Order Trombidiformes Reuter, 1909

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Title:
Order Trombidiformes Reuter, 1909. In:
Zhang, Z-Q. (ed.) Animal biodiversity: an
outline of higher-level classification and
survey of taxonomic richness.
Authors:
Zhang, Z.-Q., Fan, Q.-H., Pesic, V., Smit, H., Bochkov, A.V., Khaustov, A.A., Baker, A., Wohltmann,
A., Wen, T.-H., Amrine, J.W.,Beron, P., Lin, J.-Z., Gabrys, G. & Husband, R.
Source:
Zootaxa 3148: 129–138.
Date:
2011/12/23

Zootaxa 3148 © 2011 Magnolia Press · 129
ANIMAL BIODIVERSITY
Order Trombidiformes Reuter, 19091, 2
Suborder Sphaerolichida OConnor, 1984 (2 superfamilies)3
Superfamily Lordalychoidea Grandjean, 1939 (2 families)
Family Lordalychidae Grandjean, 1939 (1 genus, 16 species)
Superfamily Sphaerolichoidea Berlese, 1913
Family Sphaerolichidae Berlese, 1913 (1 genus, 5 species)
Suborder Prostigmata Kramer, 1877 (4 infraorders, 40 superfamilies)
Infraorder Labidostommatina Krantz, 1978 (1 superfamily)
Superfamily Labidostommatoidea Oudemans, 1904 (1 family)
Family Labidostommatidae Oudemans, 1904 (5 genera, 51 species; †0/2)4
Infraorder Eupodina Krantz, 1978 (5 superfamilies)
Superfamily Bdelloidea Dugès, 1834 (2 families)
Family Bdellidae Dugès, 1834 (15 genera, 256 Species; †0/5)5
Family Cunaxidae Thor, 1902 (27 genera, 329 species)6
Superfamily Eriophyoidea Nalepa, 1898 (3 families)7
Family Phytoptidae Murray, 1877 (20 genera, 164 species)8
Family Eriophyidae Nalepa, 1898 (274 genera, 3,790 species)9
Family Diptilomiopidae Keifer, 1944 (63 genera, 450 species)
Superfamily Eupodoidea Koch, 1842 (9 families)10
Family Cocceupodidae Jesionowska, 2010 (3 genera, 23 species)11
Family Eupodidae Koch, 1842 (10 genera, 69 species)12
Family Dendrochaetidae Olivier, 2008 (1 genus, 1 species)13
Family Eriorhynchidae Qin & Halliday, 1997 (1 genus, 5 species)
Family Pentapalpidae Olivier & Theron, 2000 (1 genus, 1 species)
Family Penthaleidae Oudemans, 1931 (5 genera, 16 species)14
Family Penthalodidae Thor, 1933 (6 genera, 35 species; ; †0/1)15
Family Rhagidiidae Oudemans, 1922 (28 genera, 157 species; ; †1/1)
Family Strandtmanniidae Zacharda, 1979 (1 genus, 2 species)16
1. BY Zhi-Qiang Zhang, Qing-Hai Fan, Vladimir Pesic, Harry Smit, Andre V. Bochkov, A. A. Khaustov, Anne Baker, Andreas Wohltmann,
Tinghuan Wen, James W. Amrine, P. Beron, Jianzhen Lin, Grzegorz Gabrys & Robert Husband (for full auhor addresses, see Contributor
roles and addresses after References). The title of this contribution should be cited as “Order Trombidiformes Reuter, 1909. In: Zhang, Z.-Q.
(Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness”. Recent Trombididformes currently
include 151 families, 2,235 genera, and 25,821 species, whereas fossil taxa include 10 genera and 24 species.
2. Higher classification and estimates of genera and species within families generally follow Walter et al. (2009), with revisions and updates as
otherwise indicated. To be consistent with Beaulieu et al. (2011) and Schatz et al. (2011) in this volume, the rank cohort is not used and older
names for higher taxa are used—also consistent with OConnor (1984) and Mironov & Bochkov (2009).
3. Based on Moraza (2008) and Walter et al. (2009), with updates.
4. Based on Dunlop & Bertrand (2011); throughout this paper, counts of genera and species are totals (Recent and fossil); †x/y refers x genera and
y species, among the totals, are based on fossil.
5. Based on van der Schyff et al. (2003), Hernandes & Feres (2006), Hernandes et al. (2007), with updates.
6. Based on Den Heyer (2011).
7. Based on unpublished database of James Amrine.
8. One species of fossil (Dunlop et al. 2011).
9. Three species of fossil galls (Dunlop et al. 2011).
10. Species numbers do not include taxa that are evidently correctly placed in the Eupodoidea but are insufficiently defined for recognition. Many of
these were described by C.L. Koch (see Thor & Willmann 1941). Jesionowska (2010) and Zacharda (1980) list species inquirendae for the
Cocceupodidae and Rhagidiidae respectively.
11. Based on Jesionowska (2010).
12. Fain (1964) synonymized Protereunetes Berlese with Ereynetes Berlese. Despite Strandtmann (1970) reassigning eupodid species described in
Protereunetes to Eupodes Koch, the generic name continued to be used by some authors. Fain’s synonymy is accepted and Protereunetes is not
here recognized as a eupodid genus.
Jesionowska (2008) transferred Hawaiieupodes Strandtmann & Goff from the Eupodidae to the Penthalodidae. However, the unsettled
diagnosis of the latter means that the new placement is also unsatisfactory. Consequently, the genus is here retained in the Eupodidae as
originally assigned (Strandtmann & Goff 1978).
Abou-Awad (1984) erected Egypteupodes on the basis of the absence of internal vertical setae (iv). Olivier and Theron (1997)
synonymized the genus with Eupodes because they observed that, although minute, iv were present. However, because it is not clear whether
these authors examined type material, Egypteupodes is included in the generic complement.
13. Published in 2008, 2009, too late to be included by Walter et al. (2009).
14. See first footnote for the Penthalodidae.

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Superfamily Halacaroidea Murray, 1877 (2 families)
Family Halacaridae Murray, 1877 (63 genera, 1,118 species)17
Family Pezidae Harvey, 1990 (1 genus, 2 species)
Superfamily Tydeoidea Kramer, 1877 (4 families)18
Family Triophtydeidae André, 198019 (3 genera, 40 species)
Family Ereynetidae Oudemans, 1931 (29 genera, 180 species)
Family Tydeidae Kramer, 1877 (30 genera, 340 species; †1/2)
Family Iolinidae Pritchard, 1956 (36 genera, 125 species)
Infraorder Anystina van der Hammen, 1972 (2 hyporders, 22 superfamilies)
Hyporder Anystae Krantz, 1978 (5 superfamilies)
Superfamily Adamystoidea Cunliffe, 1957 (1 family)
Family Adamystidae Cunliffe, 1957 (4 genera, 20 species)20
Superfamily Anystoidea Oudemans, 1936 (3 families)
Family Anystidae Oudemans, 1936 (18 genera, 107 species; †2/5)21
Family Pseudocheylidae Oudemans, 1909 (3 genera, 15 species)22
Family Teneriffiidae Thor, 1911 (5 genera, 20 species)23
Superfamily Caeculoidea Berlese, 1883 (1 family)
Family Caeculidae Berlese, 1883 (7 genera, 73 species; †1/2)24
Superfamily Pomerantzioidea Baker, 1949 (1 family)
Family Pomerantziidae Baker, 1949 (2 genera, 6 species)25
Superfamily Paratydeoidea Baker, 1949 (1 family)
Family Paratydeidae Baker, 1949 (6 genera, 10 species)26
Family Stigmocheylidae Kethley, 1990 (1 genus, 4 species)27
Hyporder Parasitengona Oudemans, 1909 (17 superfamilies)28
Superfamily Calyptostomatoidea Oudemans, 1923 (1 family)29
Family Calyptostomatidae Oudemans, 1923 (1 genus, 6 species)
Superfamily Erythraeoidea Robineau-Desvoidy, 1828 (2 families)30
Family Erythraeidae Robineau-Desvoidy, 1828 (55 genera, 771 species)31
Family Smarididae Kramer, 1878 (10 genera, 53 species)32
15. Penthalodidae: Qin (1997) considered that Protopenthalodes Jesionowska would be better accommodated in either the Eupodidae or
Penthaleidae, rather than the Penthalodidae as originally placed (Jesionowska 1989). However, he retained the genus in the latter until familial
monophyly was resolved and a correct taxonomic position could be established.
Jesionowska (2008) reassessed the importance of the epirostrum as a diagnostic familial character state and concluded that
Callipenthalodes Qin, Turanopenthalodes Barilo and Stereotydeus Berlese should be removed from the Penthalodidae. Because she neither
reassigned them to existing families nor erected new ones for them, these genera have been treated here as penthalodids.
16. Strandtmanniidae is considered a possible junior synonym of Rhagidiidae by Qin (1996).
17. Based on Bartsch (2009).
18. After Walter et al. (2009); we regret to note here that the data were not checked by a specialist.
19. H.M. André of Belgium (not M. André of France).
20. Based on Fuangarworn & Lekprayoon (2010).
21. Based on Meyer & Ueckermann (1987), Otto (1999, 2000), with updates.
22. Based on Ueckermann & Khanjani (2004).
23. Based on Judson (1994), Ueckermann & Khanjani (2002), Khanjani et al. (2011), with updates.
24. Based on Franz (1952), Coineau & Poinar (2001), with updates.
25. Based on Bochkov & Walter (2007).
26. Based on Seeman et al. (2000).
27. Based on Bochkov (2008a).
28. The classification above the family level is highly unsettled for this group (Zhang & Fan, 2007). One view is that water mites (Hydracarina)
form a sister group to the rest of the terrestrial Parasitengona (Krantz 1978; Smith & Cook 1991; Witte 1991). A radically different view
(Welbourn (1991), is that water mites and Trombidiina are considered sister groups, and together they form a sister group to Erythraeina
(Erythraeoidea + Calyptostomatoidea). Walter et al. (2009) proposed Stygothrombiae for Stygothrombidioidea, which was not analysed in Witte
(1991) and Welbourn (1991). Future analyses sampling all superfamilies of both water mites and terrestrial Parasitengona are much needed
before any reliable classification of the Parasitengona at the ordinal level can be reached. Therefore we provide a list of superfamilies without
choosing any higher classification.
29. Based on unpublished list of Beron, who also provided data for Erythraeoidea.
30. †Pararainbowia martilli Dunlop, 2007 is unplaced within the superfamily.
31. Based on Southcott (1961), Beron (2008) with updates. Dunlop et al. (2011) listed 13 species described from Baltic amber in 1854; these require
checking.
32. Based on Southcott (1961), Zhang (2003), Beron (2008), Wohltmann (2010) with updates.

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Family †Proterythraeidae Vercammen-Grandjean, 1973 (1 genus, 1 species; †1/1)
Superfamily Amphotrombioidea Zhang, 1998 (1 family)33
Family Amphotrombiidae Zhang, 1998 (1 genus, 1 species)
Superfamily Allotanaupodoidea Zhang & Fan, 2007 (1 family)34
Family Allotanaupodidae Zhang & Fan, 2007 (3 genera, 6 species)
Superfamily Chyzerioidea Womersley, 1954 (1 family)
Family Chyzeriidae Womersley, 1954 (11 genera, 30 species)35
Superfamily Tanaupodoidea Thor, 1935 (1 family)
Family Tanaupodidae Thor, 1935 (7 genera, 23 species; †1/1)36
Superfamily Trombiculoidea Ewing, 1929 (7 families)
Family Johnstonianidae Thor, 1935 (10 genera, 53 species)37
Family Neotrombidiidae Feider, 1959 (5 genera, 24 species)38
Family Trombellidae Leach, 1815 (19 genera, 41 species)39
Family Leeuwenhoekiidae Womersley, 1944 (33 genera, 230 species)
Family Trombiculidae Ewing, 1929 (152 genera, 3,100 species)40
Family Walchiidae Ewing, 1946 (19 genera, 298 species)
Family Audyanidae Southcott, 1987 (1 genus, 1 species)41
Superfamily Trombidioidea Leach, 1815 (4 families)
Family Achaemenothrombiidae Saboori, Wohltmann & Hakimitabar, 2010 (1 genus, 2 species)42
Family Neothrombiidae Feider, 1959 (16 genera, 25 species)43
Family Microtrombidiidae Thor, 1935 (126 genera, 437 species)44
Family Trombidiidae Leach, 1815 (25 genera, 253 species)45
Superfamily Yurebilloidea Southcott, 1996 (1 family)46
Family Yurebillidae Southcott, 1996 (1 genus, 1 species)47
Superfamily Hydryphantoidea Piersig, 1896 (8 families)48
Family Hydryphantidae Piersig, 1896 (51 genera, 329 species)
Family Hydrodromidae Viets, 1936 (2 genera, 28 species)
Family Rhynchohydracaridae Lundblad, 1936 (5 genera, 13 species)
Family Teratothyadidae Viets, 1929 (2 genera, 9 species)
Family Ctenothyadidae Lundblad, 1936 (2 genera, 3 species)
Family Thermacaridae Sokolow, 1927 (1 genera, 4 species)
Family Zelandothyadidae Cook, 1983 (2 genera, 3 species)
Family Malgasacaridae Tuzovskij, Gerecke & Goldschmidt 2007 (1 genus, 1 species)
Superfamily Eylaoidea Leach, 1815 (4 families)
Family Eylaidae Leach, 1815 (2 genera, 96 species)
Family Limnocharidae Grube, 1859 (5 genera, 44 species)
33. Based on larval stage only with similarities to both early-derivative water mites and Trombidina (Zhang 1998),
34. This superfamily was missed in the chapter on Trombidiformes in the Manual of Acarology, third edition (Walter et al. 2009). Zhang & Fan
(2007) provided a key to superfamilies of terrestrial Parasitengona (based on post-larval stages).
35. Based on Zhang & Saboori (1995); Haitlinger (1999), Saboori et al. (2005), with updates.
36. Based on Newell (1957), Welbourn, W.C. (1991), Judson & Makol (2009) with updates.
37. Based on Newell (1957, 1960), Wohltmann et al. (2004), with updates.
38. Based on Lindquist & Vercammen-Grandjean (1971), Vercammen-Grandjean (1973), Feider (1977), Beron (1995), with updates.
39. Based on Southcott (1987), Welbourn (1991), Harvey (1996), Saboori et al. (2005) with updates.
40. An estimate to be varified.
41. Also placed by some authors in Trombellidae (Southcott 1987).
42. Known from larvae only; Saboori et al. (2010) provided a key to families of Trombidioidea based on larvae.
43. Based on Zhang (1994), Zhang & Fan (2005), Haitlinger (2007), Saboori et al. (2008) and M kol et al. (2009), with updates.
44. Based on Southcott (1993, 1994), Gabry (1999), Wohltmann (2006), M kol & Laydanowicz (2010) with updates; Eutrombidiidae is merged
into Microtrombidiidae based on phylogenetic work of Wohltmann (2006).
45. Updated based on M kol (2007) and M kol & Sevsay (2011). Includes Podothrombiidae recognized in M kol (2007) and some earlier papers
cited by M kol (2007). Dunlop et al. (2011) listed 5 species described from Baltic amber in 1854; these require checking.
46. Updated from Southcott (1993) and M kol & Gabry (2005).
47. Described by Southcott (1996) from larval stage only, initially placed in Trombidioidea s.l., but was excluded from Trombidioidea s.s. by
Saboori et al. (2010). Here it is placed in Yurebilloidea Southcott, 1996; superfamily rank is used here for the first time; decision by Z. Zhang.
Based on larval characters, this superfamily is very closely related to Trombidoidea sin its strict sense.
48. Davids et al. (2006) with updates.

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Family Piersigiidae Oudemans, 1902 (4 genera, 9 species)
Family Apheviderulicidae Gerecke, Smith & Cook, 1999 (1 genus, 3 species)
Superfamily Hydrovolzioidea Thor, 1905 (2 families)
Family Hydrovolziidae Thor, 1905 (3 genera, 16 species)
Family Acherontacaridae Cook, 1967 (2 genera, 19 species)
Superfamily Hydrachnoidea Leach, 1815 (1 family)
Family Hydrachnidae Leach, 1815 (1 genus, 131 species)
Superfamily Lebertioidea Thor, 1900 (10 families)
Family Bandakiopsidae Panesar, 2004 (3 genera, 4 species)
Family Stygotoniidae Cook, 1992 (1 genus, 1 species)
Family Sperchontidae Thor, 1900 (5 genera, 173 species)
Family Rutripalpidae Sokolow, 1934 (1 genus, 2 species)
Family Teutonidae Koenike, 1910 (2 genera, 6 species)
Family Anisitsiellidae Koenike, 1910 (29 genera, 159 species)
Family Lebertiidae Thor, 1900 (3 genera, 138 species)
Family Acucapitidae Wiles, 1996 (1 genus, 2 species)
Family Oxidae Viets, 1926 (2 genera, 126 species)
Family Torrenticolidae Piersig, 1902 (6 genera, 397 species)
Superfamily Hygrobatoidea Koch, 1842 (13 families)
Family Pontarachnidae Koenike, 1910 (2 genera, 40 species)
Family Limnesiidae Thor, 1900 (29 genera, 227 species)
Family Omartacaridae Cook, 1963 (2 genera, 14 species)
Family Wettinidae Cook, 1956 (4 genera, 9 species)
Family Frontipodopsidae Viets, 1931 (1 genus, 9 species)
Family Ferradasiidae Cook, 1980 (1 genus, 1 species)
Family Lethaxonidae Cook, Smith & Harvey, 2000 (3 genera, 16 species)
Family Hygrobatidae Koch, 1842 (82 genera, 846 species)
Family Aturidae Thor, 1900 (81 genera, 800 species)
Family Feltriidae Viets, 1926 (1 genus, 115 species)
Family Unionicolidae Oudemans, 1909 (18 genera, 477 species)
Family Pionidae Thor, 1900 (15 genera, 342 species)
Family Astacocrotonidae Thor, 1927 (1 genus, 1 species)
Superfamily Arrenuroidea Thor, 1900 (20 families)
Family Momoniidae Viets, 1926 (13 genera, 55 species)
Family Nudomideopsidae Smith, 1990 (3 genera, 31 species)
Family Mideopsidae Koenike, 1910 (8 genera, 113 species)
Family Athienemanniidae Viets, 1922 (16 genera, 43 species)
Family Chappuisididae Motas & Tanasachi, 1946 (2 genera, 13 species)
Family Gretacaridae Viets, 1978 (1 genus, 15 species)
Family Neoacaridae Motas & Tanasachi, 1947 (2 genera, 19 species)
Family Mideidae Thor, 1911 (2 genera, 6 species)
Family Acalyptonotidae Walter, 1911 (2 genera, 4 species)
Family Kantacaridae Imamura, 1959 (1 genus, 1 species)
Family Nipponacaridae Imamura, 1959 (1 genus, 3 species)
Family Harpagopalpidae Viets, 1924 (1 genus, 7 species)
Family Arenohydracaridae Cook, 1974 (1 genus, 3 species)
Family Amoenacaridae Smith & Cook, 1997 (1 genus, 3 species)
Family Laversiidae Cook, 1955 (1 genus, 1 species)
Family Krendowskiidae Viets, 1926 (4 genera, 52 species)
Family Arrenuridae Thor, 1900 (7 genera, 950 species)
Family Bogatiidae Motas & Tanasachi, 1948 (2 genera, 3 species)
Family Hungarohydracaridae Motas & Tanasachi, 1959 (5 genera, 17 species)
Family Uchidastygacaridae Imamura, 1956 (3 genera, 15 species)
Superfamily Stygothrombidioidea Thor, 1935 (1 family)

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Family Stygothrombiidae Thor, 1935 (5 genera, 16 species)
Infraorder Eleutherengona Oudemans , 1909 (12 superfamilies)
Hyporder Raphignathina Kethley, 1982 (5 superfamilies)
Superfamily Cheyletoidea Leach, 1815 (5 families)
Family Cheyletidae Leach, 1815 (75 genera, 438 species)49
Family Demodicidae Nicolet, 1855 (7 genera, 108 species)50
Family Harpirhynchidae Dubinin, 1957 (14 genera, 93 species)51
Family Psorergatidae Dubinin, 1955 (3 genera, 73 species)52
Family Syringophilidae Lavoipierre, 1953 (63 genera, 259 species)53
Superfamily Myobioidea Mégnin, 1877 (1 family)
Family Myobiidae Mégnin, 1877 (54 genera, 503 species)54
Superfamily Pterygosomatoidea Oudemans, 1910 (1 family)
Family Pterygosomatidae Oudemans, 1910 (10 genera, 166 species)55
Superfamily Raphignathoidea Kramer 1877 (11 families)56
Family Barbutiidae Robaux, 1975 (1 genus, 5 species)57
Family Caligonellidae Grandjean, 1944 (5 genera, 50 species)58
Family Camerobiidae Southcott, 1957 (7 genera, 146 species; †0/1)59
Family Cryptognathidae Oudemans, 1902 (3 genera, 52 species)60
Family Dasythyreidae Walter & Gerson, 1998 (2 genera, 5 species)61
Family Eupalopsellidae Willmann, 1952 (5 genera, 40 species)62
Family Homocaligidae Wood, 1970 (2 genera, 8 species)63
Family Mecognathidae Gerson & Walter, 1998 (2 genera, 7 species)64
Family Raphignathidae Kramer, 1877 (3 genera, 57 species)65
Family Stigmaeidae Oudemans, 1931 (30 genera, 502 species)66
Family Xenocaligonellididae Gonzalez, 1978 (2 genera, 5 species)67
Superfamily Tetranychoidea Donnadieu 1875 (5 families)
Family Allochaetophoridae Reck, 1959 (1 genus, 2 species)68
Family Linotetranidae Baker & Pritchard, 1953 (4 genera, 16 species)69
Family Tenuipalpidae Berlese, 1913 (34 genera, 895 species)70
Family Tetranychidae Donnadieu 1875 (95 genera, 1,270 species)71
Family Tuckerellidae Baker & Pritchard, 1953 (1 genus, 28 species)72
Hyporder Heterostigmata Berlese, 1899 (7 superfamilies)
49. Based on Bochkov (2005) with additions.
50. Based on Bochkov (2009) with modifications.
51. Based on a unpublished database of A.V. Bochkov.
52. Based on Giesen (1990).
53. Based on Skoracki (2011) with additions.
54. Based on Bochkov (2009).
55. Based on Bochkov (2008b) with additions.
56. Based on a unpublished database of Q.-H. Fan.
57. Based on Fan et al. (2003) with additions.
58. Based on Fan (2005).
59. Based on Fan (2005) and Fan & Walter (2011) with additions.
60. Based on Do an & Dönel (2010) and Fan (2005).
61. Based on Walter & Gerson (1998) with additions.
62. Based on Fan (2005) with additions.
63. Based on Fan (1997, 2005).
64. Based on Fan & Zhang (2005) and Fan (2005).
65. Based on Fan (2005) with additions.
66. Based on Fan & Zhang (2005), Meyer & Ueckermann (1989) and Fan (2005) with additions.
67. Based on Fan (2005).
68. Based on Meyer & Ueckermann (1997).
69. Baed on Bagheri et al. (2008) and Khanjani et al. (2011), with updates.
70. Based on Mesa et al. (2009) and Beard & Gerson (2009), with updates by Q.-H. Fan.
71. Based on Spider Mites Web (http://www.ensam.inra.fr/CBGP/spmweb/index.php). Dunlop et al. (2011) listed 2 species described from Baltic
amber in 1854; these require checking.
72. Based on Meyer & Ueckermann (1997) with additions.

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Superfamily Dolichocyboidea Mahunka, 1970 (1 family)
Family Dolichocybidae Mahunka, 1970 (6 genera, 37 species)73
Family Crotalomorphidae Lindquist & Krantz, 2002 (1 genus, 1 species)
Superfamily Heterocheyloidea Trägårdh, 1950 (1 family)
Family Heterocheylidae Trägårdh, 1950 (1 genus, 25 species)
Superfamily Pyemotoidea Oudemans, 1937 (4 families)
Family Acarophenacidae Cross, 1965 (7 genera, 31 species; †1/1)
Family Caraboacaridae Mahunka, 1970 (2 genera, 7 species)
Family Resinacaridae Mahunka, 1976 (2 genera, 2 species; †1/1)
Family Pyemotidae Oudemans, 1937 (1 genus, 24, species; †1/1)74
Superfamily Pygmephoroidea Cross, 1965 (4 families)75
Family Pygmephoridae Cross, 1965 (27 genera, 310 species)
Family Scutacaridae Oudemans, 1916 (24 genera, 802 species)
Family Microdispidae Cross, 1965 (17 genera, 109 species)
Family Neopygmephoridae Cross, 1965 (17 genera, 248 species)
Superfamily Tarsocheyloidea Atyeo & Baker, 1964 (1 family)
Family Tarsocheylidae Atyeo & Baker, 1964 (2 genera, 3 species)
Superfamily Tarsonemoidea Kramer, 1877 (2 families)
Family Podapolipidae Ewing, 1922 (30 genera, 239 species)
Family Tarsonemidae Kramer, 1877 (41 genera, 566 species)76
Superfamily Trochometridioidea Mahunka 1970 (2 families)
Family Trochometridiidae Mahunka 1970 (2 genera, 6 species)77
Family Athyreacaridae Lindquist, Kaliszewski & Rack, 1990 (1 genus, 1 species)
Hyporder incertae sedis
Superfamily Cloacaroidea Camin, Moss, Oliver & Singer, 1967 (2 families)78
Family Cloacaridae Camin, Moss, Oliver & Singer, 1967 (6 genera, 14 species)
Family Epimyodicidae Fain, Lukoschus & Rosmalen, 1982 (1 genus, 4 species)
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77. Based on Mortazavi et al. (2011).
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Contributor roles and addresses (order of authors based on the amount of data contributed)
Zhi-Qiang Zhang (coordination, higher taxonomy and superfamilies/families not covered by others) New Zealand Arthropod Collec-
tion, Landcare Research, Private Bag 92170, Auckland, New Zealand; zhangz@landcareresearch.co.nz
Qing-Hai Fan (Raphignathoidea & Tetranychoidea) Plant Health & Environment Laboratory, MAF Biosecurity New Zealand, 231
Morrin Road, St Johns, PO Box 2095, Auckland 1072, New Zealand; qinghai.fan@maf.govt.nz
Vladimir Pesic (fresh water mite families except Limnesiidae, Unionicolidae, Arrenuridae, Pionidae) Department of Biology, Univer-
sity of Montenegro, Cetinjski put b.b., 81000 Podgorica, Montenegro; vladopesic@gmail.com
Harry Smit (fresh water mite families specially Limnesiidae, Unionicolidae, Arrenuridae, Pionidae) Netherlands Centre for Biodiver-
sity Naturalis, P.O. Box 9517, 2300 RA Leiden, The Netherlands; Harry.Smit@ncbnaturalis.nl
Andre V. Bochkov (Cheyletoidea, Myobioidea, Pomerantizioidea, Pterygosomatoidea & Stygmochelidae) Zoological Institute of the
Russian Academy of Sciences, Universitetskaya Embankment 1, 199034 St. Petersburg, Russia; andrevbochkov@gmail.com
A. A. Khaustov (Pygmephoroidea, Acarophenacidae, Caraboacaridae & Trochometridiidae) Nikita Botanical Gardens, National Sci-
entific Center, Yalta, 98648 Ukraine; alkhaustov@mail.ru
Anne Baker (Eupodoidea) Department of Entomology, The Natural History Museum, London SW7 5BD, UK
Andreas Wohltmann (Tanaupodidae, Chyzeriidae, Johnstonianidae, Trombellidae, Microtrombidiidae) Findorffstr. 11, D-27721 Ritter-
hude, Germany; wohltman@uni-bremen.de
Tinghuan Wen (Trombiculidae, Leeuwenhoekiidae & Walchiidae) Shanghai Medical School, Fudan University, Shanghai, China
James W. Amrine (Eriophyoidea) Division of Plant and Soil Sciences, P.O. Box 6108, West Virginia University, Morgantown, WV
26506-6018, USA
P. Beron (Calyptostomatidae, Erythraeidae & Smarididae) National Museum of Natural History, Sofia, Hungary; beron@mail.bg
Jianzhen Lin (Tarsonemidae & Bdellidae) Research Institute of Plant Protection, Fujian Academy of Agricultural Sciences, Fuzhou
350013, China
Grzegorz Gabrys (Microtrombidiidae) Department of Zoology, University of Zielona Góra, Szafrana 1, 65-516 Zielona Góra, Poland;
g.gabrys@wnb.uz.zgora.pl
Robert Husband (Podapolipidae) Biology Department, Adrian College, Adrian, Michigan 49221, U.S.A.; husbandadrian@aol.com