The MUSORSTOM-TDSB deep-sea benthos exploration programme (1976-2012): An overview of crustacean discoveries and new perspectives on deep-sea zoology and biogeography

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The MusorsToM-TDSB deep-sea crusTaceans 13
RICHER DE FORGES B., CHAN T.Y., CORBARI L., LEMAITRE R., MACPHERSON E., AHYONG S. T. & NG P. K. L. —
The Musorstom-TDSB deep-sea benthos
exploration program (1976-2012): An overview of crustacean discoveries and new perspectives on deep-sea zoology and biogeography
, in AHYONG S. T., CHAN
T.-Y., CORBARIL. & NG P. K. L.(eds), Tropical Deep-Sea Benthos 27. Muséum national d’Histoire naturelle, Paris : 13-66 (Mémoires du Muséum national
d’Histoire naturelle ; 204). ISBN : 978-2-85653-692-6.
The MUSORSTOM-TDSB deep-sea benthos exploration
programme (1976-2012): An overview of crustacean
discoveries and new perspectives on deep-sea zoology and
biogeography
Bertrand Richer de Forges (1), Tin-Yam Chan (2), Laure Corbari (3), Rafael Lemaitre (4),
Enrique Macpherson (5), Shane T. Ahyong (6) & Peter K. L. Ng (7)
(1) Kiwa Consulting - 5, rue Félix Franchette - 98800 Nouméa - New Caledonia
b.richerdeforges@gmail.com
(2) Institute of Marine Biology and Center of Excellence for the Oceans
National Taiwan Ocean University, Keelung 20224 - Taiwan, R.O.C
tychan@mail.ntou.edu.tw
(3) Muséum national d’Histoire naturelle - Département Systématique & Évolution - CP 26 - 43, rue Cuvier - 75005 Paris, France
corbari@mnhn.fr
(4) Smithsonian Institution - National Museum of Natural History - Department of Invertebrate Zoology
MRC 4210 Silver Hill Road, Suitland, MD 20746, USA
lemaitrr@si.edu
(5) Centro de Estudios Avanzados de Blanes (CEAB-CSIC) - C. Acces Cala Sant Francesc 14 - 17300 Blanes, Girona, Spain
macpherson@ceab.csic.es
(6) Australian Museum - 6 College St. - Sydney, NSW 2010, Australia
Shane.Ahyong@austmus.gov.au
(7) Tropical Marine Science Institute and Department of Biological Sciences - National University of Singapore
Kent Ridge - Singapore 119260, Republic of Singapore
peterng@nus.edu.sg
INTRODUCTION
In 1976, the French research vessel “Vauban” was scheduled to travel from France to New Caledonia. rough the
efforts of Alain Crosnier (Director of the Oceanography, ORSTOM), it was decided that the craft take a detour through
the Philippines —this became the first MUSORSTOM cruise. e principal objective of this cruise was to find new

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
14
specimens of the living fossil glypheid
Neoglyphea inopinata
described one year earlier by Jacques Forest and Michèle de
Saint Laurent from a single specimen collected by the U.S. Fish Commission Steamer research vessel “Albatross” in the
begining of the 20th century (Forest 1981, 2006; Richer de Forges & Justine 2006; Bouchet
et al
. 2008). is first French
deep-sea cruise in the Indo-West Pacific was a great success for two reasons. Firstly, nine fresh specimens of
Neoglyphea
inopinata
were collected near Lubang island in a very small area (Figure 1A). e detailed description of this only extant
species of glypheid contributed significantly to our knowledge on the evolution of decapod crustaceans (Schram & Ahyong
2002). Secondly, the trawls in the Philippine archipelago also collected a very rich benthic fauna including many rare and
new species, demonstrating that the exploration of the deep-sea fauna was far from complete.
In the last quarter of the 20th century, our knowledge of the deep-sea fauna was mainly based on the results of
the so-called “Great Expeditions” of preceding decades. ese expeditions, which took place in the era between the
CHALLENGER in 1873 and the GALATHEA in 1952, sampled the oceans on a world-wide scale. ey studied and
mapped the geography and bathymetry of the sea, and brought up a totally new fauna, resulting in the re-writing of
marine zoology (omson 1878; Bruun 1953). However, despite their many significant discoveries, these expeditions
only sampled about 1000 deep-sea stations across the planet. Considering the size of Earth’s oceans, huge areas remained
totally unexplored.
FIGURE 1
The species of extant glypheids
were considered as “living fossils”
supposedly extinct 50 million years
ago.
A
, Neoglyphea inopinata, male,
cl 120 mm, Philippines (photograph
J. Forest).
B
, Laurentaeglyphaea
neocaledonica, female, cl 60.0 mm,
Chestereld Island (photograph J. Lai).
B
A

The MusorsToM-TDSB deep-sea crusTaceans 15
is first MUSORSTOM cruise in 1976 was to be the starting point of a long series of deep-sea cruises in the Indo-
Pacific (Figure 2). Under the leadership of Alain Crosnier, a modern pioneer in deep-sea biodiversity research, these
cruises rewrote the annals of deep-sea biodiversity. e saga of the MUSORSTOM cruises is also intricately linked to
the late Jacques Forest and Michèle de Saint Laurent. eir discovery of, and search for,
Neoglyphea
was the catalyst
that started the whole exploration process. To appreciate the significance of these cruises and explorations, one must
remember that they took place 10 years before the invention of the word “biodiversity” and 30 years before the “CENSUS
OF MARINE LIFE” (CoML) began. With the publication of the 27th volume of scientific results of the MUSORSTOM-
TDSB cruises in 2013, after 37 years of exploration across the Indo-West Pacific, it is timely for a preliminary assessment
of its impact. As noted above, the cruises first started in 1976. As these were guided and inspired by carcinologists like
Crosnier, Forest and de Saint Laurent, it is not surprising that a “carcinological slant” continued over the years. Of all the
animals collected by these cruises, the best studied and best reported on were the crustaceans. Some 33% of all papers
resulting from these cruises were on these crustaceans (Figure 3). It is thus appropriate to summarize how our increased
knowledge of crustaceans resulting from the MUSORSTOM-TDSB program contributed to knowledge of deep-sea biology.
CONTRIBUTIONS OF THE TDSB PROGRAMME TO ZOOLOGY
e programme of Tropical Deep-Sea Benthos (TDSB), formerly MUSORSTOM, is ongoing and very ambitious:
to describe the deep-sea fauna of the tropical Indo-Pacific. e first three cruises in the Philippines (1976, 1980, 1985)
mainly devoted to the catch of
Neoglyphea
showed that even the deep-sea fauna from the area sampled by the U.S. Fish
Commission Steamer research vessel “Albatross”, was still poorly known. e question then was: what about the other
archipelagoes never sampled because they stayed outside of the cruise track of the “Great Expeditions”? What started
in 1976 with the research vessel “Vauban” was soon expanded to other parts of the western Pacific (Richer de Forges
1990). Between 1984 to 1994, 12 cruises totally devoted to the zoological exploration (after 1988 using the R.V. “Alis”),
7 cruises targeting chemical products in the deep-sea animals and 3 cruises focused on resources were realized in the
New Caledonia EEZ (Exclusive Economic Zone). ese cruises captured a huge amount of deep-sea material. From this
FIGURE 2
Updated map of the MUSORSTOM-TDSB cruises.

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FIGURE 3
Publications from the MUSORSTOM-
TDSB programme between 1977 and
2012: total number 1411 of which 468
(33 %) concern Crustacea (grey bars).
100
80
60
40
20
0
Numbers of TDSB publications
1977
1980
1990
2000
2010
FIGURE 4
Histogram from CoML showing the
species discovered during the 10 years
of the program. Crustaceans dominate
(after Ausubel et al. 2010: g. 12).
Average new species
Crustacea crabs, lobsters, shrimps, barnacles
Mollusca squid, octopus, clams, snails, slugs
Pisces sh including sharks
Platyhelminthes atworms
Bacteria & Archea microbes
Annelida segmented worms
Cnidaria sea anemones, corals, jellysh
Porifera sponges
Other invertebrates
Algae
Protists microbial eukaryotes
Tunicata sea squirts
Echinodermata sea stars, sea urchins, sea cucumbers
Nematoda roundworms
Bryozoa mat or ’moss animals‘
Nemertea ribbon worms
Fungi
Mammalia whales
0 50 100 150 200 250 300 350 400 450 500
452
379
136
97
94
84
71
59
56
55
47
31
30
30
26
7
4
<1
Taxonomic group

The MusorsToM-TDSB deep-sea crusTaceans 17
material, deposited primarily in Muséum national d’Histoire naturelle, actively managed by Alain Crosnier for many
years, and later by Philippe Bouchet, more than 1400 scientific articles were published, totally renewing knowledge of the
Tropical Deep-Sea Benthos (Figure 3). For New Caledonia alone, of the 1619 species reported in 2000, 60.7 % were new
to science (Richer de Forges
et al.
2000, 2005). Since 1994 other areas have been extensively sampled: Wallis & Futuna,
Fiji, Tonga, Marquesas, Australes, Vanuatu, Solomons, Indonesia, Papua New Guinea, Taiwan, Philippines, Mozambique
and Madagascar. e collections have covered the Central Pacific, the Melanesian Arc in the Southern Hemisphere,
part of the Southeast Asia in the Northern Hemisphere targeting the “Coral Triangle” and in the Indian Ocean. It has
made possible broad geographical taxonomic and systematic revisions, and for the first time, resulted in well grounded
hypothesis about the origin and evolution of the deep-sea fauna.
Among the deep-sea fauna, crustaceans are dominant across all depths, sometimes with high diversity. e TDSB
programme overlapped and contributed significantly to the CoML initiative. Ausubel
et al.
(2010) observed that crustaceans
dominated the number of new species found during the decade of the CoML, with 452 species named (Figure 4) —
some 20% of all discovered species. Molluscs and fish came out as second and third in diversity. Much of this crustacean
diversity was reported by the carcinologists working on the TDSB collections. Below, we provide a summary of some of
the accomplishments of the TDSB programme.
DISCOVERY SUMMARY AND HIGHLIGHTS OF KEY CRUSTACEAN GROUPS
TRUE CRABS (BRACHYURA)
— e brachyuran crabs are the most diverse of all decapods, with half the known taxa (Ng
et
al.
2008; De Grave
et al.
2009) and over 7000 species. e MUSORSTOM-TDSB collections have been invaluable in the
discovery of hundreds of species, not just in obtaining specimens of new species but also providing valuable comparative
material to resolve the taxonomy of many others. e collections from the first few expeditions were first reported on by
Serène & Vadon (1981) but the few new species described belies this paper’s significance as it was just a preliminary study.
A series of papers by Guinot and others demonstrated that the deep-sea brachyuran fauna was not just rich, but extremely
rich (
e.g.
, see Guinot & Richer de Forges 1981a, b). is was not restricted to the Philippines but in waters off New Caledonia
(see Richer de Forges & Ng 2006) and even Taiwan (
e.g.
, see Ho
et al
. 2004). e expedition material has been the basis of
a large number of studies in the following families: Atelecylidae (Guinot 1989b); Cancridae (Davie 1991), Calappidae and
Leucosiidae (Chen 1989, 1990; Tan & Ng 1993, 1995; Galil 1997, 2003; Galil & Ng 2007, 2009, 2010; Ng & Richer de Forges
2007), Dorippidae (Chen 1986, 1993), Dromiidae (McLay 1993, 2001; McLay & Ng 2005, 2007), Ethusidae (Chen 1997,
2000; Ng & Ho 2003; Castro 2005; Naruse
et al
. 2009), Euryplacidae (Ng & Castro 2007; Castro & Ng 2010a), Goneplacidae
(Guinot 1989a, 1990; Castro 2007, 2009, 2013; Castro & Ng 2010b), Latreilliidae (Castro
et al
. 2003), various families in the
Majoidea (Guinot & Richer de Forges, 1982a, b, 1986; Tavares 1991a; Poupin 1995; Ahyong & Ng 2007; Richer de Forges
& Ng 2007b, 2008a, 2009a, b, 2012, 2013; Richer de Forges 2010; Richer de Forges & Corbari 2012; Ng & Richer de Forges
2013), Mathildellidae (Crosnier & Ng 2004), Parthenopidae (Tan & Ng 2007a, b; McLay & Tan 2009), Pilumnidae (Chia &
Ng 1995, 2000; Komai & Motoh 2012), Plagusiidae (Crosnier 2001), Pinnotheridae (Ahyong & Ng 2008), Portunidae (Moosa
1996; Crosnier 2002; Davie & Crosnier 2006; Nguyen & Ng 2010), Progeryonidae (Castro & Ng 2008), Trapeziidae (Castro
et al
. 2004), Trichopeltarionidae (Tavares & Cleva 2010), and Xanthidae (Davie 1993, 1997; Ng & Chia 1994; Ng & Naruse
2007b; Manuel-Santos & Ng 2007; Mendoza & Ng 2008a, b, 2010; Mendoza 2012; Mendoza & Manuel-Santos 2012).
Among the Brachyura, members of the family Homolidae are interesting, with some deep-sea taxa even displaying what
seems like “a human face” on the dorsal shell (
e.g.
,
Lamoha personata
, Figure 5A). Almost all are deep-water genera, with only
the genus
Latreillopsis
having some representatives in waters shallower than 100 m. It took five years just to gather the few
specimens collected by the “Great Expeditions” before they could be compared with the MUSORSTOM-TDSB collections.
Some new taxa from the study were written up (Guinot & Richer de Forges 1981c) but a world-wide revision of the family was
eventually published in the MUSORSTOM series (Guinot & Richer de Forges 1995). Before 1995, the family contained 14 genera
and some 40 species. eir 1995 revision added four new genera and 17 new species! is group shows a very special shape of

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
18
A B
C D
F
H
E
G

The MusorsToM-TDSB deep-sea crusTaceans 19
the 5th ambulatory legs, forming a forceps-like structure to carry other animals like sponge for camouflage. e morphological
diversity of this supplementary structure was commented in relation with the behavior of these animals (Guinot & Richer de
Forges 1995). e mysterious pigmented spot situated on the chelae of some species of
Lamoha
(formerly
Hypsophrys
, see
Ng 1998a) and interpreted as a luminescent organ were also documented. Since this key revision, many more species were
described from new cruises in the Philippines, Salomon and Vanuatu archipelago. e family now has 68 species (Ng 1998b; Ng
& Chen 1999; Ng & Wang 2002; Richer de Forges & Ng 2007a, 2008b; Naruse & Richer de Forges 2010; Ng & Eldredge 2012).
Associated with the homolids were several essentially deep-water families that many carcinologists regard as “primitive
crabs”. e Homolodromiidae was revised by Guinot (1993, 1995), and although no new genera were established, the
affinities of this enigmatic family were substantially clarified. Even so, eight new species were described. In subsequent years,
expeditions have obtained two more species from TDSB cruises (Ng & McLay 2005; Ng & Naruse 2007a). e small family
Dynomenidae was revised by McLay (1999), and in the process, he named 13 species in five genera, two of which were newly
described. Using new collections from the subsequent expeditions, seven more species have since been added by McLay &
Ng (2004, 2005) and Ng & McLay (2010). Recently, in a major reappraisal of the systematics of the family, Guinot (2008)
recognised three new subfamilies in the Dynomenidae. Perhaps one of the most exciting discoveries to French Polynesia was
a new family, Poupiniidae, described by Guinot (1991). A revision of Cyclodorippidae and Cymonomidae from the Indo-West
Pacific conducted by Tavares (1991b, 1992, 1993), which had previously only nine genera and 48 species, resulted in eight new
genera and 21 new species described! In a follow-up study (Tavares 1998), several genera were removed into their own family,
the Phyllotymolinidae Tavares, 1998. Since then, using these studies as the basis, seven other species have been described
from the Indo-West Pacific as a result of new MUSORSTOM-TDSB collections (Ahyong & Ng 2009, 2011; Naruse 2013)!
From the crabs collected in the Philippines, a new superfamily was described by de Saint Laurent (1989): Retroplumoidea.
Retroplumid crabs live in very muddy habitats of the deep-sea. ey have a very flat carapace and with the fifth ambulatory
legs greatly modified and sometimes feather-like. ere are only two genera,
Retropluma
and
Bathypluma,
in the Indo-West
Pacific. Description on the newly collected MUSORSTOM-TDSB material allowed detailed comparisons with the crab fossils
of this group. Based on morphological characters, de Saint Laurent (1989) proposed a phylogeny of this group of crabs as
separated from the Eubrachyura since the upper Cretaceous. Using a larger collection of Retroplumidae, McLay (2006) was
able to provide full descriptions on these animals and added more paleogeographical comments. ere are now seven species
of
Retropluma
and three species of
Bathypluma
. Another poorly known group are the palicids. Studying the large Indo-
Pacific collection, Castro (2000) completed a revision of the family Palicidae. ere are now two subfamilies, 10 genera and
43 species, with four genera and 17 species newly described. More species were treated by Castro (2010) from additional
collections. A strange crab, presumably associated with deep-water wood and debri, collected from recent expeditions to the
Philippines,
Vultocinus anfractus
, proved to belong to a new family, Vultocinidae (Ng & Manuel-Santo 2007). Surprisingly, it
was later also discovered in New Caledonia from among the older MUSORSTOM collections (Ng & Richer de Forges 2009).
SHRIMPS AND LOBSTERS
— e MUSORSTOM-TDSB collection has contributed significant knowledge that has often
led to a world-wide or Indo-West Pacific revision of many shrimp and lobster groups (Figures 1, 5E, 6). ese include
Solenoceridae (Crosnier 1978, 1985a, 1988a, 1989, 1994c; Crosnier & Dall 2004; Chan 2012), Sicyoniidae (Crosnier 2003),
Parapenaeus
(Crosnier 1985b, 2005),
Metapenaeopsis
(Crosnier 1987, 1991, 1994a, b),
Trachypenaeopsis
(Crosnier 2007),
Stenopodidae (de Saint Laurent & Cleva 1981; Saito
et al
. 2009), Spongicolidae (de Saint laurent & Cleva 1981; Komai &
Saito 2006; Saito & Komai 2008), Stylodactylidae (Cleva 1990, 1997, 2004), Processidae (Noël 1985), Pasiphaeidae (Crosnier
FIGURE 5
Remarkable crustaceans collected by the MUSORSTOM-TDSB programme.
A
, Lamoha personata, New Caledonia (photograph B. Richer de Forges, carapace
size about 40 x 50 mm).
B
, Lithodes
richeri, Vanuatu (photograph B. Richer de Forges, carapace size about 70 x 100 mm).
C
, Giant ostracod Azygocypridina
brynmawria, measuring 17 mm across, Coral Sea (photograph B. Richer de Forges).
D
, The living fossil barnacle Waikalasma
boucheti, New Caledonia
(photograph B. Richer de Forges, size about 20 mm).
E
, Glyphocrangon
panglao, Philippines (photograph T.-Y. Chan, cl 15.0 mm).
F
, Sympagurus
brevipes,
Philippines (photograph T.-Y. Chan, size about 10 mm).
G
, Babamunida
callista, New Caledonia (photograph B. Richer de Forges, cl 12.0 mm).
H
, Eurysquilloides
sibogae, Philippines (photograph T.-Y. Chan, size about 50 mm).

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
20
1988b; Hayashi 1999, 2004, 2006), Bathypalaemonellidae (Cleva 2001, 2004), deep-sea Pontoniinae (Bruce 1981, 1990, 1991,
1996; Li
et al
. 2008; Mitsuhashi & Chan 2006, 2009; Mitsuhashi
et al
.
2010, 2012),
Heterocarpus
(Crosnier 1988c, 1999b;
Cleva & Crosnier 2006; Li 2006; Yang
et al
. 2010; Li & Chan 2013),
Plesionika
(Crosnier 1986; Chan & Crosnier 1991, 1997;
Chan & Yu 1991c; Chan 2004; Komai & Chan 2010; Li & Chan 2013),
Eugonatonotus
(Chan & Yu 1991b), Nematocarcini-
dae (Burukovsky 2013),
Glyphocrangon
(Komai 2004, 2006b, 2007, 2011; Komai & Chan 2008, 2013), many Crangonidae
genera (
Aegaeon
,
Pontocaris
,
Parapontocaris
(Chan 1996),
Parapontophilus
(Komai 2008),
Prionocrangon
(Kim & Chan
2005),
Lissosabinea
(Komai 2006a),
Metacrangon
(Komai 1997, 2012)), Glypheidae (Forest & de Saint Laurent 1981, 1989;
Richer de Forges 2006; Forest 2006a, b; Boisselier-Dubayle
et al
. 2010),
Enoplometopus
(de Saint Laurent 1988; Chan &
Yu 1998; Chan & Ng 2008),
Nephropsi
s (Macpherson 1990b; Chan 1997),
Metanephr
o
ps
(Macpherson 1990b; Chan & Yu
1991a; Chan 1997), aumastochelidae (Chan & de Saint Laurent 1999; Ahyong
et al
. 2010),
Justitia
(Poupin 1994),
Pali-
nustus
(Chan & Yu 1995),
Puerulus
(Chan
et al.
2013), Scyllarinae (Holthuis 2002; Yang
et al
. 2008, 2011; Yang & Chan 2010,
2012) and Polychelidae (Galil 2000; Ahyong & Galil 2006; Ahyong & Chan 2008). Altogether,29 genera and263 species
were described from these works, and many of these reports have become the essential taxonomic references for deep-sea
shrimps and lobsters. e high diversity for several classical deep-sea caridean groups such as Pasiphaeidae,
Glyphocrangon
and
Plesionika
has certainly exceeded all expectations.
Of the numerous crustacean species collected during the MUSORSTOM-TDSB programme, the most famous are certainly
the glypheids. e discovery of extant species by Forest of this group, previously known only as fossils and supposedly
extinct at the end of the Cenozoic era, was a major event in the taxonomy of crustaceans. e glypheids, well known from
the Mezozoic, was considered to be extinct in the Eocene, 50 million years ago. ey were originally placed under the
infraorder Palinura and considered as probable ancestors of part of the remaining Decapoda Reptantia. However, detailed
studies on the first extant species
N. inopinata
showed that the glypheids are closer to the Astacidae (Forest 2006). In 2005,
the cruise EBISCO discovered a second species of glypheid (Figure 1B) living in a very different habitat in the Coral Sea
(Richer de Forges 2006). is new species was so different from
Neoglyphea inopinata
that Forest decided to establish a
new genus
Laurentaeglyphea
for it. is lobster-like animal,
Laurentaeglyphea neocaledonica
soon became famous under
the nickname “Jurassic shrimp” (the first species discovered was called the“Phoenix lobster”, Trew Crist
et al
. 2009). e
second species,
L. neocaledonica,
is even closer to those glypheids known from the Mezozoic and Eocene (Forest 2006).
FIGURE 6
Dinochelus ausubeli, male, cl 31.1 mm,
Philippines (photograph T.-Y. Chan). An icon
of the Census of Marine Life.

The MusorsToM-TDSB deep-sea crusTaceans 21
Another bizarre lobster
Dinochelus ausubeli
(Figure 6), recently collected from the Philippines AURORA expedition
(Ahyong
et al
. 2010) shows all the ‘monster-like’ features associated with deep-sea animals. is species was chosen as
an icon of CoML and has been used by popular media to promote deep-sea biodiversity and conservation (
e.g.
, Trew
Crist
et al
. 2009; Snelgrove 2010).
ANOMURANS
— Perhaps no other group of decapods has been as substantially impacted by the MUSORSTOM-TDSB
expeditions than the Paguroidea (sensu McLaughlin
et al
. 2007). Prior to 1976 when collecting efforts began on cruises
of the VAUBAN, the extent of deep-water paguroid specimens from the Indo-West and Central Pacific available in
museums, was minimal and insufficient for scientists to decipher the morphological boundaries and distributions of
many species. Taxonomic research was confined mostly to studying type material, with only scant additional preserved
or fresh specimens accessible. e first major work on paguroids that was produced with MUSORSTOM-TDSB material
was that by Forest (1987), who expanded exponentially our knowledge of the intriguing and mysterious “symmetrical”
hermit crabs of the family Pylochelidae, a presumably ancient group considered key to understanding hermit crab evolu-
tion (Tsang
et al
. 2011). Before Forest’s work, only 19 species were known, based on 60 specimens obtained at 30 stations
from around the world. He added 24 new species and subspecies using 400 specimens from about 200 sites, primarily
from MUSORSTOM-TDSB cruises, and complemented by specimens from earlier deep-sea expeditions on the ALBA-
TROSS. Subsequent MUSORSTOM-TDSB and other French cruises obtained considerably more material which was
used to further increase our knowledge of the diversity of pylochelids, resulting in revisions of their classification and
phylogeny (McLaughlin & Lemaitre 2008; Lemaitre
et al.
2009; McLaughlin
et al.
2009).
Among the hermit crabs of the family Diogenidae, the genus
Trizopagurus
benefitted greatly from a major revision by
Forest (1995) of their taxonomy and morphology, largely based on specimens obtained during MUSORSTOM-TDSB. is
genus contained only 10 species before the study by Forest, who expanded it to contain (in addition to
Trizopagurus
) two
new genera (
Ciliopagurus
Forest, 1995,
Strigopagurus
Forest, 1995) and 24 species, 14 of them new. Particularly fascinating
are the stridulatory structures in these three genera, the morphology of which Forest (1995) described for the first time in
great detail. He also discussed their anatomy in an evolutionary context. From the extraordinary collections in Polynesia,
another new
Ciliopagurus
was described by Poupin (2001), as well as new species and colour data of
Calcinus
species
(Poupin & McLaughlin 1998; Poupin & Lemaitre 2003). e taxonomy, diversity and morphology of what is probably
the most speciose and complex diogenid (indeed paguroid) genus,
Paguristes
, and other taxonomically difficult diogenid
genera, were significantly improved with a number of important taxonomic and morphological studies by Rahayu (2005,
2006, 2007, 2008, 2010) and Rahayu & Forest (2009). Even a new species of burrowing barnacle of the genus
Tomlinsonia
that is a parasite on three species of
Calcinus
was documented by Williams & Boyko (2006).
A seminal paper on the Paguridae from Indonesian waters was published by McLaughlin (1997). at paper was based
on specimens obtained during the KARUBAR expedition, and revealed an unexpected wealth of hermit crabs of this
family in the Indo-West Pacific. Seven new genera and 26 new species were discovered. Previous to McLaughlin’s paper,
most hermit crabs inhabiting the tropical environs of the Indo-West Pacific were thought to be species of Diogenidae. In
fact, McLaughlin’s paper showed that the Paguridae are as rich or even richer in this region of the world oceans than the
Diogenidae, and opened the door to a complete re-evaluation of the Paguroidea in general from the Indian and Pacific
Oceans or indeed the world over. Whereas brachyurans are certainly the most diverse in number of species among
the Decapoda, the anomurans, and in particular the Paguroidea, are the most diverse in body shape and morphology.
Additional discoveries based on MUSORSTOM-TDSB include: a new species of
Xylopagurus
A. Milne-Edwards, 1880,
a genus previously known only from the western Atlantic and eastern Pacific (Forest 1997); several more new genera
and new species (Lemaitre & McLaughlin 1995; McLaughlin & Forest 1997); new species of
Porcellanopagurus
and
Solitariopagurus
, two genera that include remarkably carcinized forms (McLaughlin 2000); a remarkable new genus and
species with reduced corneas (Lemaitre 2003); two astounding and minute new species of a new genus,
Pteropagurus
McLaughlin & Rahayu, 2006, found living in empty planktonic pteropod shells accumulated on the seafloor in a very
unusual hermit crab-mollusc association; new species or records, and taxonomic clarifications of species of
Cestopagurus
,

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
22
Pagurixus
,
Pseudopagurus
(Rahayu & Komai 2013) and
Pagurus
(Komai & Chan 2006; Komai 2010; Komai & Poupin
2012); and a new monotypic genus with a dwarf new species,
Pumilopagurus tuberculomanus
McLaughlin & Rahayu,
2008b. In addition, a major taxonomic and morphological review of
Nematopagurus
, a genus with 26 species distributed
primarily in the Indo-West Pacific, was published by McLaughlin (2004b).
Within the paguroids, no other family has been impacted as much by the rich collections from the MUSORSTOM-
TDSB projects, than the deep-water hermit crabs of the family Parapaguridae. In the last 40 years, primarily because of
the wealth of deep-water material collected by French expeditions, the number of species has increased from 31 to 86,
and the number of genera from 4 to 10, an increase of 277% and 250%, respectively (Figure 5F). Numerous parapagurid
specimens were collected from many deep-water habitats and regions that had never been sampled before. Before 1986,
when French expeditions using the S.M.C.B. “Marara” began to collect in French Polynesia including the Marquesas,
there were no specimens or records of parapagurids from that immense region of the central South Pacific. Works on
those collections resulted in the description of several remarkable new species, and more importantly a better picture of
the anomurans from that region emerged. e Polynesian parapagurids as well as other paguroids and decapods, were
summarized in a number of catalogues or atlases, including fisheries data and many color photographs (Poupin
et al
.
1990; Poupin, 1996a, b). Furthermore, a database of the species of decapods (shrimps, lobsters, crabs, hermit crabs) and
stomatopods (mantis shrimps), recorded from the French Polynesian Islands (Marquesas, Society, Tuamotu, Austral, and
Gambier), Pitcairn, Easter Island and Clipperton, is available through a free database in the internet (http://decapoda.free.
fr/). e numerous studies on the taxonomy and systematics of parapagurids based on specimens from all these French-
sponsored faunistic studies, greatly expanded and clarified the world-wide picture of diversity and biogeography of the
parapagurids, an evolutionary key group of hermit crabs. Taxonomic and systematic information of this family can be
found in a number of convenient, broad generic reviews or revisions (Lemaitre 1999, 2004a, b, 2013), as well is in a number
papers reporting on parapagurids from particular regions and describing new species, or discussing morphological and
biological aspects (Lemaitre 1993, 1994, 1997, 1998, 2004a, b, 2006; Lemaitre & Poupin 2003).
e specialized burrowing crabs of the superfamily Hippoidea from the tropical Indo-West Pacific region had been
the subject of few systematic studies until many specimens of this poorly known group of crabs became available from
collections obtained by MUSORSTOM-TDSB. As result, three landmark and detailed studies on the systematics of the
families Albuneidae and Hippidae, including descriptions of several new species, have been published (Boyko & Harvey
1999; Boyko 2000, 2002). In those papers, the diagnostic morphology of the group was radically redefined and deciphered
to the benefit of future studies on their biology and evolution.
Squat lobsters are a major component of the deep-sea crustacean fauna (Baba
et al.
2008). ey are divided into
two main groups. e Galatheoidea (with four families Galatheidae, Munididae, Munidopsidae and Porcellanidae) has
a large number of free-living species living on the ocean bottom, and generally have numerous eggs with planktonic
larvae. e Chirostyloidea (with three families Chirostylidae, Eumunididae and Kiwaidae) are often associated with other
benthic invertebrates like sponges, antipatharians or gorgonians, and produce few eggs with advanced larvae and limited
dispersion. e abundant galatheoid material obtained in the MUSORSTOM-TDSB collection has resulted in intensive
taxonomic studies on these animals that have revolutionised our understanding of their diversity. After the TDSB studies
initiated in 1976, these families have been enriched by 13 new genera (Baba 1991, 2004; Baba & de Saint Laurent 1996;
Macpherson 1998, 2006; Macpherson & Machordom 2000; Cabezas
et al
. 2008). e total number of known species
has increasednearly 2.5 fold since 1976, with 937 taxa today (Baba
et al
. 2008; Appeltans
et al
.
2012). Most of the newly
described species have been based on the MUSORSTOM-TDSB material. Some of them living in the upper part of the
bathyal zone (200-600 m) can be surprisingly colourful, although some live colours and patterns have to be seen to be
believed (Figure 5G).
e first contribution towards galatheoid systematics using these collections was published by Baba (1990) on
specimens collected by Alain Crosnier on board of the R.V. “Vauban”, in Madagascar. Subsequently, Baba (1991a, b),
using material from some expeditions carried out around New Caledonia, described new genera and species of chirostylids
and galatheids of the genera
Alainius, Leiogalathea, Phylladiorhynchus, Gastroptychus
and
Chirostylus
. ese papers,

The MusorsToM-TDSB deep-sea crusTaceans 23
together with the review of the genus
Eumunida
(de Saint Laurent & Macpherson 1990a) along the Indian and Pacific
Oceans, were the starting point of the many subsequent studies of the group. e genus
Eumunida
was revised again by
de Saint Laurent & Poupin (1996), adding numerous specimens from the MUSORSTOM expeditions, and describing
six new species from the Indian and Pacific Oceans. Baba & de Saint Laurent (1996) described some new genera of small
galatheids (
Anoplonida, Neonida, Plesionida
, among others), closely related to the genus
Bathymunida
. Furthermore,
they divide the galatheids in two large groups (
i.e.
having one or two pairs of male gonopods) and reorganizing the
taxonomy of the family. e phylogenetic value of this division was later validated by molecular tools (Machordom &
Macpherson 2004).
Baba (2004) erected the genus
Uroptychodes
from
Uroptychus
, using material collected in New Caledonia, Indonesia
and adjacent waters. An additional study on the genus
Uroptychus
from Taiwan was published by Baba & Lin (2008),
Cabezas
et al
. (2012), and a large and complete revision of the genus including more than 100 species will be published
soon (Baba in preparation). is major taxonomic revision of the Chirostylidae (Baba in preparation) will also form the
basis for new hypotheses on seamount endemism.
Macpherson (1993a, b) and Macpherson & Baba (1993) studied the material collected in the early MUSORSTOM-TDSB
cruises carried out in the Philippines and Indonesia, adding numerous new species belonging to the genera
Agononida,
Munida
and
Paramunida
. In addition, they solved the confusing status of
Munida japonica
and their relatives. In 1994,
a study on the collections obtained mostly in New Caledonia, Chesterfield, Matthew and Hunter Islands, demonstrated
the existence of 56 new species of
Agononida
and
Munida
(Macpherson 1994). e existence of a very rich fauna in the
western Pacific was also supported by the results obtained in MUSORSTOM-TDSB expeditions from Wallis and Futuna
(Macpherson 1996), Kei Islands (Indonesia) (Macpherson 1997), Vanuatu (Macpherson 1999a), Marquesas Islands
(Macpherson 2000), Fiji and Tonga (Macpherson 2004), Austral Islands (Macpherson 2006a), Solomon Islands (Cabezas
et
al.
2009) and Taiwan (Lin
et al.
2004, 2005; Cabezas
et al.
2011b). Other studies using specimens from recent expeditions
are completing our knowledge of the group along the continental shelf and slope of the western and central Pacific, and
eastern Indian Ocean (Macpherson 1999b, 2006b, 2009; Macpherson & Baba 2006).
e deep-sea fauna, mostly belonging to the genera
Munidopsis
and
Galacantha
, was also studied using numerous
specimens collected in all MUSORSTOM-TDSB cruises, from Madagascar to French Polynesia (Baba & de Saint
Laurent 1992, Macpherson 2007, 2012), adding more than 25 new species. Furthermore, numerous new species have
been collected in recent expeditions along the coasts of Taiwan and the Philippines (Lin
et al.
2007, 2011; Osawa
et al.
2006a, b, 2007, 2008, 2013).
e shallow-water genera (
e.g.
,
Allogalathea
,
Galathea
,
Lauriea
,
Sadayoshia
) have also been enriched by the
MUSORSTOM-TDSB expeditions confirming the existence of a highly diverse fauna with similar patterns than in the
continental shelf, slope and abyssal basins (Macpherson & Baba 2010; Macpherson & Cleva 2010; Cabezas
et al
. 2011),
and a major revision of the diverse genus
Galathea
is in preparation (Macpherson in preparation).
e results obtained from MUSORSTOM-TDSB programme have demonstrated that squat lobsters are abundant,
speciose and widely distributed. ey occur in all marine habitats at depths ranging from the surface of the sea to more
than 5000 m (Baba
et al.
2008). Squat lobsters are most diverse in the western Pacific area and, at present, the most speciose
genera are
Munida
and
Munidopsis
. e highest numbers of squat lobsters are found in the Coral Sea (Solomon-Vanuatu-
New Caledonia), and Indo-Malay-Philippines Archipelago (Macpherson
et al.
2010; Osawa
et al.
2013). Most species of
Munida
are from shelf and slope depths, whereas those of
Munidopsis
and
Galacantha
are found mainly on continental
slopes and the continental rise zones. e distribution of endemism along the Pacific Ocean indicates that there are
several major centers of diversity, for example in the Coral Sea, Indo-Malay-Philippines Archipelago and French Polynesia.
e squat lobsters are becoming one of the best-known groups of deep-sea crustaceans. As a summary of the work that
started in 1976, the different authors have produced many important papers that have formed the basis for other major,
synthetic works. For instance, studies on the fauna of some basins,
e.g.
, Taiwan (Baba
et al.
2009), the checklist of squat
lobsters of the world (Baba
et al.
2008), and a book on the biology of squat lobsters (Poore
et al.
2011).

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
24
e king crabs (family Lithodidae) are also well represented in MUSORSTOM-TDSB collections (Figure 5B). e
first contribution using specimens collected by Alain Crosnier in Madagascar was published by Macpherson (1988).
Other studies covering the area of New Caledonia and French Polynesia were published by Macpherson (1990) and
(1991), respectively. e hydrothermal vents also provide some interesting new species (de Saint Laurent & Macpherson
1997), and the most recent MUSORSTOM-TDSB expeditions collected additional material, including interesting new
occurrences and species from New Caledonia, Vanuatu, Solomon, Fiji and Tonga areas (Macpherson 2001, 2003). More
recently, some interesting species have been described in expeditions carried out in Taiwan (Macpherson & Chan 2008;
Ahyong & Chan 2010).
OTHER REMARKABLE CRUSTACEANS
— Not all the work done on crustaceans during the MUSORSTOM-TDSB exercise
was on decapods. From the crustaceans collected in the deep-sea some were remarkable by their giant size such as the
isopods
Bathynomus
and ostracod
Azygocypridina
. Other than “living fossils” like the glypheids
Neoglyphea inopinata
and
Laurentaeglyphea neocaledonica
, the ancient cirriped
Waikalasma boucheti
also merits mention. ese have since
been used as “icons” for deep-sea biodiversity research.
Bathynomus
is an isopod genus containing the largest known isopod species, its members living between 100 and
1000 m deep. Based mainly on the MUSORSTOM-TDSB collection, Lowry & Dempsey (2006) published a revision of
this group of giant scavengers from the Indo-West Pacific. Six species are redescribed and six new species were added.
e authors separated the genus into two groups, the giants and the super-giants having an adult size between 150 and
500 mm long (Figure 7).
FIGURE 7
A supergiant of the isopod genus Bathynomus — B. kensleyi, Philippines (photograph P. K. L. Ng). A saga of the MUSORSTOM-TDSB deep-sea expeditions.

The MusorsToM-TDSB deep-sea crusTaceans 25
Another giant described from the MUSORSTOM-TDSB material was the ostracod Myodocopa in the family Cyprinidae,
Azygocypridina brynmawria
. It was collected in the Coral Sea at about 400 m deep (Diamond, Richer de Forges & Kornicker
2008). e majority of ostracods have adult sizes of only a few millimetres, but this species has a diameter of 17 mm
(Figure 5C).
Azygocypridina
ostracods possess iridescence on the first antennae. e description of this phenomenon by
Andrew Parker on a species of the same genus was the starting point of the fascinating speculations on the color vision
in fossil crustaceans and its role in the so called “Cambrian Explosion”of life (Parker 1998, 2003; Nielsen & Parker 2010).
Cirripeds of the genus
Waikalasma
was known only from fossils in the Miocene of New Zealand. However, during
the cruise MUSORSTOM 8 in Vanuatu a living species was collected:
Waikalasma boucheti
. In 2011, the cruise EXBODI
discovered another specimen of this species from New Caledonia. is genus is characterised by an extra layer of plates
around the body (Figure 5D).
Although a much smaller group than the decapods, the mantis shrimps (Order Stomatopoda) are generally best
known from shallow waters. Knowledge of their deep-water diversity, however, has been significantly increased through
MUSORSTOM-TDSB sampling programmes, with discovery of many new species, genera and even a new family,
Alainosquillidae, from New Caledonia (Moosa 1991). Further discoveries are ongoing with many more new species under
study. Many uncommon species were often collected by the MUSORSTOM-TDSB cruises, such as
Bathysquilla microps
,
B. crassispinosa
and
Gonodactyloideus cracens
, and others that were until that time thought particularly rare, such as the
enigmatic,
Eurysquilloides sibogae
(Figure 5H), which is now known to be common in the Philippines. Studies dealing
with Stomatopoda include major studies from the Philippines (Moosa 1986; Ahyong 2010, 2013), New Caledonia (Moosa
1991; Ahyong 2007), Indonesia and the Marquesas (Ahyong 2002a, b).
OTHER CONTRIBUTIONS OF THE MUSORSTOM-TDSB PROGRAMME
Biogeography
e science of biogeography synthesises knowledge from taxonomy, phylogeny and genetics, and helps us understand
how the marine environment has evolved, the impacts of sea level variations and plate tectonics on biodiversity. All these
different aspects of science contribute to reconstructing the spatio-temporal scenario explaning the origin and current
distribution of species (Heads 2005, 2008; Hoeksema 2007). e sparseness of deep-sea samplings often impairs a meaningful
interpretation of the deep-sea biogeography. Hypotheses of deep-sea biogeography, inferred primarily from shallow-water
patterns are far from accurate. One exception is among the squat lobsters (Galatheoidea and Chirostyloidea) because
they have been extensively sampled by the MUSORSTOM-TDSB programme from a wide range of depths (littoral to
1500 m deep), having a large number of species supported by solid taxonomy.
Using the taxonomic results of squat lobsters coming largely from the MUSORSTOM-TDSB collections, two papers
were published in 2010 marking the end of the first phase of the Census of Marine Life operations (Macpherson
et al.
2010; Rowden
et al.
2010). e first paper was trying to define through statistical analysis the characteristics of each
biogeographical province in the Pacific Ocean. e data set composed of 54 deep-sea cruises and 3200 stations with
402 species. e multivariate analyses clearly showed a gradient of decreasing specific richness from Western to Central
Pacific. is fundamental concept, well known for the littoral marine fauna, was demonstrated for the first time in the
bathyal zone. e richest areas are the Coral triangle (Indo-Malayan-Philippines) and the Coral Sea.
e second work focused on the seamounts of the southwest Pacific and was trying to determine if the faunistic
composition is different between seamounts and the slopes at comparable depths. ey used a data set from a total of
502 species (Rowden
et al.
2010) and concluded that the different assemblages of squat lobsters between seamounts and
non-seamounts are mainly a consequence of hydrological conditions and especially the availability of food.
Both studies have clearly demonstrated that good taxonomic work at species level is a prerequisite for any meaningful
analysis on ecology or biogeography.

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
26
SpeCiaL eCoSySTeMS
During the MUSORSTOM-TDSB explorations two unusual habitats were discovered, representing rich oasis in the deep-
sea desert: the summit of seamounts and sunken woods. Both have a special assemblage of life with remarkable crustaceans.
SUNKEN WOOD FAUNA
— e presence of animals on plant remains at the deep-sea floor was first documented by the
CHALLENGER expedition (1872-1876). e GALATHEA expedition (1950-1952) also recorded the abundant presence
of plant remains on the deep-sea floor (Bruun 1959). e importance of plant material in the deep sea was first empha-
sised by Wolf (1979). Associated with these remains is a unique fauna, which has mainly been examined by the taxono-
mists. e ecological aspects are less well studied until recently. Apart from sporadic taxonomic considerations, plant-
associated organisms were seen as zoological and/or ecological curiosities and thus only anecdotally studied. Sunken
wood and plant material has long been considered a mere curiosity, but this habitat was recently emphasized as deep-sea
environment of major evolutionary importance. Many of the species living on sunken wood are phylogenetically related
to those living on hydrothermal vents, cold seeps and other organic substrates (
e.g.
, whale falls).
Distel
et al.
(2000) underlined the evolutionary affinities of the giant hydrothermal vent mussels with those found on
sunken wood or whale bones (so-called whale falls). is result was then confirmed by several studies (Samadi
et al.
2007;
Lorion
et al.
2009; Fujita
et al
. 2009; Kyuno
et al.
2009). e zoological affinities of organisms associated with sunken woods
and whale falls with those from hydrothermal vent or cold seep ecosystems suggests that these ecosystems, developing
on organic substrata in the deep-sea, could be an evolutionary step toward the colonization of hydrothermal vents (the
so-called “wooden steps” hypothesis of Distel
et al.
(2000). However, few data are available for sunken wood organisms.
Recent results suggest that the evolutionary history of deep-sea mussels associated with organic falls is completely mixed up
with that of vent and seep mussels, with evolutionary scenarios probably more complex than the “wooden steps” hypothesis
(Lorion
et al.
2010; ubaut 2012). Moreover, geography, rather than habitat specificity or chemosymbiotic requirements,
seems to significantly structure diversification patterns in sunken wood-associated mytilids (Lorion
et al.
2009, 2010).
In 2004, the TDSB programme started specific series of cruises particularly devoted to the exploration of sunken-
wood environments (Biodiversité des Organismes Associés aux bois coulés — BOA). Since the first MUSORSTOM
cruises in the Philippines (see Forest 1981) and Indonesia (see Moosa 1984), it became clear that the slopes of oceanic
islands within these regions accumulate large amounts of decomposing vegetation. Subsequent cruises in the Solomon
Islands, Vanuatu Archipelago, and Papua New Guinea specifically targeted sunken vegetation to further characterize its
fauna. Several cruises provided specimens associated with sunken wood in the depth range between 100-1500 m from
different locations corresponding to the most important bathyal area where sunken woods accumulate in the Indo-Pacific
region: (i) Philippines (PANGLAO, 2005; AURORA, 2007; LUMIWAN, 2008), (ii) Solomon Islands (SALOMON 2, 2004;
SALOMONBOA 3, 2007), (iii) Vanuatu Archipelago (BOA 0, 2004; BOA 1, 2005 and SANTOBOA, 2006) and Papua
New Guinea (BIOPAPUA, 2010). Additional samples from other cruises, far from these basins, also provided specimens
associated with organic remains. Indeed, for example, on seamounts, small pieces of wood and seeds (
e.g.
, coconuts) are
variously pulled up by the trawls or the dredges and these erratic substrates, are usually also colonized by the organisms
typically found on pieces of wood in accumulation areas.
is sampling is, however, focussed on substrates that are easy to locate on the deep-sea floor (such as large pieces
of wood off the mouth of the rivers). erefore, to more accurately catch the diversity of the organisms associated with
a larger diversity of organic substrates that can be found on the deep-sea floor, pots covered by netting with mesh of
3 mm were deployed (Samadi
et al.
2010). e “bait” placed in the pots were seeds and pieces of wood of various tree
species, sugar cane stems, whale bones, green turtle shell, stag horn, feathers, cephalopod beaks and cuttlefish-bones.
ese lines of pots were deployed in 2003 off the barrier of New Caledonia and off Santo island in Vanuatu and left for
periods ranging from 10 to 30 months.
ese studies demonstrated that such organic substrates were associated with a rich and unique fauna (Samadi
et
al
. 2007) and its confirmation catalyzed new research interests for the TDSB team (reviewed in Samadi
et al
. 2010).
With the large scale sampling conducted by TDSB cruises, new recent taxonomic and phylogenetic studies have been

The MusorsToM-TDSB deep-sea crusTaceans 27
launched, mainly focused on different key-groups like molluscs such as Mytilidae (Lorion
et al
. 2010, 2011; ubaut
2012) and polyplacophoran chitons (Duperron
et al.
2012). Several studies have also been done on the feeding behaviour
of echinoderms (Becker
et al
. 2009) and molluscs (Zbinden
et al
. 2010), botanical identification of the plant-remains
(Pailleret
et al
. 2007), prokaryotic diversity (Palacios
et al
. 2009), fungal diversity (Dupont
et al
. 2009) and the association
of mytilids with chemosynthetic bacteria (reviewed in Duperron
et al
. 2009).
For crustaceans, isopods, amphipods and decapods as well as barnacles have been found among plants debri in
the trawls or dredges. It is, however, difficult to know to what extent they depend on the decaying plant substrates, in
particular for the vagrant fauna. Among the decapods, the main groups repeatedly caught with such substrata include
squat lobsters (
e.g.
,
Munidopsis andamanica
,
M. nitida
,
M. bispinosculata
,
M. similior
), two different families of
symmetrical hermit crabs (all the species of the genus
Xylopagurus
and several species of
Pylocheles
inhabit centimetre-
sized pieces of wood), ghost shrimp (
e.g.
,
Rayllianassa amboinensis
), some Axiidae and a number of snapping shrimps
(Alpheidae) (Figure 8A-E).
FIGURE 8
A
-
D
, Wood-associated crustaceans.
A
, Galacantha sp. (photograph B. Richer de Forges).
B
, Parapylocheles
scorpio (photograph T.-Y. Chan).
C
, Bathyceradocus sp.
D
, Onesimoides sp.
E
-
H
, Bathyceradocus sp.
E
, General view, total length 18 mm.
F
, SEM picture (magnication x 40) of the mouthparts illustrating the dense
bacterial colonisation.
G
,
H
, SEM pictures (respectively x800 and x 6000) of the diversity of bacterial morphotypes (rod-shaped and lamentous bacteria)
xed on the mouthpart setae (photographs
C
-
H
, L. Corbari).
A B
CD E
F G H

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
28
With regard to peracarid crustaceans, the most recurrent specimens collected in association with wood were: Isopoda
(Sphaeromatidae,
Cassidias
; Limnoriidae,
Limnoria
sp.; Gnathiidae,
Elaphognathia
sp.), Amphipoda (Lysianassidae,
Onesimoides
sp.; Maeridae,
Bathyceradocus
sp.). e same organisms were also collected in the pots that were submerged
off New Caledonia or Vanuatu.
e main challenge is now to understand the interactions between crustaceans and organic substrates (
e.g.
, habitat,
feeding relationship or even dependence in relation to reduced compounds produced by the wood-degradation via
microorganisms such as fungi). Preliminary studies on the diet of wood-associated decapods have been conducted on squat
lobsters, ghost shrimp and pylochelid hermit crabs (TDSB samples from Vanuatu, New Caledonia, Solomon Archipelago,
Philippines). A multidisciplinary approach based on electron microscopy (SEM, TEM); fluorescence in situ hybridization
(FISH) and stable isotopic analyses revealed their affiliation to different diets and emphasizes their trophic relationships
by the presence (or absence) of bacterial symbioses (Hoyoux 2010). e results revealed that two main diet-groups can
be identified for the studied wood-associated decapods: (i) predators such as
Xylocheles macrops
,
Bathycheles incisus
,
B. integer
and
Xylopagurus caledonicus
; and (ii) detritivorous species such as
Munidopsis bispinoculata
,
M. pilosa
,
M.
tida
,
M. andamanica
and the thalassinid
Rayllianassa amboinensis
. e most striking results deal with the detrivorous
groups, especially
M. andamanica
and
M. nitida
. Both species have detritivorous and xylophagous diets and harbour
symbiotic digestive microflora, indicating their close trophic interactions with wood and organic substrates. Indeed, a
careful examination of the feeding appendages, gut contents and gut lining of
M. andamanica
caught with wood falls
revealed this species to be a truly detritivorous species which uses wood and the biofilm-covering as its two main food
sources (Hoyoux
et al
. 2009).
Munidopsis andamanica
also possesses a resident gut microflora consisting of bacteria and
fungi possibly involved in the digestion of the wood fragments. is species can therefore be regarded as a xylophagous
specialist. In this work (Hoyoux
et al
. 2012), authors also examined the feeding appendages, gut contents and gut lining
of
M. nitida
,
M. bispinoculata
and
M. pilosa
from natural sunken woods and compared them with specimens of the
same species having survived and grown on different hard to-digest substrates (
i.e.
, woods, turtle shells and whale bones)
experimentally submerged in the deep South Pacific. In both cases, all three species directly ingest large wood fragments
that have been substantially degraded by micro-organisms. However,
M. nitida
also feeds on experimentally submerged
whale bone and turtle shell fragments.
Munidopsis nitida
is also the only species known to host a resident gut microflora,
although the bacterial morphotypes vary according to the ingested substrate. e results suggest that the three species
are most probably opportunistic, bacterivorous detritivores and
M. nitida
could be at the beginning of an evolutionary
process leading towards xylophagy within the genus
Munidopsis
.
New investigations are in progress on wood-associated amphipods. Taxonomic and phylogenetic studies have been
conducted on several groups. Identified as a key-group is the genus
Bathyceradocus
(family Maeridae; Figure 8E-H) that
revealed an unsuspected diversity.
Bathyceradocus
has only three species:
B. iberiensis
, described from Mediterranean
Sea;
B. stephenseni
described from three specimens collected during cruises of the SIBOGA in the Celebes sea (Pirlot
1934) and
B. wuzzae
recently described by Larsen & Krapp-Schickel (2007) from the vicinity of hydrothermal vents (Juan
de Fuca). In all these descriptions, the specimens were collected from dropped wood or plant remains but without more
ecological information. About 80 specimens of
Bathyceradocus
sp. have been collected during MUSORSTOM-TDSB
cruises from Madagascar to Fiji islands through Philippines and Papua New Guinea. e first morphological hypothesis
suggests that these specimens are representative of
B. stephenseni
. Phylogenetic analyses (COI and 18S) have revealed
that this group is much more diversified than expected: 12 species (based on morphological and molecular data) have
been differentiated, revealing the cryptic diversity in this genus (L. Corbari unpublished data). Moreover, microscopic
and ultra-structural investigations on the digestive tract have revealed the presence of wood-fragments in the digestive
content and the occurrence of micro-organisms (bacteria and fungi) in the posterior part of the intestine. ese results
confirm that the diet of the
Bathyceradocus
group as strictly wood-feeders (L. Corbari unpublished data). More interesting
is the recurrent presence of bacterial ectosymbiosis on the mouthparts of these specimens that suggests an original
pathway of nutrition that could involve bacteria in wood degradation (L. Corbari unpublished data). e same approach

The MusorsToM-TDSB deep-sea crusTaceans 29
of integrative taxonomy (combining morphological and molecular characters) is also being conducted for amphipods in
the
Onesimoides
group.
SEAMOUNTS
— Seamounts are mountains under the sea level and elevated above the sea bottoms more than 1000 m.
Seamounts are major features of the oceans. Recent mapping estimated that their number is at least to 14,000 but less
than 400 of them had been biologically sampled (Richer de Forges
et al
. 2000; Samadi
et al
. 2007; McClain 2007; Koslow
2007; Richer de Forges & Tessier 2012). ey form a benthic habitat of exceptional richness, both in diversity and abun-
dance, and they became a theme of a special section in the CoML called CenSeam, Census of Seamounts (2005-2010).
Several MUSORSTOM-TDSB cruises were devoted to this rich habitat in the Pacific (CHALCAL 2, AZTEQUE, NOR-
FOLK 1 & 2, EBISCO, TERRASSE, TARASOC). A preliminary hypothesis on the endemism of the seamount fauna
was proposed after the first Norfolk cruises (Richer de Forges
et al
. 2000). More recent studies of population genetics
on several animal groups such as molluscs and crustaceans have shown that seamounts are functioning as oases in the
deep-sea (Samadi
et al
. 2006; Castelin
et al
. 2012). On the other hand, some species generally considered to be very “rare”
were also caught on the Norfolk Ridge seamounts like
Palinustus unicornutus
;
Homola ranunculus
;
Sphaerodromia
lamellata
, and
Takedromia longispina
. But the notion of rarity in ecology is a complex issue because sampling in the
sea is generally not representative and with some habitats largely neglected, particularly at steep slopes of islands and
seamounts (see Mendoza
et al
. 2009).
EVOLUTION
— Traditional phylogenetic studies based on morphology have been complicated by the identification of
homologies, which is often difficult. Many modern techniques have been recently developed to overcome these difficul-
ties. However, all these modern techniques require fresh specimens or specimens preserved in a particularly way when
fresh. Most museum specimens are or have been preserved in formalin and unsuitable for these modern techniques.
Taxa from the deep-sea are generally not easy to collect because of the access of ship-time and very high running cost of
research vessels. Unfortunately, many species and even the whole genus or family in various animal groups are entirely
from the deep-sea. us, phylogenetic studies using modern techniques often encounter difficulties in sourcing enough
taxa to be included in the analysis.
e abundant and diverse specimens collected during the MUSORSTOM-TDSB programme from a very broad
geographical area are almost all preserved in ethanol or with the specimens specially treated immediately after collection
for later molecular or histological work. For example, a fruitful collaboration was established amongst Bertrand Richer
de Forges of the IRD, Danièle Guinot of the Muséum national d’Histoire naturelle and Barrie Jamieson of the University
of Queensland on the ultrastructure of spermatozoa in decapod crustaceans. Morphology of reproductive cells is very
conservative and can be used to reconstruct phylogeny. Before the MUSORSTOM programme very little was known
on the morphology of sperm in decapod crustaceans. Having large access to fresh material from the deep-sea new data
were rapidly obtained for the following families: Brachyura (Homolidae, Homolodromiidae, Dromiidae, Dynomeniidae,
Latreilliidae, Cyclodorippidae, Dorippidae, Calappidae, Atelecyclidae, Raninidae, Palicidae, Retroplumidae, Leucosiidae,
Geryonidae, Goneplacidae, Cancridae, Grapsidae, Xanthidae, Portunidae, Majidae, Hymenosomatidae),Anomura
(Paguridae, Pylochelidae, Parapaguridae, Chirostylidae, Galatheidae, Lithodidae, Hippidae, Porcellanidae),and Macrura
(Palinuridae, Polychelidae, Nephropidae). A large part of these results has been published (Guinot
et al
. 1994; Richer de
Forges
et al
.
1997; Tudge
et al.
2012). Using the sperm characters a new phylogeny of Brachyura was proposed (Jamieson
et al
.
1995). e phylogeny trees obtained are coherent with the classification of Guinot (1977) and distinguishing the
three groups Podotremata, Heterotremata and oracotremata.
During the last decade, the fast development of molecular biology has provided supplementary information that
can strengthen the outcomes of traditional morphology-based studies. e vast and diverse samples collected by
the MUSORSTOM-TDSB programme are now a major focus of the international BarCode of Life programme on
crustaceans. On the other hand, the availability of many deep-sea taxa that can be used for molecular sequencing has
allowed comprehensive studies on the higher classification and evolution of decapod crustaceans (
e.g.
, Tong
et al
. 2000;

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
30
Ahyong
et al
. 2007; Chan
et al
. 2008, 2009, 2010; Tsang, Lin
et al
.
2008; Tsang, Ma
et al
. 2008; Ahyong
et al
. 2009; ; Chu
et al
. 2009; Lai
et al
. 2009; Ma
et al
. 2009; Sin
et al.
2009; Tshudy
et al
. 2009; Tsang
et al
. 2009, 2011; De Grave
et al
. 2010;
Boisselier-Dubayle
et al
. 2010; Lai
et al
. 2011; Li
et a
l. 2011; Schnabel
et al
. 2011; Tsoi
et al
. 2011; Yang
et al
. 2012). Some
studies have shown the importance of the southwest Pacific area, specifically the island arc of Fiji, Tonga, Vanuatu, Wallis
and Futuna, for diversification of squat lobsters, probably associated with a global warming, high tectonic activity, and
changes in oceanic currents that took place in this region during the Oligocene-Miocene period (Cabezas
et al
. 2012).
e results of these molecular studies have revolutionized the classification scheme of decapod crustaceans; such
as the hermit crabs, squat lobsters, the mud shrimps alassinidea, the subfamilies in Nephropidae and many caridean
superfamilies are not likely to be monophyletic groups, while the family or generic status of Sicyoniidae, aumastochelidae,
and Synaxidae may not be valid. On the other hand, the status of Enoplometopidae is settled and considered as a superfamily
in Astacidea. Moreover, the molecular data provide insights into the evolution within some of the infraorders, such as
parallel carcinization within the Anomura (Tsang
et al
. 2011).
Phylogenetics and molecular studies of paguroids in particular, and anomurans or decapods in general, have taken
advantage of the new availability of specimens as well as the greatly expanded morphological and distributional knowledge
generated during nearly 40 years of publications based on sampling programs in the deep-seas of the Indo-West Pacific.
Discoveries of remarkable new forms of what may be considered primitive or advanced taxa, have provided improved insight
into the evolution of paguroids. Cladistic and genetic studies have incorporated the data or molecular information extracted
from the vast MUSORSTOM-TDSB collections. Significantly improved evolutionary scenarios have been produced, and
although theories are still controversial and hotly debated, we are much closer than before to the understanding of the
true relationships and origins of paguroids in particular, and anomurans in general (
e.g.
, McLaughlin
et al
. 2007; Tsang
et al
. 2008, 2011; Ahyong
et al
. 2009; Lemaitre & McLaughlin 2009; Lemaitre
et al
. 2009).
e molecular techniques have been providing new findings on the taxonomy, phylogeny and classification of squat
lobsters,
e.g.
, Machordom & Macpherson (2001, 2004), Chu
et al.
(2009), Ahyong
et al.
(2009), Cabezas
at al.
(2008, 2010),
Puillandre
et al.
(2011). e most recent and complete studies have proposed a new classification of the squat lobsters
(Ahyong
et al
. 2010; Schnabel & Ahyong 2010), with two superfamilies (Galatheoidea and Chirostyloidea). e material
obtained along the Indian and Pacific Oceans has allowed some studies on the phylogeny and origin of this rich fauna.
ese studies suggest that the galatheid diversification in the southwestern Pacific may have begun some 7-14 million
years ago, during the Middle or Late Miocene (Machordom & Macpherson 2004), and the Coral Sea area seems to be the
origin of several genera of squat lobsters like
Paramunida
and
Plesionida
(Cabezas
et al
. 2009, 2012).
All in all, the MUSORSTOM-TDSB collection now forms an important part in specimen sourcing for the molecular
component in the international Assembling the Tree of Life project on decapod crustaceans.
THE ALAIN CROSNIER EFFECT
All crustacean researchers who have worked on the MUSORSTOM-TDSB collections were invariably influenced by
the “Alain Crosnier effect”. His energy, passion and dedication were legendary. In the years between the first cruise (1976)
to 2008 (when he decided to fully retire), he was instrumental in arranging expeditions, having the material centralized in
the Muséum national d’Histoire naturelle where he sorted it out, then invited specialists from around the world to work
on them (or occasionally sending it to them for study) and then assiduously arranging to have them published. e series
of MUSORSTOM volumes for which he personally edited and readied for press exceeds 10,000 printed pages, and with
hundreds of new genera and species described within.
More significantly, many of the papers published as a result of Crosnier’s efforts were not just straightforward new
genera and new species descriptions. A good number were major revisions or reviews of genera, subfamilies and families
that are extremely important and useful for biologists. Crosnier used the extensive MUSORSTOM-TDSB collections as
a nexus and a catalyst for such revisions that brought together types and material from all over the world. Crosnier was
directly or indirectly responsible for more such major revisions than any other period in carcinological science.

The MusorsToM-TDSB deep-sea crusTaceans 31
Overall, the efforts he put in to secure the collections and the funds to have the experts work in Paris, and then to have
them published were nothing short of phenomenal. And he was a tough taskmaster, driving the researchers hard to have the
job done and done well. He was a “one-man army”, almost single-handedly pushing carcinological and deep-sea research
to heights it has never seen before. It was a Herculean effort that has never been exceeded and may never be equaled.
rough all these years, Alain also created a special community of carcinologists devoted to the MUSORSTOM
enterprise and the advancement of crustacean systematic knowledge (see Macpherson
et al
. 1998). is included the
many researchers he invited to the museum to work on his material (the first was the late Chen Hui-Lian from Qingdao,
China, who worked on Leucosiidae and Dorippidae; the last being Tomoyuki Komai from Chiba, Japan, a specialist of
crangonids and glyphocrangonids) as well as those he generously sent material to. Affectionately known as “Crosnier’s
cronies”, this community continues their works to this day across almost all continents and over a dozen countries. He was
even honoured by his peers and “cronies” in a special volume of
Zoosystema
in 1998 (Volume 20 number 2). Alain was an
influential carcinologist himself, specialising in penaeoid shrimps, but also publishing on a wide variety of other decapods.
Even today, several years after his retirement from science, many researchers of this “fellowship of the MUSORSTOM”
still correspond with him. For us, he is a classic demonstration of the French proverb “la foi soulève des montagnes” (faith
that moves mountains). And he did move “many large mountains” in advancing carcinology and deep-sea research. e
Alain Crosnier effect is now deeply entrenched in carcinological history.
THE INDISPENSABLE ROLE OF TAXONOMY
e endeavours of the MUSORSTOM-TDSB rammed home the importance of taxonomy as a science. Nowadays,
with the rapid development of powerful new molecular tools, there is a tendency for many people, including respected
colleagues and scientific leaders, to decry morphological taxonomy as “old fashioned” and obsolete, dated, unimportant and
irrelevant. ey think new techniques like DNA barcoding and fingerprinting will replace morphologists. ey are wrong.
e MUSORSTOM-TDSB programme is a clear demonstration that taxonomy is at the heart of good biological science.
Exploration and surveys can collect extensive material for study, but for this material to be useful for science, the right
questions must be asked and hypotheses tested. To have a solid knowledge of zoology, the species in this material have
to be formally described and named. Detractors who argue that a COI gene sequence alone is enough to characterise
a species are fooling themselves. A genetic barcode is not unlike a person’s identification number — it indicates your
position in a list, but not who you are (see also Kwong
et al.
2012; Taylor & Harris 2012).
It is often forgotten (or ignored) that taxonomic classifications and species names, represent hypotheses of biological
relationship. e population to which a species name has been applied has unique biological properties, shared within
the group, and different from the next species. e information embodied in the species’ classification tells us about its
diagnostic and distinctive features, and provides indications as to what the animals may do in life.
ere can be no denying that molecular methods are powerful aids to the work of taxonomists, but they cannot be used
in isolation. Moreover, genetic tools are also limited in their ability to distinguish species, without reference to morphology,
as recently demonstrated for deep-sea corals (Baco & Cairns 2012). Ultimately, knowledge of the morphology is integral to
the human understanding of what animals are and do. Far from being “old-fashioned”, morphological taxonomy remains
a timeless and very essential discipline.
e strong taxonomic foundation deriving from the MUSORSTOM-TDSB programme has led to numerous advances
in our knowledge of deep-sea biology. e many named taxa (families, genera and species) have been used for dozens of
important phylogenetic, biogeographic and ecological questions, many of which have been reviewed above.
NEW PARADIGMS
In the middle of the 19th century, our deep-sea knowledge was very limited and the dominant idea at this time was
that the bathyal world is an extremely harsh environment that was primarily azoic. In the second half of the 19th century,

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
32
especially after the brilliant success of the deep-sea dredgings by Charles Wyville omson, the scientific community
began to realise that there is a high diversity of animals in the deep-sea. England started the race to discover and sample
the world oceans with the CHALLENGER expedition (1872-1876) funded by the Royal Society. Other countries quickly
followed suit: e Netherlands (SIBOGA 1899-1900), Germany (VALDIVIA 1898-1899), British India (INVESTIGATOR
1884-1897), United States (ALBATROSS 1891-1905) and Denmark (GALATHEA 1950-1952). As a result of these surveys,
by the middle of the 20th century, the revised thinking was that there is life in the deep-sea at all depths, but the diversity
is relatively poor with the animals widely distributed.
From 1970 onwards, ORSTOM, under the leadership of Alain Crosnier, started sampling the deep-sea fauna in
Madagascar, originally aimed at exploring fish and crustacean resources. But these explorations were boosted by the
discovery of
Neoglyphea
in 1975 and its rediscovery in 1976 in the Philippines; the same year the American submersible
“Alvin” discovered the astonishing fauna living in and around hydrothermal vents. is extraordinary discovery completely
changed our concept of life in the deep-sea. Several decades of deep-sea studies mainly using “high technology” submersibles
have since focused on this remarkable fauna that is dependent on chemosynthetic ecosystems along the oceanic ridges
(Desbruyères & Segonzac 1997; Laubier 2008; www.noc.soton.ac.uk/chess).
At the same time, the MUSORSTOM-TDSB programme used “low technology” efforts to sample the bathyal zone
around tropical Islands and started to document the “more ordinary fauna”. ese studies have opened several new
paradigms in our understanding of the deep-sea environment: (i) the deep-sea benthos in tropical areas is very diverse in
the upper part of the bathyal zone; (ii) some ecosystems like steep hard cliffs are neglected and very hard to sample but
nevertheless very species rich; (iii) even if some species have a wide distribution, each archipelago generally has its own
unique faunal assemblage; (iv) for the deep-sea epi-benthos in the Pacific, there is a decreasing gradient of diversity from
West to East; with a similar pattern from the West Pacific to West Indian Ocean; (v) organic substrates such as sunken
wood provides a key food resource in the deep-sea, and acts as stepping stones for the dispersion of chemosynthetic
species; and (vi) the thousands of seamounts in the world oceans are generally oases that are concentrate with deep-sea
species and play an important role in the global dispersion of species.
CONCLUSIONS
With the rapid changes of environment, the huge MUSORSTOM-TDSB collection of deep-sea fauna that spans the
last
40
years
constitutes a very valuable archive to understand deep-sea evolution on our planet. Climate change is a hot
topic nowadays. With the Arctic ice disappearing rapidly, sea level rise may increase faster than expected. e increasing
erosion of the coast and discharge of anthropogenically impacted rivers bringing huge amounts of sediments and organic
material into the oceans. e changing hydrological conditions on the surface modify the plankton productivity and may
also modify oceanic currents and deep-sea habitats. All this will ultimately influence the deep-sea fauna significantly.
In only 160 years, our scientific knowledge of the deep-sea has transitioned from a trickle to a flood, giving scientists
an almost “panoramic vision” of the great diversity of life existing on the sea floor of our planet. Not surprisingly, this
diversity is mostly localized at ridges, seamounts and continental slopes. e richest deep-sea areas appear to be situated
in the western tropical Pacific Ocean. However, we have to remain very humble about our knowledge of the deep-sea,
remembering that a large part of it remains unexplored (Observatory Biodiversity Information System: www.iobis.org).
For example, less than 3% of large seamounts have been sampled, and a large part of the zoological collections already
obtained from seamounts and deep-water systems are still unstudied because of an acute lack of taxonomic specialists.
Even for studied species very little is known about the reproduction, growth and behavior of these deep-sea animals. Deep-
sea explorations now mainly focus on the role of chemosynthesis, methane seeps, hydrothermal vents and the origin of
chemosynthetic fauna. To understand, conserve and manage the deep-sea environment, deep-sea biodiversity survey
programs such as TDSB are more important than ever.

The MusorsToM-TDSB deep-sea crusTaceans 33
ACKNOWLEDGEMENTS
A great number of people have contributed to the MUSORSTOM saga in discovering and describing the deep-sea
tropical fauna. is short assessment on crustaceans is an opportunity to thank themall: the crews of the oceanographic
and fishing boats involved in the hundreds of cruises; the technical teams associate with specimen collection, preservation
and curation; the artists for making photographs and drawings; the editors and publishersof the MUSORSTOM-TDSB
volumes; the administrations supporting the researches; the sponsors helping to fund the boats expenses; the network
of international researchers spending time to study the fauna. A special mention for Alain Crosnier who was a pioneer
in many ways animated the research groups (not limited to Crustacea), from the beginning of obtaining funding for the
cruises till the end of publications of the results.
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The MusorsToM-TDSB deep-sea crusTaceans 47
APPENDIX 1
Cruises in the Tropical Deep-Sea Benthos programme and satellite programmes that have yielded deep-sea zoological material, and relevant published cruise
narratives or station lists (updated list since Bouchet et al. (2008).
CRUISE NAME
OR ACRONYM YEAR SHIP AREA NO. OF STATIONS
STATION NUMBERS
(CONSECUTIVE
NUMBERING
CRUISE REPORT IF ANY
ATIMO VATAE 2010 Nosy bé 11 South Madagascar 119 3508-3627
AURORA 2007 2007 DA-BFAR Philippines 117 2653-2764
AZTEQUE 1990 Alis New Caledonia 11 Grandperrin et al. 1990
BATHUS 1 1993 Alis New Caledonia 75 639-713
Richer de Forges & Chevillon 1996
BATHUS 2 1993 Alis New Caledonia 58 714-771
Richer de Forges & Chevillon 1996
BATHUS 3 1993 Alis New Caledonia 78 773-849
Richer de Forges & Chevillon 1996
BATHUS 4 1994 Alis New Caledonia 74 882-955
Richer de Forges & Chevillon 1996
BENTHAUS 2002 Alis Austral Islands 161 1861-2021
BERYX 11 1992 Alis New Caledonia 60 Lehodey et al. 1992
BERYX 2 1991 Alis New Caledonia 19 Grandperrin & Lehodey 1992
BIOCAL 1985 Jean-Charcot New Caledonia 68 Lévi, 2006
BIOGEOCAL 1987 Coriolis New Caledonia 152 Cotillon P. & Monniot C. 1987
BIOPAPUA 2010 Alis Papua New-Guinea 156 3628-3783
BOA0 2004 Alis Vanuatu 27 2304-2330
BOA1 2005 Alis Vanuatu 71 2411-2480
BORDAU 1 1999 Alis Fiji 117 1391-1507 Richer de Forges 2000
BORDAU 2 2000 Alis Tonga 138 1508-1645
CALSUB 1989 Cyana New Caledonia 22 Grandperrin & Richer de Forges
1989; Roux 1994
CHALCAL 1 1984 Coriolis Coral Sea 27 Richer de Forges & Pianet 1984
CHALCAL 2 1986 Coriolis New Caledonia 38 Richer de Forges et al. 1987
CONCALIS 2008 Alis New Caledonia 102 2929-3030
CORAIL 2 1988 Coriolis Coral Sea 19 Richer de Forges et al. 1988
CORINDON 2 1980 Coriolis Indonesia 70 Moosa 1985
EBISCO 2005 Alis Coral Sea 187 1-14 and 2492-2652
EXBODI 2011 Alis New Caledonia 162 3784-3945
GEMINI 1989 Alis New Hebrides Arc 15 Bargibant et al. 1989
HALICAL 1 1994 Alis New Caledonia 4 Grandperr in et al. 1995a
HALIPRO 1 1994 Alis New Caledonia 32 850-881 Grandperrin et al., 1995b
HALIPRO 2 1996 Tangaroa New Caledonia 106 Grandperrin et al. 1997
KARUBAR 1991 Baruna Jaya 1 Indonesia 91 Crosnier et al. 1997
LAGON 1984-89 Vauban, then Alis New Caledonia 62 Richer de Forges 1991
LITHIST 1999 Alis New Caledonia 18
MADANG 2012 Alis Papua New-Guinea 137 3946-4082
MAINBAZA 2009 Vizconde de Eza Mozambique Channel 46 3130-3175
MIRIKY 2009 Miriky Madagascar 119 3176-3294
MUSORSTOM 1 1976 Vauban Philippines 75 1-73 Forest 1981
MUSORSTOM 2 1980 Coriolis Philippines 82 1-83 Forest 1986
MUSORSTOM 3 1985 Coriolis Philippines 59 86-145 Forest 1989
MUSORSTOM 4 1985 Vauban New Caledonia 148 146-249 Richer de Forges 1986
MUSORSTOM 5 1986 Coriolis Coral Sea 137 250-390 Richer de Forges et al. 1986
MUSORSTOM 6 1989 Alis New Caledonia 103 391-493 Richer de Forges & Laboute 1989
MUSORSTOM 7 1992 Alis Wallis & Futuna 147 494-638 Richer de Forges & Menou 1993
MUSORSTOM 8 1994 Alis Vanuatu 186 956-1141 Richer de Forges et al. 1996
MUSORSTOM 9 1997 Alis Marquesas 166 1142-1307 Richer de Forges et al. 1999
MUSORSTOM 10 1998 Alis Fiji 83 1308-1390 Richer de Forges et al. 2000
NORFOLK 1 2001 Alis New Caledonia 88 1651-1739
NORFOLK 2 2003 Alis New Caledonia 142 2022-2161

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48
CRUISE NAME
OR ACRONYM YEAR SHIP AREA NO. OF STATIONS
STATION NUMBERS
(CONSECUTIVE
NUMBERING
CRUISE REPORT IF ANY
PALEO-SURPRISE 1999 Alis New Caledonia 29 Garrigue et al. 2000
PANGLAO 2005 2005 DA-BFAR Philippines 80 2331-2409 Richer de Forges et al. 2009
SALOMON 1 2001 Alis S olomon Islands 121 1740-1860
SALOMON 2 2004 Alis S olomon Islands 142 2164-2303
SALOMONBOA 3 2007 Alis Solomon Islands 97 2765-2860
SANTO 2006 2006 Alis Vanuatu 112
SMCB
1990-91
Marara French Polynesia 80 Poupin 1991
SMIB 1 1986 Vauban New Caledonia 15 Richer de Forges 1990
SMIB 2 1986 Vauban New Caledonia 31 Richer de Forges 1990
SMIB 3 1987 Vauban New Caledonia 32 Richer de Forges 1990
SMIB 4 1989 Alis New Caledonia 36 Richer de Forges 1990
SMIB 5 1989 Alis New Caledonia 36 Richer de Forges 1993
SMIB 6 1990 Alis New Caledonia 32 Richer de Forges 1993
SMIB 8 1993 Alis New Caledonia 55 Richer de Forges & Chevillon 1996
SMIB 9 1993 Alis New Caledonia 1
SMIB 10 1995 Alis New Caledonia 17 Richer de Forges 1990
TAIWAN 2000 2000 Fisheries Researcher 1 Taiwan 62
TAIWAN 2001 2001 Chung Tung Long 26 Taiwan 80
TAIWAN 2002 2002 Cheng-Ming Fa and
Ocean Researcher 1
Taiwan 35
TAIWAN 2004 2004 Rih-Jheng 101 and Ocean
Researcher 2 Taiwan 23
TARASOC 2009 Alis French Polynesia 213 3295-3507
TERRASSES 2008 Alis New Caledonia 99 3031-3129
Vauban 1978-79 1978-79
Vauban New Caledonia 47
VOLSMAR 1989 Alis New Hebrides Arc 25 Laboute et al. 1989

The MusorsToM-TDSB deep-sea crusTaceans 49
PENAEOIDEA
ARISTEIDAE
Aristaeomorpha foliacea (Risso, 1827)
Aristaeopsis edwardsiana (Johnson, 1868)
Aristeus mabahissae Ramadan, 1938
Aristeus semidentatus Bate, 1881
Aristeus virilis (Bate, 1881)
Austropenaeus nitidus (Barnard, 1947)
Hemipenaeus carpenteri Wood-Mason & Alcock, 1891
Plesiopenaeus armatus (Bate, 1881)
BENTHESICYMIDAE
Benthesicymus howensis Dall, 2001
Benthesicymus investigatoris Alcock & Anderson, 1899
Benthesicymus urinator Burkenroad, 1936
Gennadas bouvieri Kemp, 1909
Gennadas capensis Calman, 1925
Gennadas clavicarpus de Man, 1907
Gennadas crassus Tirmizi, 1960
Gennadas gilchristi Calman, 1925
Gennadas incertus (Balss, 1927)
PENAEIDAE
Funchalia villosa (Bouvier, 1905)
Metapenaeopsis dicilis Crosnier, 1991
Metapenaeopsis evermanni (Rathbun, 1906)
Metapenaeopsis provocatoria Racek & Dall, 1965
Metapenaeopsis sibogae (De Man, 1907)
Metapenaeopsis velutina (Dana, 1852)
Parapenaeus cayrei Crosnier, 2005
Parapenaeus ssurus (Bate, 1881)
Parapenaeus investigatoris Alcock & Anderson, 1899
Parapenaeus kensleyi Crosnier, 2005 *
Parapenaeus murrayi Ramadan, 1938
Parapenaeus perezfarfantae Crosnier, 1986
Parapenaeus sextuberculatus Kubo, 1949
Penaeopsis balssi Ivanov & Hassan, 1976
Penaeopsis challengeri De Man, 1911
Penaeopsis eduardoi Pérez Farfante, 1977
Penaeopsis jerryi Pérez Farfante, 1979
Penaeopsis mclaughlinae Crosnier, 2006 *
Penaeopsis rectacuta (Bate, 1881)
SICYONIIDAE
Sicyonia adunca Crosnier, 2003 *
Sicyonia altirostrum Crosnier, 2003 *
Sicyonia australiensis Hanamura & Wadley, 1998
Sicyonia benthophila De Man, 1907
Sicyonia curvirostris Balss, 1913
Sicyonia dejouanneti Crosnier, 2003 *
Sicyonia fallax De Man, 1907
Sicyonia inexa (Kubo, 1949)
Sicyonia laevis Bate, 1881
Sicyonia lancifer (Olivier, 1811)
Sicyonia rotunda Crosnier, 2003 *
Sicyonia trispinosa De Man, 1907
Sicyonia truncata (Kubo, 1949)
Sicyonia vitulans (Kubo, 1949)
SOLENOCERIDAE
Cryptopenaeus clevai Crosnier, 1985
Cryptopenaeus crosnieri Pérez Farfante & Kensley, 1985
Gordonella kensleyi Crosnier, 1988a *
Gordonella paravillosa Crosnier, 1988a *
Hadropenaeus lucasii (Bate, 1881)
Hadropenaeus spinicaudatus Liu & Zhong, 1983
Haliporoides cristatus Kensley, Tranter & Grin, 1987
Haliporoides sibogae sibogae (De Man, 1907)
Haliporus thetis Faxon, 1893
Hymenopenaeus debilis Smith, 1882
Hymenopenaeus equalis (Bate, 1888)
Hymenopenaeus halli Bruce, 1966
Hymenopenaeus methalli Crosnier & Dall, 2004 *
Hymenopenaeus neptunus (Bate, 1881)
Hymenopenaeus obliquirostris (Bate, 1881)
Hymenopenaeus propinquus (De Man, 1907)
Mesopenaeus brucei Crosnier, 1986
Solenocera comata Stebbing, 1915
SERGESTOIDEA
SERGESTIDAE
Sergia talismani (Barnard, 1947)
STENOPODIDEA
SPONGICOLIDAE
Globospongicola spinulatus Komai & Saito, 2006
Paraspongicola inatus (de Saint Laurent & Cleva, 1981)
Spongicola andamanicus Alcock, 1901
Spongicola depressus Saito & Komai, 2008 *
Spongicola goyi Saito & Komai, 2008 *
Spongicola levigatus Hayashi & Ogawa, 1987
CARIDEA
ACANTHEPHYRIDAE
Acanthephyra armata A. Milne- Edwards, 1881
Acanthephyra curtirostris Wood-Mason & Alcock, 1891
Acanthephyra eximia Smith, 1884
Acanthephyra quadrispinosa Kemp, 1939
Acanthephyra sanguinea Wood-Mason [in Wood-Mason &
Alcock, 1892]
Acanthephyra smithi Kemp, 1939
Acanthephyra tenuipes (Bate, 1888)
Heterogenys microphthalma (Smith, 1885)
Meningodora vesca (Smith, 1886)
ALPHEIDAE
Batella praecipua De Grave, 2004 *
ANCHISTIOIDIDAE
Anchistioides willeyi (Borradaile, 1900)
BATHYPALAEMONELLIDAE
Bathypalaemonella hayashii Komai, 1995
Bathypalaemonella serratipalma Pequegnat, 1970
Bathypalaemonetes brevirostris (Bruce, 1986)
Bathypalaemonetes pilosipes (Bruce, 1986)
BRESILIIDAE
Bresilia antipodarum Bruce, 1990
CRANGONIDAE
Aegaeon lacazei (Gourret, 1887)
Aegaeon rathbuni De Man, 1918
Lissosabinea armata Komai, 2006 *
Lissosabinea indica (De Man, 1918)
Lissosabinea unispinosa Komai, 2006 *
Parapontocaris aspera Chace, 1984
Parapontophilus caledonicus Komai, 2008 *
Parapontophilus cyrton Komai, 2008 *
Parapontophilus demani (Chace, 1984)
Parapontophilus dicilis Komai, 2008
Parapontophilus junceus (Bate, 1888)
Parapontophilus longirostris Komai, 2008
Parapontocaris levigata Chace, 1984
Pontocaris propensalata Bate, 1888
Pontocaris sibogae (De Man, 1918)
Pseudopontophilus serratus Komai, 2004a *
EUGONATONOTIDAE
Eugonatonotus chacei Chan & Yu, 1991 *
GLYPHOCRANGONIDAE
Glyphocrangon armata Komai, 2004b *
Glyphocrangon conodactylus Komai, 2004b *
Glyphocrangon cornuta Komai, 2004b *
Glyphocrangon dimorpha Komai, 2004b *
Glyphocrangon musorstomia Komai, 2006
Glyphocrangon parviocullus Komai, 2006 *
Glyphocrangon punctata Komai, 2004 *
Glyphocrangon regalis Bate, 1888
Glyphocrangon richeri Komai, 2004b
Glyphocrangon similior Komai, 2004
Glyphocrangon speciosa Komai, 2004b *
HIPPOLYTIDAE
Gelastreutes crosnieri Bruce, 1990b *
NEMATOCARCINIDAE
Nematocarcinus combensis Burukovsky, 2000
Nematocarcinus crosnieri Burukovsky, 2000 *
Nematocarcinus gracilis Bate, 1888
Nematocarcinus parvus Burukovsky, 2000
APPENDIX 2
Check-list of deep-sea decapod crustaceans (recorded below 100 m depth) from New Caledonia. An asterisk * indicates that the species was originally
described from New Caledonia. References to the original descriptions are listed in Appendix 4.The classication and the succession of the families follow
De Grave et al. (2009).

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
50
Nematocarcinus paucidentatus Bate, 1888
Nematocarcinus productus Bate, 1888
Nematocarcinus richeri Burukovsky, 2000
Nematocarcinus tenuipes Bate, 1888
Nematocarcinus tenuirostris Bate, 1888
Nigmatullinus acanthitelsonis (Pequegnat, 1970)
OPLOPHORIDAE
Janicella spinicauda (A. Milne-Edwards, 1883)
Oplophorus gracilirostris A. Milne-Edwards, 1881
Oplophorus spinosus (Brullé, 1839)
Systellaspis debilis (A.Milne- Edwards, 1881)
Systellaspis lanceocaudata Bate, 1888
Systellaspis pellucida (Filhol, 1884)
PALAEMONIDAE
Altopontonia disparostris Bruce, 1990a *
Amphipontonia kanak Bruce, 1991 *
Anchistus pectinis Kemp, 1925
Ancylomenes tenuirostris (Bruce, 1991) *
Cuapetes tenuipes (Borradaile, 1898)
Dactylonia monnioti (Bruce, 1990a) *
Mesopontonia gorgoniophila Bruce, 1967
Mesopontonia gracilicarpus Bruce, 1990a *
Mesopontonia monodactylus Bruce, 1991 *
Palaemonella dolichodactylus Bruce, 1991 *
Palaemonella rotumana (Borradaile, 1898)
Paraclimenes franklini (Bruce, 1990a) *
Paraclimenes setirostris (Bruce, 1991) *
Periclimenaeus jeancharcoti Bruce, 1991 *
Periclimenes acanthimerus Bruce, 2006 *
Periclimenes albatrossae Chace & Bruce, 1993
Periclimenes alcocki Kemp, 1922
Periclimenes aleator Bruce, 1991 *
Periclimenes brevirostris Bruce, 1991 *
Periclimenes forcipulatus Bruce, 1991 *
Periclimenes forgesi Li & Bruce, 2006 *
Periclimenes foveolatus Bruce, 1981
Periclimenes fujinoi Bruce, 1990a *
Periclimenes hertwigi Balss, 1913
Periclimenes laccadivensis (Alcock & Anderson, 1894)
Periclimenes latipollex Kemp, 1922
Periclimenes leptodactylus Bruce, 1991 *
Periclimenes loyautensis Li & Bruce, 2006 *
Periclimenes ordinarius Bruce, 1991 *
Periclimenes paraleator Li & Bruce, 2006 *
Periclimenes parvispinatus Bruce, 1990a *
Periclimenes pectinipes Bruce, 1991 *
Periclimenes platyrhynchus Bruce, 1991 *
Periclimenes rectirostris Bruce, 1981
Periclimenes richeri Bruce, 1990a *
Periclimenes uniunguiculatus Bruce, 1990a *
Periclimenes vaubani Bruce, 1990a *
Urocaridella urocaridella (Holthuis, 1950)
PANDALIDAE
Bitias brevis (Rathbun, 1906)
Calipandalus elachys Komai & Chan, 2003
Chelonika macrochela Fransen, 1997 *
Heterocarpus amacula Crosnier, 1988b
Heterocarpus dorsalis Bate, 1888
Heterocarpus ensifer A. Milne-Edwards, 1881
Heterocarpus hayashii Crosnier, 1988b *
Heterocarpus intermedius Crosnier, 1999 *
Heterocarpus laevigatus Bate, 1888
Heterocarpus lepidus De Man, 1917
Heterocarpus longirostris MacGilchrist, 1905
Heterocarpus parvispina De Man, 1917
Heterocarpus sibogae De Man, 1917
Heterocarpus tricarinatus Alcock & Anderson, 1894
Heterocarpus woodmasoni Alcock, 1901
Pandalus (Parapandalus) longirostris Borradaile, 1900
Plesionika alcocki (Anderson, 1896)
Plesionika bimaculata Chan, 2004
Plesionika echinicola Chan & Crosnier, 1991 *
Plesionika ensis (A. Milne-Edwards, 1881)
Plesionika erythrocyclus Chan & Crosnier, 1997
Plesionika avicauda Chan & Crosnier, 1991
Plesionika hsuehyui Chan, 2004
Plesionika laurentae Chan & Crosnier, 1991 *
Plesionika macropoda Chace, 1939
Plesionika reexa Chace, 1985
Plesionika rufomaculata Chan, 2004 *
Plesionika scopifera Chan, 2004 *
Plesionika semilaevis Bate, 1888
Plesionika serratifrons (Borradaile, 1900)
Plesionika spinipes Bate, 1888
Plesionika suusa Chan, 2004 *
Pseudopandalus curvirostris Crosnier, 1997a *
PASIPHAEIDAE
Leptochela (Leptochela) japonica Hayashi & Miyake, 1969
Pasiphaea fragilis Hayashi, 1999 *
Pasiphaea gracilis Hayashi, 1999 *
Pasiphaea kapala Kensley, Tranter & Grin, 1987
Pasiphaea major Hayashi, 2006 *
Pasiphaea mclaughlinae Hayashi, 2006 *
Pasiphaea planidorsalis Hayashi, 2004 *
Pasiphaea sirenkoi Burukovsky, 1987
Pasiphaea telacantha Hayashi, 2004 *
PSALIDOPODIDAE
Psalidopus huxleyi Wood-Mason [in Wood-Mason & Alcock, 1892]
STYLODACTYLIDAE
Neostylodactylus anis Hayashi & Miyake, 1968
Parastylodactylus bimaxillaris Bate, 1888)
Parastylodactylus richeri Cleva, 1990 *
Parastylodactylus semblatae Cleva, 1990 *
Parastylodactylus tranterae Cleva, 1990 *
Stylodactyloides crosnieri Cleva, 1990 *
Stylodactylus discissipes Bate, 1888
Stylodactylus laurentae Cleva, 1990 *
Stylodactylus libratus Chace, 1983
Stylodactylus licinus Chace, 1983
Stylodactylus macropus Chace, 1983
Stylodactylus multidentatus multidentatus Kubo, 1942
Stylodactylus profundus Cleva, 1990 *
Stylodactylus tokarensis Zarenkov, 1968
ASTACIDEA
ENOPLOMETOPIDAE
Enoplometopus gracilipes (Saint Laurent, 1988)
NEPHROPIDAE
Acanthacaris tenuimana Bate, 1888
Nephropsis acanthura Macpherson, 1990
Nephropsis suhmi Bate, 1888
Nephropsis sulcata Macpherson, 1990
Thaumastocheles japonicus Calman, 1913
GLYPHEIDEA
GLYPHEIDAE
Laurentaeglyphea neocaledonica (Richer de Forges, 2006) *
AXIIDEA
AXIIDAE
Acanthaxius miyazakiensis (Yokoya, 1933)
Ambiaxius alcocki (McArdle, 1900)
Bouvieraxius rudis (Rathbun, 1906)
Oxyrhynchaxius tricarinatus Lin, 2006 *
Spongiaxius holthuisi Poore & Collins, 2010 *
CALLIANASSIDAE
Callianassa joculatrix de Man, 1905
Cheramus propinquus (De Man, 1905)
MICHELEIDAE
Meticonaxius dentatus Lin, 2006 *
Meticonaxius noumea Poore, 1997 *
Tethisea indica Poore, 1994
STRAHLAXIIDAE
Neaxius trondlei Ngoc-Ho, 2005
ACHELATA
PALINURIDAE
Linuparus meridionalis Tsoi, Chan & Chu, 2011 *
Linuparus sordidus Bruce, 1965
Nupalirus chani (Poupin, 1994) *
Nupalirus japonicus Kubo, 1955
Palinustus unicornutus Berry, 1979
Puerulus mesodontus, Chan, Ma & Chu, this volume
Puerulus quadridentis, Chan, Ma & Chu, this volume *
Puerulus richeri, Chan, Ma & Chu, this volume
Puerulus sericus, Chan, Ma & Chu this volume *
SCYLLARIDAE
Bathyarctus chani Holthuis, 2002 *
Bathyarctus rubens (Alcock & Anderson, 1894)
Eduarctus martensii (Pfeer, 1881)
Galearctus aurora (Holthuis, 1982)
Galearctus avulsus Yang, Chen & Chan, 2011 *
Galearctus timidus (Holthuis, 1960)
Ibacus brevipes Bate, 1888
Ibacus brucei Holthuis, 1977
Petrarctus brevicornis (Holthuis, 1946)
Remiarctus bertholdii (Paulson, 1875)
Scammarctus batei (Holthuis, 1946)

The MusorsToM-TDSB deep-sea crusTaceans 51
POLYCHELIDA
POLYCHELIDAE
Pentacheles laevis Bate, 1878
Pentacheles obscurus Bate, 1878
Pentacheles validus A. Milne-Edwards, 1880
Polycheles baccatus Bate, 1878
Polycheles coccifer Galil, 2000
Polycheles enthrix (Bate, 1878)
Polycheles typhlops Heller, 1862
Stereomastis aculeata (Galil, 2000) *
Stereomastis auriculata (Bate, 1878)
Stereomastis helleri (Bate, 1878)
Stereomastis nana (Smith, 1884)
Stereomastis phosphorus (Alcock, 1894)
Stereomastis surda (Galil, 2000)
ANOMURA
ALBUNEIDAE
Austrolepidopa caledonia Boyko & Harvey, 1999 *
Paralbunea dayriti (Serène & Umali, 1965)
Zygopa nortoni Serène & Umali, 1965
CHIROSTYLIDAE
Chirostylus novaecaledoniae Baba, 1991a *
Gastroptychus brevipropodus Baba, 1991a *
Gastroptychus hendersoni (Alcock & Anderson, 1899)
Gastroptychus paucispina Baba, 1991a *
Gastroptychus sternoornatus (Van Dam, 1933)
Uroptychodes epigaster Baba, 2004 *
Uroptychodes musorstomi Baba, 2004 *
Uroptychodes spinimarginatus (Henderson, 1885)
Uroptychus cavirostris Alcock & Anderson, 1899
Uroptychus tridentatus (Henderson, 1885)
DIOGENIDAE
Areopaguristes micheleae (Rahayu, 2005)
Areopaguristes ngochoae (Rahayu, 2005)
Ciliopagurus alcocki Forest, 1995 *
Ciliopagurus kremp (Forest, 1952)
Ciliopagurus pacicus Forest, 1995
Paguristes aciculus Grant, 1905
Paguristes alcocki McLaughlin & Rahayu, 2005
Paguristes ortmanni Miyake, 1978
Paguristes palythophilus Ortmann, 1892
Paguristes puniceus Henderson, 1896
Paguristes versus Komai, 2001
Pseudopaguristes laurentae (Morgan & Forest, 1991)
Pseudopaguristes monoporus (Morgan, 1987)
Strigopagurus boreonotus Forest, 1995 *
EUMUNIDIDAE
Eumunida annulosa de Saint Laurent & Macpherson, 1990 *
Eumunida capillata de Saint Laurent & Macpherson, 1990 *
Eumunida karubar de Saint Laurent & Poupin, 1996
Eumunida keijii de Saint Laurent & Macpherson, 1990 *
Eumunida marginata de Saint Laurent & Macpherson, 1990 *
Eumunida minor de Saint Laurent & Macpherson, 1990 *
Eumunida pacica Gordon, 1930
Eumunida parva de Saint Laurent & Macpherson, 1990 *
Eumunida spinosa Macpherson, 2006b *
Eumunida sternomaculata de Saint Laurent & Macpherson,
1990 *
Pseudomunida fragilis Haig, 1979
GALATHEIDAE
Alainius crosnieri Baba, 1991b *
Allogalathea babai Cabezas, Macpherson & Machordom, 2011 *
Allogalathea elegans (Adams & White, 1848)
Allogalathea inermis Cabezas, Macpherson & Machordom, 2011 *
Galathea barbellata Macpherson, 2012 *
Galathea echinata Macpherson, 2012 *
Galathea profunda Macpherson, 2012
Galathea sanctae Macpherson, 2012
Leiogalathea laevirostris (Balss, 1913)
Phylladiorhynchus ikedai (Miyake & Baba, 1965)
Phylladiorhynchus pusillus (Henderson, 1885)
LITHODIDAE
Lithodes richeri Macpherson, 1990 *
Neolithodes bronwynae Ahyong, 2010
Paralomis dawsoni Macpherson, 2001 *
Paralomis haigae Eldredge, 1976
MUNIDIDAE
Agononida andrewi (Macpherson, 1994) *
Agononida callirrhoe (Macpherson, 1994) *
Agononida eminens (Baba, 1988)
Agononida incerta (Henderson, 1888)
Agononida laurentae (Macpherson, 1994) *
Agononida marini (Macpherson, 1994) *
Agononida norfocerta Poore & Andreakis, 2012
Agononida normani (Henderson, 1885)
Agononida ocyrhoe (Macpherson, 1994) *
Agononida rubrizonata Macpherson & Baba, 2009 *
Agononida sabatesae (Macpherson, 1994) *
Agononida soelae (Baba, 1986)
Agononida sphecia (Macpherson, 1994) *
Agononida squamosa (Henderson, 1885)
Anoplonida inermis (Baba, 1994)
Babamunida callista (Macpherson, 1994) *
Babamunida javieri (Macpherson, 1994) *
Bathymunida balssi Van Dam, 1938
Bathymunida eurybregma Baba & de Saint Laurent, 1996 *
Bathymunida nebulosa Baba & de Saint Laurent, 1996 *
Bathymunida ocularis Baba & de Saint Laurent, 1996 *
Bathymunida rudis Baba & de Saint Laurent, 1996 *
Bathymunida sibogae Van Dam, 1938
Crosnierita dicata Macpherson, 1998 *
Crosnierita urizae (Macpherson, 1994) *
Crosnierita yante (Macpherson, 1994)
Enriquea leviantennata (Baba, 1988)
Heteronida aspinirostris (Khodkina, 1981)
Munida acantha Macpherson, 1994 *
Munida alonsoi Macpherson, 1994 *
Munida amblytes Macpherson, 1994 *
Munida angusta Macpherson, 2004
Munida armilla Macpherson, 1994 *
Munida asprosoma Ahyong & Poore, 2004
Munida aulakodes Macpherson, 2006b *
Munida barangei Macpherson, 1994 *
Munida brachytes Macpherson, 1994 *
Munida clinata Macpherson, 1994 *
Munida congesta Macpherson, 2000
Munida delicata Macpherson, 2004 *
Munida devestiva Macpherson, 2006b *
Munida distiza Macpherson, 1994 *
Munida eclepsis Macpherson, 1994 *
Munida elachia Macpherson, 1994 *
Munida erato Macpherson, 1994 *
Munida gilii Macpherson, 1993
Munida gordoae Macpherson, 1994 *
Munida guttata Macpherson, 1994 *
Munida hyalina Macpherson, 1994 *
Munida idyia Macpherson, 1994 *
Munida inornata Henderson, 1885
Munida japonica Stimpson, 1858
Munida leagora Macpherson, 1994 *
Munida leptitis Macpherson, 1994 *
Munida lineola Macpherson, 1994 *
Munida magniantennulata Baba & Türkay, 1992
Munida masi Macpherson, 1994 *
Munida microps Alcock, 1894
Munida moliae Macpherson, 1994 *
Munida notata Macpherson, 1994 *
Munida ommata Macpherson, 2004 *
Munida oritea Macpherson & Baba, 1993
Munida pagesi Macpherson, 1994 *
Munida parca Macpherson, 1996a *
Munida parile MacPherson & Machordom, 2005 *
Munida pectinata MacPherson & Machordom, 2005 *
Munida pontoporea Macpherson, 1994 *
Munida proto Macpherson, 1994 *
Munida psamathe Macpherson, 1994 *
Munida pseliophora Macpherson, 1994 *
Munida psylla Macpherson, 1994 *
Munida pygmaea Macpherson, 1996a *
Munida rhodonia Macpherson, 1994 *
Munida rogeri Macpherson, 1994 *
Munida rosula Macpherson, 1994 *
Munida rubridigitalis Baba, 1994
Munida ruantennulata Baba, 1969
Munida runcinata Macpherson, 1994 *
Munida sacksi Macpherson, 1993
Munida sao Macpherson, 1994 *
Munida semoni Ortmann, 1894
Munida simulatrix MacPherson & Machordom, 2005 *
Munida spilota Macpherson, 1994 *
Munida stia Macpherson, 1994 *
Munida stigmatica Macpherson, 1994 *
Munida striola Macpherson & Baba, 1993
Munida taenia Macpherson, 1994 *
Munida thoe Macpherson, 1994 *
Munida tiresias Macpherson, 1994 *
Munida tuberculata Henderson, 1885
Munida tyche Macpherson, 1994 *
Munida typhle Macpherson, 1994 *
Munida zebra Macpherson, 1994 *
Onconida alaini Baba & de Saint Laurent, 1996 *
Onconida tropis Baba & de Saint Laurent, 1996
Paramunida amphitrita Macpherson, 1996b

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52
Paramunida antares Cabezas, Macpherson & Machordom, 2010 *
Paramunida belone Macpherson, 1993 *
Paramunida granulata (Henderson, 1885)
Paramunida labis Macpherson, 1996b
Paramunida longior Baba, 1988
Paramunida microrhina Cabezas, Macpherson & Machordom,
2010 *
Paramunida parvispina Cabezas, Macpherson & Machordom,
2010 *
Paramunida pictura Macpherson, 1993 *
Paramunida pronoe Macpherson, 1993 *
Paramunida setigera Baba, 1988
Paramunida stichas Macpherson, 1993 *
Paramunida tenera Cabezas, Macpherson & Machordom, 2010 *
Paramunida thalie Macpherson, 1993 *
Plesionida aliena (Macpherson, 1996a) *
Plesionida psila Baba & de Saint Laurent, 1996 *
Raymunida bellior (Miyake & Baba, 1967)
Raymunida confundens Macpherson & Machordom, 2001 *
Raymunida dextralis Macpherson & Machordom, 2001 *
Torbenella calvata (Macpherson, 2006b) *
Torbenella insolita (Macpherson, 2004) *
Torbenella orbis (Baba, 2005)
MUNIDOPSIDAE
Galacantha bellis Henderson, 1885
Galacantha quiquei Macpherson, 2007
Galacantha rostrata A. Milne Edwards, 1880
Leiogalathea laevirostris (Balss, 1913)
Munidopsis analoga Macpherson, 2007 *
Munidopsis andamanica MacGilchrist, 1905
Munidopsis arenula Macpherson, 2007 *
Munidopsis carinimarginata Baba, 1988
Munidopsis centrina Alcock & Anderson, 1894
Munidopsis ceres Macpherson, 2007 *
Munidopsis cornuata Macpherson, 2007 *
Munidopsis cylindrophthalma (Alcock, 1894)
Munidopsis debilis (Henderson, 1885)
Munidopsis kensleyi Ahyong & Poore, 2004
Munidopsis latimana Miyake & Baba, 1966
Munidopsis nitida (A. Milne Edwards, 1880)
Munidopsis orcina McArdle, 1901
Munidopsis sinclairi McArdle, 1901
Munidopsis tafrii Osawa, Lin & Chan, 2006
Munidopsis ternaria Macpherson, 2007 *
Munidopsis treis Ahyong & Poore, 2004
Munidopsis trida Henderson, 1885
PAGURIDAE
Alainopaguroides megalophthalmus McLaughlin, 2006 *
Alainopagurus crosnieri Lemaitre & McLaughlin, 1995 *
Anapagurus bonnieri Nobili, 1905
Catapagurus danida McLaughlin, 2002
Catapagurus franklinae McLaughlin, 2004a *
Catapagurus imperialis (Asakura, 2001)
Catapagurus spinicarpus de Saint Laurent & McLaughlin, 2000
Catapagurus tanimbarensis McLaughlin, 1997
Diacanthurus ecphyma McLaughlin & Forest, 1997 *
Diacanthurus richeri McLaughlin & Forest, 1997 *
Icelopagurus crosnieri McLaughlin, 1997
Icelopagurus undulatus McLaughlin, 2006 *
Michelopagurus limatulus (Henderson, 1888)
Micropagurus Polynesiansis (Nobili, 1906)
Micropagurus spinimanus Asakura, 2005 *
Nematopagurus alcocki McLaughlin, 1997
Nematopagurus australis (Henderson, 1888)
Nematopagurus diadema Lewinsohn, 1969
Nematopagurus gardineri Alcock, 1905
Nematopagurus indicus Alcock, 1905
Nematopagurus kosiensis McLaughlin, 1998
Nematopagurus lepidochirus (Doein, 1902)
Nematopagurus lewinsohni Türkay, 1986
Nematopagurus meiringae McLaughlin, 1998
Nematopagurus ricei McLaughlin, 2004b *
Nematopagurus richeri McLaughlin, 2004b *
Nematopagurus scutelliformis McLaughlin, 1997
Nematopagurus spinulosensoris McLaughlin & Brock, 1974
Nematopagurus spongioparticeps McLaughlin, 2004b *
Nematopagurus vallatus (Melin, 1939)
Pagurojacquesia polymorpha (de Saint Laurent & McLaughlin,
1999)
Porcellanopagurus chiltoni de Saint Laurent & McLaughlin, 2000
Porcellanopagurus lholi de Saint Laurent & McLaughlin, 2000
Porcellanopagurus haptodactylus McLaughlin, 2000 *
Porcellanopagurus tridentatus Whitelegge, 1900
Propagurus haigae (McLaughlin, 1997)
Pteropagurus inermis McLaughlin & Rahayu, 2006 *
Pteropagurus spina McLaughlin & Rahayu, 2006 *
Pteropagurus spinulocarpus McLaughlin, 2007a *
Solitariopagurus triprobolus Poupin & McLaughlin, 1996
Solitariopagurus trullirostris McLaughlin, 2000 *
Solitariopagurus tuerkayi McLaughlin, 1997
Tomopaguroides valdividae (Balss, 1911)
Turleania boucheti McLaughlin, 2007b
Turleania multispina McLaughlin, 1997
Turleania senticosa (McLaughlin & Haig, 1996)
Xylopagurus caledonicus Forest, 1997 *
PARAPAGURIDAE
Oncopagurus conicus Lemaitre, 2006 *
Paragiopagurus acutus (de Saint Laurent, 1972)
Paragiopagurus bicarinatus (de Saint Laurent, 1972)
Paragiopagurus boletifer (de Saint Laurent, 1972)
Paragiopagurus diogenes (Whitelegge, 1900)
Paragiopagurus hirsutus (de Saint Laurent, 1972)
Paragiopagurus orthotenes, Lemaitre, this volume *
Paragiopagurus oxychelos, Lemaitre, this volume *
Paragiopagurus ruticheles (A. Milne-Edwards, 1891)
Paragiopagurus schnauzer Lemaitre, 2006 *
Paragiopagurus trilineatus, Lemaitre, this volume *
Paragiopagurus umbonatus, Lemaitre, this volume *
Parapagurus furici Lemaitre, 1999 *
Parapagurus latimanus Henderson, 1888
Parapagurus richeri Lemaitre, 1999 *
Strobopagurus breviacus Lemaitre, 2004a *
Strobopagurus gracilipes (A. Milne-Edwards, 1891)
Strobopagurus sibogae (de Saint Laurent, 1972)
Sympagurus acinops Lemaitre, 1989 *
Sympagurus anis (Henderson, 1888)
Sympagurus aurantium Lemaitre, 2004b *
Sympagurus brevipes (de Saint Laurent, 1972)
Sympagurus burkenroadi Thompson, 1943
Sympagurus doeini (Balss, 1912)
Sympagurus planimanus (de Saint Laurent, 1972)
Sympagurus poupini Lemaitre, 1994
Sympagurus soela Lemaitre, 1996 *
Sympagurus symmetricus Lemaitre, 2004b *
Sympagurus trispinosus (Balss, 1911)
Sympagurus villosus Lemaitre, 1996 *
Tylaspis anomala Henderson, 1885
PYLOCHELIDAE
Cheiroplatea pumicicola Forest, 1987
Trizocheles caledonicus Forest, 1987 *
Trizocheles pilgrimi Forest & McLaughlin, 2000
Trizocheles pulcher Forest, 1987 *
Trizocheles spinosus (Henderson, 1888)
Trizocheles vaubanae McLaughlin & Lemaitre, 2008 *
BRACHYURA
AETHRIDAE
Aethra scruposa (Linnaeus, 1764)
CALAPPIDAE
Mursia armata De Haan, 1837
Mursia australiensis Campbell, 1971
Mursia longispina Crosnier, 1997b *
Mursia microspina Davie & Short, 1989
Mursia musorstomia Galil, 1993 *
Mursia trispinosa Parisi, 1914
CANCRIDAE
Platepistoma nanum Davie, 1991 *
CRYPTOCHIRIDAE
Cecidocarcinus zibrowii Manning, 1991 *
CYCLODORIPPIDAE
Ketamia depressa (Ihle, 1916)
Krangalangia orstom Tavares, 1993 *
Krangalangia spinosa (Zarenkov, 1970)
Tymolus daviei Tavares, 1997
Xeinostoma richeri Tavares, 1993 *
CYMONOMIDAE
Elassopodus stellatus Tavares, 1993 *
DROMIIDAE
Cryptodromia coronata Stimpson, 1858
Cryptodromia hilgendor de Man, 1888
Eodromia denticulata McLay, 1993 *
Epigodromia areolata (Ihle, 1913)
Epigodromia ebalioides (Alcock, 1899)
Epigodromia nodosa Sakai, 1936
Epigodromia rotunda McLay, 1993 *
Frodromia atypica (Sakai, 1936)
Lewindromia unidentata (Rüppell, 1830)
Metadromia wilsoni (Fulton & Grant, 1902)
Petalomera pulchra Miers, 1884
Sphaerodromia lamellata Crosnier, 1994 *

The MusorsToM-TDSB deep-sea crusTaceans 53
Stimdromia foresti (McLay, 1993) *
Takedromia cristatipes (Sakai, 1969)
Takedromia longispina McLay, 1993 *
DYNOMENIDAE
Dynomene hispida (Latreille, in Milbert, 1812)
Dynomene pilumnoides Alcock, 1900
Metadynomene tanensis (Yokoya, 1933)
Paradynomene demon McLay & Ng, 2004 *
Paradynomene diablo McLay & Ng, 2004 *
Paradynomene quasimodo McLay & Ng, 2004
Paradynomene tuberculata Sakai, 1963
EPIALTIDAE
Guinotinia cordis Richer de Forges & Ng, 2009a *
Oxypleurodon alaini Richer de Forges & Ng, 2009b *
Oxypleurodon mammatum (Guinot & Richer de Forges, 1986b) *
Oxypleurodon orbiculatum (Guinot & Richer de Forges, 1986a) *
Oxypleurodon stuckiae (Guinot & Richer de Forges, 1986a) *
Stegopleurodon planirostrum Richer de Forges & Ng, 2009b *
ETHUSIDAE
Ethusa abbreviata Castro, 2005a
Ethusa crosnieri Chen, 1993 *
Ethusa curvipes Chen, 1993 *
Ethusa furca Chen, 1993 *
Ethusa granulosa Ihle, 1916
Ethusa indica Alcock, 1894
Ethusa izuensis Sakai, 1937
Ethusa magnipalmata Chen, 1993 *
Ethusa minuta Sakai, 1937
Ethusa obliquedens Chen, 1993 *
Ethusa orientalis Miers, 1886
Ethusa parapygmaea Chen, 1993 *
Ethusa pygmaea Alcock, 1894
Ethusa quadrata Sakai, 1937
Ethusina brevidentata Chen, 1993 *
Ethusina ciliacirrata Castro, 2005a
Ethusina coronata Castro, 2005a
Ethusina dilobotus Chen, 1993 *
Ethusina microspina Chen, 2000
Ethusina paralongipes Chen, 1993 *
Ethusina pubescens Chen, 1993 *
Ethusina robusta (Miers, 1886)
Ethusina stenommata Castro, 2005a
Ethusina vanuatuensis Chen, 2000
EURYPLACIDAE
Eucrate laevis (Borradaile, 1903)
GERYONIDAE
Chaceon bicolor Manning & Holthuis, 1989 *
GONEPLACIDAE
Carcinoplax cooki Rathbun 1906
Carcinoplax cracens Castro, 2007 *
Carcinoplax ischurodous (Stebbing, 1923)
Carcinoplax specularis Rathbun, 1914
Carcinoplax tenuidentata Castro, 2007
Carcinoplax tuberosa Castro, 2007 *
Carcinoplax uncinata Castro, 2009 *
Goneplacoides marivenae (Komatsu & Takeda, 2004)
Menoplax longispinosa (Chen, 1984)
Microgoneplax elegans (Chen, 1998)
Microgoneplax prion Castro, 2007 *
Neogoneplax renoculis (Rathbun, 1914)
Neogoneplax serratipes Castro, 2007 *
Paragoneplax serenei (Zarenkov, 1972)
Psopheticus stridulans Wood- Mason, 1892
Psopheticus vocans Guinot, 1985
Pycnoplax aspera Castro, 2009 *
Pycnoplax suruguensis (Rathbun, 1932)
Thyraplax cooki (Rathbun, 1906)
Thyraplax crosnieri (Guinot & Richer de Forges, 1981a) *
Thyraplax truncata Castro, 2007
HEXAPODIDAE
Hexaplax megalops Doein, 1904
HOMOLIDAE
Dagnaudus petterdi (Grant, 1905)
Homola coriolisi Guinot & Richer de Forges, 1995 *
Homola mieensis Sakai, 1979
Homola orientalis Henderson, 1888
Homola ranunculus Guinot & Richer de Forges, 1995 *
Homolochunia kullar Grin & Brown, 1976
Homolochunia valdiviae Doein, 1904
Homologenus levii Guinot & Richer de Forges, 1995 *
Homolomannia sibogae Ihle, 1912
Ihlopsis tirardi Guinot & Richer de Forges, 1995 *
Lamoha inata (Guinot & Richer de Forges, 1981b) *
Latreillopsis antennata Guinot & Richer de Forges, 1995 *
Latreillopsis bispinosa Henderson, 1888
Latreillopsis gracilipes Guinot & Richer de Forges, 1981b *
Moloha majora (Kubo, 1936)
Paromola bathyalis Guinot & Richer de Forges, 1995 *
Paromolopsis boasi Wood-Mason & Alcock, 1891
HOMOLODROMIIDAE
Dicranodromia foersteri Guinot, 1993 *
Dicranodromia spinulata Guinot, 1995 *
Homolodromia kai Guinot, 1993
HYPOTHALASSIIDAE
Hypothalassia armata (De Haan, 1835)
INACHIDAE
Cyrtomaia coriolisi Richer de Forges & Guinot, 1988 *
Cyrtomaia cornuta Richer de Forges & Guinot, 1988 *
Cyrtomaia ericina Guinot & Richer de Forges, 1982 *
Cyrtomaia furici Richer de Forges & Guinot, 1988 *
Cyrtomaia grini Richer de Forges & Guinot, 1990
Cyrtomaia horrida Rathbun, 1916
Cyrtomaia ihlei Guinot & Richer de Forges, 1982 *
Cyrtomaia platypes Yokoya, 1933
Grypachaeus hyalinus Alcock & Anderson, 1894
Platymaia mbriata Rathbun, 1916
Platymaia rebierei Guinot & Richer de Forges, 1986a
Platymaia wyvillethomsoni Miers, 1886
Pleistacantha cervicornis Ihle & Ihle-Landenberg, 1931
Pleistacantha exophthalmus Guinot & Richer de Forges, 1982 *
Pleistacantha cf japonica (Yokoya, 1933)
LATREILLIIDAE
Eplumula australiensis (Henderson, 1888)
Latreillia metanesa Williams, 1982
Latreillia pennifera Alcock, 1900
Latreillia valida De Haan, 1839
LEUCOSIIDAE
Alox ornatum (Ihle, 1918)
Ancylodactyla elata (Zarenkov, 1994)
Ancylodactyla elongata (Zarenkov, 1969)
Ancylodactyla nana (Zarenkov, 1990)
Arcania elongata Yokoya, 1933
Arcania tuberculata Bell, 1855
Galilia narusei Ng & Richer de Forges, 2007 *
Myra curtimana Galil, 2001a *
Myra eudactylus (Bell, 1855)
Myra fugax (Fabricius, 1798)
Tanaoa serenei (Richer de Forges, 1983) *
Toru septimus Galil, 2003 *
Urnalana granulimera Galil, 2005
MATHILDELLIDAE
Intesius pilosus Guinot & Richer de Forges, 1981a *
Intesius richeri Crosnier & Ng, 2004 *
Mathildella maxima Guinot & Richer de Forges, 1981a *
Platypilumnus jamiesoni Richer de Forges, 1996 *
OZIIDAE
Eupilumnus laciniatus (Sakai, 1980)
PALICIDAE
Miropalicus vietnamensis (Zarenkov, 1968)
Paliculus kyusyuensis (Yokoya, 1933)
Parapalicus ambonensis Moosa & Serène, 1981
Parapalicus clinodentatus Castro, 2000
Parapalicus denticulatus Castro, 2000 *
Parapalicus inanis Castro, 2000 *
Parapalicus piruensis Moosa & Serène, 1981
Parapalicus trispiralis Castro, 2000
Pseudopalicus acanthodactylus Castro, 2000 *
Pseudopalicus amadaibai (Sakai, 1963)
Pseudopalicus declivis Castro, 2000 *
Pseudopalicus glaber Castro, 2000 *
Pseudopalicus investigatoris (Alcock, 1900)
Pseudopalicus oahuensis (Rathbun, 1906)
Rectopalicus amphiceros Castro, 2000 *
Rectopalicus ampullatus Castro, 2000 *
Rectopalicus woodmasoni (Alcock, 1900)
PARTHENOPIDAE
Garthambrus poupini (Garth, 1993)
Pseudolambrus guinotae Tan, 2010 *
PHYLLOTYMOLINIDAE
Genkaia keijii Tavares, 1993 *
Phyllotymolinum crosnieri Tavares, 1993 *

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54
PILUMNIDAE
Eumedonus brevirhynchus Chia & Ng, 2000 *
PLAGUSIIDAE
Euchirograpsus timorensis Türkay, 1975
Miersiograpsus australiensis Türkay, 1978
POLYBIIDAE
Benthochascon hemingi Alcock & Anderson, 1899
Brusinia elongata (Sakai, 1969)
Brusinia profunda Moosa, 1996 *
Ovalipes iridescens (Miers, 1886)
Parathranites granosus Crosnier, 2002b
Parathranites intermedius Crosnier, 2002b *
Parathranites orientalis (Miers, 1886)
PORTUNIDAE
Charybdis (Charybdis) rufodactylus Stephenson & Rees, 1968
Cycloachelous orbitosinus (Rathburn, 1911)
Echinolatus caledonicus (Moosa, 1996) *
Liocarcinus vernalis (Risso, 1816)
Lissocarcinus polybiodes Adams & White, 1849
Lupocyclus philippinensis Semper, 1880
Lupocyclus tugelae Barnard, 1950
Nectocarcinus pubescens Moosa, 1996 *
Portunus (Monomia) gladiator Fabricius, 1798
Portunus (Monomia) lecromi Moosa, 1996 *
Portunus (Xiphonectes) hastatoides Fabricius, 1798
Portunus (Xiphonectes) stephensoni Moosa, 1981 *
Thalamita spinifera Borradaile, 1902
PROGERYONIDAE
Progeryon vaubani Guinot & Richer de Forges, 1981a *
Rhadinoplax microphthalmus (Guinot & Richer de Forges,
1981a) *
RANINIDAE
Cosmonotus mclaughlinae Tavares, 2006
Lyreidus brevifrons Sakai, 1937
Ranina ranina (Linnaeus, 1758)
RETROPLUMIDAE
Retropluma serenei Saint Laurent, 1989
Retropluma solomonensis McLay, 2006
TRAPEZIIDAE
Calocarcinus africanus Calman, 1909
Calocarcinus crosnieri Galil & Clark, 1990 *
Quadrella maculosa Alcock, 1898
TRICHOPELTARIIDAE
Trichopeltarion vanuatuensis Tavares & Cleva, 2010
XANTHIDAE
Alainodaeus alis Davie, 1997 *
Alainodaeus rimatara Davie, 1993
Antrocarcinus petrosus Ng & Chia, 1994 *
Demania cultripes (Alcock, 1898)
Demania garthi Guinot & Richer de Forges, 1981a *
Demania intermedia Guinot, 1969
Demania mortenseni (Odhner, 1925)
Demania wardi Garth & Ng, 1985
Euryxanthops latifrons Davie, 1997 *
Gaillardiellus bathus Davie, 1997 *
Glyptocarcinus politus Ng & Chia, 1994 *
Liomera nigrimanus Davie, 1997 *
Lophozozymus bertonciniae Guinot & Richer de Forges, 1981a *
Medaeops gemini Davie, 1997 *
Medaeops merodontos Davie, 1997 *
Medaeus aztec Davie, 1997 *
Meractaea multidentata Davie, 1997 *
Miersiella haswelli (Miers, 1886)
Palatigum trichostoma Davie, 1997 *
Paramedaeus globosus Serène & Vadon, 1981 *
Paraxanthodes cumatodes (MacGilchrist, 1905)
Rata chalcal Davie, 1997 *
Xanthias teres Davie, 1997 *

The MusorsToM-TDSB deep-sea crusTaceans 55
APPENDIX 3
New species of decapod crustaceans (recorded below 100 m depth) described from South Pacic island groups (other than New Caledonia) based on the
Tropical Deep-Sea Benthos expeditions material. For references, see Appendix 4.
PENAEOIDEA
PENAEIDAE
Parapenaeus cayrei Crosnier, 2005
..............................................................
Tonga
SOLENOCERIDAE
Maximiliaeus odoceros Chan, 2012
................................................................
PNG
STENOPODIDEA
SPONGICOLIDAE
Globospongicola spinulatus Komai & Saito, 2006
.................................
Vanuatu
CARIDEA
CRANGONIDAE
Metacrangon clevai Komai, 2012
............................................................
Solomon
Parapontophilus cornutus Komai, 2008
.....................................
French
Polynesia
Parapontophilus dicilis Komai, 2008
....................................................
Vanuatu
Parapontophilus longirostris Komai, 2008
.................................
French
Polynesia
Parapontophilus stenorhinus Komai, 2008
.................................................
Tonga
GLYPHOCRANGONIDAE
Glyphocrangon arduus Komai, 2007
...........................................
French
Polynesia
Glyphocrangon boa Komai, 2011
..............................................................
Vanuatu
Glyphocrangon musorstomia Komai, 2006
...................................
Wallis & Futuna
Glyphocrangon prostrata Komai, 2011
...................................................
Solomon
Glyphocrangon richeri Komai, 2004b
...........................................
Wallis & Futuna
Glyphocrangon rubricinctuta Komai, 2004b
...............................
Wallis & Futuna
Glyphocrangon similior Komai, 2004b
....................................................
Vanuatu
HYPPOLYTIDAE
Eumanninigia pliarthron Crosnier, 2000
......................................
Wallis & Futuna
NEMATOCARCINIDAE
Nemato carcinus machaerophorus Burukovsky, 2003
.............
French Polynesia
PALAEMONIDAE
Izucaris crosnieri Li, 2008
..............................................................
French Polynesia
Palaemonella komaii Li & Bruce, 2006
........................................................
Tonga
Periclimenes paralcocki Li & Bruce, 2006
.....................................
Wallis & Futuna
Periclimenes poupini Bruce, 1989
................................................
French Polynesia
PANDALIDAE
Heterocarpus amacula Crosnier, 1988b
.....................................
French Polynesia
Heterocarpus corona Yang, Chan & Chu, 2010
......................................
Vanuatu
Heterocarpus tenuidentatus Cleva & Crosnier, 2006
............................
Solomon
Plesionika bimaculata Chan, 2004
...........................................................
Vanuatu
Plesionika carsini Crosnier, 1986a
...............................................
French Polynesia
Plesionika curvata Chan & Crosnier, 1991
.................................
French Polynesia
Plesionika erythrocyclus Chan & Crosnier, 1997
.......................
French Polynesia
Plesionika fenneri Crosnier, 1986b
..............................................
French Polynesia
Plesionika avicauda Chan & Crosnier, 1991
............................
French Polynesia
Plesionika payeni Chan & Crosnier, 1997
...................................
French Polynesia
Plesionika picta Chan & Crosnier, 1997
......................................
French Polynesia
Plesionika poupini Chan & Crosnier, 1997
.................................
French Polynesia
Plesionika protati Chan & Crosnier, 1997
...................................
French Polynesia
Plesionika rubrior Chan & Crosnier, 1991
...................................
French Polynesia
STYLODACTYLIDAE
Stylodactylus brucei Cleva, 1994
...................................................
Wallis & Futuna
ASTACIDEA
ENOPLOMETOPIDAE
Enoplometopus gracilipes (de Saint Laurent, 1988)
.................
French Polynesia
AXIIDEA
AXIIDAE
Acanthaxius clevai Ngoc-Ho, 2006
.........................................................
Solomon
STRAHLAXIIDAE
Neaxius trondlei Ngoc-Ho, 2005
.................................................
French Polynesia
THOMASSINIIDAE
Crosniera dayrati Ngoc-Ho, 2005
................................................
French Polynesia
ACHELATA
PALINURIDAE
Nupalirus vericeli (Poupin, 1994)
.................................................
French Polynesia
Puerulus richeri Chan, Ma & Chu, this volume
..........................
French Polynesia
SCYLLARIDAE
Eduarctus marginatus Holthuis, 2002
..............................................................
Fiji
Galearctus rapanus (Holthuis, 1993)
..........................................
French Polynesia
Chelarctus crosnieri Holthuis, 2002
..............................................................
Tonga
POLYCHELIDA
POLYCHELIDAE
Stereomastis alis (Ahyong & Galil, 2006)
....................................
French Polynesia
ANOMURA
ALBUNEIDAE
Albunea marquisiana Boyko, 2000
......................................................... French Polynesia

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
56
DIOGENIDAE
Calcinus albengai Poupin & Lemaitre, 2003
..................................... French Polynesia
Calcinus anani Poupin & McLaughlin, 1998
.................................... French Polynesia
Calcinus gouti Poupin, 1997
....................................................................... French Polynesia
Ciliopagurus major Forest, 1995
.............................................................. French Polynesia
Ciliopagurus pacicus Forest, 1995
........................................................ French
Polynesia
Ciliopagurus plessisi Forest, 1995
............................................................. French
Polynesia
Strigopagurus poupini Forest, 1995
....................................................... French
Polynesia
EUMUNIDIDAE
Eumunida treguieri de Saint Laurent & Poupin, 1996
................. French
Polynesia
GALATHEIDAE
Galathea profunda Macpherson, 2012
................................................................. Vanuatu
Galathea raventosae Macpherson, 2012
............................................................. Vanuatu
Galathea sanctae Macpherson, 2012
.................................................................... Vanuatu
LITHODIDAE
Lithodes megacantha Macpherson, 1991
......................................... French
Polynesia
Paralomis arae Macpherson, 2001
..................................................................................... Fiji
Paralomis mendagnai Macpherson, 2003
........................................................ Solomon
MUNIDIDAE
Agononida aequabilis Macpherson, 2006a
....................................... French
Polynesia
Agononida alisae Macpherson, 1999
.................................................................... Vanuatu
Agononida garciai Macpherson, 2004
............................................................................. Fiji
Agononida imitata Macpherson, 2006a
............................................. French
Polynesia
Agononida isabelensis Cabezas, Macpherson & Machordom, 2009
..... Solomon
Agononida simillima Macpherson, 2006a
......................................... French
Polynesia
Anoplonida patae MacPherson & Baba, 2006
...................................................... Tonga
Babamunida corniculata Macpherson, this volume
................... French
Polynesia
Babamunida hystrix (Macpherson & de Saint Laurent, 1991)
.... French Polynesia
Babamunida plexaura (Macpherson & de Saint Laurent, 1991)
... French Polynesia
Bathymunida avatea MacPherson & Baba, 2006
.......................... French
Polynesia
Bathymunida dissimilis Baba & de Saint Laurent, 1996
................ Wallis & Futuna
Bathymunida recta Baba & de Saint Laurent, 1996
......................... Wallis & Futuna
Crosnierita tucanae Macpherson, 2004
........................................................................... Fiji
Crosnierita yante (Macpherson, 1994)
................................................. French
Polynesia
Heteronida clivicola MacPherson & Baba, 2006
.............................. French
Polynesia
Munida abelloi Macpherson, 1994
........................................................... Wallis & Futuna
Munida amathea Macpherson & de Saint Laurent, 1991
........ French Polynesia
Munida angusta Macpherson, 2004
.......................................................................... Tonga
Munida antliae Macpherson, 2006a
..................................................... French
Polynesia
Munida apheles Macpherson, 2006a
.................................................... French
Polynesia
Munida apodis Macpherson, 2004
............................................................................. Tonga
Munida arae Macpherson, 2006a
........................................................... French
Polynesia
Munida atarapa Macpherson, this volume
...................................... French
Polynesia
Munida caeli Cabezas, Macpherson & Machordom, 2009
..................... Solomon
Munida clevai Macpherson, 2009
............................................................................ Vanuatu
Munida columbae Macpherson, 2006a
.............................................. French
Polynesia
Munida congesta Macpherson, 1999
.................................................................... Vanuatu
Munida cornuta Macpherson, 1994
.................................................................................. Fiji
Munida descensa Macpherson, 2006a
................................................. French
Polynesia
Munida erugata Macpherson, 2006a
................................................... French
Polynesia
Munida fasciata Macpherson, 2006a
................................................... French
Polynesia
Munida fornacis Macpherson, 2006a
................................................... French
Polynesia
Munida galaxaura Macpherson, 1996b
................................................. Wallis & Futuna
Munida glabella Macpherson, 2000
..................................................... French Polynesia
Munida ignea Macpherson, 2006a
........................................................ French
Polynesia
Munida lailai Cabezas, Macpherson & Machordom, 2009
.................... Solomon

Munida lenticularis Macpherson & de Saint Laurent, 1991
..... French
Polynesia
Munida limatula Macpherson, 2004
.......................................................................... Tonga
Munida llenasi Macpherson, 2006a
....................................................... French
Polynesia
Munida longicheles Macpherson & de Saint Laurent, 1991
.... French
Polynesia
Munida masoae Macpherson, 1996b
..................................................... Wallis & Futuna
Munida mendagnai Cabezas, Macpherson & Machordom, 2009
..... Solomon
Munida micula Macpherson, 1996b
........................................................ Wallis & Futuna
Munida miniata Macpherson, 1996b
...................................................... Wallis
& Futuna
Munida oblonga Macpherson, 2006a
.................................................. French Polynesia
Munida oblongata Cabezas, Macpherson & Machordom, 2009
........ Solomon
Munida ocellata Macpherson & de Saint Laurent, 1991
........... French
Polynesia
Munida oella Macpherson, 1996b
......................................................... Wallis
& Futuna
Munida pasithea Macpherson & de Saint Laurent, 1991
......... French Polynesia
Munida pavonis Macpherson, 2004
........................................................................... Tonga
Munida polynoe Macpherson & de Saint Laurent, 1991
........... French
Polynesia
Munida profunda Macpherson & de Saint Laurent, 1991
........ French
Polynesia
Munida pumila Macpherson, 2004
............................................................................ Tonga
Munida rona Macpherson, this volume
............................................. French
Polynesia
Munida rubella Macpherson & de Saint Laurent, 1991
............. French
Polynesia
Munida rubrovata Macpherson & de Saint Laurent, 1991
...... French
Polynesia
Munida spissa Macpherson, 1996b
.......................................................... Wallis & Futuna
Munida squarrosa Macpherson, 2009
.................................................................. Vanuatu
Munida volantis Macpherson, 2004
.................................................................................. Fiji
Neonida grandis Baba & de Saint Laurent, 1996
............................................. Vanuatu
Onconida gemini Baba & de Saint Laurent, 1996
........................................... Vanuatu
Onconida modica Baba & de Saint Laurent, 1996
........................... Wallis & Futuna
Onconida prostrata Baba & de Saint Laurent, 1996
........................ Wallis & Futuna
Paramunida achernar Cabezas, Macpherson & Machordom, 2010
....... Tonga
Paramunida amphitrita Macpherson, 1996b
..................................... Wallis & Futuna
Paramunida ascella Cabezas, Macpherson & Machordom, 2010
........ Vanuatu
Paramunida cretata Macpherson, 1996b
............................................. Wallis & Futuna
Paramunida cristata Macpherson, 2004
......................................................................... Fiji
Paramunida curvata Macpherson, 2004
........................................................................ Fiji
Paramunida echinata Macpherson, 2000
.......................................... French Polynesia
Paramunida labis Macpherson, 1996b
................................................... Wallis & Futuna
Paramunida lophia Cabezas, Macpherson & Machordom, 2009
....... Solomon
Paramunida luminata Macpherson, 1996b
......................................... Wallis & Futuna
Paramunida poorei Cabezas, Macpherson & Machordom, 2010
... French Polynesia
Paramunida salai Cabezas, Macpherson & Machordom, 2009
.......... Solomon
Paramunida spatula Macpherson, 2006a
.......................................... French Polynesia
Paramunida spica Cabezas, Macpherson & Machordom, 2010
........... Vanuatu
Plesionida concava Cabezas, Macpherson & Machordom, 2009
....... Solomon
Raymunida cagnetei Macpherson & Machordom, 2000
.......... French Polynesia
Raymunida erythrina Macpherson & Machordom, 2001
............ Wallis & Futuna
Raymunida limbata Macpherson, 2006a
........................................... French Polynesia
Setanida cristata Macpherson, 2006a
.................................................. French
Polynesia
MUNIDOPSIDAE
Galacantha quiquei Macpherson, 2007
................................................. Wallis & Futuna
Munidopsis austellus Macpherson, 2007
............................................ French
Polynesia
Munidopsis bruta Macpherson, 2007
.................................................................. Solomon

The MusorsToM-TDSB deep-sea crusTaceans 57
Munidopsis concava Macpherson, 2007
........................................................................ Fiji
Munidopsis demeter Macpherson, 2007
............................................................ Solomon
Munidopsis denudata Macpherson, 2007
........................................................ Solomon
Munidopsis keijii Macpherson, 2007
..................................................................... Solomon
Munidopsis pericalla Macpherson, 2007
........................................................... Solomon
Munidopsis strigula Macpherson, 2007
.............................................................. Solomon
Munidopsis zarazagai Macpherson, 2007
.......................................................... Vanuatu
PAGURIDAE
Lithopagurus boucheti McLaughlin & Lemaitre, 2004
............................................ Fiji
Micropagurus bijdeleyi Lemaitre, 2010
................................................. French Polynesia
Pagurojacquesia polymorpha (de Saint Laurent & McLaughlin, 1999) ....
Vanuatu
Solitariopagurus triprobolus Poupin & McLaughlin, 1996
........ French Polynesia
Turleania boucheti McLaughlin, 2007b
.................................................... Wallis&Futuna
PARAPAGURIDAE
Oncopagurus oimos Lemaitre, 1998
..................................................... French Polynesia
Oncopagurus tuamotu Lemaitre, 1994
............................................... French Polynesia
Paragiopagurus bougainvillei (Lemaitre, 1994)
............................... French Polynesia
Paragiopagurus fasciatus Lemaitre & Poupin, 2003
..................... French Polynesia
Paragiopagurus trilineatus Lemaitre, this volume
.............................................. Tonga
Paragiopagurus wallisi (Lemaitre, 1994)
............................................. French
Polynesia
Sympagurus poupini Lemaitre, 1994
..................................................... French
Polynesia
PYLOCHELIDAE
Bathycheles phenax McLaughlin & Lemaitre, 2009
.................................................. Fiji
Trizocheles mendanai McLaughlin & Lemaitre, 2009
................................. Solomon
BRACHYURA
CALAPPIDAE
Calappa sebastieni Galil, 1997
................................................................... French
Polynesia
Mursia diwata Galil & Takeda, 2004
........................................................................ Vanuatu
Mursia poupini Galil, 2001
........................................................................... French
Polynesia
CYCLODORIPPIDAE
Ketamia limatula Tavares, 1993
................................................................................. Vanuatu
Tymolus daviei Tavares, 1997
...................................................................................... Vanuatu
DYNOMENIDAE
Metadynomene tuamotu Ng & McLay, 2010
.................................... French
Polynesia
DROMIIDAE
Cryptodromia erioxylon McLay, 2001
.................................................... French
Polynesia
Cryptodromia marquesas McLay, 2001
................................................ French
Polynesia
Sphaerodromia ducoussoi McLay, 1991
.............................................. French
Polynesia
EPIALTIDAE
Garthinia disica Richer de Forges & Ng, 2009a
.............................................. Solomon
Guinotinia lehouarnoi Richer de Forges & Ng, 2009a
............................................. Fiji
Naxioides teatui Poupin, 1995
................................................................... French
Polynesia
Naxioides vaitahu Poupin, 1995
.............................................................. French
Polynesia
Oxypleurodon barazeri Richer de Forges & Ng, 2009b
.............................. Solomon
Oxypleurodon christiani Richer de Forges & Corbari, 2012
............................... PNG
Oxypleurodon parallelum Richer de Forges & Ng, 2009b
........................ Solomon
Oxypleurodon tavaresi Richer de Forges, 1995
.................................. Wallis & Futuna
Rochinia boucheti Richer de Forges & Ng, this volume
........................... Solomon
ETHUSIDAE
Ethusa abbreviata Castro, 2005a
.............................................................................. Vanuatu
Ethusa barbata Castro, 2005a
.................................................................................. Solomon
Ethusa crassipodia Castro, 2005a
........................................................................... Solomon
Ethusina ciliacirrata Castro, 2005a
........................................................................... Vanuatu
Ethusina coronata Castro, 2005a
............................................................................ Solomon
Ethusina crenulata Castro, 2005a
............................................................ French
Polynesia
Ethusina exophthalma Castro, 2005a
............................................................................... Fiji
Ethusina huilianae Castro, 2005a
............................................................ French
Polynesia
Ethusina microspina Chen, 2000
.............................................................................. Vanuatu
Ethusina ocellata Castro, 2005a
.............................................................................. Solomon
Ethusina stenommata Castro, 2005a
....................................................... Wallis & Futuna
Ethusina vanuatuensis Chen, 2000
.......................................................................... Vanuatu
Parethusa hylophora Castro, 2005a
...................................................................... Solomon
Serpenthusa brucei Naruse, Castro & Ng, 2009
................................................ Vanuatu
GERYONIDAE
Chaceon australis Manning, 1993
.......................................................... French
Polynesia
GONEPLACIDAE
Carcinoplax tenuidentata Castro, 2007
............................................................................ Fiji
Carcinoplax velutina Castro, 2007
........................................................... French
Polynesia
Guinoplax apheles Castro & Ng, 2010
............................................................................... Fiji
Microgoneplax cope Castro, 2007
.............................................................. Wallis & Futuna
Thyraplax cristata Castro, 2007
................................................................. French
Polynesia

Thyraplax digitodentata Castro, 2007
................................................... French
Polynesia

Thyraplax truncata Castro, 2007
.......................................................................................... Fiji
HOMOLIDAE
Homola poupini Richer de Forges & Ng, 2007
................................ French
Polynesia
Homolochunia menezi Richer de Forges & Ng, 2008b
............................. Solomon
Homologenus broussei Guinot & Richer de Forges, 1981b
...... French
Polynesia
Homologenus wallis Guinot & Richer de Forges, 1995
................. Wallis
&
Futuna
Lamoha longirostris (Chen, 1986)
.............................................................. Wallis
&
Futuna
Lamoha personata (Guinot & Richer de Forges, 1981b)
............................ Vanuatu
Yaldwynopsis guinotae Richer de Forges & Ng, 2007
................. French
Polynesia
INACHIDAE
Cyrtomaia polynesica Richer de Forges & Ng, 2008a
.................. French
Polynesia
Platymaia rebierei Guinot & Richer de Forges, 1986a
.................................. Vanuatu
Pleisticanthoides piccardorum Ng & Richer de Forges, 2012
........................... PNG
LEUCOSIIDAE
Euclosiana guinotae Galil & Ng, 2010
.................................................................... Vanuatu
Euclosiana vella (Galil, 2007)
..................................................................................... Solomon
Tanaoa nanus Galil, 2003
.............................................................................................. Vanuatu
Tokoyo cirrata Galil, 2003
.............................................................................................. Vanuatu
Urnalana granulimera Galil, 2005
....................................................................................... Fiji
MATHILDELLIDAE
Beuroisia manquenei Guinot & Richer de Forges, 1981a
.......... French
Polynesia
Intesius crosnieri Davie, 1998
..................................................................... French
Polynesia
PALICIDAE
Neopalicus simulus Castro, 2010
.............................................................. French
Polynesia
Parapalicus armatus Castro, 2000
............................................................................ Vanuatu
Parapalicus clinodentatus Castro, 2000
................................................................ Vanuatu
Parapalicus inermis Castro, 2000
......................................................................................... Fiji

BerTrand richer de Forges, Tin-YaM chan, Laure corBari, raFaeL LeMaiTre, enrique Macpherson, shane T. ahYong & peTer K. L. ng
58
Pseudopalicus pictus Castro, 2000
........................................................................... Vanuatu
PLAGUSIIDAE
Euchirograpsus tuerkayi Crosnier, 2001
............................................... French
Polynesia
POLYBIIDAE
Parathranites parahexagonus Crosnier, 2002b
................................ French
Polynesia
Parathranites tuberosus Crosnier, 2002b
............................................. French
Polynesia
PORTUNIDAE
Echinolatus proximus Davie & Crosnier, 2006
.................................. French
Polynesia
Laleonectes stridens Crosnier & Moosa, 2002
................................... French
Polynesia
Thalamita dicilis Crosnier, 2002a
......................................................... French
Polynesia
PROGERYONIDAE
Progeryon mararae Guinot & Richer de Forges, 1981a
.............. French
Polynesia
PSEUDOZIIDAE
Euryozius danielae Davie, 1993
................................................................. French
Polynesia
RETROPLUMIDAE
Retropluma solomonensis McLay, 2006
.............................................................. Solomon
TRAPEZIIDAE
Hexagonalia unidentata Castro, 2005b
............................................................... Salomon
TRICHOPELTARIIDAE
Trichopeltarion dejouanneti Tavares & Cleva, 2010
...................................... Solomon
Trichopeltarion parvum Tavares & Cleva, 2010
............................................... Solomon
Trichopeltarion vanuatuensis Tavares & Cleva, 2010
..................................... Vanuatu
Sphaeropeltarion edentatum Tavares & Cleva, 2010
....................... Wallis & Futuna
XANTHIDAE
Alainodaeus akiaki Davie, 1993
................................................................ French
Polynesia
Alainodaeus nuku Davie, 1997
.................................................................. French
Polynesia
Alainodaeus rimatara Davie, 1993
.......................................................... French
Polynesia
Banareia fatuhiva Davie, 1993
.................................................................. French
Polynesia
Epistocavea mururoa Davie, 1993
........................................................... French
Polynesia
Lipkemera rufomaculata (Davie, 1993)
................................................ French
Polynesia
Medaeus grandis Davie, 1993
.................................................................... French
Polynesia
Meractaea tafai Davie, 1993
....................................................................... French
Polynesia
Paramedaeus megagomphios Davie, 1997
.......................................... Wallis
&
Futuna
Paraxanthodes polynesiensis Davie, 1993
........................................... French
Polynesia
Rata tuamotense Davie, 1993
................................................................... French
Polynesia

The MusorsToM-TDSB deep-sea crusTaceans 59
APPENDIX 4
References to taxonomical publications containing original descriptions (listed in Appendix 2 & 3) of decapod crustaceans collected by the Tropical Deep-
Sea Benthos programme and satellite programmes.
AHYONG S. T. 2010 — The marine fauna of New Zealand: King crabs of New Zealand, Australia and the Ross Sea (Crustacea: Decapoda: Lithodidae) NIWA
Biodiversity Memoir 123, Wellington, 196 p.
AHYONG S. T. & GALIL B. S. 2006 — Polychelidae from the southern and western Pacic (Decapoda, Polychelida). Zoosystema 28 (3): 757-767.
ASAKURA A. 2005 — A review of the genus Micropagurus (Crustacea Decapoda Anomura Paguridae). Zootaxa 1090: 1-34.
BABA K. 1991a — Crustacea Decapoda: Chirostylus Ortmann, 1892, and G astroptychus Caullery, 1896 (Chirostylidae) from New Caledonia, in CROSNIER A.
(ed.), Résultats des Campagnes MUSORSTOM, vol. 9. Mémoires du Muséum national d’Histoire naturelle 152: 463-477.
BABA K. 1991b — Crustacea Decapoda: Alainius gen. nov., Leiogalathea Baba, 1969, and Phylladiorhynchus Baba, 1969 (Galatheidae) from New Caledonia, in
CROSNIER A. (ed.), Résultats des Campagnes MUSORSTOM, vol. 9. Mémoires du Muséum national d’Histoire naturelle152: 479-491.
BABA K. 2004 — Uroptychodes, new genus of Chirostylidae (Crustacea: Decapoda: Anomura), with description of three new species. Scientia Marina 68: 97-116.
BABA K. & SAINT LAURENT DE M. 1996 — Crustacea Decapoda: Revision of the genus Bathymunida Balss, 1914, and description of six new related genera
(Galatheidae), in CROSNIER A. (ed.), Résultats des Campagnes MUSORSTOM, vol. 15. Mémoires du Muséum national d’Histoire naturelle 168: 433-502.
BOYKO C. B. & HARVEY A. W. 1999 — Crustacea Decapoda: Albuneidae and Hippidae of the tropical Indo-West Pacic region, in CROSNIER A. (ed.), Résultats
des Campagnes MUSORSTOM, vol. 20. Mémoires du Muséum national d’Histoire naturelle 180: 379-406.
BOYKO C. B. 2000 — The Hippoidea (Decapoda, Anomura) of the Marquises Islands, with description of a new species of Albunea. Zoosytema 22: 107-116.
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