New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange, Southern Turkey

Abstract

The lower Tuvalian sequence of the Sorgun Ophiolitic Mélange (S Turkey) harbors one of the most diverse and best preserved Late Triassic (Carnian) radiolarian faunas of the world. Here we describe a new genus (Spinostylosphaera n. gen.) and 7 new species from the Tavusçay?r? block north of Erdemili. The fauna belongs to the Spongotortilispinus moixi radiolarian Zone (Paragondolella postinclinata - Paragondolella noah conodont zone). ©2015 E. Schweizerbart?sche Verlagsbuchhandlung, Stuttgart, Germany.

Full text

New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic
Mélange, southern Turkey
Péter Ozsvárt, Patrice Moix, and Heinz W. Kozur (†)
With 6 figures
Abstract: The lower Tuvalian sequence of the Sorgun Ophiolitic Mélange (S Turkey) harbors one of
the most diverse and best preserved Late Triassic (Carnian) radiolarian faunas of the world. Here we
describe a new genus (Spinostylosphaera n. gen.) and 7 new species from the Tavusçayırı block north
of Erdemili. The fauna belongs to the Spongotortilispinus moixi radiolarian Zone (Paragondolella
postinclinata – Paragondolella noah conodont zone).
Key words: Radiolaria, Spumellaria, Entactinaria, Carnian, Early Tuvalian, Sorgun Ophiolitic Mé-
lange, Southern Turkey.
1. Introduction
The Tavusçayırı Block sensu Masset & Moix (2004) is
one of the most important and well developed tectonic
blocks belonging to the Sorgun Ophiolitic Mélange,
southern Turkey (Moix et al. 2011). This tectonic block
is comparable to the Huğlu unit described elsewhere
in the Taurides. Part of the investigated sequence con-
tains a very well-preserved and particularly diverse
lower Tuvalian (Carnian, Triassic) radiolarian fauna,
which was already discovered in 2002 by Masset &
Moix. Several lower Tuvalian radiolarians were re-
ported in the MSc thesis of Masset & Moix (2004),
and more than 2000 SEM pictures were taken by
Kozur in 2003 and 2004. In 2007, Heinz W. Kozur,
in collaboration with Péter ozsvárt and Patrice
Moix began to study this assemblage in more detail.
Consequently, Moix et al. (2007) established the lower
Tuvalian Spongotortilispinus moixi radiolarian zone
and described a few of the radiolarian species from
this zone. Numerous additional radiolarian taxa were
subsequently described from the type locality of the S.
moixi zone by Kozur et al. (2007a, b, c, 2009). Thus
far, 3 new families, 10 new genera, and 106 new spe-
cies and subspecies were described. Unfortunately, on
the 20th of December 2014, Heinz W. Kozur passed
away. Shortly before, he managed to handover the
complete lower Tuvalian radiolaria material to Péter
ozsvárt including digital and printed SEM pictures.
In this paper, we intend to continue his work and we
would like to dedicate this paper to the memory of
Heinz W. Kozur. The illustrated type material has
been deposited in the Hungarian Natural History
Museum, Budapest.
2. Geological setting
The Mersin Mélanges are situated in southern Turkey
and belong to the South-Taurides Exotic Units defined
by Moix et al. (2008). Moix et al. (2011) introduced
the concept of Mersin Mélanges to describe two major
independent units identified in the Mersin Ophiolitic
Complex, i.e. the Late Cretaceous Sorgun Ophiolitic
Mélange (SOM) and the Middle to Upper Triassic
Hacialanı Mélange (Figs. 1-2). Both mélanges consist
©2015 E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, Germany www.schweizerbart.de
DOI: 10.1127/njgpa/2015/0509 0077-7749/2015/0509 $ 4.00
N. Jb. Geol. Paläont. Abh. 277/3 (2015), 337–352 Article
Stuttgart, September 2015
E
eschweizerbart_xxx

338 Péter Ozsvárt et al.
of a chaotic accumulation of blocks and rocks in a tec-
tonic and sedimentary mixture (olistostrome) of clas-
tics, ophiolitic material, and oceanic and exotic blocks
of various ages. Blocks in the mélanges typically range
in size from meters to hundreds of meters. Individual
formations are represented by elongated bodies rang-
ing in size from hundreds of meters to kilometers.
The Sorgun Ophiolitic Mélange is part of the infra-
ophiolitic mélanges associated to the Mersin Ophiolitic
Complex (ParlaK 1996) and was emplaced on the
Bolkardağ during the Late Cretaceous (Campanian-
Maastrichtian). The Mersin ophiolite formed in a su-
pra-subduction zone tectonic setting during the Late
Cretaceous (ParlaK & Delaloye 1996). Many coher-
ent series were identified within the Sorgun Ophiolitic
Mélange. One of them is represented by the Tavusçayırı
block which is presented in detail below. In addition to
these km-sized blocks, there is a multitude of smaller
blocks: carbonates ranging from Early Carboniferous
to Late Triassic; radiolarites ranging from Ladinian to
late Turonian–early Coniacian; rare blocks of amphi-
bolites; blocks of partially serpentinous peridotites,
gabbros and pillow-lavas, and blocks of debris-flows
(including ophiolitic debris).
3. Stratigraphy
The section investigated (Tavusçayırı block) is situ-
ated north of Erdemli (Mersin, southern Turkey) and
is bounded by the villages of Gâvuruçtuĝu to the north
and Sorg un to the sout h (Fig. 3). The Tavusçayırı block
Fig. 1. Tectonic map of Turkey and surrounding region. Modified from Moix et al. (2008). Location of Fig. 2.
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New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange 339
(Fig. 4) is an isolated 600-m-thick formation. Its lat-
eral extent is large (km-scale), and its lithological com-
ponents are sometimes reproduced on a smaller scale
elsewhere in the SOM. This succession represents a
typical transgressive sequence marking the break-up
of a platform and the opening of a rift basin during
the Late Triassic. The sequence starts with 15 to 20 m
polymictic breccia in a red micritic matrix. The ele-
ments are heterogeneous in size, varying from mm-
to m-size, and are mostly composed of white neritic
limestones, pink micritic limestones and red calcar-
enites. Some elements yielded Middle Triassic pelagic
faunas. Above an erosional contact, the breccia is fol-
lowed by a 60 m polygenic clast-supported conglomer-
ate with a red silty matrix. The conglomerate shows
cross-bedding and is composed of cm- to dm-size ele-
ments, such as black and white, partly reefal limestone,
beige and red micritic limestones, and sandstones.
Fig. 2. Composite section and simplified geological map of the Mersin Ophiolitic Complex (MOC). Modified from Moix
et al. (2011). Location of Fig. 3.
eschweizerbart_xxx

340 Péter Ozsvárt et al.
This conglomerate is interpreted to be the product of
the dismantling of a platform. It is followed by 15 m
of black Middle Triassic (Anisian?) calciturbidites,
reworked foraminifers, corals (Permian?), algae and
megalodontid-type bivalves. The series continues with
a brownish medium-bedded Upper Triassic (Carnian?)
wackestone containing echinoderms, ostracods and fo-
raminifera typical for reef environments which could
be compared to assemblages described from Cyprus
by Martini et al. (2009). The platform ends with an
irregular surface made of reef limestone showing syn-
sedimentary faulting.
Fig. 3. Schematic geological map of the studied area, showing the location of the Sorgun and Hacialanı mélanges, and the
main blocks. Modified from Moix et al. (2011).
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New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange 341
The palaeo-surface is covered by discontinuous
pink micritic nodular limestones in Hallstatt Limestone
facies which yielded ammonoids (Austrotrachyceras
austriacum Zone), foraminifera, echinoderms, cri-
noids, fish remains, brachiopods and conodonts of
middle Carnian age. Near Tavusçayırı Tepe in the
SOM, these levels were erroneously assigned to the
Early Jurassic by ParlaK & robertson (2004).
On the reference section, the Hallstatt Limestone is
conformably overlain by 130 m of thin bedded Huğlu-
type re-deposited green tuffites showing flute-casts,
load-casts and also Bouma sequences. The geochemi-
cal signature is of arc-type (VAB), suggesting a pos-
sible derivation from an eroding arc. The thickness
of these deposits varies from section to section, and
they may be associated with highly altered brownish
lavas and tuffs separated by numerous faults. The tuf-
fites may have acted as a preferential level for inter-
slicing or thrusting. The tuffitic series is interspersed
with alternations of micritic limestone and calcitur-
bidites. One micritic limestone level (Sample G11)
contains conodonts, sponge spicules, ostracods and a
well-preserved radiolarian fauna of the early Tuvalian
Spongotortilispinus moixi Zone (Kozur et al. 2007a,
b, c, 2009; Moix et al. 2007b).
The tuffitic episode is followed by 300 m of pelagic
limestone, calciturbidites, bioclastic limestone and de-
bris-flows. The pelagic limestone sedimentation usu-
ally started during the upper Carnian, continues dur-
ing the Norian and most probably ended during the
early Rhaetian (?). The early Rhaetian (?) limestone
is overlain by a breccia, followed by late Bajocian
brownish radiolarian cherts.
The succession of the Tavusçayırı block corre-
sponds to the Huğlu-type sequences described by
Özgul (1976) in the Bozkır Units and by MonoD
(1977) in the Beyşehir-Hoyran Nappes. All the taxa
described in the next section are from a single ra-
diolarian-rich sample (G11) belonging to the lower
Tuvalian Spongotortilispinus moixi radiolarian zone
(Moix et al. 2007).
4. The radiolarian fauna
Sample G11 contained new material of the genus
Triassobullasphaera. Due to the excellent preserva-
tion, all specimens present have a relatively large,
circular opening surrounded by a small tubular exten-
sion (pylome?). This opening may represent a relevant
structure of the skeleton for the axoflagellum shaping
Fig. 4. Synthetic lithostratigraphic section of the Tavusçayırı
block in the Sorgun Ophiolitic Mélange. Adapted from
Moix et al. (2011).
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342 Péter Ozsvárt et al.
Fig. 5.
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New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange 343
the general skeletal morphology. Kozur & Mostler
(2006) assumed that Triassobullasphaera might be-
long to the collodarians, and that they appeared in
the fossil record much earlier than previously thought
(Tertiary). Recent collodarians are the only colonial
lifestyle radiolarians. This group wears distinct globu-
lar latticed shells with relatively large pores surround-
ed by tubular extensions or spines (De Wever et al.
2001). This group is particularly important in the fossil
record because its large biomass and bioproductivity
may have effected organic carbon production in an-
cient oceans (isHitani et al. 2012).
The oldest known unequivocal collodarian organ-
isms appeared in the fossil record around the Middle
Eocene. Kozur & Mostler (2006) documented a new
putative collodarian species (Triassobullasphaera
hemisphaerica Kozur & Mostler, 2006) from the
well preserved and diverse Longobardian (Ladinian,
Middle Triassic) radiolarian fauna from the Varoški
Creek, Fojnica (Bosnia and Herzegovina). This un-
certain classification is based on the incomplete open-
ing of the test. They assumed that this opening was
filled, but it cannot be clearly seen on the illustrated
pictures. More recently, this genus was classified as
a member of the family Xiphostylidae HaecKel, 1881
(Spumellaria) by o’DogHerty et al. (2009).
Additional important species from sample G11
are represented by Parentactinia pugnax DuMitrica
and Pentactinorbis pessagnoi Kozur & Mostler, be-
cause these species were only known from the Anisian
(Illyrian). Originally, these species were reported
from the “Buchenstein-Schichten” of the Vicentinian
Alps and from Felsőörs, Balaton Highland (Hungary).
However, many comprehensive studies have been
published from Anisian, Ladinian and lower Carnian
around the world, and these “Lazarus taxa” disap-
peared from the palaeontological record until the
lower Tuvalian. They appeared again in the lower
Tuvalian assemblage from sample G11. Consequently,
their range can be updated and now expands from
the lower Illyrian (Tetraspinocyrtis laevis Radiolaria
Zone) to the lower Tuvalian (Spongotortilispinus moixi
Radiolaria Zone).
An additional important and interesting palaeon-
tological element among the radiolarians from sam-
ple G11 is the Hindeosphaeridae group. Indeed, the
secondary spines appear on the polar spines at the
same time and are present in two different taxa (e.g.,
Spinostylosphaera n. gen., Spinoparasepsagon n.
gen.). It is the first time that this feature is recorded in
this group. The presence of secondary spines could be
related to environmental stress (e.g. salinity or other
chemical parameters or palaeoclimatic changes such
as the Middle Carnian wet intermezzo) or to genetic
modifications.
In addition, some taxa from the Hindeosphaeridae
group (e.g., Spinostylosphaera andrasi n. gen. n. sp.)
represent good examples for rapid evolution (see Fig.
5.18-5.23) where the size of the secondary spines in-
crease quickly. This is demonstrated by the fact that
all specimens illustrated were extracted from the same
sample (G11).
Fig. 5. 1 – Archaeocenosphaera clathrata (Parona, 1890), INV 2014.78.5. 2 – Archaeocenosphaera parvispinosa (Kozur,
Krainer & Mostler, 1996), INV 2014.79. 3 – Triassobullasphaera hemisphaerica Kozur & Mostler, 2006, INV 2014.80.1.
4 – Triassobullasphaera hemisphaerica Kozur & Mostler, 2006, INV 2014.80.2. 5 – Triassobullasphaera miriae n. sp.,
holotype, PAL 2014.125.1. 6 – Triassobullasphaera miriae n. sp., paratype, PAL 2014.125.2. 7 – Parentactinia pugnax
DuMitr ica, 1978, INV 2014.81. 8 – Pentactinocarpus acanthicus DuMitrica, 1978, INV 2014.82.1. 9 – Pentactinocarpus
acanthicus Du Mitrica, 1978, INV 2014.82.2. 10: Pentactinocarpus acanthicus DuMitrica, 1978, INV 2014.82.3. 11 –
Pentactinocarpus tetracanthus DuMitrica, 1978, INV 2014.83.1. 12 – Pentactinocarpus tetracanthus DuMitrica, 1978,
INV 2014.83.2. 13 – Pentactinocarpus tetracanthus DuMitrica, 1978, INV 2014.83.3. 14 – Pentactinocarpus magnus
(Kozur & Mostler, 1979), INV 2014.84. 15 – Pentactinorbis pessagnoi Kozur & Mostler, 1994, INV 2014.85. 16 –
Pentactinorbis cf. megasphaera (sugiyaMa, 1997), INV 2014.86 17 – Hexapylomella carnica Kozur & Mostler, 1979, INV
2014.87. 18 – Hindeosphaera foremanae Kozur & Mostler, 1979, INV 2014.89. 19 – Hindeosphaera naomiae n. sp., holo-
type, PAL 2014.134. 20 – Hindeosphaera djani n. sp., holotype, PAL 2014.135. 21 – Hindeosphaera burrii n. sp., paratype,
PAL 2014.136.3.22 – Hindeosphaera burrii n. sp., paratype, PAL 2014.136.2. 23 – Hindeosphaera burrii n. sp., holotype,
PAL 2014.136.1.24 – Hindeosphaera cf. goestlingensis Kozur & Mostler, 1979, INV 2014.90.2. 25 – Hindeosphaera sp.,
INV 2014.91.1.
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344 Péter Ozsvárt et al.
5. Systematic palaeontology
Order Spumellaria eHrenberg, 1875
Family Xiphostylidae HaecKel, 1881
Genus Archaeocenosphaera Pessagno & yang, 1989
Type species: Archaeocenosphaera ruesti Pessagno &
yang, 1989.
Archaeocenosphaera clathrata (Parona, 1890)
Fig. 5.1
1880 ?Heliosphaera echinoidites n. sp. – Pantanelli, p.
46, fig. 8.
1890 Cenosphaera clathrata n. sp. – Parona, p. 19, pl. 1,
fig. 5.
1892 Cenosphaera clathrata Parona. – Parona, pl. 1, fig.
2.
1892 Cenosphaera apiaria n. sp. – rüst, p. 135, pl. 6, fig.
6.
1979 Cenosphaera clathrata Parona. – Kozur &
Mostler, p. 69, pl. 4, fig. 1.
1984 Cenosphaera clathrata Parona. – laHM, p. 15, pl. 1,
figs. 1-2.
1989 Archaeocenosphaera ruesti n. sp. – Pessagno &
yang, p. 203, pl. 1, fig. 9; pl. 9, fig. 23.
1996 Cenosphaera clathrata Parona. – cHiari et al., pl. 1,
fig. 19.
1996 Cenosphaera parvispinosa n. sp. – Kozur et al., pp.
222-223, pl. 9, fig. 12.
1997 Cenosphaera sp. A. – yao, pl. 1, figs. 2, 4-5, 11.
1997 Archaeocenosphaera clathrata (Parona). –
DuMitr ica et al., p. 17, pl. 1, fig. 2.
2000 Cenosphaera igoi n. sp. – sasHiDa et al., p. 804, fig.
10.7-10.8.
2009 Archaeocenosphaera clathrata (Parona). – Feng et
al., p. 585, fig. 3(1-4).
2011 Archaeocenosphaera sp. – tHassanaPaK et al., p.
188. fig. 4C, D.
Occurrence: Lower Devonian of the southern Ural, Anisian
to Ladinian from Italy, Austria, Hungary and Thailand;
Carnian of Albania, Lower Tuvalian of the Sorgun
Ophiolitic Mélange, Turkey and Middle Late Triassic of
Tibet. The species is also recorded from the Berriasian in
Masirah Island, Oman and from the Eocene in Italy.
Remarks: The original illustration of A. clathrata (Parona,
1890) is very schematic. Therefore, it is almost impossible
to identify the original structure of the cortical shell and
the pore frames of the holotype. The specimen illustrated
by Parona wears relatively large and hexagonal pores. The
specimen from the Sorgun Ophiolitic Mélange differs in
having circular, hexagonal and polygonal pores.
Archaeocenosphaera parvispinosa (Kozur, Krainer
& Mostler, 1996)
Fig. 5.2
1996 Cenosphaera parvispinosa n. sp. – Kozur et al., pp.
222-223, pl. 9, fig. 13.
2002 Cenosphaera clathrata Parona. – suzuKi et al., p.
170, fig. 4C.
Occurrence: Basal Tuvalian of the Sorgun Ophiolitic
Mélange, southern Turkey, Upper Anisian and Lower
Ladinian of Austria, Hungary and Italy, Lower Jurassic of
N Peru.
Genus Triassobullasphaera Kozur & Mostler, 2006
Type species: Triassobullasphaera hemisphaerica Kozur
& Mostler, 2006.
Triassobullasphaera hemisphaerica Kozur &
Mostler, 2006
Fig. 5. 3 -5.4
2006 Triassobullasphaera hemisphaerica n. sp. – Kozur
& Mostler, p. 42, pl. 1, figs. 5-6, 8, 11-12.
Remarks: The original description of Triassobullasphaera
hemisphaerica Kozur & Mostler (2006) stated clearly
that the opening of the test is completely missing, although
these authors supposed the presence of an opening (ap-
erture). In contrast, all the investigated specimens from
the Sorgun Ophiolitic Mélange present a relatively large,
well-defined circular opening. While the specimens of
Triassobullasphaera miriae n. sp. have small tubular ex-
tensions around the opening, T. hemisphaerica Kozur &
Mostler, 2006 has no such extensions.
Occurrence: Upper Longobardian (Bosnia and
Herzegovina) to Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
Triassobullasphaera miriae n. sp.
Fig. 5.5-5.6
Etymology: In honor of natHalie Miria scHMiDHauser.
Holotype: The specimen illustrated in Fig. 5.5.
Material: More than 50 specimens.
Diagnosis: Spherical or slightly ellipsoidal, single lay-
ered shell with polygonal to mostly circular pore frames.
Meshwork with elevated nodes (Diameter = 20-30 μm). The
hemisphere-like nodes are positioned uniformly and rela-
tively densely on the test. 12 to 15 nodes can be seen on one
half of the test. Shell bearing relatively large (diameter =
30-40 μm), well-defined, circular opening surrounded by a
small tubular extension.
Occurrence: Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
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New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange 345
Remarks: Triassobullasphaera miriae n. sp. differs from
T. hemisphaerica Kozur & Mostler, 2006 by the existing
opening, the smaller sized and the more densely located
nodes.
Order Entactinaria Kozur & Mostler, 1982
Family Palaeoscenidiidae rieDel, 1967
Genus Parentactinia DuMitrica, 1978
Type species: Parentactinia pugnax DuMitr ica, 1978.
Parentactinia pugnax DuMitrica, 1978
Fig. 5.7
1978 Parentactinia pugnax n. sp. – DuMitrica, p. 50, pl. 4,
figs. 4?, 5, pl. 5, figs. 1-3.
1990 Parentactinia pugnax DuMitrica. – yeH, p. 13, pl. 4.
fig.19.
1990 Parentactinia pugnax Dumitrica. – Goričan &
buser, p. 149, pl. 7, fig. 6.
1994 Parentactinia pugnax DuMitrica. – Kozur &
Mostler, p. 45, pl. 1, figs. 11-12.
1995 Parentactinia pugnax DuMitrica. – raMovs &
Goričan, p. 187, pl. 4, fig. 7.
Occurrence: ?Olenekian of New Zealand, Middle Triassic
(Illyrian – ?Fassanian) in the southern Alps, Hungary and
the Philippines. Upper Triassic (lower Tuvaluan) of the
Sorgun Ophiolitic Mélange, S Turkey.
Family Pentactinocarpidae DuMitrica, 1978
Genus Pentactinocarpus DuMitrica, 1978
1979 Praedruppatractylis n. gen. – Kozur & Mostler, p.
82.
1979 Oertlisphaera n. gen. – Kozur & Mostler, p. 53.
Type species: Pentactinocarpus fusiformis DuMitrica,
1978.
Pentactinocarpus acanthicus DuMitrica, 1978
Fig. 5. 8-5.10
1978 Pentactinocarpus acanthicus n. sp. – DuMitrica, p.
44, pl. 3, fig. 3.
1980 Pentactinocarpus acanthicus DuMitrica. –
DuMitr ica et al., p. 7, pl. 4, fig. 7.
1984 Pentactinocarpus acanthicus DuMitrica. – laHM, p.
22, pl. 2, figs. 9-10.
1990 Pentactinocarpus acanthicus Dumitrica. – Goričan
& buser, p. 149, pl. 7, fig. 12.
1994 Pentactinocarpus acanthicus DuMitrica. – Kozur
& Mostler, p. 46, pl. 2, figs. 3, 5.
1999 Pentactinocarpus acanthicus DuMitrica. – teKin, p.
133, pl. 27, fig. 6.
2000 Pentactinocarpus acanthicus DuMitr ica. – carter
& orcHarD, pl. 2, fig. 4.
2012 Pentactinocarpus acanthicus DuMitr ica. – stocKar
et al., p. 388, pl. 1, figs. 8-10.
Occurrence: Middle to Late Triassic in the Tethyan Realm.
Pentactinocarpus tetracanthus DuMitrica, 1978
Fig. 5.11-5.13
1978 Pentactinocarpus tetracanthus n. sp. – DuMitrica,
p. 43, pl. 1, fig. 1.
1979 Sethophaena (?) sp. A. – naKaseKo & nisHiMu ra, p.
79, pl. 8, fig. 7.
1980 Pentactinocarpus tetracanthus DuMitrica. –
DuMitr ica et al., p. 8, pl. 4, figs. 1-4
1981 Pentactinocarpus austriacus n. sp. – Kozur &
Mostler, p. 20, pl. 53, fig. 4.
1984 Pentactinocarpus tetracanthus DuMit rica. – laHM,
p. 23, pl. 2, fig. 11;
1990 Pentactinocarpus tetracanthus DuMitrica. –
Goričan & Buser, p. 150, pl. 7, figs. 8-10;
1994 Pentactinocarpus tetracanthus DuMit rica. – Kozur
& Mostler, p. 46, pl. 2, figs. 6-7, 13.
1995 Pentactinocarpus tetracanthus DuMitrica. –
Kellici & De Wever, p. 153, pl. 3, fig. 20.
1997 Pentactinocarpus tetracanthus DuMitrica. –
sugiyaMa, p. 184. fig. 49-23.
1999 Pentactinocarpus sp. af f. P. tetracanthus DuMitrica.
– bragin & Krylov, p. 544. fig. 7D.
1999 Pentactinocarpus tetracanthus DuMitrica. – teKin,
p. 134, pl. 27, figs. 9-10.
2006 Pentactinocarpus tetracanthus DuMitrica. – et al.,
pl. 2, fig. 30.
2007 Pentactinocarpus tetracanthus DuMitrica. –
bragin, p. 967, pl. 1, fig. 1.
2010 Pentactinocarpus tetracanthus DuMitrica. – teKin
& sÖnMez, fig. 8N-O.
Occurrence: Middle to Late Triassic of the Tethyan Realm.
Pentactinocarpus magnus (Kozur & Mostler, 1979)
Fig. 5.14
1979 Oertlisphaera magna n. sp. – Kozur & Mostler, p.
53, pl. 10, fig. 1.
1990 Pentactinocarpus magnus (Kozur & Mostler). –
yeH, p. 13, pl. 4, fig. 22, pl. 5, fig. 17, pl. 11, fig. 19.
Occurrence: Middle (Ladinian) to Late (Carnian) Triassic
of the Philippines, Carnian in the Tethyan Realm.
Genus Pentactinorbis DuMitrica, 1978
Type species: Pentactinorbis kozuri DuMitrica, 1978.
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346 Péter Ozsvárt et al.
Pentactinorbis pessagnoi Kozur & Mostler, 1994
Fig. 5.15
1994 Pentactinorbis pessagnoi n. sp. – Kozur & Mostler,
p. 48, pl. 4, figs. 3-4.
Occurrence: Illyrian of the Balaton Highland, Hungary and
Upper Triassic (lower Tuvalian) from the Sorgun Ophiolitic
Mélange, S Turkey.
Pentactinorbis cf. P. megasphaera (sugiyaMa, 1997)
Fig. 5.16
Remarks: The specimen resembles P. megasphaera but it
has larger circular to polygonal pore frames. Small tricari-
nate (?) spines connected to cortical shell. Inner structure
is hidden.
Family Hexapylomellidae Kozur & Mostler, 1979
Genus Hexapylomella Kozur & Mostler, 1979
Type species: Hexapylomella carnica Kozur & Mostler,
1979.
Hexapylomella carnica Kozur & Mostler, 1979
Fig. 5.17
1979 Hexapylomella carnica n. sp. – Kozur & Mostler,
p. 69. pl. 3, fig. 2.
Occurrence: Carnian in the Tethyan Realm.
Family Hindeosphaeridae Kozur & Mostler, 1981
Genus Hindeosphaera Kozur & Mostler, 1979
Type species: Hindeosphaera foremanae Kozur &
Mostler, 1979.
Hindeosphaera burrii n. sp.
Fig. 5. 21-5. 23
Etymology: In honor of georges burri.
Holotype: The specimen illustrated in Fig. 5.23.
Material: More than 100 specimens.
Diagnosis: Large, spherical or ellipsoidal, double-layered
cortical shell with circular and mostly polygonal pore
frames on outer layer and very small, circular pores on in-
ner layer. Thick tricarinate, straight or twisted lateral spines
with wide ridges and relatively deep grooves. In addition,
4-5 long (200-250 μm), tricarinate and 2-3 needle-like,
pointed spines connected to the cortical shell. Spherical or
subspherical medullary shell with large mostly polygonal
or circular pore frames. Reduced median bar with three
needle-like or tricarinate apical spines. Two lateral spines,
one of them tricarinate that continues into large spine on the
cortical shell.
Occurrence: Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
Remarks: Hindeosphaera burrii n. sp. is distinguished
from all other species of the genera Hindeosphaera by hav-
ing long spines.
Hindeosphaera djani n. sp.
Fig. 5.20
Etymology: In honor of Djan Moix.
Holotype: The specimen illustrated in Fig 5.20.
Material: More than 5 specimens.
Diagnosis: Spherical or slightly compressed, double lay-
ered cortical shell with circular to polygonal pore frames
on the outer layer and very small pores on the inner layer.
Medullary shell spherically connected with 5 to 6 spines
Fig. 6. 1 – Pseudostylosphaera dumitricai n. sp., holotype, PAL 2014.128.1. 2 – Pseudostylosphaera dumitricai n. sp.,
paratype, PAL 2014.128.2. 3 – Pseudostylosphaera dumitricai n. sp., paratype, PAL 2014.128.3. 4 – Pseudostylosphaera
dumitricai n. sp., paratype, PAL 2014.128.4. 5 – Presumably pathological specimen of Pseudostylosphaera cf. P. dumi-
tricai n. sp., paratype, PAL 2014.128.5. 6 – Pseudostylosphaera longispinosa Kozur & Mostler, 1981, INV 2014.88. 7 –
Spinostylosphaera andrasi n. gen. n. sp., holotype, PAL 2014.126.1. 8 – Spinostylosphaera andrasi n. gen. n. sp., paratype,
PAL 2014.127.1. 9 – Spinostylosphaera andrasi n. gen. n. sp., paratype, PAL 2014.126.2. 10 – Spinostylosphaera andrasi
n. gen. n. sp., paratype, PAL 2014.126.3. 11 – Spinostylosphaera andrasi n. gen. n. sp., paratype, PAL 2014.127.2. 12.1 –
Spinostylosphaera andrasi n. gen. n. sp., paratype, PAL 2014.127.3.6. 12.2 – Spinostylosphaera andrasi n. gen. n. sp., para-
type, PAL 2014.127.3. 13.1 – Spinostylosphaera vachardi n. gen. n. sp., paratype, PAL 2014.131.2. 13.2 – Spinostylosphaera
vachardi n. gen. n. sp., paratype, PAL 2014.131.2. 14 – Spinostylosphaera vachardi n. gen. n. sp., paratype, PAL 2014.131.2.
15 – Spinostylosphaera vachardi n. gen. n. sp., holotype, PAL 2014.131.1.
eschweizerbart_xxx

New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange 347
Fig. 6.
eschweizerbart_xxx

348 Péter Ozsvárt et al.
to the cortical shell, which continue in outer spines. Two
lateral spines are three or four-bladed and one of them
continues in a massive, large, twisted spine. The others are
needle-like and continue in smaller, pointed spines (Fig.
6.22). Arch of medullary shell connected in nodes around
lateral and apical spines. Sagittal ring formed in central part
of the medullary shell and its plan is perpendicular to the
direction of lateral spines. The most characteristic element
of the species is the very large, massive spine connected to
the cortical shell. It is four-bladed with deep grooves and
high ridges. Terminal part of the spine is expanded and the
connecting edges of ridges are slightly curved.
Occurrence: Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
Remarks: Hindeosphaera djani n. sp. is distinguished from
all other species of the genera Hindeosphaera by the mas-
sive spines with four ridges, deep grooves and expanded
terminal part.
Hindeosphaera foremanae Kozur & Mostler, 1979
Fig. 5.18
1979 Hindeosphaera foremanae n. sp. – Kozur &
Mostler, p. 62, pl. 5, fig. 6.
1984 Hindeosphaera foremanae Kozur & Mostler. –
laHM, p. 38, pl. 5, figs. 11-12.
Occurrence: Carnian of the Tethyan Realm.
Hindeosphaera naomiae n. sp.
Fig. 5.19
Etymology: In honor of naoMi taFelMacHer.
Holotype: The specimen illustrated in Fig. 5.19.
Material: More than 5 specimens.
Diagnosis: Spherical or subspherical double-layered corti-
cal shell. Small circular or polygonal pore frames on inner
layer which is covered by spongy mantle. The outer spongy
mantle is in nodes. Two approximately equal, tricarinate,
twisted spines and 3-4 significantly shorter, tricarinate,
pointed spines connected to the cortical shell.
Occurrence: Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
Remarks: Hindeosphaera naomiae n. sp. is distinguished
from all other species of Hindeosphaera by having two
identical tricarinate and twisted lateral spines.
Hindeosphaera cf. H. goestlingensis Kozur &
Mostler, 1979
Fig. 5.24
Remarks: The cortical shell is spherical with polygonal
pore frames and one significantly larger, tricarinate, twisted
spine and three smaller pointed, tricarinate spines. Due to
the good preservation, the inner structure can be studied.
Very small median bar at the pointed part of the medul-
lary shell with three, needle-like apical spines (Fig. 6.18).
Two lateral spines, one of them tricarinate and continues
in a large, strongly twisted spine which is connected to the
cortical shell. The other spine is significantly thinner, nee-
dle-like and continues into a smaller, tricarinate outer spine.
The medullary shell is ellipsoidal with large polygonal pore
frames.
Hindeosphaera sp.
Fig. 5.25
Remarks: This specimen resembles H. austriaca Kozur &
Mostler, 1979 and H. goestlingensis Kozur & Mostler,
1979. Because of the broken shell, it cannot be assigned ac-
curately. The cortical shell is globular and double-layered.
The outer layer bears mostly polygonal pore frames, the
inner layer has very small circular pores. The medullary
shell is ellipsoidal or irregular shaped, with large polygonal
pore frames. Three slender, slightly curved apical spines.
Two massive, tricarinate, lateral spines, which continue into
larger spines that are connected to the cortical shell.
Genus Pseudostylosphaera Kozur & Mostler, 1981
Type species: Pseudostylosphaera gracilis Kozur &
Mostler, 1981.
Pseudostylosphaera dumitricai n. sp.
Fig. 6.1-6.5.
Etymology: In honor of Dr. Paulia n DuMitrica, for his out-
standing work on radiolarian taxonomy.
Holotype: The specimen illustrated in Fig. 6.1.
Material: More than 100 specimens.
Diagnosis: Test is composed of spherical cortical shell
and two, equal, twisted polar spines. The cortical shell is
double-layered: outer layer with polygonal pore frames, in-
ner layer contains small, circular pores. The shell includes
many, radial, pointed, needle-like spines (length 40-70 μm).
Relatively small (diameter = 50-55 μm), ellipsoidal med-
ullary shell with large, circular or polygonal pore frames.
In the specimen illustrated, the median bar is hidden by
sediment or calcite (Fig. 6.2). Poorly preserved apical and
ventral spines. Three or four rod-shaped basal spines. Two,
three-bladed, massive lateral spines connected to the med-
ullary shell which continue in polar spines. Pointed polar
spines are long, straight or slightly curved and twisted, tri-
carinate with wide ridges and deep or narrow grooves.
eschweizerbart_xxx

New Carnian (Upper Triassic) radiolarians from the Sorgun Ophiolitic Mélange 349
Occurrence: Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
Remarks: The new species describe here differs from
Pseudostylosphaera multispinata teKin & Mostler, 2005
in the shape of the tricarinate polar and radial spines
(strongly twisted, thin ridges, less radial).
Pseudostylosphaera longispinosa Kozur & Mostler,
1981
Fig. 6.6
1981 Pseudostylosphaera longispinosa n. sp. – Kozur &
Mostler, p. 32, pl. 1, fig. 6.
1981 Pseudostylosphaera longobardica n. sp. – Kozur &
Mostler, p. 33, pl. 49, fig. 3.
1984 Pseudostylosphaera longispinosa Kozur &
Mostler. – laHM, pp. 34-35, pl. 4, figs. 11-12.
1986 Pseudostylosphaera longispinosa Kozur &
Mostler. – Kozur & réti, fig. 6D
1990 Pseudostylosphaera longispinosa Kozur &
mostler. – Goričan & Buser, p. 155, pl. 5, fig. 4.
1990 Pseudostylosphaera longispinosa Kozur &
Mostler. – Kolar-jurKovšeK, p. 76, pl. 5, fig. 6a-b.
1990 Pseudostylosphaera longispinosa Kozur &
Mostler. – yeH, p. 15, pl. 4, fig. 2.
1997 Pseudostylosphaera longispinosa Kozur &
Mostler. – sugiyaMa, p. 188, fig. 48.16.
1998 Pseudostylosphaera longispinosa Kozur &
Mostler. – taKaHasHi et al., pl. 1, fig. 4.
1999 Pseudostylosphaera longispinosa Kozur &
Mostler. – teKin, p. 129., pl. 25, fig. 14.
2001 Pseudostylosphaera longispinosa Kozur &
Mostler. – Feng et al. p. 190, pl. 5, figs. 22-24.
2006 Pseudostylosphaera longispinosa Kozur &
Mostler. – Marquez et al., pl. 2, fig. 13.
2007 Pseudostylosphaera longispinosa Kozur &
Mostler. – teKin & gÖncüoglu, pl. 2, fig. 25.
2010 Pseudostylosphaera longispinosa Kozur &
Mostler. – teKin & sÖnMez, fig. 7R.
2011 Pseudostylosphaera longispinosa Kozur &
Mostler. – tHassanaPaK et al., p. 195, fig. 6T.
2012 Pseudostylosphaera longispinosa Kozur &
Mostler. – ozsvárt et al., fig. 9.7.
2012 Pseudostylosphaera longobardica Kozur &
Mostler. – ozsvárt et al., fig. 9.8.
2012 Pseudostylosphaera longispinosa Kozur &
Mostler. – stocKar et al., p. 397, pl. 4, figs. 16-24.
Occurrence: Anisian to Norian of the Tethyan Realm.
Remarks: The original description of P. longispinosa
Kozur & Mostler, 1981 does not contain any information
about tiny, pointed and needle-like spines on the outer pore
frames. This specimen bears several very short spines (15-
20 μm), but most of them are probably broken.
Genus Spinostylosphaera nov.
Etymology: In allusion to its similarities to the genera
Pseudostylosphaera, but with secondary spines on the po-
lar spines.
Type species: Spinostylosphaera andrasi n. gen. n. sp.
Included species: Spinostylosphaera andrasi n. gen. n. sp.,
Spinostylosphaera mesotriassica (DuMitrica, Kozur &
Mostler, 1980), Spinostylosphaera vachardi n. gen. n. sp.
Diagnosis: Relatively small spherical to ellipsoidal, single
layered cortical shell. From tiny to relatively large-sized
pentagonal to mostly polygonal pore frames. Two long,
three bladed polar spines are broad or wear furrows. They
can be straight or twisted. Polar spines wear three pointed,
straight or distally slightly curved secondary spines. Short
to long terminal spine present, slightly curved, proximally
tricarinate, distally pointed.
Remarks: The genus Spinostylosphaera differs from the
Pseudostylosphaera by its single layered cortical shell and
by having three secondary spines on the polar spines. These
secondary spines are unknown among the species of the ge-
nus Pseudostylosphaera.
Occurrence: From Middle Triassic (?) of the southern Alps
to Late Triassic (Julian to lower Tuvalian) in the northern
Alps and the Sorgun Ophiolitic Mélange, S-Turkey.
Spinostylosphaera andrasi n. gen. n. sp.
Fig. 6.7-6.12
Etymology: For anDrás ozsvárt, Luxembourg.
Holotype: The specimen illustrated in Fig. 6.7
Material: More than 50 specimens.
Diagnosis: Relatively small, spherical to ellipsoidal, sin-
gle layered cortical shell. Pentagonal to mostly polygonal
pore frames. Presumably, with a small spherical or ellip-
soidal medullary shell. Initial spicule hardly visible, very
small median bar with one or two apical and basal spines.
Two, equal, tricarinate polar spines with three, pointed,
straight or distally slightly upward curved secondary spines.
Terminal spine very long, slightly curved, proximally tri-
carinate, distally pointed.
Occurrence: Lower Tuvalian of the Sorgun Ophiolitic
Mélange, S Turkey.
Spinostylosphaera vachardi n. gen., n. sp.
Fig. 6.13-6.15
2011 Dumitricasphaera simplex teKin. – bragin, p. 755,
pl. 10, fig. 6.
eschweizerbart_xxx

350 Péter Ozsvárt et al.
Etymology: In honor of Dr. Daniel vacHarD, Lille, for his
outstanding work on Palaeozoic microbiotas.
Holotype: The specimen in Fig 6.15.
Material: More than 50 specimens.
Diagnosis: Large, spherical single-layered cortical shell
with mostly polygonal pore frames. Two equal, tricarinate
polar spines with furrows and with three, pointed, straight
or distally slightly downward curved secondary spines.
Terminal spine long, slightly curved, proximally tricarinate,
distally pointed.
Occurrence: Upper Julian (Tetraporobrachia haeck-
eli Radiolaria Zone) of the Northern Calcareous Alps
(Großreifling section) and Lower Tuvalian of the Sorgun
Ophiolitic Mélange.
Remarks: Spinostylosphaera vachardi n. gen., n. sp. has
significantly longer polar spines and terminal spines than
Spinostylosphaera mesotriassica (DuMitrica, Kozur &
Mostler, 1980).
Acknowledgements
This paper is dedicated to the memory of Heinz W. Kozur,
who was an outstanding expert on radiolarians, conodonts,
ostracods, conchostracans and on the Triassic System
in general. This work was supported by the Hungarian
Scientific Research fund (OTKA K81298). Special thanks
are due to the anonymous reviewers for comments, to Dr.
noritosHi suzuKi for discussions on radiolarian taxonomy
and to Gloria Mouanga and the editor for reading and cor-
recting the manuscript. This is MTA-MTM-ELTE Paleo
Contribution No. 211.
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Manuscript received: September 8th, 2014.
Revised version accepted by the Bonn editor: March 21st,
2015.
Addresses of the authors:
Péter ozsvárt, MTA-MTM-ELTE, Research Group for
Paleontology, P. O. Box 137 H-1431, Budapest, Hungary;
e-mail: ozsi@nhmus.hu
Patrice Moix, Rue de la Combe 55, 1969 Eison, Switzerland;
e-mail: patricemoix@gmail.com
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