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ISSN 1313-2970 (online) ISSN 1313-2989 (print)
A peer-reviewed open-access journal
Launched to accelerate biodiversity research
ZooKeys 441 2014 CHECKLIST OF THE DIPTERA OF FINLAND
Edited by Jere Kahanpää & Jukka Salmela
http://www.zookeys.org
ZooKeys 441
2014
Special Issue
CHECKLIST OF THE
DIPTERA OF FINLAND
EditEd by Jere Kahanpää & JuKKa Salmela
!http://www.zookeys.org
This special issue of ZooKeys forms a checklist of all true ies (Insecta:
Diptera) found in Finland. Twenty-four authors contributed 31 manu-
scripts covering the whole order. Each paper treats one or more Diptera
families and includes a brief introduction, a systematic checklist, a list of
excluded species (if any), and explanatory notes.
Diptera is a large order of insects commonly known as the true ies. With
some 150,000 described extant species and many more still undescribed,
Diptera is one of the most successful groups of animals. Flies, midges and
their relatives are important actors in most land and freshwater ecosys-
tems, particularly so in the arctic and alpine zones. The economic relevance
of true ies is also considerable: they are key pollinators and biological
control agents, but also – less benecially – pests and important vectors of
human and domestic animal diseases.
The new checklist records 6920 Diptera species from Finland. 2932 species
belong to Nematocera or lower ies and 3989 to suborder Brachycera. In
comparison, the previous Finnish checklist – published in 1980 – involved
just 4852 species. Our understanding of the Finnish fauna has grown
across the whole order, but especially among Nematocera: for example,
more than 400 non-biting midges (Chironomidae) and 219 gall midges
(Cecidomyiidae) have been added since 1980 and the number of moth
ies (Psychodidae) known from the country has more than tripled. Among
Brachycera, the big-headed ies (Pipunculidae, with a 98 % increase) and
the parasitic family Tachinidae with 118 newly recorded species stand out.
Proceedings of a symposium honoring the careers of Ross and Joyce Bell... i
Checklist of the Diptera of Finland
by
Jere Kahanpää, Jukka Salmela
Sofia–Moscow
2014
Terry Erwin / ZooKeys 147 (2011)
ii
Pensoft Publishers
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ZK 441 (S I)
C D F
by Jere Kahanpää, Jukka Salmela
First published 2014
ISBN 978-954-642-@@@-@ (paperback)
Printed in Bulgaria, September 2014
Proceedings of a symposium honoring the careers of Ross and Joyce Bell... iii
Contents
1 Checklist of the Diptera (Insecta) of Finland:
an introduction and a summary of results
Jere Kahanpää
21 Checklist of the infraorder Tipulomorpha
(Trichoceridae, Tipuloidea) (Diptera) of Finland
Jukka Salmela, Andrius Petrašiūnas
37 Checklist of the familes Chaoboridae, Dixidae, Thaumaleidae
Psychodidae and Ptychopteridae (Diptera) of Finland
Jukka Salmela, Lauri Paasivirta, Gunnar M. Kvifte
47 Checklist of the family Culicidae (Diptera) in Finland
Larry Huldén, Lena Huldén
53 Checklist of the family Ceratopogonidae (Diptera) of Finland
Larry Huldén, Lena Huldén
63 Checklist of the family Chironomidae (Diptera) of Finland
Lauri Paasivirta
91 Checklist of the family Simuliidae (Diptera) of Finland
Jari Ilmonen
97 Checklist of the families Anisopodidae, Bibionidae, Canthyloscelididae,
Mycetobiidae, Pachyneuridae and Scatopsidae (Diptera) of Finland
Antti Haarto
103 Checklist of the Cecidomyiidae (Diptera) of Finland
Mathias Jaschhof, Marcela Skuhravá, Jouni Penttinen
119 Checklist of the fungus gnats of Finland: Bolitophilidae, Diadocidiidae,
Ditomyiidae, Keroplatidae and Mycetophilidae (Diptera)
Jevgeni Jakovlev
151 Checklist of the family Sciaridae (Diptera) of Finland
Pekka Vilkamaa
165 Checklist of the ‘lower Brachycera’ of Finland:
Tabanomorpha, Asilomorpha and associated families (Diptera)
Jere Kahanpää, Kaj Winqvist, eo Zeegers
183 Checklist of the Empidoidea of Finland (Insecta, Diptera)
Jere Kahanpää
Terry Erwin / ZooKeys 147 (2011)
iv
209 Checklist of the families Opetiidae and
Platypezidae (Diptera) of Finland
Gunilla Ståhls
213 Checklist of the families Lonchopteridae and
Phoridae of Finland (Insecta, Diptera)
Jere Kahanpää
225 Checklist of the family Pipunculidae (Diptera) of Finland
Christian Kehlmaier
233 Checklist of the family Syrphidae (Diptera) of Finland
Antti Haarto, Sakari Kerppola
251 Checklist of the superfamilies Conopoidea, Diopsoidea and
Nerioidea of Finland (Insecta, Diptera)
Jere Kahanpää, Jens-Hermann Stuke
259 Checklist of the Diptera superfamilies Tephritoidea and
Sciomyzoidea of Finland (Insecta)
Jere Kahanpää, Kaj Winqvist
277 Checklist of the fly families Chamaemyiidae and
Lauxaniidae of Finland (Insecta, Diptera)
Jere Kahanpää
285 Checklist of the smaller families of Opomyzoidea, Anthomyzidae,
Asteiidae, Aulacigastridae, Clusiidae, Odiniidae, Opomyzidae and
Periscelididae (Diptera) of Finland
Jere Kahanpää
291 Checklist of the leaf-mining flies (Diptera, Agromyzidae) of Finland
Jere Kahanpää
305 Checklist of the Diptera families Acartophthalmidae, Canacidae
(including Tethinidae), Carnidae and Milichiidae of Finland (Insecta)
Jere Kahanpää
311 Checklist of the family Chloropidae (Diptera) of Finland
Emilia Nartshuk, Jere Kahanpää
319 Checklist of the fly families Chyromyidae and
Heleomyzidae (Diptera) of Finland
Jere Kahanpää
325 Checklist of the family Sphaeroceridae (Diptera) of Finland
Antti Haarto, Jere Kahanpää
Proceedings of a symposium honoring the careers of Ross and Joyce Bell... v
333 Checklist of the Braulidae, Camillidae, Diastatidae and
Drosophilidae of Finland (Diptera, Ephydroidea)
Jere Kahanpää
339 Checklist of the family Ephydridae of Finland (Insecta, Diptera)
Tadeusz Zatwarnicki, Jere Kahanpää
347 Checklist of the families Scathophagidae, Fanniidae and
Muscidae of Finland (Insecta, Diptera)
Jere Kahanpää, Antti Haarto
369 Checklist of the family Anthomyiidae (Diptera) of Finland
Verner Michelsen
389 Checklist of the superfamilies Oestroidea and Hippoboscoidea of
Finland (Insecta, Diptera)
Jaakko Pohjoismäki, Jere Kahanpää
Terry Erwin / ZooKeys 147 (2011)
vi
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 1
Checklist of the Diptera (Insecta) of Finland:
an introduction and a summary of results
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: Jukka Salmela|Received 14 April 2014|Accepted 3 September 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results. In:
Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 1–20. doi: 10.3897/zookeys.441.7620
Abstract
Nearly thirty-ve years have passed since Hackman published his “Check list of the Finnish Diptera”
(1980). e number of true ies (Diptera) known from Finland has increased by more than two thousand
species since then. At the same time, hundreds of erroneous records have been recognized and purged
from the checklist.
ZooKeys issue 441 provides a new checklist of the Diptera species of the Republic of Finland. is
introductory paper presents the rationale behind the project, provides technical documentation on the
checklist format and sources used, and summarizes the results. e remaining papers in this issue cover
one or more Diptera families in detail.
Two electronic appendices are provided: supporting data (additional references to rst published
records and the previous checklist) and a complete list of Finnish Diptera taxa in Darwin Core compliant
format for easy computer access and processing.
e new checklist records 6920 y species from Finland, 2932 belonging to the nematoceran or
lower ies and 3989 to the suborder Brachycera. e changes since 1980 are most prominent in the Lower
Diptera. For example, more than 400 non-biting midges (Chironomidae) have been added since 1980,
and the number of moth ies (Psychodidae) known from Finland has more than tripled. Among the
larger families, large increases in known Finnish species are also seen in Cecidomyiidae (161% increase),
Pipunculidae (98%), and Chironomidae (90%).
ZooKeys 441: 1–20 (2014)
doi: 10.3897/zookeys.441.7620
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
RESEARCH ARTICLE
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
2
Keywords
Finland, Diptera, ies, biodiversity, faunistics
Introduction
e Diptera is a large order of holometabolous insects commonly known as ies and
midges. With some 150,000 described extant species (Pape et al. 2011) and many
more still undescribed, the Diptera is one of the most successful groups of animals by
any measure. Flies and midges are also an important part of food webs in most land
and freshwater ecosystems, particularly so in the arctic and alpine zones (Mani 1968,
Pape 2009a). e economic relevance of ies is also considerable (see Pape 2009b for a
recent summary): they are key pollinators and biological control agents, but also – less
benecially – important vectors of human and domestic animal diseases.
Why a new checklist?
e history of Finnish Diptera catalogs starts with Evert J. Bonsdor (1861, 1866),
who reviewed a part of the brachyceran fauna known from the country at the time,
totaling 657 species. Systematic maintenance and publishing of faunistic data began
with the start of Wolter Hellén’s amazing series of summary papers covering new coun-
try records (and deletions) of all Insecta and running uninterrupted for six decades
(Hellén 1922, 1926, 1931, 1936, 1941, 1946, 1952, 1956, 1961, 1966, 1971, 1976).
Richard Frey, Lauri Tiensuu, and Ragnar Storå (1941) published the rst complete list
of Diptera recorded from Finland. It included 3824 species.
e era of Hellén culminated in a revised list of Finnish Diptera by Walter Hack-
man (1980a, 1980b, 1980c) in cooperation with Bernhard Lindeberg and Rauno
Väisänen. e number of species had increased to 4852. Hans Silfverberg succeeded
Hellén as the author of regular updates on additions and deletions to the Finnish insect
fauna (Silfverberg 1981, 1986, 1991, 1996, 2007, 2012).
Nearly thirty-ve years have passed since Hackman’s checklist. e number of
Diptera species known from Finland has increased by more than two thousand spe-
cies since 1980 (an increase of more than 40%). Several hundred erroneous records
have been recognized during the same period. Many new names have been introduced
and others found invalid, incorrectly used, or synonymized. e Diptera fauna of the
whole Palaearctic Region has been catalogued (Soós and Papp 1984–1993) and many
new local, regional, or global checklists have been published.
At this point, an updated checklist is urgently needed to provide a current refer-
ence to the Diptera fauna of Finland as a fresh starting point for further studies into
the taxonomy, ecology, and other aspects of ies in Northern Europe.
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 3
Methods and format
Denition of a checklist
ere is no universally accepted denition for an entomological checklist. e word
is often used for lists of species with more information than merely valid names, but
without comprehensive details on nomenclature or distribution and incomplete lit-
erature references. is new checklist of Finnish Diptera falls into category 5 on the
comprehensiveness scale of ompson and Knutson (1987). It does include full names
with authorships and some but not all synonyms. Some literature references are pro-
vided in an Suppl. material 2.
National insect checklists have traditionally included the species recorded at least
once from the country. Exotic imports without locally reproducing populations are
usually excluded, but vagrants are included. is is by no means the only possible ap-
proach: the latest Danish list (Petersen and Meier 2001) use a predictive approach: it
lists not only the species actually recorded from the country, but also those that are like-
ly to be present based on the fauna of neighboring countries. e latter method works
best when the fauna of the surrounding areas is at least as well known as the study area.
Taxonomic, geographical, and temporal limits
is checklist covers the Diptera fauna of the Republic of Finland. Only species re-
corded at least once within the current (i.e. post-1944) borders are included.
Many species were originally reported from Finland on the basis of specimens col-
lected from areas ceded to Russia in 1944. If no reliable records (preferably voucher
specimens) from within the current borders of the country exist, species were excluded
from the checklist. In some cases – e.g. the muscid Coenosia comita (Huckett, 1936) –
it has been impossible to determine whether the purported Finnish collecting localities
fall within the current borders of the country. ese species are indicated with a ques-
tion mark on the checklist and are usually accompanied by a comment in the Notes
section of the relevant paper.
No fossil or subfossil records were considered during the preparation of the check-
list. Nationally extinct species are included. In practice, the oldest collected Diptera
specimens from Finland are from the early 19th century, so all of the species in this
checklist have been found in the country at least once during the last two hundred years.
Data sources and validation
Draft checklists for each family were created in the latter part of 2012 by combining
data from the most recent checklist of the Finnish fauna of the family in question, the
Palaearctic Catalog (Soós and Papp 1984–1993), Fauna Europaea (Beuk and Pape
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
4
2012), Silfverberg’s updates to Hackman’s checklist and references therein, other rel-
evant scientic literature, and the Finnish Insect Record Database (Finnish Museum
of Natural History 2013). From here onwards, the individual authors of each checklist
paper were responsible for data validation.
For a great majority of families, the next step was examining the major Diptera
collections in Finland (see the Acknowledgments). Fortunately, these collections have
voucher specimens for most Diptera species reported from the country. If no recently
identied specimens from the country could be found, the reliability of the record was
judged by section authors based on details provided in the literature. For example, the
record of Alliopsis longiceps (Ringdahl, 1935) from Finland was rejected. While Ring-
dahl correctly recorded this species from Finland in 1935, the type location (“Kuusamo
bei Paanajärvi”) is now Russian territory, and no later observations have been published.
Exceptions to the procedure include Trichoceridae, Ceratopogonidae, Cecidomyi-
idae (subfamily Cecidomyiinae), and Phoridae (genus Megaselia Rondani, 1856), for
which the checklists are largely based on a critical literature review. For more details on
sources and validation, see the introductions and notes of individual checklist papers.
Checklist structure and presentation
e new checklist of the Diptera of Finland is presented in three formats: a series of
papers each covering one or more families, a comma separated values (CSV) le with
the full taxon list, and a PDF (portable document format) le with literature references
for species records. e latter two are included as electronic appendices to this paper.
All checklist papers follow the same general format. An introduction to the families
covered is followed by a table or list of number of species recorded from the world,
Europe, and Finland. An estimate of the faunistic level of knowledge is also given on a
simple three-step scale (poor–average–good). is estimate is admittedly very subjective,
being based on individual authors’ judgment. e following factors were considered:
the number of species known from Finland in comparison with neighboring countries,
taking into account known habitat preferences, etc., of absent species; the number of spe-
cialists who have worked with Finnish fauna; the number of publications on the Finnish
fauna; and the quantity and quality of identied material collected from Finland.
e Checklist section of each paper starts with the systematic position of the family
or families treated. As an example, the checklist of lauxanoid ies starts with:
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Lauxanoidea Macquart, 1835
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 5
e family-level classication used follows mostly Marshall (2012), with exceptions
noted in the Introductions of individual papers. e presentation order of subfamilies,
tribes, genera, and subgenera can be either alphabetical or systematic depending on the
author’s preference. Species are always listed alphabetically within a genus or subgenus.
Each species record starts with the name of the taxon, the author’s name, and the
year of description. Doubtful records are indicated with a question mark (?) before the
species name. e valid name may be followed by one or more additional names used
for the same taxon. ese names can be younger synonyms, preoccupied names, misi-
dentications, or common misuses in Finnish or international literature. As an exam-
ple, the following entry lists three additional names used for Rhamphomyia trilineata.
Rhamphomyia trilineata Zetterstedt, 1859
= sulcatina Collin, 1926
= tibialis auct. nec Meigen, 1822
= propinqua misid.
R. sulcatina Collin is a younger synonym of R. trilineata. e name R. tibialis Meigen
was erroneously used for this species by Frey (1956) and others; the genuine R. tibialis of
Meigen has not been found in the Nordic countries. Frey et al. (1941) misidentied this
species as R. propinqua de Meijere (in reality a junior synonym name for R. sulcata Meigen).
Table 1 lists abbreviations used in the checklist papers.
e checklist section is followed by a list of species not included on the checklist
for various reasons. Excluded species comprise species recorded only from areas ceded
to Russia before 1945, exotic species occasionally imported to Finland by man without
locally reproducing populations, records based on misidentications, etc. e Notes
section presents authors’ comments on individual taxa.
Table 1. Common abbreviations used in the checklist papers.
Abbreviation Word or term Interpretation
a. anis aned to, near
auct. nec auctorum, nec wrong interpretation, literally ‘of authors, not’
cf. confer compare with (may be identical with)
coll. collective collective name for sister species not easily separable by morphology
emend. emendation an intentional alternative spelling
misid. misidentied
nom. dubium nomen dubium a name with uncertain meaning
nom. nudum nomen nudum a name without a proper scientic denition
pr. prope near
preocc. preoccupied preoccupied by an older homonymous name
sg. subgenus
sp. species
suppr. suppressed a name made unavailable by an ICZN decision
var. variety a described variety
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
6
Discussion
Table 2 presents a summary of the results for each family of Diptera recorded from
Finland, which includes the number of species known from the country now; in the
previous checklist (Hackman 1980a, 1980b, 1980c); and the dierence between the
two checklists in species counts and as a fraction of the fauna known in 1980.
e Diptera has traditionally been split into two suborders, Nematocera and
Brachycera. It is now generally agreed that while Brachycera is a monophyletic line-
age, Nematocera is not (see Yeates et al. 2007 and references therein). e Brachycera
may have evolved from a bibionomorphan ancestor, but this has so far proved dicult
to conrm (Lambkin et al. 2013). e systematic order of Table 2 follows Marshall
(2012), with one exception: Conopoidea is given superfamily status. Families are listed
alphabetically within each superfamily.
Two nematocerous families dominate by absolute numbers of species: the non-
biting midges (Chironomidae) with 780 species, and the true fungus gnats (Mycet-
ophilidae, 691 spp.). ese two families also show the largest number of new species
reported since 1980 (369 and 260 species respectively). Hoveries (Syrphidae, 362
spp.) is the largest brachyceran family, followed closely by tachinid parasitic ies (Ta-
chinidae, 319 spp.). e largest absolute increases are seen in Tachinidae (118 spp.)
and Anthomyiidae (112 spp.).e relative number of moth y (Psychodidae) species
has more than tripled since Hackman’s checklist, mostly due to the work of Jukka
Salmela. Among the larger families, major increases are also seen in the Pipunculidae
(98% increase), Cecidomyiidae (161%), and Chironomidae (90%).
Most Diptera families show a decreasing trend in the number of species with lati-
tude in Europe, but some are genuinely more diverse in the boreal zone (see Kjærand-
sen et al. 2007). e number of species known from Finland compares favorably with
the results from neighboring countries (see Table 3). With 6920 species, Finland has
the highest reported Diptera diversity among the Nordic countries. Only Germany
(9544 species, Schumann 2009) and the Czech Republic (7917 spp., Jedlička et al.
2009) of all the North and Central European countries report signicantly higher na-
tional Diptera faunas. is must, however, be at least partially attributed to dierences
between surveying intensity and the history of various countries. Most countries have
only a few (if any) active dipterologists. Up until the last decade, access to the taxo-
nomic literature required for Diptera identication was restricted to those working in
close cooperation with major taxonomic institutes.
e number of species present in an area does usually increase with the size of the
area (see Ulrich and Buszko 2003 for an example involving European insects). Peter-
son and Meier (2001) presented a species-area curve (of type IV, see Scheiner 2003)
for Diptera species of European countries. Figure 1, based on Table 3, is an updated
version of their gure. Two simple models (linear and logarithmic) were tted to the
data using least squares ts. One should, however, not draw too many conclusions
from these models: the true number of Diptera species present in each country is likely
to be signicantly larger than the known number of species. Furthermore, large – or at
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 7
Table 2. Number of species recorded from Finland for each Diptera family. Systematic order follows Marshall (2012).
Family Finland 2014 Hackman (1980) increase in species #
species # with and without doubtful records species # including doubtful records spp. %
LOWER DIPTERA (NEMATOCERAN FLIES)
Tipulomorpha
Tipuloidea
Cylindrotomidae 7 5 2 40%
Limoniidae 196 144 52 36%
Pediciidae 19 13 646%
Tipulidae 114–115 98 16 16%
Trichoceridae 15–17 13 215%
Psychodomorpha
Pscyhodidae 61–63 14 47 336%
Ptychopteromorpha
Ptychopteridae 7 6 1 17%
Culicomorpha
Culicoidea
Chaoboridae 8 8 0 0%
Culicidae 38 37 13%
Dixidae 16 7 9 129%
Chironomoidea
Ceratopogonidae 97 69 28 41%
Chironomidae 780 411 369 90%
Simuliidae 56 35 21 60%
aumaleidae 1 0 1 --
Bibionomorpha
Anisopodoidea
Anisopodidae 7 5 2 40%
Bibionoidea
Bibionidae 17 13 431%
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
8
Family Finland 2014 Hackman (1980) increase in species #
species # with and without doubtful records species # including doubtful records spp. %
Canthyloscelidae 3 3 0 0%
Mycetobiidae 1 1 0 0%
Pachyneuridae 1 1 0 0%
Scatopsidae 30 26 415%
Sciaroidea
Bolitophilidae 21 17 424%
Cecidomyiidae 355–356 136 219 161%
Diadocidiidae 5 3 2 67%
Ditomyiidae 2 1 1 100%
Keroplatidae 46–47 37 924%
Mycetophilidae 691–692 431 260 60%
Sciaridae 337 207 130 63%
Sciarosoma 1 0 1 --
BRACHYCERA
lower Brachycera
Tabanomorpha
Xylophagoidea
Xylophagidae 5 4 1 25%
Rhagionoidea
Athericidae 1 1 0 0%
Rhagionidae 16–17 14 214%
Tabanoidea
Tabanidae 38–39 37 13%
Stratiomyioidea
Stratiomyidae 29 28 14%
Xylomyidae 1 1 0 0%
Asilomorpha
Asiloidea
Asilidae 35 35 00%
Bombyliidae 18–19 22 -4 -18%
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 9
Family Finland 2014 Hackman (1980) increase in species #
species # with and without doubtful records species # including doubtful records spp. %
Mythicomyiidae 1 1 0 0%
Scenopinidae 3 2 1 50%
erevidae 17 20 -3 -15%
unplaced in Asilomorpha
Acroceridae 5 5 0 0%
Empidoidea
Atelestidae 2 1 1 100%
Brachystomatidae 4 4 0 0%
Dolichopodidae 260 219 41 19%
Hybotidae 143–144 132 11 8%
Empididae 172 154 18 12%
Iteaphila group 3 4 -25%
higher Brachycera, Cyclorrhapha
Phoroidea
Opetiidae 1 1 0 0%
Lonchopteridae 8 3 5 167%
Phoridae 224–234 202 22 11%
Platypezidae 39 20 19 95%
Syrphoidea
Pipunculidae 107 54 53 98%
Syrphidae 362 271 91 34%
Conopoidea
Conopidae 19 19 00%
Diopsoidea
Psilidae 29 26 312%
Tanypezidae 1 1 0 0%
Strongyloph-
thalmyiidae 2 2 0 0%
Megamerinidae 1 1 0 0%
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
10
Family Finland 2014 Hackman (1980) increase in species #
species # with and without doubtful records species # including doubtful records spp. %
Nerioidea
Micropezidae 6 5 1 20%
Pseudopomyzidae 1 1 0 0%
Tephritoidea
Eurygnathomyiidae 1 1 0 0%
Lonchaeidae 41–44 37 411%
Neottiophilidae 2 0 2 --
Pallopteridae 13 10 330%
Piophilidae 15 13 215%
Platystomatidae 2 2 0 0%
Tephritidae 69 61 813%
Ulidiidae 16 14 214%
Lauxanoidea
Chamaemyiidae 27–28 16 11 69%
Lauxaniidae 45 42 37%
Sciomyzoidea
Coelopidae 1 1 0 0%
Dryomyzidae 5 3 2 67%
Heterocheilidae 1 1 0 0%
Phaeomyiidae 2 2 0 0%
Sciomyzidae 73–74 68 710%
Sepsidae 32 24 938%
Opomyzoidea
Agromyzidae 280–281 221 58 26%
Anthomyzidae 15 12 325%
Asteiidae 6 5 1 20%
Aulacigastridae 2 1 1 100%
Clusiidae 12 10 220%
Odiniidae 5 5 0 0%
Opomyzidae 16 13 323%
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 11
Family Finland 2014 Hackman (1980) increase in species #
species # with and without doubtful records species # including doubtful records spp. %
Periscelididae 4 3 1 33%
Carnoidea
Acartophthalmidae 3 3 0 0%
Canacidae 1 1 0 0%
Carnidae 13 12 18%
Chloropidae 150 91 59 65%
Milichiidae 12–13 11 218%
Sphaeroceroidea
Chyromyidae 4 4 0 0%
Heleomyzidae 61 53 815%
Sphaeroceridae 118 97 21 22%
Ephydroidea
Braulidae 1 1 0 0%
Camillidae 3 3 0 0%
Diastatidae 10 5 5 100%
Drosophilidae 67 51 16 31%
Ephydridae 112 106 66%
Hippoboscoidea
Hippoboscidae s. lat.
incl. Nycteribiidae 12 12 00%
Muscoidea
Anthomyiidae 289 177 112 63%
Fanniidae 61 40 22 55%
Muscidae 307–309 253 54 21%
Scathophagidae 85–86 83 22%
Oestroidea
Calliphoridae 45 36 925%
Oestridae 8–9 8 0 0%
Rhiniidae 0–1 1 0 0%
Rhinophoridae 5 4 1 25%
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
12
Family Finland 2014 Hackman (1980) increase in species #
species # with and without doubtful records species # including doubtful records spp. %
Sarcophagidae 64 58 610%
Tachinidae 319 201 118 59%
Nematoceran, total 2932–2942 1741 1191 67%
Brachycera, total 3989–4015 3166 823 26%
Diptera, total 6920–6956 4907 2013 41%
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 13
1000
2000
3000
4000
5000
6000
7000
8000
9000
10000
0100200300400500
Number of species
Country area (1000 km²)
Figure 1. e species-area curve for some Northern, Western and Central European countries based
on Table 3, excluding the Baltic states (see also Fig. 1 in Petersen and Meier 2001). Data for Finland is
marked by a circle, other countries by rectangles. e two lines show linear and logarithmic least-squares
t models for the data.
1000
2000
3000
4000
5000
6000
7000
8000
9000
10000
0 20 40 60 80 100
Number of species
Country populaon (million²)
Figure 2. e species-population curve for some Northern, Western and Central European countries
based on Table 3. Data for Finland is marked by a circle, other countries by rectangles. e two lines show
linear and logarithmic least-squares t models for the data.
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
14
Table 3. Recent Diptera checklists from north and central European countries. e species numbers of
nematoceran, brachyceran, and all Diptera are listed for each country. Species of doubtful occurrence are
not included in the counts.
Country Reference # of species recorded
nematoceran brachyceran all Diptera
Nordic countries
Finland 2932 3989 6920
Sweden Cederberg et al. (2010) 2260 4410 6670
Norway Gammelmo et al. (2010) 1936 3116 5052
Denmark Petersen and Meier (2001) 1327 3034 4361
Baltic countries
Latvia Karpa (2008) – – 1654
Lithuania Pakalniškis et al. (2006) – – 3311
Western and Central Europe
Poland Zatwarnicki (2001) – – 6721
Great Britain Chandler (1998, 2013a) 2844 4210 7054
Ireland Chandler et al. (2008), Chandler (2013b) 1479 1907 3386
the Netherlands Beuk (2002) 1640 3324 4964
Belgium Grootaert et al. (1991) – – 4474
Germany Schumann et al. (1999), Schumann
(2009) – – 9544
Switzerland Merz et al. (1998, 2006) – – 6813
Czech Republic Jedlička et al. (2009) 5162 2755 7917
Slovakia Jedlička et al. (2009) 4460 2380 6840
Hungary Papp et al. (2001) ~1460 ~4090 ~5550
2000
3000
4000
5000
6000
7000
8000
Diptera species (Finland)
Year
1980 checklist
1941 checklist
2014 checklist
Figure 3. Total number of Diptera species known from Finland over time based on the 5-year sum-
maries by Hellén and Silfverberg and the three checklists of Finnish Diptera. e publishing years of
the three checklists are noted.
Checklist of the Diptera (Insecta) of Finland: an introduction and a summary of results 15
least populous countries—countries are more likely to harbor dipterists, who contrib-
ute faunistic records (see Figure 2).
From an accumulation curve of new records over the last century (Figure 3), one
can immediately see that the Finnish fauna is far from completely known. If almost all
species present in the country had already been found, one would expect the rate of
new records to diminish (but not fall to zero as genuine expansive species would still
occasionally arrive). What has actually happened is the opposite; the rate of new dis-
coveries has increased in the last decade. e number of species found as new to Fin-
land during each ve-year period seems to reect the number of active dipterologists
in the country. To a degree, this may represent a backlog from the 1980s and 1990s,
when the number of dipterists collecting in Finland and publishing new faunistic re-
cords was smaller than during the previous 70 years, or in the 21st century. Still, one
could predict that the true number of Diptera species present in Finland may be well
over eight thousand species, including hundreds of still unknown and undescribed
species (Fontaine et al. 2012).
Acknowledgments
Where to begin? Creating this checklist has very much been a collaborative eort, with
many amateur and professional dipterists providing insightful commentary and snip-
pets of data since this project began in the year 2002.
First and foremost, I want to thank all authors of the papers in this issue of ZooK-
eys, and Jukka Salmela, the coeditor of this checklist, for taking responsibility for the
papers dealing with the lower ies. is project would not have been possible without
the existence of Finnish entomological museums and their sympathetic sta: the Finn-
ish Museum of Natural History (MZH), the Zoological Museums of the University of
Turku, University of Oulu, and University of East Finland, the Kuopio Natural Histo-
ry Museum, the Forssa Nature Museum, and the Tampere Museum of Natural Histo-
ry (Vapriikki). Equally critical was the support of Finnish amateur entomologists with
signicant private Diptera collection, including Jari Flinck (Helsinki), Antti Haarto
(Turku), Jari Ilmonen (Karkkila), Jevgeni Jakovlev (Helsinki), Jere Kahanpää (Hel-
sinki), Iiro Kakko (Hämeenlinna), Sakari Kerppola (Helsinki), Lauri Paasivirta (Salo),
Jouni Penttinen (Jyväskylä), Jukka Salmela (Rovaniemi), and Kaj Winqvist (Turku).
In addition, Miroslav Barták, Hans Bartch, Christer Bergström, Peter Chandler, Mi-
lan Chvála, Henry Disney, Neal Evenhuis, Steve Gaimari, Jean-Paul Haenni, Chris-
tian Kehlmaier, Valery Korneyev, Iain McGowan, Bernhard Merz, Verner Michelsen,
Lorenzo Munari, Alexei Polevoi, Marc Pollet, Adrian Pont, Jindřich Roháček, Michael
von Tschirnhaus, Andrzej Woźnica, and eo Zeegers, among others, contributed by
examining collection specimens, providing literature, commenting on preliminary lists
or species, and being supportive and friendly when pestered for answers.
e Finnish Expert Group for Diptera (http://dipteratyoryhma.myspecies.info/)
provided funds and other support, allowing the author and Kaj Winqvist to spend a
Jere Kahanpää / ZooKeys 441: 1–20 (2014)
16
few months examining Finnish Diptera fulltime in 2003–2007. Finally, I would per-
sonally like to thank my friends K. Winqvist and A. Haarto for introducing me to the
fascinating and curious world of ies.
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Jere Kahanpää / ZooKeys 441: 1–20 (2014)
20
Supplementary material 1
Electronic appendix 1: CVS taxon le
Authors: Jere Kahanpää
Data type: (measurement/occurence/multimedia/etc.)
Explanation note: is le is encoded as UTF-8 text with a semicolon as the eld sepa-
rator. In Excel, the CSV le is best opened using the Data/From Text functionality.
e data is formatted as Darwin Core (Darwin Core Task Group 2009) compliant.
e following elds are provided: taxonID [Fauna Europaea URN if available],
scienticName, taxonRank, higherClassication, order, family, genus, subgenus,
specicEpithet, scienticNameAuthorship, identicationQualier, identication-
Remarks, bibliographicCitation, taxonConceptID.
Copyright notice: is dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). e Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Supplementary material 2
Electronic appendix 2: PDF literature references le
Authors: Jere Kahanpää
Data type: References list.
Explanation note: A PDF le with additional references to rst published records is
also provided. Cross-references to Hackman’s 1980 checklist are included. is le
includes an index to genus- and species-level names.
Copyright notice: is dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). e Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland 21
Checklist of the infraorder Tipulomorpha
(Trichoceridae, Tipuloidea) (Diptera) of Finland
Jukka Salmela1, Andrius Petrašiūnas2
1 Metsähallitus, Natural Heritage Services, P.O. Box 8016, FI-96101 Rovaniemi, Finland 2 Vilnius uni-
versity Faculty of Natural Sciences Department of Zoology Ciurlionio st. 21/27 LT 03101 Vilnius, Lithuania
Corresponding author: Jukka Salmela (jukka.e.salmela@gmail.com)
Academic editor: Jere Kahanpää|Received 17 March 2014|Accepted 30 July 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Salmela J, Petrašiūnas A (2014) Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera)
of Finland. In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 21–36. doi: 10.3897/
zookeys.441.7533
Abstract
A checklist of the infraorder Tipulomorpha: families Trichoceridae, Pediciidae, Limoniidae, Cylindroto-
midae and Tipulidae (Diptera) recorded from Finland.
Keywords
Checklist, Finland, Diptera, Trichoceridae, Pediciidae, Limoniidae, Cylindrotomidae, Tipulidae
Introduction
Trichoceridae is a small family of medium sized gnats, mostly of Holarctic distribu-
tion. Larvae live in dierent decaying matter like leaves, wood, fungi or dead mam-
mals. Adults form numerous swarms that are encountered during colder fall and spring
months or warmer winter days in warmer climates or during the short summer months
in northern latitudes.
No comprehensive and up to date key exists for north European adults or larvae,
but a combination of Shtackelberg (1988), Starý (1999) and Petrašiūnas (2009) could
be used. However, Shtackelberg (1988) must be used with caution because several spe-
cies have since been published. List of trichocerids presented here is based on literature
review (see Dahl 1968, Dahl and Alexander 1976), no new material has been studied.
ZooKeys 441: 21–36 (2014)
doi: 10.3897/zookeys.441.7533
www.zookeys.org
Copyright J. Salmela, A. Petrašiˉunas . This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
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Jukka Salmela & Andrius Petrašiūnas / ZooKeys 441: 21–36 (2014)
22
Tipuloidea crane ies (Pediciidae, Limoniidae, Cylindrotomidae and Tipulidae)
belong to the infraorder Tipulomorpha together with trichocerids (Wiegmann et al.
2011). Following Starý (1992), most European authors distinguish four families:
Limoniidae, Tipulidae, Pediciidae and Cylindrotomidae. North American authors,
strongly inuenced by C.P. Alexander, usually prefer a single family, Tipulidae; other
families, sensu Starý, are there subfamilies or tribes (Byers 1992, Petersen et al. 2010).
According to a recent phylogenetic analysis (Petersen et al. 2010), two families are
recognized, Tipulidae and Pediciidae, but the consensus tree of that analysis had many
unresolved polytomies. Crane y larvae occur usually in moist or (semi)aquatic habi-
tats, but larvae may also be found on mosses, in dry soil, fungal fruiting bodies and
dead wood. Cylindrotomids are herbivorous, feeding on vascular plants or mosses.
Most larvae of (semi)aquatic species have open spiracles and are dependent on aerial
oxygen supply, but truly aquatic species are able to withdraw dissolved oxygen through
the cuticle of tracheal gills.
An annotated list of Finnish crane ies was provided by Salmela (2012a) and a few
species were later added in subsequent papers (Salmela 2012b, 2013). Species richness of
Finnish crane ies is negatively correlated with latitude but mire-dwelling crane ies have
anomal species richness gradient (Salmela 2012). Finnish crane ies are rather well known,
but perhaps up to 20 more species could be found from Finland (Salmela 2013).
Checklist
Nematocera Dumeril, 1805
infraorder Tipulomorpha Rohdendorf, 1961
TRICHOCERIDAE Róndani, 1841
CLADONEURA Scudder, 1894
= Diazosma Bergroth, 1913
? Cladoneura hirtipenne (Siebke, 1863)
TRICHOCERA Meigen, 1803
sg. Metatrichocera Dahl, 1966
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Oosterbroek 2014) Europe (Pape et al. 2011) Finland
Tipulomorpha:
Trichoceridae 183 50 15–17 poor-average
Pediciidae 490 63 19 good
Limoniidae 10528 545 196 good
Cylindrotomidae 71 7 7 good
Tipulidae 4280 456 114–115 good
Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland 23
Trichocera gigantea (Dahl, 1967)
Trichocera lutea Becher, 1886
Trichocera mackenziei (Dahl, 1967)
sg. Saltrichocera Krzeminska, 2002
Trichocera annulata Meigen, 1818
? Trichocera arctica Lundström, 1915
Trichocera implicata Dahl, 1976
Trichocera maculipennis Meigen, 1818
Trichocera parva Meigen, 1804
Trichocera regelationis (Linnaeus, 1758)
Trichocera rufescens Edwards, 1921
Trichocera saltator (Harris, 1776)
= fuscata Meigen, 1818
Trichocera sparsa Starý & Martinovský, 1996
sg. Trichocera Meigen, 1803
Trichocera hiemalis (De Geer, 1776)
Trichocera inexplorata (Dahl, 1967)
Trichocera major Edwards, 1921
Trichocera sibirica Edwards, 1920
PEDICIIDAE Osten Sacken, 1860
PEDICIINAE Osten Sacken, 1860
DICRANOTA Zetterstedt, 1838
sg. Dicranota Zetterstedt, 1838
Dicranota bimaculata (Schummel, 1829)
Dicranota crassicauda Tjeder, 1972
Dicranota guerini Zetterstedt, 1838
sg. Paradicranota Alexander, 1934
Dicranota gracilipes Wahlgren, 1905
Dicranota pavida (Haliday, 1833)
Dicranota robusta Lundström, 1912
Dicranota subtilis Loew, 1871
sg. Rhaphidolabis Osten Sacken, 1869
Dicranota exclusa (Walker, 1848)
PEDICIA Latreille, 1809
sg. Crunobia Kolenati, 1859
Pedicia straminea (Meigen, 1838)
sg. Pedicia Latreille, 1809
Pedicia rivosa (Linnaeus, 1758)
TRICYPHONA Zetterstedt, 1837
sg. Tricyphona Zetterstedt, 1837
Tricyphona immaculata (Meigen, 1804)
Tricyphona livida Madarassy, 1881
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Tricyphona schummeli Edwards, 1921
Tricyphona unicolor (Schummel, 1829)
ULINAE Alexander, 1920
ULA Haliday, 1833
sg. Ula Haliday, 1833
Ula bolitophila Loew, 1869
Ula kiushiuensis Loew, 1869
Ula mixta Starý, 1983
Ula mollissima Haliday, 1833
= inconclusa (Walker, 1856)
Ula sylvatica (Meigen, 1818)
LIMONIIDAE Speiser, 1909
CHIONEINAE Róndani, 1841
ARCTOCONOPA Alexander, 1955
Arctoconopa forcipata (Lundström, 1915)
Arctoconopa obscuripes (Zetterstedt, 1851)
Arctoconopa zonata (Zetterstedt, 1851)
CHEILOTRICHIA Rossi, 1848
sg. Cheilotrichia Rossi, 1848
Cheilotrichia imbuta (Meigen, 1818)
sg. Empeda Osten Sacken, 1869
Cheilotrichia areolata (Lundström, 1912)
Cheilotrichia cinerascens (Meigen, 1804)
= nubila (Schummel, 1829)
Cheilotrichia neglecta (Lackschewitz, 1927)
CHIONEA Dalman, 1816
sg. Chionea Dalman, 1816
Chionea araneoides Dalman, 1816
= minuta Tahvonen, 1932
Chionea crassipes Boheman, 1846
sg. Sphaeconophilus Becker, 1912
Chionea lutescens Lundström, 1907
= brevirostris Tahvonen, 1932
CRYPTERIA Bergroth, 1913
Crypteria limnophiloides Bergroth, 1913
ERIOCONOPA Stary, 1976
Erioconopa diuturna (Walker, 1848)
Erioconopa trivialis (Meigen, 1818)
ERIOPTERA Meigen, 1803
sg. Erioptera Meigen, 1803
Erioptera beckeri Kuntze, 1914
= fuscipennis misid.
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Erioptera divisa (Walker, 1848)
Erioptera avata (Westho, 1882)
= gemina Tjeder, 1967
Erioptera griseipennis Meigen, 1838
Erioptera lutea Meigen, 1804
Erioptera nielseni Meijere, 1921
Erioptera pederi Tjeder, 1969
Erioptera sordida Zetterstedt, 1838
Erioptera squalida Loew, 1871
Erioptera tordi Tjeder, 1973
GNOPHOMYIA Osten Sacken, 1860
Gnophomyia acheron Alexander, 1950
Gnophomyia lugubris (Zetterstedt, 1838)
Gnophomyia viridipennis (Gimmerthal, 1847)
GONEMPEDA Alexander, 1924
Gonempeda ava (Schummel, 1829)
GONOMYIA Meigen, 1818
sg. Gonomyia Meigen, 1818
Gonomyia abscondita Lackschewitz, 1935
= lucidula misid.
Gonomyia bida Tonnoir, 1920
Gonomyia dentata Meijere, 1920
Gonomyia simplex Tonnoir, 1920
Gonomyia stackelbergi Lackschewitz, 1935
Gonomyia tenella (Meigen, 1818)
sg. Teuchogonomyia Alexander, 1968
Gonomyia edwardsi Lackschewitz, 1925
HOPLOLABIS Osten Sacken, 1869
sg. Parilisia Savchenko, 1976
Hoplolabis areolata (Siebke, 1872)
Hoplolabis vicina (Tonnoir, 1920)
MOLOPHILUS Curtis, 1833
sg. Molophilus Curtis, 1833
Molophilus appendiculatus (Staeger, 1840)
Molophilus ater (Meigen, 1804)
Molophilus bidus Goetghebuer, 1920
Molophilus bihamatus Meijere, 1918
Molophilus cinereifrons Meijere, 1920
Molophilus corniger Meijere, 1920
Molophilus crassipygus Meijere, 1918
= ochrescens Edwards, 1938
Molophilus avus Goetghebuer, 1920
Molophilus griseus (Meigen, 1804)
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Molophilus medius Meijere, 1918
Molophilus obscurus (Meigen, 1818)
Molophilus occultus Meijere, 1918
Molophilus ochraceus (Meigen, 1818)
Molophilus propinquus (Egger, 1863)
Molophilus pullus Lackschewitz, 1927
NEOLIMNOPHILA Alexander, 1920
Neolimnophila carteri (Tonnoir, 1921)
Neolimnophila placida (Meigen, 1830)
ORMOSIA Rondani, 1856
sg. Oreophila Lackschewitz, 1935
Ormosia sootryeni (Lackschewitz, 1935)
sg. Ormosia Rondani, 1856
Ormosia brevinervis (Lundström, 1907)
Ormosia clavata (Tonnoir, 1920)
Ormosia depilata Edwards, 1938
Ormosia fascipennis (Zetterstedt, 1838)
Ormosia hederae (Curtis, 1835)
Ormosia lineata (Meigen, 1804)
Ormosia loxia Starý, 1983
Ormosia pseudosimilis (Lundström, 1912)
Ormosia rucauda (Zetterstedt, 1838)
Ormosia staegeriana Alexander, 1953
PHYLLOLABIS Osten Sacken, 1877
Phyllolabis macroura (Siebke, 1863)
RHABDOMASTIX Skuse, 1890
Rhabdomastix borealis Alexander, 1924
Rhabdomastix laeta (Loew, 1873)
Rhabdomastix parva (Siebke, 1863)
RHYPHOLOPHUS Kolenati, 1860
Rhypholophus haemorrhoidalis (Zetterstedt, 1838)
Rhypholophus varius (Meigen, 1818)
SCLEROPROCTA Edwards, 1938
Scleroprocta pentagonalis (Loew, 1973)
Scleroprocta sororcula (Zetterstedt, 1851)
= danica (Nielsen, 1923)
SYMPLECTA Meigen, 1830
sg. Psiloconopa Zetterstedt, 1838
Symplecta lindrothi (Tjeder, 1955)
Symplecta meigeni (Zetterstedt, 1838)
Symplecta stictica (Meigen, 1818)
sg. Symplecta Meigen, 1830
Symplecta chosenensis (Alexander, 1940)
Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland 27
Symplecta hybrida (Meigen, 1804)
Symplecta mabelana (Alexander, 1955)
Symplecta scotica (Edwards, 1938)
sg. Trimicra Osten Sacken, 1861
Symplecta pilipes (Fabricius, 1787)
TASIOCERA Skuse, 1890
sg. Dasymolophilus Goetghebuer, 1920
Tasiocera exigua Savchenko, 1973
Tasiocera fuscescens (Lackschewitz, 1940)
Tasiocera murina (Meigen, 1818)
LIMNOPHILINAE Bigot, 1854
ADELPHOMYIA Bergroth, 1891
Adelphomyia punctum (Meigen, 1818)
AUSTROLIMNOPHILA Alexander, 1920
sg. Archilimnophila Alexander, 1934
Austrolimnophila harperi (Alexander, 1926)
Austrolimnophila unica (Osten Sacken, 1869)
DICRANOPHRAGMA Osten Sacken, 1860
sg. Brachylimnophila Alexander, 1966
Dicranophragma adjunctum (Walker, 1848)
Dicranophragma separatum (Walker, 1848)
= nemorale misid.
= leucophaea misid.
ELOEOPHILA Rondani, 1856
Eloeophila maculata (Meigen, 1804)
Eloeophila mundata (Loew, 1971)
Eloeophila submarmorata (Verrall, 1887)
Eloeophila trimaculata (Zetterstedt, 1838)
Eloeophila verralli (Bergroth, 1912)
EPIPHRAGMA Osten Sacken, 1860
sg. Epiphragma Osten Sacken, 1860
Epiphragma ocellare (Linnaeus, 1761)
EUPHYLIDOREA Alexander, 1972
Euphylidorea dispar (Meigen, 1818)
= lineola misid.
Euphylidorea meigeni (Verrall, 1886)
Euphylidorea phaeostigma (Schummel, 1829)
EUTONIA van der Wulp, 1874
Eutonia barbipes (Meigen, 1804)
HEXATOMA Latreille, 1809
sg. Hexatoma Latreille, 1809
Hexatoma fuscipennis (Curtis, 1836)
= nubeculosa misid.
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IDIOPTERA Macquart, 1834
Idioptera linnei Oosterbroek, 1992
= fasciata (Linnaeus, 1767) preocc.
Idioptera pulchella (Meigen, 1830)
= pulchella var. macropteryx (Tjeder, 1955)
LIMNOPHILA Macquart, 1834
sg. Limnophila Macquart, 1834
Limnophila pictipennis (Meigen, 1818)
Limnophila schranki Oosterbroek, 1992
= punctata (Schrank, 1781)
NEOLIMNOMYIA Seguy, 1937
sg. Neolimnomyia Seguy, 1937
Neolimnomyia batava (Edwards, 1938)
PARADELPHOMYIA Alexander, 1936
sg. Oxyrhiza de Meijere, 1946
Paradelphomyia fuscula (Loew, 1873)
Paradelphomyia nigrina (Lackschewitz, 1940)
PHYLIDOREA Bigot, 1854
sg. Macrolabina Savchenko, 1986
Phylidorea nigronotata (Siebke, 1870)
sg. Paraphylidorea Savchenko, 1986
Phylidorea fulvonervosa (Schummel, 1829)
= Euphylidorea lineola misid.
sg. Phylidorea Bigot, 1854
Phylidorea abdominalis (Staeger, 1840)
Phylidorea bicolor (Meigen, 1804)
Phylidorea ferruginea (Meigen, 1818)
Phylidorea heterogyna (Bergroth, 1913)
Phylidorea longicornis (Schummel, 1829)
= glabricula (Meigen, 1830)
Phylidorea nervosa (Schummel, 1829)
= nigricollis (Meigen, 1830)
Phylidorea squalens (Zetterstedt, 1838)
Phylidorea umbrarum (Krogerus, 1937)
PILARIA Sintenis, 1889
Pilaria decolor (Zetterstedt, 1851)
Pilaria discicollis (Meigen, 1818)
Pilaria meridiana (Staeger, 1840)
Pilaria nigropunctata (Agrell, 1945)
= fuscipennis misid.
Pilaria scutellata (Staeger, 1840)
PSEUDOLIMNOPHILA Alexander, 1919
sg. Pseudolimnophila Alexander, 1919
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Pseudolimnophila lucorum (Meigen, 1818)
LIMONIINAE Speiser, 1909
ACHYROLIMONIA Alexander, 1965
Achyrolimonia decemmaculata (Loew, 1873)
ANTOCHA Osten Sacken, 1860
sg. Antocha Osten Sacken, 1860
Antocha vitripennis (Meigen, 1830)
ATYPOPHTHALMUS Brunetti, 1911
sg. Atypophthalmus Brunetti, 1911
Atypophthalmus inustus (Meigen, 1818)
DICRANOMYIA Stephens, 1829
sg. Dicranomyia Stephens, 1829
Dicranomyia aperta Wahlgren, 1904
Dicranomyia autumnalis (Staeger, 1840)
Dicranomyia consimilis (Zetterstedt, 1838)
Dicranomyia didyma (Meigen, 1804)
Dicranomyia distendens Lundström, 1912
Dicranomyia frontalis (Staeger, 1840)
Dicranomyia halterata Osten Sacken, 1869
Dicranomyia handlirschi Lackschewitz, 1928
Dicranomyia hyalinata (Zetterstedt, 1851)
Dicranomyia longipennis (Schummel, 1829)
Dicranomyia mitis (Meigen, 1830)
Dicranomyia modesta (Meigen, 1818)
Dicranomyia moniliformis Doane, 1900
Dicranomyia omissinervis Meijere, 1918
Dicranomyia patens Lundström, 1907
Dicranomyia radegasti Starý, 1993
Dicranomyia sera (Walker, 1848)
Dicranomyia terraenovae Alexander, 1920
Dicranomyia ventralis (Schummel, 1829)
Dicranomyia zernyi Lackschewitz, 1928
sg. Glochina Meigen, 1830
Dicranomyia liberta Osten Sacken, 1860
Dicranomyia tristis (Schummel, 1829)
= subtristis Alexander, 1924
= schineri misid.
= schineriana misid.
sg. Idiopyga Savchenko, 1987
Dicranomyia danica Kuntze, 1919
Dicranomyia esbeni (Nielsen, 1940)
Dicranomyia halterella Edwards, 1921
Dicranomyia intricata Alexander, 1927
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Dicranomyia klefbecki (Tjeder, 1941)
Dicranomyia lulensis (Tjeder, 1969)
Dicranomyia magnicauda Lundström, 1912
Dicranomyia murina (Zetterstedt, 1851)
Dicranomyia ponojensis Lundström, 1912
Dicranomyia stigmatica (Meigen, 1830)
sg. Melanolimonia Alexander, 1965
Dicranomyia caledonica Edwards, 1926
Dicranomyia morio (Fabricius, 1787)
Dicranomyia occidua Edwards, 1926
Dicranomyia ruventris (Strobl, 1901)
Dicranomyia stylifera Lackschewitz, 1928
sg. Numantia Bigot, 1854
Dicranomyia fusca (Meigen, 1804)
DICRANOPTYCHA Osten Sacken, 1860
Dicranoptycha cinerascens (Meigen, 1818)
Dicranoptycha fuscescens (Schummel, 1829)
DISCOBOLA Osten Sacken, 1865
Discobola annulata (Linnaeus, 1758)
Discobola caesarea (Osten Sacken, 1854)
ELEPHANTOMYIA Osten Sacken, 1860
sg. Elephantomyia Osten Sacken, 1860
Elephantomyia edwardsi Lackschewitz, 1932
Elephantomyia krivosheinae Savchenko, 1976
HELIUS Lepeletier & Serville, 1828
sg. Helius Lepeletier & Serville, 1828
Helius avus (Walker, 1856)
Helius longirostris (Meigen, 1818)
Helius pallirostris Edwards, 1921
LIBNOTES Westwood, 1876
sg. Afrolimonia Alexander, 1965
Libnotes ladogensis (Lackschewitz, 1940)
LIMONIA Meigen, 1803
sg. Limnobia Meigen, 18018
Limonia badia (Walker, 1848)
Limonia avipes (Fabricius, 1787)
Limonia macrostigma (Schummel, 1829)
Limonia maculicosta (Coquillett, 1905)
Limonia messaurea Mendl, 1971
Limonia nubeculosa Meigen, 1804
Limonia phragmitidis (Schrank, 1781)
= tripunctata Fabricius, 1781
Limonia stigma (Meigen, 1818)
Limonia sylvicola (Schummel, 1829)
Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland 31
Limonia trivittata (Schummel, 1829)
LIPSOTHRIX Loew, 1873
Lipsothrix ecucullata Edwards, 1938
Lipsothrix errans (Walker, 1848)
METALIMNOBIA Matsumura, 1911
sg. Metalimnobia Matsumura, 1911
Metalimnobia bifasciata (Schrank, 1781)
Metalimnobia charlesi Salmela & Starý, 2008
Metalimnobia quadrimaculata (Linnaeus, 1761)
Metalimnobia quadrinotata (Meigen, 1818)
Metalimnobia tenua Savchenko, 1976
Metalimnobia zetterstedti (Tjeder, 1968)
= elegans Zetterstedt, 1838 preocc.
NEOLIMONIA Alexander, 1964
Neolimonia dumetorum (Meigen, 1804)
ORIMARGA Osten Sacken, 1869
sg. Orimarga Osten Sacken, 1869
Orimarga attenuata (Walker, 1848)
Orimarga juvenilis (Zetterstedt, 1851)
RHIPIDIA Meigen, 1818
sg. Rhipidia Meigen, 1818
Rhipidia maculata Meigen, 1818
= duplicata (Doane, 1900)
Rhipidia uniseriata Schiner, 1864
CYLINDROTOMIDAE Schiner, 1863
CYLINDROTOMA Macquart, 1834
Cylindrotoma borealis Peus, 1952
Cylindrotoma distinctissima (Meigen, 1818)
Cylindrotoma nigriventris Loew, 1849
DIOGMA Edwards, 1938
Diogma caudata Takahashi, 1960
Diogma glabrata (Meigen, 1818)
PHALACROCERA Schiner, 1863
Phalacrocera replicata (Linnaeus, 1758)
TRIOGMA Schiner, 1863
Triogma trisulcata (Schummel, 1829)
TIPULIDAE Latreille, 1802
CTENOPHORINAE Osten Sacken, 1887
tribe Ctenophorini Osten Sacken, 1887
CTENOPHORA Meigen, 1803
sg. Ctenophora Meigen, 1803
Ctenophora aveolata (Fabricius, 1794)
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Ctenophora guttata Meigen, 1818
Ctenophora nigriceps (Tjeder, 1949)
Ctenophora pectinicornis (Linnaeus, 1758)
DICTENIDIA Brulle, 1833
Dictenidia bimaculata (Linnaeus, 1761)
PHOROCTENIA Coquillett, 1910
Phoroctenia vittata (Meigen, 1830)
tribe Tanypterini Savchenko, 1966
TANYPTERA Latreille, 1804
sg. Tanyptera Latreille, 1804
Tanyptera atrata (Linnaeus, 1758)
Tanyptera nigricornis (Meigen, 1818)
DOLICHOPEZINAE Osten Sacken, 1887
DOLICHOPEZA Curtis, 1825
sg. Dolichopeza Curtis, 1825
Dolichopeza albipes (Strom, 1768)
Dolichopeza bida Osterbrook & Lantsov, 2011
= nitida of authors
TIPULINAE Latreille, 1802
tribe Prionocerini Savchenko, 1966
PRIONOCERA Loew, 1844
Prionocera abscondita Lackschewitz, 1933
Prionocera chosenicola Alexander, 1945
= dimidiata misid.
= absentiva misid.
Prionocera pubescens Loew, 1844
Prionocera recta Tjeder, 1948
= lapponica Tjeder, 1948
= lackschewitzi Mannheims, 1951
Prionocera ringdahli Tjeder, 1948
Prionocera serricornis (Zetterstedt, 1838)
Prionocera subserricornis (Zetterstedt, 1851)
= proxima Lackschewitz, 1933
Prionocera turcica (Fabricius, 1787)
Prionocera woodoorum Brodo, 1987
tribe Tipulini Latreille, 1802
ANGAROTIPULA Savchenko, 1961
Angarotipula tumidicornis (Lundström, 1907)
NEPHROTOMA Meigen, 1803
Nephrotoma aculeata (Loew, 1871)
Nephrotoma analis (Schummel, 1833)
Nephrotoma appendiculata (Pierre, 1919)
Nephrotoma cornicina (Linnaeus, 1758)
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Nephrotoma crocata (Linnaeus, 1758)
Nephrotoma dorsalis (Fabricius, 1781)
Nephrotoma avescens (Linnaeus, 1758)
Nephrotoma lundbecki (Nielsen, 1907)
Nephrotoma lunulicornis (Schummel, 1833)
Nephrotoma pratensis (Linnaeus, 1758)
Nephrotoma quadristriata (Schummel, 1833)
Nephrotoma relicta (Savchenko, 1973)
Nephrotoma scurra (Meigen, 1818)
Nephrotoma submaculosa Edwards, 1928
Nephrotoma tenuipes (Riedel, 1910)
NIGROTIPULA Hudson & Vane-Wright, 1969
Nigrotipula nigra (Linnaeus, 1758)
TIPULA Linnaeus, 1758
sg. Acutipula Alexander, 1924
Tipula fulvipennis De Geer, 1776
Tipula maxima Poda, 1761
sg. Arctotipula Alexander, 1934
Tipula salicetorum Siebke, 1870
= nigricornis Zetterstedt, 1851
sg. Beringotipula Savchenko, 1961
Tipula unca Wiedemann, 1817
= hortensis Meigen, 1818
sg. Dendrotipula Savchenko, 1964
Tipula avolineata Meigen, 1804
sg. Emodotipula Alexander, 1966
Tipula obscuriventris Strobl, 1900
sg. Lindnerina Mannheims, 1965
Tipula bistilata Lundström, 1907
Tipula subexcisa Lundström, 1907
sg. Lunatipula Edwards, 1931
Tipula anis Schummel, 1833
Tipula circumdata Siebke, 1863
= livida misid.
Tipula fascipennis Meigen, 1818
Tipula humilis Staeger, 1840
Tipula laetabilis Zetterstedt, 1838
= dilatata Schummel, 1833
Tipula limitata Schummel, 1833
Tipula lunata Linnaeus, 1758
= luna mistake
? Tipula peliostigma Schummel, 1833
Tipula recticornis Schummel, 1833
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Tipula selene Meigen, 1830
Tipula trispinosa Lundström, 1907
Tipula vernalis Meigen, 1804
sg. Odonatisca Savchenko, 1956
Tipula nodicornis Meigen, 1818
= juncea Meigen, 1818
sg. Platytipula Matsumura, 1916
Tipula luteipennis Meigen, 1830
Tipula melanoceros Schummel, 1833
sg. Pterelachisus Róndani, 1842
Tipula cinereocincta Lundström, 1907
Tipula crassicornis Zetterstedt, 1838
Tipula irrorata Macquart, 1826
Tipula jutlandica Nielsen, 1947
Tipula kaisilai Mannheims, 1954
Tipula laetibasis Alexander, 1934
Tipula luridorostris Schummel, 1833
Tipula matsumuriana ssp. pseudohortensis Lackschewitz, 1932
Tipula mutila Wahlgren, 1905
Tipula octomaculata Savchenko, 1964
Tipula pabulina Meigen, 1818
Tipula pauli Mannheims, 1964
Tipula pseudoirrorata Goetghebuer, 1921
Tipula recondita Pilipenko & Salmela, 2012
Tipula stenostyla Savchenko, 1964
Tipula submarmorata Schummel, 1833
= meigeni Mannheims, 1966
Tipula truncorum Meigen, 1830
Tipula varipennis Meigen, 1818
= pseudovariipennis misid.
Tipula wahlgreni Lackschewitz, 1925
Tipula winthemi Lackschewitz, 1932
sg. Savtshenkia Alexander, 1965
Tipula alpium Bergroth, 1888
Tipula benesignata Mannheims, 1954
Tipula confusa van der Wulp, 1883
= marmorata Meigen, 1818
Tipula gimmerthali Lackschewitz, 1925
Tipula grisescens Zetterstedt, 1851
Tipula interserta Riedel, 1913
Tipula invenusta Riedel, 1919
Tipula limbata Zetterstedt, 1838
Tipula obsoleta Meigen, 1818
Checklist of the infraorder Tipulomorpha (Trichoceridae, Tipuloidea) (Diptera) of Finland 35
Tipula pagana Meigen, 1818
Tipula signata Staeger, 1840
Tipula subnodicornis Zetterstedt, 1838
sg. Schummelia Edwards, 1931
Tipula variicornis Schummel, 1833
sg. Tipula Linnaeus, 1758
Tipula paludosa Meigen, 1830
= oleracea misid.
Tipula subcunctans Alexander, 1921
= czizeki de Jong, 1925
sg. Vestiplex Bezzi, 1924
Tipula excisa Schummel, 1833
Tipula hortorum Linnaeus, 1758
Tipula laccata Lundström & Frey, 1916
Tipula montana Curtis, 1834
Tipula nubeculosa Meigen, 1804
Tipula pallidicosta Pierre, 1924
Tipula scripta Meigen, 1830
Tipula sintenisi Lackschewitz, 1933
Tipula tchukchi Alexander, 1934
= bo Mannheims, 1967
sg. Yamatotipula Matsumura, 1916
Tipula chonsaniana Alexander, 1945
Tipula coerulescens Lackschewitz, 1923
Tipula couckei Tonnoir, 1921
Tipula fendleri Mannheims, 1963
Tipula freyana Lackschewitz, 1936
Tipula lateralis Meigen, 1804
Tipula marginella eowald, 1980
= marginata Meigen, 1818
Tipula moesta Riedel, 1919
Tipula montium Egger, 1863
Tipula pierrei Tonnoir, 1921
= solstitialis Westho, 1879
Tipula pruinosa Wiedemann, 1817
Tipula quadrivittata Staeger, 1840
Excluded species
Trichocera japonica Matsumura, 1915
Twenty-two previously reported crane y species were deleted from the Finnish list by
Salmela (2012a); these are not repeated here.
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References
Byers GW (1992) Crane ies – three families or one. Acta Zoologica Cracoviensia 35: 37–41.
Dahl C (1968) Notes on the taxonomy and distribution of Trichoceridae (Dipt. Nemat.) from
Finland and adjacent areas of the USSR. Opuscula Entomologica 33(3): 365–370.
Dahl C, Alexander CP (1976) A world catalogue of Trichoceridae Kertész, 1902 (Diptera).
Entomologica scandinavica 7(1): 7–18. doi: 10.1163/187631276X00027.
Oosterbroek P (2014) Catalogue of the Craneies of the World. Online available at http://
nlbif.eti.uva.nl/ccw/ (latest version 21 March 2014)
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Petersen MJ, Bertone MA, Wiegmann BM, Courtney GW (2010) Phylogenetic synthesis of
morphological and molecular data reveals new insights into the higher-level classication
of Tipuloidea (Diptera). Systematic Entomology 35: 526–545. doi: 10.1111/j.1365-
3113.2010.00524.x
Petrašiūnas A (2009) Taxonomic and phylogenetic review of the genus Trichocera Meigen, 1803
(Diptera, Trichoceridae). PhD esis, Vilniaus Universitetas. http://vddb.library.lt/fedora/
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A Journal of Entomology 2012: 913710. doi: 10.1155/2012/913710
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Trichoceridae). Entomological problems 30(1): 1–10.
Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA,
Cassel BK, Bayless KM, Heimberg AM,Wheeler BM, Peterson KJ, Pape T, Sinclair BJ,
Skevington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Otto U, Kampmeier GE,
ompson FC, Grimaldi DA, Beckenbach AT, Courtney GW, Friedrich M, Meier R, Yeats
DK (2011) Episodic radiations in the y tree of life. Proceedings of the National Academy
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Checklist of the familes Chaoboridae, Dixidae, aumaleidae Psychodidae... 37
Checklist of the familes Chaoboridae, Dixidae,
Thaumaleidae Psychodidae and Ptychopteridae
(Diptera) of Finland
Jukka Salmela1, Lauri Paasivirta2, Gunnar M. Kvifte3
1 Metsähallitus, Natural Heritage Services, P.O. Box 8016, FI-96101 Rovaniemi, Finland 2 Ruuhikosken-
katu 17 B 5, 24240 Salo, Finland 3 Department of Limnology, University of Kassel, Heinrich-Plett-Str. 40,
34132 Kassel-Oberzwehren, Germany
Corresponding author: Jukka Salmela (jukka.e.salmela@gmail.com)
Academic editor: J. Kahanpää|Received 17 March 2014|Accepted 22 May 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Salmela J, Paasivirta L, Kvifte GM (2014) Checklist of the familes Chaoboridae, Dixidae, aumaleidae
Psychodidae and Ptychopteridae (Diptera) of Finland. In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of
Finland. ZooKeys 441: 37–46. doi: 10.3897/zookeys.441.7532
Abstract
A checklist of the families Chaoboridae, Dixidae, aumaleidae, Psychodidae and Ptychopteridae (Diptera)
recorded from Finland is given. Four species, Dixella dyari Garret, 1924 (Dixidae), reticus tridactilis
(Kincaid, 1899), Panimerus albifacies (Tonnoir, 1919) and P. przhiboroi Wagner, 2005 (Psychodidae) are
reported for the rst time from Finland.
Keywords
Finland, Diptera, species list, biodiversity, faunistics
Introduction
Psychodidae or moth ies are an intermediately diverse family of nematocerous ies,
comprising over 3000 species world-wide (Pape et al. 2011). Its taxonomy is still very
unstable, and multiple conicting classications exist (Duckhouse 1987, Vaillant 1990,
Ježek and van Harten 2005). e nomenclature used in the Finnish check-list follows the
ZooKeys 441: 37–46 (2014)
doi: 10.3897/zookeys.441.7532
www.zookeys.org
Copyright Jukka Salmela et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jukka Salmela et al. / ZooKeys 441: 37–46 (2014)
38
Norwegian one (Kvifte et al. 2011). For dierences between the taxonomy used herein
and that used in previous works on Finnish Psychodidae (e.g. Hackman 1980, Salmela
2003, 2005, Salmela et al. 2007, Autio and Salmela 2010) see Kvifte et al. (2011). Two
species, Pneumia nubila and Pneumia palustris, listed in the previous check-lists by Hack-
man (1980) and Wagner (2013) have not been conrmed through study of material. Of
these, P. palustris have been suggested to be synonymous with Pneumia gracilis (Eaton,
1893) (Withers 1989). Both these species have been linked to some taxonomic confusion
in the past, and thus the record of P. nubila may represent Pneumia trivialis. e present
check-list contains the rst Finnish records of the species reticus tridactilis (Kincaid,
1899), Panimerus albifacies (Tonnoir, 1919) and P. przhiboroi Wagner, 2005 (see Notes).
Finnish Psychodidae are very well studied compared to other Nordic countries, how-
ever new species are still being discovered at a regular rate. Most of these are new country
records, but in recent years the fauna has also yielded some species new to science (Salmela
et al. 2012, Kvifte unpubl.). is trend is likely to continue. Probably between twelve and
thirty further species can be expected to be found in the Finnish fauna. ere are no good
identication sources available for Psychodids, as all works suer from taxonomic or no-
menclatural errors and/or insucient descriptions. e best works relevant to the Finnish
fauna are Vaillant (1971-1983), Withers (1989) and Wagner (1997c, genera only).
Phantom midges (Chaoboridae) are close relatives to mosquitoes (Culicidae) and
meniscus midges (Dixidae) (Saether 1970, 1997). ere are around 50 known species
of phantom midges in the world and 15 in the palearctic region, of which ve are
holoarctic species. In Finland, there are eight species in three genera (Hirvenoja 1960,
Hackman 1980, in which the species in the genus Mochlonyx were still doubtful).
e only strictly lacustrine species in Finland is Chaoborus avicans. e other
species live in small shless lakes, ponds and springs where their predators are mainly
dragon ies, dytiscid beetles and frogs. Chaoborus nyblaei and Cryophila lapponica are
found from small shless ponds on fells and ponds created by melting water. However,
in 2013 larvae of C. nyblaei were found from the boreal zone, NE Lapland (Savukoski)
from a small but permanent pond. Chaoborus pallidus lives in eutrophic lakes and small
lakes in southern and central Finland (Hirvenoja 1960, 1963, Paasivirta 2002). How-
ever, there have not been any systematic surveys of the fauna across the biogeographic
provinces and the conservation statuses of the species have not been evaluated. Finnish
species can be identied by using Saether (1997, 2002).
Meniscus midges (Dixidae) belong to infraorder Culicomorpha. ere are nearly
200 known species of which 53 occur in the Palaearctic region. In Finland two genera
and 16 species are present (Hackman 1980, Salmela 2003, Salmela et al. 2007). One
species (Dixella dyari Garret, 1924) is reported here for the rst time from Finland (see
Notes). Larvae of meniscus midges dwell in or slightly above water meniscus, having
a characteristic U-shaped position (Wagner 1997a). Members of the genus Dixella are
associated with lentic or slow owing waters, such as ponds, lake shores and bog pools.
Species of the genus Dixa are lotic, dwelling in running waters of varying size. Two
Finnish species, Dixa dilatata and D. submaculata prefer springs and spring brooks.
e Finnish fauna is generally rather well known, although three species have hitherto
Checklist of the familes Chaoboridae, Dixidae, aumaleidae Psychodidae... 39
been collected from single localities. One northern Dixa species close to D. dilatata is
still undescribed. A majority of the species can be identied by using Martini (1928),
Peus (1934, 1936), Vaillant (1967) and Disney (1999).
Trickle midges (aumaleidae) is a small family belonging to infraorder Culicomor-
pha. Globally, nearly 200 species are known of which 76 are European. Central European
mountains harbor a rich assemblage of trickle midges (Wagner 2002), at least if compared
to the poor diversity (5 spp.) in Fennoscandian area (Wagner 1997b, Andersen et al.
2013). Only one species, aumalea truncata, has been hitherto recorded from Finland
(see the discussion in Andersen et al. 2013). Larvae of trickle midges dwell on hygropetric
substrates or splash zones of running waters (Wagner 1997b). aumalea truncata occurs
mainly in northern Finland, both boreal and subarctic ecoregions, and adult specimens
have been caught around springs and headwater streams (Salmela 2003, Salmela 2008).
Phantom crane ies (Ptychopteridae) belong to infraorder Ptychopteromorpha,
being a sister group of Psychdomorpha and Culicomorpha (Wiegmann et al. 2011).
Ptychopteridae is a small family with over 69 species in the world and 14 in Europe.
Seven species are known from Finland, all belonging to the genus Ptychoptera. Only
one Finnish species, Ptychoptera minuta, is widespread, locally abundant and almost
ubiquitous. Other species are much less frequent, occurring in peatlands, headwa-
ter streams, springs and swamps. Adult Ptychoptera supercially resemble Tipuloidea
crane ies, being slender and having long legs. Larvae of Ptychoptera live in shallow
aquatic habitats, usually in soft substrates (Andersson 1997). Finnish species can be
identied by using Andersson (1997) and Krzeminski (1986).
Checklist
Nematocera Dumeril, 1805
infraorder Psychodomorpha Hennig, 1968
PSYCHODIDAE Newman, 1834
PSYCHODINAE Newman, 1834
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Psychodomorpha:
Psychodidae 2958 493 61–63 average
Culicomorpha:
Chaoboridae 54 9 8 good
Dixidae 186 32 16 average
aumaleidae 182 76 1 good
Ptychopteromorpha:
Ptychopteridae 69 (Wagner et al. 2008) 14 7 good
Jukka Salmela et al. / ZooKeys 441: 37–46 (2014)
40
tribe Pericomaini Enderlein, 1935
PARABAZARELLA Vaillant, 1983
Parabazarella subneglecta (Tonnoir, 1922)
BERDENIELLA Vaillant, 1976
Berdeniella freyi (Berdén, 1954)
CLYTOCERUS Eaton, 1904
sg. Boreoclytocerus Duckhouse, 1978
Clytocerus ocellaris (Meigen, 1818)
Clytocerus rivosus (Tonnoir, 1919)
Clytocerus tetracorniculatus Wagner, 1977
PERICOMA Walker, 1856
sg. Pachypericoma Vaillant, 1978
Pericoma blandula Eaton, 1893
Pericoma nielseni Kvifte, 2010
= formosa Nielsen, 1964 (preocc. Meunier, 1905)
sg. Pericoma Walker, 1856
Pericoma rivularis Berdén, 1954
PNEUMIA Enderlein, 1935
Pneumia borealis (Berdén, 1954)
Pneumia bucegica (Vaillant, 1981)
Pneumia mutua (Eaton, 1893)
? Pneumia nubila (Meigen, 1818)
? Pneumia palustris (Meigen, 1818)
Pneumia pilularia (Tonnoir, 1940)
Pneumia stammeri (Jung, 1954)
Pneumia trivialis (Eaton, 1893)
Pneumia ussurica (Wagner, 1994)
ULOMYIA Walker, 1856
Ulomyia cognata (Eaton, 1893)
Ulomyia fuliginosa (Meigen, 1804)
tribe Psychodini Newman, 1834
PHILOSEPEDON Eaton, 1904
Philosepedon balkanicus Krek, 1970
Philosepedon humeralis (Meigen, 1818)
Philosepedon soljani Krek, 1971
PSYCHODA Latreille, 1796
Psychoda albipennis Zetterstedt, 1850
= parthenogenetica Tonnoir, 1940
Psychoda alternata Say, 1824
Psychoda brevicornis Tonnoir, 1940
Psychoda buxtoni Withers, 1988
Psychoda cinerea Banks, 1894
Checklist of the familes Chaoboridae, Dixidae, aumaleidae Psychodidae... 41
Psychoda crassipenis Tonnoir, 1940
Psychoda cultella Salmela, Kvifte & More, 2012
Psychoda erminea Eaton, 1898
Psychoda gemina (Eaton, 1904)
Psychoda grisescens Tonnoir, 1922
Psychoda itoco Tokunaga & Komyo, 1955
Psychoda lativentris Berdén, 1952
Psychoda lobata Tonnoir, 1940
Psychoda minuta Banks, 1894
Psychoda phalaenoides Linnaeus, 1758
Psychoda satchelli Quate, 1955
Psychoda setigera Tonnoir, 1922
Psychoda trinodulosa Tonnoir, 1922
Psychoda uniformata Haseman, 1907
THRETICUS Eaton, 1904
reticus tridactilis (Kincaid, 1899)
TRICHOPSYCHODA Tonnoir, 1922
Trichopsychoda hirtella (Tonnoir, 1919)
tribe Maruinini Enderlein, 1937
LOBULOSA Szabo, 1960
Lobulosa pollex (Berdén, 1954)
TONNOIRIELLA Vaillant, 1971
Tonnoiriella nigricauda (Tonnoir, 1919)
tribe Mormiini Enderlein, 1937
MORMIA Enderlein, 1935
Mormia niesiolowskii Wagner, 1985
Mormia strobli (Ježek, 1986)
tribe Paramormiini Enderlein, 1937
PARAJUNGIELLA Vaillant, 1972
Parajungiella consors (Eaton, 1893)
Parajungiella ellisi (Withers, 1987)
Parajungiella pseudolongicornis Wagner, 1975
LEPISEODINA Enderlein, 1937
Lepiseodina rothschildi (Eaton, 1912)
PANIMERUS Eaton, 1913
sg. Panimerus Eaton, 1913
Panimerus albomaculatus (Wahlgren, 1904)
Panimerus notabilis (Eaton, 1893)
sg. Psycmera Ježek, 1984
Panimerus albifacies (Tonnoir, 1919)
Panimerus intellegus (Jung, 1956)
Panimerus przhiboroi Wagner, 2005
Jukka Salmela et al. / ZooKeys 441: 37–46 (2014)
42
PARAMORMIA Enderlein, 1935
Paramormia polyscoidea (Krek, 1970)
Paramormia ustulata (Walker, 1856)
PERIPSYCHODA Enderlein, 1935
Peripsychoda auriculata (Curtis, 1839)
Peripsychoda fusca (Macquart, 1826)
TELMATOSCOPUS Eaton, 1904
Telmatoscopus advena (Eaton, 1893)
Telmatoscopus similis Tonnoir, 1922
SYCORACINAE Rondani, 1856
SYCORAX Haliday in Curtis, 1839
Sycorax silacea Haliday in Curtis, 1839
infraorder Culicomorpha Hennig, 1948
superfamily Culicoidea Meigen, 1818
CHAOBORIDAE Newman, 1834
CHAOBORUS Lichtenstein, 1800
sg. Chaoborus Lichtenstein, 1800
Chaoborus crystallinus (De Geer, 1776)
Chaoborus avicans (Meigen, 1830)
Chaoborus obscuripes (van der Wulp, 1859)
sg. Peusomyia Sæther, 1970
Chaoborus pallidus (Fabricius, 1794)
sg. Schadanophasma Dyar & Shannon, 1924
Chaoborus nyblaei (Zetterstedt, 1838)
CRYOPHILA Edwards, 1930
Cryophila lapponica (Martini, 1928)
MOCHLONYX Loew, 1844
Mochlonyx fuliginosus Felt, 1905
= M. martinii Edwards, 1930
= M. velutinus of authors
Mochlonyx velutinus (Ruthe, 1831)
= M. culiciformis (De Geer, 1776) preocc.
DIXIDAE Schiner, 1868
DIXA Meigen, 1818
Dixa dilatata Strobl, 1900
Dixa nebulosa Meigen, 1830
Dixa puberula Loew, 1849
Dixa submaculata Edwards, 1920
DIXELLA Dyar & Shannon, 1924
Dixella aestivalis (Meigen, 1818)
Dixella amphibia (De Geer, 1776)
Checklist of the familes Chaoboridae, Dixidae, aumaleidae Psychodidae... 43
Dixella autumnalis (Meigen, 1838)
Dixella borealis (Martini, 1928)
Dixella dyari (Garret, 1924)
Dixella licornis (Edwards, 1926)
Dixella hyperborea (Bergroth, 1889)
Dixella laeta (Loew, 1849)
Dixella naevia (Peus, 1934)
Dixella nigra (Staeger, 1840)
Dixella obscura (Loew, 1849)
Dixella serotina (Meigen, 1818)
THAUMALEIDAE Bezzi, 1913
THAUMALEA Ruthe, 1831
aumalea truncata Edwards, 1929
infraorder PtychopteromorphaWood & Borkent, 1986
superfamily Ptychopteroidea Osten Sacken, 1862
PTYCHOPTERIDAE Osten Sacken, 1862
PTYCHOPTERA Meigen, 1803
sg. Paraptychoptera Tonnoir, 1919
Ptychoptera lacustris Meigen, 1830
Ptychoptera paludosa Meigen, 1804
sg. Ptychoptera Meigen, 1803
Ptychoptera albimana (Fabricius, 1787)
Ptychoptera contaminata (Linnaeus, 1758)
Ptychoptera hugoi Tjeder, 1968
Ptychoptera minuta Tonnoir, 1919
Ptychoptera scutellaris Meigen, 1818
Excluded species
Dixa maculata Meigen, 1818. ere is one female specimen collected from Russia,
Kantalaks (=Kandalaksha) that belongs to Dixella borealis.
Notes
reticus tridactilis: Ostrobotthnia borealis pars borealis, Rovaniemi, Nammalik-
kokivalo, 13.6.–11.7.2004, J. Salmela leg.; Regio kuusamoensis, Taivalkoski,
Poikaloukusanoja, 1.6.–1.7.2006, J. Salmela leg.; Regio kuusamoensis, Taivalko-
ski, Syväoja 1.6.-1.7.2006, J. Salmela leg.
Jukka Salmela et al. / ZooKeys 441: 37–46 (2014)
44
Panimerus albifacies: Regio aboensis, Turku, Pomponrahka-Isosuo mire conserva-
tion area, 1.8.–28.9.2011, J. Salmela & E. Nummela leg.
Panimerus przhiboroi: Ostrobotthnia borealis pars borealis, Tervola, Ruuttulammi
conservation area, 4.6.–2.7.2012 J. Salmela leg. (New for European fauna)
Psychoda albipennis Zetterstedt has been conrmed to occur in Finland by Bo-W.
Svensson (female specimen from Tavastia australis, South Finland, Tampere,
2009, J. Salmela leg.).
Dixella dyari: Lapponia enontekiensis, Enontekiö, Pallas-Yllästunturi National Park,
Lumikero E, 25.8.2010, J. Salmela leg.
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Checklist of the Culicidae (Diptera) in Finland 47
Checklist of the family Culicidae (Diptera) in Finland
Larry Huldén1, Lena Huldén2
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
2 University of Helsinki, Department of Agricultural Sciences, P.O. Box 27, FI-00014 University of Helsinki,
Finland
Corresponding author: Larry Huldén (larry.hulden@helsinki.)
Academic editor: J. Kahanpää|Received 17 April 2014|Accepted 22 August 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Huldén L, Huldén L (2014) Checklist of the Culicidae (Diptera) in Finland. In: Kahanpää J, Salmela J (Eds)
Checklist of the Diptera of Finland. ZooKeys 441: 47–51. doi: 10.3897/zookeys.441.7743
Abstract
A checklist of the Culicidae (Diptera) recorded from Finland is provided.
Keywords
Checklist, Finland, Diptera, Culicidae, mosquitoes
Introduction
e family Culicidae comprises 3539 extant species world-wide. e Finnish fauna is
relatively small in relation to world fauna. Because of medical importance this family
has been extensively taxonomically studied but still new species are discovered. e
generic and subgeneric classications of Culicidae follow Reinert et al. 2009. Because of
some confusing use of names in the rst half of the twentieth century in the Finnish lit-
erature, all available names are included in the list for clarity. e most important work
on the Finnish mosquitoes was done by Pirkka Utrio (1975, 1976, 1977, 1978, 1979).
Number of species:
World: 3539 extant species + 25 fossil species (Harbach 2014)
Europe: 104 species (Snow and Ramsdale 2004)
Finland: 38 species
ZooKeys 441: 47–51 (2014)
doi: 10.3897/zookeys.441.7743
www.zookeys.org
Copyright Larry Huldén, Lena Huldén. This is an open access article distributed under the terms of the Creative Commons Attribution License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Larry Huldén & Lena Huldén / ZooKeys 441: 47–51 (2014)
48
Checklist
suborder Nematocera Dumeril, 1805
infraorder Culicomorpha Hennig, 1948
superfamily Culicoidea Meigen, 1818
CULICIDAE Meigen, 1818
ANOPHELINAE Grassi, 1900
Anopheles Meigen, 1818
sg. Anopheles Meigen, 1818
Anopheles beklemishevi Stegni & Kabanova, 1976
= maculipennis auct. in part
Anopheles claviger (Meigen, 1804)
Anopheles messeae Falleroni, 1926
= claviger (of Fabricius, 1805) auct. nec Meigen, 1804
= maculipennis auct. in part
CULICINAE Meigen, 1818
Aedes Meigen, 1818
Aedes cinereus Meigen, 1818
Aedimorphus eobald, 1903
Aedes vexans (Meigen, 1830)
Coquillettidia Dyar, 1905
Coquillettidia richiardii (Ficalbi, 1889)
Culex Linnaeus, 1758
sg. Culex Linnaeus, 1758
Culex pipiens Linnaeus, 1758
Culex torrentium Martini, 1925
sg. Neoculex Dyar, 1905
Culex territans Walker, 1856
= apicalis auct. nec Adams, 1903
Culiseta Felt 1904
= eobaldia Neveu-Lemaire, 1902 preocc.
sg. Culicella Felt, 1904
Culiseta morsitans (eobald, 1901)
Culiseta ochroptera Peus, 1935
sg. Culiseta Felt, 1904
Culiseta alaskaensis (Ludlow, 1906)
Culiseta annulata (Schrank, 1776)
Culiseta bergrothi (Edwards, 1921)
= glaphyroptera auct. nec (Shiner, 1864)
Culiseta subochrea (Edwards, 1921)
Dahliana Reinert, Harbach & Kitching, 2006
Dahliana geniculata (Olivier, 1791)
Ochlerotatus Lynch Arribalzaga, 1881
Checklist of the Culicidae (Diptera) in Finland 49
sg. Ochlerotatus Lynch Arribalzaga, 1881
Ochlerotatus annulipes (Meigen, 1830)
Ochlerotatus euedes (Howard, Dyar & Knab, 1913)
= beklemishevi Denisova, 1955
Ochlerotatus cantans (Meigen, 1818)
= waterhousei (eobald in Waterhouse, 1905)
= maculatus auct. nec (Meigen, 1804)
= rusticus auct. nec (Rossi,1790)
Ochlerotatus caspius (Pallas, 1771)
Ochlerotatus cataphylla Dyar, 1916
Ochlerotatus communis (De Geer, 1776)
= nemorosus (Meigen, 1818)
Ochlerotatus cyprius Ludlow, 1920
= freyi Edwards, 1921
Ochlerotatus diantaeus Howard, Dyar & Knab, 1912
Ochlerotatus dorsalis (Meigen, 1830)
Ochlerotatus excrucians (Walker, 1848)
Ochlerotatus avescens (Müller, 1776)
= lutescens (Fabricius, 1775)
= variegatus (Schrank, 1781)
Ochlerotatus hexodontus Dyar, 1919
Ochlerotatus impiger (Walker, 1848)
= parvulus (Edwards 1921)
= nearcticus auct. nec (Dyar, 1922)
Ochlerotatus intrudens Dyar, 1919
Ochlerotatus leucomelas (Meigen, 1804)
Ochlerotatus nigrinus (Eckstein, 1918)
= sticticus auct. nec Meigen 1828
Ochlerotatus nigripes (Zetterstedt, 1838)
= alpinus auct. nec (Linnaeus) nomen dubium
Ochlerotatus pionips Dyar, 1919
Ochlerotatus pullatus (Coquillett, 1904)
Ochlerotatus punctodes Dyar, 1922
Ochlerotatus punctor (Kirby, 1837)
Ochlerotatus riparius Dyar & Knab, 1907
Notes
Anopheles maculipennis Meigen, 1818. Not recorded in Finland. All previous reports
refer to An. messeae or An. beklemishevi.
Anopheles claviger (of Fabricius, 1805). Mistaken name usage for Anopheles macu-
lipennis s. lat. in the early 20th century Finnish literature which then followed the
Italian tradition.
Larry Huldén & Lena Huldén / ZooKeys 441: 47–51 (2014)
50
Culex pipiens f. molestus Forskåhl, 1775. is is not a valid species level taxon. It is a
man biting form of the mainly bird biting Culex pipiens. Both have been reported
from Finland.
Dahliana geniculata (Oliver, 1791). A possibly occasional specimen of this species
was collected in Western Finland, veried by Christine Dahl (Itämies 1981).
Ochlerotatus alpinus L. Frey (1921) included this invalid species name (see Harbach
2014) in the Finnish list with a question mark. Hellén (1931) withdrew it from the
Finnish list. According to Utrio (1977), Frey refers to nigripes (Zetterstedt, 1838).
Ochlerotatus behningi Martini, 1926. e species is listed for both Finland and Swe-
den in Dahl (1997). Lundström et al. (2013) do not accept the report for Sweden
because of absence of documented locality. Based on the same argument, we do
not include this species in the Finnish list.
Ochlerotatus (Rusticoidus) rusticus (Rossi, 1790). Dahl (1997) listed this spe-
cies for Finland. ere are, however, no conrmed collections of the species
in Finland. Frey (1921) listed Aedes maculatus Meigen (with A. waterhousei
eobald as a presumed synonym) from Finland. According to Natvig (1948),
Meigen made a mistake in 1830 when he stated that “maculatus is the male
of cantans” instead of “reptans is the male of cantans”, leading to a switch in
synonyms. According to Minar (1990) the correct combinations should be as
follows: maculatus Meigen is a synonym of rusticus (Rossi) and both reptans
(Meigen) and waterhousei (eobald) are synonyms of cantans (Meigen). From
this context, it is obvious that Frey refers to the common species cantans and
not to the southern species rusticus.
Ochlerotatus sticticus (Meigen, 1838). e species was reported from Finland (Kilpis-
järvi) by Frey (1932) and listed for Finland in Frey and Storå (1941). According
to Natvig (1948) and Utrio (1979) this record refers to Aedes nigrinus (Eckstein).
In addition, it was reported by Syrjämäki (1960) in Finland (Hanko: Tvärminne)
but later withdrawn by the author according to Utrio (1979, personal communi-
cation). It was also included for Finland in Dahl (1997), probably based on the
former reports. According to Gutsevich et al. (1970) this species occurs in Russian
Karelia. us, A. sticticus is not included in the Finnish list.
Culex apicalis Adams, 1903. Originally reported by Martini (1928) from Finland. All
European reports of this Nearctic species before 1950 represent the closely related
Holarctic species Culex territans (Becker et al. 2003).
References
Becker N, Petrić D, Zgomba M, Boase C, Dahl C, Lane J, Kaiser A (2003) Mosquitoes and their
control. Kluwer Academic / Plenum Publishers, New York, Boston, Dordrecht, London,
Moscow, 498 pp. doi: 10.1007/978-1-4757-5897-9
Dahl C (1997) Diptera Culicidae, Mosquitoes. In: Nilsson AN (Ed) Aquatic Insects of North
Europe – A taxonomic Handbook. Volume 2. Apollo Books, Stenstrup, 163–186.
Checklist of the Culicidae (Diptera) in Finland 51
Frey R (1921) Provisorisk förteckning over Finlands culicider. Meddelanden af Societas pro
Fauna et Flora Fennica 47: 98–102.
Frey R (1932) [Aedes sticticus Meigen, ny för Finland]. Notulae Entomologicae 12: 64–65.
Frey R, Storå R (1941) Enumeratio Insectorum Fenniae. VI. Diptera: Culicidae. Helsingin
Hyönteisvaihtoyhdistys, Helsinki, 52 pp.
Gutsevich AV, Monchadskii AS, Stackelberg AA (1970) Fauna USSR. Diptera. III. 4. Mosquitoes.
Family Culicidae. Nauka, Leningrad, 384 pp. [in Russian]
Hackman W (1980) Checklist of Finnish Diptera I. Nematocera and Brachycera. Notulae
Entomologicae 60: 17–48.
Harbach R (2014) Mosquito Taxonomic Inventory. http://mosquito-taxonomic-inventory.
info/valid-species-list [accessed July 29th 2014]
Itämies J (1981) Aedes geniculatus Suomelle uusi hyttynen (Diptera, Culicidae). Notulae Ento-
mologicae 61: 207–208.
Lundström JO, Schäfer ML, Hesson JC, Blomgren E, Lindström A, Wahlqvist P, Halling A, Ha-
gelin A, Ahlm C, Evander M, Broman T, Forsman, Persson Vinnersten TZ (2013) e geo-
graphic distribution of mosquito species in Sweden. Journal of the European Mosquito Control
Association 31: 21–35. http://e-m-b.org/sites/e-m-b.org/les/JEMCA%2831%2921-35.pdf
Martini E (1928) Dixinae und Corethrinae des Museums zu Helsingfors. Notulae Entomo-
logicae 8: 33–35.
Minar J (1990) Family Culicidae. In: Soos A, Papp L, Oosterbroek P (Eds) Catalogue of
palaearctic Diptera 2. Akademiai Kiadó, Budapest, 74–113.
Natvig LR (1948) Contributions to the knowledge of the Danish and Fennoscandian mosqui-
toes, Culicini. Norsk entomologisk Tidsskrift. Supplement 1: 1–567. http://www.mosqu-
itocatalog.org/les/pdfs/094600-2.pdf
Reinert JF, Harbach RE, Kitching IJ (2009) Phylogeny and classication of Aedini (Diptera:
Culicidae). Zoological Journal of the Linnean Society 157: 700−794. doi: 10.1111/j.1096-
3642.2009.00570.x
Snow KR, Ramsdale CD (2004) Culicidae. In: Beuk P, Pape T, de Jong H (Eds) Fauna Europaea:
Diptera. Fauna Europaea version 1.0. http://www.faunaeur.org [accessed July 29th 2014]
Syrjämäki A (1960) Preliminary observations on the daily rhythm of biting activity of some
Aedes species (Dipt., Culicidae). Annales Entomologici Fennici 26: 92–97.
Utrio P (1975) Anopheles claviger (Meig.), Aedes pionips Dyar and A. beklemishevi Den. (Diptera,
Culicidae) found in Finland. Notulae Entomologicae 55: 63–64. http://www.mosquito-
catalog.org/les/pdfs/134700-0.pdf
Utrio P (1976) Identication key to Finnish mosquito larvae (Diptera, Culicidae). Annales
Agriculturae Fenniae 15: 128–136. http://www.mosquitocatalog.org/les/pdfs/13470.pdf
Utrio P (1977) A checklist of Finnish mosquitoes (Diptera, Culicidae, including Aedes annulipes,
new to Finland. Notulae Entomologicae 57: 130–132. http://www.mosquitocatalog.org/
les/pdfs/134700-2.pdf
Utrio P (1978) Distribution of mosquitoes (Diptera, Culicidae) attracted by man in Finland in early
July. Notulae Entomologicae 58: 107–114.
Utrio P (1979) Geographic distribution of mosquitoes (Diptera, Culicidae) in eastern Fennoscandia.
Notulae Entomologicae 59: 105–123. http://www.mosquitocatalog.org/les/pdfs/134700-3.pdf
Larry Huldén & Lena Huldén / ZooKeys 441: 47–51 (2014)
52
Checklist of the family Ceratopogonidae (Diptera) of Finland 53
Checklist of the family Ceratopogonidae
(Diptera) of Finland
Larry Huldén1, Lena Huldén2
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
2 University of Helsinki, Department of Agricultural Sciences, P.O. Box 27, FI-00014 University of Helsinki,
Finland
Corresponding author: Larry Huldén (larry.hulden@helsinki.)
Academic editor: J. Kahanpää|Received 17 April 2014|Accepted 22 August 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Huldén L, Huldén L (2014) Checklist of the family Ceratopogonidae (Diptera) of Finland. In: Kahanpää J,
Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 53–61. doi: 10.3897/zookeys.441.7742
Abstract
A checklist of the family Ceratopogonidae (Diptera) recorded from Finland is provided.
Keywords
Checklist, Finland, Diptera, Ceratopogonidae
Introduction
e biting midges (Ceratopogonidae) have only occasionally been studied by Finnish
entomologists. Most notable are the studies by Lundström (1910, 1916), Munster-
hjelm (1920), Storå (1939) and Hämäläinen (1988). e last mentioned reference is
not published in print but included because the determinations have been checked
by Peter Havelka. Other species reports were given by Frey (1918), Forsius (1924),
Krogerus (1960), Meinander (1977) and Hildén et al. (2010). A considerable part of
the species information has been contributed by foreign specialists, Edwards (1924),
Clastrier (1961, 1991), Delecolle et al. (1983) Borkent and Bissett (1990), Szadziewski
(2001), Ashe et al. (2007), Dominiak et al. (2007), Dominiak and Szadziewski (2010)
and Dominiak (2012).
ZooKeys 441: 53–61 (2014)
doi: 10.3897/zookeys.441.7742
www.zookeys.org
Copyright Larry Huldén, Lena Huldén. This is an open access article distributed under the terms of the Creative Commons Attribution License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Larry Huldén & Lena Huldén / ZooKeys 441: 53–61 (2014)
54
A World Catalogue of Ceratopogonidae is maintained by Art Borkent (2014),
updated Jan. 20, 2014.
Number of species:
World: 6180 extant species + 274 fossil species (Borkent 2014)
Europe: 567 species (Fauna Europaea, Szadzievski, Borkent and Dominiak 2014)
Finland: 97 species
Faunistic knowledge level in Finland: medium
Checklist
suborder Nematocera Dumeril, 1805
infraorder Culicomorpha Hennig, 1948
superfamily Chironomoidea Newman, 1834
CERATOPOGONIDAE Newman, 1834
FORCIPOMYIINAE Lenz, 1934
ATRICHOPOGON Kieer, 1906
sg. Atrichopogon Kieer, 1906
Atrichopogon brunnipes (Meigen, 1804)
Atrichopogon fuscus (Meigen, 1804)
= fossicola (Kieer, 1922)
Atrichopogon longicalcar Remm, 1961
Atrichopogon minutus (Meigen, 1830)
Atrichopogon pavidus (Winnertz, 1852)
sg. Meloehelea Wirth, 1956
Atrichopogon lucorum (Meigen, 1818)
Atrichopogon meloesugans Kieer, 1922
= winnertzi auct. nec Goetghebuer, 1922
Atrichopogon oedemerarum Storå, 1939
sg. Psilokempia Enderlein, 1936
Atrichopogon appendiculatus (Goetghebuer, 1920)
Atrichopogon forcipatus (Winnertz, 1852)
Atrichopogon maculatus (Lundström, 1910)
= minutus var. maculatus Lundström, 1910
Atrichopogon rostratus (Winnertz, 1852)
FORCIPOMYIA Meigen, 1818
sg. Caloforcipomyia Saunders, 1957
Forcipomyia glauca Mace, 1934
sg. Forcipomyia Meigen, 1818
Forcipomyia bipunctata (Linnaeus, 1767)
Forcipomyia ciliata (Winnertz, 1852)
Checklist of the family Ceratopogonidae (Diptera) of Finland 55
Forcipomyia costata (Zetterstedt, 1838)
= picea (Winnertz, 1852)
Forcipomyia kaltenbachi (Winnertz, 1852)
Forcipomyia myrmecophila (Egger, 1863)
Forcipomyia nigra (Winnertz, 1852)
sg. Microhelea Kieer, 1917
Forcipomyia fuliginosa (Meigen, 1818)
= brevimana (Lundström, 1910)
sg. yridomyia Saunders, 1925
Forcipomyia monilicornis (Coquillett, 1905)
= hirta (Lundström, 1910)
sg. Trichohelea Goetghebuer, 1920
Forcipomyia eques (Johanssen, 1908)
DASYHELEINAE Lenz, 1934
DASYHELEA Kieer, 1911
sg. Dasyhelea Kieer, 1911
Dasyhelea bensoni Edwards, 1933
= vernalis Remm, 1979
Dasyhelea bilineata Goetghebuer, 1920
= saxicola (Edwards, 1929)
= obscura auct. nec (Winnertz, 1852)
= versicolor auct. nec (Winnertz, 1852)
Dasyhelea pallidiventris (Goetghebuer, 1931)
sg. Dicryptoscena Enderlein, 1936
Dasyhelea modesta (Winnertz, 1852)
= aestiva (Winnertz, 1852)
= longipalpis Kieer, 1913
Dasyhelea notata Goetghebuer, 1920
sg. Pseudoculicoides Malloch, 1915
Dasyhelea corinneae Gosseries, 1991
= scutellata (Meigen, 1830), preocc.
Dasyhelea avoscutellata (Zetterstedt, 1850)
= egens (Winnertz, 1852)
sg. Sebessia Remm, 1979
Dasyhelea holosericea (Meigen, 1804)
CERATOPOGONINAE Newman 1834
tribe Culicoidini Kieer, 1911
Culicoides Latreille, 1809
Culicoides achrayi Kettle & Lawson, 1955
Culicoides albicans (Winnertz, 1852)
= vexans auct. nec (Staeger, 1839)
Culicoides chiopterus (Meigen, 1830)
Culicoides circumscriptus Kieer, 1918
Larry Huldén & Lena Huldén / ZooKeys 441: 53–61 (2014)
56
Culicoides dewul Goetghebuer, 1936
Culicoides fascipennis (Staeger 1839)
Culicoides festivipennis Kieer, 1914
= odiblis Austen, 1921
Culicoides grisescens Edwards, 1939
Culicoides impunctatus Goetghebuer, 1920
Culicoides kibunensis Tokunaga 1937
= cubitalis Edwards in Edwards et al. 1939
Culicoides nubeculosus (Meigen, 1830)
Culicoides obsoletus (Meigen, 1818)
= varius (Winnertz, 1852)
Culicoides pallidicornis (Kieer 1919)
Culicoides pictipennis (Staeger, 1839)
Culicoides pseudoheliophilus Callot & Kremer, 1961
Culicoides pulicaris (Linnaeus, 1758)
Culicoides punctatus (Meigen, 1804)
Culicoides riethi Kieer, 1914
Culicoides riouxi Callot & Kremer, 1961
Culicoides salinarius Kieer, 1914
Culicoides scoticus Downes & Kettle, 1952
Culicoides segnis Campbell & Pelham-Clinton 1960
Culicoides sphagnumensis Williams, 1955
= carjalaensis Glukhova, 1957
Culicoides stigma (Meigen, 1818)
Culicoides subfasciipennis Kieer, 1919
tribe Ceratopogonini Newman, 1834
ALLUAUDOMYIA Kieer, 1913
Alluaudomyia needhami omsen, 1935
= pentaspila Remm & Glukhova, 1971
Alluaudomyia quadripunctata (Goetghebuer, 1934)
BRACHYPOGON Kieer, 1899
sg. Isohelea Kieer, 1917
Brachypogon aquilonalis (Clastrier, 1961)
Brachypogon hyperboreus (Clastrier 1961)
Brachypogon incompletus (Kieer, 1925)
= lapiae Clastrier, 1961
Brachypogon nitidulus (Edwards, 1921)
= nniae Clastrier, 1961
Brachypogon sociabilis (Goetghebuer, 1920)
CERATOPOGON Meigen, 1800
Ceratopogon lacteipennis Zetterstedt, 1838
Ceratopogon niveipennis Meigen, 1818
= candidatus Winnertz, 1852
Checklist of the family Ceratopogonidae (Diptera) of Finland 57
SCHIZOHELEA Kieer, 1917
Schizohelea leucopeza (Meigen, 1804)
= copiosa Winnertz, 1852
= xanthopeza (Clastrier, 1963)
SERROMYIA Meigen, 1818
Serromyia atra (Meigen, 1818)
Serromyia femorata (Meigen, 1804)
Serromyia subinermis Kieer, 1919
tribe Heteromyiini Wirth, 1962
CLINOHELEA Kieer, 1917
Clinohelea unimaculata (Macquart, 1826)
= variegata (Winnertz, 1852)
tribe Sphaeromiini Newman 1834
MALLOCHOHELEA Wirth, 1962
Mallochohelea inermis (Kieer, 1909)
Mallochohelea munda (Loew, 1864)
= dentata (Kieer, 1909)
Mallochohelea nitida (Macquart, 1826)
Mallochohelea setigera (Loew, 1864)
Mallochohelea vernalis Remm, 1965
PROBEZZIA Kieer, 1906
= Dicrobezzia Kieer, 1919
Probezzia concinna (Meigen, 1818)
= venusta v. concinna (Meigen, 1818)
Probezzia seminigra (Panzer, 1798)
= venusta (Meigen, 1818)
= borealis (Clastrier, 1962)
SPHAEROMIAS Curtis, 1829
Sphaeromias fasciatus (Meigen, 1804)
Sphaeromias pictus (Meigen, 1818)
= candidatus (Loew, 1856) preocc.
tribe Palpomyiini Enderlein, 1936
BEZZIA Kieer, 1899
Bezzia annulipes (Meigen, 1830)
Bezzia bicolor (Meigen, 1804)
Bezzia circumdata (Staeger, 1839)
= solstitialis (Winnertz, 1852)
Bezzia coracina (Zetterstedt, 1850)
= albipes (Winnertz, 1852)
Bezzia fascispinosa Clastrier, 1962
Bezzia avicornis (Staeger, 1839)
= avipalpis (Winnertz, 1852)
Bezzia fuscifemoris Remm, 1971
Larry Huldén & Lena Huldén / ZooKeys 441: 53–61 (2014)
58
Bezzia leucogaster (Zetterstedt, 1850)
= xanthocephala Goetghebuer, 1911
Bezzia nigritula (Zetterstedt, 1838)
Bezzia nobilis (Winnertz, 1852)
= leucogaster var. punctata Lundström, 1910
Bezzia winnertziana Kieer, 1919
= gracilis (Winnertz, 1852) nec Haliday, 1833
PALPOMYIA Meigen, 1818
Palpomyia armipes (Meigen, 1838)
= rupecta (Winnertz, 1852)
Palpomyia distincta (Haliday, 1833)
Palpomyia avipes (Meigen, 1804)
= hortulana Meigen, 1818
Palpomyia avitarsis (Meigen, 1838)
Palpomyia lineata (Meigen, 1804)
= stagnalis Clastrier, 1962
Palpomyia lundstroemi Remm, 1981
= bispinosa Lundström, 1916, preocc.
Palpomyia serripes (Meigen, 1818)
= transfuga (Staeger, 1839)
Palpomyia tibialis (Meigen, 1818)
Notes
Atrichopogon meloesugans Kieer, 1922. is species was restored from synonymy
with A. winnertzi Goetghebuer, 1922 by Szadziewski et al. (2007).
Dasyhelea modesta (Winnertz, 1852). Old records reported before 1988 of this species (as
D. aestiva (Winnertz, 1852) or D. longipalpis (Winnertz, 1852)) are questionable ac-
cording to Dominiak and Szadziewski (2010). Records by Hämäläinen (1988) are valid.
Ceratopogon neglectus Winnertz, 1852. Reported by Lundström (1910) as Culicoides
neglectus from Finland. Szadziewski (1984) transferred it to Dasyhelea. Remm
(1988) considered it a doubtful Ceratopogonidae. Because the type specimen is
lost and the original description is insucient it is considered nomen dubium by
Dominiak and Szadziewski (2010). e species is removed from the Finnish list.
Culicoides vexans (Staeger, 1839). Reported from Finland by Lundström (1910)
who listed Culicoides albicans as a probable synonym. C. vexans was omitted in
Frey and Storå (1941) and Havelka (1978) but included in Hackman (1980) and
Szadziewski et al. (1997) (citing Hackman). According to Delecolle et al. (1983)
Lundström’s C. vexans represents C. albicans. C. albicans is included in the Finnish
list and C. vexans is removed from the Finnish fauna.
Culicoides reconditus (Campbell & Pelham-Clinton, 1960) and C. riouxi Callot &
Kremer, 1961. C. riouxi was treated as a synonym of C. reconditus in Borkent and
Checklist of the family Ceratopogonidae (Diptera) of Finland 59
Wirth (1997) but restored from synonymy in Borkent (2012). Delecolle et al.
(1983) reported C. riouxi as a valid species from Finland (N: Tvärminne). e
record of C. reconditus by Hämäläinen (1988) from Finland (Ta: Riihimäki) was
checked by Havelka but because he considered C. riouxi and C. reconditus syno-
nyms the identity of Hämäläinen’s record remains unclear (see Havelka and Aqui-
lar 1999 and Huldén et al. 2008 in accordance with Borkent and Wirth 1997). As
a consequence C. reconditus is not included in the Finnish list.
Culicoides salinarius Kieer, 1914. According to Delecolle et al. (1983) this species
is indistinguishable from C. manchuriensis Tokunaga, 1941 which is reported from
Russian Karelia by Glukhova (1957).
Probezzia concinna (Meigen, 1818). e record listed by Krogerus (1960) from
Kuusamo district (by mistake as ‘coracina’) is tentatively interpreted to be in
the same locality in Finland as Probezzia seminigra, Kuusamo: Isosuo (Storå
1937). P. concinna was listed as a variety of P. seminigra in Frey and Storå
(1941). Storå obviously identied both taxa. e species is here interpreted as
a Finnish species.
Palpomyia avipennis Meigen, 1804. Misspelled species name for P. avipes (Frey
and Storå 1941).
Palpomyia aterrima Goetghebuer, 1921. e record listed by Krogerus (1960) from
Kuusamo district is from the Russian part according to locality data in Storå
(1939). Not recorded in Finland.
Palpomyia spinipes (Meigen) in Panzer, 1806. e record listed by Krogerus (1960)
from Kuusamo district is from the Russian part according to the bog type informa-
tion. Not recorded in Finland.
Acknowledgements
Art Borkent (Canada) and Patrycja Dominiak (Poland) for corrections and updated
faunistic information on species.
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Checklist of the family Chironomidae (Diptera) of Finland 63
Checklist of the family Chironomidae
(Diptera) of Finland
Lauri Paasivirta1
1 Ruuhikoskenkatu 17 B 5, FI-24240 Salo, Finland
Corresponding author: Lauri Paasivirta (lauri.paasivirta@suomi24.)
Academic editor: J. Kahanpää|Received 10 March 2014|Accepted 26 August 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Paasivirta L (2014) Checklist of the family Chironomidae (Diptera) of Finland. In: Kahanpää J, Salmela J
(Eds) Checklist of the Diptera of Finland. ZooKeys 441: 63–90. doi: 10.3897/zookeys.441.7461
Abstract
A checklist of the family Chironomidae (Diptera) recorded from Finland is presented.
Keywords
Finland, Chironomidae, species list, biodiversity, faunistics
Introduction
ere are supposedly at least 15 000 species of chironomid midges in the world
(Armitage et al. 1995, but see Pape et al. 2011) making it the largest family among the
aquatic insects. e European chironomid fauna consists of 1262 species (Sæther and
Spies 2013). In Finland, 780 species can be found, of which 37 are still undescribed
(Paasivirta 2012).
e species checklist written by B. Lindeberg on 23.10.1979 (Hackman 1980)
included 409 chironomid species. Twenty of those species have been removed from
the checklist due to various reasons. e total number of species increased in the 1980s
to 570, mainly due to the identication work by me and J. Tuiskunen (Bergman and
Jansson 1983, Tuiskunen and Lindeberg 1986). ere has been a new rapid increase
in the total number of species since 1998. I have reported 187 additional species, of
which 8 have been new to science, and other researchers have reported 24 additional
ZooKeys 441: 63–90 (2014)
doi: 10.3897/zookeys.441.7461
www.zookeys.org
Copyright Lauri Paasivirta. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
64
species. e chironomid fauna of all biogeographical provinces has been relatively well
surveyed (Paasivirta 2012, including the trapping eciency and the collectors).
Holotypes and paratypes exist for 87 species described from Finland. 23 species
have been found only in Finland, thus making them putatively endemic species. Six of
these species have only been found from the original type locality. 42 species have been
found only from one collecting site in Finland.
Larvae of chironomid midges are found in all types of aquatic habitats. In addi-
tion, some species (130 species, 17%) specialize in semi-aquatic environments like wet
moss on wetlands and shores of lotic waters. A total of 367 species have been identied
from springs and the surrounding habitats in Finland. Sixteen percent of these species
are associated with or specialized in springs and 25% are semi-aquatic species (Paasi-
virta 2007). One hundred and seventy aquatic species have been found from marine
habitats, excluding river estuaries (Paasivirta 2000 and later additions). Sixty ve ad-
ditional species live in the mildly brackish water in the northern part of Bothnian Bay.
ere are only four strictly marine species.
e conservation statuses have been estimated for 667 Finnish chironomid spe-
cies. Eleven species are classied as vulnerable (VU), 21 as near threatened (NT) and
6 species as lacking sucient information (DD) (Penttinen et al. 2010). Since the chi-
ronomid midges have not been extensively studied and no long-term studies have been
performed in Finland, the only useful criteria for the assessment of the conservation
statuses have been either extent of occurrence or area of occupancy.
e nomenclature of the species list is consistent with Fauna Europaea (Sæther and
Spies 2013). Undescribed species are specied with appellations derived from the rst col-
lecting locality. For example Procladius sp. 2 “Palsa” was rst found at a site called Palsa.
e list of Finnish chironomids (with bioprovincial distribution data) was last
updated by Paasivirta (2012).
Number of species:
World: 7053 species (Pape et al. 2011)
Europe: 1262 species (Sæther and Spies 2013)
Finland: 780 species, incl. 37 undescribed species
Faunistic knowledge level in Finland: good
Checklist
Nematocera Dumeril, 1805
infraorder Culicomorpha Hennig, 1948
superfamily Chironomoidea Newman, 1834
CHIRONOMIDAE Newman, 1834
PODONOMINAE ienemann & Edwards, 1937
tribe Boreochlini Brundin, 1966
Checklist of the family Chironomidae (Diptera) of Finland 65
BOREOCHLUS Edwards, 1938
Boreochlus thienemanni Edwards, 1938
LASIODIAMESA Kieer, 1924
Lasiodiamesa armata Brundin, 1966
Lasiodiamesa gracilis (Kieer, 1924)
Lasiodiamesa sphagnicola (Kieer, 1925)
tribe Podonomini ienemann & Edwards, 1937
PAROCHLUS Enderlein, 1912
Parochlus kieeri (Garrett, 1925)
TRICHOTANYPUS Kieer, 1906
Trichotanypus mariae Wirth & Sublette, 1970
Trichotanypus posticalis (Lundbeck, 1898)
TANYPODINAE Skuse, 1889
tribe Anatopyniini Fittkau, 1962
ANATOPYNIA Johannsen, 1905
Anatopynia plumipes (Fries, 1823)
tribe Coelotanypodini Coman, 1978
= Clinotanypodini Lipina, 1928
CLINOTANYPUS Kieer, 1913
Clinotanypus nervosus (Meigen, 1818)
tribe Macropelopiini Zavřel, 1929
APSECTROTANYPUS Fittkau, 1962
Apsectrotanypus trifascipennis (Zetterstedt, 1838)
MACROPELOPIA ienemann, 1916
Macropelopia adaucta Kieer, 1916
= goetghebueri (Kieer, 1918)
Macropelopia nebulosa (Meigen, 1804)
Macropelopia notata (Meigen, 1818)
PSECTROTANYPUS Kieer, 1909
Psectrotanypus varius (Fabricius, 1787)
tribe Natarsiini Roback & Moss, 1978
NATARSIA Fittkau, 1962
Natarsia nugax (Walker, 1856)
Natarsia punctata (Fabricius, 1805)
tribe Pentaneurini Hennig, 1950
ABLABESMYIA Johannsen, 1905
Ablabesmyia longistyla Fittkau, 1962
Ablabesmyia monilis (Linnaeus, 1758)
Ablabesmyia phatta (Egger, 1864)
ARCTOPELOPIA Fittkau, 1962
Arctopelopia barbitarsis (Zetterstedt, 1850)
Arctopelopia griseipennis (van der Wulp, 1859)
Arctopelopia melanosoma (Goetghebuer, 1933)
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
66
CONCHAPELOPIA Fittkau, 1957
Conchapelopia aagaardi Murray, 1987
Conchapelopia hittmairorum Michiels & Spies, 2002
Conchapelopia intermedia Fittkau, 1962
Conchapelopia melanops (Meigen, 1818)
Conchapelopia pallidula (Meigen, 1818)
GUTTIPELOPIA Fittkau, 1962
Guttipelopia guttipennis (van der Wulp, 1861)
HAYESOMYIA Murray & Fittkau, 1986
Hayesomyia tripunctata (Goetghebuer, 1922)
KRENOPELOPIA Fittkau, 1962
Krenopelopia binotata (Wiedemann, 1817)
Krenopelopia nigropunctata (Staeger, 1839)
LABRUNDINIA Fittkau, 1962
Labrundinia longipalpis (Goetghebuer, 1921)
LARSIA Fittkau, 1962
Larsia atrocincta (Goetghebuer, 1942)
MONOPELOPIA Fittkau, 1962
Monopelopia tenuicalcar (Kieer, 1918)
NILOTANYPUS Kieer, 1923
Nilotanypus dubius (Meigen, 1804)
PARAMERINA Fittkau, 1962
Paramerina cingulata (Walker, 1856)
Paramerina divisa (Walker, 1856)
PENTANEURELLA Fittkau & Murray, 1983
Pentaneurella katterjokki Fittkau & Murray, 1983
RHEOPELOPIA Fittkau, 1962
Rheopelopia maculipennis (Zetterstedt, 1838)
Rheopelopia ornata (Meigen, 1838)
TELMATOPELOPIA Fittkau, 1962
Telmatopelopia nemorum (Goetghebuer, 1921)
TELOPELOPIA Roback, 1971
Telopelopia fascigera (Verneaux, 1970)
THIENEMANNIMYIA Fittkau, 1957
ienemannimyia carnea (Fabricius, 1805)
ienemannimyia fusciceps (Edwards, 1929)
ienemannimyia laeta (Meigen, 1818)
ienemannimyia lentiginosa (Fries, 1823)
ienemannimyia pseudocarnea Murray, 1976
ienemannimyia vitellina (Kieer, 1916)
TRISSOPELOPIA Kieer, 1923
Trissopelopia longimana (Staeger, 1839)
XENOPELOPIA Fittkau, 1962
Checklist of the family Chironomidae (Diptera) of Finland 67
Xenopelopia falcigera (Kieer, 1911)
Xenopelopia nigricans (Goetghebuer, 1927)
ZAVRELIMYIA Fittkau, 1962
Zavrelimyia barbatipes (Kieer, 1911)
Zavrelimyia hirtimana (Kieer, 1918)
Zavrelimyia melanura (Meigen, 1804)
tribe Procladiini Roback, 1971
PROCLADIUS Skuse, 1889
sg. Holotanypus Roback, 1982
Procladius appropinquatus (Lundström, 1917)
Procladius choreus (Meigen, 1804)
Procladius culiciformis (Linnaeus, 1767)
= crassinervis (Zetterstedt, 1838)
Procladius mbriatus Wülker, 1959
Procladius fuscus Brundin, 1949
? Procladius islandicus (Goetghebuer, 1931)
Procladius nudipennis Brundin, 1947
Procladius pectinatus (Kieer, 1909)
Procladius signatus (Zetterstedt, 1850)
Procladius simplicistylus Freeman, 1948
Procladius suecicus Brundin, 1949
Procladius cf. vesus Roback, 1971
Procladius sp. 1 “Valassaaret”
Procladius sp. 2 “Palsa”
Procladius sp. 3 “Kaldoaivi”
sg. Psilotanypus Kieer, 1906
Procladius avifrons Edwards, 1929
Procladius imicola Kieer, 1922
Procladius lugens Kieer, 1915
Procladius rufovittatus (van der Wulp, 1874)
tribe Tanypodini Skuse, 1889
TANYPUS Meigen, 1803
Tanypus kraatzi (Kieer, 1912)
Tanypus punctipennis Meigen, 1818
Tanypus vilipennis (Kieer, 1918)
DIAMESINAE Kieer, 1922
tribe Diamesini Kieer, 1922
DIAMESA Meigen, 1835
Diamesa aberrata Lundbeck, 1898
Diamesa arctica (Boheman, 1865)
Diamesa bertrami Edwards, 1935
Diamesa bohemani Goetghebuer, 1932
Diamesa hyperborea Holmgren, 1869
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
68
Diamesa incallida (Walker, 1856)
Diamesa insignipes Kieer, 1908
Diamesa latitarsis (Goetghebuer, 1921)
Diamesa permacra (Walker, 1856)
Diamesa serratosioi Willassen, 1986
Diamesa tonsa (Haliday, 1856)
= thienemanni Kieer, 1909
POTTHASTIA Kieer, 1922
Potthastia gaedii (Meigen, 1838)
Potthastia longimanus Kieer, 1922
Potthastia pastoris (Edwards, 1933)
PSEUDODIAMESA Goetghebuer, 1939
Pseudodiamesa branickii (Nowicki, 1873)
Pseudodiamesa nivosa (Goetghebuer, 1928)
PSEUDOKIEFFERIELLA Zavřel, 1941
= Diplomesa Zavřel 1941
Pseudokieeriella parva (Edwards, 1932)
SYMPOTTHASTIA Pagast, 1947
Sympotthastia fulva (Johannsen, 1921)
Sympotthastia huldeni Tuiskunen, 1986
SYNDIAMESA Kieer, 1918
Syndiamesa sp., indet. larva
tribe Protanypini Brundin, 1956
PROTANYPUS Kieer, 1906
Protanypus caudatus Edwards, 1924
Protanypus morio (Zetterstedt, 1838)
TELMATOGETONINAE Wirth, 1949
TELMATOGETON Schiner, 1866
Telmatogeton japonicus Tokunaga, 1933
PRODIAMESINAE Sæther, 1976
MONODIAMESA Kieer, 1922
Monodiamesa bathyphila (Kieer, 1918)
Monodiamesa ekmani (Brundin, 1949)
ODONTOMESA Pagast, 1947
Odontomesa fulva (Kieer, 1919)
PRODIAMESA Kieer, 1906
Prodiamesa olivacea (Meigen, 1818)
ORTHOCLADIINAE Kieer, 1911
AAGAARDIA Sæther, 2000
Aagaardia protensa Sæther, 2000
Aagaardia sivertseni (Aagaard, 1979)
Aagaardia(?) sp. 1
ABISKOMYIA Edwards, 1937
Checklist of the family Chironomidae (Diptera) of Finland 69
Abiskomyia paravirgo Goetghebuer, 1940
Abiskomyia virgo Edwards, 1937
ACAMPTOCLADIUS Brundin, 1956
Acamptocladius reissi Cranston & Sæther, 1982
Acamptocladius submontanus (Edwards, 1932)
ACRICOTOPUS Kieer, 1921
Acricotopus lucens (Zetterstedt, 1850)
ALLOCLADIUS Kieer, 1913
Allocladius bothnicus (Tuiskunen, 1984)
BOREOSMITTIA Tuiskunen, 1986
Boreosmittia inariensis Tuiskunen, 1986
Boreosmittia karelioborealis Tuiskunen, 1986
BRILLIA Kieer, 1913
Brillia bida (Kieer, 1909)
= modesta (Meigen, 1830) preocc.
Brillia longifurca Kieer, 1921
BRYOPHAENOCLADIUS ienemann, 1934
Bryophaenocladius aestivus (Brundin, 1947)
Bryophaenocladius dentatus (Karl, 1937)
Bryophaenocladius exidens (Brundin, 1947)
Bryophaenocladius ictericus (Meigen, 1830)
Bryophaenocladius illimbatus (Edwards, 1929)
Bryophaenocladius cf. impectinus Sæther, 1976
Bryophaenocladius inconstans (Brundin, 1947)
Bryophaenocladius cf. laticaudus Sæther, 1973
Bryophaenocladius muscicola (Kieer, 1906)
Bryophaenocladius nidorum (Edwards, 1929)
Bryophaenocladius nigrus Albu, 1974
Bryophaenocladius nitidicollis (Goetghebuer, 1913)
Bryophaenocladius pectinatus Albu, 1974
Bryophaenocladius psilacrus Sæther, 1982
Bryophaenocladius saanae Tuiskunen, 1986
Bryophaenocladius scanicus (Brundin, 1947)
Bryophaenocladius cf. sclerus Wang, Liu & Epler, 2004
Bryophaenocladius subparallelus (Malloch, 1915)
Bryophaenocladius subvernalis (Edwards, 1929)
Bryophaenocladius tuberculatus (Edwards, 1929)
Bryophaenocladius vernalis (Goetghebuer, 1921)
Bryophaenocladius xanthogyne (Edwards, 1929)
Bryophaenocladius sp., pr. xanthogyne
Bryophaenocladius sp. 1 ”Syöte”
Bryophaenocladius sp. 2 ”Syöte”
Bryophaenocladius sp. 3 ”Malla”
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
70
Bryophaenocladius sp. 4 ”Tsarmi”
Bryophaenocladius sp. 5 ”Pallas”
Bryophaenocladius sp. 6 ”Rommas”
CAMPTOCLADIUS van der Wulp, 1874
Camptocladius stercorarius (De Geer, 1776)
CARDIOCLADIUS Kieer, 1912
Cardiocladius capucinus (Zetterstedt, 1850)
Cardiocladius fuscus Kieer, 1924
CHAETOCLADIUS Kieer, 1911
Chaetocladius acuminatus Brundin, 1956
Chaetocladius binotatus (Lundström, 1915)
Chaetocladius britae Säwedal, 1976
Chaetocladius crassisaetosus Tuiskunen, 1986
Chaetocladius dentiforceps (Edwards, 1929)
Chaetocladius dissipatus (Edwards, 1929)
Chaetocladius glacialis (Lundström, 1915)
Chaetocladius gracilis Brundin, 1956
Chaetocladius grandilobus Brundin, 1956
Chaetocladius laminatus Brundin, 1947
Chaetocladius maeaeri Brundin, 1947
Chaetocladius sp., pr. melaleucus (Meigen, 1818)
Chaetocladius muliebris Tuiskunen, 1986
Chaetocladius perennis (Meigen, 1830)
Chaetocladius piger (Goetghebuer, 1913)
Chaetocladius cf. rusticus (Goetghebuer, 1932)
Chaetocladius suecicus (Kieer, 1916)
Chaetocladius tenuistylus Brundin, 1947
CLUNIO Haliday, 1855
Clunio balticus Heimbach, 1978
CORYNONEURA Winnertz, 1846
Corynoneura arctica Kieer, 1923
Corynoneura brundini Hirvenoja & Hirvenoja, 1988
Corynoneura carriana Edwards, 1924
Corynoneura celeripes Winnertz, 1852
Corynoneura celtica Edwards, 1924
Corynoneura coronata Edwards, 1924
Corynoneura edwardsi Brundin, 1949
Corynoneura ttkaui Schlee, 1968
Corynoneura gratias Schlee, 1968
Corynoneura gynocera Tuiskunen, 1983
Corynoneura lacustris Edwards, 1924
Corynoneura lobata Edwards, 1924
Corynoneura magna Brundin, 1949
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Corynoneura scutellata Winnertz, 1846
= longipennis auct. nec Tokunaga, 1936
Corynoneura sp. 1 “Tvärminne”
Corynoneura sp. 2
CORYNONEURELLA Brundin, 1949
Corynoneurella paludosa Brundin, 1949
CRICOTOPUS van der Wulp, 1874
sg. Cricotopus van der Wulp, 1874
Cricotopus albiforceps (Kieer, 1916)
Cricotopus annulator Goetghebuer, 1927
Cricotopus bicinctus (Meigen, 1818)
Cricotopus caducus Hirvenoja, 1973
Cricotopus coronatus Hirvenoja, 1973
Cricotopus cumulatus Hirvenoja, 1973
Cricotopus cylindraceus (Kieer, 1908)
Cricotopus ephippium (Zetterstedt, 1838)
Cricotopus festivellus (Kieer, 1906)
Cricotopus avocinctus (Kieer, 1924)
Cricotopus fuscus (Kieer, 1909)
Cricotopus magus Hirvenoja, 1973
Cricotopus pallidipes Edwards, 1929
Cricotopus patens Hirvenoja, 1973
Cricotopus pilidorsum Hirvenoja, 1973
Cricotopus pilosellus Brundin, 1956
Cricotopus pirifer Hirvenoja, 1973
Cricotopus polaris Kieer, 1926
Cricotopus sp., pr. polaris
Cricotopus pulchripes Verrall, 1912
Cricotopus septentrionalis Hirvenoja, 1973
Cricotopus similis Goetghebuer, 1921
Cricotopus slossonae Malloch, 1915
= claripes Hirvenoja, 1973
Cricotopus tibialis (Meigen, 1804)
Cricotopus tremulus (Linnaeus, 1758)
Cricotopus triannulatus (Macquart, 1826)
Cricotopus trifascia Edwards, 1929
Cricotopus tristis Hirvenoja, 1973
Cricotopus vierriensis Goetghebuer, 1935
Cricotopus villosus Hirvenoja, 1973
sg. Isocladius Kieer, 1909
Cricotopus arcuatus Hirvenoja, 1973
Cricotopus brevipalpis Kieer, 1909
Cricotopus intersectus (Staeger, 1839)
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Cricotopus laetus Hirvenoja, 1973
Cricotopus laricomalis Edwards, 1932
Cricotopus maurii Spies & Sæther, 2004
= polychaetus Hirvenoja, 1989 preocc.
Cricotopus obnixus (Walker, 1856)
Cricotopus sp. cf. obnixus Hirvenoja, 1973
Cricotopus obtusus Hirvenoja, 1973
Cricotopus ornatus (Meigen, 1818)
Cricotopus perniger (Zetterstedt, 1850)
Cricotopus pilicauda Hirvenoja, 1973
Cricotopus pilitarsis (Zetterstedt, 1850)
Cricotopus reductus Hirvenoja, 1973
Cricotopus relucens Hirvenoja, 1973
Cricotopus reversus Hirvenoja, 1973
Cricotopus suspiciosus Hirvenoja, 1973
Cricotopus sylvestris (Fabricius, 1794)
Cricotopus tricinctus (Meigen, 1818)
Cricotopus trifasciatus (Meigen, 1810)
sg. Nostococladius Ashe & Murray, 1980
Cricotopus lygropis Edwards, 1929
sg. ?
Cricotopus(?) sp. 1 “Tapionaho”
DIPLOCLADIUS Kieer, 1908
Diplocladius cultriger Kieer, 1908
DONCRICOTOPUS Sæther, 1981
Doncricotopus dentatus Tuiskunen, 1985
DRATNALIA Sæther & Halvorsen, 1981
Dratnalia potamophylaxi (Fittkau & Lellak, 1971)
EPOICOCLADIUS Sulc & Zavřel, 1924
Epoicocladius ephemerae (Kieer, 1924)
= avens auct. nec (Malloch, 1915)
EUKIEFFERIELLA ienemann, 1926
Eukieeriella boevrensis Brundin, 1956
Eukieeriella brevicalcar (Kieer, 1911)
Eukieeriella claripennis (Lundbeck, 1898)
Eukieeriella clypeata (ienemann, 1919)
Eukieeriella devonica (Edwards, 1929)
Eukieeriella dittmari Lehmann, 1972
Eukieeriella gracei (Edwards, 1929)
Eukieeriella sp., pr. gracei
Eukieeriella ilkleyensis (Edwards, 1929)
Eukieeriella minor (Edwards, 1929)
EURYCNEMUS van der Wulp, 1874
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Eurycnemus crassipes (Meigen, 1810)
GEORTHOCLADIUS Strenzke, 1941
Georthocladius luteicornis (Goetghebuer, 1941)
Georthocladius platystylus Sæther & Sublette, 1983
?= Parachaetocladius retezati Albu, 1972
GYMNOMETRIOCNEMUS Edwards, 1932
sg. Gymnometriocnemus Edwards, 1932
Gymnometriocnemus subnudus (Edwards, 1929)
sg. Rhaphidocladius Sæther, 1983
Gymnometriocnemus brumalis (Edwards, 1929)
Gymnometriocnemus volitans (Goetghebuer, 1940)
HALOCLADIUS Hirvenoja, 1973
Halocladius variabilis (Staeger, 1839)
HELENIELLA Gowin, 1943
Heleniella ornaticollis (Edwards, 1929)
Heleniella serratosioi Ringe, 1976
HETEROTANYTARSUS Spärck, 1923
Heterotanytarsus apicalis (Kieer, 1921)
Heterotanytarsus brundini Fittkau, 1956
Heterotanytarsus sp. 1
HETEROTRISSOCLADIUS Spärck, 1923
Heterotrissocladius brundini Sæther & Schnell, 1988
Heterotrissocladius grimshawi (Edwards, 1929)
Heterotrissocladius maeaeri Brundin, 1949
Heterotrissocladius marcidus (Walker, 1856)
Heterotrissocladius subpilosus (Kieer, 1911)
Heterotrissocladius sp. 1 ”Kolmperä”
HYDROBAENUS Fries, 1830
Hydrobaenus conformis (Holmgren, 1869)
Hydrobaenus fusistylus (Goetghebuer, 1933)
Hydrobaenus lapponicus (Brundin, 1956)
Hydrobaenus martini Sæther, 1976
Hydrobaenus pilipes (Malloch, 1915)
Hydrobaenus spinnatis Sæther, 1976
Hydrobaenus(?) sp. 1
HYDROSMITTIA Ferrington & Sæther, 2011
Hydrosmittia oxoniana (Edwards, 1922)
= recta (Edwards, 1929)
Hydrosmittia ruttneri (Strenzke & ienemann, 1942)
= brevitarsis (Brundin, 1947)
KRENOSMITTIA ienemann & Krüger, 1939
Krenosmittia boreoalpina (Goetghebuer, 1944)
Krenosmittia camptophleps (Edwards, 1929)
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Krenosmittia halvorseni (Cranston & Sæther, 1986)
LAPPOKIEFFERIELLA Tuiskunen, 1986
Lappokieeriella platytarsus Tuiskunen, 1986
LIMNOPHYES Eaton, 1875
Limnophyes aagaardi Sæther, 1990
Limnophyes angelicae Sæther, 1990
Limnophyes asquamatus Soegaard Andersen, 1937
= smolandicus Brundin, 1947
Limnophyes bidumus Sæther, 1990
Limnophyes sp., pr. bidumus
Limnophyes brachytomus (Kieer, 1922)
Limnophyes dicilis Brundin, 1947
Limnophyes edwardsi Sæther, 1990
Limnophyes er Sæther, 1985
Limnophyes habilis (Walker, 1856)
= truncorum Goetghebuer, 1921
Limnophyes margaretae Sæther, 1975
Limnophyes minimus (Meigen, 1818)
= exiguus (Goetghebuer, 1913)
Limnophyes natalensis (Kieer, 1914)
Limnophyes ninae Sæther, 1975
Limnophyes pentaplastus (Kieer, 1921)
Limnophyes pumilio (Holmgren, 1869)
= globifer Lundström, 1915
Limnophyes sp., pr. pumilio
Limnophyes schnelli Sæther, 1990
Limnophyes spinigus Sæther, 1990
Limnophyes torulus Sæther, 1990
MESOCRICOTOPUS Brundin, 1956
Mesocricotopus thienemanni (Goetghebuer, 1940)
MESOSMITTIA Brundin, 1956
Mesosmittia exuella (Edwards, 1929)
METRIOCNEMUS van der Wulp, 1874
Metriocnemus acutus Sæther, 1995
Metriocnemus albolineatus (Meigen, 1818)
= atratulus (Zetterstedt, 1850)
Metriocnemus atriclava Kieer, 1921
Metriocnemus beringensis (Cranston & Oliver, 1988)
Metriocnemus caudigus Sæther, 1995
Metriocnemus corticalis Strenzke, 1950
Metriocnemus eurynotus (Holmgren, 1883)
= hygropetricus Kieer, 1912
Metriocnemus exilacies Sæther, 1995
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Metriocnemus fuscipes (Meigen, 1818)
Metriocnemus intergerivus Sæther, 1995
Metriocnemus picipes (Meigen, 1818)
Metriocnemus ursinus (Holmgren, 1869)
NANOCLADIUS Kieer, 1913
= Microcricotopus ienemann & Harnisch, 1932
Nanocladius balticus (Palmén, 1959)
Nanocladius dichromus (Kieer, 1906)
= bicolor (Zetterstedt, 1838) nom. nudum
Nanocladius parvulus (Kieer, 1909)
Nanocladius rectinervis (Kieer, 1911)
ORTHOCLADIUS van der Wulp, 1874
sg. Eudactylocladius ienemann, 1935
Orthocladius fuscimanus (Kieer, 1908)
Orthocladius gelidorum (Kieer, 1923)
Orthocladius gelidus Kieer, 1922
Orthocladius musester Sæther, 2004
Orthocladius olivaceus (Kieer, 1911)
= mixtus auct., partim
Orthocladius priomixtus Sæther, 2004
= mixtus auct., partim
sg. Euorthocladius ienemann, 1935
Orthocladius abiskoensis ienemann & Krüger, 1937
Orthocladius cf. annellae Sæther, 2005
Orthocladius ashei Soponis, 1990
Orthocladius rivicola Kieer, 1911
Orthocladius rivulorum Kieer, 1909
Orthocladius saxosus (Tokunaga, 1939)
Orthocladius telochaetus Langton, 1985
sg. Mesorthocladius Sæther, 2005
Orthocladius frigidus (Zetterstedt, 1838)
sg. Orthocladius van der Wulp, 1874
Orthocladius decoratus (Holmgren, 1869)
Orthocladius dentifer Brundin, 1947
Orthocladius excavatus Brundin, 1947
Orthocladius lapponicus Goetghebuer, 1940
Orthocladius nitidoscutellatus Lundström, 1915
Orthocladius oblidens (Walker, 1856)
Orthocladius pedestris Kieer, 1909
Orthocladius rhyacobius Kieer, 1911
Orthocladius rubicundus (Meigen, 1818)
= saxicola Kieer, 1911
Orthocladius wetterensis Brundin, 1956
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sg. Pogonocladius Brundin, 1956
Orthocladius consobrinus (Holmgren, 1869)
sg. Symposiocladius Cranston, 1982
Orthocladius holsatus Goetghebuer, 1937
Orthocladius lignicola Kieer, 1914
Orthocladius ruoi Rossaro & Prato, 1991
Orthocladius schnelli Sæther, 2004
Orthocladius smolandicus Brundin, 1947
PARACHAETOCLADIUS Wülker, 1959
Parachaetocladius abnobaeus (Wülker, 1959)
PARACLADIUS Hirvenoja, 1973
Paracladius alpicola (Zetterstedt, 1850)
Paracladius conversus (Walker, 1856)
Paracladius quadrinodosus Hirvenoja, 1973
PARACRICOTOPUS Brundin, 1956
Paracricotopus niger (Kieer, 1913)
Paracricotopus uliginosus (Brundin, 1947)
PARAKIEFFERIELLA ienemann, 1936
Parakieeriella bathophila (Kieer, 1912)
Parakieeriella bilobata Tuiskunen, 1986
Parakieeriella coronata (Edwards, 1929)
Parakieeriella fennica Tuiskunen, 1986
Parakieeriella nnmarkica Tuiskunen, 1986
Parakieeriella gynocera (Edwards, 1937)
Parakieeriella minuta Tuiskunen, 1986
Parakieeriella nigra Brundin, 1949
Parakieeriella scandica Brundin, 1956
Parakieeriella smolandica (Brundin, 1947)
Parakieeriella subaterrima (Malloch, 1915)
= torulata Sæther, 1969
PARALIMNOPHYES Brundin, 1956
Paralimnophyes longiseta (ienemann, 1919)
= hydrophilus (Goetghebuer, 1921)
PARAMETRIOCNEMUS Goetghebuer, 1932
Parametriocnemus boreoalpinus Gowin & ienemann, 1942
Parametriocnemus stylatus (Spärck, 1923)
Parametriocnemus sp. 1
PARAPHAENOCLADIUS ienemann, 1924
Paraphaenocladius exagitans (Johannsen, 1905)
Paraphaenocladius sp., pr. exagitans
Paraphaenocladius impensus (Walker, 1856)
Paraphaenocladius intercedens Brundin, 1947
Paraphaenocladius sp., pr. intercedens
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Paraphaenocladius irritus (Walker, 1856)
Paraphaenocladius pseudirritus Strenzke, 1950
Paraphaenocladius triangulus Sæther & Wang, 1995
PARASMITTIA Strenzke, 1950
Parasmittia carinata Strenzke, 1950
Parasmittia sp. 1 ”Luukki”
PARATRICHOCLADIUS Santos Abreu, 1918
Paratrichocladius ruventris (Meigen, 1830)
Paratrichocladius skirwithensis (Edwards, 1929)
PARATRISSOCLADIUS Zavřel, 1937
Paratrissocladius excerptus (Walker, 1856)
PROPSILOCERUS Kieer, 1923
= Synpsilocerus Kieer, 1923
= Tokunagayusurika Sasa, 1978
Propsilocerus jacuticus (Zvereva, 1950)
Propsilocerus komensis (Zvereva, 1950)
?= paradoxus (Lundström, 1915)
Propsilocerus saetheri Wang, Liu & Paasivirta, 2007
PROSMITTIA Brundin, 1956
Prosmittia jemtlandica (Brundin, 1947)
Prosmittia rectangularis Tuiskunen, 1985
PSECTROCLADIUS Kieer, 1906
sg. Allopsectrocladius Wülker, 1956
Psectrocladius conjungens (Brundin, 1947)
Psectrocladius obvius (Walker, 1856)
Psectrocladius platypus (Edwards, 1929)
sg. Mesopsectrocladius Laville, 1971
Psectrocladius barbatipes Kieer, 1923
sg. Monopsectrocladius Wülker, 1956
Psectrocladius calcaratus (Edwards, 1929)
sg. Psectrocladius Kieer, 1906
Psectrocladius barbimanus (Edwards, 1929)
Psectrocladius bisetus Goetghebuer, 1942
Psectrocladius fennicus Storå, 1939
Psectrocladius limbatellus (Holmgren, 1869)
= edwardsi Brundin, 1949
Psectrocladius octomaculatus Wülker, 1956
Psectrocladius oligosetus Wülker, 1956
Psectrocladius oxyura Langton, 1985
Psectrocladius psilopterus (Kieer, 1906)
Psectrocladius schlienzi Wülker, 1956
Psectrocladius sordidellus (Zetterstedt, 1838)
Psectrocladius ventricosus Kieer, 1925
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Psectrocladius zetterstedti Brundin, 1949
PSEUDORTHOCLADIUS Goetghebuer, 1943
Pseudorthocladius curtistylus (Goetghebuer, 1921)
Pseudorthocladius liformis (Kieer, 1908)
Pseudorthocladius pilosipennis Brundin, 1956
PSEUDOSMITTIA Edwards, 1932
Pseudosmittia albipennis (Goetghebuer, 1921)
Pseudosmittia angusta (Edwards, 1929)
Pseudosmittia danconai (Marcuzzi, 1947)
Pseudosmittia forcipata (Goetghebuer, 1921)
Pseudosmittia sp., pr. forcipata
Pseudosmittia gracilis (Goetghebuer, 1913)
Pseudosmittia mathildae Albu, 1968
Pseudosmittia obtusa Strenzke, 1960
Pseudosmittia trilobata (Edwards, 1929)
PSILOMETRIOCNEMUS Sæther, 1969
Psilometriocnemus europaeus Tuiskunen, 1985
RHEOCRICOTOPUS Brundin, 1956
sg. Psilocricotopus Sæther, 1986
Rheocricotopus atripes (Kieer, 1913)
= foveatus (Edwards, 1929)
Rheocricotopus chalybeatus (Edwards, 1929)
Rheocricotopus chapmani (Edwards, 1935)
Rheocricotopus glabricollis (Meigen, 1830)
sg. Rheocricotopus Brundin, 1956
Rheocricotopus eusus (Walker, 1856)
Rheocricotopus fuscipes (Kieer, 1909)
= dispar (Goetghebuer, 1913)
Rheocricotopus reduncus Sæther & Schnell, 1988
Rheocricotopus sp., pr. unidentatus Sæther & Schnell, 1988
RHEOSMITTIA Brundin, 1986
Rheosmittia languida (Brundin, 1956)
Rheosmittia spinicornis (Brundin, 1956)
SMITTIA Holmgren, 1869
Smittia alpilonga Rossaro & Lencioni, 2000
Smittia cf. amoena Caspers, 1988
Smittia aterrima (Meigen, 1818)
Smittia sp., pr. aterrima
Smittia betuletorum Edwards, 1941
Smittia contingens (Walker, 1856)
Smittia edwardsi Goetghebuer, 1932
Smittia foliacea (Kieer, 1921)
Smittia leucopogon (Meigen, 1804)
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Smittia nudipennis (Goetghebuer, 1913)
Smittia paranudipennis Brundin, 1947
Smittia pratorum (Goetghebuer, 1927)
Smittia scutellosetosa Caspers, 1988
Smittia stercoraria Rossaro & Lencioni, 2000
Smittia sp. 1 s. Tuiskunen
Smittia sp., pr. sp. 1
Smittia sp. 2 ”Jehkats”
STACKELBERGINA Shilova & Zelentsov, 1978
Stackelbergina praeclara Shilova & Zelentsov, 1978
STILOCLADIUS Rossaro, 1979
Stilocladius intermedius Wang, 1998
SYNORTHOCLADIUS ienemann, 1935
Synorthocladius semivirens (Kieer, 1909)
TAVASTIA Tuiskunen, 1985
Tavastia australis Tuiskunen, 1985
Tavastia yggdrasilia Brodin, Lundström & Paasivirta, 2008
THIENEMANNIA Kieer, 1909
ienemannia gracilis Kieer, 1909
ienemannia paasivirtai Tuiskunen, 1986
THIENEMANNIELLA Kieer, 1911
ienemanniella acuticornis (Kieer, 1912)
ienemanniella clavicornis (Kieer, 1911)
ienemanniella majuscula (Edwards, 1924)
ienemanniella minuscula (Brundin, 1949)
ienemanniella obscura Brundin, 1947
ienemanniella vittata (Edwards, 1924)
ienemanniella sp., pr. vittata
TOKUNAGAIA Sæther, 1973
Tokunagaia excellens (Brundin, 1956)
Tokunagaia parexcellens Tuiskunen, 1986
Tokunagaia rectangularis (Goetghebuer, 1940)
Tokunagaia scutellata (Brundin, 1956)
Tokunagaia tonollii (Rossaro, 1983)
TRISSOCLADIUS Kieer, 1908
Trissocladius brevipalpis Kieer, 1908
TVETENIA Kieer, 1922
Tvetenia bavarica (Goetghebuer, 1934)
Tvetenia calvescens (Edwards, 1929)
Tvetenia discoloripes (Goetghebuer & ienemann, 1936)
Tvetenia duodenaria Kieer, 1922
= saanensis (Wülker, 1959)
Tvetenia verralli (Edwards, 1929)
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?= tshernovskii (Pankratova, 1968 )
VIVACRICOTOPUS Schnell & Sæther, 1988
Vivacricotopus ablusus Schnell & Sæther, 1988
ZALUTSCHIA Lipina, 1939
Zalutschia cf. humphriesiae Dowling & Murray, 1980
Zalutschia mallae Tuiskunen, 1986
Zalutschia mucronata (Brundin, 1949)
Zalutschia tatrica (Pagast, 1935)
Zalutschia tornetraeskensis (Edwards & ienemann, 1941)
Zalutschia zalutschicola Lipina, 1939
genus unknown
Orthocladiinae sp. 1 ”Outajärvi”
Orthocladiinae sp. 2 ”Pallas”
Orthocladiinae sp. 3 ”Kaldoaivi”
CHIRONOMINAE Newman, 1834
tribe Chironomini Newman, 1834
BENTHALIA Lipina, 1939
Benthalia carbonaria (Meigen, 1804)
= dissidens (Walker, 1856)
CHIRONOMUS Meigen, 1803
sg. Chaetolabis Townes, 1945
Chironomus macani Freeman, 1948
sg. Chironomus Meigen, 1803
Chironomus acerbus Hirvenoja, 1962
Chironomus acutiventris Wülker, Ryser & Scholl, 1983
Chironomus agilis Shobanov & Dyomin, 1988
Chironomus annularius Meigen, 1818
Chironomus anthracinus Zetterstedt, 1860
Chironomus aprilinus Meigen, 1818
Chironomus beljaninae Wülker, 1991
Chironomus borokensis Kerkis, Filippova, Shobanov, Gunderina & Kiknadze, 1988
Chironomus brevidentatus Hirvenoja & Michailova, 1998
Chironomus cingulatus Meigen, 1830
Chironomus clarus Hirvenoja, 1962
Chironomus coaetaneus Hirvenoja, 1998
Chironomus dorsalis Strenzke, 1959
Chironomus entis Shobanov, 1989
Chironomus esai Wülker, 1997
Chironomus fraternus Wülker, 1991
Chironomus heteropilicornis Wülker, 1996
? Chironomus hyperboreus Staeger, 1845
Chironomus inermifrons Goetghebuer, 1921
Chironomus islandicus (Kieer, 1913)
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Chironomus jonmartini Lindeberg, 1979) coll.
= neglectus Lindeberg, 1960 preocc.
Chironomus longistylus Goetghebuer, 1921
Chironomus lugubris Zetterstedt, 1850
Chironomus luridus Strenzke, 1959
Chironomus melanescens Keyl, 1961
Chironomus melanotus Keyl, 1961
Chironomus muratensis Ryser, Scholl & Wülker, 1983
Chironomus neocorax Wülker & Butler, 1983
Chironomus pallidivittatus Edwards, 1929
Chironomus piger Strenzke, 1956
Chironomus pilicornis (Fabricius, 1787)
Chironomus plumosus (Linnaeus, 1758)
Chironomus pseudothummi Strenzke, 1959 coll.
Chironomus riihimakiensis Wülker, 1973
Chironomus riparius Meigen, 1804
Chironomus salinarius Kieer, 1915
Chironomus saxatilis Wülker, Ryser & Scholl, 1981 coll.
Chironomus sollicitus Hirvenoja, 1962
Chironomus sororius Wülker, 1973
? Chironomus staegeri Lundbeck, 1898
Chironomus tentans Fabricius, 1805
Chironomus tenuistylus Brundin, 1949
sg. Lobochironomus Ryser, Wülker & Scholl, 1985
Chironomus dorsalis Meigen, 1818
= longipes Staeger, 1839 as Einfeldia
Chironomus improvidus Hirvenoja, 1998
Chironomus mendax Storå, 1936
Chironomus storai Goetghebuer, 1954
= luctuosus Storå, 1936 preocc.
CLADOPELMA Kieer, 1921
Cladopelma bicarinatum (Brundin, 1947)
Cladopelma edwardsi (Kruseman, 1933)
Cladopelma goetghebueri Spies & Sæther, 2004
= lateralis (Goetghebuer, 1934) preocc.
Cladopelma virescens (Meigen, 1818)
Cladopelma viridulum (Linnaeus, 1767)
CRYPTOCHIRONOMUS Kieer, 1918
Cryptochironomus albofasciatus (Staeger, 1839)
Cryptochironomus denticulatus (Goetghebuer, 1921)
Cryptochironomus obreptans (Walker, 1856)
Cryptochironomus psittacinus (Meigen, 1830)
Cryptochironomus redekei (Kruseman, 1933)
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Cryptochironomus rostratus Kieer, 1921
Cryptochironomus supplicans (Meigen, 1830)
Cryptochironomus ussouriensis (Goetghebuer, 1933)
= nigridens Chernovskij, 1949
CRYPTOTENDIPES Beck & Beck, 1969
Cryptotendipes darbyi (Sublette, 1960)
Cryptotendipes pugfelderi Reiss, 1964
Cryptotendipes pseudotener (Goetghebuer, 1922)
Cryptotendipes usmaensis (Pagast, 1931)
DEMEIJEREA Kruseman, 1933
Demeijerea rupes (Linnaeus, 1761)
DEMICRYPTOCHIRONOMUS Lenz, 1941
sg. Demicryptochironomus Lenz, 1941
Demicryptochironomus vulneratus (Zetterstedt, 1838)
Demicryptochironomus sp. Pe 1 Langton, 1991
sg. Irmakia Reiss, 1988
Demicryptochironomus neglectus Reiss, 1988
DICROTENDIPES Kieer, 1913
= Limnochironomus Kieer, 1920
Dicrotendipes lobiger (Kieer, 1921)
Dicrotendipes nervosus (Staeger, 1839)
Dicrotendipes notatus (Meigen, 1818)
Dicrotendipes pulsus (Walker, 1856)
= objectans (Walker, 1856)
Dicrotendipes tritomus (Kieer, 1916)
EINFELDIA Kieer, 1924
Einfeldia pagana (Meigen, 1838)
Einfeldia pectoralis Kieer, 1924
ENDOCHIRONOMUS Kieer, 1918
Endochironomus albipennis (Meigen, 1830)
Endochironomus stackelbergi Goetghebuer, 1935
Endochironomus tendens (Fabricius, 1775)
GLYPTOTENDIPES Kieer, 1913
sg. Caulochironomus Heyn, 1993
Glyptotendipes aequalis (Kieer, 1922)
Glyptotendipes caulicola (Kieer, 1913)
Glyptotendipes imbecilis (Walker, 1856)
= severini (Goetghebuer, 1923)
Glyptotendipes scirpi (Kieer, 1915)
= fodiens (Kieer, 1924)
= mancunianus (Edwards, 1929 )
Glyptotendipes viridis (Macquart, 1834)
sg. Glyptotendipes Kieer, 1913
Checklist of the family Chironomidae (Diptera) of Finland 83
Glyptotendipes barbipes (Staeger, 1839)
Glyptotendipes cauliginellus (Kieer, 1913)
= gripekoveni (Kieer, 1913)
Glyptotendipes glaucus (Meigen, 1818)
Glyptotendipes pallens (Meigen, 1804)
Glyptotendipes paripes (Edwards, 1929)
sg. Heynotendipes Spies & Sæther, 2004
Glyptotendipes signatus (Kieer, 1909)
HARNISCHIA Kieer, 1921
Harnischia curtilamellata (Malloch, 1915)
Harnischia fuscimana Kieer, 1921
KIEFFERULUS Goetghebuer, 1922
Kieerulus tendipediformis (Goetghebuer, 1921)
KLOOSIA Kruseman, 1933
Kloosia pusilla (Linnaeus, 1767)
LAUTERBORNIELLA ienemann & Bause, 1913
Lauterborniella agrayloides (Kieer, 1911)
LIPINIELLA Shilova, 1961
? Lipiniella araenicola Shilova, 1961 (larval ident.)
Lipiniella prima Shilova, Kerkis & Kiknadze, 1993
MICROCHIRONOMUS Kieer, 1918
Microchironomus tener (Kieer, 1918)
MICROTENDIPES Kieer, 1915
Microtendipes brevitarsis Brundin, 1947
Microtendipes chloris (Meigen, 1818)
Microtendipes connis (Meigen, 1830)
Microtendipes nigellus Hirvenoja, 1963
Microtendipes pedellus (De Geer, 1776)
Microtendipes rydalensis (Edwards, 1929)
NILOTHAUMA Kieer, 1921
Nilothauma brayi (Goetghebuer, 1921)
OMISUS Townes, 1945
Omisus caledonicus (Edwards, 1932)
PAGASTIELLA Brundin, 1949
Pagastiella orophila (Edwards, 1929)
PARACHIRONOMUS Lenz, 1921
Parachironomus biannulatus (Staeger, 1839)
Parachironomus digitalis (Edwards, 1929)
Parachironomus frequens (Johannsen, 1905)
Parachironomus gracilior (Kieer, 1918)
= arcuatus (Goetghebuer, 1919)
Parachironomus monochromus (van der Wulp, 1874)
Parachironomus paradigitalis Brundin, 1949
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
84
Parachironomus parilis (Walker, 1856)
Parachironomus cf. pseudovarus Zorina, 2003
Parachironomus siljanensis Brundin, 1949
Parachironomus subalpinus (Goetghebuer, 1932)
Parachironomus tenuicaudatus (Malloch, 1915)
Parachironomus varus (Goetghebuer, 1921)
Parachironomus vitiosus (Goetghebuer, 1921)
Parachironomus sp. 1 “Iijoki”
PARACLADOPELMA Harnisch, 1923
Paracladopelma camptolabis (Kieer, 1913)
Paracladopelma galaptera (Townes, 1945)
Paracladopelma laminatum (Kieer, 1921)
Paracladopelma nereis (Townes, 1945)
Paracladopelma nigritulum (Goetghebuer, 1942)
Paracladopelma undine (Townes, 1945)
PARALAUTERBORNIELLA Lenz, 1941
Paralauterborniella nigrohalteralis (Malloch, 1915)
PARATENDIPES Kieer, 1911
Paratendipes albimanus (Meigen, 1818)
Paratendipes subaequalis (Malloch, 1915)
Paratendipes conn. no. 3 Lipina, 1926 (larval type)
PHAENOPSECTRA Kieer, 1921
Phaenopsectra avipes (Meigen, 1818)
Phaenopsectra punctipes (Wiedemann, 1817)
POLYPEDILUM Kieer, 1912
sg. Pentapedilum Kieer, 1913
Polypedilum sordens (van der Wulp, 1874)
Polypedilum tritum (Walker, 1856)
? = uncinatum (Goetghebuer, 1921)
sg. Polypedilum Kieer, 1912
Polypedilum acutum Kieer, 1915
Polypedilum albicorne (Meigen, 1838)
Polypedilum amoenum Goetghebuer, 1930
Polypedilum arundineti (Goetghebuer, 1921)
Polypedilum laetum (Meigen, 1818)
Polypedilum nubeculosum (Meigen, 1804)
Polypedilum pedestre (Meigen, 1830)
Polypedilum trigonus Townes, 1945
sg. Tripodura Townes, 1945
Polypedilum aegyptium Kieer, 1925
Polypedilum bicrenatum Kieer, 1921
Polypedilum pullum (Zetterstedt, 1838)
Polypedilum quadriguttatum Kieer, 1921
Checklist of the family Chironomidae (Diptera) of Finland 85
Polypedilum scalaenum (Schrank, 1803)
Polypedilum tetracrenatum Hirvenoja, 1962
sg. Uresipedilum Sasa & Kikuchi, 1995
Polypedilum convictum (Walker, 1856)
Polypedilum cultellatum Goetghebuer, 1931
ROBACKIA Sæther, 1977
Robackia pilicauda Sæther, 1977
? Robackia demeijerei (Kruseman, 1933) (larval ident.)
SAETHERIA Jackson, 1977
Saetheria reissi Jackson, 1977
SERGENTIA Kieer, 1922
Sergentia baueri Wülker, Kiknadze, Kerkis & Nevers, 1999
Sergentia coracina (Zetterstedt, 1850)
Sergentia prima Proviz & Proviz, 1997
STENOCHIRONOMUS Kieer, 1919
sg. Petalopholeus Borkent, 1984
Stenochironomus fascipennis (Zetterstedt, 1838)
sg. Stenochironomus Kieer, 1919
Stenochironomus gibbus (Fabricius, 1794)
Stenochironomus hibernicus (Edwards, 1929)
STICTOCHIRONOMUS Kieer, 1919
Stictochironomus crassiforceps (Kieer, 1921)
Stictochironomus maculipennis (Meigen, 1818)
Stictochironomus pictulus (Meigen, 1830)
Stictochironomus rosenschoeldi (Zetterstedt, 1838)
Stictochironomus sticticus (Fabricius, 1781)
SYNENDOTENDIPES Grodhaus, 1987
Synendotendipes dispar (Meigen, 1830)
Synendotendipes impar (Walker, 1856)
Synendotendipes lepidus (Meigen, 1830)
TRIBELOS Townes, 1945
Tribelos cf. donatoris (Shilova, 1974 )
?= subatrum Grodhaus, 1987
Tribelos intextum (Walker, 1856)
XENOCHIRONOMUS Kieer, 1921
Xenochironomus xenolabis (Kieer, 1916)
ZAVRELIELLA Kieer, 1920
Zavreliella marmorata (van der Wulp, 1859)
tribe Pseudochironomini Sæther, 1977
PSEUDOCHIRONOMUS Malloch, 1915
Pseudochironomus prasinatus (Staeger, 1839)
tribe Tanytarsini Zavřel, 1917
CLADOTANYTARSUS Kieer, 1921
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
86
sg. Cladotanytarsus Kieer, 1921
Cladotanytarsus atridorsum Kieer, 1924
Cladotanytarsus cyrylae Giłka, 2001
Cladotanytarsus dicilis Brundin, 1947
Cladotanytarsus gedanicus Giłka, 2001
Cladotanytarsus iucundus Hirvenoja, 1962
Cladotanytarsus lepidocalcar Krüger, 1938
Cladotanytarsus mancus (Walker, 1856)
Cladotanytarsus matthei Giłka, 2001
Cladotanytarsus molestus Hirvenoja, 1962
Cladotanytarsus nigrovittatus (Goetghebuer, 1922)
Cladotanytarsus pallidus Kieer, 1922
Cladotanytarsus teres Hirvenoja, 1962
Cladotanytarsus vanderwulpi (Edwards, 1929)
sg. Lenziella Kieer, 1922
Cladotanytarsus amandus Hirvenoja, 1962
Cladotanytarsus bicornutus Kieer, 1922
= wexionensis Brundin, 1947
CONSTEMPELLINA Brundin, 1947
Constempellina brevicosta (Edwards, 1937)
CORYNOCERA Zetterstedt, 1837
Corynocera ambigua Zetterstedt, 1837
Corynocera oliveri Lindeberg, 1970
MICROPSECTRA Kieer, 1908
= Lauterbornia Kieer, 1911
= Lundstroemia Kieer, 1921
= Parapsectra Reiss, 1969
Micropsectra appendica Stur & Ekrem, 2006
Micropsectra apposita (Walker, 1856)
Micropsectra atrofasciata (Kieer, 1911)
Micropsectra attenuata Reiss, 1969
Micropsectra calcifontis Stur & Ekrem, 2006
Micropsectra chionophila (Edwards, 1933)
Micropsectra insignilobus Kieer, 1924
Micropsectra junci (Meigen, 1818)
Micropsectra klinki Stur & Ekrem, 2006
Micropsectra lacustris Säwedal, 1975
Micropsectra lindebergi Säwedal, 1976
Micropsectra logani (Johannsen, 1928)
= groenlandica Soegaard Andersen, 1937
Micropsectra malla Giłka & Paasivirta, 2008
Micropsectra nana (Meigen, 1818)
Micropsectra notescens (Walker, 1856)
Checklist of the family Chironomidae (Diptera) of Finland 87
Micropsectra pallidula (Meigen, 1830)
= bidentata auct.
Micropsectra radialis Goetghebuer, 1939
= coracina (Kieer, 1911)
Micropsectra recurvata Goetghebuer, 1928
Micropsectra rilensis Giłka, 2001
Micropsectra roseiventris (Kieer, 1909)
= fusca auct. nec (Meigen, 1804)
Micropsectra schrankelae Stur & Ekrem, 2006
Micropsectra soae Stur & Ekrem, 2006
Micropsectra styriaca Reiss, 1969
NEOZAVRELIA Goetghebuer & ienemann, 1941
Neozavrelia cuneipennis (Edwards, 1929)
PARATANYTARSUS ienemann & Bause, 1913
Paratanytarsus abiskoensis Reiss & Säwedal, 1981
Paratanytarsus austriacus (Kieer, 1924)
Paratanytarsus bituberculatus (Edwards, 1929)
Paratanytarsus brevicalcar (Kieer, 1909)
= intricatus (Goetghebuer, 1921)
Paratanytarsus dimorphis Reiss, 1965
Paratanytarsus dissimilis (Johannsen, 1905)
= confusus Palmén, 1960
Paratanytarsus grimmii (Schneider, 1885)
= boiemicus Bause, 1913
Paratanytarsus hyperboreus Brundin, 1949
Paratanytarsus inopertus (Walker, 1856)
Paratanytarsus laccophilus (Edwards, 1929)
Paratanytarsus laetipes (Zetterstedt, 1850)
Paratanytarsus lauterborni (Kieer, 1909)
Paratanytarsus natvigi (Goetghebuer, 1933)
Paratanytarsus paralaccophilus Giłka & Paasivirta, 2008
Paratanytarsus penicillatus (Goetghebuer, 1928)
Paratanytarsus tenellulus (Goetghebuer, 1921)
Paratanytarsus tenuis (Meigen, 1830)
RHEOTANYTARSUS ienemann & Bause, 1913
Rheotanytarsus curtistylus (Goetghebuer, 1921)
Rheotanytarsus muscicola ienemann, 1929
Rheotanytarsus pellucidus (Walker, 1848)
= distinctissimus (Brundin, 1947)
Rheotanytarsus pentapoda (Kieer, 1909)
Rheotanytarsus photophilus (Goetghebuer, 1921)
Rheotanytarsus ringei Lehmann, 1970
STEMPELLINA ienemann & Bause, 1913
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
88
Stempellina almi Brundin, 1947
Stempellina bausei (Kieer, 1911)
Stempellina subglabripennis (Brundin, 1947)
Stempellina tervolae Giłka, 2005
STEMPELLINELLA Brundin, 1947
Stempellinella brevis (Edwards, 1929)
Stempellinella edwardsi Spies & Sæther, 2004
= minor (Edwards, 1929) preocc.
Stempellinella avidula (Edwards, 1929)
Stempellinella saltuum (Goetghebuer, 1921)
TANYTARSUS van der Wulp, 1874
Tanytarsus aberrans Lindeberg, 1970
Tanytarsus aculeatus Brundin, 1949
Tanytarsus anderseni Reiss & Fittkau, 1971
Tanytarsus bathophilus Kieer, 1911
= tripunctatus Reiss, 1968
Tanytarsus brundini Lindeberg, 1963
Tanytarsus buchonius Reiss & Fittkau, 1971
Tanytarsus chinyensis Goetghebuer, 1934
Tanytarsus curticornis Kieer, 1911
Tanytarsus debilis (Meigen, 1830)
Tanytarsus desertor Giłka & Paasivirta, 2007
Tanytarsus dibranchius Kieer, 1926
= separabilis Brundin, 1947
Tanytarsus dispar Lindeberg, 1967
Tanytarsus eminulus (Walker, 1856)
Tanytarsus excavatus Edwards, 1929
Tanytarsus fennicus Lindeberg, 1970
Tanytarsus gibbosiceps Kieer, 1922
Tanytarsus glabrescens Edwards, 1929
Tanytarsus gracilentus (Holmgren, 1883)
Tanytarsus gregarius Kieer, 1909
Tanytarsus heusdensis Goetghebuer, 1923
Tanytarsus inaequalis Goetghebuer, 1921
Tanytarsus innarensis Brundin, 1947
Tanytarsus lactescens Edwards, 1929
Tanytarsus lapponicus Lindeberg, 1970
Tanytarsus latiforceps Edwards, 1941
Tanytarsus lestagei Goetghebuer, 1922
= decipiens Lindeberg, 1967
= palmeni Lindeberg, 1967
Tanytarsus longitarsis Kieer, 1911
Checklist of the family Chironomidae (Diptera) of Finland 89
Tanytarsus lugens (Kieer, 1916)
Tanytarsus mancospinosus Ekrem & Reiss, 1999
Tanytarsus medius Reiss & Fittkau, 1971
Tanytarsus mendax Kieer, 1925
= holochlorus Edwards, 1929
Tanytarsus miriforceps (Kieer, 1921)
Tanytarsus multipunctatus Brundin, 1947
Tanytarsus nemorosus Edwards, 1929
Tanytarsus niger Soegaard Andersen, 1937
Tanytarsus norvegicus (Kieer, 1924)
Tanytarsus occultus Brundin, 1949
Tanytarsus palettaris Verneaux, 1969
Tanytarsus pallidicornis (Walker, 1856)
Tanytarsus paraniger Giłka & Paasivirta, 2008
Tanytarsus quadridentatus Brundin, 1947
Tanytarsus recurvatus Brundin, 1947
Tanytarsus salmelai Giłka & Paasivirta, 2009
Tanytarsus signatus (van der Wulp, 1859)
Tanytarsus smolandicus Brundin, 1947
Tanytarsus striatulus Lindeberg, 1976
Tanytarsus sylvaticus (van der Wulp, 1859)
= aptus Hirvenoja, 1963
Tanytarsus telmaticus Lindeberg, 1959
= simulans Lindeberg, 1967
= socialis Lindeberg, 1967
Tanytarsus trux Giłka & Paasivirta, 2007
Tanytarsus usmaensis Pagast, 1931
Tanytarsus verralli Goetghebuer, 1928
Tanytarsus volgensis Miseiko, 1967
= mbriatus Reiss & Fittkau, 1971
VIRGATANYTARSUS Pinder, 1982
Virgatanytarsus arduennensis (Goetghebuer, 1922)
ZAVRELIA Kieer, ienemann & Bause, 1913
Zavrelia pentatoma Kieer & Bause, 1913
= atrofasciata Kieer, 1921
Excluded species
e numerous species that were reported by Hackman (1980), but are excluded from
the current check-list, are not listed here. Reasons for deletions of those species are vari-
ous, mainly due to misidentications and nomenclatural changes.
Lauri Paasivirta / ZooKeys 441: 63–90 (2014)
90
Notes
Chironomus pallidivittatus Edwards, 1929. is name is preoccupied by C. pallidi-
vittatus Malloch, 1915. e European species needs a new name.
References
Armitage PD, Cranston PS, Pinder LCV (1995) e Chironomidae: biology and ecology of
non-biting midges. Chapman and Hall, London, 572 pp. doi: 10.1007/978-94-011-0715-0
Bergman G, Jansson A (1983) Accession to the Zoological Museum, University of Helsinki/Hels-
ingfors 14.5–31.12.1982. Memoranda Societatis pro Fauna et Flora Fennica 59: 165–166.
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Paasivirta L (2000) Chironomidae (Diptera) of the northern Baltic Sea. In: Horichter O (Ed)
Late 20th century research on Chironomidae: an anthology from the 13th international
symposium on Chironomidae. Freiburg, Germany, September 5-9, 1997. Shaker Verlag,
Aachen, Germany, 589–598.
Paasivirta L (2007) Chironomid species in Finnish springs and their surroundings. In: An-
dersen T (Ed.) Contributions to the systematics and ecology of aquatic Diptera – a tribute
to Ole A. Sæther. e Caddis Press, Columbus, 235–246.
Paasivirta L (2012) Finnish Chironomidae. Chironomus 25: 47–70. http://www.ntnu.no/ojs/
index.php/chironomus/article/view/1546/1439
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Penttinen J, Ilmonen J, Jakovlev J, Salmela J, Kuusela K, Paasivirta L (2010) read-horned ies
(Diptera: Nematocera). In: Rassi P, Hyvärinen E, Juslen A, Mannerkoski I (Eds) e 2010 Red
List of Finnish Species. Ympäristöministeriö & Suomen ympäristökeskus, Helsinki, 477–489.
Sæther OA, Spies M (2013) Fauna Europaea: Chironomidae. In: Pape T, Beuk P (Eds) Fauna
europaea: Diptera, version 2.6. http://www.faunaeur.org/
Tuiskunen J, Lindeberg B (1986) Chironomidae (Diptera) from Fennoscandia north of 68 deg.
N, with a description of ten new species and two new genera. Annales zoologici fennici
23(4): 361–393. http://www.sekj.org/PDF/anzf23/anz23-361-393.pdf
Checklist of the family Simuliidae (Diptera) of Finland 91
Checklist of the family Simuliidae (Diptera) of Finland
Jari Ilmonen1
1 Metsähallitus Natural Heritage Services, Vantaa, Finland
Corresponding author: Jari Ilmonen (jari.ilmonen@metsa.)
Academic editor: J. Kahanpää|Received 27 March 2014|Accepted 19 June 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Ilmonen J (2014) Checklist of the family Simuliidae (Diptera) of Finland. In: Kahanpää J, Salmela J (Eds)
Checklist of the Diptera of Finland. ZooKeys 441: 91–95. doi: 10.3897/zookeys.441.7600
Abstract
A checklist of the family Simuliidae (Diptera) is provided for Finland and recognizes 56 species. One new
record has been added (Simulium latipes) and one name sunken in synonymy (Simulium carpathicum).
Furthermore, Simulium tsheburovae is treated as a doubtful record.
Keywords
Checklist, Finland, Diptera, Simuliidae
Introduction
Simuliidae is a relatively small family of nematoceran ies, comprised of 2,163 species
(2,151 living and 12 fossil) world-wide (Adler and Crosskey 2014). Numerous taxo-
nomical confusions and dierent views on nomenclature between eastern and western
scientists have created confusion in blacky studies for many decades. Some taxonomi-
cal issues concerning species occurring in North Europe have been resolved by recent
synonymisations (Raastad et al. 2010, Adler and Crosskey 2014). Systematics and
nomenclature of the enumeration follow that of Adler and Crosskey (2014), and only
the most relevant synonyms used in previous checklists for Finland are listed.
e faunistic level of knowledge in Finland is good, only few changes were made
since compilation of the previous checklist (Ilmonen and Adler 2008). Few additional
species can be expected to be found, and many taxonomical issues have been resolved
within the last two decades.
ZooKeys 441: 91–95 (2014)
doi: 10.3897/zookeys.441.7600
www.zookeys.org
Copyright Jari Ilmonen. This is an open access article distributed under the terms of the Creative Commons Attribution International License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jari Ilmonen / ZooKeys 441: 91–95 (2014)
92
Number of species:
World: 2151 species (Adler and Crosskey 2014)
Europe: 230 species (Adler and Crosskey 2014)
Finland: 56 species
Faunistic knowledge level in Finland: good
Checklist
Nematocera Dumeril, 1805
infraorder Culicomorpha Hennig, 1948
superfamily Chironomoidea Newman, 1834
SIMULIIDAE Newman, 1834
SIMULIINAE Newman, 1834
tribe Prosimuliini Enderlein, 1921
HELODON Enderlein, 1921
Helodon ferrugineus (Wahlberg, 1844)
PROSIMULIUM Roubaud, 1906
Prosimulium hirtipes (Fries, 1824)
Prosimulium luganicum Rubtsov, 1956
Prosimulium macropyga (Lundström, 1911)
Prosimulium ursinum (Edwards, 1935)
tribe Simuliini Newman, 1834
CNEPHIA Enderlein, 1921
Cnephia eremites Shewell, 1952
Cnephia pallipes (Fries, 1824)
= lapponica (Enderlein, 1921)
GRENIERA Doby & David, 1959
Greniera ivanovae Ivashchenko, 1970
METACNEPHIA Crosskey, 1969
Metacnephia bilineata (Rubtsov, 1940)
= saileri (Stone, 1952)
Metacnephia lyra (Lundström, 1911)
= tabescentifrons (Enderlein, 1929)
= trigoniformis Yankovsky, 2003
Metacnephia tredecimata (Edwards, 1920)
SIMULIUM Latreille, 1802
sg. Boophthora Enderlein, 1921
Simulium erythrocephalum (De Geer, 1776)
sg. Eusimulium Roubaud, 1906
Simulium angustipes Edwards, 1915
= securiforme (Rubtsov, 1956)
Checklist of the family Simuliidae (Diptera) of Finland 93
Simulium aureum Fries, 1824
Simulium velutinum (Santos Abreu, 1922)
sg. Boreosimulium Rubzov & Yankovsky, 1982
Simulium annulus (Lundström, 1911)
Simulium banense Twinn, 1936
Simulium crassum (Rubtsov, 1956)
sg. Hellichiella Rivosecchi & Cardinali, 1975
Simulium dogieli (Rubtsov, 1956)
Simulium latipes (Meigen, 1804)
Simulium usovae (Golini, 1987)
sg. Nevermannia Enderlein, 1921
= Cnetha Enderlein, 1921
Simulium angustitarse (Lundström, 1911)
Simulium beltukovae (Rubtsov, 1956)
= carpathicum (Knoz, 1961)
Simulium bicorne Dorogostaisky, Rubtsov & Vlasenko, 1935
Simulium cryophilum (Rubtsov, 1959)
= carthusiense f. brevicaulis Dorier & Grenier, 1961
Simulium curvans (Rubtsov & Carlsson, 1965)
Simulium dendrolum (Patrusheva, 1962)
Simulium fontinale Radzivilovskaya, 1948
Simulium juxtacrenobium Bass & Brockhouse, 1990
Simulium lundstromi (Enderlein, 1921)
Simulium silvestre (Rubtsov, 1956)
Simulium vernum Macquart, 1826
= pritzkowi (Enderlein, 1926)
sg. Schoenbaueria Enderlein, 1921
Simulium pusillum Fries, 1824
Simulium subpusillum Rubtsov, 1940
sg. Simulium Latreille, 1802
Simulium annulitarse Zetterstedt, 1838
Simulium argyreatum Meigen, 1838
Simulium frigidum Rubtsov, 1940
Simulium intermedium Roubaud, 1906
Simulium longipalpe Beltyukova, 1955
= curvistylus Rubtsov, 1957
Simulium monticola Friederichs, 1920
Simulium morsitans Edwards, 1915
Simulium murmanum Enderlein, 1935
= forsi (Carlsson, 1962)
Simulium noelleri Friederichs, 1920
Simulium ornatum Meigen, 1818
Simulium paramorsitans Rubtsov, 1956
Jari Ilmonen / ZooKeys 441: 91–95 (2014)
94
Simulium posticatum Meigen, 1838
= austeni Edwards, 1915
= verecundum misid.
Simulium reptans (Linnaeus, 1758)
Simulium rostratum (Lundström, 1911)
= sublacustre Davies, 1966
Simulium rubtzovi Smart, 1945
Simulium transiens Rubtsov, 1940
Simulium truncatum (Lundström, 1911)
Simulium tuberosum (Lundström, 1911)
Simulium tumulosum Rubtsov, 1956
Simulium vulgare Dorogostaisky, Rubtsov & Vlasenko, 1935
sg. Wilhelmia Enderlein, 1921
Simulium equinum (Linnaeus, 1758)
= zetlandense (Davies, 1966)
STEGOPTERNA Enderlein, 1930
Stegopterna trigonium (Lundström, 1911)
= richteri (Enderlein, 1930)
Excluded species
Simulium fuscipes (Fries, 1824) see Notes
Simulium lineatum (Meigen, 1804) misidentied, most likely S. equinum
Simulium meigeni Rubzov & Carlsson, 1965 see Notes
= pygmaeum of some authors
Simulium tsheburovae (Rubtsov, 1956) see Notes
Notes
Simulium fuscipes (Fries, 1824). Adler and Crosskey (2014) treat fuscipes as a valid
species of S. rucorne species group in the subgenus Nevermannia, but no known
records exist from Finland.
Simulium meigeni Rubzov & Carlsson, 1965. Adler and Crosskey (2014) recognize
the identity of S. meigeni within the subgenus Hellichiella, but there are no known
records of this species in Finland. e name pygmaeum in Kuusela (1971) refers to
the form of pygmaeum that is a synonym for pusillum, not meigeni (see Adler and
Crosskey 2014).
Simulium tsheburovae (Rubtsov, 1956). No conrmed records of this species exist
from Finland (Raastad et al. 2010), and the name probably subsumes one or more
of the existing species within the subgenus Hellichiella (Adler and Crosskey 2014).
Checklist of the family Simuliidae (Diptera) of Finland 95
References
Adler PH, Crosskey RW (2014) World blackies (Diptera: Simuliidae): a comprehensive revision
of the taxonomic and geographical inventory. http://www.clemson.edu/cas/biomia/pdfs/
blackyinventory.pdf
Ilmonen J, Adler P (2008) Updated checklist of the black ies (Diptera: Simuliidae) of Finland.
British Simuliid Group Bulletin 29: 9–15.
Kuusela K (1971) Preliminary notes on the blacky species (Dipt., Simuliidae) of Finland.
Annales Entomologici Fennici 37: 190–194.
Raastad JE, Usova ZV, Kuusela K (2010) Blackies of Northern Europe (Diptera: Simuliidae).
CD-ROM. ETI Bioinformatics, Amsterdam.
Jari Ilmonen / ZooKeys 441: 91–95 (2014)
96
Checklist of the families Anisopodidae, Bibionidae, Canthyloscelididae, Mycetobiidae... 97
Checklist of the families Anisopodidae, Bibionidae,
Canthyloscelididae, Mycetobiidae, Pachyneuridae
and Scatopsidae (Diptera) of Finland
Antti Haarto1
1 Zoological Museum, Section of Biodiversity and Environmental Science, Department of Biology, FI-20014
University of Turku, Finland
Corresponding author: Antti Haarto (ahaarto@gmail.com)
Academic editor: J. Kahanpää|Received 23 February 2014|Accepted 28 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Haarto A (2014) Checklist of the families Anisopodidae, Bibionidae, Canthyloscelididae, Mycetobiidae,
Pachyneuridae and Scatopsidae (Diptera) of Finland. In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of
Finland. ZooKeys 441: 97–102. doi: 10.3897/zookeys.441.7358
Abstract
A checklist of the families Anisopodidae, Bibionidae, Canthyloscelididae, Mycetobiidae, Pachyneuri-
dae and Scatopsidae (Diptera) recorded from Finland. e Finnish species of these families were last
listed in 1980. After that two species of Anisopodidae, four species of Bibionidae and six species of
Scatopsidae have been added to the Finnish fauna and two species of Scatopsidae have been removed
from the Finnish fauna.
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
Six families (Anisopodidae, Bibionidae, Canthyloscelididae, Mycetobiidae, Pach-
yneuridae and Scatopsidae) are included in this paper. All six families have been rather
poorly collected and studied in Finland. Lundström (1910) published a study of the
family Bibionidae. During the last one hundred years, foreign specialists have con-
ZooKeys 441: 97–102 (2014)
doi: 10.3897/zookeys.441.7358
www.zookeys.org
Copyright Antti Haarto. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
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A peer-reviewed open-access journal
Antti Haarto / ZooKeys 441: 97–102 (2014)
98
tributed the majority of the new records and taxonomic treatises. Some Anisopodidae
were treated by Krivosheina and Menzel (1998, 2002) and Bibionidae by Greve and
Haenni (1994) and Skartveit (1999). Finnish scatopsid records were included in the
synopsis of Palaearctic Scatopsidae by Cook (1969, 1972, 1974). e genera Anapau-
sis Enderlein (Chandler 1999, Haenni 1984), Colobostema Enderlein (Haenni 2013,
Krivosheina 2001) and Aspistes Meigen (Krivosheina 2000) have been examined in
more detail. Recently Anisopodidae (Haarto 2011) and Bibionidae (Haarto 2012)
have attracted some attention in the country. e only Finnish species of Pachyneu-
ridae, Pachyneura fasciata Zetterstedt, 1838, was studied by Salmela and Ilmonen
(2004) and removed from the Finnish Red List. e Finnish species of all six families
were last listed by Hackman (1980). e Finnish faunae of anisopodids and bibio-
nids have been recently reviewed by Haarto (2011, 2012). e nomenclature and the
number of European species are presented according to the current listing of Fauna
Europaea (Beuk and Pape 2013).
Checklist
Nematocera Duméril, 1805
infraorder Bibionomorpha Hennig, 1954
superfamily Anisopodoidea Knab, 1912
ANISOPODIDAE Knab, 1912
SYLVICOLA Harris, 1780
= Phryne Meigen, 1800 suppr.
= Rhyphus Latreille, 1804
Sylvicola cinctus (Fabricius, 1787)
Sylvicola fenestralis (Scopoli, 1763)
Sylvicola fuscatoides Michelsen, 1999
= fuscatus sensu Andersson 1967, nec Fabricius
Sylvicola fuscatus (Fabricius, 1775)
Sylvicola punctatus (Fabricius, 1787)
Sylvicola stackelbergi Krivosheina & Menzel, 1998
Sylvicola zetterstedti (Edwards, 1923)
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Anisopodidae 77 10 7 average
Mycetobiidae 35 4 1 average
Bibionidae 760 50 17 average
Scatopsidae 390 102 30 poor
Canthyloscelididae 16 3 3 average
Pachyneuridae 5 1 1 average
Checklist of the families Anisopodidae, Bibionidae, Canthyloscelididae, Mycetobiidae... 99
MYCETOBIIDAE Winnertz, 1863
MYCETOBIA Meigen, 1818
Mycetobia pallipes Meigen, 1818
superfamily Bibionoidea Fleming, 1821
BIBIONIDAE Fleming, 1821
BIBIONINAE Fleming, 1821
BIBIO Georoy, 1762
Bibio brunnipes (Fabricius, 1794)
= fulvipes (Zetterstedt, 1838)
Bibio clavipes Meigen, 1818
Bibio ferruginatus (Linnaeus, 1767)
Bibio fulvicollis Gimmerthal, 1842
= festinans (Zetterstedt, 1850)
Bibio johannis (Linnaeus, 1767)
Bibio lautaretensis Villeneuve, 1925
= crassipes Duda, 1930
Bibio longipes Loew, 1864
= lepidus Loew, 1871
Bibio marci (Linnaeus, 1758)
Bibio nigriventris Haliday, 1833
Bibio pomonae (Fabricius, 1775)
Bibio rupes (Zetterstedt, 1838)
Bibio siebkei Mik, 1887
= femoralis (Siebke, 1863) preocc.
Bibio varipes Meigen, 1830
DILOPHUS Meigen, 1803
Dilophus borealis Skartveit, 1993
Dilophus febrilis (Linnaeus, 1758)
Dilophus femoratus Meigen, 1804
PLECIINAE Fleming, 1821
PENTHETRIA Meigen, 1803
Penthetria funebris Meigen, 1804
= holosericea Meigen, 1818
superfamily Scatopsoidea Newman, 1834
SCATOPSIDAE Newman, 1834
ASPISTINAE Rondani, 1840
ARTHRIA Kirby in Richardson, 1837
Arthria analis (Kirby in Richardson, 1837)
ASPISTES Meigen, 1818
Aspistes berolinensis Meigen, 1818
Antti Haarto / ZooKeys 441: 97–102 (2014)
100
Aspistes freyi Cook, 1965
Aspistes helleni Krivosheina, 2000
ECTAETIINAE Enderlein, 1936
ECTAETIA Enderlein, 1912
Ectaetia clavipes (Loew, 1846)
Ectaetia platyscelis (Loew, 1869)
PSECTROSCIARINAE Cook, 1963
ANAPAUSIS Enderlein, 1912
Anapausis oricola Chandler, 1999
= soluta misid.
Anapausis rectinervis Duda, 1928
SCATOPSINAE Newman, 1834
tribe Colobostematini Amorim, 1994
COLOBOSTEMA Enderlein, 1926
Colobostema infumatum (Haliday, 1833)
Colobostema nigripenne (Meigen, 1830)
Colobostema triste (Zetterstedt, 1850)
EFCOOKELLA Haenni, 1998
Efcookella albitarsis (Zetterstedt, 1850)
FERNEIELLA Cook in Freeman, 1985
Ferneiella incompleta (Verrall, 1886)
HOLOPLAGIA Enderlein, 1912
Holoplagia bullata (Edwards, 1925)
Holoplagia transversalis (Loew, 1846)
tribe Rhegmoclematini Cook, 1955
RHEGMOCLEMINA Enderlein, 1936
Rhegmoclemina vaginata (Lundström, 1910)
THRIPOMORPHA Enderlein, 1905
= Rhegmoclema Enderlein, 1912
ripomorpha freyi (Cook, 1969)
ripomorpha halteratum (Meigen, 1838)
ripomorpha paludicola Enderlein, 1905
= edwardsi (Collin, 1954)
ripomorpha verralli (Edwards, 1934)
tribe Scatopsini Newman, 1834
APILOSCATOPSE Cook, 1974
Apiloscatopse avicollis (Meigen, 1818)
Apiloscatopse avocincta (Duda, 1928)
REICHERTELLA Enderlein, 1912
Reichertella geniculata (Zetterstedt, 1850)
SCATOPSE Georoy, 1762
Scatopse lapponica Duda, 1928
Scatopse notata (Linnaeus, 1758)
Checklist of the families Anisopodidae, Bibionidae, Canthyloscelididae, Mycetobiidae... 101
tribe Swammerdamellini Cook, 1972
COBOLDIA Melander, 1916
Coboldia fuscipes (Meigen, 1830)
RHEXOZA Enderlein, 1936
Rhexoza subnitens (Verrall, 1886)
SWAMMERDAMELLA Enderlein, 1912
Swammerdamella adercotris Cook, 1972
Swammerdamella brevicornis (Meigen, 1830)
Swammerdamella genypodis Cook, 1972
CANTHYLOSCELIDIDAE Enderlein, 1912
CANTHYLOSCELIDINAE Enderlein, 1912
HYPEROSCELIS Hardy & Nagatomi, 1960
Hyperoscelis eximia (Boheman, 1858)
Hyperoscelis veternosa Mamaev & Krivosheina, 1969
SYNNEURINAE Enderlein, 1936
SYNNEURON Lundström, 1910
Synneuron annulipes Lundström, 1910
superfamily Pachyneuroidea Schiner, 1864
PACHYNEURIDAE Schiner, 1864
PACHYNEURA Zetterstedt, 1838
Pachyneura fasciata Zetterstedt, 1838
Excluded species
Anapausis soluta (Loew, 1846): Specimens under the name A. soluta in the earlier checklist
(Hackman 1980) represent the later described species A. oricola Chandler, 1999.
Swammerdamella acuta Cook, 1956: Has not been found within present borders of
Finland.
Notes
Aspistes helleni Krivosheina, 2000 in this checklist based on other material than the one
mentioned by Krivosheina (2000).
References
Beuk P, Pape T (2013) Fauna Europaea: Diptera. Fauna Europaea version 2.6. faunaeur.org
Antti Haarto / ZooKeys 441: 97–102 (2014)
102
Chandler PJ (1999) e British species of Anapausis Enderlein (Diptera, Scatopsidae). Dipter-
ists Digest 6: 1–21.
Cook EF (1969) A synopsis of the Scatopsidae of the Palaearctic Part I. Rhegmoclematini.
Journal of Natural History 3(3): 393–407. doi: 10.1080/00222936900770341
Cook EF (1972) A synopsis of the Scatopsidae of the Palaearctic Part II. Swammerdamellini.
Journal of Natural History 6(6): 625–634. doi: 10.1080/00222937200770561
Cook EF (1974) A Synopsis of the Scatopsidae of the palaearctic Part III. e Scatopsini. Jour-
nal of Natural History 8(1): 61–100. doi: 10.1080/00222937400770061
Greve L, Haenni J-P (1994) Revision of the European species of the Bibio lautaretensis-group
(Diptera, Bibionidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft
67: 385–392.
Haarto A (2011) Suomen ikkunasääsket – Finnish window gnats (Diptera, Anisopodidae). w-
album 10: 3–13. http://org.utu./harrastus/TEKS/w-album/w_album_10.pdf
Haarto A (2012) Suomen karvasääsket ja maasääsket (Diptera, Bibionidae ja Pleciidae). w-al-
bum 12: 3–31. http://org.utu./harrastus/TEKS/w-album/w_album_12.pdf
Hackman W (1980) A Check List of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Haenni J-P (1984) Contribution à la connaissance de la faune des Scatopsidae (Diptera) de Sui-
sse. II. Le genre Anapausis Enderlein. Mitteilungen der Schweizerischen Entomologischen
Gesellschaft 57: 273–285.
Haenni J-P (2013) A revision of the West Palaearctic species of Colobostema Enderlein, 1926
(Diptera, Scatopsidae). Part I. European subregion. Mitteilungen der Schweizerischen
Entomologischen Gesellschaft 86: 199–242.
Krivosheina NP (2000) A new and little known species of the genus Aspistes Meig. (Diptera,
Scatopsidae) from Finland. Entomologica Fennica 11: 119–123.
Krivosheina NP (2001) To the fauna of scatopsid ies of the genus Colobostema Enderlein of
Finland, with the description of a new species (Diptera, Scatopsidae). An international
Journal of Dipterological Research 12: 73–77.
Krivosheina NP, Menzel F (1998) e Palaearctic species of the genus Sylvicola Harris, 1776
(Diptera, Anisopodidae). Beiträge zur Entomologie 48(1): 201–217.
Krivosheina NP, Menzel F (2002) K морфологии самок ксило-мицетофильных видов рода
Sylvicola Harris (Diptera, Anisopodidae) [e morphology of females of the xylo-mycetophilous
species of the genus Sylvicola Harris]. Entomologitscheskoje obozrenije 81: 424–438.
Lundström C (1910) Beiträge zur Kenntnis der Dipteren Finlands. V. Bibionidae. Acta Societatis
pro Fauna et Flora Fennica 33: 4–15.
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Salmela J, Ilmonen J (2004) New records of the threatened y Pachyneura fasciata Zetterstedt,
1838 in eastern Fennoscandia (Diptera, Nematocera). Sahlbergia 9: 123–126.
Skartveit J (1999) Two species of Bibionidae (Diptera) new to the Finnish fauna. Entomologica
Fennica 10: 115–116.
Checklist of the Cecidomyiidae (Diptera) of Finland 103
Checklist of the Cecidomyiidae (Diptera) of Finland
Mathias Jaschhof1, Marcela Skuhravá2, Jouni Penttinen3
1 älmann-Ring 64, D-17491 Greifswald, Germany 2 Bítovská 1227/9, 140 00 Praha 4, Czech Republic
3Metsähallitus, Natural Heritage, P.O. Box 36, FIN-40101 Jyväskylä, Finland
Corresponding author: Mathias Jaschhof (mjaschhof@yahoo.de)
Academic editor: J. Kahanpää|Received 13 March 2014|Accepted 3 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Jaschhof M, Skuhravá M, Penttinen J (2014) Checklist of the Cecidomyiidae (Diptera) of Finland. In: Kahanpää
J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 103–118. doi: 10.3897/zookeys.441.7503
Abstract
A list of the 356 species of Cecidomyiidae (Diptera) recorded from Finland is presented, which comprises
6 Lestremiinae, 156 Micromyinae, 16 Winnertziinae, 69 Porricondylinae, and 109 Cecidomyiinae. e
faunistic knowledge of Finnish Winnertziinae, Porricondylinae and Cecidomyiinae is regarded as particu-
larly poor. Based on species numbers known from other countries in Europe, a conservative estimate is
700–800 species of Cecidomyiidae actually occurring in Finland.
Keywords
Gall midges, Finland, species list
Introduction
With a world total of more than 6200 species described, Cecidomyiidae are among
the largest families of Diptera, although adult cecidomyiids are mostly delicate midges
with body lengths of less than 5 mm. e family has been catalogued (Gagné and
Jaschhof 2014) and that publication provides bibliographic and other information
on all known species. Six subfamilies are recognised: the fungivorous Catotrichinae,
Lestremiinae, Micromyinae, Winnertziinae and Porricondylinae, and the mostly her-
bivorous Cecidomyiinae. e latter subfamily, which also includes fungivores and pre-
dators, comprises the largest gall-making group of insects (hence the family name that
translates as gall midges).
ZooKeys 441: 103–118 (2014)
doi: 10.3897/zookeys.441.7503
www.zookeys.org
Copyright Mathias Jaschhof et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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Mathias Jaschhof et al. / ZooKeys 441: 103–118 (2014)
104
In the fungivorous subfamilies, many larvae are associated with dead wood, rende-
ring those a biodiverse, albeit little studied group of saproxylic insects. In Cecidomyii-
nae, larvae associated with living owering plants are usually restricted to development
on one, or a few closely-related host species; predators are also generally assumed to be
highly host-specic, as far as is known.
e Palaearctic has more named Cecidomyiidae species than any other zoo-
geographical region, and the best known gall midge fauna is that of Europe, with
more than 1800 species (Skuhravá 2006). Even so, European Cecidomyiidae are far
from being completely known, and the Finnish fauna provides a good example. To
illustrate, while the previous checklist of Finnish Cecidomyiidae (Hackman 1980)
comprised only four species of Lestremiinae and Micromyinae, a recent taxonomic
study increased that number to 162, including 34 new species (Jaschhof and Jaschhof
2009). As for Winnertziinae and Porricondylinae, 27 species in Hackman’s (1980)
list compare with 69 species recorded here (Penttinen and Siitonen 2005, Penttinen
and Spungis 2007, Penttinen and Jaschhof 2009; Jaschhof, unpublished; Penttinen,
unpublished), yet much remains to be discovered, as is testied by specimens in the
Penttinen Collection belonging to a number of unnamed species. In Cecidomyiinae,
only three species (Hulden 2003; Skuhravá, unpublished) are here added to the
Hackman list, a meager increase that is obviously explained by the slight research
eort in recent years.
Identication of cecidomyiids, both larvae and adults, is generally dicult and
usually left to a few taxonomic specialists of the family. An adult key to the Palaearctic
genera of Cecidomyiidae was provided by Skuhravá (1997); to the Holarctic genera
of Lestremiinae and Micromyinae by Jaschhof and Jaschhof (2009), and to the Hol-
arctic genera of Winnertziinae and Porricondylinae by Jaschhof and Jaschhof (2013).
Adult keys and illustrations by Jaschhof and Jaschhof (2009, 2013) can be used to
identify the Finnish species of the fungivorous subfamilies. Adult keys to identify
species of Cecidomyiinae in Finland are only available for a few small groups, so any
attempts in this regard rely largely on descriptions and illustrations scattered in an ex-
tensive literature. In general, many Cecidomyiinae species cannot be identied unless
a bundle of information (regarding host, gall, immature stages, adults of both sexes)
are known. Illustrated keys to galls, including those caused by gall midges, have been
published for Central and Northern Europe by Buhr (1964, 1965), a resource that
is currently being revised and updated for publication in English in 2015 (see http://
www.panzengallen.de).
The arrangement of taxa in this list largely follows the recent edition of the
world catalogue (Gagné and Jaschhof 2014) except that genera are here arranged
under their pertinent tribes and supertribes, if known. All Finnish records of
species of the fungivorous subfamilies were revised in recent years (Jaschhof and
Jaschhof 2009, 2013; Penttinen, unpublished), so can be related to actual voucher
specimens deposited in museum collections. The list of Cecidomyiinae, which is
not updated to the same extent, is compiled from published locality records based
on varying evidence, including galls alone.
Checklist of the Cecidomyiidae (Diptera) of Finland 105
Number of species:
World: 6203 species (Gagné and Jaschhof 2014)
Europe: 1800 species (Skuhravá 2006)
Finland: 356 species (this paper)
Faunistic knowledge level in Finland: poor
Checklist
Nematocera Dumeril, 1805
infraorder Bibionomorpha Hennig, 1954
superfamily Sciaroidea Billberg, 1820
CECIDOMYIIDAE Newman, 1834
LESTREMIINAE Rondani, 1840
tribe Lestremiini Rondani, 1840
ALLARETE Pritchard, 1951
Allarete nigra Mamaev, 1994
ANARETE Haliday, 1833
Anarete sp. 2 [see Note below]
ANARETELLA Enderlein, 1911
Anaretella defecta (Winnertz, 1870)
= magnicornis Mamaev, 1964
Anaretella iola Pritchard, 1951
LESTREMIA Macquart, 1826
Lestremia cinerea Macquart, 1826
Lestremia leucophaea (Meigen, 1818)
MICROMYINAE Rondani, 1856
tribe Acoenoniini Pritchard, 1960
ACOENONIA Pritchard, 1947
Acoenonia europaea Mamaev, 1964
tribe Aprionini Jaschhof, 1998
APRIONUS Kieer, 1894
Aprionus accipitris Jaschhof, 1997
Aprionus acutus Edwards, 1938
Aprionus adventitius Jaschhof, 2009
Aprionus aquilonius Jaschhof, 2009
Aprionus arcticus Mamaev, 2001
Aprionus aviarius Mamaev & Berest, 1990
Aprionus betulae Jaschhof, 1996
Aprionus bidentatus (Kieer, 1894)
Aprionus bidus Mamaev, 1963
Aprionus bispinosus Edwards, 1938
Mathias Jaschhof et al. / ZooKeys 441: 103–118 (2014)
106
Aprionus brachypterus Edwards, 1938
Aprionus brevitegminis Jaschhof, 2009
Aprionus cardiophorus Mamaev, 1963
Aprionus carinatus Jaschhof, 1996
Aprionus complicatus Mamaev & Berest, 1986
Aprionus confusus Mamaev, 1969
Aprionus corniculatus Mamaev, 1963
Aprionus cornutus Berest, 1986
Aprionus dalarnensis Mamaev, 1998
Aprionus dentifer Mamaev, 1965
Aprionus dispar Mamaev, 1963
Aprionus dissectus Mamaev & Berest, 1990
Aprionus duplicatus Mamaev, 1998
Aprionus ensiferus Jaschhof, 1996
Aprionus fennicus Jaschhof, 2009
Aprionus avidus (Winnertz, 1870)
Aprionus foliosus Jaschhof, 2009
Aprionus gladiator Jaschhof, 2009
Aprionus halteratus (Zetterstedt, 1852)
Aprionus heothinos Jaschhof, 2009
Aprionus hintelmannorum Jaschhof, 2009
Aprionus inquisitor Mamaev, 1963
Aprionus insignis Mamaev, 1963
Aprionus laevis Mohrig, 1967
Aprionus lapponicus Jaschhof & Mamaev, 1997
Aprionus laricis Mamaev & Jaschhof, 1997
Aprionus latitegminis Jaschhof, 2009
Aprionus longicollis Mamaev, 1963
Aprionus longitegminis Yukawa, 1967
Aprionus marginatus Mamaev, 1963
Aprionus miki Kieer, 1895
Aprionus oligodactylus Jaschhof, 2009
Aprionus piceae Jaschhof, 1997
Aprionus praecipuus Jaschhof, 2009
Aprionus pyxidiifer Mamaev, 1998
Aprionus reduncus Jaschhof, 2009
Aprionus separatus Mamaev & Jaschhof, 1997
Aprionus sievertorum Jaschhof, 2009
Aprionus similis Mamaev, 1963
Aprionus spiniferus Mamaev & Berest, 1990
Aprionus spiniger (Kieer, 1894)
Aprionus stiktos Jaschhof, 2009
Aprionus stylifer Mamaev, 1998
Checklist of the Cecidomyiidae (Diptera) of Finland 107
Aprionus styloideus Mamaev & Berest, 1990
Aprionus subacutus Jaschhof, 1997
Aprionus svecicus Jaschhof, 1996
Aprionus taigaensis Jaschhof, 2009
Aprionus tiliamcorticis Mamaev, 1963
Aprionus victoriae Jaschhof, 2009
tribe Bryomyiini Berest, 1993
BRYOMYIA Kieer, 1895
= Tomonomyia Berest, 1993
Bryomyia apsectra Edwards, 1938
Bryomyia bergrothi Kieer, 1895
Bryomyia gibbosa (Felt, 1907)
Bryomyia producta (Felt, 1908)
HETEROGENELLA Mamaev, 1963
= Cervuatina Berest, 1993
Heterogenella nitima Mamaev, 1998
Heterogenella hybrida Mamaev, 1963
Heterogenella linearis Yukawa, 1971
SKUHRAVIANA Mamaev, 1963
Skuhraviana triangulifera Mamaev, 1963
tribe Campylomyzini Kieer, 1898
CAMPYLOMYZA Meigen, 1818
Campylomyza abbreviata Jaschhof, 2009
Campylomyza aemula Mamaev, 1998
Campylomyza alpina Siebke, 1863
Campylomyza arcuata Jaschhof, 2009
Campylomyza armata Mamaev, 1963
Campylomyza cavitata Mamaev, 1998
Campylomyza cingulata Jaschhof, 2009
Campylomyza dilatata Felt, 1907
Campylomyza falcifera Jaschhof, 2009
Campylomyza avipes Meigen, 1818
= pallipes Zetterstedt, 1850
Campylomyza furva Edwards, 1938
Campylomyza fusca Winnertz, 1870
Campylomyza inornata Jaschhof, 2009
Campylomyza insolita Jaschhof, 2009
Campylomyza ormerodi (Kieer, 1913)
Campylomyza serrata Jaschhof, 1998
Campylomyza spatulata Mamaev, 1998
Campylomyza stegetfore Jaschhof, 2009
EXCRESCENTIA Mamaev & Berest, 1991
Excrescentia mutuata Mamaev & Berest, 1991
Mathias Jaschhof et al. / ZooKeys 441: 103–118 (2014)
108
NEUROLYGA Rondani, 1840
= Cordylomyia Felt, 1911
Neurolyga acuminata Jaschhof, 2009
Neurolyga bilobata (Mamaev & Rozhnova, 1982)
Neurolyga excavata (Yukawa, 1967)
Neurolyga interrupta Jaschhof, 2009
Neurolyga lonsdalensis Jaschhof, 2009
Neurolyga paludosa Jaschhof, 2009
Neurolyga sylvestris (Felt, 1907)
Neurolyga verna (Mamaev, 1963)
tribe Catochini Edwards, 1938
CATOCHA Haliday, 1833
Catocha incisa Jaschhof, 2009
Catocha latipes Haliday, 1833
tribe Micromyini Rondani, 1856
MICROMYA Rondani, 1840
Micromya lucorum Rondani, 1840
MONARDIA Kieer, 1895
= Pezomyia Kieer, 1913
sg. Monardia Kieer, 1895
Monardia abnormis Mamaev, 1963
Monardia armata Jaschhof, 2003
Monardia lignivora (Felt, 1907)
Monardia obsoleta Edwards, 1938
Monardia pediculata (Mamaev, 1993)
Monardia stirpium Kieer, 1895
Monardia yasumatsui Yukawa, 1967
sg. Trichopteromyia Williston, 1896
Monardia magnica Mamaev, 1963
Monardia relicta Jaschhof, 2009
sg. Xylopriona Kieer, 1913
Monardia adentis Jaschhof, 1998
Monardia atra (Meigen, 1804)
Monardia monotheca Edwards, 1938
Monardia radiella Mamaev, 1993
Monardia toxicodendri (Felt, 1907)
Monardia unguifera Berest & Mamaev, 1997
POLYARDIS Pritchard, 1947
Polyardis adela Pritchard, 1947
Polyardis bispinosa (Mamaev, 1963)
Polyardis silvalis (Róndani, 1840)
tribe Peromyiini Kleesattel, 1979
PEROMYIA Kieer, 1894
Checklist of the Cecidomyiidae (Diptera) of Finland 109
Peromyia abdita Jaschhof, 2009
Peromyia abnormis Mamaev & Berest, 1990
Peromyia albicornis (Meigen, 1830)
Peromyia anatina Mamaev & Berest, 1990
Peromyia angellifera Jaschhof, 1997
Peromyia anisotoma Mamaev, 1994
Peromyia apposita Jaschhof, 1997
Peromyia bicolor (Edwards, 1938)
Peromyia bidentata Berest, 1998
Peromyia boreophila Jaschhof, 2001
Peromyia caricis (Kieer, 1901)
Peromyia concitata Mamaev & Berest, 1994
Peromyia cornuta (Edwards, 1938)
Peromyia curta Jaschhof, 1997
Peromyia diadema Mamaev, 1963
Peromyia edwardsi Berest, 1994
Peromyia fagiphila Jaschhof, 1997
Peromyia fungicola (Kieer, 1901)
Peromyia intermedia (Kieer, 1895)
Peromyia mitrata Jaschhof, 1997
Peromyia modesta (Felt, 1907)
Peromyia monilis Mamaev, 1965
Peromyia muscorum (Kieer, 1895)
Peromyia ovalis (Edwards, 1938)
Peromyia palustris (Kieer, 1895)
Peromyia perpusilla (Winnertz, 1870)
Peromyia photophila (Felt, 1907)
Peromyia pumila Jaschhof, 2001
Peromyia ramosa (Edwards, 1938)
Peromyia ramosoides Jaschhof, 2009
Peromyia sanguinea (Kieer, 1894)
Peromyia scirrhosa Jaschhof, 2009
Peromyia scutellata Mamaev, 1990
Peromyia semotoides Jaschhof, 2009
Peromyia subbicolor Jaschhof, 2009
Peromyia subborealis Jaschhof, 1997
Peromyia sylteordensis Jaschhof, 1996
Peromyia truncata Yukawa, 1967
Peromyia tundrae Jaschhof, 1996
Peromyia upupoides Jaschhof, 1997
WINNERTZIINAE Panelius, 1965
tribe Diallactiini Jaschhof, 2013
DIALLACTIA Gagné, 2004
Mathias Jaschhof et al. / ZooKeys 441: 103–118 (2014)
110
= Diallactes Kieer, 1894 preocc.
Diallactia crocea (Kieer, 1894)
= obscuripes (Spungis, 1985)
SYLVENOMYIA Mamaev & Zaitzev, 1998
Sylvenomyia fennica Penttinen & Jaschhof, 2009
Sylvenomyia spinigera (Spungis, 1985)
tribe Heteropezini Schiner, 1868
HETEROPEZA Winnertz, 1846
Heteropeza pygmaea Winnertz, 1846
LEPTOSYNA Kieer, 1894
Leptosyna similis Jaschhof, 2013
MIASTOR Meinert, 1864
Miastor metraloas Meinert, 1864
tribe Winnertziini Panelius, 1965
KRONOMYIA Felt, 1911
Kronomyia ovalis (Mamaev, 1964)
RHIPIDOXYLOMYIA Mamaev, 1964
Rhipidoxylomyia brevicornis Mamaev, 1964
WINNERTZIA Rondani, 1860
Winnertzia bulbifera Mamaev, 1963
Winnertzia curvata Panelius, 1965
Winnertzia fusca Kieer, 1901
Winnertzia globifera Mamaev, 1963
Winnertzia graduata Spungis, 1992
Winnertzia nigripennis Kieer, 1896
Winnertzia rotundata Spungis, 1992
Winnertzia solidaginis Felt, 1907
= calciequina Felt, 1907
PORRICONDYLINAE Kieer, 1913
tribe Asynaptini Rübsaamen & Hedicke, 1926
ASYNAPTA Loew, 1850
Asynapta breviata Spungis, 1988
Asynapta inata Spungis, 1988
Asynapta magdalini Panelius, 1965
Asynapta pectoralis (Winnertz, 1853)
Asynapta rufomaculata Panelius 1965
Asynapta saliciperda Felt, 1908
= populnea Panelius, 1965
Asynapta strobi (Kieer, 1920)
CAMPTOMYIA Kieer, 1894
Camptomyia abnormis Mamaev, 1961
Camptomyia calcarata Mamaev, 1964
Camptomyia corticalis (Loew, 1851)
Checklist of the Cecidomyiidae (Diptera) of Finland 111
Camptomyia avocinerea Panelius, 1965
Camptomyia gigantea Spungis, 1989
Camptomyia pinicola Mamaev, 1961
Camptomyia piptopori Panelius, 1965
Camptomyia regia Spungis, 1989
Camptomyia salicicola Mamaev, 1961
= populicola Mamaev, 1961
Camptomyia spinifera Mamaev, 1961
COLOMYIA Kieer, 1892
Colomyia clavata Kieer, 1892
PARASYNAPTA Panelius, 1965
Parasynapta intermedia Panelius, 1965
tribe Dicerurini Mamaev, 1966
= Solntseviini Mamaev, 1966
DICERURA Kieer, 1898
Dicerura furculata Mamaev, 1968
Dicerura mixta Spungis, 1987
Dicerura rossica (Mamaev, 1960)
Dicerura triangularis Mamaev, 1966
Dicerura unidentata Spungis, 1987
DIRHIZA Loew, 1850
Dirhiza lateritia Loew, 1850
HILVERSIDIA Mamaev, 1966
Hilversidia autumnalis Mamaev, 1966
PARATETRANEUROMYIA Spungis, 1987
Paratetraneuromyia vernalis Spungis, 1987
SOLNTSEVIA Mamaev, 1965
Solntsevia nigripes Mamaev, 1965
TETRANEUROMYIA Mamaev, 1964
Tetraneuromyia lenticularis (Spungis, 1987)
tribe Porricondylini Kieer, 1913
= Holoneurini Enderlein, 1936
ARCTEPIDOSIS Mamaev, 1990
Arctepidosis paneliusi Mamaev & Zaitzev, 1998
CASSIDOIDES Mamaev, 1960
Cassidoides fulvus (Kieer, 1896)
= pini Mamaev, 1960
CLASPETTOMYIA Grover, 1964
= Pachylabis Panelius, 1965
Claspettomyia hamata (Felt, 1907)
Claspettomyia niveitarsis (Zetterstedt, 1850)
Claspettomyia rossica Mamaev, 1998
Claspettomyia ussuriensis Mamaev, 1998
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112
COCCOPSILIS Harris, 2004
= Coccopsis Meijere preocc.
Coccopsilis marginata (Meijere, 1901)
Coccopsilis obscura (Mamaev, 1964)
Coccopsilis paneliusi (Yukawa, 1971)
DENDREPIDOSIS Mamaev, 1990
Dendrepidosis longipennis (Spungis, 1981)
DIVELLEPIDOSIS Fedotova & Sidorenko, 2007
Divellepidosis armilla (Mamaev, 1994)
Divellepidosis fuscostriata (Panelius, 1965)
Divellepidosis hypoxantha (Panelius, 1965)
Divellepidosis lutescens (Spungis, 1981)
Divellepidosis pallescens (Panelius, 1965)
Divellepidosis taigacola Jaschhof, 2013
JAMALEPIDOSIS Mamaev, 1990
Jamalepidosis spungisi Jaschhof, 2013
MONEPIDOSIS Mamaev, 1966
Monepidosis furcata Mamaev, 1966
Monepidosis pectinata Mamaev, 1966
PANELIUSIA Jaschhof, 2013
Paneliusia albimanoides Jaschhof, 2013
Paneliusia aurantiaca (Panelius, 1965)
= modesta (Spungis, 1981)
PAREPIDOSIS Kieer, 1913
Parepidosis arcuata Mamaev, 1964
= longinodis Panelius, 1965
Parepidosis venustior Gagné, 2004
PAURODYLA Jaschhof, 2013
Paurodyla tyresta Jaschhof, 2013
PORRICONDYLA Rondani, 1840
Porricondyla fulvescens Panelius, 1965
Porricondyla neglecta Mamaev in Mamaev & Krivosheina, 1965
Porricondyla nigripennis (Meigen, 1830)
Porricondyla rufescens Panelius, 1965
PSEUDEPIDOSIS Mamaev, 1966
Pseudepidosis trida Mamaev, 1966
ROSTELLATYLA Jaschhof, 2013
Rostellatyla rostellata (Panelius, 1965)
ROSTRATYLA Jaschhof, 2013
Rostratyla globosa (Spungis, 1981)
SCHISTONEURUS Mamaev, 1964
Schistoneurus impressus Mamaev, 1964
Schistoneurus irregularis Mamaev, 1964
Checklist of the Cecidomyiidae (Diptera) of Finland 113
SERRATYLA Jaschhof, 2013
Serratyla furcata (Mamaev, 2001)
Serratyla pubescens (Walker, 1856)
Serratyla spinosa Jaschhof, 2013
SPUNGISOMYIA Mamaev & Zaitzev, 1996
Spungisomyia media (Spungis, 1981)
ZAITZEVIOLA Fedotova & Sidorenko, 2007
Zaitzeviola latistylata Jaschhof, 2013
Zaitzeviola pilosistylata Jaschhof, 2013
Zaitzeviola rufocinerea (Panelius, 1965)
CECIDOMYIINAE Newman, 1834
supertribe Cecidomyiidi Rübsaamen & Hedicke, 1925
tribe Aphidoletini Harris, 1966
APHIDOLETES Kieer, 1904
Aphidoletes aphidimyza (Rondani, 1847)
Aphidoletes urticaria (Kieer, 1895)
MONOBREMIA Kieer, 1912
Monobremia subterranea (Kieer, 1898)
tribe Asphondyliini Rübsaamen & Hedicke, 1925
ASPHONDYLIA Loew, 1850
Asphondylia serpylli Kieer, 1898
= thymi Kieer, 1898
KIEFFERIA Mik, 1895
Kieeria pericarpiicola (Bremi, 1847)
SCHIZOMYIA Kieer, 1889
Schizomyia galiorum Kieer, 1889
tribe Cecidomyiini Rübsaamen & Hedicke, 1925
ANISOSTEPHUS Rübsaamen, 1917
Anisostephus betulinus (Kieer, 1889)
CECIDOMYIA Meigen, 1803
Cecidomyia pini (De Geer, 1776)
CONTARINIA Rondani, 1860
Contarinia coryli (Kaltenbach, 1859)
Contarinia craccae Loew, 1850
Contarinia gei Kieer, 1909
= geicola Rübsaamen, 1917
Contarinia kanervoi Barnes, 1958
Contarinia lonicerearum (F. Löw, 1877)
Contarinia loti (De Geer, 1776)
Contarinia merceri Barnes, 1930
Contarinia petioli (Kieer, 1898)
Contarinia pisi (Loew, 1850)
Contarinia quercina (Rübsaamen, 1890)
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114
Contarinia sambuci (Kaltenbach, 1873)
= lonicerearum (F. Löw, 1874)
Contarinia tiliarum (Kieer, 1890)
Contarinia tritici (Kirby, 1798)
Contarinia vincetoxici Kieer, 1909
MACRODIPLOSIS Kieer, 1895
Macrodiplosis pustularis (Bremi, 1847)
= dryobia (Löw, 1877)
Macrodiplosis roboris (Hardy, 1854)
= volvens Kieer, 1895
PLEMELIELLA Seitner, 1908
Plemeliella abietina Seitner, 1908
STENODIPLOSIS Reuter, 1895
Stenodiplosis geniculati Reuter, 1895
THECODIPLOSIS Kieer, 1895
ecodiplosis brachyntera (Schwägrichen, 1835)
tribe Clinodiplosini Enderlein, 1936
AMETRODIPLOSIS Rübsaamen, 1910
Ametrodiplosis thalictricola (Rübsaamen, 1895)
CLINODIPLOSIS Kieer, 1894
Clinodiplosis cilicrus (Kieer, 1889)
SITODIPLOSIS Kieer, 1913
Sitodiplosis mosellana (Géhin, 1857)
tribe Hormomyiini Rübsaamen & Hedicke, 1925
PLANETELLA Westwood, 1840
Planetella ?grandis (Meigen, 1804)
tribe Lestodiplosini Harris, 1966
LESTODIPLOSIS Kieer, 1894
Lestodiplosis pallidicornis Kieer, 1898
unplaced Cecidomyiidi
HARMANDIOLA Skuhravá, 1997
= Harmandia Kieer, 1896 preocc.
Harmandiola globuli (Rübsaamen, 1889)
Harmandiola tremulae (Winnertz, 1853)
= loewi (Rübsaamen, 1917)
HYGRODIPLOSIS Kieer, 1912
Hygrodiplosis vaccinii (Kieer, 1897)
MASSALONGIA Kieer, 1897
Massalongia rubra (Kieer, 1890)
RESSELIELLA Seitner, 1906
= omasiniana Strand, 1927
Resseliella ribis (Marikovskij, 1956)
Resseliella theobaldi (Barnes, 1927)
Checklist of the Cecidomyiidae (Diptera) of Finland 115
XYLODIPLOSIS Kieer, 1894
Xylodiplosis nigritarsis (Zetterstedt, 1850)
supertribe Lasiopteridi Rübsaamen & Hedicke, 1925
tribe Dasineurini Rübsaamen & Hedicke, 1925
DASINEURA Rondani, 1840
Dasineura anis (Kieer, 1886)
Dasineura alopecuri (Reuter, 1895)
Dasineura angelicae Rübsaamen, 1916
Dasineura cardaminis (Winnertz, 1853)
Dasineura engstfeldi (Rübsaamen, 1889)
Dasineura epilobii (F. Löw, 1889)
Dasineura fraxinea Kieer, 1907
Dasineura fraxini (Bremi, 1847)
Dasineura galiicola (F. Löw, 1880)
Dasineura gentneri Pritchard, 1953
Dasineura hygrophila (Mik, 1883)
Dasineura hyperici (Bremi, 1847)
Dasineura jaapi Rübsaamen, 1914
Dasineura kieeriana (Rübsaamen, 1891)
Dasineura leguminicola (Lintner, 1879)
Dasineura lotharingiae (Kieer, 1888)
Dasineura mali (Kieer, 1904)
Dasineura napi (Loew, 1850)
= brassicae (Winnertz, 1853)
Dasineura populeti (Rübsaamen, 1889)
Dasineura pteridicola (Kieer, 1901)
Dasineura pteridis (Muller, 1871)
= licina (Kieer, 1889)
Dasineura pustulans (Rübsaamen, 1889)
Dasineura pyri (Bouché, 1847)
Dasineura ribis Barnes, 1940
Dasineura rosae (Bremi, 1847)
= Wachtliella rosarum (Hardy, 1854)
Dasineura similis (F. Löw, 1888)
Dasineura sisymbrii (Schrank, 1803)
Dasineura tetensi (Rübsaamen, 1892)
Dasineura thomasiana (Kieer, 1888)
Dasineura tiliae (Schrank, 1803)
= tiliamvolvens (Rübsaamen, 1889)
Dasineura trifolii (F. Löw, 1874)
Dasineura ulmaria (Bremi, 1847)
Dasineura urticae (Perris, 1840)
Dasineura viciae (Kieer, 1888)
Mathias Jaschhof et al. / ZooKeys 441: 103–118 (2014)
116
Dasineura violae (F. Löw, 1880)
GEOCRYPTA Kieer, 1913
Geocrypta galii (Loew, 1850)
GEPHYRAULUS Rübsaamen, 1915
Gephyraulus raphanistri (Kieer, 1886)
GIRAUDIELLA Rübsaamen, 1915
Giraudiella inclusa (Frauenfeld, 1862)
JAAPIELLA Rübsaamen, 1915
Jaapiella loticola (Rübsaamen, 1889)
Jaapiella veronicae (Vallot, 1827)
KALTENBACHIOLA Hedicke, 1938
Kaltenbachiola strobi (Winnertz, 1853)
MACROLABIS Kieer, 1892
Macrolabis cirsii (Rübsaamen, 1890)
Macrolabis orobi (F. Löw, 1877)
RABDOPHAGA Westwood, 1847
Rabdophaga dubiosa Kieer, 1913
Rabdophaga gemmicola (Kieer, 1896)
Rabdophaga heterobia (Loew, 1850)
Rabdophaga iteobia (Kieer, 1890)
Rabdophaga marginemtorquens (Bremi, 1847)
Rabdophaga nervorum (Kieer, 1895)
Rabdophaga rosaria (Loew, 1850)
Rabdophaga salicis (Schrank, 1803)
Rabdophaga terminalis (Loew, 1850)
WACHTLIELLA Rübsaamen, 1915
Wachtliella caricis (Loew, 1850)
= riparia (Winnertz, 1853)
= muricatae (Meade, 1886)
Wachtliella persicariae (Linnaeus, 1767)
tribe Lasiopterini Rübsaamen & Hedicke, 1925
LASIOPTERA Meigen, 1818
Lasioptera populnea Wachtl, 1883
Lasioptera rubi (Schrank, 1803)
tribe Oligotrophini Rübsaamen & Hedicke, 1925
OLIGOTROPHUS Latreille, 1805
Oligotrophus juniperinus (Linnaeus, 1758)
Oligotrophus panteli Kieer, 1898
tribe Poomyini Rübsaamen & Hedicke, 1925
MAYETIOLA Kieer, 1896
Mayetiola avenae (Marchal, 1895)
Mayetiola destructor (Say, 1817)
Mayetiola joannisi Kieer, 1896
tribe Rhopalomyiini Harris, 1966
Checklist of the Cecidomyiidae (Diptera) of Finland 117
RHOPALOMYIA Rübsaamen, 1892
Rhopalomyia chrysanthemi (Ahlberg, 1939)
Rhopalomyia millefolii (Loew, 1850)
Rhopalomyia ptarmicae (Vallot, 1849)
Rhopalomyia tanaceticola (Karsch, 1879)
Rhopalomyia tubifex (Bouché, 1847)
unplaced Lasiopteridi
CYSTIPHORA Kieer, 1892
Cystiphora sanguinea (Bremi, 1847)
= hieracii (F. Löw, 1874)
= pilosellae Kieer, 1892
Cystiphora sonchi (Vallot, 1827)
ITEOMYIA Kieer, 1913
Iteomyia capreae (Winnertz, 1853)
SEMUDOBIA Kieer, 1913
Semudobia betulae (Winnertz, 1853)
Excluded species
Anarete lacteipennis Kieer, 1906. Misidentication.
Camptomyia antennata Felt, 1920. Misidentication.
Camptomyia maxima Mamaev, 1961. Presumptive misidentication; Finnish record
cannot be veried.
Camptomyia piceae Panelius, 1965. Presumptive misidentication; Finnish record can-
not be veried.
Catarete brevinervis (Zetterstedt, 1851). Misidentication.
Cecidomyia aurora Mannerheim, 1823. Nomen dubium.
Epidosis avescens F. Löw, 1874. Nomen dubium.
Note
Anarete sp. 2. is species was described by Jaschhof and Jaschhof (2009) but not
named. Identication of Anarete species is practically impossible using the keys and
descriptions available in the literature.
Acknowledgements
e subfamilies Lestremiinae and Micromyinae were edited by M.J., Winnertzi-
inae and Porricondylinae by M.J. and J.P., and Cecidomyiinae by M.S. e manu-
script was read and commented on by Keith M. Harris, Ripley, UK, and by two
reviewers unknown to us.
Mathias Jaschhof et al. / ZooKeys 441: 103–118 (2014)
118
References
Buhr H (1964) Bestimmungstabellen der Gallen (Zoo- und Phytocecidien) an Panzen Mittel- und
Nordeuropas. Band I. VEB Gustav Fischer Verlag, Jena, xv + 761 pp.
Buhr H (1965) Bestimmungstabellen der Gallen (Zoo- und Phytocecidien) an Panzen Mit-
tel- und Nordeuropas. Band II. VEB Gustav Fischer Verlag, Jena, 763–1572. [25 pls. with
unnumberered facing pages bearing gure legends]
Gagné RJ, Jaschhof M (2014) A Catalog of the Cecidomyiidae (Diptera) of the World. 3rd
Edition. Digital version 2. http://www.ars.usda.gov/SP2UserFiles/Place/12454900/Gag-
ne_2014_World_Cecidomyiidae_Catalog_3rd_Edition.pdf
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48.
Hulden L (2003) Dasineura pustulans (Rübs.) (Cecidomyiidae) discovered in Finland. Sahlbergia
8(2): 100.
Jaschhof M, Jaschhof C (2009) e Wood Midges (Diptera: Cecidomyiidae: Lestremiinae) of
Fennoscandia and Denmark. Studia dipterologica Supplement 18. Ampyx-Verlag, Halle
(Saale), 333 pp.
Jaschhof M, Jaschhof C (2013) e Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with
notes on extralimital species. Studia dipterologica Supplement 20. Ampyx-Verlag, Halle
(Saale), 392 pp.
Penttinen J, Jaschhof M (2009) On the systematics of Sylvenomyia Mamaev & Zaitzev (Diptera,
Cecidomyiidae, “Porricondylinae”), with the description of a new species from Finland.
Zootaxa 2032: 48–54.
Penttinen J, Siitonen J (2005) Five species of log midges (Diptera, Cecidomyiidae: Porricon-
dylinae) new to Finland. Sahlbergia 10(2): 50–53.
Penttinen J, Spungis V (2007) Additions to the Finnish fauna of log midges (Diptera, Cecido-
myiidae: Porricondylinae). Sahlbergia 12(1): 36–42.
Skuhravá M (1997) Family Cecidomyiidae. In: Papp L, Darvas B (Eds) Contributions to a
Manual of Palaearctic Diptera (with Special Reference to Flies of Economic Importance).
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Checklist of the fungus gnats of Finland: Bolitophilidae, Diadocidiidae, Ditomyiidae... 119
Checklist of the fungus gnats of Finland:
Bolitophilidae, Diadocidiidae, Ditomyiidae,
Keroplatidae and Mycetophilidae (Diptera)
Jevgeni Jakovlev1
1 Finnish Environment Institute, Helsinki, Finland
Corresponding author: Jevgeni Jakovlev (jevgeni.jakovlev@ymparisto.)
Academic editor: J. Kahanpää|Received 4 April 2014|Accepted 30 May 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Jakovlev J (2014) Checklist of the fungus gnats of Finland: Bolitophilidae, Diadocidiidae, Ditomyiidae,
Keroplatidae and Mycetophilidae (Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland.
ZooKeys 441: 119–149. doi: 10.3897/zookeys.441.7646
Abstract
A checklist of the ’fungus gnat’ group of families in Sciaroidea: Bolitophilidae, Diadocidiidae, Ditomyi-
idae, Keroplatidae and Mycetophilidae (Diptera) recorded from Finland. At present, the known Finnish
fauna comprises 767 recognized species in 78 genera; 21 species of the family Bolitophilidae, 5 of the
family Diadocidiidae, 2 of the family Ditomyiidae, 47 of the family Keroplatidae and 692 of the family
Mycetophilidae. e unassigned genus Sciarosoma Chandler, 2002 is not included in this paper because
it is listed with Sciaridae.
e most relevant synonyms, on which species were ever listed from Finland in Hackman (1980) and
subsequent authors are indicated in the checklist. Eighteen species names of wrong records were removed.
Keywords
Species list, Finland, Diptera, biodiversity, faunistics
Introduction
Fungus gnats are a rich assemblage of nematocerous ies (Diptera, Nematocera) placed
in the infraorder Bibionomorpha and superfamily Sciaroidea, currently divided into
eight families (Blagoderov and Grimaldi 2004). Five of these families (Bolitophilidae,
Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae) are called informally
ZooKeys 441: 119–149 (2014)
doi: 10.3897/zookeys.441.7646
www.zookeys.org
Copyright Jevgeni Jakovlev. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jevgeni Jakovlev / ZooKeys 441: 119–149 (2014)
120
‘fungus gnats’ by most European authors. Species in these families are morphologically
closely related and the group is also ecologically uniform.
Adults of fungus gnats are generally quite easily recognized by their habitus. ey
are slender to moderately robust ies between 2 and 10 mm (a few up to 15 mm) in
body length, with long antenna, a high and often strongly arched thorax made dis-
tinctly higher by the long, stout and tight-sitting coxae of their legs. eir wings have
reduced and variable venation that is quite distinctive for the dierent families and
much used also to classify genera. eir body and legs are often covered with strong
bristles. As larvae the majority of fungus gnat species are mycetophagous, associated
with fungi, either fruiting bodies or mycelia in dead wood and soil litter.
Here I cover all the Finnish fungus gnat fauna of the families Bolitophilidae, Dia-
docidiidae, Ditomyiidae, Keroplatidae, and Mycetophilidae. In Europe, the superfamily
Sciaroidea further includes two large families, the Sciaridae – black-winged fungus gnats
and the Cecidomyiidae – gall midges (Søli et al. 2000) although some authors, e.g. Hippa
and Vilkamaa (2006) consider the Cecidomyiidae as the sister group of all other Sciar-
oidea. ey dier markedly in habitus and larval biology and are presented elsewhere.
Finally, the unassigned genus Sciarosoma Chandler, 2002 is not included in this checklist
since it is listed with the family Sciaridae according Hippa and Vilkamaa (2006), but see
also opinion by Jaschhof et al. (2006).
Systematics and nomenclature of the enumeration follow the online checklist of
Nordic fungus gnats (Kjærandsen 2012). Only the most relevant synonyms, on which
species were ever listed from Finland, are included in the checklist.
Adults of the North European species can largely be identied using Hutson et al.
(1980) and Zaitzev (1994) − Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplati-
dae, and Mycetophilidae in part (Mycomyiinae, Sciophilinae, Gnoristinae and Leii-
nae), and Zaitzev (2003) – Mycetophilidae in part (Mycetophilinae and Manotinae).
However, since many new taxa are added, and corrections, synonyms and revisions
continually published, one needs a complete set of the fragmented taxonomic litera-
ture to cover all the species. A good source for obtaining most important literature on
the Sciaroidea is the ViBrant/Scratchpad module Fungus Gnats Online (2014). Spe-
cies identication of fungus gnats based on examination of larvae is currently impos-
sible because the larvae seemingly lack unique morphological characteristics and search
for species-specic molecular markers are yet to be implemented.
Fungus gnats are distributed all over the world with about 5,300 known species
(Table 1), more than 1,450 of them are known from the Palaearctic region (Søli et al.
2000). According to the current version of the Fauna Europaea Web-database (Chan-
dler 2004), 1,098 species of fungus gnats had then been recorded from Europe. Taking
into account literature published subsequently these gures could be increased in ca 100
additional species, i.e. some 1,200 species are known in Europe at present. As opposed
to many other insect groups, fungus gnats, at least Mycetophilidae, seem to display an
increasing diversity towards the north in Europe, and in the European perspective, they
are especially diverse in the boreal zone. e current version of the checklist of North
European (Sweden, Norway, Finland, Russian Karelia, Russian Lapland, Denmark,
e Faroes, and Iceland) fungus gnats (Kjærandsen 2012) includes 896 species.
Checklist of the fungus gnats of Finland: Bolitophilidae, Diadocidiidae, Ditomyiidae... 121
In Finland, 269 species of the fungus gnats (of which ca. 40 currently recognized spe-
cies were described by Carl Lundström) were rst listed in the Enumeration Insectorum
Fenniae (Frey et al. 1941). Hackman (1980) presented the second checklist of Finnish
Diptera that incorporates 486 species of fungus gnats including ve with a question mark.
is list of species, with minor additions and corrections concerning Mycetophilidae af-
ter systematical revisions of the genera Trichonta (Gagné 1981) and Mycomya (Väisänen
1984) in the Holarctic region was included into Catalogue of the Palaearctic Diptera
(Hackman et al. 1988). During the recent two decades, large-scale inventories (e.g. Polevoi
and Jakovlev 2004, Polevoi et al. 2006, Jakovlev et al. 2006, 2014, Jakovlev and Penttinen
2007) have increased the total number of Finnish fungus gnats with ca. 300 species. e
2010 Finnish Red List assessment of fungus gnats was based on a regional species pool
consisting on 734 species (Penttinen et al. 2010). At present, the Finnish fungus gnat
fauna incorporates 767 recognized species (except Sciarosoma) and ca. 20 species awaiting
descriptions. Altogether, 18 species names of wrong records from Finland were removed
from the checklist. ese species and 3 doubtful species names are listed in the end.
Checklist
Nematocera Dumeril, 1805
infraorder Bibionomorpha Hennig, 1954
superfamily Sciaroidea Schiner, 1864
BOLITOPHILIDAE Winnertz, 1864
BOLITOPHILA Meigen, 1818
sg. Bolitophila Meigen, 1818
Bolitophila austriaca (Mayer, 1950)
Bolitophila basicornis (Mayer, 1951)
Bolitophila caspersi Plassmann, 1956
Bolitophila cinerea Meigen, 1818
Bolitophila saundersii (Curtis, 1836)
Bolitophila tenella Winnertz, 1864
sg. Cliopisa Enderlein, 1936
Bolitophila aperta Lundström, 1915
Table 1. Number of species by family.
Family
Number of species (except Sciarosoma) in Level of
knowledge
World
(Pape et al. 2011)
Europe
(Chandler 2004)
Nordic Region
(Kjærandsen 2012) Finland
Bolitophilidae 59 36 31 21 average
Diadocidiidae 34 5 5 5 average
Ditomyiidae 94 4 3 2 average
Keroplatidae 953 110 62 47 average
Mycetophilidae 4150 942 794 692 average
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122
Bolitophila bimaculata Zetterstedt, 1838
Bolitophila dubia Siebke, 1863
= disjuncta Loew, 1869
Bolitophila fumida Edwards, 1941
Bolitophila glabrata Loew, 1869
Bolitophila hybrida (Meigen, 1804)
Bolitophila ingrica Stackelberg, 1969
Bolitophila limitis Polevoi, 1996
Bolitophila maculipennis Walker, 1836
= coronata Mayer, 1951
Bolitophila modesta Lackschewitz, 1937
= tarsata Mayer, 1951
Bolitophila nigrolineata Landrock, 1912
Bolitophila obscurior Stackelberg, 1969
Bolitophila occlusa Edwards, 1913
Bolitophila pseudohybrida Landrock, 1912
Bolitophila rossica Landrock, 1912
DIADOCIDIIDAE Winnertz, 1864
DIADOCIDIA Ruthe, 1831
sg. Adidocidia Laštovka & Matile 1972
Diadocidia trispinosa Polevoi, 1996
Diadocidia valida Mik, 1874
sg. Diadocidia Ruthe, 1831
Diadocidia ferruginosa (Meigen, 1830)
Diadocidia ssa Zaitzev, 1994
Diadocidia spinosula Tollet, 1948
DITOMYIIDAE Keilin, 1919
SYMMERUS Walker, 1848
Symmerus annulatus (Meigen, 1830)
Symmerus nobilis Lackschewitz, 1937
KEROPLATIDAE Rondani, 1856
KEROPLATINAE Rondani, 1856
tribe Keroplatini Rondani, 1856
CEROTELION Rondani, 1856
Cerotelion striatum (Gmelin, 1790)
= lineatum (Fabricius, 1775) preocc.
KEROPLATUS Bosc, 1792
Keroplatus testaceus Dalman, 1818
Keroplatus tipuloides Bosc, 1792
= sesioides Wahlberg, 1839
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* Keroplatus tuvensis Zaitzev, 1991 (see Notes)
ROCETELION Matile, 1988
Rocetelion humerale (Zetterstedt, 1850)
tribe Orfeliini Matile, 1990
ISONEUROMYIA Brunetti, 1912
Isoneuromyia semirufa (Meigen, 1818)
MACRORRHYNCHA Winnertz, 1846
Macrorrhyncha ava Winnertz, 1846
Macrorrhyncha rostrata (Zetterstedt, 1851)
MONOCENTROTA Edwards, 1925
Monocentrota lundstroemi Edwards, 1925
NEOPLATYURA Malloch, 1928
Neoplatyura ava (Macquart, 1826)
Neoplatyura modesta (Winnertz, 1864)
ORFELIA Costa, 1857
Orfelia discoloria (Meigen, 1818)
Orfelia falcata Zaitzev, 1994
Orfelia fasciata (Meigen, 1804)
Orfelia krivosheinae Zaitzev, 1994
Orfelia lugubris (Zetterstedt, 1851)
= tristis (Lundström, 1911)
Orfelia nemoralis (Meigen, 1818)
Orfelia nigricornis (Fabricius, 1805)
Orfelia ochracea (Meigen, 1818)
= unicolor (Staeger, 1840)
Orfelia pallida (Staeger, 1840)
PYRATULA Edwards, 1929
Pyratula perpusilla (Edwards, 1913)
Pyratula subcanariae Chandler, 2001
Pyratula zonata (Zetterstedt, 1855)
URYTALPA Edwards, 1929
Urytalpa atriceps (Edwards, 1913)
Urytalpa dorsalis (Staeger, 1840)
= ochracea (Meigen, 1818)
Urytalpa galdes Hedmark & Kjærandsen, 2009
Urytalpa macrocera (Edwards, 1913)
Urytalpa trivittata (Lundström, 1914)
MACROCERINAE Rondani, 1856
tribe Macrocerini Rondani, 1856
MACROCERA Meigen, 1803
Macrocera angulata Meigen, 1818
Macrocera centralis Meigen, 1818
Macrocera crassicornis Winnertz, 1864
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124
Macrocera estonica Landrock, 1924
Macrocera fasciata Meigen, 1804
Macrocera fascipennis Staeger, 1840
Macrocera grandis Lundström, 1912
= magna Landrock, 1917
Macrocera lutea Meigen, 1804
Macrocera maculata Meigen, 1818
Macrocera nigricoxa Winnertz, 1864
Macrocera parva Lundström, 1914
Macrocera phalerata Meigen, 1818
Macrocera pilosa Landrock, 1917
Macrocera pumilio Loew, 1869
Macrocera pusilla Meigen, 1830
Macrocera stigma Curtis, 1837
Macrocera stigmoides Edwards, 1925
Macrocera vittata Meigen, 1830
Macrocera zetterstedti Lundström, 1914
= nana Zetterstedt, 1860
MYCETOPHILIDAE Newman, 1834
MYCOMYINAE Edwards, 1925
MYCOMYA Rondani, 1856
sg. Calomycomya Väisänen, 1984
Mycomya avala Väisänen, 1984
Mycomya pulchella (Dziedzicki, 1885)
sg. Coheromyia Väisänen, 1984
Mycomya branderi Väisänen, 1984
= tantilla auct. nec (Loew, 1869) misid.
sg. Cymomya Väisänen, 1984
Mycomya circumdata (Staeger, 1840)
sg. Lycomya Väisänen, 1984
Mycomya pectinifera Edwards, 1924
sg. Mycomya Rondani, 1856
Mycomya annulata (Meigen, 1818)
= incisurata (Zetterstedt, 1838)
Mycomya bialorussica Landrock, 1925
Mycomya bicolor (Dziedzicki, 1885)
Mycomya bisulca Lackschewitz, 1937
Mycomya britteni Kidd, 1955
Mycomya brunnea (Dziedzicki, 1885)
Mycomya cinerascens (Macquart, 1826)
= hyalinata (Meigen, 1830)
Mycomya collini Edwards, 1941
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Mycomya danielae Matile, 1972
Mycomya denmax Väisänen, 1979
Mycomya digitifera Edwards, 1925
Mycomya disa Väisänen, 1984
Mycomya dziedzickii Väisänen, 1984
= fulva (Dziedzicki, 1885) preocc.
Mycomya egregia (Dziedzicki, 1885)
Mycomya festivalis Väisänen, 1984
Mycomya avicollis (Zetterstedt, 1852)
Mycomya forestaria Plassmann, 1978
Mycomya fuscata (Winnertz, 1864)
Mycomya griseovittata (Zetterstedt, 1852)
= fasciata (Zetterstedt, 1838) preocc.
= clavigera (Lundström, 1912)
Mycomya hackmani Väisänen, 1984
Mycomya heydeni Plassmann, 1970
Mycomya hians (Lundström, 1912)
Mycomya hiisi Väisänen, 1979
Mycomya humida Garrett, 1924
= curvata Fisher, 1937
Mycomya indistincta Polevoi, 1995
Mycomya insignis (Winnertz, 1864)
= wrzesniowskii (Dziedzicki, 1885)
Mycomya islandica Väisänen, 1984
Mycomya karelica Väisänen, 1979
Mycomya kaurii Väisänen, 1979
Mycomya kuusamoensis Väisänen, 1979
Mycomya lambi Edwards, 1941
Mycomya leporina Väisänen, 1984
Mycomya levis (Dziedzicki, 1885)
Mycomya livida (Dziedzicki, 1885)
Mycomya maculata (Meigen, 1804)
Mycomya marginata (Meigen, 1818)
= limbata (Winnertz, 1864)
Mycomya mituda Väisänen, 1980
Mycomya neohyalinata Väisänen, 1984
Mycomya nigricornis (Zetterstedt, 1852)
= melanoceras Edwards, 1924
Mycomya nitida (Zetterstedt, 1852)
= exigua (Winnertz, 1864)
Mycomya norna Väisänen, 1984
Mycomya occultans (Winnertz, 1864)
Mycomya ornata (Meigen, 1818)
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126
Mycomya parva (Dziedzicki, 1885)
Mycomya prominens (Lundström, 1913)
Mycomya pseudoapicalis Landrock, 1925
Mycomya pseudocurvata Väisänen, 1979
Mycomya punctata (Meigen, 1804)
Mycomya rucollis (Zetterstedt, 1852)
Mycomya safena Väisänen, 1984
Mycomya shermani Garrett, 1924
= kingi Edwards, 1941
Mycomya shewelli Väisänen, 1984
Mycomya siebecki (Landrock, 1912)
Mycomya sieberti (Landrock, 1930)
Mycomya sigma Johannsen, 1910
= duplicata Edwards, 1925
Mycomya simulans Väisänen, 1984
Mycomya spinicoxa Väisänen, 1979
Mycomya storai Väisänen, 1979
Mycomya subarctica Väisänen, 1979
Mycomya tenuis (Walker, 1856)
Mycomya thula Väisänen, 1984
Mycomya trivittata (Zetterstedt, 1838)
Mycomya tumida (Winnertz, 1864)
Mycomya vittiventris (Zetterstedt, 1852)
= elegans (Lundström, 1912)
Mycomya wankowiczii (Dziedzicki, 1885)
Mycomya winnertzi (Dziedzicki, 1885)
sg. Mycomyopsis Väisänen, 1984
Mycomya anis (Staeger, 1840)
= ava (Winnertz, 1864)
Mycomya confusa Väisänen, 1977
Mycomya fennica Väisänen, 1979
Mycomya maura (Walker, 1856)
Mycomya neolittoralis Väisänen, 1984
Mycomya paradentata Väisänen, 1984
= dentata auct. nec Fisher, 1937 misid.
Mycomya penicillata (Dziedzicki, 1885)
Mycomya permixta Väisänen, 1984
Mycomya trilineata (Zetterstedt, 1838)
= fusca (Meigen, 1818)
sg. Neomycomya Väisänen, 1984
Mycomya mbriata (Meigen, 1818)
NEOEMPHERIA Osten Sacken, 1878
Neoempheria bimaculata (von Roser, 1840)
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Neoempheria pictipennis (Haliday, 1833)
Neoempheria striata (Meigen, 1818)
Neoempheria tuomikoskii Väisänen, 1982
Neoempheria winnertzi Edwards, 1913
SCIOPHILINAE Rondani, 1840
ACNEMIA Winnertz, 1864
Acnemia amoena Winnertz, 1864
Acnemia angusta Zaitzev, 1982
Acnemia falcata Zaitzev, 1982
Acnemia longipes Winnertz, 1864
Acnemia nitidicollis (Meigen, 1818)
Acnemia trida Zaitzev, 1982
ALLOCOTOCERA Mik, 1886
Allocotocera pulchella (Curtis, 1837)
ANACLILEIA Meunier, 1904
Anaclileia dispar (Winnertz, 1864)
Anaclileia dziedzickii (Landrock, 1911)
AZANA Walker, 1856
sg. Azana Walker, 1856
Azana anomala (Staeger, 1840)
COELOPHTHINIA Edwards, 1941
Coelophthinia thoracica (Winnertz, 1864)
EUDICRANA Loew, 1869
Eudicrana nigriceps (Lundström, 1909)
LEPTOMORPHUS Curtis, 1831
sg. Leptomorphus Curtis, 1831
Leptomorphus forcipatus Landrock, 1918
Leptomorphus subforcipatus Zaitzev & Ševčík, 2002
= quadrimaculatus auct. nec Matsumura, 1910 misid. (see Notes)
Leptomorphus walkeri Curtis, 1831
MEGALOPELMA Enderlein, 1911
Megalopelma nigroclavatum (Strobl, 1910)
MONOCLONA Mik, 1886
Monoclona braueri (Strobl, 1895)
Monoclona rulatera (Walker, 1837)
Monoclona silvatica Zaitzev, 1983
= mikii Kertesz,1898 misid. (see Notes)
NEURATELIA Rondani, 1856
Neuratelia nemoralis (Meigen, 1818)
Neuratelia sintenisi Lackschewitz, 1937
PARATINIA Mik, 1874
Paratinia sciarina Mik, 1874
PHTHINIA Winnertz, 1864
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128
Phthinia congenita Plassmann, 1984
Phthinia humilis Winnertz, 1864
Phthinia mira (Ostroverkhova, 1977)
= plassmanni Caspers, 1984
Phthinia setosa Zaitzev, 1994
Phthinia winnertzi Mik, 1869
POLYLEPTA Winnertz, 1864
Polylepta borealis Lundström, 1912
Polylepta guttiventris (Zetterstedt, 1852)
SCIOPHILA Meigen, 1818
Sciophila adamsi Edwards, 1925
Sciophila antiqua Chandler, 1987
= hebes auct. nec Johannsen, 1910 misid. (see Notes)
Sciophila bicuspidata Zaitzev, 1982
Sciophila buxtoni Freeman, 1956
Sciophila caesarea Chandler, 2001
Sciophila dziedzickii Edwards, 1925
Sciophila fenestella Curtis, 1837
Sciophila fuliginosa Holmgren, 1883
Sciophila geniculata Zetterstedt, 1838
Sciophila hirta Meigen, 1818
Sciophila karelica Zaitzev, 1982
Sciophila krysheni Polevoi, 2001
Sciophila limbatella Zetterstedt, 1852
Sciophila lutea Macquart, 1826
Sciophila minuta Zaitzev, 1982
Sciophila modesta Zaitzev, 1982
Sciophila nigronitida Landrock, 1925
Sciophila nonnisilva Hutson, 1979
Sciophila persubtilis Polevoi, 2001
Sciophila plurisetosa Edwards, 1921
Sciophila pomacea Chandler, 2006
= ochracea Stephens in Walker, 1856 preocc.
Sciophila pseudoexuosa Kurina, 1991
Sciophila rufa Meigen, 1830
Sciophila salassea Matile, 1983
Sciophila setosa Garrett, 1925
Sciophila thoracica Staeger, 1840
= quadriterga (Hutson, 1979)
Sciophila varia (Winnertz, 1864)
Sciophila yakutica Blagoderov, 1992
GNORISTINAE Edwards, 1925
ACOMOPTERA Vockeroth, 1980
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Acomoptera dicilis (Dziedzicki 1885)
Acomoptera spinistylus (Søli, 1993)
APOLEPHTHISA Grzegorzek, 1885
Apolephthisa subincana (Curtis, 1837)
BOLETINA Staeger, 1840
Boletina atridentata Polevoi & Hedmark, 2004
Boletina basalis (Meigen, 1818)
Boletina bidenticulata Sasakawa & Kimura, 1974
Boletina borealis Zetterstedt, 1852
Boletina brevicornis Zetterstedt, 1852
Boletina cincticornis (Walker, 1848)
Boletina cordata Polevoi & Hedmark, 2004
Boletina cornuta Zaitzev, 1994
Boletina digitata Lundström, 1914
Boletina dispecta Dziedzicki, 1885
Boletina dispectoides Jakovlev & Penttinen, 2007
Boletina dissipata Plassmann, 1986
Boletina dubia (Meigen, 1804)
Boletina edwardsi Chandler, 1992
Boletina falcata Polevoi & Hedmark, 2004
Boletina fennoscandica Polevoi & Hedmark, 2004
Boletina gripha Dziedzicki, 1885
Boletina griphoides Edwards, 1925
Boletina groenlandica Staeger, 1845
Boletina gusakovae Zaitzev, 1994
Boletina hedstroemi Polevoi & Hedmark, 2004
Boletina intermedia Lundström, 1915
Boletina jamalensis Zaitzev, 1994
= struthioides Polevoi & Hedmark, 2004
Boletina kivachiana Polevoi & Hedmark, 2004
Boletina kurilensis Zaitzev, 1994
Boletina landrocki Edwards, 1924
Boletina lapponica Polevoi & Hedmark, 2004
Boletina lundbecki Lundström, 1912
Boletina lundstroemi Landrock, 1912
Boletina maculata Holmgren, 1870
= apicalis (Walker, 1848) preocc.
Boletina minuta Polevoi in Zaitzev & Polevoi, 1995
Boletina moravica Landrock, 1912
Boletina nasuta (Haliday, 1839)
Boletina nigricans Dziedzicki, 1885
Boletina nigricoxa Staeger, 1840
Boletina nigrofusca Dziedzicki, 1885
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130
Boletina nitida Grzegorzek, 1885
Boletina nitiduloides Zaitzev, 1994
Boletina onegensis Polevoi in Zaitzev & Polevoi, 1995
Boletina pallidula Edwards, 1925
Boletina palmata Polevoi, 2013
Boletina pectinunguis Edwards, 1932
Boletina pinusia Maximova, 2001
Boletina plana (Walker, 1856)
Boletina polaris Lundström, 1915
Boletina populina Polevoi in Zaitzev & Polevoi, 1995
Boletina pseudonitida Zaitzev, 1994
Boletina rejecta Edwards, 1941
Boletina sciarina Staeger, 1840
Boletina silvatica Dziedzicki, 1885
Boletina subtriangularis Polevoi & Hedmark, 2004
Boletina takagii Sasakawa & Kimura, 1974
Boletina tiroliensis Plassmann, 1980
Boletina triangularis Polevoi in Zaitzev & Polevoi, 1995
Boletina trispinosa Edwards, 1913
Boletina trivittata (Meigen, 1818)
Boletina verticillata Stackelberg, 1943
Boletina villosa Landrock, 1912
COELOSIA Winnertz, 1864
Coelosia bicornis Stackelberg, 1946
Coelosia ava (Staeger, 1840)
Coelosia fusca Bezzi, 1892
= silvatica Landrock, 1918
Coelosia gracilis Johannsen, 1912
Coelosia limpida Plassmann, 1986
Coelosia tenella (Zetterstedt, 1852)
Coelosia truncata Lundström, 1909
DZIEDZICKIA Johannsen, 1909
Dziedzickia marginata (Dziedzicki, 1885)
ECTREPESTHONEURA Enderlein, 1911
Ectrepesthoneura colyeri Chandler, 1980
Ectrepesthoneura hirta (Winnertz, 1846)
Ectrepesthoneura nigra Zaitzev, 1984
Ectrepesthoneura ovata Ostroverkhova, 1977
= bucera Plassmann, 1980
Ectrepesthoneura pubescens (Zetterstedt, 1860)
Ectrepesthoneura referta Plassmann, 1976
Ectrepesthoneura tori Zaitzev & Økland, 1994
GNORISTE Meigen, 1818
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Gnoriste apicalis Meigen, 1818
Gnoriste bilineata Zetterstedt, 1852
GRZEGORZEKIA Edwards, 1941
Grzegorzekia collaris (Meigen, 1818)
HADRONEURA Lundström, 1906
Hadroneura palmeni Lundström, 1906
IMPLETA Plassmann, 1978
Impleta consorta Plassmann, 1978
KATATOPYGIA Martinsson & Kjærandsen, 2012
Katatopygia erythropyga (Holmgren, 1883)
Katatopygia sahlbergi (Lundström, 1906)
PALAEODOCOSIA Meunier, 1904
Palaeodocosia vittata (Coquillett, 1901)
= janickii (Dziedzicki, 1923)
= alpicola auct. nec (Strobl, 1894) misid. (see Notes)
SAIGUSAIA Vockeroth, 1980
Saigusaia aviventris (Strobl, 1894)
SPEOLEPTA Edwards, 1925
Speolepta leptogaster (Winnertz, 1864)
SYNAPHA Meigen, 1818
Synapha fasciata Meigen, 1818
Synapha vitripennis (Meigen, 1818)
SYNTEMNA Winnertz, 1864
Syntemna daisetsuzana Okada, 1938
Syntemna elegantia Plassmann, 1978
Syntemna hungarica (Lundström, 1912)
Syntemna morosa Winnertz, 1864
Syntemna nitidula Edwards, 1925
Syntemna oulankaensis Polevoi, 2003
Syntemna penicilla Hutson, 1979
Syntemna relicta (Lundström, 1912)
Syntemna setigera (Lundström, 1914)
= haagvari Økland, 1995
Syntemna stylata Hutson, 1979
Syntemna stylatoides Zaitzev, 1994
TETRAGONEURA Winnertz, 1846
Tetragoneura ambigua Grzegorzek, 1885
Tetragoneura obirata Plassmann, 1990
Tetragoneura pudogensis Polevoi & Jakovlev, 2011
Tetragoneura ruuhijarvii Polevoi & Jakovlev, 2011
Tetragoneura sylvatica (Curtis, 1837)
LEIINAE Edwards, 1925
CLASTOBASIS Skuse, 1890
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Clastobasis alternans (Winnertz, 1864)
DOCOSIA Winnertz, 1864
Docosia expectata Laštovka & Ševčík, 2006
Docosia avicoxa Strobl, 1900
= pallipes Edwards, 1941
Docosia gilvipes (Haliday in Walker, 1856)
Docosia landrocki Laštovka & Ševčík, 2006
Docosia muelleri Plassmann, 1986
Docosia tibialis Laštovka & Ševčík, 2006
GREENOMYIA Brunetti, 1912
Greenomyia baikalica Zaitzev, 1994
Greenomyia borealis (Winnertz, 1864)
Greenomyia mongolica Laštovka & Matile, 1974
LEIA Meigen, 1818
Leia bilineata (Winnertz, 1864)
Leia bimaculata (Meigen, 1804)
Leia crucigera Zetterstedt, 1838
Leia cylindrica (Winnertz, 1864)
Leia fascipennis Meigen, 1818
Leia longiseta Barendrecht, 1938
Leia picta Meigen, 1818
Leia subfasciata (Meigen, 1818)
Leia winthemii Lehmann, 1822
RONDANIELLA Johannsen, 1909
Rondaniella dimidiata (Meigen, 1804)
MANOTINAE Edwards, 1925
MANOTA Williston, 1896
Manota unifurcata Lundström, 1913
MYCETOPHILINAE Newman, 1834
tribe Exechiini Edwards, 1925
ALLODIA Winnertz, 1864
sg. Allodia Winnertz, 1864
Allodia anglofennica Edwards, 1921
Allodia confusa Zaitzev, 2003
= simplex Zaitzev, 1983 preocc.
Allodia embla Hackman, 1971
Allodia lugens (Wiedemann, 1817)
Allodia lundstroemi Edwards, 1921
Allodia ornaticollis (Meigen, 1818)
Allodia pyxidiiformis Zaitzev, 1983
Allodia septentrionalis Hackman, 1971
Allodia truncata Edwards, 1921
Allodia tuomikoskii Hackman, 1971
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Allodia zaitzevi Kurina, 1998
sg. Brachycampta Winnertz, 1864
Allodia adunca Zaitzev, 1992
Allodia alternans (Zetterstedt, 1838)
Allodia angulata (Lundström, 1913)
Allodia barbata (Lundström, 1909)
Allodia bohemica Ševčík, 2004
Allodia czernyi (Landrock, 1912)
Allodia foliifera (Strobl, 1910)
Allodia grata (Meigen, 1830)
Allodia huggerti Kjærandsen, 2007
Allodia neglecta Edwards, 1925
Allodia penicillata (Lundström, 1912)
Allodia pistillata (Lundström, 1911)
Allodia protenta Laštovka & Matile, 1974
= mendli Plassmann, 1977
Allodia silvatica (Landrock, 1912)
Allodia subpistillata Ševčík, 1999
ALLODIOPSIS Tuomikoski, 1966
Allodiopsis domestica (Meigen, 1830)
Allodiopsis gracai Ševčík & Papp 2003
Allodiopsis korolevi Zaitzev, 1982
Allodiopsis pseudodomestica (Lackschewitz, 1937)
Allodiopsis rustica (Edwards, 1941)
ANATELLA Winnertz, 1864
Anatella ankeli Plassmann, 1977
Anatella aquila Zaitzev, 1989
Anatella bremia Chandler, 1994
Anatella ciliata Winnertz, 1864
Anatella crispa Zaitzev, 1994
Anatella damp Landrock, 1924
Anatella dentata Zaitzev, 1989
Anatella emergens Caspers, 1987
Anatella avicauda Winnertz, 1864
Anatella avomaculata Edwards, 1925
Anatella gibba Winnertz, 1864
Anatella lenis Dziedzicki, 1923
Anatella maritima Ostroverkhova, 1979
Anatella minuta (Staeger, 1840)
Anatella novata Dziedzicki, 1923
Anatella setigera Edwards, 1921
Anatella simpatica Dziedzicki, 1923
Anatella turi Dziedzicki, 1923
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Anatella unguigera Edwards, 1921
BRACHYPEZA Winnertz, 1864
sg. Brachypeza Winnertz, 1864
Brachypeza armata Winnertz, 1864
Brachypeza bisignata Winnertz, 1864
= hilaris Winnertz, 1864 preocc.
sg. Paracordyla Tuomikoski, 1966
Brachypeza obscura Winnertz, 1864
BREVICORNU Marshall, 1896
Brevicornu arcticoides Caspers, 1985
Brevicornu arcticum (Lundström, 1913)
Brevicornu auriculatum (Edwards, 1925)
Brevicornu beatum (Johannsen, 1912)
Brevicornu bellum (Johannsen, 1912)
Brevicornu bipartitum Laštovka & Matile, 1974
Brevicornu canescens (Zetterstedt, 1852)
= griseolum auct. nec (Zetterstedt, 1852)
Brevicornu cognatum Ostroverkhova, 1979
Brevicornu fasciculatum (Lackschewitz, 1937)
Brevicornu fennicum (Landrock, 1927)
Brevicornu ssicauda (Lundström, 1911)
Brevicornu foliatum (Edwards, 1925)
Brevicornu fuscipenne (Staeger, 1840)
Brevicornu glandis Laštovka & Matile, 1974
Brevicornu griseicolle (Staeger, 1840)
Brevicornu griseolum (Zetterstedt, 1852)
= boreale (Lundström, 1914)
Brevicornu improvisum Zaitzev, 1992
Brevicornu kingi (Edwards, 1925)
Brevicornu luteum (Landrock, 1925)
Brevicornu melanderi Zaitzev, 1988
Brevicornu nigrofuscum (Lundström, 1909)
Brevicornu occidentale Zaitzev, 1988
Brevicornu parafennicum Zaitzev in Zaitzev & Polevoi, 1995
Brevicornu proximum (Staeger, 1840)
Brevicornu rosmellitum Chandler, 2001
Brevicornu rucorne (Meigen, 1838)
Brevicornu serenum (Winnertz, 1864)
Brevicornu sericoma (Meigen, 1830)
Brevicornu setulosum Zaitzev, 1988
Brevicornu spathulatum (Lundström, 1911)
Brevicornu verralli (Edwards, 1925)
CORDYLA Meigen, 1803
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Cordyla brevicornis (Staeger, 1840)
Cordyla crassicornis Meigen, 1818
Cordyla fasciata Meigen, 1830
Cordyla ssa Edwards, 1925
Cordyla aviceps (Staeger, 1840)
Cordyla fusca Meigen, 1804
Cordyla insons Laštovka & Matile, 1974
Cordyla murina Winnertz, 1864
Cordyla nitens Winnertz, 1864
Cordyla nitidula Edwards, 1925
= bergensis (Barendrecht, 1938)
Cordyla parvipalpis Edwards, 1925
Cordyla pusilla Edwards, 1925
= sixi (Barendrecht, 1938)
Cordyla semiava (Staeger, 1840)
EXECHIA Winnertz, 1864
Exechia bicincta (Staeger, 1840)
Exechia borealis Lundström, 1912
Exechia cincta Winnertz, 1864
Exechia connis Winnertz, 1864
Exechia contaminata Winnertz, 1864
Exechia cornuta Lundström, 1914
Exechia dentata Lundström, 1916
Exechia dizona Edwards, 1924
Exechia dorsalis (Staeger, 1840)
Exechia exigua Lundström, 1909
Exechia festiva Winnertz, 1864
Exechia frigida (Boheman, 1865)
Exechia fusca (Meigen, 1804)
Exechia lucidula (Zetterstedt, 1838)
Exechia lundstroemi Landrock, 1923
Exechia macula Chandler, 2001
= maculipennis (Stannius, 1831) preocc.
Exechia micans Laštovka & Matile, 1974
Exechia nigra Edwards, 1925
Exechia nigrofusca Lundström, 1909
Exechia nigroscutellata Landrock, 1912
Exechia papyracea Stackelberg, 1948
Exechia parva Lundström, 1909
Exechia parvula (Zetterstedt, 1852)
= nana Staeger, 1840 preocc.
Exechia pectinivalva Stackelberg, 1948
Exechia pseudocincta Strobl, 1910
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Exechia pseudofestiva Lackschewitz, 1937
Exechia repanda Johannsen, 1912
Exechia repandoides Caspers, 1984
Exechia separata Lundström, 1912
Exechia seriata (Meigen, 1830)
= pallida Stannius, 1831
Exechia similis Laštovka & Matile, 1974
Exechia spinigera Winnertz, 1864
Exechia spinuligera Lundström, 1912
Exechia styriaca Strobl, 1898
= sororcula Lackschewitz, 1937
Exechia subfrigida Laštovka & Matile, 1974
Exechia unifasciata Lackschewitz, 1937
Exechia unimaculata (Zetterstedt, 1860)
EXECHIOPSIS Tuomikoski, 1966
sg. Exechiopsis Tuomikoski, 1966
Exechiopsis aemula Plassmann, 1984
Exechiopsis clypeata (Lundström, 1911)
Exechiopsis distendens (Lackschewitz, 1937)
Exechiopsis dumitrescae (Burghele-Balacesco, 1972)
Exechiopsis mbriata (Lundström, 1909)
Exechiopsis forcipata Lackschewitz, 1937)
Exechiopsis grassatura (Plassmann, 1978)
Exechiopsis hammi (Edwards, 1925)
Exechiopsis indecisa (Walker, 1856)
Exechiopsis ingrica Stackelberg, 1948
Exechiopsis intersecta (Meigen, 1818)
Exechiopsis januarii (Lundström, 1913)
Exechiopsis lackschewitziana (Stackelberg, 1948)
Exechiopsis landrocki (Lundström, 1912)
Exechiopsis ligulata (Lundström, 1913)
Exechiopsis pseudindecisa Laštovka & Matile, 1974
Exechiopsis pseudopulchella (Lundström, 1912)
Exechiopsis pulchella (Winnertz, 1864)
Exechiopsis sagittata Laštovka & Matile, 1974
Exechiopsis subulata (Winnertz, 1864)
sg. Xenexechia Tuomikoski, 1966
Exechiopsis crucigera (Lundström, 1909)
Exechiopsis davatchii Matile, 1969
Exechiopsis leptura (Meigen, 1830)
Exechiopsis membranacea (Lundström, 1912)
Exechiopsis perspicua (Johannsen, 1912)
Exechiopsis pollicata (Edwards, 1925)
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Exechiopsis praedita (Plassmann, 1976)
Exechiopsis seducta (Plassmann, 1976)
Exechiopsis stylata Laštovka & Matile, 1974
MYROSIA Tuomikoski, 1966
Myrosia maculosa (Meigen, 1818)
NOTOLOPHA Tuomikoski, 1966
Notolopha brachycera (Zetterstedt, 1852)
= tuomikoskii (Zaitzev & Maximova, 2000)
Notolopha cristata (Staeger, 1840)
Notolopha sibirica (Zaitzev & Maximova, 2000)
PSEUDEXECHIA Tuomikoski, 1966
Pseudexechia aurivernica Chandler, 1978
Pseudexechia canalicula (Johannsen, 1912)
Pseudexechia parallela (Edwards, 1925)
Pseudexechia pectinacea (Ostroverkhova, 1979)
Pseudexechia tristriata Stackelberg, 1969
Pseudexechia trivittata (Staeger, 1840)
PSEUDOBRACHYPEZA Tuomikoski, 1966
Pseudobrachypeza helvetica (Walker, 1856)
PSEUDORYMOSIA Tuomikoski, 1966
Pseudorymosia fovea (Dziedzicki, 1910)
= optiva auct. nec (Dziedzicki, 1910) misid. (see Notes)
RYMOSIA Winnertz, 1864
Rymosia acta Dziedzicki, 1910
Rymosia anis Winnertz, 1864
= gracilipes Dziedzicki, 1910
Rymosia batava (Barendrecht, 1938)
Rymosia bida Edwards, 1925
Rymosia britteni Edwards, 1925
Rymosia connexa Winnertz, 1864
Rymosia fasciata (Meigen, 1804)
Rymosia fraudatrix Dziedzicki, 1910
? = spiniforceps Matile, 1963
Rymosia guttata Lundström, 1912
Rymosia pinnata Ostroverkhova, 1979
Rymosia placida Winnertz, 1864
Rymosia setiger Dziedzicki, 1910
Rymosia signatipes (van der Wulp, 1859)
= winnertzi Barendrecht, 1938
STIGMATOMERIA Tuomikoski, 1966
Stigmatomeria crassicornis (Stannius, 1831)
SYNPLASTA Skuse, 1890
= GYMNOGONIA Tuomikoski, 1966
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Synplasta bayardi Matile, 1971
Synplasta dulcia (Dziedzicki, 1910)
Synplasta exclusa (Dziedzicki, 1910)
= sintenisi (Lackschewitz, 1937)
Synplasta gracilis Winnertz, 1864
= excogitata auct. nec (Dziedzicki, 1910) misid. (see Notes)
Synplasta ingeniosa (Kidd 1969)
Synplasta praeformida (Dziedzicki, 1910)
Synplasta pseudingeniosa Zaitzev, 1993
Synplasta rulatera (Edwards, 1941)
TARNANIA Tuomikoski, 1966
Tarnania fenestralis (Meigen, 1838)
Tarnania tarnanii (Dziedzicki, 1910)
tribe Mycetophilini Newman, 1834
DYNATOSOMA Winnertz, 1864
Dynatosoma cochleare Strobl, 1895
Dynatosoma dihaeta Polevoi, 1995
= schachti Plassmann, 1999
Dynatosoma fuscicorne (Meigen, 1818)
Dynatosoma majus Landrock, 1912
Dynatosoma nigromaculatum Lundström, 1913
= abdominale auct. nec Staeger, 1840
Dynatosoma nobile Loew, 1873
Dynatosoma reciprocum (Walker, 1848)
Dynatosoma rufescens (Zetterstedt, 1838)
= bukowskii Zaitzev, 1986
= lutescens (Zetterstedt, 1852)
Dynatosoma silesiacum Ševčík, 2001
Dynatosoma thoracicum (Zetterstedt, 1838)
= norwegiense Zaitzev & Økland, 1994
EPICYPTA Winnertz, 1864
Epicypta aterrima (Zetterstedt, 1852)
Epicypta fumigata (Dziedzicki, 1923)
Epicypta limnophila Chandler, 1981
Epicypta scatophora (Perris, 1849)
MYCETOPHILA Meigen, 1803
= FUNGIVORA Meigen, 1800 suppr.
Mycetophila abbreviata Landrock, 1914
Mycetophila abiecta (Laštovka, 1963)
Mycetophila adumbrata Mik, 1884
Mycetophila alea Laoon, 1965
= guttata Dziedzicki, 1884
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Mycetophila attonsa (Laoon, 1957)
= uncta Plassmann, 1999
Mycetophila autumnalis Lundström, 1909
Mycetophila bialorussica Dziedzicki, 1884
* Mycetophila biformis Maximova, 2002 preocc. (see Notes)
Mycetophila biusta Meigen, 1818
Mycetophila blanda Winnertz, 1864
Mycetophila bohemica (Laštovka, 1963)
Mycetophila boreocruciator Ševčík, 2003
Mycetophila brevitarsata (Laštovka, 1963)
Mycetophila caudata Staeger, 1840
Mycetophila cingulum Meigen, 1830
Mycetophila conuens Dziedzicki, 1884
= fulva Winnertz, 1864
Mycetophila confusa Dziedzicki, 1884
= auctata Edwards, 1941
Mycetophila curviseta Lundström, 1911
Mycetophila deexa Chandler, 2001
Mycetophila dentata Lundström, 1915
Mycetophila devioides Bechev, 1988
Mycetophila distigma Meigen, 1830
= w-fuscum Dziedzicki, 1884
Mycetophila dziedzickii Chandler, 1977
= obscura Dziedzicki, 1884 nec Walker, 1848
Mycetophila edwardsi Lundström, 1913
Mycetophila estonica Kurina, 1992
Mycetophila evanida Laštovka, 1972
Mycetophila exstincta Loew, 1869
= mikii Dziedzicki, 1884
Mycetophila nlandica Edwards, 1913
Mycetophila ava Winnertz, 1864
Mycetophila forcipata Lundström, 1913
Mycetophila formosa Lundström, 1911
Mycetophila freyii Lundström, 1909
Mycetophila fungorum (De Geer, 1776)
Mycetophila gibbula Edwards, 1925
Mycetophila hetschkoi Landrock, 1918
Mycetophila ichneumonea Say, 1823
Mycetophila idonea Laštovka, 1972
Mycetophila immaculata (Dziedzicki, 1884)
Mycetophila laeta Walker, 1848
= calva Lundström, 1912
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Mycetophila lapponica Lundström, 1906
Mycetophila lobulata Zaitzev, 1999
Mycetophila lubomirskii Dziedzicki, 1884
Mycetophila luctuosa Meigen, 1830
= modesta Winnertz, 1864
Mycetophila magnicauda Strobl, 1895
Mycetophila marginata Winnertz, 1864
Mycetophila mitis (Johannsen, 1912)
Mycetophila mohilevensis Dziedzicki, 1884
Mycetophila monstera Maximova, 2002
Mycetophila morata Zaitzev, 1999
Mycetophila moravica Landrock, 1925
Mycetophila morosa Winnertz, 1864
Mycetophila nigrofusca Dziedzicki, 1884
Mycetophila occultans Lundström, 1913
= tarsata Winnertz, 1864 preocc.
Mycetophila ocellus Walker, 1848
Mycetophila ornata Stephens, 1846
Mycetophila ostentanea Zaitzev, 1998
Mycetophila pecinai (Laštovka, 1963)
Mycetophila perpallida Chandler, 1993
Mycetophila pictula Meigen, 1830
= bimaculata Fabricius, 1805 nec Meigen, 1804
Mycetophila pseudoforcipata Zaitzev, 1998
Mycetophila pumila Winnertz, 1864
Mycetophila pyrenaica Matile, 1967
Mycetophila quadra Lundström, 1909
Mycetophila rucollis Meigen, 1818
Mycetophila schnabli (Dziedzicki, 1884)
Mycetophila sigmoides Loew, 1869
Mycetophila signata Meigen, 1830
Mycetophila signatoides Dziedzicki, 1884
= assimilis Matile, 1967
Mycetophila sinuosa Plassmann & Schacht, 1999
Mycetophila sordida van der Wulp, 1874
Mycetophila stolida Walker, 1856
Mycetophila stricklandi (Laoon, 1957)
Mycetophila strigata Staeger, 1840
Mycetophila strigatoides Landrock, 1927
Mycetophila strobli Laštovka, 1972
Mycetophila stylata (Dziedzicki, 1884)
Mycetophila sublunata Zaitzev, 1998
Mycetophila subsigillata Zaitzev, 1999
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Mycetophila sumavica (Laštovka, 1963)
Mycetophila triangularis Lundström, 1912
Mycetophila trinotata Staeger, 1840
Mycetophila uliginosa Chandler, 1988
Mycetophila unguiculata Lundström, 1913
Mycetophila uninotata Zetterstedt, 1852
Mycetophila unipunctata Meigen, 1818
Mycetophila xanthopyga Winnertz, 1864
Mycetophila zetterstedtii Lundström, 1906
PHRONIA Winnertz, 1864
Phronia aviculata Lundström, 1914
Phronia avidoides Jakovlev & Polevoi, 2009
Phronia biarcuata (Becker, 1908)
= johannae Steenberg, 1924
= praecox Edwards, 1925
Phronia bicolor Dziedzicki, 1889
= fusciventris van Duzee, 1928
Phronia borealis Hackman, 1970
Phronia braueri Dziedzicki, 1889
= annulata Winnertz, 1864
Phronia caliginosa Dziedzicki, 1889
= trivittata Dziedzicki, 1889
Phronia cinerascens Winnertz, 1864
Phronia conformis (Walker, 1856)
= girschneri Dziedzicki, 1889
Phronia cordata Lundström, 1914
Phronia coritanica Chandler, 1992
Phronia cornuta Lundström, 1914
Phronia crassitarsus Hackman, 1970
Phronia digitata Hackman, 1970
Phronia disgrega Dziedzicki, 1889
Phronia distincta Hackman, 1970
Phronia dziedzickii Lundström, 1906
Phronia egregia Dziedzicki, 1889
Phronia electa Dziedzicki, 1889
Phronia elegans Dziedzicki, 1889
Phronia elegantula Hackman, 1970
Phronia exigua (Zetterstedt, 1852)
Phronia fennica Jakovlev & Polevoi, 2009
Phronia avipes Winnertz, 1864
Phronia forcipata Winnertz, 1864
Phronia forcipula Winnertz, 1864
Phronia gagnei Chandler, 1992
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Phronia gracilis Hackman, 1970
Phronia humeralis Winnertz, 1864
Phronia interstincta Dziedzicki, 1889
Phronia longelamellata Strobl, 1898
= minuta Landrock, 1928
Phronia lutescens Hackman, 1970
Phronia maculata Dziedzicki, 1889
Phronia mutabilis Dziedzicki, 1889
Phronia mutila Lundström, 1911
Phronia nigricornis (Zetterstedt, 1852)
= dubia Dziedzicki, 1889
Phronia nigripalpis Lundström, 1909
Phronia nitidiventris (van der Wulp, 1859)
= squalida Winnertz, 1864
= vitiosa Winnertz, 1864
Phronia notata Dziedzicki, 1889
Phronia obscura Dziedzicki, 1889
Phronia obtusa Winnertz, 1864
Phronia peculiaris Dziedzicki, 1889
Phronia persimilis Hackman, 1970
Phronia petulans Dziedzicki, 1889
Phronia portschinskyi Dziedzicki, 1889
Phronia siebeckii Dziedzicki, 1889
Phronia signata Winnertz, 1864
= austriaca Winnertz, 1864
Phronia spinigera Hackman, 1970
Phronia strenua Winnertz, 1864
= avicollis Winnertz, 1864 nom. dubium
Phronia subsilvatica Hackman, 1970
Phronia sudetica Dziedzicki, 1889
Phronia sylvatica Dziedzicki, 1889
Phronia taczanowskyi Dziedzicki, 1889
Phronia tenuis Winnertz, 1864
Phronia tiei Dziedzicki, 1889
= marginata Dziedzicki, 1889
Phronia triangularis Winnertz, 1864
Phronia unica Dziedzicki, 1889
Phronia vitrea Plassmann, 1999
= carli Chandler, 2001
Phronia willistoni Dziedzicki, 1889
PLATUROCYPTA Enderlein, 1910
Platurocypta punctum (Stannius, 1831)
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Platurocypta testata (Edwards, 1925)
SCEPTONIA Winnertz, 1864
Sceptonia concolor Winnertz, 1864
Sceptonia costata (van der Wulp, 1859)
Sceptonia demeijerei Bechev, 1997
Sceptonia avipuncta Edwards, 1925
Sceptonia fumipes Edwards, 1925
Sceptonia fuscipalpis Edwards, 1925
Sceptonia hamata Ševčík, 2004
Sceptonia longisetosa Ševčík, 2004
Sceptonia membranacea Edwards, 1925
Sceptonia nigra (Meigen, 1804)
Sceptonia pughi Chandler, 1991
Sceptonia regni Chandler, 1991
Sceptonia tenuis Edwards, 1925
Sceptonia thaya Ševčík, 2004
TRICHONTA Winnertz, 1864
Trichonta apicalis Strobl, 1898
Trichonta atricauda (Zetterstedt, 1852)
= adunca Edwards, 1925
Trichonta beata Gagné, 1981
Trichonta bicolor Landrock, 1912
Trichonta bida Lundström, 1909
Trichonta brevicauda Lundström, 1909
Trichonta canora Gagné, 1981
Trichonta clara Gagné, 1981
Trichonta comica Gagné, 1981
Trichonta comis Gagné, 1981
Trichonta concinna Gagné, 1981
Trichonta conjungens Lundström, 1909
Trichonta delicata Gagné, 1981
Trichonta eximia Gagné, 1981
Trichonta facilis Gagné, 1981
Trichonta falcata Lundström, 1911
Trichonta ssicauda (Zetterstedt, 1852)
= claripennis Lundström, 1914
Trichonta avicauda Lundström, 1914
Trichonta fragilis Gagné, 1981
Trichonta fusca Landrock, 1918
Trichonta generosa Gagné, 1981
Trichonta girschneri Landrock, 1912
Trichonta hamata Mik, 1880
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Trichonta icenica Edwards, 1925
Trichonta melanura (Staeger, 1840)
Trichonta nigritula Edwards, 1925
Trichonta palustris Maximova, 2002
Trichonta patens Johannsen, 1912
Trichonta perspicua van der Wulp, 1881
= mediastinalis (Lundström, 1909)
Trichonta subfusca Lundström, 1909
Trichonta submaculata (Staeger, 1840)
Trichonta subterminalis Zaitzev & Menzel, 1996
Trichonta terminalis (Walker, 1856)
* Trichonta tristis (Strobl, 1898) (see Notes)
Trichonta trivittata Lundström, 1916
Trichonta venosa (Staeger, 1840)
Trichonta vitta (Meigen, 1830)
Trichonta vulcani (Dziedzicki, 1889)
= appropinquata (Strobl, 1900)
? = trida Lundström, 1909 (see Notes)
Trichonta vulgaris Loew, 1869
= nigricauda Lundström, 1906
ZYGOMYIA Winnertz, 1864
Zygomyia angusta Plassmann, 1977
Zygomyia humeralis (Wiedemann, 1817)
Zygomyia kiddi Chandler, 1991
Zygomyia notata (Stannius, 1831)
Zygomyia pictipennis (Staeger, 1840)
Zygomyia pseudohumeralis Caspers, 1980
Zygomyia semifusca (Meigen, 1818) (see Notes)
Zygomyia valida Winnertz, 1864
Zygomyia vara (Staeger, 1840)
Zygomyia zaitzevi Chandler, 1991
Excluded species
Bolitophilidae
Bolitophila spinigera Edwards, 1925 misidentied
Keroplatidae
Macrocera nigropicea Lundström, 1906 not found within present borders
Macrocera sudetica Landrock, 1924 only females, unidentiable (see Notes)
Orfelia bicolor (Macquart, 1826) only females, unidentiable (see Notes)
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Mycetophilidae
Allodia triangularis (Strobl, 1895) misidentied
Cordyla styliforceps (Bukowski, 1934) misidentied (see Notes)
Exechiopsis unguiculata (Lundström, 1911) not found within present borders
Leptomorphus quadrimaculatus Matsumura, 1916 misidentied (see Notes)
Monoclona mikii Kertesz, 1898 misidentied (see Notes)
Mycetophila gratiosa Winnertz, 1864 misidentied (see Notes)
Mycetophila lunata Meigen, 1804 misidentied (see Notes)
Mycetophila vittipes Zetterstedt, 1852 misidentied (see Notes)
Palaeodocosia alpicola (Strobl, 1895) misidentied (see Notes)
Pseudexechia trisignata (Edwards, 1913) not found within present borders
Pseudorymosia optiva (Dziedzicki, 1910) misidentied (see Notes)
Sciophila hebes Johannsen, 1910 misidentied (see Notes)
Synplasta excogitata (Dziedzicki, 1910) misidentied (see Notes)
Trichonta delis Gagné, 1981 mistake (see Notes)
Notes
Cordyla styliforceps (Bukowski, 1934) was added to the list of Finnish insects by Sil-
fverberg (2007) with a reference to Polevoi (2001) which most probably based on
misidentied species (A.Polevoi, pers. comm).
Leptomorphus quadrimaculatus Matsumura, 1916 was added to the list of Finnish
insects by Silfverberg (2001) with a reference to Polevoi (1995). is record has
later shown to belong to Leptomorphus subforcipatus (Zaitzev and Ševčík 2002).
Macrocera sudetica Landrock, 1924 and Orfelia bicolor (Macquart, 1826) (= basalis
Winnertz, 1864). Listed in Hackman (1980) by the only female specimen located
in the collection of the Finnish Museum of Natural History “Diptera Fennica”
that cannot be recognised. No further records from Finland.
Monoclona mikii Kertesz, 1898. Listed in Hackman (1980) by erroneously identied
male specimen of Monoclona braueri and by the female specimen (both are located
in “Diptera Fennica”) that cannot be recognized. Later this species has been recorded
from Finland under its valid name, Monoclona silvatica (Jakovlev et al. 2014).
Mycetophila gratiosa Winnertz, 1864. Listed in Hackman (1980) by erroneously
identied male specimens of Mycetophila luctuosa located in “Diptera Fennica”.
Mycetophila lunata Meigen, 1804 was interpreted broadly in old sources, and al-
though it was conrmed by Silfverberg (2001) with a reference to Polevoi (1995)
this record has later shown to belong to another related species, M. sublunata Za-
itzev, 1999. Hence, M. lunata still needs to be conrmed from Finland.
Mycetophila vittipes Zetterstedt, 1852. Listed in Hackman (1980) by erroneously identi-
ed male specimens of Mycetophila vittipes-group located in “Diptera Fennica”.
Palaeodocosia alpicola (Strobl, 1895) Listed in Hackman (1980) by erroneously
identied male specimen of P. vittata located in “Diptera Fennica”.
Jevgeni Jakovlev / ZooKeys 441: 119–149 (2014)
146
Pseudorymosia optiva (Dziedzicki, 1910). e small genus Pseudorymosia contains only
two known Palaearctic species, P. fovea (Dziedzicki, 1910) and P. optiva (Dziedzicki,
1910), that in fact might be only one species (Zaitzev (2003). Re-examination of
specimens in the “Diptera Fennica” collections formerly identied as P. optiva and
published by Hackman (1980) revealed that all of them are in fact the widely dis-
tributed P. fovea as interpreted in newer determination sources (Jakovlev et al. 2006).
Sciophila hebes Johannsen, 1910. Recorded by Komonen (2001) by erroneously
identied specimens of Sciophila antiqua.
Synplasta excogitata (Dziedzicki, 1910). Listed in Hackman (1980) by erroneously
identied male specimen of Synplasta gracilis.
Trichonta delis Gagné, 1981. Listed in Hackman (1980) with the note that T. delis
Gagné was only a manuscript of Gagné (1981) at the time. Apparently Gagné
changed his mind before publication and used this name for an oriental species
instead (Hackman 1982).
Zygomyia semifusca (Meigen, 1818). Listed in Hackman (1980) as Mycetophila semifusca.
Doubtful and unrecognized species
* Keroplatus tuvensis Zaitzev, 1991. is species might be a junor synonym of Keroplatus
testaceus (Kjærandsen et al. 2007). With this reference, it was deleted from the list of
Finnish insects by Silfverberg (2012), but this synonym has not been adopted in the
checklist of North European fungus gnats (Kjærandsen 2012). Hereby we keep K.
tuvensis in the Finnish checklist pending revision of the K. testaceus complex of species.
* Mycetophila biformis Maximova, 2002 preocc. Added by Jakovlev et al. (2014). is
is a junior primary homonym of Mycetophila biformis Duret, 1992. A replacement
name will be given by the author in the near future (Yulia Maximova, pers. comm.).
* Trichonta tristis (Strobl, 1898). Judging from illustration of Dziedzicki (1889), he
actually illustrated this species as Trichonta vulcani. is makes vulcani a junior
synonym of tristis. In this case, Trichonta trida Lundström, 1909 becomes the
valid species that we now call vulcani (J. Kjærandsen, pers.comm.). Here we list T.
vulcani and T. tristis as valid species sensu Zaitzev (2003), gures 107,1 and 109,5.
Species listed by Hackman (1980) with a question marks
Five species, Docosia avicoxa Strobl, 1900, Cordyla bergensis (Barendrecht, 1938),
Cordyla murina Winnertz, 1864, Mycetophila lunata Meigen, 1804 and Phronia
petulans Dziedzicki, 1889 were listed in the former checklist of Finnish Diptera (Hack-
man 1980) with a question marks. Later, Finnish records were conrmed for D. avicoxa
by Jakovlev et al. (2014), for C. murina by Jakovlev et al. (2006), and for P. petulans by
Jakovlev and Polevoi (2009). Cordyla bergensis was excluded as a junior synonym of Cor-
dyla nitidula (Jakovlev et al. 2006). Mycetophila lunata – see above.
Checklist of the fungus gnats of Finland: Bolitophilidae, Diadocidiidae, Ditomyiidae... 147
Acknowledgements
e author thanks Dr. Jukka Salmela (Finland) who has discovered tens of fungus gnat
species new to Finland resulting from his studies of under recorded fauna of wetlands,
mires and forests along brooks in northern boreal and oroarctic zones of Finnish Lap-
land. Jere Kahanpää and Dr. Jukka Salmela, the editors of the Checklist of Finnish
Diptera, Dr. Jostein Kjærandsen (Norway), the editor of the the online checklist of
Nordic fungus gnats (Kjærandsen 2012), Dr. Pekka Vilkamaa (Finland), and two ref-
erees: Dr. Jukka Salmela and Dr. Alexei Polevoi (Petrozavodsk, Russia) are thanked for
useful comments on the manuscript.
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Checklist of the family Sciaridae (Diptera) of Finland 151
Checklist of the family Sciaridae (Diptera) of Finland
Pekka Vilkamaa1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Pekka Vilkamaa (pekka.vilkamaa@helsinki.)
Academic editor: J. Salmela|Received 26 February 2014|Accepted 21 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Vilkamaa P (2014) Checklist of the family Sciaridae (Diptera) of Finland. In: Kahanpää J, Salmela J (Eds)
Checklist of the Diptera of Finland. ZooKeys 441: 151–164. doi: 10.3897/zookeys.441.7381
Abstract
A checklist of the family Sciaridae (Diptera) recorded from Finland is provided. e genus Sciarosoma
Chandler with a disputed family placement is also included in the list.
Keywords
Checklist, Finland, Diptera, Sciaridae, Sciarosoma
Introduction
Sciaridae, the black fungus gnats, is one of the large families in Sciaroidea, little stud-
ied and notoriously dicult in its taxonomy. Up-to-date keys are not available for all
European species, but various publications must be consulted for identication.Our
knowledge of Finnish fauna stems from two classic publications: Frey (1948) and Tuo-
mikoski (1960) but Menzel and Mohrig (2000) made fundamental nomenclatorial
changes in their study of the type materials of all Palaearctic species.
As no records of Sciaridae in Finland were published between the publication of
Tuomikoski’s (1960) monograph on the family and Hackman’s (1980) check-list
ZooKeys 441: 151–164 (2014)
doi: 10.3897/zookeys.441.7381
www.zookeys.org
Copyright Pekka Vilkamaa. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Pekka Vilkamaa / ZooKeys 441: 151–164 (2014)
152
of Finnish nematoceran Diptera, the species lists in both works are essentially un-
changed, despite some obvious lapses in the latter. Our knowledge of Finnish fauna
has since grown mainly in the taxonomic treatment of various genera and species
groups (see Vilkamaa et al. 2013). All species recorded in Finland in publications
thus far, besides the most recent ones, appear in H. Silfverberg’s compilations of
changes to the Finnish insect fauna published every ve years (for the latest one, see
Silfverberg 2012).
With regard to nomenclature and generic classication, the present list follows
the revisions of Menzel and Mohrig (2000) and Mohrig et al. (2013), although on
the basis of phylogenetic analyses Baeosciara Tuomikoski, Dolichosciara Tuomiko-
ski, Peyerimhoa Kieer, Prosciara Frey and Mouetina Frey are regarded as genera
(see Vilkamaa 2000, 2003, Vilkamaa and Hippa 2005, Shin et al. 2013). For names
currently understood as junior synonyms, mostly only the names listed as valid in
Tuomikoski (1960) are mentioned. For complete synonym lists, one can consult
Menzel and Mohrig (2000) and Mohrig et al. (2013). Although some genera can
be regarded as relatively well-known for the Finnish species they contain, for some
others, such as the species-rich Bradysia Winnertz, many new species including un-
described ones may be present in the native fauna. Even so, together with Germany,
Sweden and the United Kingdom, Finland can be considered one of the countries
best known for its sciarid fauna.
Number of species:
World: 2600 species
Europe: 700 species
Finland: 338 species
Faunistic knowledge level in Finland: average
Checklist
Nematocera Dumeril, 1805
infraorder Bibionomorpha Hennig, 1954
superfamily Sciaroidea Billberg, 1820
SCIARIDAE Billberg, 1820
BAEOSCIARA Tuomikoski, 1960
Baeosciara discolor (Lengersdorf, 1928)
= pusillima (Frey, 1948)
Baeosciara sinuata (Menzel & Mohrig, 1997)
Baeosciara scotica (Edwards, 1925)
BRADYSIA Winnertz, 1867
Bradysia anis (Zetterstedt, 1838)
= pratincola Tuomikoski, 1960
Checklist of the family Sciaridae (Diptera) of Finland 153
Bradysia albanensis (Lengersdorf, 1926)
Bradysia alpicola (Winnertz, 1867)
= mutabilis (Lengersdorf, 1926)
= morio auct. nec (Fabricius, 1794)
Bradysia angustata Tuomikoski, 1960
Bradysia angustostylata Menzel, 2005
Bradysia aprica (Winnertz, 1867)
Bradysia arcana Menzel & Mohrig, 1998
= fenestralis auct. nec (Zetterstedt, 1838)
Bradysia arcula Vilkamaa, Salmela & Hippa, 2007
Bradysia ascenda Rudzinski, 1994
Bradysia bicolor (Meigen, 1818)
Bradysia bispinifera Mohrig & Krivosheina, 1983
Bradysia brevispina Tuomikoski, 1960
Bradysia browni (Shaw, 1935)
= diversiabdominalis (Lengersdorf, 1941)
= laurencei Menzel & Mohrig, 2000
Bradysia cinerascens (Grzegorzek, 1884)
= lanicauda Tuomikoski, 1960
Bradysia connis (Winnertx, 1867)
Bradysia excelsa Menzel & Mohrig, 1998
Bradysia avipila Tuomikoski, 1960
Bradysia forcipulata (Lundbeck, 1898)
Bradysia forculata (Bezzi, 1914)
= nocturna Tuomikoski, 1960
Bradysia fungicola (Winnertz, 1867)
Bradysia giraudii (Egger, 1862)
Bradysia globulifera (Lengersdorf, 1934)
Bradysia hilariformis Tuomikoski, 1960
Bradysia hilaris (Winnertz, 1867)
Bradysia holsatica Heller, 2004
Bradysia hortensis Heller, 2000
Bradysia impatiens (Johannsen, 1912)
= diormis (Frey, 1948)
= paupera Tuomikoski, 1960
Bradysia inusitata Tuomikoski, 1960
Bradysia iridipennis (Zetterstedt, 1838)
Bradysia lapponica (Lengersdorf, 1926)
Bradysia latiterga Tuomikoski, 1960
Bradysia leptoptera Tuomikoski, 1960
Bradysia lilienthalae Mohrig & Menzel, 1990
Bradysia lobulifera Frey, 1948
Bradysia longicauda Mohrig & Menzel, 1990
Pekka Vilkamaa / ZooKeys 441: 151–164 (2014)
154
Bradysia longicubitalis (Lengersdorf, 1924)
= cinereovittata Frey, 1948
Bradysia minima Mohrig & Mamaev, 1989
Bradysia moesta Frey, 1948
= albosetosa Frey, 1948
Bradysia moestula Tuomikoski, 1960
Bradysia nervosa (Meigen, 1818)
Bradysia nitidicollis (Meigen, 1818)
= atroparva Frey, 1948
Bradysia normalis Frey, 1948
Bradysia ocellaris (Comstock, 1882)
Bradysia pallipes (Fabricius, 1787)
= brunnipes (Meigen, 1804)
= picipes (Zetterstedt, 1838)
Bradysia pauperata (Winnertz, 1867)
Bradysia peranis Tuomikoski, 1960
Bradysia pilistriata Frey, 1948
Bradysia placida (Winnertz, 1867)
= mbricauda Tuomikoski, 1960
Bradysia polonica (Lengersdorf, 1929)
Bradysia praecox (Meigen, 1818)
Bradysia rectinervis Frey, 1948
Bradysia reexa Tuomikoski, 1960
Bradysia regularis (Lengersdorf, 1934)
= subnervosa Frey, 1948
Bradysia rufescens (Zetterstedt, 1852)
Bradysia scabricornis Tuomikoski, 1960
= subscabricornis Mohrig & Menzel, 1990
Bradysia siberica Komarova, 2001
Bradysia spinostyla Mohrig & Menzel, 1990
Bradysia strigata (Staeger, 1840)
Bradysia subalpina Frey, 1948
Bradysia subamoena Mohrig & Krivosheina, 1989
Bradysia submoesta Mohrig & Krivosheina, 1989
Bradysia tilicola (Loew, 1850)
= amoena (Winnertz, 1867)
= cellarum Frey, 1948
Bradysia trivittata (Staeger, 1840)
Bradysia vagans (Winnertz, 1868)
= callicera Frey, 1948
Bradysia vernalis (Zetterstedt, 1851)
Bradysia zonata Rudzinski, 1993
BRADYSIOPSIS Tuomikoski, 1960
Checklist of the family Sciaridae (Diptera) of Finland 155
Bradysiopsis vittata (Meigen, 1830)
= leucotricha (Tuomikoski, 1960)
Bradysiopsis vittigera (Zetterstedt, 1851)
CAMPTOCHAETA Hippa & Vilkamaa, 1994
Camptochaeta austriaca Heller, 2012
Camptochaeta bournei (Shaw, 1941)
Camptochaeta camptochaeta (Tuomikoski, 1960)
Camptochaeta consimilis (Holmgren, 1869)
Camptochaeta delicata (Lengersdorf, 1935)
Camptochaeta duplicata Hippa & Vilkamaa, 1994
Camptochaeta fallax Hippa & Vilkamaa, 1994
Camptochaeta hirtula (Lengersdorf, 1934)
= fulvicollis (Tuomikoski, 1960)
Camptochaeta propria Hippa & Vilkamaa, 1994
Camptochaeta scanica Hippa & Vilkamaa, 1994
Camptochaeta sicilicula Hippa & Vilkamaa, 1994
Camptochaeta simulator Hippa & Vilkamaa, 1994
Camptochaeta stammeri (Lengersdorf, 1940)
Camptochaeta tenuipalpalis (Mohrig & Antonova, 1978)
Camptochaeta uniformis (Mohrig & Menzel, 1990)
Camptochaeta vivax (Frey, 1948)
Camptochaeta xystica Hippa & Vilkamaa, 1994
CHAETOSCIARA Frey, 1942
Chaetosciara estlandica (Lengersdorf, 1929)
CLAUSTROPYGA Hippa, Vilkamaa & Mohrig, 2003
Claustropyga acanthostyla (Tuomikoski, 1960)
Claustropyga brevichaeta (Mohrig & Antonova, 1978)
Claustropyga clausa (Tuomikoski, 1960)
Claustropyga corticis (Mohrig & Antonova, 1978)
Claustropyga ctenophora Hippa, Vilkamaa & Mohrig, 2003
Claustropyga heteroclausa (Rudzinski, 1991)
Claustropyga refrigerata (Lengersdorf, 1930)
Claustropyga subcorticis (Mohrig & Krivosheina, 1985)
CORYNOPTERA Winnertz, 1867
Corynoptera barbata Tuomikoski, 1960
Corynoptera bicuspidata (Lengersdorf, 1926)
= gymnops Tuomikoski, 1960
Corynoptera bipartita Mohrig & Krivosheina, 1985
Corynoptera blanda (Winnertz, 1867)
Corynoptera boletiphaga (Lengersdorf, 1940)
= geogenia Tuomikoski, 1960
Corynoptera breviformis Mohrig & Krivosheina, 1983
Corynoptera cracentis Vilkamaa, Hippa & Heller, 2013
Pekka Vilkamaa / ZooKeys 441: 151–164 (2014)
156
Corynoptera cuniculata (Lengersdorf, 1942)
= caldariorum Tuomikoski, 1960
Corynoptera defecta (Frey, 1948)
Corynoptera dentata (Bukowski & Lengersdorf, 1936)
Corynoptera deserta Heller & Menzel, 2006
= minutula (Bukowski & Lengersdorf, 1936)
Corynoptera dubitata Tuomikoski, 1960
Corynoptera fera Mohrig & Heller, 1992
Corynoptera avicauda (Zetterstedt, 1855)
Corynoptera forcipata (Winnertz, 1867)
Corynoptera furcifera Mohrig & Mamaev, 1987
Corynoptera globiformis (Frey, 1945)
Corynoptera hypopygialis (Lengersdorf, 1926)
= piniphila (Lengersdorf, 1940)
Corynoptera inari Vilkamaa, Hippa & Heller, 2013
Corynoptera inexspectata Tuomikoski, 1960
Corynoptera irmgardis (Lengersdorf, 1930)
Corynoptera levis Tuomikoski, 1960
Corynoptera luteofusca (Bukowski & Lengersdorf, 1936)
Corynoptera marinae Mohrig & Krivosheina, 1986
Corynoptera melanochaeta Mohrig & Menzel, 1992
Corynoptera membranigera (Kieer, 1903)
= trispina Tuomikoski, 1960
Corynoptera montana (Winnertz, 1869)
Corynoptera ninae Hippa, Vilkamaa & Heller, 2010
Corynoptera obscuripila Tuomikoski, 1960
Corynoptera parvula (Winnertz, 1867)
= uncinata (Hippa & Vilkamaa, 1994)
Corynoptera parvulaformis Mohrig, 1985
Corynoptera penna (Pettey, 1918)
= alneti Hippa, Vilkamaa & Heller, 2010
Corynoptera perochaeta (Mohrig & Menzel, 1990)
Corynoptera plusiochaeta Hippa, Vilkamaa & Heller, 2010
Corynoptera polana Rudzinski, 2009
Corynoptera postforcipata Rudzinski, 1993
Corynoptera postglobiformis Mohrig, 1993
Corynoptera praeforcipata Mohrig & Mamaev, 1987
Corynoptera quantula (Hippa & Vilkamaa, 1994)
Corynoptera saccata Tuomikoski, 1960
Corynoptera saetistyla Mohrig & Krivosheina, 1985
Corynoptera salmelai Vilkamaa, Hippa & Heller, 2013
Corynoptera sphenoptera Tuomikoski, 1960
Corynoptera spiciforceps Vilkamaa, Hippa & Heller, 2013
Checklist of the family Sciaridae (Diptera) of Finland 157
Corynoptera spinifera Tuomikoski, 1960
Corynoptera subblanda Tuomikoski, 1960
Corynoptera subdentata Mohrig, 1985
Corynoptera subparvula Tuomikoski, 1960
Corynoptera subsedula Mohrig & Mamaev, 1987
Corynoptera subtetrachaeta Komarova, 1995
Corynoptera subtilis (Lengersdorf, 1929)
= longicornis (Bukowski & Lengersdorf, 1936)
Corynoptera subvariegata Rudzinski, 1992
Corynoptera tetrachaeta Tuomikoski, 1960
Corynoptera trepida (Winnertz, 1867)
= clinochaeta Tuomikoski, 1960
Corynoptera triacantha Tuomikoski, 1960
Corynoptera tumidula Hippa, Vilkamaa & Heller, 2010
Corynoptera tuomikoskii Hippa, Vilkamaa & Heller, 2013
Corynoptera unidentata (Hippa & Vilkamaa, 1994)
Corynoptera vagula Tuomikoski, 1960
Corynoptera verrucifera (Lengersdorf, 1952)
Corynoptera voluptuosa Mohrig & Mamaev, 1987
Corynoptera waltraudis Mohrig & Mamaev, 1987
Corynoptera winnertzi Mohrig, 1993
COSMOSCIARA Frey, 1942
Cosmosciara perniciosa (Edwards, 1922)
CRATYNA Winnertz, 1867
= Plastosciara Berg, 1899
= Decembrina Frey, 1948
= Dendrosciara Frey, 1948
sg. Cratyna Winnertz, 1867
Cratyna ambigua (Lengersdorf, 1934)
= latiforceps (Bukowski & Lengersdorf, 1936)
Cratyna atra Winnertz, 1867
= pictiventris (Kieer, 1898)
Cratyna betulae (Mohrig & Mamaev, 1992)
Cratyna breviagellata (Mohrig & Mamaev, 1985)
Cratyna monumenta Rudzinski, 2009
Cratyna pernitida (Edwards, 1915)
Cratyna schineri (Winnertz, 1867)
Cratyna sicata Vilkamaa, Hippa & Heller, 2013
Cratyna symplecta (Rudzinski, 1991)
Cratyna uliginosa (Lengersdorf, 1929)
Cratyna vaporariorum (Frey, 1948)
sg. Diversicratyna Menzel & Mohrig, 1998
Cratyna spiculosa (Rudzinski, 1993)
Pekka Vilkamaa / ZooKeys 441: 151–164 (2014)
158
sg. Spathobdella Frey, 1948
Cratyna colei (Freeman, 1990)
= brachialis auct. nec (Winnertz, 1871)
Cratyna falcata (Tuomikoski, 1960)
Cratyna falcifera (Lengersdorf, 1933)
Cratyna longispina (Pettey, 1918)
= tuberculata (Tuomikoski, 1960)
Cratyna nobilis (Winnertz, 1867)
= brachialis (Winnertz, 1867)
Cratyna perplexa (Winnertz, 1867)
= socialis (Winnertz, 1871)
= brevicornis (Tuomikoski, 1957)
CTENOSCIARA Tuomikoski, 1960
Ctenosciara exigua Salmela & Vilkamaa, 2005
Ctenosciara hyalipennis (Meigen, 1804)
DICHOPYGINA Vilkamaa, Hippa & Komarova, 2004
Dichopygina aculeata Vilkamaa, Hippa & Komarova, 2004
Dichopygina intermedia (Mohrig & Krivosheina, 1982)
Dichopygina nigrohalteralis (Frey, 1948)
Dichopygina ramosa Vilkamaa, Hippa & Komarova, 2004
DOLICHOSCIARA Tuomikoski, 1960
Dolichosciara avipes (Meigen, 1804) 185
Dolichosciara hippai Komarova & Vilkamaa, 2006
Dolichosciara nigrovittata (Strobl, 1910)
Dolichosciara orcina (Tuomikoski, 1960)
Dolichosciara ornata (Winnertz, 1867)
Dolichosciara saetosa (Lengersdorf, 1929)
Dolichosciara spissispina Vilkamaa, Hippa & Heller, 2013
EPIDAPUS Haliday, 1851
sg. Epidapus Haliday, 1851
Epidapus alnicola (Tuomikoski, 1957)
Epidapus atomarius (De Geer, 1778)
Epidapus gracilis (Walker, 1848)
Epidapus ignotus (Lengersdorf, 1942)
= gracilior (Tuomikoski, 1960)
Epidapus microthorax (Börner, 1903)
= gracilicornis (Lengersdorf, 1926)
Epidapus schillei (Börner, 1903)
= titan Frey, 1948
= intermittens Tuomikoski, 1959
sg. Pseudoaptanogyna Vimmer, 1926
Epidapus abieticola Frey, 1948
Epidapus bispinulosus Mohrig & Kauschke, 1994
Checklist of the family Sciaridae (Diptera) of Finland 159
Epidapus echinatum Mohrig & Kozánek, 1992
Epidapus ignavus (Lengersdorf, 1941)
KEILBACHIA Mohrig, 1987
Keilbachia ferrata (Hippa & Vilkamaa, 1994)
LEPTOSCIARELLA Tuomikoski, 1960
sg. Hirtipennia Mohrig & Menzel, 1997
Leptosciarella hirtipennis (Zetterstedt, 1838)
Leptosciarella holotricha Mohrig & Menzel, 1997
sg. Leptosciarella Tuomikoski, 1960
Leptosciarella brevior (Tuomikoski, 1960)
Leptosciarella brevipalpa (Mohrig & Menzel, 1992)
Leptosciarella claviforceps (Tuomikoski, 1960)
Leptosciarella dimera (Tuomikoski, 1960)
Leptosciarella fuscipalpa (Mohrig & Mamaev, 1979)
Leptosciarella helvetica (Rudzinski, 1992)
Leptosciarella hispida (Winnertz, 1871)
Leptosciarella ignis Heller, 2012
Leptosciarella krille Heller, 2012
Leptosciarella melanoma (Mohrig & Menzel, 1990)
Leptosciarella nudinervis (Tuomikoski, 1960)
Leptosciarella pilosa (Staeger, 1840)
Leptosciarella reducta Heller & Menzel, 2013
Leptosciarella rejecta (Winnertz, 1867)
Leptosciarella scutellata (Staeger, 1840)
= elegans (Winnertz, 1867)
Leptosciarella subcoarctata Mohrig & Menzel, 1997
Leptosciarella subpilosa (Edwards, 1925)
Leptosciarella subspinulosa (Edwards, 1925)
Leptosciarella subviatica Mohrig & Menzel, 1997
Leptosciarella trochanterata (Zetterstedt, 1851)
= coarctata (Winnertz, 1867)
Leptosciarella truncata (Tuomikoski, 1960)
Leptosciarella viatica (Winnertz, 1867)
Leptosciarella viaticella Mohrig & Krivosheina, 1979
Leptosciarella yerburyi (Freeman, 1983)
sg. Leptospina Mohrig & Menzel, 1997
Leptosciarella atricha (Tuomikoski, 1960)
sg. Trichosiopsis Tuomikoski, 1960
Leptosciarella tuberculigera (Tuomikoski, 1960)
LYCORIELLA Frey, 1942
sg. Coelostylina Tuomikoski, 1960
Lycoriella eagellata Tuomikoski, 1960
Lycoriella freyi Tuomikoski, 1960
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160
sg. Hemineurina Tuomikoski, 1960
Lycoriella algida (Frey, 1948)
Lycoriella cochleata (Rübsaamen, 1898)
Lycoriella conspicua (Winnertz, 1867)
Lycoriella inata (Winnertz, 1867)
= venosa auct. nec (Staeger, 1840)
Lycoriella modesta (Staeger, 1840)
Lycoriella piristylata Vilkamaa, Hippa & Heller, 2013
Lycoriella thuringiensis Menzel & Mohrig, 1991
Lycoriella vitticollis (Holmgren, 1883)
= permutata (Lundbeck, 1900)
sg. Lycoriella Frey, 1942
Lycoriella aberrans Tuomikoski, 1960
Lycoriella acutostylia Mohrig & Menzel, 1990
Lycoriella agraria (Felt, 1897)
= cellaris (Lengersdorf, 1934)
Lycoriella approximatonervis (Frey, 1948)
Lycoriella brevipila Tuomikoski, 1960
Lycoriella inconspicua Tuomikoski, 1960
Lycoriella ingenua (Dufour, 1839)
= solani (Winnertz, 1871)
Lycoriella latilobata Menzel & Mohrig, 2000
Lycoriella lundstromi (Frey, 1948)
Lycoriella micria Mohrig & Menzel, 1990
Lycoriella minutula Mohrig & Krivosheina, 1987
Lycoriella pallidior Tuomikoski, 1960
Lycoriella parva (Holmgren, 1869)
= obscuratipes (Frey, 1948)
= curvispina Tuomikoski, 1960
Lycoriella sativae (Johannsen, 1912)
=castanescens (Lengersdorf, 1940)
= fucorum (Frey, 1948)
Lycoriella subterranea (Märkel, 1844)
= vanderwieli (Schmitz, 1920)
Lycoriella tenera Vilkamaa, Hippa & Heller, 2013
Lycoriella weberi Menzel & Heller, 2013
MOUFFETINA Frey, 1942
Mouetina expolita (Coquillett, 1900)
Mouetina pulchricornis (Edwards, 1925)
Mouetina silvestris (Mohrig & Antonova, 1978)
PEYERIMHOFFIA Kieer, 1903
Peyerimhoa crassistylata (Frey, 1948)
Peyerimhoa infera Vilkamaa & Hippa, 2005
Checklist of the family Sciaridae (Diptera) of Finland 161
Peyerimhoa menzeli Vilkamaa & Hippa, 2005
Peyerimhoa quadrifera Vilkamaa, Hippa & Heller, 2013
Peyerimhoa sepei Hippa & Vilkamaa, 2005
Peyerimhoa thula Vilkamaa & Hippa, 2005
Peyerimhoa vagabunda (Winnertz, 1867)
= brachyptera Kieer, 1903
PHYTOSCIARA Frey, 1942
Phytosciara halterata (Lengersdorf, 1926)
Phytosciara macrotricha (Lengersdorf, 1926)
PNYXIA Johannsen, 1912
Pnyxia scabiei (Hopkins, 1895)
PNYXIOPSIS Tuomikoski, 1960
Pnyxiopsis aliger Tuomikoski, 1960
Pnyxiopsis degener (Tuomikoski, 1957)
PROSCIARA Frey, 1942
Prosciara furtiva Vilkamaa & Hippa, 1996
Prosciara plusiochaeta Hippa & Vilkamaa, 1991
Prosciara porrecta (Lengersdorf, 1929)
Prosciara producta (Tuomikoski, 1960)
Prosciara prosciaroides (Tuomikoski, 1960)
Prosciara ungulata (Winnertz, 1867)
PSEUDOLYCORIELLA Menzel & Mohrig, 1998
Pseudolycoriella brunnea (Bukowski & Lengersdorf, 1936)
Pseudolycoriella japonensis (Mohrig & Menzel, 1992)
Pseudolycoriella koreensis (Mohrig & Menzel, 1992)
Pseudolycoriella monticula (Mohrig & Menzel, 1992)
Pseudolycoriella nodulosa (Mohrig & Krivosheina, 1985)
Pseudolycoriella paludum (Frey, 1948)
Pseudolycoriella subbruckii (Mohrig & Hövemeyer, 1992)
SCATOPSCIARA Edwards, 1927
sg. Scatopsciara Edwards, 1927
Scatopsciara atomaria (Zetterstedt, 1851)
= vivida (Winnertz, 1867)
Scatopsciara bucera Rudzinski, 1994
Scatopsciara calamophila Frey, 1948
Scatopsciara curviforceps (Bukowski & Lengersdorf, 1936)
= myrmecophila Frey, 1948
Scatopsciara edwardsi Freeman, 1983
Scatopsciara uviatilis (Lengersdorf, 1940)
Scatopsciara geophila (Tuomikoski, 1960)
Scatopsciara multispina (Bukowski & Lengersdorf, 1936)
Scatopsciara neglecta Menzel & Mohrig, 1998
Scatopsciara pusilla (Meigen, 1818)
Pekka Vilkamaa / ZooKeys 441: 151–164 (2014)
162
Scatopsciara subcalamophila Menzel & Mohrig, 1991
Scatopsciara subciliata Tuomikoski, 1960
Scatopsciara tricuspidata (Winnertz, 1867)
= degenerans (Frey, 1948)
Scatopsciara vitripennis (Meigen, 1818)
sg. Xenopygina Frey, 1948
Scatopsciara gabyae (Heller, 1998)
Scatopsciara obliqua Vilkamaa, Hippa & Heller, 2013
Scatopsciara paradoxa (Frey, 1948)
Scatopsciara simillima (Tuomikoski, 1960)
SCHWENCKFELDINA Frey, 1942
Schwenckfeldina carbonaria (Meigen, 1830)
Schwenckfeldina pectinea Menzel & Mohrig, 1991
SCIARA Meigen, 1803
Sciara avimana Zetterstedt, 1851
Sciara hebes (Loew, 1869)
= mendax Tuomikoski., 1960
= nursei Freeman, 1983
= ulrichi Menzel & Mohrig, 1998
Sciara helvola Winnertz, 1867
Sciara hemerobioides (Scopoli, 1763)
= thomae (Linnaeus, 1767)
Sciara humeralis Zetterstedt, 1851
Sciara lackschewitzi (Lengersdorf, 1934)
SCYTHROPOCHROA Enderlein, 1911
Scythropochroa quercicola (Winnertz, 1869)
Scythropochroa radialis Lengersdorf, 1926
TRICHOSIA Winnertz, 1867
= Leptosciara Frey, 1942
= Lestremioides Frey, 1942
Trichosia Winnertz, 1867
Trichosia acrotricha Tuomikoski, 1960
Trichosia borealis (Frey, 1942)
Trichosia confusa Menzel & Mohrig, 1997
Trichosia avicoxa Tuomikoski, 1960
Trichosia diota (Garrett, 1925)
Trichosia glabra (Meigen, 1830)
Trichosia habilis (Johannsen, 1912)
= edwardsi (Lengersdorf, 1930)
Trichosia morio (Fabricius, 1794)
= caudata (Walker, 1848)
Trichosia splendens Winnertz, 1867
Trichosia ussurica Mohrig & Antonova, 1978
Checklist of the family Sciaridae (Diptera) of Finland 163
XYLOSCIARA Tuomikoski, 1957
sg. Protoxylosciara Tuomikoski, 1960
Xylosciara longiforceps (Bukowski & Lengersdorf, 1936)
sg. Xylosciara Tuomikoski, 1957
Xylosciara heptacantha Tuomikoski, 1957
Xylosciara lignicola (Winnertz, 1867)
Xylosciara microdon (Frey, 1948)
Xylosciara misella (Frey, 1948)
Xylosciara phryganophila (Frey, 1948)
Xylosciara senta Vilkamaa, Hippa & Heller, 2013
Xylosciara spinata (Pettey, 1918)
= betulae Tuomikoski, 1960
Xylosciara steleocera Tuomikoski, 1960
Xylosciara trimera Tuomikoski, 1960
Xylosciara validinervis Tuomikoski, 1960
ZYGONEURA Meigen, 1830
sg. Allozygoneura Menzel & Mohrig, 1998
Zygoneura calthae Tuomikoski, 1960
sg. Zygoneura Meigen, 1830
Zygoneura sciarina Meigen, 1830
UNPLACED IN SCIAROIDEA
SCIAROSOMA Chandler, 2002
Sciarosoma nigriclava (Strobl, 1898)
= borealis Chandler, 2002
Excluded species
Lycoriella auripila (Winnertz, 1867), misidentied in Tuomikoski (1960).
Bradysia betuleti (Lengersdorf, 1940), misidentied in Tuomikoski (1960).
Bradysia subbetuleti Mohrig & Krivosheina, 1989, misidentied in Salmela and Vilkamaa
(2005).
Corynoptera concinna (Winnertz, 1867), misidentied in Tuomikoski (1960).
Corynoptera bistrispina (Bukowski & Lengersdorf, 1936), misidentied in Tuomikoski
(1960).
Trichosia trochanterata (Zetterstedt, 1851), misidentied in Tuomikoski (1960).
Acknowledgements
I thank my colleagues Kai Heller, Heikki Hippa, Frank Menzel, Werner Mohrig and
Jukka Salmela for their long-term help in identifying Sciaridae.
Pekka Vilkamaa / ZooKeys 441: 151–164 (2014)
164
References
Frey R (1948) Entwurf einer neuen Klassikation de Mückenfamilie Sciaridae. II. Die nor-
deuropäishen Arten. Notulae Entomologique 27: 1–92.
Hackman W (1980) A check list of the Finnish Diptera I. Nematocera and Brachycera (s.str.).
Notulae Entomologique 60: 17–48.
Menzel F, Mohrig W (2000) Revision der paläarktischen Trauermücken (Diptera: Sciaridae).
Studia dipterologica Supplement 6: 1–761.
Mohrig W, Heller K, Hippa H, Vilkamaa P, Menzel F (2013) Revision of the black fungus
gnats (Diptera: Sciaridae) of North America. Studia dipterologica 19: 141–286.
Salmela J, Vilkamaa, P (2005) Sciaridae (Diptera) from central Finland: faunistics and taxonomy.
Entomologica Fennica 16: 287–300.
Shin S, Jung S, Menzel F, Heller K, Lee H, Lee S (2013) Molecular phylogeny of black fungus
gnats (Diptera: Sciaroidea: Sciaridae) and the evolution of larval habitats. Molecular Phy-
logenetics and Evolution 66: 833–846. doi: 10.1016/j.ympev.2012.11.008
Silfverberg H (2012) Changes 2006–2010 in the list of Finnish insects. Entomologica Fennica
23: 18–41.
Tuomikoski R (1960) Zur Kenntnis der Sciariden (Dipt.) Finnlands. Annales Zoologici Socie-
tatis Zoologicae Botanicae Fennicae ’Vanamo’ 21: 1–164.
Vilkamaa P (2000) Phylogeny of Prosciara Frey and related genera (Diptera: Sciaridae). Systematic
Entomology 25: 47–72. doi: 10.1046/j.1365-3113.2000.00094.x
Vilkamaa P (2003) Review of the genus Baeosciara Tuomikoski (Diptera: Sciaridae). Entomologica
Fennica 14: 220–226.
Vilkamaa P, Hippa H (2005) Phylogeny of Peyerimhoa Kieer, with the revision of the species.
Insect Systematics and Evolution 35: 457–480. doi: 10.1163/187631204788912445
Vilkamaa P, Heller K, Hippa H (2013) New species of Sciaridae (Diptera) to the Finnish
fauna. Sahlbergia 19: 20–29.
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 165
Checklist of the ‘lower Brachycera’ of Finland:
Tabanomorpha, Asilomorpha and
associated families (Diptera)
Jere Kahanpää1, Kaj Winqvist2, eo Zeegers3
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
2 Mikonkatu 3 C 52, FI–20100 Turku, Finland 3 Eikenlaan 24, 3768 EV Soest, the Netherlands
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 5 February 2014|Accepted 9 June 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J, Winqvist K, Zeegers T (2014) Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha,
Asilomorpha and associated families (Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland.
ZooKeys 441: 165–181. doi: 10.3897/zookeys.441.7198
Abstract
A checklist of the ‘lower Brachycera’ of Finland is presented. is part of the complete checklist of Finnish
Diptera covers the families Acroceridae, Asilidae, Athericidae, Bombyliidae, Mythicomyiidae, Rhagioni-
dae, Scenopinidae, Stratiomyidae, Tabanidae, erevidae, Xylomyidae and Xylophagidae.
Keywords
Species list, Finland, Diptera, biodiversity, faunistics
Introduction
is part of the checklist of the Diptera of Finland covers non-eremoneuran true ies
(Diptera: Brachycera). e brachyceran ies excluded from the clade Eremoneura are
often called the ‘lower Brachycera’ due to their basal position in the true y tree of life.
It remains unclear whether this assemblage of families is a monophyletic clade. ere
are also several models for the relative relationships of the various superfamilies and
families. A simple classication scheme following Marshall (2012) is adopted for this
ZooKeys 441: 165–181 (2014)
doi: 10.3897/zookeys.441.7198
www.zookeys.org
Copyright Jere Kahanpää et al. This is an open access ar ticle distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää et al. / ZooKeys 441: 165–181 (2014)
166
checklist. Only two infraorders, Tabanomorpha and Asilomorpha, are recognized. e
presentation order of families follows Woodley et al. (2009).
World catalogues have recently been published for Stratiomyidae (Woodley
2001, 2011b), Xylomyidae (Woodley 2011a), Xylophagidae (Woodley 2011c),
Bombyliidae (Evenhuis and Greathead 1999, 2003) and Mythicomyiidae (Even-
huis 2002). e Finnish species were last listed by Kahanpää and Winqvist (2005).
Five species have been added since the last checklist: Haematopota italica Meigen,
1804, Lasiopogon septentrionalis Lehr, 1984, Nemotelus infortunatus Kahanpää,
2010, Xylophagus inermis Krivosheina & Krivosheina, 2000 and Zabrachia tenella
(Jaennicke, 1866) (see Kahanpää 2013, Cannings and Kahanpää 2013, Kahanpää
2010, this paper, and Krivosheina and Rozkošný 1990 respectively). Table 1 sum-
marizes the current family species counts for the world, Europe (based on Fauna
Europaea), and Finland.
Tabanomorpha
e stratiomyoid and xylophagoid lineages are often treated as infraorders (Woodley
et al. 2009).
e soldieries (Stratiomyidae) are very diverse in the tropics but the species diver-
sity decreases sharply towards the higher latitudes. e wood soldier ies (xylomyids)
is a small y family associated with dead wood. e Fauna Entomologica Scandinavica
series has a volume on stratiomyoid ies (Rozkošný 1973). A new Nemotelus species
was recently described from Finland (Kahanpää 2010a).
e Finnish rhagionids are relatively well known but a few additional species could
occur in the country. Ptiolina is a problematic genus and the number of recognized
species in Northern Europe has varied from two to seven during the last century.
Athericidae was traditionally placed as a subfamily of Rhagionidae, but it seems more
closely associated with Tabanidae (Marshall 2012). Itämies et al. (1990, 1993) have
studied the distribution of Atherix ibis in Finland.
e Finnish xylophagid fauna is relatively well sampled. Adults of the North Euro-
pean species can be identied using Nartshuk (1988) or Kahanpää (2009). e larvae
can also be identied at least at the last larval stage (Stubbs and Drake 2001, Kriv-
osheina and Krivosheina 1966).
e tabanid nomenclature (especially Hybomitra) is quite convoluted and records
in older publications must be taken with a grain of salt. Karvonen (1969) summa-
rized the distribution of tabanids in Finland, but this work is now partially obsolete
due to the diculties in identifying Hybomitra and Haematopota before Chvála et al.
(1972) was published. For identication of North European tabanids Chvála et al.
(1972) complemented with pictures in Zeegers and van Haaren (2000) or Krčmar et
al. (2011) is recommended. An illustrated guide to the Finnish species is in preparation
(A. Haarto, unpublished).
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 167
Asilomorpha
e asilids and bombyliids of Finland are rather well known from a faunistic point of view
but little is known about their ecology. Most of the North European species are easy to iden-
tify but problems with Villa resulted in a cascade of name changes in the late 20th century.
Falck (2009) and Blöchlinger (2008) are good starting points for identifying Villa adults.
François (1969) has male genitalia gures for some of the more dicult Villa species. e
Mythicomyiidae or micro bee ies were long seen as a subfamily of Bombyliidae.
Identifying ereva species was also fraught with diculties in the past but by
the end of the 20th century the North European fauna was pretty well understood.
A review of the Finnish therevid fauna with keys has recently been published (Haarto
and Winqvist 2006). e window ies, Scenopinidae, is a smallish asiloid lineage as-
sociated with the therevids. It has even been proposed they are a specialized subgroup
of the erevidae (Woodley 2009).
Acroceridae
e small-headed ies (Acroceridae) are a y family of obscure origin. Anities with
Nemestrinidae, Tabanoidea, Stratiomyoidea, Bombyliidae and Asilomorpha have been
proposed (see Marshall 2012 for further discussion). Finnish acrocerid records are
mostly of single adults caught by sweep-netting, although Storå (1956) found groups
of 20–40 Acrocera orbiculus swarming on a coastal meadow. e acrocerid species seem
to have declined in abundance during the 20th century. Four of our ve Finnish spe-
cies are now on the national red list (Kahanpää 2010b).
Table 1. Number of species in tabanomorph and asilomorph families plus Acroceridae.
Family Number of species in Level of knowledge
World Europe Finland
Tabanomorpha:
Stratiomyidae 2715 (Woodley 2001, 2011b) 141 29 good
Xylomyidae 138 (Woodley 2011a) 13–14 1 good
Xylophagidae 134 (Woodley 2011c) 8 5 average–good
Rhagionidae 694 (Pape et al. 2011) 85 15–16 average–good
Athericidae 124 (Pape et al. 2011) 10 1 good
Tabanidae 4405 (Pape et al. 2011) 213 38–39 good
Asilomorpha:
Asilidae 7513 (Pape et al. 2011) 524 35 good
Bombyliidae ~5000 (Evenhuis and Greathead 1999, 2003) 335 18–19 good
Mythicomyiidae ~330 (Evenhuis 2002, Pape et al. 2011) 30 1 average
Scenopinidae 416 (Pape et al. 2011) 17 3 good
erevidae 1129 (Pape et al. 2011) 99 17 average–good
unplaced:
Acroceridae 392 (Pape et al. 2011) 34 5 average
Jere Kahanpää et al. / ZooKeys 441: 165–181 (2014)
168
Checklist part 1: Tabanomorpha (sensu lato)
suborder Brachycera Macquart, 1834
clade Orthorrapha Brauer, 1863
superfamily Stratiomyoidea Latreille, 1802
STRATIOMYIDAE Latreille, 1802
BERIDINAE Westwood, 1838
BERIS Latreille, 1802
Beris chalybata (Forster, 1771)
Beris clavipes (Linnaeus, 1767)
Beris fuscipes Meigen, 1820
Beris hauseri Stuke, 2004
= strobli auct. nec. Dušek & Rozkošný, 1968
Beris morrisii Dale, 1841
NEMOTELINAE Kertész, 1912
NEMOTELUS Georoy, 1762
sg. Camptopelta Williston, 1917
Nemotelus nigrinus Fallén, 1817
sg. Nemotelus Georoy, 1762
Nemotelus infortunatus Kahanpää, 2010
Nemotelus notatus Zetterstedt, 1842
Nemotelus uliginosus (Linnaeus, 1767)
PACHYGASTRINAE Loew, 1856
BERKSHIRIA Johnson, 1914
= Pseudowallacea Kertész, 1921
Berkshiria hungarica (Kertesz, 1921)
= albistylum misid.
= barovskii misid.
NEOPACHYGASTER Austen, 1901
Neopachygaster meromelas (Dufour, 1841)
= orbitalis (Wahlberg, 1854)
ZABRACHIA Coquillett, 1901
Zabrachia minutissima (Zetterstedt, 1838)
Zabrachia tenella (Jaennicke, 1866) see Notes
SARGINAE Walker, 1834
CHLOROMYIA Duncan, 1837
Chloromyia formosa (Scopoli, 1763)
MICROCHRYSA Loew, 1855
Microchrysa cyaneiventris (Zetterstedt, 1842)
Microchrysa avicornis (Meigen, 1822)
Microchrysa polita (Linnaeus, 1758)
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 169
SARGUS Fabricius, 1798
Sargus cuprarius (Linnaeus, 1758)
Sargus avipes Meigen, 1822
= nigripes Zetterstedt, 1842
= splendens auct. nec. Meigen, 1804
Sargus iridatus (Scopoli, 1763)
Sargus rupes Wahlberg, 1854
STRATIOMYINAE Latreille, 1802
tribe Oxycerini Enderlein, 1914
OXYCERA Meigen, 1803
Oxycera centralis Loew, 1863
= centralis Frey, 1911 preocc.
= freyi Lindner, 1938
Oxycera dives Loew, 1845
Oxycera trilineata (Linnaeus, 1767)
tribe Stratiomyini Latreille, 1802
ODONTOMYIA Meigen, 1803
Odontomyia angulata (Panzer, 1798)
Odontomyia argentata (Fabricius, 1794)
Odontomyia microleon (Linnaeus, 1758)
OPLODONTHA Rondani, 1863
Oplodontha viridula (Fabricius, 1775)
STRATIOMYS Georoy, 1762
Stratiomys singularior (Harris, 1776)
= furcata Fabricius, 1794
XYLOMYIDAE Verrall, 1901
XYLOMYA Rondani, 1861
Xylomya czekanovskii Pleske, 1925
= interrupta auct. nec. (Pleske, 1926)
= maculata auct. nec. (Meigen, 1804)
superfamily Xylophagoidea Fallén, 1810
XYLOPHAGIDAE Fallén, 1810
XYLOPHAGUS Meigen, 1803
= Erinna Meigen, 1800 suppr.
Xylophagus ater Meigen, 1804 see Notes
= compeditus Wiedemann, 1820
Xylophagus cinctus (De Geer, 1776)
Xylophagus inermis Krivosheina & Krivosheina, 2000 see Notes
= matsumurae misid.
Xylophagus junki (Szilády, 1932)
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170
Xylophagus kowarzi (Pleske, 1925) see Notes
= ater auct. nec. Meigen, 1804
superfamily Rhagionoidea Latreille, 1802
RHAGIONIDAE Latreille, 1802
RHAGIONINAE Latreille, 1802
RHAGIO Fabricius, 1775
Rhagio annulatus (De Geer, 1776)
Rhagio lineola Fabricius, 1794
Rhagio maculatus (De Geer, 1776)
Rhagio notatus (Meigen, 1820)
Rhagio scolopaceus (Linnaeus, 1758)
Rhagio tringarius (Linnaeus, 1758)
CHRYSOPILINAE Bezzi, 1903
CHRYSOPILUS Macquart, 1826
Chrysopilus auratus (Fabricius, 1805)
?= cristatus (Fabricius, 1775) nom. dubium
Chrysopilus luteolus (Fallén, 1814)
Chrysopilus nubecula (Fallén, 1814)
? Chrysopilus suomianus (Szilády, 1934) see Notes
SPANIINAE Rondani, 1856
OMPHALOPHORA Becker, 1900
Omphalophora oculata Becker, 1900
= lapponica Frey, 1911
PTIOLINA Zetterstedt, 1842
Ptiolina nigra Zetterstedt, 1842
Ptiolina nigrina Wahlgren, 1854 see Notes
Ptiolina nitida Wahlgren, 1854
Ptiolina obscura (Fallén, 1814)
SPANIA Meigen, 1830
Spania nigra Meigen, 1830
SYMPHOROMYIA Frauenfeld, 1867
sg. Paraphoromyia Becker, 1921
Symphoromyia crassicornis (Panzer, 1806)
ATHERICIDAE Nowicki, 1873
ATHERIX Meigen, 1803
Atherix ibis (Fabricius, 1798)
superfamily Tabanoidea Latreille, 1802
TABANIDAE Latreille, 1802
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 171
CHRYSOPSINAE Lutz, 1905
tribe Chrysopsini Lutz, 1905
CHRYSOPS Meigen, 1803
sg. Chrysops Meigen, 1803
Chrysops caecutiens (Linnaeus, 1758)
Chrysops divaricatus Loew, 1858
Chrysops nigripes Zetterstedt, 1838
= lapponicus Loew, 1858
Chrysops relictus Meigen, 1820
= melanopleurus Wahlberg, 1848
Chrysops rupes Meigen, 1820
Chrysops sepulcralis (Fabricius, 1794)
Chrysops viduatus (Fabricius, 1794)
= pictus Meigen, 1820
TABANINAE Latreille, 1802
tribe Haematopotini Enderlein, 1922
HAEMATOPOTA Meigen, 1803
Haematopota crassicornis Wahlberg, 1848
Haematopota italica Meigen, 1804
Haematopota pluvialis (Linnaeus, 1758)
= italica misid.
? Haematopota subcylindrica Pandellé, 1883 see Notes
HEPTATOMA Meigen, 1803
Heptatoma pellucens (Fabricius, 1776)
tribe Tabanini Latreille, 1802
ATYLOTUS Osten Sacken, 1876
Atylotus fulvus (Meigen, 1820)
Atylotus plebeius (Fallén, 1817)
Atylotus rusticus (Linnaeus, 1767)
Atylotus sublunaticornis (Zetterstedt, 1842)
HYBOMITRA Enderlein, 1922
Hybomitra arpadi (Szilády, 1923)
Hybomitra astuta (Osten Sacken, 1876) see Notes
= polaris (Frey, 1915)
Hybomitra auripila (Meigen, 1820) see Notes
= aterrima (Meigen, 1820)
Hybomitra bimaculata (Macquart, 1826)
= tropica misid.
?= solstitialis (Meigen, 1820) see Notes
Hybomitra borealis (Fabricius, 1781)
= lapponicus (Wahlberg, 1848)
Hybomitra ciureai (Séguy, 1937)
= schineri Lyneborg, 1959
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172
Hybomitra distinguenda (Verrall, 1909)
Hybomitra kaurii Chvála & Lyneborg, 1970
= borealis misid.
Hybomitra lundbecki Lyneborg, 1959
= fulvicornis misid.
Hybomitra lurida (Fallén, 1817)
Hybomitra montana (Meigen, 1820)
Hybomitra muehlfeldi (Brauer, 1880)
= aviceps (Zetterstedt, 1842)
Hybomitra nigricornis (Zetterstedt, 1842)
Hybomitra nitidifrons (Szilády, 1914)
= connis misid.
Hybomitra sexfasciata (Hine, 1923)
= borealis anderi Kauri, 1951
Hybomitra tarandina (Linnaeus, 1758)
Hybomitra tropica (Linnaeus, 1758)
TABANUS Linnaeus, 1758
Tabanus autumnalis Linnaeus, 1761
Tabanus bovinus Linnaeus, 1758
Tabanus bromius Linnaeus, 1758
Tabanus cordiger Meigen, 1820
Tabanus maculicornis Zetterstedt, 1842
Tabanus sudeticus Zeller, 1842
Checklist part 2: Asilomorpha
suborder Brachycera Macquart, 1834
clade Orthorrapha Brauer, 1863
superfamily Asiloidea Latreille, 1802
ASILIDAE Latreille, 1802
ASILINAE Latreille, 1802
ASILUS Linnaeus, 1758
Asilus crabroniformis Linnaeus, 1758
DIDYSMACHUS Lehr, 1996
Didysmachus picipes (Meigen, 1820)
DYSMACHUS Loew, 1860
Dysmachus trigonus (Meigen, 1804)
MACHIMUS Loew, 1849
Machimus setibarbis Loew, 1849
NEOITAMUS Osten Sacken, 1878
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 173
Neoitamus cothurnatus (Meigen, 1820)
Neoitamus cyanurus (Loew, 1849)
Neoitamus socius (Loew, 1871)
NEOMOCHTHERUS Osten Sacken, 1878
Neomochtherus pallipes (Meigen, 1820)
PAMPONERUS Loew, 1849
Pamponerus germanicus (Linnaeus, 1758)
PHILONICUS Loew, 1849
Philonicus albiceps (Meigen, 1820)
RHADIURGUS Loew, 1849
Rhadiurgus variabilis (Zetterstedt, 1838)
TOLMERUS Loew, 1849
Tolmerus atricapillus (Fallén, 1814)
Tolmerus pyragra (Zeller, 1840)
LAPHRINAE Macquart, 1838
tribe Andrenosomatini Hull, 1962
ANDRENOSOMA Rondani, 1856
Andrenosoma albibarbe (Meigen, 1820)
tribe Laphrini Macquart, 1838
CHOERADES Walker, 1851
Choerades fuliginosus (Panzer, 1798)
Choerades gilvus (Linnaeus, 1758)
Choerades igneus (Meigen, 1820)
Choerades lapponicus (Zetterstedt, 1842)
Choerades marginatus (Linnaeus, 1758)
LAPHRIA Meigen, 1803
Laphria ava (Linnaeus, 1761)
Laphria gibbosa (Linnaeus, 1758)
LEPTOGASTRINAE Schiner, 1862
LEPTOGASTER Meigen, 1803
Leptogaster cylindrica (De Geer, 1776)
Leptogaster guttiventris Zetterstedt, 1842
STENOPOGONINAE Hull, 1962
tribe Dioctriini Hendel, 1936
DIOCTRIA Meigen, 1803
Dioctria atricapilla Meigen, 1804
Dioctria cothurnata Meigen, 1820
Dioctria hyalipennis (Fabricius, 1794)
Dioctria oelandica (Linnaeus, 1758)
Dioctria rupes (De Geer, 1776)
tribe Stegopogonini Hull, 1962
CYRTOPOGON Loew, 1847
Cyrtopogon avimanus (Meigen, 1820)
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174
Cyrtopogon lapponicus (Zetterstedt, 1838)
Cyrtopogon lateralis (Fallén, 1814)
Cyrtopogon luteicornis (Zetterstedt, 1842)
= luteicornis var. pollinosus Frey, 1911
Cyrtopogon pulchripes Loew, 1871
tribe Stichopogonini Hardy, 1930
LASIOPOGON Loew, 1847
Lasiopogon cinctus (Fabricius, 1781)
Lasiopogon septentrionalis Lehr, 1984
BOMBYLIIDAE Latreille, 1802
PHTHIRIINAE Becker, 1913
tribe Phthiriini Becker, 1913
PHTHIRIA Meigen, 1803
Phthiria pulicaria (Mikan, 1796)
BOMBYLIINAE Latreille, 1802
tribe Bombyliini Latreille, 1802
BOMBYLIUS Linnaeus, 1758
sg. Bombylius Linnaeus, 1758
Bombylius discolor Mikan, 1796
Bombylius major Linnaeus, 1758
Bombylius minor Linnaeus, 1758
= allibarbis Zetterstedt, 1842
= albibarbis emend.
SYSTOECHUS Loew, 1855
Systoechus ctenopterus (Mikan, 1796)
= sulphureus (Mikan, 1796)
Systoechus gradatus (Wiedemann, 1820)
ANTHRACINAE Latreille, 1804
tribe Anthracini Latreille, 1804
ANTHRAX Scopoli, 1763
Anthrax anthrax (Schrank, 1781)
Anthrax trifasciatus Meigen, 1804
= leucogaster Wiedemann, 1820
Anthrax varius Fabricius, 1794
tribe Exoprosopini Becker, 1913
EXOPROSOPA Macquart, 1840
Exoprosopa capucina (Fabricius, 1781)
MICOMITRA Bowden, 1964
Micomitra stupida (Rossi, 1790)
tribe Villini Hull, 1973
HEMIPENTHES Loew, 1869
Hemipenthes maura (Linnaeus, 1758)
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 175
Hemipenthes morio (Linnaeus, 1758)
THYRIDANTHRAX Osten Sacken, 1886
yridanthrax fenestratus (Fallén, 1814)
VILLA Lioy, 1864
Villa cingulata (Meigen, 1804)
? Villa halteralis (Kowarz, 1883) see Notes
Villa hottentotta (Linnaeus, 1758)
Villa modesta (Meigen, 1820)
Villa occulta (Wiedemann, 1820)
MYTHICOMYIIDAE Melander, 1902
GLABELLULINAE Cockerell, 1914
GLABELLULA Bezzi, 1902
Glabellula arctica (Zetterstedt, 1838)
SCENOPINIDAE Burmeister, 1835
SCENOPINUS Latreille, 1802
Scenopinus fenestralis (Linnaeus, 1758)
Scenopinus niger (De Geer, 1776)
Scenopinus sp. A see Notes
= vitripennis misid.
THEREVIDAE Newman, 1834
THEREVINAE Newman, 1834
ACROSATHE Irwin & Lyneborg, 1981
Acrosathe annulata (Fabricius, 1805)
DIALINEURA Rondani, 1856
Dialineura anilis (Linnaeus, 1761)
DICHOGLENA Irwin & Lyneborg, 1981
Dichoglena nigripennis (Ruthe, 1831)
PANDIVIRILIA Irwin & Lyneborg, 1981
Pandivirilia eximia (Meigen, 1820)
PSILOCEPHALA Zetterstedt, 1838
Psilocephala imberbis (Fallén, 1814)
SPIRIVERPA Irwin & Lyneborg, 1981
Spiriverpa lunulata (Zetterstedt, 1838)
= clausa (Frey, 1911)
THEREVA Latreille, 1796
ereva cinifera Meigen, 1830
= subfasciata Schummel, 1830
ereva fuscinervis Zetterstedt, 1838
ereva handlirschi Kröber, 1912
= praestans Collin, 1948
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176
ereva inornata Verrall, 1909
ereva lanata Zetterstedt, 1838
ereva microcephala Loew, 1847
ereva nobilitata (Fabricius, 1775)
ereva plebeja (Linnaeus, 1758)
ereva strigata (Fabricius, 1794)
ereva unica (Harris, 1780)
= bipunctata Meigen, 1820
ereva valida Loew, 1847
= circumscripta auct. nec. Loew, 1847
Checklist part 3: families of uncertain position (incertae sedis)
suborder Brachycera Macquart, 1834
clade Orthorrapha Brauer, 1863
? superfamily Nemestrinoidea Grith & Pidgeon, 1832
ACROCERIDAE Leach, 1815
ACROCERA Meigen, 1803
= Paracrocera Mik, 1886
sg. Acrocera Meigen, 1803
Acrocera orbiculus (Fabricius, 1787)
= globulus (Panzer, 1804)
= borealis Zetterstedt, 1838
OGCODES Latreille, 1796
sg. Ogcodes Latreille, 1796
Ogcodes borealis Cole, 1919 see Notes
Ogcodes gibbosus (Linnaeus, 1758)
Ogcodes nigripes (Zetterstedt, 1838) see Notes
Ogcodes pallipes Latreille in Olivier, 1812
Excluded species
Anastoechus nitidulus (Fabricius, 1794) labeling mistake
Beris geniculata Curtis, 1830 misidentied
Cliorismia ardea (Fabricius, 1794) not found within present borders
Cliorismia rustica (Panzer, 1804) not found within present borders
Choerades mbriata (Meigen, 1820) mistake
Choerades ursulus (Loew, 1851) misidentied see Notes
Chrysopilus splendidus (Meigen, 1820) mistake
Cyrtopogon maculipennis (Macquart, 1834) labeling mistake
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha... 177
Epitriptus arthriticus (Zeller, 1840) mistake
Machimus gonatistes (Zeller, 1840) not found within present borders
Odontomyia hydroleon (Linnaeus, 1758) not found within present borders
Pandivirilia nigroanalis (Kröber, 1928) misidentied
Phthiria canescens Loew, 1846 not found within present borders
Tabanus miki Brauer, 1880 misidentied
Tolmerus cingulatus (Fabricius, 1781) mistake
Villa fasciata (Meigen, 1804) not found within present borders
= circumdata (Meigen, 1820)
= venusta (Meigen, 1820)
Villa longicornis Lyneborg, 1965 not found within present borders
Villa panisca (Rossi, 1790) not found within present borders
= circumdata auct. nec. (Meigen, 1820)
Xylophagus matsumurae Miyatake, 1965 misidentied
Notes
Choerades ursulus (Loew, 1851) is a poorly known taxon. It was synonymized with
C. fuliginosus by Lehr (1991) but later considered valid by Bosák and Hradský
(2001). Kahanpää and Winqvist (2005) accepted it as a Finnish species but upon
re-examination we consider it most likely that the single Finnish specimen previ-
ously identied as C. ursulus is a dark male of C. fuliginosus.
Chrysopilus suomianus (Szilády, 1934). e type locality of this species is Enontekiö,
Finland (Szilády 1934). Unfortunately the type material seems lost and the name
is probably best treated as a nomen dubium. Based on Szilády’s original description
it may be a dark form of C. nubecula.
Hybomitra astuta (Osten Sacken, 1876). Kahanpää and Winqvist (2005) could not
locate any material in Finnish collections. Several new records of this species have
since been made and its presence in Finland is now conrmed.
Hybomitra auripila (Meigen, 1820). e synonymy of Hybomitra auripila (Meigen,
1820) with H. aterrima (Meigen, 1820) was established by Schacht (1994) and is
accepted here. Schiner (1862) already mentioned H. aterrima as synonym to H.
auripila. Since we consider him to be the rst revisor, the name H. auripila is valid
under the current Code.
Hybomitra solstitialis (Meigen, 1820) has long been known to be a problematic tax-
on. It is separated from H. bimaculata (Macquart, 1826) based on color characters
alone. e examined Finnish material includes a range of intermediates between
typical H. bimaculata and H. solstitialis forms. It seems likely that the two names are
synonymous, but types should be consulted before synonymy is formally published.
Haematopota subcylindrica Pandellé, 1883. First recorded from Finland by Vuori-
mies (1984). Unfortunately the specimens listed in his paper could not be found
and their identication remains somewhat doubtful.
Jere Kahanpää et al. / ZooKeys 441: 165–181 (2014)
178
Ogcodes borealis Cole, 1919. A single Finnish specimen collected in the mid-19th
century is the sole Palearctic record of this species. Originally identied and pub-
lished by Hackman (1970), the record was later conrmed by Kahanpää and Win-
qvist (2005). O. borealis Cole sensu Schlinger (1960) may be a species complex.
Ogcodes nigripes (Zetterstedt, 1838) is probably a senior synonym of O. zonatus
Erichson, 1840.
Ptiolina nigrina Wahlgren, 1854 may be a synonym of P. nigra Zetterstedt, 1842.
Scenopinus sp. A is an apparently undescribed species near S. fenestralis with black
femora. It occurs widely in Finland in association with bird nests.
Villa halteralis (Kowarz, 1883). See Kahanpää and Winqvist (2005) for a discussion
of the single supposed Finnish record of this species.
Xylophagus ater Meigen, 1804. is name has widely been used for two species. Old
Finnish checklists (Frey et al. 1941, Hackman 1980) followed the model also used
in the world checklist Woodley (2011c) and used this name for the species also
known as Xylophagus kowarzi (Pleske, 1925). On the British Isles the name X. ater
is used as a senior name for X. compeditus Wiedemann in Meigen, 1820. Accord-
ing to Alexander and Clements (1991) and Chandler (1998a, b) the British usage
is correct and it is followed here. us, X. ater is the common species with females
easily identied by the three stripes of dusting on the mesonotum.
Xylophagus inermis Krivosheina & Krivosheina, 2000 was described as a subspecies
of X. matsumurae Miyatake, 1965 = maculatus Matsumura, 1916 (preoccupied by
X. maculatus Meigen, 1804) (Krivosheina and Krivosheina 2000). It was raised to
a full species status in the recent world catalogue (Woodley 2011c). All collected
Finnish specimens formerly identied as X. matsumurae were examined and they
belong to X. inermis.
Zabrachia tenella (Jaennicke, 1866). First recorded from Finland by Krivosheina
and Rozkošný (1990). We have examined the Finnish Zabrachia material and
conrmed the presence of both Z. tenella and Z. minutissima in the country.
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Checklist of the Empidoidea of Finland (Insecta,Diptera)183
Checklist of the Empidoidea of Finland
(Insecta, Diptera)
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 28 January 2014|Accepted 7 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the Empidoidea of Finland (Insecta,Diptera). In: Kahanpää J, Salmela J (Eds)
Checklist of the Diptera of Finland. ZooKeys 441: 183–207. doi: 10.3897/zookeys.441.7154
Abstract
An updated checklist of the Atelestidae, Brachystomatidae, Dolichopodidae, Empididae and Hybotidae
(Diptera) recorded from Finland is presented. e genera with uncertain placement within superfamily
Empidoidea (= the Iteaphila group) are also included in this paper.
Keywords
Checklist, Finland, Diptera, Atelestidae, Brachystomatidae, Dolichopodidae, Hybotidae, Empididae
Introduction
Empidoidea (dance ies and long-legged ies) is one of the most diverse lineages of
Diptera with some 13,000 species currently recognized worldwide (Pape et al. 2011)
and many more still undescribed. e superfamily is generally accepted as monophy-
letic, but its subdivision into families, subfamilies and tribes is still being debated.
e families Atelestidae, Brachystomatidae, Hybotidae and Empididae were tradi-
tionally all included in Empididae sensu lato. Chvála (1983) split the empidids into ve
ZooKeys 441: 183–207 (2014)
doi: 10.3897/zookeys.441.7154
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 183–207 (2014)
184
families: Empididae sensu stricto, Hybotidae, Atelestidae, Dolichopodidae and Micro-
phoridae. Sinclair and Cumming (2006) analyzed the whole superfamily Empidoidea,
elevated Brachystomatidae to a full family and transferred Microphorinae to the long-
legged ies (Dolichopodidae). Two of the Finnish genera, Anthepiscopus Becker and
Iteaphila Zetterstedt, remain unassigned on family level (Sinclair and Cumming 2006,
Shamshev and Sinclair 2009). e classication proposed by Sinclair and Cumming
(2006) is adopted for this checklist.
e world fauna of the Empidoidea has recently been catalogued by Yang et al.
(2006, 2007). e Finnish species were last listed by Kahanpää and Winqvist (2005)
for Atelestidae, Brachystomatidae, Hybotidae and Empididae (plus Microphorinae)
and by Kahanpää and Grichanov (2004) for Dolichopodidae. Two updates to the
dolichopodid checklist have since been published (Kahanpää 2006, 2013).
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
superfamily Empidoidea Latreille, 1809
Uncertain family position within Empidoidea (incertae familiae)
ITEAPHILA Zetterstedt, 1838
Iteaphila macquarti Zetterstedt, 1838
Iteaphila nitidula Zetterstedt, 1838
= obscura Zetterstedt, 1849
ANTHEPISCOPUS Becker, 1891
Anthepiscopus oedalinus (Zetterstedt, 1838)
EMPIDIDAE Latreille, 1809
HEMERODROMIINAE Schiner, 1862
Chelipodini Hendel, 1936
CHELIPODA Macquart, 1823
Chelipoda albiseta (Zetterstedt, 1838)
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Atelestidae 11 4 2 average
Brachystomatidae 147 13 4 average
Dolichopodidae 7235 790 265 good
Hybotidae 1972 442 143–144 average
Empididae 3049 803 175 average
Checklist of the Empidoidea of Finland (Insecta,Diptera)185
Chelipoda inexspectata Tuomikoski, 1966
Chelipoda vocatoria (Fallén, 1816)
PHYLLODROMIA Zetterstedt, 1837
Phyllodromia melanocephala (Fabricius, 1794)
Hemerodromiini Schiner, 1862
CHELIFERA Macquart, 1823
Chelifera concinnicauda Collin, 1927
= lapponica Frey, 1950
= stigmatica misid.
Chelifera avella (Zetterstedt, 1838)
Chelifera frigelii (Zetterstedt, 1838)
Chelifera precabunda Collin, 1961
Chelifera precatoria (Fallén, 1816)
Chelifera subangusta Collin, 1961
Chelifera trapezina (Zetterstedt, 1838)
HEMERODROMIA Meigen, 1822
Hemerodromia adulatoria Collin, 1927
Hemerodromia oratoria (Fallén, 1816)
Hemerodromia raptoria Meigen, 1830
EMPIDINAE Latreille, 1809
Empidini Latreille, 1809
EMPIS Linnaeus, 1758
sg. Anacrostichus Bezzi, 1909
Empis lucida Zetterstedt, 1838
sg. Coptophlebia Bezzi, 1909
Empis hyalipennis Fallén, 1816
sg. Empis Linnaeus, 1758
Empis acinerea Chvala, 1985
= cinerea Zetterstedt, 1855 preocc.
Empis bicuspidata Collin, 1927
Empis caudatula Loew, 1867
Empis chioptera Meigen, 1804
Empis laminata Collin, 1927
Empis nigripes Fabricius, 1794
= pennaria Fallén, 1816
= vernalis Meigen, 1822
Empis pennipes Linnaeus, 1758
Empis prodromus Loew, 1867
Empis staegeri Collin, 1963
= planetica misid.
Empis syrovatkai Chvála, 1985
= plumipes Zetterstedt, 1842 preocc.
sg. Euempis Frey, 1953
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Empis picipes Meigen, 1804
= maculipes Zetterstedt, 1842
Empis tessellata Fabricius, 1794
sg. Kritempis Collin, 1926
Empis livida Linnaeus, 1758
sg. Leptempis Collin, 1926
Empis grisea Fallén, 1816
sg. Platyptera Meigen, 1803
Empis borealis Linnaeus, 1758
sg. Xanthempis Bezzi, 1909
Empis aemula Loew, 1873
Empis digramma Meigen in Gistl, 1835
= diagramma misspelling
Empis laetabilis Collin, 1926
Empis stercorea Linnaeus, 1761
Empis univittata Loew, 1867
RHAMPHOMYIA Meigen, 1822
sg. Aclonempis Collin, 1926
Rhamphomyia galactoptera Strobl, 1893
sg. Amydroneura Collin, 1926
Rhamphomyia gibba (Fallén, 1816)
sg. Holoclera Schiner, 1860
Rhamphomyia bohemica Barták & Kubík, 2012
= caliginosa misid.
= trigemina misid.
Rhamphomyia culicina (Fallén, 1816)
Rhamphomyia nigripennis (Fabricius, 1794)
Rhamphomyia sciarina (Fallén, 1816)
Rhamphomyia umbripennis Meigen, 1822
Rhamphomyia variabilis (Fallén, 1816)
= tenuirostris (Fallén, 1816)
sg. Lundstroemiella Frey, 1922
Rhamphomyia dudai Oldenberg, 1927
Rhamphomyia hybotina Zetterstedt, 1838
sg. Megacyttarus Bigot, 1880
Rhamphomyia anomalina Zetterstedt, 1838
Rhamphomyia anomalipennis Meigen, 1822
= anomala misid.
Rhamphomyia crassirostris (Fallén, 1816)
= nigripes auct. nec (Fabricius, 1794)
Rhamphomyia gutar Frey, 1922
Rhamphomyia maculipennis Zetterstedt, 1842
= tephraea misid.
Checklist of the Empidoidea of Finland (Insecta,Diptera)187
= poissoni misid.
Rhamphomyia nodipes (Fallén, 1816)
= spissirostris (Fallén, 1816)
Rhamphomyia paradoxa Wahlberg, 1844
sg. Pararhamphomyia Frey, 1922
Rhamphomyia albidiventris Strobl, 1898
= woldstedti Frey, 1913
Rhamphomyia albipennis (Fallén, 1816)
Rhamphomyia albitarsis Collin, 1926
Rhamphomyia albissima Frey, 1913
Rhamphomyia alpina Zetterstedt, 1838
Rhamphomyia amoena Loew, 1840
Rhamphomyia angulifera Frey, 1913
Rhamphomyia breviventris Frey, 1913
Rhamphomyia caesia Meigen, 1822
= lata Zetterstedt, 1842
Rhamphomyia caudata Zetterstedt, 1838
= aethiops Zetterstedt, 1838
Rhamphomyia chibinensis Frey, 1922
Rhamphomyia cribrata Oldenberg, 1927
Rhamphomyia curvula Frey, 1913
Rhamphomyia dentata Oldenberg, 1910
Rhamphomyia fascipennis Zetterstedt, 1838
Rhamphomyia licaudula Frey, 1949
Rhamphomyia fuscipennis Zetterstedt, 1838
Rhamphomyia fuscula Zetterstedt, 1838
Rhamphomyia geniculata Meigen, 1830
= plumipes auct. nec (Meigen, 1804)
Rhamphomyia griseola (Zetterstedt, 1838)
= dispar Zetterstedt, 1838
= aperta Zetterstedt, 1859
Rhamphomyia helleni Frey, 1922
Rhamphomyia lividiventris Zetterstedt, 1838
Rhamphomyia longestylata Frey, 1916
= caudata auct. nec Zetterstedt, 1838
Rhamphomyia lucidula Zetterstedt, 1842
Rhamphomyia marginata (Fabricius, 1787)
Rhamphomyia modesta Wahlberg, 1844
Rhamphomyia murina Collin, 1926
Rhamphomyia niveipennis Zetterstedt, 1838
Rhamphomyia obscura Zetterstedt, 1838
Rhamphomyia obscuripennis Meigen, 1830
= nitidicollis Frey, 1913
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Rhamphomyia physoprocta Frey, 1913
Rhamphomyia pilifer Meigen, 1838
= dentipes Zetterstedt, 1842
= glaucella Zetterstedt, 1842
= intermedia Frey, 1922
Rhamphomyia plumifera Zetterstedt, 1838
Rhamphomyia poplitea Wahlberg, 1844
Rhamphomyia praestans Frey, 1913
Rhamphomyia pusilla Zetterstedt, 1838
Rhamphomyia rupes Zetterstedt, 1838
= lapponica Frey, 1955
Rhamphomyia simplex Zetterstedt, 1849
Rhamphomyia subglaucella Frey, 1922
Rhamphomyia tibiella Zetterstedt, 1842
Rhamphomyia tipularia (Fallén, 1816)
Rhamphomyia unguiculata Frey, 1913
Rhamphomyia sp. a. albipennis
sg. Rhamphomyia Meigen, 1822
Rhamphomyia albosegmentata Zetterstedt, 1838
Rhamphomyia cinerascens (Meigen, 1804)
Rhamphomyia coracina Zetterstedt, 1849
Rhamphomyia dorsata Becker, 1915
= phanerostigma Frey, 1918
Rhamphomyia hambergi Frey, 1916
Rhamphomyia ignobilis Zetterstedt, 1859
= attenuata Frey, 1913
Rhamphomyia laevipes (Fallén, 1816)
= tephraea Meigen, 1822
Rhamphomyia latifrons Frey, 1913
Rhamphomyia morio Zetterstedt, 1838
Rhamphomyia nitidula Zetterstedt, 1842
Rhamphomyia palmeni Frey, 1913
Rhamphomyia plumipes (Meigen, 1804)
= vespertilio Zetterstedt, 1842
Rhamphomyia reexa Zetterstedt, 1838
Rhamphomyia spinipes (Fallén, 1816)
Rhamphomyia stigmosa Macquart, 1827
= conformis auct. nec Kowarz, 1867
Rhamphomyia sulcata (Meigen, 1804)
= tibialis auct. nec Meigen, 1822
Rhamphomyia trilineata Zetterstedt, 1859
= sulcatina Collin, 1926
= tibialis auct. nec Meigen, 1822
Checklist of the Empidoidea of Finland (Insecta,Diptera)189
= propinqua misid.
Rhamphomyia vesiculosa (Fallén, 1816)
Hilarini Collin, 1961
HILARA Meigen, 1822
Hilara abdominalis Zetterstedt, 1838
= obscuritarsis Zetterstedt, 1859
Hilara albitarsis von Roser, 1840
Hilara anglodanica Lundbeck, 1913
Hilara barbipes Frey, 1908
Hilara beckeri Strobl, 1892
Hilara biseta Collin, 1927
Hilara bistriata Zetterstedt, 1842
Hilara brevistyla Collin, 1927
Hilara campinosensis Niesiolowski, 1986
Hilara canescens Zetterstedt, 1849
Hilara chorica (Fallén, 1816)
Hilara clavipes (Harris, 1776)
= spinimana Zetterstedt, 1838
Hilara clypeata Meigen, 1822
Hilara coracina Oldenberg, 1916
= quadrifaria Strobl, 1892
Hilara cornicula Loew, 1873
Hilara discoidalis Lundbeck, 1910
Hilara diversipes Strobl, 1892
= germanica Engel, 1941
Hilara eviana Straka, 1976
Hilara femorella Zetterstedt, 1842
Hilara gallica (Meigen, 1804)
Hilara griseola Zetterstedt, 1838
Hilara hirta Strobl, 1892
= hirtella misid.
Hilara hybrida Collin, 1961
Hilara hyposeta Straka, 1976
Hilara implicata Collin, 1927
Hilara intermedia (Fallén, 1816)
= pubipes Loew, 1873
Hilara interstincta (Fallén, 1816)
Hilara monedula Collin, 1927
= longifurca auct. nec Strobl, 1892
Hilara lapponica Chvála, 2002
Hilara litorea (Fallén, 1816)
Hilara longivittata Zetterstedt, 1842
Hilara lurida (Fallén, 1816)
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Hilara nigritarsis Zetterstedt, 1838
= infans Zetterstedt, 1842
Hilara nitidula Zetterstedt, 1838
Hilara pilipes Zetterstedt, 1838
Hilara pseudochorica Strobl, 1892
Hilara pulchripes Frey, 1913
Hilara quadrifasciata Chvála, 2002
= quadrivittata auct. nec Meigen, 1822
Hilara sturmii Meigen, 1822
= cingulata misid.
Hilara submaura Collin, 1927
Hilara tanythrix Frey, 1913
Hilara tenuinervis Zetterstedt, 1838
Hilara woodiella Chvála, 1999
CLINOCERINAE Collin, 1928
CLINOCERA Meigen, 1803
Clinocera appendiculata (Zetterstedt, 1838)
= simplicinervis Frey, 1913
Clinocera aucta (Zetterstedt, 1849)
Clinocera nivalis (Zetterstedt, 1838)
Clinocera stagnalis (Haliday, 1833)
Clinocera wesmaeli (Macquart, 1835)
DOLICHOCEPHALA Macquart, 1823
Dolichocephala guttata (Haliday, 1833)
Dolichocephala irrorata (Fallén, 1815)
Dolichocephala thomasi Wagner, 1983
= ocellata misid.
WIEDEMANNIA Zetterstedt, 1838
sg. Eucelidia Mik, 1881
Wiedemannia zetterstedti (Fallén, 1826)
sg. Philolutra Mik, 1881
Wiedemannia bohemani (Zetterstedt, 1838)
Wiedemannia simplex (Loew, 1862)
= fallaciosa auct. nec Loew, 1873
sg. Wiedemannia Zetterstedt, 1838
Wiedemannia bistigma (Curtis, 1834)
Uncertain subfamily position within Empididae (incertae sedis)
HORMOPEZA Zetterstedt, 1838
Hormopeza copulifera Melander, 1927
Hormopeza obliterata Zetterstedt, 1838
RAGAS Walker, 1837
Ragas unica Walker, 1837
Checklist of the Empidoidea of Finland (Insecta,Diptera)191
ATELESTIDAE Hennig, 1970
ATELESTINAE Hennig, 1970
ATELESTUS Walker, 1837
Atelestus pulicarius (Fallén, 1816)
MEGHYPERUS Loew, 1850
Meghyperus sudeticus Loew, 1850
HYBOTIDAE Meigen, 1820
TRICHININAE Chvála, 1983
TRICHINA Meigen, 1830
Trichina bilobata Collin, 1926
Trichina clavipes Meigen, 1830
Trichina elongata Haliday, 1833
Trichina opaca Loew, 1864
= picipes Tuomikoski, 1935
Trichina pallipes (Zetterstedt, 1838)
TRICHINOMYIA Tuomikoski, 1959
Trichinomyia avipes (Meigen, 1830)
Trichinomyia fuscipes (Zetterstedt, 1838)
OCYDROMIINAE Schiner, 1862
Ocydromiini Schiner, 1862
CHVALAEA Papp & Földvári, 2001
Chvalaea sopianae Papp & Földvári, 2001
LEPTODROMIELLA Tuomikoski, 1936
Leptodromiella crassiseta Tuomikoski, 1936
LEPTOPEZA Macquart, 1834
Leptopeza borealis Zetterstedt, 1842
Leptopeza avipes (Meigen, 1820)
OCYDROMIA Meigen, 1820
Ocydromia glabricula (Fallén, 1816)
Ocydromia melanopleura Loew, 1840
OEDALEINAE Chvála, 1983
ALLANTHALIA Melander, 1927
Allanthalia pallida (Zetterstedt, 1838)
ANTHALIA Zetterstedt, 1838
Anthalia schoenherri Zetterstedt, 1838
EUTHYNEURA Macquart, 1836
Euthyneura albipennis (Zetterstedt, 1842)
Euthyneura gyllenhali (Zetterstedt, 1838)
Euthyneura myrtilli Macquart, 1836
= myricae Haliday, 1851 misid.
OEDALEA Meigen, 1820
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Oedalea freyi Chvála, 1983
Oedalea holmgreni Zetterstedt, 1852
Oedalea hybotina (Fallén, 1816)
Oedalea stigmatella Zetterstedt, 1842
Oedalea tibialis Macquart, 1827
Oedalea zetterstedti Collin, 1926
TACHYDROMIINAE Meigen, 1822
Symballophthalmini Sinclair & Cumming, 2006
SYMBALLOPHTHALMUS Mecker, 1889
Symballophthalmus dissimilis (Fallén, 1815)
Symballophthalmus fuscitarsis (Zetterstedt, 1859)
= scapularis Collin, 1961
Tachydromiini Meigen, 1822
PLATYPALPUS Macquart, 1827
Platypalpus agilis (Meigen, 1822)
Platypalpus albicornis (Zetterstedt, 1842)
Platypalpus albiseta (Panzer, 1806)
Platypalpus albocapillatus (Fallén, 1815)
Platypalpus alpinus Chvála, 1971
Platypalpus alter (Collin, 1961)
Platypalpus annulatus (Fallén, 1815)
= fulvipes (Meigen, 1822)
Platypalpus annulipes (Meigen, 1822)
Platypalpus articulatoides (Frey, 1918)
? Platypalpus articulatus Macquart, 1827
Platypalpus ater (Wahlberg, 1844)
= atra misspelling
Platypalpus boreoalpinus Frey, 1943
Platypalpus brachystylus (Bezzi, 1892)
= brunneitibia (Strobl, 1899)
Platypalpus brevicornis (Zetterstedt, 1842)
Platypalpus calceatus (Meigen, 1822)
Platypalpus candicans (Fallén, 1815)
Platypalpus ciliaris (Fallén, 1816)
Platypalpus conformis Chvála, 1971
Platypalpus connis (Zetterstedt, 1842)
Platypalpus cothurnatus Macquart, 1827
Platypalpus cryptospina (Frey, 1909)
= tantulus (Collin, 1961)
Platypalpus cursitans (Fabricius, 1775)
= bicolor (Meigen, 1804)
Platypalpus ecalceatus (Zetterstedt, 1838)
Platypalpus excavatus Yang & Yao, 2007
Checklist of the Empidoidea of Finland (Insecta,Diptera)193
= excisus (Becker, 1907) preocc.
Platypalpus exilis (Meigen, 1822)
Platypalpus fenestella Kovalev, 1971
Platypalpus avicornis (Meigen, 1822)
Platypalpus fuscicornis (Zetterstedt, 1842)
Platypalpus hackmani Chvála, 1972
Platypalpus infectus (Collin, 1926)
Platypalpus interstinctus (Collin, 1926)
Platypalpus laestadianorum (Frey, 1913)
Platypalpus lapponicus Frey, 1943
Platypalpus longicornis (Meigen, 1822)
Platypalpus longiseta (Zetterstedt, 1842)
= extricatus (Collin, 1926)
Platypalpus luteicornis (Meigen, 1838)
= dicilis (Frey, 1907)
= interjectus (Lundbeck, 1910)
Platypalpus luteus (Meigen, 1804)
Platypalpus maculus (Zetterstedt, 1842)
Platypalpus maculimanus (Zetterstedt, 1842)
= articulatus auct. nec Macquart, 1827
Platypalpus maculipes (Meigen, 1822)
Platypalpus major (Zetterstedt, 1842)
Platypalpus melancholicus (Collin, 1961)
Platypalpus minutus (Meigen, 1804)
Platypalpus nigricoxa (Mik, 1884)
Platypalpus nigritarsis (Fallén, 1816)
Platypalpus nigrosetosus (Strobl, 1893)
Platypalpus nonstriatus Strobl, 1901
Platypalpus notatus (Meigen, 1822)
Platypalpus pallidicornis (Collin, 1926)
Platypalpus pallidicoxa (Frey, 1913)
Platypalpus pallidiventris (Meigen, 1822)
= avipes (Fabricius, 1794)
Platypalpus pallipes (Fallén, 1815)
Platypalpus pectoralis (Fallén, 1815)
Platypalpus pseudofulvipes (Frey, 1909)
= coarctatus (Collin, 1926)
Platypalpus pseudorapidus Kovalev, 1971
Platypalpus pulicarius (Meigen, 1830)
Platypalpus rapidus (Meigen, 1822)
Platypalpus sahlbergi (Frey, 1909)
Platypalpus scandinavicus Chvála, 1972
Platypalpus sordidus (Zetterstedt, 1838)
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Platypalpus stabilis (Collin, 1961)
Platypalpus stackelbergi Kovalev, 1971
Platypalpus stigmatellus (Zetterstedt, 1842)
Platypalpus strigifrons (Zetterstedt, 1849)
Platypalpus subbrevis (Frey, 1913)
Platypalpus subtilis (Collin, 1926)
Platypalpus sylvicola (Collin, 1926)
Platypalpus tuomikoskii Chvála, 1972
Platypalpus unguiculatus (Zetterstedt, 1838)
Platypalpus verralli (Collin, 1926)
Platypalpus vividus (Meigen, 1838)
= albisetoides Chvála, 1973
Platypalpus zetterstedti Chvála, 1971
TACHYDROMIA Meigen, 1803
Tachydromia aemula (Loew, 1864)
Tachydromia arrogans (Linnaeus, 1761)
Tachydromia connexa Meigen, 1822
Tachydromia incompleta (Becker, 1900)
Tachydromia lundstroemi (Frey, 1913)
Tachydromia morio (Zetterstedt, 1838)
Tachydromia punctifera (Becker, 1900)
Tachydromia sabulosa Meigen, 1830
Tachydromia terricola Zetterstedt, 1819
Tachydromia umbrarum Haliday, 1833
TACHYPEZA Meigen, 1830
Tachypeza fennica Tuomikoski, 1932
Tachypeza fuscipennis (Fallén, 1815)
Tachypeza heeri (Zetterstedt, 1838)
Tachypeza nubila (Meigen, 1804)
Tachypeza truncorum (Fallén, 1815)
Tachypeza winthemi (Zetterstedt, 1838)
Drapetini Collin, 1961
CHERSODROMIA Haliday, 1851
Chersodromia arenaria (Haliday, 1833)
Chersodromia cursitans (Zetterstedt, 1819)
CROSSOPALPUS Bigot, 1857
Crossopalpus curvinervis (Zetterstedt, 1842)
Crossopalpus curvipes (Meigen, 1822)
Crossopalpus humilis (Frey, 1913)
Crossopalpus nigritellus (Zetterstedt, 1842)
Crossopalpus setiger (Loew, 1859)
DRAPETIS Meigen, 1822
Drapetis arcuata Loew, 1859
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Drapetis assimilis (Fallén, 1815)
Drapetis exilis Meigen, 1822
Drapetis intialis Collin, 1961
Drapetis ingrica Kovalev, 1972
Drapetis parilis Collin, 1926
Drapetis pusilla Loew, 1859
Drapetis simulans Collin, 1961
Elaphropeza Macquart, 1827
Elaphropeza ephippiata (Fallén, 1815)
STILPON Loew, 1859
Stilpon graminum (Fallén, 1815)
HYBOTINAE Meigen, 1820
Bicellariini Sinclair & Cumming, 2006
BICELLARIA Macquart, 1823
= Cyrtoma Meigen, 1824
Bicellaria austriaca Tuomikoski, 1955
Bicellaria intermedia Lundbeck, 1910
Bicellaria nigra (Meigen, 1824)
Bicellaria pilosa Lundbeck, 1910
Bicellaria simplicipes (Zetterstedt, 1842)
Bicellaria spuria (Fallén, 1816)
Bicellaria subpilosa Collin, 1926
Bicellaria sulcata (Zetterstedt, 1842)
Bicellaria uvens Melander, 1928
= bisetosa Tuomikoski, 1936
Hybotini Meigen, 1820
HYBOS Meigen, 1803
Hybos culiciformis (Fabricius, 1775)
Hybos femoratus (Müller, 1776)
= femoralis misspelling
Hybos grossipes (Linnaeus, 1767)
SYNDYAS Loew, 1857
Syndyas nigripes (Zetterstedt, 1842)
BRACHYSTOMATIDAE Melander, 1908
TRICHOPEZINAE Vaillant, 1981
GLOMA Meigen, 1822
Gloma fuscipennis Meigen, 1822
HELEODROMIA Haliday, 1833
sg. Heleodromia Haliday, 1833
Heleodromia immaculata Haliday, 1833
TRICHOPEZA Rondani, 1856
Trichopeza albocincta (Boheman, 1863)
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= albicincta Frey, 1913
Trichopeza longicornis (Meigen, 1822)
DOLICHOPODIDAE Latreille, 1809
MICROPHORINAE Collin, 1960
MICROPHOR Macquart, 1827
= Microphorus misspelling
Microphor anomalus (Meigen, 1824)
Microphor crassipes Macquart, 1827
Microphor holosericeus (Meigen, 1804)
= velutinus Macquart, 1827
PARATHALASSIINAE Chvála, 1981
MICROPHORELLA Becker, 1909
Microphorella praecox (Loew, 1864)
DOLICHOPODINAE Latreille, 1809
DOLICHOPUS Latreille, 1796
= Hygroceleuthus Loew, 1857
sg. Dolichopus Latreille, 1796
Dolichopus acuticornis Wiedemann, 1817
Dolichopus annulitarsis Ringdahl, 1920
Dolichopus apicalis Zetterstedt, 1849
Dolichopus argyrotarsis Wahlberg, 1850
Dolichopus armillatus Wahlberg, 1850
= stenhammari var. b Zetterstedt, 1843
Dolichopus atripes Meigen, 1824
Dolichopus austriacus Parent, 1927
Dolichopus bonsdor Frey, 1915
Dolichopus brevipennis Meigen, 1824
Dolichopus caligatus Wahlberg, 1850
= avipes misid.
= albifrons misid.
Dolichopus calinotus Loew, 1871
Dolichopus campestris Meigen, 1824
Dolichopus cilifemoratus Macquart, 1827
= pseudocilifemoratus Stackelberg, 1930
Dolichopus cinctipes Wahlberg, 1850
Dolichopus claviger Stannius, 1831
Dolichopus clavipes Haliday, 1832
Dolichopus costalis Frey, 1915
Dolichopus discifer Stannius, 1831
?= nigricornis Meigen, 1824
Dolichopus discimanus Wahlberg, 1851
Dolichopus fraterculus Zetterstedt, 1843
Checklist of the Empidoidea of Finland (Insecta,Diptera)197
Dolichopus griseipennis Stannius, 1831
Dolichopus gubernator Mik, 1878
Dolichopus hilaris Loew, 1862
Dolichopus lancearius Hedström, 1966
Dolichopus latilimbatus Macquart, 1827
Dolichopus latipennis Fallén, 1823
Dolichopus lepidus Staeger, 1842
= cruralis Wahlberg, 1850
= lapponicus Becker, 1917
Dolichopus linearis Meigen, 1824
Dolichopus lineatocornis Zetterstedt, 1843
Dolichopus longicornis Stannius, 1831
Dolichopus longitarsis Stannius, 1831
Dolichopus maculipennis Zetterstedt, 1843
Dolichopus mannerheimi Zetterstedt, 1838
Dolichopus migrans Zetterstedt, 1843
Dolichopus nigripes Fallén, 1823
Dolichopus nitidus Fallén, 1823
Dolichopus notatus Staeger, 1842
= notabilis Zetterstedt, 1843
Dolichopus nubilus Meigen, 1824
Dolichopus pennatus Meigen, 1824
Dolichopus picipes Meigen, 1824
= consimilis Wahlberg, 1850
Dolichopus planitarsis Fallén, 1823
Dolichopus plumipes (Scopoli, 1763)
= parvicaudatus Zetterstedt, 1843
= pectinitarsis Stenhammar, 1851
Dolichopus popularis Wiedemann, 1817
Dolichopus pseudomigrans Ringdahl, 1928
Dolichopus punctum Meigen, 1824
Dolichopus remipes Wahlberg, 1839
Dolichopus rupestris Haliday, 1833
Dolichopus ruthei Loew, 1847
Dolichopus sabinus Haliday, 1838
Dolichopus setiger Negrobov, 1973
Dolichopus signatus Meigen, 1824
Dolichopus signifer Haliday, 1832
Dolichopus simplex Meigen, 1824
Dolichopus stenhammari Zetterstedt, 1843
= annulipes Zetterstedt, 1838 invalidated
Dolichopus subpennatus d’Assis-Fonseca, 1976
Dolichopus trivialis Haliday, 1832
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= cilifemoratus auct. nec Macquart, 1827
Dolichopus ungulatus (Linnaeus, 1758)
Dolichopus urbanus Meigen, 1824
Dolichopus vitripennis Meigen, 1824
Dolichopus wahlbergi Zetterstedt, 1843
Dolichopus zetterstedti Stenhammar, 1852
sg. Macrodolichopus Stackelberg, 1933
Dolichopus diadema Haliday, 1832
ETHIROMYIA Brooks, 2005
Ethiromyia chalybeus (Wiedemann, 1817)
GYMNOPTERNUS Loew, 1857
Gymnopternus aerosus (Fallén, 1823)
Gymnopternus angustifrons (Staeger, 1842)
Gymnopternus brevicornis (Staeger, 1842)
Gymnopternus celer (Meigen, 1824)
Gymnopternus metallicus (Stannius, 1831)
HERCOSTOMUS Loew, 1857
Hercostomus germanus (Wiedemann, 1817)
Hercostomus nigrilamellatus (Macquart, 1827)
Hercostomus nigriplantis (Stannius, 1831)
Hercostomus sahlbergi (Zetterstedt, 1838)
SYBISTROMA Meigen, 1824
Sybistroma discipes (Germar, 1817)
Sybistroma obscurellum (Fallén, 1823)
TACHYTRECHUS Haliday, 1851
= Ammobates Stannius, 18312 preocc.
Tachytrechus ammobates (Haliday, 1851)
= plumipes (Fallén, 1823) preocc.
Tachytrechus hamatus Loew, 1871
Tachytrechus notatus (Stannius, 1831)
SCIAPODINAE Becker, 1917
SCIAPUS Zeller, 1842
= Sciopus misspelling
Sciapus albifrons (Meigen, 1830)
Sciapus basilicus Meuels & Grootaert, 1990
Sciapus lobipes (Meigen, 1824)
Sciapus longulus (Fallén, 1823)
Sciapus maritimus Becker, 1918
= avomaculatus Ringdahl, 1949
Sciapus platypterus (Fabricius, 1805)
Sciapus wiedemanni (Fallén, 1823)
Sciapus zonatulus (Zetterstedt, 1843)
= contristans misid.
Checklist of the Empidoidea of Finland (Insecta,Diptera)199
SYMPYCNINAE Aldrich, 1905
CAMPSICNEMUS Haliday, 1851
= Ectomus Mik, 1878
Campsicnemus alpinus (Haliday, 1833)
Campsicnemus armatus (Zetterstedt, 1849)
Campsicnemus articulatellus (Zetterstedt, 1843)
= pilosellus (Zetterstedt, 1843)
= dasycnemus Loew, 1857
Campsicnemus compeditus Loew, 1857
Campsicnemus curvipes (Fallén, 1823)
Campsicnemus femoratus Ringdahl, 1949
Campsicnemus loripes (Haliday, 1832)
Campsicnemus lumbatus Loew, 1857
Campsicnemus marginatus Loew, 1857
Campsicnemus paradoxus (Wahlberg, 1844)
Campsicnemus picticornis (Zetterstedt, 1843)
Campsicnemus pumilio (Zetterstedt, 1843)
= pectinulatus Loew, 1864
Campsicnemus pusillus (Meigen, 1824)
Campsicnemus scambus (Fallén, 1823)
LAMPROCHROMUS Mik, 1878
Lamprochromus strobli Parent, 1925
SYMPYCNUS Loew, 1857
Sympycnus aeneicoxa (Meigen, 1824)
Sympycnus pulicarius (Fallén, 1823)
?= annulipes (Meigen, 1824)
?= desoutteri Parent, 1925
SYNTORMON Loew, 1857
= Bathycranium Strobl, 1892
Syntormon bicolorellus (Zetterstedt, 1843)
Syntormon liger Verrall, 1912
= rupes misid.
= simplicipes Frey, 1915
Syntormon freymuthae Loew, 1873
= S. denticulatus misid.
Syntormon metathesis (Loew, 1850)
Syntormon pallipes (Fabricius, 1794)
Syntormon pumilus (Meigen, 1824)
Syntormon tarsatus (Fallén, 1823)
= aulicus misid.
TELMATURGUS Mik, 1874
Telmaturgus tumidulus (Raddatz, 1873)
TEUCHOPHORUS Loew, 1857
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= Teucophorus misspelling
Teuchophorus monacanthus Loew, 1859
Teuchophorus nigricosta (von Roser, 1840)
= pectinifer Kowarz, 1868
= signatus Zetterstedt, 1849
Teuchophorus spinigerellus (Zetterstedt, 1843)
DIAPHORINAE Schiner, 1864
ARGYRA Macquart, 1834
= Leucostola Loew, 1857
Argyra argentina (Meigen, 1824)
Argyra argyria (Meigen, 1824)
Argyra auricollis (Meigen, 1824)
Argyra diaphana (Fabricius, 1775)
Argyra elongata (Zetterstedt, 1843)
Argyra ilonae Geories, 1989
= connis (Zetterstedt, 1849) preocc.
Argyra leucocephala (Meigen, 1824)
Argyra magnicornis (Zetterstedt, 1838)
Argyra setulipes Becker, 1918
Argyra setimana Loew, 1859
= subarctica misid.
Argyra spoliata Kowarz, 1879
Argyra vestita (Wiedemann, 1817)
ASYNDETUS Loew, 1869
Asyndetus latifrons (Loew, 1857)
CHRYSOTUS Meigen, 1824
Chrysotus angulicornis Kowarz, 1874
Chrysotus cilipes Meigen, 1824
Chrysotus cupreus (Macquart, 1827)
Chrysotus femoratus Zetterstedt, 1843
Chrysotus gramineus (Fallén, 1823)
= microcerus Kowarz, 1874
= varians Kowarz, 1874
Chrysotus laesus (Wiedemann, 1817)
= amplicornis Zetterstedt, 1849
Chrysotus neglectus (Wiedemann, 1817)
Chrysotus obscuripes Zetterstedt, 1838
= kowarzi Lundbeck, 1912
Chrysotus pulchellus Kowarz, 1874
Chrysotus suavis Loew, 1857
DIAPHORUS Meigen, 1824
Diaphorus homannseggi Meigen, 1830
Diaphorus nigricans Meigen, 1824
Checklist of the Empidoidea of Finland (Insecta,Diptera)201
Diaphorus oculatus (Fallén, 1823)
MELANOSTOLUS Kowarz, 1884
Melanostolus melancholicus (Loew, 1869)
MEDETERINAE Lioy, 1864
DOLICHOPHORUS Lichtward, 1902
Dolichophorus kerteszi Lichtward, 1902
MEDETERA Fischer von Waldheim, 1819
Medetera abstrusa uneberg, 1955
Medetera acanthura Negrobov & uneberg, 1970
Medetera adjaniae Gosseries, 1989
= breviseta Parent, 1927 preocc.
Medetera ambigua (Zetterstedt, 1843)
Medetera apicalis (Zetterstedt, 1843)
Medetera belgica Parent, 1936 sensu Negrobov & Stackelberg, 1972
Medetera betulae Ringdahl, 1949
Medetera borealis uneberg, 1955
Medetera cuspidata Collin, 1941
Medetera dichrocera Kowarz, 1877
Medetera excellens Frey, 1909
Medetera fasciata Frey, 1915
Medetera freyi uneberg, 1955
Medetera fumida Negrobov, 1967
Medetera impigra Collin, 1941
Medetera incrassata Frey, 1909
Medetera infumata Loew, 1857
Medetera inspissata Collin, 1952
Medetera jacula (Fallén, 1823)
Medetera jugalis Collin, 1941
Medetera melancholica Lundbeck, 1912
Medetera muralis Meigen, 1824
Medetera nitida (Macquart, 1834)
= stackelbergi Parent, 1927
Medetera obscura (Zetterstedt, 1838)
Medetera pallipes (Zetterstedt, 1843)
Medetera parenti Stackelberg, 1925
= collini uneberg, 1955
Medetera pinicola Kowarz, 1877
= nuortevai uneberg, 1955
Medetera plumbella Meigen, 1824
Medetera prjachinae Negrobov & Stackelberg, 1974
Medetera protuberans Negrobov, 1967
Medetera pseudoapicalis uneberg, 1955
Medetera seguyi Parent, 1926
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Medetera senicula Kowarz, 1877
Medetera setiventris uneberg, 1955
Medetera signaticornis Loew, 1857
Medetera striata Parent, 1927
Medetera tristis (Zetterstedt, 1838)
Medetera vagans Becker, 1917
= fennica uneberg, 1955
Medetera veles Loew, 1861
= bilineata Frey, 1915
Medetera zinovjevi Negrobov, 1967
SYSTENUS Loew, 1857
Systenus bipartitus (Loew, 1850)
Systenus pallipes (von Roser, 1840)
= adpropinquus (Loew, 1857)
Systenus scholtzi (Loew, 1850)
THRYPTICUS Gerstäcker, 1864
rypticus atomus Frey, 1915
rypticus bellus Loew, 1869
rypticus cuneatus (Becker, 1917)
rypticus divisus Strobl, 1880
= fennicus Becker, 1917
rypticus intercedens Negrobov, 1967
rypticus laetus Verrall, 1912
rypticus nigricauda Wood, 1913
rypticus pollinosus Verrall, 1912
rypticus pruinosus Parent, 1932
rypticus smaragdinus Gerstäcker, 1864
rypticus tarsalis Parent, 1932
rypticus sp. A
HYDROPHORINAE Lioy, 1864
HYDROPHORUS Fallén, 1823
Hydrophorus albiceps Frey, 1915
Hydrophorus alpinus Wahlberg, 1844
Hydrophorus altivagus Aldrich, 1911
= wahlgreni Frey, 1915
Hydrophorus bipunctatus (Lehmann, 1822)
Hydrophorus borealis Loew, 1857
Hydrophorus brunnicosus Loew, 1857
Hydrophorus callosoma Frey, 1915
= albosignatus Ringdahl, 1919
Hydrophorus freyi Storå, 1954
Hydrophorus geminus Frey, 1915
Hydrophorus litoreus Fallén, 1823
Checklist of the Empidoidea of Finland (Insecta,Diptera)203
Hydrophorus nebulosus Fallén, 1823
Hydrophorus norvegicus Ringdahl, 1928
Hydrophorus pectinatus Gerstäcker, 1864
= forcipatus Frey, 1915
Hydrophorus pilipes Frey, 1915
Hydrophorus praecox (Lehmann, 1822)
Hydrophorus rubarbis Gerstäcker, 1864
= micans Frey, 1915
Hydrophorus signiferus Coquillett, 1899
= magnicornis Frey, 1915
Hydrophorus viridis (Meigen, 1824)
LIANCALUS Loew, 1857
Liancalus virens (Scopoli, 1763)
PEODES Loew, 1857
Peodes petsamoensis Frey, 1930
SCELLUS Loew, 1857
Scellus spinimanus (Zetterstedt, 1843)
THINOPHILUS Wahlberg, 1844
sg. inophilus Wahlberg, 1844
inophilus avipalpis (Zetterstedt, 1843)
inophilus rucornis (Haliday, 1838)
NEURIGONINAE Aldrich, 1905
NEURIGONA Rondani, 1856
Neurigona abdominalis (Fallén, 1823)
Neurigona pallida (Fallén, 1823)
Neurigona quadrifasciata (Fabricius, 1781)
Neurigona suturalis (Fallén, 1823)
Neurigona sp. A
RHAPHIINAE Bigot, 1852
RHAPHIUM Meigen, 1803
= Xiphandrium Loew, 1857
= Porphyrops auctt.
Rhaphium albifrons (Zetterstedt, 1843)
Rhaphium appendiculatum (Zetterstedt, 1849)
= macrocerum auct. nec Meigen, 1824
Rhaphium basale Loew, 1850
Rhaphium caliginosum Meigen, 1824
= zetterstedti (Parent, 1925)
Rhaphium commune (Meigen, 1824)
= spinicoxa Loew, 1850
Rhaphium conne Zetterstedt, 1843
Rhaphium crassipes (Meigen, 1824)
Rhaphium discolor Zetterstedt, 1838
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= consobrinum Zetterstedt, 1843
= riparium auct. nec (Meigen, 1824: 53)
Rhaphium elegantulum (Meigen, 1824)
Rhaphium fasciatum Meigen, 1824
Rhaphium fascipes (Meigen, 1824)
Rhaphium ssum Loew, 1850
Rhaphium glaciale (Ringdahl, 1920)
Rhaphium holmgreni (Mik, 1878)
= luteipenne (Frey, 1915)
Rhaphium lanceolatum Loew, 1850
= caliginosum auct. nec Meigen, 1824
Rhaphium laticorne (Fallén, 1823)
= nemorum Meigen, 1830
Rhaphium latimanum Kahanpää, 2007
Rhaphium longicorne (Fallén, 1823)
Rhaphium micans (Meigen, 1824)
Rhaphium monotrichum Loew, 1850
= auctum misid.
Rhaphium nasutum (Fallén, 1823)
Rhaphium nigribarbatum (Becker, 1900)
Rhaphium patulum (Raddatz, 1873)
= antennatum misid.
Rhaphium penicillatum Loew, 1850
Rhaphium riparium (Meigen, 1824)
= praerosum Loew, 1850
Rhaphium rivale (Loew, 1869)
Rhaphium tridactylum (Frey, 1915)
Rhaphium umbripenne (Frey, 1915)
ACHALCINAE Grootaert & Meuels, 1997
ACHALCUS Haliday, 1857
Achalcus cinereus (Haliday, 1851)
Achalcus avicollis (Meigen, 1824)
Achalcus nigropunctatus Pollét & Brunhues, 1996
Achalcus vaillanti Brunhues, 1987
PELOROPEODINAE Robinson, 1970
CHRYSOTIMUS Loew, 1857
Chrysotimus molliculus (Fallén, 1823)
MICROMORPHUS Mik, 1878
Micromorphus claripennis (Strobl, 1899)
XANTHOCHLORINAE Aldrich, 1905
XANTHOCHLORUS Loew, 1857
Xanthochlorus ornatus (Haliday, 1832)
Xanthochlorus tenellus (Wiedemann, 1817)
Checklist of the Empidoidea of Finland (Insecta,Diptera)205
Excluded species
Chelifera astigma Collin, 1927 misidentied
Chrysotus arcticus Frey, 1915 not found within present borders
Chrysotus longipalpus Aldrich, 1896 imported
= pallidipalpus van Duzee, 1933
Chrysotus ringdahli Parent, 1929 not found within present borders
Clinocera nigra Meigen, 1804 misidentied
Crossopalpus abditus Kovalev, 1972 not found within present borders
Dolichopus angustipennis Kertesz, 1901 not found within present borders
= adustus Frey, 1915
Dolichopus grandicornis Wahlberg, 1850 not found within present borders
Dolichopus mediicornis Verrall, 1875 not found within present borders
Dolichopus plumitarsis Fallén, 1823 misidentied
Dolichopus propinquus Zetterstedt, 1852 misidentied
Drapetis incompleta Collin, 1926 misidentied
Empis punctata Meigen, 1804 not found within present borders
Gymnopternus assimilis (Stæger, 1842) misidentied
Hilara avipes Meigen, 1822 misidentied
Hilara maura (Fabricius, 1776) not found within present borders
Hilara scrobiculata Loew, 1873 misidentied
Hilara tetragramma Loew, 1873 misidentied
Hydrophorus balticus (Meigen, 1824) not found within present borders
Hydrophorus ponojensis Frey, 1915 not found within present borders
Iteaphila furcata (Zetterstedt, 1842) not found within present borders
Medetera annulitarsus von Roser, 1840 misidentied
Medetera glauca Loew, 1869 misidentied
Medetera feminina Negrobov, 1967 misidentied
Medetera truncorum Meigen, 1824 imported
Oedalea avipes Zetterstedt, 1842 misidentied
Parathalassius krogerusi Frey, 1927 nomen nudum
Rhamphomyia tibialis Meigen, 1822 misidentied
Rhaphium antennatum (Carlier, 1835) misidentied
Rhaphium obscuripes Zetterstedt, 1849 misidentied
Rhaphium suave (Loew, 1859) not found within present borders
Tachytrechus insignis (Stannius, 1831) misidentied
Trichoclinocera lapponica (Ringdahl, 1933) not found within present borders
Notes
Dolichopus caligatus Wahlberg, 1850 was synonymized with D. avipes Stannius,
1831 by Becker (1918) but Collin (1940) reinstated D. caligatus as a valid species.
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206
True D. avipes (if it is a valid species) has not been found in Finland but some
specimens have been recorded from Northwestern European Russia.
Dolichopus stenhammari Zetterstedt, 1843. Zetterstedt (1838) rst described this
species as Dolichopus annulipes. Later he proposed D. stenhammari Zett., 1843 as
an (unnecessary) replacement name for the taxon: Porphyrops annulipes Meigen,
1824 was recognized in Dolichopus at the time, but only as a junior synonym of
what is now Sympycnus pulicarius (Fallén, 1823) (Zetterstedt 1843). Nonetheless
this act means, according to article 59.3 of e Code (ICZN 1999), that D. an-
nulipes Zett., 1838 is permanently invalid.
Hilara pseudochorica Strobl, 1892. Erroneously deleted from the Finnish checklist
by Kahanpää and Winqvist (2005).
Iteaphila furcata (Zetterstedt 1842). Sinclair and Shamshev (2012) list a specimen
from Kuusamo with code label 1546. According to R. Frey’s notes this specimen
was collected in 1917 from eastern Kuusamo, an area ceded to Russia in 1944.
Neurigona sp. A is probably N. uralensis Becker, 1918.
Platypalpus articulatus and P. maculimanus. According to Allen (1986) these two
names are not synonymous and the species identied as P. articulatus Macquart by
Chvála (1975, 1989) is in fact P. maculimanus (Zett.). Upon re-examination, all Finn-
ish material previously identied as P. articulatus belonged to P. maculimanus and P.
articulatoides Frey. It remains unclear whether the real P. articulatus occurs in Finland.
Rhamphomyia sp. nr. albipennis. An apparently undescribed species near R. albipen-
nis (Fallén, 1816) (K. Winqvist & M. Barták, priv. comm.).
Sympycnus pulicarius (Fallén, 1823). Pollet (2004) (Fauna Europaea) treats S. an-
nulipes (Meigen, 1824) and S. desoutteri Parent, 1925 as valid species and lists S.
pulicarius (Fallén, 1823) as a synonym of the latter but notes that it might actually
be a synonym of S. annulipes.
rypticus sp. A is probably T. incanus Negrobov, 1967 described from the Leningrad
Oblast, Russia (Negrobov 1967).
Trichoclinocera lapponica (Ringdahl, 1933). e type locality of this species, ”Lapp-
land nära Tjuonajaure”, (Ringdahl 1933) is in Sweden, not Finland as stated in
Chvála and Wagner (1989).
References
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mologist’s Record and Journal of Variation 98: 177–179.
Becker T (1918) Dipterologische Studien. Dolichopodidae. A. Paläarktische Region. Nova Acta
Academiae Caesareae Leopoldinisch-Carolinae Germanicae Naturae Curiosorum 104: 35–214.
Chvála M (1975) e Tachydromiinae (Dipt. Empididae) of Fennoscandia and Denmark. Fau-
na Entomologica Scandinavica 3. Scandinavian Science Press Ltd., Klampenborg, 336 pp.
Chvála M (1983) e Empidoidea (Diptera) of Fennoscandia and Denmark. II. General Part.
e Families Hybotidae, Atelestidae and Microphoridae. Fauna Entomologica Scandi-
navica 12. Scandinavian Science Press Ltd., Klampenborg, 279 pp.
Checklist of the Empidoidea of Finland (Insecta,Diptera)207
Chvála M (1989) Monograph of Northern and Central European species of Platypalpus (Diptera,
Hybotidae), with data on the occurrence in Czechoslovakia. Acta Universitatis Carolinae–
Biologica 32: 209–376.
Chvála M, Wagner R (1989) Family Empididae. In: Soós A, Papp L (Eds) Catalogue of Palae-
arctic Diptera. erevidae-Empididae. Volume 6. Akademiai Kiado, Budapest & Elsevier,
Amsterdam, 228–336.
Collin JE (1940) Critical notes on some recent synonymy aecting British species of Dolicho-
podidae (Diptera). Entomologist’s Monthly Magazine 76: 261–271.
Kahanpää J (2006) First update to the checklist of Finnish Dolichopodidae (Diptera). Interna-
tional Journal of Dipterological Research 17(2): 121–125.
Kahanpää J (2013) A second update to the checklist of Finnish long-legged ies (Diptera:
Dolichopodidae), with a re-evaluation of the status of Hydrophorus callosoma Frey, 1915.
Biodiversity Data Journal 1: e976. doi: 10.3897/BDJ.1.e976
Kahanpää J, Grichanov IY (2004) A check-list of Finnish long-legged ies (Diptera: Dolicho-
podidae). International Journal of Dipterological Research 15(1): 57–62.
Kahanpää J, Winqvist K (2005) Check-list of Finnish ies: families Xylophagidae–Microphoridae.
Sahlbergia 10(1): 10–27.
Negrobov OP (1967) New Palaearctic species of the subfamily Medeterinae (Diptera, Dolicho-
podidae). Entomologicheskoe Obozrenie 46(4): 890–908.
Negrobov OP, Stackelberg AA (1972) 29. Dolichopodidae. In: Lindner E (Ed) Die Fliegen
der palaearktischen Region 4(5). E. Schweizerbart‘sche Verlagsbuchhandlung, Stuttgart,
257–302.
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed)
Animal biodiversity: An outline of higher-level classication and survey of taxonomic richness.
Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
Pollet M (2004) Fauna Europaea: Dolichopodidae. In: Pape T, Beuk P (Eds) Fauna Europaea:
Diptera. Fauna Europaea version 1.0. http://www.faunaeur.org
Ringdahl O (1933) Översikt av de i Sverige funna arterna av släktet Atalanta Meig. (Clinocera
Meig.) inom fam. Empididae. Entomologisk Tidskrift 54(3–4): 277–282.
Sinclair BJ, Shamshev IV (2012) World revision of Iteaphila macquarti group (Diptera: Empididae).
Zootaxa 3561: 1–61.
Shamshev IV, Sinclair BJ (2009) Revision of the Iteaphila setosa group (Diptera: Empididae).
European Journal of Entomology 106: 441–450. doi: 10.14411/eje.2009.055
Sinclair BJ, Cumming JM (2006) e morphology, higher-level phylogeny and classica-
tion of the Empidoidea (Diptera). Zootaxa 1180: 1–172. http://www.mapress.com/
zootaxa/2006f/zt01180p140.pdf
Yang D, Zhang K, Yao G, Zhang J (2007) World catalog of Empididae (Insecta: Diptera).
China Agricultural University Press, Beijing, 599 pp.
Yang D, Zhu YJ, Wang MQ, Zhang L (2006) World catalog of Dolichopodidae (Insecta: Diptera).
China Agricultural University Press, Beijing, 704 pp. [+44 pl.]
Zetterstedt JW (1838) Insecta Lapponica. Dipterologis Scandinaviae. Sect. 3: Diptera. Leopold
Voss, Lipsiae, 477–868.
Zetterstedt JW (1843) Diptera Scandinaviae. Disposita et descripta. Tomus primus. Ocina
Lundbergiana, Lund, 894 pp.
Jere Kahanpää / ZooKeys 441: 183–207 (2014)
208
Checklist of the families Opetiidae and Platypezidae(Diptera) of Finland 209
Checklist of the families Opetiidae and
Platypezidae (Diptera) of Finland
Gunilla Ståhls1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Gunilla Ståhls (gunilla.stahls@helsinki.)
Academic editor: J. Kahanpää|Received 3 April 2014|Accepted 11 June 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Ståhls G (2014) Checklist of the families Opetiidae and Platypezidae (Diptera) of Finland. In: Kahanpää J,
Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 209–212. doi: 10.3897/zookeys.441.7639
Abstract
A checklist of the Opetiidae and Platypezidae (Diptera) recorded from Finland.
Keywords
Checklist, Finland, Diptera, Opetiidae, Platypezidae
Introduction
Opetiidae and Platypezidae are small families of small-sized ies. Platypezidae are prin-
cipally forest insects, and all known larvae develop in fungi. New host fungi records
were recently reported in Reemer et al. (2014), Ståhls et al. (submitted) and Ståhls and
Rättel (2014).
e Finnish species were last listed by Hackman (1980) and Chandler (2001).
Chandler (2001) included one and 21 species of Opetiidae and Platypezidae, respec-
tively. Ståhls and Kahanpää (2006), Winqvist (2011), Ståhls et al. (2013), Reemer et
al. (2014) and Ståhls and Rättel (2014) added altogether 12 species of family Platypezi-
dae. Bolopus furcatus was included in Hackman (1980) but not in Chandler (2001),
Ståhls et al. (2012) reported the taxon from Finland again. Agathomyia (subfamily Cal-
lomyiinae) is the largest genus of the Platypezidae. A recent study reinstated two Agath-
omyia species from synonymy (A. boreella Zetterstedt and A. scutellaris Zetterstedt) and
two species were described as new to science (Ståhls et al. submitted). e taxonomic
ZooKeys 441: 209–212 (2014)
doi: 10.3897/zookeys.441.7639
www.zookeys.org
Copyright Gunilla Ståhls. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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Gunilla Ståhls / ZooKeys 441: 209–212 (2014)
210
status of A. dahlbomi (Zetterstedt), presently listed as synonym of A. elegantula (Zet-
terstedt), remains unclear. Callomyia elegans and Protoclythia modesta are only known
from previous single records, their occurrence has not been veried with new ndings.
Chandler and Shatalkin (1998) and Chandler (2001) provided keys to subfamilies,
genera and species of adult Palaearctic Platypezidae, and key to genera of known early
stages of Platypezidae. Ståhls et al. (submitted) provided a key to Northern European
Agathomyia spp.
Number of species (Opetiidae, Platypezidae):
World: 102, 252 species (Pape et al. 2011)
Europe: 1, 46 species
Finland: 1, 38 species
Faunistic knowledge level in Finland: average
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Aschiza Becher, 1882
superfamily Phoroidea Curtis, 1833
OPETIIDAE Rondani, 1856
OPETIA Meigen, 1830
Opetia nigra Meigen, 1830
PLATYPEZIDAE Fallén, 1815
CALLOMYIINAE Rondani, 1841
AGATHOMYIA Verrall, 1901
Agathomyia alneti Ståhls & Rättel, in prep.
Agathomyia antennata (Zetterstedt, 1819)
Agathomyia boreella (Zetterstedt, 1838)
Agathomyia cinerea (Zetterstedt, 1852)
Agathomyia elegantula (Fallén, 1815)
= dahlbomi (Zetterstedt, 1838)
Agathomyia falleni (Zetterstedt, 1819)
Agathomyia lundbecki Chandler in Shatalkin, 1985
Agathomyia sexmaculata (von Roser, 1840)
Agathomyia scutellaris (Zetterstedt, 1838)
Agathomyia shatalkini Ståhls, in prep.
Agathomyia unicolor Oldenberg, 1928
Agathomyia vernalis Shatalkin, 1981
Checklist of the families Opetiidae and Platypezidae(Diptera) of Finland 211
Agathomyia viduella (Zetterstedt, 1838)
Agathomyia wankowiczii (Schnabl, 1884)
Agathomyia woodella Chandler in Shatalkin, 1985
= elegantula auct. nec (Fallén, 1815)
Agathomyia zetterstedti (Wahlberg in Zetterstedt, 1844)
CALLOMYIA Meigen, 1804
Callomyia amoena Meigen, 1824
Callomyia elegans Meigen, 1804
= leptiformis (Fallén, 1810)
Callomyia krivosheinae Shatalkin, 1982
Callomyia speciosa Meigen, 1824
= humeralis Loew, 1869
MICROSANIINAE Enderlein, 1936
MICROSANIA Zetterstedt, 1837
Microsania capnophila Shatalkin, 1985
Microsania collarti Chandler, 2001
Microsania pallipes (Meigen, 1830)
Microsania pectipennis (Meigen, 1830)
= pectinipennis emend.
Microsania straeleni Collart, 1954
Microsania vrydaghi Collart, 1954
PLATYPEZINAE Fallén, 1815
BOLOPUS Enderlein, 1932
Bolopus furcatus (Fallén, 1826)
PARAPLATYPEZA Kessel & Maggioncalda, 1968
Paraplatypeza atra (Meigen, 1804)
Paraplatypeza bicincta (Szilády, 1941)
PLATYPEZA Meigen, 1803
Platypeza aterrima Walker, 1836
Platypeza consobrina Zetterstedt, 1844
Platypeza fasciata Meigen, 1804
Platypeza hirticeps Verrall, 1901
PLATYPEZINA Wahlgren, 1910
Platypezina connexa (Bohemann, 1858)
POLYPORIVORA Kessel & Maggioncalda, 1968
Polyporivora boletina (Fallén, 1815)
Polyporivora ornata (Meigen, 1838)
= infumata (Haliday, 1838)
Polyporivora picta (Meigen, 1830)
PROTOCLYTHIA Kessel, 1950
Protoclythia modesta (Zetterstedt, 1844)
SERI Kessel & Maggioncalda, 1966
Seri obscuripennis (Oldenberg, 1916)
Gunilla Ståhls / ZooKeys 441: 209–212 (2014)
212
References
Chandler PJ (2001) e at-footed ies (Diptera: Opetiidae and Playtypezidae) of Europe.
Fauna Entomologica Scandinavica 36: 1–276.
Hackman W (1980) A check cist of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Shatalkin AI (1985) A review of the at-footed ies (Diptera, Platypezidae) of USSR. Sbornik
Trudy zoologicheskii Muzei Moskovskii Gasudarstvennyi Universitet 23: 69–136. [in
Russian]
Reemer M, Ståhls G, Rättel E (2014) Host fungus of Agathomyia sexmaculata (Diptera: Platy-
pezidae) and notes on breeding biology. Sahlbergia, submitted.
Ståhls G, Kahanpää J (2006) New data on Platypezidae and Opetiidae (Diptera) of Finland.
Sahlbergia 11: 1–6.
Ståhls G, Rättel E, Muona J (2012) Bolopus furcatus (Fallén, 1826) (Diptera, Platypezidae)
rediscovered in Finland. Sahlbergia 18: 29–32.
Ståhls G, Rättel E (2013) e genus Microsania (Platypezidae, Microsaninae) in Finland. Sahl-
bergia 19: 50–52.
Ståhls G, Rättel E, Miettinen O (2014) New data on genus Callomyia of Finland (Diptera:
Platypezidae). Sahlbergia, in press.
Ståhls G, Miettinen O, Rättel E (2014) MtDNA COI in ecient use: revealing hidden species
diversity and clarifying taxonomy, linking morphologically discordant sexes and identify-
ing the immature stages of Agathomyia at-footed ies (Diptera). Journal of Zoological
Systematics and Evolutionary Research, submitted.
Winqvist K (2011) Ruissalon kärpäsistä (Diptera, Brachycera). w-album 11: 3–23. [in Finnish
with summary in English]
Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)213
Checklist of the families Lonchopteridae and
Phoridae of Finland (Insecta, Diptera)
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 5 February 2014|Accepted 26 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera). In:
Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 213–223. doi: 10.3897/zookeys.441.7197
Abstract
A checklist of the Lonchopteridae and Phoridae recorded from Finland is presented.
Keywords
Checklist, Finland, Diptera, Lonchopteridae, Phoridae
Introduction
e superfamily Phoroidea includes at least the y families Phoridae, Lonchopteridae,
and the small family Ironomyiidae known only from Australia. e at-footed y
families Platypezidae and Opetiidae, treated in a separate paper in this volume, are
placed either in their own basal superfamily Platypezoidea or included in Phoroidea
(see Cumming et al. 1995, Woodley et al. 2009 and Wiegmann et al. 2011).
Lonchoptera Meigen, 1803 is the only currently recognized genus in Lonchopteri-
dae. Five Lonchoptera species have been added to the Finnish fauna after the checklist
of Hackman (1980) (Andersson 1991, Kahanpää 2013).
e scuttle ies (family Phoridae) may be the largest single family in Diptera. e true
number of species in this family has been estimated to be over 30000, the vast majority still
undescribed (Marshall 2012). e ‘megagenus’ Megaselia Rondani, 1856 contains about
ZooKeys 441: 213–223 (2014)
doi: 10.3897/zookeys.441.7197
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
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A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 213–223 (2014)
214
50% of the species (Marshall 2012). Other phorid genera are relatively better known in
Finland but none of them can be called well studied. It is likely that at least a hundred (or
hundreds of) Finnish species remain unnoticed, most of them in Megaselia.
e checklist for Megaselia is mostly based on previously published records. e
collections of Finnish Museum of Natural History (MZH) were examined by the
author during the preparation of this checklist and a few obvious mistakes were taken
into account. Most of the material in MZH was identied by H. Schmitz (between
years 1915 and 1940) or E. Beyer (in 1958). e lists for other phorid genera are based
on recently examined specimens from both MZH and private collections. Due to the
diculties encountered in identifying Megaselia, the list for this genus should de-
nitely be considered preliminary. A revision of the Finnish material is sorely needed.
e Finnish species of these families were last listed by Hackman (1980).
Number of species (Lonchopteridae, Phoridae):
World: 63, 4105 species (Pape et al. 2011)
Europe: 13, 656 species
Finland: 8, ~224–234 species
Faunistic knowledge level in Finland: good, poor
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
superfamily Phoroidea Curtis, 1833
LONCHOPTERIDAE Macquart, 1835
LONCHOPTERA Meigen, 1803
= Dipsa Fallén, 1810
Lonchoptera bifurcata (Fallén, 1810)
= furcata (Fallén, 1823)
Lonchoptera fallax de Meijere, 1906
Lonchoptera impicta Zetterstedt, 1848
Lonchoptera lutea Panzer, 1809
Lonchoptera meijerei Collin, 1938
Lonchoptera nigrociliata Duda, 1927
Lonchoptera nitidifrons Strobl, 1898
Lonchoptera scutellata Stein, 1890
PHORIDAE Curtis, 1833
AENIGMATIAS Meinert, 1890
Aenigmatias lubbockii (Verrall, 1877)
Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)215
ANEVRINA Lioy, 1864
Anevrina thoracica (Meigen, 1804)
Anevrina unispinosa (Zetterstedt, 1860)
= fennica (Becker, 1901)
Anevrina urbana (Meigen, 1830)
BOROPHAGA Enderlein, 1924
Borophaga agilis (Meigen, 1830)
Borophaga carinifrons (Zetterstedt, 1848)
Borophaga femorata (Meigen, 1830)
Borophaga incrassata (Meigen, 1830)
Borophaga subsultans (Linnaeus, 1767)
= okellyi Schmitz, 1937
CHAETOPLEUROPHORA Schmitz, 1922
Chaetopleurophora erythronota (Strobl, 1892)
Chaetopleurophora spinosissima (Strobl, 1892)
CONICERA Meigen, 1830
Conicera dauci (Meigen, 1830)
= atra Meigen, 1830
Conicera oricola Schmitz, 1938
= similis auct. nec (Haliday, 1833)
= minuscula Schmitz, 1953
Conicera schnittmanni Schmitz, 1926
Conicera similis (Haliday, 1833)
= pauxilla Schmitz, 1920
? Conicera tarsalis Schmitz, 1920
Conicera tibialis Schmitz, 1925
= fallens Schmitz, 1948
DIPLONEVRA Lioy, 1864
= Diploneura emend.
Diplonevra orescens (Turton, 1801)
= orea (Fabricius, 1794) preocc.
= abdominalis (Fallén, 1823)
Diplonevra concinna (Meigen, 1830)
Diplonevra freyi Schmitz, 1927
Diplonevra funebris (Meigen, 1830)
= rostralis (Schmitz, 1918)
Diplonevra glabra Schmitz, 1927
= parcepilosa Schmitz, 1927
Diplonevra nitidula (Meigen, 1830)
Diplonevra oldenbergi Schmitz, 1920
DOHRNIPHORA Dahl, 1898
Dohrniphora cornuta (Bigot, 1857)
GYMNOPHORA Macquart, 1835
Jere Kahanpää / ZooKeys 441: 213–223 (2014)
216
Gymnophora arcuata (Meigen, 1830)
Gymnophora healeyae Disney, 1980
Gymnophora nigripennis Schmitz, 1926
Gymnophora quartomollis Schmitz, 1920
HYPOCERA Lioy, 1864
Hypocera mordellaria (Fallén, 1823)
MEGASELIA Rondani, 1856
= Aphiochaeta Brues, 1903
= Plastophora Brues, 1905
Megaselia aequalis (Wood, 1909)
Megaselia anis (Wood, 1909)
= proxima (Lundbeck, 1920)
Megaselia albicaudata (Wood, 1910)
Megaselia albiclava (Schmitz, 1926)
Megaselia altifrons (Wood, 1909)
Megaselia analis (Lundbeck, 1920)
? Megaselia angelicae (Wood, 1910)
Megaselia angularis (Schmitz, 1924)
Megaselia angusta (Wood, 1909)
= angustata emend.
= pulicaria auct. nec (Fallén, 1923)
Megaselia annulipes (Schmitz, 1921)
Megaselia armata (Wood, 1909)
Megaselia atrosericea Schmitz, 1927
Megaselia auricoma Schmitz, 1927
Megaselia baltica (Schmitz, 1924)
Megaselia barbulata (Wood, 1909)
= depilata (Lundbeck, 1921)
? Megaselia basispinata (Lundbeck, 1920)
Megaselia basiveluta Schmitz, 1935
Megaselia beckeri (Wood, 1909)
= laticrus Schmitz, 1927
Megaselia berndseni (Schmitz, 1919)
= pygmaeoides (Lundbeck, 1921)
Megaselia bovista (Gimmerthal, 1848)
Megaselia brevicostalis (Wood, 1910)
Megaselia breviseta (Wood, 1912)
Megaselia breviterga (Lundbeck, 1920)
= similata (Lundbeck, 1921)
Megaselia campestris (Wood, 1908)
Megaselia ciliata (Zetterstedt, 1848)
Megaselia cinereifrons (Strobl, 1910)
Megaselia clara (Schmitz, 1921)
Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)217
Megaselia coaetanea Schmitz, 1929
Megaselia coccyx Schmitz, 1965
? Megaselia collini (Wood, 1909)
Megaselia conformis (Wood, 1909)
Megaselia costalis (von Roser, 1840)
Megaselia curvivenia Schmitz, 1928
Megaselia dahli (Becker, 1901)
Megaselia discreta (Wood, 1909)
= nudiventris (Wood, 1909)
Megaselia diversa (Wood, 1909)
Megaselia dubitalis (Wood, 1908)
Megaselia eccoptomera Schmitz, 1927
Megaselia eisfelderae Schmitz, 1948
Megaselia elongata (Wood, 1914)
Megaselia erecta (Wood, 1910)
Megaselia exarcuata Schmitz, 1927
Megaselia excavata Schmitz, 1927
Megaselia excorticata Disney, 2009
Megaselia fennicola Beyer, 1958
Megaselia ammula Schmitz, 1928
Megaselia ava (Fallén, 1823)
Megaselia avicoxa (Zetterstedt, 1848)
Megaselia fumata (Malloch, 1909)
Megaselia funeralis Schmitz, 1928
Megaselia fungivora (Wood, 1909)
Megaselia fusca (Wood, 1909)
Megaselia fuscipalpis (Lundbeck, 1920)
Megaselia fuscoides Schmitz, 1934
Megaselia fuscovariana Schmitz, 1933
Megaselia giraudii (Egger, 1862)
Megaselia glabrifrons (Wood, 1909)
Megaselia groenlandica (Lundbeck, 1901)
Megaselia halterata (Wood, 1910)
= plurispinosa (Lundbeck, 1920)
Megaselia hectochaeta Schmitz, 1974
Megaselia hilaris Schmitz, 1927
Megaselia hirsuta (Wood, 1910)
Megaselia hirticaudata (Wood, 1910)
Megaselia hirticrus (Schmitz, 1918)
Megaselia humeralis (Zetterstedt, 1838)
= cubitalis (Becker, 1901)
Megaselia hyalipennis (Wood, 1912)
Megaselia infraposita (Wood, 1909)
Jere Kahanpää / ZooKeys 441: 213–223 (2014)
218
Megaselia ignobilis (Schmitz, 1919)
Megaselia intonsa Schmitz, 1948
Megaselia involuta (Wood, 1910)
Megaselia latifemorata (Becker, 1901)
Megaselia limburgensis (Schmitz, 1918)
Megaselia longicostalis (Wood, 1912)
Megaselia longifurca (Lundbeck, 1921)
= spinolabella Disney, 1989
Megaselia longipalpis (Wood, 1910)
Megaselia longiseta (Wood, 1909)
Megaselia lucifrons (Schmitz, 1918)
= subnitida (Lundbeck, 1920)
Megaselia luminifrons (Schmitz, 1926)
Megaselia lutea (Meigen, 1830)
Megaselia major (Wood, 1912)
Megaselia mallochi (Wood, 1909)
Megaselia manicata (Wood, 1910)
Megaselia maura (Wood, 1910)
Megaselia meconicera (Speiser, 1925)
Megaselia meigeni (Becker, 1901)
? Megaselia miki Schmitz, 1929
Megaselia minor (Zetterstedt, 1848)
= angustifrons (Wood, 1912)
Megaselia minuta (Aldrich, 1892)
= luminosa Schmitz, 1952
Megaselia mixta (Schmitz, 1918)
Megaselia nasoni (Malloch, 1914)
= coaequalis (Schmitz, 1919)
Megaselia nigra (Meigen, 1830)
= albidihalteralis (Felt, 1896)
Megaselia nigriceps (Loew, 1866)
= projecta (Becker, 1901)
Megaselia nigripalpis (Lundbeck, 1920)
Megaselia nudipleura (Beyer, 1958)
Megaselia obscuripennis (Wood, 1909)
Megaselia ouscata (Schmitz, 1921)
Megaselia opacicornis Schmitz, 1949
Megaselia parnassia Disney, 1986
Megaselia parva (Wood, 1909)
Megaselia pectoralis (Wood, 1910)
Megaselia pectunculata Schmitz, 1927
Megaselia picta (Lehmann, 1822)
Megaselia pleuralis (Wood, 1909)
Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)219
Megaselia plurispinulosa (Zetterstedt, 1860)
Megaselia posticata (Strobl, 1898)
Megaselia praeacuta (Schmitz, 1919)
Megaselia prodroma (Lundbeck, 1921)
Megaselia producta (Schmitz, 1921)
= sordescens auct. nec Schmitz, 1927
Megaselia propinqua (Wood, 1909)
Megaselia pubecula Schmitz, 1927
Megaselia pulicaria (Fallén, 1923)
= sinuata Schmitz, 1926
Megaselia pumila (Meigen, 1830)
= atripes (Brues, 1915)
Megaselia pusilla (Meigen, 1830)
Megaselia pygmaea (Zetterstedt, 1848)
Megaselia quadriseta (Schmitz, 1918)
= badia Schmitz, 1938
? Megaselia raetica Schmitz, 1934
Megaselia robertsoni Disney, 2008
Megaselia robusta Schmitz, 1928
Megaselia rubricornis (Schmitz, 1919)
Megaselia rudis (Wood, 1909)
Megaselia rucornis (Meigen, 1830)
Megaselia rupes (Meigen, 1804)
Megaselia scalaris (Loew, 1866)
Megaselia scutellaris (Wood, 1909)
= scutellariformis (Schmitz, 1926)
Megaselia sepulchralis (Lundbeck, 1920)
Megaselia setulipalpis Schmitz, 1938
Megaselia sordida (Zetterstedt, 1838)
= eminens Schmitz, 1953
= semiscaura Schmitz, 1927
? Megaselia specularis Schmitz, 1935
Megaselia spinicincta (Wood, 1910)
Megaselia spinigera (Wood, 1908)
Megaselia styloprocta (Schmitz, 1921)
Megaselia subcarpalis (Lundbeck, 1920)
Megaselia subpalpalis (Lundbeck, 1920)
Megaselia subpleuralis (Wood, 1909)
Megaselia subtumida (Wood, 1909)
Megaselia sulphuripes (Meigen, 1830)
Megaselia superciliata (Wood, 1910)
Megaselia superfurcata Schmitz, 1928
Megaselia sylvatica (Wood, 1910)
Jere Kahanpää / ZooKeys 441: 213–223 (2014)
220
= impolluta (Schmitz, 1920)
Megaselia tarsalis (Wood, 1910)
Megaselia tenuiventris Schmitz, 1927
Megaselia tignorum Disney, 2009
Megaselia tumida (Wood, 1909)
= setifer Lundbeck, 1920
Megaselia unicolor (Schmitz, 1919)
Megaselia valvata Schmitz, 1935
Megaselia variana Schmitz, 1926
Megaselia vestita (Wood, 1914)
Megaselia winqvisti Disney, 2011
Megaselia woodi (Lundbeck, 1922)
Megaselia xanthozona (Strobl, 1892)
= euryprocta Schmitz, 1957
Megaselia zonata (Zetterstedt, 1838)
MENOZZIOLA Schmitz, 1927
Menozziola obscuripes (Schmitz, 1927)
METOPINA Macquart, 1835
Metopina galeata (Haliday, 1833)
= inaequalis Schmitz, 1927
Metopina oligoneura (Mik, 1867)
Metopina pileata Schmitz, 1936
MICROSELIA Schmitz, 1934
Microselia forsiusi (Schmitz, 1927)
PHALACROTOPHORA Enderlein, 1912
Phalacrotophora berolinensis Schmitz, 1920
Phalacrotophora beuki Disney in Disney & Beuk, 1997
Phalacrotophora fasciata (Fallén, 1823)
PHORA Latreille, 1796
= Trineura Meigen, 1803
Phora artifrons Schmitz, 1920
Phora atra (Meigen, 1804)
= aterrima (Fabricius, 1794) preocc.
Phora bullata Schmitz, 1927
Phora convallium Schmitz, 1928
Phora convergens Schmitz, 1920
Phora dubia (Zetterstedt, 1848)
= schineri (Becker, 1901)
Phora edentata Schmitz, 1920
Phora hamata Schmitz, 1927
Phora holosericea Schmitz, 1920
Phora hyperborea Schmitz, 1927
Phora obscura (Zetterstedt, 1848)
Phora occidentata Malloch, 1912
Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)221
= zetterstedti Schmitz, 1927
Phora penicillata Schmitz, 1920
Phora praepandens Schmitz, 1927
Phora pubipes Schmitz, 1920
Phora stictica Meigen, 1830
Phora tincta Schmitz, 1920
PLECTANOCNEMA Schmitz, 1926
Plectanocnema nudipes (Becker, 1901)
PSEUDACTEON Coquillett, 1907
Pseudacteon fennicus Schmitz, 1927
Pseudacteon formicarum (Verrall, 1877)
SPINIPHORA Malloch, 1909
Spiniphora bergenstammi Mik, 1864
Spiniphora dorsalis (Becker, 1901)
Spiniphora excisa (Becker, 1901)
Spiniphora maculata (Meigen, 1830)
= helicivora (Dufour, 1841)
TRIPHLEBA Rondani, 1856
= Citrago Schmitz, 1924
Triphleba admirabilis Schmitz, 1927
? Triphleba aequalis (Schmitz, 1919)
? Triphleba autumnalis (Becker, 1901)
Triphleba bicornuta (Strobl, 1910)
= uncinata (Schmitz, 1918)
Triphleba citreiformis (Becker, 1901)
Triphleba distinguenda (Strobl, 1892)
= unicalcarata (Becker, 1901)
Triphleba excisa (Lundbeck, 1921)
Triphleba gracilis (Wood, 1907)
Triphleba hyalinata (Meigen, 1830)
? Triphleba inaequalis Schmitz, 1943
Triphleba intermedia (Malloch, 1908)
Triphleba lugubris (Meigen, 1830)
Triphleba luteifemorata (Wood, 1906)
Triphleba nudipalpis (Becker, 1901)
Triphleba opaca (Meigen, 1830)
Triphleba pachyneurella (Schmitz, 1919)
Triphleba palposa (Zetterstedt, 1848)
Triphleba renidens Schmitz, 1927
Triphleba subcompleta Schmitz, 1927
Triphleba trinervis (Becker, 1901)
VERUANUS Schmitz, 1927
Veruanus oldenbergi Schmitz, 1919
= memorabilis Schmitz, 1927
Jere Kahanpää / ZooKeys 441: 213–223 (2014)
222
Excluded species
Megaselia cirriventris Schmitz, 1929 mistake?
Megaselia crassipes (Wood, 1909) misidentied
Megaselia digitalis Schmitz, 1957 mistake?
Megaselia indierens (Lundbeck, 1920) not found within present borders
Megaselia lapponica Schmitz, 1928 not found within present borders
Megaselia luteipes (Schmitz, 1918) misidentied
Megaselia palmeni (Becker, 1901) not found within present borders
Megaselia septentrionalis (Schmitz, 1919) not found within present borders
Megaselia simulans (Wood, 1912) not found within present borders ?
Triphleba gilvipes Schmitz, 1943 not found within present borders
Notes
Conicera tarsalis, Megaselia collini. Records of these species are based on single fe-
males identied by H. Schmitz.
Megaselia basispinata, M. miki, M. raetica. Species recorded from Finland by Hack-
man (1980) and Disney (1991) but no Finnish specimens or published records
with details could be located.
Megaselia cirriventris, M. digitalis. ese species were recorded from ‘Fennia’ and
‘Lappland’ by Schmitz (1958), but all specimens listed are from the Abisko region
in northen Sweden. It seems likely that Schmitz erroneously assigned the material
from Swedish Lappland to Finland.
Megaselia luteipes. Examined specimens belong to Megaselia sp. not luteipes (females)
(Espoo, leg. W. Hackman) and possibly Megaselia sp. 24 of Disney (2013) (Fin-
land, Muonio, leg. J. Sahlberg and Russia, Kola, Kantalahti, leg. R. Frey).
Megaselia pulicaria agg. is dicult complex of species was revised by Disney
(1999). e Finnish material has not been re-examined. e Finnish species in-
volved are M. angusta, M. costalis, M. longicostalis, M. longifurca, M. pulicaria, M.
subtumida and M. tumida.
Megaselia simulans. Recorded only(?) from Kuusamo, a municipality split between
Finland and Russia in 1944. Most records preceding the split are from the Russian
side. No material was found in MZH.
Megaselia specularis. Recorded, with some doubt, from Finland by Hackman (1963).
No material could be found in MZH.
Triphleba aequalis. Schmitz (1943) mention a Finnish male stored in MZH, but no
such specimen could be found.
Triphleba autumnalis, T. inaequalis. Finnish according to Disney (1991), but no
specimens or detailed records could be located.
Checklist of the families Lonchopteridae and Phoridae of Finland (Insecta, Diptera)223
References
Andersson H (1991) Family Lonchopteridae. In: Soós Á, Papp L (Eds) Catalogue of Palaearctic
Diptera, Volume 7, Dolichopodidae–Platypezidae. Akademiai Kiado, Budapest & Elsevier,
Amsterdam, 139–142.
Cumming JM, Sinclair BJ, Wood DM (1995) Homology and phylogenetic implications of
male genitalia in Diptera - Eremoneura. Entomologica scandinavica 26: 120–151. doi:
10.1163/187631295X00143
Disney RHL (1991) Family Phoridae. In: Soós Á, Papp L (Eds) Catalogue of Palaearctic
Diptera, Volume 7, Dolichopodidae–Platypezidae. Akademiai Kiado, Budapest & Else-
vier, Amsterdam, 143–204.
Disney RHL (1999) A troublesome sibling species complex of scuttle ies (Diptera: Phoridae)
revisited. Entomologica Scandinavica 33(8): 1159–1216. doi: 10.1080/002229399299987
Disney RHL (2013) An unusually rich scuttle y fauna (Diptera, Phoridae) from north of the
Arctic Circle in the Kola Peninsula, N.W. Russia. ZooKeys 342: 45–74. doi: 10.3897/
zookeys.342.5772
Hackman W (1963) Studies on the dipterous fauna in burrows of voles (Microtus, Clethrionomys)
in Finland. Acta Zoologica Fennica 102: 1–64.
Hackman W (1980) A Check List of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Kahanpää J (2013) Viisikymmentä Suomelle uutta kärpäslajia (Diptera: Brachycera). Sahlbergia
19(1–2): 63–71.
Marshall SA (2012) Flies: e Natural History and Diversity of Diptera. Firey Books, Richmond
Hill, Ontario, 616 pp.
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Schmitz H (1943) 33. Phoridae. In: Lindner E (Ed) Die Fliegen der Palaearktischen Region
4(7). E. Schweizerbart‘sche Verlagsbuchhandlung, Stuttgart, 673–712.
Schmitz H (1958) 33. Phoridae. Lieferung 202. In: Lindner E (Ed) Die Fliegen der Palaearktischen
Region 4(7). E. Schweizerbart‘sche Verlagsbuchhandlung, Stuttgart, 465–512.
Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA,
Cassel BK, Bayless KM, Heimberg AM, Wheeler BM, Peterson KJ, Pape T, Sinclair BJ, Ske-
vington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Ott U, Kampmeier GE, omp-
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(2011) Episodic radiations in the y tree of life. Proceedings of the National Academy of Sci-
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Woodley NE, Borkent A, Wheeler TA (2009) Phylogeny of the Diptera. In: Brown BV, Bor-
kent A, Cumming JM, Wood DM, Woodley NE, Zumbado MA (Eds) Manual of Central
American Diptera. Volume 1. NRC Research Press, Ottawa, 79–94.
Jere Kahanpää / ZooKeys 441: 213–223 (2014)
224
Checklist of the family Pipunculidae (Diptera) of Finland 225
Checklist of the family Pipunculidae (Diptera) of Finland
Christian Kehlmaier1
1 c/o Senckenberg Natural History Collections Dresden, Museum of Zoology, Dresden, Germany
Corresponding author: Christian Kehlmaier (kehlmaier@web.de)
Academic editor: J. Kahanpää|Received 17 March 2014|Accepted 25 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kehlmaier C (2014) Checklist of the family Pipunculidae (Diptera) of Finland. In: Kahanpää J, Salmela J
(Eds) Checklist of the Diptera of Finland. ZooKeys 441: 225–232. doi: 10.3897/zookeys.441.7278
Abstract
A checklist of the Pipunculidae (Diptera) cited from Finland is presented. At present, 106 species have
been recorded.
Keywords
Checklist, Finland, Diptera, Pipunculidae
Introduction
Pipunculidae or big-headed ies can be readily identied at family level by their large
compound eyes occupying almost the entire globular head. Additional morphological
autapomorphies at the family rank of this group are their enlarged anterior ommatidial
facets in the female including a so far unique retina1 pattern among Diptera, the pierc-
er-like shape of the female ovipositor and the presence of a chitinized postspiracular
plate of their last instar larvae (Wada 1991, Rafael and De Meyer 1992). Due to the
small to medium sized adults (2–12 mm), their uniform dark appearance and their
rather cryptic lifestyle, it takes some eort to detect them in the eld. erefore, Pi-
punculidae can best be collected by Malaise traps, although a trained eye and patience
can yield species rich catches by hand-netting as well. Big-headed ies can be found in
a variety of habitats, where they are hovering in between herbal vegetation or among
leaves of bushes and trees, searching for food sources (mainly honeydew) and mating
ZooKeys 441: 225–232 (2014)
doi: 10.3897/zookeys.441.7578
www.zookeys.org
Copyright Christian Kehlmaier. This is an open access article distributed under the terms of the Creative Commons Attribution International License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Christian Kehlmaier / ZooKeys 441: 225–232 (2014)
226
partners or for suitable hosts for their parasitic larvae, which are important endopara-
sitoids of nymphal and adult Auchenorrhyncha (Chalarinae and Pipunculinae) and
adult Tipulidae (Nephrocerinae) (Koenig and Young 2007). is highly specialized
life-style is unique to this family of Diptera. Hardly any larvae have been described so
far and the knowledge about their host specicity is fragmentary. For the European
fauna, host records are available for 30% of the 209 known species, i.e., 62 pipunculid
species have been reared from 59 host species in the past (Kehlmaier unpublished).
e knowledge of the Finnish fauna can be considered as good, with the latest re-
view published only recently by Kehlmaier and Ståhls (2007). Whereas about a dozen
additional species can be expected to be found, the distribution of the individual taxa
within Finland is largely unknown.
Identication: Pipunculidae taxonomy largely depends on male genitalic features.
erefore, dissection of the male genital apparatus is essential and a high magnication of
at least 50 times should be used for identication. Females can best be identied through
a combination of outer anatomical features primarily based on the shape of the oviposi-
tor. Most pipunculid genera present in Europe have been taxonomically reviewed within
the past three decades. As no comprehensive key exists for northern Europe, adults have
to be identied with the following set of publications: Chalarus (Kehlmaier and Assmann
2008, Kehlmaier 2010), Jassidophaga and Verrallia (Kuznetzov 1992, Kehlmaier 2006),
Nephrocerus (Grootaert and De Meyer 1986), Cephalopsini (De Meyer 1989, Ackland
1993, Kehlmaier and De Meyer 2005, Kehlmaier 2008), Eudorylini (Kehlmaier 2005,
Kuznetzov 1990), Pipunculini (Kehlmaier 2008), Dorylomorpha (Albrecht 1990), Tomo-
svaryella (Földvári and De Meyer 1999, Kehlmaier 2008).
Number of species:
World: 1428 species (Pape et al. 2011)
Europe: 209 species
Finland: 106 species
Faunistic knowledge level in Finland: good
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Aschiza Becher, 1882
superfamily Syrphoidea Latreille, 1802
PIPUNCULIDAE Walker, 1834
CHALARINAE Aczél, 1939
CHALARUS Walker, 1834
Chalarus basalis Loew, 1873
Chalarus brevicaudis Jervis, 1992
Checklist of the family Pipunculidae (Diptera) of Finland 227
Chalarus decorus Jervis, 1992
Chalarus elegantulus Jervis, 1992
= absconditus Kehlmaier in Kehlmaier & Assmann, 2008
Chalarus mbriatus Coe, 1966
Chalarus gynocephalus Jervis, 1992
Chalarus holosericeus (Meigen, 1824)
= perplexus Jervis, 1992
Chalarus immanis Kehlmaier, 2008
Chalarus indistinctus Jervis, 1992
Chalarus juliae Jervis, 1992
Chalarus latifrons Hardy, 1943
Chalarus pughi Coe, 1966
Chalarus spurius (Fallén, 1816)
= obscurus (Zetterstedt, 1838)
= argenteus misid.
JASSIDOPHAGA Aczél, 1939
Jassidophaga beatricis (Coe, 1966)
Jassidophaga fasciata (von Roser, 1840)
= setosa (Verrall, 1901)
Jassidophaga pilosa (Zetterstedt, 1838)
Jassidophaga villosa (von Roser, 1840)
Jassidophaga spec. A
VERRALLIA Mik, 1899
Verrallia aucta (Fallén, 1817)
NEPHROCERINAE Aczél, 1939
NEPHROCERUS Zetterstedt, 1838
Nephrocerus avicornis Zetterstedt, 1844
Nephrocerus lapponicus Zetterstedt, 1838
Nephrocerus scutellatus (Macquart, 1834)
PIPUNCULINAE Walker, 1834
tribe Cephalopsini Macquart, 1834
CEPHALOPS Fallén, 1810
sg. Cephalops Fallén, 1810
Cephalops aeneus Fallén, 1810
Cephalops vittipes (Zetterstedt, 1844)
= annulipes (Zetterstedt, 1838) in part
sg. Parabeckerias De Meyer, 1994
Cephalops obtusinervis (Zetterstedt, 1844)
sg. Semicephalops De Meyer, 1994
Cephalops carinatus (Verrall, 1901)
Cephalops straminipes (Becker, 1900)
= chlorionae (Frey, 1945)
Cephalops subultimus Collin, 1956
Christian Kehlmaier / ZooKeys 441: 225–232 (2014)
228
Cephalops varipes (Meigen, 1824)
= semifumosus (Kowarz, 1887)
CEPHALOSPHAERA Enderlein, 1936
Cephalosphaera furcata (Egger, 1860)
Cephalosphaera germanica Aczél, 1940
tribe Eudorylini Rafael & De Meyer, 1992
CLISTOABDOMINALIS Skevington, 2001
Clistoabdominalis doczkali Kehlmaier, 2005
EUDORYLAS Aczél, 1940
Eudorylas angustimembranus Kozánek & Kwon, 1991
= kozaneki De Meyer, 1993
Eudorylas arcanus Coe, 1966
Eudorylas barkalovi Kuznetzov, 1990
Eudorylas carpathicus Kozánek, 1993
Eudorylas coloratus (Becker, 1897)
Eudorylas elephas (Becker, 1897)
Eudorylas fascipes (Zetterstedt, 1844)
Eudorylas furvulus Collin, 1956
Eudorylas fuscipes (Zetterstedt, 1844)
= roseri misid.
= trochanteratus misid.
Eudorylas fusculus (Zetterstedt, 1844)
Eudorylas goennersdorfensis Dempewolf & Dunk, 1996
Eudorylas inferus Collin, 1956
Eudorylas jenkinsoni Coe, 1966
Eudorylas johnenae Dempewolf, 1996
Eudorylas kowarzi (Becker, 1897)
Eudorylas montium (Becker, 1897)
Eudorylas obscurus Coe, 1966
Eudorylas restrictus Coe, 1966
= pannonicus misid.
Eudorylas slovacus Kozánek, 1993
Eudorylas stackelbergi Kuznetzov, 1990
Eudorylas subfascipes Collin, 1956
Eudorylas subterminalis Collin, 1956
Eudorylas terminalis (omson, 1870)
Eudorylas unicolor (Zetterstedt, 1844)
Eudorylas vonderdunki Dempewolf, 1998
Eudorylas zermattensis (Becker, 1897)
Eudorylas zonatus (Zetterstedt, 1849)
Eudorylas zonellus Collin, 1956
tribe Microcephalopsini Rafael & De Meyer, 1991
MICROCEPHALOPS De Meyer, 1989
Checklist of the family Pipunculidae (Diptera) of Finland 229
Microcephalops opacus (Fallén, 1816)
= vestitus (Becker, 1900)
tribe Pipunculini Walker, 1834
PIPUNCULUS Latreille, 1802
Pipunculus calceatus von Roser, 1840
Pipunculus campestris Latreille, 1802
= ater Meigen, 1824
= spinipes Meigen, 1830
= thomsoni Becker, 1897
Pipunculus dimi Kuznetzov, 1991
Pipunculus elegans Egger, 1860
= spinipes auct. nec Meigen, 1830
Pipunculus fonsecai Coe, 1966
Pipunculus lenis Kuznetsov, 1991
= thomsoni auct. nec Becker, 1897
Pipunculus lichtwardti Kozanek, 1981
Pipunculus oldenbergi Collin, 1956
Pipunculus omissinervis Becker, 1889
Pipunculus tenuirostris Kozanek, 1981
= balticus Kuznetzov, 1991
Pipunculus violovitshi Kuznetzov, 1991
= varipes auct. nec Meigen, 1824
Pipunculus zugmayeriae Kowarz, 1887
tribe Tomosvaryellini Hardy, 1943
DORYLOMORPHA Aczél, 1939
sg. Dorylomorpha Aczél, 1939
Dorylomorpha aczeli (Hardy, 1947)
Dorylomorpha confusa (Verrall, 1901)
Dorylomorpha extricata (Collin, 1937)
Dorylomorpha imparata (Collin, 1937)
Dorylomorpha rupes (Meigen, 1824)
= xanthoceroides (Aczél, 1939)
Dorylomorpha spinosa Albrecht, 1979
sg. Dorylomyia Albrecht, 1990
Dorylomorpha beckeri (Aczél, 1939)
Dorylomorpha xanthocera (Kowarz, 1887)
sg. Dorylomyza Albrecht, 1990
Dorylomorpha albitarsis (Zetterstedt, 1844)
Dorylomorpha anderssoni Albrecht, 1979
Dorylomorpha canadensis Hardy, 1943
Dorylomorpha clavata Albrecht, 1979
Dorylomorpha clavifemora Coe, 1966
Dorylomorpha fennica Albrecht, 1979
Christian Kehlmaier / ZooKeys 441: 225–232 (2014)
230
Dorylomorpha hackmani Albrecht, 1979
Dorylomorpha haemorrhoidalis (Zetterstedt, 1838)
Dorylomorpha inrmata (Collin, 1937)
Dorylomorpha lautereri Albrecht, 1990
Dorylomorpha occidens (Hardy, 1939)
Dorylomorpha onegensis Albrecht, 1990
Dorylomorpha platystylis Albrecht, 1979
Dorylomorpha praetermissa Albrecht, 1979
Dorylomorpha xanthopus (omson, 1870)
sg. Pipunculina Albrecht, 1990
Dorylomorpha borealis (Wahlgren, 1910)
Dorylomorpha maculata (Walker, 1834)
TOMOSVARYELLA Aczél, 1939
= Alloneura Rondani, 1856 nomen nudum
Tomosvaryella cilitarsis (Strobl, 1910)
= forsiusi (Frey, 1932)
Tomosvaryella coquilletti (Kertész, 1907)
Tomosvaryella geniculata (Meigen, 1824)
= nigritula (Zetterstedt, 1844)
Tomosvaryella kalevala Kehlmaier, 2008
Tomosvaryella kuthyi Aczél, 1944
Tomosvaryella minuscula (Collin, 1956)
= magyarica Földvári & De Meyer, 1999
Tomosvaryella palliditarsis (Collin, 1931)
Tomosvaryella rossica Kuznetzov, 1993
Tomosvaryella sylvatica (Meigen, 1824)
Excluded species (as discussed in Kehlmaier and Ståhls 2007)
Claraeola halterata (Meigen, 1838) misidentied
Dorylomorpha incognita (Verrall, 1901) misidentied
Clistoabdominalis trochanteratus (Becker, 1900) misidentied
Dasydorylas roseri (Becker, 1897) misidentied
Eudorylas pannonicus (Becker, 1897) misidentied
Notes
Chalarus argenteus Coe, 1966. e single Finnish specimen previously identied as
C. argenteus was re-examined and was found to belong to C. spurius (Kaj Winqvist,
pers. comm.).
Checklist of the family Pipunculidae (Diptera) of Finland 231
Jassidophaga spec. A. e specimens summarised under this place holder in Kehlmaier
and Ståhls (2008) might represent an additional species. Using the key in Kuznetzov
(1992), males run towards the Eastern Palaearctic J. kurilensis (Kuznetzov, 1992),
whereas females are closest to J. beatricis.
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Checklist of the family Syrphidae (Diptera) of Finland 233
Checklist of the family Syrphidae (Diptera) of Finland
Antti Haarto1, Sakari Kerppola2
1 Zoological Museum, Section of Biodiversity and Environmental Science, Department of Biology, FI-20014
University of Turku, Finland 2 Hiihtomäentie 44 A 6, FI-00800 Helsinki, Finland
Corresponding author: Antti Haarto (ahaarto@gmail.com)
Academic editor: J. Kahanpää|Received 12 February 2014|Accepted 7 May 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Haarto A, Kerppola S (2014) Checklist of the family Syrphidae (Diptera) of Finland. In: Kahanpää J, Salmela
J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 233–249. doi: 10.3897/zookeys.441.7251
Abstract
A checklist of the Syrphidae (Diptera) recorded from Finland. ree species of Syrphidae, Platycheirus
modestus Ide, 1926, Cheilosia barovskii (Stackelberg, 1930) and Mallota tricolor Loew, 1871, are published
as new to the Finnish fauna. P. modestus is also new to the Palaearctic.
Keywords
Checklist, Finland, Diptera, Syrphidae, faunistics
Introduction
Adults of Syrphidae (hoveries or ower ies) are mimics of bees, bumblebees, wasps
and sawies. Syrphid ies are active visitors of owers for nectar and pollen and
therefore they are important pollinators. e syrphid adults are excellent iers and
often seen hovering.
e larvae of hoveries exploit a wide range of food sources. e majority of the
species are predators and a large number of them feed on aphids. Almost half of the
syrphid species are phytophagous or saprophagous. e larvae of phytophagous spe-
cies tunnel in roots and stems of plants and some mine in leaves, but this rarely causes
ZooKeys 441: 233–249 (2014)
doi: 10.3897/zookeys.441.7251
www.zookeys.org
Copyright A. Haarto, S. Kerppola. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Antti Haarto & Sakari Kerppola / ZooKeys 441: 233–249 (2014)
234
signicant economic damage. Aquatic larvae are saprophagous in decaying organic
material and feed on detritus or lter bacteria. Also larvae living in dead wood and
in sap runs probably lter bacteria from the decaying mass. (Marshall 2012, Van
Veen2004).
Only a few active Diptera researchers, Richard Frey, Wolter Hellén and Erkki
Kanervo, studied the fauna of Finnish syrphids from the 1910s to the 1960s. ey cre-
ated a very good base for the knowledge of the Finnish syrphid fauna. Hackman(1980)
published a list of the Finnish ies including 270 species of hoveries. e beginning
of the 2000s was an active period for faunistic research of the hovery fauna and the
number of recorded hovery species had increased to 347 as the book on Finnish
hoveries (Haarto and Kerppola 2007a) was published. After that the knowledge has
still been increasing thanks to working on the taxonomy and faunistics of the Finnish
hovery fauna so that the distribution can be considered as well-known for over 90 per
cent of the Finnish hovery species (Kahanpää 2010).
e Finnish hovery species were last listed by Kerppola (2013) with additions
and omissions of species and with some nomenclature changes.
Identication: e Finnish syrphid fauna is best covered for identication by
Haarto and Kerppola (2007a) and Bartsch (2009) although some species are missing
in these books. Most of the species can be identied by external characters but some
species require male genitalic characters for reliable determination.
Number of species:
World: 6015 species (Pape et al. 2011)
Europe: 830 species
Finland: 362 species
Faunistic knowledge level in Finland: good
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Aschiza Becher, 1882
superfamily Syrphoidea Latreille, 1802
SYRPHIDAE Latreille, 1802
SYRPHINAE Latreille, 1802
tribe Bacchini Bigot, 1883
BACCHA Fabricius, 1805
Baccha elongata (Fabricius, 1775)
= obscuripennis Meigen, 1822
tribe Melanostomini Williston, 1885
MELANOSTOMA Schiner, 1860
Checklist of the family Syrphidae (Diptera) of Finland 235
Melanostoma dubium (Zetterstedt, 1837)
= freyi (Hellén, 1950)
Melanostoma mellinum (Linnaeus, 1758)
Melanostoma scalare (Fabricius, 1794)
PLATYCHEIRUS Le Peletier & Serville, 1828
sg. Platycheirus Le Peletier & Serville, 1828
Platycheirus aeratus Coquillet, 1900
= angustitarsis Kanervo, 1934
Platycheirus albimanus (Fabricius, 1781)
Platycheirus ambiguus (Fallén, 1817)
Platycheirus amplus Curran, 1927
Platycheirus angustatus (Zetterstedt, 1843)
Platycheirus aurolateralis (Stubbs, 2002)
Platycheirus brunnifrons Nielsen, 2004
Platycheirus carinatus (Curran, 1927)
? = hirtipes Kanervo, 1938
Platycheirus clypeatus (Meigen, 1822)
Platycheirus discimanus Loew, 1871
Platycheirus europaeus Goeldlin, Maibach & Speight, 1990
Platycheirus fulviventris (Macquart, 1829)
Platycheirus goeldlini Nielsen, 2004
Platycheirus groenlandicus Curran, 1927
= boreomontanus Nielsen, 1981
= monticolus Nielsen, 1972 preocc.
Platycheirus holarcticus Vockeroth, 1990
Platycheirus hyperboreus (Staeger, 1844)
Platycheirus immarginatus (Zetterstedt, 1849)
Platycheirus jaerensis Nielsen, 1971
Platycheirus kittilaensis Dušek & Láska, 1982
= complicatus misid.
Platycheirus laskai Nielsen, 1999
Platycheirus latimanus (Wahlberg, 1844)
Platycheirus lundbecki (Collin, 1931)
Platycheirus magadanensis Mutin, 1999
Platycheirus manicatus (Meigen, 1822)
Platycheirus modestus Ide, 1926
Platycheirus nielseni Vockeroth, 1990
Platycheirus nigrofemoratus Kanervo, 1934
Platycheirus occultus Goeldlin, Maibach & Speight, 1990
Platycheirus parmatus Rondani, 1857
Platycheirus peltatus (Meigen, 1822)
Platycheirus perpallidus Verral, 1901
Platycheirus podagratus (Zetterstedt, 1838)
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236
Platycheirus ramsarensis Goeldlin, Maibach & Speight, 1990
Platycheirus scambus (Staeger, 1843)
Platycheirus scutatus (Meigen, 1822)
Platycheirus splendidus Rotheray, 1998
Platycheirus sticticus (Meigen, 1822)
Platycheirus subordinatus Becker, 1915
Platycheirus tarsalis (Schummel, 1836)
Platycheirus transfugus (Zetterstedt, 1838)
Platycheirus urakawensis (Matsumura, 1919)
Platycheirus varipes Curran, 1923
sg. Pyrophaena Schiner, 1860
Platycheirus granditarsus (Forster, 1771)
Platycheirus rosarum (Fabricius, 1787)
XANTHANDRUS Verrall, 1901
Xanthandrus comtus (Harris, 1780)
tribe Paragini Glumac, 1961
PARAGUS Latreille, 1804
Paragus constrictus Šimić, 1986
Paragus nitimus Goeldlin, 1971
Paragus haemorrhous Meigen, 1822
= sigillatus Curtis, 1836
Paragus pecchiolii Rondani, 1857
= majoranae auct. nec Rondani, 1857
= albifrons misid.
Paragus tibialis (Fallén, 1817)
tribe Chrysotoxini Newman, 1834
CHRYSOTOXUM Meigen, 1803
Chrysotoxum arcuatum (Linnaeus, 1758)
= fasciatum (Müller, 1764)
Chrysotoxum bicinctum (Linnaeus, 1758)
Chrysotoxum cautum (Harris, 1776)
Chrysotoxum fasciolatum (De Geer, 1776)
Chrysotoxum festivum (Linnaeus, 1758)
= arcuatum auct. nec (Linnaeus, 1758)
Chrysotoxum octomaculatum Curtis, 1937
= elegans misid.
Chrysotoxum vernale Loew, 1841
tribe Syrphini Latreille, 1802
DASYSYRPHUS Enderlein, 1838
Dasysyrphus albostriatus (Fallén, 1817)
Dasysyrphus friuliensis (van der Goot, 1960)
= claviger (Frey, 1930)
= carpathicus Stys & Moucha, 1962
Checklist of the family Syrphidae (Diptera) of Finland 237
Dasysyrphus hilaris (Zetterstedt, 1843)
Dasysyrphus nigricornis (Verrall, 1873)
= obscuratus (Ringdahl, 1928)
Dasysyrphus pauxillus (Williston, 1887)
Dasysyrphus pinastri (De Geer, 1776)
= lunulatus (Meigen, 1822)
Dasysyrphus postclaviger Stys & Moucha, 1962
= claviger (Frey, 1930) preocc.
Dasysyrphus tricinctus (Fallén, 1817)
Dasysyrphus venustus (Meigen, 1822)
= arcuatus (Fallén, 1817)
DIDEA Macquart, 1834
Didea alneti (Fallén, 1817)
Didea fasciata Macquart, 1834
Didea intermedia Loew, 1854
DOROS Meigen, 1803
Doros profuges (Harris, 1780)
= conopseus auct. nec (Linnaeus, 1758)
EPISTROPHE Walker, 1852
Epistrophe annulitarsis Stackelberg, 1918
Epistrophe cryptica Doczkal & Schmid, 1994
Epistrophe diaphana (Zetterstedt, 1843)
Epistrophe eligans (Harris, 1780)
= bifasciata (Fabricius, 1794) preocc.
Epistrophe ava Doczkal & Schmid, 1994
= melanostomoides misid.
Epistrophe grossulariae (Meigen, 1822)
Epistrophe melanostoma (Zetterstedt, 1843)
= melanostomoides Strobl, 1910
Epistrophe nitidicollis (Meigen, 1822)
Epistrophe obscuripes (Strobl, 1910)
= similis Doczkal & Schmid, 1994
Epistrophe ochrostoma (Zetterstedt, 1849)
Epistrophe olgae Mutin, 1990
EPISTROPHELLA Dušek & Láska, 1967
Epistrophella euchroma (Kowarz, 1885)
EPISYRPHUS Matsumura & Adachi, 1917
Episyrphus balteatus (De Geer, 1776)
ERIOZONA Schiner, 1860
Eriozona syrphoides (Fallén, 1817)
EUPEODES Osten Sacken, 1877
= Metasyrphus Matsumura, 1917
Eupeodes abiskoensis (Dušek & Láska, 1973)
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238
Eupeodes biciki (Nielsen, 2003)
Eupeodes bucculatus (Rondani, 1857)
= latilunulatus (Collin, 1931)
Eupeodes corollae (Fabricius, 1794)
Eupeodes curtus (Hine, 1922)
Eupeodes duseki Mazánek, Láska & Bicik, 1999
Eupeodes goeldlini Mazánek, Láska & Bicik, 1999
Eupeodes latifasciatus (Macquart, 1829)
Eupeodes lundbecki (Soot-Ryen, 1946)
Eupeodes luniger (Meigen, 1822)
Eupeodes nielseni (Dušek & Láska, 1976)
Eupeodes nitens (Zetterstedt, 1843)
Eupeodes punctifer (Frey, 1934)
= borealis misid.
Eupeodes tirolensis (Dušek & Láska, 1973)
FAGISYRPHUS Dušek & Láska, 1967
Fagisyrphus cinctus (Fallén, 1817)
LAPPOSYRPHUS Dušek & Láska, 1967
Lapposyrphus lapponicus (Zetterstedt, 1838)
LEUCOZONA Schiner, 1860
sg. Leucozona Schiner, 1860
Leucozona inopinata Doczkal, 2000
Leucozona lucorum (Linnaeus, 1758)
sg. Ischyrosyrphus Bigot, 1882
Leucozona glaucia (Linnaeus, 1758)
Leucozona laternaria (Müller, 1776)
MEGASYRPHUS Duššek & Láska, 1967
Megasyrphus erraticus (Linnaeus, 1758)
= annulipes (Zetterstedt, 1838)
MELANGYNA Verrall, 1901
Melangyna arctica (Zetterstedt, 1838)
Melangyna barbifrons (Fallén, 1817)
Melangyna coei Nielsen, 1971
Melangyna compositarum (Verrall, 1873)
Melangyna lasiophthalma (Zetterstedt, 1843)
Melangyna lucifera Nielsen, 1980
Melangyna quadrimaculata (Verrall, 1873)
Melangyna umbellatarum (Fabricius, 1794)
MELIGRAMMA Frey, 1946
Meligramma guttata (Fallén, 1817)
Meligramma triangulifera (Zetterstedt, 1843)
MELISCAEVA Frey, 1946
Meliscaeva auricollis (Meigen, 1822)
Checklist of the family Syrphidae (Diptera) of Finland 239
Meliscaeva cinctella (Zetterstedt, 1843)
PARASYRPHUS Matsumura, 1917
Parasyrphus annulatus (Zetterstedt, 1838)
Parasyrphus groenlandicus (Nielsen, 1910)
Parasyrphus lineolus (Zetterstedt, 1843)
Parasyrphus macularis (Zetterstedt, 1843)
Parasyrphus malinellus (Collin, 1952)
Parasyrphus nigritarsis (Zetterstedt, 1843)
Parasyrphus proximus Mutin, 1990
Parasyrphus punctulatus (Verrall, 1873)
Parasyrphus tarsatus (Zetterstedt, 1838)
= dryadis (Holmgren, 1869)
Parasyrphus vittiger (Zetterstedt, 1843)
SCAEVA Fabricius, 1805
Scaeva pyrastri (Linnaeus, 1758)
Scaeva selenitica (Meigen, 1822)
SPHAEROPHORIA Lepeletier & Serville, 1828
Sphaerophoria abbreviata Zetterstedt, 1859
Sphaerophoria bankowskae Goeldlin, 1989
Sphaerophoria batava Goeldlin, 1974
Sphaerophoria boreoalpina Goeldlin, 1989
Sphaerophoria chongjini Bankowska, 1964
Sphaerophoria fatarum Goeldlin, 1989
Sphaerophoria interrupta (Fabricius, 1805)
= menthastri (Linnaeus, 1758)
Sphaerophoria kaa Violovitsh, 1960
Sphaerophoria laurae Goeldlin, 1989
Sphaerophoria loewi Zetterstedt, 1843
Sphaerophoria pallidula Mutin, 1999
Sphaerophoria philantha (Meigen, 1822)
= sarmatica Bankowska, 1964
Sphaerophoria rueppelli (Wiedemann, 1830)
Sphaerophoria scripta (Linnaeus, 1758)
Sphaerophoria taeniata (Meigen, 1822)
Sphaerophoria virgata Goeldlin, 1974
SYRPHUS Fabricius, 1775
Syrphus admirandus Goeldlin, 1996
Syrphus attenuatus Hine, 1922
= pilisquamus Ringdahl, 1928
Syrphus ribesii (Linnaeus, 1758)
= brevicinctus Kanervo, 1938
Syrphus sexmaculatus Zetterstedt, 1838
Syrphus torvus Osten Sacken, 1875
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Syrphus vitripennis Meigen, 1822
= rectus bretoletensis Goeldlin, 1996
XANTHOGRAMMA Schiner, 1860
Xanthogramma citrofasciatum (De Geer, 1776)
= festivum auct. nec (Linnaeus, 1758)
Xanthogramma pedissequum (Harris, 1776)
Xanthogramma stackelbergi Violovitsh, 1975
MILESIINAE Rondani, 1845
tribe Cerioidini Wahlgren, 1909
CERIANA Ranesque, 1815
Ceriana conopsoides (Linnaeus, 1758)
SPHIXIMORPHA Rondani, 1850
Sphiximorpha subsessilis (Illiger in Rossi, 1807)
tribe Cheilosini Williston, 1885
CHEILOSIA Meigen, 1822
Cheilosia alba Vujić & Claussen, 2000
Cheilosia albipila Meigen, 1838
Cheilosia albitarsis (Meigen, 1822)
Cheilosia alpina (Zetterstedt, 1838)
Cheilosia angustigenis Becker, 1894
Cheilosia barbata Loew, 1857
= honesta Rondani, 1868
Cheilosia barovskii (Stackelberg, 1930)
Cheilosia carbonaria Egger, 1860
Cheilosia chrysocoma (Meigen, 1822)
Cheilosia cynocephala Loew, 1840
Cheilosia avipes (Panzer, 1798)
Cheilosia avissima Becker, 1894
= pallipes auct. nec Loew, 1863
Cheilosia fraterna (Meigen, 1830)
Cheilosia frontalis Loew, 1857
Cheilosia gigantea (Zetterstedt, 1838)
= gracilis Hellén, 1914
Cheilosia grossa (Fallén, 1817)
Cheilosia illustrata (Harris, 1780)
Cheilosia impressa Loew, 1840
Cheilosia ingerae Nielsen & Claussen, 2001
Cheilosia lasiopa Kowarz, 1885
= honesta auct. nec Rondani, 1868
Cheilosia latifrons (Zetterstedt, 1843)
= intonsa Loew, 1857
Cheilosia longula (Zetterstedt, 1838)
= plumulifera Loew, 1857
Checklist of the family Syrphidae (Diptera) of Finland 241
= soror misid.
Cheilosia melanopa (Zetterstedt, 1843)
Cheilosia morio (Zetterstedt, 1838)
Cheilosia mutabilis (Fallén, 1817)
= ruralis (Meigen, 1822)
Cheilosia naruska Haarto & Kerppola, 2007
Cheilosia nebulosa Verrall, 1871
= langhoeri Becker, 1894
Cheilosia nigripes (Meigen, 1822)
Cheilosia pagana (Meigen, 1822)
Cheilosia proxima (Zetterstedt, 1843)
Cheilosia psilophthalma Becker, 1894
Cheilosia pubera (Zetterstedt, 1838)
Cheilosia reniformis Hellén, 1930
Cheilosia rumana Becker, 1894
Cheilosia sahlbergi Becker, 1894
Cheilosia scutellata (Fallén, 1817)
Cheilosia semifasciata Becker, 1894
Cheilosia sootryeni Nielsen, 1970
Cheilosia urbana (Meigen, 1822)
= ruralis (Meigen, 1822)
= praecox (Zetterstedt, 1843)
= globulipes Becker, 1894
Cheilosia uviformis Becker, 1894
= argentifrons Hellén, 1914
Cheilosia variabilis (Panzer, 1798)
Cheilosia velutina Loew, 1840
Cheilosia vernalis (Fallén, 1817)
= rotundicornis Hellén, 1914
= rotundiventris Becker, 1894
= rucollis Becker, 1894
Cheilosia vicina (Zetterstedt, 1849)
= nasutula Becker, 1894
Cheilosia vulpina (Meigen, 1822)
Cheilosia sp. A in Haarto & Kerppola, 2007
FERDINANDEA Rondani, 1844
Ferdinandea cuprea (Scopoli, 1763)
RHINGIA Scopoli, 1763
Rhingia borealis Ringdahl, 1928
= austriaca auct. nec Meigen, 1830
Rhingia campestris Meigen, 1822
tribe Chrysogasterini Shannon, 1922
BRACHYOPA Meigen, 1822
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Brachyopa cinerea Wahlberg, 1844
Brachyopa dorsata Zetterstedt, 1837
Brachyopa obscura ompson & Torp, 1982
Brachyopa pilosa Collin, 1939
= bicolor misid.
Brachyopa testacea (Fallén, 1817)
= conica misid.
Brachyopa vittata Zetterstedt, 1843
Brachyopa zhelochovtsevi Mutin, 1998
CHRYSOGASTER Meigen, 1803
Chrysogaster coemiteriorum (Linnaeus, 1758)
= cemiteriorum emend.
= chalybeata Meigen, 1822
Chrysogaster solstitialis (Fallén, 1817)
Chrysogaster virescens Loew, 1854
CHRYSOSYRPHUS Sedman, 1965
= Helleniola Stackelberg, 1965
Chrysosyrphus nasutus (Zetterstedt, 1838)
Chrysosyrphus niger (Zetterstedt, 1843)
= Myolepta nigra mistake
HAMMERSCHMIDTIA Schummel, 1834
Hammerschmidtia ferruginea (Fallén, 1817)
Hammerschmidtia ingrica Stackelberg, 1952
LEJOGASTER Rondani, 1857
Lejogaster metallina (Fabricius, 1781)
Lejogaster tarsata (Meigen, 1822)
= splendida (Meigen, 1822)
MELANOGASTER Rondani, 1857
Melanogaster aerosa (Loew, 1843)
= macquarti (Loew, 1843)
= viduata misid.
Melanogaster parumplicata (Loew, 1840)
NEOASCIA Williston, 1886
sg. Neoascia Williston, 1886
Neoascia podagrica (Fabricius, 1775)
Neoascia tenur (Harris, 1780)
= dispar (Meigen, 1822)
= lapponica Kanervo, 1934
= splendida Kanervo, 1934
sg. Neoasciella Stackelberg, 1970
Neoascia geniculata (Meigen, 1822)
Neoascia interrupta (Meigen, 1822)
Neoascia meticulosa (Scopoli, 1763)
Checklist of the family Syrphidae (Diptera) of Finland 243
= aenea (Meigen, 1822)
Neoascia obliqua Coe, 1940
Neoascia subchalybea Curran, 1925
= petsamoensis Kanervo, 1934
ORTHONEVRA Macquart, 1829
Orthonevra elegans (Meigen, 1822)
Orthonevra erythrogona (Malm, 1863)
Orthonevra geniculata (Meigen, 1830)
= linnaniemii Kanervo, 1934
Orthonevra intermedia Lundbeck, 1916
= rossica Stackelberg, 1953
Orthonevra nobilis (Fallén, 1817)
Orthonevra plumbago (Loew, 1840)
= brevicornis misid.
Orthonevra stackelbergi ompson & Torp, 1982
= intermedia misid.
SPHEGINA Meigen, 1822
sg. Sphegina Meigen, 1822
Sphegina clunipes (Fallén, 1816)
Sphegina elegans Schummel, 1843
= kimakowiczi misid.
Sphegina montana Becker, 1921
= violovitshi misid.
= verecunda misid.
Sphegina spheginea (Zetterstedt, 1838)
sg. Asiosphegina Stackelberg, 1975
Sphegina sibirica Stackelberg, 1953
tribe Eristalini Newman, 1834
ANASIMYIA Schiner, 1864
= Eurimyia Bigot, 1883
Anasimyia contracta Claussen & Torp, 1980
Anasimyia interpuncta (Harris, 1776)
Anasimyia lineata (Fabricius, 1787)
Anasimyia lunulata (Meigen, 1822)
Anasimyia transfuga (Linnaeus, 1758)
ERISTALINUS Rondani, 1845
Eristalinus aeneus (Scopoli, 1763)
Eristalinus sepulchralis (Linnaeus, 1758)
ERISTALIS Latreille, 1804
Eristalis abusiva Collin, 1931
Eristalis alpina (Panzer, 1798)
Eristalis anthophorina (Fallén, 1817)
Eristalis arbustorum (Linnaeus, 1758)
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= nemorum auct. nec (Linnaeus, 1758)
Eristalis cryptarum (Fabricius, 1794)
Eristalis fratercula Zetterstedt, 1838
= vallei Kanervo, 1934
Eristalis gomojunovae Violovitsh, 1977
= fratercula auct. nec Zetterstedt, 1838
Eristalis hirta Loew, 1866
= E. tundrarum Frey, 1932
Eristalis horticola (De Geer, 1776)
= lineata (Harris, 1776)
Eristalis intricaria (Linnaeus, 1758)
Eristalis nemorum (Linnaeus, 1758)
= interrupta (Poda, 1761)
Eristalis obscura Loew, 1866
= pseudorupium Kanervo, 1938
= vitripennis Strobl, 1893
Eristalis oestracea (Linnaeus, 1758)
Eristalis pertinax (Scopoli, 1763)
Eristalis picea (Fallén, 1817)
Eristalis rupium (Fabricius, 1805)
Eristalis similis (Fallén, 1817)
= pratorum (Meigen, 1822)
Eristalis tenax (Linnaeus, 1758)
HELOPHILUS Meigen, 1822
Helophilus anis Wahlberg, 1844
Helophilus bottnicus Wahlberg, 1844
Helophilus groenlandicus (Fabricius, 1780)
Helophilus hybridus Loew, 1846
Helophilus lapponicus Wahlberg, 1844
= borealis Staeger, 1845
Helophilus pendulus (Linnaeus, 1758)
Helophilus trivittatus (Fabricius, 1805)
= parallelus (Harris, 1776)
MALLOTA Meigen, 1822
Mallota megilliformis (Fallén, 1817)
Mallota tricolor Loew, 1871
MYATHROPA Rondani, 1845
Myathropa orea (Linnaeus, 1758)
PARHELOPHILUS Girschner, 1897
Parhelophilus consimilis (Malm, 1863)
Parhelophilus frutetorum (Fabricius, 1775)
Parhelophilus versicolor (Fabricius, 1794)
tribe Eumerini Smirnov, 1924
Checklist of the family Syrphidae (Diptera) of Finland 245
EUMERUS Meigen, 1822
Eumerus avitarsis Zetterstedt, 1843
Eumerus funeralis Meigen, 1822
= tuberculatus Rondani, 1857
Eumerus grandis Meigen, 1822
= annulatus (Panzer, 1798)
Eumerus rucornis Meigen, 1822
Eumerus sabulonum (Fallén, 1817)
Eumerus strigatus (Fallén, 1817)
tribe Merodontini Edwards, 1915
MERODON Meigen, 1803
Merodon equestris (Fabricius, 1794)
tribe Pelecocerini Williston, 1887
PELECOCERA Meigen, 1822
= Chamaesyrphus Mik, 1895
Pelecocera caledonica (Collin, 1940)
Pelecocera lusitanica (Mik, 1898)
Pelecocera scaevoides (Fallén, 1817)
Pelecocera tricincta Meigen, 1822
tribe Pipizini Williston, 1885
CRYPTOPIPIZA Mutin, 1998
Cryptopipiza notabila (Violovitsh, 1985)
HERINGIA Rondani, 1856
Heringia heringi (Zetterstedt, 1843)
NEOCNEMODON Goe, 1944
= Cnemodon Egger, 1865 preocc.
Neocnemodon fulvimanus (Zetterstedt, 1843)
Neocnemodon larusi (Vujić, 1999)
= fulvimanus auct. nec (Zetterstedt, 1843)
Neocnemodon latitarsis (Egger, 1865)
Neocnemodon pubescens (Delucchi & Pschorn-Walcher, 1955)
Neocnemodon verrucula (Collin, 1931)
Neocnemodon vitripennis (Meigen, 1822)
PIPIZA Fallén, 1810
Pipiza accola Violovitsh, 1985
Pipiza austriaca Meigen, 1822
Pipiza fasciata Meigen, 1822
Pipiza festiva Meigen, 1822
Pipiza lugubris (Fabricius, 1775)
Pipiza luteitarsis Zetterstedt, 1843
Pipiza noctiluca (Linnaeus, 1758)
Pipiza notata Meigen, 1822
= bimaculata Meigen, 1822
Antti Haarto & Sakari Kerppola / ZooKeys 441: 233–249 (2014)
246
Pipiza quadrimaculata (Panzer, 1804)
PIPIZELLA Rondani, 1856
Pipizella certa Violovitsh, 1981
= brevis auct. nec Lucas, 1976
Pipizella obscura Van Steenis & Lucas, 2011
Pipizella viduata (Linnaeus, 1758)
= virens misid.
TRICHOPSOMYIA Williston, 1888
Trichopsomyia avitarsis (Meigen, 1822)
Trichopsomyia joratensis Goeldlin, 1997
TRIGLYPHUS Loew, 1840
Triglyphus primus Loew, 1840
tribe Sericomyiini Rondani, 1845
SERICOMYIA Meigen, 1803
Sericomyia arctica Schirmer, 1913
Sericomyia jakutica (Stackelberg, 1927)
Sericomyia lappona (Linnaeus, 1758)
Sericomyia nigra Portschinsky, 1872
Sericomyia silentis (Harris, 1776)
tribe Volucellini Newman, 1834
VOLUCELLA Georoy, 1762
Volucella bombylans (Linnaeus, 1758)
Volucella inanis (Linnaeus, 1758)
Volucella pellucens (Linnaeus, 1758)
tribe Xylotini Bigot, 1883
BLERA Billberg, 1820
Blera eoa (Stackelberg 1928)
Blera fallax (Linnaeus, 1758)
BRACHYPALPOIDES Hippa, 1978
Brachypalpoides lentus (Meigen, 1822)
BRACHYPALPUS Macquart, 1834
Brachypalpus laphriformis (Fallén, 1816)
CHALCOSYRPHUS Curran, 1925
Chalcosyrphus jacobsoni (Stackelberg, 1921)
Chalcosyrphus nemorum (Fabricius, 1805)
Chalcosyrphus nigripes (Zetterstedt, 1838)
Chalcosyrphus piger (Fabricius, 1794)
Chalcosyrphus rupes (Loew, 1873)
Chalcosyrphus valgus (Gmelin, 1790)
= femoratus auct. nec Linnaeus, 1758
CRIORHINA Meigen, 1822
Criorhina asilica (Fallén, 1816)
LEJOTA Rondani, 1857
Checklist of the family Syrphidae (Diptera) of Finland 247
Lejota rucornis (Zetterstedt, 1843)
SPHECOMYIA Latreille, 1829
Sphecomyia vespiformis Gorski, 1852
SPILOMYIA Meigen, 1803
Spilomyia diophthalma (Linnaeus, 1758)
SYRITTA Lepeletier & Serville, 1828
Syritta pipiens (Linnaeus, 1758)
TEMNOSTOMA Lepeletier & Serville, 1828
Temnostoma angustistriatum Krivosheina, 2002
= bombylans auct. nec (Fabricius, 1805)
Temnostoma apiforme (Fabricius, 1794)
Temnostoma carens Gaunitz, 1936
Temnostoma sericomyiaeforme (Portschinsky, 1886)
Temnostoma vespiforme (Linnaeus, 1758)
TROPIDIA Meigen, 1822
Tropidia fasciata Meigen, 1822
Tropidia scita (Harris, 1780)
XYLOTA Meigen, 1822
Xylota caeruleiventris Zetterstedt, 1838
Xylota orum (Fabricius, 1805)
Xylota ignava (Panzer, 1798)
Xylota jakutorum Bagatshanova, 1980
= caeruleiventris auct. nec Zetterstedt, 1838
Xylota meigeniana Stackelberg, 1964
Xylota segnis (Linnaeus, 1758)
Xylota suecica (Ringdahl, 1943)
Xylota sylvarum (Linnaeus, 1758)
Xylota tarda Meigen, 1822
Xylota triangularis Zetterstedt, 1838
Xylota xanthocnema Collin, 1939
MICRODONTINAE Rondani, 1845
tribe Microdontini Rondani, 1845
MICRODON Meigen, 1803
Microdon analis (Macquart, 1842)
= eggeri Mik, 1897
Microdon miki Doczkal & Schmid, 1999
= latifrons auct. nec (Loew, 1856)
Microdon mutabilis (Linnaeus, 1758)
Excluded species
Microdon devius (Linnaeus, 1761) was not found within present borders.
Antti Haarto & Sakari Kerppola / ZooKeys 441: 233–249 (2014)
248
Neoascia annexa (Müller, 1776) was included in the Finnish species list in Fauna Eu-
ropaea (Speight 2004). Authors have not found any specimens or publications that
reliably show N. annexa to belong in the Finnish fauna.
Paragus punctulatus (Zetterstedt, 1838) was included in the Finnish species list in Fau-
na Europaea (Speight 2004). Authors have not found any specimens or publica-
tions that reliably show P. punctulatus to belong in the Finnish fauna.
Parasyrphus relictus (Zetterstedt, 1838): e status of P. relictus is unclear and the type
material is lost (Speight 2013). P. relictus was excluded from Finnish checklist by
Haarto and Kerppola (2007b).
Sphaerophoria sp. B was discovered as a deformed specimen of Sphaerophoria laurae
Goeldlin, 1989.
Temnostoma meridionale (Krivosheina & Mamayev, 1962) was included in the Finnish
species list in Fauna Europaea (Speight 2004). Authors have not found any specimens
or publications that reliably show T. meridionale to belong in the Finnish fauna.
Notes
Melanostoma mellinum (Linnaeus, 1758) and Melanostoma dubium (Zetterstedt,
1837) form a species complex and it is very doubtful whether present species concepts
provide an accurate reection of the number of Melanostoma species (Speight 2013).
Platycheirus modestus Ide, 1926. A previously unpublished record. A new species to the
Palaearctic. One male was collected in 4.7.2008 from Obb: Tornio (leg. I. Kakko).
Dasysyrphus spp. Both D. venustus (Meigen, 1822) and D. hilaris (Zetterstedt, 1843)
are known to contain several species (Haarto and Kerppola 2007a, Bartsch 2009a).
Cheilosia barovskii (Stackelberg, 1930). A previously unpublished record. Five fe-
males were collected in 30.5.1937 from Ta: Sääksmäki (leg. K.E. Kivirikko) and
one female in 20.5.2013 from N: Sipoo (leg. T. Järveläinen).
Myolepta nigra (Loew, 1872) was erroneously included in the list of Finnish ies
(Hackman 1980) instead of Helleniola nigra (Zetterstedt, 1843) = Chrysosyrphus
niger (Hackman 1982).
e used nomenclature of the species E. nemorum (Linnaeus, 1758), E. arbustorum
(Linnaeus, 1758) and E. horticola (De Geer, 1776) follows the proposal in Case
3259 by Chandler et al. (2004) and the usage of these names is conrmed in Opinion
2153 by ICZN (International Commission on Zoological Nomeclature) (2006).
Mallota tricolor Loew, 1871. A previously unpublished record. One male was col-
lected in 4.8.1977 from Ta: Hämeenlinna (leg. M. Raekunnas) and another male
in 13.6.–6.7.2013 from Sa: Joutseno (leg. J. Vilén).
Microdon mutabilis (Linnaeus, 1758) and Microdon myrmicae Schönrogge, Barr,
Wardlaw, Napper, Gardner, Breen, Elmes & omas, 2002 form a species
complex the species of which can be separate only by characters of the larvae and
the puparia (Bartsch 2009b). In the Finnish checklist M. mutabilis represents ei-
ther of the two species or include both of the species.
Checklist of the family Syrphidae (Diptera) of Finland 249
References
Bartsch H (2009a) Nationalnyckeln till Sveriges ora och fauna. Tvåvingar: Syrphidae: Syrphinae.
Artdatabanken, SLU, Uppsala.
Bartsch H (2009b) Nationalnyckeln till Sveriges ora och fauna. Tvåvingar: Syrphidae:
Eristalinae & Microdintinae. Artdatabanken, SLU, Uppsala.
Chandler PJ, Wakeham-Dawson A, McCullogh A (2004) Eristalis Latreille, 1804 (Insecta,
Diptera): proposed conrmation that the gender is feminine; Musca nemorum Linnaeus,
1758, M. arbustorum Linnaeus, 1758 and M. horticola De Geer, 1776 (currently Eristalis
nemorum, E. arbustorum and E. horticola): proposed conservation of usage of the specic
names by designation of neotypes. Bulletin of Zoological Nomenclature 61(4): 241–245.
Case 3259. http://biodiversityheritagelibrary.org/page/34353951
Haarto A, Kerppola S (2007a) Suomen kukkakärpäset ja lähialueiden lajeja. Suomen ympäris-
töministeriö, Helsinki, 647 pp.
Haarto A, Kerppola S (2007b) Muutoksia Suomen kukkakärpäsfaunaan (Diptera: Syrphidae).
Sahlbergia 13: 34–40. [in Finnish]
Hackman W (1980) A Check List of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Hackman W (1982) Tillägg och rättelser till förteckningen över Finlands Diptera, Smärre
meddelanden. Notulae entomologicae 62: 159.
ICZN (2006) Opinion 2153 (Case 3259) Eristalis Latreille, 1804 (Insecta, Diptera): conr-
mation that the gender is feminine; Musca nemorum Linnaeus, 1758, M. arbustorum Lin-
naeus, 1758 and M. horticola De Geer, 1776 (currently Eristalis nemorum, E. arbustorum
and E. horticola): usage of the specic names conserved by the designation of neotypes.
Bulletin of Zoological Nomenclature 63(2): 146–147. http://www.biodiversitylibrary.
org/page/34353034
Kerppola S (2013) Muutoksia Suomen kukkakärpäsfaunaan (Diptera, Syrphidae). Sahlbergia
19(1-2): 30–38. [in Finnish]
Kahanpää J (2010) Kärpäset. In: Rassi P, Hyvärinen E, Juslén A, Mannerkoski I (Eds) Suomen
lajien uhanalaisuus – Punainen kirja 2010. Ympäristöministeriö & Suomen ympäristökes-
kus, Helsinki, 490–504.
Marshall SA (2012) Flies: e Natural History and Diversity of Diptera. Firey, 305–310.
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Speight MCD (2004) Fauna Europaea: Syrphidae. In: Pape T, Beuk P. Fauna Europaea:
Diptera. Fauna Europaea version 1.0. http://www.faunaeur.org
Speight MCD (2013) Species accounts of European Syrphidae (Diptera), 2013. Syrph the Net,
the database of European Syrphidae, vol. 72. Syrph the Net publications, Dublin, 316 pp.
Van Veen MP (2004) Hoveries of Northwest Europe, Identication Keys to the Syrphidae.
KNNV Publishing, Utrecht, 254 pp.
Antti Haarto & Sakari Kerppola / ZooKeys 441: 233–249 (2014)
250
Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of Finland... 251
Checklist of the superfamilies Conopoidea, Diopsoidea
and Nerioidea of Finland (Insecta, Diptera)
Jere Kahanpää1, Jens-Hermann Stuke2
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
2 Roter Weg 22, 26789 Leer, Germany
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 28 February 2014|Accepted 28 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J, Stuke J-H (2014) Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of
Finland (Insecta, Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 251–257.
doi: 10.3897/zookeys.441.7227
Abstract
A checklist of the Diptera superfamilies Conopoidea (Conopidae), Nerioidea (Micropezidae, Pseudopo-
myzidae) and Diopsoidea (Megamerinidae, Psilidae, Strongylophthalmyiidae, Tanypezidae) from Finland
in presented. Myopa vicaria Walker, 1849 is formally recorded for the rst time from the country.
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
Four y families form the superfamily Nerioidea. Two of them, Micropezidae and
Pseudopomyzidae, occur in Finland. Neriids are mostly tropical and cypselomatids
(sensu stricto) have only been found in the Australasian and Oriental regions. e pseu-
dopomyzids were included in Cypselosomatidae Hendel, 1931 in the previous check-
list (Hackman 1980) and are still often included in Cypselosomatidae sensu lato. e
ZooKeys 441: 251–257 (2014)
doi: 10.3897/zookeys.441.7227
www.zookeys.org
Copyright J. Kahanpää, J.-H. Stuke. This is an open access article distributed under the terms of the Creative Commons Attribution International License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää & Jens-Hermann Stuke / ZooKeys 441: 251–257 (2014)
252
Finnish micropezid fauna is relatively well known. One or two species of Cnodacophora
Czerny, 1930 and Rainieria Rondani, 1843 could be present but are so far undetected.
Diopsoidea consists of seven to ten families (depending on the source), none par-
ticularly species-rich. Only four are found in Finland: Psilidae, Tanypezidae, Stron-
gylophthalmyiidae and Megamerinidae. Tanypezidae and Strongylophthalmyiidae
are here treated as separate families following Lonsdale (2013). Megamerinidae has
been assigned to several superfamilies, most often Diopsoidea following McAlpine
(1989), but also Nerioidea, Sciomyzoidea, and recently Opomyzoidea (Wiegmann et
al. 2011). ese families have raised no particular interest among Finnish dipterists.
e genus Chamaepsila Hendel, 1917 in particular needs more attention. It is likely
that a few Finnish species remain unrecognised in this genus.
e systematic position of the Conopidae is unclear. It has traditionally been con-
sidered basal within Schizophora. Wiegmann et al. (2011) listed Conopidae within
Sciomyzoidea while others (see Skevington et al. 2010) place the family near Tephri-
toidea. Within the family the recent classication of Gibson and Skevington (2013)
is followed. e Finnish conopid fauna is now relatively well-understood and few ad-
ditional species are expected to occur in the country.
e Finnish species of these families were last listed by Hackman (1980).
Checklist part 1: Conopoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
? clade Archischiza Enderlein, 1936
superfamily Conopoidea Latreille, 1802
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Conopidae ~796 83 19 good
Nerioidea:
Micropezidae 573 22 6 average
Pseudopomyzidae 18 1 1 good
Diopsoidea:
Psilidae 320 47 29 average
Tanypezidae 27 (Lonsdale 2013) 1 1 good
Strongylophthalmyiidae 47 (Lonsdale 2013) 2 2 good
Megamerinidae 15 1 1 good
Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of Finland... 253
CONOPIDAE Latreille, 1802
CONOPINAE Latreille, 1802
CONOPS Linnaeus, 1758
sg. Conops Linnaeus, 1758
Conops quadrifasciatus De Geer, 1776
Conops strigatus Wiedemann, 1824
Conops vesicularis Linnaeus, 1761
PHYSOCEPHALA Schiner, 1861
Physocephala nigra (De Geer, 1776)
ZODIONINAE Rondani, 1856
ZODION Latreille, 1796
Zodion cinereum (Fabricius, 1794)
= notatum Meigen, 1804
SICINAE Zimina, 1960
SICUS Scopoli, 1763
Sicus ferrugineus (Linnaeus, 1761)
MYOPINAE Macquart, 1834
MYOPA Fabricius, 1775
Myopa buccata (Linnaeus, 1758)
Myopa fasciata Meigen, 1804
Myopa hirsuta Stuke & Clements, 2008
= strandi auct. nec Duda, 1940
Myopa occulta Wiedemann, 1824
= nigrifrons von Bonsdor, 1866
Myopa tessellatipennis Motschulsky, 1859
= polystigma auct. nec Rondani, 1857
Myopa testacea (Linnaeus, 1767)
Myopa vicaria Walker, 1849
= villosa Ringdahl, 1945
= strandi Duda, 1940
MYOPOTTA Zimina, 1969
Myopotta pallipes (Wiedemann, 1824)
THECOPHORA Rondani, 1845
= Occemyia Robineau-Desvoidy, 1853
ecophora cinerascens (Meigen, 1804)
= pusilla (Meigen, 1824)
= atra misid.
ecophora distincta (Wiedemann, 1824)
= melanopa misid.
ecophora fulvipes (Robineau-Desvoidy, 1830)
= sundewalli (Zetterstedt, 1844)
ecophora jakutica Zimina, 1974
= atra misid.
Jere Kahanpää & Jens-Hermann Stuke / ZooKeys 441: 251–257 (2014)
254
DALMANNIINAE Hendel, 1916
DALMANNIA Robineau-Desvoidy, 1830
Dalmannia dorsalis (Fabricius, 1794)
= punctata misid.
Checklist part 2: Nerioidea and Diopsoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Nerioidea Westwood, 1840
MICROPEZIDAE Blanchard, 1840
CALOBATINAE Bigot, 1853
CALOBATA Meigen, 1803
Calobata petronella (Linnaeus, 1761)
NERIA Robineau-Desvoidy, 1830
Neria cibaria (Linnaeus, 1761)
Neria commutata (Czerny, 1930)
= nigricornis auct. nec Zetterstedt, 1838
Neria ephippium (Fabricius, 1794)
Neria nigricornis (Zetterstedt, 1838)
= helleni (Frey, 1918)
= nitidicollis (Frey, 1947)
MICROPEZINAE Blanchard, 1840
MICROPEZA Meigen, 1803
Micropeza corrigiolata (Linnaeus, 1767)
PSEUDOPOMYZIDAE McAlpine, 1966
PSEUDOPOMYZA Strobl, 1893
Pseudopomyza atrimana (Meigen, 1830)
superfamily Diopsoidea Billberg, 1820
MEGAMERINIDAE Hendel, 1913
MEGAMERINA Rondani, 1861
= Lissa Meigen, 1826 preocc.
Megamerina dolium (Fabricius, 1805)
= loxocerina (Fallén, 1820)
Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of Finland... 255
PSILIDAE Macquart, 1835
CHYLIZINAE Rondani, 1856
CHYLIZA Fallén, 1820
sg. Chyliza Fallén, 1820
Chyliza annulipes Macquart, 1835
= fuscipennis auct. nec Robineau-Desvoidy, 1830
Chyliza leptogaster (Panzer, 1798)
= scutellata (Fabricius, 1798)
Chyliza nova Collin, 1944
Chyliza vittata Meigen, 1826
PSILINAE Macquart, 1835
tribe Loxocerini Macquart, 1835
IMANTIMYIA Frey, 1925
Imantimyia albiseta (Schrank, 1803)
= ichneumonea (Linnaeus, 1758) nom. dubium (see Notes)
Imantimyia fulviventris (Meigen, 1826)
Imantimyia nigrifrons (Macquart, 1835)
Imantimyia sylvatica (Meigen, 1826)
LOXOCERA Meigen, 1803
sg. LOXOCERA Meigen, 1803
Loxocera aristata (Panzer, 1801)
= ichneumonea (Linnaeus, 1758) nom. dubium (see Notes)
tribe Psilini Macquart, 1835
CHAMAEPSILA Hendel, 1917
= Tetrapsila Frey, 1925
Chamaepsila atra (Meigen, 1826)
Chamaepsila bicolor (Meigen, 1826)
= nigromaculata (Strobl, 1909)
Chamaepsila buccata (Fallén, 1826)
= gracilis (Meigen, 1826)
Chamaepsila humeralis (Zetterstedt, 1847)
Chamaepsila limbatella (Zetterstedt, 1847)
Chamaepsila morio (Zetterstedt, 1835)
Chamaepsila nigra (Fallén, 1820)
Chamaepsila nigricornis (Meigen, 1826)
Chamaepsila nigrosetosa Frey, 1925
Chamaepsila obscuritarsis (Loew, 1856)
Chamaepsila pallida (Fallén, 1820)
Chamaepsila pectoralis (Meigen, 1826)
Chamaepsila rosae (Fabricius, 1794)
Chamaepsila rufa (Meigen, 1826)
Chamaepsila unilineata (Zetterstedt, 1847)
PSILA Meigen, 1803
Jere Kahanpää & Jens-Hermann Stuke / ZooKeys 441: 251–257 (2014)
256
sg. FREYOPSILA Shatalkin, 1986
Psila sibirica (Frey, 1925)
sg. PSILA Meigen, 1803
Psila metaria (Linnaeus, 1761)
Psila merdaria Collin, 1944
PSILOSOMA Zetterstedt, 1860
Psilosoma audouini (Zetterstedt, 1835)
Psilosoma lefebvrei (Zetterstedt, 1835)
STRONGYLOPHTHALMYIIDAE Hendel, 1917
STRONGYLOPHTHALMYIA Heller, 1902
Strongylophthalmyia pictipes Frey, 1935
Strongylophthalmyia ustulata (Zetterstedt, 1847)
TANYPEZIDAE Rondani, 1856
TANYPEZA Fallén, 1820
Tanypeza longimana Fallén, 1820
Excluded species
Dalmannia punctata (Fabricius, 1794) not found within present borders.
Notes
Chamaepsila unilineata (Zetterstedt, 1847) and C. nigrosetosa Frey, 1925 were
synonymized with C. pallida but they have recently been reinstated as valid species
(Shatalkin and Merz 2010).
Loxocera ichneumonea (Linnaeus, 1758). is name has been used for L. aristata
(Panzer) and L. albiseta (Schrank). Unfortunately no type material is known. Linné’s
original description is insucient and may in fact belong to a hovery (see Buck and
Marshall 2006).
Myopa vicaria Walker, 1849. One of the common Myopa species through Finland,
but previously unreported due to diculties in identication. Recorded from
provinces Ab, N, Ka, St, Ta, Sa, Kl, Om, Ok, Lkoc and Li.
References
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genetic redenition of Holarctic subgenera and species groups. European Journal of Ento-
mology 103: 193–219. doi: 10.14411/eje.2006.022
Checklist of the superfamilies Conopoidea, Diopsoidea and Nerioidea of Finland... 257
Gibson JF, Skevington JH (2013) Phylogeny and taxonomic revision of all genera of Conopidae
(Diptera) based on morphological data. Zoological Journal of the Linnean Society 167(1):
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McAlpine JF (1989) Phylogeny and classication of the Muscomorpha. In: McAlpine JF,
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Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)259
Checklist of the Diptera superfamilies Tephritoidea
and Sciomyzoidea of Finland (Insecta)
Jere Kahanpää1, Kaj Winqvist2
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
2 Mikonkatu 3 C 52, FI–20100 Turku, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 30 January 2014|Accepted 6 March 2014|Published 19 Septembers 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J, Winqvist K (2014) Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of
Finland (Insecta). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 259–275. doi:
10.3897/zookeys.441.7143
Abstract
A revised checklist of the ies of superfamilies Tephritoidea and Sciomyzoidea of Finland is provided. e
following families are covered: Eurygnathomyiidae, Lonchaeidae, Neottiophilidae, Pallopteridae, Piophi-
lidae, Platystomatidae, Tephritidae, Ulidiidae (Tephritoidea); Coelopidae, Dryomyzidae, Heterocheilidae,
Phaeomyiidae, Sciomyzidae, Sepsidae (Sciomyzoidea).
Keywords
Finland, Diptera, checklist, Tephritoidea, Sciomyzoidea
Introduction
With over 7800 known extant species (Pape et al. 2011), Tephritoidea is one of the
larger Diptera superfamilies. e nominotypical family of Tephritoidea, the fruit ies
(Tephritidae), includes over half of the currently known species in the superfamily. e
highest diversity of tephritoids occurs in the tropics. Six of the nine extant tephritoid
families have been found in Finland: no richardiid, ctenostylid or pyrgotid have been
found in the country. e Tephritidae, Ulidiidae and Platystomatidae (plus Pyrgoti-
dae) form a well-dened, probably monophyletic group. e other tephritoid families
are more basal in their phylogenetic position (Korneyev 1999). e exact relationships
ZooKeys 441: 259–275 (2014)
doi: 10.3897/zookeys.441.7143
www.zookeys.org
Copyright J. Kahanpää, K.Winqvist. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää & Kaj Winqvist / ZooKeys 441: 259–275 (2014)
260
between the families are far from settled and some authors exclude Pallopteridae and
Eurygnathomyiidae from the superfamily (Wiegmann et al. 2011). Several subfamily
schemes have been proposed for Tephritidae. e classication by Korneyev (2000)
is adopted here. For Pallopteridae, Piophilidae (with Neottiophilidae) and Ulidiidae
we follow McAlpine (1981), Ozerov (2005) and Kameneva and Korneyev (2006) re-
spectively. Norrbom et al. (1999) provide a catalogue of world tephritid names. e
Finnish tephritoid ies were last listed by Winqvist and Kahanpää (2007). Hackman
(1956) reviewed the Lonchaeidae of Eastern Fennoscandia. is paper is now some-
what outdated, but still valuable.
Sciomyzoidea is a smaller superfamily with some 1150 described species (Pape et
al. 2011). It includes two medium-sized families (Sciomyzidae and Sepsidae) and nine
smaller families comprising at most 30 species. e millipede parasitoids of family
Phaeomyiidae are sometimes classied as a subfamily of Sciomyzidae. e coelopids
and heterocheilids, with just one species each in Finland, are closely associated with
stranded marine seaweed. e family placement of Heterocheila has been rather un-
stable: during the last 40 years it has been placed in Helcomyzidae, Dryomyzidae
and Coelopidae or regarded as having a separate family status. Here we follow the
recent world catalogue (Mathis 2011) and keep Heterocheilidae as a separate family.
e family name Heterocheilidae is preoccupied by Heterocheilidae Railliet & Henry,
1915 in Nematoda (Mathis 2011). is homonymy has not yet been resolved.
World catalogues have recently been published for Coelopidae (Mathis and
McAlpine 2011) Dryomyzidae (Mathis and Sueyoshi 2011), Heterocheilidae (Mathis
2011), Sciomyzidae (Rozkošný 1995) and Sepsidae (Ozerov 2005). e Finnish scio-
myzoid ies were last listed by Winqvist and Kahanpää (2007).
Table 1. Number of species in tephritoid and sciomyzoid families.
Family Number of species in Level of
knowledge
World (Pape et al. 2011) Europe (Pape and Beuk 2013) Finland
Tephridoidea:
Eurygnathomyiidae 1 1 1 good
Lonchaeidae 504 96 41–44 average
Neottiophilidae 4 2 2 average
Pallopteridae 70 22 13 good
Piophilidae 78 27 16 average
Platystomatidae 1162 21 2 good
Tephritidae 4712 267 69 average–good
Ulidiidae 675 106 16 good
Sciomyzoidea:
Coelopidae 35 3 1 average
Dryomyzidae 22 5 5 average
Heterocheilidae 2 1 1 average
Phaeomyiidae 3 3 2 good
Sciomyzidae 605 136 73–74 good
Sepsidae 340 48 32 good
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)261
Checklist part 1: Tephritoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Tephritoidea Newman, 1834
EURYGNATHOMYIIDAE Czerny, 1934
EURYGNATHOMYIA Czerny, 1904
Eurygnathomyia bicolor (Zetterstedt, 1837)
LONCHAEIDAE Rondani, 1856
DASIOPINAE Morge, 1963
Dasiopini Morge, 1963
DASIOPS Rondani, 1856
Dasiops appendiculus Morge, 1959
Dasiops facialis Collin, 1953
Dasiops mucronatus Morge, 1959
= hennigi misid.
= latiterebra misid.
Dasiops occultus Collin, 1953
= albiceps (Frey, 1930) preocc.
Dasiops perpropinquus Morge, 1959
Dasiops spatiosus (Becker, 1895)
Dasiops trichosternalis Morge, 1959
EAROMYIINAE Morge, 1963
CHAETOLONCHAEA Czerny, 1934
Chaetolonchaea pallipennis (Zetterstedt, 1855)
= dasyops misid.
EAROMYIA Zetterstedt, 1842
Earomyia lonchaeoides Zetterstedt, 1848
Earomyia schistopyga Collin, 1953
Earomyia viridana (Meigen, 1826)
PROTEAROMYIA McAlpine, 1962
Protearomyia nigra (Meigen, 1826)
LONCHAEINAE Rondani, 1856
Lonchaeini Rondani, 1856
LONCHAEA Fallén, 1820
Lonchaea anis Malloch, 1920
= laxa auct. nec Collin, 1953
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Lonchaea albigena Collin, 1953
Lonchaea albitarsis Zetterstedt, 1838
? Lonchaea bruggeri Morge, 1967
Lonchaea bukowskii Czerny, 1934
Lonchaea caledonica MacGowan & Rotheray, 2000
= laticornis auct. nec Meigen, 1826
Lonchaea carpathica Kovalev, 1974
Lonchaea chorea (Fabricius, 1781)
Lonchaea collini Hackman, 1956
? Lonchaea contigua Collin, 1953
Lonchaea corusca Czerny, 1934
= alni Ringdahl, 1947
= lauta Collin, 1953
= britteni Collin, 1953
Lonchaea defecta McAlpine, 1964
Lonchaea deutschi Zetterstedt, 1838
= sarekensis Frey, 1916
? Lonchaea dicilis Hackman, 1956
Lonchaea fraxina MacGowan & Rotheray, 2000
Lonchaea freyi Czerny, 1934
Lonchaea fugax Becker, 1895
= cariecola Czerny, 1934
Lonchaea hackmani Kovalev, 1981
= peregrina auct. nec Becker, 1895
Lonchaea ipsiphaga McAlpine, 1964
= maniola misid.
Lonchaea limatula Collin, 1953
= avidipennis auct. nec Zetterstedt, 1847
Lonchaea nitens (Bigot, 1885)
= krogerusi Czerny, 1934
Lonchaea palposa Zetterstedt, 1847
Lonchaea patens Collin, 1953
Lonchaea ragnari Hackman, 1956
Lonchaea scutellaris Rondani, 1874
Lonchaea sororcula Hackman, 1956
Lonchaea stackelbergi Czerny, 1934
Lonchaea subneatosa Kovalenko, 1974
Lonchaea sylvatica Beling, 1873
= lucidiventris Becker, 1895
Lonchaea tarsata Fallen, 1820
Lonchaea ultima Collin, 1953
Lonchaea zetterstedti Becker, 1902
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)263
NEOTTIOPHILIDAE Hendel, 1916
ACTENOPTERA Czerny, 1904
Actenoptera hilarella (Zetterstedt, 1847)
NEOTTIOPHILUM Frauenfeld, 1868
Neottiophilum praeustum (Meigen, 1826)
PALLOPTERIDAE Loew, 1862
PALLOPTERA Fallén, 1820
Palloptera anderssoni Rotheray & MacGowan, 1999
Palloptera formosa Frey, 1930
Palloptera marginata (Meigen, 1826)
= costalis Loew, 1873
Palloptera umbellatarum (Fabricius, 1775)
= arcuata auct. nec (Fabricius, 1781)
Palloptera ustulata Fallén, 1820
TEMNOSIRA Enderlein, 1936
Temnosira ambusta (Meigen, 1826)
Temnosira saltuum (Linnaeus, 1758)
TOXONEURA Macquart, 1835
Toxoneura ephippium (Zetterstedt, 1860)
Toxoneura laetabilis (Loew, 1873)
Toxoneura modesta (Meigen, 1830)
= umbellatarum auct. nec (Fabricius, 1775)
Toxoneura trimacula (Meigen, 1826)
Toxoneura usta (Meigen, 1826)
Toxoneura venusta (Loew, 1858)
= atriventris (Ringdahl, 1947)
PIOPHILIDAE Macquart, 1835
PIOPHILINAE Macquart, 1835
ALLOPIOPHILA Hendel, 1917
= Arctopiophila Duda, 1924
= Boreopiophila Frey, 1930
= Parapiophila McAlpine, 1977
Allopiophila calceata (Duda, 1924)
Allopiophila avipes (Zetterstedt, 1847)
Allopiophila lonchaeoides (Zetterstedt, 1838)
Allopiophila luteata (Haliday, 1833)
Allopiophila pectiniventris (Duda, 1924)
Allopiophila tomentosa Frey, 1930
Allopiophila vulgaris (Fallén, 1820)
Allopiophila sp. A
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AMPHIPOGON Wahlberg, 1845
Amphipogon avus (Zetterstedt, 1838)
= spectrum Wahlberg, 1845
MYCETAULUS Loew, 1845
Mycetaulus bipunctatus (Fallén, 1823)
PIOPHILA Fallén, 1810
Piophila casei (Linnaeus, 1758)
PROCHYLIZA Walker, 1849
= Liopiophila Duda, 1924
Prochyliza nigrimana (Meigen, 1826)
Prochyliza varipes (Meigen, 1830)
= lundbecki misid.
PSEUDOSEPS Becker, 1902
Pseudoseps signata (Fallén, 1820)
STEARIBIA Lioy, 1864
Stearibia nigriceps (Meigen, 1826)
= foveolata (Meigen, 1826)
= coerulescens (Zetterstedt, 1847)
PLATYSTOMATIDAE Schiner, 1862
PLATYSTOMA Meigen, 1803
Platystoma seminationis (Fabricius, 1775)
RIVELLIA Robineau-Desvoidy, 1830
Rivellia syngenesiae (Fabricius, 1781)
TEPHRITIDAE Newman, 1834
TRYPETINAE Loew, 1861 sensu lato
Adramini Hendel, 1914
EUPHRANTA Loew, 1862
sg. Euphranta Loew, 1862
Euphranta connexa (Fabricius, 1794)
sg. Rhacochlaena Loew, 1862
Euphranta toxoneura (Loew, 1846)
Trypetini Loew, 1861
ACIDIA Robineau-Desvoidy, 1830
Acidia cognata (Wiedemann, 1817)
ANOMOIA Walker, 1835
= Phagocarpus Rondani, 1870
Anomoia purmunda (Harris, 1776)
CHETOSTOMA Rondani, 1856
Chetostoma stackelbergi (Rohdendorf, 1955)
CORNUTRYPETA Han, Wang & Kim, 1993
Cornutrypeta spinifrons (Schroeder, 1913)
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)265
Cornutrypeta superciliata (Frey, 1935)
EULEIA Walker, 1835
= Cryptaciura Hendel, 1927
Euleia heraclei (Linnaeus, 1758)
Euleia rotundiventris (Fallén, 1814)
MYOLEJA Rondani, 1856
Myoleja lucida (Fallén, 1826)
PHILOPHYLLA Rondani, 1870
Philophylla caesio (Harris, 1776)
RHAGOLETIS Loew, 1862
Rhagoletis alternata (Fallén, 1814)
Rhagoletis cerasi (Linnaeus, 1758)
Rhagoletis meigenii (Loew, 1844)
TRYPETA Meigen, 1803
Trypeta artemisiae (Fabricius, 1794)
Trypeta immaculata (Macquart, 1835)
= hamifera Loew, 1846
Trypeta zoe Meigen, 1826
TEPHRITINAE Newman, 1834
Terellini Hendel, 1927
CHAETORELLIA Hendel, 1927
Chaetorellia jaceae (Robineau-Desvoidy, 1830)
CHAETOSTOMELLA Hendel, 1927
Chaetostomella cylindrica (Robineau-Desvoidy, 1830)
= onotrophes (Loew, 1848)
ORELLIA Robineau-Desvoidy, 1830
Orellia falcata (Scopoli, 1763)
TERELLIA Robineau-Desvoidy, 1830
sg. Cerajocera Rondani, 1856
Terellia ceratocera (Hendel, 1913)
Terellia plagiata (Dahlbom, 1850)
Terellia tussilaginis (Fabricius, 1775)
sg. Terellia Robineau-Desvoidy, 1830
Terellia colon (Meigen, 1826)
Terellia rucauda (Fabricius, 1794)
Terellia serratulae (Linnaeus, 1758)
Terellia winthemi (Meigen, 1826)
Xyphosiini Hendel, 1927
XYPHOSIA Robineau-Desvoidy, 1830
Xyphosia miliaria (Schrank, 1781)
Noeetini Norrbom & Korneyev, 1999
ENSINA Robineau-Desvoidy, 1830
Ensina sonchi (Linnaeus, 1767)
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NOEETA Robineau-Desvoidy, 1830
= Oplocheta Rondani, 1856
Noeeta pupillata (Fallén, 1814)
Myopitini Bezzi, 1910
EURASIMONA Korneyev & White, 1991
Eurasimona stigma (Loew, 1840)
UROPHORA Robineau-Desvoidy, 1830
Urophora aprica (Fallén, 1820)
Urophora cardui (Linnaeus, 1758)
Urophora cuspidata (Meigen, 1826)
Urophora jaceana (Hering, 1935)
Urophora solstitialis (Linnaeus, 1758)
= sonderupi (Hering, 1940)
Urophora stylata (Fabricius, 1775)
Dithrycini Hendel, 1927
DITHRYCA Rondani, 1856
Dithryca guttularis (Meigen, 1826)
Eutretini Munro, 1952
XANTHOMYIA Phillips, 1923
= Paracarphotricha Hendel, 1927
Xanthomyia alpestris (Pokorny, 1887)
= pseudoradiata (Becker, 1900)
Tephritini Newman, 1834
CAMPIGLOSSA Rondani, 1870
Campiglossa absinthii (Fabricius, 1805)
= parvula (Loew, 1862)
Campiglossa argyrocephala (Loew, 1844)
Campiglossa dicilis (Hendel, 1927)
= tessellata misid.
Campiglossa grandinata (Rondani, 1870)
= borealis Portschinsky, 1875
Campiglossa guttella (Rondani, 1870)
= achyrophori misid.
= producta misid.
Campiglossa loewiana (Hendel, 1927)
Campiglossa plantaginis (Haliday, 1833)
Campiglossa punctella (Fallén, 1814)
Campiglossa solidaginis (White, 1986)
Campiglossa sp. A
DIOXYNA Frey, 1945
Dioxyna bidentis (Robineau-Desvoidy, 1830)
= sororcula auct. nec (Wiedemann, 1830)
HERINGINA Aczél, 1940
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)267
Heringina guttata (Fallén, 1814)
OXYNA Robineau-Desvoidy, 1830
Oxyna avipennis (Loew, 1844)
Oxyna nebulosa (Wiedemann, 1817)
Oxyna parietina (Linnaeus, 1758)
SPHENELLA Robineau-Desvoidy, 1830
Sphenella marginata (Fallén, 1814)
TEPHRITIS Latreille, 1804
Tephritis angustipennis (Loew, 1844)
Tephritis bardanae (Schrank, 1803)
Tephritis cometa (Loew, 1840)
Tephritis conura (Loew, 1844)
Tephritis dilacerata (Loew, 1846)
Tephritis fallax (Loew, 1844)
Tephritis hyoscyami (Linnaeus, 1758)
Tephritis leontodontis (De Geer, 1776)
Tephritis mutabilis Merz, 1992
Tephritis neesii (Meigen, 1830)
= nesii misspelling
Tephritis ruralis (Loew, 1844)
Tephritis sp. cf. rydeni Hering, 1956
= dioscurea misid.
TRUPANEA Schrank, 1795
Trupanea stellata (Fuessly, 1775)
Unplaced in Tephritinae (incertae sedis)
ACINIA Robineau-Desvoidy, 1830
Acinia corniculata (Zetterstedt, 1819)
ULIDIIDAE Macquart, 1835
OTITINAE Aldrich, 1932
Myennidini Kameneva & Korneyev, 2006
PSEUDOTEPHRITIS Johnson, 1802
Pseudotephritis trypetoptera (Hennig, 1939)
= corticalis auct. nec (Loew, 1873)
Otitini Aldrich, 1932
CEROXYS Macquart, 1835
Ceroxys urticae (Linnaeus, 1758)
HERINA Robineau-Desvoidy, 1830
Herina frondescentiae (Linnaeus, 1758)
Herina paludum (Fallén, 1820)
Herina palustris (Meigen, 1826)
MELIERIA Robineau-Desvoidy, 1830
sg. Melieria Robineau-Desvoidy, 1830
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268
Melieria crassipennis (Fabricius, 1794)
Melieria omissa (Meigen, 1826)
= obscuripes auct. nec (Loew, 1873)
TETANOPS Fallén, 1820
sg. Eurycephalomyia Hendel, 1907
Tetanops sintenisi Becker, 1909
sg. Tetanops Fallén, 1820
Tetanops myopinus Fallén, 1820
ULIDIINAE Macquart, 1835
Seiopterini Kameneva & Korneyev, 1994
HOMALOCEPHALA Zetterstedt, 1838
Homalocephala albitarsis Zetterstedt, 1838
= bipunctata (Loew, 1854)
Homalocephala angustata (Wahlberg, 1839)
Homalocephala apicalis (Wahlberg, 1839)
= biseta Frey, 1908
Homalocephala bimaculata (Wahlberg, 1839)
Homalocephala biumbrata (Wahlberg, 1838)
= albitarsis auct. nec Zetterstedt, 1838
SEIOPTERA Kirby, 1817
Seioptera vibrans (Linnaeus, 1758)
Ulidiini Macquart, 1835
PHYSIPHORA Fallén, 1810
Physiphora alceae (Preyssler, 1791)
= demandata (Fabricius, 1798)
Checklist part 2: Sciomyzoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Sciomyzoidea Fallén, 1820
COELOPIDAE Hendel, 1910
COELOPA Meigen, 1830
sg. Fucomyia Haliday, 1838
Coelopa frigida (Fabricius, 1805)
DRYOMYZIDAE Schiner, 1862
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)269
DRYOMYZA Fallén, 1820
= Neuroctena Rondani, 1868
Dryomyza anilis Fallén, 1820
DRYOPE Robineau-Desvoidy, 1830
Dryope decrepita (Zetterstedt, 1838)
Dryope aveola (Fabricius, 1794)
PARADRYOMYZA Ozerov, 1987
Paradryomyza spinigera Ozerov, 1987
PSEUDONEUROCTENA Ozerov, 1987
Pseudoneuroctena senilis (Zetterstedt, 1838)
HETEROCHEILIDAE McAlpine, 1991
HETEROCHEILA Rondani, 1857
Heterocheila buccata (Fallén, 1820)
PHAEOMYIIDAE Verbeke, 1950
PELIDNOPTERA Rondani, 1856
Pelidnoptera fuscipennis (Meigen, 1830)
= fumipennis (Zetterstedt, 1846)
Pelidnoptera nigripennis (Fabricius, 1794)
SCIOMYZIDAE Fallén, 1820
SCIOMYZINAE Fallén, 1820
Sciomyzini Fallén, 1820
COLOBAEA Zetterstedt, 1837
Colobaea bifasciella (Fallén, 1820)
Colobaea distincta (Meigen, 1830)
Colobaea nigroaristata Rozkošný, 1984
Colobaea pectoralis (Zetterstedt, 1847)
Colobaea punctata (Lundbeck, 1923)
DITAENIELLA Sack, 1939
Ditaeniella grisescens (Meigen, 1830)
PHERBELLIA Robineau-Desvoidy, 1830
Pherbellia albocostata (Fallén, 1820)
Pherbellia alpina (Frey, 1930)
Pherbellia argyra Verbeke, 1967
Pherbellia brunnipes (Meigen, 1838)
Pherbellia cinerella (Fallén, 1820)
Pherbellia dubia (Fallén, 1820)
Pherbellia goberti (Pandellé, 1902)
= stylifera Rozkošný, 1982
Pherbellia griseicollis (Becker, 1900)
= lapponica (Ringdahl, 1948)
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270
Pherbellia griseola (Fallén, 1820)
Pherbellia hackmani Rozkošný, 1982
Pherbellia nana (Fallén, 1820)
Pherbellia obscura (Ringdahl, 1948)
Pherbellia obtusa (Fallén, 1820)
Pherbellia pallidiventris (Fallén, 1820)
Pherbellia rozkosnyi Verbeke, 1967
= scutellaris misid.
Pherbellia schoenherri (Fallén, 1826)
Pherbellia sordida (Hendel, 1902)
Pherbellia stackelbergi Elberg, 1965
Pherbellia ventralis (Fallén, 1820)
PTEROMICRA Lioy, 1864
Pteromicra angustipennis (Staeger, 1845)
Pteromicra glabricula (Fallén, 1820)
Pteromicra leucopeza (Meigen, 1838)
Pteromicra oldenbergi (Hendel, 1902)
Pteromicra pectorosa (Hendel, 1902)
SCIOMYZA Fallén, 1820
Sciomyza dryomyzina Zetterstedt, 1846
Sciomyza sebezhica Przhiboro, 2001
Sciomyza simplex Fallén, 1820
TETANURA Fallén, 1820
Tetanura pallidiventris Fallén, 1820
Tetanocerini Newman, 1834
ANTICHETA Haliday, 1839
Anticheta analis (Meigen, 1830)
Anticheta atriseta (Loew, 1849)
Anticheta brevipennis (Zetterstedt, 1846)
Anticheta nigra Karl, 1921
= nigroaenea Frey, 1935
COREMACERA Rondani, 1856
Coremacera marginata (Fabricius, 1775)
= tristis (Harris, 1780) preocc.
DICHETOPHORA Rondani, 1868
Dichetophora nlandica Verbeke, 1964
DICTYA Meigen, 1803
Dictya umbrarum (Linnaeus, 1758)
ECTINOCERA Zetterstedt, 1838
Ectinocera borealis Zetterstedt, 1838
ELGIVA Meigen, 1838
Elgiva cucularia (Linnaeus, 1767)
Elgiva divisa (Loew, 1845)
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)271
Elgiva solicita (Harris, 1780)
= sundewalli Kloet & Hincks, 1945
EUTHYCERA Latreille, 1829
Euthycera chaerophylli (Fabricius, 1798)
Euthycera fumigata (Scopoli, 1763)
HYDROMYA Robineau-Desvoidy, 1830
Hydromya dorsalis (Fabricius, 1775)
ILIONE Haliday in Curtis, 1837
= Tumidicerus Knutson & Berg, 1967
sg. Ilione Haliday in Curtis, 1837
Ilione lineata (Fallén, 1820)
sg. Knutsonia Verbeke, 1964
Ilione albiseta (Scopoli, 1763)
LIMNIA Robineau-Desvoidy, 1830
Limnia paludicola Elberg, 1965
Limnia unguicornis (Scopoli, 1763)
PHERBINA Robineau-Desvoidy, 1830
Pherbina coryleti (Scopoli, 1763)
PSACADINA Enderlein, 1939
Psacadina zernyi (Mayer, 1953)
= punctata misid.
RENOCERA Hendel, 1900
Renocera pallida (Fallén, 1820)
Renocera striata (Meigen, 1830)
Renocera stroblii Hendel, 1900
= fuscinervis auct. nec (Zetterstedt, 1838)
SEPEDON Latreille, 1804
sg. Sepedon Latreille, 1804
Sepedon sphegea (Fabricius, 1775)
Sepedon spinipes (Scopoli, 1763)
TETANOCERA Duméril, 1800
sg. Chaetotetanocera Mayer, 1953
Tetanocera robusta Loew, 1847
sg. Tetanocera Duméril, 1800
? Tetanocera amurensis Hendel, 1909
Tetanocera arrogans Meigen, 1830
Tetanocera elata (Fabricius, 1781)
Tetanocera ferruginea Fallén, 1820
= brunnipennis Frey, 1924
Tetanocera freyi Stackelberg, 1963
Tetanocera fuscinervis (Zetterstedt, 1838)
= unicolor Loew, 1847
Tetanocera hyalipennis von Roser, 1840
Jere Kahanpää & Kaj Winqvist / ZooKeys 441: 259–275 (2014)
272
Tetanocera kerteszi Hendel, 1901
= griseicollis Frey, 1924
= ornatifrons Frey, 1924
Tetanocera lapponica Frey, 1924
Tetanocera latibula Frey, 1924
Tetanocera montana Day, 1881
= borealis Frey, 1924
Tetanocera phyllophora Melander, 1920
= nigricosta misid.
Tetanocera silvatica Meigen, 1830
TRYPETOPTERA Hendel, 1900
Trypetoptera punctulata (Scopoli, 1763)
SEPSIDAE Walker, 1833
ORYGMATINAE Frey, 1921
ORTALISCHEMA Frey, 1925
Ortalischema albitarse (Zetterstedt, 1847)
SEPSINAE Walker, 1833
THEMIRA Robineau-Desvoidy, 1830
emira annulipes (Meigen, 1826)
emira arctica (Becker, 1915)
emira biloba Andersson, 1975
emira germanica Duda, 1926
emira gracilis (Zetterstedt, 1847)
emira leachi (Meigen, 1826)
emira lucida (Staeger, 1844)
emira malformans Melander & Spuler, 1917
emira minor (Haliday, 1833)
emira nigricornis (Meigen, 1826)
emira paludosa Elberg, 1963
emira pusilla (Zetterstedt, 1847)
emira putris (Linnaeus, 1758)
emira superba (Haliday, 1833)
SALTELLA Robineau-Desvoidy, 1830
Saltella sphondylii (Schrank, 1803)
NEMOPODA Robineau-Desvoidy, 1830
Nemopoda nitidula (Fallén, 1820)
= cylindrica (Fabricius, 1794) preocc.
Nemopoda pectinulata Loew, 1873
Nemopoda speiseri (Duda, 1926)
MEROPLIUS Rondani, 1874
Meroplius fukuharai (Iwasa, 1984)
Meroplius minutus (Wiedemann, 1830)
Checklist of the Diptera superfamilies Tephritoidea and Sciomyzoidea of Finland (Insecta)273
= stercorarius (Robineau-Desvoidy, 1830)
SEPSIS Fallén, 1810
Sepsis biexuosa Strobl, 1893
Sepsis cynipsea (Linnaeus, 1758)
Sepsis duplicata Haliday, 1838
= pilipes van der Wulp, 1871
Sepsis avimana Meigen, 1826
= borealis Frey, 1825
Sepsis fulgens Meigen, 1826
= communis Frey, 1925
Sepsis luteipes Melander & Spuler, 1917
= lamellifera Frey, 1925
Sepsis nigripes Meigen, 1826
Sepsis orthocnemis Frey, 1908
Sepsis punctum (Fabricius, 1794)
= luteipes misid.
Sepsis thoracica (Robineau-Desvoidy, 1830)
Sepsis violacea Meigen, 1826
Excluded species
Allopiophila dudai (Frey, 1930) not found within present borders
Bactrocera sp. imported with fruit
Campiglossa irrorata (Fallén, 1814) not found within present borders
Ceratitis capitata (Wiedemann, 1824) imported with fruit
Earomyia crystallophila (Becker, 1895) not found within present borders
= nigroviolacea (Frey in Lundström & Frey, 1913)
Lonchaea hirticeps Zetterstedt, 1838 misidentied
Orygma luctuosum Meigen, 1830 not found within present borders
Pherbellia dorsata (Zetterstedt, 1846) misidentied
Tephritis nigricauda (Loew, 1856) misidentied
Notes
Allopiophila dudai (Frey, 1930). is species was described by Frey (1930) from
the Kola peninsula (Murmansk Oblast, Russia). Frey et al. (1941) and Hackman
(1980) listed it from Finland, but to our knowledge it has never been reliably re-
corded from this country.
Allopiophila sp. A. An apparently undescribed arctic species near A. vulgaris.
Campiglossa sp. A is a distinct, possibly undescribed species of Campiglossa. It is common
on Saussurea alpina in Northern Finland (Juhani Itämies, priv. comm.).
Jere Kahanpää & Kaj Winqvist / ZooKeys 441: 259–275 (2014)
274
Lonchaea bruggeri Morge, 1967, L. contigua Collin, 1953 and L. dicilis Hackman,
1956. No males have been conrmed from the country. See Winqvist and Kahanpää
(2007) for more details.
Lonchaea bukowskii Czerny, 1934. No males have been conrmed from the country.
Tephritis sp. cf. rydeni Hering, 1956. Finnish material formerly assigned to T. dioscurea
(Loew, 1856) belongs to a currently unrecognised species that may be T. rydeni Hering
(Söderman et al. 2007, Winqvist and Kahanpää 2007).
Tetanocera amurensis Hendel, 1909. Recorded from Finland in Fauna Europaea
(Rozkošný and Knutson 2011) and known from Estonia and Northwestern Euro-
pean Russia (Elberg 1988). We have been unable to nd any Finnish material or
primary published records.
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Jere Kahanpää & Kaj Winqvist / ZooKeys 441: 259–275 (2014)
276
Checklist of the y families Chamaemyiidae and Lauxaniidae of Finland (Insecta, Diptera)277
Checklist of the fly families Chamaemyiidae and
Lauxaniidae of Finland (Insecta, Diptera)
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 13 March 2014|Accepted 14 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the y families Chamaemyiidae and Lauxaniidae of Finland (Insecta, Diptera). In:
Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 277–283. doi: 10.3897/zookeys.441.7506
Abstract
A revised checklist of the Chamaemyiidae and Lauxaniidae (Diptera) recorded from Finland is presented.
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
ree families are currently recognized in Lauxanoidea, two of which are present in
Finland. e exception is the beetle ies (Celyphidae), which are restricted to the Ori-
ental and Afrotropical Regions.
e silvery white ies of the family Chamaemyiidae are poorly studied in Finland.
Reliable identication of most species is, at least for now, only possible by the exami-
nation of male genitalia. Unfortunately many species have not been adequately (re)
described which has resulted in much confusion about the proper use of species names.
It is likely that several additional chamaemyiid species will be found in Finland. Of the
ZooKeys 441: 277–283 (2014)
doi: 10.3897/zookeys.441.7506
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution International License
(CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 277–283 (2014)
278
two subfamilies of chamaemyiids, Cremifaniinae is absent from Finland. Chamaemyi-
inae is split into two tribes, Chamaemyiini and Leucopini, although the monophyly of
the former has not yet been demonstrated (Gaimari 2010).
e lauxaniids are better known both in Finland and worldwide. Additional species
are still likely to be found in the country, especially among Homoneura and Sapromyza.
Of the three subfamilies, the Eurychoromyiinae, a strictly Neotropical group (Gaimari
and Silva 2010a), is absent from the Palaearctic. e other two subfamilies, Homoneu-
rinae and Lauxaniinae, are both present in Finland e subfamily Lauxaniinae is prob-
ably paraphyletic as currently dened (Gaimari and Silva 2010b).
e Finnish species of these families were last listed by Hackman (1980). Kahan-
pää (2013) reviewed the changes in the Finnish lauxaniid fauna since 1980.
Number of species (Chamaemyiidae, Lauxaniidae):
World: 336 (S. Gaimari, pers. comm. March 18, 2014), 1895 species (Pape et al. 2011)
Europe: 107, 157 species (Fauna Europaea)
Finland: 27–28, 45 species
Faunistic knowledge level in Finland: poor, average
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Lauxanoidea Macquart, 1835
CHAMAEMYIIDAE Hendel, 1910
CHAMAEMYIINAE Hendel, 1910
tribe Chamaemyiini Hendel, 1910
ACROMETOPIA Schiner, 1862
Acrometopia wahlbergi (Zetterstedt, 1846)
CHAMAEMYIA Meigen, 1803
Chamaemyia aestiva Tanasijtshuk, 1970
Chamaemyia aridella (Fallén, 1823)
Chamaemyia elegans (Panzer, 1809)
Chamaemyia emiliae Tanasijtshuk, 1970
Chamaemyia avipalpis (Haliday, 1838)
Chamaemyia geniculata (Zetterstedt, 1838)
? Chamaemyia herbarum (Robineau-Desvoidy, 1830) sensu Coe, 1943
Chamaemyia juncorum (Fallén, 1823)
Chamaemyia paludosa Collin, 1966
Checklist of the y families Chamaemyiidae and Lauxaniidae of Finland (Insecta, Diptera)279
Chamaemyia polystigma (Meigen, 1830)
Chamaemyia sylvatica Collin, 1966
PAROCHTHIPHILA Czerny, 1904
sg. Euestelia Enderlein, 1927
Parochthiphila coronata (Loew, 1858)
sg. Parochthiphila Czerny, 1904
Parochthiphila spectabilis (Loew, 1858)
tribe Lecopini Hendel, 1928
ANCHIOLEUCOPIS Tanasijtshuk 1997
Anchioleucopis geniculata (Zetterstedt, 1855)
LEUCOPIS Meigen, 1830
Leucopis annulipes Zetterstedt, 1848
Leucopis argentata Heeger, 1848
= conciliata McAlpine & Tanasijtshuk, 1972
= argenticollis auct. nec Zetterstedt, 1848
Leucopis atritarsis Tanasijtshuk, 1958
Leucopis glyphinivora Tanasijtshuk, 1958
Leucopis griseola (Fallén, 1823)
Leucopis sorbi Tanasijtshuk, 1986
Leucopis sp. cf. szepligetii Aczel, 1937
NEOLEUCOPIS Malloch, 1921
Neoleucopis atratula (Ratzeburg, 1844)
Neoleucopis freyi (McAlpine, 1971)
Neoleucopis obscura (Haliday, 1833)
Neoleucopis orbiseta (McAlpine, 1971)
LEUCOPOMYIA Malloch, 1921
Leucopomyia silesiaca (Egger, 1862)
= alticeps misid.
LIPOLEUCOPIS de Meijere, 1928
Lipoleucopis praecox de Meijere, 1928
LAUXANIIDAE Macquart, 1835
HOMONEURINAE Stuckenberg, 1971
HOMONEURA van der Wulp, 1891
Homoneura biumbrata (Loew, 1873)
= tesquae misid.
Homoneura lamellata (Becker, 1895)
Homoneura mediospinosa Merz, 2003
= interstincta auct. nec (Fallén, 1820)
Homoneura tenera (Loew, 1846)
LAUXANIINAE Macquart, 1835
AULOGASTROMYIA Hendel, 1925
Aulogastromyia anisodactyla (Loew, 1845)
CALLIOPUM Strand, 1928
Jere Kahanpää / ZooKeys 441: 277–283 (2014)
280
Calliopum aeneum (Fallén, 1820)
Calliopum elisae (Meigen, 1826)
= nitens auct. nec (Loew, 1858)
CNEMACANTHA Macquart, 1835
Cnemacantha muscaria (Fallén, 1823)
LAUXANIA Latreille, 1804
sg. Czernushka Shatalkin, 2000
Lauxania albomaculata Strobl, 1909
sg. Lauxania Latreille, 1804
Lauxania cylindricornis (Fabricius, 1794)
MEIOSIMYZA Hendel, 1925
= Lycia Robineau-Desvoidy, 1830 preocc.
= Lyciella Collin, 1948
Meiosimyza anis (Zetterstedt, 1847)
Meiosimyza decempunctata (Fallén, 1820)
Meiosimyza decipiens (Loew, 1847)
Meiosimyza illota (Loew, 1847)
Meiosimyza laeta (Zetterstedt, 1838)
Meiosimyza platycephala (Loew, 1847)
Meiosimyza rorida (Fallén, 1820)
Meiosimyza subfasciata (Zetterstedt, 1838)
MINETTIA Robineau-Desvoidy, 1830
sg. Frendelia Collin, 1948
Minettia longipennis (Fabricius, 1794)
sg. Minettia Robineau-Desvoidy, 1830
Minettia desmometopa (de Meijere, 1907)
Minettia lupulina (Fabricius, 1787)
sg. Plesiominettia Shatalkin, 2000
Minettia helvola (Becker, 1895)
Minettia loewi (Schiner, 1864)
Minettia lia (Becker, 1895)
sg. unplaced in ?Minettia (see Notes)
Minettia styriaca (Strobl, 1892)
PACHYCERINA Macquart, 1835
Pachycerina pulchra (Loew, 1850)
Pachycerina seticornis (Fallén, 1820)
PEPLOMYZA Haliday, 1837
Peplomyza discoidea (Meigen, 1830)
POECILOLYCIA Shewell, 1986
Poecilolycia vittata (Walker, 1849)
= quadrivittata (Loew, 1861)
PSEUDOLYCIELLA Shatalkin, 2000
Pseudolyciella pallidiventris (Fallén, 1820)
Checklist of the y families Chamaemyiidae and Lauxaniidae of Finland (Insecta, Diptera)281
Pseudolyciella stylata (Papp, 1978)
SAPROMYZA Fallén, 1810
sg. Nannomyza Frey, 1941
Sapromyza basalis Zetterstedt, 1847
sg. Sapromyza Fallén, 1810
Sapromyza albiceps (Fallén, 1820)
Sapromyza amabilis Frey, 1930
Sapromyza apicalis Loew, 1847
= obsoleta misid.
Sapromyza opaca Becker, 1895
= imitatrix Czerny, 1932
= leningradensis misid.
Sapromyza schnabli Papp, 1987
Sapromyza setiventris Zetterstedt, 1847
Sapromyza sexpunctata Meigen, 1826
= atechna Becker, 1895
= pellucida Becker, 1895
Sapromyza simplicior Hendel, 1908
Sapromyza zetterstedti Hendel, 1908
sg. Schumannimyia Papp, 1978
Sapromyza hyalinata (Meigen, 1826)
SAPROMYZOSOMA Lioy, 1864
Sapromyzosoma quadripunctata (Linnaeus, 1767)
TRICHOLAUXANIA Hendel, 1925
Tricholauxania praeusta (Fallén, 1820)
TRIGONOMETOPUS Macquart, 1835
Trigonometopus frontalis (Meigen, 1830)
Excluded species
Homoneura dilecta (Rondani, 1868) misidentication
Leucopis impunctata von Roser, 1840 nomen dubium
Leucopis puncticornis Meigen, 1830 nomen dubium
Sapromyza thoracica Becker, 1895 nomen dubium
Neoparoecus signatipes (Loew, 1856) not found within present borders
Sapromyza leningradensis Czerny, 1932 nomen dubium ?
Notes
Several chamaemyiid species mentioned in this checklist have not been previously re-
corded from Finland. A paper detailing the new records is in preparation.
Jere Kahanpää / ZooKeys 441: 277–283 (2014)
282
Chamaemyia herbarum (Robineau-Desvoidy, 1830). A poorly known species.
e name has been widely used for various Chamaemyia species in the past. Coe (1943)
and Collin (1966) applied this name (without seeing types) to a species which may
be C. subjuncorum Tanasijtshuk, 1980 (sensu Beschovski, 1995). Czerny (1936) and
Tanasijtshuk (1986) listed this species as a junior synonym of Chamaemyia juncorum
(Fallén). Most Finnish specimens under this name are either unidentiable females or
belong to other species, most frequently C. aestiva Tanasijtshuk, 1970, but some speci-
mens probably belonging to C. herbarum R.-D. sensu Coe have been found in Finland.
Minettia(?) styriaca (Strobl, 1892). is species, originally described in Sapro-
myza, has traditionally been placed in Minettia. Unfortunately Strobl’s description,
which was based on a single female, is rather short and unillustrated. e holotype is
lost according to Chvála (2008). e Finnish and Russian material in the Finnish Mu-
seum of Natural History (MZH) identied as Minettia styriaca (see Kahanpää 2013
for data and a photograph) matches Strobl’s description, but these specimens lack the
postsutural supra-alar setae found in Minettia (and many other genera).
Acknowledgements
I would like to thank Dr. Stephen Gaimari for his valuable comments and suggestions
on improving this paper.
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Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
Tanasijtshuk VN (1986) Chamaemyiidae. In: Fauna of the USSR n.s., 134: Diptera 14(7).
Nauka, Leningrad, 335 pp. + 16 pls. [in Russian]
Jere Kahanpää / ZooKeys 441: 277–283 (2014)
284
Checklist of the smaller families of Opomyzoidea, Anthomyzidae, Asteiidae... 285
Checklist of the smaller families of Opomyzoidea,
Anthomyzidae, Asteiidae, Aulacigastridae, Clusiidae,
Odiniidae, Opomyzidae and Periscelididae
(Diptera) of Finland
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 5 February 2014|Accepted 6 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the smaller families of Opomyzoidea, Anthomyzidae, Asteiidae, Aulacigastridae,
Clusiidae, Odiniidae, Opomyzidae and Periscelididae (Diptera) of Finland. In: Kahanpää J, Salmela J (Eds) Checklist of
the Diptera of Finland ZooKeys 441: 285–290. doi: 10.3897/zookeys.441.7141
Abstract
A species checklist is presented for Finland covering seven smaller families of Opomyzoidea: Anthomyzidae,
Asteiidae, Aulacigastridae, Clusiidae, Odiniidae, Opomyzidae and Periscelididae (Diptera).
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
e superfamily Opomyzoidea is likely to be para- or polyphyletic (see Wiegmann
et al. 2011). It has long been used as a dumping grounds for acalyptrate taxa that do
not t well elsewhere. As such, the superfamily is poorly dened and hardly any two
authors agree on its denition. Opomyzoidea includes one large family (Agromyzidae,
the leaf-mining ies, with over 3000 described species) and more than a dozen small
families. e exact number of recognized families varies. e Agromyzidae are treated
ZooKeys 441: 285–290 (2014)
doi: 10.3897/zookeys.441.7141
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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Jere Kahanpää / ZooKeys 441: 285–290 (2014)
286
elsewhere in this volume of ZooKeys. e remaining opomyzoid families are presented
here in alphabetical order.
Hackman (1980) last listed the Finnish species of these families. World catalogues
have recently been published for Periscelididae (Mathis and Rung 2011) and Odinii-
dae (Gaimari and Mathis 2011). For Anthomyzidae the Revision of West Palearctic
Anthomyzidae by Roháček (2006, 2009) is followed. ree of the four Palearctic Aul-
acigaster species have been found in Europe (Kassebeer 2001, Rung and Mathis 2011).
Papp and Withers (2011) reviewed the Palearctic Periscelidinae and proposed some
nomenclatorial changes. Hackman’s (1959) review of the Finnish opomyzid fauna is
still valuable but several additional species have since been recognized.
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Opomyzoidea Fallén, 1820
ANTHOMYZIDAE Czerny, 1903
ANTHOMYZA Fallén, 1810
Anthomyza collini Andersson, 1976
Anthomyza dissors Collin, 1944
Anthomyza elbergi Andersson, 1976
= sordidella auct. nec (Zetterstedt, 1848)
Anthomyza gracilis Fallén, 1823
Anthomyza macra Czerny, 1928
Anthomyza pallida (Zetterstedt, 1838)
= unguicella (Zetterstedt, 1838)
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Anthomyzidae 93 28 15 average
Asteiidae 136 18 6 average
Aulacigastridae 18 4 2 good
Clusiidae 360 14 12 good
Odiniidae 64 14 5 average
Opomyzidae 59 33 16 average
Periscelididae 83 6 4 average
Checklist of the smaller families of Opomyzoidea, Anthomyzidae, Asteiidae... 287
Anthomyza paraneglecta Elberg, 1968
Anthomyza pleuralis Czerny, 1928
ARGANTHOMYZA Roháček, 2009
Arganthomyza socculata (Zetterstedt, 1847)
= ungulata (Loew, 1873)
CAREXOMYZA Roháček, 2009
Carexomyza caricis (Roháček, 1999)
FUNGOMYZA Roháček, 1999
Fungomyza albimana (Meigen, 1830)
PARANTHOMYZA Czerny, 1902
Paranthomyza nitida (Meigen, 1838)
STIPHROSOMA Czerny, 1928
= Ptenotaenia Enderlein, 1936
Stiphrosoma laetum (Meigen, 1830)
= cingulatum misid.
Stiphrosoma sabulosum (Haliday, 1837)
TYPHAMYZA Roháček, 1992
Typhamyza bifasciata (Wood, 1911)
ASTEIIDAE Rondani, 1856
ASTEIINAE Rondani, 1856
ASTEIA Meigen, 1830
sg. Asteia Meigen, 1830
Asteia amoena Meigen, 1830
Asteia concinna Meigen, 1830
Asteia elegantula Zetterstedt, 1847
SIGALOESSINAE Sabrosky, 1956
LEIOMYZA Macquart, 1835
Leiomyza dudai Sabrosky, 1956
Leiomyza laevigata (Meigen, 1830)
Leiomyza scatophagina (Fallén, 1823)
AULACIGASTRIDAE Duda, 1924
AULACIGASTER Macquart, 1835
= Aulacogaster emend.
Aulacigaster leucopeza (Meigen, 1830)
Aulacigaster pappi Kassebeer, 2001
CLUSIIDAE Handlirsch, 1884
CLUSIINAE Handlirsch, 1884
CLUSIA Haliday, 1838
= Paraclusia Czerny, 1903
Clusia ava (Meigen, 1830)
Jere Kahanpää / ZooKeys 441: 285–290 (2014)
288
Clusia tigrina (Fallén, 1820)
CLUSIODINAE
CLUSIODES Coquillett, 1904
= Heteroneura Fallén, 1823 preocc.
Clusiodes albimanus (Meigen, 1830)
Clusiodes apicalis (Zetterstedt, 1848)
Clusiodes caledonicus (Collin, 1912)
= gentilis misid.
Clusiodes freyi Tuomikoski, 1933
Clusiodes geomyzinus (Fallén, 1823)
Clusiodes microcercus Stackelberg, 1955
Clusiodes pictipes (Zetterstedt, 1855)
Clusiodes rucollis (Meigen, 1830)
Clusiodes verticalis (Collin, 1912)
HENDELIA Czerny, 1903
Hendelia beckeri Czerny, 1903
ODINIIDAE Hendel, 1920
ODINIINAE Hendel, 1920
NEOALTICOMERUS Hendel, 1903
Neoalticomerus formosus (Loew, 1844)
ODINIA Robineau-Desvoidy, 1830
Odinia boletina (Zetterstedt, 1848)
Odinia czernyi Collin, 1952
= meijerei misid.
Odinia ornata (Zetterstedt, 1838)
Odinia xanthocera Collin, 1952
OPOMYZIDAE Fallén, 1820
ANOMALOCHAETA Frey, 1921
Anomalochaeta guttipennis (Zetterstedt, 1838)
GEOMYZA Fallén, 1810
Geomyza angustipennis Zetterstedt, 1847
Geomyza apicalis (Meigen, 1830)
Geomyza consobrina Zetterstedt, 1847
Geomyza hackmani Nartshuk, 1984
= combinata auct. nec (Linnaeus, 1767)
= balachowskyi misid.
Geomyza majuscula (Loew, 1864)
= breviforceps Hackman, 1959
Geomyza martineki Drake, 1992
Geomyza paganettii (Strobl, 1909)
Geomyza pilosula Czerny, 1928
Checklist of the smaller families of Opomyzoidea, Anthomyzidae, Asteiidae... 289
Geomyza tripunctata Fallén, 1823
OPOMYZA Fallén, 1820
Opomyza orum (Fabricius, 1794)
Opomyza germinationis (Linnaeus, 1758)
Opomyza lineatopunctata von Roser, 1840
Opomyza nigriventris Loew, 1865
Opomyza punctata Haliday, 1833
Opomyza punctella Fallén, 1820
PERISCELIDIDAE Oldenberg, 1914
PERISCELIDINAE Oldenberg, 1914
MYODRIS Lioy, 1864
= Microperiscelis Oldenberg, 1914
Myodris annulata (Fallén, 1813)
PERISCELIS Loew, 1858
Periscelis annulipes Loew, 1858
Periscelis nigra (Zetterstedt, 1860)
Periscelis winnertzii Egger, 1862
Excluded species
Clusiodes gentilis (Collin, 1912) misidentied
Geomyza balachowskyi Mesnil, 1934 misidentied
Odinia meijerei Collin, 1952 misidentied
Notes
Aulacigaster pappi Kassebeer, 2001. Most of the Finnish Aulacigaster material belong
to this species, including all specimens currently stored in the Finnish Museum on
Natural History (MZH). Storå (1956, 1957) and Hackman (1980) misidentied
A. pappi as A. leucopeza. A few Finnish specimens of the real A. leucopeza were
found in private collections.
Clusiodes gentilis (Collin, 1912). Lonsdale and Marshall (2007) record this species
from Finland on a map. e data is apparently taken from Frey (1928). As Tuo-
mikoski (1936) has pointed out, Frey misidentied C. caledonicus as C. gentilis.
References
Frey R (1928) Förteckning över Finlands Clusiider (Dipt., Haplost.). Notulae entomologicae
8: 108–109.
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Gaimari SD, Mathis WN (2011) World catalog and conspectus on the family Odiniidae (Diptera:
Schizophora). In: Brake I, ompson FC (Eds) Contributions to the Systema Dipterorum
(Insecta: Diptera). Myia 12: 291–339. http://hdl.handle.net/10088/18930
Hackman W (1959) e Opomyzidae (Dipt.) of Eastern Fennoscandia. Notulae entomologi-
cae 38[1958](4): 114–126.
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Kassebeer CF (2001) Die einheimischen Arten der Gattung Aulacigaster Macquart, 1835
(Diptera, Aulacigastridae). Dipteron 4(1): 23–32.
Lonsdale O, Marshall SA (2007) Redenition of the genera Clusiodes and Hendelia (Diptera:
Clusiidae: Clusiodinae), with a review of Clusiodes. Studia dipterologica 14(1): 117–159.
Mathis WN, Rung A (2011) World catalog and conspectus on the Family Periscelididae (Diptera:
Schizophora). In: Brake I, ompson FC (Eds) Contributions to the Systema Dipterorum
(Insecta: Diptera). Myia 12: 341–377. http://hdl.handle.net/10088/18931
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed)
Animal biodiversity: An outline of higher-level classication and survey of taxonomic richness.
Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
Papp L, Withers P (2011) A revision of the Palaearctic Periscelidinae with notes on some
New World species (Diptera: Periscelididae). Annales historico-naturales Musei Nationalis
Hungarici 103: 345–371.
Roháček J (2006) A monograph of Palaearctic Anthomyzidae (Diptera) Part 1. Časopis Slezského
zemského muzea Opava (A) 55, Supplement 1. Slezske zemske muzeum, Opava, 328 pp.
Roháček J (2009) A monograph of Palaearctic Anthomyzidae (Diptera) Part 2. Časopis Slezského
zemského muzea Opava (A) 58, Supplement 1. Slezske zemske muzeum, Opava, 180 pp.
Rung A, Mathis WN (2011) A revision of the genus Aulacigaster Macquart (Diptera: Aul-
acigastridae). Smithsonian Contributions to Zoology, vol. 633. Smithsonian Institution
Scholarly Press, Washington D.C., 132 pp. http://hdl.handle.net/10088/15584
Storå R (1956) Fynd av sällsynta dipterer i Österbotten (Om). Notulae entomologicae 36: 17–22.
Storå R (1957) Einige bemerkenswerte Dipteren-Funde an einem saftenden Birkenstamme.
Notulae entomologicae 37(1): 23–26.
Tuomikoski R (1933) Notizen über die Clusiiden Finnlands. Notulae entomologicae 13(1): 15–19.
Tuomikoski R (1936) Bemerkungen über die Clusiiden (Dipt.) Finnlands. Annales Entomologici
Fennici 2(4): 182–186.
Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA,
Cassel BK, Bayless KM, Heimberg AM, Wheeler BM, Peterson KJ, Pape T, Sinclair BJ, Ske-
vington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Ott U, Kampmeier GE, omp-
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Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 291
Checklist of the leaf-mining flies
(Diptera, Agromyzidae) of Finland
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 25 March 2014|Accepted 28 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland. In: Kahanpää J,
Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 291–303. doi: 10.3897/zookeys.441.7586
Abstract
A checklist of the Agromyzidae (Diptera) recorded from Finland is presented. 279 (or 280) species are
currently known from the country. Phytomyza linguae Lundqvist, 1947 is recorded as new to Finland.
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
e Agromyzidae are called the leaf-miner or leaf-mining ies and not without reason,
although a substantial fraction of the species feed as larvae on other parts of living
plants. While Agromyzidae is traditionally placed in the superfamily Opomyzoidea,
its exact relationships with other acalyptrate Diptera are poorly understood (see for
example Winkler et al. 2010).
Two subfamilies are recognised within the leaf-mining ies: Agromyzinae and
Phytomyzinae. Both are now recognised as natural groups (Dempewolf 2005, Scheer
et al. 2007). Unfortunately the genera are not as well dened: at least Ophiomyia, Phy-
toliriomyza and Aulagromyza are paraphyletic in DNA sequence analyses (see Scheer
et al. 2007). e same studies and Winkler et al. (2009) suggest that Chromatomyia
ZooKeys 441: 291–303 (2014)
doi: 10.3897/zookeys.441.7586
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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Jere Kahanpää / ZooKeys 441: 291–303 (2014)
292
consists of at least two separate groups of species within Phytomyza, and Napomyza
(which is given a status of a subgenus of Phytomyza by Winkler et al. (2009)) also needs
re-evaluation. For the purposes of this checklist, a conservative approach is adopted
with Chromatomyia and Napomyza as separate genera.
Richard Frey was the rst Finnish dipterist to study the agromyzid fauna of the coun-
try in detail (Frey 1937, 1946). L. Tiensuu and E. uneberg continued Frey’s work but
most of their work remained unpublished until Nowakowski (1967) and Spencer (1976)
reviewed the major Finnish agromyzid collections. Spencer’s book remains the primary
identication tool for North European Agromyzidae. Interest in Finnish agromyzids has
remained sporadic. e Finnish species were last listed by Hackman (1980).
Number of species:
World: 3004 spp. (M. Černý & M. von Tschirnhaus, pers. comm., March 2014)
Europe: 914 species (von Tschirnhaus, pers. comm., March 2014)
Finland: 279–280 species
Faunistic knowledge level in Finland: average
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Opomyzoidea Newman, 1834
AGROMYZIDAE Fallén, 1823
AGROMYZINAE Fallén, 1823
AGROMYZA Fallén, 1810
Agromyza abiens Zetterstedt, 1848
Agromyza albipennis Meigen, 1830
Agromyza albitarsis Meigen, 1830
Agromyza alnibetulae Hendel, 1931
Agromyza alnivora Spencer, 1969
Agromyza alunulata (Hendel, 1931)
Agromyza ambigua Fallén, 1823
= niveipennis Zetterstedt, 1848
Agromyza anthracina Meigen, 1830
= freyi Hendel, 1931
Agromyza bicaudata (Hendel, 1931)
Agromyza cinerascens Macquart, 1835
Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 293
Agromyza demeijerei Hendel, 1920
Agromyza erythrocephala Hendel, 1920
Agromyza aviceps Fallén, 1823
Agromyza graminicola Hendel, 1931
Agromyza idaeiana Hardy, 1853
= potentillae (Kaltenbach, 1864)
= spiraeae Kaltenbach, 1867
= stackelbergi (Frey, 1946)
Agromyza igniceps Hendel, 1920
= lathyri misid.
Agromyza lapponica Hendel, 1931
Agromyza lucida Hendel, 1920
Agromyza luteitarsis (Rondani, 1875)
= intermittens misid.
Agromyza marionae Griths, 1963
= alandensis Spencer, 1976
Agromyza mobilis Meigen, 1830
Agromyza nana Meigen, 1830
Agromyza nigrella (Rondani, 1875)
Agromyza nigrescens Hendel, 1920
Agromyza nigripes Meigen, 1830
Agromyza nigrociliata Hendel, 1931
Agromyza orobi Hendel, 1920
Agromyza phragmitidis Hendel, 1922
Agromyza pseudoreptans Nowakowski, 1967
= urticae Nowakowski, 1964 preocc.
Agromyza quadriseta Zlobin, 2001
Agromyza reptans Fallén, 1823
Agromyza rupes (Meigen, 1830)
Agromyza salicina Hendel, 1922
Agromyza sulfuriceps Strobl, 1898
Agromyza vicifoliae Hering, 1932
HEXOMYZA Enderlein, 1936
Hexomyza schineri (Giraud, 1861)
Hexomyza simplicoides (Hendel, 1920)
MELANAGROMYZA Hendel, 1920
Melanagromyza aenea (Meigen, 1830)
Melanagromyza aeneoventris (Fallén, 1823)
Melanagromyza angeliciphaga Spencer, 1969
Melanagromyza chaerophylli Spencer, 1969
Melanagromyza lappae (Loew, 1850)
Melanagromyza nigrissima Spencer, 1976
Melanagromyza oligophaga Spender, 1990
Jere Kahanpää / ZooKeys 441: 291–303 (2014)
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Melanagromyza pubescens Hendel, 1923
Melanagromyza submetallescens Spencer, 1966
OPHIOMYIA Braschnikov, 1897
Ophiomyia beckeri (Hendel, 1923)
= goniaea (Hendel, 1931)
Ophiomyia cunctata (Hendel, 1920)
Ophiomyia curvipalpis (Zetterstedt, 1848)
Ophiomyia fennoniensis Spencer, 1976
?= eucodonus Hering, 1960
= melandryi misid.
Ophiomyia heringi Starý, 1930
= penicillata misid.
Ophiomyia labiatarum Hering, 1937
= persimilis misid.
Ophiomyia longilingua (Hendel, 1920)
= rostrata misid.
Ophiomyia maura (Meigen, 1838)
Ophiomyia nasuta (Melander, 1913)
Ophiomyia orbiculata (Hendel, 1931)
= hexachaeta (Hendel, 1931)
= nostradamus (Hering, 1933)
= paracelsus (Hering, 1933)
Ophiomyia pinguis (Fallén, 1820)
Ophiomyia pulicaria (Meigen, 1830)
Ophiomyia ranunculicaulis Hering, 1949
Ophiomyia vitiosa Spencer, 1964
?= alliariae Hering, 1954
PHYTOMYZINAE Fallén, 1823
AMAUROMYZA Hendel, 1931
sg. Amauromyza Hendel, 1931
Amauromyza morionella (Zetterstedt, 1848)
sg. Cephalomyza Hendel, 1931
= Trilobomyza Hendel, 1931
Amauromyza chenopodivora Spencer, 1971
= abnormalis misid.
Amauromyza avifrons (Meigen, 1830)
Amauromyza gyrans (Fallén, 1823)
Amauromyza karli (Hendel, 1927)
= Dizygomyza facialis Frey, 1941 nom. nudum
Amauromyza labiatarum (Hendel, 1920)
Amauromyza luteiceps (Hendel, 1920)
Amauromyza monfalconensis (Strobl, 1909
AULAGROMYZA Enderlein, 1936
Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 295
= Paraphytomyza Enderlein, 1936
Aulagromyza buhri (de Meijere, 1938)
Aulagromyza fulvicornis (Hendel, 1935)
Aulagromyza hendeliana (Hering, 1926)
Aulagromyza heringii (Hengel, 1920)
Aulagromyza lucens (de Meijere, 1924)
Aulagromyza luteoscutellata (de Meijere, 1924)
= xylostei misid.
Aulagromyza similis (Brischke, 1880)
Aulagromyza tremulae (Hering 1957)
Aulagromyza tridentata (Loew, 1858)
Aulagromyza trivittata (Loew, 1873)
CALYCOMYZA Hendel, 1931
Calycomyza artemisiae (Kaltenbach, 1856)
Calycomyza subapproximata (Sasakawa, 1955)
CERODONTHA Rondani, 1861
sg. Butomomyza Nowakowski, 1967
Cerodontha caricivora (Groschke, 1954)
Cerodontha eucaricis Nowakowski, 1967
Cerodontha rohdendor Nowakowski, 1967
Cerodontha scirpi (Karl, 1926)
= scutellaris misid.
= semiposticata misid.
Cerodontha staryi (Starý, 1930)
sg. Cerodontha Rondani, 1861
Cerodontha anis (Fallén, 1823)
Cerodontha denticornis (Panzer, 1806)
Cerodontha fulvipes (Meigen, 1830)
= femoralis (Meigen, 1838)
Cerodontha hennigi Nowakowski, 1967
= lateralis (Zetterstedt, 1848) preocc.
Cerodontha stackelbergi Nowakowski, 1972
sg. Dizygomyza Hendel, 1920
Cerodontha bimaculata (Meigen, 1830)
Cerodontha fasciata (Strobl, 1880)
Cerodontha ireos (Robineau-Desvoidy, 1851)
Cerodontha luctuosa (Meigen, 1830)
Cerodontha morosa (Meigen, 1830)
sg. Icteromyza Hendel, 1931
Cerodontha bohemani (Rydén, 1951)
Cerodontha capitata (Zetterstedt, 1848)
Cerodontha churchillensis Spencer, 1969
Cerodontha geniculata (Fallén, 1823)
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Cerodontha lineella (Zetterstedt, 1838)
sg. Phytagromyza Hendel, 1920
Cerodontha avocingulata (Strobl, 1909)
= storai (Frey, 1946)
sg. Poemyza Hendel, 1931
Cerodontha atra (Meigen, 1830)
Cerodontha calamagrostidis Nowakowski, 1967
= spenceri Nowakowski, 1967
= tschirnhausi Nowakowski, 1973
Cerodontha calosoma (Hendel, 1931)
Cerodontha imbuta (Meigen, 1838)
= deschampsiae (Spencer, 1957)
Cerodontha incisa (Meigen, 1830)
Cerodontha lateralis (Macquart, 1835)
Cerodontha muscina (Meigen, 1830)
Cerodontha phragmitidis Nowakowski, 1967
Cerodontha pygmaea (Meigen, 1830)
Cerodontha pygmella (Hendel, 1931)
= lapplandica (Rydén, 1956)
Cerodontha spencerae Zlobin, 1993
= inconspicua auct. nec (Malloch, 1914)
Cerodontha thunebergi Nowakowski, 1967
sg. Xenophytomyza Frey, 1946
Cerodontha atronitens (Hendel, 1920)
Cerodontha biseta (Hendel, 1920)
= crassinervis (Frey, 1946)
Cerodontha venturii Nowakowski, 1967
CHROMATOMYIA Hardy, 1849
Chromatomyia asteris (Hendel, 1934)
Chromatomyia ciliata (Hendel, 1935)
Chromatomyia farfarella (Hendel, 1935)
Chromatomyia fuscula (Zetterstedt, 1838)
Chromatomyia glacialis (Griths, 1964)
Chromatomyia horticola (Goureau, 1851)
Chromatomyia isicae (Hering, 1962)
Chromatomyia linnaeae Griths, 1974
Chromatomyia lonicerae (Robineau-Desvoidy, 1851)
= xylostei (Kaltenbach, 1862) preocc.
Chromatomyia luzulae (Hering, 1924)
Chromatomyia milii (Kaltenbach, 1864)
Chromatomyia nigra (Meigen, 1830)
Chromatomyia norwegica (Rydén, 1957)
Chromatomyia ochracea (Hendel, 1920)
Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 297
Chromatomyia opacella (Hendel, 1935)
Chromatomyia periclymeni (Hendel, 1922)
Chromatomyia primulae (Robineau-Desvoidy, 1851)
Chromatomyia ramosa (Hendel, 1923)
= nigriventris (Hendel, 1935)
Chromatomyia styriaca Griths, 1980
Chromatomyia syngenesiae Hardy, 1849
GALIOMYZA Spencer, 1981
Galiomyza morio (Brischke, 1880)
LIRIOMYZA Mik, 1894
Liriomyza amoena (Meigen, 1830)
Liriomyza angulicornis (Malloch, 1918)
= triglochinae Hendel, 1931
Liriomyza approximata (Hendel, 1920)
Liriomyza artemisicola de Meijere, 1924
Liriomyza bryoniae (Kaltenbach, 1858)
Liriomyza buhri Hering, 1937
= adolescens Frey, 1946
Liriomyza bulbata Hendel, 1931
Liriomyza canescens Spencer, 1976
= graminicola misid.
Liriomyza cannabis Hendel, 1931
Liriomyza congesta (Becker, 1903)
= minima Hendel, 1931
= parva Hendel, 1931
Liriomyza demeijerei Hering, 1930
Liriomyza equiseti de Meijere, 1924
Liriomyza eupatorii (Kaltenbach, 1873)
= orbitella Hendel, 1931
Liriomyza aveola (Fallén, 1823)
Liriomyza avopicta Hendel, 1931
Liriomyza freyella Spencer, 1976
Liriomyza hieracii (Kaltenbach, 1862)
Liriomyza infuscata Hering, 1926
Liriomyza lutea (Meigen, 1830)
Liriomyza occipitalis Hendel, 1931
Liriomyza orbona (Meigen, 1830)
Liriomyza pedestris Hendel, 1931
Liriomyza pseudopygmina (Hering, 1933)
Liriomyza ptarmicae de Meijere, 1925
Liriomyza pusilla (Meigen, 1830)
= fasciola (Meigen, 1838)
Liriomyza pusio (Meigen, 1830)
Jere Kahanpää / ZooKeys 441: 291–303 (2014)
298
= graminicola de Meijere, 1924
= breviseta Frey, 1946
Liriomyza richteri Hering, 1927
Liriomyza sonchi Hendel, 1931
Liriomyza strigata (Meigen, 1830)
= pumila (Meigen, 1830)
Liriomyza tanaceti de Meijere, 1924
Liriomyza taraxaci Hering, 1927
Liriomyza valerianae Hendel, 1932
Liriomyza virgo (Zetterstedt, 1838)
Liriomyza virgula Frey, 1946
Liriomyza wachtlii Hendel, 1920
METOPOMYZA Enderlein, 1936
Metopomyza avonotata (Haliday, 1833)
Metopomyza interfrontalis (Melander, 1913)
= xanthaspida (Hendel, 1920)
Metopomyza scutellata (Fallén, 1823)
= avoscutellaris misid.
Metopomyza xanthaspioides (Frey, 1946)
Metopomyza xanthaspis (Loew, 1858)
NAPOMYZA Westwood, 1840
Napomyza achilleanella von Tschirnhaus, 1992
Napomyza bellidis Griths, 1967
Napomyza carotae Spencer, 1966
Napomyza cichorii Spencer, 1966
Napomyza elegans (Meigen, 1830)
Napomyza hirticornis (Hendel, 1932)
Napomyza lateralis (Fallén, 1823)
Napomyza maritima von Tschirnhaus, 1981
Napomyza merita Zlobin, 1993
Napomyza plumea Spencer, 1969
NEMORIMYZA Frey, 1946
Nemorimyza posticata (Meigen, 1830)
PHYTOBIA Lioy, 1864
= Dendromyza Hendel, 1931
Phytobia aucupariae (Kangas, 1949)
Phytobia cambii (Hendel, 1931)
= betulae (Kangas, 1935)
= tremulae (Kangas, 1949)
Phytobia mallochi (Hendel, 1924)
PHYTOLIRIOMYZA Hendel, 1931
Phytoliriomyza arctica (Lundbeck, 1901)
Phytoliriomyza dorsata (Siebke, 1863)
= Liriomyza fasciata misid.
Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 299
= pectoralis misid.
Phytoliriomyza hilarella (Zetterstedt, 1848)
Phytoliriomyza melampyga (Loew, 1869)
Phytoliriomyza mikii (Strobl, 1898)
Phytoliriomyza ornata (Meigen, 1830)
= elegantula (Zetterstedt, 1848)
Phytoliriomyza perpusilla (Meigen, 1830)
PHYTOMYZA Fallén, 1810
Phytomyza abdominalis Zetterstedt, 1848
= gentianae misid.
Phytomyza aconitophila Hendel, 1927
Phytomyza actaeae Hendel, 1922
Phytomyza adjuncta Hering, 1928
Phytomyza albiceps Meigen, 1830
= rydeniana Hering, 1949
Phytomyza albipennis Fallén, 1823
Phytomyza anemones Hering, 1925
Phytomyza angelicae Kaltenbach, 1872
Phytomyza angelicastri Hering, 1932
Phytomyza aquilegiae Hardy, 1849
Phytomyza aquilonia Frey, 1946
Phytomyza artemisivora Spencer, 1971
Phytomyza buhriella Spencer, 1969
Phytomyza calthophila Hering, 1931
Phytomyza campanulae Hendel, 1920
Phytomyza chaerophylli Kaltenbach, 1856
= anthrisci Hendel, 1924
= carvi Hering, 1931
Phytomyza cirsii Hendel, 1923
= cirsicola Hendel, 1927
Phytomyza continua Hendel, 1920
= polyarthrocera Frey, 1946
Phytomyza crassiseta Zetterstedt, 1860
Phytomyza dasyops Hendel, 1920
Phytomyza diversicornis Hendel, 1927
Phytomyza enigmoides Hering, 1937
= enigma Hering, 1936 preocc.
Phytomyza erigerophila Hering, 1927
Phytomyza eumorpha Frey, 1946
Phytomyza evanescens Hendel, 1920
= calthivora misid.
= opaca misid.
Phytomyza fallaciosa Brischke, 1880
= pseudohellebori Hendel, 1920
Jere Kahanpää / ZooKeys 441: 291–303 (2014)
300
= bonsdor Hendel, 1935
Phytomyza avicornis Fallén, 1823
Phytomyza avofemorata Strobl, 1893
= distantipila Frey, 1950
= aristata misid.
= pratensis de Meijere, 1926
Phytomyza glabra Hendel, 1935
Phytomyza glechomae Kaltenbach, 1862
? Phytomyza gymnostoma Loew, 1858
Phytomyza hellebori Kaltenbach, 1872
Phytomyza hendeli Hering, 1923
Phytomyza heracleana Hering, 1937
Phytomyza hirsuta Spencer, 1976
Phytomyza isais Hering, 1936
Phytomyza krygeri Hering, 1949
Phytomyza lappae Goureau, 1851
Phytomyza linguae Lundqvist, 1947
Phytomyza marginella Fallén, 1823
Phytomyza minuscula Goureau, 1851
Phytomyza murina Hendel, 1935
Phytomyza mylini Hering, 1954
Phytomyza nigrifemur Hering, 1934
= semitenella Hendel, 1935
Phytomyza nigripennis Fallén, 1823
Phytomyza nigritula Zetterstedt, 1838
Phytomyza notata Meigen, 1830
Phytomyza obscurella Fallén, 1823
Phytomyza pauliloewii Hendel, 1920
Phytomyza pimpinellae Hendel, 1924
Phytomyza plantaginis Robineau-Desvoidy, 1851
Phytomyza ptarmicae Hering, 1937
Phytomyza pubicornis Hendel, 1920
Phytomyza pullula Zetterstedt, 1848
Phytomyza ranunculi (Schrank, 1803)
Phytomyza ranunculivora Hering, 1932
Phytomyza rapunculi Hendel, 1927
= campanulivora Spencer, 1971
Phytomyza rhabdophora Griths, 1964
Phytomyza rostrata Hering, 1934
Phytomyza rufescens von Roser, 1840
Phytomyza rupes Meigen, 1830
Phytomyza rydeni Hering, 1934
Phytomyza sedicola Hering, 1924
Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 301
Phytomyza selini Hering, 1922
Phytomyza socia Brischke, 1880
Phytomyza soenderupi Hering, 1941
Phytomyza solidaginis Hendel, 1920
Phytomyza spinaciae Hendel, 1935
= anis misid.
Phytomyza spoliata Strobl, 1906
Phytomyza spondylii Robineau-Desvoidy, 1851
= pastinacae misid.
Phytomyza subrostrata Frey, 1946
Phytomyza tanaceti Hendel, 1923
Phytomyza tenella Meigen, 1830
Phytomyza thysselini Hendel, 1923
Phytomyza trollii Hering, 1930
Phytomyza tussilaginis Hendel, 1925
Phytomyza varipes Macquart, 1835
Phytomyza virgaureae Hering, 1926
Phytomyza virosae Pakalniškis, 2000
Phytomyza wahlgreni Rydén, 1944
= robustella misid.
PSEUDONAPOMYZA Hendel, 1920
Pseudonapomyza atra (Meigen, 1830)
Pseudonapomyza europaea Spencer, 1973
SELACHOPS Wahlberg, 1844
Selachops avocinctus Wahlberg, 1844
Excluded species
Cerodontha silvatica (Groschke, 1957) misidentication
Dizygomyza facialis Frey, 1941 nomen nudum
Liriomyza huidobrensis (Blanchard, 1926) imported
Liriomyza sativae Blanchard, 1938 imported
Liriomyza trifolii (Burgess in Comstock, 1880) imported
Phytomyza atricornis Meigen, 1838 nomen dubium
Phytomyza brischkei Hendel, 1922 misidentication
Notes
Agromyza nigrociliata. First recorded from Finland by uneberg in Kaisila (1960).
e species have since been found at several sites on the south coast (leg. L. Tien-
suu, K. Winqvist & J. Kahanpää).
Jere Kahanpää / ZooKeys 441: 291–303 (2014)
302
Cerodontha pygmella. First described from the Russian Far East, then redescribed
from northern Sweden as C. lapplandica (Rydén) (synonymized by Zlobin 2001)
and from Poland as C. tatrica Nowakowski (syn. Spencer 1976). e status of this
species as distinct from C. pygmaea has been doubtful, but Zlobin (2001) pointed
out that while C. pygmella and C. pygmaea are dicult to identify reliably as adults,
the larvae are clearly dierent.
Liriomyza pedestris Hendel, 1931 is a valid species according to Zlobin (2003). e
Finnish material of the L. aveola-group needs re-examination.
Ophiomyia fennoniensis Spencer is probably a synonym of O. eucodonus Hering.
Likewise, Ophiomyia vitiosa Spencer is probably a synonym of O. alliariae Her-
ing, 1954 (M. von Tschirnhaus, pers. comm.)
Phytomyza angelicae and P. aegopodii. Spencer (1976) synonymized P. aegopodii
Hendel, 1923 with P. angelicae Kaltenbach. Pakalniškis (2000) reinstated P. aego-
podii as a valid species but opinions on its status are not unanimous. e Finnish
material examined so far represents typical P. angelicae sensu Pakalniškis (2000).
Phytomyza brischkei. Recorded by Hackman (1980) from Finland, probably on the
basis of Frey (1937). As Spencer (1976) points out, this is a misidentication.
Phytomyza gymnostoma. Recorded from Finland in Fauna Europaea (Martinez
2011), but I have been unable to trace the original records. Possibly imported to
Finland with leeks or onions (Allium), its host plants. P. gymnostoma is an expand-
ing species with Central Europe and has lately damaged crops of Allium species
(Darvas et al. 1988).
Phytomyza linguae. Spencer (1976) synonymised this species with P. ranunculivora
Hering. Pakalniškis (1997) reinstated it as a valid species. Both species occur in
Finland: L. Tiensuu reared P. ranunculivora from Ranunculus acer and P. linguae
from Ranunculus lingua from mines collected from Vehkalahti in 1972–3.
Acknowledgements
I would like to heartily thank Michael von Tschirnhaus for all the help he has given
Finnish dipterists interested in Agromyzidae over the last decade and especially for his
numerous valuable comments on an early version of this paper.
References
Darvas B, Szarukán I, Papp L (1988) [Napomyza gymnostoma (Loew) (Dipt.: Agromyzidae), an
agromyzid pest on leek in Hungary]. Növényvédelem 24(10): 450–455. [in Hungarian]
Dempewolf M (2005) Dipteran leaf miners. In: Raman A, Schaefer CW, Withers TM (Eds)
Biology, Ecology, and Evolution of Gall-Inducing Arthropods. Volume 1. Science Pub-
lishers Inc., Plymouth, 407–429.
Frey R (1937) Finlands agromyzider, bestämda av Fr. Hendel. Notulae entomologicae 17(3): 82–96.
Checklist of the leaf-mining ies (Diptera, Agromyzidae) of Finland 303
Frey R (1946) Anteckningar om Finlands agromyzider. Notulae entomologicae 26(1–2): 13–55.
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Kaisila J (1960) Kokousselostuksia – Sitzungsberichte. Annales entomologici fennici 26(4):
291–299.
Martinez M (2011) Agromyzidae. In: Pape T, Beuk P (Eds) Fauna europaea: Diptera. Fauna
europaea version 2.4. http://www.faunaeur.org
Nowakowski JT (1973) Monographie der europäischen Arten der Gattung Cerodontha Rond.
Annales zoologici (Warszawa) 31(1): 1–327.
Pakalniškis S (1997) e Lithuanian Agromyzidae (Diptera). Descriptions of 4 new species and
other notes. In: Jonaitis V (Ed) Lietuvos entomologų darbai (Lietuvos entomologų draugijos
30-mečiui, 1996). Institute of Ecology & Lithuanian entomological Society, Vilnius, 17–34.
Pakalniškis S (2000) New data on the bionomics and distribution of Agromyzidae (Diptera)
with the description of a new species. Acta zoologica lituanica 10(1): 59–62.
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed)
Animal biodiversity: An outline of higher-level classication and survey of taxonomic richness.
Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
Scheer S, Winkler I, Wiegmann B (2007) Phylogenetic relationships within the leaf-mining
ies (Diptera: Agromyzidae) inferred from sequence data from multiple genes. Molecular
Phylogenetics and Evolution 42(3): 756–775. doi: 10.1016/j.ympev.2006.12.018
Spencer KA (1976) e Agromyzidae (Diptera) of Fennoscandia and Denmark. Fauna ento-
mologica Scandinavica 5(1–2). Scandinavian Science Press, Klampenborg, 606 pp.
Winkler I, Sheer S, Mitter C (2009) Molecular phylogeny and taxonomy in leafmining ies
(Diptera: Agromyzidae): Delimitation of Phytomyza Fallén s. lat. and its species groups, with
new insights on genitalic and host-use evolution. Systematic Entomology 34(2): 260–292.
doi: 10.1111/j.1365-3113.2008.00462.x
Winkler IS, Rung A, Scheer SJ (2010) Hennig’s orphans revisited: Testing morphological
hypotheses in the “Opomyzoidea” (Diptera: Schizophora). Molecular Phylogenetics and
Evolution 54(3): 746–762. doi: 10.1016/j.ympev.2009.12.016
Zlobin VV (2001) Review of Mining Flies of the Genus Cerodontha Rond. XI. Some remarks
on the Oriental Poemyza Species (Diptera: Agromyzidae). International Journal of Dip-
terological Research 12(2): 107–112.
Zlobin VV (2003) Review of Mining Flies of the Genus Liriomyza Mik (Diptera: Agromyzidae).
I. e Palaearctic aveola-Group Species. International Journal of Dipterological Research
13(3): 145–178.
Jere Kahanpää / ZooKeys 441: 291–303 (2014)
304
Checklist of the Diptera families Acartophthalmidae, Canacidae (including Tethinidae)... 305
Checklist of the Diptera families Acartophthalmidae,
Canacidae (including Tethinidae), Carnidae and
Milichiidae of Finland (Insecta)
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 27 January 2014|Accepted 6 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the Diptera families Acartophthalmidae, Canacidae (including Tethinidae),
Carnidae and Milichiidae of Finland (Insecta). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland.
ZooKeys 441: 305–309. doi: 10.3897/zookeys.441.7144
Abstract
A checklist of 29 species in the smaller carnoid families Acartophthalmidae, Canacidae, Carnidae and
Milichiidae (Diptera) recorded from Finland is presented. Tethinidae are also included as a subfamily
of Canacidae. Phyllomyza tetragona Hendel is removed from the list as no reliable records of this species
within the post-1944 borders of Finland could be found.
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
e superfamily Carnoidea (sensu Buck 2006 but excluding Cryptochetidae, see Wieg-
mann et al. 2011) includes seven families. e largest by far is Chloropidae, treated in
a separate article in this issue. Four of the remaining six are found in Finland: Acar-
ZooKeys 441: 305–309 (2014)
doi: 10.3897/zookeys.441.7144
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 305–309 (2014)
306
tophthalmidae, Canacidae, Carnidae and Milichiidae. e remaining two families,
Inbiomyiidae and Australimyzidae, are respectively restricted to the Neotropical and
Australasian Regions.
All four families are rather poorly studied in Finland. e presence of additional,
even undescribed, species is very likely in Carnidae and Milichiidae. e identication
of all species of Meoneura was veried from male genitalia during the preparation of
this checklist with one exception: the male of Meoneura elongella (Zetterstedt, 1838)
remains undescribed. It is the only North European Meoneura with black halteres and
as such easily recognised.
World catalogues have recently been published for Canacidae (Munari and Mathis
2010), Carnidae (Brake 2011), and Milichiidae (Brake 2000). e Finnish species of
these families were last listed in Hackman (1980).
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Carnoidea Newman, 1834
ACARTOPHTHALMIDAE Czerny, 1928
ACARTOPHTHALMUS Czerny, 1902
Acartophthalmus bicolor Oldenberg, 1910
Acartophthalmus nigrinus (Zetterstedt, 1848)
Acartophthalmus pusio Frey, 1947
CANACIDAE Jones, 1906
PELOMYIINAE Foster, 1976
PELOMYIELLA Hendel, 1934
Pelomyiella cinerella (Haliday, 1837)
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Acartophthalmidae 5 3 3 average
Canacidae 322 38 1average
Carnidae 93 (Brake 2011) 39 13 poor
Milichiidae 278 43 12–13 average
Checklist of the Diptera families Acartophthalmidae, Canacidae (including Tethinidae)... 307
CARNIDAE Newman, 1834
CARNUS Nitzsch, 1818
Carnus hemapterus Nitzsch, 1818
MEONEURA Rondani, 1856
Meoneura anceps Frey, 1935
Meoneura elongella (Zetterstedt, 1838)
Meoneura avifacies Collin, 1930
Meoneura glaberrima Becker, 1907
= neglecta Collin, 1930
Meoneura lacteipennis (Fallén, 1823)
Meoneura lamellata Collin, 1930
Meoneura minutissima (Zetterstedt, 1860)
Meoneura neottiophila Collin, 1930
Meoneura obscurella (Fallén, 1823)
Meoneura prima (Becker, 1903)
= seducta Collin, 1937
Meoneura triangularis Collin, 1930
Meoneura vagans (Fallén, 1823)
MILICHIIDAE Schiner, 1862
MADIZINAE Czerny, 1909
DESMOMETOPA Loew, 1866
Desmometopa m-nigrum (Zetterstedt, 1848)
Desmometopa sordida (Fallén, 1820)
Desmometopa varipalpis Malloch, 1927
LEPTOMETOPA Becker, 1903
Leptometopa latipes (Meigen, 1830)
MADIZA Fallén, 1810
Madiza glabra Fallén, 1820
MILICHIINAE Schiner, 1862
MILICHIA Meigen, 1830
Milichia ludens (Wahlberg, 1847)
PHYLLOMYZINAE Curran, 1934
NEOPHYLLOMYZA Melander, 1913
Neophyllomyza acyglossa (Villeneuve, 1920)
PHYLLOMYZA Fallén, 1810
? Phyllomyza equitans (Hendel, 1919)
Phyllomyza formicae Schmitz, 1923
Phyllomyza longipalpis (Schmitz, 1924)
Phyllomyza rubricornis Schmitz, 1923
Phyllomyza securicornis Fallén, 1823
Phyllomyza silesiaca (Duda, 1935)
Jere Kahanpää / ZooKeys 441: 305–309 (2014)
308
Excluded species
Phyllomyza tetragona Hendel, 1924 not found within present borders
Tethina grisea (Fallén, 1823) misidentication?
Tethina illota (Haliday, 1838)
Notes
Meoneura. At least two additional, possibly undescribed species occur in Finland.
Phyllomyza equitans (Hendel, 1919). Only females have been found in Finland.
Tethina grisea (Fallén, 1823) and T. illota (Haliday, 1838). Munari and Mathis
(2010) list two Tethina species from Finland. Krogerus (1932) is mentioned as the
source for the T. grisea record; there is no listed source for the record of T. illota.
An older version of the world catalogue (Mathis and Munari 1996) mentions only
T. grisea from Finland. No Finnish specimens of these species could be found in
the Finnish Museum of Natural History (MZH) where most of the material col-
lected by Krogerus is kept, or in any other Finnish insect collection. Unfortunately
the identications in Krogerus (1932) are rather unreliable. It is possible that Teth-
ina species occur in Finland, but none are included in this checklist because the
evidence for their presence is, in my opinion, insucient.
References
Brake I (2000) Phylogenetic systematics of the Milichiidae (Diptera, Schizophora). Entomo-
logica Scandinavica Supplement 57: 1–120.
Buck M (2006) A new family and genus of acalypterate ies from the Neotropical region, with
a phylogenetic analysis of Carnoidea family relationships (Diptera, Schizophora). System-
atic Entomology 31(3): 377–404. doi: 10.1111/j.1365-3113.2006.00328.x
Brake I (2011) World catalog of the family Carnidae (Diptera, Schizophora). In: Brake I,
ompson FC (Eds) Contributions to the Systema Dipterorum (Insecta: Diptera). Myia
vol. 12. Pensoft Publishers & North America Dipterists Society, Soa, Moscow & Wash-
ington D.C., 113–169. http://diptera.myspecies.info/sites/diptera.myspecies.info/les/
Carnidae_catalog_0.pdf
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Krogerus R (1932) Über die Ökologie und Verbreitung der Arthropoden der Triebsandgebiete
an den Küsten Finnlands. Acta Zoologica Fennica 12: 1–308.
Mathis WN, Munari L (1996) World catalog of the family Tethinidae (Diptera). Smithsonian
Contributions to Zoology 584: 1–27. doi: 10.5479/si.00810282.584
Checklist of the Diptera families Acartophthalmidae, Canacidae (including Tethinidae)... 309
Munari L, Mathis WN (2010) World catalog of the family Canacidae (including Tethini-
dae), with keys to the supraspecic taxa. Zootaxa 2471: 1–84. http://www.mapress.com/
zootaxa/2010/f/zt02471p084.pdf
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang Z-Q (Ed.)
Animal biodiversity: An outline of higher-level classication and survey of taxonomic richness.
Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/zt03148p229.pdf
Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA,
Cassel BK, Bayless KM, Heimberg AM, Wheeler BM, Peterson KJ, Pape T, Sinclair BJ, Sk-
evington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Ott U, Kampmeier GE, omp-
son FC, Grimaldi DA, Beckenbach AT, Courtney GW, Friedrich M, Meier R, Yeates DK
(2011) Episodic radiations in the y tree of life. Proceedings of the National Academy of Sci-
ences of the United States of America 108(14): 5690–5695. doi: 10.1073/pnas.1012675108
Jere Kahanpää / ZooKeys 441: 305–309 (2014)
310
Checklist of the family Chloropidae (Diptera) of Finland 311
Checklist of the family Chloropidae
(Diptera) of Finland
Emilia Nartshuk1, Jere Kahanpää2
1 Zoological Institute of the Russian Academy of Sciences, Universitetskaya nab. 1 St.-Petersburg 199034 Russia
2 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Emilia Nartshuk (chlorops@zin.ru)
Academic editor: J. Salmela|Received 24 March 2014|Accepted 10 June 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Nartshuk E, Kahanpää J (2014) Checklist of the family Chloropidae (Diptera) of Finland. In: Kahanpää J,
Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 311–318. doi: 10.3897/zookeys.441.7505
Abstract
A checklist of 147 species the Chloropidae (Diptera) recorded from Finland. Centorisoma elegantulum
Becker is recorded for the rst time from Finland.
Keywords
Finland, Chloropidae, species list, biodiversity, faunistics
Introduction
e Chloropidae is a large family of acalyptrate Diptera. It belongs to superfamily
Carnoidea with Milichiidae as the closest relative. e classication of the family used
follows Andersson (1977) and Nartshuk (1983).
e North European chloropid fauna has recently been revised by Nartshuk and
Andersson (2013). Details of Finnish chloropid literature, species distribution and
ecology, and other details can be found in their book. In comparison with the neigh-
bouring countries, Finland ranks second in the number of chloropid species after Swe-
den (189 species) but well ahead of Denmark (119 species) and Norway (97 species).
e genera Gaurax, Incertella and Rhopalopterum need more attention as there are sev-
eral species in these genera known from neighbouring countries but not from Finland.
ZooKeys 441: 311–318 (2014)
doi: 10.3897/zookeys.441.7505
www.zookeys.org
Copyright E. Nartshuk, J. Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Emilia Nartshuk & Jere Kahanpää / ZooKeys 441: 311–318 (2014)
312
is checklist is based mostly on museum material in the Finnish Natural History
museum (MZH) studied by E. Nartshuk. e Finnish species were last listed in Nart-
shuk and Andersson (2013).
Number of species:
World: 2880 species (Pape et al. 2011)
Europe: 394 species
Finland: 149–150 species
Faunistic knowledge level in Finland: average
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Carnoidea Newman, 1834
CHLOROPIDAE Rondani, 1856
CHLOROPINAE Rondani, 1856
CENTORISOMA Hendel, 1907
Centorisoma elegantulum Becker, 1910 (see Notes)
CETEMA Hendel, 1907
= Centor Loew, 1866 preocc.
sg. Cetema Hendel, 1907
Cetema cereris (Fallén, 1820)
Cetema elongatum (Meigen, 1830)
Cetema myopinum (Loew, 1866)
Cetema neglectum Tonnoir, 1921
Cetema simile Ismay, 1985
CHLOROPS Meigen, 1803
= Oscinis Latreille, 1804
sg. Chlorops Meigen, 1803
Chlorops anthracophagoides Strobl, 1900 (see Notes)
Chlorops calceatus Meigen, 1830
Chlorops centromaculatus Duda, 1933
Chlorops geminatus Meigen, 1830
Chlorops gracilis Meigen, 1830
Chlorops hypostigma Meigen, 1830
Chlorops kirigaminensis Kanmiya, 1978
Checklist of the family Chloropidae (Diptera) of Finland 313
= zonulatus auct. nec Wahlgren, 1913
Chlorops laetus Meigen, 1830
Chlorops meigenii Loew, 1866
= rufescens Oldenberg, 1923
Chlorops nigripalpis (Duda, 1933)
= crassipalpis Smirnov, 1958
Chlorops obscurellus (Zetterstedt, 1838)
= brunnipes auct. nec (Zetterstedt, 1848)
Chlorops palpatus Smirnov, 1959
Chlorops pannonicus Strobl, 1893
Chlorops planifrons (Loew, 1866)
= triangularis Becker, 1910
Chlorops pumilionis (Bjerkander, 1778)
Chlorops ringens Loew, 1866
Chlorops riparius Smirnov, 1958
Chlorops rossicus Smirnov, 1955
Chlorops runus (Zetterstedt, 1848)
= citrinellus (Zetterstedt, 1848)
Chlorops scalaris Meigen, 1830
Chlorops scutellaris (Zetterstedt, 1838)
= laevicollis Becker, 1910
= freyi Duda, 1933
Chlorops serenus Loew, 1866
Chlorops speciosus Meigen, 1830
Chlorops troglodytes (Zetterstedt, 1848)
sg. Sclerophallus Beschovski, 1978
Chlorops limbatus Meigen, 1830
= brevimanus Loew, 1866
Chlorops varsoviensis Becker, 1910
CHLOROPSINA Becker, 1911
Chloropsina distinguenda (Frey, 1909)
CRYPTONEVRA Lioy, 1864
Cryptonevra diadema (Meigen, 1830)
Cryptonevra avitarsis (Meigen, 1830)
DIPLOTOXA Loew, 1863
Diplotoxa messoria (Fallén, 1820)
DIPLOTOXOIDES Andersson, 1977
Diplotoxoides dalmatina (Strobl, 1900)
EPICHLOROPS Becker, 1910
Epichlorops puncticollis (Zetterstedt, 1848)
EUTROPHA Loew, 1866
Eutropha fulvifrons (Haliday, 1833)
Eutropha variegata Loew, 1866
Emilia Nartshuk & Jere Kahanpää / ZooKeys 441: 311–318 (2014)
314
LASIOSINA Becker, 1910
Lasiosina albipila (Loew, 1866)
Lasiosina brevisurstylata Dely-Draskovits, 1977
Lasiosina herpini (Guérin-Méneville, 1843)
= cinctipes auct. nec (Meigen, 1830)
Lasiosina parvipennis Duda, 1933
MELANUM Becker, 1910
Melanum laterale (Haliday, 1833)
MEROMYZA Meigen, 1830
sg. Meromyza Meigen, 1830
Meromyza curvinervis (Zetterstedt, 1848)
= hybrida Péter, 1961
Meromyza elbergi Fedoseeva, 1979
Meromyza ingrica Nartshuk, 1992
Meromyza mosquensis Fedoseeva, 1960
Meromyza nigriseta Fedoseeva, 1960
Meromyza nigriventris Macquart, 1835
= rostrata Hubicka, 1966
Meromyza ornata (Wiedemann, 1817)
= sororcula Fedoseeva, 1962
Meromyza palposa Fedoseeva, 1960
Meromyza pluriseta Péter, 1961
Meromyza pratorum Meigen, 1830
Meromyza rohdendor Fedoseeva, 1974
Meromyza saltatrix (Linnaeus, 1761)
Meromyza sibirica Fedoseeva, 1961
Meromyza triangulina Fedoseeva, 1960
Meromyza variegata Meigen, 1830
= lidiae Nartshuk, 1992
Meromyza zimzerla Nartshuk, 1992
NEOHAPLEGIS Beschovski, 1981
Neohaplegis tarsata (Fallén, 1820)
PLATYCEPHALA Fallén, 1820
Platycephala planifrons (Fabricius, 1798)
Platycephala umbraculata (Fabricius, 1794)
PSEUDOPACHYCHAETA Strobl, 1902
Pseudopachychaeta approximatonervis (Zetterstedt, 1848)
Pseudopachychaeta oscinina (Fallén, 1813)
= heleocharis (Nartshuk, 1964)
Pseudopachychaeta ruceps (Zetterstedt, 1838)
THAUMATOMYIA Zenker, 1833
aumatomyia glabra (Meigen, 1830)
aumatomyia hallandica Andersson, 1966
Checklist of the family Chloropidae (Diptera) of Finland 315
aumatomyia notata (Meigen, 1830)
aumatomyia rufa (Macquart, 1835)
aumatomyia trifasciata (Zetterstedt, 1848)
TRICHIEURINA Duda, 1933
Trichieurina pubescens (Meigen, 1830)
OSCINELLINAE Becker, 1910
APHANOTRIGONUM Duda, 1932
Aphanotrigonum cinctellum (Zetterstedt, 1848)
= fasciella (Zetterstedt, 1855)
Aphanotrigonum hungaricum Dely-Draskovits, 1981
Aphanotrigonum nigripes (Zetterstedt, 1848)
Aphanotrigonum trilineatum (Meigen, 1830)
= beschovskii Dely-Draskovits, 1981
ASPISTYLA Duda, 1933
Aspistyla plumiger (Meigen, 1830)
CALAMONCOSIS Enderlein, 1911
sg. Calamoncosis Enderlein, 1911
Calamoncosis aprica (Meigen, 1830)
Calamoncosis duinensis (Strobl, 1909)
Calamoncosis minima (Strobl, 1893)
Calamoncosis oscinella (Becker, 1910)
sg. Rhaphiopyga Nartshuk, 1971
Calamoncosis glyceriae Nartshuk, 1958
COLLINIELLA Nartshuk & Andersson, 2013
Colliniella meijerei (Duda, 1932) (see Notes)
CONIOSCINELLA Duda, 1929
Conioscinella frontella (Fallén, 1820)
Conioscinella gallarum (Duda, 1933)
Conioscinella livida Nartshuk, 1970
Conioscinella mimula Collin, 1946
Conioscinella sordidella (Zetterstedt, 1848)
Conioscinella zetterstedti Andersson, 1966
DICRAEUS Loew, 1873
sg. Dicraeus Loew, 1873
Dicraeus tibialis (Macquart, 1835)
sg. Oedesiella Becker, 1910
Dicraeus fennicus Duda, 1932
Dicraeus rossicus Stackelberg, 1955
sg. Paroedesiella Anonymous in Imperial Institute of Entomology, 1937
Dicraeus nitidus Wahlgren, 1913
= napaeus Collin, 1946
Dicraeus styriacus (Strobl, 1898)
= vallaris Collin, 1946
Emilia Nartshuk & Jere Kahanpää / ZooKeys 441: 311–318 (2014)
316
Dicraeus vagans (Meigen, 1838)
ELACHIPTERA Macquart, 1835
Elachiptera cornuta (Fallén, 1820)
Elachiptera diastema Collin, 1946
Elachiptera scrobiculata (Strobl, 1901)
= tenuiseta (Frey, 1947)
Elachiptera tuberculifera (Corti, 1909)
ERIBOLUS Becker, 1910
Eribolus gracilior (de Meijere, 1918)
Eribolus hungaricus Becker, 1910
Eribolus nana (Zetterstedt, 1838)
= sudeticus Becker, 1910
Eribolus slesvicensis Becker, 1910
GAMPSOCERA Schiner, 1862
Gampsocera numerata (Heeger, 1858)
GAURAX Loew, 1863
Gaurax borealis (Duda, 1933)
Gaurax dubius (Macquart, 1835)
Gaurax ephippium (Zetterstedt, 1848)
= strobilum Karps, 1981
Gaurax fascipes Becker, 1910
Gaurax macrocerus (Nartshuk, 1962)
Gaurax maculipennis (Zetterstedt, 1848)
HAPLEGINELLA Duda, 1933
Hapleginella laevifrons (Loew, 1858)
INCERTELLA Sabrosky, 1980
Incertella albipalpis (Meigen, 1830)
Incertella kerteszi (Becker, 1910)
Incertella nigrifrons (Duda, 1933)
Incertella scotica (Collin, 1946)
Incertella zuercheri (Duda, 1933)
LASIAMBIA Anonymous in Imperial Instutute of Entomology, 1937
= Lasiambia Enderlein, 1936
= Lasiambia Sabrosky, 1941
Lasiambia brevibucca (Duda, 1932)
Lasiambia coxalis (von Roser, 1840)
= oophila (Hennig, 1941)
Lasiambia palposa (Fallén, 1820)
LIPARA Meigen, 1830
Lipara lucens Meigen, 1830
Lipara pullitarsis Doskočil & Chvála, 1971
= rutarsis misid.
MICROCERCIS Beschovski, 1978
Checklist of the family Chloropidae (Diptera) of Finland 317
Microcercis trigonella (Duda, 1933)
OSCINELLA Becker, 1909
sg. Oscinella Becker, 1909
Oscinella angularis Collin, 1946
Oscinella cariciphila Collin, 1946
Oscinella frit (Linnaeus, 1758)
Oscinella maura (Fallén, 1820)
= albiseta (Meigen, 1830)
Oscinella nigerrima (Macquart, 1835)
Oscinella nitidissima (Meigen, 1838)
Oscinella pusilla (Meigen, 1830)
Oscinella ventricosi Nartshuk, 1956
Oscinella vindicata (Meigen, 1830)
= hortensis Collin, 1946
OSCINIMORPHA Lioy, 1864
Oscinimorpha minutissima (Strobl, 1900)
Oscinimorpha sordidissima (Strobl, 1893)
OSCINISOMA Lioy, 1864
Oscinisoma cognatum (Meigen, 1830)
POLYODASPIS Duda, 1933
Polyodaspis rucornis (Macquart, 1835)
Polyodaspis sulcicollis (Meigen, 1838)
RHOPALOPTERUM Duda, 1929
Rhopalopterum anthracinum (Meigen, 1930)
Rhopalopterum atricillum (Zetterstedt, 1838)
= atripes (Duda, 1933)
Rhopalopterum atricorne (Zetterstedt, 1838)
= platythorax (Nartshuk, 1958)
Rhopalopterum brunneipenne Beschovski & Lansbury, 1987
Rhopalopterum fasciolum (Meigen, 1830)
Rhopalopterum femorale (Collin, 1946)
SIPHONELLA Macquart, 1835
Siphonella oscinina (Fallén, 1820)
SIPHUNCULINA Rondani, 1856
Siphunculina aenea (Macquart, 1835)
SPECCAFRONS Sabrosky, 1980
Speccafrons halophila (Duda, 1933)
TRACHYSIPHONELLA Enderlein, 1936
Trachysiphonella ruceps (Macquart, 1835)
= pygmaea (Meigen, 1838)
Trachysiphonella scutellata (von Roser, 1840)
TRICIMBA Lioy, 1864
sg. Nartshukiella Beschovski, 1981
Emilia Nartshuk & Jere Kahanpää / ZooKeys 441: 311–318 (2014)
318
Tricimba cincta (Meigen, 1830)
= sulcella (Zetterstedt, 1848)
sg. Tricimba Lioy, 1864
Tricimba lineella (Fallén, 1820)
Excluded species
Chlorops guratus (Zetterstedt, 1848) not found within present borders.
Notes
Centorisoma elegantulum Becker, 1910. First records from Finland: 1 ex Ta: Heinola,
Lusi, July 2013, leg. J. Kahanpää; 6 exx N: Loviisa, 2013, leg. Jari Flinck.
Chlorops anthracophagoides Strobl, 1900. Recorded in Nartshuk (1998) from
Eriksberg, which is a Finnish locality.
Colliniella meijerei (Duda, 1932). Recently added to the list of Finnish Diptera
(Kahanpää 2013).
References
Andersson H (1977) Taxonomic and phylogenetic studies on Chloropidae (Diptera) with spe-
cial reference to Old World genera. Entomologica scandinavica Supplement 8. Berlingska
Boktryckeriet, Lund, 200 pp.
Kahanpää J (2013) Viisikymmentä Suomelle uutta kärpäslajia (Diptera: Brachycera). Sahlbergia
19(1–2): 63–71.
Nartshuk EP (1983) A system of the superfamily Chloropoidea (Diptera, Cyclorrhapha). En-
tomologicheskoe Obozrenie 62: 638–648.
Nartshuk EP (1998) A revision of grassies of the tribe Chloropini (Diptera, Chloropidae) of
Finland, Estonia and North-West Russia. Entomologica fennica 9(3): 153–183.
Nartshuk EP, Andersson H (2013) e Frit Flies (Chloropidae, Diptera) of Fennoscandia and
Denmark. Fauna Entomologica Scandinavica Brill, Leiden, Vol. 43(2012), 282 pp. doi:
10.1163/9789004190665
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Checklist of the y families Chyromyidae andHeleomyzidae (Diptera) of Finland 319
Checklist of the fly families Chyromyidae
and Heleomyzidae (Diptera) of Finland
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: Jukka Salmel|Received 13 March 2014|Accepted 6 May 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the y families Chyromyidae and Heleomyzidae (Diptera) of Finland. In:
Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 319–324. doi: 10.3897/zookeys.441.7507
Abstract
A Finnish checklist of the sphaeroceroid y families Chyromyidae and Heleomyzidae is provided.
Keywords
Species list, Finland, Diptera, biodiversity, faunistics
Introduction
e superfamily Sphaeroceroidea is a medium-sized one, with two families of moder-
ate diversity, Sphaeroceridae (1550 species) and Heleomyzidae (~720 species), and
the small family Chyromyidae. e enigmatic afrotropical Mormotomyia hirsuta Aus-
ten, 1936 was once placed near Sphaeroceridae but it is now seen as an ephydroid
y (Kirk-Spriggs et al. 2011). McAlpine (2007) has proposed an alternative concept
for Sphaeroceroidea with Sphaeroceridae and Heleomyzidae united as a single family
called Heteromyzidae. is proposal has not gained signicant support and for the
purposes of this checklist the traditional concept of family Sphaeroceridae is retained.
ZooKeys 441: 319–324 (2014)
doi: 10.3897/zookeys.441.7507
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 319–324 (2014)
320
ere is no general agreement on the relationships of various heleomyzid tribes. Sev-
eral dierent schemes for subfamilies have been proposed (see McAlpine 2007, McAlp-
ine and Woodley 2010). Some taxa treated here as heleomyzids (primarily Trixosce-
lidinae, Chiropteromyzinae, Heteromyzinae and Borboropsini) may deserve full family
status. As a conservative approach this checklist follows Marshall (2012) and keeps them
as subfamilies and tribes. e heleomyzid subfamilies and tribes are listed alphabetically.
e Finnish chyromyids are small yellow ies with (at least while alive) iridescent
blue or green eyes. Chyromyids are rarely collected and little is known about their ecol-
ogy or the proper place of the family within Sphaeroceroidea. ey may actually be a
specialized lineage arising from within Heleomyzidae sensu lato.
Two of the three sphaeroceroid families are treated in this paper. e largest,
Sphaeroceridae, is covered in a separate paper in this issue of ZooKeys. e Finnish
species of Heleomyzidae and Chyromyidae were last listed by Hackman (1980).
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Sphaeroceroidea Macquart, 1835
CHYROMYIDAE Hendel, 1916
CHYROMYINAE Hendel, 1916
CHYROMYA Robineau-Desvoidy, 1830
Chyromya ava (Linnaeus, 1758)
Chyromya oppidana (Scopoli, 1763)
GYMNOCHIROMYIA Hendel, 1933
Gymnochiromyia avella (Zetterstedt, 1848)
= minima (Becker, 1904)
Gymnochiromyia inermis (Collin, 1933)
HELEOMYZIDAE Westwood, 1840
BORBOROPSINAE Griths, 1972
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Chyromyidae 138 59 4 poor–average
Heleomyzidae 727 175 61 average
Checklist of the y families Chyromyidae andHeleomyzidae (Diptera) of Finland 321
BORBOROPSIS Czerny, 1902
Borboropsis puberula (Zetterstedt, 1838)
= fulviceps (Strobl, 1898)
CHIROPTEROMYZINAE Frey, 1952
CHIROPTEROMYZA Frey, 1952
Chiropteromyza broersei (de Meijere, 1946)
= wegelii Frey, 1952
NEOSSOS Malloch, 1927
= Ornitholeria Frey, 1930
Neossos nidicola (Frey, 1930)
HETEROMYZINAE Fallén, 1820
HETEROMYZA Fallén, 1820
Heteromyza atricornis Meigen, 1830
Heteromyza oculata Fallén, 1820
Heteromyza rotundicornis (Zetterstedt, 1846)
TEPHROCHLAMYS Loew, 1862
Tephrochlamys avipes (Zetterstedt, 1838)
Tephrochlamys ruventris (Meigen, 1830)
= lapponica (Czerny, 1924)
Tephrochlamys steniusi Frey, 1930
Tephrochlamys tarsalis (Zetterstedt, 1847)
HELEOMYZINAE Westwood, 1840
tribe Heleomyzini Westwood, 1840
GYMNOMUS Loew, 1863
Gymnomus amplicornis (Czerny, 1924)
HELEOMYZA Fallén, 1810
= Helomyza Fallén, 1820 emend.
= Leria Robineau-Desvoidy, 1830
sg. Heleomyza Fallén, 1810
Heleomyza borealis Boheman, 1865
= czernyi Collart, 1933
= modesta misid.
Heleomyza hackmani Frey, 1950
Heleomyza pleuralis (Becker, 1907)
Heleomyza serrata (Linnaeus, 1758)
MORPHOLERIA Garrett, 1921
sg. Spanoparea Czerny, 1924
Morpholeria dudai (Czerny, 1924)
Morpholeria kerteszii Czerny, 1924
Morpholeria obscuriventris (Zetterstedt, 1847)
Morpholeria rucornis (Meigen, 1830)
NEOLERIA Malloch, 1919
Neoleria inscripta (Meigen, 1830)
Jere Kahanpää / ZooKeys 441: 319–324 (2014)
322
= minuta (Zetterstedt, 1838)
Neoleria prominens (Becker, 1897)
= tibialis misid.
Neoleria rucauda (Zetterstedt, 1847)
Neoleria ruceps (Zetterstedt, 1838)
SCOLIOCENTRA Loew, 1862
sg. Chaetomus Czerny, 1924
Scoliocentra confusa (Wahlgren, 1918)
Scoliocentra avotestacea (Zetterstedt, 1838)
sg. Leriola Gorodkov, 1962
Scoliocentra brachypterna (Loew, 1873)
Scoliocentra nigrinervis (Wahlgren, 1918)
sg. Scoliocentra Loew, 1862
Scoliocentra dupliciseta (Strobl, 1894)
Scoliocentra scutellaris (Zetterstedt, 1838)
Scoliocentra villosa (Meigen, 1830)
tribe Oecotheini Gorodkov, 1972
ECCOPTOMERA Loew, 1862
Eccoptomera infuscata Wahlgren, 1918
Eccoptomera longiseta (Meigen, 1830)
Eccoptomera marginicornis Czerny, 1924
Eccoptomera microps (Meigen, 1830)
Eccoptomera obscura (Meigen, 1830)
Eccoptomera ornata Loew, 1862
Eccoptomera pallescens (Meigen, 1830)
OECOTHEA Haliday, 1837
Oecothea fenestralis (Fallén, 1820)
tribe Orbelliini Gorodkov, 1972
ORBELLIA Robineau-Desvoidy, 1830
Orbellia nivicola Frey, 1913
SUILLIINAE Wahlgren, 1917
SUILLIA Robineau-Desvoidy, 1830
= Allophyla Loew, 1862
Suillia anis (Meigen, 1830)
Suillia apicalis (Loew, 1862)
Suillia atricornis (Meigen, 1830)
Suillia bicolor (Zetterstedt, 1838)
Suillia femoralis (Loew, 1862)
Suillia ava (Meigen, 1830)
Suillia avifrons (Zetterstedt, 1838)
= nudipes (Czerny, 1932)
Suillia fuscicornis (Zetterstedt, 1847)
Suillia humilis (Meigen, 1830)
Checklist of the y families Chyromyidae andHeleomyzidae (Diptera) of Finland 323
= inornata (Loew, 1862)
Suillia laevifrons (Loew, 1862)
Suillia lineitergum (Pandellé, 1901)
= stroblii (Czerny, 1904)
Suillia lurida (Meigen, 1830)
Suillia mikii (Pokorny, 1886)
Suillia nemorum (Meigen, 1830)
Suillia pallida (Fallén, 1820)
Suillia parva (Loew, 1862)
= collini Hackman, 1972
= avifrons auct. nec (Zetterstedt, 1838)
Suillia quadrilineata Czerny, 1924
Suillia vaginata (Loew, 1862)
TRIXOSCELIDINAE Hendel, 1916
TRIXOSCELIS Rondani, 1856
Trixoscelis frontalis (Fallén, 1823)
? = canescens misid. (see Notes)
Trixoscelis marginella (Fallén, 1823)
Trixoscelis obscurella (Fallén, 1823)
Trixoscelis similis Hackman, 1970
Notes
Chyromya oppidana (Scopoli, 1763). Found only inside houses and farm buildings
in Finland.
Orbellia nivicola Frey, 1913. is species was synonymized with O. myiopiformis R.-
D. by Storå (1958), but Frey (1958) defended its validity. e status of O. nivicola
as a species needs verication.
Trixoscelis canescens (Loew, 1865). is species was originally described on the basis of
a single female. Soós (1979) examined the type and revived the name from synonymy
with T. frontalis. Woźnica (2008) provided an illustrated diagnosis for T. canescens
and synonymized T. gigans Carles-Tolrá, 2001 and T. fumipennis Papp, 2005 with
it. e species was recently recorded from Finland by Flinck and Kahanpää (2013).
Specimens with darkened costal veins and dorsal abdominal surfaces, both proposed
diagnostic characters of T. canescens, are common among Finnish T. frontalis mate-
rial (see Fig. 8 in Flinck and Kahanpää 2013). Finnish males with these features
have genitalia identical with those illustrated for T. frontalis by Hackman (1970)
and quite unlike the genitalia of T. gigans (= fumipennis Papp). e male specimen
mentioned in Flinck and Kahanpää (2013) was later dissected and it belongs to T.
frontalis. e external characters (darkened costa and dorsum of abdomen) can not
be used to reliably separate T. canescens from T. frontalis. e Finnish records of T.
canescens are probably all misidentications of T. frontalis.
Jere Kahanpää / ZooKeys 441: 319–324 (2014)
324
References
Flinck J, Kahanpää J (2013) Suomelle uusia kärpäslajeja (Diptera: Brachycera). Sahlbergia
19(1–2): 53–62.
Frey R (1958) Über Orbellia nivicola Frey (Dipt., Helomyzidae). Notulae entomologicae
38: 97–100.
Hackman W (1970) Trixoscelidae (Diptera) from Southern Spain and description of a new
Trixoscelis species from Northern Europe. Entomologica Scandinavica 1(2): 127–134. doi:
10.1163/187631270X00168
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Kirk-Spriggs AH, Kotrba M, Copeland RS (2011) Further details of the morphology of the
enigmatic African y Mormotomyia hirsuta Austen (Diptera: Mormotomyiidae). African
Invertebrates 52(1): 145–165. doi: 10.5733/an.052.0107
Marshall SA (2012) Flies: e Natural History and Diversity of Diptera. Firey Books, Rich-
mond Hill, Ontario, 616 pp.
McAlpine DK (2007) Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae),
with reconsideration of the status of families Heleomyzidae and Sphaeroceridae, and de-
scriptions of femoral gland-baskets. Records of the Australian Museum 59(3): 143–219.
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zt03148p229.pdf
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Storå R (1958) Über einige Helomyziden-Arten (Diptera) Finnlands. Notulae entomologicae
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Checklist of the family Sphaeroceridae (Diptera) of Finland 325
Checklist of the family Sphaeroceridae
(Diptera) of Finland
Antti Haarto1, Jere Kahanpää2
1 Zoological Museum, Section of Biodiversity and Environmental Science, Department of Biology, FI-20014
University of Turku, Finland 2 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014
University of Helsinki, Finland
Corresponding author: Antti Haarto (ahaarto@gmail.com)
Academic editor: J.Salmela|Received 12 February 2014|Accepted 23 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Haarto A, Kahanpää J (2014) Checklist of the family Sphaeroceridae (Diptera) of Finland. In: Kahanpää J,
Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 325–332. doi: 10.3897/zookeys.441.7250
Abstract
A checklist of the family Sphaeroceridae (Diptera) recorded from Finland.
Keywords
Checklist, Finland, Diptera, Sphaeroceridae
Introduction
Sphaeroceridae or lesser dung ies are mostly small and dull dark brown to grey aca-
lyptrate ies. ey are easily distinguished from other acalyptratae because of the short
and thickened basal tarsomeres of hind legs. Larvae of Sphaeroceridae develop in de-
caying organic matter such as decaying vegetation, dung and rotten fungi.
e family has gathered little interest among entomologists in Finland. Walter
Hackman concentrated on the taxonomy of the subfamily Copromyzinae (Hackman
1965) and investigated the dipterous fauna, including sphaerocerids, in the burrows
of small mammals (Hackman 1963). Hackman (1980) published the latest list of the
Finnish sphaerocerid species. Further species were added in several revisions of Eu-
ZooKeys 441: 325–332 (2014)
doi: 10.3897/zookeys.441.7250
www.zookeys.org
Copyright A. Haarto, J. Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited..
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Antti Haarto & Jere Kahanpää / ZooKeys 441: 325–332 (2014)
326
ropean taxa by Roháček (1982, 1983, 1991). ere has been increasing interest in
sphaerocerids in Finland after the year 2000 (see Haarto and Kahanpää 2013).
e classication follows the world catalogue of Sphaeroceridae (Roháček et al.
2001, Marshall et al. 2011).
Number of species:
World: 1571 species (Pape et al. 2011)
Europe: 257 species
Finland: 118 species
Faunistic knowledge level in Finland: average
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Sphaeroceroidea Macquart, 1835
SPHAEROCERIDAE Macquart, 1835
COPROMYZINAE Stenhammar, 1854
ALLOBORBORUS Duda, 1923
Alloborborus pallifrons (Fallén, 1820)
= avipennis (Haliday, 1836)
BORBORILLUS Duda, 1923
Borborillus uncinatus (Duda, 1923)
Borborillus vitripennis (Meigen, 1830)
= longipennis (Haliday, 1836)
COPROMYZA Fallén, 1810
Copromyza borealis Zetterstedt, 1847
Copromyza equina Fallén, 1820
Copromyza nigrina (Gimmerthal, 1847)
= similis (Collin, 1930)
Copromyza stercoraria (Meigen, 1830)
CRUMOMYIA Macquart, 1835
= Apterina Macquart, 1835
Crumomyia metaria (Meigen, 1830)
Crumomyia gelida Hackman, 1965
Crumomyia glabrifrons (Meigen, 1830)
Crumomyia nigra (Meigen, 1830)
Checklist of the family Sphaeroceridae (Diptera) of Finland 327
Crumomyia nitida (Meigen, 1830)
Crumomyia notabilis (Collin, 1902)
= glacialis misid.
Crumomyia pedestris (Meigen, 1830
Crumomyia pruinosa (Richards, 1932)
= annulus misid.
= rufoannulata misid.
Crumomyia setitibialis (Spuler, 1925)
= freyi (Hackman, 1965)
= annulipes misid.
= roseri misid.
LOTOPHILA Lioy, 1864
Lotophila atra (Meigen, 1830)
NORRBOMIA Papp, 1988
Norrbomia costalis (Zetterstedt, 1847)
Norrbomia fumipennis (Stenhammar, 1855)
Norrbomia hispanica (Duda, 1923)
Norrbomia sordida (Zetterstedt, 1847)
SPHAEROCERINAE Macquart, 1835
ISCHIOLEPTA Lioy, 1864
Ischiolepta crenata (Meigen, 1838)
Ischiolepta denticulata (Meigen, 1830)
= paracrenata (Duda, 1920)
Ischiolepta micropyga (Duda, 1938)
Ischiolepta nitida (Duda, 1920)
= denticulata auct. nec (Meigen, 1830)
Ischiolepta pusilla (Fallén, 1820)
Ischiolepta scabricula (Haliday, 1836)
Ischiolepta vaporariorum (Haliday, 1836)
LOTOBIA Lioy, 1864
Lotobia pallidiventris (Meigen, 1830)
SPHAEROCERA Latreille, 1804
= Cypsela Meigen, 1800 suppr.
Sphaerocera curvipes Latreille, 1805
Sphaerocera monilis Haliday, 1836
LIMOSININAE Frey, 1921
APTEROMYIA Vimmer, 1929
Apteromyia claviventris (Strobl, 1909)
BIFRONSINA Roháček, 1983
Bifronsina bifrons (Stenhammar, 1855)
CHAETOPODELLA Duda, 1920
sg. Chaetopodella Duda, 1920
Chaetopodella scutellaris (Haliday, 1836)
Antti Haarto & Jere Kahanpää / ZooKeys 441: 325–332 (2014)
328
COPROICA Rondani, 1861
Coproica acutangula (Zetterstedt, 1847)
Coproica ferruginata (Stenhammar, 1855)
Coproica hirticula Collin, 1965
Coproica hirtula (Rondani, 1880)
Coproica lugubris (Haliday, 1835)
Coproica pusio (Zetterstedt, 1847)
= pseudolugubris (Duda, 1924)
Coproica vagans (Haliday, 1833)
ELACHISOMA Rondani, 1880
Elachisoma aterrimum (Haliday, 1833)
EULIMOSINA Roháček, 1983
Eulimosina ochripes (Meigen, 1830)
GIGALIMOSINA Roháček, 1983
Gigalimosina aviceps Zetterstedt, 1847
GONIONEURA Rondani, 1880
= Halidayina Duda, 1918
Gonioneura spinipennis (Haliday, 1836)
HERNIOSINA Roháček, 1983
Herniosina bequaerti (Villeneuve, 1917)
LEPTOCERA Olivier, 1813
= Paracollinella Duda, 1924
Leptocera caenosa (Rondani, 1880)
Leptocera nalis (Collin, 1956)
Leptocera fontinalis (Fallén, 1826)
Leptocera nigra Olivier, 1813
LIMOSINA Macquart, 1835
Limosina silvatica (Meigen, 1830)
MINILIMOSINA Roháček, 1983
sg. Minilimosina Roháček, 1983
Minilimosina baculum Marshall, 1985
Minilimosina bicuspis Roháček, 1993
= trogeri misid.
Minilimosina fungicola (Haliday, 1836)
= exigua (Rondani, 1880)
Minilimosina parvula Stenhammar, 1855
Minilimosina tenera Roháček, 1983
sg. Svarciella Roháček, 1983
Minilimosina guestphalica (Duda, 1918)
= v-atrum misid.
Minilimosina v-atrum (Villeneuve, 1917)
= splendens (Duda, 1928)
Minilimosina vitripennis (Zetterstedt, 1847)
Checklist of the family Sphaeroceridae (Diptera) of Finland 329
OPACIFRONS Duda, 1918
Opacifrons coxata (Stenhammar, 1855)
OPALIMOSINA Roháček, 1983
sg. Dentilimosina Roháček, 1983
Opalimosina denticulata (Duda, 1924)
sg. Opalimosina Roháček, 1983
Opalimosina collini (Richards, 1929)
Opalimosina mirabilis (Collin, 1902)
Opalimosina simplex Richards 1929
sg. Pappiella Roháček, 1983
Opalimosina liliputana (Rondani, 1880)
= appendiculata Villeneuve, 1918
PARALIMOSINA Papp, 1973
Paralimosina kaszabi Papp, 1973
PHILOCOPRELLA Richards, 1929
Philocoprella quadrispina (Laurence, 1952)
PHTHITIA Enderlein, 1938
sg. Alimosina Roháček, 1983
Phthitia empirica (Hutton, 1901)
= cadaverina (Duda, 1918)
sg. Collimosina Roháček, 1983
Phthitia spinosa Collin, 1930
sg. Kimosina Roháček, 1983
Phthitia longisetosa (Dahl, 1909)
Phthitia plumosula (Rondani, 1880)
PSEUDOCOLLINELLA Duda, 1924
Pseudocollinella avilabris (Hackman, 1968)
Pseudocollinella humida (Haliday, 1836)
Pseudocollinella septentrionalis (Stenhammar, 1855)
PTEREMIS Rondani, 1856
Pteremis fenestralis (Fallén, 1820)
= nivalis (Haliday, 1833)
= subaptera Frey, 1946
PULLIMOSINA Roháček, 1983
sg. Dahlimosina Roháček, 1983
Pullimosina dahli (Duda, 1918)
sg. Pullimosina Roháček, 1983
Pullimosina heteroneura (Haliday, 1836)
Pullimosina meijerei (Duda, 1918)
Pullimosina moesta (Villeneuve, 1918)
= antennata (Duda, 1918)
Pullimosina pullula (Zetterstedt, 1847)
Pullimosina vulgesta Roháček, 2001
Antti Haarto & Jere Kahanpää / ZooKeys 441: 325–332 (2014)
330
= moesta auct. nec (Villeneuve, 1918)
RACHISPODA Lioy, 1864
Rachispoda anceps (Stenhammar, 1855)
Rachispoda breviceps (Stenhammar, 1855)
Rachispoda fuscipennis (Haliday, 1833)
Rachispoda hostica Villeneuve, 1917
Rachispoda intermedia (Duda, 1918)
Rachispoda limosa (Zetterstedt, 1820)
Rachispoda lugubrina (Zetterstedt, 1847)
Rachispoda lutosa (Stenhammar, 1855)
= palustris (Collin, 1930)
Rachispoda lutosoidea (Duda, 1938)
= lutosa auct. nec (Stenhammar, 1855)
SPELOBIA Spuler, 1924
Spelobia belanica Roháček, 1983
Spelobia cambrica (Richards, 1929)
Spelobia clunipes (Meigen, 1830)
= crassimana (Haliday, 1836)
Spelobia ibrida Roháček, 1983
Spelobia luteilabris (Rondani, 1880)
Spelobia manicata (Richards, 1927)
Spelobia nana (Rondani, 1880)
Spelobia palmata (Richards, 1927)
Spelobia pappi Roháček, 1983
Spelobia parapusio (Dahl, 1909)
Spelobia pseudonivalis (Dahl, 1909)
Spelobia pseudosetaria (Duda, 1918)
= penetralis (Collin, 1925)
Spelobia rulabris (Stenhammar, 1855)
Spelobia talparum (Richards, 1927)
Spelobia ulla Roháček, 1983
SPINILIMOSINA Roháček, 1983
Spinilimosina brevicostata (Duda, 1918)
TELOMERINA Roháček, 1983
Telomerina eburnea Roháček, 1983
Telomerina avipes (Meigen, 1830)
Telomerina pseudoleucoptera (Duda, 1924)
TERRILIMOSINA Roháček, 1983
Terrilimosina racovitzai (Bezzi, 1911)
Terrilimosina schmitzi (Duda, 1918)
THORACOCHAETA Duda, 1918
oracochaeta brachystoma (Stenhammar, 1855)
oracochaeta zosterae (Haliday, 1833)
Checklist of the family Sphaeroceridae (Diptera) of Finland 331
TRACHYOPELLA Duda, 1918
sg. Nudopella Roháček & Marshall, 1986
Trachyopella leucoptera (Haliday, 1836)
sg. Trachyopella Duda, 1918
Trachyopella atomus (Rondani, 1880)
Trachyopella bovilla Collin, 1954
= coprina misid.
Trachyopella lineafrons (Spuler, 1925)
= atomus misid.
Trachyopella melania (Haliday, 1836)
= villeneuvii (Duda, 1924)
Excluded species
Rachispoda cilifera (Rondani, 1880) not found within present borders
Minilimosina unica (Papp, 1973) not found within present borders
= hackmani (Roháček, 1977)
Notes
Spelobia manicata (Richards, 1927) is considered a valid species although Pitkin
(1988) synonymized it with Spelobia clunipes (Meigen, 1830).
References
Haarto A, Kahanpää J (2013) Notes of Finnish Sphaeroceridae (Diptera) with description of
the female of Minilimosina tenera Rohacek, 1983. Entomologica Fennica 24: 228–233.
http://ojs.tsv./index.php/entomolfennica/article/download/9385/6695
Hackman W (1963) Studies on the dipterous fauna in burrows of voles (Microtus, Clethriono-
mys) in Finland. Acta Zoologica Fennica 102: 1–64.
Hackman W (1965) On the genus Copromyza Fall. (Dipt., Sphaeroceridae), with special refer-
ence to the Finnish species. Notulae Entomologicae 45: 33–46.
Hackman W (1980) A Check List of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Marshall SA, Roháček J, Dong H, Buck M (2011) e state of Sphaeroceridae (Diptera: Aca-
lyptratae): a world catalog update covering the years 2000-2010, with new generic syn-
onymy, new combinations, and new distributions. Acta Entomologica Musei Nationalis
Pragae 51(1): 217–298. http://www.aemnp.eu/PDF/51_1/51_1_217.pdf
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed) Animal biodiversity: An outline of higher-level classication and survey of
Antti Haarto & Jere Kahanpää / ZooKeys 441: 325–332 (2014)
332
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Pitkin BR (1988) Lesser dung ies, Diptera: Sphaeroceridae. Handbooks for the Identication of
British Insects, Vol 10, Part 5e. Royal Entomological Society of London, London, 175 pp.
Roháček J (1982) Revision of the subgenus Leptocera (s.str.) of Europe. Entomologische Ab-
handlungen Staatliches Museum für Tierkunde in Dresden 46(1): 1–44.
Roháček J (1983) A monograph and re-classication of the previous genus Limosina Macquart
(Diptera, Sphaeroceridae) of Europe. Part II. Beiträge zur Entomologie, Berlin 33: 3–195.
Roháček J (1991) A monograph of Leptocera (Rachispoda Lioy) of the West Palaearctic area
(Diptera, Sphaeroceridae). Časopis Slezského zemského Muzea, Opava (A) 40: 97–288.
Roháček J, Marshall SA, Norrbom AL, Buck M, Quiros DI, Smith I (2001) World Catalog
of Sphaeroceridae. Slezské Zemské Muzeum, Opava, 300 pp. ISBN 808622421X. http://
www.uoguelph.ca/debu/catalog.htm
Checklist of the Braulidae, Camillidae, Diastatidae and Drosophilidae of Finland... 333
Checklist of the Braulidae, Camillidae, Diastatidae and
Drosophilidae of Finland (Diptera, Ephydroidea)
Jere Kahanpää1
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 5 February 2014|Accepted 2 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J (2014) Checklist of the Braulidae, Camillidae, Diastatidae and Drosophilidae of Finland (Diptera,
Ephydroidea). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 333–338. doi:
10.3897/zookeys.441.7057
Abstract
A checklist of the Braulidae (1 species), Camillidae (3 spp.), Diastatidae (10 spp.) and Drosophilidae (67
spp.) of Finland is presented. Campichoetinae is included as a subfamily of Diastatidae.
Keywords
Checklist, Finland, Diptera, biodiversity, faunistics
Introduction
e Ephydroidea is now recognized as a well-supported, probably monophyletic clade
of acalyptrate ies (Wiegmann et al. 2011), although the correct placement of some
of the included families – especially the very unsual bee louses of family Braulidae–
is still being discussed. e relationships of the families within the superfamily are
less understood and the families are listed alphabetically in the checklist below. Seven
or eight families are currently included in Ephydroidea. Five of them are found in
Finland. Four families are covered in this paper; the fth (Ephydridae itself) has its
own treatment in this volume of ZooKeys. e families missing from Finland are the
Curtonotidae, Cryptochetidae, and Nannodastiidae with uncertain anities (Papp
ZooKeys 441: 333–338 (2014)
doi: 10.3897/zookeys.441.7057
www.zookeys.org
Copyright Jere Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää / ZooKeys 441: 333–338 (2014)
334
and Mathis 2001). Griths (1972) proposed a new family, Campichoetidae, for
Campichoeta Macquart, 1835. We follow the world catalogue of Diastatidae (Mathis
and Barraclough 2011) and keep Campichoetinae as a subfamily of Diastatidae.
World catalogues have been recently published for Diastatidae (Mathis and Barr-
aclough 2011) and Drosophilidae (Brake and Bächli 2008). e North European
Drosophilidae were treated in detail by Bächli et al. (2004) and the review of Finnish
Drosophila by Hackman (1954) is now obsolete. e Finnish species of Braulidae
and Camillidae were last listed by Hackman (1980); the diastatids were treated by
Chandler (1987).
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Ephydroidea Zetterstedt, 1837
BRAULIDAE Egger, 1853
BRAULA Nitzsch, 1818
Braula coeca Nitzsch, 1818
CAMILLIDAE Frey, 1921
CAMILLA Haliday, 1838
Camilla atrimana Strobl, 1910
= atripes Duda, 1934
= acutipennis misid.
Camilla avicauda Duda, 1922
= glabrata Collin, 1956
Camilla glabra (Fallén, 1823)
= subfuscipes Collin, 1933
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Braulidae 7 3 1 poor
Camillidae 40 8 3 average
Diastatidae 49 (Mathis and Barraclough 2011) 16 10 good
Drosophilidae 4003 121 67 good
Checklist of the Braulidae, Camillidae, Diastatidae and Drosophilidae of Finland... 335
DIASTATIDAE Hendel, 1917
CAMPICHOETINAE Griths, 1972
CAMPICHOETA Macquart, 1835
Campichoeta griseola (Zetterstedt, 1855)
Campichoeta obscuripennis (Meigen, 1830)
DIASTATINAE Hendel, 1917
DIASTATA Meigen, 1830
Diastata adusta Meigen, 1830
= unipunctata Zetterstedt, 1847
Diastata boreonigra Chandler, 1987
Diastata costata Meigen, 1830
Diastata avicosta Chandler, 1987
Diastata fuscula (Fallén, 1820)
Diastata nebulosa (Fallén, 1823)
Diastata ornata Meigen, 1830
Diastata vagans Loew, 1864
DROSOPHILIDAE Rondani, 1856
STEGANINAE Hendel, 1917
AMIOTA Loew, 1862
sg. Amiota Loew, 1862
Amiota albilabris (Roth in Zetterstedt, 1860)
Amiota alboguttata (Wahlberg, 1839)
Amiota rufescens (Oldenberg, 1914)
Amiota subtusradiata Duda, 1934
CACOXENUS Loew, 1858
sg. Paracacoxenus Hardy & Wheeler, 1960
Cacoxenus argyreator Frey, 1932
LEUCOPHENGA Mik, 1886
sg. Neoleucophenga Oldenberg, 1915
Leucophenga quinquemaculata Strobl, 1893
PHORTICA Schiner, 1862
sg. Phortica Schiner, 1862
Phortica variegata Fallén, 1823
STEGANA Meigen, 1830
sg. Stegana Meigen, 1830
Stegana furta (Linnaeus, 1766)
= curvipennis (Fallén, 1823)
sg. Steganina Wheeler, 1960
Stegana baechlii Laštovka & Máca, 1982
Stegana coleoptrata (Scopoli, 1763)
Stegana hypoleuca Meigen, 1830
Stegana longibula Takada, 1968
Jere Kahanpää / ZooKeys 441: 333–338 (2014)
336
Stegana mehadiae Duda, 1924
Stegana nigrithorax Strobl, 1898
Stegana similis Laštovka & Máca, 1982
DROSOPHILINAE Rondani, 1856
CHYMOMYZA Czerny, 1903
Chymomyza amoena (Loew, 1862)
Chymomyza caudatula Oldenberg, 1914
Chymomyza costata (Zetterstedt, 1838)
Chymomyza distincta (Egger, 1862)
Chymomyza fuscimana (Zetterstedt, 1838)
DROSOPHILA Fallén, 1823
sg. Dorsilopha Sturtevant, 1942
Drosophila busckii Coquillett, 1901
sg. Drosophila Fallén, 1823
Drosophila ezoana Takada & Okada, 1957
Drosophila funebris (Fabricius, 1787)
Drosophila histrio Meigen, 1830
Drosophila hydei Sturtevant, 1921
= repleta misid.
Drosophila immigrans Sturtevant, 1921
= tripunctata Becker, 1908 preocc.
Drosophila limbata von Roser, 1840
Drosophila littoralis Meigen, 1830
Drosophila lummei Hackman, 1972
Drosophila montana Stone, Grien & Patterson, 1941
= ovivororum Lakovaara & Hackman, 1973
Drosophila phalerata Meigen, 1830
Drosophila picta Zetterstedt, 1847
Drosophila repleta Wollaston, 1858
Drosophila subarctica Hackman, 1969
Drosophila testacea von Roser, 1840
Drosophila transversa Fallén, 1823
Drosophila vireni Bächli, Vilela & Haring, 2002
sg. Sophophora Sturtevant, 1942
Drosophila alpina Burla, 1948
Drosophila ambigua Pomini, 1940
Drosophila bifasciata Pomini, 1940
Drosophila eskoi Lakovaara & Lankinen, 1974
Drosophila ingrica Hackman, 1957
Drosophila melanogaster Meigen, 1830
Drosophila obscura Fallén, 1823
Drosophila simulans Sturtevant, 1919
Drosophila subobscura Collin, 1936
Checklist of the Braulidae, Camillidae, Diastatidae and Drosophilidae of Finland... 337
Drosophila subsilvestris Hardy & Kaneshiro, 1968
= silvestris Basden, 1954 preocc.
Drosophila tristis Fallén, 1823
HIRTODROSOPHILA Duda, 1924
Hirtodrosophila cameraria (Haliday, 1833)
Hirtodrosophila confusa (Staeger, 1844)
Hirtodrosophila lundstroemi (Duda, 1935)
Hirtodrosophila oldenbergi (Duda, 1924)
Hirtodrosophila trivittata (Strobl, 1893)
LORDIPHOSA Basden, 1961
Lordiphosa fenestrarum (Fallén, 1823)
Lordiphosa nigricolor (Strobl, 1898)
MICRODROSOPHILA Malloch, 1921
sg. Microdrosophila Malloch, 1921
Microdrosophila congesta (Zetterstedt, 1847)
sg. Oxystyloptera Duda, 1924
Microdrosophila zetterstedti Wheeler, 1959
= nigriventris (Zetterstedt, 1847) preocc.
SCAPTODROSOPHILA Duda, 1923
Scaptodrosophila deexa (Duda, 1924)
= guyenoti (Burla, 1948)
SCAPTOMYZA Hardy, 1849
sg. Hemiscaptomyza Hackman, 1959
Scaptomyza trochanterata Collin, 1953
Scaptomyza unipunctum (Zetterstedt, 1847)
sg. Parascaptomyza Duda, 1924
Scaptomyza pallida (Zetterstedt, 1847)
= disticha (Duda, 1921)
sg. Scaptomyza Hardy, 1849
Scaptomyza consimilis Hackman, 1955
Scaptomyza ava (Fallén, 1823)
= aveola (Meigen, 1830)
= apicalis Hardy, 1849
Scaptomyza graminum (Fallén, 1823)
Scaptomyza griseola (Zetterstedt, 1847)
Scaptomyza montana Wheeler, 1949
Scaptomyza teinoptera Hackman, 1955
Excluded species
Stegana strobli Mik, 1898 misidentication
Lordiphosa hexasticha (Papp, 1971) not found within present borders
Jere Kahanpää / ZooKeys 441: 333–338 (2014)
338
References
Brake I, Bächli G (2008) World catalogue of insects. Vol. 9: Drosophilidae (Diptera). Apollo
Books, Stenstrup, 412 pp.
Bächli G, Vilela C, Escher SA, Saura A (2004) e Drosophilidae (Diptera) of Fennoscandia
and Denmark. Fauna Entomologica Scandinavica 39, Brill NV, Leiden & Boston, 362 pp.
Chandler PJ (1987) e families Diastatidae and Campichoetidae (Diptera, Drosophiloidea)
with a revision of Palaearctic and Nepalese species of Diastata Meigen. Entomologica
Scandinavica 18(1): 1–50. doi: 10.1163/187631287X00016
Griths GCD (1972) e phylogenetic classication of Diptera Cyclorrhapha, with special
references to the structure of the male postabdomen. Series entomologica 8. W. Junk,
N.V., Haag, 340 pp.
Hackman W (1954) Die Drosophila-Arten Finnlands. Notulae entomologicae 34: 130–139.
Hackman W (1980) A check list of the Finnish Diptera. Notulae entomologicae 60: 17–48,
117–162.
Mathis WN, Barraclough DA (2011) World catalog and conspectus on the family Diastati-
dae (Diptera: Schizophora). In: Brake I, ompson FC (Eds) Contributions to the Sys-
tema Dipterorum (Insecta: Diptera). Myia vol. 12. Pensoft Publishers & North America
Dipterists Society, Soa, Moscow & Washington D.C., 235–266. http://hdl.handle.
net/10088/18925
Pape T, Blagoderov V, Mostovski MB (2011) Order Diptera Linnaeus, 1758. In: Zhang
Z-Q (Ed.) Animal biodiversity: An outline of higher-level classication and survey of
taxonomic richness. Zootaxa 3148: 222–229. http://www.mapress.com/zootaxa/2011/f/
zt03148p229.pdf
Papp L, Mathis WN (2001) A review of the family Nannodastiidae (Diptera). Proceedings of
the Entomological Society of Washington 103: 337–348. http://biodiversitylibrary.org/
page/16279583
Wiegmann BM, Trautwein MD, Winkler IS, Barr NB, Kim J-W, Lambkin C, Bertone MA,
Cassel BK, Bayless KM, Heimberg AM, Wheeler BM, Peterson KJ, Pape T, Sinclair BJ,
Skevington JH, Blagoderov V, Caravas J, Kutty SN, Schmidt-Ott U, Kampmeier GE,
ompson FC, Grimaldi DA, Beckenbach AT, Courtney GW, Friedrich M, Meier R,
Yeates DK (2011) Episodic radiations in the y tree of life. Proceedings of the National
Academy of Sciences of the United States of America 108(14): 5690–5695. doi: 10.1073/
pnas.1012675108
Checklist of the family Ephydridae of Finland (Insecta, Diptera)339
Checklist of the family Ephydridae of Finland
(Insecta, Diptera)
Tadeusz Zatwarnicki1, Jere Kahanpää2
1 Department of Biosystematics, Opole University, ul. Oleska 22, 45-052 Opole, Poland 2 Finnish Museum of
Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
Corresponding author: Tadeusz Zatwarnicki (zatwar@uni.opole.pl)
Academic editor: J. Salmela|Received 7 March 2014|Accepted 24 April 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Zatwarnicki T, Kahanpää J (2014) Checklist of the family Ephydridae of Finland (Insecta, Diptera). In:
Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 339–346. doi: 10.3897/zookeys.441.7448
Abstract
A checklist of 112 species of shore ies (Ephydridae, Diptera) recorded from Finland is presented. Com-
paring this to the list of Hackman (1980), 52 changes are made: 25 species are added (all but one recorded
after 1980), 18 misidentications are deleted, 5 junior synonyms are replaced and 5 updated generic
combinations are given.
Keywords
Checklist, Finland, Diptera, Ephydridae
Introduction
Shore ies (Ephydridae) are dark coloured, dull to shining, 1–6 mm long ies. Most
species develop in aquatic or semi-aquatic conditions, feeding of algae, bacteria, yeasts
or even decaying animal matter and excrements; or they are phytophagous and mining
in leaves or stems. e group is divided into 5 subfamilies and 19 tribes (Zatwarnicki
1992), of which 15 could potentially be found in Finland.
Hackman (1980) originally listed 105 species. Zatwarnicki (1997) corrected the
synonymy and generic placement of some species. Compared to Hackman (1980)
ZooKeys 441: 339–346 (2014)
doi: 10.3897/zookeys.441.7448
www.zookeys.org
Copyright T. Zatwarnicki, J. Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY
4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
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A peer-reviewed open-access journal
T. Zatwarnicki & J. Kahanpää / ZooKeys 441: 339–346 (2014)
340
the present list adds 25 species and removes 18 misidentied species (mostly Hydrel-
lia). Five junior synonyms in Hydrellia, Philygria, Scatella and Scatophila are replaced;
four species of Ditrichophora are combined with Gymnoclasiopa and one species of
Lamproscatella with Haloscatella. Two species are endemic for Finland: Psilopa nitidi-
facies Frey, 1930 (note that this was omitted by Hackman 1980), and Gymnoclasiopa
psilopina (Frey, 1933).
Nomenclature follows the current version of the Ephydridae section of Fauna Eu-
ropaea (Zatwarnicki 2013). e subfamilies and the tribes are ordered systematically;
genera within tribes and species within genera are listed alphabetically.
Number of species:
World: 1927 species and 63 nomina dubia (as of February 2014)
Europe: 340 species and 3 subspecies (Zatwarnicki 2013)
Finland: 112 species
Faunistic knowledge level in Finland: average
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Acalyptratae Macquart, 1835
superfamily Ephydroidea Zetterstedt, 1837
EPHYDRIDAE Zetterstedt, 1837
DISCOMYZINAE Acloque, 1897
Discomyzini Acloque, 1897
DISCOMYZA Meigen, 1830
Discomyza incurva (Fallén, 1823)
Psilopini Cresson, 1925
PSILOPA Fallén, 1823
Psilopa compta (Meigen, 1830)
Psilopa leucostoma (Meigen, 1830)
Psilopa marginella Fallén, 1823
Psilopa nigritella Stenhammar, 1844
Psilopa nitidifacies Frey, 1958
Psilopa nitidula (Fallén, 1813)
Psilopa polita (Macquart, 1835)
TRIMERINA Macquart, 1835
Trimerina madizans (Fallén, 1813)
Checklist of the family Ephydridae of Finland (Insecta, Diptera)341
Trimerina microchaeta Hendel, 1932
= indistincta Krivosheina 2004
HYDRELLIINAE Robineau-Desvoidy, 1830
Atissini Cresson, 1942
ATISSA Haliday, 1839
Atissa limosina Becker, 1896
Atissa pygmaea Haliday, 1839
Hydrelliini Robineau-Desvoidy, 1830
HYDRELLIA Robineau-Desvoidy, 1830
Hydrellia albifrons Fallén, 1813
Hydrellia albilabris (Meigen, 1830)
Hydrellia cardamines Haliday, 1839
= baltica Frey, 1930
Hydrellia cochleariae Haliday, 1839
= avicornis misid.
Hydrellia aviceps (Meigen, 1830)
= discors Collin, 1966
= lapponica misid.
Hydrellia fulviceps (Stenhammar, 1844)
= chrysostoma misid.
Hydrellia fusca (Stenhammar, 1844)
Hydrellia griseola (Fallén, 1813)
= chrysostoma (Meigen, 1830)
Hydrellia laticeps (Stenhammar, 1844)
Hydrellia mutata (Zetterstedt, 1846)
Hydrellia obscura (Meigen, 1830)
= fascitibia misid.
Hydrellia pilitarsis (Stenhammar, 1844)
Hydrellia subalbiceps Collin, 1966
Hydrellia tarsata Haliday, 1839
Hydrellia thoracica Haliday, 1839
= modesta misid.
Notiphilini Bigot, 1853
DICHAETA Meigen, 1830
Dichaeta caudata (Fallén, 1813)
= brevicauda Loew, 1860
NOTIPHILA Fallén, 1810
sg. Agrolimna Cresson, 1917
Notiphila uliginosa Haliday, 1839
sg. Notiphila Fallén, 1810
Notiphila annulipes Stenhammar, 1844
Notiphila aquatica Becker, 1896
Notiphila brunipes (Robineau-Desvoidy, 1830)
T. Zatwarnicki & J. Kahanpää / ZooKeys 441: 339–346 (2014)
342
= brunnipes emend.
Notiphila cinerea Fallén, 1813
Notiphila dorsata Stenhammar, 1844
Notiphila graecula Becker, 1926
= maculata misid.
Notiphila major Stenhammar, 1844
Notiphila pollinosa Krivosheina, 1998
Notiphila riparia Meigen, 1830
GYMNOMYZINAE Latreille, 1829
Gymnomyzini Latreille, 1829
ATHYROGLOSSA Loew, 1860
sg. Athyroglossa Loew, 1860
Athyroglossa glabra (Meigen, 1830)
MOSILLUS Latreille, 1804
Mosillus subsultans (Fabricius, 1794)
Lipochaetini Becker, 1896
GLENANTHE Haliday, 1839
Glenanthe fuscinervis Becker, 1896
= ripicola auct. nec Haliday, 1839
Hecamedini Mathis, 1991
ALLOTRICHOMA Becker, 1896
Allotrichoma bezzii Becker, 1896
= laterale misid.
Ochtherini Dahl, 1959
OCHTHERA Latreille, 1802
Ochthera manicata (Fabricius, 1794)
Ochthera mantis (De Geer, 1776)
Discocerinini Cresson, 1925
DISCOCERINA Macquart, 1835
Discocerina obscurella (Fallén, 1813)
DITRICHOPHORA Cresson, 1924
Ditrichophora calceata (Meigen, 1830)
Ditrichophora fuscella (Stenhammar, 1844)
GYMNOCLASIOPA Hendel, 1930
Gymnoclasiopa aurivillii (Becker, 1896)
Gymnoclasiopa bohemanni (Becker, 1896)
Gymnoclasiopa cinerella (Stenhammar, 1844)
= pulchella misid.
Gymnoclasiopa nigerrima (Strobl, 1893)
Gymnoclasiopa psilopina (Frey, 1933)
HECAMEDOIDES Hendel, 1917
Hecamedoides glaucellus (Stenhammar, 1844)
Checklist of the family Ephydridae of Finland (Insecta, Diptera)343
Hecamedoides unispinosus (Collin, 1943)
POLYTRICHOPHORA Cresson, 1924
Polytrichophora duplosetosa (Becker, 1896)
ILYTHEINAE Cresson, 1943
Hyadinini Philips et al. in Cresson, 1949
AXYSTA Haliday, 1839
Axysta cesta (Haliday, 1833)
HYADINA Haliday, 1839
Hyadina guttata (Fallén, 1813)
Hyadina humeralis Becker, 1896
Hyadina rupes (Meigen, 1830)
= nitida (Macquart, 1835)
Hyadina scutellata (Haliday, 1839)
LYTOGASTER Becker, 1896
Lytogaster abdominalis (Stenhammar, 1844)
NOSTIMA Coquillett, 1900
Nostima picta (Fallén, 1813)
PELINA Haliday, 1839
Pelina aenea (Fallén, 1813)
Pelina aenescens (Stenhammar, 1844)
PHILYGRIA Stenhammar, 1844
Philygria femorata (Stenhammar, 1844)
= posticata misid.
Philygria avipes (Fallén, 1823)
Philygria interstincta (Fallén, 1813)
= maculipennis (Robineau-Desvoidy, 1830)
= sexmaculata Becker, 1896
Philygria obtecta Becker, 1896
Philygria vittipennis (Zetterstedt, 1838)
= nigricauda Stenhammar, 1844
= trilineata De Meijere, 1907
Ilytheini Cresson, 1943
ILYTHEA Haliday, 1839
Ilythea spilota (Haliday in Curtis, 1832)
EPHYDRINAE Zetterstedt, 1837
Parydrini Wirth & Stone, 1956
EUTAENIONOTUM Oldenberg, 1923
Eutaenionotum guttipenne (Stenhammar, 1843)
PARYDRA Stenhammar, 1844
= Napaea Robineau-Desvoidy, 1830 preocc.
sg. Chaetoapnaea Hendel, 1930
Parydra arctica Clausen, 1971
T. Zatwarnicki & J. Kahanpää / ZooKeys 441: 339–346 (2014)
344
Parydra fossarum (Haliday, 1833)
Parydra mitis (Cresson, 1930)
Parydra nigritarsis Strobl, 1893
Parydra pusilla (Meigen, 1830)
Parydra quadripunctata (Meigen, 1830)
sg. Parydra Stenhammar, 1844
Parydra aquila (Fallén, 1813)
Parydra coarctata (Fallén, 1813)
Parydra nubecula (Becker, 1896)
Ephydrini Zetterstedt, 1837
CALOCOENIA Mathis, 1975
sg. Leptocoenia Mathis, 1975
Calocoenia paurosoma (Sturtevant & Wheeler, 1954)
COENIA Robineau-Desvoidy, 1830
Coenia curvicauda (Meigen, 1830)
Coenia palustris (Fallén, 1823)
Coenia vulgata Krivosheina, 2001
EPHYDRA Fallén, 1810
Ephydra macellaria Egger, 1862
Ephydra riparia Fallén, 1813
Ephydra scholtzi Becker, 1896
= krogerusi Frey, 1930
PARACOENIA Cresson, 1935
Paracoenia fumosa (Stenhammar, 1844)
SETACERA Cresson, 1930
Setacera aurata (Stenhammar, 1844)
Setacera micans (Haliday, 1833)
Scatellini Wirth & Stone, 1956
HALOSCATELLA Mathis, 1979
Haloscatella dichaeta (Loew, 1860)
LAMPROSCATELLA Hendel, 1917
Lamproscatella sibilans (Haliday, 1833)
LIMNELLIA Malloch, 1925
Limnellia fallax (Czerny, 1903)
Limnellia quadrata (Fallén, 1813)
Limnellia stenhammari (Zetterstedt, 1846)
PHILOTELMA Becker, 1896
Philotelma defectum (Haliday, 1833)
Philotelma nigripenne (Meigen, 1930)
SCATELLA Robineau-Desvoidy, 1830
sg. Neoscatella Malloch, 1933
Scatella crassicosta Becker, 1896
Checklist of the family Ephydridae of Finland (Insecta, Diptera)345
Scatella silacea Loew, 1860
Scatella subguttata (Meigen, 1830)
sg. Scatella Robineau-Desvoidy, 1830
Scatella obsoleta Loew, 1861
= callosicosta Bezzi, 1895
Scatella paludum (Meigen, 1830)
Scatella stagnalis (Fallén, 1813)
Scatella tenuicosta Collin, 1930
SCATOPHILA Becker, 1896
Scatophila caviceps (Stenhammar, 1844)
Scatophila contaminata (Stenhammar, 1844)
= halterata Becker, 1896
Scatophila cribrata (Stenhammar, 1844)
Scatophila despecta (Haliday, 1839)
= hamifera Becker, 1896
Scatophila iowana Wheeler, 1961
Scatophila mesogramma (Loew, 1869)
Scatophila noctula (Meigen, 1830)
= avitarsis (Zetterstedt, 1846)
= laevigata (Loew, 1860)
= silesiaca Becker, 1896
Scatophila quadriguttata (Meigen, 1830)
= variegata (Loew, 1860)
Excluded species
Gymnoclasiopa aurifacies (Strobl, 1893) misidentied
Gymnoclasiopa plumosa (Fallén, 1923) misidentied
Hyadina nigricornis Frey, 1930 not found within present borders
Hydrellia argyrogenis Becker, 1896 misidentied
Hydrellia concolor (Stenhammar, 1844) misidentied
Hydrellia avicornis (Fallén, 1823) misidentied
Hydrellia incana (Stenhammar, 1844) misidentied
Hydrellia meigeni Zatwarnicki, 1988 misidentied
= albiceps (Meigen, 1830) preocc.
Hydrellia nympheae (Stenhammar, 1844) misidentied
Hydrellia tibialis Cresson, 1917 not found within present borders
= diadema Frey, 1930
Notiphila stagnicola (Robineau-Desvoidy, 1830) misidentied
Psilopa pulicaria (Haliday, 1839) misidentied
Scatella lutosa Haliday, 1833 misidentied
T. Zatwarnicki & J. Kahanpää / ZooKeys 441: 339–346 (2014)
346
Notes
Gymnoclasiopa aurivillii (Becker, 1896) is probably a junior synonym of Gymnoclasiopa
nigerrima (Strobl, 1893).
Hydrellia. At least two additional species occur in Finland but their proper names are
still under investigation.
References
Hackman W (1980) A check list of the Finnish Diptera II. Cyclorrhapha. Notulae entomo-
logicae 60: 117–162.
Zatwarnicki T (1992) A new classication of Ephydridae based on phylogenetic reconstruction
(Diptera Cyclorrhapha). Genus 3(2): 65–119.
Zatwarnicki T (1997) Diptera Ephydridae, Shore Flies. In: Nilsson A (Ed) e Aquatic Insects
of North Europe, Vol 2. Apollo Books, Stenstrup, 383–399.
Zatwarnicki T (2013) Family Ephydridae (Animalia: Eumetazoa: Arthropoda: Hexapoda: In-
secta: Diptera: Brachycera). In: Pape T, Beuk P (Eds) Fauna Europaea: Diptera, version
2.6.2. http://www.faunaeur.org
Checklist of the families Scathophagidae, Fanniidae and Muscidae of Finland (Insecta, Diptera)347
Checklist of the families Scathophagidae, Fanniidae
and Muscidae of Finland (Insecta, Diptera)
Jere Kahanpää1, Antti Haarto2
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
2 Zoological Museum, Section of Biodiversity and Environmental Science, Department of Biology, University
of Turku, FI–20014 Turku, Finland
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.)
Academic editor: J. Salmela|Received 5 February 2014|Accepted 19 May 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Kahanpää J, Haarto A (2014) Checklist of the families Scathophagidae, Fanniidae and Muscidae of Finland
(Insecta, Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 347–367. doi:
10.3897/zookeys.441.7142
Abstract
A revised checklist of the Scathophagidae, Fanniidae and Muscidae recorded from Finland is presented.
Phaonia amicula Villeneuve, 1922 is noted from Finland for the rst time.
Keywords
Species list, Finland, Diptera, biodiversity, faunistics
Introduction
Four families make up the traditional superfamily Muscoidea: Scathophagidae, An-
thomyiidae, Fanniidae and Muscidae. e monophyly of the superfamily has been
strongly questioned (Nirmala et al. 2001, Kutty et al. 2010) on the basis of DNA
sequence analyses.
ree of the four families of Muscoidea are treated in this paper, the fourth (An-
thomyiidae) is covered in a separate paper in this issue of ZooKeys. All three are im-
portant components of the Finnish y fauna as can be seen from the large number of
species involved and the fact that the species recorded from the country represent up
to one fth of the world fauna in the case of Scathophagidae.
ZooKeys 441: 347–367 (2014)
doi: 10.3897/zookeys.441.7142
www.zookeys.org
Copyright J. Kahanpää, A. Haarto. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jere Kahanpää & Antti Haarto / ZooKeys 441: 347–367 (2014)
348
Unfortunately the muscoid ies have been poorly collected and studied in Finland
in comparison with many other Diptera families. Lauri Tiensuu concentrated on these
families during the 1930s (1935, 1938, 1939) and Walter Hackman (1956) reviewed
the Finnish Scathophagidae. During the last ten years, both the fanniids (Kahanpää
and Haarto 2013) and the muscids (Kahanpää 2013) have attracted some attention in
the country. e number of species known from the country has increased markedly
since the year 2000.
e Finnish muscoid species were last listed by Hackman (1980).
Checklist
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Calyptratae Robineau-Desvoidy, 1830
superfamily Muscoidea Latreille, 1802
SCATHOPHAGIDAE Robineau-Desvoidy, 1830
DELININAE Séguy, 1952
DELINA Robineau-Desvoidy, 1830
Delina nigrita (Fallén, 1819)
LEPTOPA Zetterstedt, 1838
Leptopa liformis Zetterstedt, 1838
MICROPSELAPHA Becker, 1894
Micropselapha liformis (Zetterstedt, 1846)
PARALLELOMMA Strobl, 1894
= Chylizosoma Hendel, 1924
Parallelomma medium Becker, 1894
? Parallelomma paridis Hering, 1923 (see Notes)
Parallelomma sellatum (Hackman, 1956)
Parallelomma vittatum (Meigen, 1826)
PHROSIA Robineau-Desvoidy, 1830
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Scathophagidae 419 158 85–86 good
Fanniidae 359 83 61 average
Muscidae 5218 572 307–309 average
Checklist of the families Scathophagidae, Fanniidae and Muscidae of Finland (Insecta, Diptera)349
Phrosia albilabris (Fabricius, 1805)
SCATHOPHAGINAE Robineau-Desvoidy, 1830
ACANTHOCNEMA Becker, 1894
sg. Clinoceroides Hendel, 1917
Acanthocnema glaucescens (Loew, 1864)
= nigripes Ringdahl, 1936
ACEROCNEMA Becker, 1894
Acerocnema macrocera (Meigen, 1826)
= tie Becker, 1894
= pokornyi Becker, 1894
ALLOMYELLA Malloch, 1923
Allomyella albipennis (Zetterstedt, 1838)
Allomyella frigida (Holmgren, 1883)
= portenkoi (Stackelberg, 1952) misid.
BOSTRICHOPYGA Becker, 1894
Bostrichopyga crassipes (Zetterstedt, 1838)
CHAETOSA Coquillett, 1898
Chaetosa punctipes (Meigen, 1826)
CLEIGASTRA Macquart, 1835
= Cnemopogon Rondani, 1856
Cleigastra apicalis (Meigen, 1826)
CONISTERNUM Strobl, 1894
= Coniosternum Becker, 1894
Conisternum lapponicum (Ringdahl, 1920)
Conisternum obscurum (Fallén, 1819)
Conisternum tinctinerve (Becker, 1894)
CORDILURA Fallén, 1810
= Cordylura Meigen, 1826 emend.
sg. Cordilura Fallén, 1810
Cordilura aberrans Becker, 1894
Cordilura aemula Collin, 1958
Cordilura atrata Zetterstedt, 1846
Cordilura ciliata Meigen, 1826
Cordilura picipes Meigen, 1826
Cordilura picticornis Loew, 1864
Cordilura proboscidea Zetterstedt, 1838
Cordilura pubera (Linnaeus, 1758)
Cordilura pudica Meigen, 1826
Cordilura rumana Meigen, 1826
Cordilura socialis (Becker, 1894)
= freyi Hackman, 1956
sg. Cordilurina James, 1955
= Parallelomma Becker, 1894 preocc.
Jere Kahanpää & Antti Haarto / ZooKeys 441: 347–367 (2014)
350
Cordilura albipes Fallén, 1819
Cordilura fuscipes Zetterstedt, 1838
sg. Scoliaphleps Becker, 1894
Cordilura ustulata Zetterstedt, 1838
= melanacra Loew, 1873
COSMETOPUS Becker, 1894
Cosmetopus dentimana (Zetterstedt, 1838)
= fulvipes (Zetterstedt, 1838)
Cosmetopus longa (Walker, 1849)
= bergrothi Becker, 1900 in part
= fulvipes auct. nec (Zetterstedt, 1838)
Cosmetopus ringdahli Andersson, 1974
= bergrothi Becker, 1900 in part
ERNONEURA Becker, 1894
Ernoneura argus (Zetterstedt, 1838)
GONARCTICUS Becker, 1894
Gonarcticus abdominalis (Zetterstedt, 1846)
GONATHERUS Rondani, 1856
Gonatherus planiceps (Fallén, 1826)
GIMNOMERA Rondani, 1866
= Cochliarium Becker, 1894
Gimnomera albipila (Zetterstedt, 1846)
Gimnomera cuneiventris (Zetterstedt, 1846)
Gimnomera dorsata (Zetterstedt, 1838)
Gimnomera hirta Hendel, 1930
Gimnomera tarsea (Fallén, 1819)
HEXAMITOCERA Becker, 1894
Hexamitocera loxoceratum (Fallén, 1826)
HYDROMYZA Fallén, 1823
Hydromyza livens (Fabricius, 1794)
MEGAPHTHALMA Becker, 1894
Megaphthalma pallida (Fallén, 1819)
MEGAPHTHALMOIDES Ringdahl, 1936
Megaphthalmoides unilineatus (Zetterstedt, 1838)
MICROPROSOPA Strobl, 1894
Microprosopa haemorrhoidalis (Meigen, 1826)
Microprosopa lacteipennis Ringdahl, 1920
Microprosopa lineata (Zetterstedt, 1838)
Microprosopa pallidicauda (Zetterstedt, 1838)
NANNA Strobl, 1894
= Amaurosoma Becker, 1894
Nanna armillata (Zetterstedt, 1846)
Nanna articulata (Becker, 1894)
Checklist of the families Scathophagidae, Fanniidae and Muscidae of Finland (Insecta, Diptera)351
Nanna bispinosa (Malloch, 1920)
Nanna brevifrons (Zetterstedt, 1838)
Nanna fasciata (Meigen, 1826)
Nanna avipes (Fallén, 1819)
= minuta (Becker, 1894)
?= multisetosa (Hackman, 1956)
Nanna inermis (Becker, 1894)
Nanna leucostoma (Zetterstedt, 1846)
Nanna tibiella (Zetterstedt, 1838)
= nigripes (Zetterstedt, 1846)
NORELLIA Robineau-Desvoidy, 1830
Norellia tipularia (Fabricius, 1794) (see Notes)
NORELLISOMA Hendel, 1910
Norellisoma lituratum (Wiedemann, 1826)
Norellisoma spinimanum (Fallén, 1819)
OKENIELLA Hendel, 1907
Okeniella caudata (Zetterstedt, 1838)
Okeniella dasyprocta (Loew, 1864)
ORTHACHETA Becker, 1894
Orthacheta pilosa (Zetterstedt, 1838)
PLEUROCHAETELLA Vockeroth, 1965
Pleurochaetella simplicipes (Becker, 1900)
POGONOTA Zetterstedt, 1860
sg. Pogonota Zetterstedt, 1860
Pogonota barbata (Zetterstedt, 1838)
sg. Lasioscelus Becker, 1894
Pogonota immunda (Zetterstedt, 1838)
= clavatus (Zetterstedt, 1846)
Pogonota sahlbergi Becker, 1900
SCATHOPHAGAMeigen, 1803
= Scatophaga Frabricius, 1805 emend.
= Scopeuma Meigen, 1800 suppr.
= Scatomyza Fallén, 1810
Scathophaga apicalis Curtis in Ross, 1835
Scathophaga furcata (Say, 1823)
Scathophaga incola Becker, 1900
Scathophaga inquinata (Meigen, 1826)
= analis Meigen, 1826
Scathophaga litorea (Fallén, 1819)
Scathophaga lutaria (Fabricius, 1794)
Scathophaga obscurinervis Becker, 1900
Scathophaga pictipennis Oldenberg, 1923
Scathophaga scybalaria (Linnaeus, 1758)
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Scathophaga stercoraria (Linnaeus, 1758)
Scathophaga suilla (Fabricius, 1794)
SPAZIPHORA Rondani, 1856
Spaziphora hydromyzina (Fallén, 1819)
= fascipes (Becker, 1894)
STAEGERIA Rondani, 1856
Staegeria kunzei (Zetterstedt, 1821)
TRICHOPALPUS Rondani, 1856
Trichopalpus fraternus (Meigen, 1826)
Trichopalpus nigribasis Curran, 1927
= pilirostris (Ringdahl, 1936)
Trichopalpus obscurellus (Zetterstedt, 1846)
= subarcticus (Ringdahl, 1936)
FANNIIDAE Schnabl & Dziezicki, 1911
FANNIA Robineau-Desvoidy, 1830
= Homalomyia Bouché, 1834
= Coelomyia Haliday, 1840
Fannia aethiops Malloch, 1913
Fannia alpina Pont, 1970
Fannia armata (Meigen, 1826)
Fannia atra (Stein, 1895)
Fannia canicularis (Linnaeus, 1761)
Fannia carbonaria (Meigen, 1826)
Fannia carbonella (Stein, 1895)
Fannia coracina (Loew, 1873)
Fannia corvina (Verrall, 1892)
= halterata Ringdahl, 1918
Fannia cothurnata (Loew, 1873)
Fannia dicilis (Stein, 1895)
Fannia fuscitibia Stein, 1920
Fannia fuscula (Fallén, 1825)
Fannia genualis (Stein, 1895)
Fannia gotlandica Ringdahl, 1926
Fannia hirticeps (Stein, 1892)
Fannia hirundinis Ringdahl, 1948
Fannia immutica Collin, 1939
Fannia incisurata (Zetterstedt, 1838)
Fannia latifrontalis Hennig, 1955
Fannia latipalpis (Stein, 1892)
Fannia lepida (Wiedemann, 1817)
= mutica (Zetterstedt, 1845)
Fannia leucosticta (Meigen, 1838)
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Fannia limbata (Tiensuu, 1938)
= berolinensis Hennig, 1955
Fannia lucidula (Zetterstedt, 1860)
= glaucescens auct. nec (Zetterstedt, 1845)
Fannia lugubrina (Zetterstedt, 1838)
Fannia lustrator (Harris, 1780)
= hamata (Macquart, 1835)
Fannia manicata (Meigen, 1826)
Fannia melania (Dufour, 1839)
= ciliata (Stein, 1895)
Fannia metallipennis (Zetterstedt, 1838)
= kowarzi (Verrall, 1892)
Fannia minutipalpis (Stein, 1895)
Fannia mollissima (Haliday, 1840)
= spathulata (Zetterstedt, 1845)
Fannia monilis (Haliday, 1838)
Fannia pallitibia (Rondani, 1866)
Fannia parva (Stein, 1895)
Fannia pauli Pont, 1997
= nitida (Stein, 1895) preocc.
Fannia polychaeta (Stein, 1895)
Fannia postica (Stein, 1895)
Fannia posticata (Meigen, 1826)
= pretiosa (Schiner, 1862)
Fannia rabdionata Karl, 1940
Fannia ringdahlana Collin, 1939
= umbrosa auct. nec (Stein, 1895)
Fannia rondanii (Strobl, 1893)
= carbonaria (Rondani, 1871) preocc.
= aerea misid.
Fannia scalaris (Fabricius, 1794)
= subscalaris Zimin, 1946
Fannia serena (Fallén, 1825)
Fannia similis (Stein, 1895)
Fannia slovaca Gregor & Rozkošný, 2005
Fannia sociella (Zetterstedt, 1845)
Fannia spathiophora Malloch 1918
= nodulosa Ringdahl, 1926
Fannia speciosa (Villeneuve, 1898)
Fannia stigi Rognes, 1982
Fannia subatripes d’Assis-Fonseca, 1967
Fannia subpellucens (Zetterstedt, 1845)
Fannia subpubescens Collin, 1958
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Fannia tuberculata (Zetterstedt, 1849)
Fannia umbratica Collin, 1939
Fannia umbrosa (Stein, 1895)
= subumbrosa Ringdahl, 1934
Fannia verrallii (Stein, 1895)
Fannia vesparia (Meade, 1891)
Fannia vespertilionis Ringdahl, 1934
PIEZURA Rondani, 1844
= Platycoenosia Strobl, 1894
Piezura graminicola (Zetterstedt, 1846)
= boletorum (Rondani, 1866)
= mikii (Strobl, 1894)
Piezura pardalina (Rondani, 1866)
= graminicola auct. nec (Zetterstedt, 1846)
MUSCIDAE Latreille, 1802
ACHANTHIPTERINAE Hennig, 1962
ACHANTHIPTERA Rondani, 1856
Achanthiptera rohrelliformis (Robineau-Desvoidy, 1830)
= inanis (Fallén, 1825) preocc.
COENOSIINAE Verrall, 1888
tribe Coenosiini Verrall, 1888
COENOSIA Meigen, 1826
= Caricea Robineau-Desvoidy, 1830
= Oplogaster Rondani, 1856
= Dexiopsis Pokorny, 1893
Coenosia acuminata Strobl, 1898
= annulipes Ringdahl, 1932
Coenosia agromyzina (Fallén, 1825)
Coenosia ambulans Meigen, 1826
Coenosia bilineella (Zetterstedt, 1838)
Coenosia campestris (Meigen, 1830)
= sexnotata auct. nec Meigen, 1826
? Coenosia comita (Huckett, 1936) (see Notes)
= ovulifera Tiensuu, 1938
Coenosia dealbata (Zetterstedt, 1838)
= fulvicornis (Zetterstedt, 1845)
Coenosia avimana (Zetterstedt, 1845)
= albatella (Zetterstedt, 1849)
Coenosia humilis Meigen, 1826
Coenosia intermedia (Fallén, 1825)
Coenosia lacteipennis (Zetterstedt, 1845)
Coenosia lineatipes (Zetterstedt, 1845)
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= albicornis misid.
Coenosia means Meigen, 1826
Coenosia mollicula (Fallén, 1825)
Coenosia octopunctata (Zetterstedt, 1838)
Coenosia paludis Tiensuu, 1939
Coenosia pedella (Fallén, 1825)
= decipiens Meigen, 1826
Coenosia perpusilla Meigen, 1826
Coenosia pudorosa Collin, 1953
Coenosia pulicaria (Zetterstedt, 1845)
Coenosia pumila (Fallén, 1825)
Coenosia pygmaea (Zetterstedt, 1845)
Coenosia rucornis Macquart, 1835
= litoralis (Zetterstedt, 1846)
Coenosia rupalpis Meigen, 1826
= avicauda Ringdahl, 1937
Coenosia sallae Tiensuu, 1938
Coenosia testacea (Robineau-Desvoidy, 1830)
= tricolor (Zetterstedt, 1845)
= alleni d’Assis-Fonseca, 1966
Coenosia tigrina (Fabricius, 1775)
Coenosia trilineella (Zetterstedt, 1838)
= trilineata emend.
Coenosia verralli Collin, 1953
= steini Verrall, 1912 preocc.
LIMNOSPILA Schnabl, 1902
Limnospila albifrons (Zetterstedt, 1849)
LISPOCEPHALA Pokorny, 1893
Lispocephala alma (Meigen, 1826)
Lispocephala erythrocera (Robineau-Desvoidy, 1830)
Lispocephala falculata Collin, 1963
Lispocephala fuscitibia Ringdahl, 1944
Lispocephala pallipalpis (Zetterstedt, 1845)
Lispocephala spuria (Zetterstedt, 1838)
= vitripennis Ringdahl, 1951
Lispocephala verna (Fabricius, 1794)
MACRORCHIS Rondani, 1877
Macrorchis meditata (Fallén, 1825)
PSEUDOCOENOSIA Stein, 1916
Pseudocoenosia abnormis Stein, 1916
Pseudocoenosia solitaria (Zetterstedt, 1838)
= longicauda (Zetterstedt, 1860)
SCHOENOMYZA Haliday, 1833
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Schoenomyza litorella (Fallén, 1823)
tribe Limnophoriini Villeneuve, 1902
LIMNOPHORA Robineau-Desvoidy, 1830
Limnophora nigripes (Robineau-Desvoidy, 1830)
Limnophora pandellei Séguy 1923 (see Notes)
Limnophora riparia (Fallén, 1824)
Limnophora rotundata Collin, 1930
Limnophora tigrina (Am Stein, 1860)
= notata (Fallén, 1823) preocc.
Limnophora triangula (Fallén, 1825)
Limnophora uniseta Stein, 1916
LISPE Latreille, 1796
Lispe consanguinea Loew, 1858
Lispe hydromyzina Fallén, 1825
Lispe litorea Fallén, 1825
Lispe melaleuca Loew, 1847
Lispe pygmaea Fallén, 1825
Lispe tentaculata (De Geer, 1776)
Lispe uliginosa Fallén, 1825
SPILOGONA Schnabl, 1911
Spilogona aerea (Fallén, 1825)
Spilogona albisquama (Ringdahl, 1932)
Spilogona alpica (Zetterstedt, 1845)
Spilogona arenosa (Ringdahl, 1918)
Spilogona atrisquamula Hennig, 1959
Spilogona baltica (Ringdahl, 1918)
Spilogona brunneifrons Ringdahl, 1931
Spilogona brunneisquama (Zetterstedt, 1845)
Spilogona carbonella (Zetterstedt, 1845)
Spilogona contractifrons (Zetterstedt, 1838)
Spilogona denigrata (Meigen, 1826)
Spilogona depressiuscula (Zetterstedt, 1838)
Spilogona depressula (Zetterstedt, 1845)
Spilogona dispar (Fallén, 1823)
= funeralis Rondani, 1866
Spilogona falleni Pont, 1984
= litorea auct. nec (Fallén, 1823)
Spilogona krogerusi (Ringdahl, 1941)
= micans misid.
Spilogona leucogaster (Zetterstedt, 1838)
Spilogona malaisei (Ringdahl, 1920)
Spilogona marginifera Hennig, 1959
= marginalis (Fallén, 1824) preocc.
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Spilogona meadei (Schnabl, 1915)
Spilogona megastoma (Boheman, 1866)
Spilogona nigriventris (Zetterstedt, 1845)
Spilogona nitidicauda (Schnabl, 1911)
Spilogona novemmaculata (Zetterstedt, 1860)
Spilogona opaca (Schnabl, 1915)
Spilogona pacica (Meigen, 1826)
= vana (Zetterstedt, 1845)
Spilogona palmeni (Ringdahl, 1935)
Spilogona pseudodispar (Frey, 1915)
= spinitibia (Ringdahl, 1918)
Spilogona pusilla (Huckett, 1932)
Spilogona quinquelineata (Zetterstedt, 1838)
Spilogona semiglobosa (Ringdahl, 1916)
Spilogona sororcula (Zetterstedt, 1845)
= zetterstedti (Ringdahl, 1918) preocc.
Spilogona spectabilis (Tiensuu, 1938)
Spilogona surda (Zetterstedt, 1845)
Spilogona tenuis Hennig, 1959
Spilogona tornensis (Ringdahl, 1926)
Spilogona triangulifera (Zetterstedt, 1838)
Spilogona trianguligera (Zetterstedt, 1838)
= insularis (Collin, 1921)
Spilogona trigonata (Zetterstedt, 1838)
Spilogona tundrae (Schnabl, 1915)
= macropyga (Frey, 1915)
Spilogona tundrica (Schnabl & Dziedzicki, 1911)
Spilogona varsaviensis (Schnabl & Dziedzicki, 1911)
= glauca (Stein, 1916)
Spilogona veterrima (Zetterstedt, 1845)
VILLENEUVIA Schnabl & Dziedzicki, 1911
Villeneuvia aestuum (Villeneuve, 1902)
AZELIINAE Robineau-Desvoidy, 1830
tribe Azeliini Robineau-Desvoidy, 1830
AZELIA Robineau-Desvoidy, 1830
Azelia aterrima (Meigen, 1826)
Azelia cilipes (Haliday, 1838)
Azelia gibbera (Meigen, 1826)
Azelia monodactyla Loew, 1874
Azelia nebulosa Robineau-Desvoidy, 1830
= macquarti (Staeger in Schiødte, 1843)
Azelia trigonica Hennig, 1956
= nuda Hennig, 1956
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Azelia triquetra (Wiedemann, 1817)
Azelia zetterstedtii Rondani, 1866
DRYMEIA Meigen, 1826
= Pogonomyia Rondani, 1871
= Trichopticoides Ringdahl, 1931
Drymeia hamata (Fallén, 1823)
Drymeia tetra (Meigen, 1826)
Drymeia vicana (Harris, 1780)
= decolor (Fallén, 1824)
HYDROTAEA Robineau-Desvoidy, 1830
= Ophyra Robineau-Desvoidy, 1830
= Lasiops Meigen, 1838
Hydrotaea aenescens (Wiedemann, 1830)
Hydrotaea albipuncta (Zetterstedt, 1845)
Hydrotaea anxia (Zetterstedt, 1838)
= bispinosa (Zetterstedt, 1845)
Hydrotaea armipes (Fallén, 1825)
= occulta (Meigen, 1826)
Hydrotaea basdeni Collin, 1939
Hydrotaea borussica Stein, 1899
Hydrotaea cyrtoneurina (Zetterstedt, 1845)
Hydrotaea dentipes (Fabricius, 1805)
Hydrotaea diabolus (Harrris, 1780)
= ciliata (Fabricius, 1794) preocc.
= bimaculata (Meigen, 1826)
Hydrotaea occosa Macquart, 1835
= armipes auct. nec (Fallén, 1825)
Hydrotaea ignava (Harrris, 1780)
= leucostoma (Wiedemann, 1817)
Hydrotaea irritans (Fallén, 1823)
Hydrotaea meridionalis Portschinsky, 1882
Hydrotaea meteorica (Linnaeus, 1758)
Hydrotaea militaris (Meigen, 1826)
Hydrotaea nidicola Malloch, 1925
Hydrotaea palaestrica (Meigen, 1826)
Hydrotaea pandellei Stein, 1899
Hydrotaea parva Meade, 1889
Hydrotaea pellucens Portschinsky, 1879
Hydrotaea pilipes Stein, 1903
Hydrotaea pilitibia Stein, 1916
Hydrotaea ringdahli Stein, 1916
Hydrotaea scambus (Zetterstedt, 1838)
Hydrotaea similis Meade, 1887
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Hydrotaea tuberculata Rondani, 1866
Hydrotaea velutina Robineau-Desvoidy, 1830
POTAMIA Robineau-Desvoidy, 1830
= Dendrophaonia Malloch, 1923
Potamia littoralis Robineau-Desvoidy, 1830
= querceti (Bouché, 1834)
THRICOPS Rondani, 1856
= Alleostylus Schnabl, 1888
ricops aculeipes (Zetterstedt, 1838)
ricops albibasalis (Zetterstedt, 1849)
= sudeticus auct. nec (Schnabl, 1888)
ricops coquilletti (Malloch, 1920)
ricops cunctans (Meigen, 1826)
= hirsutulus auct. nec (Zetterstedt, 1838)
ricops diaphanus (Wiedemann, 1817)
ricops foveolatus (Zetterstedt, 1845)
ricops genarum (Zetterstedt, 1838)
= sundewalli (Zetterstedt, 1845)
ricops hirtulus (Zetterstedt, 1838)
= subrostratus (Zetterstedt, 1845)
ricops innocuus (Zetterstedt, 1838)
ricops lividiventris (Zetterstedt, 1845)
ricops longipes (Zetterstedt, 1845)
ricops nigrifrons (Robineau-Desvoidy, 1830)
ricops nigritellus (Zetterstedt, 1838)
ricops rostratus (Meade, 1882)
ricops rusquamus (Schnabl, 1915)
= penicillatus (Ringdahl, 1926)
ricops semicinereus (Wiedemann, 1817)
ricops separ (Zetterstedt, 1845)
ricops simplex (Wiedemann, 1817)
tribe Reinwardtiini Brauer & Bergenstamm, 1889
MUSCINA Robineau-Desvoidy, 1830
Muscina angustifrons (Loew, 1858)
Muscina levida (Harris, 1780)
= assimilis (Fallén, 1823)
Muscina pascuorum (Meigen, 1826)
Muscina prolapsa (Harris, 1780)
= pabulorum (Fallén, 1817)
Muscina stabulans (Fallén, 1817)
MUSCINAE Latreille, 1802
tribe Muscini Latreille, 1802
EUDASYPHORA Townsend, 1911
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Eudasyphora cyanicolor (Zetterstedt, 1845)
Eudasyphora zimini (Hennig, 1963)
MESEMBRINA Meigen, 1826
= Hypodermodes Townsend, 1912
Mesembrina intermedia Zetterstedt, 1848?
Mesembrina meridiana (Linnaeus, 1758)
Mesembrina mystacea (Linnaeus, 1758)
Mesembrina resplendens Wahlberg, 1844
MORELLIA Robineau-Desvoidy, 1830
sg. Morellia Robineau-Desvoidy, 1830
Morellia aenescens Robineau-Desvoidy, 1830
Morellia hortorum (Fallén, 1817)
Morellia podagrica (Loew, 1857)
sg. Ziminiella Nihei & Carvalho, 2007
Morellia simplex (Loew, 1857)
MUSCA Linnaeus, 1758
Musca autumnalis De Geer, 1776
= corvina Fabricius, 1781
Musca domestica Linnaeus, 1758
Musca tempestiva Fallén, 1817
NEOMYIA Walker, 1859
= Orthellia Robineau-Desvoidy, 1863
Neomyia cornicina (Fabricius, 1781)
= caesarion (Meigen, 1826)
= fennica (Frey, 1909)
Neomyia viridescens (Robineau-Desvoidy, 1830)
= cornicina auct. nec (Fabricius, 1781)
POLIETES Rondani, 1866
Polietes domitor (Harris, 1780)
= albolineatus (Fallén, 1823)
Polietes lardarius (Fabricius, 1781)
Polietes nigrolimbatus (von Bonsdor, 1866)
Polietes steinii (Ringdahl, 1913)
PYRELLIA Robineau-Desvoidy, 1830
Pyrellia vivida Robineau-Desvoidy, 1830
= cadaverina auct. nec (Linnaeus, 1758)
tribe Stomoxyini Meigen, 1824
HAEMATOBIA Le Peletier & Serville, 1828
= Lyperosia Rondani, 1856
Haematobia irritans (Linnaeus, 1758)
HAEMATOBOSCA Bezzi, 1907
Haematobosca alcis Snow, 1891
= crassipalpis (Ringdahl, 1920)
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Haematobosca stimulans (Meigen, 1824)
STOMOXYS Georoy, 1762
Stomoxys calcitrans (Linnaeus, 1758)
MYDAEINAE Verrall, 1888
GRAPHOMYA Robineau-Desvoidy, 1830
Graphomya maculata (Scopoli, 1763)
Graphomya minor Robineau-Desvoidy, 1830
GYMNODIA Robineau-Desvoidy, 1863
= Brontaea Kowarz, 1873
Gymnodia humilis (Zetterstedt, 1860)
HEBECNEMA Schnabl, 1889
Hebecnema fumosa (Meigen, 1826)
Hebecnema nigra (Robineau-Desvoidy, 1830)
= vespertina auct. nec (Fallén, 1823)
Hebecnema nigricolor (Fallén, 1825)
Hebecnema umbratica (Meigen, 1826)
Hebecnema vespertina (Fallén, 1823)
= anis Malloch, 1921
MYDAEA Robineau-Desvoidy, 1830
Mydaea anis Meade, 1891
= discimana Malloch, 1920
Mydaea ancilla (Meigen, 1826)
Mydaea anicula (Zetterstedt, 1860)
Mydaea corni Scopoli, 1763
= pagana (Fabricius, 1794) preocc.
= scutellaris Robineau-Desvoidy, 1830
Mydaea deserta (Zetterstedt, 1845)
Mydaea detrita (Zetterstedt, 1845)
= electa (Zetterstedt, 1860)
Mydaea humeralis Robineau-Desvoidy, 1830
= tincta (Zetterstedt, 1845)
Mydaea nebulosa (Stein, 1893)
Mydaea obscurella Malloch, 1921
= bengtssoni Ringdahl, 1924
Mydaea orthonevra (Macquart, 1835)
= detrita auct. nec (Zetterstedt, 1845)
Mydaea palpalis Stein, 1916
Mydaea setifemur Ringdahl, 1924
Mydaea sootryeni Ringdahl, 1928
Mydaea urbana (Meigen, 1826)
MYOSPILA Rondani, 1856
Myospila bimaculata (Macquart, 1834)
Myospila meditabunda (Fabricius, 1781)
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OPSOLASIA Coquillett, 1910
Opsolasia orichalcea (Zetterstedt, 1849)
PHAONIINAE Malloch, 1917
tribe Phaoniini Malloch, 1917
HELINA Robineau-Desvoidy, 1830
Helina abdominalis (Zetterstedt, 1846)
Helina allotalla (Meigen, 1830)
Helina annosa (Zetterstedt, 1838)
Helina atricolor (Fallén, 1825)
= denudata (Zetterstedt, 1845)
= denutata emend.
Helina bohemani (Ringdahl, 1916)
Helina celsa (Harris, 1780)
= quadrimaculata (Fallén, 1823) preocc.
= quadrimaculella Hennig, 1957
? Helina ciliata Karl, 1929 (see Notes)
Helina ciliatocosta (Zetterstedt, 1845)
= ciliatocostata emend.
Helina cilipes (Schnabl, 1902)
Helina cinerella (van der Wulp, 1867)
= vanderwulpi (Schnabl, 1888)
= calceata misid.
Helina connis (Fallén, 1825)
= anceps (Zetterstedt, 1838)
Helina consimilis (Fallén, 1825)
Helina cothurnata (Rondani, 1866)
= obscuripes auct. nec (Zetterstedt, 1845)
Helina daicles (Walker, 1849)
= binotata (Zetterstedt, 1845)
Helina depuncta (Fallén, 1825)
Helina evecta (Harris, 1780)
= lucorum (Fallén, 1823) preocc.
= laetica (Robineau-Desvoidy, 1830)
Helina avisquama (Zetterstedt, 1849)
Helina fratercula (Zetterstedt, 1845)
Helina fulvisquama (Zetterstedt, 1845)
Helina impuncta (Fallén, 1825)
Helina latitarsis Ringdahl, 1924
Helina laxifrons (Zetterstedt, 1860)
Helina longicornis (Zetterstedt, 1838)
Helina luteisquama (Zetterstedt, 1845)
Helina maculipennis (Zetterstedt, 1845)
= obscuripes (Zetterstedt, 1845)
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Helina obscurata (Meigen, 1826)
Helina pertusa (Meigen, 1826)
Helina protuberans (Zetterstedt, 1845)
Helina pubiseta (Zetterstedt, 1845)
Helina quadrinotata (Meigen, 1826)
Helina quadrum (Fabricius, 1805)
Helina reversio (Harris, 1780)
= duplicata (Meigen, 1826)
= duplaris auct. nec (Zetterstedt, 1845)
= communis (Robineau-Desvoidy, 1830)
Helina setiventris Ringdahl, 1924
Helina sexmaculata (Preyssler, 1791)
= uliginosa (Fallén, 1825) preocc.
= punctata (Robineau-Desvoidy, 1830)
Helina spinicosta (Zetterstedt, 1845)
Helina squalens (Zetterstedt, 1838)
= borealis (Zetterstedt, 1838)
Helina subvittata (Séguy, 1923)
= rothi Ringdahl, 1939
= marmorata auct. nec (Zetterstedt, 1860)
Helina tetrastigma (Meigen, 1826)
= agripes (Rondani, 1866)
Helina trivittata (Zetterstedt, 1860)
= atripes (Meade, 1889)
Helina veterana (Zetterstedt, 1838)
Helina vicina (Czerny, 1900)
= suecica Ringdahl, 1924
LOPHOSCELES Ringdahl, 1922
Lophosceles cinereiventris (Zetterstedt, 1845)
= cristata (Zetterstedt, 1845)
Lophosceles frenatus (Holmgren, 1872)
Lophosceles hians (Zetterstedt, 1838)
Lophosceles mutatus (Fallén, 1825)
PHAONIA Robineau-Desvoidy, 1830
= Wahlgrenia Ringdahl, 1929
= Dialytina Ringdahl, 1945
Phaonia aeneiventris (Zetterstedt, 1845)
= cinctinervis (Zetterstedt, 1860)
Phaonia alpicola (Zetterstedt, 1845)
Phaonia amabilis (Meigen, 1826)
Phaonia amicula Villeneuve, 1922 (see Notes)
Phaonia angelicae (Scopoli, 1763)
= basalis (Zetterstedt, 1838)
Jere Kahanpää & Antti Haarto / ZooKeys 441: 347–367 (2014)
364
Phaonia angulicornis (Zetterstedt, 1838)
= erinacea (Fallén, 1824)
Phaonia apicalis Stein, 1914
Phaonia atriceps (Loew, 1858)
Phaonia atrocyanea Ringdahl, 1916
Phaonia canescens Stein, 1916
Phaonia consobrina (Zetterstedt, 1838)
Phaonia czernyi Hennig, 1963
= steinii Czerny, 1900 preocc.
Phaonia errans (Meigen, 1826)
= erratica (Fallén, 1825) preocc.
Phaonia erronea (Schnabl, 1887)
Phaonia falleni (Michelsen, 1977)
= vagans (Fallén, 1825) preocc.
Phaonia fugax Tiensuu, 1946
Phaonia fuscata (Fallén, 1825)
Phaonia gobertii (Mik, 1881)
Phaonia gracilis Stein, 1916
Phaonia grandaeva (Zetterstedt, 1845)
Phaonia halterata (Stein, 1893)
Phaonia hybrida (Schnabl, 1888)
Phaonia incana (Wiedemann, 1817)
Phaonia jaroschewskii (Schnabl, 1888)
= crinipes Stein, 1913
Phaonia kowarzii (Schnabl, 1886)
= fulvicornis Tiensuu, 1936
Phaonia laeta (Fallén, 1823)
Phaonia latipalpis Schnabl, 1911
= umbraticola d’Assis-Fonseca, 1957
Phaonia longicornis Stein, 1916
Phaonia lugubris (Meigen, 1826)
= morio (Zetterstedt, 1845)
Phaonia magnicornis (Zetterstedt, 1845)
Phaonia meigeni Pont, 1986
= lugubris auct. nec (Meigen, 1826)
Phaonia mystica (Meigen, 1826)
= vittifera (Zetterstedt, 1845)
Phaonia nymphaearum (Robineau-Desvoidy, 1830)
= nitida (Macquart, 1835)
Phaonia pallida (Fabricius, 1787)
Phaonia pallidisquama (Zetterstedt, 1849)
Phaonia palpata (Stein, 1897)
Phaonia perdita (Meigen, 1830)
Phaonia pratensis (Robineau-Desvoidy, 1830)
Checklist of the families Scathophagidae, Fanniidae and Muscidae of Finland (Insecta, Diptera)365
= laeta auct. nec (Fallén, 1923)
Phaonia rupalpis (Macquart, 1835)
Phaonia ruventris (Scopoli, 1763)
= populi (Meigen, 1826)
Phaonia serva (Meigen, 1826)
Phaonia steinii (Strobl, 1898)
Phaonia subfuscinervis (Zetterstedt, 1838)
Phaonia subventa (Harris, 1780)
= variegata (Meigen, 1826)
Phaonia taigensis Zinovjev, 1987
Phaonia tiei Schnabl, 1888
Phaonia trimaculata (Bouché)
= servaeformis Ringdahl, 1916
Phaonia tuguriorum (Scopoli, 1763)
= signata (Meigen, 1826)
Phaonia valida (Harris, 1780)
= viarum (Robineau-Desvoidy, 1830)
= erratica auct. nec (Fallén, 1825)
Phaonia villana Robineau-Desvoidy, 1830
= mystica auct. nec (Meigen, 1826)
Phaonia vivida (Rondani, 1870)
= austriaca (Czerny, 1900)
Phaonia wahlbergi Ringdahl, 1930
Phaonia zugmayeriae (Schnabl, 1888)
?= humeralis (Zetterstedt, 1845)
= humerella (Stein, 1900)
Excluded species
Coenosia femoralis (Robineau-Desvoidy, 1830) misidentied
Fannia barbata (Stein, 1892) not found within present borders
Fannia lineata (Stein, 1895) not found within present borders
Fannia nigra Malloch, 1910 not found within present borders
Hydrotaea glabricula (Fallén, 1825)
Limnophora pollinifrons Stein, 1916 not found within present borders
Phaonia bitincta (Rondani, 1866) misidentied
Scathophaga calida Haliday in Curtis, 1832 not found within present borders
Spilogona acrostichalis (Stein, 1916)
Spilogona norvegica (Ringdahl, 1932)
Spilogona obscuripennis (Stein, 1916) not found within present borders
Spilogona septemnotata (Zetterstedt, 1845) misidentied
Spilogona setigera (Stein, 1907)
Spilogona spininervis (Villeneuve, 1922)
Jere Kahanpää & Antti Haarto / ZooKeys 441: 347–367 (2014)
366
Notes
Coenosia comita (Huckett, 1936). Collected only from Salla. e type locality of C. ovuli-
vera Tiensuu, Vuorijärvi, is now Russian territory (Tiensuu 1938). A second specimen,
also taken in 1936 by R. Krogerus (in MZH), bears only the label ’Salla’ and probably
also originates from the vicinity of Vuorijärvi, but it is possible that it was collected from
the Finnish part of Salla, a municipality split between Russia and Finland in 1944.
Helina ciliata Karl, 1929. Ringdahl (1952), quoting ”Tiensuu’s papers”, lists this
species from Finland. We could not locate the original record in Tiensuu’s pub-
lished works or among the available collection specimens. It seems possible Ring-
dahl make a mistake in his tables. e species is also listed as Finnish by Hackman
(1980), again without any details.
Limnophora pandellei Séguy, 1923. Only females have been found in Finland. e
identication remains slightly doubtful (Kahanpää 2013).
Norellia tipularia (Fabricius, 1794). is species was rst recorded from Finland
by Winqvist (2011). Šifner (2003, 2008) considers N. tipularia a nomen dubium
and probably synonymous with N. spinipes Meigen, but Ozerov and Krivosheina
(2011) disagree and provide a detailed diagnosis.
Parallelomma paridis Hering, 1923. is name was synonymized with P. vittatum (Mei-
gen) by Šifner (1978). Ozerov (2010, 2014) examined the types of P. paridis and up-
held the synonymy. Not everyone agrees with this synonymy (Nelson 1990). Finnish
CO1 DNA ngerprints (Kahanpää, unpublished) of larvae on Paris (Liliaceae) and
adults caught among Dactylorhiza (Orchidaceae) suggest that two species are involved.
Phaonia amicula Villeneuve, 1922. New to Finland. Recorded as Phaonia sp. nr.
halterata by Winqvist (2011): Ab: Turku, Ruissalo (67093:32336), 3.vii.2008, leg
K. Winqvist, 1 male.
Acknowledgements
I would like to express my appreciation to Dr. Verner Michelsen for his constructive
feedback and for identifying the rst Finnish specimen of Phaonia amicula. Mr. Kaj
Winqvist kindly allowed us to include his record of P. amicula in this paper.
References
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Tiensuu L (1935) Die bisher aus Finnland bekannten Musciden. Acta Societatis pro Fauna et
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Checklist of the family Anthomyiidae (Diptera) of Finland 369
Checklist of the family Anthomyiidae (Diptera) of Finland
Verner Michelsen1
1 Natural History Museum of Denmark (Zoological Museum), Universitetsparken 15, DK-2100, Copenhagen Ø,
Denmark
Corresponding author: Verner Michelsen (vmichelsen@snm.ku.dk)
Academic editor: J. Kahanpää|Received 15 March 2014|Accepted 8 May 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Michelsen V (2014) Checklist of the family Anthomyiidae (Diptera) of Finland. In: Kahanpää J, Salmela J
(Eds) Checklist of the Diptera of Finland. ZooKeys 441: 369–382. doi: 10.3897/zookeys.441.7527
Abstract
An updated checklist of the the genera and species of Anthomyiidae (Diptera) found in Finland is
provided.
Keywords
Checklist, Finland, Diptera, Anthomyiidae
Introduction
e family Anthomyiidae is a large and taxonomically dicult group of ies that has
for the same reason suered from unstable taxonomy and nomenclature. A checklist
of the anthomyiid species known from pre-war Finland was compiled by their leading
regional specialist of calyptrate ies Lauri Tiensuu (1906−1980) and published in Frey
et al. (1941). e Anthomyiidae were then not recognized as a separate family but
combined with the fanniid and true muscid ies in a comprehensive Muscidae fam-
ily equivalent of the present Muscoidea less Scathophagidae. Tiensuu’s list included
conrmed records of 199 anthomyiid species classied in 41 genera and subgenera. No
less than 34% of the species names and 58% of the genus-group names in that list are
ZooKeys 441: 369–382 (2014)
doi: 10.3897/zookeys.441.7527
www.zookeys.org
Copyright Verner Michelsen. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Verner Michelsen / ZooKeys 441: 369–382 (2014)
370
presently replaced by other names for taxonomic and/or nomenclatorial reasons. Most
changes were introduced by Hennig (1966−1976) and Michelsen (1985).
e next Finnish checklist of Anthomyiidae by Hackman (1980) was prepared
under the inuence by two major events. Firstly, 10% of the area of pre-war Finland,
notably a major part of Finnish Karelia, had been ceded to the Soviet Union. In con-
sequence, Hackman’s list included fewer species: 175 (too few actually, as he missed a
few species recorded by Tiensuu (1936) from present Finland). Secondly, the appear-
ance of Hennig’s (1966−1976) major revision of Palearctic Anthomyiidae had resulted
in numerous name changes and redenitions of both species and genera. Hackman’s
list was updated according to Hennig’s proposals except for the splitting of newly rec-
ognized species complexes.
After the appearance of the 1941-checklist, L. Tiensuu had stopped publishing
taxonomic and faunistic papers on Finnish Diptera altogether, although he stayed
active all his life as a skilled collector of Finnish Diptera. Apart from W. Hennig’s
study of some type specimens in the Helsinki Museum and a project on mushroom
feeding Pegomya (Hackman, 1976, 1979; Hackman & Meinander, 1979) Finnish an-
thomyiids were given little attention until the present author in the early 1980’s began
a long-term study of the species composition and distributionof Anthomyiidae in Fen-
noscandia and Denmark.
e web-based database Fauna Europaea ver. 1.0 released in 2004 included an-
thomyiid checklists (Michelsen, 2004) for all parts of Europe. It was based extensively
on literature records, but in respect to Finland and the other Scandinavian countries
almost exclusively on personally veried records. e Diptera collection of the Finnish
Museum of Natural History, University of Helsinki (MZH) provided most of the data
on the Finnish anthomyiid fauna. After several updates, most recently ver. 2.4 from
January 2011, the Finnish list of Anthomyiidae (Michelsen 2011) had grown sub-
stantially to 273 species (http://www.faunaeur.org/). e anthomyiid checklist given
below has been further expanded and now contains 289 species. Comparisons with
the better known anthomyiid fauna of Sweden and the presence of some species await-
ing description suggest that the anthomyiid fauna of Finland may totally comprise
315−325 species.
All nomenclatorial and taxonomic adjustments (new synonyms, misidenti-
cations, changed circumscriptions of species and genera) made since the appearance
of Hackman’s (1980) checklist are indicated in the following, updated checklist.
In agreement with Fauna Europaea the genera are listed alphabetically, as existing
attempts to split the family into subfamilies and tribes are preliminary and in part
poorly substantiated.
Number of described species:
World: Presently c. 2000.
Europe: 508 (Michelsen 2011)
Finland: 289 (present paper)
Faunistic knowledge level in Finland: average
Checklist of the family Anthomyiidae (Diptera) of Finland 371
Checklist
superfamily Muscoidea Latreille, 1802
family ANTHOMYIIDAE Robineau-Desvoidy, 1830
ADIA Robineau-Desvoidy, 1830
= Paregle auct. p. p.
Adia cinerella (Fallén, 1825)
ALLIOPSIS Schnabl & Dziedzicki, 1911
= Paraprosalpia Villeneuve, 1922
Alliopsis albipennis (Ringdahl, 1928)
Alliopsis aldrichi (Ringdahl, 1934)
Alliopsis atronitens (Strobl, 1893)
Alliopsis benanderi (Ringdahl, 1926)
Alliopsis billbergi (Zetterstedt, 1838)
Alliopsis brunneigena (Schnabl, 1915)
= incisa Ringdahl, 1926
Alliopsis conifrons (Zetterstedt, 1845)
Alliopsis denticauda (Zetterstedt, 1838)
Alliopsis dentiventris (Ringdahl, 1918)
Alliopsis fractiseta (Stein, 1908)
Alliopsis glacialis (Zetterstedt, 1845)
Alliopsis longipennis (Ringdahl, 1918)
Alliopsis maculifrons (Zetterstedt, 1838)
Alliopsis moerens (Zetterstedt, 1838)
Alliopsis sepiella (Zetterstedt, 1845)
Alliopsis silvestris (Fallén, 1824)
Alliopsis teriolensis (Pokorny, 1893)
= borealis Stein, 1916
ANTHOMYIA Meigen, 1803
= Craspedochoeta Macquart, 1851
= Craspedochaeta Scudder, 1884 (emend.)
= Chelisia Rondani, 1856
Anthomyia canningsi Griths, 2001
Anthomyia confusanea Michelsen in Michelsen & Báez, 1985
= pullula auct. p. p.
Anthomyia imbrida Rondani, 1866
= pluvialis auct. p. p.
Anthomyia liturata (Robineau-Desvoidy, 1830)
= pullula Zetterstedt, 1845
Anthomyia mimetica (Malloch, 1918)
= angulata Tiensuu, 1938
Anthomyia monilis (Meigen, 1826)
Verner Michelsen / ZooKeys 441: 369–382 (2014)
372
Anthomyia plurinotata Brullé, 1832
Anthomyia pluvialis (Linnaeus, 1758)
Anthomyia procellaris Rondani, 1866
= pluvialis auct. p. p.
BOREOPHORBIA Michelsen, 1987
Boreophorbia hirtipes (Stein, 1907)
BOTANOPHILA Lioy, 1864
= Pegohylemyia Schnabl, 1911
Botanophila apiciseta (Ringdahl, 1933)
Botanophila betarum (Lintner, 1883)
= macra Karl, 1940
Botanophila biciliaris (Pandellé, 1900)
Botanophila bidens (Ringdahl, 1933)
Botanophila brunneilinea (Zetterstedt, 1845)
Botanophila cognata Suwa, 2011
= subnitida Malloch, 1920?
Botanophila discreta (Meigen, 1826)
= striolata auct. p. p.
Botanophila dissecta (Meigen, 1826)
Botanophila estonica (Elberg, 1970)
= varicolor auct. p. p.
Botanophila fugax (Meigen, 1826)
Botanophila gemmata (Zetterstedt, 1860)
Botanophila gnava (Meigen, 1826)
Botanophila hucketti (Ringdahl, 1935)
Botanophila impudica (Rondani, 1866)
= varicolor auct. p. p.
Botanophila laterella (Collin, 1967)
Botanophila latifrons (Zetterstedt, 1845)
= humeralis Hennig, 1970
Botanophila lobata (Collin, 1967)
Botanophila maculipes (Zetterstedt, 1845)
= pseudomaculipes Strobl, 1893
Botanophila miniatura (Huckett, 1965)
Botanophila nigra Michelsen, 2009
Botanophila petrophila (Ringdahl, 1926)
Botanophila phrenione (Séguy, 1937)
Botanophila profuga (Stein, 1916)
Botanophila relativa (Huckett, 1965)
Botanophila rubrifrons (Ringdahl, 1933)
Botanophila rubrigena (Schnabl, 1915)
Botanophila salicis (Ringdahl, 1918)
Botanophila sericea (Malloch, 1920)
Checklist of the family Anthomyiidae (Diptera) of Finland 373
= obscura auct. nec Macquart, 1835
Botanophila silvatica (Robineau-Desvoidy, 1830)
Botanophila sonchi (Hardy, 1872)
Botanophila spinosa (Rondani, 1866)
Botanophila striolata (Fallén, 1824)
Botanophila trapezina (Zetterstedt, 1845)
= varicolor auct. p. p.
Botanophila unicolor (Ringdahl, 1932)
Botanophila varicolor (Meigen, 1826)
Botanophila verticella (Zetterstedt, 1838)
= lineatula Karl, 1928
CALYTHEA Schnabl & Dziedzicki, 1911
Calythea nigricans (Robineau-Desvoidy, 1830)
CHIASTOCHETA Pokorny, 1889
Chiastocheta dentifera Hennig, 1953
Chiastocheta inermella (Zetterstedt, 1838)
= trollii auct. p. p.
Chiastocheta macropyga Hennig, 1953
Chiastocheta trollii (Zetterstedt, 1845)
CHIROSIA Rondani, 1856
= Craspedochaeta auct. p. p.
Chirosia albitarsis (Zetterstedt, 1845)
Chirosia betuleti (Ringdahl, 1935)
Chirosia bisinuata (Tiensuu, 1939)
Chirosia cinerosa (Zetterstedt, 1845)
Chirosia crassiseta Stein, 1908
Chirosia avipennis (Fallén, 1823)
Chirosia griseifrons (Séguy, 1923)
Chirosia grossicauda Strobl, 1901
= parvicornis auct. nec Zetterstedt, 1845
Chirosia histricina (Rondani, 1866)
= hystrix Brischke, 1880
Chirosia nigripes Bezzi, 1895
= albifrons Tiensuu, 1938
Chirosia similata (Tiensuu, 1939)
DELIA Robineau-Desvoidy, 1830
Delia abruptiseta (Ringdahl, 1935)
Delia albula (Fallén, 1825)
Delia angustaeformis (Ringdahl, 1933)
Delia angustifrons (Meigen, 1826)
Delia antiqua (Meigen, 1826)
Delia brassicaeformis (Ringdahl, 1926)
Delia brunnescens (Zetterstedt, 1845)
Verner Michelsen / ZooKeys 441: 369–382 (2014)
374
Delia cardui (Meigen, 1826)
Delia carduiformis (Schnabl in Schnabl & Dziedzicki, 1911)
Delia coarctata (Fallén, 1825)
Delia coarctoides Michelsen, 2007
Delia cregyoglossa (Huckett, 1965)
= turcica Hennig, 1974
Delia criniventris (Zetterstedt, 1860)
Delia cuneata Tiensuu, 1946
Delia diluta (Stein, 1916)
Delia echinata (Séguy, 1923)
Delia fabricii (Holmgren, 1872)
Delia avogrisea (Ringdahl, 1926)
Delia oralis (Fallén, 1824)
Delia orilega (Zetterstedt, 1845)
Delia frontella (Zetterstedt, 1838)
Delia hirtitibia (Stein, 1916)
Delia judicariae (Pokorny, 1893)
= avifrons auct. p. p.
Delia lamelliseta (Stein, 1900)
Delia linearis (Stein, 1898)
= abellifera Pandellé, 1900
Delia lineariventris (Zetterstedt, 1845)
Delia longicauda (Strobl, 1898)
Delia lophota (Pandellé, 1900)
= nuda Strobl, 1901
Delia martini Griths, 1993
Delia nigrescens (Rondani, 1877)
Delia nudicosta (Ringdahl, 1949)
Delia pallipennis (Zetterstedt, 1838)
= candens Zetterstedt, 1845
Delia penicilliventris Ackland, 2010
= penicillaris auct. nec Rondani, 1866
Delia pilifemur (Ringdahl, 1933)
Delia piliventris (Pokorny, 1889)
Delia planipalpis (Stein, 1898)
Delia platura (Meigen, 1826)
Delia pruinosa (Zetterstedt, 1845)
= avifrons Zetterstedt, 1860
Delia radicum (Linnaeus, 1758)
= brassicae Wiedemann, 1817
Delia rimiventris Michelsen, 2007
Delia rondanii (Ringdahl, 1918)
Delia setigera (Stein, 1920)
Checklist of the family Anthomyiidae (Diptera) of Finland 375
Delia sileni Michelsen, 2012
= avifrons auct. p. p.
Delia subalpina (Ringdahl, 1926)
Delia tarsata (Ringdahl, 1918)
Delia tenuiventris (Zetterstedt, 1860)
= angustitarsis Malloch, 1920
Delia tumidula (Ringdahl, 1949)
Delia uniseriata (Stein, 1914)
EGLE Robineau-Desvoidy, 1830
Egle atomaria (Zetterstedt, 1845)
Egle brevicornis (Zetterstedt, 1838)
Egle ciliata (Walker, 1849)
= muscaria auct. nec Fabricius, 1794
Egle concomitans (Pandellé, 1900)
Egle ignobilis Michelsen, 2009
Egle inermis Ackland, 1970
Egle lyneborgi Ackland & Griths in Griths, 2003
Egle minuta (Meigen, 1826)
Egle parva Robineau-Desvoidy, 1830
Egle parvaeformis Schnabl in Schnabl & Dziedzicki, 1911
Egle pilitibia (Ringdahl, 1918)
Egle rhinotmeta (Pandellé, 1900)
Egle steini Schnabl in Schnabl & Dziedzicki, 1911
Egle subarctica (Huckett, 1965)
EMMESOMYIA Malloch, 1917
Emmesomyia grisea (Robineau-Desvoidy, 1830)
= socia auct. p. p.
Emmesomyia socia (Fallén, 1825)
EUSTALOMYIA Kowarz, 1873
Eustalomyia festiva (Zetterstedt, 1845)
Eustalomyia hilaris (Fallén, 1823)
Eustalomyia histrio (Zetterstedt, 1838)
Eustalomyia vittipes (Zetterstedt, 1845)
EUTRICHOTA Kowarz, 1893
= Eremomyia Stein, 1898
= Pegomyza Schnabl & Dziezicki, 1911
= Arctopegomyia Ringdahl, 1938
Eutrichota frigida (Zetterstedt, 1845)
Eutrichota labradorensis (Malloch, 1920)
Eutrichota longimana (Pokorny, 1887)
Eutrichota pilimana (Ringdahl, 1918)
Eutrichota praepotens (Wiedemann, 1817)
Eutrichota schineri (Schnabl, 1910)
Verner Michelsen / ZooKeys 441: 369–382 (2014)
376
= socculata auct. nec Zetterstedt, 1845
Eutrichota socculata (Zetterstedt, 1845)
= consanguinea Tiensuu, 1938
Eutrichota tunicata (Zetterstedt, 1846)
FUCELLIA Robineau-Desvoidy, 1842
Fucellia fucorum (Fallén, 1819)
Fucellia griseola (Fallén, 1819)
Fucellia tergina (Zetterstedt, 1845)
HETEROSTYLODES Hennig, 1967
Heterostylodes macrura (Schnabl in Schnabl & Dziedzicki, 1911)
Heterostylodes nominabilis (Collin, 1947)
= pratensis auct. p. p.
Heterostylodes obscura (Macquart, 1835)
= pratensis auct. p. p.
Heterostylodes pilifera (Zetterstedt, 1845)
= pratensis auct. p. p.
Heterostylodes pratensis (Meigen, 1826)
HYDROPHORIA Robineau-Desvoidy, 1830
Hydrophoria lancifer (Harris, 1780)
= conica Wiedemann, 1817
Hydrophoria linogrisea (Meigen, 1826)
Hydrophoria silvicola (Robineau-Desvoidy, 1830)
= annulata auct. nec Pandellé, 1899
HYLEMYA Robineau-Desvoidy, 1830
Hylemya nigrimana (Meigen, 1826)
Hylemya urbica Wulp, 1896
= latifrons Schnabl in Schnabl & Dziedzicki, 1911
Hylemya vagans (Panzer, 1798)
= strenua Robineau-Desvoidy, 1830
Hylemya variata (Fallén, 1823)
HYLEMYZA Schnabl & Dziedzicki, 1911
= Hylemya auct. p. p.
Hylemyza partita (Meigen, 1826)
LASIOMMA Stein, 1916
= Crinurina Karl, 1928
= Acrostilpina Ringdahl, 1929
Lasiomma anthomyinum (Rondani, 1866)
Lasiomma atricauda (Zetterstedt, 1845)
Lasiomma collini (Ringdahl, 1929)
Lasiomma craspedodontum (Hsue, 1980)
Lasiomma cuneicorne (Zetterstedt, 1838)
Lasiomma iwasai Suwa, 1978
Lasiomma japonicum Suwa, 1971
Checklist of the family Anthomyiidae (Diptera) of Finland 377
Lasiomma latipenne (Zetterstedt, 1838)
Lasiomma monticola Suh & Kwon, 1985
Lasiomma morionellum (Zetterstedt, 1838)
Lasiomma picipes (Meigen, 1826)
= octoguttata Zetterstedt, 1845
Lasiomma quinquelineatum (Ringdahl, 1926)
Lasiomma seminitidum (Zetterstedt, 1845)
Lasiomma strigilatum (Zetterstedt, 1838)
= nitidicauda Zetterstedt, 1855
LEUCOPHORA Robineau-Desvoidy, 1830
Leucophora cinerea Robineau-Desvoidy, 1830
Leucophora grisella Hennig, 1967
Leucophora obtusa (Zetterstedt, 1838)
Leucophora sericea Robineau-Desvoidy, 1830
Leucophora tavastica (Tiensuu, 1939)
Leucophora unilineata (Zetterstedt, 1838)
Leucophora unistriata (Zetterstedt, 1838)
MYCOPHAGA Rondani, 1856
Mycophaga testacea (Gimmerthal, 1834)
MYOPINA Robineau-Desvoidy, 1830
Myopina myopina (Fallén, 1824)
= reexa Robineau-Desvoidy, 1830
Myopina scoparia (Zetterstedt, 1845)
PARADELIA Ringdahl, 1933
= Pseudonupedia Ringdahl, 1959 (nomen nudum)
= Pseudonupedia Huckett, 1971
Paradelia brunneonigra (Schnabl in Schnabl & Dziedzicki, 1911)
Paradelia intersecta (Meigen, 1826)
Paradelia lunatifrons (Zetterstedt, 1846)
Paradelia lundbeckii (Ringdahl, 1918)
PAREGLE Schnabl, 1911
= Chionomyia Ringdahl, 1933
Paregle atrisquama (Ringdahl, 1948)
Paregle audacula (Harris, 1780)
= radicum auct. nec Linnaeus, 1758
Paregle vetula (Zetterstedt, 1838)
PEGOMYA Robineau-Desvoidy, 1830
Pegomya atricauda Ringdahl, 1944
Pegomya avida Hennig, 1973
Pegomya betae (Curtis, 1847)
= hyoscyami auct. p. p.
Pegomya bicolor (Wiedemann, 1817)
Pegomya caesia Stein, 1906
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378
Pegomya calyptrata (Zetterstedt, 1846)
Pegomya circumpolaris Ackland & Griths in Griths, 1983
Pegomya conformis (Fallén, 1825)
Pegomya cunicularia (Rondani, 1866)
= hyoscyami auct. p. p.
Pegomya curviphallis Michelsen, 2006
Pegomya deprimata (Zetterstedt, 1845)
Pegomya exilis (Meigen, 1826)
= hyoscyami auct. p. p.
Pegomya avifrons (Walker, 1849)
Pegomya avoscutellata (Zetterstedt, 1838)
Pegomya fulgens (Meigen, 1826)
Pegomya furva Ringdahl, 1938
Pegomya fuscinata Tiensuu, 1939
Pegomya geniculata (Bouché, 1834)
Pegomya grahami Michelsen & Ackland, 2009
Pegomya haemorrhoum (Zetterstedt, 1838)
= haemorrhoa (misspelling)
Pegomya holmgreni (Boheman, 1859)
Pegomya holosteae (Hering, 1924)
Pegomya hyoscyami (Panzer, 1809)
Pegomya incisiva Stein, 1906
Pegomya interruptella (Zetterstedt, 1855)
Pegomya lurida (Zetterstedt, 1846)
Pegomya maculata Stein, 1906
= atricauda auct. p. p.
Pegomya meridiana (Villeneuve, 1923)
Pegomya notabilis (Zetterstedt, 1846)
= zonata auct. nec Zetterstedt, 1838
Pegomya pallidoscutellata (Zetterstedt, 1852)
Pegomya pulchripes (Loew, 1857)
= avipes Fallén, 1825
Pegomya rubivora (Coquillett in Slingerland, 1897)
Pegomya ruceps (Zetterstedt, 1838)
Pegomya runa (Fallén, 1825)
Pegomya scapularis (Zetterstedt, 1846)
= pilosa Stein, 1900
Pegomya seitenstettensis (Strobl, 1880)
Pegomya setaria (Meigen, 1826)
Pegomya solennis (Meigen, 1826)
= nigritarsis Zetterstedt, 1838
Pegomya steini Hendel, 1925
Checklist of the family Anthomyiidae (Diptera) of Finland 379
Pegomya tabida (Meigen, 1826)
Pegomya tenera (Zetterstedt, 1838)
Pegomya testacea (De Geer, 1776)
= silacea Meigen, 1830
Pegomya transgressa (Zetterstedt, 1846)
Pegomya vanduzeei Malloch, 1919
= versicolor auct. nec Meigen, 1826
Pegomya vittigera (Zetterstedt, 1838)
Pegomya winthemi (Meigen, 1826)
Pegomya zonata (Zetterstedt, 1838)
= tenera auct. nec Zetterstedt, 1838
PEGOPLATA Schnabl & Dziedzicki, 1911
= Nupedia Karl, 1930
Pegoplata aestiva (Meigen, 1826)
Pegoplata annulata (Pandellé, 1899)
= virginea auct. nec Meigen, 1826
Pegoplata inrma (Meigen, 1826)
Pegoplata nigroscutellata (Stein, 1920)
Pegoplata palposa (Stein, 1897)
Pegoplata patellans (Pandellé, 1900)
Pegoplata tundrica (Schnabl, 1915)
PHORBIA Robineau-Desvoidy, 1830
Phorbia atrogrisea Tiensuu, 1936
Phorbia curvicauda (Zetterstedt, 1845)
Phorbia fascicularis Tiensuu, 1936
Phorbia fumigata (Meigen, 1826)
= securis Tiensuu, 1936
Phorbia genitalis (Schnabl in Schnabl & Dziedzicki, 1911)
Phorbia longipilis (Pandellé, 1900)
Phorbia melania Ackland & Michelsen, 1987
Phorbia moliniaris (Karl, 1917)
Phorbia penicillaris (Stein, 1916)
Phorbia singularis Tiensuu, 1938
Ringdahlia Michelsen, 2014
Ringdahlia curtigena (Ringdahl, 1935)
STROBILOMYIA Michelsen, 1988
= Lasiomma auct. p. p.
Strobilomyia anthracina (Czerny, 1906)
Strobilomyia infrequens (Ackland, 1965)
Strobilomyia laricicola (Karl, 1928)
Strobilomyia sibirica Michelsen, 1988
Strobilomyia svenssoni Michelsen, 1988
Verner Michelsen / ZooKeys 441: 369–382 (2014)
380
SUBHYLEMYIA Ringdahl, 1933
Subhylemyia longula (Fallén, 1824)
ZAPHNE Robineau-Desvoidy, 1830
= Acroptena Pokorny, 1893
= Hydrophoria auct. p. p.
Zaphne ambigua (Fallén, 1823)
Zaphne barbiventris (Zetterstedt, 1845)
Zaphne brunneifrons (Zetterstedt, 1838)
Zaphne caudata (Zetterstedt, 1855)
Zaphne divisa (Meigen, 1826)
Zaphne fasciculata (Schnabl, 1915)
Zaphne frontata (Zetterstedt, 1838)
Zaphne ignobilis (Zetterstedt, 1845)
Zaphne inuncta (Zetterstedt, 1838)
= hyalipennis Zetterstedt, 1855
Zaphne lineatocollis (Zetterstedt, 1838)
= laticornis Ringdahl, 1916
Zaphne nuda (Schnabl in Schnabl & Dziedzicki, 1911)
Zaphne proxima (Malloch, 1920)
Zaphne verticina (Zetterstedt, 1838)
Zaphne wierzejskii (Mik, 1867)
Zaphne zetterstedtii (Ringdahl, 1918)
Excluded species
Botanophila rotundivalva (Ringdahl, 1937). Misdentication of closely related, still
undescribed species rst recorded from Ks by Frey (1956).
Delia angustiventris (Zetterstedt, 1845). Misdentication of Delia nudicosta (Ringdahl)
by Tiensuu (1936).
Delia oricola Robineau-Desvoidy, 1830. An erroneous record introduced by Hennig
(1974).
Delia penicillosa Hennig, 1974. An erroneous record introduced by Michelsen (2004).
Lasiomma multisetosum (Ringdahl, 1926). According to Tiensuu (1936) seen from
”nnischen Lappland” by O. Ringdahl, but no specimens have been found in the
Museum of Zoology, Lund University.
Pegomya glabroides Michelsen, 2008. Listed from Finland (Kuusamo: Kuolajärvi) by
Michelsen (2008), but Kuolajärvi (now Kuoloyarvi) is presently situated in Rus-
sian Murmansk Oblast close to the Finnish border.
Pegomya laticornis (Fallén, 1825) (= genupuncta Stein, 1906). Identication inferred
by Hendel (1922) from nds of leaf mines on Arctium by Linnaniemi (1913).
Supposed Pegomya-leaf mines on Arctium have been reported from Finland several
times since then, but no ies have been recovered so far.
Checklist of the family Anthomyiidae (Diptera) of Finland 381
Pegomya nigrisquama (Stein, 1888). Identication inferred by Hering (1925) from
nds of leaf-mines on Solidago virgaurea by Linnaniemi (1913).
Pegomya sociella Stein, 1906. Only recorded from Le: Kilpisjärvi (Tiensuu, 1933).
Probably a misidentication.
Phorbia bartaki Ackland & Michelsen, 1987. Records by Kahanpää (2013) refer to P.
melania Ackland & Michelsen.
Acknowledgements
I am grateful to Jere Kahanpää and Jukka Salmela for inviting me to compile the An-
thomyiidae part of the new Checklist of the Diptera of Finland. Antti Haarto, Jere Ka-
hanpää and Kaj Winqvist helped me with valuable information about Finnish records
of species and Finnish literature.
References
Frey R (1956) Samlingarnas tillväxt vid Helsingfors Universitets Entomologiska museum
under verksamhetsåret 1954−1955. Memoranda societatis pro fauna et ora fennica
31: 127−131.
Frey R, Tiensuu L, Storå R (1941) Enumeratio Insectorum Fenniae. VI. Diptera. Helsingin
hyönteisvaihtoyhdistys r.y., Helsinki, 63 pp.
Hackman W (1976) De som larver i hattsvampar levande anthomyiidernas biologi (Diptera).
Notulae entomologicae 56: 129–134.
Hackman W (1979) Pegomya pallidoscutellata (Zetterstedt) ny för Finland (Anthomyiidae).
Notulae entomologicae 59: 172–173.
Hackman W, Meinander M (1979) Diptera feeding as larvae on macrofungi in Finland. Annales
zoologici fennici 16: 50–83.
Hackman W (1980) A check list of the Finnish Diptera II. Cyclorrhapha. Notulae entomologicae
60: 117–162.
Hendel F (1922) Blattminierende Fliegen (Musciden). Wiener entomologische Zeitung 39: 65–72.
Hennig W (1966−1976) Anthomyiidae. In: Lindner E (Ed) Die Fliegen der palaearktischen
Region. E. Schweizerbart, Stuttgart, 7(2), 974 + 78 pp.
Hering M (1925) Minenstudien V. Zeitschrift für wissenschaftliche Insekten-Biologie 20:
125–136, 161–174.
Kahanpää J (2013) Viisikymmentä Suomelle uutta kärpäslajia (Diptera: Brachycera). Sahlbergia
19: 63–71.
Linnaniemi WM (1913) Zur Kenntnis der Blattminierer, speziell derjenigen Finnlands. I. Acta
societatis pro ora et fauna fennica 37(4): 1–137. [8 plates, 1 map]
Michelsen V (1985) A revision of the Anthomyiidae (Diptera) described by J. W. Zetterstedt.
Steenstrupia 11: 37−65.
Michelsen V (2004) Anthomyiidae. In: Pape T (Ed) Diptera Brachycera. Fauna Europaea, ver. 1.0.
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Michelsen V (2008) e Palaearctic species of the Pegomya terminalis species group (Diptera:
Anthomyiidae), with descriptions of two new species. Zootaxa 1781: 31–46.
Michelsen V (2011) Anthomyiidae. In: Pape T (Ed) Diptera Brachycera. Fauna Europaea, ver. 2.4.
[Current ver. 2.6.2 available from http://www.faunaeur.org]
Tiensuu L (1933) Tietoja Suomessa tavatuista sukakärpäsistä (Muscidae). Notulae entomologicae
13: 78–80.
Tiensuu L (1936) Die bisher aus Finnland bekannten Musciden. Acta societatis pro fauna et
ora fennica 58(4)[1935]: 1–56.
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)383
Checklist of the superfamilies Oestroidea and
Hippoboscoidea of Finland (Insecta, Diptera)
Jaakko Pohjoismäki1, Jere Kahanpää2
1 University of Eastern Finland, Department of Biology, P.O.Box 111, FI-80101 Joensuu, Finland 2 Finnish
Museum of Natural History, Zoology Unit, P.O. Box 17, FI-00014 University of Helsinki, Finland
Corresponding author: Jaakko Pohjoismäki (Jaakko.Pohjoismaki@uef.)
Academic editor: J. Salmela|Received 13 February 2014|Accepted 24 March 2014|Published 19 September 2014
urn:lsid:zoobank.org:pub:
Citation: Pohjoismäki J, Kahanpää J (2014) Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland
(Insecta, Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland. ZooKeys 441: 383–408. doi:
10.3897/zookeys.441.7252
Abstract
An updated checklist of the superfamilies Oestroidea and Hippoboscoidea recorded from Finland is pre-
sented. e checklist covers the following families: Calliphoridae, Rhiniidae, Sarcophagidae, Rhinophori-
dae, Tachinidae, Oestridae and Hippoboscidae.
Keywords
Checklist, Finland, Diptera, Oestroidea, Hippoboscoidea
Introduction
Oestroidea is a large and diverse superfamily of ies with a variety of life history strate-
gies ranging from saprophages to parasitoids or endoparasites, sometimes even within
the family. Some groups, such as certain subfamilies of the Calliphoridae, have also sig-
nicant veterinary, medical and economical impact. Oestroidea are regarded as mono-
phyletic clade, however the same does not apply to the family of Calliphoridae within
Oestroidea (Marinho et al. 2012). Besides Calliphoridae, Oestroidea currently includes
ZooKeys 441: 383–408 (2014)
doi: 10.3897/zookeys.441.7252
www.zookeys.org
Copyright J. Pohjoismäki, J. Kahanpää. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY
4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited..
CHECKLIST
Launched to accelerate biodiversity research
A peer-reviewed open-access journal
Jaakko Pohjoismäki & Jere Kahanpää / ZooKeys 441: 383–408 (2014)
384
six other monophyletic families: Rhiniidae, Sarcophagidae, Rhinophoridae, Tachini-
dae, Oestridae and the non-nnish Mystacinobiidae. Additionally, the subfamilies of
Oestridae (Oestrinae, Gasterophilinae) are sometimes treated as full families.
Hippoboscoidea contains the louse ies Hippoboscidae as well as the tropical
Glossinidae or tsetse ies. Hippoboscidae is divided to the mammal or bird parasitic
louse ies (Hippoboscinae) and baties (Nycterbiinae, Streblinae). Although baties
have been treated as full families, they seem to be subordinate to the other Hippobosci-
dae (Kutty et al. 2010). e checklist will therefore follow the family division of Pape
et al. 2011.
While world catalogues has been published for Sarcophagidae (Pape 1996) and
Hippoboscidae (Dick 2006) only regional (Herting and Dely-Draskovits 1993) or
genus-level (O’Hara 2012) catalogues exist for the other oestroid and hippoboscoid
families. e Finnish species of Oestroidea and Hippoboscoidea were last listed by
Hackman (1980). e sarcophagid checklist was updated by Pape (1987a,b) and the
calliphorids by Rognes (1991). e updated checklists for Oestridae, Rhinophoridae
and Tachinidae follow the taxonomic order of Pape (2001), Herting (1993) and Hert-
ing and Dely-Draskovits (1993), respectively.
Checklist part 1: Oestroidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Calyptratae Robineau-Desvoidy, 1830
superfamily Oestroidea Leach, 1815
CALLIPHORIDAE Brauer & Bergenstamm, 1889
Table 1. Number of species by family.
Family Number of species in Level of knowledge
World (Pape et al. 2011) Europe Finland
Oestroidea:
Calliphoridae 1525 102 45 good
Rhiniidae 376 12 0–1 good
Sarcophagidae 3094 309 64 average
Rhinophoridae 174 45 5 average
Tachinidae 9626 877 319 average
Oestridae 176 22 8 average
Hippoboscoidea:
Hippoboscidae s. lat. 782 45 12 average
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)385
CALLIPHORINAE Brauer & Bergenstamm, 1889
BELLARDIA Robineau-Desvoidy, 1863
= Pseudonesia Villeneuve, 1920
Bellardia bayeri (Jacentkovsky, 1937)
Bellardia pandia (Walker, 1849)
= biseta (Kramer, 1917)
Bellardia pubicornis (Zetterstedt, 1838)
= puberula (Zetterstedt, 1838)
Bellardia stricta (Villeneuve, 1926)
= polita auct. nec (Mik, 1883)
Bellardia viarum (Robineau-Desvoidy, 1830)
= pusilla (Meigen, 1826)
Bellardia vulgaris (Robineau-Desvoidy, 1830)
= agilis (Meigen, 1826) preocc.
CALLIPHORA Robineau-Desvoidy, 1830
= Steringomyia Pokorny, 1889
= Acrophaga Brauer & Bergenstamm, 1891
= Abonesia Villeneuve, 1927
= Acronesia Hall, 1948
Calliphora genarum (Zetterstedt, 1838)
= alpina (Zetterstedt, 1845)
Calliphora loewi Enderlein, 1903
Calliphora stelviana (Brauer & Bergenstamm, 1891)
Calliphora subalpina (Ringdahl, 1931)
Calliphora uralensis Villeneuve, 1922
Calliphora vicina Robineau-Desvoidy, 1830
= erythrocephala (Meigen, 1826) preocc.
Calliphora vomitoria (Linnaeus, 1758)
CYNOMYA Robineau-Desvoidy, 1830
Cynomya mortuorum (Linnaeus, 1761)
ONESIA Robineau-Desvoidy, 1830
Onesia oralis Robineau-Desvoidy, 1830
CHRYSOMYINAE Shannon, 1923
PROTOCALLIPHORA Hough, 1899
Protocalliphora azurea (Fallén, 1817)
Protocalliphora nuortevai Grunin, 1972
Protocalliphora peusi Gregor & Povolny, 1959
Protocalliphora proxima Grunin, 1966
Protocalliphora rognesi ompson & Pont, 1993
= chrysorrhoea Meigen, 1826 preocc.
PROTOPHORMIA Townsend, 1908
= Boreellus Aldrich & Schannon, 1923
Protophormia atriceps (Zetterstedt, 1845)
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386
Protophormia terraenovae (Robineau-Desvoidy, 1830)
TRYPOCALLIPHORA Peus, 1960
Trypocalliphora braueri (Hendel, 1901)
= lindneri Peus, 1960
HELICOBOSCINAE Verves, 1980
EURYCHAETA Brauer & Bergenstamm, 1891
= Helicobosca Bezzi, 1906
Eurychaeta palpalis (Robineau-Desvoidy, 1830)
LUCILIINAE Shannon, 1923
LUCILIA Robineau-Desvoidy, 1830
Lucilia ampullacea Villeneuve, 1922
Lucilia bufonivora Moniez, 1876
Lucilia caesar (Linnaeus, 1758)
Lucilia illustris (Meigen, 1826)
Lucilia magnicornis (Siebke, 1863)
= fuscipalpis (Zetterstedt, 1845)
Lucilia richardsi Collin, 1926
Lucilia sericata (Meigen, 1826)
Lucilia silvarum (Meigen, 1826)
MELANOMYINAE Townsend, 1919
ANGIONEURA Brauer & Bergenstamm, 1893
Angioneura acerba (Meigen, 1838)
MELANOMYA Rondani, 1856
Melanomya nana (Meigen, 1826)
MELINDA Robineau-Desvoidy, 1830
Melinda gentilis Robineau-Desvoidy, 1830
= coerulea (Meigen, 1826) preocc.
= cognata (Meigen, 1830)
Melinda viridicyanea (Robineau-Desvoidy, 1830)
POLLENIINAE Brauer & Bergenstamm, 1889
MORINIA Robineau-Desvoidy, 1830
Morinia doronici (Scopoli, 1763)
= melanoptera (Fallén, 1817)
= Angioneura mbriata misid.
POLLENIA Robineau-Desvoidy, 1830
Pollenia amentaria (Scopoli, 1763)
= vespillo auct. nec (Fabricius, 1794)
Pollenia angustigena Wainwright, 1940
Pollenia griseotomentosa (Jacentkowský, 1944)
Pollenia hungarica Rognes, 1987
Pollenia labialis Robineau-Desvoidy, 1863
= excarinata Wainwright, 1940
= intermedia misid.
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)387
Pollenia pediculata Macquart, 1834
Pollenia rudis (Fabricius, 1794)
= varia (Meigen, 1826)
Pollenia vagabunda (Meigen, 1826)
RHINIIDAE Brauer & Bergenstamm, 1889
STOMORHINA Rondani, 1861
Stomorhina lunata (Fabricius, 1805)
SARCOPHAGIDAE Macquart, 1834
MILTOGRAMMINAE Lioy, 1864
AMOBIA Robineau-Desvoidy, 1830
Amobia oculata (Zetterstedt, 1844)
= signata misid.
Amobia signata (Meigen, 1824)
HILARELLA Rondani, 1856
Hilarella hilarella (Zetterstedt, 1844)
MACRONYCHIA Rondani, 1859
= Macronichia orig. emend.
sg. Macronychia Rondani, 1859
Macronychia striginervis (Zetterstedt, 1838)
sg. Moschusa Robineau-Desvoidy, 1863
Macronychia agrestis (Fallén, 1820)
Macronychia griseola (Fallén, 1820)
Macronychia polyodon (Meigen, 1824)
METOPIA Meigen, 1803
Metopia argentata Macquart 1850
= roserii Rondani, 1859
= stackelbergi Rohdendorf, 1955
Metopia argyrocephala (Meigen, 1824)
= leucocephala (Rossi, 1790) preocc.
Metopia campestris (Fallén, 1810)
Metopia grandii Venturi, 1953
Metopia staegerii Rondani, 1859
= rondaniana Venturi, 1953
Metopia tshernovae Rohdendorf, 1955
MILTOGRAMMA Meigen, 1803
Miltogramma germari Meigen, 1824
Miltogramma iberica (Villeneuve, 1912)
Miltogramma oestracea (Fallén, 1820)
Miltogramma punctata Meigen, 1824
Miltogramma villeneuvei Verves, 1982
OEBALIA Robineau-Desvoidy, 1863
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Oebalia cylindrica (Fallén, 1810)
Oebalia minuta (Fallén, 1810)
PHROSINELLA Robineau-Desvoidy, 1863
Phrosinella sannio (Zetterstedt, 1838)
= nasuta misid.
PTERELLA Robineau-Desvoidy, 1830
Pterella grisea (Meigen, 1824)
SENOTAINIA Macquart, 1846
Senotainia albifrons (Rondani, 1859)
Senotainia conica (Fallén, 1810)
Senotainia puncticornis (Zetterstedt, 1859)
= imberbis (Zetterstedt, 1838) preocc.
= crabronum (Kramer, 1920)
TAXIGRAMMA Perris, 1852
Taxigramma elegantulum (Zetterstedt, 1844)
Taxigramma heteroneurum (Meigen, 1830)
PARAMACRONYCHIINAE Brauer & Bergenstamm, 1889
AGRIA Robineau-Desvoidy, 1830
Agria anis (Fallén, 1817)
= punctata Robineau-Desvoidy, 1830
Agria mamillata (Pandellé, 1896)
ANGIOMETOPA Brauer & Bergenstamm, 1889
Angiometopa falleni Pape, 1986
= ruralis (Fallén, 1817) preocc.
BRACHICOMA Rondani, 1856
Brachicoma devia (Fallén, 1820)
Brachicoma borealis Ringdahl, 1932
SARCOPHAGINAE Macquart, 1834
BLAESOXIPHA Loew, 1861
sg. Blaesoxipha Loew, 1861
Blaesoxipha lapidosa Pape, 1994
= agrestis auct. nec (Robineau-Desvoidy, 1863)
= campestris auct. nec (Robineau-Desvoidy, 1863)
Blaesoxipha laticornis (Meigen, 1826)
Blaesoxipha plumicornis (Zetterstedt, 1859)
= laticornis misid.
= gladiatrix (Pandellé, 1896)
sg. Servaisia Robineau-Desvoidy, 1863
Blaesoxipha erythrura (Meigen, 1826)
RAVINIA Robineau-Desvoidy, 1830
Ravinia pernix (Harris, 1780)
= striata (Fabricius, 1794) preocc.
SARCOPHAGA Meigen, 1826
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sg. Bellieriomima Rohdendorf, 1937
Sarcophaga subulata Pandellé, 1896
sg. Discacheata Enderlein, 1928
Sarcophaga pumila Meigen, 1826
sg. Helicophagella Enderlein, 1928
Sarcophaga crassimargo Pandellé, 1896
Sarcophaga melanura Meigen, 1826
Sarcophaga rosellei Böttcher, 1912
sg. Heteronychia Brauer & Bergenstamm, 1889
Sarcophaga depressifrons Zetterstedt, 1845
= ouscata auct. nec Meigen, 1826
Sarcophaga haemorrhoa Meigen, 1826
Sarcophaga proxima (Rondani, 1860)
Sarcophaga vagans (Meigen, 1826)
= frenata Pandellé, 1896
Sarcophaga vicina (Macquart, 1835)
sg. Liosarcophaga Enderlein, 1928
Sarcophaga emdeni (Rohdendor, 1969)
= teretirostris auct. nec Pandellé, 1896
Sarcophaga portschinskyi Rohdendorf, 1937
= tuberosa misid.
sg. Mehria Enderlein, 1928
Sarcophaga sexpunctata (Fabricius, 1805)
= clathrata (Meigen, 1826)
Sarcophaga nemoralis Kramer, 1908
sg. Myorhina Robineau-Desvoidy, 1830
= Pierretia Robineau-Desvoidy, 1863
Sarcophaga socrus Rondani, 1860
= rostrata Pandellé, 1896
Sarcophaga villeneuvei Böttcher, 1912
sg. Pandelleisca Rohdendorf, 1937
Sarcophaga similis Meade, 1876
sg. Parasarcophaga Johnston & Tiegs, 1921
Sarcophaga albiceps Meigen, 1826
= privigna Rondani, 1860
sg. Robineauella Enderlein, 1928
Sarcophaga caerulescens Zetterstedt, 1838
= scoparia Pandellé, 1896
sg. Rosellea Rohdendorf, 1937
Sarcophaga aratrix Pandellé, 1896
sg. Sarcophaga Meigen, 1826
Sarcophaga carnaria (Linnaeus, 1758)
= schulzi Müller, 1922
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= lehmanni auct. nec Müller, 1922
Sarcophaga lehmanni Müller, 1922
= lasiostyla auct. nec Macquart, 1843
= cognata Rondani, 1860 preocc.
Sarcophaga subvicina Rohdendorf, 1937
Sarcophaga variegata (Scopoli, 1763)
= carnaria auct. nec (Linnaeus, 1758)
sg. Sarcotachinella Townsend, 1892
Sarcophaga sinuata Meigen, 1826
sg. yrsocnema Enderlein, 1928
Sarcophaga incisilobata Pandellé, 1896
Sarcophaga kentejana (Rohdendorf, 1937)
= lapponica (Tiensuu, 1939)
RHINOPHORIDAE Robineau-Desvoidy, 1863
MELANOPHORA Meigen, 1803
Melanophora roralis (Linnaeus, 1758)
PAYKULLIA Robineau-Desvoidy, 1830
Paykullia brevicornis (Zetterstedt, 1844)
Paykullia maculata (Fallén, 1815)
STEVENIA Robineau-Desvoidy, 1830
Stevenia atramentaria (Meigen, 1824)
TRICOGENA Rondani, 1856
= Frauenfeldia Egger, 1865
Tricogena rubricosa (Meigen, 1824)
TACHINIDAE Robineau-Desvoidy, 1830
EXORISTINAE Robineau-Desvoidy, 1863
tribe Exoristini Robineau-Desvoidy, 1863
EXORISTA Meigen, 1803
sg. Adenia Robineau-Desvoidy, 1863
Exorista mimula (Meigen, 1824)
Exorista rustica (Fallén, 1810)
Exorista sp. A
sg. Exorista Meigen, 1803
Exorista fasciata (Fallén, 1820)
Exorista larvarum (Linnaeus, 1758)
sg. Podotachina Brauer & Bergenstamm, 1891
Exorista grandis (Zetterstedt, 1844)
= sorbillans auct. nec Wiedemann, 1830
sg. Ptilotachina Brauer & Bergenstamm 1891
Exorista deligata Pandellé, 1896
CHETOGENA Rondani, 1856
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Chetogena tschorsnigi Ziegler, 1999
DIPLOSTICHUS Brauer & Bergenstamm, 1889
Diplostichus janitrix (Hartig, 1838)
PHOROCERA Brauer & Bergenstamm, 1889
Phorocera assimilis (Fallén, 1810)
Phorocera obscura (Fallén, 1810)
PHORINIA Robineau-Desvoidy, 1830
Phorinia aurifrons Robineau-Desvoidy, 1830
BESSA Robineau-Desvoidy, 1830
Bessa selecta (Meigen, 1824)
tribe Blondeliini Robineau-Desvoidy, 1863
BELIDA Robineau-Desvoidy, 1863
= Aporotachina Meade, 1894
Belida angelicae (Meigen, 1824)
MEIGENIA Robineau-Desvoidy, 1830
Meigenia dorsalis (Meigen, 1824)
Meigenia mutabilis (Fallén, 1810)
= bisignata (Meigen, 1824)
ZAIRA Robineau-Desvoidy, 1830
= Viviania Rondani, 1861
Zaira cinerea (Fallén, 1810)
MEDINA Robineau-Desvoidy, 1830
Medina collaris (Fallén, 1820)
Medina luctuosa (Meigen, 1824)
Medina separata (Meigen, 1824)
POLICHETA Rondani, 1856
= Perichaeta preocc.
Policheta unicolor (Fallén, 1820)
ISTOCHETA Rondany, 1859
= Hyperecteina Schiner, 1862
Istocheta longicornis (Fallén, 1810)
STAUROCHAETA Brauer & Bergenstamm, 1889
Staurochaeta albocingulata (Fallén, 1820)
ADMONTIA Brauer & Bergenstamm, 1889
= Trichopareia Brauer & Bergenstamm, 1889
Admontia blanda (Fallén, 1820)
Admontia grandicornis (Zetterstedt, 1849)
Admontia seria (Meigen, 1824)
OSWALDIA Robineau-Desvoidy, 1863
Oswaldia muscaria (Fallén, 1810)
Oswaldia eggeri (Brauer & Bergenstamm, 1889)
Oswaldia reducta (Villeneuve, 1930)
Oswaldia spectabilis (Meigen, 1824)
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= albisquama (Zetterstedt, 1844)
PARACRASPEDOTHRIX Villeneuve, 1919
Paracraspedothrix montivaga Villeneuve, 1919
LIGERIA Robineau-Desvoidy, 1863
Ligeria angusticornis (Loew, 1847)
= zetterstedti (Ringdahl, 1945)
LIGERIELLA Mesnil, 1961
Ligeriella aristata (Villeneuce, 1911)
BLONDELIA Robineau-Desvoidy, 1830
Blondelia nigripes (Fallén, 1810)
tribe Acemyini Brauer & Bergenstamm, 1889
ACEMYA Robineau-Desvoidy, 1830
Acemya acuticornis (Meigen, 1824)
Acemya rutibia (von Roser, 1840)
tribe Ethillini Mesnil, 1944
PARATRYPHERA Brauer & Bergenstamm, 1891
Paratryphera barbatula (Rondani, 1859)
Paratryphera bisetosa (Brauer & Bergenstamm, 1891)
tribe Winthemiini Townsend, 1913
RHAPHIOCHAETA Brauer & Bergenstamm, 1889
Rhaphiochaeta breviseta (Zetterstedt, 1838)
SMIDTIA Robineau-Desvoidy, 1830
= Timavia Robineau-Desvoidy, 1863
Smidtia amoena (Meigen, 1824)
Smidtia conspersa (Meigen, 1824)
WINTHEMIA Robineau-Desvoidy, 1830
Winthemia cruentata (Rondani, 1859)
Winthemia erythrura (Meigen, 1838)
Winthemia quadripustulata (Fabricius, 1794)
Winthemia venusta (Meigne, 1824)
NEMORILLA Rondani, 1856
Nemorilla oralis (Fallen, 1810)
Nemorilla maculosa (Meigen, 1824)
= oralis misid.
tribe Eryciini Robineau-Desvoidy, 1830
APLOMYA Robineau-Desvoidy, 1830
Aplomya connis (Fallén, 1820)
PHEBELLIA Robineau-Desvoidy, 1846
Phebellia clavellariae (Brauer & Bergenstamm, 1891)
Phebellia glauca (Meigen, 1824)
Phebellia glaucoides Herting, 1961
Phebellia glirina (Rondani, 1859)
Phebellia margaretae Bergström, 2005
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Phebellia nigripalpis (Robineau-Desvoidy, 1847)
= fuscipennis (Robineau-Desvoidy, 1830)
Phebellia pauciseta (Villeneuve, 1908)
Phebellia strigifrons (Zetterstedt, 1838)
Phebellia stulta (Zetterstedt, 1844)
Phebellia triseta (Pandellé, 1896)
Phebellia vicina Wainwright, 1940
Phebellia villica (Zetterstedt, 1838)
Phebellia sp. A
NILEA Robineau-Desvoidy, 1863
Nilea hortulana (Meigen, 1824)
Nilea innoxia Robineau-Desvoidy, 1863
Nilea ruscutellaris (Zetterstedt, 1859)
TLEPHUSA Robineau-Desvoidy, 1863
Tlephusa cincinna (Meigen, 1859)
= diligens auct. nec (Zetterstedt, 1844)
EPICAMPOCERA Macquart, 1849
Epicampocera succincta (Meigen, 1824)
PHRYXE Robineau-Desvoidy, 1830
Phryxe erythrostoma (Hartig, 1838)
Phryxe magnicornis (Zetterstedt, 1838)
Phryxe nemea (Meigen, 1824)
Phryxe vulgaris (Fallén, 1810)
PERIARCHICLOPS Villeneuve, 1924
Periarchiclops scutellaris (Fallén, 1820)
BACTROMYIA Brauer & Bergenstamm, 1891
Bactromyia aurulenta (Meigen, 1824)
= ecocarcelia acutangula misid.
PSEUDOPERICHAETA Brauer & Bergenstamm, 1899
Pseudoperichaeta nigrolineata (Stephens, 1853)
LYDELLA Robineau-Desvoidy, 1830
Lydella ripae (Brischke, 1885)
= grisescens misid.
= lepida misid.
Lydella stabulans (Meigen, 1824)
CADURCIELLA Villeneuve, 1927
Cadurciella tritaeniata (Rondani, 1859)
DRINO Robineau-Desvoidy, 1863
Drino bohemica Mesnil, 1949
Drino galii (Brauer & Bergenstamm, 1891)
Drino gilva (Hartig, 1838)
Drino inconspicua (Meigen, 1830)
Drino lota (Meigen, 1824)
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Drino vicina (Zetterstedt, 1849)
HUBNERIA Robineau-Desvoidy, 1848
= Huebneria orig. emend.
Hubneria anis (Fallén, 1810)
CARCELIA Robineau-Desvoidy, 1830
sg. Carcelia Robineau-Desvoidy, 1830
Carcelia atricosta Herting, 1961
= bombylans misid.
Carcelia bombylans Robineau-Desvoidy, 1830
Carcelia gnava (Meigen, 1824)
= excavata (Zetterstedt, 1844)
Carcelia laxifrons Villeneuve, 1912
Carcelia lucorum (Meigen, 1824)
Carcelia rasa (Macquart, 1849)
= amphion Robineau-Desvoidy, 1863
Carcelia tibialis (Robineau-Desvoidy, 1863)
SENOMETOPIA Macquart, 1834
Senometopia excisa (Fallén, 1820)
Senometopia intermedia (Herting, 1960)
Senometopia pollinosa Mesnil, 1941
= rutilla auct. nec (Villeneuve, 1912)
= obesa auct. nec (Zetterstedt, 1859)
Senometopia separata (Rondani, 1859)
= bombycivora auct. nec (Robineau-Desvoidy, 1830)
ERYCIA Robineau-Desvoidy, 1830
Erycia fatua (Meigen, 1824)
= festinans misid.
Erycia furibunda (Zetterstedt, 1844)
XYLOTACHINA Brauer & Bergenstamm, 1891
Xylotachina diluta (Meigen, 1824)
tribe Goniini Lioy, 1864
PLATYMYA Robineau-Desvoidy, 1830
Platymya mbriata (Meigen, 1824)
EUMEA Robineau-Desvoidy, 1863
= Platymya Robineau-Desvoidy, 1830 in part
Eumea linearicornis (Zetterstedt, 1844)
= westermanni (Zetterstedt, 1844)
Eumea mitis (Meigen, 1824)
MYXEXORISTOPS Townsend, 1911
Myxexoristops abietis Herting, 1964
Myxexoristops arctica (Zetterstedt, 1838)
Myxexoristops bonsdor (Zetterstedt, 1859)
Myxexoristops stolida (Stein, 1924)
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)395
= blondeli misid.
ZENILLIA Robineau-Desvoidy, 1830
Zenillia fulva (Fallén, 1820)
= libatrix (Panzer, 1798) preocc.
CLEMELIS Robineau-Desvoidy, 1863
Clemelis pullata (Meigen, 1824)
PALES Robineau-Desvoidy, 1830
Pales pavida (Meigen, 1824)
CYZENIS Robineau-Desvoidy, 1863
Cyzenis albicans (Fallén, 1810)
Cyzenis jucunda (Meigen, 1838)
BOTRIA Rondani, 1856
= Bothria orig. emend.
Botria frontosa (Meigen, 1824)
Botria subalpina Villeneuve, 1910
CEROMASIA Rondani, 1856
Ceromasia rubrifrons (Macquart, 1834)
ALLOPHOROCERA Hendel, 1901
= Erycilla Mesnil, 1957
Allophorocera ferruginea (Meigen, 1824)
Allophorocera lapponica Wood, 1974
OCYTATA Gistel, 1848
Ocytata pallipes (Fallén, 1820)
ERYNNIA Robineau-Desvoidy, 1830
Erynnia ocypterata (Fallén, 1810)
= nitida (Robineau-Desvoidy, 1830)
ELODIA Robineau-Desvoidy, 1863
Elodia ambulatoria (Meigen, 1824)
= convexifrons (Zetterstedt, 1844)
STURMIA Robineau-Desvoidy, 1830
Sturmia bella (Meigen, 1824)
BLEPHARIPA Rondani, 1856
Blepharipa pratensis (Meigen, 1824)
PROSOPEA Rondani, 1861
Prosopea nigricans (Egger, 1861)
HEBIA Robineau-Desvoidy, 1830
Hebia avipes Robineau-Desvoidy, 1830
FRONTINA Meigen, 1838
Frontina laeta (Meigen, 1824)
BRACHICHETA Rondani, 1861
Brachicheta strigata (Meigen, 1824)
GONIA Meigen, 1803
Gonia capitata (De Geer, 1776)
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Gonia divisa Meigen, 1826
Gonia ornata Meigen, 1826
Gonia picea (Robineau-Desvoidy, 1830)
= sicula auct. nec (Robineau-Desvoidy, 1830)
ONYCHOGONIA Brauer & Bergenstamm, 1889
Onychogonia cervini (Bigot, 1881)
Onychogonia aviceps (Zetterstedt, 1838)
= interrupta (Rondani, 1859)
SPALLANZANIA Robineau-Desvoidy, 1830
Spallanzania hebes (Fallén, 1820)
TACHININAE Robineau-Desvoidy, 1830
tribe Polideini Brauer & Bergenstamm, 1889
LYDINA Robineau-Desvoidy, 1830
Lydina aenea (Meigen, 1824)
LYPHA Robineau-Desvoidy, 1830
Lypha dubia (Fallén, 1810)
Lypha rucauda (Zetterstedt, 1838)
tribe Tachinini Robineau-Desvoidy, 1830
TACHINA Meigen, 1803
= Larvaevora Meigen, 1800 suppr.
sg. Eudoromyia Bezzi, 1906
Tachina fera (Linnaeus, 1761)
Tachina sp. a. magnicornis
Tachina magnicornis (Zetterstedt, 1844)
sg. Servillia Robineau-Desvoidy, 1830
Tachina ursina (Meigen, 1824)
sg. Tachina Meigen, 1803
Tachina grossa (Linnaeus, 1758)
NOWICKIA Wachtl, 1894
Nowickia alpina (Zetterstedt, 1844)
Nowickia ferox (Panzer, 1809)
Nowickia marklini (Zetterstedt, 1838)
PELETERIA Robineau-Desvoidy, 1830
= Cuphocera Macquart, 1845
Peleteria ferina (Zetterstedt, 1844)
Peleteria rubescens Robineau-Desvoidy, 1830
= nigricornis (Meigen, 1838)
Peleteria rucornis (Macquart, 1835)
GERMARIA Robineau-Desvoidy, 1830
= Atractochaeta Brauer & Bergenstamm, 1889
Germaria angustata (Zetterstedt, 1844)
Germaria ruceps (Fallén, 1820)
tribe Linnaemyini Townsend, 1919
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LINNAEMYA Robineau-Desvoidy, 1830
Linnaemya haemorrhoidalis (Fallén, 1810)
Linnaemya olsuevi Zimin, 1954
= haemorrhoidalis misid.
Linnaemya rossica Zimin, 1954
= haemorrhoidalis misid.
Linnaemya vulpina (Fallén, 1810)
Linnaemya sp. A
CHRYSOSOMOPSIS Townsend, 1916
Chrysosomopsis aurata (Fallén, 1820)
PANZERIA Robineau-Desvoidy, 1830
= Ernestia Robineau-Desvoidy, 1830
Panzeria puparum (Fabricius, 1794)
Panzeria rudis (Fallén, 1810)
Panzeria vagans (Meigen, 1824)
APPENDICIA Stein, 1924
Appendicia truncata (Zetterstedt, 1838)
EURITHIA Robineau-Desvoidy, 1844
Eurithia anthophila (Robineau-Desvoidy, 1830)
= radicum auct. nec Linnaeus, 1758
Eurithia caesia (Fallén, 1810)
Eurithia connivens (Zetterstedt, 1844)
Eurithia consobrina (Meigen, 1824)
Eurithia intermedia (Zetterstedt, 1844)
= conjugata (Zetterstedt, 1855)
Eurithia vivida (Zetterstedt, 1838)
HYALURGUS Brauer & Bergenstamm, 1893
Hyalurgus crucigerus (Zetterstedt, 1838)
Hyalurgus lucidus (Meigen, 1824)
NEMORAEA Robineau-Desvoidy, 1830
Nemoraea pellucida (Meigen, 1824)
GYMNOCHETA Robineau-Desvoidy, 1830
Gymnocheta magna (Fallén, 1810)
Gymnocheta viridis (Fallén, 1810)
ZOPHOMYIA Macquart, 1835
Zophomyia temula (Scopoli, 1763)
CLEONICE Robineau-Desvoidy, 1863
= Steiniella Berg, 1898 preocc.
Cleonice callida (Meigen, 1824)
Cleonice keteli Ziegler, 2000
Cleonice nitidiuscula (Zetterstedt, 1859)
LOEWIA Egger, 1856
= Fortisia Rondani, 1861
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Loewia erecta Bergström, 2007
= phaeoptera misid.
Loewia foeda (Meigen, 1824)
ELOCERIA Robineau-Desvoidy, 1863
Eloceria delecta (Meigen, 1824)
tribe Brachymerini Mesnil, 1939
PSEUDOPACHYSTYLUM Mik, 1891
Pseudopachystylum gonioides (Zetterstedt, 1838)
tribe Pelatachinini Mesnil, 1966
PELATACHINA Meade, 1894
Pelatachina tibialis (Fallén, 1810)
tribe Macquartiini Robineau-Desvoidy, 1830
MACQUARTIA Robineau-Desvoidy, 1830
Macquartia dispar (Fallén, 1820)
Macquartia nudigena Mesnil, 1972
= buccalis auct. nec Robineau-Desvoidy, 1830
Macquartia tenebricosa (Meigen, 1824)
Macquartia viridana Robineau-Desvoidy, 1863
ANTHOMYIOPSIS Townsend, 1916
= Ptilopsina Villeneuve, 1920
Anthomyiopsis nigrisquamata (Zetterstedt, 1838)
tribe Triarthriini Belshaw, 1993
TRIARTHRIA Stephens, 1829
Triarthria setipennis (Fallén, 1810)
tribe Neaerini Mesnil, 1966
PHYTOMYPTERA Rondani, 1845
= Ela Robineau-Desvoidy, 1850
Phytomyptera bohemica Kramer, 1907
Phytomyptera cingulata (Robineau-Desvoidy, 1830)
Phytomyptera nigrina (Meigen, 1824)
Phytomyptera minutissima (Zetterstedt, 1844)
Phytomyptera riedeli (Villeneuve, 1930)
Phytomyptera zonella (Zetterstedt, 1844)
GRAPHOGASTER Rondani, 1868
= Anurogyna Brauer & Bergenstamm, 1889
Graphogaster brunnescens Villeneuve, 1907
Graphogaster buccata Herting, 1971
Graphogaster dispar (Brauer & Bergenstamm, 1889)
= punctiventris (Ringdahl, 1942)
Graphogaster nigrescens Herting, 1971
tribe Siphonini Rondani, 1844
ENTOMOPHAGA Lioy, 1864
Entomophaga nigrohalterata (Villeneuve, 1921)
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Entomophaga suerta (Villeneuve, 1942)
CEROMYA Robineau-Desvoidy, 1830
Ceromya bicolor (Meigen, 1824)
Ceromya dorsigera Herting, 1967
Ceromya aviceps Ratzeburg, 1844
Ceromya silacea (Meigen, 1824)
ACTIA Robineau-Desvoidy, 1830
Actia crassicornis (Meigen, 1824)
Actia infantula (Zetterstedt, 1844)
Actia lamia (Meigen, 1838)
Actia maksymovi Mesnil, 1952
Actia nigroscutellata Lundbeck, 1927
Actia pilipennis (Fallén, 1810)
Actia resinellae (Schrank, 1781)
= nudibasis Stein, 1924
PERIBAEA Robineau-Desvoidy, 1863
Peribaea hertingi Andersen, 1996
= ssicornis misid.
Peribaea longirostris Andersen, 1996
CERANTHIA Robineau-Desvoidy, 1830
Ceranthia abdominalis Robineau-Desvoidy, 1830
Ceranthia lichtwardtiana (Villeneuve, 1931)
Ceranthia pallida Herting, 1959
Ceranthia tenuipalpis (Villeneuve, 1921)
Ceranthia tristella Herting, 1966
Ceranthia verneri Andersen, 1996
APHANTORHAPHOPSIS Townsend, 1926
Aphantorhaphopsis samarensis (Villeneuve, 1921)
Aphantorhaphopsis siphonoides (Strobl, 1898)
Aphantorhaphopsis verralli (Wainwright, 1928)
SIPHONA Meigen, 1803
Siphona boreata Mesnil, 1960
Siphona collini Mesnil, 1960
Siphona confusa Mesnil, 1961
= mesnili Andersen, 1982
Siphona cristata (Fabricius, 1805)
= geniculata auct. nec (De Geer, 1776)
Siphona avifrons (Staeger, 1849)
Siphona geniculata (De Geer, 1776)
= urbana (Harris, 1776)
Siphona grandistyla Pandellé, 1894
Siphona immaculata Andersen, 1996
Siphona ingerae Andersen, 1982
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Siphona hokkaidensis Mesnil, 1957
= silvarum Herting, 1967
= martini Andersen, 1982
Siphona maculata (Staeger, 1849)
Siphona nigricans (Villeneuve, 1930)
Siphona paludosa Mesnil, 1960
Siphona pauciseta Rondani, 1865
Siphona rossica Mesnil, 1961
Siphona setosa Mesnil, 1960
Siphona subarctica Andersen, 1996
Siphona variata Andersen, 1982
tribe Leskiini Townsend, 1919
APHRIA Robineau-Desvoidy, 1830
Aphria longilingua Rondani, 1861
Aphria longirostris (Meigen, 1824)
DEMOTICUS Macquart, 1854
Demoticus plebejus (Fallén, 1810)
LESKIA Robineau-Desvoidy, 1830
Leskia aurea (Fallén, 1820)
SOLIERIA Robineau-Desvoidy, 1848
Solieria inanis (Fallén, 1810)
Solieria pacica (Meigen, 1824)
= tibialis (von Roser, 1840)
tribe Minthonini Brauer & Bergenstamm, 1889
MINTHO Robineau-Desvoidy, 1830
Mintho ruventris (Fallén, 1816)
MINTHODES Brauer & Bergenstamm, 1899
Minthodes picta (Zetterstedt, 1844)
tribe Microphthalmini Crosskey, 1976
DEXIOSOMA Rondani, 1856
Dexiosoma caninum (Fabricius, 1794)
DEXIINAE Macquart, 1834
tribe Dexiini Macquart, 1834
TRIXA Meigen, 1824
= Murana Meigen, 1824
Trixa caerulescens Meigen, 1824
= alpina Meigen, 1824
Trixa conspersa (Harris, 1776)
= variegata Meigen, 1824
= oestroidea (Robineau-Desvoidy, 1830)
BILLAEA Robineau-Desvoidy, 1830
Billaea irrorata (Meigen, 1826)
Billaea kolomyetzi Mesnil, 1970
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)401
Billaea triangulifera (Zetterstedt, 1844)
DINERA Robineau-Desvoidy, 1830
Dinera ferina (Fallén, 1816)
Dinera grisescens (Fallén, 1816)
Dinera carinifrons (Fallén, 1817)
ESTHERIA Robineau-Desvoidy, 1830
Estheria petiolata (Bonsdor, 1866)
DEXIA Meigen, 1826
Dexia vacua (Fallén, 1816)
PROSENA Le Peletier & Serville, 1828
Prosena siberita (Fabricius, 1775)
tribe Voriini Townsend, 1912
ERIOTHRIX Meigen, 1803
Eriothrix argyreatus (Meigen, 1824)
= apenninus misid.
Eriothrix prolixa (Meigen, 1824)
Eriothrix rufomaculata (De Geer, 1776)
= monochaeta (Wainwright, 1928)
TRAFOIA Brauer & Bergenstamm, 1893
Trafoia monticola Brauer & Bergenstamm, 1893
CAMPYLOCHETA Rondani, 1859
= Elpe Robineau-Desvoidy, 1863
Campylocheta fuscinervis (Stein, 1924)
Campylocheta inepta (Meigen, 1824)
Campylocheta praecox (Meigen, 1824)
BLEPHAROMYIA Brauer & Bergenstamm, 1889
Blepharomyia angustifrons Herting, 1971
Blepharomyia pagana (Meigen, 1824)
= amplicornis (Zetterstedt, 1844)
Blepharomyia piliceps (Zetterstedt, 1859)
PETEINA Meigen, 1838
Peteina erinaceus (Fabricius, 1794)
RAMONDA Robineau-Desvoidy, 1863
Ramonda prunaria (Rondani, 1861)
= carbonaria misid.
Ramonda ringdahli (Villeneuve, 1922)
Ramonda spathulata (Fallén, 1820)
WAGNERIA Robineau-Desvoidy, 1830
Wagneria alpina (Villeneuve, 1910)
Wagneria costata (Fallén, 1820)
ATHRYCIA Robineau-Desvoidy, 1830
= Blepharigena Rondani, 1856
Athrycia curvinervis (Zetterstedt, 1844)
Jaakko Pohjoismäki & Jere Kahanpää / ZooKeys 441: 383–408 (2014)
402
Athrycia impressa (van der Wulp, 1869)
Athrycia trepida (Meigen, 1824)
Athrycia sp. A
VORIA Robineau-Desvoidy, 1830
= Plagia Meigen, 1838
Voria ruralis (Fallén, 1810)
CYRTOPHLEBA Rondani, 1856
Cyrtophleba ruricola (Meigen, 1824)
Cyrtophleba vernalis (Kramer, 1917)
KLUGIA Robineau-Desvoidy, 1863
Klugia marginata (Meigen, 1824)
CHAETOVORIA Villeneuve, 1920
Chaetovoria antennata Villeneuve, 1920
PHYLLOMYA Robineau-Desvoidy, 1830
Phyllomya volvulus (Fabricius, 1794)
THELAIRA Robineau-Desvoidy, 1830
elaira nigrina (Fallén, 1817)
= nigripes (Fabricius, 1794) preocc.
HALIDAYA Egger, 1856
Halidaya aurea Egger, 1856
STOMINA Robineau-Desvoidy, 1830
Stomina tachinoides (Fallén, 1816)
tribe Dufouriini Robineau-Desvoidy, 1830
DUFOURIA Robineau-Desvoidy, 1830
Dufouria chalybeata (Meigen, 1824)
Dufouria nigrita (Fallén, 1810)
RONDANIA Robineau-Desvoidy, 1850
Rondania dimidiata (Meigen, 1824)
Rondania dispar (Dufour, 1851)
Rondania fasciata (Macquart, 1834)
MICROSOMA Macquart, 1855
= Campogaster Rondani, 1856
Microsoma exiguum (Meigen, 1824)
FRERAEA Robineau-Desvoidy, 1830
Freraea gagatea Robineau-Desvoidy, 1830
= albipennis (Zetterstedt, 1838)
PHASIINAE Robineau-Desvoidy, 1830
tribe Phasiini Robineau-Desvoidy, 1830
CLYTIOMYA Rondani, 1861
Clytiomya continua (Panzer, 1798)
ELIOZETA Rondani, 1856
Eliozeta pellucens (Fallén, 1820)
SUBCLYTIA Pandellé, 1894
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)403
Subclytia rotundiventris (Fallén, 1820)
GYMNOSOMA Meigen, 1803
Gymnosoma clavatum (Rohdendorf, 1947)
Gymnosoma dolycoridis Dupuis, 1961
Gymnosoma nudifrons Herting, 1966
CISTOGASTER Latreille, 1804
= Pallasia Robinaeu-Desvoidy, 1830
Cistogaster globosa (Fabricius, 1775)
PHASIA Latreille, 1804
= Alophora Robineau-Desvoidy, 1830
sg. Hyalomya Robineau-Desvoidy, 1830
Phasia pusilla (Meigen, 1824)
sg. Phasia Latreille, 1804
Phasia aurulans (Meigen, 1824)
Phasia barbifrons (Girschner, 1887)
Phasia hemiptera (Fabricius, 1794)
Phasia obesa (Fabricius, 1798)
Phasia subcoleoptrata (Linnaeus, 1767)
tribe Catharosiini Townsend, 1936
CATHAROSIA Rondani, 1868
Catharosia pygmaea (Fallén, 1815)
tribe Strongygastrini Townsend, 1936
STRONGYGASTER Macquart, 1834
= Tamiclea Macquart, 1836
Strongygaster celer (Meigen, 1838)
tribe Leucostomatini Townsend, 1908
LEUCOSTOMA Meigen, 1803
Leucostoma simplex (Fallén, 1815)
BRULLAEA Robineau-Desvoidy, 1863
Brullaea ocypteroidea Robineau-Desvoidy, 1863
CINOCHIRA Zetterstedt, 1845
Cinochira atra Zetterstedt, 1945
tribe Cylindromyiini Townsend, 1912
LOPHOSIA Meigen, 1824
sg. Lophosia Meigen, 1824
Lophosia fasciata Meigen, 1824
CYLINDROMYIA Meigen, 1803
sg. Cylindromyia Meigen, 1803
Cylindromyia brassicaria (Fabricius, 1775)
sg. Neocyptera Townsend, 1916
Cylindromyia interrupta (Meigen, 1824)
sg. Ocypterula Rondani, 1856
Cylindromyia pusilla (Meigen, 1824)
Jaakko Pohjoismäki & Jere Kahanpää / ZooKeys 441: 383–408 (2014)
404
HEMYDA Robineau-Desvoidy, 1830
Hemyda obscuripennis (Meigen, 1824)
Hemyda vittata (Meigen, 1824)
BESSERIA Robineau-Desvoidy, 1830
Besseria anthophila (Loew, 1871)
Besseria melanura (Meigen, 1824)
PHANIA Meigen, 1824
Phania curvicauda (Fallén, 1820)
Phania funesta (Meigen, 1824)
= pseudofunesta (Villeneuve, 1931)
Phania thoracica Meigen, 1824
OESTRIDAE Leach, 1815
HYPODERMATINAE Rondani, 1856
HYPODERMA Latreille, 1818
Hypoderma bovis (Linnaeus, 1758)
Hypoderma lineatum (De Villers, 1789)
Hypoderma tarandi (Linnaeus, 1758)
CEPHENEMYIINAE Patton, 1921
tribe Cephenemyiini Patton, 1921
CEPHENEMYIA Latreille, 1818
Cephenemyia trompe (Modeer, 1786)
Cephenemyia ulrichii Brauer, 1862
GASTEROPHILINAE Girschner, 1896
GASTEROPHILUS Leach, 1817
Gasterophilus haemorrhoidalis (Linnaeus, 1758)
Gasterophilus intestinalis (De Geer, 1776)
Gasterophilus nasalis (Linnaeus, 1758)
Checklist part 2: Hippoboscoidea
suborder Brachycera Macquart, 1834
clade Eremoneura Lameere, 1906
clade Cyclorrhapha Brauer, 1863
infraorder Schizophora Becher, 1882
clade Muscaria Enderlein, 1936
parvorder Calyptratae Robineau-Desvoidy, 1830
superfamily Hippoboscoidea Samouelle, 1819
HIPPOBOSCIDAE Samouelle, 1819
HIPPOBOSCINAE Samouelle, 1819
tribe Ornithomyini Costa, 1846
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)405
CRATAERINA von Olfers, 1816
= Stenepteryx Leach, 1817
Crataerina hirundinis (Linnaeus, 1758)
Crataerina pallida (Olivier, 1812)
tribe Olfersiini Maa, 1969
OLFERSIA Wiedemann, 1830
= Feronia Leach, 1817 preocc.
Olfersia fumipennis (Sahlberg, 1886)
ORNITHOMYA Latreille, 1802
Ornithomya avicularia (Linnaeus, 1758)
Ornithomya chloropus Bergroth, 1901
Ornithomya fringillina Curtis, 1836
ORNITHOPHILA Rondani, 1879
Ornithophila metallica (Schiner, 1864)
tribe Hippoboscini Samouelle, 1819
HIPPOBOSCA Linnaeus, 1758
Hippobosca equina Linnaeus, 1758
tribe Lipoptenini Speiser, 1908
LIPOPTENA Nitzsch, 1818
Lipoptena cervi (Linnaeus, 1758)
MELOPHAGUS Latreille, 1802
Melophagus ovinus (Linnaeus, 1758)
NYCTERIBIINAE Samouelle, 1819
tribe Nycteribiini Samouelle, 1819
NYCTERIBIA Latreille, 1796
sg. Nycteribia Latreille, 1796
Nycteribia kolenatii eodor & Moscona, 1954
= pedicularia auct. nec Latreille, 1805
= latreillei misid.
= blasii (Kolenati, 1863) preocc.
PENICILLIDIA Kolenati, 1863
Penicillidia monoceros Speiser, 1900
Excluded species
Aphantorhaphopsis (Ceranthia) selecta (Pandellé, 1894) misidentied
Baumhaueria goniaeformis (Meigen, 1824) misidentied
Bithia geniculata (Zetterstedt, 1844) see Notes
Blepharipoda scutellata (Robineau-Desvoidy, 1830) misidentied
Blondelia piniariae (Hartig, 1838). Specimens reared from the correct host are mentioned
in the notes of Lauri Tiensuu. See Bergström and Bystrowski (2011) for a recent
revalidation of this species.
Jaakko Pohjoismäki & Jere Kahanpää / ZooKeys 441: 383–408 (2014)
406
Billaea fortis (Rondani, 1862) a likely misidentication, the specimen cannot be located
Dexia rustica (Fallén, 1816) misidentied
Drino atropivora Robineau-Desvoidy, 1830 not found within present borders
Entomophaga exoleta (Meigen, 1824) misidentied
Graphogaster vestita Rondani, 1868 misidentied
Gymnosoma rotundatum (Linnaeus, 1758) misidentied
Istocheta subcinerea (Borisova-Zinoveva, 1966) misidentied
Leiophora innoxia (Meigen, 1824) specimen lost, but likely a misidentication due to
the host record (Diprion sp. Symphyta: Diprionidae)
Linnaemya comta (Fallén, 1810) misidentied
Lydella grisescens Robineau-Desvoidy, 1830 misidentied
Macronychia alpestris Rondani, 1865 labeling mistake
Macronychia conica Robineau-Desvoidy, 1830 labeling mistake
Macroprosopa atrata (Fallén, 1810) misidentied
Masicera silvatica (Fallén, 1810) misidentied
Oestrus ovis Linnaeus, 1758 extinct?
Opesia cana (Meigen, 1824) not found within present borders
Pseudogonia rufrons (Wiedemann, 1830) imported
Pseudoperichaeta palesoidea (Robineau-Desvoidy, 1830) misidentied
Sarcophaga agnata Rondani, 1860 misidentied
Sarcophaga dissimilis Meigen, 1826 misidentied
Sarcophaga granulata Kramer, 1908
Sarcophila latifrons (Fallén, 1817) not found within present borders
Senometopia confundens Rondani, 1859 misidentied
elymorpha marmorata (Fabricius, 1805), record cannot be conrmed – specimen lost?
Notes
Athrycia sp. A. Probably an undescribed species (J. Flinck leg., T. Zeegers priv. comm.)
Bithia geniculata (Zetterstedt, 1844). Erroneously included in the previous checklist
(Hackman 1980, 1981) on the basis of misintepreted records of Siphona geniculata
(De Geer, 1776).
Dexia vacua (Fallén, 1816) and Dinera carinifrons (Fallén, 1817). Conspiguous large
species with many historical records, however they have not been seen for decades in
Finland (last record for D. carinifrons from 1945). While the species have certainly
become rarer, their complete extinction cannot be conrmed due to the regional pau-
city of recording. For example, Linnaemya rossica Zimin, 1954 was similarly missing
for 50 years until rediscovered in 2013 from Lieksa, eastern Finland.
Exorista sp. A. Probably an undescribed species, near to E. tubulosa Herting, 1967 (C.
Bergström, pers. comm.)
Linnaemya sp. A. An undescribed species (C. Bergström, pers. comm.). A highly
unusual, shiny black Linneamya species from Lapland.
Checklist of the superfamilies Oestroidea and Hippoboscoidea of Finland (Insecta, Diptera)407
Nemorilla oralis (Fallen, 1810). N. maculosa (Meigen, 1824) has been mixed with
this species on Hackman´s (1980) list. True oralis was found new to Finland in
2011 from Loviisa (J. Flinck leg.).
Phebellia sp. A. Probably an undescribed species (J. Flinck leg. , T. Zeegers priv. comm.)
Stomorhina lunata (Fabricius, 1805). Recorded once from the University Botanical
Gardens in Helsinki (2 exx). It is quite possible that the ies were imported to Finland,
but the species is also a known vagrant and the possibility of natural dispersal cannot
be excluded.
Tachina sp. a. magnicornis. Close to the real T. magnicornis of Zetterstedt. e
proper name of this species still under investigation, but may be T. tetramera
(Zetterstedt, 1849). It is common and widespread in Finland. Some records have
been conrmed by the DNA barcoding project (Pohjoismäki et al. 2013). See
Tschorsnig et al. (2002) and Novotna et al. (2009).
References
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garter Beiträge zur Naturkunde A, Neue Serie 4: 321–334.
Dick CW (2006) Checklist of world Hippoboscidae (Diptera: Hippoboscidae). Department of
Zoology, Field Museum Natural History, Chicago, IL.
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117–162.
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Diptera. Notulae entomologicae 61: 225–226.
Herting B (1993) Family Rhinophoridae. In: Soós Á, Papp L (Eds) Catalogue of Palaearctic
Diptera 13. Hungarian Natural History Museum, Budapest, 102–117.
Herting B, Dely-Draskovits A (1993) Family Tachinidae. In: Soós Á, Papp L (Eds) Catalogue
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tion of Mystacinobiidae and McAlpine’s y. Systematic Entomology 35: 614–635. doi:
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(2012) Molecular phylogenetics of Oestroidea (Diptera: Calyptratae) with emphasis on
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O’Hara JE (2012) World genera of the Tachinidae (Diptera) and their regional occurrence.
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Entomology, International American Entomological Institute, Gainsville, 558 pp.
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København-Köln, 272 pp.
Tschorsnig H-P, Ziegler J, Herting B (2002) Tachinid ies (Diptera: Tachinidae) from the
Hautes-Alpes, France. Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie) 656: 1–62.
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