An annotated list of the Lepidoptera of Honduras

Abstract

A biodiversity inventory of the Lepidoptera of Pico Bonito National Park and vicinity, in the Department of Atlantida of northern Honduras, was initiated in 2009 to obtain baseline data. We present a revised checklist of Honduran butterfly species (updated from the initial 1967 lists), as well as the first comprehensive list of Honduran moths. Our updated list includes 550 species of Papilionoidea, 311 Hesperioidea, and 1,441 moth species.

Full text

Center for SyStematiC entomology, inC., Gainesville, FL
Insecta
MundI A Journal of World Insect Systematics
0205
An annotated list of the Lepidoptera of Honduras
Jacqueline Y. Miller, Deborah L. Matthews, Andrew D. Warren
McGuire Center for Lepidoptera and Biodiversity
Florida Museum of Natural History
University of Florida
P. O. Box 112710, Gainesville, Florida 32611-2710, USA
jmiller@mnh.u.edu, dlott@mnh.u.edu, andy@butteriesofamerica.com
M. Alma Solis
Systematic Entomology Laboratory, PSI, ARS, USDA
c/o National Museum of Natural History, Washington, D.C., 20560-0168, USA
Alma.Solis@ARS.USDA.GOV
Donald J. Harvey, Patricia Gentili-Poole
Department of Entomology
Smithsonian Institution, Washington, D.C., 20560-0127, USA
HARVEYD@si.edu, GENTILIP@si.edu
Robert Lehman
Museo de Mariposas y Insectos
Box 720, La Ceiba, Honduras
robertdlt42@gmail.com
Thomas C. Emmel and Charles V. Covell, Jr.
McGuire Center for Lepidoptera and Biodiversity
Florida Museum of Natural History
University of Florida
P. O. Box 112710, Gainesville, Florida 32611-2710, USA
tcemmel@mnh.u.edu, ccovell@mnh.u.edu
Date of Issue: February 29, 2012

Jacqueline Y. Miller, Deborah L. Matthews, Andrew D. Warren, M. Alma Solis, Donald
J. Harvey, Patricia Gentili-Poole, Robert Lehman, Thomas C. Emmel and Charles V.
Covell, Jr.
An annotated list of the Lepidoptera of Honduras
Insecta Mundi 0205: 1-72
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An annotated list of the Lepidoptera of Honduras
Jacqueline Y. Miller, Deborah L. Matthews, Andrew D. Warren
McGuire Center for Lepidoptera and Biodiversity
Florida Museum of Natural History
University of Florida
P. O. Box 112710, Gainesville, Florida 32611-2710, USA
jmiller@mnh.u.edu, dlott@mnh.u.edu, andy@butteriesofamerica.com
M. Alma Solis
Systematic Entomology Laboratory, PSI, ARS, USDA
c/o National Museum of Natural History, Washington, D.C., 20560-0168, USA
Alma.Solis@ARS.USDA.GOV
Donald J. Harvey, Patricia Gentili-Poole
Department of Entomology
Smithsonian Institution, Washington, D.C., 20560-0127, USA
HARVEYD@si.edu, GENTILIP@si.edu
Robert Lehman
Museo de Mariposas y Insectos
Box 720, La Ceiba, Honduras
robertdlt42@gmail.com
Thomas C. Emmel and Charles V. Covell, Jr.
McGuire Center for Lepidoptera and Biodiversity
Florida Museum of Natural History
University of Florida
P. O. Box 112710, Gainesville, Florida 32611-2710, USA
tcemmel@mnh.u.edu, ccovell@mnh.u.edu
Abstract. A biodiversity inventory of the Lepidoptera of Pico Bonito National Park and vicinity, in the Department
of Atlantida of northern Honduras, was initiated in 2009 to obtain baseline data. We present a revised checklist
of Honduran buttery species (updated from the initial 1967 lists), as well as the rst comprehensive list of
Honduran moths. Our updated list includes 550 species of Papilionoidea, 311 Hesperioidea, and 1,441 moth
species.
Keywords. Mesoamerica, Neotropical, Central America, biodiversity, faunal composition, Lepidoptera, Honduras.
Introduction
A comprehensive inventory of the Lepidoptera of Honduras has been initiated by the McGuire
Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, in collaboration with
the Centro Zamorano de Biodiversidad, Escuela Agrícola Panamericana–Zamorano, The Lodge at Pico
Bonito, and the Centro Universitario Regional del Litoral Atlantico. Together with the support of
taxonomic specialists at various institutions, students, research associates, and ecotourism citizen-
scientist participants, we seek to expand the knowledge and appreciation of the faunal diversity and
natural history of Honduras. In addition, our objectives are to provide research and reference material
for current and future studies on the phylogeny, systematics, and zoogeography of Lepidoptera. To
date, our survey efforts have concentrated on the northern coast, while at the same time assembling
data from literature records and museum specimens from areas throughout Honduras. The present
1
0205: 1-72 2012

2 • Insecta MundI 0205, February 2012 Miller et al.
publication summarizes the results of the rst two years of our surveys (2009-2010) in the form of an
annotated list, and gives an introduction to the biodiversity, biogeography, and climate of the region.
In addition, we provide a brief description of the principal study area and a general review of the
history of previous biodiversity studies on Lepidoptera of Mesoamerica.
Biodiversity, Historical Geology, and Biogeography
Mesoamerica is a biodiversity hotspot that includes all tropical and subtropical ecosystems from
eastern Panama north and west through Costa Rica, Nicaragua, Honduras, El Salvador, Belize,
and Guatemala, in addition to portions of southern and coastal Mexico. Approximately 40% of the
Mesoamerican forests are theoretically still present today but they are often highly fragmented.
Although the Mesoamerican corridor does not have the breadth or the high elevations of the Andes,
there are numerous metamorphic mountain chains throughout the region with peaks reaching 4000
m, in addition to a series of active volcanoes (Mittermeier et al. 1999). The intervening lowlands and
valleys throughout these cordilleras have acted as corridors for animal and human migrations and
dispersal over millions of years. Historical geological evidence suggests a rather wide separation of
the two continents from the Cretaceous through the Oligocene and into the late Miocene (Raven and
Axelrod 1974; Gose 1985; Pindell and Dewey 1982; Smith 1985; Smith and Bredin 1977; Miller and
Miller 2001). It has been suggested that once the contiguous land connection was complete during the
Pliocene (3 mybp) (Donnelly 1992; Iturralde-Vinent and MacPhee 1999) to 5.7 mybp (Webb 1978, 1985,
1991, 2006), an extensive faunal exchange began. The species that inhabit these areas today are relicts
of the fauna and ora from both North and South America. Thus, the Mesoamerican biodiversity
hotspot exhibits a unique geological and evolutionary history with a patchwork of habitats supporting
some of the highest levels of biodiversity and endemism on earth.
Climate and Habitats
The Caribbean lowlands of Honduras are among the wettest sections of Central America, with
2,500-6,500 mm of annual rainfall (occurring throughout the year), consistently warm temperatures,
and high humidity. Alternatively, the Pacic lowlands and interior of Honduras have pronounced wet
and dry seasons, with rain shadow effects associated with various mountain ranges. The rainy season
commences in May or June and extends to November, followed by a markedly severe dry season. In
both the Caribbean and Pacic lowlands, the average daytime temperatures are 28-33°C. The average
temperature in the interior Serranía habitats (see Carr 1950) decreases with elevation and reaches
highs of 30°C during the warmest months with 25°C during the coolest month. In montane areas,
daytime temperatures range from 16-25°C with frost occurring occasionally above 2,000 m. At higher
elevations, dense pine-oak forests predominate.
Ratcliffe and Cave (2006a) reviewed the different forest types of Mesoamerica, detailing more
than 20 life zones found in El Salvador, Honduras and Nicaragua alone. In Honduras, there are six
general forest types based on the Holdridge System (Holdridge 1967) that vary according to climate,
associated vegetation and elevation. These include the montane broadleaf forest, montane pine-oak
forest, tropical dry broadleaf forest, lowland broadleaf rain forest, lowland pine forest, and mangrove
forest. These forest types vary with elevation and many unique microhabitats occur throughout
Honduras. The forests in the Pico Bonito area are fragmentary with a mixture of primary and secondary
rainforests including montane broadleaf, lowland broadleaf and pine forests. Montane broadleaf forest
is characteristic of the intermediate and higher elevations of Pico Bonito National Park, with canopy
trees at 25-30 m and an upper layer comprised of Quercus that may reach 50 m. Buttressed trees are
common with tree ferns (Cyatheaceae) common in the understory. Palms are uncommon but species
of Ericaceae and Melastomataceae are abundant as well as a number of epiphytes (gesneriads, aroids,
bromeliads, orchids) and mosses covering rocks and tree trunks or limbs. Lower elevations in the park
are dominated by lowland broadleaf forest, with a notable transition to palms and hardwoods with
Magnoliaceae more frequently encountered.

Insecta MundI 0205, February 2012 • 3Lepidoptera of Honduras
Parque Nacional Pico Bonito and La Ceiba Vicinity
Our primary eld site thus far is Parque Nacional Pico Bonito, with boundaries extending within
both Atlantida and Yoro departments. It is part of the Sierra Nombre de Dios, which runs parallel to
the northern (Caribbean) coast. Located to the west of La Ceiba, this is the second largest national
park in Honduras, comprised of 1,314 km2 and named for its third highest peak (Pico Bonito, 2436
m). However, there are two higher peaks in the park within the largely unexplored core area of 500
km2, currently off limits for hiking. The rugged terrain is rocky with dense forests, rivers, waterfalls
and abundant wildlife, including various mammals, a diverse herpetofauna and a very extensive
avifauna (Perlo 2006). At lower elevations, the vegetation is tropical: mainly a mixture of sub-montane
broadleaf evergreen forest of low to intermediate height with the canopy trees reaching 25-30 m.
In some lowland areas, the landscape is dominated by agriculture with pineapple and oil palm as
principal crops. However, there are areas with lush vegetation at lower elevations of the mountains
that have been preserved.
History of Biodiversity Studies on Lepidoptera in Mesoamerica
Although Linnaeus, Walker, Butler, Schaus, Druce, and others described Lepidoptera from Central
America in their respective works, it is the extraordinary work of Godman and Salvin (1877, 1878a,b,c,
1879-1901, 1889) which forms the basis for the Lepidoptera known at the time with descriptions of new
species and superb illustrations. In their introduction, Godman (1901) summarized the various areas
collected in each country and documented the primary collecting localities and/or those associated with
each major collection, as in the case of Schaus. They noted the topography and major mountain ranges
in each country and provided a general idea of the height of major summits, gave information on the
location of the principal forests, and also recorded locations and conditions of the plains and drier
areas. It is rather interesting that their description of Honduras is very limited, mentioning that the
“general aspect of the country is varied by mountains and hills radiating from the common base of the
Cordilleras…From the Pacic, it presents the appearance of a great natural wall, with a lower range
of mountains bristling with volcanic peaks.” They state further that “the greater part of the country is
well watered and extensive forests are said to exist in the interior, which, however, is but imperfectly
known.”
Despite the expanse of time that has passed since Godman and Salvin, our voucher-based
knowledge of the Lepidoptera fauna of Honduras remains incomplete and fragmentary at best.
Within the last forty years, great advances have been made in recording distributions of Lepidoptera
in other Mesoamerican countries surrounding Honduras. Austin et al. (1996) produced a checklist
of butteries of the Tikal National Park area, Guatemala, and Austin et al. (1998) and Barrios et
al. (2006) inventoried the Hesperiidae of Guatemala. Salinas–Gutiérrez et al. (2009) published a
historical synopsis of Lepidoptera collecting in Guatemala and summarized the current knowledge
of species records and analyses of the Papilionoidea based on collection records, literature, eldwork,
and databases. Other recent studies have provided extensive information on the Lepidoptera fauna of
southern Mexico and the Yucatan Peninsula (Pozo et al. 2003, 2008).
For Belize, Barnes (2002) has provided numerous moth records and images, and Meerman and
Boomsma (1993) produced an initial checklist of butteries. The latter was subsequently expanded by
Meerman (1999). For the past 15 years, John Shuey and colleagues have been involved in studies on
the biodiversity and conservation of the butteries of the Rio Bravo Conservation and Management
Area and other regions in Belize, resulting in many new buttery records for the country. Shuey et
al. (2005) reported 85 new national records for butteries. Distributional information for Lepidoptera
in El Salvador is limited, with an annotated list of the Hesperiidae of the country (Steinhauser 1975)
being the only comprehensive faunal survey. Jean-Michel Maes (Museo Entomologico, Leon ) and
colleagues have been completing various inventories of insects in Nicaragua over the last 15 years
with recent publications on the Papilionidae (Maes 2006), Pieridae (Maes 2007), and Hesperiidae
(Anderson 2007).
Many studies have been completed on the Lepidoptera of Costa Rica. Based on eldwork throughout
Costa Rica, DeVries (1983, 1987, 1997) produced two excellent illustrated volumes on butteries,

4 • Insecta MundI 0205, February 2012 Miller et al.
including detailed information on the distribution, life history, ecology, behavior and diversity of
Costa Rican Papilionidae, Pieridae, Nymphalidae and especially Riodinidae (DeVries et al. 1994).
More recently, Chacon and Montero (2007) published an introduction to the diversity of the Costa
Rican Lepidoptera fauna, with descriptions of key characters for the identication of families, both
immatures and adults. Janzen and Hallwachs (2010) have continued to improve our knowledge of Costa
Rican Lepidoptera through mass rearings of many species in the Area de Conservación Guanacaste,
including biological notes and illustrations of larval foodplants, immatures and adults. Janzen,
Hallwachs and collaborators have combined molecular techniques with traditional morphological and
ecological studies to identify a number of potentially new taxa (e.g., Burns and Janzen 2001, 2005a,b;
Hebert et al. 2004; Burns et al. 2007, 2008; Janzen et al. 2009).
Although the diversity of plants in Honduras is relatively well known (Molina 1975; Sutherland
1986, 1996; Davidse et al. 1994, 1995, 2009), the Lepidoptera fauna of Honduras is the least known
of any Mesoamerican country. Monroe et al. (1967) published the rst list of the Papilionoidea of
Honduras, followed by a list of Hesperiidae by Monroe and Miller (1967). Fieldwork for these studies
was restricted to areas near San Pedro Sula, from Tela along the northern coast, southeast to areas
north of Tegucigalpa, with two collecting sites in the Department of Choluteca. Subsequent to these
publications, there are scattered records from Honduras in revisionary studies completed by Miller
(1972, 1974, 1976), Jenkins (1983, 1984, 1985a,b, 1986, 1989, 1990), Hall (1999), Willmott (2003) and
many others. Hall noted Honduran records for ve taxa of Theope Doubleday (Riodinidae). Willmott
recorded 20 species of Adelpha Hübner (Nymphalidae) from Honduras in museum collections throughout
the world; based on information derived from countries surrounding Honduras, he expected a total
of 32 Adelpha species for the country. Warren et al. (2011) list and gure essentially all described
Mesoamerican buttery taxa, including all taxa known from Honduras, and provide images of live and
pinned specimens from Honduras and surrounding countries.
Recent publications on Honduran moths have involved agricultural pests such as Spodoptera
Guenée (Escribano et al. 2000a,b) and other members of the Noctuoidea (Gregory 1987; Sponagel and
Díaz 1994; Cave 1992; Caballero et al. 1994; Ovalle and Cave 1989). Passoa worked in Honduras for
a few years and produced several papers describing the hosts and immature stages for various moth
taxa (Eichlin and Passoa 1983; Passoa 1982, 1983, 1985, 1991; Passoa and Habeck 1987; Passoa and
Steinly 1992). Studies by Solis on the mahogany webworm (Howard and Solis 1989) and a pest of
avocados (Solis and Styer 2003) have included data on immature stages of economically important
Honduran Pyraloidea. Recent revisionary studies and catalogues on Pterophoridae (Gielis 2003, 2006),
Pyraloidea (Munroe et al. 1995), Tortricidae (Brown 1996; Brown et al. 2005; Heppner 1995, 1996;
Razowski 1986; Razowski and Becker 2007), Limacodidae (Epstein and Corrales 2004), Notodontidae
(Thiaucourt 2002, 2007, 2008; J. S. Miller 2009a,b), and Arctiinae (de Toulgoët 1996, 1997) suggest
that there should be additional species of those families represented in Honduras. Lehman (1971)
published an initial list of the Sphingidae known from Honduras. Samayoa and Cave (2008) published
an updated list for this family in Honduras, documenting 107 species. This publication increased the
Honduran representation within the family by 33%, with new records represented in three subfamilies:
Macroglossinae (80), Sphinginae (22), and Smerinthinae (5).
In 1996, Dr. Ronald Cave, then of Escuela Agricultura Panamerica en Zamorano (EAPZ), initiated
a major project to survey all insects in the protected forests of the Pico Bonito National Park and
in other areas of the Sierra Nombre de Dios. This collection has been completely databased and is
deposited in the Collection of Arthropods at EAPZ. Cave subsequently collaborated with Ratcliffe on
revisionary studies of Mesoamerican Scarabaeidae (Ratcliffe 2000, 2001, 2003; Ratcliffe and Cave 2002,
2006a,b, 2008). Although major progress has been made, a comprehensive voucher-based inventory
of the Lepidoptera within Honduras has not been completed. Among us, author Robert Lehman has
been a resident and collected in and around La Ceiba for many years. His specimens, in part at the
National Museum of Natural History, Washington, DC (USNM) and in the Museo de Mariposas, La
Ceiba (RLMM), together with material at EAPZ, have comprised the most complete representation of
the Honduran Lepidoptera fauna to date. Both EAPZ and RLMM are strongest in macrolepidoptera
holdings. The microlepidopteran fauna of Honduras has, however, remained relatively unsampled and
unstudied. Even within macrolepidoptera, for butteries there has not been an updated species list for
Honduras in almost a half a century (Monroe et al. 1967, Monroe and Miller 1967).

Insecta MundI 0205, February 2012 • 5Lepidoptera of Honduras
Our knowledge of the life history, ecology, and biogeography of Honduran Lepidoptera is woefully
inadequate, compared to recent publications and information currently available on websites covering
adjacent countries. Given the massive deforestation occurring throughout Mesoamerica on a daily
basis (Tucker et al. 2005), we urgently need to improve our knowledge of the diversity and distribution
of Lepidoptera species here before we miss the opportunity, especially with regard to potential endemic
and key bioindicator species. The provisional list presented below is far from complete; however, we
feel this is an important rst step in establishing a foundation for our continuing inventory of the
Lepidoptera of Honduras.
Acronyms and Museum Codens
Along with references to the authors by initials [JYM, DLM, TCE, CVC, RL], numerous
institutional and locality acronyms are used in the text and list below. Four letter museum codens
are: EAPZ = Escuela Agricultura Panamerica in Zamorano, Honduras; MGCL = McGuire Center for
Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville;
USNM = National Museum of Natural History, Washington, DC; RLMM = Robert Lehman’s Museo
de Mariposas y Insectos, La Ceiba, Honduras. The following locality acronyms are also used in the text
and list: PBL = The Lodge at Pico Bonito, La Ceiba, Honduras; CURLA = reserve property of Parque
Nacional Pico Bonito managed by Centro Universitario Regional del Litoral Atlantico, Honduras; ER
= The Honduran Emerald Hummingbird Reserve, Rio Aguan Valley, Olanchito, Honduras; KC = The
Kellogg Center at Zamorano, EAPZ; UYUCA = Uyuca biological reserve of EAPZ; CA = Parque Nacional
Cerro Azul, Meambar, Honduras. Reference is likewise made to records based on type localities [TL]
and species included based on type localities of their current junior synonyms [TLS].
Methods
In June, 2009, a project was initiated by MGCL to survey the butteries and moths along the
northern coast of Honduras, concentrating in the area of Parque National Pico Bonito. Primary
collection sites include habitats in the vicinity of the Lodge at Pico Bonito, Estación CURLA, and the
Emerald Hummingbird Reserve, in the Rio Aguan Valley south of Pico Bonito (Map 1). Six trips to the
area were made by MGCL staff, students, and eld associates, with ve trips in 2009-2010 devoted to
collecting activities. Results of collecting trips are summarized in the present list by month and year.
These included an initial collection based trip (23–30 June, 2009) by JYM and DLM with collections
and/or species observed at the Kellogg Center of the EAPZ campus, Uyuca biological reserve of EAPZ,
Parque Nacional Cerro Azul Meambar, and the Lodge at Pico Bonito. A second trip (9–16 November
2009) by JYM and DLM with two students was made to PBL and included surveys at CURLA with
RL. A third trip (4–16 May 2010) by JYM and DLM with seven students/associates was made to PBL
with additional collections at CURLA. Two trips were made in June (12–20 June 2010) by TCE and
eight students/associates and (24–30 June 2010) by CVC and eight students/associates. The latter trip,
led in conjunction with Forum Herbulot VI at MGCL, focused on the Geometridae. A subsequent trip
(7–15 August 2010) by JYM and DLM and ve associates included collecting at PBL, CURLA, and ER.
Our principal collecting site, PBL (N 15° 41’ 48.00” W 86° 54’ 04.40”), is located near the base of the
mountains adjacent to Parque Nacional Pico Bonito at about 300 m (Fig 1). The property is bordered
on the east and west by two rivers, Rio Corinto and Rio Coloradito, and consists of a 400 acre preserve
with lowland tropical wet forest as well as coffee, cacao, banana and citrus groves. The CURLA reserve
(N 15° 42’ 07.00” W 86° 51’ 16.00”) is located to the east of PBL and includes areas of pristine primary
tropical forest. In contrast, ER (N 15° 25’ 44.00” W 86° 53’45.57”) is located about 24 km due south of
PBL in the rain shadow of the Sierra Nombre de Dios. This small reserve, part of a Honduran army
military base, consists of tropical dry forest characterized by thorny scrub vegetation, unique with
both cactus and orchids growing together and specialized nectar plants such as Pedilanthus Neck.
ex. Poit. (Euphorbiaceae), which support one of the few populations of the endemic and endangered
Honduran emerald hummingbird, Amazilia luciae (Lawrence).
Sampling methods involved hand collecting with nets as well as visual identication of easily
identiable diurnal species, bait traps containing fermenting fruit, and nightly collections at white

6 • Insecta MundI 0205, February 2012 Miller et al.
sheets illuminated by mercury vapor lamps or ultraviolet lights. While night collecting has thus far
been at low elevations (< 500 m), collecting at PBL was especially productive with mercury vapor lights
set at the top of a four-story bird observation tower in view of the Rio Coloradito and surrounding
mountains. All specimens were pinned, spread, and labeled, and deposited at MGCL with voucher
specimens also deposited at EAPZ and USNM. Legs have been removed from selected specimens
for subsequent DNA extraction and analysis. Tissue samples are stored at the Genetic Resources
Repository at the Florida Museum of Natural History.
Specimens were identied based on comparison with identied reference material at USNM, MGCL,
and EAPZ, as well as types or type images, primary literature accounts included in revisionary studies,
and older literature such as Cramer (1775-1782), Godman and Salvin (1879-1901), Seitz ([1907]-1924),
Hampson (1898-1913, 1914, 1920), Druce (1881-1900, 1888), Walsingham (1915), and more recent
works such as Lemaire (1971, 1973, 1974, 1978, 1980, 1988, 2002). Several internet websites have
been especially helpful with nomenclature and in obtaining initial identications and corroborating
determinations. Some examples of these include: Warren et al. (2011), Moth Photographers Group
(2011), Janzen and Hallwachs (2010), Barcode of Life Data Systems (2011), Lahousse (2011), Barnes
(2002), Smithsonian Institution (2011), Nuss et al. (2003-2011), and Phillips (2001).
In addition to records resulting from recent collecting efforts, Honduran species found in primary
literature and collection records of EAPZ, USNM, MCGL, and RLMM are incorporated into the present
list. Examples of general literature accounts yielding numerous records and type locality references
include Druce (1881-1900), Kirby (1892), Schaus (1920) and Walker (1854-1866), and some examples
of works focused on particular taxa with multiple records included Davis (1964), Pitkin (1993, 1996),
Poole (1987, 1989), Powell (1973), Scoble (1999) and numerous others.
Map 1. Natural Life Zones of Honduras (modied from Ratcliff and Cave 2006a, with permission) showing forest
types and location of collection/observation sites. See text above for location acronyms.

Insecta MundI 0205, February 2012 • 7Lepidoptera of Honduras
Annotated List of Honduran Lepidoptera
The provisional species list presented in table format below is a product of our ongoing biodiversity
inventory. Additional taxa are expected in future eld surveys. Each taxon is annotated to indicate
the source of voucher specimens, sight or collection locality records, or type localities using the
acronyms and codens listed above. Taxa included without annotations are based on records found
in the literature. Nomenclature and systematic arrangement for butteries generally follow Warren
et al. (2011), Pelham (2008), and/or Lamas (2004). Nomenclature and systematic arrangement for
the moths generally follows or is adapted from recent revisionary works and checklists including
Lafontaine and Schmidt (2010) for Noctuoidea, Watson and Goodger (1986) for Arctiinae, Scoble
(1999) for Geometroidea, and the Atlas of Neotropical Lepidoptera series (Heppner 1984, 1995, 1996)
for Pyraloidea and other groups. While we are aware of conicting nomenclature with the ongoing
systematic research of colleagues, our list includes names, subfamilial, and tribal assignments based
on published works only. Collection or sight records from recent surveys are marked with an “x” for
each species in columns to the right of the name according to the month and year: 1) 06/2009, 2)
11/2009, 3) 05/2010, 4) 06/2010, 5) 08/2010. Author names are spelled out except in one case (F. M.
Brown [FMB]) where it was necessary to use initials to maintain the single line format of the row.
Subfamilial and tribal names are included in taxa where this facilitates the understanding of the
ordering of the list.
Figure 1. Parque Nacional Pico Bonito, Pico Bonito Lodge and vicinity. a) Canopy adjacent to Rio Coloradito.
b) Canopy vegetation. c) View of forested range and lowland vegetation. d) Rio Coloradito. Figures a-c) 28 June
2009; d) 13 November 2009.

8 • Insecta MundI 0205, February 2012 Miller et al.
12345
--- HEPIALOIDEA ---
HEPIALIDAE
Druceiella sp. PBL x
--- TINEOIDEA ---
PSYCHIDAE
Cryptothelea symmicta (Dyar, 1914)
Cryptothelea gloverii (Packard, 1869)
Oiketicus geyeri Berg, 1877
Oiketicus kirbyi Guilding, 1827
Oiketicus sp. PBL x
ARRHENOPHANIDAE
Arrhenophanes perspicilla (Stoll, 1790) PBL x x
ACROLOPHIDAE
Acrolophus spp. EAPZ, PBL x x x
--- GRACILLARIOIDEA ---
GRACILLARIIDAE
Phyllocnistis citrella Stainton, 1856 EAPZ
--- YPONOMEUTOIDEA ---
YPONOMEUTIDAE
Atteva aurea (Fabricius, 1798) PBL x
PLUTELLIDAE
Plutella xylostella (Linnaeus, 1758) PBL, EAPZ x
HELIODINIDAE
Aetole bella Chambers, 1875
LYONETIIDAE
Perileucoptera coffeella (Guérin-Méneville, 1842) EAPZ
--- GELECHIOIDEA ---
ELACHISTIDAE
Ethmia catapeltica Meyrick, 1924 or nr. PBL x x x x
Ethmia lichyi Powell 1973 TL
Ethmia perpulchra Walsingham, 1912
Ethmia scythropa Walsingham, 1912
Ethmia similatella Busck, 1920
Ethmia ungulatella Busck, 1914 PBL x x x x
Stenoma catenifer Walsingham, 1912 EAPZ
Stenoma exarata (Zeller, 1854) PBL x
Stenoma impressellla (Busck, 1914) PBL x
Stenoma patens Meyrick, 1913 PBL x x x x
GLYPHIDOCERIDAE
Glyphidocera sp. 1 PBL x x
Glyphidocera sp. 2 PBL x
Glyphidocera sp. 3 PBL x
Glyphidocera sp. 4 PBL x
OECOPHORIDAE
Cerconota anonella (Sepp, 1830) EAPZ
Antaeotricha cosmoterma (Meyrick, 1930) PBL x x
Antaeotricha particularis (Zeller, 1877) PBL x x
Antaeotricha spp. 1-5 PBL x x x x

Insecta MundI 0205, February 2012 • 9Lepidoptera of Honduras
COLEOPHORIDAE
Blastobasinae
Blastobasis sp. PBL x x
COSMOPTERIGIDAE
Pyroderces badia (Hodges, 1962) EAPZ
Pyroderces rileyi (Walsingham, 1882) EAPZ
GELECHIIDAE
Keiferia lycopersicella (Walsingham, 1897) EAPZ
Phthorimaea operculella (Zeller, 1873) EAPZ
Sitotroga cerealella (Oliver, 1789) EAPZ
Trichotaphe arotrosema (Walsingham, 1911) PBL x x
--- ZYGAENOIDEA ---
MEGALOPYGIDAE
Aithorape sp. PBL x x x x
Macara nigripes (Dyar, 1909) PBL x
Megalopyge albicollis (Walker, 1855) PBL x x x
Megalopyge lanata (Cramer, 1780) PBL x
Megalopyge opercularis J.E. Smith, 1797 PBL x x x
Megalopyge sp. PBL x
Mesoscia dumilla Dyar, 1913 PBL x x x x
Mesoscia pusilla (Stoll, 1782)
Podalia contigua (Walker, 1886) PBL x
Podalia orsilochia (Cramer, 1775) PBL x
Podalia tympania (Druce, 1897) or nr. PBL x x
Trosia nigropunctigera (Fletcher, 1982) PBL xxxx
LIMACODIDAE
Acharia horrida (Dyar, 1905) PBL x x x
Euclea bidiscalis Dyar, 1926 PBL x x
Euclea mesoamericana Epstein and Corrales, 2004, PBL x
Euclea sp. 1 PBL x x
Euclea sp. 2 PBL x x x
Miresa clarissa (Stoll, 1790)
Natada cecilia Corrales and Epstein, 2000
Natada confusa Corrales and Epstein, 2003
Natada fusca (Druce, 1887) PBL xxxx
Parasa minima Schaus, 1892 PBL x
Parasa joanae Epstein, 2004 PBL x
Parasa viridogrisea Dyar, 1898 PBL x x
Perola invaria (Walker, 1855) TL
Perola sericea (Möschler, 1878) PBLx x
Phobetron hipparchia (Cramer, [1777]) EAPZ
Prolimacodes undifera (Walker, 1855) TL
Prolimacodes triangulifera Schaus, 1896 PBL x x x
Semyra nita Walker, 1855 PBL x x x
Talima aurora Dyar, 1926 PBL x
Talima postica Dyar, 1926 PBL x
DALCERIDAE
Acraga coa (Schaus, 1892) EAPZ
Acraga sp. PBL x x

10 • Insecta MundI 0205, February 2012 Miller et al.
Dalcerides alba (Druce, 1887) PBL x
Dalcerides mesoa (Druce, 1887) PBL x x x
Dalcerides sp. PBL x x
Paracraga argentea (Schaus, 1910)
ZYGAENIDAE.
Harrisina mystica (Walker, 1854) TL
--- COSSOIDEA ---
COSSIDAE
Aramos ramosa (Schaus, 1892) PBL xxx
Biocellata alfarae (Schaus, 1911) PBL x x
Brypoctia strigifer (Dyar, 1910) PBL x x
Cossula arpi Schaus, 1901 PBL x
Cossula tacita (Druce, 1898)
Givira basiplaga (Schaus, 1905) PBL x
Givira daphne Druce, 1901 PBL x
Givira juturna (Schaus, 1892) PBL x x
Givira modisma Schaus, 1921 PBL x
Inguromorpha sandelphon (Dyar, 1912) PBL x
Langsdora lunifera Dyar, 1937 PBL x x
Morpheis cognata (Walker, 1856) TL
Morpheis comisteon Schaus, 1911 PBL x x
Morpheis pyracmon (Cramer, 1780) PBL x x
--- SESIOIDEA ---
SESIIDAE
Carmenta mimosa Eichlin and Passoa, 1983 TL
Sophona hondurasensis Eichlin, 1986 TL
Melittia cucurbitae Harris, 1828 EAPZ
CASTNIIDAE
Castniinae
Telchin atymnius futilis (Walker, 1856) PBL, EAPZ, CURLA x x x
Amauta cacica procera (Boisduval, [1875]) RLMM, EAPZ
Athis i. inca (Walker, 1854) EAPZ
Athis clitarcha (Westwood, 1877) EAPZ, RLMM
Divana d. diva (Butler, [1870]) PBL, RLMM x
Telchin licus (Drury, 1773) RLMM, EAPZ
--- CHOREUTOIDEA ---
CHOREUTIDAE
Brenthia sp. PBL x
Hemerophila tristis (Felder and Rogenhoffer, 1875) or nr. PBL x
Tortyra sp. PBL x
--- TORTRICOIDEA ---
TORTRICIDAE
Mictopsichia cubae Razowski, 2009 PBL, CURLA x x
Aethesoides hondurasica Razowski, 1986 TL
Amorbia productana (Walker, 1863) TL
Cydia semicirculana (Walker, 1863) TL
Eugnosta emarcida (Razowski and Becker 1986) TL
Eugnosta fraudulenta Razowski and Becker 2007
Icteralaria idiochroma J.W. Brown, 1996

Insecta MundI 0205, February 2012 • 11Lepidoptera of Honduras
Mimeugnosta particeps Razowski, 1986 TL
Platynota rostrana (Walker, 1863)
Rhyacionia frustrana (Comstock, 1880)
Sparganothina anopla B. Landry, 2001 TL
Pseudatteria volcanica (Butler, 1872) CURLA x
--- ALUCITOIDEA ---
ALUCITIDAE
Alucita sp. PBL x x x x
--- PTEROPHOROIDEA ---
PTEROPHORIDAE
Ochyrotica sp. PBL x
Leptodeuterocopus sp. PBL x
Anstenoptilia marmarodactyla (Dyar, [1903])
Stenoptilodes brevipennis (Zeller, 1874) PBL, EAPZ x x
Amblyptilia landryi Gielis, 2006 TL
Lioptilodes albistriolatus (Zeller, 1877) EAPZ
Michaelophorus dentiger (Meyrick, 1916)
Michaelophorus nubilus (Felder and Rogenhoffer, 1875) PBL x x x x
Sphenarches anisodactylus (Walker, 1864) PBL x
Megalorhipida dulcis (Walsingham, 1915)
Megalorhipida leucodactylus (Fabricius, 1794)
Hellinsia sp. 1 UYUCA x
Hellinsia sp. 2 PBL x x
Hellinsia sp. 3 PBL x x
Adaina beckeri Gielis, 1992 PBL x
Adaina ipomoeae Bigot and Etienne, 2009 PBL x
Adaina simplicius (Grossbeck, 1917) PBL x
--- HYBLAEOIDEA ---
HYBLAEIDAE
Hyblaea puera (Cramer, 1777)
--- PYRALOIDEA ---
CRAMBIDAE
Crambinae
Microcausta sp. PBL x
Argyria centrifugens Dyar, 1914 PBL x x x
Argyria sp. KC, EAPZ x
Myelobia smerintha Hübner, [1821] PBL x
Fissicrambus minuellus (Walker, 1863) TL
Tortriculladia sp. PBL x x x
Tortriculladia eucosmella (Dyar, 1914) PBL
Microcrambus pusionellus (Zeller, 1863) PBL x
Microcrambus tactellus (Dyar, 1914) PBL x
Microcramboides meretricellus (Schaus, 1913) PBL x x
Diatraea lineolata (Walker, 1856) EAPZ
Diatraea saccharalis (Fabricius, 1794) EAPZ, TLS
Parapediasia tenuistrigata (Zeller, 1881) PBL x x
Schoenobiinae
Proschoenobius subcervinellus (Walker, 1863) TL
Rupela albina Becker and Solis, 1990 EAPZ

12 • Insecta MundI 0205, February 2012 Miller et al.
Glaphyriinae
Hellula phidilealis (Walker, 1859) EAPZ
Aethiophysa dimotalis (Walker, [1866]) TL
Cosmopterosis spatha Solis, 2009 PBL x x
Lipocosma calla (Kaye, 1901) PBL x
Pseudoligostigma enareralis (Dyar, 1914) PBL x
Dicymolomia metalophota (Hampson, 1897) PBL x
Schacontia ysticalis (Dyar, 1925) PBL x
Musotiminae
Neurophyseta camptogrammalis Hampson, 1912 PBL x
Undulambia polystichalis Capps, 1965 PBL x
Midilinae
Midila centralis Munroe, 1970 PBL x x
Midila d. daphne (Druce, 1895) PBL x
Dismidila drepanoides Monroe, 1970 PBL x
Dismidila atoca Dyar, 1914 PBL x x
Dismidila sp. PBL x
Acentropinae
Aulacodes aechmialis Guenée, 1854 KC x
Aulacodes sp. 1 PBL x
Aulacodes sp. 2 PBL x x
Parapoynx endoralis (Walker, 1859) TL
Chrysendeton romanalis (Druce, 1896) PBL x
Argyractoides nr. lucianalis (Schaus, 1924) PBL x x
Argyractoides leucogonialis (Hampson, 1906) or nr. PBL x x
Argyractoides sp. PBL x x x
Neargyractis sp. PBL x x x
Usingeriessa onyxalis (Hampson, 1897) PBL x
Petrophila mignonalis (Dyar, 1914) PBL x x x x
Petrophila iolepta (Dyar, 1914) PBL x
Petrophila argyrolepta (Dyar, 1914) PBL x x x x
Petrophila zelota (Dyar, 1914) PBL x x
Petrophila parvissimalis (Schaus, 1912) PBL xxx
Petrophila spp. 1-8 PBL x x x x
Oxyelophila puralis (Schaus, 1912) PBL x x
Oxyelophila necomalis (Dyar, 1914) PBL x x x
Oxyelophila sp. PBL x x
Odontiinae
Dicepolia rutinctalis (Hampson, 1899) PBL x
Evergestinae
Symphysa sp. PBL x x
Trischistognatha pyrenealis (Walker, 1859) PBL x x
Pyraustinae
Munroeodes thalesalis (Walker, 1859) TL
Epicorsia alvilalis Amsel, 1954 PBL x
Epicorsia sp. PBL x x
Oenobotys vinotinctalis (Hampson, 1895) PBL x x
Triuncidia ossealis (Hampson, 1899) PBL x
Anania inclusalis (Walker, [1866]) TL

Insecta MundI 0205, February 2012 • 13Lepidoptera of Honduras
Aponia minnithalis (Druce, 1895) PBL x
Pyrausta insignitalis (Guenée, 1854) TLS
Pyrausta bicoloralis (Guenée, 1854) PBL x
Hyalorista limasalis (Walker, 1886) TL
Hyalorista taeniolalis (Guenée, 1854) PBL x
Portentomorpha xanthialis (Guenée, 1854) PBL x
Spilomelinae
Phaedropsis chromalis (Guenée, 1854) PBL x
Phaedropsis maritzalis (Schaus, 1920) PBL x x x
Lygropia acastalis (Walker, 1859) TL
Lypotigris reginalis (Cramer, 1781) PBL x
Microthyris anormalis (Guenée, 1854) PBL x
Microthyris prolongalis (Guenée, 1854) PBL x
Pantographa expansalis (Lederer, 1863) PBL x x
Pantographa suffusalis Druce, 1895 PBL x x
Pantographa sp. PBL x
Phostria mapetalis (Schaus, 1912) PBL x x
Phostria tedea (Cramer, 1781) PBL x x
Phostria sp. PBL x
Phrygonodes plicatalis Guenée, 1854 PBL x
Syllepte amando (Cramer, 1779) KC x
Syllepte dioptalis (Walker, [1866]) TLS
Syllepte belialis (Walker, 1859) TLS
Cryptobotys zoilusalis (Walker, 1859) TL
Herpetogramma salbialis (Hampson, 1899) PBL x x
Herpetogramma bipunctalis (Fabricius, 1794) EAPZ
Herpetogramma cervinicosta (Hampson, 1918) PBL x x
Pilocrocis ramentalis Lederer, 1863 PBL x x x x
Pilocrocis cryptalis (Druce, 1895) PBL x x
Pilocrocis cyranoalis Schaus, 1920 PBL x
Pilocrocis sp. PBL x
Lineodes triangulalis Möschler, 1890 PBL x x
Lineodes fontella Walsingham, 1913 PBL x
Lineodes mesodonta Hampson, 1913 PBL x x
Lineodes tridentalis Hampson, 1913 EAPZ
Atomopteryx doeri Walsingham, 1891 PBL x x x
Lamprosema lunulalis Hübner, 1823 PBL x x x
Lamprosema nr. moccalis Schaus, 1920 PBL x
Lamprosema cayugalis Schaus, 1920 PBL x x
Lamprosema sp. PBL x
Neoleucinodes elegantalis (Guenée, 1854) PBL xxx
Neoleucinodes imperialis (Guenée, 1854)
Apogeshna stenialis (Guenée, 1854) PBL x x x x
Apogeshna acestealis (Walker, 1859) TLS
Blepharomastix coeneusalis (Walker, 1859) TL
Blepharomastix costaliparilis Munroe, 1995 CURLA, PBL x x
Blepharomastix benetinctalis (Dyar, 1914) PBL x x x x
Desmia albisectalis (Dognin, 1905) PBL x x
Desmia bajulalis Guenée, 1854 PBL xxx

14 • Insecta MundI 0205, February 2012 Miller et al.
Desmia cristinae Schaus, 1912 PBL x x x
Desmia daedala Druce, 1895 PBL x x x
Desmia extrema (Walker, 1856) PBL x x
Desmia odontoplaga Hampson, 1899 PBL x x
Desmia sepulchralis Guenée, 1854 PBL x x
Desmia vulcanalis (Felder and Rogenhoffer, 1875) PBL x x
Desmia sp. 1 PBL x x x
Desmia sp. 2 PBL x
Desmia sp. 3 PBL x
Desmia sp. 4 PBL x
Sacculosia isaralis (Felder and Rogenhoffer, 1875) PBL x
Hymenia perspectalis (Hübner, 1796) PBL xxxxx
Spoladea recurvalis (Fabricius, 1775) EAPZ
Spilomela discordens Dyar, 1914 PBL x x x x
Spilomela pantheralis (Geyer, 1832) PBL x
Spilomela sp. PBL x
Steniodes acuminalis (Dyar, 1914) PBL xxx
Steniodes mendica (Hedemann, 1894) EAPZ
Anarmodia nebulosalis Dognin, 1903 PBL x x
Sparagmia g. gonoptera (Latreille, 1828) PBL x x
Asturodes mbriauralis (Guenée, 1854) PBL x x x x
Diaphania nitidalis (Cramer, 1781) EAPZ, PBL x x
Diaphania arguta (Lederer, 1863) PBL x
Diaphania exclusalis (Walker, [1866]) PBL x
Diaphania latilimbalis (Guenée, 1854) PBL x x
Diaphania modialis (Dyar, 1912) PBL x
Diaphania hyalinata (Linnaeus, 1767) PBL, EAPZ x x
Diaphania costata (Fabricius, 1794) PBL xxxx
Diaphania esmeralda (Hampson, 1899) PBL x
Diaphania sp. PBL x
Glyphodes rubrocinctalis (Guenée, 1854) TLS, PBL xxx
Glyphodes s. sibillalis Walker, 1859 PBL x x
Hoterodes ausonia (Cramer, 1777) PBL x x
Leucochroma sp. nr. corope (Cramer, 1781) PBL x
Maruca vitrata (Fabricius, 1787) EAPZ, PBL xxxx
Omiodes indicata (Fabricius, 1775) EAPZ, PBL x x
Omiodes stigmosalis (Warren, 1892) PBL x
Omiodes martini Amsel, 1956 PBL x
Omiodes martyralis (Lederer, 1863) PBL x x
Omiodes simialis Guenée, 1854 PBL x
Omiodes humeralis Guenée, 1854 PBL xxxx
Omiodes pandaralis (Walker, 1858) PBL x x
Omiodes janzeni Gentili and Solis, 1998 PBL x x
Omiodes spp. 1-5 PBL x x x x
Palpita egia (Cramer, 1777) PBL x
Prenesta fenestrinalis (Guenée, 1854) PBL x x x x
Prenesta scyllalis (Walker, 1859) PBL x x x x
Synclera jarbusalis (Walker, 1859) PBL x x
Synclera chlorophasma (Butler, 1878)

Insecta MundI 0205, February 2012 • 15Lepidoptera of Honduras
Agathodes d. designalis Guenée, 1854 PBL xxx
Azochis mactalis (Felder and Rogenhoffer, 1875) PBL x
Azochis gripusalis Walker, 1859 PBL x x x x
Compacta hirtalis (Guenée, 1854) TLS
Megastes grandalis Guenée, 1854 PBL x
Megastes sp. PBL x
Polygrammodes h. hyalomaculata Dognin, 1908 PBL x x
Polygrammodes runalis Hampson, 1899 PBL x
Polygrammodes ponderalis (Guenée, 1854) PBL x x
Polygrammodes avidalis (Guenée, 1854) PBL x x
Polygrammodes elevata (Fabricius, 1794) PBL x
Syngamilyta apicolor (Druce, 1902) PBL x
Syngamilyta samarialis (Druce, 1899) PBL x
Terastia meticulosalis Guenée, 1854 PBL x
Cirrhocephalina brunneivena (Hampson, 1913) PBL x
Eulepte concordalis Hübner, [1825] 1826 or nr. PBL x x x x
Gonocausta zephyralis Lederer, 1863 PBL x x x
Leucochromodes melusinalis (Walker, 1859) PBL x x
Syllepis hortalis (Walker, 1859) PBL x x x
Ategumia ebulealis (Guenée, 1854) PBL x
Ategumia matutinalis (Guenée 1854) PBL xxx
Ategumia actealis (Walker, 1859) TLS
Ategumia dilecticolor (Dyar, 1912) PBL x x x
Ategumia sp. PBL x x
Epipagis fenestralis (Hübner, 1796) PBL x
Mimophobetron pyropsalis (Hampson, 1904) PBL x
Samea multiplicalis (Guenée, 1854) TLS
Samea nicaeusalis (Walker, 1859) TL
Samea ecclesialis Guenée, 1854 PBL xxxx
Samea delicata Kaye, 1923 PBL x
Arthromastix lauralis (Walker, 1859) PBL x
Psara prumnides (Druce, 1895) PBL x
Conchylodes salamisalis Druce, 1895 PBL x x x
Conchylodes nolckenialis (Snellen, 1875) PBL x x x x
Conchylodes arcifera Hampson, 1912 PBL x
Conchylodes platinalis (Guenée, 1854) PBL x x
Pycnarmon cecinalis (Dognin, 1897) PBL x x
Marasmia cochrusalis (Walker, 1859) PBL x x
Marasmia trapezalis (Guenée, 1854) USNM
Salbia tytiusalis (Walker, 1859) TL
Salbia haemorrhoidalis (Guenée, 1854) PBL x
Salbia pepitalis (Guenée, 1854) PBL x
Sufetula diminutalis (Walker, [1866]) TL
Syngamia orella (Cramer, 1781) EAPZ, PBL x x x x
Apilocrocis excelsalis (Schaus, 1912) PBL x x
Bocchoropsis plenilinealis (Dyar, 1917) PBL xxxx
Coenostolopsis apicalis (Lederer, 1863) PBL x x x
Deuterophysa albilunalis (Hampson, 1913) PBL x x x x
Dichocrocis clystalis Schaus, 1920 PBL x

16 • Insecta MundI 0205, February 2012 Miller et al.
Dichocrosis tlapalis Schaus, 1920 PBL x
Dichocrocis sabatalis (Druce, 1895) PBL xxxx
Maracayia chlorisalis (Walker, 1859) EAPZ, PBL x
Massepha lupa (Druce, 1899) PBL x
Metoeca foedalis (Guenée, 1854) PBL x
Microphysetica hermeasalis (Walker, 1859) PBL xxx
Nonazochis graphialis (Schaus, 1912) PBL x x
Palpusia terminalis (Dognin, 1910) PBL x
Palpusia sp. PBL x
Sisyracera subulalis (Guenée, 1854) PBL x
Trichaea pilicornis Herrich-Schäffer, 1866 PBL x x
Trichaea sp. PBL x x x x x
PYRALIDAE
Pyralinae
Hypsopygia craspedalis (Hampson, 1906) PBL x x x x
Hypsopygia resectalis (Lederer, 1863) PBL x x
Hypsopygia graaalis (Snellen, 1875) PBL x x
Chrysauginae
Azamora splendens (Druce, 1895) PBL x x
Bonchis munitalis (Lederer, 1863) TLS, PBL x x
Bonchis scoparioides Walker, 1862 PBL x x
Caphys biliniata (Stoll, 1781) TLS, PBL xxxxx
Caphys subrosealis (Walker, [1866]) TL
Carcha hersilialis Walker, 1859 TL
Carcha violalis Hampson, 1897 PBL x x x
Clydonopteron pomponius Druce, 1895 PBL x x x
Clydonopteron sp. PBL x
Dasycnema obliqualis (Hampson, 1897) PBL x x x x
Galasa strenualis Dyar, 1914 PBL x x x x
Galasa striginervalis Hampson, 1906 PBL x x
Galasa sp. 1 PBL x x x
Galasa sp. 2 PBL x x x
Galasa sp. 3 PBL x
Hyperparachma bursarialis (Walker, [1866]) TL x
Hyperparachma sp. PBL x x x x
Lophopleura eurzonalis Hampson, 1897 PBL x
Parachma meterythra Hampson, 1897 PBL x x x
Salobrena atropurpurea Hampson, 1906 PBL x
Salobrena vacuana (Walker, 1863) PBL x
Tetrachistis paula Schaus, 1904 PBL x x
Xantippe olivalis Dyar, 1914 PBL x x x
Zanclodes falculalis Ragonot, 1891 PBL x x
Epipaschiinae
Accinctapubes albifasiata (Druce, 1902) PBL x
Anarnatula sylea (Druce, 1899) PBL x x
Arnatula circumlucens Dyar, 1914 PBL x x x
Calybitia adolescens (Dyar, 1914) PBL x
Carthara albicosta Walker, 1865 PBL x x x x
Chloropaschia selecta (Schaus, 1912) PBL x

Insecta MundI 0205, February 2012 • 17Lepidoptera of Honduras
Deuterollyta cantianilla (Schaus, 1925) PBL x x
Deuterollyta majuscula Herrich-Schäffer, 1871 PBL x
Deuterollyta ragonoti Möschler, 1890 PBL xxx
Epipaschia mesoleucalis (Hampson, 1916) PBL x x x x
Macalla thyrsisalis Walker, [1859] TL, PBL x x
Macalla nstanalis Schaus, 1922 PBL x x
Milgithea melanoleuca (Hampson, 1896) PBL x x x
Pandoabella olivescens (Schaus, 1912) PBL xxx
Phidotricha erigens Ragonot, 1888 PBL, KC x x
Lepidogma modana Schaus, 1922 PBL x x x
Pococera narthusa Schaus, 1913 PBL x x x
Pococera gelidalis (Walker, [1866]) TL
Pococera sp. PBL x x x x
Tallula fovealis Hampson, 1906 PBL x x x x
Tallula melazonalis Hampson, 1906 PBL x
Tancoa crinita (Schaus, 1912) PBL x x
Tancoa calitas (Druce, 1899) PBL x x
Tancoa sp. PBL x x x
Phycitinae
Ectomyelois muriscis (Dyar, 1914) PBL x x x
Difundella sp. x
Eurythmidia ignidorsella (Ragonot, 1887) PBL x x
Caudellia declivella (Zeller, 1881) or nr. PBL x
Davara caricae (Dyar, 1913) PBL x x
Dioryctria batesella Mutuura and Neunzig, 1986 PBL x
Dioryctria erythropasa (Dyar, 1914)
Dioryctria sp. PBL x
Drescomopis soraella (Druce, 1899) PBL x
Anabasis ochrodesma (Zeller, 1881) PBL x
Piesmopoda sp. PBL x x x x
Tlascala reductella (Walker, 1863) TL
Ufa rubedinella (Zeller, 1848) TLS
--- THYRIDOIDEA ---
THYRIDIDAE
Banisia furva epasta Whalley, 1976
Draconia rusina Druce, 1895
Draconia denticulata (Pagenstecher, 1892) or nr. PBL x
Dysodia sp. 1 PBL x
Dysodia sp. 2 PBL x
Gippius sumptuosus Walker, 1855 TL
Microsca sp. CURLA x
Pseudendromis thetis (Druce, 1899) PBL x x
Rhodoneura nebulosa (Warren, 1889) or nr. PBL xxxx
Zeuzerodes fumatilis (Pagenstecher, 1892) PBL x
undet. sp. 1 PBL x
undet. sp. 2 PBL x
undet. sp. 3 PBL x
undet. sp. 4 PBL x

18 • Insecta MundI 0205, February 2012 Miller et al.
--- MIMALLONOIDEA ---
MIMALLONIDAE
Cicinnus sp. PBL x
Druentica partha (Schaus, 1905) or nr. PBL x
Roelmana maloba (Scahus, 1905) PBL x
--- LASIOCAMPOIDEA ---
LASIOCAMPIDAE
Artace cribraria (Ljungh, 1825) PBL x x
Euglyphis amathuria Druce, 1890 PBL x
Euglyphis amisena (Druce, 1890)
Euglyphis canifascia (Walker, 1869)
Euglyphis cercina (Druce, 1897)
Euglyphis guttularis (Walker, 1855)
Euglyphis poasia Schaus, 1910 PBL x x x x
Euglyphis thyatira (Druce, 1887) PBL x x
Euglyphis zurcheri Druce, 1897 PBL x x
Euglyphis sp. 1 PBL x
Euglyphis sp. 2 PBL x
Eutachyptera psidii Sallé, 1857 EAPZ
Nesara casada (Schaus, 1911) PBL x x
Nesara sp. PBL x
Prorifrons sp. PBL x
Tolype caieta Druce, 1897 PBL x
--- BOMBYCOIDEA ---
BOMBYCIDAE
Apatelodes satellitia (Walker, 1855) PBL xxxx
Apatelodes merlona Schaus, 1939 PBL x x x
Colla coelestis Schaus, 1910 PBL x x x
Epia muscosa (Butler, 1878) PBL xxx
Epia casnonia (Druce, 1887) PBL x
Olceclostera indentata Schaus, 1910 or nr. PBL x x x
Olceclostera sp. PBL x x x
SATURNIIDAE
Oxyteninae
Oxytenis modestia (Cramer, [1780]) PBL x x x
Oxytenis beprea Druce, 1886 or nr. PBL x
Arsenurinae
Arsenura armida (Cramer, 1779) PBL, EAPZ, RLMM x x
Arsenura batesii (Felder and Rogenhofer, 1874)
Caio championi (Druce, 1886) EAPZ, RLMM
Copiopteryx semiramis banghaasi Draudt 1930 EAPZ
Dysdaemonia boreas (Cramer, [1775]) PBL, EAPZ, RLMM xxxx
Titaea tamerlan nobilis (Schaus, 1912) EAPZ, RLMM x
Titaea sp. nr tamerlan (Maasen, 1869) PBL x
Rhescyntis hippodamia (Cramer, 1777) EAPZ, RLMM x x
Ceratocampinae
Adeloneivaia boisduvalii (Doûmet, 1859) PBL, EAPZ x
Adeloneivaia jason (Boisduval, 1872) PBL, EAPZ, RLMM x
Adeloneivaia subangulata (Herrich-Schäffer, [1855]) PBL, EAPZ, RLMM x x

Insecta MundI 0205, February 2012 • 19Lepidoptera of Honduras
Adelowalkeria caeca Lemaire, 1969 PBL x
Citioica anthonilis (Herrich-Schäffer, [1854]) EAPZ
Citheronia lobesis Rothschild, 1907 EAPZ
Citheronia bellavista Draudt, 1930 EAPZ, RLMM
Citheronoides collaris (Rothschild, 1907) RLMM, PBL x
Eacles imperialis (Drury, 1773) PBL x
Eacles imperialis decoris Rothschild, 1907 EAPZ, RLMM
Eacles masoni Schaus, 1896 EAPZ
Eacles ormondei Schaus, 1889 EAPZ, RLMM, PBL x
Othorene purpurascens (Schaus, 1905) EAPZ, RLMM
Othorene verana (Schaus, 1900) EAPZ, RLMM x
Schausiella santarosensis Lemaire, 1982 PBL, RLMM x
Syssphinx sp. PBL x
Syssphinx colla Dyar, 1907 PBL, EAPZ, RLMM x x
Syssphinx mexicana (Boisduval, 1872) EAPZ, RLMM
Syssphinx molina (Cramer 1780) PBL, EAPZ, RLMM x x
Syssphinx montana (Packard, 1905) PBL x
Syssphinx quadrilineata (Grote and Robinson, 1867) EAPZ, RLMM, PBL x
Hemileucinae
Automeris banus (Boisduval, 1875) EAPZ, RLMM
Automeris belti Druce 1886 PBL, EAPZ, RLMM x x x x x
Automeris excreta Draudt, 1929 RLMM
Automeris hamata Schaus, 1906 EAPZ
Automeris jucunda (Cramer, 1779)
Automeris lauta Johnson and Michener, 1948 EAPZ
Automeris metzli (Sallé, 1853) EAPZ, RLMM
Automeris moloneyi Druce, 1897 EAPZ, RLMM
Automeris zozine Druce, 1886 EAPZ, RLMM
Automeris pallidior Draudt, 1929 EAPZ, RLMM
Automeris phrynon Druce, 1897 EAPZ
Automeris postalbida Schaus, 1900 PBL, EAPZ, RLMM x
Automeris rubrescens (Walker, 1855) EAPZ
Automeris tridens (Herrich-Schäffer, 1855) EAPZ
Dirphiopsis ora (Schaus, 1911)
Hylesia lineata Druce, 1886 PBL, RLMM x
Hylesia continua (Walker, 1865) PBL, RLMM xxxx
Hylesia rubrifrons Schaus, 1911 PBL x x x
Hylesia gamelioides Michener, 1952 EAPZ, RLMM
Hylesia tinturex Schaus, 1921 EAPZ
Hylesia umbratula Dyar, 1927 EAPZ
Hylesia r. rosacea Schaus, 1911, PBL x x x
Gamelia abas (Cramer, 1775) PBL x x
Gamelia septentrionalis Bouvier, 1936 RLMM
Hyperchira nausica (Cramer, 1779) PBL, RLMM x x x x
Leucanella acutissima Walker, 1865 EAPZ, RLMM
Leucanella contempta Lemaire, 1967 EAPZ
Lonomia electra Druce 1886 EAPZ, RLMM, PBL x x
Molippa nibasa (Maassen and Weyding, 1885) PBL, EAPZ, RLMM x x
Dirphia avia (Stoll, 1780) PBL x

20 • Insecta MundI 0205, February 2012 Miller et al.
Paradirphia boudinoti Lemaire and Wolfe, 1990
Paradirphia rectilineata Wolfe, 1994 EAPZ
Paradirphia semirosea (Walker 1855) EAPZ, RLMM
Periphoba arcaei E. D. Jones, 1908 PBL, RLMM x x x
Pseudodirphia menander (Druce, 1886) EAPZ, RLMM
Pseudodirphia regia (Draudt, 1930) EAPZ, RLMM
Saturniinae
Antheraea godmani (Druce 1892) EAPZ, RLMM
Copaxa escalantei Lemaire, 1971 EAPZ, RLMM
Copaxa cydippe (Druce, 1894) EAPZ
Copaxa evelynae Wolfe and Lemaire (1993) EAPZ
Copaxa lavendera (Westwood, 1853) EAPZ
Copaxa mazaorum Lemaire, 1982 RLMM
Copaxa multifenestrata (Herrich-Schäffer, 1858) PBL, EAPZ, RLMM x
Copaxa runans Schaus, 1906 PBL, EAPZ, RLMM x
Copaxa curvilinea Schaus, 1912 EAPZ
Copaxa sophronia Schaus, 1921 EAPZ, RLMM
Eupackardia calleta (Westwood, 1853) EAPZ
Rothschildia erycina mexicana EAPZ, RLMM
Rothschildia lebeau (Guérin-Méneville 1868) PBL, EAPZ, RLMM x
Rothschildia orizaba (Westwood, 1853) EAPZ, RLMM
Rothschildia roxana Schaus, 1905 Bouvier, 1936 RLMM, PBL x
SPHINGIDAE
Macroglossinae
Aellopos ceculus (Cramer, 1777) EAPZ, PBL x
Aellopos clavipes Rothschild and Jordan, 1903
Aellopos titan (Cramer, 1777) EAPZ
Aleuron chloroptera (Perty, 1833) EAPZ
Aleuron neglectum Rothschild and Jordan, 1903 EAPZ
Callionima denticulata (Schaus, 1895), PBL, EAPZ x
Callionima falcifera (Gehlen, 1943) PBL, EAPZ xxx
Callionima nomius (Walker, 1856), PBL, EAPZ x x
Callionima pan (Cramer, 1779) EAPZ
Callionima parce (Fabricius, 1775) EAPZ, PBL x x
Enyo gorgon (Cramer, 1777) EAPZ
Enyo l. lugubris (Linnaeus, 1771) EAPZ, PBL x
Enyo ocypete (Linnaeus, 1758) EAPZ, PBL x x x
Enyo taedium Schaus, 1890 PBL
Erinnyis alope (Drury, 1773) PBL x
Erinnyis crameri (Schaus, 1898) EAPZ
Erinnyis e. ello (Linnaeus, 1758) EAPZ
Erinnyis lassauxii (Boisduval, 1859) EAPZ
Erinnyis obscura (Fabricius, 1775) EAPZ
Erinnyis oenotrus (Cramer, 1780) EAPZ, PBL x x
Erinnyis yucatana (Druce, 1888)
Hemeroplanes ornatus Rothschild, 1894 PBL x x x
Hemeroplanes triptolemus (Cramer, 1779) EAPZ
Isognathus rimosa (Grote, 1865) EAPZ
Isognathus scyron (Cramer, 1780) RLMM

Insecta MundI 0205, February 2012 • 21Lepidoptera of Honduras
Nyceryx coffaeae (Walker, 1856) EAPZ
Nyceryx eximia (Rothschild and Jordan, 1916) EAPZ
Nyceryx riscus (Schaus, 1890) EAPZ
Nyceryx tacita (Druce, 1888) EAPZ
Madoryx oiclus (Cramer, 1779) EAPZ
Madoryx plutonius (Hübner, 1819) PBL, EAPZ xxx
Pachylia darceta Druce, 1881 PBL, EAPZ x x x x x
Pachylia cus (Linnaeus, 1758) PBL xxx
Pachylia s. syces (Hübner, 1819) EAPZ
Pachylioides resumens (Walker, 1856) EAPZ, PBL xxxx
Perigonia jamaicensis Rothschild, 1894 EAPZ
Perigonia lusca (Fabricius, 1777) EAPZ, PBL x x
Perifonia pallida Rothschild and Jordan, 1903, PBL x x
Perigonia stulta Herrich-Schäffer, 1854 EAPZ, RLMM
Phryxus caicus (Cramer, 1777) EAPZ
Pseudosphinx tetrio (Linnaeus, 1771) EAPZ
Stolidoptera tachasara (Druce, 1888) EAPZ
Unzela j. japix (Cramer, 1776) EAPZ
Cautethia spuria (Boisduval, 1875) EAPZ
Cautethia yucatana Clark, 1919
Hyles l. lineata (Fabricius., 1775) EAPZ
Xylophanes acrus Rothschild and Jordan, 1910 EAPZ
Xylophanes amadis cyrene (Druce, 1881) PBL x
Xylophanes anubus (Cramer, 1777) EAPZ
Xyophanes belti (Druce, 1878) EAPZ, PBL x x
Xylophanes falco (Walker, 1856) TL
Xylophanes ceratomioides Grote and Robinson, 1867 EAPZ, PBL x x
Xylophanes chiron nechus (Cramer, 1777) PBL, EAPZ xxxx
Xylophanes cyrene (Druce, 1881)
Xylophanes damocrita (Druce, 1894) EAPZ
Xylophanes eumedon (Boisduval, 1875) EAPZ
Xylophanes g. germen (Schaus, 1890) EAPZ
Xylophanes hannemanni Closs, 1917
Xylophanes juanita Rothschild and Jordan, 1903 EAPZ
Xylophanes libya (Druce, 1878) EAPZ, PBL x x
Xylophanes loelia (Druce, 1878) EAPZ, PBL x x
Xylophanes neoptolemus (Stoll, 1780) EAPZ , PBL xxx
Xylophanes pistacina Boisduval, 1875 EAPZ
Xylophanes pluto (Fabricius, 1777) EAPZ, PBL xxx
Xylophanes porcus continentalis Rothschild and Jordan, 1903, PBL, EAPZ x x
Xylophanes pyrrhus (Rothschild and Jordan, 1906) EAPZ, PBL x
Xylophanes tersa (Linnaeus, 1771) EAPZ
Xylophanes thyelia (Linnaeus, 1758) EAPZ, PBL x x
Xylophanes titana (Druce, 1878) EAPZ
Xylophanes turbata (Edwards, 1887)
Xylophanes t. tyndarus (Boisduval, 1875) EAPZ, PBL x
Xylophanes undata Rothschild and Jordan, 1903 EAPZ, PBL x
Xylophanes zurcheri (Druce, 1894) EAPZ, PBL x
Eumorpha achemon Drury, 1773 EAPZ

22 • Insecta MundI 0205, February 2012 Miller et al.
Eumorpha anchemolus (Cramer, 1779) PBL x
Eumorpha capronneri (Boisduval, 1875)
Eumorpha fasciatus (Sulzer, 1776) EAPZ, PBL x
Eumorpha labruscae (Linnaeus, 1758) EAPZ
Eumorpha o. obliquus (Rothschild and Jordan, 1903)
Eumorpha phorbus (Cramer, 1775) EAPZ, PBL x x
Eumorpha satellitia licaon (Cramer, 1775) EAPZ, PBL x x
Eumorpha triangulum (Rothschild and Jordan, 1903) EAPZ, PBL xxx
Eumorpha typhon (Klug, 1836) EAPZ
Eumorpha v. vitis (Linnaeus, 1748) PBL, EAPZ x x x
Smerinthinae
Adhemarius dariensis (Rothschild and Jordan, 1903) EAPZ
Adhemarius donysa (Druce, 1889) EAPZ
Adhemarius g. gannascus (Stoll, 1790) EAPZ, PBL x
Adhemarius ypsilon (Rothschild and Jordan, 1903) PBL, EAPZ x x
Protambulyx strigilis (Linnaeus, 1771) EAPZ, PBL xxx
Sphinginae
Agrius cingulata (Fabricius, 1775) EAPZ, PBL x
Amphimoea walkeri (Boisduval, 1875) EAPZ
Cocytius antaeus (Drury, 1773) EAPZ
Cocytius duponchel (Poey, 1832) EAPZ, PBL x
Cocytius lucifer Rothschild and Jordan, 1903 EAPZ
Manduca albiplaga (Walker, 1856) PBL x
Manduca corallina (Druce, 1883) RLMM
Manduca extrema (Gehlen, 1926) PBL x
Manduca orestan (Stoll, 1782) PBL, EAPZ x x
Manduca incisa (Walker, 1856) EAPZ
Manduca lanuginosa (Edwards, 1887) EAPZ
Manduca lefeburei (Guèrin-Mèneville, 1844) EAPZ, PBL x
Manduca muscosa (Rothschild and Jordan, 1903) EAPZ
Manduca occulta (Rothschild and Jordan, 1903) PBL, EAPZ x x
Manduca ochus (Klug, 1836) EAPZ, PBL x
Manduca pellenia (Herrich-Schäffer, 1854) EAPZ, PBL x
Manduca r. rustica (Fabricius, 1775) EAPZ, PBL x x
Manduca sexta (Linnaeus, 1763) EAPZ
Manduca wellingi Brou, 1984
Neococytius cluentius (Cramer, 1775) EAPZ, PBL x
Sphinx leucophaeata Clemens, 1859 EAPZ
Sphinx merops (Linnaeus, 1758) EAPZ
Sphinx praelongus (Rothschild and Jordan, 1903) EAPZ
--- HEDYLOIDEA ---
HEDYLIDAE
Macrosoma bahiata (Felder and Rogenhofer, 1875) PBL x
Macrosoma cascaria (Schaus, 1901) PBL x x
Macrosoma conifera (Warren, 1897) PBL x x x
Macrosoma hyacinthina (Warren, 1905) PBL x
Macrosoma lucivittata (Walker,1863) EAPZ
Macrosoma rubedinaria (Walker, 1862) PBL x x x
Macrosoma semiermis (Prout, 1932) PBL x x

Insecta MundI 0205, February 2012 • 23Lepidoptera of Honduras
--- GEOMETROIDEA ---
SEMATURIDAE
Sematura lunus (Linnaeus, 1758) EAPZ, PBL x
Coronidia canace (Hopffer, 1856) EAPZ
URANIIDAE
Uraniinae
Urania fulgens (Walker, 1854) EAPZ
Epipleminae
Aniplecta sp. PBL x
Epiplema sp. PBL x
Philagraula slossoniae Hulst, 1896 PBL x
GEOMETRIDAE
Oenochrominae
Ametris nitocris (Cramer, 1780) PBL x
Macrotes commatica Prout, 1916 PBL x x x
Ergavia carinenta (Cramer, 1777)
Ergavia merops (Cramer, 1775) PBL x
Geometrinae
Rhodochlora brunneipalpis Warren, 1894 PBL x
Nemoria aturia (Druce, 1892)
Nemoria nr. karlae Pitkin, 1993 PBL x
Nemoria nr. parcipunta (Dognin, 1908) PBL x
Nemoria nr. priscillae Pitkin, 1993 PBL x
Nemoria scriptaria (Hübner, 1823) PBL x x x
Nemoria venezuelae (Prout, 1932) PBL x
Nemoria punctilinea (Dognin, 1902) PBL x x x
Lissochlora magnostigma (Dyar, 1912) PBL x
Synchlora expulsata (Walker, 1861) PBL x x x x
Synchlora gerularia (Hübner, [1823]) PBLxxxxx
Synchlora pulchrimbria Warren, 1907 PBL x x x x x
Synchlora venustula (Dognin, 1910) PBL x
Pyrochlora rhanis (Cramer, 1777) PBL x x
Tachyphyle sp. nr. acuta Butler, 1881 PBL x
Phrudocentra janeira (Schaus, 1897) PBL xxx
Phrudocentra pupillata Warren, 1897 PBL x
Neagathia corruptata (Felder and Rogenhofer, 1875) PBL x
Dichorda consequaria (Hy. Edwards, 1884)
Oospila concinna Warren, 1900 PBL x
Oospila rosipara Warren, 1897 PBL x x
Oospila venezuelata (Walker, 1861) PBL x
Xerochlora sp. nr. masonaria (Schaus, 1897) PBL x
Chlorochlamys sp. KC x
Chloropteryx dealbata (Warren, 1909) PBL, KC x x x x
Chloropteryx diluta (Dognin, 1911) PBL x x
Chloropteryx opalaria (Guenée, 1858) PBL x x
Chloropteryx sp. KC x
Sterrhiinae
Dithecodes distracta (Walker, 1861) PBL x
Dithecodes sp. PBL x

24 • Insecta MundI 0205, February 2012 Miller et al.
Tricentra sp. nr. fulvifera Dognin, 1908 PBL x
Tricentra quadrigata (Felder, 1875) PBL x
Tricentra sp. 1 PBL x x x x x
Tricentra sp. 2 PBL x
Semaeopus deexa (Warren, 1900)
Semaeopus caecaria (Hübner, 1825) PBL x x
Semaeopus varia (Warren, 1895) PBL x
Semaeopus sp. PBL x
Trygodes musivaria (Herrich-Schäffer, 1855) PBL xxx
Cyclophora anablemma (Prout, 1938) PBL x
Pleuroprucha sp. nr. ochrea (Warren, 1897) PBL, KC x x x
Haemalea imitans (Dognin, 1900) PBL x x
Leptostales crossii (Hulst, 1900) PBL x
Leptostales sp. PBL x x
Scopula apparitaria (Walker, 1861) TL
Scopula confertaria (Walker, 1861) TL
Scopula privata (Walker, 1861) PBL x
Scopula subquadrata (Guenée, 1858)
Scopula suffundaria (Walker, 1861) TL
Scopula umbilicata (Fabricius, 1794)
Scopula sp. PBL x
Tricentrogyna violescens (Schaus, 1901) PBL xxxxx
Tricentrogyna sp. nr. deportata (Walker, 1861) PBL xxx
Tricentrogyna sp. 1 PBL x x
Tricentrogyna sp. 2 PBL x
Lobocleta gurinata (Guenée, 1858)
Idaea complexaria (Schaus, 1901) PBL x x
Idaea contractalis (Walker, 1866) TL
Idaea sp. nr. elegentaria (Herrich-Schäffer, 1854) PBL x
Idaea helleria (Schaus, 1913) PBL x
Idaea latiferaria Walker, 1861 TL
Idaea sp. nr. pervertipennis (Hulst, 1900) PBL x
Idaea spernata (Walker, 1861)
Idaea subfervens (Prout, 1920) PBL x x
Idaea tacturata (Walker, 1861) PBL xxx
Idaea sp. PBL x x x x
Smicropus intercepta (Walker, 1854) TL
Smicropus laeta (Walker, [1865]) CURLA, PBL x x
Larentiinae
Euphyia cinerascens (Dognin, 1900) PBL x
Euphyia sp. PBL x
Disclisioprocta stellata (Guenée, 1858) PBL x
Obila occosaria (Walker, 1866) TLS
Obila pannosata (Guenée, 1858)
Obila sp. PBL x
Rheumaptera sp. PBL x
Eutrepsia dispar (Walker, 1854) TLS
Oligopluera malachitaria Herrich-Schäffer, 1855 PBL x x x
Oligopluera aulaeata Felder and Rogenhoffer, 1875 PBL x

Insecta MundI 0205, February 2012 • 25Lepidoptera of Honduras
Eubaphe conformis (Walker, 1854) PBL x
Eubaphe sp. PBL x
Eudulophasia invaria (Walker, 1854) PBL x x
Psaliodes sp. PBL x
Eupithecia spilosata (Walker, 1863) TL
Eupithecia sp. 1 PBL x x
Eupithecia sp. 2 PBL x
Eois sp. nr. binaria (Guenée [1858]) PBL x
Eois catana Druce, 1892 PBL x x
Eois nr. dorisaria (Schaus, 1913) PBL x
Eois heliadaria (Guenée, 1858)
Eois insignata (Walker, 1861) PBL x x
Eois expressaria (Walker, 1861) TL
Eois sp. 1-4 PBL x x x
Ennominae
Thysanopyga amarantha Debauche, 1937 PBL x x x x x
Thysanopyga apitruncaria Herrich-Schäffer, 1856 PBL x
Thysanopyga carnia (Druce, 1893) PBL x
Thysanopyga pygaria (Guenée, 1858) PBL x
Parilexia cermala (Druce, 1893) PBL xxxxx
Perigramma repitita Warren, 1905 PBL x
Perigramma sp. nr. guatemalaria Schaus, 1927 PBL x x x
Dyschoroneura obsolescens Warren, 1894 PBL x
Leucula abilinearia Guenée, 1858
Simena luctifera (Walker, 1856) TLS
Periclina syctaria (Walker, 1860)
Cyclomia minuta (Druce, 1892) PBL x
Cyclomia disparlis Schaus, 1911 PBL x x
Eusarca asteria (Druce, 1892) PBL, TLS x
Eusarca minucia (Druce, 1892) PBL x
Eusarca nemoria (Druce, 1892) PBL x
Polla celeraria (Walker, 1860) PBL x x x x
Cimicodes albicosta Dognin, 1913 PBL x
Pyrinia itunaria Walker, 1860 PBL x
Pyrinia sanitaria Schaus 1901 PBL x x x x x
Pyrinia sp. PBL x x
Patalene aenetusaria (Walker, 1860) PBL xxxxx
Patalene nr. asychisaria (Walker, 1860) PBL x
Patalene hamulata (Guenée, 1858)
Patalene luciata (Stoll, 1790) PBL x
Patalene trogonaria (Herrich-Schäffer, 1856) PBL x
Patalene sp. PBL x
Trotopera arrhapa (Druce, 1891) PBL x
Euclysia dentifasciata Dognin, 1910 PBL x x x
Euclysia sp. PBL x x
Synnomos rmamentaria Guenée, 1858 MGCL
Sericoptera mahometaria (Herrich-Schäffer, 1856) PBL xxx
Nepheloleuca politia (Cramer, 1777) PBL x
Nepheloleuca oridata (Grote, 1883) EAPZ

26 • Insecta MundI 0205, February 2012 Miller et al.
Paragonia cruraria (Herrich- Schäffer, 1854) PBL x x x x
Prochoerodes striata (Stoll, 1798) PBL x
Prochoerodes sp. KC x
Oxydia apidania (Cramer, 1779) PBL x x x
Oxydia vesulia (Cramer, 1779) or nr. PBLxxxxx
Microgonia rufaria Warren, 1901 PBL x x x x x
Herbita lilacina (Warren, 1897) PBL xxx
Pero afuera Poole, 1987 PBL x
Pero nr. anceta (Stoll, 1781) PBL x
Pero astapa (Druce, 1892)
Pero aurunca (Druce, 1892)
Pero clysiaria (Felder and Rogenhoffer, 1875) PBL x
Pero coronata Warren, 1904 PBL x x
Pero delauta (Warren, 1907)
Pero dularia Poole, 1987
Pero fragila Poole, 1987
Pero fusaria (Walker, 1860)
Pero incisa Dognin, 1889 PBL x x x
Pero lessema (Schaus, 1901)
Pero melissa (Druce, 1892)
Pero polygonaria (Herrich-Schäffer, 1855) PBL x x
Pero probata Poole, 1987 PBL x
Pero pumaria Felder and Rogenhoffer, 1875 PBL x x x x
Pero saturata (Walker, 1868) PBL x
Pero spina Poole, 1897
Pero verda Poole, 1987
Sphacelodes vulneraria (Hübner, 1823) PBL, EAPZ, KC xxxx
Nematocampa completa Warren, 1904 PBL x x x x
Nematocampa sp. PBL x
Erastria decrepitaria (Hübner, 1823) PBL xxx
Semiothisa carinaria Dognin, 1924 PBL x
Semiothisa divergentata (Snellen, 1894) PBL x x
Macaria aequiferaria Walker, 1861 TLS
Macaria nr. ferina (Dognin, 1924) PBL x
Macaria gambarina (Stoll, [1781]) PBL xxxxx
Macaria nr. guapilaris (Schaus, 1911) PBL x
Macaria nr. lydia (Scahus, 1912) PBL x
Macaria regulata (Fabricius, 1775) PBL x
Macaria subfulva (Warren, 1906) PBL x
Macaria sp. 1-3 PBL x x x x x
Psamatodes abydata (Guenée, [1858]) PBL, KC x x x x
Tornos pusillus Rindge, 1954 PBL x
Epimecis detexta (Walker, 1860) PBL x
Epimecis fraternaria (Guenée, 1858)
Epimecis matronaria (Guenée, 1858) PBL x
Epimecis patronaria (Walker, 1860)
Epimecis subroraria Walker, 1860 PBL x
Epimecis sp. PBL x x
Glena agria Rindge, 1967 PBL x x x x

Insecta MundI 0205, February 2012 • 27Lepidoptera of Honduras
Hymenomima camerata Warren, 1900 PBL x x
Hymenomima sp. 1 PBL x x
Hymenomima sp. 2 PBL x
Physocleora sp. PBL x
Iridopsis nr. aviceps Prout, 1932 PBL x
Iridopsis herse (Schaus, 1912) PBL x
Iridopsis validaria (Guenée, 1858) PBL xxx
Iridopsis pandarosos (Schaus, 1912) PBL xxxxx
Iridopsis oberthuri Prout, 1932 PBL x
Melanolophia exilinea (Warren, 1906)
Melanolophia sadrina (Schaus, 1901) PBL x
Melanolophia sp. PBL x
Phyle schausaria (Hy. Edwards, 1884)
Pantherodes pardalaria Hübner, 1823 EAPZ
Pantherodes sp. PBL x x
Melanchroia chephise (Stoll, 1782)
Phrygionis divitaria (Oberthür, 1916) PBL x x
Phrygionis pallicosta (Felder and Rogenhofer, 1875) PBL x
Phrygionis polita (Cramer, 1780)
Phrygionis privignaria (Guenée [1858]) PBL x x x
Pityeja histrionaria (Herrich-Schäffer, 1853) MGCL
Opisthoxia bella (Butler, 1881) TL
Opisthoxia sp. nr. cluana (Druce, 1900) PBL x x
Opisthoxia uncinata (Schaus, 1912) PBL x
Leuciris mbriaria (Stoll, [1781]) PBLxxxxx
--- NOCTUOIDEA ---
NOTODONTIDAE
Notodontinae
Cerura rarata Walker, 1865 PBL x x x x
Dudusinae
Crinodes besckei (Hübner, 1824) PBL x x
Crinodes guatemalena Druce, 1887 EAPZ, PBL, KC x x x
Crinodes schausi Rothschild, 1917 EAPZ
Crinodes striolata Schaus, 1901 PBL x x x
Hemiceratinae
Apela neobule Druce, 1905 or nr. PBL x x
Hemiceras alba Walker, 1865 TL
Hemiceras bilinea Schaus EAPZ
Hemiceras spp. 1-16 PBL x x x x
Heterocampinae
Antaea omana Schaus, 1906 PBL x x
Chilara cresus (Cramer, 1777) PBL x
Colax phocus Schaus, 1892 PBL x x x
Colax sp. PBL x
Contrebia extrema Walker, 1856 TL
Disphragis livida Schaus, 1911 PBL x x x
Disphragis remuria Druce, 1898 PBL x x
Disphragis tharis (Stoll, [1780]) PBL x x
Drugera morona Druce, 1898 or nr. PBL x x x

28 • Insecta MundI 0205, February 2012 Miller et al.
Farigia sp. nr. magniplaga Schaus, 1906 PBL x
Farigia sp. PBL x
Hapigia annulata Schaus, 1905 PBL x x
Hapigia repandens Schaus, 1906 PBL x x
Hapigia simplex (Walker, 1865) PBL x x x
Hapigia sp. 1 PBL x x
Hapigia sp. 2 PBL x
Magava multilinea Walker, 1865 PBL x
Naprepa sp. PBL x
Rhuda difcilis Schaus, 1911 PBL x x x
Rhuda tuisa Schaus, 1911 PBL x x
Rifargia gelduba Schaus, 1892 PBL x
Rifargia sp. 1 PBL x x
Rifargia sp. 2 PBL x
Rifargia sp. 3 PBL x
Rosema apicalis Walker, 1855
Rosema epigena (Stoll, 1790) PBL x x
Rosema thestia Druce, 1898 PBL x
Sericochroa arecosa Druce 1898 PBL x x x x x
Trichomaplata sp. PBL x x x x
Nystaleinae
Bardaxima meyeri Schaus, 1928 PBL x
Calledema argenta Schaus, 1905 PBL x x
Calledema jocasta (Schaus, 1901) PBL x
Didugua argentilinea Druce, 1891 EAPZ, PBL x
Didugua violascens Herrich-Schäffer EAPZ
Elasmia astuta Schaus, 1894 PBL x x
Lepasta bractea (Felder, 1874) PBL x
Lepasta grammodes Felder, 1874 PBL x x
Marthula cynrica Schaus, 1928 PBL x
Notoplusia marchiana Schaus, 1928 PBL x
Nystalea clotho Thiaucourt, 2002
Nystalea malga Schaus, 1904 PBL x x
Nystalea unguicularis Thiaucourt, 2002
Nystalea superciliosa Guenée, 1852 PBL x x
Pentobesa maya Thiaucourt, 2008
Pentobesa xylinoides (Walker, 1866) PBL x
Strophocerus albonotatus Druce, 1909 PBL x x
Symmerista meridionalis Thaiucourt, 2007
Dioptinae
Brachyglene crocearia (Schaus), 1912
Dioptis butes (Druce, 1885) USNM
Dioptis eteocles (Druce, 1885)
Ephialtias pseudena (Boisduval, 1870)
Getta turrenti Miller, 2009
Isostyla ithomeina (Butler, 1872)
Josia enoides (Boisduval, 1870)
Josia frigida Druce, 1885
Josia fusigera Walker, 1864

Insecta MundI 0205, February 2012 • 29Lepidoptera of Honduras
Josia integra Walker, 1854
Josia hyperia (Walker, 1854)
Josia megaera (Fabricius, 1787) TLS
Josia sp. CURLA x
Lyces ariaca (Druce, 1885) USNM
Lyces tamara (Hering, 1925)
Oricia truncata Walker, 1854 TL, PBL, CURLA, USNM x x x
Phanoptis cyanomelas C. and R. Felder, 1874 or nr. PBL, CURLA x x
Phaeochlaena gyon (Fabricius, 1787) or nr. PBL x x
Phaeochlaena tendinosa Hübner, 1818
Scotura annulata (Guérin-Méneville), 1844
Notodontidae (incertae sedis)
Lirimiris guatemalensis Rothchild, 1917 PBL x x x
Lirimiris lignitecta Walker, 1865 or nr. PBL x
Lirimiris meridionalis Schaus, 1904 PBL x
Pentobesa lignicolor Möschler, 1877 PBL x x
EREBIDAE
Lymantriinae
Caviria sp. PBL x x x x x
Eloria subnuda Walker, 1855 TL
Terphothrix tibialis (Walker, 1855) TL
Arctiinae
Lithosiini
Areva trigemmis (Hübner, 1827) PBL x x x
Ardonea morio Walker, 1854
Tuina cingulata (Walker, 1854) TL
Apistosia judas Hübner, [1819]
Eudesmia laetifera Walker, 1864 EAPZ
Eudesmia menea (Drury, 1782) PBL, TLS x
Euthyone theodula (Schaus, 1924) PBL x
Euthyone grisescens (Schaus 1911) or nr. PBL xxx
Euthyone simplex (Walker, 1854) TL
Cisthene polyzona Druce, 1885 PBL x x x x
Mulona phelina (Druce, 1885) PBL xxxxx
Epeiromulona roseata Field, 1952
Diarhabdosia laudamia (Druce, 1885) PBL x x
Eurylomia cordula (Boisduval, 1870)
Talara synnephela Dyar, 1916 PBL x x
Lycomorphodes sordida (Butler, 1877) PBL x x
Lycomorphodes sp. PBL x x x
Balbura fasciata Schaus, 1911 PBL x x x
Balbura intervenata Schaus, 1911 PBL x
Balbura sp. nr. dorsisigna Walker, 1854 PBL x x
Agylla pogonoda Hampson, 1900 PBL x x x
Agylla spp. 1-4 PBL x x x x x
?Metalobosia sp. nr. ducalis Schaus, 1911 PBL x
Arctiini
Arctiina
Virbia birchi (Druce, 1911) or nr. PBL x x x x

30 • Insecta MundI 0205, February 2012 Miller et al.
Virbia sp. PBL x x
Notarctia proxima (Guérin- Méneville, 1844) EAPZ
Estigmene acrea Drury PBL
Hypercompe leucarctiodes (Grote and Robinson, 1967) PBL xxxxx
Hypercompe perplexa Schaus, 1911 or nr. PBL x x x
Hypercompe caudata (Walker, 1855)
Arachnis dilecta (Boisduval, 1870) TL
Arachnis aulaea Geyer, 1837
Callimorphina
Utetheisa ornatrix (Linnaeus, 1758) PBL, TLS xxx
Pericopina
Calodesma maculifrons (Walker, 1865) TL, PBL, USNM xxx
Hypocrita albimaculata (Druce, 1897) PBL, USNM x x
Hypocrita ambigua (Hering, 1925) TL
Hypocrita arcaei (Druce, 1884) PBL, USNM x x x
Hypocrita excellens (Walker, 1854) EAPZA1
Hypocrita pylotis (Drury, 1773)
Composia delissima Herrich-Schäffer, 1866
Chetone histrio Boisduval, 1870 TLA2
Chetone ithomia (Boisduval, 1870) USNM
Chetone angulosa (Walker, 1854) USNM
Xenosoma aviceps (Walker, 1865) USNM
Hyalurga sora (Boisduval, 1870) PBL, CURLA, USNMx x
Hyalurga urioides Schaus, 1910 PBL, USNM x
Heliactinidia sitia Schaus, 1910 USNM
Dysschema lycaste (Klug, 1836) EAPZ, TL form, USNM
Dysschema sp. PBL x
Dysschema aorsa (Boisduval, 1870) USNM
Dysschema cerealis (Druce, 1884) USNM
Dysschema leucophaea (Walker, 1854) TLS, PBL, USNM x x x
Dysschema magdala (Boisduval, 1870) USNM, PBL x
Dysschema mariamne (Geyer, 1836) USNM
Phaegopterina
Parathyris cedonulli (Stoll, 1781) PBL x
Robinsonia deiopea Druce, 1895 PBL x
Robinsonia sabata Druce, 1895
Idalus dares Druce, 1894 PBL x x
Idalus crinis Druce, 1884 PBL x
Idalus alteria Schaus, 1905 PBL x
Idalus herois Schaus, 1889 PBL xxxx
Idalus vitrea (Cramer, 1780) PBL x
Phaeomolis lineatus Druce, 1884 or nr. PBL x x x x x
Symphlebia lophocampoides Felder, 1874 EAPZ
Symphlebia underwoodi (Rothschild, 1910) EAPZ
Amaxia apyga Hampson, 1901 PBL x x
Amaxia pardalis Walker, 1855 PBL x x
Amaxia beata Dognin (1909) PBL x
Amaxia sp. KC x
Evius hippia (Stoll, [1790]) PBL xxxx

Insecta MundI 0205, February 2012 • 31Lepidoptera of Honduras
Neritos samos Druce, 1896
Trichromia atta (Schaus, 1920) PBL x x x x
Trichromia sp. nr. cotes (Druce, 1896) PBL x x
Trichromia avoroseus (Walker, 1855) TL
Trichromia sp. PBL x
Parevia gurma Schaus, 1920 PBL x x
Rhipha persimilis (Rothschild, 1909) EAPZ
Rhipha chionoplaga (Dognin, 1913) EAPZ, PBL x
Cratoplastis diluta Felder, 1874 PBL, EAPZ x x x x
Gorgonidia buckleyi Druce, 1883 PBL x x x
Ormetica ataenia (Schaus, 1910) PBL x x x
Ormetica taeniata (Guérin-Méneville, [1844])
Ormetica sicilia Druce, 1884 PBL x x
Ormetica guapisa (Schaus, 1910) or nr. PBL x
Ormetica goloma (Schaus, 1920) PBL x
Ormetica orbona (Schaus, 1889) PBL x
Selenarctia elissa (Schaus, 1892) PBL x
Viviennea tegyra (Druce, 1896) PBL x x x
Castrica phalaenoides (Drury, 1773) PBL x x x
Cissura plumbea Hampson, 1901 PBL x x x x x
Melese amastris (Druce, 1884) PBL x x
Melese asana (Druce, 1884) PBL x x x
Melese avimaculata Dognin, 1899 PBL x x x
Pelochyta misera Schaus, 1911 PBL x
Elysius conspersus Walker, 1855 PBL x
Munona iridescens Schaus, 1894 PBL x x
Anaxita drucei Rodriguez 1893 RLMM
Anaxita decorata Walker, 1855
Halysidota excellens Walker EAPZ
Halysidota pectenella Watson, 1980 PBL x x x
Halysidota grandis (Rothschild, 1909) PBL xxx
Lophocampa annulata Walker EAPZ
Lophocampa citrina (Sepp, [1843]) or nr. PBL xxxxx
Lophocampa catenulata (Hübner, [1812]) or nr. PBL x x x
Lophocampa alternata (Grote, 1867) TLS
Lophocampa seruba (Herrich-Schäffer, [1855])
Pseudohemihyalea fallaciosa (Toulgoët, 1997)
Pseudohemihyalea labeculoides de Toulgoet, 1995
Pseudohemihyalea porioni de Toulgoet, 1995
Euchaetes antica (Walker, 1856) TLS
Pareuchaetes insulata (Walker, 1855) KC, PBL x x x x
Belemnia inaurata Sulzer, 1776 TLS, PBL, EAPZ x x
Euchromiina
Homoeocera gigantea (Druce, 1884) EAPZ
Sphecosoma felderi (Druce, 1883)
Myrmecopsis strigosa (Druce, 1884) EAPZ
Gymnelia sp. nr. nobilis Schaus, 1911 PBL x
Phoenicoprocta sp. sanguinea (Walker, 1854) or nr. PBL x
Pheia albisigna (Walker, 1854) PBL x

32 • Insecta MundI 0205, February 2012 Miller et al.
Loxophlebia masa (Druce, 1889) PBL x x x
Loxophlebia geminata Schaus, 1905 PBL x
Mesothen pyrrha (Schaus, 1889) PBL x
Xanthyda xanthosticta (Hampson, 1898) PBL x
Cosmosoma festiva Walker, 1854 TL
Cosmosoma braconoides (Walker, 1854) TL
Cosmosoma intensa (Walker, 1854) TL
Cosmosoma ruatana (Druce, 1888) TL
Cosmosoma sectinota Hampson, 1898 PBL x x x
Cosmosoma stibosticta (Butler, 1876) PBL xxxxx
Cosmosoma hercyna (Druce, 1884) PBL x
Cosmosoma sp. nr. teuthras (Walker, 1854) PBL x x
Cosmosoma achemon (Fabricius, 1781) PBL x x
Cosmosoma cingulatum Butler, 1876 PBL x
Pseudomya melanthoides Schaus, 1920 PBL x
Saurita temenus (Stoll, [1781]) PBLx
Saurita tipulina (Hübner, 1827) PBL x x x
Saurita myrrha Druce, 1884 PBL x
Saurita fumosa (Schaus, 1912) PBL x
Psoloptera basifulva Schaus, 1894 PBL x x
Dycladia basimacula Schaus, 1924 PBL x x
Dycladia vitrina Rothschild, 1911 PBL x x
Syntomeida melanthus (Cramer, [1779])
Macrocneme lades (Cramer, [1776])
Macrocneme chrysitis (Guérin-Méneville, 1844)
Macrocneme auripes (Walker, 1854) TL
Macrocneme cabimensis Dyar, 1914
Macrocneme adonis Druce, 1884
Macrocneme spp. incl. iole Druce, 1884 or nr. PBL x x x x x
Chrysocale principalis (Walker, [1865]) EAPZ
Poliopastea sp. PBL x x
Eunomia colombina (Fabricius, 1793)
Eunomia latenigra (Butler, 1876) TL
Horama oedippus (Boisduval, 1870) PBL x
Horama plumipes (Drury, 1773) PBL xxx
Horama panthalon (Fabricius, 1793)
Ctenuchina
Ctenucha venosa Walker, 1854
Ctenucha ruceps Walker, 1854
Dinia aeagrus (Cramer, [1779]) PBL x x x
Trichura esmeralda (Walker, 1854) TL
Trichura coarctata (Drury, 1773)
Trichura druryi Hübner, [1819]
Ecdemus obscurata Schaus, 1911 PBL x x x
Eriphioides tractipennis (Butler, 1876)
Chrysostola augusta Druce (1884) PBL x
Chrysostola moza (Druce, 1896) PBL x x
Episcepsis thetis Linnaeus 1771 or nr. PBL x
Episcepsis inornata (Walker, 1856) or nr. PBL x x x x

Insecta MundI 0205, February 2012 • 33Lepidoptera of Honduras
Episcepsis lenaeus Cramer, 1870 or nr. PBL x x
Episcepsis venata Butler, 1877 PBL x
Antichloris viridis Druce, 1884 EAPZ
Antichloris caca Hübner, 1818
Napata walkeri (Druce, 1889) EAPZ, PBL x x
Napata leucotelus Butler, 1876 TL, PBL x
Chlorostola interrupta Walker PBL x
?Desmotricha perplexa (Rothschild, 1912) PBL x x
Cyanopepla arrogans (Walker, 1854)
?Cyanopepla jalifa (Boisduval, 1870)
Aclytia sp. PBL x x
Aclytia punctata Butler, 1876 TL, PBL x
Aclytia heber (Cramer, 1780) EAPZ, PBL xxx
Aclytia albistriga Schaus, 1910 PBL x
Aclytia reducta Rothschild, 1912 or nr. PBL x x x x
Mydromera notochloris (Boisduval, 1870) CURLA x
Agyrta dux Walker, 1854 PBL x
Ptychotrichos episcepsidis Dyar, 1914 PBL x x
Delphyre testacea (Druce, 1884) PBL x x x x
Delphyre hebes Walker, 1854 TL
Neacerea ruventris Schaus, 1894 or nr. PBL x
Heliura rhodophila (Walker, 1856) PBL, TLS x x x
Heliura tetragramma (Walker, 1854)
Eucereon atriguttum Druce, 1905 EAPZ
Eucereon aurantiaca (Rothschild, 1909) PBL x
Eucereon latifascia Walker, 1856 PBL x
Eucereon pilatii Walker, 1854 TL
Eucereon varium (Walker, 1854) PBL x x
Eucereon aroa Schaus, 1894 PBL x
Eucereon rosadora Dyar, 1910 PBL x x
Eucereon rosa (Walker, 1854) TL
Eucereon balium Hampson, 1898 TL, PBL x x
Eucereon maja Druce, 1884 PBL x
Eucereon obscurum Möschler, 1872 PBL x
Eucereon pseudarchias Hampson, 1898
Correbidia elegans (Druce, 1884) PBL x x x
Correbidia germana Rothschild, 1912 or nr. PBL x x
Correbidia undulata (Druce, 1884) PBL xxxx
Correbidia costinotata Schaus, 1911 or nr. PBL x x
Hyaleucerea vulnerata Butler, 1875 PBL x x x
Hyaleucerea gigantea (Druce, 1884) PBLx
Herminiinae
Macrochilo andaca (Druce, 1891) PBL x
Macrochilo sp. PBL x
Bleptina hydrillalis Guenée, 1854 TLS
Lascoria sp. PBL x x
Palthis sp. nr. bizialis (Walker, 1859) PBL x x
Palthis orasiusalis Walker, 1859 TL

34 • Insecta MundI 0205, February 2012 Miller et al.
Hypeninae
Hypena perialis Schaus, 1904 PBL x
Hypena porrectalis Fabricius, 1794 TLS
Hypena subidalis Guenee, 1854
Hypena sp. PBL x
Trauaxa obliqualis Walker, 1866 PBL x x x
Scoliopteryginae
Anomis hemiscopsis Dyar, 1914 PBL x x
Anomis illita Guenée, 1852 TLS
Anomis editrix Guenée, 1852 TLS, PBL x
Anomis sublineata (Walker, 1869) TL
Alabama argillacea (Hübner, 1823) EAPZ
Calpinae
Ceroctena amynta (Stoll, 1782) PBL x x x x
Sosxetra grata Walker, 1862 PBL x x x x x
Lephana sp. nr. tetraphorella Walker, 1866 PBL x x
Lephana sp. PBL x
Gonodonta biarmata Guenée, 1852 PBL x x x
Gonodonta bidens Geyer, 1832 PBL x x x
Gonodonta nitidimacula Guenée, 1852 PBL x x
Gonodonta parens Guenée, 1852 PBL x
Gonodonta pyrgo (Cramer, 1777) PBL x x x
Gonodonta sinaldus Guenée, 1852 PBL xxxxx
Gonodonta uxor (Cramer, 1780) PBL x
Gonodonta syrna Guenée, 1852 PBL x x
Gonodonta indentata (Hampson, 1926) PBL x
Gonodonta immacula Guenée, 1852
Gonodonta sp. PBL x
Graphigona regina (Guenée, 1852) PBL x
Oraesia nobilis Felder and Rogenhofer, 1874 PBL x x
Obroatis distincta Butler, 1879 PBL x x
Plusiodonta clavifera (Walker, 1868) TL, PBL x x
Herminodes atrosignata (Walker, 1858) PBL xxxx
Eudocima apta (Walker, [1858]) EAPZ
Eudocima procus (Cramer, 1777) EAPZ
Dyops chromatophila Walker, 1858 PBL x
Dyops dotata (Walker, 1869) TL
Pseudbarydia severa (Druce, 1889) PBL x
Pararcte schneideriana Stoll, 1780 PBL x x
Mazacyla relata (Walker, 1858) PBL x
Lepidodes gallopavo Walker, 1856 PBL x x x
Gorgone fellearis Hübner, 1823 PBL x
Tautobriga glaucopis (Hampson, 1926) PBL x
Tautobriga euspila (Walker, 1869) TL, PBL x
Oxidercia thaumantis Hampson, 1926 PBL x x
Oxidercia toxea (Stoll, 1782) or nr. PBL xxx
Incertae sedis (Calpinae)
Arabriga bimaculata Walker, 1869 TL
Azatha retardens Walker, 1858 TL

Insecta MundI 0205, February 2012 • 35Lepidoptera of Honduras
Briarda semiplaga Walker, 1859 TL
Canatha subangulalis (Walker, [1866]) TL
Leucotela nigripalpis (Walker, [1866]) TL
Peteroma discisigna (Walker, [1858]) TL
Hypocalinae
Hypocala andremona (Stoll, [1781]) PBL, TLS x
Goniapteryx servia (Cramer, 1782) PBL x
Eublemminae
Eublemma cinnamomea (Herrich-Schäffer, 1868) PBL x x
Phytometrinae
Mursa phtisialis Guenée, 1854 TLS, PBL x x
Phytometra acillalis (Walker, 1859) TL
Cecharismena darconis (Schaus, 1913) PBL x x
Radara aquilalis Schaus, 1916 or nr. PBL x x x
Radara nezeila (Schaus, 1906) PBL x x x
Radara nealcesalis (Walker, 1859) PBL x x
Radara zoum (Dyar, 1914) PBL x
Radara sp. PBL x x
Isogona natatrix Guenée, 1852 TLS
Anobinae
Anoba pohli (Felder and Rogenhofer, 1874) PBL x x
Baniana ostia Druce, 1898 PBL x
Baniana suggesta (Walker, 1858) TL
Baniana ypita Schaus, 1901 PBL x
Erebinae
Thermesiini
Hemeroblemma opigena (Drury, 1773) PBL, TLS x x
Hemeroblemma acron (Cramer, 1779) PBL x x
Hemeroblemma dolosa Hübner, [1823] PBL x
Hemeroblemma mexicana (Guenee, 1852)
Hemeroblemma sp. PBL x
Thermesia sobria Walker, 1865 TL
Letis buteo Guenée, 1852 PBL xxxxx
Letis herilia (Stoll, 1780) PBL x x
Letis mycerina (Cramer, 1777)
Thysania agrippina (Cramer, 1776) PBL, EAPZ x
Ascalapha odorata (Linnaeus, 1758) PBL, EAPZ x x
Catocalini
Argidia sp. nr. hypoxantha (Hampson, 1926) PBL x
Hemicephalis agenoria (Druce, 1890) PBL x
Ramphia albizona (Latreille, 1817) PBL x x
Hypogrammodes balma Guenée, 1852 PBL x x
Metipotini
Melipotis perpendicularis (Guenée, 1852) EAPZ, TLS
Melipotis fasciolaris (Hübner, 1825) PBL x
Melipotis descreta (Walker, [1858]) TL
Melipotis famelica (Guenée, 1852) TLS, PBL x
Melipotis nigrobasis (Guenée, 1852) PBL x
Melipotis sp. PBL x

36 • Insecta MundI 0205, February 2012 Miller et al.
Orodesma schausi (Druce, 1890) PBL x
Euclidiini
Ptichodis bistrigata Hübner, 1818
Celiptera sp. PBL x x x
Mocis latipes (Guenée, 1852) EAPZ, PBL x x x
Mocis marcida (Guenée, 1852) EAPZ
Mocis texana (Morrison, 1875) EAPZ
Mocis difuens (Guenée, 1852) TLS
Mocis disseverans Walker, 1858 TLS
Mocis dyndima (Stoll, 1782) PBL x
Poaphilini
Dysgonia expediens (Walker, 1858) PBL xxx
Dysgonia purpurata Kaye, 1901 PBL x x
Ophiusini
Lesmone formularis (Geyer, 1837) or nr. PBL x
Epidromia tinctifera (Walker, 1858) TL
Epidromia sp. pannosa Guenée, 1852 or nr. KC x
Itomia lignaris Hubner, 1823
Itomia multilinea Walker, 1858
Itomia opistographa Guenée, 1852 or nr. PBL x
Helia celita (Schaus, 1912) or nr. PBL x x
Helia argentipes (Walker, 1869) TL
Selenisa sueroides (Guenée, 1852) PBL x x x x
Euclystis insana (Guenée, 1852) PBL, TLS x
Euclystis plusiodes (Walker, 1858) PBL x x
Coenipeta bibitrix (Hübner, 1823) EAPZ, PBL x
Coenipeta hemiplagia Felder and Rogenhoffer, 1874 EAPZ
Coenipeta damonia (Stoll, 1782) or nr. PBL x
Coenipeta tanais (Cramer,1776) PBL x
Coenipeta sp. nr. medalba Schaus 1906 PBL x
Coenipeta phasis Cramer, 1977 or nr. PBL x
Metria endopolia (Dyar, 1917) or nr. PBL x
Metria aperta (Walker, [1858]) PBL x
Metria simplicior (Walker, [1858]) TL
Metria spp. 1-4 PBL x x x
Kakopoda progenies (Guenée, 1852) TLS
Toxonprucha diffundens Walker, 1858 TLS
Homoptera brevipennis Walker, 1869 TL
Ophisma minna Guenée, 1852 PBL x
Ophisma tropicalis Guenée, 1852 PBL xxx
Achaea lienardi Boisduval, 1833 PBL x
Amolita sp. 1 PBL x
Amolita sp. 2 PBL x x
Incertae sedis (Erebinae)
Calyptis semicuprea (Walker, 1857) PBL x x
Catephiodes trinidadensis (Kaye, 1901) PBL x
Coenobela paucula (Walker, 1858) TL
Nymbis iniqua Guenée, 1852 TLS
Perasia helvina (Guenée, 1852)

Insecta MundI 0205, February 2012 • 37Lepidoptera of Honduras
Ptichodes basilans (Guenée, 1852)
Eulepidotinae
Eulepidotini
Eulepidotis addens (Walker, 1858) EAPZ, PBL x
Eulepidotis alabastraria Hübner, 1823 PBL x x x
Eulepidotis electa Dyar, 1914 PBL x x x
Eulepidotis carcistola Hampson, 1926 PBL x x
Eulepidotis hermura Schaus, 1898 PBL x x x
Eulepidotis inclyta Fabricius, 1997 PBL x x x
Eulepidotis juncida Guenée, 1852 TLS, PBL x
Eulepidotis modestula (Herrich-Schäffer, 1869) PBL x x
Eulepidotis perducens (Walker, 1858) PBL xxxx
Eulepidotis punctilinea Schaus, 1921 PBL x x x x
Eulepidotis testaceiceps Felder, 1874 PBL x x x
Eulepidotis suppura Dyar, 1914 EAPZ
Eulepidotis sp. PBL x
Obrima pyraloides Walker, 1865 TL
Panopodini
Anticarsia gemmatalis Hübner, 1818 EAPZ, TLS, PBL x
Anticarsia anisospila (Walker, 1869) TL
Azeta rhodogaster Guenée, 1852 EAPZ, PBL xxxxx
Azeta ceramina Hübner, [1821] TLS
Azeta signans Walker EAPZ
Athyrma adjutrix auth. not (Cramer, 1780) PBL x
Eulepidotinae (incertae sedis)
Antiblemma amarga (Schaus, 1911) PBL x x
Antiblemma hamilcar (Schaus, 1914) PBL x x x x
Antiblemma incarnans Felder and Rogenhoffer, 1874 PBL x x x
Antiblemma imitans (Walker, 1858) TL
Antiblemma memoranda Schaus, 1911 PBL x x x
Antiblemma neptis (Cramer, 1779) PBL x
Antiblemma runans Guenée, 1852 TLS
Antiblemma sterope (Stoll, 1780)
Antiblemma sufciens (Walker, 1858) TL
Antiblemma sp. PBL x x x x
Epitausa atriplaga (Walker, 1858) PBL x
Litoprosopus conigens (Walker, [1858]) TL
EUTELIIDAE
Euteliinae
Paectes abrostoloides Guenée, 1852 TLS
Paectes albescens Hampson, 1912 TL
Paectes fovifera Hampson, 1912 PBL x x
Paectes fuscescens (Walker, 1855) TL
Paectes glauca (Hampson, 1905) PBL x
Paectes lunodes Guenée, 1852 TL
Eutelia auratrix Walker, 1858 PBL x
Eutelia sp. PBL x
NOLIDAE
Meganola leucostola (Hampson, 1900) PBLx x x x

38 • Insecta MundI 0205, February 2012 Miller et al.
Nola turbana Schaus, 1921 or nr. PBL x
Neostictoptera nigropuncta Druce, 1900 or nr. PBL x
Iscadia argentea (Walker, 1869)
Iscadia furcifera (Walker, 1869) TLS
NOCTUIDAE
Plusiinae
Argyrogrammatini
Argyrogramma verruca (Fabricius, 1794) PBL x x
Trichoplusia ni (Hübner, [1803]) EAPZ
Chrysodeixis includens (Walker, [1858]) EAPZ
Plusiini
Autoplusia egena (Guenée, 1852) EAPZ
Bagisarinae
Bagisara patula (Druce, 1898) PBL x x x
Bagisara repanda (Fabricius, 1793) PBL x
Amyna octo Guenée, 1852 TLS
Cydosiinae
Cydosia curvinella Guenée, 1879 EAPZ, PBL x x x x
Cydosia phaedra Druce, 1897 TL
Eustrotiinae
Tripudia furcula Pogue, 2009
Marimatha botyoides (Guenée, 1852) PBL, TLS x x x
Marimatha sp. PBL x x x
Acontiinae
Acontia tetragona Walker, [1858] TL
Acidaliodes celenna Druce, 1892 PBL x
Diphtherinae
Diphthera festiva (Fabricius, 1775) EAPZ, PBL x x x
Acronictinae
Argyrosticta ditissima (Walker, [1858]) PBL xxx
Argyrosticta vauaurea (Hampson, 1908) PBL x x
Antachara rotundata Walker, 1858 PBL x x x x
Amphipyrinae
Metaponpneumata rogenhoferi Möschler, 1890 EAPZ
Cropia connecta (Smith, 1894) EAPZ
Cropia philia (Druce, 1889) PBL x
Oncocnemidinae
Neogalea sunia (Guenée, 1852)
Agaristinae
Darceta falcata (Druce, 1883) PBL x
Darceta proserpina (Stoll, [1782]) PBL x
Epithisanotia sanctijohannis Stephens, 1850 PBL x
Seirocastnia amalthea (Dalman, 1823)
Vespola caerulifera (Walker, 1867) PBL x
Condicinae
Condicini
Condica cupienta (Cramer, 1780) PBL x x
Condica mobilis Walker, [1857] TLS
Condica subaurea (Guenée, 1852)

Insecta MundI 0205, February 2012 • 39Lepidoptera of Honduras
Condica subornata (Walker, 1865) or nr. PBL x x
Condica mimica (Hampson, 1908) PBL xxxxx
Condica imitata (Druce, 1891) PBL x x
Condica sutor (Guenée, 1852) PBL x x
Condica sp. 1 PBL x x x x
Condica sp. 2 PBL x
Leuconyctini
Diastema tigris Guenée, 1852 PBL x x
Micrathetis dasarada (Druce, 1898) PBL x x
Micrathetis triplex (Walker, 1857) TL
Heliothinae
Helicoverpa zea (Boddie, 1850) EAPZ
Heliothis virescens (Fabricius, 1777) EAPZ
Eriopinae
Callopistria rivularis (Walker, [1858]) PBL x x x
Callopistria oridensis (Guenée, 1852) PBL xxxx
Noctuinae
Phosphilini
Spodoptera exigua (Hübner, [1808]) EAPZ
Spodoptera frugiperda (J.E. Smith, 1797) EAPZ
Spodoptera ornithogalli (Guenée, 1852) EAPZ
Spodoptera latifascia (Walker, 1856) EAPZ
Spodoptera dolichos (Fabricius, 1794) EAPZ
Spodoptera eridania (Cramer, 1784) EAPZ
Spodoptera albula (Walker, 1857) TL, EAPZ
Spodoptera spp. PBL x x
Elaphriini
Elaphria deltoides (Möschler, 1880) PBL x x
Elaphria agrotina (Guenée, 1852) PBL x x x
Elaphria sp. nr. perigeana (Schaus, 1911) PBL x x
Elaphria sp. 1 PBL x
Elaphria sp. 2 PBL x
Gonodes liquida (Möschler, 1886) PBL x
Dypterygiini
Neophaenis respondens (Walker, 1858)
Xylenini
Xylina subcostalis Walker, 1869 TL
Hadenini
Stauropides persimilis Hampson, 1909 PBL x x
Leucaniini
Mythimna unipuncta (Haworth, 1809) EAPZ
Leucania inconspicua Herrich-Schäffer, 1868 PBL x
Leucania multilinea Walker, 1856 TLS
Noctuini
Anicla infecta (Ochsenheimer, 1816) EAPZ, TLS
Agrotis ipsilon (Hufnagel, 1766) EAPZ
Agrotis subterranea (Fabricius, 1794) EAPZ
Stictopterinae
Nagara vitrea (Guenée, 1852) PBL x x

40 • Insecta MundI 0205, February 2012 Miller et al.
--- HESPERIOIDEA ---
HESPERIIDAE
Eudaminae
Phocides polybius lilea (Reakirt, [1867]) EAPZ, RLMM
Phocides distans licinus (Möschler, 1879)
Phocides metrodorus nigrescens E. Bell, 1938 RLMM
Phocides thermus thermus (Mabille, 1883)
Phocides p. pigmalion (Cramer, 1779)
Tarsoctenus corytus gaudialis (Hewsitson, 1876) RLMM
Phanus vitreus (Stoll, 1781)
Phanus obscurior obscurior Kaye, 1925 RLMM
Phanus marshallii (Kirby, 1880)
Udranomia kikkawai (Weeks, 1906) PBL x
Drephalys oria Evans, 1952 TL
Drephalys oriander (Hewitson, 1867)
Drephalys alcmon (Cramer, 1779)
Hyalothyrus neleus pemphigargyra (Mabille, 1888) PBL, RLMM x
Entheus crux Steinhauser, 1989 RLMM
Proteides mercurius mercurius (Fabricius, 1787)
Epargyreus exadeus cruza Evans, 1952
Epargyreus aspina Evans, 1952 PBL x x
Epargyreus spina spina Evans, 1952
Epargyreus clavicornis gaumeri Godman and Salvin, 1893 TL
Polygonus savigny savigny (Latreille, [1824]) PBL, RLMM x x
Chioides albofasciatus (Hewitson, 1867)
Chioides catillus albius Evans, 1952 RLMM
Chioides zilpa (Butler, 1872)
Aguna asander asander (Hewitson, 1867)
Aguna claxon Evans, 1952
Aguna aurunce hypozonius (Plötz, 1880) RLMM, PBL x
Aguna panama Austin and Mielke, 1998
Typhedanus undulatus (Hewitson, 1867) PBL, ER, RLMM x
Typhedanus ampyx (Godman and Salvin, 1893)
Polythrix octomaculata (Sepp, [1844]) CURLA x
Polythrix asine (Hewitson, 1867) PBL, EAPZ, RLMM
Polythrix metallescens (Mabille, 1888) EAPZ
Cephise aelius (Plötz, 1880) PBL x
Codatractus c. carlos Evans, 1952 TL, RLMM
Codatractus a. alcaeus (Hewitson, 1867)
Codatractus bryaxis (Hewitson, 1867) RLMM
Codatractus cyda (Godman, 1901) TL
Ridens mephitis (Hewitson, 1876)
Urbanus p. proteus (Linnaeus, 1758) PBL, CURLA, RLMM x
Urbanus viterboana (Ehrmann, 1907)
Urbanus belli (Hayward, 1935) PBL x x
Urbanus pronta Evans, 1952 TL
Urbanus esmeraldus (Butler, 1877) PBL x
Urbanus d. dorantes (Stoll, 1790) PBL x x x
Urbanus procne (Plötz, 1881) PBL, RLMM x

Insecta MundI 0205, February 2012 • 41Lepidoptera of Honduras
Urbanus simplicius (Stoll, 1790) PBL, RLMM xxxxx
Urbanus teleus (Hübner, 1821) PBL, CULA x x x
Urbanus tanna Evans, 1952 PBL x x x
Urbanus d. doryssus (Swainson, 1831) EAPZ, RLMM
Urbanus a. albimargo (Mabille, 1875) EAPZ, RLMM
Astraptes talus (Cramer, 1777) EAPZ, RLMM
Astraptes “fulgerator azul” (Reakirt, [1867]) Complex PBL x
Astraptes tucuti (R. Williams, 1927) RLMM
Astraptes brevicauda (Plötz, 1886)
Astraptes e. egregius (Butler, 1870)
Astraptes enotrus (Stoll, 1781)
Astraptes janeira (Schaus, 1902) PBL, RLMM x x
Astraptes alardus latia Evans, 1952 RLMM, PBL x x
Astraptes alector hopfferi (Plötz, 1881) PBL, RLMM x
Astraptes chiriquensis chiriquensis (Staudinger, 1876)
Astraptes weymeri (Plötz, 1882) PBL x
Astraptes anaphus annetta Evans, 1952 PBL x
Astraptes phalaecus (Godman and Salvin, 1893) PBL x
Narcosius parisi helen (Evans, 1952) TL
Narcosius c. colossus (Herrich-Schäffer, 1869)
Narcosius nazaraeus Steinhauser, 1986
Autochton vectilucis (Butler, 1872)
Autochton neis (Geyer, 1832) RLMM, PBL x
Autochton longipennis (Plötz, 1882) PBL, CURLA xxxx
Autochton zarex (Hübner, 1818) PBL, RLMM x
Autochton bipunctatus (Gmelin, 1790)
Achalarus a. albociliatus (Mabille, 1877) RLMM
Achalarus toxeus (Plötz, 1882) PBL x
Thessia jalapus (Plötz, 1881) PBL x
Cabares p. potrillo (Lucas, 1857) RLMM
Spathilepia clonius (Cramer, 1775) PBL xxx
Cogia c. cajeta (Herrich-Schäffer, 1869) ER x
Cogia calchas (Herrich-Schäffer, 1869) PBL, ER, CURLA, RLMM x x x
Cogia c. caicus (Herrich-Schäffer, 1869) RLMM
Telemiades avitus (Stoll, 1781)
Telemiades oiclus (Mabille, 1889)
Telemiades megallus Mabille, 1888
Telemiades des Bell, 1949
Bungalotis erythus (Cramer, 1775) RLMM, PBL x
Bungalotis midas (Cramer, 1775) PBL x
Bungalotis astylos (Cramer, 1780) PBL x x x
Bungalotis q. quadratum (Sepp, [1845]) RLMM
Nicephellus nicephorus (Hewitson, 1876)
Dyscophellus diaphorus (Magille and Boullet, 1912) PBL x
Dyscophellus phraxanor lama Evans, 1952 PBL x x
Dyscophellus ramon Evans, 1952 PBL, RLMM x
Nascus phocus (Cramer, 1777) PBL, RLMM x
Nascus solon corilla Evans, 1952
Pseudonascus paulliniae (Sepp, [1842]) EAPZ, RLMM

42 • Insecta MundI 0205, February 2012 Miller et al.
Porphyrogenes zohra (Möschler, 1879)
Porphyrogenes sula (Williams and Bell, 1940) TL
Porphyrogenes passalus (Herrich-Schäffer, 1869) PBL x
Porphyrogenes virgatus (Mabille, 1888)
Pyrginae
Myscelus amystis hages Godman and Salvin, 1893
Passova gellias (Godman and Salvin, 1893)
Pyrrhopyge zenodorus Godman and Salvin, 1893 RLMM
Chalypyge chalybea chloris (Evans, 1951) EAPZ
Yanguna cosyra (H. Druce, 1875)
Melanopyge erythrosticta (Godman and Salvin, 1879)
Elbella scylla (Ménétriés, 1855) EAPZ
Elbella p. patrobas (Hewitson, 1857)
Mysoria barcastus ambigua (Mabille and Boullet, 1908) TL
Celaenorrhinus monartus (Plötz, 1884) RLMM
Celaenorrhinus fritzgaertneri (Bailey, 1880)
Celaenorrhinus stola Evans, 1952
Celaenorrhinus c. cynapes (Hewitson, 1870)
Arteurotia t. tractipennis Butler and H. Druce, 1872
Polyctor enops (Godman and Salvin, 1894)
Nisoniades godma Evans, 1953
Nisoniades rubescens (Möschler, 1877)
Nisoniades ephora (Herrich-Schäffer, 1870) PBL x
Nisoniades macarius (Herrich-Schäffer, 1870)
Nisoniades castolus (Hewitson, 1878)
Pachyneuria licisca (Plötz, 1882) PBL, RLMM x x
Pellicia arina Evans, 1953 RLMM
Pellicia d. dimidiata Herrich-Schäffer, 1870 PBL, RLMM x
Noctuana lactifera bipuncta (Plötz, 1884)
Noctuana stator (Godman and Salvin, 1899)
Cyclosemia anastomosis Mabille, 1878 PBL x
Cyclosemia pedro R. Williams and E. Bell, 1940 TL
Gorgopas c. chlorocephala (Herrich-Schäffer, 1870) CURLA x
Bolla cylindus (Godman and Salvin, 1896)
Bolla cupreiceps (Mabille, 1891) TL, PBL, RLMM x
Bolla imbras (Godman and Salvin, 1896) [= pullata] PBL x x
Bolla clytius (Godman and Salvin, 1897) PBL x
Staphylus vulgata (Möschler, 1879) PBL x x
Staphylus ascalaphus (Staudinger, 1876) PBL x
Staphylus azteca (Scudder, 1872) TLS, PBL x x
Gorgythion begga pyralina (Möschler, 1877) PBL xxx
Gorgythion vox Evans, 1953
Sostrata nordica Evans, 1953 PBL, RLMM x x
Mylon lassia (Hewitson, 1868)
Mylon maimon (Fabricius, 1775) PBL x
Mylon pelopidas (Fabricius, 1793)
Mylon jason (Ehrmann, 1907)
Timochares t. trifasciata (Hewitson, 1868) RLMM, PBL x
Anastrus s. sempiternus (Butler and H. Druce, 1872) PBL, RLMM x x

Insecta MundI 0205, February 2012 • 43Lepidoptera of Honduras
Anastrus t. tolimus (Plötz, 1884) PBL, RLMM x
Anastrus petius peto Evans, 1953
Anastrus n. neaeris (Möschler, 1879) RLMM
Tosta platypterus (Mabille, 1895) TL
Ebrietas a. anacreon (Staudinger, 1876) PBL, EAPZ x x
Ebrietas evanidus Mabille, 1898
Helias cama Evans, 1953 PBL, RLMM xxxxx
Camptopleura theramenes Mabille, 1877
Cycloglypha t. thrasibulus (Fabricius, 1793) RLMM, PBL x
Theagenes aegides (Herrich-Schäffer, 1869) RLMM
Chiomara g. georgina (Reakirt, 1868) ER, RLMM x
Chiomara mithrax (Möschler, 1879)
Gesta invisus (Butler and H. Druce, 1872) PBL, RLMM x
Erynnis tristis tatius (W. H. Edwards, 1883)
Erynnis funeralis (Scudder and Burgess, 1870) RLMM
Aethilla lavochrea Butler, 1872
Aethilla e. echina Hewitson, 1870
Achlyodes busirus heros Ehrmann, 1909 PBL, RLMM x x
Achlyodes pallida (R. Felder, 1869) RLMM
Eantis thraso (Hübner, [1807]) PBL, CURLA, RLMM x x
Doberes anticus (Plötz, 1884) RLMM
Ouleus salvina Evans, 1953 PBL, CURLA, RLMM xxxxx
Zera h. hyacinthinus (Mabille, 1877)
Zera sp. EAPZ
Quadrus cerialis (Stoll, 1782) PBL, RLMM x x x x
Quadrus c. contubernalis (Mabille, 1883) PBL, RLMM xxx
Quadrus l. lugubris (R. Felder, 1869)
Pythonides jovianus amaryllis Staudinger, 1876 RLMM, PBL x x
Pythonides grandis assecla Mabille, 1883
Pythonides proxenus (Godman and Salvin, 1895) RLMM
Atarnes sallei (C. and R. Felder, 1867)
Paches loxus gloriosus Röber, 1925 RLMM
Carrhenes f. fuscescens (Mabille, 1891) TL
Carrhenes calidius Godman and Salvin, 1895
Carrhenes c. canescens (R. Felder, 1869) PBL x x
Xenophanes tryxus (Cramer, 1780) PBL, RLMM xxxx
Antigonus nearchus (Latreille, 1817) RLMM, PBL x x
Antigonus erosus (Hübner, [1812]) PBL, EAPZ, RLMM x
Antigonus corrosus Mabille, 1878 RLMM
Zopyrion sandace Godman and Salvin, 1896 ER x
Anisochoria bacchus Evans, 1953
Pyrgus adepta Plötz, 1884, ER, EAPZ, RLMM x
Pyrgus oileus (Linnaeus, 1767) PBL, CA, EAPZ, RLMM xxxxx
Heliopetes l. laviana (Hewitson, 1868) RLMM
Heliopetes m. macaira (Reakirt, [1867])
Heliopetes arsalte (Linnaeus, 1758) PBL, EAPZ xxxxx
Heliopetes alana (Reakirt, 1868) RLMM
Hesperiinae
Perichares adela (Hewitson, 1867) PBL, EAPZ, RLMM x

44 • Insecta MundI 0205, February 2012 Miller et al.
Perichares aurina Evans, 1955
Orses cynisca (Swainson, 1821)
Ancyloxypha arene (W. H. Edwards, 1871) EAPZ
Copaeodes minima (W. H. Edwards, 1870) ER x
Argon lota (Hewitson, 1877)
Talides sergestus (Cramer, 1775)
Talides cantra Evans, 1955
Tromba xanthura (Godman, 1901) PBL, RLMM x
Carystus p. phorcus (Cramer, 1777) RLMM
Cobalus dicula (Hewitson, 1877) RLMM
Dubiella scella belpa Evans, 1955
Damas clavus (Herrich-Schäffer, 1869) PBL x
Carystoides basoches (Latreille, [1824])
Carystoides lila Evans, 1955
Carystoides hondura Evans, 1955 TL
Lychnuchoides saptine (Godman and Salvin, 1879) RLMM
Calpodes ethlius (Stoll, 1782) RLMM, PBL x
Panoquina ocola (W. H. Edwards, 1863) RLMM
Panoquina hecebolus (Scudder, 1872) PBL, RLMM x x x
Panoquina lucas (Fabricius, 1793) EAPZ, CURLA, RLMM x
Panoquina fusina viola Evans, 1955
Panoquina p. pauper (Mabille, 1878) x
Panoquina evadnes (Stoll, 1781)
Zenis minos (Latreille, [1824])
Saliana triangularis (Kaye, 1914) RLMM, EAPZ, PBL x
Saliana e. esperi Evans, 1955
Saliana antoninus (Latreille, [1824])
Saliana longirostris (Sepp, [1840])
Saliana s. saladin Evans, 1955
Thracides phidon (Cramer, 1779)
Thracides arcalaus (Stoll, 1782)
Aides dysoni Godman, 1900 TL
Aides brino (Stoll, 1781)
Neoxeniades s. seron (Godman, 1901) TL
Neoxeniades luda (Hewitson, 1877)
Aroma aroma (Hewitson, 1867) RLMM
Falga sciras Godman, 1901 TL
Synapte pecta Evans, 1955 PBL x x x x
Synapte s. salenus (Mabille, 1893)
Anthoptus epictetus (Fabricius, 1793) PBL, CURLA, RLMM xxxx
Anthoptus insignis (Plötz, 1882)
Corticea corticea (Plötz, 1882) PBL x
Corticea l. lysias (Plötz, 1883) RLMM
Mnaseas bicolor (Mabille, 1889) TL
Zariaspes mys (Hübner, [1808]) RLMM
Callimormus r. radiola (Mabille, 1878) PBL, RLMM xxxxx
Callimormus juventus Scudder, 1872 PBL x
Callimormus saturnus (Herrich-Schäffer, 1869) PBL, CURLA x x x
Mnasicles geta Godman, 1901

Insecta MundI 0205, February 2012 • 45Lepidoptera of Honduras
Remella remus (Fabricius, 1798)
Remella vopiscus (Herrich-Schäffer, 1869) PBL xxxx
Eutocus facilis (Plötz, 1884) PBL x x
Eutocus v. vetulus (Mabille, 1883) PBL x
Eprius veleda veleda (Godman, 1900) PBL xxxxx
Methionopsis ina (Plötz, 1882) PBL x x
Methionopsis typhon Godman, 1901 PBL x
Phanes aletes (Geyer, 1832)
Monca crispinus (Plötz, 1882) PBL x
Nastra julia (H. Freeman, 1945)
Nastra l. leucone (Godman, 1900)
Cymaenes tripunctus theogenis (Capronnier, 1874)
Cymaenes trebius (Mabille, 1891) PBL, CA xxx
Cymaenes fraus (Godman, 1900)
Vehilius stictomenes illudens (Mabille, 1891) TL, PBL x x x x
Vehilius inca (Scudder, 1872) PBL x
Mnasilus allubita (Butler, 1877) PBL x
Mnasitheus chrysophrys (Mabille, 1891)
Mnasitheus cephoides Hayward, 1943
Parphorus s. storax (Mabille, 1891) PBL, CURLA x x
Papias dictys Godman, 1900 PBL x x
Papias subcostulata (Herrich-Schäffer, 1870) PBL x
Cobalopsis nero (Herrich-Schäffer, 1869) PBL, RLMM x x x
Lerema accius (J. E. Smith, 1797) PBL x
Lerema liris Evans, 1955 PBL x x
Morys valda Evans, 1955
Morys micythus (Godman, 1900) PBL x x
Morys lyde (Godman, 1900) PBL, CURLA x x x x
Niconiades incomptus Austin, 1997
Niconiades nikko Hayward, 1948 PBL x
Niconiades viridis vista Evans, 1955
Vettius lafrenaye pica (Herrich-Schäffer, 1869) RLMM
Vettius aurelius (Plötz, 1882) PBL, CURLA, RLMM x x
Vettius fantasos (Cramer, 1780)
Vettius onaca Evans, 1955
Paracarystus hypargyra (Herrich-Schäffer, 1869) RLMM
Thoon modius (Mabille, 1889)
Justinia sp. PBL x
Eutychide complana (Herrich-Schäffer, 1869)
Eutychide paria (Plötz, 1882)
Eutychide asema (Mabille, 1891) TL
Naevolus orius (Mabille, 1883)
Enosis i. immaculata (Hewitson, 1868)
Vertica verticalis coatepeca (Schaus, 1902)
Rhinthon osca (Plötz, 1882)
Hylephila p. phyleus (Drury, 1773) ER, RLMM x
Polites subreticulata (Plötz, 1883) TLS, ER x
Polites vibex praeceps (Scudder, 1872) EAPZ, RLMM, ER x
Wallengrenia o. otho (J. E. Smith, 1797)

46 • Insecta MundI 0205, February 2012 Miller et al.
Wallengrenia otho clavus (Erichson, [1849]) PBL x
Pompeius pompeius (Latreille, [1824]) PBL, RLMM x x
Atalopedes campestris huron (W. H. Edwards, 1863) RLMM
Poanes zabulon (Boisduval and Le Conte, [1837]) RLMM
Anatrytone potosiensis (H. Freeman, 1969)
Anatrytone mella (Godman, 1900)
Quasimellana eulogius (Plötz, 1882)
Quasimellana myron (Godman, 1900)
Euphyes peneia (Godman, 1900)
Euphyes ampa Evans, 1955 TL
Metron zimra (Hewitson, 1877)
Quinta cannae (Herrich-Schäffer, 1869) PBL x
Cynea irma (Möschler, 1879) RLMM
Cynea diluta (Herrich-Schäffer, 1869) PBL, RLMM xxxxx
Cynea cynea (Hewitson, 1876) PBL x
Conga chydaea (Butler, 1877) PBL xxxx
Decinea decinea derisor (Mabille, 1891) PBL x x
Tirynthia conua (Herrich-Schäffer, 1869) PBL x
Nyctelius n. nyctelius (Latreille, [1824]) ER, EAPZ, RLMM x
Thespieus dalman (Latreille, [1824]) PBL
Vacerra litana (Hewitson, 1866) PBL x x
Xeniades o. orchamus (Cramer, 1777) PBL x
--- PAPILIONOIDEA ---
PAPILIONIDAE
Parides eurimedes mylotes (H. Bates, 1861) PBL, RLMM, EAPZ xxxxx
Parides i. iphidamas (Fabricius, 1793) PBL, RLMM, EAPZ x x
Parides c. childrenae (Gray, 1832) PBL, RLMM, EAPZ xxxxx
Parides sesostris zestos (Gray, [1853]) TL
Parides panares lycimenes (Boisduval, 1870) PBL, RLMM, EAPZ x
Parides erathalion polyzelus (C. and R. Felder, 1865) PBL, RLMM, EAPZ xxx
Parides montezuma (Westwood, 1842) RLMM, EAPZ
Parides photinus (Doubleday, 1844) RLMM, EAPZ
Battus p. polydamas (Linnaeus, 1758) PBL, RLMM, EAPZ xxx
Battus laodamas copanae (Reakirt, 1863)
Battus ingenuus (Dyar, 1907), PBL, RLMM x
Battus lycidas (Cramer, 1777)
Eurytides e. epidaus (Doubleday, 1846) EAPZ
Eurytides p. philolaus (Boisduval, 1836) RLMM, EAPZ
Eurytides agesilaus neosilaus (Höpffer, 1865) EAPZ
Eurytides c. calliste (H. Bates, 1864) RLMM, EAPZ
Eurytides thyastes marchandii (Boisduval, 1836) RLMM, EAPZ
Eurytides salvini (H. Bates, 1864)
Eurytides m. macrosilaus (Gray, [1853]) TL
Eurytides dioxippus lancandones (H. Bates, 1864)
Mimoides t. thymbraeus (Boisduval, 1836) RLMM
Mimoides ilus branchus (Doubleday, 1846) TL, PBL, RLMM, EAPZ x
Mimoides p. phaon (Boisduval, 1836) RLMM, EAPZ
Papilio polyxenes asterius (Stoll, 1782) EAPZ
Papilio e. erostratus Westwood, 1847 RLMM

Insecta MundI 0205, February 2012 • 47Lepidoptera of Honduras
Papilio garamas electryon H. Bates, 1864
Papilio v. victorinus E. Doubleday, 1844 RLMM
Papilio v. vulneratus Butler, 1872
Papilio birchallii bryki Strand, 1930 RLMM
Heraclides thoas autocles (Rothschild and Jordan, 1906) PBL, CA, RLMM, EAPZ xxxx
Heraclides cresphontes (Cramer, 1777)
Heraclides astyalus pallas (G. Gray, [1853]) PBL x x
Heraclides ornythion (Boisduval, 1836) EAPZ
Heraclides androgeus epidaurus (Godman and Salvin, 1890) EAPZ
Heraclides anchisiades idaeus (Fabricius, 1793) PBL, EAPZ x
Heraclides paeon thrason (C. and R. Felder, 1865)
Heraclides torquatus tolmides (Godman and Salvin, 1890)
PIERIDAE
Dismorphiinae
Pseudopieris nehemia francisca Lamas, 1979 TL
Dismorphia amphione praxinoe (Doubleday, 1844) PBL, RLMM, EAPZ x
Dismorpha crisia steinhauseri J. and R. G. de la Maza, 1984 RLMM, EAPZ
Dismorphia theucharila fortunata (Lucas, 1854) PBL, RLMM x
Enantia a. albania (H. Bates, 1864)
Enantia jethys (Boisduval, 1836)
Enantia lina marion Godman and Salvin 1889
Lieinix nemesis atthis (Doubleday, 1842) RLMM, EAPZ
Coliadinae
Nathalis i. iole Boisduval, 1836 EAPZ
Eurema albula celata (R. Felder, 1869) PBL, CURLA, CA, RLMM, EAPZ x x
Eurema agave millerorum Llorente and Luis, 1987
Eurema daira eugenia (Wallengren, 1860) PBL, CA, RLMM, EAPZ x x
Eurema x. xanthochlora (Kollar, 1850) RLMM, EAPZ
Eurema salome jamapa (Reakirt, 1866) RLMM
Eurema boisduvaliana (C. and R. Felder, 1865) RLMM, EAPZ
Eurema m. mexicana (Boisduval, 1836) RLMM, EAPZ
Abaeis nicippe (Cramer, 1779) RLMM, EAPZ
Pyrisitia proterpia (Fabricius, 1775) PBL, RLMM, EAPZ x
Pyrisitia lisa centralis (Herrich Schäffer, 1865) PBL, CA x
Pyrisitia nise nelphe (R. Felder, 1869) PBL, RLMM, EAPZ x x
Pyrisitia dina westwoodi (Boisduval, 1836) RLMM, EAPZ
Zerene c. cesonia (Stoll, 1790) PBL, RLMM, EAPZ x
Anteos clorinde (Godart, [1824]) PBL, CA, RLMM, EAPZ x
Anteos maerula (Fabricius, 1775) RLMM
Phoebis sennae marcellina (Cramer, 1777) PBL, CA xxxxx
Phoebis p. philea (Linnaeus, 1763) PBL, CA, EAPZ xxx
Phoebis argante (Fabricius, 1775) PBL, CA, RLMM, EAPZ xxxxx
Phoebis a. agarithe (Boisduval, 1836) PBL, EAPZ x
Phoebis neocypris virgo (Butler, 1870) PBL, RLMM, EAPZ x x
Aphrissa s. statira (Cramer, 1777) PBL, RLMM x
Aphrissa boisduvalii (C. and R. Felder, 1861) RLMM
Rhabdodryas t. trite (Linnaeus, 1758)
Pierinae
Hesperocharis c. crocea H. Bates, 1866 RLMM

48 • Insecta MundI 0205, February 2012 Miller et al.
Glutophrissa drusilla tenuis (Lamas, 1981) PBL, EAPZ x
Melete lycimnia isandra (Boisduval, 1836) PBL, RLMM, EAPZ x
Catasticta nimbice ochracea (H. Bates, 1864) RLMM
Pereute charops nigricans Joicey and Talbot, 1928 RLMM
Archonias brassolis approximata (Butler, 1873) RLMM
Perrhybris pamela chajulensis J. and R. G. de la Maza, 1989
Pieriballia v. viardi (Boisduval, 1836) EAPZ
Itaballia demophile centralis Joicey and Talbot, 1928 RLMM
Itaballia pandosia kicaha (Reakirt, 1863) TL, RLMM
Leptophobia aripa elodia (Boisduval, 1836) RLMM
Ascia m. monuste (Linnaeus, 1764) PBL, RLMM, EAPZ xxxxx
Ganyra josephina josepha (Salvin and Godman, 1868) PBL, CA, RLMM, EAPZ xxx
LYCAENIDAE
Theclinae
Eumaeus childrenae (G. Gray, 1832) PBL, RLMM x x
Eumaeus toxea (Godart, [1824]) RLMM, PBL x
Brangas neora (Hewitson, 1867) PBL x
Brangas carthaea (Hewitson, 1868) RLMM
Evenus regalis (Cramer, 1775) RLMM
Evenus coronata (Hewitson, 1865) RLMM
Evenus batesii (Hewitson, 1865) RLMM
Atlides carpasia (Hewitson, 1868)
Arcas imperialis (Cramer, 1775) PBL, RLMM x
Arcas cypria (Geyer, 1837) PBL, RLMM x
Pseudolycaena damo (H. Druce, 1875) PBL, CURLA, RLMM, EAPZ x x x
Theritas mavors Hübner, 1818 PBL, RLMM x x x
Theritas theocritus (Fabricius, 1793) PBL, RLMM x x
Theritas hemon (Cramer, 1775) PBL, RLMM xxx
Theritas lisus (Stoll, 1790)
Thereus cithonius (Godart, [1824])
Rekoa meton (Cramer, 1779) RLMM
Rekoa palegon (Cramer, 1780) PBL, RLMM, EAPZ x x x
Rekoa zebina (Hewitson, 1869) PBL x x
Rekoa marius (Lucas, 1857)
Rekoa stagira (Hewitson, 1867)
Arawacus togarna (Hewitson, 1867) PBL, CURLA, RLMM, EAPZ xxxxx
Arawacus sito (Boisduval, 1836) PBL, RLMM xxxxx
Arawacus dumenilii (Godart, [1824]) RLMM
Kolana lyde (Godman and Salvin, 1887) PBL x
Chlorostrymon simaethis sarita (Skinner, 1895)
Chlorostrymon telea (Hewitson, 1868) EAPZ
Cyanophrys goodsoni (Clench, 1946) TL, RLMM
Cyanophrys amyntor (Cramer, 1775) PBL, EAPZ x
Cyanophrys herodotus (Fabricius, 1793) RLMM
Cyanophrys longula (Hewitson, 1868) RLMM
Cyanophrys agricolor (Butler and H. Druce, 1872) RLMM
Laothus oceia (Godman and Salvin, 1887) PBL, RLMM x
Laothus barajo (Reakirt, [1867]) PBL, RLMM x x x x
Janthecla janthina (Hewitson, 1867) PBL, RLMM x x

Insecta MundI 0205, February 2012 • 49Lepidoptera of Honduras
Lamprospilus collucia (Hewitson, 1877) PBL x
Camissecla charichlorus (Butler and H. Druce, 1872) RLMM
Electrostrymon hugon (Godart, [1824])
Ziegleria hesperitis (Butler and H. Druce, 1872) PBL, RLMM x
Ziegleria ceromia (Hewitson, 1877) PBL x
Calycopis calus (Godart, [1824]) PBL x
Calycopis isobeon (Butler and H. Druce, 1872) PBL, RLMM xxxxx
Calycopis cerata (Hewitson, 1877) PBL, RLMM x
Calycopis pisis (Godman and Salvin, 1887) PBL, RLMM x x
Calycopis demonassa (Hewitson, 1868) RLMM
Calycopis trebula (Hewitson, 1868) RLMM
Strymon yojoa (Reakirt, [1867]) PBL x
Strymon b. bazochii (Godart, [1824]) PBL, RLMM xxx
Strymon ziba (Hewitson, 1868) PBL, CA, RLMM x x x x
Strymon megarus (Godart, [1824]) PBL x
Strymon serapio (Godman and Salvin, 1887)
Tmolus echion echiolus (Draudt, 1920) PBL x x
Tmolus cydrara (Hewitson, 1868) PBL x
Nicolaea dolium (H. Druce, 1907) TL
Ministrymon azia Hewitson, 1873) PBL x
Ministrymon una scopas (Godman and Salvin, 1887) PBL, RLMM x x
Theclopsis mycon (Godman and Salvin, 1887) CURLA x
Theclopsis leos (Schaus, 1913) CURLA x
Ostrinotes purpuriticus (H. H. Druce, 1907) PBL x
Ostrinotes keila (Hewitson, 1869) PBL x
Strephonota tephraeus (Geyer, 1837) PBL, RLMM x x
Strephonota sphinx (Fabricius, 1775) PBL x
Panthiades bitias (Cramer, 1777) PBL, RLMM xxx
Panthiades bathildis (C. and R. Felder, 1865) PBL RLMM, EAPZ xxxxx
Panthiades phaleros (Linnaeus, 1767) PBL, RLMM x x
Oenomaus atena (Hewitson, 1867)
Oenomaus ortygnus (Cramer, 1779) RLMM
Parrhasius polibetes (Stoll, 1781) RLMM
Parrhasius orgia (Hewitson, 1867) PBL, RLMM x
Parrhasius moctezuma (Clench, 1971)
Michaelus ira (Hewitson, 1867)
Michaelus thordesa (Hewitson, 1867)
Aubergina paetus (Godman and Salvin, 1887) PBL, RLMM x
Iaspis andersoni Robbins, 2010 PBL, RLMM x x
Iaspis temesa (Hewitson, 1868) PBL x x x x
Semonina semones (Godman and Salvin, 1887) PBL x
Chalybs janias (Cramer, 1779) RLMM
Chalybs hassan (Stoll, 1790) PBL, RLMM x
Celmia celmus (Cramer, 1775) RLMM
Polyommatinae
Leptotes cassius cassidula (Boisduval, 1870) TL
Cupido comyntas texana (F. Chermock, 1945) PBL xxxxx
Celastrina echo gozora (Boisduval, 1870) TL
Hemiargus ceraunus astenidas (Lucas, 1857) PBL, CA, RLMM, EAPZ xxxxx

50 • Insecta MundI 0205, February 2012 Miller et al.
Echinargus huntingtoni hannoides (Clench, 1965) PBL x
RIODINIDAE
Euselasiinae
Euselasia cataleuca (R. Felder, 1869) RLMM
Euselasia leucophyrna (Schaus, 1913) PBL, RLMM x
Euselasia chrysippe (H. W. Bates, 1866) PBL, RLMM x
Euselasia regipennis eupepla (Godman and Salvin, 1885) RLMM
Euselasia hypophaea (Godman and Salvin, 1878)
Euselasia argentea (Hewitson, 1871) EAPZ
Euselasia eubule (R. Felder, 1869)
Euselasia h. hieronymi (Godman and Salvin, 1868) PBL x
Euselasia a. aurantiaca (Godman and Salvin, 1868) RLMM
Hades noctula Westwood, 1861 EAPZ
Riodininae
Mesosemia zonalis Godman and Salvin, 1885 PBL, RLMM x x
Mesosemia telegone telegone (Boisduval, 1836) PBL x
Mesosemia lamachus Hewitson, 1857 TL, PBL, RLMM x x x
Mesosemia gaudiolum H. W. Bates, 1865 RLMM
Leucochimona lepida nivalis (Godman and Salvin, 1885) PBL x
Leucochimona lagora (Herrich-Schäffer, [1853]) RLMM
Peropthalma tullius (Fabricius, 1787) PBL xxx
Peropthalma lasus Westwood, 1851 TL, PBL, RLMM x x x
Napaea eucharila (H. W. Bates, 1867) PBL,RLMM x
Voltinia theata Stichel, 1910 PBL x
Voltina umbra (Boisduval, 1870) TL
Hyphilaria thasus (Stoll, 1780) EAPZ, PBL x
Eurybia lycisca Westwood, 1851 TL, PBL, RLMM x x x
Eurybia patrona persona Staudinger, 1876 RLMM
Eurybia e. elvina Stichel, 1910 RLMM
Lyropteryx lyra cleadas H. Druce, 1875 PBL, RLMM x
Ancyluris i. inca (Saunders, 1850) PBL, CURLA, RLMM xxxxx
Ancyluris j. jurgensenii (Saunders, 1850) PBL, RLMM xxx
Rhetus arcius castigatus Stichel, 1909 PBL, RLMM x x
Rhetus periander Stichel, 1910 PBL x
Isapis agyrtus hera Godman and Salvin, 1886 PBL, RLMM x
Notheme erota diadema Stichel, 1910 PBL x x
Monethe albertus rudolpus Godman and Salvin, 1885
Chalodeta lypera (H. W. Bates, 1868)
Detritivora barnesi (J. Hall and Harvey, 2001) PBL xxxxx
Charis anius (Cramer, 1776) PBL xxx
Calephelis argyrodines (H. W. Bates, 1866)
Calephelis l. laverna (Godman and Salvin, 1886)
Calephelis fulmen Stichel, 1910 RLMM, PBL x x
Calephelis mexicana McAlpine, 1971
Calephelis browni McAlpine, 1971
Calephelis schausi McAlpine, 1971 PBL x x
Calephelis sixola McAlpine, 1971 PBL x
Calephelis velutina (Godman and Salvin, 1878) PBL, EAPZ, EAPZ xxx
Caria rhacotis (Godman and Salvin, 1878) TL

Insecta MundI 0205, February 2012 • 51Lepidoptera of Honduras
Baeotis z. zonata R. Felder, 1869 PBL x
Baeotis b. barce Hewitson, 1875
Baeotis macularia (Boisduval, 1870) RLMM
Lasaia s. sula Staudinger, 1888 TL, EAPZ
Lasaia agesilas callaina Clench, 1972 PBL, RLMM x
Lasaia m. maria Clench, 1972 RLMM
Exoplisia cadmeis (Hewitson, 1866)
Melanis pixe sanguinea (Stichel, 1910) PBL, RLMM, EAPZ x x x x
Melanis electron melantho (Menetries, 1865) EAPZ
Melanis cephise (Menetries, 1855)
Mesene phareus (Cramer, 1777) PBL, EAPZ x
Mesene croceella H. W. Bates, 1865 PBL x
Mesene m. margaretta (A. White, 1843) TL
Chimastrum a. argenteum (H. W. Bates, 1866)
Symmachia p. probetor (Stoll, 1782) PBL x x
Symmachia accusatrix Westwood, 1851
Pirascca tyriotes (Godman and Salvin, 1878) RLMM
Sarota subtessellata (Schaus, 1913) EAPZ
Sarota chrysus (Stoll, 1781) EAPZ
Sarota g. gamelia Godman and Salvin, 1886
Sarota acantus (Stoll, 1781) PBL x
Sarota myrtea Godman and Salvin, 1886 PBL x
Anteros formosus micon H. Druce, 1875 PBL, RLMM, EAPZ x x
Calydna sturnula (Geyer, 1837)
Emesis lucinda (Cramer, 1775) PBL xxx
Emesis aurimna (Boisduval, 1870) PBL, RLMM x
Emesis ocypore aethalia H. W. Bates, 1868 PBL, RLMM x x
Emesis tenedia C. and R. Felder, 1861 PBL, EAPZ x x x
Emesis mandana furor Butler and H. Druce, 1872 PBL, RLMM, EAPZ x
Emesis tegula Godman and Salvin, 1886 RLMM
Emesis l. lupina Godman and Salvin, 1886
Pseudonymphidia clearista (Butler, 1871) TL
Pachythone gigas nigriciliata Schaus, 1913 RLMM
Thisbe lycorias (Hewitson, [1853]) TL, EAPZ
Juditha caucana (Stichel, 1911) PBL, CA, RLMM xxxxx
Synargis mycone (Hewitson, 1865) PBL, RLMM x
Synargis phliasus velabrum (Godman and Salvin, 1878) RLMM
Synargis n. nymphidioides (Butler, 1872) RLMM
Menander menander purpurata (Godman and Salvin, 1878) PBL x
Menander pretus picta (Godman and Salvin, 1886) RLMM
Calospila cilissa (Hewitson, 1863) PBL, RLMM xxxxx
Calospila pelarge (Godman and Salvin, 1878) PBL, RLMM x x
Hypophylla zeurippa Boisduval, 1836 TL
Hypophylla s. sudias (Hewitson, [1858]) TL, RLMM
Setabis lagus jansoni (Butler, 1870) RLMM
Nymphidium a. ascolia Hewitson, [1853] RLMM
Calicosama lilina (Butler, 1870)
Theope pedias Herrich-Schäffer, [1853] PBL x x
Theope eudocia Westwood, 1851 PBL x x x

52 • Insecta MundI 0205, February 2012 Miller et al.
Theope mundula Stichel, 1926 PBL x
Theope pieridoides C. and R. Felder, 1865
Theope cratylus Godman and Salvin, 1886
Theope eupolis Schaus, 1890 RLMM
Theope virgilius (Fabricius, 1793) PBL, EAPZ x
Theope phaeo Prittwitz, 1865 RLMM
Theope publius incompositus J. Hall, 1999 PBL, RLMM x
NYMPHALIDAE
Libytheinae
Libytheana carinenta mexicana Michener, 1943 ER, RLMM, EAPZ x
Danainae
Danaus p. plexippus (Linnaeus, 1758) PBL, RLMM, EAPZ x
Danaus gilippus thersippus (H. W. Bates, 1863) RLMM, EAPZ
Danaus eresimus montezuma Talbot, 1943 PBL, RLMM, EAPZ x
Lycorea halia atergatis (E. Doubleday, [1847]) PBL, RLMM, EAPZ xxxxx
Lycorea ilione albescens (Distant, 1876) RLMM
Anetia t. thirza Geyer, [1833] RLMM, EAPZ
Ithomiini
Tithorea tarricina pinthias Godman and Salvin, 1878 PBL, RLMM, EAPZ x
Tithorea harmonia salvadoris Staudinger, 1885
Aeria eurimedia pacica Godman and Salvin, 1879 RLMM
Melinaea lilis imitata H. Bates, 1864
Thyridia psidii melantho H. Bates, 1866 PBL, RLMM x
Mechanitis menapis saturata Godman, 1901 PBL, RLMM, EAPZ x x
Mechanitis polymnia lycidice H. W. Bates, 1864 TLS, PBL, RLMM, EAPZ x x x
Mechanitis lysimnia utemaia Reakirt, 1866 TL, PBL, RLMM, EAPZ x x x
Napeogenes t. tolosa (Hewitson, 1855) RLMM
Hypothyris euclea valora (Haensch, 1909) RLMM, EAPZ
Hypothyris lycaste dionaea (Hewitson, 1854) PBL, RLMM x
Ithomia patilla Hewitson, 1852 PBL, RLMM, EAPZ x x
Hyposcada v. virginiana (Hewitson, 1855) TLS, RLMM, PBL x
Oleria paula (Weymer, 1883) RLMM, EAPZ
Oleria z. zea (Hewitson, [1855]) RLMM
Ceratinia tutia (Hewitson, 1852) RLMM, EAPZ
Callithomia hezia hedila Godman and Salvin, 1879 PBL, RLMM, EAPZ x x
Dircenna klugi (Geyer, 1837) RLMM, EAPZ
Dircenna jemina (Geyer, 1837)
Dircenna dero (Hübner, 1823) PBL, RLMM, EAPZ x
Episcada s. salvinia (H. Bates, 1864) RLMM
Pteronymia c. cotytto (Guérin-Méneville, [1844]) PBL, RLMM x
Pteronymia alcmena (Godman and Salvin, 1877) RLMM
Pteronymia a. artena (Hewitson, [1855])
Pteronymia simplex fenochioi Lamas, 1978
Godyris zavaleta sosunga (Reakirt, [1866]) PBL, RLMM, EAPZ x
Godyris nero (Hewitson, [1855]) RLMM
Hypoleria lavinia cassotis (H. W. Bates, 1864) PBL, RLMM x x
Greta andromica lyra (Salvin, 1869)
Greta a. annette (Guérin-Méneville, [1844]) RLMM
Greta morgane oto (Hewitson, [1855]) RLMM, EAPZ

Insecta MundI 0205, February 2012 • 53Lepidoptera of Honduras
Limenitidinae
Adelpha paraena massilia (C. and R. Felder, 1867) PBL, CURLA x
Adelpha serpa celerio (H. W. Bates, 1864) PBL, RLMM x x
Adelpha pithys (H. W. Bates, 1864) RLMM
Adelpha donysa albilum Steinhauser, 1974
Adelpha f. fessonia (Hewitson, 1847)
Adelpha basiloides (H. W. Bates, 1865) TLS, RLMM, EAPZ
Adelpha i. iphiclus (Linnaeus, 1758) RLMM, EAPZ
Adelpha i. iphicleola (H. W. Bates, 1864)
Adelpha cytherea marcia Fruhstorfer, 1913 TL, PBL, CURLA, RLMM, EAPZ x x x x x
Adelpha salmoneus salmonides A. Hall 1938 PBL, RLMM x
Adelpha barnesia leucas Fruhstorfer, 1913
Adelpha n. naxia (C. and R. Felder, 1867)
Adelpha m. melanthe (H. Bates, 1864) RLMM
Adelpha leuceroides Beutelspacher, 1975 RLMM
Adelpha e. erymanthis Godman and Salvin, 1884
Adelpha malea fundania Fruhstorfer, 1915
Adelpha boeotia oberthurii (Boisduval, 1870)
Adelpha delinita utina A. Hall, 1938 TL
Adelpha p. phylaca (H. W. Bates, 1866)
Adelpha phylaca pseudothalia A. Hall, 1938
Adelphsa m. messana (C. and R. Felder, 1867) PBL x
Adelpha lycorias melanthe (H. W. Bates, 1864)
Adelpha l. leuceria (H. Druce, 1874)
Adelpha cocala lorzae (Boisduval, 1870)
Adelpha felderi (Boisduval, 1870)
Adelpha milleri Beutelspacher, 1976 RLMM
Heliconiinae
Altinote ozomene nox (H. W. Bates, 1864) RLMM
Actinote anteas (Doubleday, [1847]) EAPZ
Actinote guatemalena guatemalena (H. Bates, 1864) EAPZ
Agraulis vanillae incarnata (N. Riley, 1926) PBL, CA, EAPZ x x x
Dione juno huascuma (Reakirt, 1866) PBL, EAPZ x
Dione moneta poeyii ( Butler, 1873) EAPZ
Dryadula phaetusa (Linnaeus, 1758) PBL, RLMM, EAPZ x x x
Dryas iulia moderata (N. Riley, 1926) TL, PBL, CA, RLMM, EAPZ xxxxx
Philaethria diatonica (Fruhstorfer, 1912) TL
Laparus doris viridis (Staudinger, 1885) TLS, PBL, RLMM, EAPZ x
Eueides aliphera gracilis Stichel, 1904 TL, PBL, RLMM, EAPZ x x
Eueides lineata Salvin and Godman, 1868 PBL, RLMM x x x x x
Eueides isabella eva (Fabricius, 1793) PBL, RLMM, EAPZ x
Eueides procula asidia Schaus, 1920
Heliconius charitonia vazquezae Comstock and Brown, 1950 PBL, CA, RLMM, EAPZ x x x x x
Heliconius cydno galanthus H. W. Bates, 1864 PBL, CURLA, RLMM, EAPZ x x
Heliconius erato petiverana (Doubleday, 1847) PBL, RLMM, EAPZ xxxxx
Heliconius hecalesia octavia H. W. Bates, 1866 PBL, RLMM x
Heliconius hecale discomaculatus Weymer, 1891 TL
Heliconius hecale zuleika Hewitson, 1854 PBL, EAPZ x x x
Heliconius ismenius telchinia Doubleday, 1847 TL, PBL, CURLA, RLMM x x x x

54 • Insecta MundI 0205, February 2012 Miller et al.
Heliconius sara veraepacis H. W. Bates, 1864 PBL, RLMM, EAPZ x x
Heliconius sapho leuce Doubleday, 1847 PBL, RLMM, EAPZ x x
Heliconius hortense Guérin-Méneville, [1844] RLMM
Euptoieta hegesia meridiania Stichel, 1938 PBL, RLMM, EAPZ x x
Apaturinae
Asterocampa idyja argus (H. Bates, 1864) TLS
Doxocopa pavon theodora (Lucas, 1857) TLS
Doxocopa cyane mexicana Bryk, 1953
Doxocopa laurentina cherubina (C. and R. Felder, 1867) PBL x
Doxocopa callianira (Menetries, 1855) EAPZ
Doxocopa l. laure (Drury, 1773) TL, PBL, RLMM, EAPZ x
Biblidinae
Biblis hyperia aganisa Boisduval, 1836 CA, RLMM, EAPZ x
Mestra amymone (Ménétriés, 1857) PBL, RLMM, EAPZ xxxxx
Catonephelini
Catonephele mexicana Jenkins and R. G. de la Maza, 1985 PBL, RLMM, EAPZ x x x
Catonephele numilia esite (R. Felder, 1869) PBL, RLMM, EAPZ x x
Catonephele chromis godmani Stichel, 1901 PBL x
Eunica t. tatila (Herrich-Schäffer [1885]) RLMM
Eunica caelina agusta H. W. Bates, 1866 RLMM
Eunica mygdonia omoa A, Hall, 1919
Eunica c. caralis (Hewitson, [1857])
Eunica malvina albida Jenkins, 1990
Eunica monima (Stoll, 1782) PBL, ER, RLMM, EAPZ x x
Eunica sydonia caresa (Hewitson, [1857]) RLMM
Eunica chlororhoa mira Godman and Salvin, 1877 RLMM
Eunica volumna venusia (C. and R. Felder, 1867)
Eunica pomona amata H. Druce, 1874
Eunica a. alcmena (Doubleday, [1847]) RLMM
Eunica alpais excelsa Godman and Salvin, 1877 RLMM
Myscelia c. cyaniris Doubleday, [1848] TL, RLMM, EAPZ
Myscelia ethusa pattenia Butler and H. Druce, 1872 EAPZ
Nessaea a. aglaura (Doubleday, [1848]) RLMM, EAPZ
Ectima erycinoides C. and R. Felder, 1867 PBL, CURLA, RLMM x x
Ageroniini
Hamadryas februa ferentina (Godart, [1824]) TLS, PBL, CA, RLMM, EAPZ xxxxx
Hamadryas g. glauconome (H. Bates, 1864) PBL, EAPZ x x
Hamadryas a. atlantis (H. Bates, 1864) EAPZ
Hamadryas feronia farinulenta (Fruhstorfer, 1916) TL, PBL, RLMM, EAPZ xxx
Hamadryas g. guatemalena (H. Bates, 1864) PBL, RLMM xxxx
Hamadryas iphthime joannae Jenkins, 1983 PBL, EAPZ
Hamadryas fornax fornacalia (Fruhstorfer, 1907)
Hamadryas laodamia saurites (Fruhstorfer, 1916) TL, PBL, RLMM, EAPZ xxx
Hamadryas amphinome mexicana (Lucas, 1853) PBL, RLMM, EAPZ x x
Epiphelini
Bolboneura s. sylphis (H. Bates, 1864) ER x
Epiphile a. adrasta Hewitson, 1861
Nica avilla canthara (Doubleday, 1849) PBL, RLMM, EAPZ x x
Pyrrhogyra neaerea hypensor Godman and Salvin, 1884 TLS, RLMM

Insecta MundI 0205, February 2012 • 55Lepidoptera of Honduras
Pyrrhogyra o. otolais H. W. Bates, 1864 PBL, RLMM x x
Pyrrhogyra e. edocla Doubleday, [1848] TLS
Temenis laothoe hondurensis Fruhstorfer, 1907 TL, PBL, RLMM x x
Eubagini
Dynamine postverta mexicana d’Almeida, 1952 PBL, RLMM , EAPZ x x x
Dynamine theseus (C. and R. Felder, 1861) PBL, RLMM x
Dynamine artemisia (Fabricius, 1793) RLMM, EAPZ
Dynamine ate (Godman and Salvin, 1883) RLMM
Dynamine dyonis Geyer, 1837 RLMM, EAPZ
Dynamine paulina thalassina (Boisduval, 1870) EAPZ
Callicorini
Callicore tolima peralta (Dillon, 1948) RLMM
Callicore l. lyca (Doubleday, [1847]) RLMM, UYUCA x x
Callicore astarte patelina (Hewitson, 1853) RLMM, EAPZ
Callicore texa titania (Salvin, 1869) TLS, PBL, RLMM, EAPZ x x
Callicore tolima guatemalena (H. Bates), 1866 PBL x
Callicore pitheas (Latreille, [1813]) RLMM
Diaethria a. anna (Guérin-Méneville, [1844]) PBL, CURLA, RLMM, EAPZ x x x x
Diaethria a. astala (Guérin-Méneville, [1844]) RLMM, EAPZ
Diaethria pandama (Doubleday, [1848]) RLMM
Cyrestinae
Marpesia chiron (Fabricius, 1775) PBL, EAPZ x
Marpesia merops (Doyère, [1840]) PBL, RLMM, EAPZ x
Marpesia c. corita (Westwood, 1850) RLMM
Marpesia marcella valetta (Butler and H. Druce, 1872) EAPZ
Marpesia petreus (Cramer, 1776) PBL, RLMM, EAPZ x x
Marpesia berania fruhstorferi (Seitz, 1914) TL, EAPZ
Marpesia zerynthia dentigera (Fruhstorfer, 1907) RLMM, EAPZ
Nymphalinae
Coeini
Historis odius dious Lamas, 1995 PBL, RLMM, EAPZ x x
Historis a. acheronta (Fabricius, 1775) PBL, EAPZ x
Baeotus baeotus (Doubleday, [1849]) RLMM
Pycina zamba zelys Godman and Salvin, 1884
Nymphalini
Colobura d. dirce (Linnaeus, 1758) PBL, RLMM, EAPZ x x x x x
Tigridia acesta (Linnaeus, 1758) PBL, RLMM, EAPZ x
Smyrna blomldia datis Fruhstorfer, 1908 PBL, RLMM, EAPZ x
Smyrna karwinskii Geyer, [1833] EAPZ
Hypanartia dione disjuncta Willmott et al., 2001 RLMM
Hypartnartia lethe (Fabricius, 1793) RLMM, EAPZ
Vanessa virginiensis (Drury, 1773) RLMM, EAPZ x
Vanessa cardui (Linneaus, 1758) PBL x
Victorinini
Anartia jatrophae luteipicta (Fruhstorfer, 1907) TL, PBL, CA, RLMM, EAPZ x x x x
Anartia f. fatima (Fabricius, 1793) PBL, CA, RLMM, EAPZ xxxxx
Siproeta stelenes biplagiata (Fruhstorfer, 1907) TL, PBL, CA, RLMM, EAPZ xxxxx
Siproeta e. epaphus (Latreille, 1813) PBL, CA, RLMM, EAPZ x x

56 • Insecta MundI 0205, February 2012 Miller et al.
Junoniini
Junonia evarete (Cramer, 1779) ER, EAPZ x
Melitaeini
Chlosyne g. gaudialis (H. Bates, 1864) PBL, RLMM, EAPZ x x
Chlosyne j. janais (Drury, 1782) PBL, RLMM, EAPZ x x
Chlosyne t. theona (Menetries, 1855) ER, RLMM, EAPZ x
Chlosyne ezra (Hewitson, 1864) TLS
Chlosyne l. lacinia (Geyer, 1837) PBL, RLMM, EAPZ xxxx
Microtia e. elva H. Bates, 1864 ER, CA x x
Eresia c. clio (Linnaeus, 1758) RLMM, EAPZ, PBL x
Eresia p. phillyra Hewitson, 1852 RLMM
Castilia myia (Hewitson, [1864]) x
Castilia griseobasalis (Röber, 1913)
Tegosa n. nigrella (H. Bates, 1866) RLMM
Tegosa anieta luka (H. Bates, 1866) RLMM
Tegosa claudina (Eschscholtz, 1821)
Anthanassa ardys (Hewitson, 1864) RLMM
Anthanassa drusilla lelex (H.Bates, 1864) PBL, RLMM x
Anthanassa dracaena phlegias (Godman, 1901) TL
Anthanassa drymaea (Godman and Salvin, 1878)
Anthanassa p. ptolyca (H. Bates, 1864) RLMM
Anthanassa s. sitalces (Godman and Salvin, 1882) RLMM
Anthanassa argentea (Godman and Salvin, 1882) RLMM
Anthanassa atronia (H. Bates, 1866)
Anthanassa o. otanes (Hewitson, 1864) RLMM
Anthanassa tulcis (H. Bates, 1864) PBL, RLMM x x
Charaxinae
Consul fabius cecrops (Doubleday, [1849]) PBL, RLMM x x x
Siderone galanthis (Cramer, 1775) PBL, RLMM x
Siderone s. syntyche Hewitson, [1854]) RLMM
Zaretis callidryas (R. Felder, 1869)
Zaretis ellops (Ménétriés, 1855) PBL, RLMM, EAPZ x
Zaretis i. itys (Cramer, 1777) PBL, RLMM
Anaea aidea (Guérin-Méneville, [1844]) PBL, RLMM x
Fountainea eurypyle confusa (A. Hall, 1929) PBL, RLMM, EAPZ x
Memphis aureola (H. Bates, 1866) PBL, RLMM x
Memphis beatrix (H. Druce, 1874) RLMM
Memphis proserpina (Salvin, 1869) RLMM
Memphis glauce centralis (Röber, 1916) RLMM, EAPZ
Memphis neidhoeferi (Rotger, Escalante, and Coronado, 1965)
Memphis arginussa eubaena (Boisduval, 1870) PBL x
Memphis moruus boisduvali (W. Comstock, 1961) PBL, RLMM x
Memphis lyceus (H. Druce, 1877) PBL x
Memphis oenomais (Boisduval, 1870)
Memphis artacaena (Hewitson, 1869) PBL, RLMM x
Memphis d. dia (Godman and Salvin, 1884)
Memphis p. pithyusa (R. Felder, 1869) PBL, RLMM x
Memphis mora orthesia (Godman and Salvin, 1884) RLMM
Memphis hedemanni (R. Felder, 1869) RLMM

Insecta MundI 0205, February 2012 • 57Lepidoptera of Honduras
Archaeoprepona amphimachus amphiktion (Fruhstorfer, 1916) TL, RLMM
Archaeoprepona demophon centralis (Fruhstorfer, 1905) TL, PBL, RLMM x x
Archaeoprepona demophoon gulina (Fruhstorfer, 1904) TL, PBL, EAPZ x
Archaeoprepona menander phoebus (Boisduval, 1870) TL, RLMM
Prepona dexamenus medinai Beutelspacher, 1981 PBL, RLMM x
Prepona laertes octavia Fruhstorfer, 1905 TL, PBL, RLMM, EAPZ x
Prepona pylene philetas Fruhstorfer, 1904 TL
Prepona pylene gnorima H. Bates, 1865
Agrias aedon rodriguezi Schaus, 1918 RLMM
Agrias amydon lacandona R. G. and J. de la Maza, 1999 PBL, RLMM x
Satyrinae
Morphini
Antirrhea philoctetes lindigii C. and R. Felder, 1862 CURLA, RLMM, EAPZ x x
Morpho achilles patroclus C. and R. Felder, 1861 EAPZ
Morpho helenor montezuma (Guenée, 1859) TLS, PBL, RLMM, EAPZ x x
Morpho cypris aphrodite LeMoult and Réal, 1962 PBL, CURLA, RLMM x
Morpho polyphemus catalina Corea and Chacón, 1984 EAPZ, RLMM, PBL x
Morpho polyphemus luna Butler, 1869 TLS
Morpho theseus arotos (Frustorfer, 1905) PBL, RLMM x x
Morpho theseus justitiae Salvin and Godman, 1868 TLS, PBL, RLMM x x
Brassolinae
Caligo oileus scamander (Boisduval, 1870) RLMM, EAPZ
Caligo telemonius memnon (C. and R. Felder, 1867) TLS, PBL, RLMM, EAPZ xxx
Caligo illioneus oberon Butler, 1870 RLMM
Caligo brasiliensis sulanus Fruhstorfer, 1904 TL, PBL x x x
Caligo oedipus fruhstorferi Stichel, 1904 TL, RLMM x
Caligo uranus Herrich-Schäffer, 1850 PBL, RLMM, EAPZ x x x
Catoblepia o. orgetorix (Hewitson, 1870) PBL, RLMM x x
Dynastor darius stygianus Buter, 1872 EAPZ
Dynastor macrosiris strix Bates, 1864 EAPZ
Eryphanis aesacus (Herrich-Schäffer, 1850) RLMM, EAPZ
Eryphanis lycomedon (C. and R. Felder, 1862) RLMM
Opsiphanes boisduvallii Doubleday, [1849] PBL, RLMM x x
Opsiphanes bogotanus alajuela Bristow, 1991 PBL x
Opsiphanes cassiae castaneus Stichel, 1904
Opsiphanes t. tamarindi C. and R. Felder, 1861 TLS, PBL, RLMM, EAPZ x x
Opsiphanes quiteria quirinus Godman and Salvin, 1881 PBL, RLMM, EAPZ x
Opsiphanes cassina fabricii (Boisduval, 1870) RLMM, EAPZ
Opsiphanes invirae relucens Fruhstorfer, 1907 TL
Narope testacea Godman and Salvin, 1878
Haeterini
Cithaerias p. pireta (Stoll, 1780) PBL, RLMM, EAPZ x x
Pierella helvina incanescens Godman and Salvin, 1877 PBL, CURLA, RLMM, EAPZ x
Pierella luna pallida (Salvin and Godman, 1868) PBL, EAPZ xxxxx
Pierella luna rubecula Salvin and Godman, 1868 TLS, RLMM
Melanitini
Manataria hercyna maculata (Hopffer, 1874) RLMM, EAPZ
Satyrini
Cyllopsis h. hedemanni R. Felder, 1869 RLMM

58 • Insecta MundI 0205, February 2012 Miller et al.
Cyllopsis steinhauserorum L. Miller, 1974 RLMM
Cyllopsis hilaria (Godman, 1901)
Cyllopsis suivalenoides L. Miller, 1974
Cyllopsis p. pyracmon (Butler, [1867])
Cyllopsis pephredo (Godman, 1901) RLMM
Euptychia hilaria (C. and R. Felder, 1867)
Euptychia westwoodi Butler, 1867 TL
Hermeuptychia hermes (Fabricius, 1775) PBL, CA, RLMM, EAPZ xxxxx
Cissia labe (Butler, 1870) PBL, RLMM xxxxx
Cissia similis (Butler, 1867) PBL x
Cissia pompilia (C. and R. Felder, 1867) PBL, RLMM, EAPZ x x x x
Cissia themis (Butler, 1867) PBL, RLMM x x x x
Magneuptychia libye (Linnaeus, 1767) PBL, RLMM, EAPZ xxxxx
Megeuptychia antonoe (Cramer, 1775) PBL, EAPZ x
Pareuptychia metaleuca (Boisduval, 1870) PBL, RLMM xxxxx
Pareuptychia ocirrhoe (Fabricius, 1776) PBL, CA, RLMM, EAPZ xxxxx
Satyrotaygetis satyrina (H. W. Baters, 1865) RLMM, EAPZ
Taygetis mermeria excavata Butler, 1868 TL, PBL, RLMM x
Taygetis laches (Fabricius, 1793) PBL, RLMM, EAPZ xxxxx
Taygetis rufomarginata Staudinger, 1888 PBL, RLMM, EAPZ x x
Taygetis thamyra (Cramer, 1779) PBL, RLMM x
Taygetis virgilia (Cramer, 1776) TLS, RLMM
Taygetomorpha celia (Cramer, 1779) EAPZ
Oxeoschistus tauropolis (Westwood, [1850]) UYUCA, RLMM, EAPZ x
NOTES:
A1 – Identication in EAPZ database questionable. Specimens from the TL only in USNM. These may be the very
similar H. drucei (Schaus, 1910), from Guatemala and Costa Rica.
A2 – Identication questionable: USNM has phenotype (as illustrated in Seitz) from Peru and Brazil.
Results and Discussion
While our inventory of the Lepidoptera of Honduras is far from complete, our initial results and
comparisons with previous records provide insight into the expected diversity and potential evidence of
endemism. Our eldwork to date has been based primarily at the Lodge at Pico Bonito and other areas
along the northern coast near La Ceiba. Moth collecting has also been centered at the Lodge at Pico
Bonito. Moth groups well represented in our nocturnal samples include the Pyraloidea (principally
Crambidae), Noctuoidea, especially Arctiinae and Notodontidae, Geometridae, Sphingidae, and
Saturniidae. Thus far, excluding undetermined material, more than 975 species of moths have been
collected during our surveys, with the addition of literature and museum specimen records producing
a total of 1,441 moth species included in the present list for Honduras. The highest diversity has been
within the Pyraloidea and Noctuoidea, with many species represented by single specimens. A total of
861 buttery species are now listed for Honduras, with 393 species collected during our surveys. In all,
2,302 species of Lepidoptera are recorded herein. Based on comparison with the work of Janzen and
Hallwachs (2010, pers. comm.) for Costa Rica and our current ndings, we estimate the total number
of Lepidoptera species in Honduras will exceed 6,000. Some representative taxa of butteries (Fig.
2-9) and moths (Fig. 10-36) sampled during our surveys are illustrated. However, we hope to provide
reference images for many additional species, either web-based and/or in subsequent publications.
In summarizing our preliminary results for various moth taxa, we note the Pyraloidea are
particularly diverse with 290 species listed, and many samples are still undetermined. Several new
species are present in our material, some of which are also found in other parts of Central America,
such as Area de Conservación Guanacaste, northwestern Costa Rica, and await revisionary studies

Insecta MundI 0205, February 2012 • 59Lepidoptera of Honduras
of the various genera. Not surprising, given the proximity to two rivers, the aquatic Crambidae
(Acentropinae) are especially abundant at PBL with 27 morphospecies distinguished, 11 determined
to named species thus far and 11 recognized as probable new species. While only 29 species of
Gelechioidea are indicated in our list, material representing this group includes many additional
species to be sorted and identied. In the Glyphidoceridae alone, Adamski (pers. comm.) recognized four
Figures 2-9. Honduran butteries. 2) Pareuptychia ocirrhoe. 3) Calephelis sp. 4) Charis anius. 5) Parides eurimedes
mylotes. 6) Cymaenes trebius. 7) Strymon ziba. 8) Calospila cilissa. 9) Vehilius stictomenes illudens.

60 • Insecta MundI 0205, February 2012 Miller et al.
new species that he will describe in a forthcoming publication. Other microlepidopteran groups, such
as the Tortricoidea, are poorly represented in both numbers of species and the number of individuals
collected. The Tortricidae do, however, hold some interesting surprises, with colorful, elaborately
patterned diurnal genera such as Pseudatteria Walsingham and Mictopsichia Hübner. In the later
genus, Matthews et al. (2011) reported the occurrence of M. cubae (Fig. 14) from our surveys, noting
the new country record, biogeographical signicance, and describing the previously unknown female
genitalia and evasive behavior of adults.
Some progress has been made in our knowledge of the Pterophoroidea with additional work to
be completed. Literature accounts for Pterophoridae (Walsingham 1915; Gielis 2003, 2006) indicated
Figures 10-36. A small sample of the moth diversity at Pico Bonito Lodge. 10) Hedylidae, Macrosoma hyacinthine.
11) Megalopygidae, Trosia nigropunctigera. 12) Saturniidae, Hyperchira nausica. 13) Elaschistidae, Ethmia
ungulatella. 14) Tortricidae, Mictopsichia cubae. 15) Dalceridae, Dalcerides mesoa. 16-18) Crambidae – 16)
Cosmopterosis spatha; 17) Coenostolopsis apicalis; 18) Conchylodes nolckenialis. 19) Thyrididae, Draconia
denticulate. 20-23) Geometridae – 20) Pyrinia itunaria; 21) Tricentrogyna violescens; 22) Pantherodes sp.; 23)
Phrygionis privignaria. 24-27) Noctuidae, Arctiinae – 24) Cissura plumbea; 25) Viviennea tegyra; 26) Mulona
phelina; 27) Horama oedippus. 28) Notodontidae – Calledema argenta. 29-36) Noctuidae – 29) Sosxetra grata; 30)
Eulepidotis perducens; 31) E. hermura; 32) E. punctilinea; 33) Gonodonta sinaldus; 34) Antiblemma incarnans;
35) Azeta rhodogaster; 36) Ceroctena amynta.

Insecta MundI 0205, February 2012 • 61Lepidoptera of Honduras
only six species from Honduras, while 46 and 15 species are recorded from Costa Rica and Guatemala,
respectively. Matthews and Lott (2005) reviewed and catalogued larval foodplants of the world’s
Pterophoridae. While 70 plant families are utilized, one-third of pterophorids for which larval
foodplants are known are associated with Asteraceae. Based on the diversity of the Asteraceae species
found in the Herbario Zamorano Paul C. Standley, EAPZ, from Honduras (410 species) (database
records, courtesy of Director, Dr. George Pilz) and comparison with the adjacent Central American
faunas and the Nearctic fauna (159 species, Matthews 2006), the Honduran fauna of Pterophoridae
should approach or exceed 50 species. After examination of material in the collections at EAPZ and our
recent surveys, we now recognize 17 species of Pterophoridae from Honduras with 12 named species,
and the remainder determined to genus. At least two species are known to be new, with descriptions
in preparation by Matthews and Miller. In addition, Matthews and Miller (2010) recently described
the life history of the cacao plume moth, Michaleophorus nubilus in Honduras, also noting the new
record for the country.
The major macro moth groups (Zygaenoidea, Cossoidea, Sesioidea, Lasiocampoidea, Bombycoidea,
Geometroidea, Noctuoidea) account for 1,055 or 73% of the 1,441 moth species listed, with 674 of these
represented by our recent eldwork. Among the generally diurnal taxa, the Castniidae (Castniinae)
are poorly known from Honduras. Originally, there were only a few Honduran records in collections of
Telchin atymnius futilis and Athis i. inca. No specimens of the latter have been observed during our
surveys as yet, but T. atymnius futilis has been frequently encountered along the northern coast at
PBL and CURLA. In addition, specimens of another species, Divana diva, a species widely distributed
throughout Central America, are known from PBL and also collected by RL (RLMM). Specimens of
Athis clitarcha, originally described from Panama (Salvin) and Nicaragua (Belt) (Miller 1986), have
also been collected in Honduras (RLMM and EAPZ). Amauta cacica procera was originally described
from Guatemala, with most specimens in museum collections recorded from Chiriqui in Panama
(Miller and Sourakov 2009). This species has been observed by Janzen in eastern Costa Rica (pers.
comm.) and collected in Honduras (RLMM and EAPZ). Another species, Telchin licus, has not been
previously recorded in the literature north of Panama, but it has also been collected in Honduras
(RLMM and EAPZ). This species, an agricultural pest normally associated as larvae with various
Heliconia and sugar cane in South America (especially Amazon basin), appears to be well established
in the Pico Bonito National Park and elsewhere along the northern coast of Honduras. This latter
species has obviously extended its geographic range or possibly was introduced to the area. We believe
that with continuing surveys at least another two to four species of Castniidae will be found to occur
in Honduras.
The Noctuoidea are thus far represented by 561 species, primarily Erebidae (386 species). While
most have been collected at lights, a few, such as Ascalapha odorata, were commonly found in bait
traps. We have also observed Azeta rhodogaster (Fig. 35) during the day on a regular basis, easily
spotted in ight by the red abdomen. The larvae of this species feed on the “living fence” tree Gliricidia
sepium (Fabaceae) which is very common in northern Honduras and elsewhere in Central America.
Also encountered during the day at PBL and CURLA were several dioptines (Notodontidae), including
Oricia truncata and Josia, Phanoptis, and Phaeochlanena spp., the latter along with several of the
nocturnal genera in our material containing undescribed species and in need of careful study (James
Miller, pers. comm.). Certain erebid genera such as Eulepidotis (12 species) of the Eulepidotinae and
Gonodonta (10 species) of the Calpinae are notably diverse at PBL, but the Arctiinae are overall the
most diverse with 210 species listed for Honduras and 131 collected at PBL. While some of the striking
wasp mimics of this group are frequently encountered at the sheet, such as Horama plumipes, others,
for example, the widespread species H. oedippus (Fig. 27), are represented by single specimens. Some
genera such as Macrocneme are numerous at the sheet but though rst appearing as single species,
show slight variations in the maculation and metallic hues on the black forewings, with cryptic species
likely involved. One of our most signicant nds in the Arctiinae is a single specimen of Euthyone
theodula (Lithosiini), previously known only by the Schaus holotype from Guatemala in the USNM.
The Geometroidea are another sizeable group with 215 species recorded for the country and 161
collected at PBL. Several species are more or less ubiquitous, present in large numbers at the sheet
during each sampling period. Examples include three species of Synchlora Guenée, Tricentrogyna
violescens (Fig. 21), Pyrinia sanitaria, Patalene aenetusaria, Oxydia vesulia, and Leuciris mbriaria.

62 • Insecta MundI 0205, February 2012 Miller et al.
Certain genera such as Macaria and Eois are common and will undoubtedly add named species to
the list with further identication work including dissections of genitalia and comparison with type
material. Revisionary studies such as Poole (1987) and Pitkin (1993, 1996) have greatly contributed
to the known fauna. Further collecting and identications by CVC and others as well as the results
of ongoing barcoding work by research associates visiting PBL in June 2010 in collaboration with the
Biodiversity Institute of Ontario are also expected to contribute markedly to our knowledge of the
Honduran fauna.
During the last two years, our surveys have sampled the overall lepidopteran biodiversity but
due to the limitations on the total numbers of collected specimens permitted per trip and space
considerations in transport, we have focused on many of the smaller moths. However, within the
Bombycoidea, we have collected several Bombycidae (7) and have voucher specimens and other
records for 82 Saturniidae. Although Ronald Cave and Robert Lehman have collected a number of the
Hemileucinae during their respective surveys, we have not yet collected voucher specimens for many
of these species. However, Schausiella santarosensis was a remarkable nd, collected in November
during some rather heavy rains. In reviewing the literature and other records, it is surprising that
Copiopteryx semiramis has not been reported in literature accounts for Honduras. This genus is rather
uncommon in the eld, but conspicuous because of the long tails. Meerman (1999) listed it for Belize
and there is a single specimen from Honduras in EAPZ. The diversity of another major bombycoid
lineage, Sphingidae, at PBL is remarkable with about half (50) of the total number of known Honduras
species (112) present.
Within the butteries, the number of known Mesoamerican Hesperiidae (combined species and
subspecies) is rather remarkable. Evans (1951, 1952, 1953 and 1955) listed 223 species of Hesperiidae
from Honduras, while Munroe and Miller (1967) recorded 83 species, many of which were new national
records at the time. ADW (unpublished) conducted an analysis of species represented in literature and
collections, and the results are indicative of how few biodiversity surveys have been completed in some
areas of Mesoamerica. Records for Hesperiidae in Central America were as follows: Mexico 792, Belize
282, Guatemala 391, El Salvador 283, Nicaragua 366, Costa Rica 589, and Panama 448. Including our
recent eld data, the total number of Hesperiidae now recorded for Honduras stands at 311 species.
Based on the analysis of Mesoamerican distributions (ADW), we expect up to 587 taxa of Hesperiidae
to eventually be found in Honduras, suggesting that as much as 47% of the country’s hesperiid fauna
remains to be documented. This number may at rst seem unusually high, but the discovery of 35
undescribed hesperiid taxa from Mexico within the last 15 years, coupled with at least that many new
hesperiid taxa recently discovered in Costa Rica (Burns and Janzen 2001, 2005a,b; Burns et al. 2007;
Hebert et al. 2004; Janzen and Hallwachs 2010; Pozo et al. 2008; Warren et al. 2011), reinforces the
fact that there is much to learn about the distributions and diversity of Mesoamerican Hesperiidae,
especially those of high-altitude habitats, which are very poorly known for Hesperiidae and other
groups in Honduras.
Compared to the Hesperioidea, there are fewer new additions in the Papilionoidea with a total of
550 taxa recorded for Honduras, 267 for which we have collected voucher specimens during our 2009-
2010 surveys. There are a number of potential additional species recorded for the country, but often
these are taxa which have been recorded from surrounding countries and not been previously published
elsewhere. Some of the most noteworthy new records include Ostrinotes purpuriticus, Strephannota
sphinx, Euselasia h. hieronymi, Ectima erycinoides, and Oxeoschistus tauropolis. The butteries
provide some interesting statistics with respect to the overall species biodiversity. For example, in the
Riodinidae, there are 101 vouchered or potential taxa for Honduras and of these, we have identied
and collected 50 taxa thus far. Some species such as Calospila cilissa and Juditha caucana appear to
have multiple broods throughout the year and were present at PBL during our surveys in May, June,
August and November. Other interesting species in this group include Leuchochimona lepida nivalis,
Peropthalma tullius, P. lasus, and Notheme erota diadema. While there is great diversity in the genus
Calephelis, some problematic specimens remain undetermined. Although members of this family did
perch on the under surface of plants, we observed a number taking nectar on owers in open sunny
patches both at PBL and CA. Thus far, 50 Riodinidae and 51 Lycaenidae have been identied from our
recent studies in Honduras compared with 49 Riodinidae and 101 Lycaenidae identied by Austin et
al. (1996) from Tikal, Guatemala.

Insecta MundI 0205, February 2012 • 63Lepidoptera of Honduras
Several butteries, including Lycaenidae (Eumaeini) and Hesperiidae (Eudaminae), were collected
at mercury vapor lights at night. Among these are Cyanophrys fusius, Brangas neora, Bungalotis
astylos, Dyscophellus phraxanor, and Polygonus s. savigny. Only the latter species was also collected
during the day. This source of sampling should not be overlooked since the crepuscular skippers and
canopy dwelling lycaenids, such as B. neora, might not otherwise be readily encountered.
As previously noted, our list is by no means complete, especially for the moths, with only a fraction
of the expected number of species represented for the country. Future survey work must extend to
other areas of the country, and include sampling at higher elevations and in a greater variety of
habitats. Even for PBL, it will be necessary to continue sampling with multiple techniques over the
course of several years and throughout the year, to include the variation of seasonal wet/dry cycles,
periodic heavy rainfall from Atlantic tropical systems, as well as rain shadow effects of mountains in
the vicinity. Along with more extensive sampling, there are also problems to overcome with species-
level identications. Most of our determinations of moths are based on comparisons with species
known from Guatemala, Belize, and Costa Rica, using limited literature, internet, and collection
resources. For many of the taxa listed, we have, unfortunately, had to resort to using genus plus “sp.”
or “nr.” (near) as a means of recording our results. While we have been able to determine most taxa to
genus, there are still numerous morphotypes to be sorted out and classed as numbered unknowns in
Gelechioidea, Noctuoidea, and likewise for several pyraloids, altogether representing approximately
170 species.
Although many of the low elevation species encountered thus far are broad-ranging and occur
throughout Central America and much of the rest of the Neotropics, there are still overriding questions
as to whether or not these similar phenotypes are indeed the same genotype or monophyletic species
from one area to the next. Ultimately, DNA barcoding and morphological studies may help resolve
some of these problems, including cases of cryptic sympatric species. However, especially for moths,
many of the species we have identied were described by Walker, Druce, Schaus, and others more
than 80 years ago and previously known only from the holotype specimens. Thus, while barcoding may
separate close species, unless genomic sequences can be successfully obtained from these relatively old
holotypes, nal decisions on nomenclature will have to be based on morphology alone. This problem
is further compounded by the need to examine the types of multiple synonyms already established for
many neotropical species.
As our work continues and the list of species is rened through revisionary studies, there will
be many other aspects of the natural history and biodiversity of Honduran Lepidoptera to further
explore. Janzen and Hallwachs (2010) and collaborators have produced remarkable websites and
long-term studies documenting the life histories of Guanacaste Lepidoptera in Costa Rica, and it is
hoped that similar, if less extensive, efforts can be made at PBL in collaboration with staff, students,
and parataxonomists. In many cases the best approach to life history studies may be to survey the
associates of specic larval food plants. In general, our surveys of adults have been strictly qualitative,
with the primary objective of assembling faunal lists and recognizing new records, undescribed and/
or endemic taxa. We hope that this work will provide baseline data for more quantitative studies
by future researchers investigating biodiversity patterns and trends in comparing areas based on
elevation, microhabitat, environmental quality, climate change, and other factors used in recognizing
critical areas for conservation management.
We have not yet assembled sufcient data to be able to make predictions on the number of endemic
taxa to be expected for Honduras. This is due in part to working at low elevations, and limited sampling
in isolated habitats such as the tropical dry forest habitats similar to the ER. We are aware of several
new species from Honduras, though some are recognized as also occurring in other areas of Central
America. In a few cases, such as in the Pterophoridae, though not necessarily endemic, the new species
are known from Honduras only from PBL specimens.
Honduras, as a critical part of the Mesoamerican biodiversity “hot spot” has much to offer in
the study of Lepidoptera and other organisms. In addition, studies of uctuating distributions and
evolutionary divergence patterns of taxa in Central America and the Caribbean Basin can provide
insight to our understanding of phylogenies and divergence patterns within higher taxa. The present
listing of taxa and review of previous works provide a foundation for further surveys and studies in
the area. We welcome additions and corrections to the present list for future updates and encourage

64 • Insecta MundI 0205, February 2012 Miller et al.
individual publications of faunal records, life history studies, and revisionary works including material
collected from past, present, and future surveys. This work not only contributes to science in general,
but serves to increase the value and appreciation of the natural resources of this country for future
generations of Honduran citizens.
Acknowledgments
We gratefully acknowledge Jose Manuel Mora Benavides, Director of the Biodiversity Center, and
Oliver Schlein, Escuela Agricultura Zamorano, Honduras, for their invaluable assistance in making
the facilities and the collection resources of the EAPZ available to us and aiding us with the permitting
process. Permits for collection and export of study material were granted through Instituto Nacional
de Conservaciόn y Desarrolla Forestal, Área Protegidas y Vida Silvestre, Honduras. We have beneted
from discussions with Ronald Cave, formerly of EAPZ, and now in the Department of Entomology and
Nematology, University of Florida. Ron, along with Oliver Schlein, and George Pilz graciously shared
information concerning the EAPZ collections and databases and also provided us with details on
various collecting localities throughout Honduras. We are indebted to James Adams and the staff and
directors of The Lodge at Pico Bonito, for their hospitality and for providing us logistical assistance
in completing our surveys. We also thank Juan Alberto Hernández and Kelvin W. Bodden, Centro
Universitario Regional del Litoral Atlantico for access to the CURLA reserve at Parque National Pico
Bonito. Special thanks are due to J. Court Whelan and James B. Schlachta for assistance with travel
arrangements.
In addition to the respective contributions of the authors, several colleagues, including Ulf
Eitschberger, James K. Adams, James Hayden, Terry Dickel, and David Adamski, generously
provided multiple determinations of species in their respective areas of expertise. Our communications
with Dan Janzen and Winnie Hallwachs have also given us insight into the overall diversity in the
region and the potential for undescribed species in various taxa. Authors JYM, DLM, TCE, and CVC
would also like to thank the following individuals who provided good-humored camaraderie in the
eld, assisted with our eld work, and/or supplied specimens and/or determinations resulting from
their respective collecting activities in Honduras: Eric Anderson, Eddie Atamian, David Auth, Jorge
Chavarria Bertrand, Kelvin Bodden, David G. Bridgnell, William H. Bridgnell, Richard C. Cramer,
Chad T. Douglas, Collin M. Douglas, Matt Lehnert, Ron Leuschner, James Hanlon, Lowell N. Harris,
Lawrence R. Hummer, Chris Livesay, David G. Marqua, Gerald M. McWilliams, Linda S. McWilliams,
Iva Mihoci, Carlos Montúfar, Alfred Moser, Thomas V. Myers, Ichiro Nakamura, Bedros Orchanian,
Arthur Ortegon, Tony Ortegon, John G. Pasko, Christian Salcedo, Ana Clariza Samayoa, Ian K.
Segabarth, Bradley A. Stirn, Jon D. Turner, Nancy C. Turner, Gerald S. Wagner, J. Court Whelan,
and Eckhard Wierig. MAS wishes to thank Mark Metz for support on the Pyraloidea.
Finally, and not in the least, we wish to thank James K. Adams, Dalton State College, Dalton,
Georgia, Manuel A. Balcázar-Lara, Universidad de Colima, Mexico, Terry A. Lott, Florida Museum
of Natural History, Paul E. Skelley, Florida State Collection of Arthropods, and Jennifer Zaspel,
University of Wisconsin, Oshkosh for their helpful comments in improving the text, format, and
content of the paper.
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