Two new West-Palaearctic species of Brachyopa Meigen, 1822 (Diptera, Syrphidae) with description and records of additional European species

Abstract

W.v. 2014. Two new West-Palaearctic species of Brachyopa Meigen, 1822 (Diptera, Syrphidae) with description and records of additional European species. Norwegian Journal of Entomology 61, 42–52. Brachyopa cruriscutum sp. n. (east Turkey) and B. vernalis sp. n. (Crete) are described. The new species are members of the bicolor guild sensu Doczkal & Dziock, 2004 and are compared with species of this guild. The male and the intraspecific variation of B. bimaculosa Doczkal & Dziock 2004 are described for the first time. Data on the first records of B. atlantea Kassebeer, 2000 and B. quadrimaculosa Thompson, 1981 spec. aff. Thompson in Kaplan & Thompson 1981 from Europe are given.

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Two new West-Palaearctic species of Brachyopa Meigen, 1822
(Diptera, Syrphidae) with description and records of additional
European species
JEROEN VAN STEENIS & WOUTER VAN STEENIS
Steenis, J.v. & Steenis, W.v. 2014. Two new West-Palaearctic species of Brachyopa Meigen, 1822
(Diptera, Syrphidae) with description and records of additional European species. Norwegian Journal
of Entomology 61, 42–52.
Brachyopa cruriscutum sp. n. (east Turkey) and B. vernalis sp. n. (Crete) are described. The new
species are members of the bicolor guild sensu Doczkal & Dziock, 2004 and are compared with
species of this guild. The male and the intraspecic variation of B. bimaculosa Doczkal & Dziock
2004 are described for the rst time. Data on the rst records of B. atlantea Kassebeer, 2000 and B.
quadrimaculosa Thompson, 1981 spec. aff. Thompson in Kaplan & Thompson 1981 from Europe
are given.
Key words: Brachyopa, new species, West-Palaearctic, Mediteranean, rst records, Europe.
Jeroen van Steenis, research associate Naturalis Biodiversity Center, Hof der Toekomst 48, NL-3823
HX Amersfoort, the Netherlands. E-mail: j.van.steenis@xmsnet.nl.
Wouter van Steenis, research associate Naturalis Biodiversity Center, Vogelmelk 4, NL-3621 TP
Breukelen, the Netherlands. E-mail: w.v.steenis@casema.nl
Introduction
The genus Brachyopa Meigen 1822 is a Holarctic
genus with 38 described species (12 Nearctic and
26 Palaearctic) (Pape & Thompson 2013).
The distribution of the genus is connected
with the distribution of coniferous and deciduous
broadleaved forest. The larvae live in decaying
or exuding tree sap from dead or living trees, in
tree stumps and even in rot holes. Some of the
species are generalists and can be found in both
broadleaved and coniferous trees, other species
have a narrower tree preference (McLean &
Stubbs 1990, Rotheray 1991, 1993, 1996, Sivova
et al. 1999, Krivosheina 2005, Ricarte et al. 2013).
The adults, especially the males, are regularly
observed patrolling trees, with supposed sap runs,
defending a territory and looking for females to
copulate with. Flower visiting is observed on
plants with abundant and “open” owers like
Apiacea, Prunus spp., and Crataegus spp. The
ight period in Europe is April–July (Trop 1994,
Reemer et al. 2009, Speight 2011).
The imagos of Brachyopa resemble dung-ies
(Diptera, Scatophagidae) and can be separated
from other Syrphidae by the following combination
of characters: Medium sized (6–10mm) mainly
brown, brown-red or black coloured ies.
Postpronotum pilose; basoagellomere round to
oval, often with clearly visible sensory pit; arista
pilose, from short to plumose; vein R4+5 straight;
crossvein rm before middle of discal cell; vein tm
oblique to vein R4+5 (Torp 1994, Reemer et al.
2009).
The classication of the genus as a member
of the Brachyopini with the Chrysogastrini as
© Norwegian Journal of Entomology. 30 June 2014

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sister tribe has been stable over time (Rotheray &
Gilbert 1999, Ståhls et al. 2003, Hippa & Ståhls
2005).
Material and methods
The material studied originate from: DDG – Dieter
Doczkal, Gaggenau, Germany; JSA – Jeroen van
Steenis, Amersfoort, the Netherlands; MRA –
Menno Reemer, Amsterdam, the Netherlands;
RMNH – National Museum of Natural History
(“NBC Naturalis”), Leiden, the Netherlands; WSB
– Wouter van Steenis, Breukelen, the Netherlands;
ZMAN – now incorporated in RMNH; ZMUC
– Zoological Museum, University Copenhagen,
Denmark; FSUNS – Faculty of Sciences,
Department of Biology and Ecology, University
of Novi Sad, Serbia.
The illustrations were made with the aid of
a camera Lucida attached to a stereomicroscope.
The male genitalia were dissected and treated
with potassium hydroxide (10 % KOH solution)
and drawn in glycerol. The other parts were drawn
in the dry state. The terminology used is based
on Thompson (1999) and Doczkal & Dziock
(2004). Some parts are measured with an ocular
micrometer and the absolute or relative lengths
are given. For the head of the males the relative
length of distance from occiput to anterior ocellus;
from anterior ocellus to eye contiguity, of eye
contiguity and of frons is given as A:B:C:D.
The Species
Brachyopa cruriscutum sp. n. (Figures 1–12)
Type species. Holotype ♂. TURKEY,
Hakkari, Sat Daglari, Varegös, 1600 m.a.s.l., [37°
25’ N, 43° 55’ E] 15.VI.1984, leg. J.A.W. Lucas,
(ZMAN). Paratypes 9♂♂1♀. TURKEY, Hakkari,
Sat Daglari, Varegös, 1600 m.a.s.l., 15.VI.1984,
1♀, leg. J.A.W. Lucas (ZMAN); TURKEY,
Hakkari, Sat Daglari, Varegös, 1600–1650
m.a.s.l., 17.VI.1984, leg. J.A.W. Lucas (ZMAN,
3♂♂; JSA, 2♂♂; WSB, 1♂; FSUNS, 1♂);
TURKEY, Hakkari, Sat Daglari, Varegös, 1600
m.a.s.l., 18.VI.1984, (ZMAN, 1♂; CNC, 1♂).
Diagnosis. This species is easily recognized
by the extensively shiny scutum, which in
combination with the nearly entirely black
scutellum is unlike any other species. It is similar
to an undescribed species from Lesvos, which
will be described by Dr. Vujić and co-workers
in a different paper. B. cruriscutum is compared
to a male of the Lesvos species (character states
between parentheses) and differs by: hypostomal
bridge black and entirely pollinose (yellow
with shiny macula); basoagellomere as long
as high (longer than high); proepimeron pilose
(non-pilose); abdomen entirely yellow (tergite 1
darkened laterally); posterior part of mesocoxa
pilose (bare).
Description (male). Body length: 6.5–6.8
mm. Wing length: 6.6–6.8 mm.
Head (Figures 1, 3). Face protruding H:L 95:80
(1.2–1.4); yellow with brown genae and mouth
edge; pollinose with bare shiny areas on anterior
part of genae; long sparse white pile on genae.
Paraface broad, pollinose, with a row of longer
white pile. Sides of antennal prominence yellow,
at most light brown; shiny and chagrinated. Frons
and lunula yellow, pollinose, with long white pile
along eyes; with a deep groove medially. Vertical
triangle black, anteriorly more brownish, slightly
grey pollinose. Relative length of A:B:C:D is
15:15:10:25. Hypostomal bridge black; entirely
but faintly pollinose, white pilose; subcranial
cavity L:W 60:40 (1.4–1.6). Clypeus brown, little
pollinose, sides black; L:W 10:11 (0.9–1.1). Eyes
bare. Occiput black, white pollinose with white
pile; rim behind ocellar triangle narrow. Antennae
orange, white pilose with row of black setulae on
scape and pedicel; basoagellomere as long as
high, L:H 13:13 (1.0-1.1); arista orange, white
pile about 1/3 of aristal diameter; sensory pit
hardly visible.
Thorax (Figure 4). Scutum black, subalar
callus dark brown; mostly shiny with thin white
pollinose areas; with erect to semi erect white
pile intermixed with black pile medially; sides
with longer white pile and 8–11 black setae above
wing. Postalar callus with longer white pile and
5–7 larger black setae. Postpronotum white pilose,
some black pile intermixed medially. Pleura black
with thin white pollinosity. Proepimeron with
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FIGURES 1–6. Brachyopa cruriscutum sp. n. 1. ♂ (Paratype 17.VI.1984). Head dorsal view. 2. ♀ (Paratype).
Head dorsal view. 3. ♂ (Holotype). Head lateral view. 4. ♂ (Holotype). Thorax dorsal view. 5. ♂ (Holotype).
Metafemur lateral view. 6. ♂ (Paratype 17.VI.1984). Apex metatibia and metabasitarsus ventral view.
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white pile on antero-dorsal corner; posterodorsal
part of anepisternum with some black setae;
posterior anepisternum, anepimeron and ventral
and dorsal katepisternum with white pile.
Mediotergite with lunulate pollinosity on anterior
3/4. Scutellum black, posterior ¼ dark-brown;
shiny, anterior 1/10 slighly pollinose; short semi
erect black and white pile, longer along sides; 2–6
black setae along posterior margin; with shallow
transverse sulcus; ventral scutellar fringe with
black and white pile.
Wings. Completely covered with microtrichia.
Halteres yellow.
Legs. Colour yellow; tarsi predominantly
brown, protarsus with tarsomere 1 yellow on
apical 1/10, tarsomeres 2–4 yellow on apical 1/3;
metatarsus with tarsomere 1 brownish, tarsomeres
2–5 yellow on apical 1/3. Pile on legs white,
posterior part of mesocoxa with pile. Pro- and
mesofemur with black pile apically. Black setulae
on ventral side of metafemur and tarsomeres
1–4 of mesotarsus. Lenght ratio of tarsomeres of
protarsus: 25:15:10:8:12. Metafemur (Figure 5)
L:H 85:21 (3.8–4.2). Metatibia with apicoventral
ridge as in Figure 6.
Abdomen. Colour yellow, shiny; pile white.
Sternites shiny without pollinosity. Membrane
connecting sternites with tergites pollinose
without pile.
Genitalia. See Figures 7–12.
Description (female). Similar to male except
for the usual sexual dimorphism. Body length: 5.9
mm. Wing length: 5.9 mm.
Head (Figure 2). Face H:L 70:65 (1.1); shiny
with some pollinosity below antennae. Frons
anterior 1/5 yellow, shiny; posterior 4/5 black,
white pollinose, with bare shiny medial vitta of
1/7 of width of frons; at. Ocellar plate hardly
FIGURES 7–12. Brachyopa cruriscutum sp. n. ♂ (Holotype) genitalia. 7. Hypandrium ventral view. 8.
Hypandrium lateral view. 9. Epandrium dorsal view. 10. Epandrium ventral view. 11. Epandrium lateral view.
12. Surstylus lateral view.
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pollinose. Subcranal cavity L:W 50:32. Clypeus
yellow, shiny; sides dark brown; L:W 10:9 (1.1).
Antennae: basoagellomere little longer than high
L:H 22:18 (1.2); arista greyish with grey pile;
small but distinct sensory pit.
Thorax. 1–2 black setae above wing; scutellum
posteriorly ¾ brown; with 3 setae along posterior
margin. No transverse sulcus.
Legs. little less brown on tarsi and less black
setulae on vetral side of metafemur. Metafemur
L:H 100:22 (4.5).
Etymology. The name “cruriscutum” is the
Latinised combination from the translation of the
English words shiny shield, crus scuto.
Brachyopa vernalis sp. n. (Figures 13–23)
Types species. Holotype ♂. GREECE, Crete,
[Krassio prov Iraklio East Crete], Between Krasi
& Tzermiado, 600 m.a.s.l., 8.IV.2008, leg. M.
Reemer, (ZMAN). Paratypes 7♂♂. GREECE,
Crete, [Krassio prov Iraklio East Crete], Between
Krasi & Tzermiado, 600 m.a.s.l., 8.IV.2008, leg.
M. Reemer, (MRA, 1♂; JSA, 1♂; WSB, 1♂;
FSUNS, 1♂; CNC, 1♂); GREECE, W. Crete,
[Sougia, Livadas prov Chania], Irini Gorge,
28.III.1997, 2♂♂, leg. V. Michelsen (ZMUC, 1♂;
ZMAN, 1♂).
Diagnosis. Easily recognized from other
species with grey thorax and yellow abdomen
by the dark antero-basal part of the capitulum of
the halter. Similar to an undescribed species from
Samos and Lesvos, see under B. quadrimaculosa
Thompson, 1981 spec. aff.
Description (male). Body length: 5.4–7.4
FIGURES 13–17. Brachyopa vernalis sp. n. 13. ♂ (Holotype). Head dorsal view. 14. ♂ (Holotype). Head lateral
view. 15. ♂ (Holotype). Thorax dorsal view. 16. ♂ (Holotype). Metafemur lateral view. 17. ♂ (Holotype). Apex
metatibia and metabasitarsus ventral view.
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FIGURES 18–23. Brachyopa vernalis sp. n. ♂ (Holotype) genitalia: 18. Hypandrium ventral view. 19.
Hypandrium lateral view. 20. Epandrium dorsal view. 21. Epandrium ventral view. 22. Epandrium lateral view.
23. Surstylus lateral view
mm. Wing length: 5.4–7.6 mm.
Head (Figures 13, 14). Face slightly protruding
H:L 52:40 (1.2–1.4); yellow with black maculae on
either side of antennal cavity (in some specimens
entirely yellow); pollinose, with bare shiny areas
on genae and along mouth edge; white pile on
genae and along mouth edge. Paraface pollinose,
laterally with bare shiny area. Sides of antennal
prominence bare, shiny. Frons yellow and shiny,
except for posterior ½ white pollinose. Vertical
triangle black, anterior more brownish; with white
pollinosity; pile mixed black and white; ocellar
triangle non-pilose. Relative lengths of A:B:C:D
is 12:12:3:16. Hypostomal bridge black; entirely
pollinose; white pilose; subcranial cavity L:W
62:40 (1.4–1.6). Clypeus shiny yellow, wrinkled
along edge, non-pilose; L:W 19:10 (1.8–2.0).
Eyes bare. Occiput black, white pollinose with
white pile; rim behind ocellar triangle narrow
with some black setae. Antennae orange-yellow,
predominantly black pilose; basoagellomere,
L:H 26:19 (1.4–1.5); arista black, basal part dark
orange; sensory pit absent.
Thorax (Figure 15). Scutum black covered
with dense white pollinosity except for 3 pairs of
bare maculae, 1 close to postpronotum, 1 at medial
end of transverse suture and 1 on medial part of
postalar callus. Scutum with dense, black pile, on
postpronotum, and along entire lareral margin of
scutum with white erect pile. Some black setae
laterally on both sides of transverse suture. Postalar
callus with 2–3 long black setae. Pleura black with
dense white pollinosity. Proepimeron with white
pile in antero-dorsal corner; posterodorsal part
of anepisternum with some black setae; posterior
anepisternum, anepimeron and ventral and dorsal
katepisternum with white pile. Mediotergite
with pollinosity arcuate on about anterior 1/3.
Scutellum yellow, anterior 1/2–4/7 black;
pollinose on anterior 1/4–2/5, slightly shiny; with
short semi erect white pile, medially with some
black pile; 4-8 long setae along posterior margin;
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scutellar fringe with longer white pile.
Wings. Completely covered with microtrichia.
Halteres yellow, dark on antero-basal part of
capitulum.
Legs. Colour yellow; tarsi predominantly
brown; tarsomers 4–5 always dark-brown;
protarsus with tarsomere 1–3 yellow on
respectively apical 1/10, 1/5 and 1/4; meso- and
meta tarsus with tarsomeres 1–3 variable in
colour, entirely yellow to predominantly dark-
brown. Pile on legs predominantly white; black
on tarsomeres 4–5 of pro- and metatarsus, entire
mesotarsus, basal part of mesotibia, and apical
part of pro- and mesofemur. Posterior part of
mesocoxa pilose. Black setulae on ventral side
of metafemur and tarsomeres 1–4 of mesotarsus.
Black setae on apical 1/2 of mesofemur (4–10
setae). Length ratio of tarsomeres of protarsus:
28:18:10:5:15. Metafemur (Figure 16) L:H 95:18
(5.2–5.6). Metatibia with apicoventral ridge as in
Figure 17.
Abdomen: Colour yellow, antero-lateral
corner of tergite 1 black; pile white. Sternites 1–4
completely pollinose.
Genitalia. See Figures 18–23.
Etymology. The name is from the latinized
word “vernalis” referring to the ight period,
which is in early spring.
Brachyopa bimaculosa Doczkal & Dziock, 2004
(Figures 24–37)
Material studied. GREECE, Greece, Ipiros,
Peristéri Mts., 1200–1700 m.a.s.l., 24–28.V.1994,
1♀, leg S. Andersen, Brachyopa bimaculosa
Doczkal & Dziock 2004, det Doczkal 2004
(ZMUC); Ellas, Viotia Parnassos, 1100 m.a.s.l.,
10 km NNE Del, 6.V.1998, 1♂, leg. M.J.
Gijswijt, (WSB); Hellas, Lakonia, “to the shelter
of Taigetos”, 22 km S.W. Sparti, 1000 m.a.s.l.,
7.V.1990, 1♂ leg. J.A.W. Lucas, (ZMAN); ibid,
1♀, (ZMAN); Lakonia, Anavriti ,1000 m.a.s.l.,
Megali Vrisi fountain, Greek grid 354–4097, on
Prunus sp., 22.IV.2005, 3♂♂, leg J. van Steenis,
(JSA); Greece, Peloponnesos, Arkadia Karderas,
Mt Menalo, N 37037’43” E 22018’01”, 16.V.2012,
980 m.a.s.l., 1♂1♀, leg W. van Steenis (WSB);
ibid on Acer spp, 2♀♀, leg J. van Steenis (JSA);
ibid 18.V.2012, 2♀♀, leg W. van Steenis (WSB);
Greece, Peloponnesos, Arkadia Mt Menalo,
Elati, on Acer spp, N 37037’33” E 22008’46”,
19.V.2012 1170 m.a.s.l., 1♂, leg J. van Steenis
(JSA); ibid 1♀, leg W. van Steenis (WSB);
Greece, Peloponnesos, Arkadia Mt Menalo,
Pyrgaki, on Bupleurum cf rotundifolium, N
37038’08” E 22009’27”, 19.V.2012 1090 m.a.s.l.,
1♂, leg J. van Steenis (JSA); ibid, 1♀ leg. W. van
Steenis (WSB); Greece, Peloponnesos, Arkadia
Mt Menalo, 5km South of Elati, N 37034’59” E
22009’50”, 19.V.2012 1250 m.a.s.l., 3♂♂1♀, leg
J. van Steenis (JSA); ibid 1♂2♀♀, leg W. van
Steenis (WSB); Greece, Peloponnesos, Arkadia
Mt Menalo, Vytina, road pass, on Prunus spp, N
37039’20” E 22015’56”, 19.V.2012 1620 m.a.s.l.,
1♂2♀♀, leg J. van Steenis (JSA); ibid 2♀♀ leg
W. van Steenis (WSB); Greece, Peloponnesos,
Arkadia Karderas, Mt Menalo, N 37037’43” E
22018’01”, 20.V.2012, 980 m.a.s.l., 1♀, leg W.
van Steenis (WSB); SLOVENIA Gorenjska,
Jesenica, Dom Pristava, (Javorniški Rovt) N
46°27’17” E 14°05’27”, 19.VI.2008, 970 m.a.s.l.,
1♂, leg J. van Steenis (JSA); GERMANY Bayern,
Riedberg Bolgenachtall, 47026’52”N 10008’07”E,
3.VI.2011, 1050 m.a.s.l., 1♂, leg J. van Steenis
(JSA).
Description (male). Body length: 6.3–7.8
mm. Wing length: 6.5–7.9 mm.
Head (Figures 24, 26). Face strongly pro-
truding H:L 73:75 (0.9–1.2); yellow with black
maculae on either side of antennal cavity (in
some specimens entirely yellow); pollinose, with
bare shiny areas on genae, along mouth edge and
below antennae; sparse pile on genae and along
mouth edge. Paraface pollinose, laterally with
bare shiny area. Sides of antennal prominence
yellow, bare, shiny and chagrinated. Frons yellow
and shiny, except for narrow pollinose vittae along
eye-margin, and narrow posterior margin; with a
shallow groove medially. Vertical triangle black,
anterior more brownish; with white pollinosity;
pile mixed black and white; ocellar triangle non-
pilose. Relative lengths of A:B:C:D is 14:14:9:19.
Hypostomal bridge yellow; entirely pollinose,
sometimes with very small bare maculae laterally;
white pilose; subcranial cavity L:W 58:30 (1.7–
2.0). Clypeus shiny yellow, wrinkled along edge,
non-pilose; L:W 18:8 (2.2–2.5; central European
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FIGURES 24–31. Brachyopa bimaculosa Doczkal & Dziock 2004. 24. ♂ (7.V.1990). Head dorsal view. 25. ♀
(7.V.1990). Head dorsal view. 26. ♂ (22.IV.2005). Head lateral view. 27. ♀ (7.V.1990). Head lateral view. 28.
♂ (22.IV.2005). Thorax dorsal view. 29. ♂ (22.IV.2005). Metafemur lateral view. 30. ♀ (7.V.1990). Metafemur
lateral view. 31. ♂ (22.IV.2005). Apex metatibia and metabasitarsus ventral view.
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specimens 1.6–1.8). Eyes bare. Occiput black,
white pollinose with white pile; rim behind ocellar
triangle narrow with some black setae. Antennae
orange-yellow, predominantly black pilose;
basoagellomere as long as high, L:H 16:14 (1.1–
1.3); arista black, basal part dark orange; sensory
pit minute, sometimes hardly visible.
Thorax (Figure 28). Scutum black, antero-
medial extension of postpronotum and anterior
and posterior 1/4 of postalar callus yellow;
covered with dense white pollinosity except for
two parallel vittae on anteromedial 2/3 slightly
pollinose, and 2 pair (many specimens only with
1 pair) of bare maculae, close to postpronotum
(in many specimens missing), and at medial end
of transverse suture. Scutum with dense, black,
semi erect pile, at postpronotum, notopleuron
and anterior part of scutum with white erect pile.
Some setae laterally on both sides of transverse
suture. Postalar callus with white erect pile and
two long black setae. Pleura black with dense
white pollinosity. Proepimeron with white pile
on antero-dorsal corner; posterodorsal part of
anepisternum with some black setae; posterior
anepisternum, anepimeron and ventral and dorsal
katepisternum with white pile. Mediotergite with
straight pollinosity on anterior ½. Scutellum
yellow, anterior 1/5 black; entirely but sparsely
pollinose, slightly shiny; short semi erect white
pile; 2–4 long setae along posterior margin, one
smaller on both sides laterally; scutellar fringe
with longer white pile.
Wings. Completely covered with microtrichia.
Haltere entirely yellow.
FIGURES 32–37. Brachyopa bimaculosa Doczkal & Dziock 2004 ♂ (22.IV.2005) genitalia. 32. Hypandrium
ventral view. 33. Hypandrium lateral view. 34. Epandrium dorsal view. 35. Epandrium ventral view. 36.
Epandrium lateral view. 37. Surstylus lateral view.
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Legs. Colour yellow; tarsi predominantly
brown tarsomers 4–5 always dark; protarsus
with tarsomere 1–3 yellow on at most apical
1/10, 1/6 and 1/5; meso- and meta tarsus with
tarsomeres 1–3 variable in colour, entirely yellow
to predominantly dark. Pile on legs white, black
on tarsomeres 4–5 of pro- and metatarsus, entire
mesotarsus, basal part of mesotibia, and apical
part of pro- and mesofemur. Posterior part of
mesocoxa bare. Black setulae on ventral side of
metafemur and tarsomeres 1–4 of mesotarsus.
Black setae on apical 1/2 of profemur (5–9 setae),
mesofemur (2–3 setae) and apical ends of ta: 1–3
(5 setae). Length ratio of tarsomeres of protarsus:
25:12:8:5:15. Metafemur (Figure 29) L:H 95:18
(5.1–5.4). Metatibia with apicoventral ridge as in
Figure 31.
Abdomen. Colour yellow, pile white. Sternites
1–4 completely pollinose.
Genitalia. See Figures 32–37.
Female. Similar to male, except for the usual
sexual dimorphism. Body length: 6.4–6.9 mm.
Wing length: 6.5–7.1 mm.
Head (Figures 25, 27): Face H:L 65:68 (0.9–
1.1); shiny with pollinose fascia widely interrupted
below antennae. Frons anterior 1/5 yellow, shiny;
posterior 4/5 black, white pollinose, with bare
shiny medial vitta of 1/5 of width of frons; at).
Ocellar plate pollinose. Subcranal cavity L:W
45:25 (1.7–1.9). Clypeus yellow, shiny; sides
dark brown; L:W 18:9 (1.8–2.1). Antennae:
basoagellomere little longer than high L:H 15:12
(1.2–1.4); small but distinct sensory pit.
Legs: Metafemur L:H 107:21 (4.9–5.2).
Remarks. Abundant on the Peloponnesos
(Greece), ying together with B. bicolor (Fallén),
B. grunewaldensis Kassebeer, B. insensilis Collin
and B. pilosa Collin; in Germany together with
B. dorsata Zetterstedt, B. testacea (Fallén) and
B. vittata Zetterstedt. Flowers visited: Acer spp.,
Aegopodia spp., Bupleurum cf rotundifolium,
Prunus spp and white Apiacea. In Slovenia (van
Steenis et al. 2013) and Germany, it was found
visiting owers along a river in the shade of trees.
The surrounding forest was dominated by Beech
(Fagus sylvaticus) and Fir (Abies alba) with Salix
spp., Acer spp. and Alnus spp. along the river. On
the Peloponnesos the specimens were caught on
open ower rich sub-alpine limestone meadows
within forests (dominated by Abies cephalonica
and Pinus nigra) and in rows of owering Acer
spp. trees bordering the roads close to mixed
forest.
Brachyopa atlantea Kassebeer, 2000
One of two species known from the African
continent (Kassebeer 2000, 2002). As for
other species like Platycheirus marokkanus,
Chrysotoxum volaticum, and Pipizella thapsiana
(Ssymank & Doczkal 2007, van Steenis & Lucas
2011) some of the African species tend to cross
the Mediterranean sea and turn up in the Iberian
peninsula.
Material studied. SPAIN, Espana, Grenada,
Lanjaron, 550 m.a.s.l., 13.IV.1974, [36° 55’ N, 3°
28’ E] 1♀, J.A.W. Lucas (ZMAN).
Remarks. New to Spain.
Brachyopa quadrimaculosa Thompson, 1981
spec aff.
This specimen is conspecic to a male of an
undescribed species send to us by Dr. Vujić. It
is similar to B. vernalis and B. quadrimaculosa.
Differing from both by: mediotergite with arcuate
pollinosity on anterior 1/2. Scutellum entirely
pollinose. Posterior mesocoxa bare. Metafemur
narrower L:H 82:14 (5.9). This species will be
decribed by Vujić and co-workers in a separate
paper.
Material studied. GREECE, Hellas, Samos,
Platanakia, 25.IV.1988, 1♀, leg. J.A.W. Lucas
(ZMAN).
Discussion
The genus Brachyopa is widespread in the
Holarctic region. In Europe it is a well-studied
genus. As this paper shows even within this
genus several new species can be found in the
Mediterranean sub region. Since there are several
species with a very limited distribution within
the Mediterranean sub region, it is to be expected
that more cryptic species could be found on the
Aegean islands or other isolated areas.
Norwegian Journal of Entomology 61, 42
–
52 (2014)

52
Acknowledgement. We wish to thank the following persons
for the loan of material and for other valuable help: Dieter
Doczkal (Gagenau), Menno Reemer (Amsterdam), Ben
Brugge (Leiden), Thomas Pape (Copenhagen) and Ante Vujić
(Novi Sad).
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Received: 10 December 2013
Accepted: 16 April 2014
Steenis & Steenis: Two new West-Palaearctic species of the genus Brachyopa