and newly
recorded species of the shallow-water crabs from the
Ryukyu Islands suggest that the knowledge of the
crab fauna in this region is still insufficient, probably
due to the difficulty of collecting specimens from the
topographically complicated seabed. Efforts to
collect specimens and records of these specimens are
without doubt important in bringing the faunistic
knowledge of Japan to a satisfactory level. For
example, systematically and biogeographically
interesting crabs from the vicinity of the Kii
Peninsula have been reported by Marumura (1984a,
1984b, 1985, 1994), Marumura & Manabe (1996),
Marumura et al. (2000), Takeda & Marumura (1994,
1995, 1996, 1997a, 1997b, 2000, 2002), and these
data have contributed to expanding knowledge of the
carcinological fauna of central Japan influenced by
the warm Kuroshio Current. In this paper, two
species of the family Xanthidae from the Ryukyu
Islands are recorded. One species is Hepatoporus sp.
from Tokashiki Island in the Kerama Group, Ryukyu
Islands, which is similar to H. pumex Mendoza & Ng,
2008 reported from the Philippines and also to H.
orientalis (Sakai, 1935) known from Japan and the
Philippines. In the present paper, the identification of
a female specimen at hand is restricted to the genus,
and the definite identification to the species will be
performed in due time after acquisition of a male
specimen. The other species is Gaillardiellus bathus
Davie, 1997 from Okinoerabu-jima Island in the
Ryukyu Islands, Wakayama Prefecture on the
southwestern coast of the Kii Peninsula, and the
Ogasawara Islands. This species is known from the
South Pacific, and these records extend its
distributional range further into the North Pacific.
The specimens examined are deposited in the
Department of Zoology (NSMT-Cr) and Showa
Memorial Institute (NSMT-R), Tsukuba Research
Institute of the National Museum of Nature and
Science, Tokyo, and the Wakayama Prefectural
Museum of Natural History (WMNH-Na-Cr).
Taxonomic Notes
Family Xanthidae MacLeay, 1838
Genus Hepatoporus Serène, 1984
Hepatoporus sp.
(Fig. 1)
Material examined. Ryukyu Islands. Naganita,
Tokashiki Island, Kerama Group, 1 female (15.0 ×
11.3 mm), NSMT-Cr 22975, 10 Oct. 1993, coll. S.
Nagai.
Comparative material. Hepatoporus orientalis
(Sakai, 1935). Ryukyu Islands: Oshima Passage,
Amami-Oshima Island, 1 young male (6.4 × 5.1 mm),
WMNH-Na-Cr 0713, June 1996, coll. Nagai, and
recorded by Marumura & Kosaka (2003). Sagami
Bay: Kannonzuka-dashi–Maruyama-dashi, 65 m
deep, 1 male (10.0 × 7.7 mm: Figs. 3A, B, 4A, B, D),
NSMT-R 2858, 21 Jul. 1958; SSW off Jyogashima,
15 km, 77–80 m deep, 1 male (10.8 × 8.4 mm: Figs.
3C, D, 4C), NSMT-R 3292, 25 Jul.1959; Maruyama-
dashi–Kannonzuka-dashi, 60–85 m deep, 1 male (8.1
× 6.1 mm), NSMT-R 3001, 8 Jun. 1960. Kii Penin-
2
2
Fig. 1. Hepatoporus sp., female (NSMT-Cr22975; 15.0 × 11.3 mm). A, dorsal view; B, frontal views; C, right chela; D,
subhepatic cavity; E, third maxillipeds; F, right half of carapace, with right cheliped; G, thoracic sternum.
1 (NSMT-Cr22975; 15.0 × 11.3 mm). A, ; B, ; C, ; D, ; E, 3
; F, ; G, .
sula: Off Tanabe Bay, Wakayama Pref., ca. 50m deep,
1 female (9.6 × 7.3 mm: Fig. 2), WMNH-Na-Cr,
unregistered, 20 Dec. 1997, coll. M. Marumura; Off
Cape Shiono-misaki, Wakayama Pref., 50–60m deep,
1 female (7.5 × 5.4 mm), 1 young female (5.3 × 4.1
mm), WMNH-Na-Cr, unregistered, Aug. 1988, coll.
S. Nagai.
Description. Carapace (Fig. 1A) broadly
subhexagonal, convex dorsally as a whole; dorsal
surface of carapace uneven, eroded with many pits
and irregular reticulations of variable sizes,
separated into regions by broad depressions;
epigastric (1M), protogastric (2M), mesogastric
(3M), metagastric (4M), cardiac (1P), branchial (L)
Fauna Ryukyuana, 27: 1–11.
3
[Record] Marumura & Takeda: Two xanthid crabs from the Ryukyu Islands
Fig. 2. Hepatoporus orientalis (Sakai), female (WMNH-Na-Cr, unregistered; 9.6 × 7.3 mm). A, dorsal view; B, frontal
view; C, subhepatic cavity; D, ventral view; E, third maxillipeds; F, thoracic sternum.
2 (WMHN-Na-Cr, unregistered; 9.6 × 7.3 mm). A, ; B, ; C, ; D, ; E, 3
; F, .
regions demarcated; 2M most prominent, high,
subacute at tip; L shallowly subdivided into 2
oblique subregions, posterior subregion as high as,
slightly becoming sharper than 2M; 4M linear
transversely, franked by deep depressions.
Front (Fig. 1A–D) deflexed, bilobed, thin.
Subhepatic cavity (Figs. 1B–D) very deep, sharply
edged along whole margin, not visible from above,
occupying most of subhepatic region. Anterolateral
margin of carapace narrowly cristate; anterior
4
2
onethird weakly concave anterolaterally, weakly
convex dorsally; posterior twothirds regularly
convex, with a row of minute granules;
posterolateral margin of carapace about half as long
as anterolateral margin, strongly turned to lateral end
of posterior margin of carapace, with deeply
excavated dorsal surface to receive last ambulatory
leg. Posterior margin of carapace weakly concave,
with deep concavity adjacent to lateral end for
accommodating coxa of last ambulatory leg.
Both chelipeds (Fig. 1A) equal in size and shape,
with uneven surfaces as seen in dorsal surface of
carapace; merus short, entirely disguised under
carapace, with wholly excavated inner surface;
carpus large, with depressed subtruncated tubercle at
outer margin (Fig. 1A, B, F); outer surface of palm
(Fig. 1C, D) distinctly reticulated, distal margin more
or less nodular, with prominent compressed tubercle
at upper part (Fig. 1B, C, F). Fingers (Fig. 1C, D) as
long as upper margin of palm, cutting edge with 4
sharp, subequal teeth directed obliquely outward in
parallel with tip of finger.
Ambulatory legs (Fig. 1A) tightly folded against
carapace; anterior margin of each merus thin, nearly
entire or microscopically toothed, with angulated
distal end; posterior upper and lower margins of each
merus narrowly, but distinctly ridged and granulated;
upper and lower margins of each carpus weakly
ridged, roughened with sharp granules, with two
longitudinal ridges on upper surfaces; upper and
lower margins of each propodus sharp, minutely
granulated like carpus, but granules smaller.
Color in life. Carapace, chelipeds and ambulatory
legs regularly creamy white.
Remarks. A female Hepatoporus specimen from
Tokashiki Island can be allied to H. pumex Mendoza
& Ng, 2008 and H. orientalis (Sakai, 1935) among
five congeners (Ng et al, 2008; Mendosa & Ng, 2008)
in the general shapes of the carapace with the raised
gastric, branchial and cardiac regions and the deep
subhepatic cavity.
Hepatoporus pumex was described from the
Bohol Sea, the Philippines. In the original description
of H. pumex, Mendosa & Ng (2008: 402) raised five
diagnostic characters to distinguish the new species
from morphologically most similar H. orientalis, viz.
1) broader, more truncated front (vs. triangular and
acuter), 2) more deeply excavated anterolateral
margin (vs. less concave), 3) more even posterior
two-thirds of the anterolateral margin of the carapace
(vs. more irregular and jagged), 4) the presence of a
large, distinct pit in the branchial region of the
carapace (vs. absent), 5) reticulate patterns of fused
granules and pits near the posterolateral and posterior
margins of the carapace (vs. simply granular).
Hepatoporus orientalis was originally described
by Sakai (1935) from Sagami Bay. Subsequently the
species has been repeatedly reported by Sakai (1936,
1939, 1965, 1976) with original figures, but detailed
morphological information has still not been
provided. The present study could examine several
specimens of H. orientalis collected from the type
locality and some other stations in Japanese waters
(Figs. 2–4).
Detailed comparison of the specimens of H.
orientalis and the original description of H. pumex
revealed that there are some additional differences
between the two species. They are: 1) the subhepatic
cavity of H. orientalis is deeper anteriorly with sharp
cavity margin and becomes shallower posteriorly,
without clear margin (Figs. 2C, 3B). The subhepatic
cavity of H. pumex is, however, markedly deep
throughout, with an entire sharp margin (Mendosa &
Ng, 2008: fig. 7E). 2) The outer margin of the
cheliped carpus is thick and roundly convex in H.
orientalis (Figs. 2A, 3A, C) but that of H. pumex is
crested and bilobed (Mendosa & Ng, 2008: Figs.7A,
9D). 3) The third maxilliped ischium and merus are
thickly covered with large granules in H. orientalis
(Fig. 2E), but the outer surface of the third
maxilliped is described and figured as being eroded
and pitted, with scattered granules in H. pumex
(Mendosa & Ng, 2008: Fig. 8B). 4) The exopod of
the third maxilliped gently tapers in H. orientalis
(Fig. 2C, E), but that of H. pumex tapers over distal
half (Mendosa & Ng, 2008: Figs. 7B, 8B). 5) The
male first pleopod of H. orientalis from Sagami Bay
(Fig. 4A–C) has a sharply pointed end, with a
subterminal cluster of some stout recurved setae,
differing from that of H. pumex having the
auriculiform distal end, with a subterminal flange
(Mendosa & Ng, 2008: Fig. 8D–F).
The female specimen from the Ryukyu Islands
may be indentified as H. pumex due to the basic
agreement in the regularly convex posterior two-
thirds of the anterolateral margin of the carapace,
deeply excavated subhepatic cavity, with a sharp,
entire margin, the crested and bilobed outer margin
of the cheliped carpus, and the exopod of the third
maxilliped being abruptly tapered distally. However,
a detailed comparison revealed that there are some
discrepancies: 1) The surfaces of the carapace and
chelipeds of the holotype (Mendosa & Ng, 2008: Fig.
7A) appear to be smoother than in the present
specimen (Fig. 1A); 2) The subhepatic cavity of the
holotype (Mendosa & Ng, 2008: Fig. 7C, E) seems
to be shallower than that in the present specimen (Fig.
1B, D); 3) The anterior part of the anterolateral mar-
Fauna Ryukyuana, 27: 1–11.
5
[Record] Marumura & Takeda: Two xanthid crabs from the Ryukyu Islands
Fig. 3. Hepatoporus orientalis (Sakai). A, B, male (NSMT-R2858; 10.0 × 7.7 mm); C, D, male (NSMT-R3292; 10.8 ×
8.4 mm).
3. A, B, (NSMT-R2858; 10.0 × 7.7 mm); C, D, (NSMT-R3292; (10.8 × 8.4 mm).
gin of the carapace, viz. the upper margin of the
subhepatic cavity, is concave in dorsal view on the
carapace margin in the holotype (Mendosa & Ng,
2008: Fig. 7A), but in the present specimen this part
is hardly concave and even convex (Fig. 1A, F); 4)
The third maxilliped of the holotype is figured as
being poorly granulated (Mendosa & Ng, 2008: Fig.
8B), but covered with prominent pearly granules in
the present specimen (Fig. 1E); 5) The armature of
the cheliped carpus is rather nodular in the holotype
(Mendosa & Ng, 2008: Fig. 7A), whereas tubercular
and high in the present specimen (Fig. 1E). It is
uncertain whether these discrepancies are due to
interspecific differences, or intraspecific variations.
The present report treats this Ryukyuan specimen as
Hepatoporus sp.
Genus Gaillardiellus Guinot, 1976
Gaillardiellus bathus Davie, 1997
[New Japanese name: Minami-kebuka-
awatsubugani]
(Figs. 4E, F, 5A–D)
Material examined. Ryukyu Islands: Off
Okinoerabu-jima Island, 177m deep, R/V Toyoshio
Maru cruise TY-04-05, 21 May 2004, 1 male (15.2 ×
11.5 mm), 1 female (10.4 × 7.6 mm), NSMT-Cr
16182, coll. M. Osawa. Kii Peninsula: Off Tanabe,
Wakayama Pref., ca. 100 m deep, 15 Jan. 1993, 1
male (15.5 × 12.0 mm), NSMT-Cr 23915, coll. M.
Marumura; Off Kirime, 70–80 m deep, 2 Feb. 1996,
1 ovigerous female (18.0 × 12.9 mm), NSMT-
Cr23000, coll. M. Marumura; Off Iwashiro, ca. 120
m deep, 6 Mar. 2001, 1 male (18.6 × 14.0 mm), 1
female (11.0 × 8.0 mm), NSMT-Cr 22974, coll. M.
Marumura. Ogasawara Islands: Yabe guyot (Shiba,
1979), seamount far off to the east of Ogasawara Is.
(27°15.7′ N, 145°11.4′ E, 110 m – 27°15.8′ N,
145°11.7′ E, 150 m deep), R/V Soyo Maru, 1990
cruise to Ogasawara Is., sta. 5, 8 July 1990, 1 male
(9.9 × 7.3 mm), NSMT-Cr 22990, coll. H. Saito.
Comparative material. Gaillardiellus rueppelli
(Krauss, 1843). Ryukyu Islands: Oshima Passage
between Amami-Oshima Island and Kakeroma-jima
Island, 25–40 m deep, 1 male (12.0 × 9.2 mm), 1
6
2
Fig. 4. A–D, Hepatoporus orientalis (Sakai). A, left first pleopod of male (NSMT-R 2858; 10.0 × 7.7mm) in abdominal
view; B, distal part of the same in sternal view; C, distal part of left first pleopod of male (NSMT-R3292; 10.8 × 8.4 mm)
in abdominal view; D, left second pleopod of male (NSMT-R2858; 10.0 × 7.7 mm) in abdominal view. E, F, Gaillardiellus
bathus Davie. E, distal part of left first pleopod of male (NSMT-Cr23915; 15.5 × 12.0 mm) in abdominal view; F, overall
view of the same in sternal view. Scales for A, E, F = 1 mm; B, C, D = 0.5 mm.
4. A–D, . A, B, (NSMT-R 2858; 10.0 × 7.7 mm) (A) (B)
; C, (NSMT-R 3292; 10.8 × 8.4 mm) , ; D, (NSMT-R 2858; 10.0 × 7.7
mm) 2, ; E, F, () (NSMT-Cr 23915; 15.5 × 12.0 mm)
1 (E) (F)[ A, E, F1 mm; B, C, D = 0.5 mm.
Fauna Ryukyuana, 27: 1–11.
7
[Record] Marumura & Takeda: Two xanthid crabs from the Ryukyu Islands
female (12.0 × 8.9mm), NSMT-Cr 9712, 29 June
1970, coll. Kagoshima Univ. Kii Peninsula: Off
Koza, Wakayama Pref., 20–30 m deep, 1 male (34.3
× 25.8 mm; Fig. 5E, F), NSMT-Cr10869, 13 Dec.
1987, S. Nagai leg. Boso Peninsula: Mera-se,
submarine bank off Boso Penin., R/V Tansei Maru
cruise KT-76-16, stn. C9 (35°50.5′N, 139°45.1′E,
100–102 m deep), 1 male (9.5 × 6.9 mm), NSMT-
Cr15509, Sept. 1976, R/V Tansei Maru. Seychelles
(04°29.2′S, 56°10.6′E, 63 m deep), 1 male (NSMT-
Cr4373), 22 Nov. 1968, R/V Koyo Maru.
Description. Carapace (Fig. 5A, B) transversely
subhexagonal, wider than long, weakly convex
longitudinally and transversely; regions distinctly
separated from each other by narrow, shallow
smooth furrows; each region rather flattened,
covered with regularly dispersed pearly granules of
equal size interspaced with short thick tomentum and
some scant short setae; epigastric region (1M)
weakly convex dorsally and laterally, being hardly
separated from anteromesial part of protogastric
region (2M); 2M almost, but incompletely,
subdivided into 2 by longitudinal sulcus, lateral
subregion slightly longer than mesial subregion;
mesogastric (3M) and cardiac (1P) regions
prominent, not subdivided; branchial region (L)
typically subdivided into 4 parts; anterior two (2L,
3L) on lateral part of 2M placed side by side,
posterior two (4L, 5L) on lateral to 3M placed
obliquely.
Frontal margin (Fig. 5A, B) divided into 2 lobes
by median V-shaped notch in dorsal view; each lobe
strongly produced downwards as rounded lobe in
inner half in frontal view, deeply concave upwards
sublaterally.
Anterolateral margin of carapace (Fig. 5B)
divided into 4 obtuse, convex lobes by shallow
depressions. Posterolateral margin of carapace (Fig.
5B) strongly retreated. Posterior margin of carapace
(Fig. 5B) as wide as frontal margin, with shallow
concavity at each lateral end to accommodate coxa
of last ambulatory leg.
In male (Fig. 5) and ovigerous female, both
chelipeds stout, not long, subequal in shape and size;
merus short, nearly obscured beneath carapace;
carpus prominently large, as long as palm and fingers
combined, its inner surface shallowly concave to fit
subhepatic region of carapace, with outer surface cut
into some obtuse nodules; outer surface of palm (Fig.
5C) covered with prominent pearly granules; upper
margin of palm divided into 2 humps, proximal one
rather sharp along margin, with obtusely angulated
basal end; male fingers armed with strong, subacute
teeth, 2 in immovable finger and 3 in dactylus; inner
surfaces of fingers and teeth deeply excavated in
both sexes, but especially so in male; dark color of
immovable finger extended onto half of lower outer
surface of palm in male (Fig. 5C), restricted to
immovable finger proper in female.
Ambulatory legs stout, covered with pearly
granules and short setae; both margins of each
segment fringed with larger granules and longer, but
sparse setae; dactyli covered with thick tomentum;
upper surface of each carpus sculptured with
longitudinal furrow along anterior margin.
Male telson (Fig. 5D) longer than sixth
abdominal somite, ca. 1.3 times wider than long;
sixth somite rectangular, ca.1.7 times wider than
long; third to fifth somites fused, with vestigial
sutures; lateral margins sinuous, bluntly angled at
supposed junction of fourth and fifth somites; first
somite as wide as third.
Male first pleopod (Fig. 4E, F) slender, weakly
curved outward, distal part long, sharply pointed at
tip; shaft of pleopod with many conical granules at
subdistal part of outer margin; tuft of plumose setae
at subterminal part of inner margin reaching tip of
distal part, mesial margin of shaft behind plumose
setae with line of sharp, equidistantly placed small
tubercles.
Color in life. Carapace, chelipeds and
ambulatory legs regularly brick red.
Remarks. The specimens examined in the
present study (four males, two females and one
ovigerous female from some localities in Japanese
waters) share a similar shape of the carapace that is
deeply sculptured and densely covered with short
stiff setae and pearly granules of good size on the
dorsal surface.
These carapace characters as well as the
characteristics of its male first pleopod indicate that
it belongs to the genus Gaillardiellus Guinot, 1976.
Gaillardiellus currently contains six species (five
known species listed by Ng et al., 2008, and a new
species described by Takeda & Komatsu, 2010).
The Japanese specimens in question is seemingly
close to Gaillardiellus ruepplli (Krauss, 1843),
which is widely distributed in the Indo-West Pacific
including Japanese waters, and to G. bathus Davie,
1997 known from New Caledonia and the Kermadec
Islands in the South Pacific. According to Davie
(1997: 341), G. bathus differs from G. rueppelli most
obviously by the shape of the male abdomen with its
broad telson and sixth somite, which is noticeably
broader than long and not subquadrate as in G.
rueppelli. It is otherwise mentioned that the dorsal
areolation of the carapace of G. bathus is lower and
less strongly defined, with the shallower interregion-
8
2
Fig. 5. A–D, Gaillardiellus bathus Davie. Male (NSMT-Cr22974; 18.6 × 14.0 mm). A, frontal view; B, dorsal view; C,
chelae; D, ventral view. E, F, Gaillardiellus rueppelli (Krauss). Male (NSMT-Cr 10869; 34.3 × 25.8 mm). E, dorsal
view; F, ventral view.
5A–D, (). NSMT-Cr 22974; 18.6 × 14.0 mm A, ; B, ; C,
; D, . E, F, (NSMT-Cr 10869; 34.3 × 25.8 mm)E, ; F,
al grooves, the granulation is slightly finer, the
transverse groove between sternites 3 and 4 is less
deeply marked, and the chela has a strong bluntly
rounded tooth on the cutting margin of the
immovable finger.
The specimens examined in the present study can
be identified as G. bathus as they have the above-
mentioned characteristics of the species. It is
otherwise noted that in G. rueppelli the anterolateral
teeth of the carapace are more strongly angulated and
weakly directed dorsally. These observations justify
the identification of the present specimens with G.
bathus, and warrant the occurrence of the two
species, G. rueppelli and G. bathus in Japanese
waters.
Direct comparison of the Japanese specimens of
the two species revealed that the male abdomen of G.
bathus (Fig. 5D) is apparently wider than that of G.
rueppelli (Fig. 5F), in addition to the differences of
the sixth somite and telson mentioned by Davie
Fauna Ryukyuana, 27: 1–11.
9
[Record] Marumura & Takeda: Two xanthid crabs from the Ryukyu Islands
(1997: 341). In the original figure of G. bathus
(Davie, 1997: Fig. 1d), the lateral margin of the
supposed third segment of the male abdomen was
figured as convex as a whole, with proximolateral
granulation. The lateral margin of the male abdomen
in the Japanese specimens is similarly convex, but
also extended posterolaterally as usual in other
xanthid crabs, e.g. male abdomen and sternum of G.
rueppelli given by Guinot (1976; fig. 42A).
Distribution. The type locality is New Caledonia,
270–312 m deep (Davie, 1997), and the additional
locality is the Kermadec Islands, 108–198 m deep
(Takeda & Webber, 2006). This report extends the
distributional range from the South Pacific
northward to Japan (Ryukyu Islands, Kii Peninsula
and Ogasawara Islands). The bathymetric range in
Japan is from 70 to 150 m.
Acknowledgments
We wish to thank Dr. H. Saito of the Department of
Zoology, National Museum of Nature and Science,
Tokyo, and Dr. M. Osawa of the Research Center for
Coastal Lagoon Environments, Shimane University,
who offered us the specimens for study. Our cordial
thanks are also due to the authority of the Wakayama
Prefectural Museum of Natural History, Dr. H.
Namikawa of the Showa Memorial Institute and H.
Komatsu of the Department of Zoology, National
Museum of Nature and Science, Tokyo, for the
comparative specimens under their care. Dr. T.
Naruse of the University of the Ryukyus was kind
enough to recommend us to submit the present paper
to Fauna Ryukyuana. The present paper was brushed
up by Dr. J. D. Reimer of the University of the
Ryukyus and anonymous reviewers.
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琉球列島産のオウギガニ科 2 種
丸村眞弘 1,3･武田正倫 2
1〒646-0011 和歌山県田辺市新庄町 3262
2〒305-0005 茨城県つくば市天久保 4-1-1
国立科学博物館 動物研究部
3通信著者 (mmarumura@ares.eonet.ne.jp)
. Hepatoporus sp.
, ,
Gaillardiellus bathus Davie, 1997 3.
Hepatoporus sp.
H. pumex Mendoza & Ng, 2008
H. orientalis (Sakai,1935)
,
(1) H. pumex ,
3. 33
13
3,
,
.
] G. bathus
() 3,
G. rueppelli (Krauss,
1843) 3,
3.
Fauna Ryukyuana, 27: 1–11.
11
[Record] Marumura & Takeda: Two xanthid crabs from the Ryukyu Islands
: 2014 10 22
: 2015 911
: 2015 12 24
