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Geographic distribution and changes in
population densities of waterfowl in the
Northwest Territories, Canada, 1976-2003
Bonnie J. Fournier and James E. Hines
Canadian Wildlife Service
Environmental Conservation Branch
Technical Report Series Number 433
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Geographic distribution and changes in population densities
of waterfowl in the Northwest Territories,
Canada, 1976-2003
Bonnie J. Fournier
James E. Hines
Technical Report Series No. 433
Canadian Wildlife Service
Prairie and Northern Region
March 2005
This report may be cited as:
Fournier, B. J. and J. E. Hines. 2005.
Geographic distribution and changes in population densities of waterfowl in the
Northwest Territories, Canada, 1976-2003. Technical Report Series No. 433
Canadian Wildlife Service, Yellowknife, Northwest Territories.
Published under the Authority of the
Minister of Environment
Canadian Wildlife Service
© Her Majesty the Queen in Right of Canada, represented by the
Minister of the Environment, 2005.
Catalogue No. CW69-5/433E
ISBN 0-662-39949-8
Copies may be obtained from:
Canadian Wildlife Service
Environmental Conservation Branch
Environment Canada
Suite 301, 5204-50th Ave.
Yellowknife, Northwest Territories
X1A 1E2
ABSTRACT
This report summarizes, in a series of maps, the distribution of several common
types of waterfowl throughout much of the mainland (707,589 km2) of the Northwest
Territories, Canada. The aerial survey data used in mapping represent part of a much
larger dataset collected each year by the US Fish and Wildlife Service, Canadian Wildlife
Service, and other collaborators during the Waterfowl Breeding Population and Habitat
Survey. Six strata have been delineated in the Northwest Territories and 25 transects are
surveyed by fixed-winged aircraft each year. Data from each 29-km transect segment
were geographically coded and so could be used in Geographic Information System
mapping of waterfowl densities following an Inverse Distance Weighting approach. Data
from 1976-2003, a period when the surveys had become highly standardized, were used
in production of the density maps. By comparing, earlier (1976-1980) and more recent
(1999-2003) count data, locations where numbers of waterfowl may have increased or
declined are identified. Among the common types of waterfowl considered are Mallard,
American Wigeon, Northern Pintail, Green-winged Teal, Blue-winged Teal, Northern
Shoveler, Canvasback, Ring-necked Duck, Bufflehead, Long-tailed Duck, Canada Goose,
and five general species groups (scaup, goldeneyes, scoters, mergansers, and swans).
Given the potential for industrial development in the region, the maps should be of
immediate application in wildlife management, environmental impact assessments, and
general conservation planning.
RÉSUMÉ
Grâce à une série de cartes, ce rapport résume la répartition de plusieurs espèces
de sauvagine communes dans la majeure partie continentale (707 589 km2) des
Territoires du Nord-Ouest, au Canada. Les données de relevé aérien qui ont servi à
dresser les cartes font partie d’un ensemble beaucoup plus grand de données recueillies
chaque année par le US Fish and Wildlife Service, le Service canadien de la faune et
d’autres collaborateurs dans le cadre du relevé des populations reproductrices et des
habitats de la sauvagine. Six strates ont été établies dans les Territoires du Nord-Ouest, et
le relevé a été effectué par avion sur 25 transects chaque année. Les données pour chaque
segment de transect de 29 km ont été géoréférencées afin de cartographier les densités de
sauvagine à l’aide d’un système d’information géographique et de la méthode de
pondération par distance inverse. Les cartes de densité ont été produites à partir des
données recueillies de 1976 à 2003, une période pour laquelle les relevés étaient très
normalisés. Les endroits où l’abondance de la sauvagine aurait augmenté ou diminué ont
été relevés en comparant les données de dénombrement du début (1976-1980) et de la fin
(1999-2003) de cette période. Les espèces de sauvagine étudiées comprennent le canard
malard, le canard siffleur d’Amérique, le canard pilet, la sarcelle d’hiver, la sarcelle à
ailes bleues, le canard souchet, le fuligule à dos blanc, le fuligule à collier, le petit garrot,
l’harelde kakawi, la bernache du Canada, ainsi que cinq groupes d’espèces généraux
(fuligule, garrots, macreuses, harles et cygnes). Étant donné le potentiel de
développement industriel de la région, les cartes peuvent servir immédiatement à la
gestion de la faune, à des évaluations environnementales et à la planification générale en
matière de conservation.
vii
ACKNOWLEDGEMENTS
The data summarized in this report represent the collective efforts of many
contributors, most notably the pilot-biologists and observers who carried out the surveys
and many individuals who have been involved in managing the database over the years.
We would like to specifically acknowledge the assistance of Alan Davenport, GIS
Coordinator, U.S. Fish & Wildlife Service, Division of Migratory Bird Management,
Laurel, Maryland, in clarification of stratum, transect, and segment locations.
viii
TABLE OF CONTENTS
ABSTRACT....................................................................................................................... vi
RÉSUMÉ ..........................................................................................................................vii
ACKNOWLEDGEMENTS.............................................................................................viii
LIST OF TABLES.............................................................................................................. x
LIST OF FIGURES ...........................................................................................................xi
1. INTRODUCTION ...................................................................................................... 1
2. METHODS ................................................................................................................. 1
3. RESULTS ................................................................................................................... 3
4. LITERATURE CITED............................................................................................... 4
ix
LIST OF TABLES
Table 1. Stratum number, general habitat present, stratum area, and number of transects
surveyed in the Northwest Territories as part of the Waterfowl Breeding Population
and Habitat Survey, 1976-2003. ................................................................................. 5
Table 2. Species and species groups referred to in the text or figures................................ 5
x
LIST OF FIGURES
Figure 1. Study area, strata and transects for the Waterfowl Breeding Population and
Habitat surveys within the Northwest Territories....................................................... 6
Figure 2. Geographic distribution of Mallards in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003............................. 7
Figure 3. Geographic distribution of American Wigeon in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.. ......... 7
Figure 4. Geographic distribution of Northern Pintails in the Northwest Territories during
the Waterfowl Breeding Population and Habitat Surveys, 1976-2003....................... 8
Figure 5. Geographic distribution of Green-winged Teal in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.. ......... 8
Figure 6. Geographic distribution of Blue-winged Teal in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.. ......... 9
Figure 7. Geographic distribution of Northern Shoveler in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.. ......... 9
Figure 8. Geographic distribution of scaup in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003........................... 10
Figure 9. Geographic distribution of Ring-necked Ducks in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003. ........ 10
Figure 10. Geographic distribution of Canvasbacks in the Northwest Territories during
the Waterfowl Breeding Population and Habitat Surveys, 1976-2003..................... 11
Figure 11. Geographic distribution of Buffleheads in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003........................... 11
Figure 12. Geographic distribution of Long-tailed Ducks in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003. ........ 12
Figure 13. Geographic distribution of goldeneyes in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003........................... 12
Figure 14. Geographic distribution of scoters in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003........................... 13
Figure 15. Geographic distribution of mergansers in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003........................... 13
xi
Figure 16. Geographic distribution of Canada Geese in the Northwest Territories during
the Waterfowl Breeding Population and Habitat Surveys, 1976-2003..................... 14
Figure 17. Geographic distribution of swans in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003........................... 14
Figure 18. Geographic distribution of all species of waterfowl in the Northwest
Territories during the Waterfowl Breeding Population and Habitat Surveys, 1976-
2003........................................................................................................................... 15
Figure 19. Geographic changes in population densities of Mallards in the Northwest
Territories between 1976-1980 and 1999-2003........................................................ 15
Figure 20. Geographic changes in population densities of American Wigeon in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 16
Figure 21. Geographic changes in population densities of Northern Pintails in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 16
Figure 22. Geographic changes in population densities of Green-winged Teal in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 17
Figure 23. Geographic changes in population densities of Northern Shovelers in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 17
Figure 24. Geographic changes in population densities of Canvasbacks in the Northwest
Territories between 1976-1980 and 1999-2003........................................................ 18
Figure 25. Geographic changes in population densities of scaup in the Northwest
Territories between 1976-1980 and 1999-2003........................................................ 18
Figure 26. Geographic changes in population densities of Ring-necked Ducks in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 19
Figure 27. Geographic changes in population densities of Buffleheads in the Northwest
Territories between 1976-1980 and 1999-2003........................................................ 19
Figure 28. Geographic changes in population densities of Long-tailed Ducks in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 20
Figure 29. Geographic changes in population densities of scoters in the Northwest
Territories between 1976-1980 and 1999-2003........................................................ 20
Figure 30. Geographic changes in population densities of all species of waterfowl in the
Northwest Territories between 1976-1980 and 1999-2003. ..................................... 21
xii
1. INTRODUCTION
The U.S. Fish and Wildlife Service and the Canadian Wildlife Service, in
cooperation with others, have conducted annual aerial surveys of breeding waterfowl
throughout much of central Canada, the north-central United States, and Alaska since
1955. The Waterfowl Breeding Population and Habitat Surveys provide reliable
information on the spring population size and trajectory for many common species of
waterfowl in North America. The resulting data are used in the establishment of hunting
regulations in the United States and Canada and provide long-term information that is
critical for effective conservation efforts. Most of the vast boreal and sub-arctic area
region of the Northwest Territories is included within the area surveyed each year.
The information on waterfowl numbers recorded during the surveys are
geographically referenced and thus are suitable for mapping with a Geographic
Information System. It is the primary objective of this report to summarize the long term
data in a series of maps, depicting the distribution of some common waterfowl species on
the mainland of the Northwest Territories. By comparing, earlier and more recent count
data, locations where waterfowl numbers may have increased or declined are identified as
well.
2. METHODS
For survey purposes, six strata have been delineated in the Northwest Territories
according to broad geographic and habitat characteristics (Table 1, Figure 1). Twenty-
five transects, each divided into 29-km long segments, are surveyed during late May and
early June of each year. The number of waterfowl counted along the transect route are
entered into a computer database with the transect segments serving as a basis for
recording the data. Parts of two strata (numbers 16 and 18) extend into northern Alberta.
For the purpose of this report, we have used only data from those strata when transects
were completely within the Northwest Territories. Between 1955 and 1975, the survey
design went through a period of gradual modification as a number of additional transect
segments or entire transects were added. The survey design has remained consistent
since 1976 so we chose to map only the data collected since that year.
1
The segment-level database, derived from the Waterfowl Breeding Population and
Habitat Survey was obtained online through the FWS/USGS Migratory Bird Data Center
(http://mbdcapps.fws.gov/). Variables in this database include year, stratum, transect, and
segment identifiers, species, and the number of single drakes, pairs, and flocked
waterfowl counted by the aerial survey crew. We decided to use “total indicated birds”
for this report and used data that had been corrected for visibility bias. The visibility
correction factors applied to raw segment counts have been derived from a comparison of
ground (or helicopter) counts and the counts from fixed-wing aircraft (Smith 1995).
Survey procedures for the Waterfowl Breeding Population and Habitat Survey are
described in detail by Smith (1995) and a description can be found online at
http://www.fws.gov/birddata/databases/mas/aboutmas.htm. Additional detail concerning
stratum boundaries and the location of transects and segments can be viewed interactively
at http://mbirdims.fws.gov/nbii/.
The survey area shown on our maps includes 6 strata and covers approximately
707,589 km2 (Table 1, Figure 1). Two types of maps are presented: (1) density contour
maps showing the broad distribution of each species in the survey area; and (2) trend
maps showing geographic changes in average population densities between two 5-year
periods: 1976-1980 and 1999-2003. Both map types were modeled using an Inverse
Distance Weighting (IDW) approach with a Geographic Information System (Bonham-
Carter, G. 1994). IDW has been used to map large scale data collection efforts such as
the North American Breeding Bird Survey (Sauer 2003) as well as other arctic waterfowl
surveys (Alisauskas 1997, Hines et al. 2004). We used the IDW interpolator in the Spatial
Analyst 1.1 extension for ArcView 3.2 (Environmental Research Systems Institute 2000)
to carry out the GIS analyses. Essentially, we needed to define three parameters in order
to carry out the analyses: (1) a search radius around a given cell (point on a map) which
determined the neighbouring data points to include in the analysis; (2) a cell size
reflecting the ultimate resolution the maps; and (3) an exponent or power value.
Transects were not regularly distributed, so we used a search radius that included the 15 –
20 data points that were nearest to the particular cell of interest rather than using a search
radius defined by a fixed distance (e.g., 10 km). A cell size of 2000 m was chosen as it
2
provided adequate map resolution as well as an acceptable computer processing time.
We found little guidance in the literature on what exponent value to use. We decided on
an exponent of 2 as that has been conventionally used in the past. More details on the
IDW method as applied to waterfowl surveys in the Northwest Territories are presented
in Hines et al. (2004).
3. RESULTS
The scientific names of all species of waterfowl considered in this report are
presented in Table 2 and maps indicating the distributions of 16 species or species groups
of waterfowl are presented in Figures 2 – 17. The identities of some closely related
species cannot be reliably separated from the air so these species are mapped as generic
groups such as scaup, scoters, and mergansers (Table 2).
Maps showing changes in population densities are presented for 11 species or
species groups: Mallard, American Wigeon, Northern Pintail, Green-winged Teal,
Northern Shoveler, Canvasback, scaup, Ring-necked Duck, Bufflehead, Long-tailed
Duck, and scoters. Some species (e.g. Blue-winged Teal) were not widely distributed or
occurred at low densities, and could not be effectively treated in the trend map analyses.
In general, we attempted to map all species with a common map style. Because
of their relatively high population densities, it was necessary to use different density
classes to adequately portray the changes in densities of three numerous species or
species groups (American Wigeon, scaup and scoters).
3
4. LITERATURE CITED
Alisauskas, Ray. 1997. Spatial interpolation for post-stratification of aerial survey data.
Pages 383-387 in GIS `97 Conference Proceedings. Fort Collins, Colorado.
Bonham-Carter, G. 1994. Geographic information systems for geoscientists: Modelling
with GIS. Pergamon Press. New York, N.Y.
Environmental Research Systems Institute 2000. ArcView 3.2 Spatial Analyst.
Environmental Research Systems Institute. Redlands, California.
Hines, J.E., B. Fournier, and J. O’Neill. 2004. Spring and fall distribution of waterfowl
and other aquatic birds on the mainland of the Inuvialuit Settlement Region, Western
Canadian Arctic, 1990-98. Technical Report Series No. 426. Canadian Wildlife Service,
Yellowknife, Northwest Territories.
Sauer, J.R., J.E. Hines, and J. Fallon. 2003. The North American Breeding Bird Survey,
results and analysis 1966 - 2002. Version 2003.1,
, Laurel, Maryland. USGS Patuxent Wildlife Research
Center
Smith, G.W. 1995. A critical review of the aerial and ground surveys of breeding
waterfowl in North America. Biological Science Report 5, National Biological Service,
Washington, D.C. 252 pp.
4
Table 1. Stratum number, general habitat present, stratum area, and number of
transects surveyed in the Northwest Territories as part of the Waterfowl Breeding
Population and Habitat Survey, 1976-2003.
Table 2. Species and species groups referred to in the text or figures.
Species Scientific Name Comment
Mallard Anas platyrhynchos
American Wigeon Anas americana
Northern Pintail Anas acuta
Green-winged Teal Anas crecca
Blue-winged Teal Anas discors
Northern Shoveler Anas clypeata
Canvasback Aythya valisineria
Scaup includes Lesser Scaup (Aythya affinis) and Greater
Scaup (Aythya marila).
Ring-necked Duck Aythya collaris
Bufflehead Bucephala albeola
Goldeneyes includes Common Goldeneyes (Bucephala
clangula) and Barrow’s Goldeneyes (Bucephala
islandica).
Long-tailed Duck Clangula hyemalis
Scoters includes White-winged Scoters (Melanitta fusca),
Surf Scoters (Melanitta perspicillata), and
possibly small numbers of Black Scoters
(Melanitta nigra).
Mergansers includes Common Mergansers (Mergus
merganser) and Red-breasted Mergansers (Mergus
serrator) and smaller numbers of Hooded
Mergansers (Lophodytes cucullatus).
Canada Goose Branta canadensis
Swans primarily Tundra Swans (Cygnus columbianus)
but may include very small numbers of Trumpeter
Swans (Cygnus buccinator).
Stratum
number General habitat or ecozone Stratum
area (km2) Number of
transects
13 Mackenzie Delta 12,782 4
Taiga Plains and Arctic Plains 205,172 8 14
15 Taiga Plains 130,696 2
16 Taiga Shield 149,756 4
17 Taiga Plains 178,795 5
30,388
Boreal Plains and Taiga Shield 18 2
5
Figure 1. Study area, strata and transects for the Waterfowl Breeding Population and
Habitat surveys within the Northwest Territories.
6
Figure 2. Geographic distribution of Mallards in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 3. Geographic distribution of American Wigeon in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
7
Figure 4. Geographic distribution of Northern Pintails in the Northwest Territories during
the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 5. Geographic distribution of Green-winged Teal in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
8
Figure 6. Geographic distribution of Blue-winged Teal in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 7. Geographic distribution of Northern Shoveler in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
9
Figure 8. Geographic distribution of scaup in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 9. Geographic distribution of Ring-necked Ducks in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
10
Figure 10. Geographic distribution of Canvasbacks in the Northwest Territories during
the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 11. Geographic distribution of Buffleheads in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
11
Figure 12. Geographic distribution of Long-tailed Ducks in the Northwest Territories
during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 13. Geographic distribution of goldeneyes in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
12
Figure 14. Geographic distribution of scoters in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 15. Geographic distribution of mergansers in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
13
Figure 16. Geographic distribution of Canada Geese in the Northwest Territories during
the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 17. Geographic distribution of swans in the Northwest Territories during the
Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
14
Figure 18. Geographic distribution of all species of waterfowl in the Northwest
Territories during the Waterfowl Breeding Population and Habitat Surveys, 1976-2003.
Figure 19. Geographic changes in population densities of Mallards in the Northwest
Territories between 1976-1980 and 1999-2003.
15
Figure 20. Geographic changes in population densities of American Wigeon in the
Northwest Territories between 1976-1980 and 1999-2003.
Figure 21. Geographic changes in population densities of Northern Pintails in the
Northwest Territories between 1976-1980 and 1999-2003.
16
Figure 22. Geographic changes in population densities of Green-winged Teal in the
Northwest Territories between 1976-1980 and 1999-2003.
Figure 23. Geographic changes in population densities of Northern Shovelers in the
Northwest Territories between 1976-1980 and 1999-2003.
17
Figure 24. Geographic changes in population densities of Canvasbacks in the Northwest
Territories between 1976-1980 and 1999-2003.
Figure 25. Geographic changes in population densities of scaup in the Northwest
Territories between 1976-1980 and 1999-2003.
18
Figure 26. Geographic changes in population densities of Ring-necked Ducks in the
Northwest Territories between 1976-1980 and 1999-2003.
Figure 27. Geographic changes in population densities of Buffleheads in the Northwest
Territories between 1976-1980 and 1999-2003.
19
Figure 28. Geographic changes in population densities of Long-tailed Ducks in the
Northwest Territories between 1976-1980 and 1999-2003.
Figure 29. Geographic changes in population densities of scoters in the Northwest
Territories between 1976-1980 and 1999-2003.
20
Figure 30. Geographic changes in population densities of all species of waterfowl in the
Northwest Territories between 1976-1980 and 1999-2003.
21