Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western Russia

Abstract

The recent checklist of the Ephydridae of Finland by Zatwarnicki and Kahanpää (2014) mentioned 13 ephydrid species as new for Finland without further details. This paper presents detailed records for those species and a few other species of interest. Four species are recorded for the first time from Russia. Trimerina
indistincta Krivosheina, 2004 is herein considered as a new junior synonym of Trimerina
microchaeta Hendel, 1932, syn. nov.

Full text

Biodiversity Data Journal 3: e4701
doi: 10.3897/BDJ.3.e4701
Taxonomic Paper
Notes on Shore Flies (Diptera: Ephydridae) from
Finland and north-western Russia
Jere Kahanpää , Tadeusz Zatwarnicki
‡ Zoology Unit, Museum of Natural History, University of Helsinki, Helsinki, Finland
§ Department of Biosystematics, Opole University, Opole, Poland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.fi)
Academic editor: Daniel Whitmore
Received: 09 Feb 2015 | Accepted: 25 Mar 2015 | Published: 30 Mar 2015
Citation: Kahanpää J, Zatwarnicki T (2015) Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-
western Russia. Biodiversity Data Journal 3: e4701. doi: 10.3897/BDJ.3.e4701
Abstract
The recent checklist of the Ephydridae of Finland by Zatwarnicki and Kahanpää (2014)
mentioned 13 ephydrid species as new for Finland without further details. This paper
presents detailed records for those species and a few other species of interest. Four
species are recorded for the first time from Russia. Trimerina indistincta Krivosheina, 2004
is herein considered as a new junior synonym of Trimerina microchaeta Hendel, 1932, syn.
nov.
Keywords
New records, new synonym, Trimerina, Scatophila iowana, faunistics.
Introduction
Finland is an exceptional country in having a record of regularly published checklists of the
order Diptera, including shore flies. Three of such lists have been published during the last
century: Frey (1941), Hackman (1980) and Kahanpää and Salmela (2014). The Diptera
‡ §
© Kahanpää J, Zatwarnicki T. This is an open access article distributed under the terms of the Creative Commons Attribution
License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author
and source are credited.

fauna of the country is well studied in comparison to most other countries, but far from
perfectly understood.
The most recent list of the Finnish shore fly fauna is Zatwarnicki and Kahanpää (2014) in
Kahanpää and Salmela (2014). The Finnish Ephydridae have attracted relatively little
attention: few Finnish dipterologists have studied them and no coordinated study of the
fauna has been made. The Finnish Museum of Natural History in Helsinki (MZH) holds by
far the largest ephydrid collection in the country. Existing faunal works are essentially
based on museum material collected between the years 1900 and 1960, mostly by sweep-
netting. After a long hiatus, interest in acalyptrate flies resurged in the 21st century among
Finnish amateur entomologists. The newer ephydrid material generally remains in private
collections.
Zatwarnicki (1997) included a list of the species known from the country by the mid-1990s.
Twelve species have been added to the fauna after 1997. Chronologically first is Notiphila
pollinosa Krivosheina by Krivosheina (1998), then Calocoenia paurosoma (Sturtevant &
Wheeler) and Parydra arctica Clausen by Krivosheina (2000), Coenia curvicauda (Meigen)
by Haarto and Winqvist (2002), Gymnoclasiopa pulchella (Meigen) by Winqvist (2002),
Notiphila major Stenhammar by Kahanpää and Winqvist (2003), Trimerina microchaeta
Hendel (as T. indistincta Krivosheina) by Krivosheina (2004), Philotelma defectum
(Haliday) by Mathis et al. (2009), Atissa pygmaea (Haliday) by Winqvist (2011), and
Hecamedoides unispinosus (Collin), Parydra mitis (Cresson) and P. nigritarsis Strobl by
Kahanpää (2013). The identity of the North European Glenanthe species was resolved by
Zatwarnicki and Mathis (2012).
Thirteen additional species of Ephydridae were recorded for the first time from Finland in
the recent checklist (Zatwarnicki and Kahanpää 2014). The checklist format did not allow
for the inclusion of record data. This paper provides the previously unpublished records on
which the checklist was based. Additionally, one new synonymy is proposed: Trimerina
indistincta Krivosheina, 2004 is treated as a junior synonym of Trimerina microchaeta
Hendel, 1932 (syn. nov.). After inclusion of all additions and corrections to the fauna, the
number of shore fly species recorded from Finland is now 112.
Some material from north-western Russia, adjacent to the Russian-Finnish border, was
also examined during the preparation of the Finnish checklist. Four species are recorded
from Russia for the first time.
Materials and methods
All the specimens discussed in this paper are deposited at the Zoological Museum of the
Finnish Museum of Natural History in Helsinki (MZH) unless otherwise specified. Most
MZH specimens have unique specimen identifiers in the form of a URL (Uniform Resource
Locator). The URL can be used as a web address: it points to a web page with details of
the sample in question. While we cannot guarantee that the web content is available in the
future, the address (URL) will remain unique. Standard abbreviations are used for the
2Kahanpää J, Zatwarnicki T

Finnish biogeographical provinces; see the Fauna Entomologica Scandinavica book series
or Heikinheimo and Raatikainen (1971) and Heikinheimo and Raatikainen (1981) for
details and maps.
Finnish insect collectors usually use a national uniform grid system (yhtenäiskoordinaati
sto, YKJ) for mapping and labeling purposes. The Darwin Core (DC) field
"verbatimCoordinates" is used for the YKJ coordinates printed on labels. The DC fields
"decimalLatitude" and "decimalLongitude" in the Taxon treatments section are given in
World Geodetic System 1984 (WGS84) decimal degrees. Where no collector-supplied
coordinates were available, the records were georeferenced by the senior author on the
basis of the collecting locality.
The identity of all included species is based on the study of primary types conducted by the
junior author, except for Hydrellia tarsata Haliday and Scatophila mesogramma (Loew), for
which interpretations are based on Collin (1966) and Sturtevant and Wheeler (1953)
respectively.
The order of species follows Zatwarnicki and Kahanpää (2014).
Taxon treatments
Trimerina microchaeta Hendel, 1932
Nomenclature
Trimerina microchaeta Hendel 1932: 11
= Trimerina indistincta Krivosheina 2004: 631 (syn. nov.)
Materials
a. scientificName: Trimerina microchaeta Hendel, 1932; order: Diptera; family: Ephydridae;
genus: Trimerina; specificEpithet: microchaeta; scientificNameAuthorship: Hendel, 1932;
country: Finland; stateProvince: EH; municipality: Hartola; locality: Hirtesalo;
verbatimElevation: 80; verbatimCoordinates: 6839:[3]450; decimalLatitude: 61.66;
decimalLongitude: 26.07; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 2002; month: 5; day: 15; habitat: a lake shore; individualCount: 1; sex: M; lifeStage:
adult; catalogNumber: JKA12-0653; recordedBy: Frey, Richard; associatedSequences:
BOLD:2709504; identifiedBy: Kahanpää, Jere; dateIdentified: 2012; institutionCode: Coll.
J. Kahanpää; basisOfRecord: PreservedSpecimen
b. scientificName: Trimerina microchaeta Hendel, 1932; order: Diptera; family: Ephydridae;
genus: Trimerina; specificEpithet: microchaeta; scientificNameAuthorship: Hendel, 1932;
country: Finland; stateProvince: ES; municipality: Joutsa; locality: Oravakivensalmi;
verbatimElevation: 80; verbatimCoordinates: 6845:[3]451; decimalLatitude: 61.71;
decimalLongitude: 26.08; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 2004; month: 6; day: 3; individualCount: 1; sex: F; lifeStage: adult; catalogNumber:
JKA12-0654; recordedBy: Frey, Richard; associatedSequences: BOLD:2709505;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 3

identifiedBy: Kahanpää, Jere; dateIdentified: 2012; institutionCode: Coll. J. Kahanpää;
basisOfRecord: PreservedSpecimen
Distribution
Widespread in Europe; also occurs in the Russian Far East (Kamchatka) (Zatwarnicki
2013).
Taxon discussion
After examination of primary types of Trimerina microchaeta Hendel and T. indistincta
Krivosheina we suggest to synonymize these two species. Krivosheina (2004) provided
three characters to separate T. indistincta from T. microchaeta: a gena-to-eye ratio of
0.17 (versus 0.20 for T. microchaeta), yellow mid and hind coxae (versus all coxae
partially dark), mid and hind tibiae entirely yellow (versus mid and hind tibiae with a
dark band). These characters separating the two species are not consistent and fall
within the variation range of T. microchaeta. Both nominal taxa have essentially the
same male terminalia with characteristic postgonostyli (=postsurstyli) bearing short and
apically rounded antero-ventral processes as illustrated for T. indistincta by Krivosheina
(2004).
a b
c d
Figure 1.
Ephydrid species from Finland. Scale bar = 1.0 mm.
a: Trimerina microchaeta. Holotype of Trimerina indistincta Krivosheina (specimen http://
id.luomus.fi/GV.4020)
b: Hydrellia mutata (http://id.luomus.fi/GV.16611)
c: Ditrichophora calceata (http://id.luomus.fi/GV.16561)
d: Pelina norvegica (http://id.luomus.fi/GV.19162)
4Kahanpää J, Zatwarnicki T

Notes
The holotype of Trimerina indistincta is shown in Fig. 1a.
Hydrellia albifrons Fallén, 1813
Materials
a. scientificName: Hydrellia albifrons Fallén, 1813; order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: albifrons; scientificNameAuthorship: Fallén, 1813;
country: Finland; stateProvince: V; municipality: Naantali; decimalLatitude: 60.47;
decimalLongitude: 22.47; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 20000;
georeferencedBy: Kahanpää, Jere; year: 1939; month: 8; day: 16; individualCount: 1; sex:
M; lifeStage: adult; preparations: gen. prep. on pin; catalogNumber: http://id.luomus.fi/
GV.16614; recordedBy: Tiensuu, Lauri; identifiedBy: Kahanpää, Jere; dateIdentified:
2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Hydrellia albifrons Fallén, 1813; order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: albifrons; scientificNameAuthorship: Fallén, 1813;
country: Finland; stateProvince: V; municipality: Naantali; decimalLatitude: 60.13;
decimalLongitude: 22.02; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 20000;
georeferencedBy: Kahanpää, Jere; year: 1949; month: 7; day: 1; individualCount: 1; sex:
F; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16615; recordedBy: Tiensuu,
Lauri; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as figured by Collin (1966) and
Canzoneri and Meneghini (1983).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widespread in
Europe; also known from North Africa and Pakistan (Zatwarnicki 1988, Zatwarnicki
2013).
Ecology
Larvae of this species feed on pondweeds (Potamogeton) (Zatwarnicki 1988), the
common frogbit (Hydrocharis morsus-ranae) (Hering 1926), and possibly also on water
plantains (Alisma plantago-aquatica) (see Deonier 1971).
Hydrellia cochleariae Haliday, 1839
Materials
a. scientificName: Hydrellia cochleariae Haliday, 1839; order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: cochleariae; scientificNameAuthorship: Haliday, 1839;
country: Finland; stateProvince: U; municipality: Helsinki; locality: Jollas; verbatimLocality:
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 5

Helsinge; decimalLatitude: 60.16; decimalLongitude: 25.07; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1864; month: 6; day: 20; individualCount: 4; sex:
MF; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16690; recordedBy: Palmén,
Johan Axel; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode:
MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Hydrellia cochleariae Haliday, 1839; order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: cochleariae; scientificNameAuthorship: Haliday, 1839;
country: Finland; stateProvince: U; municipality: Helsinki; locality: Jollas; verbatimLocality:
Helsinge; decimalLatitude: 60.16; decimalLongitude: 25.07; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1864; month: 6; day: 20; individualCount: 5; sex:
MF; lifeStage: adult; preparations: gen. prep. on pin; catalogNumber: http://id.luomus.fi/
GV.16691; recordNumber: 4096; recordedBy: Palmén, Johan Axel; identifiedBy:
Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
c. scientificName: Hydrellia cochleariae Haliday, 1839; order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: cochleariae; scientificNameAuthorship: Haliday, 1839;
country: Finland; stateProvince: V; municipality: Korppoo; locality: Jurmo;
decimalLatitude: 59.82; decimalLongitude: 21.60; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1960; month: 8; day: 8; individualCount: 1; sex:
MF; lifeStage: adult; preparations: gen. prep. on pin; catalogNumber: http://id.luomus.fi/
GV.16690; recordedBy: Meinander, Martin; identifiedBy: Kahanpää, Jere; dateIdentified:
2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as illustrated by Collin (1966).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Central and Western
Europe (Zatwarnicki 2013). The Finnish records are the first from Northern Europe.
Ecology
Reared from water soldiers (Stratiotes), Potamogeton (Fischer 1997), and water
starworts (Callitriche) (Hering 1957). Older, less reliable host records mention
Hydrocharis as an additional host (see Deonier 1971).
Notes
Previously misidentified as Hydrellia flavicornis (Fallén, 1823) in Finland (Hackman
1980).
Hydrellia fulviceps (Stenhammar, 1844)
Material
a. scientificName: Hydrellia fulvice ps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: fulviceps; scientificNameAuthorship:
6Kahanpää J, Zatwarnicki T

(Stenhammar, 1844); country: Finland; stateProvince: A; locality: Åland; verbatimLocality:
Aland; decimalLatitude: 60.2; decimalLongitude: 20.0; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 50000; georeferencedBy: Kahanpää, Jere;
individualCount: 1; sex: M; lifeStage: adult; preparations: gen. prep. on pin;
catalogNumber: http://id.luomus.fi/HT.808; recordNumber: 644; recordedBy: Palmén,
Johan Axel; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode:
MZH; basisOfRecord: PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as illustrated by Collin (1966).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widespread in
Europe, it is recorded also from the Russian Far East (Zatwarnicki 2013).
Ecology
No reliable records of the host plant of this species could be found.
Notes
Previously misidentified as Hydrellia chrysostoma (Meigen, 1830) (sensu Becker 1896)
in Finland (Hackman 1980).
Hydrellia laticeps (Stenhammar, 1844)
Materials
a. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 7; day: 9; individualCount: 1; sex:
M; lifeStage: adult; behavior: sitting on Juncus?; catalogNumber: http://id.luomus.fi/
GV.16624; recordNumber: 272; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 23; individualCount: 1; sex:
M; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16626; recordNumber: 1533;
recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 7

c. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 23; habitat: Phragmites
beds; individualCount: 1; sex: M; lifeStage: adult; preparations: gen. prep. on pin;
catalogNumber: http://id.luomus.fi/GV.16625; recordNumber: 1526; recordedBy: Frey,
Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
d. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 23; habitat: Phragmites
beds; individualCount: 1; sex: M; lifeStage: adult; preparations: gen. prep. on pin;
catalogNumber: http://id.luomus.fi/GV.16623; recordNumber: 1534; recordedBy: Frey,
Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
e. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 23; habitat: Phragmites
beds; individualCount: 1; sex: M; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16630; recordNumber: 1531; recordedBy: Frey, Richard; identifiedBy: Kahanpää,
Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
f. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 7; day: 6; individualCount: 1; sex:
M; lifeStage: adult; behavior: on Scirpus; catalogNumber: http://id.luomus.fi/GV.16632;
recordNumber: 127; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
g. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 22; habitat: Phragmites
beds; individualCount: 1; sex: M; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16631; recordNumber: 1496; recordedBy: Frey, Richard; identifiedBy: Kahanpää,
Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
8Kahanpää J, Zatwarnicki T

h. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 23; habitat: Phragmites
beds; individualCount: 1; sex: F; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16633; recordNumber: 1537; recordedBy: Frey, Richard; identifiedBy: Kahanpää,
Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
i. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 8; day: 15; habitat: Phragmites
beds; individualCount: 1; sex: F; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16634; recordNumber: 1204; recordedBy: Frey, Richard; identifiedBy: Kahanpää,
Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
j. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 8; day: 15; individualCount: 1; sex:
F; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16637; recordNumber: 1197;
recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
k. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 7; day: 6; individualCount: 1; sex:
F; lifeStage: adult; behavior: on Scirpus; catalogNumber: http://id.luomus.fi/GV.16628;
recordNumber: 126; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
l. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 7; day: 6; individualCount: 1; sex:
F; lifeStage: adult; behavior: on Scirpus; catalogNumber: http://id.luomus.fi/GV.16627;
recordNumber: 121; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
m. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 9

Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 7; day: 6; individualCount: 1; sex:
F; lifeStage: adult; behavior: on Scirpus; catalogNumber: http://id.luomus.fi/GV.16636;
recordNumber: 134; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
n. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 6; day: 22; habitat: Phragmites
beds; individualCount: 1; sex: F; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16635; recordNumber: 1498; recordedBy: Frey, Richard; identifiedBy: Kahanpää,
Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
o. scientificName: Hydrellia laticeps (Stenhammar, 1844); order: Diptera; family:
Ephydridae; genus: Hydrellia; specificEpithet: laticeps; scientificNameAuthorship:
(Stenhammar, 1844); country: Finland; stateProvince: U; municipality: Hanko; locality:
Tvärminne, Gloet; verbatimLocality: Tvärminne; decimalLatitude: 59.83;
decimalLongitude: 23.10; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 1000;
georeferencedBy: Kahanpää, Jere; year: 1932; month: 7; day: 9; individualCount: 1; sex:
F; lifeStage: adult; behavior: on Juncus?; catalogNumber: http://id.luomus.fi/GV.16629;
recordNumber: 269; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as illustrated by Collin (1966) and
Canzoneri and Meneghini (1983) under the name Hydrellia flaviceps (auct. nec
Meigen).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widespread in
Europe and probably across the Palaearctic Region, but poorly known (Zatwarnicki
2013).
Hydrellia mutata (Zetterstedt, 1846)
Materials
a. scientificName: Hydrellia mutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846);
country: Russia; stateProvince: Carelian Republic; municipality: Sortavala;
verbatimLocality: Sortavala; decimalLatitude: 61.7; decimalLongitude: 30.7;
geodeticDatum: wgs84; coordinateUncertaintyInMeters: 20000; georeferencedBy:
Kahanpää, Jere; year: 1936; month: 7; day: 1; verbatimEventDate: 1-VII 36;
individualCount: 1; sex: M; lifeStage: adult; behavior: on Alisma; catalogNumber: http://
10 Kahanpää J, Zatwarnicki T

id.luomus.fi/GV.16611; recordedBy: Tiensuu, Lauri; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Hydrellia mutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846);
country: Russia; stateProvince: Carelian Republic; municipality: Sortavala;
verbatimLocality: Sortavala; decimalLatitude: 61.7; decimalLongitude: 30.7;
geodeticDatum: wgs84; coordinateUncertaintyInMeters: 20000; georeferencedBy:
Kahanpää, Jere; year: 1936; month: 7; day: 1; verbatimEventDate: 1-VII 36;
individualCount: 1; sex: F; lifeStage: adult; behavior: on Alisma; catalogNumber: http://
id.luomus.fi/GV.16612; recordedBy: Tiensuu, Lauri; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
c. scientificName: Hydrellia mutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846);
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: U: Helsinki;
decimalLatitude: 60.19; decimalLongitude: 25; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 10000; georeferencedBy: Kahanpää, Jere; year: 1958;
month: 7; day: 22; verbatimEventDate: 22 VII -58; individualCount: 1; sex: F; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.16613; recordedBy: Tiensuu, Lauri;
identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
d. scientificName: Hydrellia mutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846);
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: N: Espoo,
Kaitalahti; verbatimCoordinates: 66715:33712; verbatimCoordinateSystem: YKJ;
decimalLatitude: 60.138; decimalLongitude: 24.691; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 300; georeferencedBy: Kahanpää, Jere; year: 2004;
month: 8; day: 3; individualCount: 1; sex: M; lifeStage: adult; recordedBy: Kahanpää,
Jere; identifiedBy: Kahanpää, Jere; dateIdentified: 2014-02-05; institutionCode: Coll.
Kahanpää; basisOfRecord: PreservedSpecimen
e. scientificName: Hydrellia mutata (Zetterstedt, 1846); order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: mutata; scientificNameAuthorship: (Zetterstedt, 1846);
country: Finland; stateProvince: V; municipality: Lohja; locality: Vaanila; verbatimLocality:
Ab: Lohja, Vaanila; verbatimCoordinates: 66919:33409; verbatimCoordinateSystem: YKJ;
decimalLatitude: 60.308; decimalLongitude: 24.118; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 50; georeferencedBy: Kahanpää, Jere; year: 2004;
month: 7; day: 16; individualCount: 1; sex: M; lifeStage: adult; recordedBy: Kahanpää,
Jere; identifiedBy: Kahanpää, Jere; dateIdentified: 2014-02-05; institutionCode: Coll.
Kahanpää; basisOfRecord: PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as illustrated by Collin (1966).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Northern, Western
and Central Europe (Zatwarnicki 2013).
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 11

Ecology
Larvae have recently been found feeding on Alisma (Robbins 1990) and Stratiotes,
possibly also Lemna and Alisma (see Deonier 1971).
Notes
Correctly identified by Richard Frey and Lauri Tiensuu but the record was never
published. Specimen http://id.luomus.fi/GV.16611 is illustrated in Fig. 1b.
Hydrellia subalbiceps Collin, 1966
Material
a. scientificName: Hydrellia subalbiceps Collin, 1966; order: Diptera; family: Ephydridae;
genus: Hydrellia; specificEpithet: subalbiceps; scientificNameAuthorship: Collin, 1966;
country: Finland; stateProvince: V; municipality: Salo; locality: Uskela; verbatimLocality:
Uskela; decimalLatitude: 60.38; decimalLongitude: 23.13; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 20000; georeferencedBy: Kahanpää, Jere;
individualCount: 1; sex: M; lifeStage: adult; preparations: gen. prep. on pin;
catalogNumber: http://id.luomus.fi/GV.16620; recordedBy: Ingelius, Hugo; identifiedBy:
Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as illustrated by Collin (1966) and
Canzoneri and Meneghini (1983).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widely distributed in
Europe and North Africa (Zatwarnicki 2013).
Ecology
Larval host plant(s) unknown.
Hydrellia tarsata Haliday, 1839
Material
a. scientificName: Hydrellia tarsata Haliday, 1839; order: Diptera; family: Ephydridae; genus:
Hydrellia; specificEpithet: tarsata; scientificNameAuthorship: Haliday, 1839; country:
Finland; stateProvince: V; municipality: Naantali; decimalLatitude: 60.13;
decimalLongitude: 22.02; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 10000;
georeferencedBy: Kahanpää, Jere; year: 1936; month: 7; day: 7; individualCount: 1; sex:
M; lifeStage: adult; preparations: gen. prep. on pin; behavior: on Stratiotes;
catalogNumber: http://id.luomus.fi/GV.16707; recordedBy: Tiensuu, Lauri; identifiedBy:
12 Kahanpää J, Zatwarnicki T

Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
Diagnosis
Best identified by structures of the male genitalia as illustrated by Collin (1966).
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Western and Central
Europe, also recorded from Bulgaria (Zatwarnicki 2013).
Biology
The Finnish specimen was caught on water soldiers (Stratiotes), which is a host plant
of H. tarsata (Przhiboro 2004).
Notes
Three most likely conspecific females were caught together with the male.
Allotrichoma bezzii Becker, 1896
Materials
a. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: A; municipality: Jomala; locality: Öjen; verbatimLocality:
Jomala; decimalLatitude: 60.18; decimalLongitude: 19.99; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 1000; georeferencedBy: Kahanpää, Jere; year: 1945;
month: 6; day: 19; habitat: shallow sea shore; individualCount: 1; sex: M; lifeStage: adult;
catalogNumber: http://id.luomus.fi/GV.16599; recordNumber: 7021; recordedBy: Frey,
Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
b. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: EH; municipality: Ylöjärvi; decimalLatitude: 61.55;
decimalLongitude: 23.60; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 1922; month: 4; day: 18; individualCount: 1; sex: M; lifeStage: adult; preparations:
gen. prep. on pin; catalogNumber: http://id.luomus.fi/GV.16598; recordedBy: Frey,
Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
c. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Russia; stateProvince: Carelian Republic; municipality: Paanajärvi;
decimalLatitude: 66.27; decimalLongitude: 29.80; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 17; individualCount: 1; sex:
M; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16600; recordNumber: 909;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 13

recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
d. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality:
Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day: 26; individualCount: 1; sex:
M; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16601; recordNumber: 4954;
recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
e. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi;
verbatimLocality: Paanajärvi; decimalLatitude: 66.27; decimalLongitude: 29.80;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19196; recordNumber: 989; recordedBy:
Platonoff, Stephan; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode:
MZH; basisOfRecord: PreservedSpecimen
f. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo;
decimalLatitude: 60.237; decimalLongitude: 24.007; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19197; recordNumber: 1697; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
g. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Vihti; verbatimLocality: Vichtis;
decimalLatitude: 60.424; decimalLongitude: 24.353; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19198; recordNumber: 3118; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
h. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo;
decimalLatitude: 60.237; decimalLongitude: 24.007; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19199; recordNumber: 1676; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
i. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo;
decimalLatitude: 60.237; decimalLongitude: 24.007; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19200; recordNumber: 1700; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
14 Kahanpää J, Zatwarnicki T

j. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo;
decimalLatitude: 60.237; decimalLongitude: 24.007; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19201; recordNumber: 2863; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
k. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: PPe; municipality: Hailuoto; verbatimLocality: Karlö;
decimalLatitude: 65.099; decimalLongitude: 24.750; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19202; recordNumber: 3361; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
l. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality:
Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.862; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1955; month: 7; day: 22; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19203; recordNumber: 1887;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
m. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: U: Helsinki;
decimalLatitude: 60.165; decimalLongitude: 24.915; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1949; month: 4; day: 17; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19204; recordedBy: Tiensuu, Lauri;
identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
n. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality:
Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.862; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1958; month: 6; day: 14; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19205; recordNumber: 4011;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
o. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: Helsinki;
decimalLatitude: 60.165; decimalLongitude: 24.915; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1949; month: 4; day: 17; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19206; recordedBy: Tiensuu, Lauri;
identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
p. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 15

country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality:
Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1959; month: 8; day: 12; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19207; recordNumber: 4487;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
q. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: H:fors;
decimalLatitude: 60.165; decimalLongitude: 24.915; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19208; recordNumber: 2575; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
r. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: Helsinki;
decimalLatitude: 60.165; decimalLongitude: 24.915; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19209; recordNumber: 755; recordedBy: Tuomikoski, Risto;
identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
s. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: U; municipality: Espoo; verbatimLocality: Espoo;
decimalLatitude: 60.251; decimalLongitude: 24.729; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19210; recordNumber: 41; recordedBy: Tuomikoski, Risto;
identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
t. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality:
Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.862; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day: 20; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19211; recordNumber: 4622;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
u. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality:
Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.862; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day: 13; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19212; recordNumber: 4047;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
v. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Lohja; verbatimLocality: Lojo;
decimalLatitude: 60.237; decimalLongitude: 24.007; geodeticDatum: wgs84;
16 Kahanpää J, Zatwarnicki T

georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19213; recordNumber: 2826; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
w. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality:
Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.862; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day: 20; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19214; recordNumber: 4660;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
x. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality:
Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19215; recordNumber: 4775; recordedBy: Storå, Ragnar; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
y. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality:
Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19216; recordNumber: 4764; recordedBy: Storå, Ragnar; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
z. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: V; municipality: Sammatti; verbatimLocality: Sammatti;
decimalLatitude: 60.323; decimalLongitude: 23.818; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19217; recordNumber: 1820; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
aa. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: Ks; municipality: Kuusamo; verbatimLocality: Oulankaj.;
decimalLatitude: 66.261; decimalLongitude: 29.543; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19218; recordNumber: 386; recordedBy: Frey, Richard; identifiedBy:
Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
ab. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Uusikaarlepyy; verbatimLocality:
Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1955; month: 7; day: 2; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19219; recordNumber: 1827;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 17

recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
ac. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; municipality: Kuortane; verbatimLocality: Kuortane; decimalLatitude:
62.741; decimalLongitude: 23.561; geodeticDatum: wgs84; georeferencedBy: Kahanpää,
Jere; year: 1955; month: 8; day: 17; individualCount: 1; lifeStage: adult; catalogNumber: h
ttp://id.luomus.fi/GV.19220; recordNumber: 7590; recordedBy: Storå, Ragnar;
identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
ad. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: KP; municipality: Pedersöre; verbatimLocality:
Pedersöre; decimalLatitude: 63.531; decimalLongitude: 22.862; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day: 13; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19221; recordNumber: 4588;
recordedBy: Storå, Ragnar; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
ae. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Finland; stateProvince: U; municipality: Helsinki; verbatimLocality: U: Helsinki;
decimalLatitude: 60.165; decimalLongitude: 24.915; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1949; month: 4; day: 17; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19222; recordedBy: Tiensuu, Lauri;
identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
af. scientificName: Allotrichoma bezzii Becker, 1896; order: Diptera; family: Ephydridae;
genus: Allotrichoma; specificEpithet: bezzii; scientificNameAuthorship: Becker, 1896;
country: Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi;
verbatimLocality: Paanajärvi; decimalLatitude: 66.27; decimalLongitude: 29.80;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19223; recordNumber: 888; recordedBy:
Frey, Richard; identifiedBy: Krivosheina, M.G.; dateIdentified: 1999; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Holarctic,
widespread in Europe (Zatwarnicki 2013).
Notes
All old records of Allotrichoma laterale (Loew, 1860) from Finland are misidentifications
of Allotrichoma bezzii Becker, 1896.
18 Kahanpää J, Zatwarnicki T

Ditrichophora calceata (Meigen, 1830)
Materials
a. scientificName: Ditrichophora calceata (Meigen, 1830); order: Diptera; family:
Ephydridae; genus: Ditrichophora; specificEpithet: calceata; scientificNameAuthorship:
(Meigen, 1830); country: Finland; stateProvince: U; municipality: Helsinki; locality:
University Botanical gardens; decimalLatitude: 60.17; decimalLongitude: 24.95;
geodeticDatum: wgs84; coordinateUncertaintyInMeters: 500; georeferencedBy:
Kahanpää, Jere; eventDate: 1942-7-14; year: 1942; month: 7; day: 14; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16562; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Tadeusz; dateIdentified: 09/26/2013; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
b. scientificName: Ditrichophora calceata (Meigen, 1830); order: Diptera; family:
Ephydridae; genus: Ditrichophora; specificEpithet: calceata; scientificNameAuthorship:
(Meigen, 1830); country: Russia; stateProvince: Carelian Republic; municipality:
Petrosawodsk; verbatimLocality: Äänislinna; decimalLatitude: 61.79; decimalLongitude:
34.35; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; eventDate: 1946-8-10;
year: 1946; month: 8; day: 10; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.16561; recordedBy: Tiensuu, Lauri; identifiedBy: Kahanpää, Jere;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). A West-Palaearctic
species found in almost all European countries and also in North Africa (Zatwarnicki
2013).
Notes
Specimen http://id.luomus.fi/GV.16561 is shown in Fig. 1c.
Hecamedoides unispinosus (Collin, 1943)
Materials
a. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: EH;
municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496;
decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19147;
recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
b. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: EH;
municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496;
decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19224;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 19

recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
c. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: EH;
municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496;
decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19225;
recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
d. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: EH;
municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496;
decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19226;
recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
e. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19227; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
f. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19228; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
g. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19229; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
h. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19230; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
i. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
20 Kahanpää J, Zatwarnicki T

geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19231; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
j. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19232; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
k. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19233; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
l. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19234; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
m. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Vaaseni;
verbatimLocality: Terijoki; decimalLatitude: 60.96; decimalLongitude: 34.02;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 8;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19235;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
n. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Vaaseni;
verbatimLocality: Vaaseni; decimalLatitude: 60.96; decimalLongitude: 34.02;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1942; month: 6; day:
15; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19236;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
o. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Terijoki;
verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 8;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19237;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
p. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 21

scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Terijoki;
verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 8;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19238;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
q. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Terijoki;
verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 8;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19239;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
r. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Metsäpirtti;
verbatimLocality: Metsäpirtti; decimalLatitude: 60.56; decimalLongitude: 30.52;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19240; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
s. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Terijoki;
verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 8;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19241;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
t. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Terijoki;
verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 8;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19242;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
u. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19243; recordedBy: Tiensuu, Lauri;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
v. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Vaaseni;
verbatimLocality: Vaaseni; decimalLatitude: 60.96; decimalLongitude: 34.02;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1942; month: 6; day:
15; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19244;
22 Kahanpää J, Zatwarnicki T

recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
w. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Terijoki;
verbatimLocality: Terijoki; decimalLatitude: 60.18; decimalLongitude: 29.7;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day:
11; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19245;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
x. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Russia; municipality: Vaaseni;
verbatimLocality: Vaaseni; decimalLatitude: 60.96; decimalLongitude: 34.02;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1942; month: 6; day: 1;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19246;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
y. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19247; recordedBy: Tiensuu, Lauri;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
z. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19248; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
aa. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19249; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
ab. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Lohja; verbatimLocality: Lojo; decimalLatitude: 60.237; decimalLongitude: 24.0078;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19250; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
ac. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: U; municipality:
Helsinki; locality: Botanical gardens; verbatimLocality: Helsinki, Hortus Bot.;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 23

decimalLatitude: 60.174; decimalLongitude: 24.9508; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1939; month: 6; day: 5; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19251; recordedBy: Tiensuu, Lauri;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
ad. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: KP;
municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531;
decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 1958; month: 6; day: 14; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.19252; recordedBy: Storå, Ragnar; identifiedBy: Zatwarnicki, Th.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
ae. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: KP;
municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531;
decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 1952; month: 6; day: 28; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.19253; recordedBy: Storå, Ragnar; identifiedBy: Zatwarnicki, Th.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
af. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: KP;
municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531;
decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 1952; month: 6; day: 28; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.19254; recordedBy: Storå, Ragnar; identifiedBy: Zatwarnicki, Th.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
ag. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: KP;
municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531;
decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 1952; month: 6; day: 28; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.19255; recordedBy: Storå, Ragnar; identifiedBy: Zatwarnicki, Th.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
ah. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: KP;
municipality: Pedersöre; verbatimLocality: Pedersöre; decimalLatitude: 63.531;
decimalLongitude: 22.8625; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
year: 1952; month: 6; day: 28; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.19256; recordedBy: Storå, Ragnar; identifiedBy: Zatwarnicki, Th.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
ai. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: V; municipality:
Vihti; verbatimLocality: Vichtis; decimalLatitude: 60.424; decimalLongitude: 24.3539;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage:
24 Kahanpää J, Zatwarnicki T

adult; catalogNumber: http://id.luomus.fi/GV.19257; recordedBy: Frey, Richard;
identifiedBy: Zatwarnicki, Th.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
aj. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: EH;
municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496;
decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19258;
recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
ak. scientificName: Hecamedoides unispinosus (Collin, 1943); order: Diptera; family:
Ephydridae; genus: Hecamedoides; specificEpithet: unispinosus;
scientificNameAuthorship: (Collin, 1943); country: Finland; stateProvince: EH;
municipality: Kangasala; verbatimLocality: Kangasala; decimalLatitude: 61.496;
decimalLongitude: 24.0754; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19259;
recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, Th.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
Diagnosis
Two of the North European Hecamedoides species can be identified by the number of
short, blunt anteroventral spine-like setae on the fore femur: one in H. unispinosus,
several in H. glaucellus (Stenhammar, 1844). Hecamedoides kelmorum Stuke, 2011, a
species recently described from Germany, also has a black antenna and a single
anteroventral spine (Stuke 2011). Hecamedoides unispinosus and H. kelmorum are
best identified by structures of the male genitalia. Two Finnish males (http://
id.luomus.fi/GV.19243 and http://id.luomus.fi/GV.19235) were dissected to confirm their
identification. The gonite of these specimens is clearly of the H. unispinosus type.
Distribution
A Holarctic species. Recorded from Finland by Kahanpää 2013. New for Russia.
Taxon discussion
Hecamedoides unispinosus is much more common in Finland than H. glaucellus. The
Fennica collection of MZH has only three specimens of the latter species: http://
id.luomus.fi/GV.19146 from Turku (see Fig. 2b) and two specimens from Terijoki
(=Russia, Leningrad Oblast, Zelenogorsk).
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 25

Pelina norvegica Dahl, 1975
Materials
a. scientificName: Pelina norvegica Dahl, 1975; order: Diptera; family: Ephydridae; genus:
Pelina; specificEpithet: norvegica; scientificNameAuthorship: Dahl, 1975; country: Russia;
stateProvince: Murmansk Oblast; locality: Yläluostari; verbatimLocality: Petsamo;
decimalLatitude: 69.4; decimalLongitude: 31.11; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1930; month: 7; day: 12; habitat: riverside;
individualCount: 1; sex: M; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19162;
recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Pelina norvegica Dahl, 1975; order: Diptera; family: Ephydridae; genus:
Pelina; specificEpithet: norvegica; scientificNameAuthorship: Dahl, 1975; country: Russia;
stateProvince: Murmansk Oblast; locality: Yläluostari; verbatimLocality: Petsamo;
decimalLatitude: 69.4; decimalLongitude: 31.11; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1930; month: 7; day: 12; habitat: riverside;
individualCount: 1; sex: M; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19163;
recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
Distribution
Previously recorded from Norway, Germany and Slovakia (Zatwarnicki 2013). New for
Russia.
Notes
Specimen http://id.luomus.fi/GV.19162 of Pelina norvegica is shown in Fig. 1d.
a b
Figure 2.
Hecamedoides species from Finland. Scale bar = 1.0 mm.
a: Hecamedoides glaucellus (specimen http://id.luomus.fi/GV.19146)
b: Hecamedoides unispinosus (specimen http://id.luomus.fi/GV.19147)
26 Kahanpää J, Zatwarnicki T

Parydra (Chaetoapnea) mitis (Cresson, 1930)
Materials
a. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: U; municipality: Sipoo; locality: Sjöskog; verbatimCoordinates:
6690:3385; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1975;
month: 5; day: 6; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16564; recordedBy: Albrecht, Anders; identifiedBy: Zatwarnicki, T.; institutionCode:
MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Russia; municipality: Vaaseni; geodeticDatum: wgs84; georeferencedBy: Kahanpää,
Jere; year: 1942; month: 5; day: 2; individualCount: 1; lifeStage: adult; catalogNumber: htt
p://id.luomus.fi/GV.16565; recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, T.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
c. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: U; municipality: Helsinki; locality: Herttoniemi; geodeticDatum:
wgs84; georeferencedBy: Kahanpää, Jere; year: 1945; month: 5; day: 16;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16566;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, T.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
d. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: EK; municipality: Vehkalahti; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1945; month: 4; day: 9; individualCount: 1;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16567; recordedBy: Tiensuu, Lauri;
identifiedBy: Zatwarnicki, T.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
e. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: U; municipality: Helsinki; locality: Herttoniemi; geodeticDatum:
wgs84; georeferencedBy: Kahanpää, Jere; year: 1945; month: 5; day: 16;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16568;
recordedBy: Tiensuu, Lauri; identifiedBy: Zatwarnicki, T.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
f. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: PS; municipality: Maaninka; locality: Tuovilanlahti;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1865; individualCount:
1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16569; recordedBy: Lundström,
Carl; identifiedBy: Zatwarnicki, T.; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
g. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi; locality: behind
Takalo farm; decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1939; month: 17; day: 6; habitat: lake shore;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16570;
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 27

recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, T.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
h. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: KP; municipality: Uusikaarlepyy; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954; individualCount: 1; lifeStage: adult;
catalogNumber: http://id.luomus.fi/GV.16571; recordedBy: Storå, Ragnar; identifiedBy:
Zatwarnicki, T.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
i. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Russia; stateProvince: Republic of Carelia; municipality: Paanajärvi; locality: hotel;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1939; month: 7; day: 6;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16572;
recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, T.; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
j. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: KP; municipality: Uusikaarlepyy; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1954-1955; individualCount: 1; lifeStage: adult;
catalogNumber: http://id.luomus.fi/GV.16573; recordedBy: Storå, Ragnar; identifiedBy:
Zatwarnicki, T.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
k. scientificName: Parydra mitis (Cresson, 1930); order: Diptera; family: Ephydridae; genus:
Parydra; specificEpithet: mitis; scientificNameAuthorship: (Cresson, 1930); country:
Finland; stateProvince: U; municipality: Helsinki; locality: Munkkiniemi; geodeticDatum:
wgs84; georeferencedBy: Kahanpää, Jere; year: 1941; month: 5; day: 4; habitat: a herb-
rich forest with a stream; individualCount: 1; lifeStage: adult; catalogNumber: http://
id.luomus.fi/GV.16574; recordedBy: Frey, Richard; identifiedBy: Zatwarnicki, T.;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Widespread in
Europe, it is also found in North Africa (Zatwarnicki 2013). The illustrated record by
Kahanpää (2013) represents Parydra nigritarsis Strobl, 1893.
Notes
Fig. 3c illustrates specimen http://id.luomus.fi/GV.16525.
28 Kahanpää J, Zatwarnicki T

Calocoenia paurosoma (Sturtevant & Wheeler, 1954)
Materials
a. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Finland; stateProvince:
EnL; municipality: Enontekiö; locality: Kilpisjärvi; decimalLatitude: 69.0; decimalLongitude:
20.9; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 4000; georeferencedBy:
Kahanpää, Jere; year: 1954; month: 8; day: 2; individualCount: 1; sex: M; lifeStage: adult;
catalogNumber: http://id.luomus.fi/GV.18970; recordNumber: 6643; recordedBy: Storå,
Ragnar; identifiedBy: Krivosheina, Marina G.; dateIdentified: 2004; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
b. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Finland; stateProvince:
EnL; municipality: Enontekiö; locality: Kilpisjärvi, Malla; decimalLatitude: 69.09;
decimalLongitude: 20.70; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 4000;
georeferencedBy: Kahanpää, Jere; year: 1929; month: 7; day: 15; habitat: regio
a b
c d
Figure 3.
Parydra species from Finland. Scale bar = 1.0 mm.
a: Parydra arctica, male (specimen http://id.luomus.fi/GV.16507)
b: Parydra fossarum, male (specimen http://id.luomus.fi/GV.16512)
c: Parydra mitis, female (specimen http://id.luomus.fi/GV.16525)
d: Parydra nigritarsis, male (specimen http://id.luomus.fi/GV.19145)
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 29

subalpina; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16463; recordNumber: 1106; recordedBy: Frey, Richard; identifiedBy: Kahanpää,
Jere; dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
c. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Finland; stateProvince:
Ks; municipality: Kuusamo; locality: Kuusamo town, Niilo-oja; locationRemarks: N side of
Niilo-oja; decimalLatitude: 65.97; decimalLongitude: 29.18; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 500; georeferencedBy: Kahanpää, Jere; year: 1917;
month: 6; day: 9; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/
GV.16457; recordNumber: 392; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord: PreservedSpecimen
d. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Finland; stateProvince:
KemLl; municipality: Kittilä; locality: Alakyrö; decimalLatitude: 68.07; decimalLongitude:
24.39; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1911; month: 6;
day: 19; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16459;
recordNumber: 1115; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord: PreservedSpecimen
e. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Finland; stateProvince:
PS; municipality: Nilsiä; decimalLatitude: 63.2; decimalLongitude: 28.1; geodeticDatum:
wgs84; georeferencedBy: Kahanpää, Jere; year: 1865; month: 6; day: 27;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16456;
recordNumber: 1153; recordedBy: Lundström, Carl; identifiedBy: Kahanpää, Jere;
dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord: PreservedSpecimen
f. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Russia; stateProvince:
Carelian Republic; municipality: Paanajärvi; locality: bog behind Takalo farm;
decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; year: 1937;
month: 6; day: 24; habitat: a bog; individualCount: 1; lifeStage: adult; catalogNumber: htt
p://id.luomus.fi/GV.16462; recordNumber: 712; recordedBy: Frey, Richard; identifiedBy:
Kahanpää, Jere; dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
g. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Russia; stateProvince:
Carelian Republic; municipality: Paanajärvi; locality: bog behind Takalo farm;
decimalLatitude: 66.26; decimalLongitude: 29.78; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; year: 1937;
month: 6; day: 24; habitat: a bog; individualCount: 1; lifeStage: adult; catalogNumber: htt
p://id.luomus.fi/GV.16458; recordNumber: 673; recordedBy: Frey, Richard; identifiedBy:
Kahanpää, Jere; dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
30 Kahanpää J, Zatwarnicki T

h. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Russia; stateProvince:
Murmansk Oblast; municipality: Kantalahti; decimalLatitude: 67.2; decimalLongitude:
32.4; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1913; month: 6;
day: 28; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16460;
recordNumber: 2492; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord: PreservedSpecimen
i. scientificName: Calocoenia paurosoma (Sturdevant & Wheeler, 1954); order: Diptera;
family: Ephydridae; genus: Calocoenia; specificEpithet: paurosoma;
scientificNameAuthorship: (Sturdevant & Wheeler, 1954); country: Russia; stateProvince:
Murmansk Oblast; municipality: Kantalahti; decimalLatitude: 67.2; decimalLongitude:
32.4; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1913; month: 6;
day: 27; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16461;
recordNumber: 961; recordedBy: Hellén, Wolter; identifiedBy: Kahanpää, Jere;
dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord: PreservedSpecimen
Diagnosis
At first glance this species resembles the genus Coenia, but it has five dorsocentral
setae. Specimen http://id.luomus.fi/GV.16455 is illustrated in Fig. 4b, d.
a b
c d
Figure 4.
Coenia and Calocoenia. Scale bars 0.5 mm (a-b) and 1.0 mm (c-d).
a: Coenia vulgata, face. Paratype specimen http://id.luomus.fi/GV.4185.
b: Calocoenia paurosoma, face. Specimen http://id.luomus.fi/GV.16455.
c: Coenia vulgata, lateral view. Specimen as above.
d: Calocoenia paurosoma, lateral view. Specimen as above.
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 31

Distribution
New for Russia. North Holarctic, known in Europe from Norway, Sweden and Finland
(Zatwarnicki 2013). First noted from Finland by Krivosheina (2000) from Kilpisjärvi. The
new records presented here expand the known European distribution of the species
both south- and eastwards.
Coenia vulgata Krivosheina, 2001
Materials
a. scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae;
genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001;
country: Finland; stateProvince: PPe; municipality: Oulu; decimalLatitude: 65.01;
decimalLongitude: 25.41; geodeticDatum: wgs84; coordinateUncertaintyInMeters: 10000;
georeferencedBy: Kahanpää, Jere; year: 1941; month: 4; day: 27; individualCount: 1; sex:
M; lifeStage: adult; preparations: gen. prep. on pin; catalogNumber: http://id.luomus.fi/
GV.16464; recordNumber: 174; recordedBy: Wuorentaus, Yrjö; identifiedBy: Kahanpää,
Jere; dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
b. scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae;
genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001;
country: Finland; stateProvince: PPe; municipality: Hailuoto; locality: Kirkkolahti;
decimalLatitude: 64.98; decimalLongitude: 24.72; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; year: 1947;
month: 7; day: 14; habitat: coastal meadows; individualCount: 1; sex: M; lifeStage: adult;
catalogNumber: http://id.luomus.fi/GV.16465; recordNumber: 775; recordedBy: Frey,
Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013; institutionCode: MZH;
basisOfRecord: PreservedSpecimen
c. scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae;
genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001;
country: Finland; stateProvince: PPe; municipality: Hailuoto; locality: Kirkkolahti;
decimalLatitude: 64.98; decimalLongitude: 24.72; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 2000; georeferencedBy: Kahanpää, Jere; year: 1947;
month: 7; day: 12; habitat: coastal meadows, on water; individualCount: 1; sex: F;
lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.16466; recordNumber: 606;
recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 09/26/2013;
institutionCode: MZH; basisOfRecord: PreservedSpecimen
d. scientificName: Coenia vulgata Krivosheina, 2001; order: Diptera; family: Ephydridae;
genus: Coenia; specificEpithet: vulgata; scientificNameAuthorship: Krivosheina, 2001;
country: Finland; stateProvince: PPe; municipality: Hailuoto; locality: Marjaniemi;
decimalLatitude: 65.04; decimalLongitude: 24.56; geodeticDatum: wgs84;
coordinateUncertaintyInMeters: 1000; georeferencedBy: Kahanpää, Jere; year: 1947;
month: 7; day: 13; habitat: a coastal bay and a coastal grazing area; individualCount: 1;
sex: F; lifeStage: adult; preparations: gen. prep. on pin; catalogNumber: http://
id.luomus.fi/GV.16467; recordNumber: 2557; recordedBy: Frey, Richard; identifiedBy:
Kahanpää, Jere; dateIdentified: 09/26/2013; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
32 Kahanpää J, Zatwarnicki T

Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Previously known
from Archangelsk Oblast (NW European Russia) and Kazakhstan (Krivosheina 2001).
Notes
Paratype specimen http://id.luomus.fi/GV.4185 is illustrated in Fig. 4a, c.
Scatophila iowana Wheeler, 1961
Material
a. scientificName: Scatophila iowana Wheeler, 1961; order: Diptera; family: Ephydridae;
genus: Scatophila; specificEpithet: iowana; scientificNameAuthorship: Wheeler, 1961;
country: Finland; stateProvince: KP; municipality: Pietarsaari; verbatimLocality: Fennia
Om Jakobstad; decimalLatitude: 63.69; decimalLongitude: 22.67; geodeticDatum:
WGS84; coordinateUncertaintyInMeters: 10000; georeferencedBy: Jere Kahanpää
(MZH); year: 1950; month: 7; day: 5; individualCount: 1; sex: M; lifeStage: adult;
catalogNumber: http://id.luomus.fi/HT.3120; recordedBy: Storå, Ragnar;
identificationRemarks: Identified as Scatophila modesta Becker by MG Krivosheina in
2003.; institutionCode: MZH; basisOfRecord: PreservedSpecimen
Diagnosis
Scatophila iowana Wheeler has entirely black legs, a yellow haltere, a matt brown
mesonotum and a brownish microtomentose abdomen. It lacks the pair of pronounced
facial setae typical for S. despecta (Haliday). The wing and mesonotal patterns of
Scatophila iowana resemble those of S. caviceps (Stenhammar). Males of these two
species can be identified by the shape of the central part of the face: somewhat
protruding between well-defined antennal grooves in S. iowana, weakly concave and
without antennal grooves in S. caviceps.
The anepisternum of S. iowana is rather uniformly brownish; S. caviceps has a grey
anepisternum with two brown spots, one around the base of the anepisternal bristle
and one at the ventral margin.
The male terminalia of S. iowana are characterized by the anterior margin of the
epandrium, which is slightly concave, and the aedeagus, which consists of three
sclerites. In S. caviceps the anterior margin of the epandrium bears a medial projection
and the aedeagus is a one-piece structure.
Distribution
First recorded from Finland by Zatwarnicki and Kahanpää (2014). Scatophila iowana is
a Holarctic species. It is probably widespread in Europe, but its distribution remains
poorly known (Zatwarnicki 2013).
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 33

Scatophila mesogramma (Loew, 1869)
Materials
a. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 7; day:
27; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19175;
recordNumber: 6405; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
b. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.44; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1955; month: 7; day:
20; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19176;
recordNumber: 1743; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
c. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1955; month: 7; day:
27; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19177;
recordNumber: 7133; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
d. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Russia; stateProvince: Republic of Carelia; municipality:
Paanajärvi; locality: behind Takalo farm; verbatimLocality: Paanajärvi; decimalLatitude:
66.26; decimalLongitude: 29.78; geodeticDatum: wgs84; georeferencedBy: Kahanpää,
Jere; year: 1939; month: 6; day: 17; habitat: lake shore; individualCount: 1; lifeStage:
adult; catalogNumber: http://id.luomus.fi/GV.19178; recordNumber: 885; recordedBy:
Frey, Richard; identifiedBy: Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode:
MZH; basisOfRecord: PreservedSpecimen
e. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1952; month: 8; day:
24; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19179;
recordNumber: 2005; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
f. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1952; month: 8; day:
30; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19180;
34 Kahanpää J, Zatwarnicki T

recordNumber: 2171; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
g. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day:
24; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19181;
recordNumber: 4726; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
h. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1952; month: 8; day:
30; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19182;
recordNumber: 2215; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
i. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1955; month: 8; day: 1;
individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19183;
recordNumber: 7287; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
j. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 8; day:
15; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19184;
recordNumber: 4557; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
k. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 8; day:
15; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19185;
recordNumber: 4559; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
l. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day:
26; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19186;
recordNumber: 4864; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 35

m. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: InL; municipality: Utsjoki; verbatimLocality:
Utsjoki; decimalLatitude: 69.509; decimalLongitude: 27.109; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; individualCount: 1; lifeStage: adult; catalogNumber: ht
tp://id.luomus.fi/GV.19187; recordNumber: 1308; recordedBy: Frey, Richard; identifiedBy:
Kahanpää, Jere; dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord:
PreservedSpecimen
n. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1952; month: 8; day:
24; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19188;
recordNumber: 2041; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
o. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day:
26; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19189;
recordNumber: 4859; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
p. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day:
26; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19190;
recordNumber: 4858; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
q. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Uusikaarlepyy;
verbatimLocality: Nykarleby; decimalLatitude: 63.436; decimalLongitude: 22.675;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1954; month: 5; day:
26; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19191;
recordNumber: 4856; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
r. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Russia; stateProvince: Murmansk Oblast; verbatimLocality: Kola;
decimalLatitude: 68.88; decimalLongitude: 33.04; geodeticDatum: wgs84;
georeferencedBy: Kahanpää, Jere; year: 1913; month: 7; day: 29; individualCount: 1;
lifeStage: adult; preparations: gen. prep. on pin; catalogNumber: http://id.luomus.fi/
GV.19192; recordNumber: 408; recordedBy: Hellén, Wolter; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
s. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
36 Kahanpää J, Zatwarnicki T

(Loew, 1869); country: Finland; stateProvince: EnL; municipality: Enontekiö; locality:
Palojoensuu; verbatimLocality: Enontekis; decimalLatitude: 68.296; decimalLongitude:
22.983; geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1911; month: 7;
day: 11; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19193;
recordNumber: 3871; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
t. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: EnL; municipality: Enontekiö;
verbatimLocality: Enontekis; decimalLatitude: 68.379; decimalLongitude: 22.725;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1911; month: 7; day:
14; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19194;
recordNumber: 4397; recordedBy: Frey, Richard; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
u. scientificName: Scatophila mesogramma (Loew, 1869); order: Diptera; family:
Ephydridae; genus: Scatophila; specificEpithet: mesogramma; scientificNameAuthorship:
(Loew, 1869); country: Finland; stateProvince: KP; municipality: Pietarsaari;
verbatimLocality: Jakobstad; decimalLatitude: 63.704; decimalLongitude: 22.634;
geodeticDatum: wgs84; georeferencedBy: Kahanpää, Jere; year: 1952; month: 8; day:
20; individualCount: 1; lifeStage: adult; catalogNumber: http://id.luomus.fi/GV.19195;
recordNumber: 2212; recordedBy: Storå, Ragnar; identifiedBy: Kahanpää, Jere;
dateIdentified: 2013-9-26; institutionCode: MZH; basisOfRecord: PreservedSpecimen
Distribution
New to Finland and Russia. First recorded from Finland by Zatwarnicki and Kahanpää
(2014). Holarctic; in Europe it is known from Finland, Sweden, Denmark, Great Britain,
Belgium and Germany (Zatwarnicki 2013).
Discussion
Notes on previous misidentifications
The species identified as Discocerina pulicaria [auct. nec Haliday] by Frey (1933), Frey
(1937), Frey (1949), Lindberg and Saris (1952) is actually Ditrichophora fuscella
(Stenhammar, 1844). Similarly, Allotrichoma laterale sensu Frey (1940b), Frey (1940a) is
A. bezzii Becker, 1896. Previous records of Philygria posticata by Frey 1930 represent Ph.
femorata (Stenhammar).
Acknowledgements
We would like to thank Dr. Wayne N. Mathis, Dr. Lorenzo Munari, Dr. Daniel Whitmore and
Dr. Owen Lonsdale for their constructive comments on an earlier version of this paper.
Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 37

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Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western ... 39