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41F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
FILIPPO MARIA BUZZETTI & HENDRIK DEVRIESE
TETRIGIDAE FROM ECUADOR
(Orthoptera Caelifera Tetrigoidea)
ABSTRACT - BUZZETTI F. M. & DEVRIESE H., 2007 - Tetrigidae from Ecuador (Or-
thoptera Caelifera Tetrigoidea).
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B: 41-54.
Thirty-six taxa of Tetrigidae are known from Ecuador, twenty-nine were known
previously to this study, the presence of five of these is confirmed, seven are here
reported for the first time. No tetrigids are known from Galapagos Islands. Faunisti-
cal notes on Tetrigidae recently collected in Ecuador are given. All taxa are listed and
commented. Type locality of the species is reported when in Ecuador.
KEY WORDS - Fanistics, Amazonia, Andes, Coastal Plains, Galapagos Islands,
Macropronotal, Brachypronotal, Scaria maculata Giglio Tos 1898.
RIASSUNTO - BUZZETTI F. M. & DEVRIESE H., 2007 - Tetrigidae dallEcuador (Or-
thoptera Caelifera Tetrigoidea).
Trentasei taxa di Tetrigidae sono conosciuti per lEcuador, ventinove erano co-
nosciuti precedentemente al presente studio, la presenza di cinque di questi è confer-
mata, sette sono qui riportati per la prima volta. Non sono noti tetrigidi per le Isole
Galapagos. Sono fornite note faunistiche sui Tetrigidae recentemente raccolti in Ecua-
dor. Tutti i taxa sono elencati e commentati. La località tipica è citata quando in
Ecuador.
PAROLE CHIAVE - Faunistica, Amazzonia, Ande, Pianure Costiere, Isole Galapa-
gos, Macropronotale, Brachipronotale, Scaria maculata Giglio Tos 1898.
INTRODUCTION
Tetrigidae of Ecuador are only partially known, mostly on the basis
of the two main contributions on Orthopteroid fauna of Ecuador (GI-
GLIO TOS 1898, HEBARD 1924b). Other useful references are GÜNTHER
(1939) and the contributions by Lawrence Bruner. To date a work com-
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B42
prehensive of all the species and identification key on Ecuador is lacking,
so the present paper wonts to be a first step to such a goal.
Working on Neotropical tetrigids is particularly difficult due to the
scarcity of material in Museums and private collections. This appears
rather strange, being Tetrigidae a group of insects so common along
water streams and ponds, especially in Neotropics. Further difficulty
in approaching this group is represented by the literature, mostly scat-
tered, not revisional and limited to small groups of species, and espe-
cially not using a modern systematics approach. GÜNTHER (1939) pro-
vided an identification key for subfamily Metrodorinae, to which be-
long most of the South American species, but he examined only a frac-
tion of the types so that probably misinterpretation of some descrip-
tions was unavoidable. Subfamily Batrachideinae is revised by GRANT
(1955, 1956), except for the very large genus Tettigidea, for which the
revision was never published. Subfamily Cladonotinae is also revised
by GÜNTHER (1938), while Tetriginae were never been reviewed, the
most recent work by STEINMANN (1969) listing 20 species.
Resulting from this situation is that a revision of all Neotropical
genera of Tetrigidae is strongly needed.
The present work results from recent collecting expeditions since
2002 run by the first author. Identification of the species was made by
the second author. Furthermore the collection in Museo Regionale di
Scienze Naturali di Torino has been examined.
The material examined is now in the private collection of the first
author in Arzignano, Italy, abbreviated as F.M.B., and in the Institute
Royal des Sciences Naturelles de Belgique, Brussels, Belgium, abbre-
viated in ISNB.
Species collected during expeditions in Ecuador since 2002 are li-
sted, the synonymy comprehensive of all citations concerning Ecuador
is listed for each species plus the material examined and some notes.
SPECIES COLLECTED
subfam. Batrachideinae
gen. Batrachidea Serville 1839
Batrachidea inermis Hebard 1923
Batrachidea inermis Hebard 1923: 174.
Batrachidea inermis; Grant 1956: 79.
43F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
Material Examined: Esmeraldas prov., Montalvo, La Mayronga, 95
m., 0°5327.2 N 79°132.5 W, 14/17.XI.2004, lg F. M. B. & G. Ca-
rotti, 1 , 1 , coll. F. M. B.; idem, 1 , coll. ISNB; Esmeraldas prov., San
Lorenzo, S. Francisco de Bogotà, 9/10.XII.2004, lg G. Carotti, 1 , coll.
F. M. B.
The genus Batrachidea Serville includes five species all from South
America and was revised by GRANT (1956).
B. inermis Hebard was known for Colombia only and the speci-
mens here reported are the first record of the species for Ecuador whe-
re it is present in the Coastal Plains.
All the material examined is brachypronotal.
Batrachidea mucronata (Saint-Fargeau & Serville, 1825)
Tetrix mucronata Saint-Fargeau & Serville 1825, Encyclopédie méth., Hist. nat., In-
sectes, 7: 600.
Batrachidea mucronata; Bolivar 1887: 300.
Batrachidea mucronata; Giglio Tos 1898: 36.
Batrachidea notata Bruner 1906b: 616.
Tettigidea gracilicornis Bruner 1910: 128.
Batrachidea mucronata; Grant 1956: 70.
Material Examined: Esmeraldas prov., Montalvo, La Mayronga, 95
m., 0°5327.2 N 79°132.5 W, 14/17.XI.2004, lg F. M. B. & G. Ca-
rotti, 1 , coll. F. M. B.; Esmeraldas prov., San Lorenzo, S. Francisco
de Bogotà, 9/10.XII.2004, lg G. Carotti, 1 , coll. F. M. B.; idem, 1 ,
coll. ISNB; Napo prov., Yasuni Scientific Station, 219 m., 0°4026.6 S
76°2351.5 W, 22/26.XI.2004, lg F. M. B. & G. Carotti, 1 , coll. F.
M. B.; idem, 2/5.V.2002, lg F. M. B., 2 , 1 , coll F. M. B.; idem, 1 ,
coll. ISNB; Pichincha prov., Union del Toachi, trail to Otongachi, farm,
14.IV.02, lg F. M. B., 1 , coll. F. M. B.; Morona Santiago prov., Gua-
laquiza, 851 m., lg G. Carotti & P. Tirello, 1 , coll. F. M. B.; Sucum-
bios prov., El Reventador, San Rafael Falls, 1400 m., lg G. Osella, 1 ,
coll. F. M. B.
B. mucronata (Saint-Fargeau & Serville, 1825) Fig. 1, is widely di-
stributed in South America, being present in Colombia, Ecuador, Peru,
Bolivia and Brazil (GRANT 1956). The distribution of the species in Ecua-
dor covers Amazonian part, Andes and Coastal Plains.
This species differs only slightly from B. macella Grant 1956, so
that is not clear if these are separate species.
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B44
GRANT (1956) synonymized B. notata Bruner 1906 and Tettigidea
gracilicornis Bruner 1910 with B. mucronata.
The brachypronotal specimens collected are much bigger than
macropronotal specimens, a character typical of Batrachideinae.
gen. Paurotarsus Hancock 1900
Paurotarsus ruficornis (Walker 1871)
Tettix ruficornis Walker 1871: 817.
Paurotarsus amazonus Hancock 1900: 42.
Paurotarsus rugosus Bruner 1906a: 148.
Paurotarsus ruficornis; Hebard 1924a: 92.
Paurotarsus ruficornis; Grant 1955: 4.
Material Examined: Napo prov., Yasuni Scientific Station, 219 m.,
0°4026.6 S 76°2351.5 W, 22/26.XI.2004, lg F. M. B. & G. Carotti,
1 brachypterous , coll. F. M. B.
The small Central and South American genus Paurotarsus Hancock
includes two species: P. insolitus Rehn 1916, divided in three subspe-
cies, and P. ruficornis (Walker 1871).
Hebard (1924a) synonymized P. amazonus Hancock 1900 and P.
rugosus Bruner 1906 with P. ruficornis.
P. ruficornis is distributed in Bolivia, Brazil, Peru, Trinidad (GRANT
1955) and Amazonian Ecuador.
gen. Scaria Bolivar 1887
Scaria producta Hancock 1907
Scaria producta Hancock 1907: 70.
Scaria producta; Bruner 1910: 140.
Material Examined: Napo prov., Yasuni Scientific Station, 219 m.,
0°4026.6 S 76°2351.5 W, 22/26.XI.2004, lg F. M. B. & G. Carotti,
1, coll. F. M. B.
The genus Scaria Bolivar 1887 has not been revised since 1910 and
therefore is in need of revision. Eight species have been described with
range from Southern United States to South America.
45F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
The single macropronotal male collected is the first record for Ecua-
dor of this species that was known for Peru, Bolivia and Brazil. In Ecua-
dor the distribution of S. producta Hancok is Amazonian.
Fig. 1. Batrachidea mucronata (Saint-Fargeau & Serville, 1825), male from Esmeraldas
prov., Montalvo, La Mayronga (photo G. Carotti).
Fig. 2. Eomorphopus antennatus (Bolivar 1887), female from Napo prov., Yasuni Na-
tional Park, PUCE Estacion Cientifica (photo G. Carotti).
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B46
subfam. Metrodorinae
gen. Allotettix Hancock 1899
Allotettix americanus Hancock 1907
Allotettix americanus Hancock 1907: 234.
Allotettix americanus; Bruner 1910: 116.
Allotettix americanus; Hebard 1924b: 156.
Material Examined: Pichincha, La Union del Toachi, 900 m,
10.VIII.02, lg G. Osella, 1 , coll. F. M. B.
Within the genus Allotettix Hancock 1899, eight species are actual-
ly included, all from South America except for A. peruvianus Bolivar
1887 and A. cayennensis Bolivar 1887 which are present also in Costa
Rica and West Indies respectively. The first species is cited with some
doubts sub Paratettix peruvianus by GIGLIO TOS (1898) on the basis of
a single specimen collected in the valley of Santiago River in Ecuador.
More later HEBARD (1924b) ascribes to A. peruvianus a single female
from «Allpa-yacu, Rio Pastaza, Tunguragua, 3600 feet (ca 1080 me-
ters)» probably on the eastern slopes of the Andes.
Type locality of the A. americanus is Cachabi. In Ecuador the spe-
cies ranges from the western slopes of the Andes to the coastal plains.
Gen. Chiriquia Morse 1900
Chiriquia concinna (Bolivar 1887)
Metrodora concinna Bolivar 1887: 249.
Chiriquia concinna; Hancock 1907: 39.
Chiriquia concinna; Kirby 1910: 23.
Chiriquia concinna; Bruner 1910: 99.
Chiriquia concinna; Günther 1939: 228.
Material Examined: Esmeraldas, San Lorenzo, S. Francisco de Bo-
gotà, 9/10.XII.2004, lg. G. Carotti, 1 , coll. F. M. B.
The Central and South American genus Chiriquia Morse 1900 is
very easy to recognise among Metrodorinae, by the peculiar position of
the antennae that are placed very below the base of eyes.
Four of the six described species are from Ecuador: C. compressa
Günther 1939 with type locality Ecuador, Rosario, C. mirifica Hebard
47F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
1924 with type locality Ecuador, Rio Pescado, Azuay, 480 m., C. spina-
ta Günther 1939 with unspecified Ecuadorean type locality and C. con-
cinna (Bolivar). C. concinna is known in Ecuador for the Coastal plains
and for the Western slopes of the Andes.
gen. Crimisus Bolivar 1887
Crimisus bolivianus (Bruner 1913)
Allotettix bolivianus Bruner 1913: 425.
Crimisus bolivianus; Günther 1939: 242.
Material Examined: Napo, Yasuni Scientific Station, 0°40,407
S 76°23,883 W, 2/5.V.2002, lg F. M. B., 1 , coll. F. M. B.
The genus Crimisus Bolivar 1887 has the lateral lobes of pronotum
only slightly produced, so that it could be also attributed to subfamily
Tetriginae which has the lateral lobes of pronotum not produced. This
feature is shown clearly by the present species having the lateral lobes
of pronotum not extended from the body.
This species was known before only for Bolivia and this is the first
record for Ecuador.
Crimisus tibialis (Bruner, 1910)
Sclerotettix tibialis Bruner 1910: 106.
Crimisus tibialis; Günther 1939: 246.
Material Examined: Napo prov., Yasuni Scientific Station, 219 m.,
0°4026.6 S 76°2351.5 W, 22/26.XI.2004, lg F. M. B. & G. Carotti,
1, 1 , coll. F. M. B.; idem, 1 , coll. ISNB; idem, 0,6784° S 76, 40032° W,
2/5.V.2002, lg. F. M. B., 1 , coll. F. M. B.
Twelve species have been described in the genus Crimisus Bolivar
1887 and in Sclerotettix Bruner 1910, the latter synonymized by GÜN-
THER (1939) with Crimisus.
These specimens from Ecuador have very enlarged middle femora,
this a typical character of C. tibialis. Nevertheless the present species
resembles two species very related to it: C. steinbachi (Bruner 1913)
and C. variegatus (Bruner 1910). The first differs from tibialis in having
the middle femora slightly less expanded, the latter for the comparati-
vely smoother pronotum and legs.
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B48
Known before only for Brazil, the species is likely present all along
the Amazonian Basin.
gen. Eomorphopus Hancock 1900
Eomorphopus antennatus (Bolivar 1887)
Amorphopus antennatus Bolivar 1887: 241.
Eomorphopus antennatus; Hancock 1907: 38.
Eomorphopus antennatus; Günther 1939: 262.
Material Examined: Napo prov., Yasuni Scientific Station, 219 m.,
0°4026.6 S 76°2351.5 W, 22/26.XI.2004, lg F. M. B. & G. Carotti,
12 , 7 , coll. F. M. B.; idem, 1 , 1 , coll. ISNB; idem, 0°40,407
S 76°23.883 W, 2/5.V.2002, lg. F. M. B., 1 , coll. F. M. B.
The South American genus Eomorphopus Hancock 1900 includes
3 species and can be distinguished from the related genus Amorphopus
Serville 1838 by the not dilated forelegs.
E. antennatus (Fig. 2) was known before for Brazil, Trinidad (BRU-
NER 1910), Peru and Suriname (GÜNTHER 1939), the material here li-
sted is the first record of the species for Ecuador.
gen. Metrodora Bolivar 1887
Metrodora rana Bolivar 1887
Metrodora rana Bolivar 1887: 248.
Metrodora rana; Giglio Tos 1898: 35.
Metrodora rana; Hebard 1924b: 152.
Material Examined: Napo prov., Yasuni Scientific Station, 219 m.,
0°4026.6 S 76°2351.5 W, 22/26.XI.2004, lg F. M. B. & G. Carotti,
1, coll. F. M. B.
Fourteen species belong to the genus Metrodora Bolivar 1887 and
ten of them were revised by GÜNTHER (1939).
The present species is very closely related to M. acuta Günther 1939
which was described on a single male from Loja, Ecuador.
49F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
Gen. Scabrotettix Hancock 1907
Scabrotettix bolivianus bolivianus Hancock 1907
Scabrotettix bolivianus bolivianus Hancock 1907: 47.
Scabrotettix bolivianus; Bruner 1910: 114.
Material Examined: Pichincha, Union del Toachi, pastures after
Otongachi, 868 m, S 0°2012,5 W 78°5555,7, 8.III.2004, lg. Buzzet-
ti, Carotti & Tirello, 3 , 1 , coll. F. M. B.; idem, 1 , coll. ISNB;
idem, 900 m, 20.VIII.02, lg G. Osella, 1 , coll. F. M. B.; Cotopaxi,
Otonga Nat. Res., III.2004, lg. C. Tapia, 2 B&, coll F. M. B.; Las Pam-
pas, III.2004, 2 , coll. F. M. B.; Cotopaxi, S. F. Las Pampas, Otonga
N. R. scientific station, 1906 m, S 0°2510,1 W 79°011,8, 10.III.04,
lg. Buzzetti, Carotti & Tirello, 2 , 1 , coll. F. M. B.; idem, 1 , coll.
ISNB.
The five species contained in the genus Scabrotettix Hancock 1907
are all from South America, three of them known for Ecuador: S. ama-
zonus Bolivar 1887, S. scabrosus Hancock 1907 and the species here
reported. The species of this genus are not very well distinguished.
S. bolivianus is a very variable species with two macropronotal for-
ms: one with pronotum extending behind the hind femora, the other
with pronotum as long as the femora. Even the extension of the wings
beyond the pronotal hind tip, this character separating the two subspe-
cies of bolivianus, is very variable within the same populations of the
two subspecies.
Scabrotettix bolivianus extensus Hancock 1907
Scabrotettix bolivianus extensus Hancock 1907: 47.
Scabrotettix bolivianus extensus; Bruner 1910: 114.
Scabrotettix bolivianus extensus ?; Günther 1939: 254.
Material Examined: Pichincha, Union del Toachi, 805 m, S 0°19,036
W 78°57,303, 14.IV.02, lg F. M. B., 1 , coll. F. M. B.; Cotopaxi, Otonga
Nat. Res., ponds to Sentiero Nuevo, 1682 m, S 0°259,5 W 78°5936,5,
1, coll. F. M. B.; Pichincha, S. F. Las Pampas, Otonga N. R., Sendero
Nuevo, 2.XII.04, lg G. Carotti, 1 , coll. F. M. B.
The present subspecies is distinguished from the nominal one on
the basis of the length of wings that are longer than pronotum in S. b.
extensus.
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B50
OTHER SPECIES CITED FOR ECUADOR:
Among Batrachideinae:
Scaria brevis Hancock 1909 described from unspecified locality in
Ecuador.
Scaria fasciata Hancock 1907 described from Cachabi, in Ecuado-
rean Coastal Plains (HEBARD 1923).
Scaria hamata (De Geer 1773) is cited by GIGLIO TOS (1898) from
Valley of Santiago River.
Scaria maculata Giglio Tos 1898 described on one male and one
female from Ecuador, Valley of Santiago River. The species does not
appears on Orthoptera Species File online version june 2007, but was
not synonymized with other species. No material bearing this name is
present in the material studied by Giglio Tos in the Museum of Torino.
Rehnidium perexiguum Grant 1956 described from Baños in the
Ecuadorean Andes.
Tettigidea planus Hancock 1907 described from Paramba, on Ecua-
dorean Andes (BRUNER 1910).
Among Metrodorinae:
Amorphopus notabilis Serville 1838 cited sub Paratettix cnemidotus
by GIGLIO TOS (1898) on the basis of a single female from Cuchipamba
at about 1050 meters of elevation on the eastern slopes of the Andes,
northwest of Gualaquiza. This specimen was collected by Enrico Festa
during one of his two visits to the hacienda Cuchipamba, the first the
12 November 1895, the latter the 18-20 July 1896. The travel report of
the Italian naturalist (FESTA 1909) gives no informations on collecting
of Orthoptera in Cuchipamba, but since the second visit was characte-
rized by continuous rains that compelled Festa to stop there inside the
hacienda for two days, the specimen of A. notabilis was likely collected
during November 1895. The specimen from Coca examined by GÜN-
THER (1939) is the largest of the series reported. The species is known
in Ecuador East of the Andes.
The genus Cotys Bolivar 1887 contains two species, both known
for Ecuador. Cotys gibbulosus Hancock 1913 described from Santo
Domingo del Los Colorados and C. antennatus Bolivar 1887 cited by
GÜNTHER (1939) for Amazonian Ecuador.
Otumba amazonica (Bolivar 1887) reported by GIGLIO TOS (1898)
sub Metrodora amazonica from Valley of Santiago River.
Otumba incompta Hebard 1924. As noted by HEBARD (1924b) the
genera Otumba Morse 1900 and Cotys Bolivar are very near and some
51F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
species bear character states intermediate between the two genera, so
that are not clearly ascribable to one or the other genus. Type locality
of the species in exam is Rio Pescado, about 480 m, Azuay, apparently
ranging in Ecuador from the Andes to Coastal Plains. GÜNTHER (1939)
cites material of this species from Balzapamba on the Andes.
Chiriquia mirifica Hebard 1924. Described from Rio Pescado, about
480 m, Azuay, this species bears a peculiar characteristic among other
Neotropical tetrigids in having the carinae of the vertex dorsally pro-
duced in two small horns (HEBARD 1924b). Cited but not examined by
GÜNTHER (1939).
Scabrotettix scabrosus Hancock 1907 cited by HEBARD (1924b) refer-
ring to this species the single male from «between Quito and Napo Ri-
ver» mentioned by SCUDDER (1869) as Amorphopa caiman Saussure 1861.
Crimisus contractus Bolivar 1887 cited by GIGLIO TOS (1898) for
Pun and Huaca collected on 9-11 January 1897 according to travel notes
(FESTA 1909).
Trigonofemora fossulatus Hancock 1907, described from Peru, re-
ported by GÜNTHER (1939) for Ecuador, Santa Inez.
Plesiotettix Hancock 1907 comprehends P. spinosus Hancock 1907
and P. uncinatus Hancock 1907, both from Peru. GÜNTHER (1939) cites
one single male from Ecuador, Cachabé as Plesiotettix sp.
Crimisus acutus Günther 1939 described from Ecuador, Huaca.
Among Tetriginae:
Paratettix toltecus (Saussure 1861) reported by GIGLIO TOS (1898)
for Gualaquiza. HEBARD 1933 synonymized P. borellii Giglio Tos 1897
with P. frey-gessneri Bolivar 1887. The citation by Hebard appers ra-
ther criptic since he treats P. borellii as «Ecuadorean», actually being
that species described from Argentina; furthermore the author writes
as he had previously synonymized borelli with P. toltecus (Saussure
1861), but no reference is given and we found no synonymy of borelli
with toltecus in previous publications by Hebard.
DISCUSSION
In the Orthoptera collection of Museo Regionale di Scienze Natu-
rali di Torino, where the material collected by Enrico Festa and stu-
died by Giglio Tos should be preserved, the only tetrigids present are
Otumba amazonica (Bolivar 1887), identified by Giglio Tos as Metrodo-
ra amazonica, and Metrodora rana Bolivar 1887. Likely other material
Atti Acc. Rov. Agiati, a. 257, 2007, ser. VIII, vol. VII, B52
species ocurrence reference
Batrachideinae
1. Batrachidea inermis Hebard 1923 new for Ecuador
2. Batrachidea mucronata
(Saint-Fargeau & Serville, 1825) confirmed Grant 1956
3. Paurotarsus ruficornis (Walker 1871) new for Ecuador
4. Rehnidium perexiguum Grant 1956 Grant 1956
5. Scaria brevis Hancock 1909 Hancock 1909
6. Scaria fasciata Hancock 1907 Hebard 1923
7. Scaria hamata (De Geer 1773) Giglio Tos 1898
8. Scaria maculata Giglio Tos 1898 Giglio Tos 1898
9. Scaria producta Hancock 1907 new for Ecuador
10. Tettigidea planus Hancock 1907 Bruner 1910
Metrodorinae
11. Allotettix americanus Hancock 1907 confirmed Hebard 1924b
12. Allotettix peruvianus (Bolivar 1887) Hebard 1924b
13. Amorphopus notabilis Serville 1838 Günther 1939
14. Chiriquia compressa Günther 1939 Günther 1939
15. Chiriquia concinna (Bolivar 1887) confirmed Günther 1939
16. Chiriquia mirifica Hebard 1924 Hebard 1924b
17. Chriquia spinata Günther 1939 Günther 1939
18. Cotys antennatus Bolivar 1887 Günther 1939
19. Cotys gibbulosus Hancock 1913 Hebard 1924b
20. Crimisus acutus Günther 1939 Günther 1939
21. Crimisus contractus Bolivar 1887 Giglio Tos 1898
22. Crimisus bolivianus (Bruner 1913) New for Ecuador
23. Crimisus tibialis (Bruner, 1910) New for Ecuador
24. Eomorphopus antennatus (Bolivar 1887) New for Ecuador
25. Metrodora acuta Günther 1939 Günther 1939
26. Metrodora rana Bolivar 1887 confirmed Hebard 1924b
27. Otumba amazonica (Bolivar 1887) Günther 1939
28. Otumba incompta Hancock 1913 Günther 1939
29. Otumba qudrata Hancock 1907 Hancock 1907
30. Plesiotettix sp. Günther 1939
31. Scabrotettix bolivianus bolivianus
Hancock 1907 New for Ecuador
32. Scabrotettix bolivianus extensus
Hancock 1907 confirmed Günther 1939
33. Scabrotettix scabrosus Hancock 1907 Hebard 1924b
34. Trigonofemora fossulatus Hancock 1907 Günther 1939
Tetriginae
35. Paratettix proximus (Hancock 1907) Otte 1978
36. Paratettix toltecus (Saussure 1861) Giglio Tos 1898
Tab. 1. List of Tetrigidae actually known for Ecuador.
53F. M. BUZZETTI & H. DEVRIESE: Tetrigidae from Ecuador...
has been sent to foreign specialists during the first decades of XXth
century and never returned.
After recent collectings and from literature, thirty-six specific and
subspecific taxa of Tetrigidae are known from Ecuador (Tab. 1), twen-
ty-nine were known previously to this study, the presence of five of
these is confirmed, seven are here reported for the first time. All are
from continental Ecuador since Galapagos Islands, one of World bio-
diversity hotspots, are apparently lacking of tetrigids as reported by
PECK (2001) and confirmed by recent expeditions on some islands of
the archipelago (Carotti, pers. comm.). This number is probably only a
part of the real Tetrigidae fauna of Ecuador, being this family overloo-
ked since many decades, at least in the Neotropics.
AKNOWLEDGEMENTS
We are grateful to prof. Giovanni Onore (PUCE University of Quito, Ecuador)
for his kind hospitality and support during expeditions of first author, and to Giovan-
ni Carotti (Castelplanio, Italy) for the pictures here reproduced.
REFERENCES
BOLIVAR I., 1887 - Essai sur les Acridiens de la tribu des Tettigidae - Annales de la
Société Entomologique Belgique. 31: 175-313, pl. 4-5.
BRUNER L., 1906a - Report on the Orthoptera of Trinidad, West Indies - Journal of the
New York Entomological Society. 14: 135-168.
BRUNER L., 1906b - Synoptic list of Paraguayan Acrididae or locusts with descrip-
tions of new forms - Proceedings of the United States National Museum. 30 (1461):
613-694.
BRUNER L., 1910 - South American Tetrigidae - Annals of the Carnegie Museum, VII
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Indirizzi degli Autori:
Filippo Maria Buzzetti, Università di Padova, DAAPV-Entomologia, AGRIPOLIS
Viale dellUniversità 16, I-35020 Legnaro (PD), Italia,
filippomaria.buzzetti@unipd.it
Hendrik Devriese, Willem Bernardstraat 2, 1780 Wemmel, Belgium