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Contributions to a History of Mexican Dipterology,- Part 1. Entomologists and their works before the Biologia Centrali-Americana

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The life and works of entomologists who described species of Diptera from Mexico before the publication of the Biologia Centrali-Americana are presented, including some commentaries about the collectors. Here are listed all the Mexican Diptera species-names proposed by Thomas Say (15 species), Rudolph Wilhelm Wiedemann (35 species), Pierre Justin Marie Macquart (71 species), John Obadiah Westwood (5 species), Bracy Clark (1 species), Francis Walker (91 species), Luigi Bellardi (176 species), Camilo Rondani (5species), F. Jaennicke (24 species), Edward Adolph Gerstaecker (8 species), Jaques Marie Frangille Bigot (205 species), Alfred Dugès (1 species), Friedrich Moritz Brauer (3 species), F. M. Brauer & J. F.Bergenstamm (13 species), Ermanno Giglio-Tos (177 species), and Ewald Rübsaamen (2 species). Included are a total of 832 specific names of the following 51 current families (in phylogenetic order): Tipulidae, Culicidae, Simuliidae, Anisopodidae, Bibionidae...
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Acta Zool. Mex. (n.s.) 84: 65-173 (2001)
65
CONTRIBUTIONS TO A HISTORY OF MEXICAN DIPTEROLOGY.-
PART I. ENTOMOLOGISTS AND THEIR WORKS BEFORE THE
BIOLOGIA CENTRALI-AMERICANA
.
Nelson P
APAVERO
1
and Sergio I
BÁÑEZ
-B
ERNAL
2
1
Museu de Zoologia & Instituto de Estudos Avançados,
Universidade de São Paulo, São Paulo, SP, BRAZIL. Pesquisador do Conselho Nacional
de Desenvolvimento Científico e Tecnológico (CNPq, Proc. Nº1 300994/79);
2
Departamento de Entomología, Instituto de Ecología, A.C.,
Km 2.5 Carr. Antigua a Coatepec No. 351, Congregación El Haya, 91070,
Xalapa, Veracruz, MÉXICO
RESUMEN
Se presenta la vida y obra de aquellos entomólogos quienes describieron especies de Diptera de México
antes de la publicación de la obra Biologia Centrali-Americana, incluyendo algunos comentarios respecto
a los colectores. Aquí se incluyen las listas de los nombres de especies mexicanas propuestos por Thomas
Say (15 especies), Rudolph Wilhelm Wiedemann (35 especies), Pierre Justin Marie Macquart (71 especies),
John Obadiah Westwood (5 especies), Bracy Clark (1 especie), Francis Walker (91 especies), Luigi Bellardi
(176 especies), Camilo Rondani (5 especies), F. Jaennicke (24 especies) Edward Adolph Gerstaecker (8
especies), Jaques Marie Frangille Bigot (205 especies), Alfred Dugès (1 especie), Friedrich Moritz Brauer
(3 especies), F. M. Brauer & J. F. Bergenstamm (13 especies), Ermanno Giglio-Tos (177 especies), and
Ewald Rübsaamen (2 especies), esto es, un total de 832 nombres específicos de las siguientes 51 familias
actuales (en orden filogenético): Tipulidae, Culicidae, Simuliidae, Anisopodidae, Bibionidae, Ditomyiidae,
Mycetophilidae, Sciaridae, Xylomyidae, Stratiomyidae, Tabanidae, Rhagionidae, Pantophtalmidae,
Therevidae, Mydidae, Asilidae, Nemestrinidae, Acroceridae, Bombyliidae, Empididae, Dolichopodidae,
Syrphidae, Conopidae, Micropezidae, Neriidae, Psilidae, Tanypezidae, Richardiidae, Otitidae,
Platystomatidae, Tephritidae, Dryomyzidae, Sepsidae, Sciomyzidae, Lauxaniidae, Lonchaeidae, Ephydridae,
Drosophilidae, Chloropidae, Heleomyzidae, Rhinotoridae, Anthomyiidae, Scatophagidae, Muscidae,
Hippoboscidae, Streblidae, Nycteribiidae, Calliphoridae, Sarcophagidae, Tachinidae y Cuterebridae. Se
presenta la referencia original, la localidad tipo, el museo o colección depositarios, el estado taxonómico
actual de cada nombre y referencias adicionales.
Palabras Clave: Historia, Diptera, México, taxonomía.
ABSTRACT
The life and works of entomologists who described species of Diptera from Mexico before the publication
of the Biologia Centrali-Americana are presented, including some commentaries about the collectors. Here
are listed all the Mexican Diptera species-names proposed by Thomas Say (15 species), Rudolph Wilhelm
Wiedemann (35 species), Pierre Justin Marie Macquart (71 species), John Obadiah Westwood (5 species),
Bracy Clark (1 species), Francis Walker (91 species), Luigi Bellardi (176 species), Camilo Rondani (5
species), F. Jaennicke (24 species), Edward Adolph Gerstaecker (8 species), Jaques Marie Frangille Bigot
Papavero & Ibañez-Bernal: History of Mexican Dipterology
66
(205 species), Alfred Dugès (1 species), Friedrich Moritz Brauer (3 species), F. M. Brauer & J. F.
Bergenstamm (13 species), Ermanno Giglio-Tos (177 species), and Ewald Rübsaamen (2 species). Included
are a total of 832 specific names of the following 51 current families (in phylogenetic order): Tipulidae,
Culicidae, Simuliidae, Anisopodidae, Bibionidae, Ditomyiidae, Mycetophilidae, Sciaridae, Xylomyidae,
Stratiomyidae, Tabanidae, Rhagionidae, Pantophtalmidae, Therevidae, Mydidae, Asilidae, Nemestrinidae,
Acroceridae, Bombyliidae, Empididae, Dolichopodidae, Syrphidae, Conopidae, Micropezidae, Neriidae,
Psilidae, Tanypezidae, Richardiidae, Otitidae, Platystomatidae, Tephritidae, Dryomyzidae, Sepsidae,
Sciomyzidae, Lauxaniidae, Lonchaeidae, Ephydridae, Drosophilidae, Chloropidae, Heleomyzidae,
Rhinotoridae, Anthomyiidae, Scatophagidae, Muscidae, Hippoboscidae, Streblidae, Nycteribiidae,
Calliphoridae, Sarcophagidae, Tachinidae, and Cuterebridae. The original reference, type-locality, depository
museum or collection, present taxonomic status of each name and additional references are presented.
Key Words: History, Diptera, Mexico, taxonomy.
HISTORY
1.
T
HE FIRST SPECIES
,
DESCRIBED BY
T
HOMAS
S
AY AND
R
UDOLPH
W
ILHELM
W
IEDEMANN
Before Mexican independence, in 1821, it was nearly impossible for Europeans
to settle in Mexico, being difficult for naturalists to explore a good amount of the
territory, principally due to the social and economical situation promoting the
scarcity of roads and danger.
The only exception to that rule, as is well-known, was the trip of Baron Alexander
von Humboldt (Fig. 1) and Aimé Bonpland to several Spanish colonies in South,
Central, and North America, from 1799 to 1804, thanks to special permission from
the King of Spain, Charles IV. After travelling through Venezuela, Cuba, Nueva
Granada, Ecuador, and Peru, the two naturalists left the Port of Guayaquil on
February 15, 1803, heading for Acapulco, in the Vice-kingdom of New Spain, where
they landed on March 23. They began their march to the interior of Mexico almost
immediately, through the present State of Guerrero, passing by Chilpancingo,
Taxco, and later by Cuernavaca in the present State of Morelos. In Mexico City,
they were received by the Viceroy, Count Iturrigaray. After a stay in the Capital, they
went, on August 1, 1803, to Guanajuato. On September 9, they arrived in Jorullo,
near Uruapan, proceeding thence to Toluca, via Morelia, on the 21
st
. Returning to
Mexico City, they packed their collections and on January 20, 1804, went to
Veracruz, passing through Puebla. They arrived at the former city on February 19.
On March 7, they sailed once more to Cuba, staying shortly in Havana, where they
picked up the collections made in the Orinoco (Venezuela) and left there since
1801. Through the Consul of the United States, Humboldt received a letter from
Thomas Jefferson, inviting him to visit the country. Humboldt and Bonpland sailed
to the United States and after some weeks returned to Europe, arriving there in
Acta Zool. Mex. (n.s.) 84 (2001)
67
August 1804 (Beck, 1959-1961; Chardón, 1949; Coats, 1970; Hemsley, 1887;
Kellner, 1963; Koehler, 1904; Papavero, 1971b; Penell, 1945; Sprague, 1924;
Stafleu & Cowan, 1979; Standley, 1920; Stearn, 1968; Stevens, 1956; Terra, 1955).
The travels of Humboldt and Bonpland were published in 1814 (see also Humboldt,
1861-1862).
Figure 1
Baron Alexander von Humboldt. Redrawn by S. Ibáñez-Bernal from self-portrait.
Papavero & Ibañez-Bernal: History of Mexican Dipterology
68
Almost twenty years afterwards, in 1823, William Bullock went to Mexico (Bullock,
1824) with the aim of acquiring abandoned gold and silver mines. Bullock was a
former London goldsmith, well-known as the owner of the “London Museum”, which
was sold by auction in 1819. When he returned to London at the end of 1823, he
brought back with him all sorts of Mexican curiosities which served for public show,
called “New Mexico”, in his “Egyptian Hall”. Farber (1982: 49) said:
“Bullock carefully labelled his specimens and displayed them in a scientific
manner. William Jordan, in his “Men I have known” (1866: 70-71) recalled the
opening of Bullock's Museum:
'Up to that date, there was nothing of the sort´ The British Museum was not, in
those days, a place of popular resort. The Leverian Museum, in the Blackfriars
Bridge Road, was a most heterogeneous medley of stuffed animals, without order
or classification, and savage costumes, weapons, and products from the Pacific
Ocean, or elsewhere in Asia, Africa, or America, as such curiosities were picked up
by adventurous navigators and exploring travellers. In a visit to it a few desultory
facts might be gathered; but as a means for solid or lasting instruction, its
miscellaneous and aimless character rendered it useless. Mr. Bullock's collection
was quite the reverse of this- admirably preserved and scientifically arranged-. After
three or four experimental years in its original locality, it was transported to the
Egyptian Hall, then finished for its reception, and not fewer than 32,000 subjects of
animated nature were skill-fully grouped and conveniently displayed within its walls.
The town was absolutely astonished by the individual acquisition of so vast and
marvellous a treasure and crowds soon availed themselves of the privilege of
reading its lessons. In one department were seen the quadrupeds, as natural as life,
and as they would appear in a real Indian forest, with its rocks, caverns, trees, and
all other adjuncts congenial to their habits and habitats. In another direction, 3,000
birds were set up with similar accuracy, and attended by well-selected accessories,
so as to afford sufficient ideas of their motion, food, and mode of feeding, and
peculiarities of every description'.
Bullock's Museum was also described by Mullens (1917).
In Germany, the news that mysterious Mexico had become accessible to the
ordinary traveller excited the curiosity of a wealthy nobleman, Count von Sack,
“Zweiter Ober-Jägermeister” and chamberlain to the King of Prussia (Stresemann,
1954; Papavero, 1971b). He had recently returned from a voyage to Cyprus and
Egypt, where he had made a small collection of birds, and at once felt inclined to
Acta Zool. Mex. (n.s.) 84 (2001)
69
visit Mexico, provided that there was a collecting naturalist of good reputation to go
with him.
A gardener by the name of Ferdinand Deppe (Binford, 1989; Ewan, 1955;
Hemsley, 1887; Hitchcock, 1919; Lindemann, 1884-1885; Nelson, 1922; Papavero,
1971b; Penell, 1945; Schiede, 1829-1830; Smith & Smith, 1973), appointed to the
Royal Gardens, was recommended for this task by Professor Hinrich Lichtenstein,
director of the Zoological Museum of Berlin University. Deppe was an intelligent and
energetic young man born in 1794. For a long while his connections with the
Zoological Museum had been intimate, the more so as his elderly brother Wilhelm
was accountant of the institution.
On Count von Sack's advice, Deppe gave up his job in 1821, and prepared for
the voyage to Mexico. He trained himself in skinning birds and mammals, a
technique which he soon mastered to great perfection. Besides, he studied books
on the zoology, botany and geography of South America, took lessons in drawing
and painting, and acquired English and Spanish. However, the wayward Count
remained undecided for three more years, and it was only due to the insistence of
Deppe that the plan to go Mexico was finally carried out in 1824.
The party, increased by the Count's domestic, arrived in London on August 23,
1824. There Deppe visited, besides the British Museum (the zoological section of
which he judged far inferior to that of the Berlin Museum), Mr. Bullock's Show and
Mr. Leadbeater's Shop (a dealer in natural history objects).
On October 8, they sailed from Falmouth on board a British ship to Jamaica via
Barbados. There they took another vessel bound for Alvarado, Veracruz, arriving
in mid-December 1824; shortly after the Count's footman died from a yellow fever
attack.
Two years later, in January 1827, Deppe left Mexico, after having travelled via
Mexico City to Oaxaca and Tehuantepec and back via Oaxaca to Alvarado. He and
William Bullock's son who had joined him on the trip from Mexico City to
Tehuantepec, are the first naturalists who ever collected birds for scientific purposes
in Mexico.
The following itinerary has been compiled by Stresemann (1954) from Deppe's
letters, kept in the archives of the Zoological Museum in Berlin, and from the entries
in Lichtenstein's lists of acquisitions in the same museum. From December 25, 1824
to January 1825, an excursion was made from Alvarado to the swamps and lagunas
near Tlacotalpan in Veracruz. In January 1825, he went from Alvarado to Jalapa,
Veracruz, and in February from Jalapa to Mexico City. In April, an excursion was
made from Mexico City to Temascaltepec, State of Mexico, where Deppe made the
acquaintance of Mr. Bullock Jr., who, like his father, took the hobby of collecting
birds and was an excellent shot. He had come over with his father in 1823 and lived
in Temascaltepec as an agent of a British mining company. Deppe went back to
Papavero & Ibañez-Bernal: History of Mexican Dipterology
70
Mexico City on May 10. He stayed in and near Mexico City from May 11 to the end
of the month. In June and July a trip was made to Chico and Toluca, in the State of
Mexico, including a stay at Tlalpujahua and another at Cimapán (Zimapán). On
August 25, Deppe, joined by Bullock Jr., started for a long trip to Tehuantepec.
Taking the route via Puebla and Tehuacán, they reached the city of Oaxaca by
September 6. Extensive field work was done in the vicinity. On one of these trips
they reached Villa Alta, “37 legoas” from Oaxaca, on September 22. On October 22,
Deppe and Bullock continued their journeys and six days later they reached
Tehuantepec via San Bartolo. Early in November they proceed to the Pacific shore
(San Mateo del Mar, Santa María del Mar) and were back at the city of Oaxaca by
the 22
nd
of that month. On December 5, Bullock left for Mexico City to meet his
father. Deppe left Oaxaca the next day and chose a direct route to Alvarado, which
proved very difficult. On December 19, he crossed the Cordillera Real at a place
called Valle Real (?), Veracruz, clad with luxuriant forest. He was back in Alvarado
on December 22.
Early in January 1826, Deppe paid another visit, this time an extensive one, to
Valle Real. From there he returned to Alvarado in March and proceeded via
Santuario to the city of Veracruz, which was reached by March 23. After a short
stay, he left for Mexico City at the end of March. The time from April 13 to July 16
was devoted to excursions to the environs of Mexico City, including a trip to Chico
and Ixmiquilpec (?). On July 17, Deppe left the Capital to visit his friend Bullock Jr.
at his place in Rincón de Temascaltepec. He crossed the range of Las Cruces and
proceeded via Tenancingo and Saculpán (?), Mexico. His stay in Temascaltepec
lasted from July 28 to late September, with excursions to Real (de) Arriba (?),
Mexico, and to Jantepeque (?) and Cuernavaca in Morelos. In August, in
Temascaltepec, Deppe met William Bullock (senior), who had made a business trip
to Mexico in company of his wife and daughter. After his return to London in 1823,
the senior Bullock founded a British mining company of which he was the director.
At the end of September, Deppe went back to the Capital and prepared for a quick
departure. He reached Jalapa by October 26 and embarked at Veracruz at the end
of January 1827, for Hamburg, on the German vessel “Anna Maria”.
On April 9, 1827, after almost three years of absence, Deppe was welcomed
back in Berlin.
Since the landing at Alvarado in December 1824, there was no mention of Count
von Sack in this short review of the itinerary. The Count seems to have been a very
queer and whimsical person. No longer disposed to suffer his employer's bad
manners, Deppe parted company with him in May 1825 and carried out the rest of
the expedition at his own risk. The Count returned to the coast in the autumn of
1825, sailed back to Jamaica, and reappeared in the summer of 1826 in Berlin,
where three years later he died.
Acta Zool. Mex. (n.s.) 84 (2001)
71
In the years 1825 and 1826 Deppe had gathered, all by himself, 958 bird skins,
a number of mammals, a quantity of reptiles, amphibians, fishes, snails, and
thousands of insects (but very few Diptera). Nor had botany been neglected by this
indefatigable naturalist. All his zoological material was bought by the Zoological
Museum of Berlin. But in vain, he had hoped to be rewarded with a post at one of
the scientific institutions of the Prussian capital. This induced him to return to his
beloved Mexico once more, this time in the company of a dear friend, the botanist
Dr. Christian Julius Wilheim Schiede (1798-1836) (Barnhart, 1965; Hemsley, 1887;
Knobloch, 1983; Lindemann, 1884-1885; Papavero, 1971b; Schiede, 1829-1830;
Smith & Smith, 1973; Stafleu & Cowan, 1985). They expected to make their living
in Mexico by selling zoological and botanical specimens to European museums and
dealers.
In July 1828 they settled in Jalapa, whence they made distant excursions to
various places, all within or near the boundary of the State of Veracruz; among
others were the Pico de Orizaba, which they ascended almost to the summit,
Misantla, Papantla, Veracruz, and the Laguna Huetulacán (?), west of the Cofre de
Perote. But they were very soon disappointed. Lichtenstein was no longer able to
buy quantities of Mexican materials at reasonable prices, and although some of the
specimens collected up to May 7, 1829, were acquired by the Museums of Berlin
and Vienna, the financial result of their efforts was far below what they had
expected. Ferdinand Deppe and Wilhelm Schiede were forced to give up this kind
of business in 1830. The latter died very soon afterwards. Deppe became a
commission agent to merchants located in Acapulco and Monterey, California. In
1836 he was tricked out of all he had learned and decided to sail home to Germany
once more. Back in Berlin in 1838, the unfortunate Deppe was again denied an
official appointment. He died in oblivion about 1860 (Stresemann, 1954).
*****
Another collector in Mexico in those times was the illustrious North American
entomologist Thomas Say (Fig. 2) (Stroud, 1992; Weiss & Ziegler, 1931). H. S.
Barber (1928) called attention to this very little known trip of Say. Say, in the
company of William McLure, travelled and collected along the old road between
Veracruz, Jalapa, Mexico City, and Tacuba, beginning in the autumn of 1827; and
on the approach of summer (1828) they returned to the United Sates. Barber
reproduced in his paper several extracts from Say's papers and other commentaries
by other authors confirming that Say had indeed collected in Mexico.
Papavero & Ibañez-Bernal: History of Mexican Dipterology
72
Figure 2
Thomas Say. Redrawn by S. Ibáñez-Bernal.
Say (1829) described the following Mexican species:
1. Limnobia livida, p. 150. “Mexico”. Now in Limonia (Dicranomyia) (Alexander & Alexander,
1970: 54), but identity uncertain (Tipulidae).
2. Penthetria heros, p. 154. “Mexico”. (Hardy, 1966: 2) (Bibionidae).
3. Dilophus stygius, p. 155. “Mexico”. (Hardy, 1966: 11) (Bibionidae).
Acta Zool. Mex. (n.s.) 84 (2001)
73
4. Nemotelus polyposus, p. 160. “Mexico”. (James, 1973: 48) (Stratiomyidae).
5. Stratiomys trivittatus, p. 160. “Mexico”. Now Hedriodiscus trivittatus (James, 1973: 41)
(Stratiomyidae).
6. Xilota arquata, p. 162. “Mexico”. Now Asemosyrphus arquatus (Thompson et al., 1976: 96)
(Syrphidae).
7. Syrphus stegnus, p. 163. “Mexico”. Now Carposcalis stegna (Thompson et al., 1976: 41)
(Syrphidae).
8. Syrphus mutuus, p. 164. “Mexico”. Now in Toxomerus (Thompson et al., 1976: 52)
(Syrphidae).
9. Syrphus ectypus, p. 165. “Mexico”. Now in Toxomerus (Thompson et al., 1976: 50)
(Syrphidae).
10. Chrysotus concinnarius, p. 168. “Mexico”. As unplaced species of Dolichopodidae
(Robinson, 1970: 58).
11. Medeterus punctipennis, p. 170. “Mexico”. Now Pelastoneurus punctipennis (Robinson,
1970: 51) (Dolichopodidae).
Say (1830) also described four new Mexican flies:
12. Volucella violacea, p. 166. “Mexico”. Preoccupied by Lepeletier & Serville (1825); now
Copestylum metalliferum (Walker) (Thompson et al., 1976: 79) (Syrphidae).
13. Volucella postica, p. 166. “Mexico”. Now Copestylum posticum (Thompson et al., 1976: 82)
(Syrphidae).
14. Volucella marginata, p. 167. “Mexico”. Now Copestylum marginatum (Thompson et al.,
1976: 78) (Syrphidae).
15. Ephydra hians, p. 188. “Mexico”. Now in Hydropyrus (Wirth, 1968: 24) (Ephydridae).
The first species of Mexican Diptera, collected by the travellers mentioned above,
were described by the German dipterist Christian Rudolph Wilhelm Wiedemann
(1770-1840) (cf. the excellent biography of Wiedemann published by Pont (1995)).
Wiedemann's first paper on flies was published in 1819. In 1820 he published the
first part of the first edition of his Diptera exotica (1820a) and a paper describing new
genera of Diptera (1820b). In 1821 he published the second part of the first edition
of the Diptera exotica (1821a, 1821b), and a much enlarged second edition of the
same work (1821c). In this latter work, Wiedemann described the first Mexican
species of flies, namely:
1. Culex taeniorhynchus, p. 43 (&). “Mexico”. Now Aedes (Ochlerotatus) taeniorhynchus (Wied.)
(Culicidae).
2. Culex posticatus, p. 43 (&). “Mexico (Coll. Winthem)”. Now a junior synonym of Psorophora
(Janthinosoma) ferox (Humboldt) (Belkin, 1968: 28) (Culicidae).
3. Tabanus quinquevittatus, p. 84. “Mexico”. (not included in Fairchild's (1971) catalogue or
Fairchild & Burger (1994)) (Tabanidae).
4. Bombylius helvus, p. 164. “Mexico” (Painter et al., 1978: 3) (Bombyliidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
74
5. Bombylius mexicanus, p. 166. Curiously, Painter et al. (1978: 3) mentioned “U.S.A.,
Georgia” as type locality. This species, and the foregoing, still in Bombylius, according to
Painter et al., 1978) (Bombyliidae).
6. Laphria melanogaster, p. 236. “Mexico and Savannah”. Same combination (Martin &
Papavero, 1970: 38) (Asilidae). The specimen serving for the description of this species
was also probably collected in Mexico by Humboldt, and in Savannah, Georgia, by
Thomas Say.
All these flies were most probably collected in Mexico by Humboldt.
In his Analecta entomologica (1824), Wiedemann included only one additional
Mexican species:
7. Bombylius confusus, p. 60. “Middle America”. Now in Sparnopolius, according to Painter et
al. (1978: 7) (Bombyliidae).
In the first volume of the Aussereuropäische zweiflügelige Insekten (1828) no
Mexican species were included. Only when the second volume was published
(1830) did Wiedemann include new species collected in Mexico by Deppe. It is
interesting to note that only a few species were included in the main body of the
text; most of them were only described in the appendix to that work. Why did that
happen? Why did it take such a long time for Wiedemann to obtain those
specimens collected by Deppe and deposited in the Berlin Zoological Museum?
Most of the species were from Oaxaca, where Deppe collected in 1825. It is an
unsolved mystery.
The following species were described in volume II of the Aussereuropäische
zweiflügelige Insekten (1830):
8. Xylophagus rufipalpis, p. 619. “Mexico”. Now Archistratiomys rufipalpis (Wied.) (James,
1973: 4) (Stratiomyidae).
9. Tabanus circumfusus, p. 624. “Mexico”?. Now in Catachlorops (Catachlorops). Fairchild
(1971: 67) said this species is not Mexican, but from Southern Brazil and Uruguay; the
type must have been mislabelled in the Berlin Museum (Tabanidae).
10. Pangonia aurulans, p. 620. “Mexico, Oaxaca”. Now in Scione (Fairchild, 1971: 25; Fairchild
& Burger, 1994: 49) (Tabanidae).
11. Pangonia basilaris, p. 621. “Mexico”. Now Esenbeckia (Ricardoa) wiedemanni (Bellardi). P.
basilaris was pre-occupied by Wiedemann, 1828, so Townsend (1912) proposed the new
name Pangonia tepicana (Fairchild, 1971:11; Fairchild & Burger, 1994: 29) (Tabanidae).
12. Pangonia semiflava, p. 622. “Mexico”. Now in Esenbeckia (Ricardoa) (Fairchild, 1971: 11;
Fairchild & Burger, 1994: 28) (Tabanidae).
Acta Zool. Mex. (n.s.) 84 (2001)
75
13. Mydas interruptus, p. 628; 1831: 46, pl. 53, fig. 12 (as Midas sic.). “Mexico”. Unchanged
(Papavero & Wilcox, 1968: 8) (Mydidae).
14. Mydas rubidapex, p. 626; 1931: 40, pl. 52, fig. 2 (as Midas sic.). “Mexico”. Unchanged
(Papavero & Wilcox, 1968: 9; Papavero, 1996: 632) (Mydidae).
15. Anthrax latreillei, p. 633. “Mexico, Oaxaca”. Now in Ligyra (Painter et al., 1978: 43)
(Bombyliidae).
16. Anthrax lacera, p. 634. “Mexico”. Now in Stonyx (Painter et al., 1978: 34) (Bombyliidae).
17. Anthrax clotho, p. 635. “Mexico”. Now in Stonyx (Painter et al., 1978: 34) (Bombyliidae).
18. Anthrax valida, p. 636. “Mexico”. Now in Bryodemina (Painter et al., 1978: 24)
(Bombyliidae).
19. Anthrax astarte, p. 637. “Mexico”. Now in Villa (Chrysanthrax) (Painter et al., 1978: 45)
(Bombyliidae).
20. Anthrax leucothoa, p. 638. “Mexico”. Now in Villa (Paravilla) (Painter et al., 1978: 51)
(Bombyliidae).
21. Anthrax cyanoptera, p. 638. “Mexico”. Now in Cyananthrax (Painter et al., 1978: 58)
(Bombyliidae).
22. Anthrax terminalis, p. 639. “Mexico”?. Now Ogcodocera leucoprocta (Wiedemann) (Painter
et al., 1978: 24) (Bombyliidae).
23. Anthrax disjuncta, p. 639. “Mexico”. Now in Lepidanthrax (Painter et al., 1978: 35)
(Bombyliidae).
24. Dasypogon nitidus, p. 643. “Mexico”. Now in Holcocephala (Martin & Papavero, 1970: 7)
(Asilidae).
25. Leptogaster fervens, p. 646. “Mexico”. Same combination (Martin, 1968: 3)
(Leptogastridae or Asilidae-Leptogastrinae).
26. Empis spiloptera, p. 5. “Mexico”. Still in that genus (Smith, 1967: 24) (Empididae).
27. Cyphomyia elegans, p. 58. “Mexico”. Now in Euparyphus (James, 1973: 31)
(Stratiomyidae).
28. Volucella lata, p. 195. “Mexico, Oaxaca”. Now Copestylum latum (Thompson et al., 1976:
78) (Syrphidae).
29. Tachina corpulenta, p. 280. “Mexico”. Now in Adejeania (Guimarães, 1971: 46)
(Tachinidae).
30. Trypeta scutellata, p. 494. “Mexico”. An unplaced species (Foote, 1967: 58) (Tephritidae).
31. Trypeta scutellaris, p. 484. “Mexico”. Now in Dyscrasis (Steyskal, 1968b: 4) (Otitidae).
32. Trypeta lichtensteinii, p. 497. “Mexico”. Now in Ictericodes (Foote, 1967: 29) (Tephritidae).
33. Trypeta mexicana, p. 511. “Mexico”. Now in Dyseuaresta (Foote, 1967: 22) (Tephritidae).
34. Calobata pectoralis, p. 540. “Mexico”. Now in Micropeza (Steyskal, 1968a: 4)
(Micropezidae).
35. Calobata divisa, p. 540. “Mexico”. Now in Micropeza (Steyskal, 1968a: 3) (Micropezidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
76
By 1830, Mexico was the least known part of the Americas. Consulting
Appendage 1, we see that Wiedemann had included in his Aussereuropäische
zweiflügelige Insekten (1828, 1830) 2,053 “exotic” (i.e., non-European) species; 688
were reported from South America (mainly Brazil and the Guyanas); 319 from the
United States (mostly collected by Say); the West Indies had only 48, and Mexico
28 (plus the species described by Wiedemann in 1821 and 1824 and the species
described by Say).
2.-
T
HE SPECIES DESCRIBED BY
P
IERRE
J
USTIN
M
ACQUART
Pierre Justin Marie Macquart (Fig. 3) was born in Hazebrouck, 45 km west of
Lille, France, in 1776. In his parental home there was a garden where Macquart first
became interested in natural sciences. His older brother, a fellow of the “Société des
Sciences de Lille”, was an ornithologist, and assembled a sizeable collection which,
after his death, served as the starting point for the Lille Museum. A second brother
was interested in botany, and built a botanical garden with over 3,000 species.
Macquart also became interested in natural history, choosing entomology as his
field. His first paper, published by the Lille Scientific Society, dealt with Psyllids.
At the age of 21 he had to leave his native place to join the Army of the Rhine.
He went to Mannheim, entering the Corps of Engineers. During his leaves he was
able to explore the banks of the Rhine, so rich in natural productions. He served the
commander of the Army of Rhine, General Armand Samuel, Marquis of Mariscot,
in the quality of secretary and draftsman. With the Army he visited Schwetzingen,
Heidelberg and Mainz, passing to Switzerland, to Arau and nearby places.
Afterwards the general staff moved on to Zürich, returning to Arau and Basel. While
Macquart stayed at Basel he received the sad news of his mother´s death. He
return to Lille, after an absence of 18 months (1797-1798).
From the military campaign through Germany and Switzerland he brought home
German books, a herbarium, insects and birds.
Once established again in Lille he dedicated himself entirely to natural history,
and during the winters studied in the public library of Lille. On 27 nivôse, “An” XI
(1802) he was elected fellow of the “Société des Sciences, de l'Agriculture et des Arts
de Lille”.
Macquart also travelled frequently to several regions of France, especially to
Paris, where he followed the entomological lectures of Latreille. The great
entomologist identified the collections gathered by Macquart and encouraged him
in his career.
After a trip to Holland, Macquart's “wandering life settled down by a happy
marriage, followed by a happy and numerous paternity” (Macquart, 1850a).
Acta Zool. Mex. (n.s.) 84 (2001)
77
Following his marriage, he moved from Hazebrouck to Lestrem, living in a house
made of two towers, remains of a castle from the XIV century, in the middle of a
beautiful village near the banks of the Lawe, a tributary of the Lys. Eventually, he
became the major Lestrem and a member of the “Conseil Général du Pas de Calais”.
Figure 3
Pierre Justin Marie Macquart. Photograph of his bust taken at the Museum of Natural History of Lille (N.
Papavero).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
78
He started then his studies of Diptera, made easier by Meigen's publications.
With the help of the works of the great Master he started the study of the French
Diptera, which resulted in his “Diptères du Nord de la France” (1828-1833). This work
opened to him the important collections and libraries of the time: Blainville, Geoffroy
de Saint-Hilaire, Férussac, Lepelletier de Saint-Fargeau, Audinet Serville, Audouin,
Carcel, Al. Lefebvre, Brullé, Castelnau, Winthem, etc.
Soon after the publication of this work, Latreille thought of editing a special work
on insects, in collaboration with several entomologists, and invited Macquart to take
care of the Diptera. This project, whose execution was delayed by the health of the
editor, was afterwards started again, under the editorship of N. E. Roret, and the
name changed to “Collection des Suites à Buffon, formant avec les oeuvres de cet
auteur un cours complet d'histoire naturalle” (82 volumes, 11 atlases). Macquart
worked very hard, preparing the “Histoire Naturalle des Insectes Diptères”, studying
the collections in the Paris Museum and several private collections. Once the two
volumes were published (1834-1835), he received a number of collections of exotic
flies and established relations with all the leading entomologists of his time.
In the meantime, he travelled throughout France and Belgium. In 1839 he visited
Meigen in Stolberg, where he saw the collections, 3,000 drawings of flies done by
the great dipterist and Meigen's library, which contained exclusively his own
writings. As Meigen was in straitened circumstances, Macquart offered to buy all
those collections. After obtaining permission from the authorities of the Paris
Museum, Macquart acquired all that material. The drawings made by Meigen
remained unpublished in the Paris Museum, being rediscovered by Matile (1974)
and finally published (Morge, 1975).
Returning to the Museum of Paris, and having at his disposition the great
collections brought home by the French expeditions and collecting naturalists in
almost every corner of the globe, Macquart undertook the description of the new
species in the Museum, since these had not been examined by Wiedemann while
working on exotic flies, and only a few had been dealt with before in the works of
Guerin-Méneville, Olivier, Robineau-Desvoidy and Macquart's own “Historie
Naturelle des Insectes Diptères”. In this great series, published from 1838 to 1855,
in two tomes and five supplements, he described some 1,800 species. In the first
volume of his “Diptères exotiques nouveaux ou peu connus” (1838a: 15) he gave a
list of the collections studied. As new collections arrived from the explorations of
several travelling naturalists, such as Durville, Goudot, Pilate, Sallé, d'Orbigny,
Claussen, Ghiesbreght, Saint-Hilaire, and others, Macquart published new
supplements to this work.
Acta Zool. Mex. (n.s.) 84 (2001)
79
In 1845, he visited Switzerland again, calling on Perty in Berne, and thence
passed to Germany. Returning to France, he put his newly acquired entomological
collections in order and returned to his work of classifying the exotic flies.
In 1850, he wrote a book on the “Facultés intérieures des Animaux Invertébrés
(Macquart, 1850a), in whose introduction he published an autobiography of 82
pages.
Macquart was elected fellow of the entomological and scientific societies of
France, Normandy, Bordeaux, Turin, Lyon, Zürich, Malta, Stettin, and others, such
as the Linnaean Society.
He died in 1855, the year of publication of the 5
th
Supplement of his “Diptères
exotiques nouveaux ou peu connus”. His bust was placed in the Museum of Natural
History of Lille (Fig. 3).
Macquart's types of Diptera are partly in the “Muséum National d'Histoire Naturalle
de Paris”, partly (those belonging to the Bigot collection) in Oxford University (OXF)
and the British Museum (Natural History); a few remained in the Museum of Natural
History of Lille. In the Diptera section of the “Muséum National d'Histoire Naturalle de
Paris” (MNHNP) there exists a manuscript catalogue of the Diptera collection of
Macquart still existing in Lille, organized by Julien Salmon, who saved what was left
of the collection in 1899. The introduction to that catalogue says:
“Catalogue de la collection des Diptères de Macquart (conservés au Musée
d'Histoire Naturelle de Lille (Nord).
Notice: Le 15 décembre 1854, Macquart fit don de sa bibliothèque et de ses
colections entomologiques à la Société des Sciences et Arts de Lille.
Ces précieuses richesses en reçurent malheuresement pas les soins qu' elles
méritaient: lorsque je retrouvai la collection Macquart en 1897, au Musée d'Histoire
Naturelle de Lille, les anthrènes avaient élu domicile dans tous les cartons et
continuaient leurs ravages sans être inquiétés. J'entrepris de sauver tout d'abord les
débris de la collection des Diptères. Ceux-ci étaient répartis en trois groupes: 1º une
collection renfermant un grand nombre d'espèces-types (Indiquée T dans ce
catalogue); 2º une collection composée en grande partie d'espèces exotiques
(Marquée G dans ce catalogue); 3º une collection paraissant avoir été composée par
Macquart pour le Musée; car le catalogue imprimé de 1850 lui correspond
exactement (Indiquée par M dans ce catalogue). Après avoir donné à ces Diptères
les soins de préservation que réclamait leur état, j'en ai opéré le transfert dans des
cadres neufs, en respectant rigouresement l'ordre dans lequel je les ai trouvés dans
les vieux cartons, et sans essayer aucune intercalation. Puis, à l'aide des ouvrages
de Macquart (Mémoires de la Société des Science de Lille) et du Catalogue du
Papavero & Ibañez-Bernal: History of Mexican Dipterology
80
Musée, j'ai établi l'numération des espèces qui avaient échappé à la destruction et
de celles dont il subistait des débris observables.
On remarquera que certaines espèces d'un même genre se trouvent répétées
plusieurs fois avec des annotations différentes; cela tient à ce que je me suis borné,
faute de connaissances spéciales, à retranscrire avec la plus grande exactitude, le
libellé de chaque étiquette, laissant aux spécialistes le soin d´en tirer les conclusions
qu'il leur plaira.
Il semble aussi, au premier abord, que ce catalogue soit bien restreint, en
camparaison du nombre considérable de Diptères décrits par Macquart. Mais, après
avoir fait la part des ravages causées par Antrênes et par l'humidité, il faut tenir
compte de l'observation suivante, présentée par le savant entomologista lorqu'il
légua ses collections à la Société des Sciences: ´. . . les espèces qui me furent
comuniquées pour la détermination et particulièrement la collection du Jardin des
Plantes, celles recueillies par les Commissions scientifiques de Morée et d'Algérie,
celles rapportées des Ies Canaries par MM. Webb et Berthelot, etc., me furent
simplement confiées, et je me réservau seulement par convention avec les
possesseurs, un individu sur quatre de la même espèce, ce qui m'enrichit que très
médiocrement ma collection. On y chercherait donc vainement les types d'un grand
nombre de genres nouveaux que j'ai publiés, surtout dans l'ouvrage sur les Diptères
exotiques, que la Société a bien voulu comprendre dans ses mémoires et qui,
acompagné de 186 planches, en contient pas moins de 2300 descriptions d'espèces
nouvelles.
J'espère cependant que l'énumeration des débris de cette collection célèbre,
sera de quelque utilité aux Diptèristes qui l'avaient considerée jusqu'à présent
comme entièrement détruite.
Lille, le 20 janvier 1899
Signé: Julien Salmon”
Macquart studied the collections brought from Mexico by Fontaine, Pilate,
Ghiesbreght, Linden, and Funk. Let´s examine what is known about the lives and
itineraries of those naturalists.
*****
A “Fontaine” (or “Fontaines”) was cited by du Petit-Thouars, in his telling of the
voyage of the frigate “La Vénus” (1840-1843). Pierre-Antoine Fontana, “dit
Fontaine”, was “capitaine d'armes de première classe” aboard the frigate “La Vénus”,
commanded by du Petit-Thouars. The frigate left Brest on December 29, 1836, for
Teneriffe (January 1837) and then for Fernando de Noronha and Cabo Frio
Acta Zool. Mex. (n.s.) 84 (2001)
81
(February 3) in Brazil. The frigate landed in Rio de Janeiro, staying there from
February 4 to 16, going afterwards to Florianópolis, Montevideo (February 24),
Buenos Aires, Cape Horn, and Valparaíso (March 27), whence some members of
the expedition followed to Santiago overland. Leaving Valparaíso on May 13, they
visited several localities on the Peruvian coast, and from Callao (June 14) departed
for the Sandwich Islands and other places, returning afterwards to Monterey,
California. They then explored the localities of Guadalupe (November 19, 1837),
bahía de Magdalena, San Lucas, San Pedro del Cabo, Isla Venado, Mazatlán,
Golfo de Cortés, Isla Isabel, San Blas, Tepic, and Acapulco in Mexico. Proceeding
then to the Easter Island, they returned again to the South American Pacific coast,
touching the Juan Fernández Islands, landing in Valparaíso. After visiting some
islands, they headed for the Marquesas (du Petit-Thouars, 1840-1843). However,
no reference is made by Petit-Thouars to “Fontaine” as zoological collector, credit
being given to the ship surgeon, Adolphe Simon Néboux, “chirurgien de première
classe”, who sent collections of animals, especially birds and insects, to the Paris
Museum. Thus, we cannot be sure whether the “Fontaine” cited by Macquart is the
same as the one who participated in the “La Vénus” expedition.
Louis Pilate, according to Sallé (1852), travelled through the United States
(Alabama, Louisiana, and Texas) and explored the State of Yucatan in Mexico.
Returning to France, Pilate decided to visit Mexico again, and came to Yucatan for
a second time in 1849. After a prolonged stay in that part of the country, Pilate
moved to Mexico City, where he died on March 17, 1852, at the age of 36 years, of
“a hypertrophy of the heart”. The relation of Pilate's activities and descriptions of the
regions he visited were published by himself in 1846, as follows (translation):
The province of Yucatan, situated in the southeast of the Golf of Mexico,
extends nearly from 17-29º of latitude. Even in the north, the thermometer does not
go below 11º Réamur (=13.75ºC, 56.75ºF) during the coldest part of the winter; the
heat there is considerable in the summer. The northern part down to Campeche is
general arid; it is, so to say, only a vast rock, nearly level and with a wrinkled
surface. A single chain of hills about a hundred meters or more in elevation,
originates a little south of Campeche, follows the coast northward a few kilometres
inland for about 20 leagues, and then turns south-eastward to the rocky part of the
peninsula. One finds there, at various places, especially in the Northeast, a kind of
subterranean ponds called locally senotes (sic!); otherwise there is no water during
the dry season except in wells, that is, from November to the end of May. The
vegetation is poor, the thinness of the soil permits nothing to live but shrubs, mostly
of the Mimosa family, and rarely are there trees whose highest branches attain more
than 10 meters of height. I am not speaking of the cultivated places that are
Papavero & Ibañez-Bernal: History of Mexican Dipterology
82
continually irrigated. About a dozen leagues south of Campeche, a terrain of another
nature starts -from the little river of Champoton, the land is low, flat, humid, and
inundated for several months of the year. Finally, in the southern part, rivers abound
and the vegetation is magnificent. There is never any dryness there and the country
is very unhealthy. I lived in the capital (Merida, in the Northeast) for 5 years and
made only short trips to other localities; there was hardly anything in the north that
I could secure besides insects. It is easy to imagine that I would find little, and
generally only small insects, in a dry and sterile land. I might add that except for a
dozen Coleptera, everything is rare; in 4 or 5 hours of searching I often found only
a few specimens. However, in May and June there are more. In the rainy season
diurnal Lepidoptera are abundant, but not of many kinds. Except for mosquitoes and
the housefly, Diptera are very rare, and I would say the same about all other insects.
I needed plenty of perseverance, a very determined attitude, and the aid of a
domestic for 10 years, with nothing else to do but collect, to gather here 8,000
Coleoptera comprising almost 720 species, of which 500 about are new”.
August B. Ghiesbreght (the Ghisbrecht or Ghiesbrecht of Macquart) (1810-1893;
cf. Barnhart, 1927, 1965; Breedlove, 1981; Hemsley, 1887; Knobloch, 1983; Linden,
1867; Lorence & García, 1989; McVaugh, 1972; Papavero, 1971b; Pennell, 1945;
Rovirosa, 1889; Rzedowski & Rzedowski, 1989) was the zoologist of a Belgian
commission charged by the government to undertake a scientific exploration of
Mexico and other tropical countries. The other two members were Jean Jules
Linden (1817-1898) (Barnhart, 1927, 1965; Breedlove, 1981; Hemsley, 1887;
Linden, 1867; Papavero, 1971b; Pennell, 1945; Röhl, 1938a, b; Stafleu & Cowan,
1981 (3:42); Standley, 1930), a botanist, and Nicholas Funck (1817-1896)
(Barnhart, 1927, 1965; Hemsley, 1887; Linden, 1867; Papavero, 1971b), the artist
of the expedition. Their first travel was to Brazil, where they arrived in December
1835, visiting the provinces of Rio de Janeiro, Espírito Santo, Minas Gerais, and
São Paulo. In March 1837, they returned to Belgium with the zoological and
botanical collections obtained. Their second travel was made to Cuba, where they
arrived in December 1837. The Belgians spent 3 months on the island, exploring
its northern and western districts, and left in March 1838 for Mexico. There they
visited the plateau of Anáhuac, the volcanoes Popocatépetl and Iztaccihuatl, the
peak of Orizaba, the Cofre de Perote, and all the eastern slope of the Cordillera.
From Veracruz they sailed to Campeche, crossed Yucatan and then went by sea
to visit the state of Tabasco and later Chiapas, entering also northern Guatemala.
Linden then went to Havana and the United States and the commission returned in
February 1841 to Belgium. It seems that, from 1840 on, Ghiesbreght travelled
alone, living in Tabasco, and then in Chiapas; although repeatedly visiting Europe,
Acta Zool. Mex. (n.s.) 84 (2001)
83
Ghiesbreght spent many years in Mexico. Although only the name Ghiesbreght is
cited by Macquart, it is possible that in the same collections, insects collected by
Linden and Funck were also included. According to the Paris Museum manuscript
book of accessions (catalogue des animaux sans vertèbres, vol. 3, p. 83),
Ghiesbreght sold insects to the Museum in 1842, for 30 francs per hundred
specimens.
No Mexican species was described by Macquart in his “Histoire naturalle des
insectes diptères” (1834-1835).
In his “Diptères exotiques nouveaux ou peu connus” (1838-1850), Macquart
described 794 Neotropical species, only 71 being Mexican, as follows:
1838a:
1. Pachyrhina quadrilineata, p. 50 (%,&), “Mexique (Coll. Lefebvre)”. Now Nephrotoma
ferruginea (Fabricius, 1805) (Alexander & Alexander, 1970: 17) (Tipulidae).
2. Odontomyia lefebvrei, p. 189 (%), “Mexique (Coll. Lefebvre)”. Now Hedriodiscus lefebvrei
(Macquart) (James, 1973: 41) (Stratiomyidae).
3. Odontomyia emarginata, p. 190 (%), “Mexique (Coll. Lefebvre)”. Same generic combination
(James, 1973: 44) (Stratiomyidae).
1842:
4. Volucella mexicana, p. 25 (&), “Mexique”- Type Nº 1645 (Box 53), in MNHNP; also a
specimen in Lille (M13). Now Copestylum mexicanum (Macquart) (Thompson et al., 1976:
79) (Syrphidae).
5. Helophilus mexicanus, p. 64 (&), “Mexique (Coll. Lefebvre)”- Type in Lille (T6). Now
Asemosyrphus mexicanus (Macquart) (Thompson et al., 1976: 97) (Syrphidae).
1843a:
6. Dejeania analis, p. 34 (%), “Mexique”- Type Nº 1727 in MNHNP lost. Now Adejeania analis
(Macquart) (Guimarães, 1971: 46) (Tachinidae).
7. Dejeania rufipalpis, p. 35 (%), “Mexique”. Now Adejeania rufipalpis (Macquart) (Guimarães,
1971: 47) (Tachinidae).
8. Jurinia lateralis, p. 42 (%), “Mexique”. Now Archytas lateralis (Macquart) (Guimarães, 1971:
50) (Tachinidae).
9. Hystricia amoena, p 44 (%), “Mexique”- Type Nº 1749 in MNHNP, lost. Same generic
combination (Guimarães, 1971: 41) (Tachinidae).
10. Hystricia testacea, p. 44 (%,&), “Mexique (Coll. Lefebvre) et Amérique du Nord”- Specimen
in Lille (G14). An unrecognized species of Bombyliopsis (Guimarães, 1971:40)
(Tachinidae).
11. Micropalpus ornatus, p. 48 (%), “Mexique et Colombie”- Type Nº 1753 (Box 56) in MNHNP.
Now Cryptopalpus ornatus (Macquart) (Guimarães, 1971: 61) (Tachinidae).
12. Blepharipeza rufipalpis, p. 55 (sex?), “Mexique et Cuba”- Type (from Cuba) Nº 2551 in
Coll. Sagra/Bigot, MNHNP. Now Leschenaultia leucophrys (Wiedemann, 1830)
(Guimarães, 1971: 186) (Tachinidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
84
13. Lucilia brunnicornis, p. 142 (&), “Mexique, Mozaid (?)”- Type Nº 1847 in MNHNP, lost. Now
Phaenicia (Phaenicia) purpurascens (Walker, 1837) (James, 1970: 11) (Calliphoridae).
14. Lucilia mexicana, p. 143 (%), “Mexique”- Types Nº 1850 (4 specimens) (Box 60) in
MNHNP. Now Phaenicia (Phaenicia) mexicana (Macquart) (James, 1970: 11)
(Calliphoridae).
15. Curtonevra mexicana, p. 158 (%,&), “Mexique”. Now Neomusca mexicana (Macquart) (Pont,
1972: 50) (Muscidae).
16. Herina mexicana, p. 208 (sex?), “Mexique”. Now Senopterina Macquart (Steyskal, 1965:
657) (Platystomatidae).
17. Drosophila mexicana, p. 259 (%), “Mexique”- Types Nº 1998 (Box 66) (2 wings glued to
piece of cardboard) in MNHNP. An unplaced species (Wheeler, 1970:30) (Drosophilidae).
18. Olfersia mexicana, p. 297 (sex?), “Mexique”- Types Nº 2015 (2 specimens) (Box 66) in
MNHNP. Now Lynchia nigra (Perty, 1833) (Guimarães, 1968a:6) (Hippoboscidae).
1846:
19. Pachyrhina mexicana, p. 12 (%), “Mexique (Coll. Guérin-Méneville)”- Type Nº 1240 in
MNHNP, lost. Now Nephrotoma mexicana (Macquart) (Alexander & Alexander, 1970: 17)
(Tipulidae).
20. Tabanus ruber, p. 42 (&), “Mexique (Coll. Guérin-Méneville)”- Type Nº 1345 in MNHNP.
Now Tabanus subruber Bellardi, 1859 (Fairchild, 1971: 102; Fairchild & Burger, 1994: 150)
(Tabanidae).
21. Cyphomyia fenestrata, p. 48 (%,&), “Mérida, Yucatán (Pilate)”- Specimen in Lille (M6). Now
Cyphomyia albitarsis (Fabricius, 1805) (James, 1973: 26) (Stratiomyidae).
22. Hermetia coarctata, p. 50 (&), “Mexique (Coll. Fairmaire)”. Same generic combination
(James, 1973: 37) (Stratiomyidae).
23. Hermetia planifrons, p. 50 (%), “Mérida, Yucatán (Pilate)”. Now Hermetia albitarsis
Fabricius, 1805 (James, 1973:36) (Stratiomyidae).
24. Odontomyia rubricornis, p. 53 (%), “Mérida, Yucatán (Pilate)”. Type Nº 1440 in MNHNP,
lost. Now Labostigmina rubricornis (Macquart) (James, 1973: 43) (Stratiomyidae).
25. Ephippium fenestrata, p. 54 (%,&), “Mérida, Yucatán (Pilate)”. Type Nº 1445 in MNHNP,
lost; specimen in Lille (G5). Now Adoxomyia fenestrata (Macquart) (James, 1973: 30)
(Stratiomyidae).
26. Sargus speciosus, p. 56 (&), “Mérida, Yucatán (Pilate)”. Same generic combination
(James, 1973: 20) (Stratiomyidae).
27. Dasypogon candidus, p. 67 (%), “Veracruz (Coll. Spinola)”. Now Stichopogon trifasciatus
(Say, 1823) (Martin & Papavero, 1970: 5) (Asilidae).
28. Dasypogon mexicanus, p. 68 (%), “Mexique (Coll. Guérin-Méneville)”. Type Nº 1480 in
MNHNP, lost. An unrecognized species (Martin & Papavero, 1970: 90) (Asilidae).
29. Dasypogon nigritarsis, p. 68 (%), “Mexique (Coll. Guérin-Méneville)”. Type Nº 1481 in
MNHNP, lost. Now Diogmites nigritarsis (Macquart) (Martin & Papavero, 1970: 26)
(Asilidae).
30. Megapoda cyaneiventris, p. 71 (&), “Mexique (Coll. Robyns)”. Type Nº 1507 in MNHNP.
Now Pseudorus distendens (Wiedemann, 1828) (Papavero, 1975) (Asilidae).
Acta Zool. Mex. (n.s.) 84 (2001)
85
31. Atomosia tibialis, p. 76 (%), “Mérida, Yucatán (Pilate)”. Type lost. Same generic
combination (Artigas et al., 1991: 59) (Asilidae).
32. Erax argyrogaster, p. 84 (%,&), “Mérida, Yucatán (Pilate)”. Type not in Paris. Now Efferia
argyrogaster (Macquart) (Martin & Papavero, 1970: 63) (Asilidae).
33. Asilus mexicanus, p. 94 (%), “Mexique (Coll. Guérin-Méneville)”. Type lost. Now Machimus
(Tolmerus) mexicanus (Macquart) (Martin & Papavero, 1970: 86) (Asilidae).
34. Hirmoneura brevirostris, p. 101 (%), “Mérida, Yucatán (Pilate)”. Now Hirmoneura
(Hyrmophlaeba) brevirostris Macquart (Papavero, 1968: 3) (Nemestrinidae).
35. Exoprosopa pilatei, p. 110 (%), “Mérida, Yucatán (Pilate)”. Now Ligyra pilatei (Macquart)
(Painter et al., 1978: 44) (Bombyliidae).
36. Exoprosopa limbipennis, p. 110 (%), “Mérida, Yucatán (Pilate)”. Type Nº 1595 in MNHNP.
Now Exoprosopa argentifasciata (Macquart, 1846) (Painter et al., 1978: 39) (Bombyliidae).
37. Geron rufipes, p. 119 (%,&), “Mérida, Yucatán (Pilate)”. Same generic combination (Painter
et al., 1978: 17) (Bombyliidae).
38. Psilopus incisuralis, p. 120 (&), “Mérida, Yucatán (Pilate)”. Now Condylostylus incisuralis
(Macquart) (Robinson, 1970: 8) (Dolichopodidae).
39. Mixogaster mexicanus, p. 123 (&), “Mexique (Coll. Robyns)”. Type in MRHNB. Now
Mixogaster mexicana Macquart (Thompson et al., 1976: 58) (Syrphidae).
40. Volucella tibialis, p. 123 (&), “Mérida, Yucatán (Pilate)”. Type not found. Now Copestylum
tibiale (Macquart) (Thompson et al., 1976: 85) (Syrphidae).
41. Syrphus delineatus, p. 139 (&), “Mexique (Coll. Robyns)”. Type in MRHNB. An unplaced
species (Thompson et al., 1976: 39) (Syrphidae).
42. Baccha lineata, p. 139 (&), “Texas ou Yucatán (Pilate)”. Now Ocyptamus lineatus
(Macquart) (Thompson et al., 1976: 21) (Syrphidae).
43. Trichopoda mexicana, p. 172 (&), “Mexique (Coll. Robyns)”. Same generic combination
(Guimarães, 1971: 9) (Tachinidae).
44. Dexia rubriventris, p. 189 (%), “Mérida, Yucatán (Pilate)”. Now Ptilodexia rubriventris
(Macquart) (Guimarães, 1971: 33) (Tachinidae).
45. Dexia fuscanipennis, p. 188 (&), “Mérida, Yucatán (Pilate)”. An unrecognized Dexiinae
(Guimarães, 1971: 122) (Tachinidae).
46. Lucilia meridensis, p. 199 (%,&), “Mérida, Yucatán (Pilate)”. Actually not an species of
Calliphoridae, now Morellia meridensis (Macquart) (Pont, 1972: 9) (Muscidae).
1847:
47. Dichelacera scapularis, p. 15 (&), “Mexique (Coll. Bigot)”. Same generic combination
(Fairchild, 1971: 64; Fairchild & Burger, 1994: 98) (Tabanidae).
48. Tabanus apicalis, p. 20 (sex?), “Mexique (Coll. Bigot)”. Now Tabanus bigoti Bellardi, 1859
(Fairchild, 1971: 84; Fairchild & Burger, 1994: 132) (Tabanidae).
49. Sargus nigrifemorata, p.31 (%), “Mexique (Coll. Bigot)”. Now Himantoloba nigrifemorata
(Macquart) (James, 1973: 10) (Stratiomyidae).
50. Ceraturgus rufipennis, p. 32 (&), “Mexique (Coll. Bigot)”. Type OXF. Now Taracticus
rufipennis (Macquart) (Martin & Papavero, 1970: 35) (Asilidae).
51. Dasypogon nigripennis, p. 34 (&), “Mexique (Coll. Bigot)”. Type OXF. Now Diogmites
nigripennis (Macquart) (Martin & Papavero, 1970: 26) (Asilidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
86
52. Dasypogon dimidiatus, p. 35 (%), “Mexique (Coll. Bigot)”. Type OXF. Now Taracticus
dimidiatus (Macquart) (Martin & Papavero, 1970: 35) (Asilidae).
53. Lampria mexicana, p. 37 (%), “Mexique (Coll. Bigot)”. Type OXF. Same generic
combination (Martin & Papavero, 1970: 40) (Asilidae).
54. Ommatius pumillus, p. 42 (&), “Mexique (Coll. Bigot)”. Type OXF. Same generic
combination (Martin & Papavero, 1970: 59) (Asilidae).
55. Eristalis mexicanus, p. 59 (%), “Mexique (Coll. Bigot)”. Type BMNH. Now Palpada mexicana
(Macquart) (Thompson et al., 1976: 107) (Syrphidae).
56. Lucilia violacea, p. 83 (%), “Mexique (Coll. Bigot)”. Now Phaenicia purpurescens (Walker)
(James, 1970: 11) (Calliphoridae).
1850:
57. Pangonia planiventris, p. 26 (&), “Mexique (Muséum)”. Type Nº 1322 in MNHNP. Now
Esenbeckia (Ricardoa) planiventris (Macquart) (Fairchild, 1971: 11; Fairchild & Burger,
1994: 28) (Tabanidae).
58. Pangonia nigronotata, p. 27 (%,&), “Mexique (Muséum)”. Type Nº 1324 in MNHNP. Now
Esenbeckia (Ricardoa) nigronotata (Macquart) (Fairchild, 1971: 11; Fairchild & Burger,
1994: 27) (Tabanidae).
59. Pangonia bicolor, P. 27 (&), “Mexique (Muséum)”. Type Nº 1325 in MNHNP. Now
Esenbeckia (Ricardoa) semiflava (Wiedemann, 1830) (Fairchild, 1971: 11; Fairchild &
Burger, 1994: 28) (Tabanidae).
60. Tabanus albiscutellatus, p. 34 (&), “Mexique (Ghiesbreght)”. Type Nº 1365 in MNHNP.
Now Leucotabanus exaestuans (Linnaeus, 1758) (Fairchild, 1971: 83; Fairchild & Burger,
1994: 123) (Tabanidae).
61. Odontomyia flavifasciata, p. 53 (&), “Mexique (Ghiesbreght)”. Type Nº 1442 in MNHNP.
Now Hedriodiscus lefebvrei (Macquart, 1838) (James, 1973: 41) (Stratiomyidae).
62. Mydas vittatus, p. 60 (%), “Mexique (Coll. Brémond)”. Type Nº 1460 in MNHNP. Now
Stratiomydas rufiventris (Macquart, 1850) (Papavero, 1996:632) (Mydidae).
63. Dasypogon fasciventris, p. 69 (%), “Mexique (Coll. Bigot)”. Type OXF. Now Stichopogon
trifasciatus (Say, 1823) (Martin & Papavero, 1970: 5) (Asilidae).
64. Mallophora fulviventris, p. 77 (&), “Mexique (Ghiesbreght)”. Type Nº 1101 in MNHNP.
Same generic combination (Martin & Papavero, 1970: 74) (Asilidae).
65. Mallophora fulvianalis, p. 78 (&), “Mexique (Ghiesbreght)”. Type Nº 1102 in MNHNP. A
synonym of M. fulviventris Macquart? (Martin & Papavero, 1970: 74) (Asilidae).
66. Mallophora pica, p. 78 (%), “Mexique ou Bolivie (M. d'Orbigny)”. As d'Orbigny never
collected in Mexico, this species is probably from Bolivia. Type Nº 1103 in MNHNP. Same
generic combination (Martin & Papavero, 1970: 75) (Asilidae).
67. Anisotamia eximia, p. 115 (%), “Mexique (Ghiesbreght)”. Type Nº 1615 in MNHNP. Now
Bryodemina valida (Wiedemann, 1830) (Painter et al., 1978: 24) (Bombyliidae).
68. Eristalis testaceicornis, p. 138 (&), “Mexique”. Type OXF. Now Palpada testaceicornis
(Macquart) (Thompson et al., 1976: 104) (Syrphidae).
69. Hystricia ambigua, p. 172 (&), “Mexique (Coll. Bigot)”. Now Jurinella ambigua (Macquart)
(Guimarães, 1971: 56) (Tachinidae).
70. Prosena mexicana, p. 231 (%,&), “Mexique (Ghiesbreght)”. Type Nº 1798 in MNHNP. Now
Mochlosoma mexicanum (Macquart) (Guimarães, 1971: 31) (Tachinidae).
Acta Zool. Mex. (n.s.) 84 (2001)
87
Finally, in an article from 1852, Macquart described from Mexico his last species:
71. Megistopoda pilatei, p. 332, pl. 4, fig. 4. (sex?), Mexico, Tabasco, Teapa (Pilate) (Wenzel,
1970: 9) (Streblidae).
3.
T
HE SPECIES DESCRIBED BY
J
OHN
O
BADIAH
W
ESTWOOD
John Obadiah Westwood (Fig. 4) was born at Sheffield, England, on December
22, 1805, and died shortly after completing his 87
th
year, on January 2, 1892, at
Oxford. His father was a die sinker at Sheffield, but afterwards removed to Lichfield.
When nearly 16 years of age, Westwood went to London to be articled to a solicitor,
and though he devoted his attention more to the study of natural history than of law,
he was admitted to the bar as a solicitor and became partner in a firm. Having some
private means, which he augmented by writing and drawing, he was able to neglect
his profession and devote himself entirely to entomology and archaeology.
Westwood was actively associated with the Entomological Society of London, from
its foundation in 1831, and was for many years its secretary. Subsequently, he was
elected president when the Society celebrated its jubilee in 1833. He was also a
fellow of the Linnaean Society from 1827.
Westwood's dipterological publications include short papers describing
miscellaneous new species (1835a, 1835b), a revision of the Mydidae (1841),
descriptions of a new species of Systropus (Bombyliidae) (1842), and two papers
on Acroceridae (1848, 1876). Among his other publications, special mention must
be made to his “Introduction to the modern Classification of Insects”, a systematic
arrangement of the genera of British Insects (1840).
In 1858, the Reverend F. W. Hope, a wealthy amateur, who had been for years
a warm friend and patron of Westwood, and had purchased his collection, gave the
specimens to the University of Oxford, and founded a Professorship of Invertebrate
Zoology, which bears his name. Westwood was appointed the first Hope Professor
and in consequence removed to Oxford, where he was a conspicuous figure in the
University for 35 years (Anon, 1893; Wandolleck, 1893).
Westwood described only a few species of Mexican Diptera:
1. Mydas dives, 1841: 50, pl. 13, fig. 1. “Mexico (locality unknown)”. (& OXF). Now Protomydas
rubidapex (Wiedemann) (Wilcox et al., 1989: 16; Papavero, 1996: 632) (Mydidae).
2. Mydas crassipes, 1841: 51, pl. 13, fig. 3. “North America”. Now Ceriomydas crassipes
(Westwood) (Wilcox et al., 1989: 107) (Mydidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
88
3. Mydas senilis, 1841: 52. “Mexico, Tacubaya (Coffin)”. (% OXF). Unchanged (Wilcox et al.,
1989: 41; Papavero, 1996: 631) (Mydidae).
4. Mydas basalis, 1841: 53. “Mexico, Tacubaya (Coffin)”. (& OXF). Now a junior synonym of
Mydas senilis Westwood (Wilcox et al., 1989: 41-42; Papavero, 1996: 631) (Mydidae).
5. Systropus faenoides, 1842: 3. “Mexico”. Unchanged (Painter et al., 1978: 18) (Bombyliidae).
Nothing is known about the collector “Coffin”.
Figure 4
John Obadiah Westwood. Photograph property of N. Papavero.
Acta Zool. Mex. (n.s.) 84 (2001)
89
4.
T
HE FIRST
C
UTEREBRIDAE
,
DESCRIBED BY
B
RACY
C
LARK
The oldest known reference to a Mexican Cuterebridae (now subfamily
Cuterebrinae of Oestridae, according to Wood, 1987) is found in Fray Bernardino
de Sahagun´s “Historia General de las cosas de la Nueva España”. Sahagun, who
lived in New Spain from 1529 to 1590, said in that book that there existed “gusanos
que crían en los brazos o miembros de los conejos y ratones” (worms that live in the
marms or members of rabbits and rats); those worms could be seen “metidos dentro
de la carne y miran hacia fuera” (placed within the flesh, looking to the outside).
Those larvae, certainly of Cuterebra, were called “nacaocuilin” (from nahuatl
“nacatl”= flesh, and “ocuilin”= worm) (Vogelsang & Martín del Campo, 1947: 50).
Sahagun described the opening of the wound where the larva lives and its posterior
spiracles, taken by him as “eyes”, “looking to the outside” (Guimarães et al., 1983:
241).
But the first scientific description of a Mexican Cuterebridae would be made only
in 1848, by Bracy Clark, a veterinary surgeon and Fellow of the Linnaean Society
of London. In that paper he described Cuterebra atrox from Mexico. Sabrosky (1986:
149) said:
Holotype of atrox, female, Mexico (Oxford). Clark (1848) stated that the species,
from a specimen in the Westwood collection, was “believed to inhabit Africa”
(“Habitasse creditur in Africa”), but Westwood himself crossed out those words in the
Oxford University copy of Clark´s paper and wrote in the margin “Habitat certe in
Mexico, J. O. W.”, as already noted by Austen (1895). Austen also stated that the type
is “apparently a male”, but he corrected this in 1933 to female”.
As also commented by Sabrosky (1972: 89), on November 1
st
, 1796, Bracy Clark
(for a biography see Proc. Linn. Soc. London, 1861: xxi-xxiv) read to the Linnaean
Society an important paper on “Observations on the genus Oestrus”, and the
following year published it in the third volume of the Society´s Transactions. This
was expanded and published in 1815 as his famous “An essay on the bots of horses
and other animals”. Clark´s works are particularly significant contributions to the
knowledge of oestroid flies, with their keen and direct observations on then little
known, confused and misunderstood life histories and immature stages of the bot
and warble flies. Over the next decades he published a few other papers, ending
with three short comments in 1857, in his 87
th
year. He died in 1860.
Sabrosky (1972) published a very interesting paper on the “rediscovery” of the
bot fly collection of Clark.
Papavero & Ibañez-Bernal: History of Mexican Dipterology
90
5.
T
HE SPECIES DESCRIBED BY
F
RANCIS
W
ALKER
Francis Walker (Fig. 5) was the seventh son, and the tenth and youngest child
of Mr. John Walker, a gentleman of independent fortune, residing at Arno´s Grove,
Southgate, where Francis was born on July 31, 1809. His father had a decided taste
for scientific activities, especially natural history; he was a fellow of the Royal and
Horticultural Societies and vice-president of the Linnaean, so that his son´s almost
boyish propensity for those studies could be explained by those influences.
Figure 5
Francis Walker. Photograph.
Acta Zool. Mex. (n.s.) 84 (2001)
91
In 1816 Walker´s parents were staying with their family in Geneva, then the
center of a literary coterie in which they met, among other celebrities, Lord Byron,
Madame de Staël, and the naturalists Saussure and Vernet. They spent more than
a year in Geneva and Vevey, and in 1818 proceeded to Lucerne, from which place
Francis, then a boy nine years of age, made the ascent of Mount Pilatus, in
company with his elder brother Henry; their object, in addition to mountain climbing,
being the collecting of butterflies. The family afterwards visited Neuwied and
returned to Arno´s Grove in 1820.
In 1830 the two brothers, Henry and Francis, again visited the continent, and now
it was purely an entomological tour; Mr. Curtis, the well-known author of “British
Entomology”, being their companion. This party collected the French Satyridae most
assiduously on the Island of Jersey and afterwards at Fontainebleau, Montpellier,
Nantes, Vaucluse, etc.
Walker´s career as an author commenced in 1832. He contributed to the first
number of the “Entomological Magazine”, the introductory chapter of his
Monographia Chalciditum”.
In 1834, somewhat reluctantly, he consented to undertake the editorial
management of the Entomological Magazine, but resigned this office the following
year, yet continued to be a constant contributor to its pages. The same year he
visited Lapland.
From 1837 to 1863 he was hired by the British Museum to describe the insect
collections, receiving £1 per genus and 1 shilling per species. During his lifetime,
as calculated by Horn (1937: 432), Walker prepared some 20,000 (“zahllosen und
trostlosen”= innumerable and untrustworthy) descriptions of insects in several
orders.
Walker´s first paper on neotropical flies was published in 1837, containing the
descriptions of the Diptera collected during Captain P. P. Kinks survey of the Straits
of Magellan.
In 1849 (May) Walker married May Elizabeth, the oldest daughter of Mr. Ford, of
Ellel Hall, near Lancaster, and spent the summer on the Continent, again collecting
in Switzerland.
In 1848 he had explored the Isle of Thanet, and in 1849 he went to the Island of
Wright. During those two years the first four volumes of the “List of the specimens
of Dipterous Insects in the collection of the British Museum” were published. In the
succeeding years, 1850 and 1851, he visited Geneva and Interlaken. During 1850,
he published the first part of a work on the Diptera in the collections of W. W.
Saunders (1850a), and a smaller number of new species in the “Zoologist” (1850b).
He also commenced work on Diptera for a projected series of works on British
insects, to be called “Insecta Britannica”.
Papavero & Ibañez-Bernal: History of Mexican Dipterology
92
In 1851 and 1852, the second and the third parts of his “Insecta Saundersiana
were published (1851, 1852a, b), and from 1854 to 1856 the three supplements
(=vols. 5-7) of the “List”.
In 1856 appeared the last (fourth) part of the “Insecta Saundersiana”, and in 1857
Walker published the first part of the “Characters of undescribed Diptera in the
collection of W. W. Saunders”, in the “Transactions of the Royal Entomological
Society”.
Another tour of the Continent occupied a considerable portion of 1857, with
Walker visiting Calais, Rouen, Paris, Strasbourg, Baden-Baden, Heidelberg,
Wiesbaden, Frankfurt, Mainz, Köln, Brussels, Aix-la-Chapelle and Antwerp. During
this journey he collected in the Black Forest.
In 1860, the second part of the “Characters” was published (and here he
described the great majority of his new species of Mexican flies) and the summer
of this same year was devoted to a thorough exploration of the Chennel Islands. In
1861 Walker´s excursions were confined to North Devon, and the third part of the
Characters” was published. In 1863 he toured the English lakes.
In the spring of 1865 he visited North Wales and Ireland. In the autumn he again
visited Paris, Geneva, Lucerne, Interlaken, and Altdorf, ascending the Righi, Mount
Pilatus and the Mürren, proceeding to Kanderstag, the Oeschinen See and the
Gemmi Pass.
In 1867, we find him again in France and Switzerland, ascending the Col de Voza
and examining the Jardin of the Mer de Glace; thence over the Tête Noire to
Martigny, Sion, and the Great St. Bernard, returning through St. Maurice and
Villeneuve to Geneva.
In 1868 he made the tour of the Isle of Mann and returned to Holyhead; in 1870
he paid another visit to Llanberis, as well as to all the more beautiful scenery in
North Wales, crossing over to Ireland and touring that island from south to north;
and in 1874 he examined entomologically the Scilly Islands and the districts of the
Lizard and the Land´s End.
In 1872 he turned his attention to Italy, visiting Rome, Piza, Lucca, Florence,
Naples, Sorrento, Capri, Milan and Venice, as well as the lakes Como and
Maggiore.
Finally, in 1874, he had again proceeded as far as Aberystwith, on his way to
Ireland, when his intention was frustrated by illness, which terminated fatally on the
5
th
of October, 1874 (Newman, 1874).
*****
Acta Zool. Mex. (n.s.) 84 (2001)
93
Walker used to take the insects of the British Museum home to describe them.
It is said by Horn (1937: 432) that “dem brave Fr. Walker passierte dabei nicht selten,
dass ihm “die ganze Geschichte” auf die Strasse fiel!” (Not seldom it happened to the
brave Fr. Walker that whole “kit and kaboodle” fell into the street!). His careless
descriptions aroused much argument, and it is stated by an anonymous writer at the
“Entomological Monthly Magazine” for 1874 (p. 141):
The authorities of the British Museum who permitted the scandal came under
the lash of such criticism as has happily seldom been directed against scientific men.
This criticism had no other effect than (apparently) to increase the evil. Walker, by
nature, appeared to be utterly indifferent to anything that could be hurled at him, and
the only apparent answer on the part of the ruling power at the Museum was the
commencement of catalogues of hiterto unassailed groups or orders”.
Baron Osten Sacken was especially acid when he wrote about Walker (1878: xvi-
xix):
“Mr. Walker´s writings on the order of Diptera are no better than his publications
on Lepidoptera, Hemiptera and Orthoptera, as characterized by other authors. The
same species are often found described under several different specific names and
placed in different genera; well characterized species of a certain genus are placed
in the wrong, sometimes in very distant genera, or even in the wrong family. In the
great majority of cases, the descriptions of a new species were drawn from a single,
often hardly recognizable specimen; and when new species happen to be
represented by more than one type-specimen, these are almost sure to belong to
different species. (Here Osten Sacken cites several instances of Walker´s
misidentifications and proceeds:) Mr. Walker´s identifications of the species of
former authors are often, I may say in most cases, incorrect. These facts are given
as a warning for entomologists not to trouble themselves too much about the
interpretation of Mr. Walker´s descriptions, because in most cases they will find
themselves mislead by the very data furnished by him... The authorities of the British
Museum, in a most praiseworthy, and truly scientific spirit, have bestowed a great
deal of labour upon preserving and labelling Mr. Walker´s types. But the task of
singling out the original type of the description from among the specimens added
afterwards is by no means an easy one, often hardly possible. Furthermore, it is a
well-known fact that authors are apt not to be very careful with their own types; to
remove and displace them, when made aware of an error; and Mr. Walker, in this
respect, was not an exception. Neither this, nor any other types can, therefore, be
Papavero & Ibañez-Bernal: History of Mexican Dipterology
94
implicity relied upon, and we have, ultimately, to fall back on the descriptions. In
rescuing those of Mr. Walker´s descriptions, which are available and in rejecting the
remainder, as useless, we pursue, I think, a course consistent both with justice and
scientific expediency”.
*****
Walker described over 750 species of Neotropical Diptera, 91 from Mexico (listed
below). It is very difficult, if not impossible, to say in which part of Mexico they were
collected, as Walker, with most other authors from the XIX century, was extremely
careless in citing the type-locality. We also know practically nothing about the
collectors who brought the specimens to the British Museum of Natural History.
Walker cites only the names of Coffin and Glennie, about whom nothing is known.
The species of Mexican flies described by Walker are the following:
1849a:
1. Anthrax orcus, p. 237 (&). “Mexico (Coffin)”. Now Ligyra orcus (Walker) (Painter et al., 1978:
43) (Bombyliidae).
1849b:
2. Helophilus formalis, p. 603 (&). “Mexico (Coffin)”. Now Asemosyrphus arquatus (Say, 1829)
(Thompson et al., 1976: 96) (Syrphidae).
3. Ersitalis diminutus, p. 622 (&). “Mexico (Coffin)”. Now Palpada diminuta (Walker)
(Thompson et al., 1976: 104) (Syrphidae).
4. Volucella metallifera, p. 636 (&). “Venezuela (Mr. Dyson´s collection), Mexico (Coffin)”. Now
Copestylum metalliferum (Walker). The specimen from Venezuela is a junior synonym of
Copestylum dispar Macquart (Thompson et al., 1976: 79) (Syrphidae).
5. Psilopus inficitus, p. 649 (sex?). “Mexico (Coffin)”. Now Condylostylus patibulatus (Say,
1823) (Robinson, 1970: 10) (Dolichopodidae).
1852:
6. Psilopus lepidus, p. 207 (%). “Mexico”. Now Condylostylus lepidus (Walker) (Robinson, 1970:
8) (Dolichopodidae).
7. Microdon trochilus, p. 216 (&). “Mexico”. Now Microdon (Microdon) aurifex (Wiedemann,
1830) (Thompson et al., 1976: 63) (Syrphidae).
1854a:
8. Sargus linearis, p. 318 (&). “Mexico (Mr. Glennie´s collection). Same combination nowadays
(James, 1973: 19) (Stratiomyidae).
Acta Zool. Mex. (n.s.) 84 (2001)
95
1854b:
9. Dasypogon magnificus, p. 427 (&). “Mexico (presented by Mrs. Rouquette)”. Now
Archilestris magnificus (Walker) (Martin & Papavero, 1970: 16) (Asilidae).
1855:
10. Erax aper, p. 621 (%, &). “Mexico (Mr. Glennie´s collection)”. Efferia-group (Martin &
Papavero, 1970: 63) (Asilidae).
11. Mallophora antiqua, p. 581 (&). “Mexico, Veracruz (Sallé)”. Same original combination
(Martin & Papavero, 1970: 72) (Asilidae).
1857:
12. Pipiza divisa, p. 156 (&). “Mexico, Veracruz”. Now Ocyptamus dimidiatus (Fabricius, 1781)
(Thompson et al., 1976: 16) (Syrphidae).
1860:
13. Cyphomyia simplex, p. 268 (&). “Mexico”. Same combination (James, 1973: 28)
(Stratiomyidae).
14. Stratiomys constricta, p. 269 (%). “Mexico”. Same combination (James, 1973: 45)
(Stratiomyidae).
15. Stratiomys pinguis, p. 270 (&). “Mexico”. A junior synonym of Stratiomys constricta Walker
(James, 1973: 45) (Stratiomyidae).
16. Clitellaria obesa, p. 270 (%). “Mexico”. Now Spaniomyia obesa (Walker) (James, 1973: 59)
(Stratiomyidae).
17. Chrysochlora purpurea, p. 271 (%). “Mexico”. Now Chrysochlorina purpurea (Walker)
(James, 1973: 35) (Stratiomyidae).
18. Pangonia atrifera, p. 272 (%). Not from Mexico; actually a junior synonym of the Palearctic
species Pangonia haustellata (Fabricius, 1781) (Fairchild, 1971: 164; Fairchild & Burger,
1994: 153) (Tabanidae).
19. Pangonia tenuirostris, p. 272 (%). “Mexico”. Now Esenbeckia (Ricardoa) flavohirta (Bellardi,
1859) (Fairchild, 1971: 10; Fairchild & Burger, 1994: 27) (Tabanidae).
20. Tabanus dorsifer, p. 273 (&). “Mexico”. Same combination (Fairchild, 1971: 91; Fairchild
& Burger, 1994: 136) (Tabanidae).
21. Tabanus commixtus, p. 273 (&). “Mexico”. Same combination (Fairchild & Burger, 1994:
134) (Tabanidae).
22. Tabanus alteripennis, p. 274 (&). “Mexico”. Now Philipotabanus (Philipotabanus) caliginosus
(Bellardi, 1859) (Fairchild, 1971: 178; Fairchild & Burger, 1994: 116) (Tabanidae).
23. Tabanus purus, p. 274 (&). “Mexico”. Now Phaeotabanus longiappendiculatus (Macquart,
1855) (Fairchild, 1971: 157; Fairchild & Burger, 1994: 89) (Tabanidae).
24. Dasypogon secabilis, p. 276 (&). “Mexico”. Now Blepharepium secabile (Walker) (Martin
& Papavero, 1970: 28) (Asilidae).
25. Dasypogon gelascens, p. 277 (%). “Mexico”. Now Stichopogon trifasciatus (Say, 1823)
(Martin & Papavero, 1970: 5) (Asilidae).
26. Discocephala divisa, p. 279 (&). “Mexico”. Now Holcocephala divisa (Walker) (Martin &
Papavero, 1970: 7) (Asilidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
96
27. Discocephala interlineata, p. 279 (&). “Mexico”. Now Holcocephala nitida (Wiedemann,
1830) (Martin & Papavero, 1970: 7) (Asilidae).
28. Laphria formidolosa, p. 280 (%). “Mexico”. Now Andrenosoma (Pilica) formidolosa (Walker)
(Martin & Papavero, 1970: 44) (Asilidae).
29. Laphria componens, p. 281 (%). “Mexico”. Same combination (Martin & Papavero, 1970:
38) (Asilidae).
30. Laphria triligata, p. 281 (&). “Mexico”. Same combination (Martin & Papavero, 1970: 38)
(Asilidae).
31. Atomosia sericans, p. 282 (%). “Mexico”. Same combination (Martin & Papavero, 1970:
47) (Asilidae).
32. Trupanea lateralis, p. 283 (%). “Mexico”. Now Promachus (Amblyonychus) lateralis (Walker)
(Martin & Papavero, 1970: 81) (Asilidae).
33. Asilus inamatus, p. 283 (&). “Mexico”. An unrecognized species (Martin & Papavero,
1970: 89) (Asilidae).
34. Asilus perrumpeus, p. 283 (%). “Mexico”. Now Triorla interrupta (Macquart, 1834) (Martin
& Papavero, 1970: 69) (Asilidae).
35. Chrysopila trifasciata, p. 284 (&). “Mexico”. Now Chrysopilus trifasciatus (Walker) (James,
1968: 4) (Rhagionidae).
36. Chrysopila basalis, p. 285 (%). “Mexico”. An unrecognized species (James, 1968: 6)
(Rhagionidae).
37. Psilopus solidus, p. 287 (&). Now Condylostylus solidus (Walker) (Robinson, 1970: 10)
(Dolichopodidae).
38. Psilopus peractus, p. 287 (&). “Mexico”. Condylostylus peractus (Walker) (Robinson, 1970:
10) (Dolichopodidae).
39. Psilopus haereticus, p. 287 (&). “Mexico”. Now Condylostylus haereticus (Walker)
(Robinson, 1970: 7) (Dolichopodidae).
40. Psilopus permodicus, p. 288 (%). “Mexico”. Now Condylostylus permodicus (Walker)
(Robinson, 1970: 10) (Dolichopodidae).
41. Ceria cacica, p. 288 (&). “Mexico”. Now Monoceromyia cacica (Walker) (Thompson et al.,
1976: 94) (Syrphidae).
42. Eristalis familiaris, p. 290 (%). “Mexico”. Now Palpada familiaris (Walker) (Thompson et al.,
1976: 105) (Syrphidae).
43. Eristalis expictus, p. 290 (%). “Mexico”. Now Palpada expicta (Walker) (Thompson et al.,
1976: 105) (Syrphidae).
44. Xylota subcostalis, p. 291 (&). “Mexico”. Now Quichuana subcostalis (Walker) (Thompson
et al., 1976: 99) (Syrphidae).
45. Volucella aperta, p. 292 (%). “Mexico”. Now Copestylum apertum (Walker) (Thompson et
al., 1976: 71) (Syrphidae).
46. Temnocera viridula, p. 292 (%). “Mexico”. Now Copestylum viridulum (Walker) (Thompson
et al., 1976: 87) (Syrphidae).
47. Temnocera unilecta, p. 292 (&). “Mexico”. Now Copestylum unilectum (Walker) (Thompson
et al., 1976: 86) (Syrphidae).
48. Syrphus colludens, p. 293 (&). “Mexico”. An unplaced species of Syrphini (Thompson et
al., 1976: 39) (Syrphidae).
Acta Zool. Mex. (n.s.) 84 (2001)
97
49. Jurinia debitrix, p. 296 (&). “Mexico”. Now Juriniella debitrix (Walker) (Guimarães, 1971:
57) (Tachinidae).
50. Jurinia innovata, p. 296 (&). “Mexico”. Now Archytas innovatus (Walker) (Guimarães, 1971:
50) (Tachinidae).
1861:
51. Nemoraea intrita, p. 297 (&). “Mexico”. An unrecognized species of Archytas (Guimarães,
1971: 50) (Tachinidae).
52. Eurigaster saginata, p. 298 (&). “Mexico”. Now Copecrypta nitens (Wiedemann, 1830)
(Guimarães, 1971: 75) (Tachinidae).
53. Eurigaster desita, p. 299 (&). An unrecognized Goniinae (Guimarães, 1971: 215)
(Tachinidae).
54. Eurigaster commetans, p. 299 (&). “Mexico”. Now Bolomyia commetans (Walker)
(Guimarães, 1971: 187) (Tachinidae).
55. Eurigaster fertoria, p. 300 (&). “Mexico”. An unplaced Blondeliini (Guimarães, 1971: 152)
(Tachinidae).
56. Eurigaster habilis, p. 301 (&). “Mexico”. An unplaced Sturmiini (Guimarães, 1971: 194)
(Tachinidae).
57. Eurigaster postica, p. 301 (&). “Mexico”. An unplaced Sturmiini (Guimarães, 1971: 194)
(Tachinidae).
58. Masicera disputans, p. 302 (%). “Mexico”. An unplaced Eryciini (Guimarães, 1971: 214)
(Tachinidae).
59. Masicera gentica, p. 302 (%). “Mexico”. An unplaced Blondeliini (Guimarães, 1971: 152)
(Tachinidae).
60. Masicera necopina, p. 303 (&). “Mexico”. An unplaced Tachinidae (Guimarães, 1971: 216)
(Tachinidae).
61. Masicera expergita, p. 304 (%). “Mexico”. An unplaced Blondeliini (Guimarães, 1971: 152)
(Tachinidae).
62. Lydella cessatrix, p. 305 (&). “Mexico”. An unplaced Goniinae (Guimarães, 1971: 215)
(Tachinidae).
63. Lydella indita, p. 306 (&). “Mexico”. Now Pseudeuantha indita (Walker) (Guimarães, 1971:
104) (Tachinidae).
64. Dexia pertecta, p. 307 (%). “Mexico”. Now Myiobiopsis pertecta (Walker) (Guimarães, 1971:
118) (Tachinidae).
65. Sarcophaga intermutans, p. 308 (&). “Mexico”. An unrecognized species (Lopes, 1969: 54)
(Sarcophagidae).
66. Sarcophaga perneta, p. 308 (%). “Mexico”. Now Oxysarcodexia perneta (Walker) (Lopes,
1969: 27) (Sarcophagidae).
67. Sarcophaga innota, p. 308 (&). “Mexico”. An unrecognized species (Lopes, 1969: 53)
(Sarcophagidae).
68. Sarcophaga conclausa, p. 309 (&). “Mexico”. An unrecognized species (Lopes, 1969: 53)
(Sarcophagidae).
69. Sarcophaga despensa, p. 309 (&). “Mexico”. An unrecognized species (Lopes, 1969: 53)
(Sarcophagidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
98
70. Sarcophaga effrenata, p. 309 (%). “Mexico”. Now Chaetoravinia effrenata (Walker) (Lopes,
1969: 22) (Sarcophagidae).
71. Calliphora femorata, p. 310 (&). “Mexico”. A junior synonym of Hemilucilia segmentaria
(Fabricius, 1805) (James, 1970: 7) (Calliphoridae).
72. Calliphora socors, p. 311 (&). “Mexico”. A junior synonym of Mesembrinella bicolor
(Fabricius, 1805) (James, 1970: 4) (Calliphoridae).
73. Lucilia surrepens, p. 312 (&). “Mexico”. Now Morellia basalis (Walker, 1852) (Pont, 1972:
8) (Muscidae).
74. Pyrellia suspicax, p. 312 (%). Now Morellia bipuncta (Wiedemann, 1830) (Pont, 1972: 8)
(Muscidae).
75. Pyrellia specialis, p. 313 (&). “Mexico”. Now Morellia maculipennis (Macquart, 1846) (Pont,
1972: 9) (Muscidae).
76. Pyrellia scordalus, p. 313 (&). “Mexico”. Now Hemichlora scordalus (Walker) (Pont, 1972:
49) (Muscidae).
77. Musca sensifera, p. 314 (%). “Mexico”. An unrecognized Tachinidae! (Pont, 1972: 60)
(Tachinidae).
78. Aricia rescita, p. 315 (&). “Mexico”. Now Cyrtoneurina rescita (Walker) (Pont, 1972: 53)
(Muscidae).
79. Aricia procedens, p. 315 (&). “Mexico”. Now Helina procedens (Walker) (Pont, 1972: 22)
(Muscidae).
80. Aricia circulatrix, p. 316 (&). “Mexico”. Now Helina circulatrix (Walker) (Pont, 1972: 19)
(Muscidae).
81. Anthomyia protrita, p. 317 (&). “Mexico”. An unrecognized Sarcophagidae! (Pont, 1972:
60) (Sarcophagidae).
82. Hylemyia probata, p. 318 (%). “Mexico”. An unrecognized Tachinidae! (Pont, 1972: 60)
(Tachinidae).
83. Dryomyza maculipes, p. 319 (no sex declared). “Mexico”. An unrecognized species
(Steyskal, 1977: 2) (Dryomyzidae).
84. Sapromyza apta, p. 321 (no sex declared). “Mexico”. Without additional references
(Lauxaniidae).
85. Tetanocera pectoralis, p. 321 (no sex declared). “Mexico”. An unrecognized Otitidae
(Knutson et al., 1976: 14) (Otitidae).
86. Lonchaea discrepans, p. 322 (no sex declared). “Mexico”. Without additional references
(Lonchaeidae).
87. Bricinnia flexivitta, p. 324 (&). “Mexico”. Now Senopterina flexivitta (Walker) (Steyskal,
1968f: 3) (Platystomatidae).
88. Simulium ochraceum, p. 332 (&). “Mexico”. Same combination (Dalmat, 1955: 225)
(Simuliidae).
89. Limnobia stupens, p. 333 (&). “Mexico”. Now Gnophomyia stupens (Walker) (Alexander &
Alexander, 1970: 130) (Tipulidae).
90. Tipula associans, p. 333 (%). “Mexico”. Now Zelandotipula associans (Walker) (Alexander
& Alexander, 1970: 41) (Tipulidae).
91. Tipula dispellens, p. 334 (&). “Mexico”. Now Brachypelmna dispellens (Walker) (Alexander
& Alexander, 1970: 6) (Tipulidae).
Acta Zool. Mex. (n.s.) 84 (2001)
99
6.
T
HE
“S
AGGIO DI
D
ITTEROLOGIA
M
ESSICANA
OF
L
UIGI
B
ELLARDI
Luigi Bellardi was born in Genova, Italy, on May 18, 1818. To please his family
he studied law, but as he always had been attracted to the study of natural
sciences, he soon began studies on fossil molluscs. With his friend Michelotti he
collected fossil shells in the hills of Turin. At only 20 years of age he published his
first paper on fossil molluscs. In order to increase his field action, he traveled to
Egypt, where he made a large collection of nummulitic fossils.
Between 1854 and 1874, however, he became interested in Diptera, publishing
several papers on the fauna of the Piemonte, as well as a monograph on Mexican
Diptera, the first in the history of Dipterology, the famous “Saggio di Ditterologia
Messicana” (Essay on Mexican Dipterology) (1859-1862).
For the publication of the “Saggio”, Bellardi had access to several collections:
-
Those collected by Eugenio Truqui (or Truqui) (sardinian consul in Cyprus; died in Rio de
Janeiro in April, 1860), about whose life and travels we could find nothing
else).Approximately 50 species, collected in the neighbourhood of Mexico City, by a
certain Ettore Craveri (1815-1890), about whom nothing is known (Papavero, 1973; Morisi,
1980; Passerin d´Entrèves, 1983).
- The duplicates of Mexican species in the Paris Museum, sent to Bellardi by the director of
the Museum, Milne-Edwards; some of these specimens had been studied by Macquart (cf.
part II of this work).
- The collection of the Reale Museo di Zoologia di Torino, loaned by the Director, Filippo di
Filippi.
- Specimens collected by Sallé in several parts of Mexico.
- Mexican material in the Bigot collection, which had been identified by Macquart.
- Approximately 100 specimens collected by Sumichrast and Saussure.
*****
Auguste Sallé died in Paris on May 5, 1896, in his 70
th
year. He travelled
extensively in the United States, Mexico, the West Indies, Central America, and
Venezuela, making collections in all branches of entomology. On his return to
Europe he established himself as a natural history agent in Paris. His Central
American collections were afterwards purchased by Godman and Salvin for the
Biologia Centrali-Americana”. Sallé was elected fellow of the French Entomological
Society in 1875 and of the Entomological Society of London in 1875 (Anon., 1896;
Barnhart, 1965; Binford, 1989; Sclater, 1858; Smith & Smith, 1973).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
100
Adrien Louis Jean de Sumichrast was born in Yvonne, Canton de Vaud,
Switzerland, on October 15, 1828. His studies were made in Lausanne, Geneva
and Berne. He soon became interested in the study of natural history and, as the
European fauna was not enough for him, he decided to accompany De Saussure
on his trip to Mexico.
Henri Louis Frederic De Saussure was born at Geneva, Switzerland, on
November 27, 1829, and died there on February 20, 1905. He received his
elementary education at Briquet, and his advanced training at the Institute of
Fellenberg. He studied under the entomologist François Jules Pictet de la Rive, who
directed his attention to insects. The early part of his entomological career was
spent in the study of the Hymenoptera and latter part in Orthoptera, his greatest
reputation being made with the last named order. After several years of study in
Paris, where he received the degree of licentiate of the Faculty of Paris, he began
his travels in 1854, going first to the West Indies, then to Mexico, and finally to the
United States, where he met Louis Agassiz and other scientists. He returned to
Europe in 1856 with valuable collections of insects, myriapods, crustaceans, birds
and other groups. Aside from his interest in entomology, he also studied geography
and ethnology. In 1858, he founded the Geographical Society of Geneva and was
its president from 1888 to 1889. For many years he was a member of the committee
which managed the Natural History Museum of Geneva. There he amassed the
finest collection of Hymenoptera and Orthoptera in the world. In 1892 he was
elected an Honorary Fellow of the Entomological Society of London (Essig, 1931;
Kellogg, 1932).
Sumichrast and Saussure arrived in Veracruz, Mexico, in April 1855, remaining
for a few days in that city. Proceeding thence to Córdoba (written “Cordova” by
Bellardi), they passed by tospán on April 16 (an “hacienda” not far from Córdoba),
where they visited Auguste Sallé, who was exploring Mexico with Boucard. After the
short stay in Córdoba the naturalists resumed their journey, going to Orizaba,
Puebla, Mexico City, Tampico, and other cities in the interior. Sumichrast collected
in those places with Saussure for about one year and those collections were taken
back to Geneva by Saussure, when he left Mexico, due to the troubles of travelling
and the political unrest caused by several revolutions. Sumichrast decided to stay
in Mexico. He married in Cuchitán (Juchitán), Oaxaca on August 30, 1870, and from
this time to his death he was occupied with the scientific exploration of Mexico,
visiting the States of Veracruz, Puebla, Mexico, Oaxaca (Tehuantepec), and
Chiapas. In the latter state death overtook him, in Tonalá, on September 26, 1882
(Boucard, 1884; Binford, 1989; Hemsley, 1887; Kellogg, 1932; Knobloch, 1983;
Papavero, 1973; Smith & Smith, 1973; Wauer, 1992).
Sallé´s companion, Adolphe Boucard, seems to have been a general collector,
but more especially devoted to birds and insects. He wrote an account of his
Acta Zool. Mex. (n.s.) 84 (2001)
101
extensive travels through the United States, Mexico, Central America, Colombia,
Chile and other countries (1894). In 1867, he published a catalogue of the natural
history specimens he had gathered in Mexico. In 1878, he published another
catalogue of the collections obtained in Guatemala, which were exhibited at the
Exposition Universalle de Paris” and perhaps about the same time a list of the
Coleoptera offered for sale. According to Carriker (1910), he collected in Costa Rica
in 1877. Boucard published a list of the birds collected during that trip (1878)
(Papavero, 1971b).
*****
Bellardi was also assistant of the Mineralogical Museum (since 1844) and curator
of the paleontological collections of the geological museum of Turin. He taught for
30 years at the girls college (Regina Margheritta) and in the “Liceo Reale Gilberti”
in Turin. He was elected member of several cultural and scientific societies. From
1870 on, he went back to his studies of fossil molluscs, dying in Turin on September
17, 1889 ( Lessona, 1884; Papavero, 1973; Sacco, 1889).
Bellardi´s “Saggio di Ditterologia Messicana” contains 266 species, 176 of which
were proposed as new. The collection is housed at the Istituto e Museo di Zoologia2
Sistematica, Università di Torino, together with Bellardi´s library. The list of the
species proposed by Bellardi in the “Saggio” is the following:
1859:
1. Culex mexicanus, p. 5 (&). “Messico (Sallé)”. Box 11. Apparently a distinct species, as
currently interpreted, now Psorophora (Janthinosoma) mexicana (Bellardi), according to
Belkin (1968: 27) (Culicidae).
2. Tipula nebulosa, p. 6 (%). “Messico (Sallé)”. Box 11. Now Epiphragma (Epiphragma)
nebulosa (Bellardi) (Alexander & Alexander, 1970: 97) (Tipulidae).
3. Tipula craverii, p. 7, pl. I, fig. 1 (%). “Messico (Craveri)”. Box 11. Now Tipula (Bellardina)
craverii Bellardi (Alexander & Alexander, 1970: 20) (Tipulidae).
4. Tipula edwardsi, p. 8, pl. I, fig. 2 (%). “Messico (Sallé)”. Box 11. Now Tipula (Bellardina)
edwardsi Bellardi (Alexander & Alexander, 1970: 20) (Tipulidae).
5. Tipula quadrimaculata, p. 9, pl. I, fig. 3 (&). “Messico, Cordoba (Saussure)”. Box 11. An
unplaced Tipulinae (Alexander & Alexander, 1970: 44) (Tipulidae).
6. Tipula proxima, p. 9 (&). “Messico (Craveri)”. Box 11. Now a junior synonym of Nephrotoma
ferruginea (Fabricius, 1805) (Alexander & Alexander, 1970: 17) (Tipulidae).
7. Tipula affinis, p. 10 (%). “Messico (Truqui)”. Box 11. Now Nephrotoma affinis (Bellardi)
(Alexander & Alexander, 1970: 16) (Tipulidae).
8. Tipula nigrolutea, p. 11 (%). “Messico (Sallé)”. Box 11. Now Nephrotoma nigrolutea (Bellardi)
(Alexander & Alexander, 1970: 17) (Tipulidae).
9. Sciophila popocatepetli, p. 11 (&). “Messico, Popocatépetl, 3800m (Truqui)”. Box 11. Same
combination (Papavero, 1978: 53) (Mycetophilidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
102
10. Simulium cinereum, p. 13 (%, &). “Messico, Morelia (Saussure)”. Box 11. Preocc.
Macquart, 1834). Now Simulium virgatum Coquillett, 1902 (Vulcano, 1967: 23)
(Simuliidae).
11. Simulium metallicum, p. 14 (%). “Messico (Sallé)”. Box 11. Same combination (Vulcano,
1967: 16) (Simuliidae).
12. Plecia nigerrima, p. 14 (&). “Messico, contorni di Messico (Sallé), Orizaba, Meztillán
(Saussure)”. Box 11. Now Penthetria nigerrima (Bellardi) (Hardy, 1966: 2) (Bibionidae).
13. Plecia rostrata, p. 15 (%, &). “Messico (Sallé)”. Box 11. Now Plecia rostellata Loew, 1858
(Hardy, 1966: 5) (Bibionidae).
14. Plecia bicolor (and var.), p. 16 (%, &). “Messico, Cordova (the type) (Saussure), Messico,
Orizaba (Saussure) (the variety). Box 11. Same combination (Hardy, 1966: 2)
(Bibionidae).
15. Bibio criorhinus, p. 17 (%). “Messico (Sallé)”. Box 11. Same combination (Hardy, 1966: 6)
(Bibionidae).
16. Bibio piceus, p. 17 (%). “Messico (Craveri)”. Box 11. Same combination (Hardy, 1966: 6)
(Bibionidae).
17. Bibio dubius, p. 18 (&). Messico (Craveri)”. Box 11. Now Bibio vicinus Lynch Arribálzaga,
1878 (Hardy, 1966: 6) (Bibionidae).
18. Bibio fuligineus, p. 19 (&). “Messico (Craveri e Sallé), Orizaba (Saussure)”. Box 11. Same
combination (Hardy, 1966: 6) (Bibionidae).
19. Dilophus maculatus, p. 19, pl. I, fig. 5 (&). “Messico (Truqui). Box 11. Same combination
(Hardy, 1966: 9) (Bibionidae).
20. Beris mexicana, p. 20, pl. I, fig. 6 (%, &). “Messico (Sallé)”. Box 11. Now Oplachantha
mexicana (Bellardi) (James, 1973: 8) (Stratiomyidae).
21. Cyphomyia similis, p. 23, pl. I, fig. 7 (&). “Messico (Truqui)”. Box 11. Now a junior synonym
of Cyphomyia tomentosa Gerstaeker, 1857 (James, 1973: 28) (Stratiomyidae).
22. Hermetia lativentris, p. 27, pl. I, fig. 9 (&). “Messico, Tampico (Saussure)”. Box 11. Same
combination (James, 1973: 38) (Stratiomyidae).
23. Hermetia aurata, p. 27, pl. I, fig. 8 (&, %). “Messico, Morelia (Saussure)”. Box 11. Same
combination (James, 1973: 37) (Stratiomyidae).
24. Stratiomys gerstaeckeri, p. 31, pl. I, fig. 10 (%). “Messico, contorni di Messico (Sallé)”. Box
11. Now a junior synonym of Hoplitimyia (Hoplitimyia) fasciata (Fabricius, 1787) (James,
1973: 42) (Stratiomyidae).
25. Odonthomyia truquii, p. 34, pl. I, fig. 11 (&). “Messico, Morelos, Cuernavaca (Truqui)”. Box
11. Now Hedriodiscus truquii (Bellardi) (James, 1973: 41) (Stratiomyidae).
26. Odonthomyia affinis, p. 35, pl. I, fig. 12 (&). “Messico, Puebla (Saussure)”. Box 2. Now a
junior synonym of Hedriodiscus euchlorus (Gerstaecker, 1857) (James, 1973: 40)
(Stratiomyidae).
27. Odonthomyia dissimilis, p. 35, pl. I, figs. 13-14 (&, %). “Messico, contorni di Messico
(Truqui)”. Box 2. Same combination (in Odontomyia) (James, 1973: 44) (Stratiomyidae).
28. Odonthomyia viridis, p. 36, pl. I, fig. 16 (&). “Messico, Morelos, Cuautla (as Cuantla)
(Saussure)”. Box 2. Now Labostigmina viridis (Bellardi) (James, 1973: 43) (Stratiomyidae).
29. Odonthomyia quadrimaculata, p. 37, pl. I, fig. 15 (%). “Messico (Craveri)”. Box 2. Now
Labostigmina quadrimaculata (Bellardi) (James, 1973: 43) (Stratiomyidae).
Acta Zool. Mex. (n.s.) 84 (2001)
103
30. Odonthomyia femorata, p. 37 (&). “Messico, Toluca (Saussure)”. Box 2. A junior synonym
of the preceding species (James, 1973: 43) (Stratiomyidae).
31. Odonthomyia tritaeniata, p. 38, pl. I, fig. 17 (&). “Messico, contorni di Messico (Sallé),
Cuautla (as Cuantla) (Saussure)”. Box 2. A junior synonym of Hedriodiscus trivittatus (Say,
1829) (James, 1973: 41) (Stratiomyidae).
32. Sargus caesius, p. 40, pl. I, fig. 18 (&). “Messico, Angangueo (as Angang) (Saussure)”.
Box 2. Unchanged (James, 1973: 18) (Stratiomyidae).
33. Sargus latus, p. 41, pl. I, fig. 19 (%). “Messico (Sallé)”. Box 2. Unchanged (James, 1973:
18) (Stratiomyidae).
34. Sargus aureus, p. 42, pl. I, fig. 20 (%). “Messico (Sallé)”. Box 2. Now a junior synonym of
Sargus speciosus Macquart, 1846 (James, 1973: 20) (Stratiomyidae).
35. Sargus sallei, p. 43 (%). “Messico (Sallé)”. Box 2. Now a junior synonym of Sargus
fasciatus Fabricius, 1805 (James, 1973: 19) (Stratiomyidae).
36. Sargus subinterruptus, p. 44, pl. I, fig. 22 (&). “Messico (Sallé)”. Box 2. Now Merosargus
subinterruptus (Bellardi) (James, 1973: 15) (Stratiomyidae).
37. Pangonia rhinophora, p. 46, pl. II, fig. 1 (&). “Messico (Sallé)”. Box 2. Now Fidena (Fidena)
rhinophora (Bellardi) (Fairchild, 1971: 23; Fairchild & Burger, 1994: 46) (Tabanidae).
38. Pangonia rostrifera, p. 47 (%, &). “Messico (Sallé)”. Box 2. Now a junior synonym of Scione
aurulans (Wiedemann, 1830) (Fairchild, 1971: 25; Fairchild & Burger, 1994: 49)
(Tabanidae).
39. Pangonia wiedemanni, p. 48, pl. II, fig. 3 (&). “Messico (Sallé)”. Box 2. Now Esenbeckia
(Ricardoa) wiedemanni (Bellardi) (Fairchild, 1971: 12; Fairchild & Burger, 1994: 28)
(Tabanidae).
40. Pangonia saussurei, p. 49, pl. II, fig. 4 (&). “Messico, Veracruz, Orizaba (Sallé)”. Box 2.
Now Esenbeckia (Ricardoa) planiventris saussurei (Bellardi) (Fairchild, 1971: 11; Fairchild
& Burger, 1994: 28) (Tabanidae).
41. Pangonia flavohirta, p. 49 (&). “Messico (Sallé)”. Box 2. Now Esenbeckia (Ricardoa)
flavohirta (Bellardi) (Fairchild, 1971: 10; Fairchild & Burger, 1994: 26) (Tabanidae).
42. Pangonia sallei, p. 50 (%). “Messico (Sallé)”. Box 2. A synonym of the preceding species
(Fairchild, 1971: 10; Fairchild & Burger, 1994: 27) (Tabanidae).
43. Pangonia incerta, p. 52 (&). “Messico, Veracruz, Orizaba (Saussure)”. Box 2. Now
Esenbeckia (Ricardoa) incerta (Bellardi) (Fairchild, 1971: 11; Fairchild & Burger, 1994: 27)
(Tabanidae).
44. Tabanus subruber, p. 55 (&). “Messico (Sallé, Sumichrast)”. Box 2. Unchanged (Fairchild,
1971: 102; Fairchild & Burger, 1994: 150) (Tabanidae).
45. Tabanus sumichrasti, p. 56 (&). “Messico (Sumichrast)”. Box 2. A synonym of the
preceding species (Fairchild, 1971: 102; Fairchild & Burger, 1994: 150) (Tabanidae).
46. Tabanus albonotatus, p. 56, pl. II, fig. 5 (&). “Messico (Sallé), Tamaulipas, Tampico
(Saussure)”. Box 2. Now a junior synonym of Tabanus oculus Walker, 1848 (Fairchild,
1971: 98; Fairchild & Burger, 1994: 144) (Tabanidae).
47. Tabanus defilippii, p. 57 (&). “Messico (Sumichrast)”. Box 2. Unchanged (Fairchild, 1971:
91; Fairchild & Burger, 1994: 135) (Tabanidae).
48. Tabanus bigoti, p. 58 (?&, without head). “Messico (Coll. Bigot)”. Type in Oxford.
Unchanged (Fairchild, 1971: 89; Fairchild & Burger, 1994: 132) (Tabanidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
104
49. Tabanus luteoflavus, p. 60 (&). “Messico (Sumichrast)”. Box 2. Now a junior synonym of
Phaeotabanus longiappendiculatus (Macquart, 1855) (Fairchild, 1971: 57; Fairchild &
Burger, 1994: 89) (Tabanidae).
50. Tabanus craverii, p. 60 (&). “Messico (Craveri)”. Box 2. Same combination (Fairchild &
Burger, 1994: 135) (Tabanidae).
51. Tabanus flavocinctus, p. 61 (&). “Mexico”. Types in Paris and Turin. Locality in error, as
this is a synonym of Hybomitra zonalis (Kirby, 1837), a Nearctic species (Fairchild, 1971:
105; Fairchild & Burger, 1994: 154) (Tabanidae).
52. Tabanus sallei, p. 61, pl. II, fig. 7 (%). “Messico (Sallé)”. Box 2. Now a junior synonym of
Tabanus pungens Wiedemann, 1828 (Fairchild, 1971: 100; Fairchild & Burger, 1994: 147)
(Tabanidae).
53. Tabanus carneus, p. 62 (%). “Messico (Sallé)”. Box 2. Now a junior synonym of Tabanus
occidentalis var. dorsovittatus Macquart, 1855 (Fairchild & Burger, 1994: 143) (Tabanidae).
54. Tabanus truquii, p. 64, pl. II, fig. 6 (%). “Messico (Truqui)”. Box 2. Now a junior synonym
of Tabanus colombensis Macquart, 1846 (Fairchild & Burger, 1994: 134) (Tabanidae).
55. Tabanus propinquus, p. 65 (&). “Messico (Sallé, Sumichrast)”. Types in Paris and Turin
(Box 2). Now a junior synonym of Tabanus pungens Wiedemann, 1828 (Fairchild, 1971:
100; Fairchild & Burger, 1994: 147) (Tabanidae).
56. Tabanus subsimilis, p. 66 (&). Messico (Sallé). Types in Paris and Turin (Box 2).
Unchanged (Fairchild, 1971: 102; Fairchild & Burger, 1994: 150) (Tabanidae).
57. Tabanus nigropunctatus, p. 67, pl. II, fig. 8 (&). “Messico, Cordova, Huastec (Saussure)”.
Box 2. A junior synonym of Poeciloderas quadripunctatus (Fabricius, 1805) (Fairchild,
1971: 86; Fairchild & Burger, 1994: 127) (Tabanidae).
58. Tabanus aurantiacus, p. 69, pl. II, fig. 9 (&). “Messico (Caveri), Orizaba (Saussure)”. Box
2. Now Agkistrocerus aurantiacus (Bellardi) (Fairchild & Burger, 1994: 128) (Tabanidae).
59. Tabanus caliginosus, p. 68, pl. II, fig. 10 (&). “Messico (Sallé)”. Box 3. Now Philipotabanus
(Philipotabanus) caliginosus (Bellardi) (Fairchild, 1971: 78; Fairchild & Burger, 1994: 116)
(Tabanidae).
60. Tabanus rondanii, p. 68, pl. II, fig. 11 (&). “Messico (Saussure, Sallé)”. Box 3. Now a junior
synonym of Diachlorus ferrugatus (Fabricius, 1805) (Fairchild, 1971: 53; Fairchild &
Burger, 1994: 83) (Tabanidae).
61. Chrysops subcaecutiens, p. 69, pl. II, fig. 13 (%). “Messico, contorni di Messico
(Saussure)”. Box 3. Unchanged (Fairchild, 1971: 534; Fairchild & Burger, 1994: 60)
(Tabanidae).
62. Chrysops affinis, p. 70, pl. II, fig. 14 (%). “Messico, contorni di Messico (Truqui)”. Box 3.
Unchanged (Fairchild, 1971: 30; Fairchild & Burger, 1994: 55) (Tabanidae).
63. Chrysops virgulatus, p. 71, pl. II, fig. 17 (%). “Messico, Cuautla (as Cuantla) (Saussure)”.
Box 3. Unchanged (Fairchild, 1971: 35; Fairchild & Burger, 1994: 61) (Tabanidae).
64. Chrysops latifasciatus, p. 71, pl. II, fig. 15 (&). “Messico (Sallé)”. Box 3. Unchanged
(Fairchild, 1971: 32; Fairchild & Burger, 1994: 58) (Tabanidae).
Acta Zool. Mex. (n.s.) 84 (2001)
105
65. Chrysops scalaratus, p. 72, pl. II, fig. 19 (&, without head). “Messico (Sumichrast)”. Box
3. Unchanged (Fairchild, 1971: 34; Fairchild & Burger, 1994: 60) (Tabanidae).
66. Chrysops apicalis, p. 73 (%). “Messico (Sallé). Box 3. A synonym of the preceding species
(Fairchild, 1971: 34; Fairchild & Burger, 1994: 60) (Tabanidae).
67. Chrysops pallidus, p. 73, pl. II, fig. 16 (&). “Messico (Sumichrast)”. Box 3. Now a junior
synonym of Chrysops flavidus Wiedemann, 1821 (Fairchild, 1971: 31; Fairchild & Burger,
1994: 57) (Tabanidae).
68. Chrysops megaceras, p. 74, pl. II, fig. 18 (&). “Messico (Sallé, Sumichrast). Box 3. Now
Silvius (Assipala) megaceras (Bellardi) (Fairchild, 1971: 29; Fairchild & Burger, 1994: 54)
(Tabanidae).
69. Philopota truquii, p. 77, pl. II, figs 20, 20bis (%). “Messico, Cuazimalpa (actually
Cuajimalpa, State of Mexico) (Truqui)”. Box 3. (Acroceridae).
1861:
70. Mydas bitaeniatus, p. 107, pl. I, fig. 1 (%). “Messico, Playa Vicente (Sallé)”. Box 3. Now
Stratiomydas bitaeniatus (Bellardi) (Papavero, 1996: 632) (Mydidae).
71. Mydas tricinctus, p. 108, pl. I, fig. 2 (%). “Messico, contorni di Messico (Truqui, Sallé)”. Box
3. Unchanged (Papavero, 1996: 631) (Mydidae).
72. Mydas subinterruptus, p. 110, pl. I, fig. 3 (%). “Messico, Angangueo (as Angang),
Pátzcuaro (Saussure)”. Box 3. Unchanged (Papavero, 1996: 631) (Mydidae).
73. Pseudorus bicolor, p. 111, pl. I, fig. 20 (&). “Messico, Playa Vicente (Sallé)”. Box 3. Now
Pseudoryclus bicolor (Bellardi) (Martin & Papavero, 1970: 57) (Asilidae).
74. Lampria circumdata, p. 115, pl. I, fig. 17 (&). “Messico (Sumichrast)”. Box 3. Unchanged
(Martin & Papavero, 1970: 40) (Asilidae).
75. Laphria cinerea, p. 116, pl. I, fig. 16 (&). “Messico (Sallé)”. Box 3. Now Andrenosoma
cinerea (Bellardi) (Martin & Papavero, 1970: 42) (Asilidae).
76. Laphria cincta, p. 118, pl. I, fig. 19 (%). “Messico (Sallé)”. Box 3. Now Andrenosoma cincta
(Bellardi) (Martin & Papavero, 1970: 42) (Asilidae).
77. Atomosia nigripennis, p. 119 (%, &). “Messico (Sallé)”. Box 3. Now Cerotainia nigripennis
(Bellardi) (Martin & Papavero, 1970: 52) (Asilidae).
78. Atomosia macquarti, p. 120 (&). “Messico (Craveri)”. Box 3. Unchanged (Martin &
Papavero, 1970: 46) (Asilidae).
79. Atomosia bigoti, p. 120 (%). “Messico (Sallé) (Coll. Bigot)”. Type in Oxford. Unchanged
(Martin & Papavero, 1970: 46) (Asilidae).
80. Mallophora craverii, p. 122 (&). “Messico, contorni di Messico (Craveri); Oaxaca (Sallé)”.
Box 4. Unchanged (Martin & Papavero, 1970: 73) (Asilidae).
81. Promachus cinctus, p. 125, pl. II, fig. 2 (%, &). “Messico (Sallé)”. Box 4. Unchanged (Martin
& Papavero, 1970: 80) (Asilidae).
82. Promachus magnus, p. 126 (&). “Messico (Saussure)”. Box 4. Unchanged (Martin &
Papavero, 1970: 80) (Asilidae).
83. Promachus quadratus, p. 127, pl. II, fig. 3. “Messico (Sallé)”. Box 4. Now Promachus
bellardii Martin, 1965 (Martin & Papavero, 1970: 79) (Asilidae).
84. Promachus trapezoidalis, p. 128, pl. II, fig. 4 (%, &). “Messico (Sallé)”. Box 4. Now
Promachus (Amblyonychus) trapezoidalis (Bellardi) (Martin & Papavero, 1970: 81)
(Asilidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
106
85. Promachus pulchellus, p. 129, pl. II, fig. 5 (%). “Messico (Sallé)”. Box 4. Now Promachus
(Amblyonychus) pulchellus (Martin & Papavero, 1970: 81) (Asilidae).
86. Promachus truquii, p. 130, pl. II, fig. 6 (%&). “Messico (Truqui)”. Box 4. Unchanged (Martin
& Papavero, 1970: 80) (Asilidae).
87. Erax anomalus, p. 132, pl. II, fig. 7 (%&). “Messico, Cuautla (as Cuantla) (Saussure)”. Box
4. Now Efferia anomala (Bellardi) (Martin & Papavero, 1970: 63) (Asilidae).
88. Erax comatus, p. 134 (%). “Messico (Saussure)”. Box 4. Now a junior synonym of Efferia
aper (Walker, 1855) (Martin & Papavero, 1970: 63) (Asilidae).
89. Erax parvulus, p. 135, pl. II, fig. 8 (%). “Messico (Truqui)”. Box 4. Now Efferia parvula
(Bellardi) (Martin & Papavero, 1970: 67) (Asilidae).
90. Erax carinatus, p. 136, pl. II, fig. 9 (%). “Messico (Truqui)”. Box 4. Now Efferia carinata
(Bellardi) (Martin & Papavero, 1970: 64) (Asilidae).
91. Erax unicolor, p. 137 (&). “Messico, Orizaba (Saussure)”. Box 4. Now a junior synonym
of Efferia aper (Walker, 1855) (Martin & Papavero, 1970: 63) (Asilidae).
92. Erax eximius, p. 138 (%). “Messico, contorni di Messico (Truqui)”. Box 4. Now Efferia
eximia (Bellardi) (Martin & Papavero, 1970: 64) (Asilidae).
93. Erax cinerescens, p. 139, pl. II, fig. 10 (%, &). “Messico (Sallé), Tuxpango presso Orizaba
(Sumichrast)”. Box 4. Now a junior synonym of Efferia albibarbis (Macquart, 1838) (Martin
& Papavero, 1970: 62) (Asilidae).
94. Erax tricolor, p. 140, pl. II, fig. 12 (%, &). “Messico, contorni di Messico (Truqui)”. Box 4.
Now a junior synonym of Efferia albibarbis (Macquart, 1838) (Martin & Papavero, 1970:
62) (Asilidae).
95. Erax affinis, p. 141 (%). “Messico, Cordova (Saussure)”. Box 4. Now Efferia affinis
(Bellardi) (Martin & Papavero, 1970: 62) (Asilidae).
96. Erax cingulatus, p. 142 (%). “Messico, Cuautla (as Cuantla) (Saussure)”. Box 4. Now
Efferia cingulata (Bellardi) (Martin & Papavero, 1970: 64) (Asilidae).
97. Erax quadrimaculatus, p. 144, pl. II, fig. 13 (%). “Messico, playa Vicente (Sallé); Cordova
(Saussure)”. Box 4. Now Efferia quadrimaculata (Bellardi) (Martin & Papavero, 1970: 67)
(Asilidae).
98. Erax bimaculatus, p. 145, pl. II, fig. 11 (&). “Messico (Sallé)”. Box 4. Now Efferia bimaculata
(Bellardi) (Martin & Papavero, 1970: 63) (Asilidae).
99. Erax marginatus, p. 146 (&). “Messico, Cuautla (as Cuantla) (Saussure)”. Box 4. Now
Efferia marginata (Bellardi) (Martin & Papavero, 1970: 66) (Asilidae).
100. Erax bicolor, p. 147 (&). “Messico, contorni di Messico (Saussure)”. Box 4. Now Efferia
bicolor (Bellardi) (Martin & Papavero, 1970: 63) (Asilidae).
101. Erax nigripes, p. 118 (%). “Messico (Sallé) (Coll. Bigot)”. Type in Oxford. Now Eicherax
nigripes (Bellardi) (Martin & Papavero, 1970: 70) (Asilidae).
102. Erax villosus, p. 149 (%). “Messico (Sallé)”. Box 4. Now a junior synonym of Triorla
interrupta (Macquart, 1834) (Martin & Papavero, 1970: 69) (Asilidae).
103. Proctacanthus craverii, p. 150 (%, &). “Messico, contorni di Messico (Craveri)”. Box 4.
Unchanged (Martin & Papavero, 1970: 78) (Asilidae).
104. Asilus humilis, p. 151 (&). “Messico (Truqui)”. Box 4. Now Machimus (Tolmerus) humilis
(Bellardi) (Martin & Papavero, 1970: 86) (Asilidae).
105. Asilus truquii, p. 152 (%). “Messico (Truqui)”. Box 4. Now Philonicus truquii (Bellardi)
(Martin & Papavero, 1970: 85) (Asilidae).
Acta Zool. Mex. (n.s.) 84 (2001)
107
106. Asilus fuliginosus, p. 152 (%). “Messico”. Box 5. Now Philonicus fuliginosus (Bellardi)
(Martin & Papavero, 1970: 85) (Asilidae).
107. Asilus niveibarbus, p. 153 (%, &). “Messico, Cordova (Saussure)”. Box 5. Now Machimus
niveibarbus (Bellardi) (Martin & Papavero, 1970: 86) (Asilidae).
108. Asilus albospinosus, p. 154 (%). “Messico, contorni di Messico (Truqui)”. Box 5. Now
Philonicus albospinosus (Bellardi) (Martin & Papavero, 1970: 85) (Asilidae).
109. Asilus taeniatus, p. 155 (&). “Messico (Craveri)”. Box 5. Now Philonicus taeniatus
(Bellardi) (Martin & Papavero, 1970: 85) (Asilidae).
110. Asilus infuscatus, p. 156, Pl. I, fig. 15 (%). “Messico (Sallé)”. Box 5. Now Machimus
infuscatus (Bellardi) (Martin & Papavero, 1970: 86) (Asilidae).
111. Asilus apicalis, p. 157 (&). “Messico (Sallé)”. Box 5. Now Ommatius apicalis (Bellardi)
(Martin & Papavero, 1970: 59) (Asilidae).
112. Asilus megacephalus, p. 158, pl. I, fig. 14 (%). “Messico (Sallé)”. Box 5. Now Ommatius
megacephalus (Bellardi) (Martin & Papavero, 1970: 59) (Asilidae).
113. Ceraturgus vitripennis, p. 160 (%). “Messico, Cuautla (as Cuantla) (Saussure)”. Box 5.
Now Taracticus vitripennis (Bellardi) (Martin & Papavero, 1970: 35) (Asilidae).
114. Dasypogon jalapensis, p. 165, pl. I, fig. 5 (&). “Messico, Jalapa (Truqui)”. Box 5. Now
Diogmites jalapensis (Bellardi) (Martin & Papavero, 1970: 26) (Asilidae).
115. Dasypogon goniostigma, p. 165. pl. I, fig. 6 (%, &). “Messico, Playa Vicente (Sallé)”. Box
5. Now Diogmites goniostigma (Bellardi) (Martin & Papavero, 1970: 25) (Asilidae).
116. Dasypogon cuantlensis, p. 167 (%, &). “Messico, Cuautla (as Cuantla) (Saussure)”. Box
5. Now Diogmites cuantlensis (Bellardi) (Martin & Papavero, 1970: 25) (Asilidae).
117. Dasypogon craverii, p. 168 (%, &). “Messico (Craveri)”. Box 5. Now Diogmites craverii
(Bellardi) (Martin & Papavero, 1970: 25) (Asilidae).
118. Dasypogon virescens, p. 169 (%). “Messico (Truqui)”. Box 5. Now Diogmites virescens
(Bellardi) (Martin & Papavero, 1970: 27) (Asilidae).
119. Dasypogon sallei, p. 170 (&). “Messico (Sallé), Tuxpango (Sumichrast)”. Box 5. Now
Diogmites sallei (Bellardi) (Martin & Papavero, 1970: 27) (Asilidae).
120. Dasypogon bigoti, p. 170 (%). “Mexico (Coll. Bigot)”. Type in Oxford. Lost. An
unrecognized species (Martin & Papavero, 1970: 88) (Asilidae).
121. Dasypogon rubescens, p. 171 (%). “Messico (Truqui), Tuxpango (Sumichrast)”. Box 5.
Now Diogmites rubescens (Bellardi) (Martin & Papavero, 1970: 27) (Asilidae).
122. Dasypogon tricolor, p. 172 (%, &). “Messico (Coll. Bigot)”. Box 5. Now Diogmites tricolor
(Bellardi) (Martin & Papavero, 1970: 27) (Asilidae).
123. Dasypogon affinis, p. 173 (%). “Messico (Sallé)”. Box 5. Now Diogmites affinis (Bellardi)
(Martin & Papavero, 1970: 24) (Asilidae).
124. Dasypogon dubius, p. 174 (&). “Messico, Cuautla (as Cuantla) (Saussure). Box 5. Now
Diogmites dubius (Bellardi) (Martin & Papavero, 1970: 25) (Asilidae).
125. Dasypogon nigripes, p. 175 (&). “Messico, Playa Vicente (Sallé)”. Box 5. Now Diogmites
nigripes (Bellardi) (Martin & Papavero, 1970: 26) (Asilidae).
126. Dasypogon truquii, p. 176, pl. I, fig. 10 (&). “Messico (Truqui)”. Box 5. Now Stenopogon
truquii (Bellardi) (Martin & Papavero, 1970: 13) (Asilidae).
127. Dasypogon humilis, p. 177 (%). “Messico, Cuautla (as Cuantla) (Saussure). Box 5. Now
Cophura humilis (Bellardi) (Martin & Papavero, 1970: 34) (Asilidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
108
128. Dasypogon quadrimaculatus, p. 180, pl. I, fig. 8 (%). “Messico (Saussure)”. Box 5. Now
Dizonias tristis (Walker, 1851) (Martin & Papavero, 1970: 14) (Asilidae).
129. Dasypogon lucasi, p. 181, pl. I, fig. 7 (%). “Messico (Craveri)”. Box 5. Now Dizonias lucasi
(Bellardi) (Martin & Papavero, 1970: 14) (Asilidae).
130. Dasypogon spathulatus, p. 182, pl. I, fig. 9 (%). “Messico (Truqui)”. Box 5. Now
Stenopogon (Ospriocerus) abdominalis (Say, 1824) (Martin & Papavero, 1970: 13)
(Asilidae).
131. Discocephala minuta, p. 183 (%, &). “Messico,Tuxpango presso Orizaba (Sumichrast)”.
Box 5. Now Holcocephala minuta (Bellardi) (Martin & Papavero, 1970: 7) (Asilidae).
132. Discocephala deltoidea, p. 185, pl. I, fig. 12 (&). “Messico (Sallé)”. Box 6. Now
Holcocephala deltoidea (Bellardi) (Martin & Papavero, 1970: 6) (Asilidae).
133. Discocephala longipennis, p. 186, pl. I, fig. 14 (%). “Messico, Jalapa (Truqui)”. Box 6. Now
Holcocephala longipennis (Bellardi) (Martin & Papavero, 1970: 7) (Asilidae).
134. Discocephala affinis, p. 186, pl. I, fig. 13 (%, &). “Messico (Sallé)”. Box 6. Now
Holcocephala affinis (Bellardi) (Martin & Papavero, 1970: 6) (Asilidae).
135. Leptogaster truquii, p. 187, pl. I, fig. 18 (%). “Messico, Jantepec (Truqui)”. Box 6. Now a
synonym of Leptogaster fervens Wiedemann, 1830 (Martin, 1968: 3, as Leptogastridae)
(or Asilidae).
136. Thereva crassicornis, p. 188, pl. II, fig. 16 (%). “Messico (Truqui)”. Box 6. Now
Ozodiceromya crassicornis (Bellardi) (Irwin & Lyneborg, 1980: 257) (Therevidae).
137. Thereva argentata, p. 189 (%). “Messico, contorni di Messico; Cordova (Saussure)”. Box
6. Now Ozodiceromya argentata (Bellardi) (Irwin & Lyneborg, 1980: 257) (Therevidae).
138. Psilocephala univittata, p. 190 (&). “Messico, Puebla (Saussure) (Collection Bigot)”. Type
in Oxford; specimen of Turin lost. Now Ozodiceromya univittata (Bellardi) (Irwin &
Lyneborg, 1980: 258) (Therevidae).
139. Psilocephala sumichrasti, p. 191 (%). “Messico, Tuxpango Presso Orizaba (Sumichrast)”.
Box 6. Now Cyclotelus sumichrasti (Bellardi) (Irwin & Lyneborg, 1980: 254) (Therevidae).
140. Psilocephala nigra, p. 192 (%). “Messico (Semper)”. Box 6. Now Melanothereva nigra
(Bellardi) (Irwin & Lyneborg, 1980: 193) (Therevidae).
141. Atherix latipennis, p. 193 (%). “Messico, Angangueo (as Angang) (Saussure), Oaxaca
(Sallé)”. Box 6. Now Suragina latipennis (Bellardi) (James, 1968: 6) (Rhagionidae).
142. Atherix longipes, p. 194, pl. II, fig. 17 (&). “Messico, Cordova (Saussure)”. Box 6. Now
Suragina longipes (Bellardi) (James, 1968:6) (Rhagionidae).
143. Leptis cinereus, p. 195. “Messico, Cordova (Saussure). Type lost. An unrecognized
species (James, 1968: 6) (Rhagionidae).
144. Chrysopila mexicana, p. 196 (%). “Messico, contorni di Messico (Truqui)”. Box 6. Now
Chrysopilus mexicanus Bellardi (James, 1968: 3) (Rhagionidae).
145. Hybos dimidiata, p. 197 (&). “Messico (Sallé)”. Box 6. Now Lactistomyia dimidiata
(Bellardi) (Smith, 1967: 14) (Empididae).
146. Empis bicolor, p. 198 (&). “Messico, Cuautla (as Cuantla) (Saussure)”. Box 6. Same
combination (Smith, 1967: 21) (Empididae).
147. Empis cyaneus, p. 198 (&). “Messico, Angangueo (as Angang) (Saussure)”. Box 6. Now
Lamprempis cyanea (Bellardi) (Smith, 1967: 20) (Empididae).
Acta Zool. Mex. (n.s.) 84 (2001)
109
148. Empis totipennis, p. 199 (&). “Messico, Morelia (Saussure)”. Box 6. Unchanged (Smith,
1967: 24) (Empididae).
1862:
149. Culex bigoti, p. 200, pl. III, fig. 1 (&). “Messico (Sallé)(Coll. Bigot)”. Type in Oxford.
Unchanged (Culicidae).
150. Limnophila undulata, p. 200, pl. III, fig. 2 (&). “Messico, Tuxpango presso Orizaba
(Sumichrast)”. Box 7. Unchanged (Alexander & Alexander, 1970: 108) (Tipulidae).
151. Aporosa mexicana, p. 201 (%). “Messico, Tuxpango presso Orizaba (April) (Sumichrast)”.
Box 7. Now Limonia (Geranomyia) mexicana (Bellardi) (Alexander & Alexander, 1970: 64)
(Tipulidae).
152. Leia punctata, p. 202, pl. III, fig. 3 (%). “Messico, Tuxpango presso Orizaba
(Sumichrast)”. Box 7. Unchanged (Papavero, 1978: 30) (Mycetophilidae).
153. Rhyphus taeniatus, p. 202, pl. III, fig. 15 (%). “Messico, Tuxpango presso Orizaba
(Sumichrast)”. Box 7. Now Olbiogaster taeniata (Bellardi) (Papavero, 1967a: 4)
(Anisopodidae).
154. Simulium mexicanum, p. 203 (%). “Messico, Tuxpango presso Orizaba (Sumichrast)”. Box
7. Unchanged (Vulcano, 1967: 16) (Simuliidae).
155. Plecia notata, p. 204, pl. III, fig. 4 (%, &). “Messico, Playa Vicente (Sallé)”. Box 7. Now
a junior synonym of Plecia plagiata Wiedemann, 1824 (Hardy, 1966: 4) (Bibionidae).
156. Dilophus minutus, p. 204 (%, &). “Messico, Tuxpango presso Orizaba (Sumichrast)”. Box
7. Unchanged (Hardy, 1966: 10) (Bibionidae).
157. Hermetia lativentris, p. 205 (&). “Messico, Tuxpango presso Orizaba (Sumichrast)”. Box
7. Unchanged (James, 1973: 38) (Stratiomyidae).
158. Acanthina nana, p. 206 (%, &). “Messico, Tuxpango presso Orizaba (Sumichrast)”. Box
7. Now Artemita nana (Bellardi) (James, 1973:50) (Stratiomyidae).
159. Stratiomys bimaculata, p. 207, pl. III, fig. 7 (%). “Messico, Cosamaloapan (Sallé)”. Box 7.
Now Hoplitimyia bimaculata (Bellardi) (James, 1973:42) (Stratiomyidae).
160. Chordonota fuscipennis, p. 208, pl. III, fig. 6 (%). “Messico, Playa Vicente (Sallé)”. Box 7.
Unchanged (James, 1973: 24) (Stratiomyidae).
161. Chordonota carbonaria, p. 208 (%). “Messico, Tuxpango presso Orizaba (Sumichrast)”.
Type lost. Now a junior synonym of Labocerina atrata (Fabricius, 1805) (James, 1973:29)
(Stratiomyidae).
162. Clitellaria pygmaea, p. 209, pl. III, fig. 5 (%). “Messico (Sallé), Tuxpango, presso Orizaba
(Sumichrast)”. Box 7. Now Eurineura pygmaea (Bellardi) (James, 1973:32)
(Stratiomyidae).
163. Sargus versicolor, p. 210, pl. III, fig. 8 (&). “Messico, Oaxaca (Sallé)”. Box 7. Unchanged
(James, 1973:20) (Stratiomyidae).
164. Tabanus subtilis, p. 211, pl. III, fig. 9 (&). “Messico, Oaxaca (Sallé)”. Box 7. Now
Stenotabanus subtilis (Bellardi) (Fairchild, 1971: 48; Fairchild & Burger, 1994: 76)
(Tabanidae).
165. Tabanus rubescens, p. 212 (&). “Messico, Oaxaca (Sallé)”. Box 7. Now a junior synonym
of Tabanus dorsifer Walker, 1860) (Fairchild, 1971: 91; Fairchild & Burger, 1994: 136)
(Tabanidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
110
166. Acanthomera bellardii Bigot, p. 213, pl. III, fig. 11 (&). “Messico (Sallé) (Coll. Bigot)”. Type
in Oxford. Now Pantophthalmus bellardii (Bigot) (Papavero, 1967b: 2) (Pantophthalmidae).
167. Acanthomera bigoti, p. 213, pl. III, fig. 10 (%). “Messico, Chinantla presso Oaxaca
(Sallé)”. Box 7. Now a junior synonym of Rhaphiorhynchus planiventris (Wiedemann,1821)
(Papavero, 1967b: 3) (Pantophthalmidae).
168. Apelleia vittata, p. 216, pl. III, fig. 12 (%). “Messico, Playa Vicente (Sallé)”. Box 7
(Acroceridae).
169. Laphria homopoda, p. 217, pl. III, fig. 16 (%). “Messico, Tuxpango presso Orizaba (May)
(Sumichrast)”. Box 7. Now Lampria homopoda (Bellardi) (Martin & Papavero, 1970:40)
(Asilidae).
170. Erax lowei, p. 218, pl. III, fig. 17 (%). “Messico, Tuxpango presso Orizaba (Sumichrast)”.
Box 7. Now in Efferia (Martin & Papavero, 1970: 66) (Asilidae).
171. Asilus tuxpanganus, p. 219 (%, &). “Messico, Tuxpango presso Orizaba (Sumichrast)”.
Box 7. Now Philonicus tuxpanganus (Bellardi) (Martin & Papavero, 1970: 85) (Asilidae).
172. Ommatius fuscipennis, p. 220 (%, &). Messico, Tuxpango presso Orizaba (April and May)
(Sumichrast)”. Box 7. Unchanged (Martin & Papavero, 1970: 59) (Asilidae).
173. Dasypogon pseudoialapensis, p. 222 (%, &). “Messico, Tuxpango presso Orizaba
(Sumichrast)”. Box 7. Now in Diogmites (Martin & Papavero, 1970: 26) (Asilidae).
174. Leptis bitaeniata, p. 223, pl. III, fig. 14 (%). “Messico, Tuxpango presso Orizaba
(Sumichrast)”. Box 7. Now in Rhagio (James, 1968: 6) (Rhagionidae).
175. Leptis politaeniata, p. 224, pl. III, fig. 13 (&). “Messico, Tuxpango presso Orizaba
(Sumichrast)”. Box 7. Now in Rhagio (James, 1968:6) (Rhagionidae).
176. Chrysopila nigra, p. 224, pl. III, fig. 13 (&). “Messico, Tuxpango presso Orizaba (April)
(Sumichrast)”. Box 7. Now Chrysopilus niger Bellardi (James, 1968: 4) (Rhagionidae).
7.
T
HE SPECIES
D
ESCRIBED BY
C
AMILLO
R
ONDANI
Camillo Rondani (pronounced Róndani) (Fig. 6), born in Parma on November 23,
1803, was a descendant of a very old, noble family of that city, which could boast
of having produced men of distinction as eraly as the twelfth century. Among his
ancestors was a painter of some renown, Francesco Maria Rondani (1490-1548).
His family had decided that Camilo should have an ecclesiastical career. However,
in 1818, Rondani began to show a great interest in natural history, after having read
the works of Buffon, and he discontinued his religious studies. Afterwards he
entered the University of Parma, and with Giorgio Jan, with whom he learned
botany, Rondani undertook several excursions, collecting insects.
During his life in the University, Rondani became interested in the politics of his
country. Parma was at that time a Duchy governed by Marie Louise, the ex-
Empress of France. After the fall of Napoleon in Waterloo, she obtained this
government from the Austrains, who again had dominated Italy since 1815. In 1831,
following the example given by the revolution of the preceding year in France, which
resulted in the ascension to power of Louis Philippe, the Parmese rebelled against
Acta Zool. Mex. (n.s.) 84 (2001)
111
the Austrians. Marie Louise was deposed and fled to Piacenza. The liberal party
assumed the government, and one of its members, Macedonio Melloni, offered a
chair of natural history to Rondani, with the opportunity of travelling to France,
where he was to study in order to take later the professorship in his native town.
Rondani travelled to Paris, attending several lectures of the leading naturalists,
especially Cuvier. However, with the Austrian troops having suppressed the
Parmese revolution, Rondani was forced to leave Paris and return to his native
place. In Parma, he stayed for a while helping his brother in commerce. In 1833 he
married, and the following year left Parma, going to live in Guardasone, where he
dedicated himself to entomological pursuits.
Figure 6
Camillo Rondani. Photograph.
On December 17, 1847, with the death of Marie Louise, the government of the
Duchy of Parma passed to Carlo Lodivo di Borbone, who assumed it with the title
of Carlo II. The next year another revolution took place, taking him from power. In
the same year Rondani published his first paper on exotic Diptera, a study of the
Brazilian flies collected by Ghiliani (Rondani, 1848).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
112
In 1849, Carlo Alberto di Savoia declared war on Austria and took Parma with his
army. Rondani left the countryside and again came to that city, returning to the
political life. He was elected deputy by Traversetolo. However, with the defeat of
Novara, Parma was lost again to the Austrians, and Carlo III succeeded his father
in the Duchy, exercizing a tyrannical rule. Rondani took refuge again in
Guardasone, returning to his entomological studies. In this he was rewarded by the
excellent harvest brought by Cajetano Osculati from his trip through the equatorial
regions of South America, then almost entirely unknown entomologically. The 31
species of Diptera reported by Osculati (17 of which were described as new) were
studied by Rondani, who published the results in 1850 (Rondani, 1850a). In that
same year, Rondani published another paper describing Diptera, principally from
Venezuela and from the Island of São Sebastião (Sate of São Paulo), Brazil, the
latter collected by Giovanni Casaretto (Rondani, 1850b).
In 1854, March 26, Carlo III was murdered, and his consort, Luigia Maria di
Bourbon-Artois, assumed the government of Parma. The University experienced
notable progress and Rondani was invited to assume the professorship of
agronomy, also being named director of an agronomic institution. In 1860, with the
unification of Italy and the definite defeat of the Austrians, Rondani went to teach
natural history in the Liceo di Parma, as the agronomic institution had been closed.
When it was reopened in 1865, Rondani was again appointed to it as director. In
1863, Rondani published one more paper on exotic Diptera, including mostly
specimens sent to him by R. A. Philippi, from Chile. Also included were some other
specimens caught by Ghiliani in Belém, Pará, Brazil, and some materials given to
him by Marquis Massimiliano Spinola. This paper (1863) included specimens from
Colombia and Puerto Rico, collected by Giuseppe Bertero.
In 1868, Rondani described Argentinian flies collected by Pellegrino Strobel.
In 1878, he published his last paper on Neotropical flies, describing some
Pupipara found in the Museo Civico di Genova. This is his only paper describing
Mexican flies, which includes the following:
1. Lipoptena mazamae, p. 153. “Central and South America”. Unchanged (Guimarães, 1968a:
9) (Hippoboscidae).
2. Ornithomyia bellardiana, p. 157. “Messico”. Now a junior synonym of Ornithoctona
(Ornithoctona) erythrocephala (Leach, 1817) (Guimarães, 1968a: 3) (Hippoboscidae).
3. Olfersia pallidilabris, p. 161. “Messico”. Now a junior synonym of Lynchia nigra (Perty, 1833)
(Guimarães, 1968a:6) (Hippoboscidae).
4. Olfersia obliquinervis, p. 162. “Messico”. Also a synonym of Lynchia nigra (Perty, 1833)
(Guimarães, 1968a: 6) (Hippoboscidae).
5. Strebla mexicana, p. 168. Not cited in Wenzel´s 1970 catalogue (Streblidae).
Acta Zool. Mex. (n.s.) 84 (2001)
113
In 1870, Rondani had collaborated in the foundation of the Italian Entomological
Society, of which he was the vice-president. He died on September 17, 1879. A list
of his publications was given by Baron Osten Sacken (1885). His biography was
published by Lessona (1884) (see also Bezzi, 1908a, 1908b). Rondani´s types of
exotic (i.e., Latin American) Diptera are deposited in the Museo de Instituto di
Zoologia Sistematica della Università di Torino, in the Museo Civico di Storia
Naturale di Milano, and in Naples (a list of types found in Naples, the majority of
Rondani´s types, was given by A. Costa (1866)).
8.
D
OMINIK
B
ILIMEK
Dominik Bilimek entitled himself “Kustos am National Museum in Mexiko”
(Curador del Museo Nacional de México)”. During the unfortunate reign of Emperor
Ferdinand Maximilian of Mexico, Bilimek explored the Cacahuamilpa Caves, 45
leagues from Mexico City, on January 14, 1866, leaving an account of his
explorations, and a description of Pholeomyia leucozona (1867) (Milichiidae), the
only Mexican fly described by him. According to labels in the Museum of Vienna he
also collected extensively in Orizaba and Tacubaya. Before his arrive to Mexico he
professed as Augustine monk, but being his principal fondness to collect insect
specimens for the European museums. As soon as he deserted the monastic order,
he organized a museum of natural history for Maximilian, using an abandoned
abbey of La Chroma island, in the Adriatic Sea. Later in Mexico, he collected a
great amount of insects around Mexico City and in Morelos, sometimes
accompanied by Empress Charlotte and her ladies (Hoffmann et al., 1986). No
other details are known about his life and travels (Papavero, 1973: 291-292. See
also Barnhart, 1927, 1965; Hemsley, 1887; Linden, 1867).
9.
T
HE SPECIES DESCRIBED BY
F.
J
AENNICKE
Unfortunately, we were not able to find data about the life of F. Jaennicke. In his
paper of 1867 (reprinted in 1968) he described the following species from Mexico
(most of the types are now in the Senckenberg Institution, Frankfurt a. M.):
1. Odontomyia prasina, p. 324 (16) (%). “Mexico”. Now Hedriodiscus prasinus (Jaennicke)
(James, 1973: 41) (Stratiomyidae).
2. Rondania obscura, p. 325 (17) (%, &). “Mexico”. Now Cyphomyia obscura (Jaennicke)
(James, 1973: 27) (Stratiomyidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
114
3. Anthrax castanea, p. 338 (30), pl. 44, fig. 15 (%, &). “Mexico”. Now Villa (Paravilla) castanea
(Jaennicke) (Painter et al., 1978: 50) (Bombyliidae).
4. Anthrax paradoxa, p. 339 (31), pl. 44, fig. 16 (&). “Mexico”. Now Neodiplocampta
(Neodiplocampta) paradoxa (Jaennicke) (Painter et al., 1978: 37) (Bombyliidae).
5. Antharx kaupii, p. 340 (32), pl. 44, fig. 17 (%, &). “Mexico”. Now a junior synonym of Ligyra
orcus (Walker, 1849) (Painter et al., 1978:43) (Bombyliidae).
6. Exoprosopa anthracoidea, p. 340 (32), pl. 44, fig. 18 (%, &). “Mexico”. Unchanged (Painter
et al., 1978: 39) (Bombyliidae).
7. Exoprosopa rostrifera, p. 341 (33), pl. 44, fig. 19 (%, &). “Mexico”. Unchanged (Painter et al.,
1978: 41) (Bombyliidae).
8. Exoprosopa blanchardiana, p. 341 (33), pl. 44, fig. 20 (%, &). “Mexico”. Now Villa
(Hemipenthes) sinuosa blanchardiana (Jaennicke) (Painter et al., 1978: 50) (Bombyliidae).
9. Exoprosopa pueblensis, p. 342 (34), pl. 44, fig. 21 (%, &). “Mexico”. Unchanged (Painter et
al., 1978: 41) (Bombyliidae).
10. Adelidea flava, p. 346 (38) (%). “Mexico”. Now a junior synonym of Lordotus gibbus Loew,
1863 (Painter et al., 1978: 6) (Bombyliidae).
11. Poecilognathus thlipsomyzoides, p. 351 (43), pl. 43, fig. 11 (%). “Mexico”. Now Phthiria
thlipsomyzoides (Jaennicke) (Painter et al., 1978: 15) (Bombyliidae).
12. Nicocles analis, p. 355 (47), pl. 43, fig. 13 (%). “Mexico”. Unchanged (Martin & Papavero,
1970: 31) (Asilidae).
13. Atomosia beckeri, p. 359 (51) (%, &). “Mexico”. Unchanged (Martin & Papavero, 1970: 46)
(Asilidae).
14. Micropalpus albomaculatus, p. 388 (80) (&). “Mexico”. Now Epalpus albomaculatus
(Jaennicke) (Guimarães, 1971: 64) (Tachinidae).
15. Jurinia flavifrons, p. 390 (82) (&). “Mexico”. Now in Archytas (Guimarães, 1971: 49)
(Tachinidae).
16. Jurinia apicalis, p. 390 (82) (%, &). “Mexico”. Now a junior synonym of Archytas lateralis
(Macquart, 1843) (Guimarães, 1971: 50) (Tachinidae).
17. Dejeania rutiloides, p. 394 (86) (&). “Mexico”. Now in Paradejeania (Guimarães, 1971: 59)
(Tachinidae).
18. Volucella maximiliani, p. 395 (87) (%, &). “Mexico”. Now a junior synonym of Copestylum
metalliferum (Walker, 1849) (Thompson et al., 1976: 79) (Syrphidae).
19. Volucella mellea, p. 396 (88) (%, &). “Mexico”. Now Copestylum melleum (Thompson et al.,
1976: 79) (Syrphidae).
20. Volucella haagii, p. 397 (89) (&). “Mexico”. Now in Copestylum (Thompson et al., 1976: 76)
(Syrphidae).
21. Eristalis thoracica, p. 399 (91) (%, &). “Mexico”. Now a junior synonym of Palpada ruficeps
(Macquart, 1842) (Thompson et al., 1976: 108) (Syrphidae).
22. Eristalis tricolor, p. 400 (92) (&). “Mexico”. Now a junior synonym of Palpada pusilla
(Macquart, 1842) (Thompson et al., 1976: 108) (Syrphidae).
23. Eristalis bellardii, p. 400 (92) (%). Now Eristalis (Eoseristalis) bellardii Jaennicke (Thompson
et al., 1976: 101) (Syrphidae).
24. Zodion splendens, p. 405 (97). Not cited in Papavero´s 1971a catalogue (Conopidae).
Acta Zool. Mex. (n.s.) 84 (2001)
115
10.
T
HE SPECIES DESCRIBED BY
C
ARL
E
DUARD
A
DOLPH
G
ERSTAECKER
Carl Eduard Adolph Gerstaecker died on July 20, 1895 at Greifswald, at the age
of 67. He was educated for the medical profession and took his degree, but devoted
himself to zoology, especially to entomology. For many years he was keeper of the
entomological department of the Berlin Natural History Museum and also a
professor of zoology at the University of Berlin. About the year 1876, differences
with the then director of the Berlin Museum induced him to resign his appointment
in Berlin, and he subsequently accepted the professorship of Zoology at Greifswald,
which he held until his death. Gerstaecker was an industrious and thorough worker
in all departments of entomology. Among his principal works may be noted the
“Arthropoda” in the “Handbuch der Zoologie” (1863) and the same phylum in
Bronn´s “Klassen und Ordnungen der Tierreichs”.
Gerstaecker rendered infinite help to his fellow students by compiling the
entomological portions of the German “Bericht” (the forerunner and subsequent
contemporary of the English “Zoological Record”) from the year 1853 to the year
1867.
Among Gerstaecker´s publications are those dealing with the Acroceridae (1856),
Stratiomyidae (1857), Acalyptratae (1860), parasitic flies (1868a), and his most
important paper, the revision of the Mydidae (1868b).
Gerstaecker studied the Diptera contained in the ancient collection of Tönder
Lund in Copenhagen, and that of Westermann in the Vienna Museum. Many of his
more recent specimens were collected by Natterer in Brazil (cf. Papavero, 1971b:
80-87), Lacordaire (ibid., p. 133) in French Guiana, Deppe (Ibid., pp. 103-107; part
1 of the present work) in Mexico, Olfers (cf. Papavero, 1971b: 69) in Brazil,
Burmeister (Ibid., pp. 292-293) in Brazil, Koeppen in Mexico, Moritz (Ibid., pp. 297-
298) in Colombia and the West Indies, Virmond (Ibid., pp. 110, 112) in Brazil, Appun
(Ibid., p. 291) in Colombia, Ehrenberg and Uhde in Mexico. He also studied the
collections organized by Bescke in Brazil (cf. Papavero, 1971b: 87) belonging to the
coleopterist Germar.
*****
About Koeppen and Uhde (or Uhle) nothing is known.
Carl August Ehrenberg was born in Delitsch, Province of Sachsen, Prussia, on
August 24, 1801, son of the “Stadtrat und Stadtritter” Johann Gottfried Ehrenberg.
Carl dedicated himself to commerce (Kauffmannstande) and travelled through St.
Thomas (1827-1828), Port-au-Prince in Haiti (March 1828-June 1831) and Mexico
(July 1831- April 1840), both near the Capital and 30 leagues to the interior (Mineral
Papavero & Ibañez-Bernal: History of Mexican Dipterology
116
Real del Monte). From all those places he sent to the Museums of Berlin and Halle
rich zoological and botanical collections. In Mexico he became especially interested
in the Cactaceae. Upon his return to Berlin he entered business with his brother
Ferdinand. He died there on August 13, 1849, of cholera (Urban, 1903, p. 43; see
also Barnhart, 1965; Hemsley, 1887; Knobloch, 1893; Lorence & García, 1989;
McVaugh, 1972; Papavero, 1973; Pennell, 1945; Stafleu & Cowan, 1976 (1: 727);
Urban, 1897).
Only 8 species were described by Gerstaecker from Mexico:
1857:
1. Cyphomyia varipes, p. 283 (&). “Mexico, Costa Rica, ?Venezuela”. Types in Vienna
Museum. Unchanged (James, 1973: 28) (Stratiomyidae).
2. Cyphomyia pilosissima, p. 293 (%). “Mexico (Koeppen)”. Type in Vienna Museum.
Unchanged (James, 1973: 28) (Stratiomyidae).
3. Cyphomyia tomentosa, p. 294 (&). “Mexico (Koeppen)”. Type ?. Unchanged (James, 1973:
46) (Stratiomyidae).
4. Stratiomys fenestrata, p. 327 (&). “Mexico (Deppe)”. Type ?Berlin. Unchanged (James,
1973: 46) (Stratiomyidae).
5. Stratiomys euchlorus, p. 328 (%, &). “Mexico (Deppe)”. Types ?Berlin. Now Hedriodiscus
euchlorus (Gerstaecker) (James, 1973: 40) (Stratiomyidae).
1860:
6. Diacrita costalis, p. 197, pl. II, figs. 10, 10a. “Mexico, Oaxaca (Deppe)”. Type ?Berlin.
Unchanged (Steyskal, 1968b: 4) (Otitidae).
1868b:
7. Mydas lavatus, p. 96 (%, &). “Mexico (Koeppen)”. Types Berlin. Now a junior synonym of
Mydas xanthopterus Loew, 1866 (Papavero, 1996: 631) (Mydidae).
8. Mydas annularis, p. 100 (&). “Mexico (Ehrenberg)”. Type Berlin. Unchanged (Papavero,
1996: 630) (Mydidae).
11.
T
HE SPECIES DESCRIBED BY
J
ACQUES
M
ARIE
F
RANGILLE
B
IGOT
Unfortunately, very little is known about the life of the French dipterist Jacques
Marie Frangille Bigot (Fig. 7). He was born in 1818, in 1844 was elected fellow of
the “Société Entomologique de France”, and seems to have been a very wealthy
person, who could buy large collections of exotic flies, having amassed a very
important collection of Diptera.
Acta Zool. Mex. (n.s.) 84 (2001)
117
Figure 7
Jacques Marie Frangille Bigot. Photograph property of N. Papavero.
Bigot published two large series of papers, the “Essai d´une classification générale
et synoptique de l´Ordre des Insectes Diptères”, and the “Diptères nouveaux ou peu
connus”.
In relation to the former series, Baron Osten Sacken makes the following
comments (1904: 232-233; 1978 (reprint): 232-233):
Papavero & Ibañez-Bernal: History of Mexican Dipterology
118
It remains for me now to reproduce the opinion expressed by Mr. Bigot himself, at
different times, about his own publications:
In the Annales etc. 1885, p. 225, he said: “J´avais autrefois (Ann. etc. 1852-1859)
commencé la publication d´un travail intitulé: Essai d´une classification générale et
synoptiqe de l´Ordre des Insectes Diptères etc. Cette oeuvre fut alors sévèrement
et doctoralement critiquée dans le Bericht de Gerstaecker (Berlin) par l´habile
diptèriste H. Loew. Je dois confesser que la plupart de ses objections m´ont paru
tellement fondées que, reconnaissant la médiocre valeur de mes Essais, je me
résigne à les abandonner à mi-chemin”. Whereupon Mr. Bigot proposes a new plan
for the distribution of the Diptera, into two tribes: Omalocerati and Anomalocerati etc.
A further discussion between Bigot and G. H. Verrall, about questions of orthography
and of mutual courtesy will be found in the Wien. Ent. Zeitschr., 1889, p. 168, 265 and
293. In the latter notice, Bigot renews his former recantation of his “Essai”: “Si Mr.
Verrall avait daigné lire mes opuscules, etc. il y aurait vu, maintes fois répété, que
mon ancien travail, intitulé “Essai” etc. devait être désormais, et dans son entier,
considéré comme nul et non avenu, déclaration qui m´absolut naturellement des
fautes que j´ai pu y commettre”.
Regarding the series “Diptères nouveaux ou peu connus”, Aldrich (1905: ii) points
out that:
Bigot´s main work (…) is in the form of a long series of articles in the Annales
Soc. ent. France (…) interspersed chronologically with fragments in the Bulletin of
the same society. Owing to Bigot´s peculiar, complicated and inconsistent system
of numbering his contributions, it has been almost impossible for me to find them
all, but I believe I have succeeded (…). Sometimes his Roman numerals stop and
remain the same for several papers, then again the numbering in “parts” becomes
stationary while the other goes on, and in two cases two papers have both the
same.
Bigot started publishing in 1852 and the last of his papers that interest us is dated
from 1892. He also studied the Diptera collected by Ramón de la Sagra in Cuba
(1857a), and those collected by the “Mission Scientifique du Cap Horn” (1888a).
Osten Sacken (1904: 232; 1978: 232) comments further, on Bigot´s work:
My excellent friend J. M. F. Bigot died April 14 1893 at his country-seat near Paris
at the age of 74 years (An Obituary Notice will be found in the Bolletin de la Soc. Ent.
de France, April 26 1893). My acquaintance with Bigot was of long date. When I met
Acta Zool. Mex. (n.s.) 84 (2001)
119
him again after my return from the United States in 1887, and our relations were
renewed, I felt it my duty, although treating him as a friend, to tell him the truth about
his publications. I did so, during one of my visits to Paris (the year I do not
remember), in telling him that he was doing a useful work in forming a large
collection, especially of exotic Diptera; but that he should renounce descriptive work,
for which he was not competent. I expressed my opinion in the strongest terms,
concluding with the words: “If all your publications could be suppressed, it would be
a gain for science”. He winced a little, but resumed immediately his serene
expression, and said: “Eh bien, cela m´amuse”. My frankness did not prevent us from
remaining friends up to his death. In the most generous manner, he kept his rich
collection and library at my disposal. Even when he was away from Paris (he usually
spent his winters in Algiers) he left orders to his housekeeper to admit me at any
time into his sanctum, and to light a fire for me in winter. I keep of my friend Bigot
and his family (wife and daughter) a pleasant and grateful recollection.
The great collection of flies gathered by Bigot, including many types, especially
of exotic flies, and many types described by Macquart (see part 2 of this work),
were bought after his death by G. H. Verrall, and are now deposited at the Hope
Department of Entomology, Oxford University (most of the types), some of them
also at the British Museum of Natural History. Osten Sacken (l. c.) had an important
role in that transaction:
The share I had in the sale of Bigot´s collection is explained in the Bulletin de la
Soc. Ent. de France, June 14, 1893. In this notice it should have been added that I
had written to the Authorities of the Museum in Paris, urging them to purchase this
important collection offered at a very moderate price (6000 francs). My advice,
however, was not heeded, and the collection passed into the hands of Mr. G. H.
Verrall of Newmarket, England, and will ultimately, as I understand, find its place in
the British Museum.
*****
Although he never cites the names of the collectors, it is very probable that Bigot
studied the collections gathered in Mexico by Pilate, Geisbreght, Linden, and Funck
(see part 2 of this work), as well as the material collected by Sallé, Sumichrast,
Saussure, and Boucard (see part 6 of this work).
Bigot described the following Mexican flies:
Papavero & Ibañez-Bernal: History of Mexican Dipterology
120
1875a:
1. Emphysomera pilosula, p. 243 (%). “Mexico”. Type in Oxford. Now Ommatius pilosulus
(Bigot) (Martin & Papavero, 1970: 59) (Asilidae).
2. Emphysomera bicolor, p. 244 (%, &). “Mexico”. Type in Oxford. Now in Ommatius (Martin &
Papavero, 1970: 59) (Asilidae).
3. Ommatius parvus, p. 247 (%). “Mexico”. Type in Oxford. Unchanged (Martin & Papavero,
1970: 59) (Asilidae).
1875b:
4. Volucella castanea, p. 476 (%). “Mexico, Oaxaca”. Type in Oxford. Now a junior synonym
of Copestylum posticum (Say, 1830) (Thompson et al., 1976: 82) (Syrphidae).
5. Volucella tricincta, p. 477 (&). “Mexico, Oaxaca”. Type in Oxford. Now Copestylum tricinctum
(Bigot) (Thompson et al., 1976: 85) (Syrphidae).
6. Volucella purpurifera, p. 477 (%, &). “Mexico, Oaxaca”. Type in Oxford. Now Copestylum
purpuriferum (Bigot) (Thompson et al., 1976: 82) (Syrphidae).
7. Volucella variegata, p. 478 (%, &). “Mexico”. Types in Oxford. Now Copestylum variegatum
(Bigot) (Thompson et al., 1976: 86) (Syrphidae).
8. Volucella amethistina, p. 479 (%). “Mexico”. Type in Oxford. Now Copestylum amethystinum
(Bigot) (Thompson et al., 1976: 71) (Syrphidae).
9. Volucella nigrifacies, p. 479 (%). “Mexico”. Type in BMNH. Now in Copestylum (Thompson
et al., 1976: 80) (Syrphidae).
10. Volucella pulchripes, p. 480 (%). “Mexico”. Type in Oxford. Now in Copestylum (Thompson
et al., 1976: 82) (Syrphidae).
11. Volucella varians, p. 481 (&). “Mexico”. Type in Oxford. Now in Copestylum (Thompson et
al., 1976: 86) (Syrphidae).
12. Volucella viridula, p. 481 (&). “Mexico”. Type in Oxford. Now a synonym of Copestylum
vagum (Wiedemann, 1830) (Thompson et al., 1976: 86) (Syrphidae).
13. Volucella tristis, p. 482 (&). “Mexico”. Type in Oxford. Now a junior synonym of Copestylum
subrostratum (Rondani, 1848) (Thompson et al., 1976: 85) (Syrphidae).
1875c:
14. Cyphomyia scalaris, p. 487 (%, &). “Mexico”. Unchanged (James, 1973: 27)
(Stratiomyidae).
1876a:
15. Trichopoda nigricauda, p. 395 (%). “Mexico”. Now Trichopoda (Trichopoda) nigricauda Bigot
(Guimarães, 1971: 9) (Tachinidae).
1877b:
16. Somomya pallidibasis, p. 247 (&). “Mexico”. Now Phaenicia (Phaenicia) purpurescens
(Walker, 1837) (James, 1970: 11) (Calliphoridae).
17. Somomya mutabilis, p. 248 (%, &). “Mexico”. Now Phaenicia (Phaenicia) eximia
(Wiedemann, 1819) (James, 1970: 10) (Calliphoridae).
18. Somomya flavigena, p. 249 (&). “Mexico”. Now Cochliomyia macellaria (Fabricius, 1775)
(James, 1970: 7) (Calliphoridae).
Acta Zool. Mex. (n.s.) 84 (2001)
121
19. Somomya callipes, p. 249 (%). “Mexico”. Now Myiolucilia lyrcea (Walker, 1849) (James,
1970: 8) (Calliphoridae).
20. Somomya pueblensis, p. 250 (&). “Mexico”. Now Phaenicia (Phaenicia) eximia
(Wiedemann, 1819) (James, 1970: 10) (Calliphoridae).
21. Somomya fulvinota, p. 251 (&). “Mexico”. Now in Myiolucilia (James, 1970: 7)
(Calliphoridae).
22. Somomya argentifera, p. 251 (?%). Not cited in James´s 1970 catalogue (Calliphoridae).
23. Somomya aztequina, p. 252 (%, &). “Mexico”. Now Cochliomyia macellaria (Fabricius, 1775)
(James, 1970: 7) (Calliphoridae).
1877c:
24. Carlottaemyia moerens, p. xxvii (%, &). “Mexico”. Now Diacrita costalis Gerstaecker, 1860)
(Steyskal, 1968b: 4) (Otitidae).
1878a:
25. Pyrellia scapulata, p. 35 (%, &). “Mexico”. Now Morellia basalis (Walker, 1852) (Pont, 1972:
8) (Muscidae).
26. Pyrellia iris, p. 36 (&). “Mexico”. Now Morellia flavicornis (Macquart, 1848) (Pont, 1972: 9)
(Muscidae).
1878b:
27. Ocyptera soror, p. 46 (&). “Mexico”. An unrecognized Cylindromyia (Guimarães, 1971: 16)
(Tachinidae).
28. Ocyptera simplex, p. 47 (&). “Mexico”. An unrecognized Cylindromyia (Guimarães, 1971:
16) (Tachinidae).
1878d:
29. Pogonosoma arachnoides, p. 227 (&). “Mexico”. Type in Oxford. Now in Andrenosoma
(Pogonosoma) (Martin & Papavero, 1970: 44) (Asilidae).
30. Cormansis eupoda, p. 234 (?&). “Mexico”. Type in Oxford. Now in Atomosia (Martin &
Papavero, 1970: 47) (Asilidae).
31. Aphestia nigra, p. 235 (%, &). “Mexico”. Type in Oxford. Unchanged (Martin & Papavero,
1970: 50) (Asilidae).
32. Atomosia soror, p. 236 (%, &). “Mexico”. Type in Oxford. Now Atomosia rufipes Macquart,
1847 (Martin & Papavero, 1970: 47) (Asilidae).
33. Cerotainia? nigra, p. 238 (&). “Mexico”. Type in Oxford. Unchanged (Martin & Papavero,
1970: 52) (Asilidae).
34. Cerotainia? dubia, p. 238 (%). “Mexico”. Type in Oxford. Unchanged (Martin & Papavero,
1970: 52) (Asilidae).
35. Microstylum fulvigaster, p. 410 (%). “Mexico”. Type in Oxford. Unchanged (Martin &
Papavero, 1970: 19) (Asilidae).
36. Stenopogon fuscolimbatum, p. 421 (?%, &). “Mexico”. Type in Oxford. Now Stenopogon
(Stenopogon) fuscolimbatus Bigot. Unchanged (Martin & Papavero, 1970: 12) (Asilidae).
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122
37. Ceraturgus geniculatus, p. 443 (%). “Mexico (Veracruz)”. Type in Oxford. Now in Taracticus
(Martin & Papavero, 1970: 35) (Asilidae).
1879a:
38. Macroceromys fulviventris, p. 187 (&). “Mexico”. Now Solva (Macroceromys) fulviventris
(Bigot) (James, 1975: 2) (Xylomyidae).
39. Oplacantha limbata, p. 196 (&). “Mexico, Veracruz, Orizaba”. Unchanged (but Oplachantha
Rondani) (James, 1973: 8) (Stratiomyidae).
40. Hermetia nigrifascies, p. 200 (%, &). “Mexico”. Now a junior synonym of Hermetia illucens
(Linnaeus, 1758) (James, 1973: 38) (Stratiomyidae).
41. Hermetia flavoscutata, p. 201 (%). “Mexico”. Unchanged (James, 1973: 38)
(Stratiomyidae).
42. Histiodroma flaveola, p. 205 (&). “Mexico”. Now Acrochaeta fasciata Wiedemann, 1830
(James, 1973: 9) (Stratiomyidae).
43. Sargus splendens, p. 224 (&). “Mexico”. Now Sargus speciosus Macquart, 1846 (James,
1973: 20) (Stratiomyidae).
44. Macrosargus smaragdiferus, p. 226 (&). “Mexico”. Now in Merosargus (James, 1973: 15)
(Stratiomyidae).
45. Chrysonotus flavopilosus, p. 227 (%, &). “Mexico”. Now Himantoloba flavopilosa (Bigot)
(James, 1973: 10) (Stratiomyidae).
46. Merosargus fraternus, p. 228 (&). “Mexico”. Unchanged (James, 1973: 13) (Stratiomyidae).
47. Merosargus calceolatus, p. 229 (&). “Mexico”. Unchanged (James, 1973: 12)
(Stratiomyidae).
48. Ptecticus flaviceps, p. 230 (&). “Mexico”. Unchanged (James, 1973: 17) (Stratiomyidae).
1880b:
49. Eristalomyia rufoscutata, p. 221 (%, &). “Mexico”. Types in Oxford. Now Eristalis
(Eoseristalis) bellardii Jaennicke, 1867 (Thompson et al., 1976: 101) (Syrphidae).
50. Eristalis sackenis, p. 224 (%). “Mexico”. Type in BMNH. Now Palpada testaceicornis
(Macquart, 1850) (Thompson et al., 1976: 110) (Syrphidae).
51. Eristalomyia pachypoda, p. 224 (&). “Mexico”. Type BMNH. Now Palpada obsoleta
(Wiedemann, 1830) (Thompson et al., 1976: 107) (Syrphidae).
52. Eristalomyia fulvipes, p. 225 (&). “Mexico”. Type in BMNH. Now Palpada fulvipes (Bigot)
(Thompson et al., 1976: 105) (Syrphidae).
1882c:
53. Eurhinomallota metallica, p. lxvii (&). “Mexico”. Type in Oxford. Now in Criorhina
(Thompson et al., 1976: 119) (Syrphidae).
1882e:
54. Asemosyrphus oculiferus, p. cxxviii (&). “Mexico”. Type in BMNH. Now Asemosyrphus
mexicanus (Macquart, 1842) (Thompson et al., 1976: 97) (Syrphidae).
Acta Zool. Mex. (n.s.) 84 (2001)
123
55. Asemosyrphus nigroscutatus, p. cxxviii (?&). “Mexico”. Type BMNH. Now Asemosyrphus
mexicanus (Macquart, 1842) (Thompson et al., 1976: 97) (Syrphidae).
56. Asemosyrphus flavocaudatus, p. cxxviii (?&). “Mexico”. Type BMNH. Now Asemosyrphus
mexicanus (Macquart, 1842) (Thompson et al., 1976: 97) (Syrphidae).
57. Asemosyrphus bicolor, p. cxxix (?&). “Mexico”. Type BMNH. Now Asemosyrphus arquatus
(Say, 1829) (Thompson et al., 1976: 96) (Syrphidae).
58. Romaleosyrphus villosus, p. cxxix (%). “Mexico”. Type BMNH. Now Criorhina villosa (Bigot)
(Thompson et al., 1976: 119) (Syrphidae).
1883a:
59. Volucella tau, p. 84 (&). “Mexico”. Type BMNH. Now in Copestylum (Thompson et al.,
1976: 85) (Syrphidae).
60. Phalachromyia vicina, p. 86 (%). “Mexico”. Type in BMNH. Now Copestylum vicinum (Bigot)
(Thompson et al., 1976: 86) (Syrphidae).
61. Phalachromyia melanorhina, p. 86 (&). “Mexico”. Type in BMNH. Now Copestylum
subrostratum (Rondani, 1848) (Thompson et al., 1976: 85) (Syrphidae).
1883c:
62. Sphyximorpha nigra, p. 317 (%, &). “Mexico”. Types in BMNH. Now in Polybiomyia
(Thompson et al., 1976: 95) (Syrphidae).
63. Sphyximorpha rufibasis, p. 318 (%). “Mexico”. Type in BMNH. Now in Polybiomyia
(Thompson et al., 1976: 95) (Syrphidae).
64. Microdon gracile, p. 320 (%). “Mexico”. Type in BMNH. Now Microdon (Microdon) gracilis
Bigot (Thompson et al., 1976: 65) (Syrphidae).
65. Ocyptamus infuscatus, p. 324 (%). “Mexico”. Type in BMNH. Unchanged (Thompson et al.,
1976: 20) (Syrphidae).
66. Ocyptamus fraternus, p. 324 (&). “Mexico”. Type ?BMNH. Unchanged (Thompson et al.,
1976: 18) (Syrphidae).
67. Salpingogaster anchoratus, p. 328 (%). “Mexico”. Type in ?BMNH. Now Salpingogaster
(Salpingogaster) nigra Schiner, 1868 (Thompson et al., 1976: 32) (Syrphidae).
68. Salpingogaster cothurnatus, p. 329 (&). “Mexico”. Type in ?BMNH. Now Salpingogaster
(Salpingogaster) cothurnata Bigot (Thompson et al., 1976: 31) (Syrphidae).
69. Baccha marmorata, p. 333 (%). “Mexico”. Type in BMNH. Now in Leucopodella (Thompson
et al., 1976: 47) (Syrphidae).
70. Baccha luctuosa, p. 334 (%). “Mexico”. Type in BMNH. Now Ocyptamus luctuosus (Bigot)
(Thompson et al., 1976: 21) (Syrphidae).
71. Lepidomyia cinta, p. 345 (%). “Mexico”. Type in BMNH. Now in Quichuana (Thompson et
al., 1976: 98) (Syrphidae).
72. Spilomyia pallipes, p. 352 (&). “Mexico”. Type in BMNH. Now in Blera (Thompson et al.,
1976: 119) (Syrphidae).
1883d:
73. Syritta mexicana, p. 539 (%). “Mexico”. Type in BMNH. Now a junior synonym of Neplas
vagans (Wiedemann, 1830) (Thompson et al., 1976: 115) (Syrphidae).
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124
74. Glaurotricha volucelloides, p. 548 (%). “Mexico”. Type in BMNH. Now in Copestylum
(Thompson et al., 1976: 87) (Syrphidae).
75. Cartosyrphus frontosus, p. 553 (%). “Mexico”. Type in BMNH. Now Cheilosia (Cartosyrphus)
frontosa (Bigot) (Thompson et al., 1976: 69) (Syrphidae).
1884a:
76. ?Melanostoma quadrinotata, p. 77 (%). “Mexico”. Type in Oxford. Now Xanthandrus
quadrinotatus (Bigot) (Thompson et al., 1976: 45) (Syrphidae).
77. ?Melanostoma cruciata, p. 81 (%). “Mexico”. Type in Oxford. Now Melanostoma cruciatum
Bigot (Thompson et al., 1976: 42) (Syrphidae).
78. ?Melanostoma annulifera, p. 84 (&). “Mexico”. Type in Oxford. Now Toxomerus annulifer
(Bigot) (Thompson et al., 1976: 48) (Syrphidae).
79. Syrphus eupeltatus, p. 91 (%). “Mexico”. Type in Oxford. Now Allograpta eupeltata (Bigot)
(Thompson et al., 1976: 34) (Syrphidae).
80. Syrphus agilis, p. 92 (&). “Mexico”. Type in Oxford. Now in Ocyptamus (Thompson et al.,
1976: 12) (Syrphidae).
81. Sphaerophoria rostrata, p. 102 (&). “Mexico”. Type in Oxford. Now in Allograpta
(Thompson et al., 1976: 37) (Syrphidae).
82. Sphaerophoria picticauda, p. 102 (%, &). “Mexico”. Type in Oxford. Now a junior synonym
of Allograpta micrura (Osten Sacken, 1877) (Thompson et al., 1976: 36) (Syrphidae).
83. Sphaerophoria nasuta, p. 103 (%). “Mexico”. Type in Oxford. Now in Allograpta (Thompson
et al., 1976: 36) (Syrphidae).
84. Sphaerophoria pachypyga, p. 104 (%). “Mexico”. Type in Oxford. Now a junior synonym of
Sphaerophoria sulphuripes (Thomson, 1869) (Thompson et al., 1976: 38) (Syrphidae).
85. Sphaerophoria fulcicauda, p. 104 (%). “Mexico”. Type in Oxford. Now a junior synonym of
Sphaerophoria contigua (Macquart, 1847) (Thompson et al., 1976: 38) (Syrphidae).
86. ?Mesograpta saphiridiceps, p. 105 (&). “Mexico”. Type in Oxford. Now in Toxomerus
(Thompson et al., 1976: 54) (Syrphidae).
87. ?Mesograpta mu, p. 105 (&). “Mexico”. Type in Oxford. Now in Allograpta (Thompson et
al., 1976: 36) (Syrphidae).
88. ?Mesograpta pallipes, p. 106 (%, &). “Mexico”. Type in Oxford. Now a junior synonym of
Toxomerus tridentatus (Rondani, 1868) (Thompson et al., 1976: 55) (Syrphidae).
89. ?Mesograpta circumdata, p. 107 (%, &). “Mexico”. Type in Oxford. Now Toxomerus
circumdatus (Bigot) (Thompson et al., 1976:49) (Syrphidae).
90. ?Mesograpta lacrymosa, p. 108 (%, &). “Brésil et Mexique”. Type in Oxford. Now
Toxomerus lacrymosus (Bigot) (Thompson et al., 1976: 51) (Syrphidae).
91. ?Mesograpta heraldica, p. 109 (%, &). “Mexico”. Type in Oxford. Now Toxomerus heraldicus
(Bigot) (Thompson et al., 1976: 50) (Syrphidae).
92. ?Mesograpta trilobata, p. 109 (%). “Mexico”. Type in Oxford. Now Toxomerus trilobatus
(Bigot) (Thompson et al., 1976: 55) (Syrphidae).
93. ?Mesograpta bistriga, p. 110 (&). “Mexico”. Type in Oxford. Now in Toxomerus (Thompson
et al., 1976: 48) (Syrphidae).
Acta Zool. Mex. (n.s.) 84 (2001)
125
94. ?Mesograpta maculata, p. 111 (&). “Mexique, Cuba, Brésil”. Types in Oxford. Now
Toxomerus maculatus (Bigot) (Thompson et al., 1976: 52) (Syrphidae).
95. ?Mesograpta bicincta, p. 112 (%). “Mexico”. Type in Oxford. Now in Allograpta (Thompson
et al., 1976: 34) (Syrphidae).
1885a:
96. Limnophora limbata, p. 271 (&). “Mexique, Chili”. Unchanged (Pont, 1972: 28) (Muscidae).
97. Limnophora normata, p. 272 (%). “Mexico”. Unchanged (Pont, 1972: 29) (Muscidae).
98. Limnophora rufipes, p. 273 (&). “Mexico”. Now a junior synonym of Bithoracochaeta
leucoprocta (Wiedemann, 1830) (Pont, 1972: 37) (Muscidae).
99. Limnophora anthrax, p. 274 (%). “Mexico”. Now in Spilogona (Pont, 1972: 33) (Muscidae).
100. ?Hydrophoria pictipes, p. 275 (&). “Mexico”. Now in Taeniomyia (Pont, 1974: 9)
(Anthomyiidae).
101. ?Hydrophoria calopus, p. 275 (&). “Mexico”. Now a junior synonym of Bithoracochaeta
leucoprocta (Wiedemann, 1930) (Pont, 1972: 37) (Muscidae).
102. Homalomyia mexicana, p. 284 (%). “Mexico”. Now a junior synonym of Fannia canicularis
(Linnaeus, 1761) (Pont, 1972: 4) (Muscidae).
103. Dialyta bicolor, p. 302 (%). “Mexico”. Now in Cordiluroides (Pont, 1972: 44) (Muscidae).
1885b:
104. Nycteribia mexicana, p. 245 (?sex). “?Mexico”. Now Basilia bellardii (Rondani, 1878)
(Guimarães, 1968b: 2) (Nycteribiidae).
1885d:
105. Stictomyia longicornis, p. clxvi (%, &). “Mexico”. Unchanged (Steyskal, 1968b: 22)
(Otitidae).
1885e:
106. Dimorphomyia calliphoroides, p. clxxiii (?sex). “Mexico”. Type in ?Oxford. Now
Asemosyrphus mexicanus (Macquart, 1842) (Thompson et al., 1976: 97) (Syrphidae).
1885f:
107. Melanostoma cyaneocincta, p. 251 (%). “Mexico”. Type in ?Oxford. Unplaced
Melanostomini (Thompson et al., 1976: 47) (Syrphidae).
108. Ocyptamus fenestratus, p. 251 (%). “Mexico”. Type in Oxford. Now a junior synonym of
Ocyptamus fuscipennis (Say, 1823) (Thompson et al., 1976: 19) (Syrphidae).
1886a:
109. Tetradiscus notatus, p. 371 (%). “Mexico”. Now Chyliza bigoti Melander, 1920 (Prado,
1975a: 2) (Psilidae).
110. Nerius plurivittatus, p. 372 (%). “Mexico”. Unchanged (Steyskal, 1968d: 5) (Neriidae).
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126
111. Nerius flavifrons, p. 372 (&). “Mexico”. Now in Glyphidops (Oncopsia) (Steyskal, 1968d:
3) (Neriidae).
112. Calobata callichroma, p. 373 (%). “Mexico”. Now a junior synonym of Ptilosphen
cyaneiventris (Macquart, 1846) (Steyskal, 1968a: 13) (Micropezidae).
113. Calobata calocephala, p. 375 (misprinted as 575) (%, &). “Mexico”. Now in Scipopus
(Steyskal, 1968a: 15) (Micropezidae).
114. Taeniaptera calosoma, p. 379 (%, &). “Mexico”. Now Rainieria (Rainieriella) calosoma
(Bigot) (Steyskal, 1968a: 15) (Micropezidae).
115. Taeniaptera pallidipennis, p. 380 (&). “Mexico”. Now Neotanypeza (Neotanypeza) ornatipes
(Bigot) (Steyskal, 1967: 1) (see next species) (Tanypezidae).
116. Taeniaptera ornatipes, p. 380 (%). “Mexico”. Now Neotanypeza (Neotanypeza) ornatipes
(Bigot) (Steyskal, 1967: 1) (Tanypezidae).
117. Grallomyia caloptera, p. 381 (%, &). “Mexico”. Now a junior synonym of Taeniaptera
ichneumonea Brauer, 1885 (Steyskal, 1968a: 17) (Micropezidae).
118. Micropeza obscura, p. 387 (%). “Mexico”. Unchanged (Steyskal, 1968a: 3)
(Micropezidae).
119. ?Enicopus fuscus, p. 387 (&). “Mexico”. Now Themira (Enicita) fusca (Bigot) (Steyskal,
1968c: 3, unrecognized) (Sepsidae).
1887a:
120. Conops nigrifascies, p. 40 (&). “Mexico”. Now in Physocephala (Papavero, 1971a: 8)
(Conopidae).
121. Conops carbonarius, p. 42 (%, &). “Mexico”. A junior synonym of the above (Papavero,
1971a: 8) (Conopidae).
1887b:
122. Zodion flavipenne, p. 204 (&). “Mexico”. Unchanged (Papavero, 1971a: 12) (Conopidae).
123. Zodion zebrinum, p. 204 (%). “Mexico”. Unchanged (Papavero, 1971a: 13) (Conopidae).
1887c:
124. Jurinia barbata, p. cxl (&). “Mexico”. Unchanged (Guimarães, 1971: 68) (Tachinidae).
125. Jurinia gonioides, p. cxl (&). “Mexico”. Now a junior synonym of Archytas lateralis
(Macquart, 1843) (Guimarães, 1971: 50) (Tachinidae).
126. Blepharipeza trichopsis, p. cxl (%). “Mexico”. An unrecognized species of Leschenaultia
(Guimarães, 1971: 186) (Tachinidae).
127. Echinomyia cora, p. cxl (&). “Mexico”. An unrecognized species of Peleteria (Guimarães,
1971: 44) (Tachinidae).
128. Echinomyia macrocera, p. cxl (&). “Mexico”. An unrecognized species of Peleteria
(Guimarães, 1971: 44) (Tachinidae).
129. Echinomyia notata, p. cxli (%). “Mexico”. An unrecognized species of Exopalpus
(Guimarães, 1971: 55) (Tachinidae).
130. Fabricia infumata, p. cxli (&). “Mexico”. An unrecognized species of Archytas (Guimarães,
1971: 50) (Tachinidae).
Acta Zool. Mex. (n.s.) 84 (2001)
127
131. Frontina rufostylata, p. cxli (%). “Mexico”. An unrecognized species of Chaetogaedia
(Guimarães, 1971: 178) (Tachinidae).
132. Frontina chrysopyga, p. cxli (&). “Mexico”. Now in Triachora (Guimarães, 1971: 184)
(Tachinidae).
133. Elachipalpus nigrifrons, p. cxli (&). “Mexico”. An unrecognized Tachinidae (Guimarães,
1971: 216) (Tachinidae).
1887d:
134. Calliphora xanthorrhina, p. clxxx (&). “Mexico”. Now in Mesembrinella (Guimarães, 1977:
35) (Calliphoridae).
135. Pyrellia obscuripes, p. clxxxi (%). “Mexico”. Now a junior synonym of Morellia ochricornis
(Wiedemann, 1830) (Pont, 1972: 9) (Muscidae).
136. Musca atrifrons, p. clxxxi (&). “Cuba et Mexique”. Now a junior synonym of Musca
domestica Linnaeus, 1758 (Pont, 1972: 7) (Muscidae).
137. Curtonevra fulvipes, p. clxxxii (&). “Mexico”. An unplaced Cyrtoneurininae (Pont, 1972:
58) (Muscidae).
138. Curtonevra vittigera, p. clxxxii (%). “Mexico”. Now a junior synonym of Hemichlora
scordalus (Walker, 1861) (Pont, 1972: 49) (Muscidae).
139. Curtonevra pallidicornis, p. clxxxii (&). “Mexico”. Now in Myospila (Pont, 1972: 27)
(Muscidae).
140. Curtonevra callimera, p. clxxxii (&). “Mexico”. Now in Morellia (Pont, 1972: 8) (Muscidae).
1888b:
141. Echinomyia rubrifrons, p. 80 (&). “Mexico”. An unrecognized species of Peleteria
(Guimarães, 1971: 45) (Tachinidae).
1888c:
142. Sphaerophoria nasuta, p. 253 (%). “Mexico”. Type in Oxford. Now Allograpta nasuta
(Bigot, 1884) (Thompson et al., 1976: 36) (Syrphidae).
143. Sphaerophoria trilimbata, p. 253 (%). “Mexico”. Type in Oxford. Now in Allograpta
(Thompson et al., 1976: 38) (Syrphidae).
144. Mesograpta quinquecincta, p. 254 (&). “Mexico”. Type in Oxford. Now Toxomerus
quinquecinctus (Bigot) (Thompson et al., 1976: 54) (Syrphidae).
145. Scopolia satanica, p. 254 (%). “Mexico”. Now in Penthosia (Guimarães, 1971: 17)
(Tachinidae).
146. Stevenia pallidiventris, p. 255 (%). “Mexico”. An unrecognized Phasiinae (Guimarães,
1971: 19) (Tachinidae).
147. Stevenia flavocalyptrata, p. 255 (%). “Mexico”. An unrecognized Phasiinae (Guimarães,
1971: 19) (Tachinidae).
148. Echinomyia cinerascens, p. 256 (%). “Mexico”. Now in Peleteria (Guimarães, 1971: 44)
(Tachinidae).
149. Exorista rufipalpis, p. 256 (%). “Mexico”. Now in Trafoia (Guimarães, 1971: 83)
(Tachinidae).
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128
150. Exorista rufata, p. 257 (%, &). “Mexique et Brésil (Pará)”. Now in Bolomyia (Guimarães,
1971: 187) (Tachinidae).
151. Chaetolyga erythropyga, p. 257 (&). “Mexico”. Now in Winthemia (Guimarães, 1971: 196)
(Tachinidae).
152. Chaetolyga nigriventris, p. 257 (%). “Mexico”. Not cited in Guimarães´s 1971 catalogue
(Tachinidae).
153. Chaetolyga dubia, p. 257 (%). “Mexico”. Now in Drino (Guimarães, 1971: 188)
(Tachinidae).
154. Chaetolyga nigripalpis, p. 258 (%). “Mexico”. Now in Eucelatoria (Guimarães, 1971: 133)
(Tachinidae).
155. Chaetolyga flavolimbata, p. 258 (%). “Mexico”. Now in Triodontopyga (Guimarães, 1971:
213) (Tachinidae).
156. Chaetolyga albopicta, p. 258 (&). “Mexico”. Now in Spathimeigenia (Guimarães, 1971:
148) (Tachinidae).
157. Chaetolyga occidentalis, p. 258 (%). “Mexico”. An unplaced Winthemiini (Guimarães,
1971: 198) (Tachinidae).
158. Chaetolyga nitidiventris, p. 258 (&). “Mexico”. An unplaced Winthemiini (Guimarães,
1971: 198) (Tachinidae).
159. Chaetolyga aenea, p. 259 (&). “Mexico”. An unplaced Blondeliini (Guimarães, 1971: 152)
(Tachinidae).
160. Degeeria cora, p. 259 (&). “Mexico”. Now in Eucelatoria (Guimarães, 1971: 133)
(Tachinidae).
161. Degeeria anthracina, p. 259 (&). “Mexico”. Now Oxynops anthracinus (Bigot) (Guimarães,
1971: 143) (Tachinidae).
162. Cestonia nigra, p. 259 (&). An unplaced Blondeliini (Guimarães, 1971: 153) (Tachinidae).
163. Macquartia atrifrons, p. 259 (&). “Mexico”. An unrecognized species of Pseudeuantha
(Guimarães, 1971: 104) (Tachinidae).
164. Prosopea americana, p. 260 (&). “Mexico”. An unplaced Blondeliini (Guimarães, 1971:
152) (Tachinidae).
165. Phorocera barbata, p. 260 (&). “Mexico”. Now in Metadoria (Guimarães, 1971: 140)
(Tachinidae).
166. Phorocera melanoceps, p. 260 (&). “Mexico”. A junior synonym of Metadoria barbata
(Bigot) (Guimarães, 1971: 140) (Tachinidae).
167. Ceromasia quadrivittata, p. 261 (&). “Mexico”. An unrecognized Dexiinae (Guimarães,
1971: 122) (Tachinidae).
168. Ceromasia zonata, p. 261 (&). “Mexico”. Now in Spathimeigenia (Guimarães, 1971: 149)
(Tachinidae).
169. Ceromasia pictigaster, p. 261 (&). “Mexico”. An unplaced Dexiini (Guimarães, 1971: 97)
(Tachinidae).
170. Ceromasia castanifrons, p. 261 (&). “Mexico”. An unrecognized species of Prospherysa
(Guimarães, 1971: 213) (Tachinidae).
171. Ceromasia chrysocephala, p. 261 (&). “Mexico”. An unplaced Blondeliini (Guimarães,
1971: 152) (Tachinidae).
Acta Zool. Mex. (n.s.) 84 (2001)
129
172. Ceromasia spinipes, p. 262 (&). “Mexico”. An unplaced Blondeliini (Guimarães, 1971:
153) (Tachinidae).
173. Ceromasia abbreviata, p. 262 (&). “Mexico”. An unplaced Blondeliini (Guimarães, 1971:
152) (Tachinidae).
174. ?Viviania rufopygata, p. 262 (&). “Mexico”. An unrecognized Leskiini (Guimarães, 1971:
120) (Tachinidae).
175. ?Viviania citrina, p. 262 (&). “Mexico”. Now a junior synonym of Chaetona icterica
(Wiedemann, 1830) (Guimarães, 1971: 105) (Tachinidae).
176. Micropalpus nigrifrons, p. 263 (&). “Mexico”. An unplaced Linnaemyini (Guimarães,
1971: 83) (Tachinidae).
177. Masicera flavifacies, p. 263 (&). “Mexico”. An unplaced Sturmiini (Guimarães, 1971: 193)
(Tachinidae).
178. Tricoliga caloptera, p. 263 (&). “Mexico”. Now junior synonym of Pseudeuantha indita
(Walker, 1860) (Guimarães, 1971: 104) (Tachinidae).
179. Tricoliga fulvidapex, p. 263 (%). “?Mexique ou ?Celebes”. An unrecognized species of
Winthemia (Guimarães, 1971: 196) (Tachinidae).
180. Prosena maculifera, p. 264 (&). “Mexico”. An unrecognized species of Stomatodexia
(Guimarães, 1971: 119) (Tachinidae).
181. Prosena obscura, p. 264 (%). “Mexico”. Now in Eudexia (Guimarães, 1971: 29)
(Tachinidae).
182. Prosena curvirostris, p. 264 (%,&). “Mexique ou Brésil”. Now in Prosenoides (Guimarães,
1971: 32) (Tachinidae).
183. Rhamphinina formidabilis, p. 264 (%). “Mexico”. Now in Eudexia (Guimarães, 1971: 29)
(Tachinidae).
184. Rhamphinina major, p. 265 (%). ‘Mexique et Washington Territory’. Now in Ptilodexia
(Guimarães, 1971: 33) (Tachinidae).
185. Rhamphinina anthracina, p. 265 (%,&). “Mexico”. Now in Ptilodexia (Guimarães, 1971: 33)
(Tachinidae).
186. Rhynchodexia tincticornis, p. 266 (%). “Mexico, Paso de Macho”. Now in Ptilodexia
(Guimarães, 1971: 34) (Tachinidae).
187. Microphthalma calogaster, p. 266 (%). “Mexico, Orizaba”. Now in Macrometopa
(Guimarães, 1971: 23) (Tachinidae).
188. Myocera simplex, p. 266 (&). “Mexico”. Now in Ptilodexia (Guimarães, 1971: 34)
(Tachinidae).
189. Tromodesia hoemorrhoidalis (sic), p. 267 (?sex). “Mexico”. Now a junior synonym of
Tromodesiopsis atrifrons (Wiedemann, 1830) (Guimarães, 1971: 102) (Tachinidae).
190. Homodexia longicornis, p. 267 (%). “Mexico”. Now in Pseudodexia (Guimarães, 1971:
108) (Tachinidae).
191. Homodexia vittigera, p. 267 (?&). “Mexico”. Now Thelairodes vittigerus (Bigot)
(Guimarães, 1971: 95) (Tachinidae).
192. Homodexia flavipes, p. 268 (&). “Mexico”. Now in Eucalodexia (Guimarães, 1971: 106)
(Tachinidae).
193. Homodexia spinosa, p. 268 (%). “Mexico”. Now Thelairodes spinosus (Bigot)
(Guimarães, 1971: 95) (Tachinidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
130
194. Homodexia triangulifera, p. 268 (%). “Mexico”. Now in Ptilodegeeria (Guimarães, 1971:
147) (Tachinidae).
195. Oplisa albifacies, p. 268 (&). “Mexico”. Now in Pseudodexia (Guimarães, 1971: 108)
(Tachinidae).
196. Oplisa nigrifacies, p. 268 (%, &). “Mexico”. Now in Metadoria (Guimarães, 1971: 140)
(Tachinidae).
197. Pyrrosia ochracea, p. 268 (%, &). “Mexico”. Not cited in Guimarães´s 1971 catalogue
(Tachinidae).
198. Phorostoma appendiculata, p. 269 (&). “Mexico”. An unrecognized species of Myiomima
(Guimarães, 1971: 31) (Tachinidae).
199. Dexiosoma fumipennis, p. 270 (&). “Mexico”. Now in Ptilodexia (Guimarães, 1971: 33)
(Tachinidae).
200. Dexiosoma partita, p. 270 (&). “Mexico”. An unrecognized Tachinidae (Guimarães, 1971:
216).
201. Anthracomyia pallidicornis, p. 270 (%). “Mexico”. Now in Pseudodexia (Guimarães, 1971:
108) (Tachinidae).
1888e:
202. Psilopodinus astequinus, p. xxx (%). “Mexico”. Now in Condylostylus (Robinson, 1970: 3)
(Dolichopodidae).
203. Poecilobothrus mexicanus, p. xxx (%). “Mexico”. Now in Pelastoneurus (Robinson, 1970:
51) (Dolichopodidae).
1892a:
204. Anthrax eurhinatus, p. 355 (&). “Mexico”. Now Villa (Paravilla) eurhinata (Bigot) (Painter
et al., 1978: 51) (Bombyliidae).
205. Bombylius albopenicillatus, p. 363 (%). “Mexico”. Now in Parabombylius (Painter et al.,
1978: 4) (Bombyliidae).
12.
T
HE BROTHERS
D
UGÈS
Alfred Auguste Delsescautz Dugès (Fig. 8) and his brother Eugène (Fig. 9) were
born in Montepellier, France, around 1826. Their father, Antoine Louis Delsescautz,
was a medical doctor, professor of the University of Paris, and a naturalist. Upon
their father’s death, the two brothers moved to Paris to study medicine. Alfred left
for Mexico, arriving there in 1853. As soon as his medical title was recognized in
Mexico City, he moved to Guanajuato, where he remained practicing medicine and
teaching natural history. His brother Eugène came to Mexico only in 1865, also
becoming established in Guanajuato as a medical doctor. Eugène would send many
collections of insects to Paris. Later he founded a natural history museum in Morelia
and died in that city on 13 January 1895 (Anon., 1895). Alfred also used to collect
Acta Zool. Mex. (n.s.) 84 (2001)
131
in the neighbourhoods of Guanajuato, especially on Sundays, accompanied by his
students. He died at 84 years of age on 7 January 1910 (Adler, 1989; Barnhart,
1965; Duellman, 1961; Howard, 1930; Kellogg, 1932; Martín del Campo, 1937;
McVaugh, 1972; Palmer et al., 1954; Papavero, 1971b; Smith & Smith, 1969, 1973;
Stafleu & Cowan, 1976 (1:692)).
Alfred Dugès authored the first paper on Diptera published in Mexico (1887),
describing Ornithomyia villadae (p. 20, pl. 3, figs. 3a-b), now a junior synonym of
Lynchia nigra (Perty, 1833) (Guimarães, 1968a: 6) (Hippoboscidae).
Figures 8 - 9
8. Alfred Auguste Delsescautz Dugès. Redrawn by S. Ibáñez-Bernal from a photograph published in Howard
(1930). 9. Eugène Dugès. Redran by S. Ibáñez-Bernal from a photograph poublished in Dugesiana, 6(2)
(1999).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
132
13.
T
HE SPECIES DESCRIBED BY
F
RIEDRICH
M
ORITZ
B
RAUER
Friedrich Moritz Brauer was born on 12 May 1832 in Vienna (Mariahilferstrasse
335), the son of Justus Brauer and his second wife Louise Braun, from Hannover.
The house where Brauer was born, now in the commercial center of Vienna, was
at that time the cottage of the family, which actually resided in the center of Vienna.
In the gardens and wells of his home, Brauer started to observe the biology of
insects, noting their metamorphosis; he reared mosquitoes in vials.
In 1838 he had as teacher of drawing Anton Löw, who also had the habit to
collect insects, and with him Brauer learned the collecting techniques. His father
and his uncle Daniel Braun, as well as his brother Theodor were also fond of
entomology. Thus Friedrich very early had his tastes inclined to that field of zoology.
Upon his father’s death in 1839, the family spent a last summer in
Mariahilferstrasse, as the following year they moved to Germany, where they lived
with some relatives. In 1841 they rented a cottage in Döbling. There Brauer started
excursions with Löw, especially in Krottentoch and Kallenberg. In 1842 his mother
married again, with Dr. Victor Ivanchich. Summers were then spent in Brühl, where
Friedrich frequently hunted insects in the company of Löw.
In 1845 Brauer was introduced to Gustav Gözsys, who at that time frequented the
second grade in the gymnasium. Gözsys’ father, the medical doctor of the Brauer
family since 1842, convinced them to let Friedrich frequent the public school, for
which he was prepared by private teachers since 1843. Brauer entered the school
of the ’Piaristen’, after passing examinations. He suffered bad moments, however
, for until then he had no contact with other children of his age. In 1846, being in the
third grade in the gymnasium, he was given a small collection of exotic insects,
which he proposed to identify. He was introduced to the then curator of insects in
the Hofsmuseum, Vincenz Kollar, by his uncle Daniel, with whom Kollar had
relations. From then on, Friedrich began to frequent the collections of the Vienna
Hofsmuseum, thus obtaining a good knowledge of all the orders of insects. By that
time he received Burmeister’s “Handbuch der Entomologie”, which he studied
assiduously, in spite of the fact that he could not assimilate quite a few things, due
to his age.
Thus Brauer spent his time, collecting and studying insects, until the political
crisis of 1848, which affected his family. In this same year, the gymnasium where
he studied suffered many reforms in the teaching methods, which brought him many
difficulties. Precociously interested in research, he had no taste for the regular
courses at school. By that time he also became interested in the biology of the
Odonata, and in 1850 published his first paper on the subject, in the annals of the
“Gesellschadt der Freunde der Naturwissenschaft” of Vienna, through the good
Acta Zool. Mex. (n.s.) 84 (2001)
133
efforts of Georg Frauenfeld, whom he had previously met. Enthused with his first
successes, he dedicated himself to the study of insect metamorphosis, especially
of the Neuropteroidea, a subject which he would investigate for many years to
come.
From July to September 1850 he traveled with his family to Adelsberg, Trieste,
Venice and Tyrol. As in his gymnasium, his fame of naturalist brought jealously, and
he changed schools.
In March 1852 he was not able to pass the “matura” exams; in September of the
same year he again tried, with the same result. His family decided then to take
some providences, and applied directly to the Minister of Education, quiring another
chance; a boy with published papers, accepted by the Royal Academy of Vienna
and other scientific associations, they noted, could not depend on an examination
to proceed in his career. The new chance was granted, and after passing
examinations on mathematics and Latin, Brauer entered the University, to follow the
course of medicine. He was given at that occasion Westwoods’s “Modern
classification of insects”, which induced him to study English.
On May 1854 he lost his mother, and his brother Albert became his tutor. At that
time he made the acquaintance of Schiner and Johann Egger. Schiner was not yet
a dipterist, being interested in ornithology and botany. As Brauer had a reasonable
collection of flies, Frauenfeld, Egger and Schiner became interested in the
systematics of that Order, and tried to identify them through the works of Meigen.
In 1854 and 1855 Brauer and Egger undertook frequent excursions to collect
Diptera.
In 1856 Brauer married Leontine Boschetty, whom he knew already for some
years. In the same year, Karl Lang found in the Pratter a specimen of Oestrus pictus,
and Brauer decided to start the study of the Oestroidea, beginning to compile the
literature and to make field observations. Visiting the Pratter and Imperial Zoological
Garden he was able to rear several parasitic oestrids, completing his notes and
studies, being helped by Rogenhofer, Zeller and Hermann Loew.
In 1860 he published on Oestrus hominis, based on larvae brought from Brazil by
Johann Natterer, deposited in the Vienna Museum, erecting then the genus
Dermatobia.
Finally, after 7 years of studies of medicine, Brauer passed the “Rigorosum” and
graduated. Soon afterwards he contracted typhus, and, due to his precarious
health, gave up the idea of passing a second “Rigorosum”.
In October 1861 he accepted a position in the Naturalienkabinet (Vienna
Museum), leaving medicine to dedicate himself exclusively to natural history.
However, he remained at the museum for 16 year as curator of molluscs. During
that time, with J. F. Bergenstamm, he published the famous monographs of
Papavero & Ibañez-Bernal: History of Mexican Dipterology
134
Muscoidea (1889-1894). In that series, entitled “Die Zweiflügler des Kaiserlichen
Museum zu Wien”, Brauer and Bergenstamm proposed many genera , especially
in the Family Tachinidae.
Brauer also published on the higher classification of the Diptera and on the
phylogeny of insects. In 1871 he obtained his doctor’s degree in the faculty of
philosophy, and two years latter the title of “Professor Extraordinarius”. With the
death of Frauenfeld (1873), Ferrari (1876) and Redtenbacher (1876), the number
of members of the Museum diminished and only then Brauer obtained a better
position. He dedicated himself entirely to the study of the Diptera, identifying and
organizing the collections of the Vienna Museum, which included those of
Winthem, Wiedemann, Egger and Schiner.
In 1876 he was named curator of the entomological section of the Museum and
made a member of the Academy of Science and other learned societies . From
1880 on, he started publishing his monumental work on the Diptera of the Imperial
Museum of Vienna (“Die Zweiflügler des Kaiserlichen Museums zu Wien”). In 1884
he was made “Ordenlich Professor” and in 1888 effective member of the Academy
of Sciences.
The elaboration of the “Zweiflügler” left him very little time. At that period the
Naturalienkabinet was incorporated to the Hofsmuseum. He revised the collections
of Robineau-Desvoidy, Rondani and Bigot. His collaborator, Bergenstamm, was not
an ideal partner, but as he had many important types, Brauer had to accept his
collaboration. The great work was finished only in 1893. During this period he lost
his wife and married for a second time, with Ludmila von Koenig.
With the advancement of the years he felt old and fatigued, and needed much
stimulation to prepare his works. He published less and less. In 1896 he had the
pleasure of describing the larva of Cobboldia extracted from an Indian elephant.
With the decline of his mental activities and with the death of his second wife, he
became embittered.
In 1898 he was made head of the zoological sections of the Museum, but this
position was reached too late in his life to introduce in the Museum the changes he
expected to impose. His last paper was Mik’s necrologue (1901).
He died on 29 December 1904, after much suffering, at the house of his daughter
Laura, the wife of the malacologist A. Wagner (Handlirsch, 1905).
Brauer described only a few species of Mexican Diptera, listed below. His types
are all deposited in the Vienna Natural History Museum.
1882:
1. Cynipimorpha bilimeki, p. 75, (sex?). “Mexico, Veracruz, Orizaba; Morelos, Cuernavaca” (as
bilimecki, lapsus). Unchanged (James, 1973:53) (Stratiomyidae).
Acta Zool. Mex. (n.s.) 84 (2001)
135
1885:
2. Calobata ichneumonea, p. 388, fig. 4 (pl.) (sex?). “Mexico, Veracruz, Orizaba”. Now in
Taeniaptera (Steyskal, 1968a: 17) (Micropezidae).
1897:
3. Melanota dubia, p. 357 (sex?). “Mexico”. An unrecognized Tachinidae (Guimarães, 1971:
215).
The remaining species were described with Bergenstamm:
1889:
4. Phasiopteryx bilimeki, p. 147 (1889:79). “Mexico, Veracruz, Orizaba”. Now in Ormia
(Guimarães, 1971:21) (Tachinidae).
5. Macrometopa mexicana, p. 117 (1889:49). “Mexico”. Now a junior synonym of Macrometopa
calogaster (Bigot, 1889) (Guimarães, 1971: 23) (Tachinidae).
6. Hystrisiphona pseudohystricia, p. 120 (1889: 93). “Mexico, Tacubaya”. Now in Echinodexia
(Guimarães, 1971: 29) (Tachinidae).
7. Myiopharus metopia, p. 161 (1889: 93). “Mexico, Tacubaya, and Yucatan, Merida”.
Unchanged (Guimarães, 1971: 142) (Tachinidae).
8. Masipoda geminata, p. 102 (1889: 94). “Mexico”. Now in Winthemia (Guimarães, 1971:196)
(Tachinidae).
1891:
9. Gaediopsis mexicana, p. 336 (1891:32). “Mexico”. Unchanged (Guimarães, 1971:179)
(Tachinidae).
10. Paragaedia hedemanni, p. 350 (1891:45). “Mexico”. Now in Chaetoprosopa (Guimarães,
1971: 184) (Tachinidae).
1893:
11. Cyrtosoma rufum, p. 132 (1893: 44). “Mexico, Guerrero, Omilteme”. Now in Ocyrtosoma
(Guimarães, 1971: 37) (Tachinidae).
12. Wulpia aperta, pp. 128, 188 (1893: 40, 100). “Mexico”. Now in Catemophrys (Guimarães,
1971: 114) (Tachinidae).
13. Microchira mexicana, p. 128 (1893: 40). “Mexico”. Now in Paradidyma (Guimarães, 1971:
115) (Tachinidae).
14. Paradidyma aperta, p. 127 (1893: 39). “Mexico”. A junior synonym of Paradidyma mexicana
(Brauer & Bergenstamm, 1893) (Guimarães, 1971: 115) (Tachinidae).
15. Metadoria mexicana, p. 117 (1893: 20). “Mexico”. A junior synonym of Metadoria barbata
(Bigot, 1889) (Guimarães, 1971: 140) (Tachinidae).
16. Gaediophana atra, pp. 123, 201 (1893: 35, 113). “Mexico” Unchanged (Guimarães, 1971:
179) (Tachinidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
136
14.
T
HE SPECIES DESCRIBED BY
E
RMANNO
G
IGLIO
-T
OS
Ermanno Giglio-Tos (Fig. 10) was born in Turin on August 25, 1865. He started
his studies of natural history under the guidance of M. Lessona and L. Camerano,
with whom he learned the principles of systematic zoology. Even in later years,
when he dedicated himself to studies of vertebrate anatomy, hematology, cytology,
mechanics of development and applied zoology, he never quite abandoned the
study of systematics, of which he was particularly fond.
Figure 10
Ermanno Giglio-Tos. Redrawn by S. Ibáñez-Bernal from a photograph property of N. Papavero.
Acta Zool. Mex. (n.s.) 84 (2001)
137
Entering the Museo Zoologico di Torino at the age of 20, Giglio-Tos became
interested in the systematics of insects, and began his activities with a number of
papers on Diptera, of which there was an abundant collection, left by Bellardi (see
Part 6). His first publications were on European and African flies, but his attention
turned afterwards to Mexican and Central American flies. He published short papers
from 1890 to 1893, and his great work, the “Ditteri del Messico”, was published from
1892 to 1895, in 4 parts. As Bellardi had not finished his studies of Mexican Diptera,
Giglio-Tos completed the work, describing the remaining Mexican collections of
Stratiomyidae, Aschiza, Calyptratae and Acalyptratae. In 1896, he published his last
paper on Neotropical Diptera, describing a striking new genus of Tabanidae brought
from panama by Enrico Festa.
As the Museum of Turin was receiving a great number of insects, Giglio-Tos
became gradually interested in the study of other orders of insects, and after some
essays in Coleoptera and Hemiptera, became definitely interested in the Orthoptera
(sensu lato), in which he was to become known as a specialist.
Appointed professor of zoology and comparative anatomy at the Reale Università
di Cagliari, when he was 36 years old, he founded the Instituto di Biologia Marina
in San Bartolomeo. He was next called to be professor of zoology and vertebrate
anatomy at the University of Firenze, where he remained only two years, passing
on to the chair of anatomy and physiology of the University of Turin, where he
occupied the chair of zoology, returning then to Cagliari, where he stayed for seven
years. His last position was that of professor of zoology in Turin, to which he was
appointed in December 1925. He died there on August 18, 1926 (Zavatari, 1927).
All the types of Giglio-Tos are deposited in the Turin Zoological Museum. He
described the following Mexican species:
1890:
1. Ditomyia zonata, p. 2, pl. II, fig. 6 (%, &). “Mexico, Veracruz, Orizaba (Sumichrast). Box 1.
As an unplaced Ditomyiidae (Papavero, 1977: 4) (Ditomyiidae).
2. Ditomyia mexicana, p. 3, pl. II, fig. 7 (?sex). “Mexico, Veracruz, Orizaba (Sumichrast)”. Box
1. As an unplaced Ditomyiidae (Papavero, 1977: 4) (Ditomyiidae).
1891a:
3. Heteracanthia mexicana, p. 1 (&) (1892e: 8). “Mexico (Sumichrast)”. Box 8. Now
Heteracanthia ruficornis Macquart, 1849 (James, 1973: 6) (Stratiomyidae).
4. Cyphomyia ochracea, p. 2 (&) (1892e: 10, pl. I, fig. 4). “Mexico (Sumichrast)”. Box 8.
Unchanged (James, 1973: 27) (Stratiomyidae).
5. Euparyphus carbonarius, p. 2 (%). (1892e: 12, pl. I, fig. 5). “Mexico (Sumichrast)”. Box 8.
Now Euparyphus (Euparyphus) carbonarius Giglio-Tos (James, 1973: 31) (Stratiomyidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
138
6. Clitellaria stigma, p. 2 (&) (1892e: 13). “Mexico (Sumichrast)”. Type ?lost. Now in
Dieuryneura (James, 1973: 31) (Stratiomyidae).
7. Acanthina? bellardii, p. 2 (&) (1892e: 14). “Tehuacan (Sumichrast)”. Box 8. Now in Artemita
(James, 1973: 50) (Stratiomyidae).
8. Microchrysa nova, p. 2 (&) (1892e: 16). “Mexico”. Box 8. Unchanged (James, 1973: 17)
(Stratiomyidae).
9. Merosargus hyalopterus, p. 3 (%,&) (1892e: 19). “Orizaba (Sumichrast)”. Box 8. Unchanged
(James, 1973: 13) (Stratiomyidae).
10. Merosargus dissimilis, p. 3 (%) (1892e: 20, pl. I, fig. 7). “Orizaba”. Box 8. Unchanged
(James, 1973: 12) (Stratiomyidae).
11. Merosargus orizabae, p. 3 (%) (1892e: 20). “Orizaba (Sumichrast)”. Box 9. Unchanged
(James, 1973: 14) (Stratiomyidae).
12. Merosargus coriaceus, p. 4 (%, &) (1892e: 21). “Orizaba (Sumichrast)”. Box 8. Unchanged
(James, 1973: 12) (Stratiomyidae).
13. Sargus (?Macrosargus) filiformis, p. 4 (%) (1892e: 28). “Mexico”. Box 8. Now a junior
synonym of Sargus caesius Bellardi, 1859 (James, 1973: 18) (Stratiomyidae).
1892c:
14. Mixogaster dimidiata, p. 1 (&) (1892e: 33, pl. I, figs. 9, 9a). “Tuxpango (Sumichrast)”. Box
15. Unchanged (Thompson et al., 1976: 58) (Syrphidae).
15. Ubristes chrysopyga, p. 1 (&) (1892e: 37). “Orizaba (Sumichrast)”. Box 15. Now Microdon
(Ubristes) chrysopygus (Giglio-Tos) (Thompson et al., 1976: 61) (Syrphidae).
16. Microdon aquilinus, p. 2 (&) (1892e: 37). “Tuxpango (Sumichrast)”. Box 15. Unchanged
(Thompson et al., 1976: 63) (Syrphidae).
17. Copestylum parvum, p. 2 (%, &) (1892e: 42). “Tehuacan (Sumichrast)”. Box 15. Now a
junior synonym of Copestylum simile (Giglio-Tos, 1892) (Thompson et al., 1976: 84)
(Syrphidae).
18. Copestylum simile, p. 2 (%, &) (1892e: 42). “Tehuacan (Sumichrast)”. Box 15. Unchanged
(Thompson et al., 1976: 84) (Syrphidae).
19. Volucella omochroma, p. 2 (&) (1892e: 47). “Mexico, without locality”. Box 15. Now in
Copestylum (Thompson et al., 1976: 80) (Syrphidae).
20. Volucella furens, p. 2 (%) (1892e: 48). “Mexico (Sallé)”. Box 15. Now in Copestylum
(Thompson et al., 1976: 76) (Syrphidae).
21. Volucella craverii, p. 2 (%) (1892e: 49). “Mexico (Craveri)”. Box 15. Now in Copestylum
(Thompson et al., 1976: 74) (Syrphidae).
22. Volucella flavissima, p. 3 (&) (1892e: 50). “Orizaba (Sumichrast)”. Box 15. Now Copestylum
flavissimum (Giglio-Tos) (Thompson et al., 1976: 75) (Syrphidae).
23. Volucella minima, p. 3 (%) (1892e: 53). “Orizaba (Sumichrast)”. Box 15. Now Copestylum
minimum (Giglio-Tos) (Thompson et al., 1976: 79) (Syrphidae).
24. Volucella dichroica, p. 3 (&) (1892e: 55). “Huastec (Saussure)”. Box 15. Now Copestylum
dichroicum (Giglio-Tos) (Thompson et al., 1976: 74) (Syrphidae).
25. Volucella trigona, p. 3 (%) (1892e: 52). “Orizaba (Sumichrast)”. Box 15. Now Copestylum
trigonum (Giglio-Tos) (Thompson et al., 1976: 85) (Syrphidae).
26. Volucella hyaloptera, p. 3 (&) (1892e: 57). Tampico (Saussure)”. Box 16. Now Copestylum
hyalopterum (Giglio-Tos) (Thompson et al., 1976: 77) (Syrphidae).
Acta Zool. Mex. (n.s.) 84 (2001)
139
27. Volucella hirsuta, p. 3 (%) (1892e: 60, as synon. of coesariata Williston). “Meztillan
(Saussure)”. Box 16. Now a junior synonym of Copestylum caesariatum (Williston, 1891)
(Thompson et al., 1976: 73) (Syrphidae).
28. Volucella hispida, p. 4 (%) (1892e: 61, as synon. of ornata Williston). “Orizaba
(Sumichrast)”. Box 16. Now a junior synonym of Copestylum ornatum (Williston, 1891)
(Thompson et al., 1976: 80) (Syrphidae).
29. Volucella volucris, p. 4 (%) (1892e: 61). “Orizaba”. Box 16. Now Copestylum volucre
(Giglio-Tos) (Thompson et al., 1976: 87) (Syrphidae).
30. Volucella hystrix, p. 4 (%) (1892e: 62). “Tuxpango (Sumichrast)”. Box 16. Now in
Copestylum (Thompson et al., 1976: 77) (Syrphidae).
31. Volucella brevis, p. 4 (%, &) (1892e: 63). “Meztillan (Saussure)”. Box 16. Now Copestylum
breve (Giglio-Tos) (Thompson et al., 1976: 72) (Syrphidae).
32. Volucella obesoides, p. 4 (%) (1892e: 65). “Mexico (Craveri)”. Box 16. Now a junior
synonym of Ornidia obesa (Fabricius, 1775) (Thompson et al., 1976: 70) (Syrphidae).
33. Eristalis bombusoides, p. 4 (%, &) (1893c: 3, as synon. of circe Williston). “Oaxaca
(Sumichrast)”. Box 16. Now a junior synonym of Eristalis (Eoseristalis) circe Williston, 1891
(Thompson et al., 1976: 102) (Syrphidae).
34. Eristalis trilimbata, p. 5 (&) (1893c: 8). “Tampico (Saussure)”. Box 16. Now a junior
synonym of Palpada vinetorum (Fabricius, 1798) (Thompson et al., 1976: 110) (Syrphidae).
35. Eristalis praeclarus, p. 5 (%) (1893c: 11, as synon. of rufiventris Macquart). “Tuxpango
(Sumichrast)”. Type ?lost. Now a junior synonym of Palpada rufiventris (Macquart, 1846)
(Thompson et al., 1976: 108) (Syrphidae).
36. Eristalis clarissimus, p. 5 (%) (1893c: 11). “Tuxpango (Sumichrast)”. Box 16. Now Palpada
clarissima (Giglio-Tos) (Thompson et al., 1976: 104) (Syrphidae).
37. Eristalis sallei, p. 5 (&) (1893c: 12). “Mexico (Sallé)”. Box ?. Now a junior synonym of
Palpada agrorum (Fabricius, 1787) (Thompson et al., 1976: 103) (Syrphidae).
38. Eristalis triangularis, p. 6 (&) (1893c: 9). “Cuautla”. Box 16. Now in Palpada (Thompson et
al., 1976: 110) (Syrphidae).
39. Asemosyrphus olivaceus, p. 6 (&) (1893c: 10, as synon. of bicolor Bigot). “Mexico”. Box 17.
Now a junior synonym of Asemosyrphus arquatus (Say, 1829) (Thompson et al., 1976: 96)
(Syrphidae).
40. Asemosyrphus griseus, p. 6 (%, &) (1893c: 20, as synon. of mexicanus Macquart).
“Tehuacan”. Box 17. Now a junior synonym of Asemosyrphus mexicanus (Macquart, 1842)
(Thompson et al., 1976: 97) (Syrphidae).
41. Asemosyrphus impurus, p. 6 (&) (1893c: 19, as synon. of bicolor Bigot). “Mexico”. Box 17.
Now a junior synonym of Asemosyrphus arquatus (Say, 1829) (Thompson et al., 1976: 96)
(Syrphidae).
42. Platynochaetus niger, p. 6 (&) (1893c: 20, pl. II, figs. 1, 1a). “Orizaba (Sumichrast)”. Box
17. Now a junior synonym of Quichuana championi (Williston, 1892) (Thompson et al.,
1976: 98) (Syrphidae).
43. Salpingogaster nova, p. 7 (&) (1893c: 23, pl. II, fig. 3). “No locality”. Box 17. Unchanged
(Thompson et al., 1976: 32) (Syrphidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
140
44. Crioprora arctophiloides, p. 7 (%, &) (1893c: 25, pl. II, figs. 2, 2a). “Mexico (Craveri),
Angang(ueo) (Saussure)”. Box 17. Now in Criorhina (Thompson et al., 1976: 118)
(Syrphidae).
1892d:
45. Eristalis sumischrasti (sic), p. 1 (%) (1893c: 6, corrected to sumichrasti). “Tehuacan
(Sumichrast)”. Box 16. Now a junior synonym of Palpada testaceicornis (Macquart, 1850)
(Thompson et al., 1976: 110) (Syrphidae).
46. Eristalis atropos, p. 1 (%) (1893c: 14, pl. II, fig. 23). “Mexico (Sallé)”. Box 17. Now a junior
synonym of Palpada trigona (Williston, 1891) (Thompson et al., 1976: 110) (Syrphidae).
47. Syrphus lautus, p. 2 (%) (1893c: 29). “Tuxpango (Sumichrast)”. Box 17. Now in Ocyptamus
(Thompson et al., 1976: 21) (Syrphidae).
48. Syrphus saussurii, p. 2 (%) (1893c: 30). “Orizaba (Saussure)”. Box 17. Now in Allograpta
(Thompson et al., 1976: 37) (Syrphidae).
49. Sphaerophoria syrphica, p. 2 (%) (1893c: 32, pl. II, figs. 5, 5a). “Orizaba (Sumichrast)”. Box
17. Now in Allograpta (Thompson et al., 1976: 37) (Syrphidae).
50. Melanostoma elegans, p. 2 (%, &) (1893c: 39, pl. II, fig. 21). “Orizaba (Sumichrast)”. Box
18. Now in Ocyptamus (Thompson et al., 1976: 17) (Syrphidae).
51. Melanostoma bellum, p. 3 (&). (1893c: 38, pl. II, figs. 7, 7a). “Mexico, without locality”. Box
?. Unchanged (Thompson et al., 1976: 42) (Syrphidae).
52. Mesogramma rombicum, p. 3 (%, &) (1893c: 46, pl. II, fig. 13). “Orizaba (Sumichrast)”. Box
18. Now Toxomerus rombicus (Giglio-Tos) (Thompson et al., 1976: 54) (Syrphidae).
53. Mesogramma diversum, p. 3 (&) (1893c: 48, pl. II, fig. 13). “Mexico, without locality”. Box
18. Now Toxomerus diversus (Giglio-Tos) (Thompson et al., 1976: 50) (Syrphidae).
54. Mesogramma ciliatum, p. 3 (%, &) (1893c: 50, pl. II, figs. 16, 16a-b). “Tuxpango
(Smichrast)”. Box 18. Now Toxomerus ciliatus (Giglio-Tos) (Thompson et al., 1976: 49)
(Syrphidae).
55. Mesogramma comma, p. 4 (%) (1893c: 53). “Mexico (Sallé, Truqui)”. Box 18. Now in
Toxomerus (Thompson et al., 1976: 49) (Syrphidae).
56. Baccha spatulata, p. 4 (&) (1893c: 56, pl. II, figs. 18, 18a). “Orizaba (Sumichrast)”. Box 18.
Now Ocyptamus spatulatus (Giglio-Tos) (Thompson et al., 1976: 27) (Syrphidae).
57. Chilosia aurotecta, p. 4 (%, &) (1893c: 58, pl. II, fig. 22). “Orizaba (Sumichrast)”. Box 19.
Now Cheilosia aurotecta Giglio-Tos (Thompson et al., 1976: 68) (Syrphidae).
58. Physocephala maxima, p. 5 (&) (1893c: 64). “Mexico”. Box 19. Unchanged (Papavero,
1971a: 9) (Conopidae).
59. Conops ocellatus, p. 5 (%). (1893c: 67, as synon. of parvus Williston). “Mexico
(Sumichrast)”. Box 19. Now in Physoconops (Papavero, 1971a: 3) (Conopidae).
1892e:
60. Berismyia fusca, p. 7, pl. I, fig. 2 (%). “Mexico (Sallé)”. Box. 7. Unchanged (James, 1973:
5) (Stratiomyidae).
61. Hermetia flavipes var. aeneipennis, p. 11 (&). “Mexico (Sumichrast)”. Box 8. Now Hermetia
aeneipennis Giglio-Tos (James, 1973: 36) (Stratiomyidae).
Acta Zool. Mex. (n.s.) 84 (2001)
141
62. Chrysonotus aeneiventris, p. 25 (%, &). “Mexico, Orizaba (Sumichrast)”. Box 9. Now a
junior synonym of Himantoloba flavopilosa (Bigot, 1879) (James, 1973: 10)
(Stratiomyidae).
63. Copestylum distinctum, p. 41, pl. I, fig. 15 (%). “Tehuacan (Sumichrast)”. Box 15. Now a
junior synonym of Copestylum marginatum (Say, 1830) (Thompson et al., 1976: 78)
(Syrphidae).
64. Camerania macrocephala, p. 45, pl. I, fig. 13 (%). “Mexico (Sallé). Box ?. Now Copestylum
macrocephalum (Giglio-Tos) (Thompson et al., 1976: 78) (Syrphidae).
1893a (Giglio-Tos did not cite the localities in this paper, only in the 1894 paper):
65. Hystrisiphona (sic) bicolor, p. 1 (%) (1894: 54, fig. 17). “Mexico (Truqui)”. Box 24. Now in
Jurinodexia (Guimarães, 1971: 30) (Tachinidae).
66. Mochlosoma anale, p. 1 (?sex) (1894: 55 (%)). “Mexico (Truqui)”. Box ?. Unchanged
(Guimarães, 1971: 30) (Tachinidae).
67. Mochlosoma sericeum, p. 2 (&) (1894: 56). “Mexico (Truqui)”. Box 24. Unchanged
(Guimarães, 1971: 31) (Tachinidae).
68. Hystrichodexia mellea, p. 2 (%) (1894: 58). “Oaxaca (Sallé)”. Box 24. Unchanged
(Guimarães, 1971: 30) (Tachinidae).
69. Hystrichodexia aurea, p. 2 (&) (1894: 59). “Mexico, without locality (Sumichrast)”. Box 24.
Now in Sumichrastia (Guimarães, 1971: 35) (Tachinidae).
70. Scotiptera cyanea, p. 2 (%) (1894: 61). “Angang(ueo) (Saussure)”. Box 25. Unchanged
(Guimarães, 1971: 35) (Tachinidae).
71. Myioscotiptera cincta, p. 2 (%) (1894: 62, fig. 14). “Solco”. Box 25. Unchanged (Guimarães,
1971: 31) (Tachinidae).
72. Microphthalmum sordida, p. 3 (%) (1894: 63). “Mexico (Truqui), Toluca (Saussure,
Boucard)”. Box 25. Now a junior synonym of Microphthalma obsoleta (Wulp, 1890)
(Guimarães, 1971: 24) (Tachinidae).
73. Thelairodes basalis, p. 3 (&) (1894: 65). “Mexico, no locality (Boucard)”. Box 25. Now a
junior synonym of Cholomyia inaequipes Bigot, 1884 (Guimarães, 1971: 106) (Tachinidae).
74. Chaetona cruenta, p. 3 (&) (1894: 65). “Mexico, no locality (Boucard)”. Box 25. Unchanged
(Guimarães, 1971: 105) (Tachinidae).
75. Aporia elegans, p. 3 (%) (1894: 66, fig. 15). “Tuxpango (Sumichrast)”. Box 25. Now in
Paraporia (Guimarães, 1971: 104) (Tachinidae).
76. Calliphora praepes, p. 3 (%) (1895:1). “Mexico (Saussure)”. Box 26. An unplaced species
of Calliphoridae (James, 1970: 16) (Calliphoridae).
77. Lucilia praescia, p. 4 (%) (1894: 3). “Tampico (Saussure)”. Box?. Now a junior synonym
of Phaenicia purpurescens (Walker, 1837) (James, 1970: 11) (Calliphoridae).
78. Lucilia quieta, p. 4 (&) (1894: 4). “Mexico, no locality”. Box 26. An unplaced species of
Calliphoridae (James, 1970: 16) (Calliphoridae).
79. Mesembrinella bicolor, p. 4 (%) (1895: 11, figs. 1, 1bis). “Chinantla, Oaxaca (Sumichrast)”.
Box 26. Now a synonym (and homonym) of Mesembrinella bicolor (Fabricius, 1805)
(James, 1970: 4) (Calliphoridae).
80. Hyadesimyia grisea, p. 5 (&) (1895: 8, fig. 2). “Mexico, Oaxaca (Sallé)”. Box 26. Now a
junior synonym of Synthesiomyia nudiseta (Wulp, 1883) (Pont, 1972: 12) (Muscidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
142
81. Graphomyia mexicana, p. 5 (?sex) (1895: 9 (%)). “Tuxpango (Sumichrast, Boucard)”. Box
26. Unchanged but in Graphomya (Pont, 1972: 28) (Muscidae).
82. Cyrtoneurina parilis, p. 6 (?sex) (1895: 14 (%, &)). “Oaxaca (Sumichrast)”. Box 27. Now in
Neomuscina (Pont, 1972: 50) (Muscidae).
83. Cyrtoneurina vecta, p. 6 (?sex) (1895: 14, fig. 4 (%)). “Tuxpango (Sumichrast)”. Box 27.
Now in Neomuscina (Pont, 1972: 51) (Muscidae).
84. Cyrtoneurina inuber, p. 6 (%) (1895: 15). “Solco (Sumichrast)”. Box 27. Unchanged (Pont,
1972: 53) (Muscidae).
85. Cyrtoneurina gluta, p. 6 (%, &) (1895: 15). “Tuxpango (Sumichrast)”. Box 27. Unchanged
(Pont, 1972: 53) (Muscidae).
86. Cyrtoneurina pellex, p. 6 (&) (1895: 16). “Tuxpango (Sumichrast)”. Box 27. Now a junior
synonym of Cyrtoneurina inuber Giglio-Tos (Pont, 1972: 53) (Muscidae).
87. Cyrtoneurina uber, p. 7 (%) (1895: 17). “Tuxpango (Sumichrast)”. Box 27. Unchanged
(Pont, 1972: 54) (Muscidae).
88. Leucomelina garrula, p. 7 (%) (1895: 18). “Tuxpango (Sumichrast)”. Box 27. Now in
Limnophora (Pont, 1972: 28) (Muscidae).
89. Leucomelina corvina, p. 7 (?sex) (1895: 19 (%, &)). “Mexico (sallé), Solco (Sumichrast)”.
Box 27. Now in Limnophora (Pont, 1972: 28) (Muscidae).
90. Leucomelina strigata, p. 7 (&) (1895: 19). “Tuxpango (Sumichrast)”. Box 27. Now in
Limnophora (Pont, 1972: 29) (Muscidae).
91. Hyetodesia mulcata, p. 7 (%) (1895: 20, fig. 5). “Mexico, no locality (Sumichrast)”. Box 27.
Now in Helina (Pont, 1972: 21) (Muscidae).
92. Hyetodesia parsura, p. 8 (%) (1895: 20). “Mexico, no locality (Truqui)”. Box 27. Now in
Helina (Pont, 1972: 22) (Muscidae).
93. Hyetodesia abacta, p. 8 (%) (1895: 21). “Mexico, no locality (Boucard)”. Box 27. Now a
junior synonym of Helina laetifica (Robineau-Desvoidy, 1830) (Pont, 1972: 21) (Muscidae).
94. Hyetodesia abdicta, p. 8 (&) (1895: 21). “Mexico, no locality (Craveri)”. Box 27. Now a
junior synonym of Helina laetifica (Robineau-Desvoidy, 1830) (Pont, 1972: 21) (Muscidae).
95. Hyetodesia insons, p. 8 (%) (1895:21). “Mexico, no locality (Boucard)”. Box 27. Now in
Mydaea (Pont, 1972: 25) (Muscidae).
96. Spilogaster refusa, p. 8 (%, &) (1895: 22). “Tuxpango (Sumichrast)”. Box 27. Now in Helina
(Pont, 1972: 22) (Muscidae).
97. Spilogaster abdita, p. 8 (&) (1895: 23). “Mexico, no locality (Boucard)”. Box 27. Now in
Bigotomyia (Pont, 1972: 18) (Muscidae).
98. Spilogaster etesia, p. 8 (&) (1895: 23). “Tuxpango (Sumichrast)”. Box 27. Now in Helina
(Pont, 1972: 20) (Muscidae).
99. Spilogaster meracula, p. 9 (&) (!895: 24). “Mexico, no locality (Boucard)”. Box 27. Now in
Pegoplata (Pont, 1974: 9) (Anthomyiidae).
100. Spilogaster pansa, p. 9 (&) (1895: 24, fig. 3). “Mexico, no locality (Sumichrast)”. Box 27.
Now in Mydaea (Pont, 1972: 25) (Muscidae).
101. Spilogaster sera, p. 9 (%) (1895: 25). “Mexico (Truqui)”. Box 27. Now in Helina (Pont,
1972: 23) (Muscidae).
Acta Zool. Mex. (n.s.) 84 (2001)
143
102. Spilogaster scabra, p. 9 (%, &) (1895: 25). “Mexico (Craveri)”. Box 27. Now in
Dendrophaonia (Pont, 1972: 13) (Muscidae).
103. Brachiophyra effrons, p. 9 (%, &) (1895: 26, fig. 6). “Tuxpango (Sumichrast)”. Box 27. Now
a junior synonym of Limnophora normata Bigot, 1885 (Pont, 1972: 29) (Muscidae).
104. Lasiops mexicana, p. 10 (?sex) (1895: 28 (%, &)). “Orizaba (Sumichrast, Boucard)”. Box
27. Now a junior synonym of Calythea crenata (Bigot, 1885) (Pont, 1974: 2)
(Anthomyiidae).
105. Anthomyia trifilis, p. 10 (?sex) (1895: 28 (%, &)). “Coscom(atepec) (Sumichrast)”. Box 27.
Now a junior synonym of Delia platura (Meigen, 1826) (Pont, 1974: 4) (Anthomyiidae).
106. Chortophila stlemba, p. 10 (&) (1895: 29). “Mexico (Craveri)”. Box 27. Now in Pegomya
(Pont, 1974: 9) (Anthomyiidae).
107. Hylemyia levipes, p. 10 (&) (1895: 30). “Mexico, no locality (Boucard)”. Box 27. An
unplaced Anthomyiidae (Pont, 1974: 10) (Anthomyiidae).
108. Hylemyia abrepta, p. 10 (&) (1895: 30). “Mexico, no locality”. Box 27. An unplaced
Anthomyiidae (Pont, 1974: 10) (Anthomyiidae).
109. Hylemyia rhodina, p. 10 (&) (1895: 31). “Tuxpango (Sumichrast)”. Box 27. Now in
Pegoplata (Pont, 1974: 9) (Anthomyiidae).
110. Coenosia pacifera, p. 11 (&) (1895: 31). “Txpango (Sumichrast)”. Box 28. Now in
Bithoracochaeta (Pont, 1972: 37) (Muscidae).
111. Coenosia vitilis, p. 11 (&) (1895: 32 (%, &)). “Tuxpango (Sumichrast)”. Box 28. Now in
Neodexiopsis (Pont, 1972: 48) (Muscidae).
112. Coenosia uvens, p. 11 (&) (1895: 32 (%, &)). “Mexico, no locality (Boucard)”. Box 28. A
junior synonym of Bithoracochaeta leucoprocta (Wiedemann, 1830) (Pont, 1972: 37)
(Muscidae).
1893b (Also in this paper Giglio-Tos did not cite the localities, which were only given in his
1894 and 1895 papers):
113. Gymnomma novum, p. 1 (&) (1894: 12, fig. 2). “Mexico (Sumichrast)”. Box 20.
Unchanged (Guimarães, 1971: 67) (Tachinidae).
114. Nemochaeta incerta, p. 2 (%) (1894: 14, fig. 3). “Oaxaca (Sumichrast)”. Box 21 (under
Tachinodes). Now Archytas incertus (Giglio-Tos) (Guimarães, 1971: 50) (Tachinidae).
115. Nemochaeta dubia, p. 2 (%) (1894: 15, fig. 8). “Mexico, no locality”. Box 21 (under
Tachinodes). Now Archytas dubius (Giglio-Tos) (Guimarães, 1971: 49) (Tachinidae).
116. Nemochaeta crucia, p. 2 (?sex) (1894: 16 (%, &)). “Mexico (Truqui), Tuxpango
(Sumichrast), Huastec”. Box 21 (under Tachinodes). Now Archytas crucius (Giglio-Tos)
(Guimarães, 1971: 49) (Tachinidae).
117. Nemochaeta pernox, p. 2 (?sex) (1894: 16) “Mexico (Boucard), Orizaba (Sumichrast)”.
Box 21 (under Tachinodes). Now in Archytas (Guimarães, 1971: 51) (Tachinidae).
118. Nemochaeta jurinoides, p. 2 (%) (1894: 18, fig. 5). “Oaxaca (Sallé)”. Box 21 (under
Tachinodes). Now an unrecognized species of Archytas (Guimarães, 1971: 50)
(Tachinidae).
119. Nemochaeta? aberrans, p. 2 (&) (1894: 18, fig. 9). “Meztillan”. Box 21 (under Tachinodes).
Now in Archytas (Guimarães, 1971: 48) (Tachinidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
144
120. Dejeania aurea, p. 3 (%) (1894: 20). “Solco (Sumichrast)”. Box?. Now in Adejeania
(Guimarães, 1971: 46) (Tachinidae).
121. Saundersia aurea, p. 3 (?sex) (1894: 21 (%, ?&). “Mexico (Craveri), Angang(ueo)”. Box
21. Now Epalpus aureus (Giglio-Tos) (Guimarães, 1971: 64) (Tachinidae).
122. Saundersia picea, p. 3 (?sex) (1894: 25, fig. 10 (%, &)). “Mexico (Sumichrast)”. Box 20.
Now Epalpus piceus (Giglio-Tos) (Guimarães, 1971: 65) (Tachinidae).
123. Belvosia bella, p. 3 (&) (1894: 30, figs. 6, 6a). “Mexico, no locality”. Box 22. Unchanged
(Guimarães, 1971: 181) (Tachinidae).
124. Chaetogena carbonaria, p. 4 (%) (1894: 31, fig. 19). “Orizaba (Sumichrast)”. Box 23. An
unplaced species of Dexiinae (Guimarães, 1971: 122) (Tachinidae).
125. Chaetogena cincta, p. 4 (?sex) (1894: 32 (%, &)). “Orizaba (Sumichrast)”. Box 23. An
unplaced Dexiinae (Guimarães, 1971: 122) (Tachinidae).
126. Chaetogena gracilis, p. 4 (&) (1894: 33, fig. 6). “Mexico, no locality (Boucard)”. Box 23.
An unplaced Dexiinae (Guimarães, 1971: 122) (Tachinidae).
127. Acroglossa tesselleta, p. 5 (&) (1894: 35). “Oaxaca (Sumichrast)”. Box 23. Now in
Chaetogaedia (Guimarães, 1971: 178) (Tachinidae).
128. Plagia mexicana, p. 5 (&) (1894: 42, fig. 13). “Mexico, no locality (Boucard)”. Box 23. Now
a junior synonym of Voria ruralis (Fallén, 1810) (Guimarães, 1971: 93) (Tachinidae).
129. Plagia dicta, p. 5 (&) (1894: 42). “Mexico, no locality”. Box 23. Now in Voria (Guimarães,
1971: 93) (Tachinidae).
130. Tricholyga gracilens, p. 5 (%) (1894: 39, fig. 16). “Mexico, no locality (Boucard)”. Box 23.
An unplaced Goniinae (Guimarães, 1971: 215) (Tachinidae).
131. Tricholyga insita, p. 5 (%) (1894: 39). “Mexico, no locality (Boucard)”. Box 23. An
unplaced Goniinae (Guimarães, 1971: 215) (Tachinidae).
132. Blepharipoda mexicana, p. 6 (&) (1894: 33, fig. 18). “Tehuacan”. Box 23. Now in Drino
(Guimarães, 1971: 189) (Tachinidae).
133. Cyrtophloeba horrida, p. 6 (%) (1894: 40). “Mexico, no locality (Sumichrast)”. Box 23. Now
in Eucyrtophloeba (Guimarães, 1971: 90) (Tachinidae).
134. Masicera sesquiplex, p. 6 (&) (1894: 43). “Mexico, no locality (Boucard)”. Box 23. An
unrecognized Tachinidae (Guimarães, 1971: 216) (Tachinidae).
135. Masicera usta, p. 6 (&) (1894: 44). “Mexico (Boucard)”. Box 23. An unrecognized
Tachinidae (Guimarães, 1971: 216) (Tachinidae).
136. Masicera glauca, p. 6 (&) (1894: 46). “Mexico, no locality (Boucard)”. Box 23. An
unrecognized Tachinidae (Guimarães, 1971: 216) (Tachinidae).
137. Degeeria mexicana, p. 7 (%) (1894: 46). “Orizaba (Sumichrast)”. Box 23. An unplaced
Blondeliini (Guimarães, 1971: 152) (Tachinidae).
138. Degeeria dicax, p. 7 (%) (1894: 49). “Mexico, no locality (Boucard)”. Box 23. An unplaced
Blondeliini (Guimarães, 1971: 152) (Tachinidae).
139. Degeeria insecta, p. 7 (?&) (1894: 48). “Mexico, no locality (Boucard). Box 23. An
unplaced Blondeliini (Guimarães, 1971: 152) (Tachinidae).
140. Deggeria cruralis, p. 7 (&) (1894: 49). “Mexico, no locality (Sumichrast)”. Box 23. An
unplaced Blondeliini (Guimarães, 1971: 152) (Tachinidae).
141. Clistomorpha ochracea, p. 7 (&) (1894: 53). “Mexico (Sumichrast)”. Box 23. Without
additional references (Tachinidae).
Acta Zool. Mex. (n.s.) 84 (2001)
145
142. Scatophaga reses, p. 7 (?sex) (1895: 34 (%, &)). “Mexico (Truqui)”. Box 28. Unchanged
(Albuquerque, 1984: 3) (Scatophagidae).
143. Scatophaga coenosa, p. 8 (1895: 34 (&)). “Popocatepetl, 3800 m”. Box 28. Without
references (Scatophagidae).
144. Tauromyia pachyneura, p. 8 (&) (1895: 37, figs. 7-8). “Mexico (Sallé)”. Box 28. No more
information.
145. Helomyza iniens, p. 8 )?sex) (1895: 35, figs. 9-10 (%, &). “Mexico, no locality
(Sumichrast)”. Box 28 Now in Suillia (Gill, 1968: 7) (Heleomyzidae).
146. Sepedon praemiosus, p. 8 (%) (1895: 38). “Morelia (Saussure)”. Box 28. Unchanged
(Knutson et al., 1976: 10) (Sciomyzidae).
147. Sapromyza stata, p. 9 (&) (1895: 47). “Orizaba (Sumichrast)”. Box 28. (Lauxaniidae).
148. Sapromyza plagosa, p. 9 (?sex) (1895: 47, as synon. of geminata Fabricius). “Mexico”.
Box 28. (Lauxaniidae).
149. Sapromyza sonax, p. 9 (?sex) (1895: 48). “Tuxpango (Sumichrast)”. Box 28.
(Lauxaniidae).
150. Sapromyza vinnula, p. 9 (?sex) (1895: 48). “Tuxpango (Sumichrast)”. Box 28.
(Lauxaniidae).
151. Sapromyza urina, p. 9 (?sex) (1895: 50, as Physogenua). “Mexico, Tuxpango
(Sumichrast)”. Box 28. (Lauxaniidae).
152. Sapromyza innuba, p. 9 (?sex) (1895: 49). “Mexico, no locality (Boucard)”. Box 28.
(Lauxaniidae).
153. Euaresta audax, p. 9 (&) (1895: 55, fig. 22). “Tuxpango (Sumichrast)”. Box 29. A junior
synonym of Dyseuaresta mexicana (Wiedemann, 1830) (Foote, 1967: 23) (Tephritidae).
154. Blepharoneura regina, p. 9 (%) (1895: 56, fig. 16). “Mexico, no locality”. Box 28.
Unchanged (Foote, 1967: 18) (Tephritidae).
155. Blepharoneura saga, p. 10 (%) (1895: 56, fig. 19). “Cordova (sic) (Saussure)”. Box 28. A
junior synonym of Blepharoneura poecilosoma (Schiner, 1868) (Foote, 1967: 18)
(Tephritidae).
156. Blepharoneura io, p. 10 (%) (1895: 57, fig. 18). “Tuxpango (Sumichrast)”. Box 29.
Unchanged (Foote, 1967: 18) (Tephritidae).
157. Blepharoneura diva, p. 10 (&) (1895: 57, fig. 17). “Tuxpango (Sumichrast)”. Box 29.
Unchanged (Foote, 1967: 18) (Tephritidae).
158. Blepharoneura splendida, p. 10 (%) (1895: 58, fig. 20). “Mexico, no locality (Boucard)”.
Box 29. Now in Hexachaeta (Foote, 1967: 28) (Tephritidae).
159. Acrotoxa mucida, p. 10 (%) (1895: 58, fig. 21). “Mexico, no locality (Boucard)”. Box 29.
Now in Polionota (Foote, 1967: 36) (Tephritidae).
160. Ostracocoelia mirabilis, p. 11 (&) (1895: 44, fig. 13). “Tampico (Saussure)”. Box 29.
Unchanged (Foote, 1967: 32) (Tephritidae).
161. Chaetocoelia palans, p. 11 (%) (1895: 51, fig. 14). “Orizaba (Sumichrast)”. Box 29.
Without additional references.
162. Molynocoelia lutea, p. 11 (%) (1895: 60, fig. 23). “Tuxpango (Sumichrast)”. Box 29.
Unchanged (Foote, 1967: 31) (Tephritidae).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
146
163. Platystoma mexicanum, p. 12 (%) (1895: 46, fig. 24). “Tuxpango (Sumichrast)”. Box 29.
Now a synonym of Automola atomaria (Wiedemann, 1830) ( Steyskal, 1968e: 2)
(Richardiidae).
164. Paragorgopis maculata, p. 12 (&) (1895: 41, fig. 12). “Tuxpango (Sumichrast)”. Box 29.
Unchanged (Steyskal, 1968b: 7) (Otitidae).
165. Oedopa elegans, p. 12 (&) (1895: 42). “Oaxaca (Sallé)”. Box 29. Unchanged (Steyskal,
1968b: 20) (Otitidae).
166. Rhinotora diversa, p. 13 (%) (1895: 43). “Tuxpango (Sumichrast)”. Box 29. Now in
Neorhinotora (Papavero, 1967c: 2) (Rhinotoridae).
167. Bricinniella cyanea, p. 13 (?%) (1895: 45). “Cuernavaca (Boucard)”. Box 29. Now in
Senopterina (Steyskal, 1968f: 2) (Platystomatidae).
168. Chyliza enthea, p. 13 (&) (1895: 64). “Tuxpango (Sumichrast)”. Box 29. Unchanged
(Prado, 1975a: 2) (Psilidae).
169. Tanypeza mexicana, p. 14 (?sex) (1895: 61 (%, &)). “Orizaba; Tuxpango (Sumichrast)”.
Box 29. Now a junior synonym of Neotanypeza (Neotanypeza) ornatipes (Bigot, 1886)
(Steyskal, 1967: 1) (Tanypezidae).
170. Calobata plectilis, p. 14 (&) (1895: 62). “Mexico, no locality”. Now in Rainieria (Rainieria)
(Steyskal, 1968a: 14) (Micropezidae).
171. Drosophila excita, p. 14 (&) (1895: 66). “Mexico, no locality (Boucard)”. Box 29. Not
included in Wheeler´s 1970 catalogue (Drosophilidae).
172. Chlorops unicornis, p. 14 (?sex) (1895: 65). “Coscom(atepec) (Sumichrast); Puebla
(Saussure)”. Box 30. Unchanged (Sabrosky & Paganelli, 1984: 35) (Chloropidae).
1893c:
173. Sphaerophoria willistoni, p. 31, pl. II, figs. 6, 6a (%, &). “Mexico, Orizaba (Sumichrast)”.
Box 17. Now in Allograpta (Thompson et al., 1976: 38) (Syrphidae).
174. Sphaerophoria forreri, p. 32 (%). “Mexico, Ciudad in Durango; Solco; Cuernavaca. Box
17. Now in Allograpta (Thompson et al., 1976: 35 (Syrphidae).
175. Mesogramma bidentatum, p. 49, pl. II, fig. 12 (%, &). “Mexico: Acahuizotla, Chilpancingo,
Tepetlapa, Medellin near Veracruz, Teapa in Tabasco, Orizaba (Sumichrast), Tampico”.
Box 18. Now in Toxomerus (Thompson et al., 1976: 48) (Syrphidae).
1895:
176. Pterocalla bella, p. 39, fig. 11 (&). “Orizaba (Sumichrast, Boucard)”. Box 29. Unchanged
(Steyskal, 1968b: 8) (Otitidae).
177. Chaetocoelia vergens, p. 51, fig. 15 (%). Tuxpango (Sumichrast)”. Box 29. Without
additional references.
15.
T
HE SPECIES DESCRIBED BY
E
WALD
R
ÜBSAAMEN
Ewald Rübsaamen was born on May 20, 1857 in Haardt (now Weidenau,
Province of Siegen), on the margins of the Sieg, as the son of Frans and Mathilde
Acta Zool. Mex. (n.s.) 84 (2001)
147
Rübsaamen. His basic studies were made at the public school in Wiedenau and
later at the Royal Gymnasium of Siegen, until the end of the “Ober Sekunda”. The
father, owner of a factory of instruments for mountain engeneering and metallurgy,
wanted him to study metallurgy, and for that reason, in 1875, Rübsaamen entered
the Superior Technical School in Karlsruhe, to study mathematics. However, after
4 semesters, he gave up the idea and, with his parents´ consent, decided to
become professor of drawing, studying for that purpose at Hilchenbach (Siegen).
In 1878, he became a teacher in a public school in Steinhauserberg, in Schwen,
and during the fall of the same year he was the main teacher of that school. In the
spring of 1879 he was granted the directorship of a private school in Siegen.
He learned by himself botany and zoology and became especially interested in
plant-galls, very little known at that time, and in a special way in the dipterous family
Cecidomyiidae. Upon his father´s death, he and his family moved to Berlin, so he
could finish his botanical studies. His mother stimulated him to complete those
studies.
In Berlin he found a job at the “Pfanzenphysiologische Institut der
Landwirtschaftlichen Hochschule”. From 1892 to 1893 he frequented the “Königlich
Kunstschule”, and after two years passed examination to graduate. Natural history
enthusmed him so much that he could not think of becoming a teacher any more,
and he decided to become a researcher. He found a position as a volunteer in the
Berliner Museum, with Professor Möbius. Later on, as a research assistant, in the
section of entomology, he received materials collected in several regions around the
world. In 1884 he started the preparation of several water-colors depicting the plant
galls and their agents. A collection of 100 water-colors was exposed during the
International Exposition of Gardens in Hamburg, in 1897, and he received the Great
Golden Medal from the Government of Hamburg. He also received a Golden Medal
from the “Vereins zur Föderung des Gartenbaues in den Kgl. preussischen Staates”,
by the occasion of the 75
th
anniversary and jubilaeum of that Society.
From 1897 on, he occupied several important positions in the Phylloxera
commission. He published some interesting papers on zoocecidiae, some with
beautiful colored plates.
Towards the end of his life he suffered much from heart and lung diseases,
remaining sick for many years; he also suffered from dropsy. In 1912, he received
the title of Professor, through the Ministry of Agriculture, and was elected in 1917
a member of the Leopoldinischen-Carolinischen Akademie der Wissenschaften in
Halle.
Rübsaamen died on March 17, 1919, in Metternich bei Koblenz (Schaffnit, 1927;
Papavero, 1973: 327-328).
Papavero & Ibañez-Bernal: History of Mexican Dipterology
148
Rübsaamen published only seven papers dealing with Neotropical Cecidomyiidae
and Sciaridae (1892, 1894, 1895, 1905, 1908a, 1908b, 1916). Only two Mexican
species, both Sciaridae, were described by him:
1. Sciara nigra, 1884: 26, pl. 1, fig. 7, pl. 3, figs. 12. 19, 22. “Mexico, Oaxaca”. Types: Berliner
Museum. Now in Odontosciara (Amorim, 1992: 63) (Sciaridae).
2. Sciara striata, 1894: 37, pl. 2, fig. 12, pl. 3, fig. 4. “Mexico, Oaxaca”. Type: Berliner
Museum. Now in Pseudosciara (Amorim, 1992: 65) (Sciaridae).
ACKNOWLEDGMENTS
This work was partially supported by Department of Entomology, Instituto de Ecología, A.
C. (902-02).
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__________. 1859. Dipterorum aliquot nova genera. Revue Mag. Zool. (2) 11: 305-315, pl. 11.
__________. 1875a. Diptères nouveaux ou peu connus. 4e. partie. V. Asilides exotiques
nouveaux. Ann. Soc. Entomol. France, 5(5): 237-248.
__________. 1875b. Diptères nouveaux ou peu connus. 5e. partie. VI. Espèces nouvelles des
genres Sphixea (Rondani) et Volucella (auctorum). Ann. Soc. Entomol. France, 5(5): 469-
482.
__________. 1875c. Diptères nouveaux ou peu connus. 5e. partie. VII. Espèces nouvelles
du genre Cyphomyia. Ann. Soc. Entomol. France, 5(5): 483-488.
__________. 1876a. Diptères nouveaux ou peu connus. 6e. partie. VIII. Curie des Phasides
(Phasidae, mihi). Genres Trichopoda (Macq.) et Bogosia (Rond.). Ann. Soc. Entomol.
France, (5) 6: 389-400.
__________. 1876b. Diagnose d´un nouveau genre de Diptères: Pharanthrax). Ann. Soc.
Entomol. France, (5) 6 (Bull.): lxvi.
Papavero & Ibañez-Bernal: History of Mexican Dipterology
150
__________. 1876c. (Diagnoses de quatre Diptères exotiques nouveaux: Ommatius et
Emphysomera). Ann. Soc. Entomol. France, (5) 6 (Bull.): lxxxv-lxxxvi.
__________. 1877a. Diptères nouveaux ou peu connus. 7e. partie. IX. Genre Somomyia
(Rondani), Lucilia (Rob.-Desv.), Calliphora, Phormia, Chrysomyia (id.). Ann. Soc. Entomol.
France,(5)7: 35-48.
__________. 1877b. Diptères nouveaux ou peu connus. 8e. partie. X. Genre Somomyia
(Rondani), Calliphora, Melinda, Mufetia, Lucilia, Chrysomyia (alias Microchrysa Rond.)
Robineau-Desvoidy. Ann. Soc. Entomol. France, (5) 7: 243-259.
__________. 1877c. Genus novum Ortalidarum (Macq. Schin. Et auctor) vel Tanipezidarum
(Rond.) Vel Trypetidarum (J. Bigot). Ann. Soc. Entomol. France, (5) 7: xxvi-xxvii.
__________. 1878a. Diptères nouveaux ou peu connus. 9e. partie. XII. Genus Phumosia,
Cosmina, Ochromyia et Curtonevra. Ann. Soc. Entomol. France,(5)8: 31-40.
__________. 1878b. Diptères nouveaux ou peu connus. 9e. partie. XIII. Genres Ocyptera
(Latr.), Ocypterula, Exogaster (Rond.). Ann. Soc. Entomol. France,(5)8: 40-47.
__________. 1878c. Diptères nouveaux ou peu connus. 9e. partie. XIV. Notes et mélanges.
Ann. Soc. Entomol. France, (5) 8: 48.
__________. 1878d. Diptères nouveaux ou peu connus. 10e. partie. XV. Tribu des Asilidi,
curies des Laphridae et Dasypogonidae. Ann. Soc. Entomol. France,(5)8: 213-240, 401-
446.
__________. 1879a. Diptères nouveaux ou peu connus. 11e. partie. XVI. Curiae
Xylophagidarum et Stratiomydarum (Bigot). Ann. Soc. Entomol. France,(5)9: 183-234.
__________. 1879b. (Diagnoses de trois genres nouveaux de Diptères). Ann. Soc. Entomol.
France,(5) 9(Bull.): lxvii-lxviii.
__________. 1880a. Diptères nouveaux ou peu connus. 12e. partie. XVIII. Genres Plagiocera
(Macq.), Formosia (Guérin) et Rutilia (Rob.-Desv.). Ann. Soc. Entomol. France, (5)10: 85-
89.
__________. 1880b. Diptères nouveaux ou peu connus. 14e. partie. XXI. Syrphidi (mihi)-
Genre Eristalis (Fabr.). Ann. Soc. Entomol. France,(5)10: 213-230.
__________. 1880c. (Description d´un genre nouveau et de nouvelles espèces
d´Acanthomeridae Weidemann (sic)). Ann. Soc. Entomol. France, (5) 10 (Bull.): v-vi.
__________. 1880d. (Diagnoses de trois genres nouveaux des Diptères). Ann. Soc. Entomol.
France, (5) 10(Bull.): xlvi-xlviii.
__________. 1880e. (Diagnose d´un nouveau genre de Diptères de la tribu Tachinidi). Ann.
Soc. Entomol. France,(5)10 (Bull.): liii.
__________. 1881. Diptères nouveaux ou peu connus. 16e. partie. XXIII. Tribus
Nemestrinidorum. Ann. Soc. Entomol. France, (6)1: 13-21.
__________. 1882a. Diptères nouveaux ou peu connus. 18e. partie. XXVIII. Acanthomeridae
(Wiedemann, Dipt. Exot.). Ann. Soc. Entomol. France,(6)1 (1881): 453-460.
__________. 1882b. Diptères nouveaux ou peu connus. 19e. partie. XXX. Genre Ctenostylum.
Ann. Soc. Entomol. France,(6)2: 21-22.
__________. 1882c. (Descriptions de quatre genres nouveaux de la trubu des Syrphides
(Syrphidae, auctorum), ainsi que celles de deux nouvelles spèces). Ann. Soc. Entomol.
France,(6)2 (Bull.): lxvii-lxviii.
Acta Zool. Mex. (n.s.) 84 (2001)
151
__________. 1882d. (Description de deux nouvelles espèces de diptères propres à la
Californie, dont l´une est le type d´un genre nouveau). Ann. Soc. Entomol. France,(6)2
(Bull.): xci-xcii.
__________. 1882e. (Diagnoses de genres et espèces inédits de Syrphides 1re. partie; 2e.
partie; 3e. partie). Ann. Soc. Entomol. France,(6)2 (Bull.): cxiv-cxv; cxx-cxxi; cxxviii-cxxix.
__________. 1883a. Diptères nouveaux ou peu connus. 20e. partie. XXXI. Genres Volucella
(Geoffr., Hist. des Insectes, 1764) et Phalachromyia (Rondani, Esame d. var. spec.
d´Insetti Ditteri Brasiliani, Torino, 1848). Ann. Soc. Entomol. France,(6)3: 61-88.
__________. 1883b. Diptères nouveaux ou peu connus. 21e. partie. XXXII. Syrphidi (1e.
partie). Ann. Soc. Entomol. France, (6)3: 221-258.
__________. 1883c. Diptères nouveaux ou peu connus. 22e. partie. XXXII. Syrphidi (2e.
partie). Espèces nouvelles, nº I. Ann. Soc. Entomol. France, (6) 3: 315-356.
__________. 1883d. Diptères nouveaux ou peu connus. 23e. partie. XXXII. Syrphidi (2e.
partie). Espèces nouvelles, nº II. Ann. Soc. Entomol. France, (6)3: 535-560.
__________. 1883e. (Description d’un nouveau genre de diptères de la tribu des Syrphidaes).
Ann. Soc. ent. France (6) 3(Bull.): xx-xxi.
__________. 1883f. (Diagnose d’un nouveau genre et d’une nouvelle espèce de diptères de
la tribu des Antomyzides /sic/). Ann. Soc. ent. France (6) 3(Bull.): xxx.
__________. 1884a. Diptères nouveaux ou peu connus. 4e. partie. XXXII. Syrphidi (2e.
partie). Espèces nouvelles, no. III. Ann. Soc. ent. France (6) 4: 73-116.
__________. 1884b. (Diagnoses d’un genre et d’une espèce de Diptères). Ann. Soc. ent.
France (6) 3(Bull.) (1883): cviii-cix.
__________. 1884c. (Descriptions d’un nouveau genre et d’une nouvelle espèce de diptères
de la famille des Ortalidae). Ann. Soc. ent. France (6) 4(Bull.): xxix.
__________. 1884d. (Descriptions d’un nouveau genre et d’une nouvelle espèce de diptères
de la famille des Dexidae). Ann. Soc. ent. France (6) 4(Bull.): xxxvii.
__________. 1885a. Diptères nouveaux ou peu connus. 25e. partie. XXXIII. Anthomyzides
nouvelles. Ann. Soc. ent. France (6) 4 (1884): 263-304.
__________. 1885b. Diptères nouveaux ou peu connus. 27e. partie. XXXV. Famille des
Anomalocerati (mihi) (Coriacea, Pupipara, Nycteribidae, auctor). Ann. Soc. ent. France (6)
5: 225-246.
__________. 1885c. (Diagnoses de deux genres nouveaux de diptèes du groupe des
Déxiaires). Ann. Soc. ent. France (6) 5(Bull.): xxv-xxvi.
__________. 1885d. (Diagnoses de deux genres nouveaux de diptères appartenant à la
famille des Ortalides). Ann. Soc. ent. France (6) 5(Bull.): clxv-clxvi.
__________. 1885e. (Description d’un nouveau genre de diptères). Ann. Soc. ent. France (6)
5(Bull.): clxxiii-clxxiv.
__________. 1885f. Diptères nouveaux ou peu connus. 28e.partie. XXXVI. Syrphidi. Addenda
au mémoire publié dans les Annales de la Société Entomologique de France (Années
1883-1884). Ann. Soc. ent. France (6) 5: 24-252.
__________. 1886a. Diptères nouveaux ou peu connus. 29e. partie. XXXVII, 1er. Essai d’une
classification synoptique du grupe des Tanypezidi (mihi) et descriptions de genres et
d’espèces inédits; 2e. (Suite). Ann. Soc. ent. France (6) 6: 287-302, 369-392.
Papavero & Ibañez-Bernal: History of Mexican Dipterology
152
__________. 1886b. (Diagnoses nouvelles d’un genre et d’une espèce de l’ordre des
Diptères). Ann. Soc. ent. France (6) 6 (Bull.): ciii-civ.
__________. 1887a. Diptères nouveaux ou peu connus. 31e. partie XXXIX. Descriptions de
nouvelles espèces de Stratiomydi et de Conopsidi. Ann. Soc. ent. France (6) 7: 20-46.
__________. 1887b. Diptères nouveaux ou peu connus. 32e. partie. XL. Descriptions de
nouvelles espèces de Myopidi. Ann. Soc. ent. France (6) 7: 203-208.
__________. 1887c. (Diagnoses de quelques espèces nouvelles de diptères). Ann. Soc. ent.
France (6) 7(Bull.): cxxxix-cxlii.
__________. 1887d. (Diagnoses abrégées de quelques diptères nouveaux, provenant de
l’Amerique du Nord). Ann. Soc. ent. France (6) 7(Bull.): clxxx-clxxxii.
__________. 1888a. Diptères, pp. 1-45, pls. 1-4, in: Ministère de la Marine et de l’Instruction
Publique /de France/, Mission scientifique du Cap Horn, 1882-1883, 6 (Zoologie), 2me.
Partie. Paris., ‘1891'.
__________. 1888b. Diptères nouveaux ou peu connus. 33e. partie. XLI. Tachinidae. Ann.
Soc. ent. France (6) 8: 77-101.
__________. 1888c. Diptères nouveaux ou peu connus. 34e. partie. XLII. Diagnoses de
nouvelles espèces. Ann. Soc. ent. France (6) 8: 253-270.
__________. 1888d. (Notes critiques sur les Diptères). Ann. Soc. ent. France (6) 8(Bull.): xxiv.
__________. 1888e. (Diagnoses d’espèces nouvelles de Dolichopodi). Ann. Soc. ent. France
(6) 8(Bull.): xxix-xxx.
__________. 1889a. Diptères nouveaux ou peu connus. 34e. partie. XLII. Empidi. Ann. Soc.
ent. France (6) 9: 111-134.
__________. 1889b. Diptères nouveaux ou peu connus. 35e. partie. XLIII. Cyrtidi, XLIV.
Therevidi. Ann. Soc. ent. France (6) 9: 313-320, 321-328.
__________. 1889c. (Novum genus dipterorum ex Trypetidis, genus Chetostomae (Rondani)
sat vicium). Ann. Soc. ent. France (6) 9(Bull.): xxix-xxx.
__________. 1889d. (Change of generic name). Ann. Soc. ent. France (6) 9(Bull.): xciii.
__________. 1889e. (Note on relationships of Archilestes and synonymy; Change of generic
name). Ann. Soc. ent. France (6) 9(Bull.): clxxxiii.
__________. 1890a. Diptères nouveaux ou peu connus. 46e. partie. XLV. Dolichopi. Essai
d’une classification générale. Ann. Soc. ent. France (6) 10: 261-296.
__________. 1890b. (Change of generic name). Wien. Ent. Ztg. 9:96.
__________. 1892a. Diptères nouveaux ou peu connus. 37e partie. XLVI. Bombylidi (mihi).
Ann. Soc. ent. France 61: 321-376.
__________. 1892b. Descriptions de diptères nouveaux. Tabanidi. Mém. Soc. zool. France 5:
602-691.
__________. 1892c. Nova genera dipterorum. Wien. Ent. Ztg. 11: 161-162.
__________. 1892d. Description d’une espèce nouvelle de diptères parasite de Costa Rica.
The Hummingbird 2(7): 49.
Bilimek, D. 1867. Fauna der Grotte Cacahuamilpa in Mexiko. Verh. Zool.-bot. Ges. Wien 17
(Abhandl.): 901-908.
Binford, L. C. 1989. A distributional survey of the birds of the Mexican State of Oaxaca.
(American Ornithological Union, Washington, D.C.). Ornithol. Mon., 43:1-418.
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Recibido: 27 de noviembre 2000
Aceptado: 21 de mayo 2001
Acta Zool. Mex. (n.s.) 84 (2001)
167
APPENDIX 1
Exotic species of Diptera treates by C. E. W. Wiedeman in his “Aussereuropäische
Zweiflügelige Insekten” (1828-1830) (Abbr. NA-North America; WI-West Indies; SA-South
America (mostly from Brazil).
Family Total
Genera number of NA WI SA Mexico
species
I. Fam. TIPULARIAE (Mücken)
1. Culex 26 5 1 11 2
2. Anopheles 541
3. Corethra 11
4. Chironomus 11 6 3
5. Tanypus 42 1
6. Macropeza 1
7. Lasioptera 11
8. Cecidomyia 22
9. Campylomyza 11
10. Psychoda 11
11. Erioptera 11
12. Limnobia 38 7 22
13. Ctenophora 52
14. Tipula 25 11 6
15. Megistocera 41
16. Polymera 22
17. Ptychoptera 22
18. Trichocera 11
19. Platyura 32 1
20. Sciophila 66
21. Leia 21 1
22. Mycetophila 33
23. Sciara 10 7 3
24. Simulia 11
25. Scatopse 11
26. Penthetria 2
27. Plecia 76
28. Dilophus 65 1
29. Bibio 10 9
30. Rhyphus 33
Total in the family 185 85 2 58 2
Papavero & Ibañez-Bernal: History of Mexican Dipterology
168
Appendix 1.
Family Total
Genera number of NA WI SA Mexico
species
II. Fam. XYLOPHAGI (Holzfliege)
1. Beris 32 1
2. Xylophagus 42 1
3. Coenomyia 11
Total in the family 8 5 0 1 1
III. Fam. TABANII (Bremsen)
1. Pangonia 37 1 14 3
2. Rhinomyza 2
3. Rhaphiorhynchus 11
4. Acanthomera 615
5. Silvius 41 21
6. Tabanus 137 21 28 1
7. Chrysops 27 7 12
8. Haematopota 11 5
Total in the family 225 30 1 67 5
IV. Fam. LEPTIDES (Schepfenfliegen)
1. Leptis 18 11 0 4 0
V. Fam. XYLOTOMAE
1. Thereva 18 5 2
2. Chiromyza 33
3. Ruppelia 1
Total in the family 22 5 0 5 0
VI. Fam. MIDASII (Midasfliegen)
1. Midas 23 2 1 11 2
VII. Fam BOMBYLIARII (Schwebfliegen)
1. Hirmoneura 11
2. Nemestrina 15 2
3. Lomatia 11
4. Anthrax 118 17 4 23 10
5. Tomomyza 1
6. Xestomyza 2
Acta Zool. Mex. (n.s.) 84 (2001)
169
Appendix 1.
Family Total
Genera number of NA WI SA Mexico
species
VII. Fam BOMBYLIARII (continued)
7. Apatomyza 1
8. Corsomyza 4
9. Lasia 11
10. Mulio 2
11. Bombylius 43 5 8
12. Amictus 2
13. Thlipsomyza 1
14. Phthiria 31
15. Geron 11
16. Cyllenia 2
17. Systropus 21
18. Toxophora 41
19. Dolichomyia 11
20. Usia 1
Total in the family 206 23 5 39 10
VIII. Fam. ASILICI (Räuberfliegen)
1. Dioctria 61 1
2. Dasypogon 84 8 1 32 1
3. Ceraturgus 11
4. Damalis 4
5. Ommatius 10 1 1
6. Asilus 122 18 2 54
7. Laphria 64 13 27 1
8. Leptogaster 71 2
Total in the family 298 43 4 116 2
IX. Fam. HYBOTINAE
1. Hybos 71 3
X. Fam. EMPIDIAE
1. Empis 12 1 2 1
2. Rhamphomyia 74 1
3. Drapetis 1
Total in the family 20 4 1 3 1
Papavero & Ibañez-Bernal: History of Mexican Dipterology
170
Appendix 1.
Family Total
Genera number of NA WI SA Mexico
species
XI. Fam. TACHYDROMIDES
1. Hemerodromia 11
2. Tachydromia 21
Total in the family 3 2 0 0 0
XII. Fam INFLATAE
1. Cyrtus 4
2. Acrocera 21 1
3. Philopota 11
Total in the family 7 1 0 2 0
XIII. Fam. STRATIOMYDAE
1. Hermetia 76
2. Sargus 28 4 1 17
3. Acrochaeta 11
4. Eudmeta 1
5. Nemotelus 21
6. Clitellaria 511
7. Acanthina 33
8. Platyna 1
9. Cyphomyia 871
10. Ptilocera 1
11. Oxycera 11
12. Stratiomys 28 7 12
Total in the family 86 12 3 47 1
XIV. Fam. SYRPHIDIAE
1. Ceratophya 22
2. Ceria 2
3. Microdon 11 2 7
4. Chrysotoxum 21
5. Paragus 4
6. Ascia 1
7. Psarus 22
8. Baccha 912
9. Xylota 11 5 3
Acta Zool. Mex. (n.s.) 84 (2001)
171
Appendix 1.
Family Total
Genera number of NA WI SA Mexico
species
XIV. Fam SYRPHIDIAE (continued)
10. Milesia 83 1
11. Pipiza 1
12. Eumerus 7
13. Rhingia 11
14. Chrysogaster 11
15. Syrphus 53 12 1 18
16. Platynochaetus 1
17. Merodon 61 1
18. Eristalis 64 6 2 18
19. Volucella 20 2 14 1
20. Graptomyza 4
Total in the family 210 33 7 66 1
XV. Fam. MEGACEPHALI (Grossköpfe)
1. Pipunculus 11
XVI. Fam. DOLICHOPODII (Dünnbeinfliegen)
1. Rhaphium 1
2. Diaphorus 1
3. Chrysetus 1
4. Psilopus 33 10 1 11
5. Porphyrops 1
6. Medeterus 1
7. Dolichopus 92 2
8. Scenopinus 1
Total in the family 48 12 1 13 0
XVII. Fam. CONOPSARIAE (Kopffliegen)
1. Conops 10 3 3
2. Zodion 32 1
3. Myopa 63
Total in the family 19 8 0 4 0
XVIII. Fam. STOMOXYDIAE
1. Stomoxys 12 1 7
2. Glossina 1
Total in the family 13 1 0 7 0
Papavero & Ibañez-Bernal: History of Mexican Dipterology
172
Appendix 1.
Family Total
Genera number of NA WI SA Mexico
species
XIX. Fam. OESTRACIDAE (Stumpfmaulfliegen)
1. Oestrus 21
2. Trypoderma 54
3. Colax 21
Total in the family 9 5 0 1 0
XX. Fam. MUSCIDIAE (Urfliegen)
1. Phasia 31
2. Ocyptera 61
3. Phania 11
4. Trichopoda 13 6 7
5. Trixa 1
6. Miltogramma 2
7. Tachina 117 4 7 45 1
8. Gonia 61
9. Musca 73 1 15
10. Mesembrina 11
11. Idia 12 1
12. Sarcophaga 31 1 5 17
13. Dexia 27 1 16
14. Anthomyia 36 1 1 17
15. Coenosia 13
16. Lispe 7
17. Cordylura 21
18. Ochtera 11
19. Scatophaga 52 1
20. Dryomyza 11
21. Sapromyza 16 1 1 7
22. Ortalis 16 2 2 7
23. Sepsis 71 1
24. Cephalia 33
25. Lauxania 16 1 13
26. Lonchaea 312
27. Trypeta 58 3 17 4
28. Dacus 20 6
29. Tetanops 11
30. Psila 1
31. Loxocera 11
Acta Zool. Mex. (n.s.) 84 (2001)
173
Appendix 1.
Family Total
Genera number of NA WI SA Mexico
species
XX. Fam. MUSCIDIAE (continued)
32. Tanypeza 33
33. Chyliza 41
34. Calobata 26 1 1 17 2
35. Micropeza 41 2
36. Nerius 85
37. Longina 11
38. Achias 32
39. Diopsis 81
40. Timia 1
41. Ulidia 21
42. Dictya 11
43. Platystoma 3
44. Ropalomera 44
45. Homalura 41
46. Sciomyza 43
47. Sepedon 5
48. Oscinis 2
49. Pyrgota 11
50. Tetanocera 55
51. Helomyza 41 1
52. Notiphila 11 2
53. Ephydra 42
54. Chlorops 11 1 6
55. Borborus 2
56. Stegana 11
57. Trineura 1
58. Celyphus 2
59. Hippobosca 4
60. Olfersia 31
61. Ornithomyia 94 1
62. Strebla 11
63. Melophaga 31
Total in the family 645 36 24 240 7
GRAND TOTAL 2053 319 48 688 32
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