New and Little-Known Ulidiidae (Diptera, Tephritoidea) from Europe

Abstract

New and Little-Known Ulidiidae (Diptera, Tephritoidea) from Europe
New and Little-Known Ulidiidae (Diptera, Tephritoidea) from Europe. Kameneva E. P. — Improved keys to European species of Tetanops and Ulidia are provided. The shape of projections of the phallus glans is found to be essential to distinguish species in the genus Ulidia. Ulidia erythrophthalma (Meigen, 1826), U. albidipennis Loew, 1845, U. nigripennis Loew, 1845, U. parallela Loew, 1845 and U. atrata Loew, 1868 are shown to be separate species and redescribed with the use of genitalic characters. The following synonymy is established: Homalocephala albitarsis Zetterstedt, 1838 = Ortalis diopsides Walker, 1849, syn. n. = Ortalis costalis Walker, 1849, syn. n. Homalocephala apicalis (Wahlberg, 1838) = Psairoptera biseta Frey, 1909, syn. n. = Psairoptera similis Cresson, 1924, syn. n.; Homalocephala mamaevi Krivosheina et Krivosheina, 1995 is recorded for the first time from Europe (Austria, Sweden). Euxesta stigmatias Loew, 1873 is recorded for the first time in the Palaearctic Region based on a single unintentionally introduced specimen from Bulgaria. New finds of other ulidiid species in European countries are listed. Several species previously known only from Europe are recorded from Asia.

Full text

UDC 595.773.4(292.4)
NEW AND LITTLE-KNOWN ULIDIIDAE
(DIPTERA, TEPHRITOIDEA) FROM EUROPE
E. P. Kameneva
Schmalhausen Institute of Zoology, NAS of Ukraine,
Bogdan Chmielnicki str., 15, Kyiv, 01601 Ukraine
E-mail: seioptera@yandex.ru
Accepted 4 August 2008
New and Little-Known Ulidiidae (Diptera, Tephritoidea) from Europe. Kameneva E. P. — Improved keys
to European species of Tetanops and Ulidia are provided. The shape of projections of the phallus glans
is found to be essential to distinguish species in the genus Ulidia. Ulidia erythrophthalma (Meigen,
1826), U. albidipennis Loew, 1845, U. nigripennis Loew, 1845, U. parallela Loew, 1845 and U. atrata
Loew, 1868 are shown to be separate species and redescribed with the use of genitalic characters. The
following synonymy is established: Homalocephala albitarsis Zetterstedt, 1838 = Ortalis diopsides
Walker, 1849, syn. n. = Ortalis costalis Walker, 1849, syn. n. Homalocephala apicalis (Wahlberg, 1838)
= Psairoptera biseta Frey, 1909, syn. n. = Psairoptera similis Cresson, 1924, syn. n.; Homalocephala
mamaevi Krivosheina et Krivosheina, 1995 is recorded for the first time from Europe (Austria, Sweden).
Euxesta stigmatias Loew, 1873 is recorded for the first time in the Palaearctic Region based on a single
unintentionally introduced specimen from Bulgaria. New finds of other ulidiid species in European
countries are listed. Several species previously known only from Europe are recorded from Asia.
Key words: Diptera, Ulidiidae, Europe, new synonymy, new records.
Íîâûå è ìàëîèçâåñòíûå Ulidiidae (Diptera, Tephritoidea) èç Åâðîïû. Êàìåíåâà Å. Ï. — Ïðèâåäåíû
äîïîëíåííûå òàáëèöû äëÿ îïðåäåëåíèÿ åâðîïåéñêèõ âèäîâ Tetanops è Ulidia. Îáíàðóæåíî, ÷òî
ôîðìà âûðîñòîâ ãëàíñà ôàëëþñà ÿâëÿåòñÿ ñóùåñòâåííî âàæíûì ïðèçíàêîì äëÿ ðàçëè÷åíèÿ
âèäîâ ðîäà Ulidia. Ïîêàçàíî, ÷òî Ulidia erythrophthalma Meigen, 1826, U. albidipennis Loew, 1845,
U. nigripennis Loew, 1845, U. parallela Loew, 1845 è U. atrata Loew, 1868 — îòäåëüíûå âèäû; îíè
ïåðåîïèñàíû ñ èñïîëüçîâàíèåì ãåíèòàëüíûõ ïðèçíàêîâ. Óñòàíîâëåíà ñèíîíèìèÿ: Homalocepha-
la albitarsis Zetterstedt, 1838 = Ortalis diopsides Walker, 1849, syn. n. = Ortalis costalis Walker, 1849,
syn. n. Homalocephala apicalis (Wahlberg, 1838) = Psairoptera biseta Frey, 1909, syn. n. = Psairoptera
similis Cresson, 1924, syn. n. Âïåðâûå èç Åâðîïû (Àâñòðèÿ è Øâåöèÿ) îòìå÷åíà Homalocephala
mamaevi Krivosheina et Krivosheina, 1995. Èç Áîëãàðèè âïåðâûå äëÿ Ïàëåàðêòèêè îòìå÷åíà
Euxesta stigmatias Loew, 1873 íà îñíîâàíèè ñëó÷àéíî çàâåçåííîãî åäèíñòâåííîãî ýêçåìïëÿðà.
Ïåðå÷èñëåíû íîâûå íàõîäêè äðóãèõ âèäîâ óëèäèèä èç ñòðàí Åâðîïû. Äëÿ Àçèè âïåðâûå
îòìå÷åí ðÿä âèäîâ, ðàíåå èçâåñòíûõ òîëüêî èç Åâðîïû.
Êëþ÷åûå ñëîâà: Diptera, Ulidiidae, Åâðîïà, íîâàÿ ñèíîíèìèÿ, íîâûå íàõîäêè.
Introduction
The picture-winged flies (Ulidiidae) is a widespread family that includes approximately 110 species in
Europe (about 20% of the World fauna). The most diverse fauna is in southern regions of West and Central
Europe and in the Mediterranean region (Kameneva, Greve, 2004).
In 1998–2008, while preparing and improving the checklist of European Ulidiidae (Kameneva, Greve,
2004) a vast additional material from many countries of Europe was examined. Comparison of the
distributional data against the material deposited in collections added a few new data based on the material
listed below. This is the second article in the series of publications on the ulidiid fauna of Europe. The first
paper (Kameneva, 2007) considered species of the genus Herina
Robineau-Desvoidy, whereas the third
(Kameneva, in prep.) will concern the genus Otites Latreille.
The genus Ulidia Meigen, 1826 was found to include several species indistinguishable from external
characters alone, but differing by the structure of male genitalia (phallus glans). Use of these characters has
resulted in improvements in the key to European species.
Vestnik zoologii, 42(5): 427–454, 2008

Material and methods
The specimens listed in this paper are deposited in the following institutions (ordered alphabetically by
the abbreviations):
American Museum of Natural History, New York City, U.S.A. (AMNH); Academy of Natural
Sciences, Philadelphia’s Natural History Museum, U.S.A. (ANSP); Natural History Museum, London, U.K.
(BMNH); German Entomological Institute (Deutsches Entomologisches Institut), Muncheberg, Germany
(DEI); Museum of Natural History (Muse´um d’histoire naturelle), Gene`ve, Switzerland (MHNG); Museum
of Natural History (Muse´um d’histoire naturelle), Lille, France (MHNLille); Natural History Museum of
Vienna, Austria (Naturhistorisches Museum Wien) (NHMW); National Museum of Natural History
(Muse´um National d’histoire naturelle), Paris, France (MNHNP); National Museum, Prague, Czech
Republic (NMP); Royal Belgian Institute of Natural History, Brussels, Belgium (RBINH); Naturalis, or
Museum of Natural History in Leiden, the Netherlands (Rijksmuseum van Natuurlijke Historie) (RMNH);
I.I.Schmalhausen Institute of Zoology, Kyiv, Ukraine (SIZK); State Museum of Natural History (Staatliches
Museum fur Naturkunde), Karlsruhe (SMNK); State Museum of Natural History (Staatliches Museum fur
Naturkunde), Stuttgart, Germany (SMNS); Tel Aviv University, Israel (TAU); National Museum of Natural
History, Washington, D.C., U.S.A. (USNM); Alexander Koenig Research Museum (Forschungsmuseum
Alexander Koenig), Bonn, Germany (ZFIB); Museum of Natural History of the Humboldt University in
Berlin, Germany (Museum fur Naturkunde der Humboldt-Universitat zu Berlin) (ZMHB); Zoological
Museum University of Copenhagen (Zoologisk Museum, Universitets Copenhagen), Denmark (ZMUC);
Zoological Museum, University of Helsinki, Finland (ZMUH); Museum of Zoology, Lund University,
Sweden (ZIL); Zoological Museum of Russian Academy of Sciences, St. Petersburg (ZISP); Zoological
Museum of M. V. Lomonosov University, Moscow, Russia (ZMUM); Zoologische Staatssammlung,
Munchen, Germany (ZSSM).
Morphological terminology generally follows J. F. McAlpine (1981).
Taxa are listed according to the classification proposed by Kameneva and Korneyev (2006); genera and
species within each tribe are listed alphabetically.
Labels of type specimens are quoted verbatim. The slash character (/) is used to separate lines. The
non-type material is arranged alphabetically by country names, then from the West to the East and from the
North to the South within each country, and finally by the year, month and day of collecting; the collector(s)
name(s) and the abbreviation of depositary follows the list only once if repeated.
Series of photos were taken directly from a dissecting or compound microscope with Nikon 5200 digital
camera and then composed with the use of CombineZM software (Hadley, 2007).
Subfamily Otitinae
Tribe Otitini
Ceroxys baneai Gheorghiu, 1994
No material was available.
Notes. Possibly a synonym of C. urticae (Linnaeus) based on melanistic specimens from Romania.
Ceroxys cinifera Loew, 1846
Material examined. Russia: “Sarepta” [Volgograd], { (DEI); Ukraine: Kherson Region: The
Black Sea Natural Reserve, Ivanivka cordon”, 2.07.1985 (V. Korneyev) (SIZK).
Notes. The material from Ukraine was recently recorded by Kameneva (2002 c) in connection with
general distribution of this species.
Ceroxys fraudulosa Loew, 1864
Loew (1864): Bulgaria; Rondani (1869); Rivosecchi (1995): Italy.
Material examined. Greece: Axios River near Axiopolis, 40°59,298’N, 22°33,299’E, h = 30 m,
swept from reed, 4.06.2002, } (Kameneva) (SIZK); Loutra — Langadas, oestl. Saloniki, 4.05.1942, } (Babiy)
(ZSSM).
Notes. According to Rivosecchi (1995), rather common in mainland Italy, from Venice to Campania,
and also in Sicily. First record from Greece.
Ceroxys hortulana (Rossi, 1790)
Jaroszewski (1880): Ukraine (“Kharkov and vicinity”, as “Ceroxys hyalinata Panzer”).
Material examined. Bosnia: “oriental.”, 1895, { (Mik); Kupres, { (NHMW); Croatia:
“Kroatien”, with number 19034, { (Becker), and label “Ceroxys / hyalinata / J. Panzer” (Becker det.);
428
E. P. Kameneva

“Dalmatien”, with numbers 22984, 22985, 22987, 22990, 22993, 22995, 25.05.[year?], 3 {, 3 } (Becker)
(ZMHB); France: “Korsika / 54680. V”, 2 { (Becker) (ZMHB); Romania: “Orsova / V. 37600” (Becker)
(ZMHB); Moldova: Ceobruci [46°36’N, 39°41’E], 2.04.1918, { (Paramonov) (SIZK); Ukraine: Cherkasy
Region:mouth of Ros’ River, 9.04.1988, { (Zrazhevsky); “manast. Korsunskij, cursus inf. fl. Dniepr”,
15.10.1930, { (S. Medvedev); Kirovograd Region: “Znamenka, distr. Alexandria”, 11.04.1927, }
(Paramonov); Mykolaiv Region: Chichikleya River, near Veselinovo, 15.08.1991, 2 { (V. Korneyev);
Kherson Region, Yagorlyk Bay shore nr. Ivanivka, 2.07.1985, } (V. Korneyev); Kyiv Region: Shandra (as
“Øàíäîðîâêà”), 12.05.1922, } [Paramonov] (SIZK); Odessa Region: Belgorod-Dnestrovsky (as “Belgrad
Bessarab.”), 6.08.1937, 2 } (Gotz) (SMNS); “Valegotsulovo, d. Balta, g. Odessa”, 17.05.1925, {; “Ananjev
prope Odessa”, 20.07.1932, } (Paramonov); Kuchurgan [46°44’N, 29°58’E], 10.04.1919, 14 {, 4 }; Yasski,
[46°30’N, 30°05’E], 11.01[sic!].1921, } [Paramonov]; Zaporizha Region: Vodyanoye, sands [47°29’N,
34°29’E], 25.09.1930, 2 { (S. Medvedev); Krul’man River below Kamenskoye, [47°32’N, 35°20’E],
30.09.1992, } (V. Korneyev); Altahir near Melitopol, 4.06.1977, { (Djafarov) (SIZK).
Notes. First record from Bosnia.
Ceroxys munda (Loew, 1868)
Hennig (1939): European Russia (Rostov and Volgograd Regions); Ukraine; Kazakhstan; China (Xingjian).
Material examined. Ukraine: Vinnytsya Region: Popelyukhy, 7.07.1930, } (Pasyuchnyk); Kyiv
Region: Boyarka, 10.06.1928, {; Malyutinka, 14, 19.06.1927, 2 } (Panocini); Kyiv: Kyrylivskyi Hay,
11.06.2003, 2 } (V. Korneyev); Baykove Cemetery, 7.08.2003, { (Verves); Cherkasy Region: Trakhtemyriv,
24.06.1987, } (Verves); Kaniv, Natural Reserve, cesspool, 21.06.1961, 2 }; at pig farm, 22.06.1961, 2 }
(O. Viktorov); Kaniv Natural Reserve: 26.06.1985, } (Verves); 28.06.1988, } (Zrazhevsky); Tyasmyn River
45 km SE of Cherkasy, 5.07.1988, { (Zrazhevsky); Poltava Region, Bugaevka, [49°23’N, 32°55’E],
18.06.1989, { (Zrazhevsky); Odessa Region: Ananyev, 1, 4, 11–15, 19.06.1931, 6 {, 15 } (Paramonov);
Kherson Region: Askania Nova», 14.06.1927, 2 {, 3 } (Charleman and Schepe); Zaporizha Region: Altahir
near Melitopol, 4.06.1977, } (Djafarov); Berdyansk (as “Osipenko”), 4.06.1940, 2 { (Gulinov); Lugansk
Region: Lugansk, 28.06.1929, 1 specimen (abdomen lost) (Talitsky) (SIZK); Russia: Rostov Region:
Taganrog, 12.07.1923, {; 26.05.1921, } (collector unknown) (SIZK).
Ceroxys urticae (Linnaeus, 1758)
Jaroszewski (1877): Ukraine (“Kharkov and vicinity”); Hennig (1939): Sweden, Finland, England, France,
Hungary, Italy, European Russia (Volgograd and Astrakhan Regions); Kazakhstan, Turkmenistan, China;
Egypt.
Material examined. Bosnia: “oriental”, 1895, { (Mik) (NHMW); Romania: Iasi Co., David Vy.,
11 km W of Iasi, 60 m, 20–22.06.1980, } (Hepper) (USNM); Ukraine: Odessa Region: Valegotsulovo,
26.05.1925, {; Ananjev, 4.06.1931, } (Paramonov); Kuchurgan [46°44’N, 29°58’E], 12.08.1920, {; Odessa,
24.08.1918, } [Paramonov]; Mykolayiv Region: Chichikleya River nr. Veselinovo, 15.08.1991, 4 {
(V. Korneyev); Kherson Region: the Black Sea Natural Biosphere Reserve, Solonoozerna area, 3.06.1981, {
(Rayevsky); Yagorlyk Gulf, shore near Ivanivka vill., 2.07.1985, } (V. Korneyev); Donetsk Region: “Bezy-
mennoje, sea shore”, } (no date, no collector) (SIZK). Russia: Murmansk Region: Olenitsa nr. Kandalaksha,
12–14.08.1980, 6 {, 2 } (Kameneva) (SIZK).
Notes. First record from Bosnia and North of European Russia.
Dorycera graminum graminum (Fabricius, 1794)
Hennig (1939): France (Corsica incl.), Austria, Croatia, England, Italy, Portugal, Spain, Turkey (Asia
Minor). Soo´s (1980): Hungary; Roha´c

ek (2006): Slovakia.
Material examined. Austria: “Wien”, { (Schiner) (Mik det.) (NHMW); Germany: “Scatoph.
gr. Fb. German.”, “Coll. Wiedem.”, { (collector unknown) (NHMW); Hungary: “Ungarische Flugsand-
stegg”[Hungarian sand dune belt], { (Sa´lo`) (SMNS); France: Hautes-Alpes, 1 km NW Upaix, 25.05.2002,
} (Tschorsnig) (SMNS); Spain: “Arragonia / Albarrazin”, 23.05.1925, {; 27. 05.1925, 2 }; 10.06.1925, }
(collector unknown) (SIZK); Croatia: Rovinj, 4.06.1961, } (ZFIB); “Dalmatien / Alte Sammlung”, 4 }
(collector unknown) (Schiner det.) (NHMW).
Notes. This species is listed in the German and Polish Checklists of Diptera (Martinek, 1999: 169;
Nowakowski, 1991: 174) without further details; so far, I have not seen any specimens from Poland and consider
this record unsubstantiated. The male from Wiedemann’s collection is the only known German specimen.
Dorycera hybrida Loew, 1862
Hennig (1939): Bulgaria (terra typica: Varna), Greece Mainland and Crete, France, Turkey (European Part),
Ukraine (North of Odessa); Lebanon.
429
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

Material examined. Non-type. Greece: “Lassethe [Lassiti?] Plain, Crete, circa 2.820 ft. 1904”,
“Miss D.M.A.Bate, 1905–25”, “Coll. Hendel”, {, } (NHMW). Ukraine: Odessa Region, Velykodolynske
[46°19’ N 30°34’ E], 3.05.1986, 2 {; } (Korneyev) (SIZK). Lebanon: “Beyrouth”, “Coll. Hendel”, 2 }, {
(NHMW).
Notes. Some specimens from Loew’s collection (ZMHB) labelled as “Varna” were possibly
collected in Turkey, as they are marked with the colour labels, which refer to Asia Minor coast of the Black
Sea, where Loew also collected (Korneyev, pers. comm.). The Bulgarian record (and terra typica!) needs no
further confirmation, as it lies within the area of confirmed distribution of this species. Material with cyrillic
label from “Krupaz, Gouv. Charkow [Ukraine] oder Kursk [Russia]” mentioned by Hennig (1939) has not
been located so far, and the record from Russia is therefore unsubstantiated.
Melieria acuticornis (Loew, 1854)
Loew, 1854 (Ortalis). Jaroszewski (1877): Ukraine (“Kharkov and vicinity”, as “Ceroxys acuticornis”).
Material examined. Type. Syntypes Ortalis acuticornis: [Hungary?:] “Ungarn / v. Friv.”, “Coll. /
H. Loew”, 2 } (ZMHB), with labels: “Type [red label]” and “Syntypus / des. Kameneva”. Non-type. Austria:
Wien, “Coll. / H. Loew”, { with labels: “Type [red label]” and “acuti / cornis / Lw.” (Schiner) (ZMHB)
[not a type of O. acuticornis]; Croatia: “Dalmatien / 30077”, “Sammlung / Dr. Th. Becker”, { with labels:
“Type [red label]” and “obscuricornis Beck.”, “Melieria acuticornis Loew / nicht Typus von obscuricotnis /
Beck. (Typus sammte / aus Tibet!)” (ZMHB) [not a type of M. obscuricornis]; Hungary: “Ungarn / 53201”,
“Sammlung / Dr. Th. Becker”, }; Budapest / 40788”, “Sammlung / Dr. Th. Becker } (ZMHB); Italy:
“Sicilien”, “Coll. / H. Loew”, }, with labels: “Type” [red label], “Ortalis / bifasciata / Lw.”, “nicht Typus
von / Tepronota bifasciata / son vorn eine / Melieria-Art!” [Hennig’s remark]; [Fasedo], “20”, “Coll. /
H. Loew } (ZMHB); Poland: “Breslau / 7.05” [Wroclaw] “Coll. H. Loew” { (ZMHB); Romania: “Ba´zia´s,
Banat, Holtz”, “S. Ungarn / 61567. V.”, “Sammlung / Dr. Th. Becker { (ZMHB); Russia: Volgograd:
“Sarepta / Becker”, “Coll. / H. Loew”, {; “Sarepta / Christoph”, “Coll. / H. Loew”, {; “Sarepta /
Christoph”, “Coll. / H. Loew” {, } (ZMHB) [all these specimens from Sarepta also bear red labels “Type”,
and the latter two also Loew’s handwriting “Ortalis / acuticornis / m.”, but none of them actually is a type
specimen]; “Sarepta”, “Coll. / H. Loew”, {, “varie / tas”; “Sarepta / 36628”, “Sammlung / Dr. Th. Becker”
{, 2 }; “Orenburg / 58043. VI.”, “Sammlung / Dr. Th. Becker”, 2 { (ZMHB); Ukraine: Kherson Region:
Yagorlyk Gulf, shore near Ivanivka vill., 2.07.1985, } (Korneyev); Askania-Nova, 14, 27.06.1927, {, 3 }
(Charleman, Schepe) (SIZK); Odessa, 1893, { (Horvath) (NHMW).
Notes. First record from Italy (Sicily).
Melieria cana Loew, 1858
Karpa et al. (2005): Latvia.
Material examined. Type. Syntype {: [Italy:]“Friuli-Venezia Giuli, Zaule” [Zante prope
Tergestum = Aquilinia nr. Trieste], “Coll. / H. Loew”, “Type” [red label], “Ortalis / cana / Lw.”,
“Syntypus” [red label] (ZMHB). Non-type. Austria: “Mont. / Majgen / Stiria”, { (Mann) (NHMW); Italy:
Venezia, Valle morosina, 3.06.1932, { (Gridelli) (ZMHB); Greece: Paralia near Epanomi, 23,1 km S of
Thessaloniki, 40°24,699’N, 22°53,383’E, sea shore, h = 0 m, 29.05.2002, 3 { (Kameneva, V. Korneyev, S.
Korneyev) (SIZK); Moldova: Comrat, 6.06.1988, 17 {, 14 { (V. Korneyev) (SIZK); Romania:
“Transylvania, Siebenburgen”, “7. 39092”, “Sammlung / Dr. Th. Becker”, 5 {, 6 } (ZMHB); Histria,
Salicornia, 10.06.1973, } (Zwolfer) (SMNS); Russia: Orenburg Region, “Tolstinskij Distr. Troitsk, prov.
Ural”, 20–28.07.1926, } (Argyropulo) (ZMUM); Volgograd, “Sarepta”, “ Coll. / H. Loew”, 2 } (Becker)
(ZMHB); “Sarepta / 33587” and “Sarepta / 30327”, “Sammlung / Dr. Th. Becker”, {, }, with red label
“Typus” and “sareptae / Beck.” (ZMHB); Spain: Pr. Zaragoza, Pina de Ebro, 15 km W Trockental,
19.05.1992, 2 } (Osten) (SMNS); Ukraine: Kherson Region, Yagorlyk Bay shore nr. Ivanivka , 2.07.1985, {
(V. Korneyev); the Black Sea Natural Biosphere Reserve, Ivano-Rybalche area, 7.08.1985, {, } (Kameneva);
Donetsk Region: sea shore nr. Bezimennoye [47°06’N 37°55’E] (no date, no collector’s name); Crimea:
Theodosia, 19.05.1989, 3 {, 2 } (Bilashivsky) (SIZK).
Notes. First record from Romania. The name “sareptae” is an unpublished manuscript name.
Melieria crassipennis (Fabricius, 1794)
Jaroszewski (1877): Ukraine (“Kharkov Gouvern.”, as “Ceroxys crassipennis”)
Material examined. Belarus: “Lithuania / Minsk”, 1909, 2 } (NHMW); Russia: “Sarepta /
33581”, “Sammlung / Dr. Th. Becker”, {, 2 } (ZSSM) (as “Melieria etrusca Rondani”, Becker det.);
Moscow Region: Moskva River nr. Tuchkovo, 7–9.07.1983, 2 { (A. Rasnitsyn) (SIZK); Ukraine: Zhytomyr
Region: Zhytomyr, 19.08.1922, { (Prozhyga); Glezno [49°58’N 27°51’E] 2.07.1926, { (Gensicky);
Vyshevichy, Radomyshl Distr. [50°38’N 29°24’E] 16–17.06.1917, 3 { [Paramonov?]; Kyiv Region: Kontsha-
Zaspa near Kyiv, 15.061991, 2 {, } (V. Korneyev), Khodosievka, meadow, 16.07.1985, {, 2 } (V. Korneyev);
430
E. P. Kameneva

Vasylkiv Distr.: Zvonkovaya, 22.06.1925, { [Paramonov?]; Cherkasy Region: Trakhtemyriv 30 km N of Kaniv
[49°29’N 31°20’E], 4.07.1988, } (Zrazhevsky); Tubiltsi 20 km SE of Kaniv, 19.06.1988, } (Zrazhevsky);
Krapivna [49°38’N 32°09’E], swampy meadow, 16.06.1988, } (Zrazhevsky); Poltava Region: “Chatki, distr
Poltava.” 28.07.1926, {, 2 } (Kistiakovsky); Dnepropetrovsk Region: Sinelnikovo, 15.06.1930, } (Talitsky)
(SIZK).
Notes. First record from Belarus. The specimens from Sarepta certainly do not belong to “M. etrusca”,
which is the junior synonym of M. omissa (Meigen) (see below).
Melieria nana (Loew, 1873)
Material examined. Type. Holotype }: Spain: “Spanien”, “Coll. / H. Loew”, “Type” [red label],
“na- / nus / Lw.” (ZMHB).
Notes. This species is still known from the holotype only.
Melieria nigritarsis Becker, 1903
Material examined. Ukraine: Zaporizha Region: Berdyansk, 08.1954, } (no collector); Donetsk
Region: “Bezymennoje, sea shore”, } (no collector) (SIZK).
Notes. First record from Ukraine. Additional non-European specimens have been examined in this
study, among them, a syntype from Egypt and a non-type specimen from Kaschgar (Western China:
Xingjian) (ZMHB) and Shadegan (Iran: Khuzistan) (SMNS).
Melieria omissa (Meigen, 1826)
Jaroszewski (1877): Ukraine (“Kharkov and vicinity”, as “Ceroxys omissa”). — Melieria etrusca Rondani,
1869; Soo´s (1983): synonymy.
Material examined. Greece: Apollonia, 40°39,036’N, 23°29,667’E, relict forest at Volvi Lake,
h = 60 m, 30.05.2002, } (Kameneva, V. Korneyev, S. Korneyev) (SIZK); Latvia: Randu, 12.06.1990,
28.06.1994, {, 2 } (Karpa) (SIZK); Moldova: Cimis´eni nr. Vadul-lui-Vode, 22.07.1988, 2 } (V. Korneyev)
(SIZK); Ukraine: Cherkasy Region: Trakhtemyriv 30 km N of Kaniv [49°29’N 31°20’E], 20.06.1980, {;
Kaniv, 20.06.1980, } (Verves); Kherson Region, Yagorlyk Bay shore nr. Ivanivka, 2.07.1985, {
(V. Korneyev) (SIZK).
Notes. First records from Greece and Moldova.
Melieria parmensis Rondani, 1869
Material examined. France: [Basses Alpes Frankreich] “Digne VI / 56106”, { (ZMHB);
Germany: Baden-Wurttemberg, “Innufer bei Nussdorf”, 30.06.1917, {, idem, 14.07.1917, 9 specimens, idem,
2.08.1919, 18 specimens (collector unknown) (SMNS); Thuringen: “Artern, P. — Sach.[sen], 13.06.1915 3 {,
2 } (ZMHB); Austria: Innsbruck, 14.06 “8 943” [Becker’s manuscript catalogue of collection: “Uper / zun
/ Anhohem” (???)], 6 {, 2 } (Becker) (ZMHB); Salzburg, 20.06.1885, 12 {, 5 } (Mik) (MLUH), idem,
same data, 2 {, } (ZMHB); Switzerland: Allondon, 20.09.1974, 4 specimens (Tournier) (RBINH); Italy:
“Bozen” [= Bolzano], 10.06.1914, 8 specimens (SMNS), idem, 13.06.1915, 4 {, 4 } (Duda) (ZMHB);
Sondrio, 1.05.1898, 2 {, } (Becker) (ZMHB).
Notes. B. Merz (1996) recorded M. parmensis from France and Switzerland (as Hypochra parmensis).
For general distribution (including Transcaucasia and Central Asia) and taxonomic remarks see
E. P. Kameneva (1997). It was recorded from Germany by Dunk (1996) and included into the “Katalog der
Dipteren Bayerns” (Schacht, 2007) based on that publication. Here, two additional occurrences, from
Baden-Wurttemberg and Thuringen, are recorded.
Melieria picta (Meigen, 1826)
Material examined. Austria: “Austria / Coll. Egger”, { (NHMW); Bulgaria: Kiten, 42°14’N /
27°48’E, 17.07.1987, 3 { (Bartak) (ZSSM); Croatia: “Dalm., Ins. Arbe / Sta, Eufemia”, 2.07.1934, }
(Zerny); France: “Paris / Fairm.”, “Coll. H. Loew”, { (ZMHB); Languedoc-Roussillon: Aude, Etang de
Gruissan S of Narbonne, 0 m, 7–10.06.1980, { (Schacht) (ZSSM); Germany: Halle, “2752”, 2 { (Erichson)
(ZMHB); Italy: Zaule, “Coll. / H. Loew”, {, 06.1858 (ZMHB); Moldova: Comrat, 6.06.1988, {
(Kameneva); Ialpujeli River nr. Comrat, 12.07.1988, {, 3 }; Borotani, 6.06.1988, 4 {, 4 }; Taraklia, 5 {
(Korneyev) (SIZK); Romania: “Moldavie / vall. du Berlad”, } (Montandon) (NHMW); idem, 2 specimens
(DEI); Russia: “Sarepta / Christoph”, “Coll. / H. Loew”, 4 { (ZMHB); Ukraine: “Podolien, Krywcze kr.
Borszczow” [Krivche nr. Borschiv], 5.05–15.06.1936, } (Toll) (ZMHB); Mykolaiv Region: Elanets Distr.,
Kalinovka, 22.06.1984, 2 { (Karachevskaya); Chichikleya River nr. Veselinovo, 15.08.1991, {; Kherson
Region: the Black Sea Natural Biosphere Reserve, Ivano-Rybalche area, 7.08.1985, 2 { (V. Korneyev);
Crimea: Theodosia, 19.05.1989, 6 {, 9 } (Bilashivsky) (SIZK).
431
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

Notes. First record from Bulgaria. Widespread throughout the Palaearctic Region to Alaska and
Canada (Nearctic).
Melieria unicolor Loew, 1854
Material examined. Type. Holotype: {: “Ungarn / v. Friv. ”, “Coll. / H. Loew”, “Type” [red
label], “uni- / color / Lw. ”, “Ortalis / unicolor / m. ” (ZMHB). Non-type. Spain: C. B. Estartit, 9.05.1973,
{ (Harde) (SMNS); Italy: Sicily, 1858, } (Mann) (NHMW).
Notes. Type locality (“Ungarn”) at the time of description also included western Romania and a small
part of Serbia. This record might belong to anyone of them. Soo´s (1980) did not mention it. This species was
recorded for the first time from Spain based on material listed above (Carles-Tolra´, Kameneva, 2008). First
record from Italy (Sicily).
Tetanops Falle´n, 1820
Seven species are known in Europe. W. Hennig (1939) keyed 5 of them, except
T. psammophila Loew and T. corsicana Becker, which he considered subspecies of
T. flavescens Macquart. As the two latter taxa are found to occur sympatrically, further
revision of their status is necessary. Here, meanwhile, they are treated as species.
The European species of Tetanops can be recognized with the following key mo-
dified from W. Hennig (1939).
Key to species of Palaearctic Tetanops
Òàáëèöà äëÿ îïðåäåëåíèÿ ïàëåàðêòè÷åñêèõ âèäîâ ðîäà Tetanops
1. Wing entirely hyaline, short (not exceeding abdominal apex in female); thorax and abdomen black,
densely microtrichose, shiny black dotted. .................................................. T. (s. str.) laticeps Loew
— Wing at least with faint brown spot, long (far exceeding abdomen apex); abdomen yellow or black,
shiny or microtrichose, but not dotted. .......................................................................................... 2.
2. Two pairs of dorsocentral setae; frons and mesonotum uniformly gray microtrichose, non-dotted.....
.......................................................................................................................................................... 3.
— Only one pair of dorsocentral setae; frons and mesonotum gray microtrichose, with numerous bare
dots around setulae. ........................................................................................................................ 5.
3. Abdomen with non-microtrichose areas yellow; male surstylus with 3–4 prensisetae. ........................
................................................................................................................ T. (s. str.) corsicana Becker
— Abdomen with non-microtrichose areas dark brown to black. ........................................................ 4.
4. Wing apex with entire brown apical band; male surstylus with 2 prensisetae. ....................................
............................................................................................................ T. (s. str.) flavescens Macquart
— Wing apex with 2 brown spots at R
2+3
and R
4+5
apices more or less separated by hyaline area; male
surstylus with 5–8 prensisetae. ............................................................ T. (s. str.) psammophila Loew
5. Parafacial smooth, entirely microtrichose; frons conspicuously protruding anteriorly, mostly gray
microtrichose with numerous bare dots; wings with several faint brown spots, at least in pterostigma
and at R
2+3
and R
4+5
apices; abdomen shining black, with gray microtrichia on some parts. ....... 6.
— Parafacial wrinkled or pitted, shining in anterior half, frons only slightly protruding anteriorly, mostly
shiny and bare; wing with one faint spot at radial fork, posterior to pterostigma; abdomen shiny black,
non-tomentose. ........................................................................ T. (Eurycephalomyia) sintenisi Becker
6. Frons without longitudinal bare vittae; eye wide oval, 0.75–0.80 times as long as wide. ..................
.................................................................................................................. T. (s. str.) myopina Falle´n
— Frons with pair of bare vittae in anterior portion; eye narrow oval, 0.60–0.65 times as long as wide.
.............................................................................................................. T. (s. str.) contarinii Rondani
Tetanops contarinii Rondani, 1869
Hennig (1939): Italy.
Material examined. Non-type. Spain: C. B. Estartid, 12.09.1973, } (Harde) (SMNS); Pr. Sevilla:
El Rompido b. Huelva: Ortsrand, Sanddune, 0–10 m, 17.09.1974, }; Camping “Catapum”, 50 m,
15–22.09.1974, { (Amsel, Roesler) (SMNK).
Notes. This species was recorded for the first time from Spain based on material listed above (Carles-
Tolra´, Kameneva, 2008).
Tetanops corsicana Becker, 1909
Hennig (1939): France: Corsica and southern mainland; Italy: Sicily.
Material examined. Type. Holotype {: “Corsica / 39357”, “TYPUS” (ZMHB). Non-type. Spain:
C. B. Estartid, 28.05.1974, {, 3 } (Harde) (SMNS); Mallorca: Aleudia bantal, 28.05.–9.06.1956, 23 {, 12 }
432
E. P. Kameneva

(Bequaert) (RBINH); France: Bouches-du-Rh
^
one, Salin de Giraud, 29.05.1995, { (Merz, Eggenberger)
(MHNG); [Languedoc-Raussellon:] St.-Cyprien, 12.06.1954, 2 {, 2 } (Bequaert) (RBINH); Tunisia:
Tabarca area, 7–18.05.1988, 3 {, } (ZMUC).
Notes. The species was recorded for the first time from Spain based on material listed above (Carles-
Tolra´, Kameneva, 2008). The specimen from Tunisia represents the first record beyond Europe.
All the males listed have non-tomentose parts of abdomen yellow, contrary to the black abdomen found
in T. flavescens Macquart (and T. psammophila Loew). The coastal distribution of T. corsicana largely
coincides with that of T. flavescens Macquart. Although its “subspecies” rank must be abandoned, the status
of T. corsicana and T. flavescens need further revision to clarify the position of these previously synonymous
species. For taxonomic comments on the new status of T. corsicana see L. Rivosecchi (1995).
Tetanops flavescens Macquart, 1835
Syn.: Tetanops impunctata Loew, 1854; Rondani (1869).
Material examined. Type. Syntype Tetanops impunctata: Switzerland: “Zurich / Bremi”, “Coll. /
H. Loew”, “Type”, { (ZMHB). Non-type. Portugal: “Lusitan. Hffnsg” [= Portugal, Hoffmannsegg leg.],
“2747”, “Type”, “impunctata / Loew* / buccata N. ant.”, {; “T. Eryngii / Ldf. / Ztt. occ.”, “Coll. /
H. Loew”, “Type”, 2 {, 2 }; “Terellia / eryngii / [unreadible: Dufour?], “impun / ctata / Lw.”, “Coll. /
H. Loew”, ‘Type”, {, } (ZMHB); “T. impunctata / det. Becker”, “T. Eryngii / Ztt. occ. 201”, 3 {, 3 };
France: “Provence / Frejns (Var)”, 9.06.1924, 3 } (Zerny) (NHMW).
Notes. Type } of Tetanops flavescens has not been located either in the collection of MNHN Paris or
in MHN Lille (V. A. Korneyev, pers. comm.). J. P. M. Macquart (1835: 423) recorded its type locality as
“environs de Paris”, but no additional material from Northern France is known to me. E. Se´guy (1934: 54)
lists Bayonne (Southern France, Atlantic coast), Lyon, “Switzerland” (after Pandelle´, no further details) and
Tanger (Morocco) as additional records of this species. According to B. Merz (1996: 410), the origin of the
T. impunctata type from “Zurich” is dubious; it might simply indicate the origin of that specimen from
Bremi’s collection, as he lived in Zurich. L. Rivosecchi (1995) redescribed this species (including male
genitalia) based on specimens from Toscana (Italy) from Rondani’s collection, and B. Merz (1996) recorded
it from Bouches-du-Rh
^
one, and St. — Cyprien Plage (as “Plague”), the two localities, from which also
T. corsicana is recorded above. The record from Bouches-du-Rh
^
one is based, indeed, on a misidentified
specimen of T. corsicana (B. Merz, pers. comm.).
All the specimens examined have non-tomentose parts of the abdomen black and without a bright
yellow spot ventral to the eye margin, differing in that way from T. corsicana, as figured by L. Rivosecchi (1995).
Tetanops laticeps Loew, 1868
Material examined. Type. }: Russia: “Sarepta / Christoph”, “Coll. / H. Loew”, “Type” [red
label], “lati- / ceps / m.” (ZMHB). Non-type. Non-European. Iran: Khuzistan, Shadegan, 15–23.02.1936, }
(Richter, Schaufelle), (SMNS); Turkmenistan: Ashhabat, 22.04.1926, } (Paramonov) (SIZK).
Notes. First records from Iran and Turkmenistan.
Tetanops myopina Falle´n, 1820
Jaroszewski (1876): Ukraine (“Kharkov and vicinity”). Hennig (1939): Northern Russia (Leningrad Region;
Karelia). Kabos, van Aartsen, 1984: the Netherlands.
Material examined. Estonia: “Dorpat” [= Tartu], 3.07.1886, 3 {, 3 } (NHMW); Ukraine: Kyiv:
“Zhukiv Khutir”, 30.05.1972, } (Verves); Cherkasy Region: Kaniv, vicinity, Keleberda, 12.06.1986, 5 {
(Korneyev); Korobivka, 4.06.1989, } (Zrazhevsky) (SIZK).
Notes. A common species in the sand dunes in all the countries along the North and Baltic Seas, and
also along rivers and lakes, including Northern Ukraine. Dutch, Lithuanian and Russian records were omitted
and must be added to the Fauna Europaea checklist.
Tetanops psammophila Loew, 1862
Material examined. Type. {: Bulgaria: “Varna”, “Coll. / H. Loew”, “Type” [red label],
“psammo- / phila / Lw.”; “Nesebar, diunite”, 31.09.1972, 2 {, 2 } (Beschovsky); “Kam. Perla / Elymus”,
27.05.1973, 4 {, 3 } (Beschovsky) (ZMHB).
Notes. For taxonomic comments see V. L. Beschovsky (1974).
Tetanops sintenisi Becker, 1909
Hennig (1939): Russia (Yaroslavl; vicinity of St. Petersburg; Far East); Poland; Latvia; Ukraine (“Kurjach.,
Gouv. Charkov” = Kurjachivka, northern part of Lugansk Region); China; Lobanov (1972): European
Russia (Ivanovo); Smit (2005): the Netherlands; Stuke, Merz (2006): Germany.
433
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

Material examined. Ukraine: Kyiv: “Prope Kyiv”, 16.06.1928, { [Paramonov]; near Lysa Hora,
19.06.1989, } (Korneyev) (SIZK).
Notes. This species belongs to the subgenus Eurycephalomyia Hendel, which includes one other
species, T. myopaeformis (Roder) known to be a sugar-beet pest in the United States of America; the
morphological differences between specimens from the New and Old World populations need further study.
Ulidiopsis mirabilis Hennig, 1939
Material examined. Type. Holotype {: [Greece: ] “Saloniki / 26 303”, “HOLOTYPUS” [red
label], “Ulidiopsis / mirabilis n. g. / n. sp. / det. Dr. W. Hennig 1939” (ZMHB). Non-type. Non-European.
Turkey: Pozanti-Konya, Affyon, 13.05.1968, } (Lindner) (SMNS); W Turkey: Miletus, nr Lake Bafa,
saltmarsh, 28.04.1998, { (Ackland) (MHNG).
Notes. This species was known from the holotype only. Here it is recorded, for the first time since
description, from Asian Turkey.
Tribe Cephaliini
Cephalia rufipes Meigen, 1826
Becker (1902): remark on the absence of types both in Paris and Vienna collection; Schiner (1864): Austria
(Prater, the park on Danube islands in Vienna); Hennig (1939): France, Germany, Italy, Spain; Soo´s (1957):
Hungary; Roha´c

ek (2006): Czech Republic (Bohemia), Slovakia; van Aartsen, Beuk (2002): Netherlands;
Myrmecomyia rufipes: Se´guy (1934). — Cephalia nigripes Meigen, 1826; Se´guy (1934): as “variation” of
rufipes.
Material examined. Type. Syntype (?) } Cephalia rufipes “Meigen \ 24[6?]0” [paper circle],
“2247”, “Cephalia / rufipes” [old paper rectangle, ink handwriting] (MNHNP). Syntype } Cephalia nigripes
[Germany:] “Aachen”, “Alte Sammlung”, “Cephalia / nigripes / M. / v. 29 / 7 A” [paper square, ink
handwriting] [Bamhauer] (NHMW). Non-type. Austria: “Austria / coll. Egger”, {, } “rufipes // det.
Schiner”; “Schin. 1869” }; “Alte Sammlung”, 19 specimens (C. rufipes det. Schiner and Hendel) (NHMW);
Wien. 08.1861, “coll. H. Loew”, {; “Austria, Brauer”, “coll. H. Loew”, } (ZMHB); France: Rambuiett,
4.07.1900, {, 2.07.1946, } (RBINH); “Cephalia // rufipes // Lyon”, “rufipes // coll. Winthem”, }
(NHMW); Pyrene´es-Orientales, 610m, Can Baills, 10km SW Thuir, 42.34N/02.39E, 8.06.2007, } (Merz)
(MHNG). France/Spain(?): “Pyrenaei Keitel”, “6626”, } (“rufipes / Meig.”) (ZMHB); Germany:
Karlsruhe, 30.07.72, } (Stritt) (SMNK); Spain: Pr. Cadiz, Hozgarganta-Tal bei Jimena 200m, 17.07.1979,
} (Schacht) (ZSSM); Pr. Salamanca, Villar de Ciervo, Las Coronas, 18.06–8.07.1995, } (Tschorsnig)
(SMNS); Switzerland: “Basel Imhot / v. Roser Coll.”, 2 {, } (“Myrmecomyia rufipes Mg.” [det. Roser]
(SMNS).
Notes. J. W. Meigen (1826: 294) wrote concerning the syntype females of C. rufipes: “Ich erhielt das
Weibchen von Herrn Medizinalrath Klug in Berlin; ein anderes schifte Dr. Megerle v. Muhlfeld als osterreich.
Produkt”. The female in Meigen’s collection (MNHNP) placed under C. rufipes (No. 2247) does not fit the
original description of that species; it has a uniformly black mesonotum, and Becker (1902) noted that it has
entirely black legs (like in “nigripes”), but determined it as a female C. nigripes type specimen, which is
obviously an error. The holotype female of C. nigripes (Meigen, 1826: “Herr Baumhauer fing das Weibchen
im August am Lustberge bei Aachen”) is in NHMW collection, but the only difference from the original data
is the month of collecting. The specimen from Lyon (Winthem’s coll., NHMW) clearly has nothing to do
with the original syntypes.
Tribe Myennidini
Callopistromyia annulipes (Loew, 1873)
Kameneva, Korneyev (2006): Canada, USA; Merz (2008): Switzerland; Merz, van Gyseghem (2008):
Germany.
Notes. This species is widespread in the Nearctic Region and was unintentionally introduced to
Europe recently.
Myennis octopunctata (Coquebert, 1798)
Musca octopunctata Coquebert, 1798; Coquebert (1804); Myennis octopunctata:Se´guy (1934): France
mainland; Hennig (1939): England, France (Corsica), Germany, Italy, Poland, Romania, European Russia
(Sarepta), Ukraine (records from Middle Asia and Far East Russia partly belong to other species); Soo´s
(1957): Hungary, Transpolar Russia (Franz-Josef Land), Turkey (Asia Minor: Konia); Merz (1996):
Switzerland; Roha´c

ek (2006): Czech Republic, Slovakia; Krivosheina, Krivosheina (1997): European
Kazakhstan (right bank of the Ural); European Russia: Central (Moscow); Asian Russia: West Siberia, Tyva;
van Aartsen, Beuk (2002): Netherlands; Carles-Tolra´, Baez (2002): Andorra, Spain. — Scatophaga fasciata
434
E. P. Kameneva

Fabricius, 1805; Ortalis fasciata: Macquart (1835); Myennis fasciata: Schiner (1864): Austria; Se´guy
(1934). — Myennis fasciata Robineau-Desvoidy, 1830.
Material examined. Type. Possible syntype [sex?] of both Musca octopunctata and Scatophaga
fasciata: [France:] “Gallia” [paper piece of irregular shape; handwriting hardly readable; 2 wings on a piece
of carton] (ZMUC). Non-type. France: Corsica: Ajaccio, 06.1899, 4 } (MTD); Greece: Apollonia,
40°39,036’N, 23°29,667’E, relict forest at Volvi Lake, h = 60 m, on fallen poplar, 30.05.2002, {, }; Ioanina,
39°41,296’N, 20°50,319’E, h = 450 m, 12.06.2002, 5 {, 33 } (Kameneva, V. Korneyev, S. Korneyev)
(SIZK); Russia: Orenburg, woods in the flood plain of the Ural River, 2.06.1985, 7 {, 6 } (Ermolenko);
Rostov Region: Taganrog, 26.05.1928, 2 { (collector’s name unreadible) (SIZK); Switzerland: Chamcy near
Geneva, 25.07.2004, 10 } (Kameneva) (SIZK); Ukraine: Kyiv: Lysa Hora [50°23’50”N 30°33’16”E],
4.07.2000, 10 {, 10 } (V. Korneyev, Kameneva); Kyrylivs’kyi Hay [50°28’39”N 30°28’21”E], 11.06.2003,
11 }, 27.06.2003, {, } (V. Korneyev, Kameneva); Lukyanivka [50°28’09”N 30°28’22”E], 28.06.1995, }
(S. Korneyev); Hydropark [50°26’36”N 30°34’47”E], on poplar and willow logs, 2.06.2007, {, } (V. Korne-
yev) (SIZK); Kyiv Region: Irpin [50°31’40”N 30°15’54”E], 9.06.1996, 2 } (V. Korneyev); Bilychi-Irpin
[50°29’06”N 30°19’30”E], 31.05.2003, 4 {, 4 }, 25.07.2003, {, } (V. Korneyev); Crimea: Karadagh, 22.06,
2.07.1929, 5 {, 4 } (S. P.[aramonov]) (SIZK); Non-European. Armenia: “Eriwan, a. s.”, 8.06, 1.07.1924, 2 }
[Paramonov] (SIZK); Israel: Panyas, 14.06.1996, 5 {, 5 } (Freidberg) (TAU); Russia: Primorskiy Kray,
Kamenushka, 16.07.1984, {, } (Shatalkin) (ZMUM).
Notes. The list of material on this species was omitted in the revision of the tribe Myennidini
(Kameneva, Korneyev, 2006) and I place it here.
From the original diagnoses (Coquebert, 1798; 1804; Fabricius, 1805) it is obvious that Coquebert’s
series of M. octopunctata was examined by Fabricius, and Fabricius based his description of S. fasciata on
Coquebert’s specimens from “Gallia”. Hence, these specimens were syntypes of both nominal species, but
only 2 wings remain in the Fabricius collection (ZMUC) belonging to one of those syntypes.
First record from Greece. The single record from Franz-Josef Land (Soo´s, 1957) is apparently based
on unintentionally introduced specimens.
Pseudotephritis corticalis (Loew, 1873)
Kameneva, Korneyev (2006): synonymy and distribution.
Notes. Along with the record from Denmark (Borgersen, Greve, 1989) (omitted in Kameneva, Greve,
2004), it was recorded from Norway (Greve, 1997).
Subfamily Ulidiinae
Tribe Lipsanini
Euxesta pechumani Curran, 1938
Euxesta nitidiventris (misidentification): Hennig (1940): Italy. — Euxesta pechumani Curran: Zuska (1967):
Slovakia; Soo´s (1980): Hungary; Kameneva (1992): Ukraine, Middle Asia; Rivosecchi (1995): Spain; Merz
(1996): Switzerland. — Euxesta stackelbergi Krivosheina & Krivosheina, 1995. — Euxesta freyi Krivosheina &
Krivosheina, 1997. Kameneva (2000): synonymy.
Material examined. Type. Holotype {: USA: “Bronx / New York City / 24 Aug. 1935 /
L. L. Pechuman”, and paratype (allotype) }, with same labels except “26 Aug.” and “Euxesta / pechumani }
/ Curran / Holotype” (red handwritten latest labels with the word “Holotype” on both [sic!] specimens)
(AMNH). Non-type. Bulgaria: Pastra, near Rila, 850 m, 26.07–22.07.1998, {, } (Achterberg, de Vries,
Atanassova) (MHNG); Greece: Ioanina, 39°41,296’N, 20°50,319’E, h = 450 m, on a dead tree, 12.06.2002,
7 } (Kameneva, V. Korneyev, S. Korneyev) (SIZK); Ukraine: Zaporizha Region: Berdyansk, 08.1954, } [no
collector] (SIZK); Non-European: Russia: North Caucasus: Northern Ossetia, 15 km N of Ardon,
18.07.1988, } (Ozerov) (ZMUM); Turkmenistan: W Kopetdagh Range, Ay-Dere Ravin, 30.04.1981, {
(Ozerov) (ZMUM); Israel: Ramat Gan, 20.01.1978, 15 {, 17 } (Kaplan); Tel Aviv, Savion, ex bark
16.09–7.10.1982», 11 {, 10 } (Zvik) (TAU).
Notes. This species was unintentionally introduced with plant material from the USA to Europe during
World War I before it was described. Later, it was recognized as a new species based on material reared from
damaged elm trees in New York City (Curran, 1938). The first correct identification of European material
was made by J. Zuska (1967). Later, N. P. Krivosheina and M. G. Krivosheina (1995) described 2 additional
species also based on Palaearctic material; however, they were compared only with E. nitidiventris Loew, not
with E. pechumani, which was shown to be a senior synonym (Kameneva, 2000). The species seems to be
rather widespread in south-western areas of the Palaearctic Region, where elm trees are planted. In Greece,
however, it was collected on a dead willow log. First records from Bulgaria and Greece.
Euxesta stigmatias Loew, 1873
Kameneva (2004): USA, Mexico, Central America, West Indies, Trinidad, South America.
435
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

Material examined. Bulgaria: Pastra, near Rila, 850 m, 11–31.05.1998, } (Achterberg, de Vries,
Atanassova) (MHNG).
Notes. Single specimen from Bulgaria was apparently unintentionally introduced with plant matter
(palms, fruits, etc.).
Tribe Seiopterini
Homalocephala albitarsis Zetterstedt, 1838
Homalocephala albitarsis Zetterstedt, 1838, nec Hennig (1940); Andersson (1991): Finland, Germany,
Norway, Poland, European Russia, Sweden; type data, identity, synonymy; Krivosheina, Krivosheina (1995):
European Russia (Vologda, Kostroma), Asian Russia (Buryatia, Amur Region); Rotheray, Robertson (1998):
Great Britain (England, Scotland); Greve, Nielsen (2001): Norway. — Psairoptera bipunctata Loew, 1854;
Homalocephala bipunctata: Hennig (1940): Poland; Steyskal (1965): USA (Oregon), Canada; Belcari et al.
(1995): Northern Italy. — Ortalis? diopsides Walker, 1849, syn. n.; Homalocephala diopsides (as synonym of
H. bipunctata): Steyskal (1965); Cole (1969). — Ortalis? costalis Walker, 1849, syn. n.; Seioptera costalis:
Steyskal (1965).
Material examined. Type. Holotype } Ortalis diopsides: [Canada: Ontario]: “Huds.[ons] Bay”
(handwritten on white circle), “St. Martin’s Falls” [“Albany River”]; “Type” (printed on green-bordered
circle), “One of Walkers / series so named [printed] / EAW [handwritten]”, “Holotype” (printed on red-
bordered circle), “Holotype Homalocephala diopsides Walker verified by J.E.Chainey, 2002” (BMNH).
Holotype { [sic!] Ortalis costalis: [Canada: Ontario]: “St. Martin’s Falls” [“Albany River”], “One of Walkers
/ series so named [printed] / EAW [handwritten]”, “Homalocephala det. J.F.Malloch”, “Holotype” (printed
on red-bordered circle), “Holotype Ortalis costalis Walker verified by J.E.Chainey, 2002” (BMNH). Non-
type. Germany: “Berlin”, “Homalocephala bipunctata”, { (?), } (DEI); Psairoptera bipunctata [sex?]: Poland:
“Tatra / 1.8.69”, “Coll. / H. Loew”, “Typus” [red label], “bipun / ctata / Lw.”, “Psairoptera / Wahlb.”
(ZMHB). European Russia: Moscow Region: Golitsyno, 20.06.1981, } (Shatalkin) (ZMUM); Sweden:
“Gellivara [= Gallivare] 435557”, { (ZMHB). Non-European: Russia: Tyumen Region: Labytnagi Near
Salekhard, 6.07.1973, } (“Homalocephala bipunctata Lw. Stackelberg det.”), 4.09.1978, {, } (Sychevskaya)
(“Homalocephala bipunctata Lw.Sychevskaja det.”) (ZISP); Far East: Amur Region: Zeya, 3006, 2,
15.07.1981, 3 { (Shatalkin); Primorskiy Kray: Kamenushka 32 km SE of Ussuriysk, 24.07.1986, {
(Antropov); Kedrovaya Pad’ Nature Reserve, 22.08, 27.09.1980, 2 {, } (Shatalkin) (ZMUM).
Notes. Both holotypes of O. diopsides and O. costalis are found to belong to the same series and to be
certainly conspecific, though the latter is in poor condition: the head is missing, and the mesonotum is almost
entirely destroyed by the overly large pin on which it was mounted. Examination of the holotype of
O. diopsides confirms its conspecificity with the lectotype of H. albitarsis as redescribed by Andersson (1991),
and I therefore synonymize these names.
The specimen labelled as a “type” of Psairoptera bipunctata in ZMHB was collected 15 years after the
species was described, and obviously is the specimen H. Loew substituted for the missing type from “Posen”
[= Poznan], but it certainly is not a syntype. Type material of P. bipunctata is therefore considered lost.
A puparium was found under bark of aspen (Greve, Nielsen, 2001).
Homalocephala apicalis (Wahlberg, 1838)
Hennig (1940): Sweden, Finland; Andersson (1991): Sweden; type data, lectotype designation, diagnosis;
Krivosheina, Krivosheina (1995): Asian Russia; Belcari et al. (1995): Northern Italy. — Psairoptera biseta
Frey, 1908, syn. n.; Homalocephala biseta: Hennig (1940), Zaitzev (1984), Winqvist (2004): Finland. —
Psairoptera similis Cresson, 1924, syn. n.; Homalocephala similis: Steyskal (1965); Cole (1969).
Material examined. Type. Holotype { P. biseta: [Finland:] “Messuby”, “R. Frey”, “4603” (blue
label), “Mus. Zool. H:fors / Spec. typ. No 4366/ Psairoptera biseta Frey”, “Mus. Zool. Helsinki / Loan Nr
D00–360” (yellow label) (ZMUH). Holotype { P. similis: [Canada:] “Star City / Sask.[atchewan] Can[ada]”
(handwritten), “H. Skinner / 20.VI.1907”, “Type No. / 6285” [red label], “9200”, “Holo- type / Psairoptera
/ similis / E. T. Cresson Jr.” (ANSP). Paratype } P. similis: [Canada:] “Star City / Sask.[atchewan]
Can[ada]” (handwritten), “H. Skinner / 20.VI.1907”, “Type No. / 6285a” [blue label], “Allo-/ type /
Psairoptera / similis / E. T. Cresson Jr.” (ANSP). Non-type. Sweden: Karlstad, 2.05.1915, {, 4.05.1915, }
(SMNS); Canada: Quebec, Mistassini, 23, 29.07.1944, {, } (Robert) (DEI).
Notes. The holotype of P. biseta is an aberrant specimen of H. apicalis with the basal scutellar setae
lacking and wing pattern somewhat pale, but otherwise keeping all the key characters of that species, so
I consider these names synonyms. The North American specimens previously determined as Homalocephala
similis (Cresson), including the types of the latter species, are conspecific with Palaearctic H. apicalis, and
I synonymize both names here. Most North American records of “H. apicalis” actually belong to H. mamaevi
(see below). Larvae under bark of aspen (Krivosheina, Krivosheina, 1995).
436
E. P. Kameneva

Homalocephala biumbrata (Wahlberg, 1838)
Psairoptera biumbrata Wahlberg, 1838; Homalocephala biumbrata: Andersson (1991): Sweden; type data,
lectotype designation; diagnosis; Krivosheina, Krivosheina (1995): European Russia (Kostroma, Moscow,
Vologda Regions, Krasnodar Kray); Rotheray, Robertson (1998): Great Britain (Scotland); Kameneva
(2002 a): Ukraine; Merz, Roha´c

ek (2005): Czech Republic, France. — Psairoptera albitarsis: Schiner (1864):
Austria; Homalocephala albitarsis: Hennig (1940): Sweden, Finland, Latvia; Belcari et al. (1995): Northern
Italy (misidentification).
Material examined. Non-type. Austria: “Alte Sammlung”, “albitarsis / det. Schiner”, }
(NHMW); Germany: “Berlin / Piche[l]sberg”, 30.08.1904, 1 specimen (DEI); Estonia: Tartu (“Dorpat”),
07.1887, } (Mik); idem, 31.07.1887, } (Sintenis) (ZMHB); idem, 1.08.1887, 2 {, 10 } (Sintenis) (NHMW);
Latvia: Liepaja (“Libau”): 3 {, 4 } (Siebert) (DEI); Sweden: “ Suecia. Boh.[eman]”, “6368”, } (ZMHB);
Ukraine: Kyiv, Lukjanivka, on poplar log, 9.05.2002, } (S. Korneyev, V. Korneyev), idem, 11.05.2002, }
(V. Korneyev) (SIZK).
Notes. Larvae under bark of dead aspens, poplars and willows (Krivosheina, Krivosheina, 1995;
Rotheray, Robertson, 1998). First records from Estonia and Latvia.
Homalocephala mamaevi Krivosheina et Krivosheina, 1995
Krivosheina, Krivosheina (1995): Asian Russia (Tyva). — Homalocephala apicalis: Steyskal (1965); Cole
(1969) (misidentification).
Material examined. Austria: Lechtaler Alpen, Tirol, “Boden–Hahnrenn, 1. VII”, 1 specimen
[abdomen lost], “Homalocephala apicalis Wahlb. Lindner det.” (SMNS); Sweden: Karlstad, “et faulem
ulmen” [on fallen elms?] 4.05.1915, }, 19.05.1915, {, } (“a. Mulin”) (SMNS).
Notes. The first European records from Austria and Sweden. Examination of Nearctic material shows
the records of “H. apicalis” from the Nearctic Region to be at least partly based on misidentification of this
species (Kameneva, pers. observation). Larvae were found under bark of larch (Krivosheina, Krivosheina,
1995).
Pseudoseioptera demonstrans (Hennig, 1941)
Seioptera demonstrans Hennig, 1941; Pseudoseioptera demonstrans: Krivosheina, Kovalev (1972); Kameneva
(2002 b): Ukraine. — Pseudoseioptera ingrica Stackelberg, 1955; Krivosheina, Kovalev (1972): European
Russia, Siberia, description of larva; Kameneva, Korneyev (1995): synonymy.
Material examined. Type. See Kameneva, Korneyev (1995). Non-type. European Russia: Moscow
Region: Malinki near Krasnaya Pakhra, 7.06.1973, } (Kovalev) (ZMUM); Ukraine: Kyiv, Lukyanivka,
14.05.2002, }; Kyrylivskyi Hay, 11.06.2003, }, 16.06.2006, 3 } (Kameneva, V. Korneyev) (SIZK).
Notes. Larvae under bark of fallen deciduous trees (Krivosheina, Kovalev, 1973). In Europe it is
associated with poplar logs (Kameneva, unpublished data).
Seioptera vibrans (Linnaeus, 1758)
Myodina vibrans: Schiner (1864): Austria; Jaroszewski (1876): Ukraine (“Kharkov and vicinity”); Seioptera
vibrans: Se´guy (1934): France, Spain; Hennig (1939): British Is., Croatia, Germany, Norway, Poland,
Romania, Sweden; (Soo´s, 1957): Albania, Hungary, Russia (Franz-Joseph Land, Belgorod Region);
Slovakia; Turkey (Asia Minor); Lobanov (1958): Russia, description of larva; Rivosecchi (1995): Italy; Merz
(1996): Switzerland; Greve (1997): Norway; Roha´c

ek (2006): Czech Republic; van Aartsen, Beuk (2002):
Netherlands; Pakalnis

kis, Pod
.
enas (1992): Lithuania; Winqvist (2004): Finland; Karpa et al. (2005): Latvia.
Material examined. Greece: Apollonia, 40°39,036’N, 23°29,667’E, relict forest at Volvi Lake,
h = 60 m, 30.05.2002, } (Kameneva, V. Korneyev, S. Korneyev) (SIZK); Russia: Moscow Region:
Zvenigorod, 9.07.1973, 1 specimen (abdomen lost) (Sakharova) (ZMUM); Ukraine: Zhitomir Region:
Novograd-Volynsky, 35.07.2003, {, 2 } (V. Korneyev); Korostyshiv, 30.05.1903, }; Kyiv Region: Kyiv:
25.06.1920, } [Paramonov], 16.06.1923, { (no collector); 28.06.1995, } (S. Korneyev); Kyrylivskyi Hay,
27.06.2007, on flowers of Daucus carota, } (V. Korneyev); Golosiyeve, 6.10.1978, }; Kontsha-Zaspa,
meadow, 4.06.1985, }; Irpin: trapped in glasshouse, 18.05.1996, 13 {, 5 }, 9.06.1996, 2 {, 5} (Kameneva
and V. Korneyev); Kyiv Region: Malyutinka, 12.06.1927, } (Panocini); Obukhiv, vicinity, 11.06.1983, }
(V. Korneyev); Vinnitsa Region: Vinnitsa, 14.07.1930, {, } (Paramonov); Cherkasy Region: Trakhtemyriv,
30.06.1983, } (Verves); Kaniv, Nature Reserve, 22.05.1965, 1.06.1967, {, 2 } (O. Viktorov-Nabokov);
30.05.1981, { (Verves); 7.07.1984, } (V. Korneyev); Cherkasy, 1.06.1989, { (Zrazhevsky); Kirovograd
Region: Oleksandrivka, } [no date; Paramonov]; Dnipropetrovsk Region: Lozovatka, 20.06.1918, 3 }; Odessa
Region: Valegotsulovo, 22.05.1925, }; Ananyev, 4, 6, 11.06.1931, 5 {, 6 } (Paramonov); Mykolayiv Region:
Peski, 06.1991, {, } (V. Korneyev); Zaporizha Region: Berdyansk, 4.06.1940, 3 } (Gulinov) (SIZK);
437
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

Lugansk Region: Lugansk, 7.07.1929, } (Talitsky) (ZMUM); Crimea: Miskhor, 6.08.1926, { (Muzytshenko)
(SIZK). Non-European: Armenia: Erevan, 17.05.1924, {, 18, 20.06.1924, 2 } (no collector) (SIZK).
Notes. A common saprophylic species, often found in association with rotting plant matter
(vegetables, hay, compost), sometimes attracted to dung. First record from Greece. Single record from
Franz-Josef land (Soo´s, 1957) is apparently based on unintentionally introduced specimens.
Tribe Ulidiini
Physiphora alceae (Preyssler, 1791)
Musca alceae Preyssler, 1791: Czech Republic; Physiphora alceae: Karpa et al. (2005): Latvia. — Musca
demandata Fabricius, 1795; Chloria demandata: Schiner (1864): Austria, France, Germany, Sweden;
Jaroszewski (1876): Ukraine (Kharkiv and vicinity); Physiphora demandata: Hennig (1940): England,
European Russia, Spain (Mallorca, Canary Is.); Siberia; Iran; Egypt, Libya, Morocco, Tunisia; Soo´s (1957):
Hungary; Gosseries (1991): Belgium; Merz (1996): Switzerland; van Aartsen, Beuk (2002): the Netherlands.
Material examined. Bulgaria: Sliven: “5 km N, along brook”, 42°45N 26°17E, 400 m, 21.07.1987,
{, }; Kiten: “oak wood + pig farm”, 42°14N 27°48E, 21.07.1987, {, } (Barta´k) (NMP); Croatia:
“Dalmatia”, } [collector unknown] (NHMW); Rovinj, 28.05.1961, } (ZFIB); France: Mont Dauphin, “pine
wood”, 44°41N 6°37E, 800 m, 11.07.1990, { (Barta´k) (NMP); Digne, 06.1908, {; Corsica, Ajaccio
24.06.1899, { [collector unknown] (MTD); Greece: Corfu, {, } [collector unknown] (DEI); Litohoro,
40°06,067’N, 22°29,020’E, h = 340 m, on a window, 11.06.2002, } (V. Korneyev) (SIZK); Italy: Ferrara:
“along river”, 44°54N 11°38 E, 6.08.1988, } (Barta´k); Castiglione: “D’Orca, along river”, 43°00N 11°37E,
800 m, 7.08.1988, { (Barta´k) (NMP); Moldova: Chisina˘u: Institute of Biocontrol for Plant Protection,
13.07.1987, 2 { (V. Korneyev); idem, in the room, 12.09.1987, } (Kameneva); Balabanesti near Vadu-lui-
Voda˘, 22–26.07.1988, { (Korneyev) (SIZK); Romania: Bucuresti, 2 { (Montandon) (ZSSM); Spain:
Tordera, “along river”, 41°42N 2°45E, 5.07.1990, { (Barta´k) (NMP); Russia: Rostov Region: Taganrog, 3,
31.08, 3.09.1923, 3 } (no collector) (SIZK); Ukraine: Kyiv: “in the room”, “Korolenko 55”, 17.09.1931, 2 }
(Belanovsky); Kyiv Region: Malyutinka, 5.09.1927, } (Panotshini); Irpin, 28.06.1995, } (S. Korneyev);
Cherkasy Region: Kaniv Nature Reserve, 30.03.1957, { (Viktorov-Nabokov); Moshny, 30.08.1988, }
(Zrazhevsky); Kherson Region: Ananyev, 11.06.1921, } (Paramonov); Kalanchak, Novooleksandrivka,
23.07.1987, 2 {, 3 } (V. Korneyev); Zaporizhya Region: Berdyansk, vicinity, 06.1964, } (Gulinov) (SIZK).
Notes. First records from Bulgaria, Moldova and Croatia.
Timia abstersa Loew, 1873
Hennig (1940): Russia (Orenburg, Volgograd, Astrakhan Regions), Lebanon, Middle Asia; Kameneva (2002
c): Ukraine, Moldova, Transcaucasia, Kazakhstan, Iran, Mongolia.
Material examined. Type. Holotype {: [Tadjikistan: Zeravshan Range or valley:] “Sarawsch. /
Fedtsch”, “Coll./ H. Loew”, “Type” [red label] (ZMHB). Non-type. Moldova: Ghidighici [Gidigich] NW
of Chisina˘u [47°06’N 28°52’E], 5.07.1988, {, }; Chisina˘u: Institut of Biocontrol for Plant Protection
[46°58’N 28°53’E], 14.06.1986, }; Comrat, vicinity: Tigheci River near Bujac [46°21’N 28°40’E], 17.06.1987,
6 {; } (Korneyev) (SIZK); Ukraine: Kyiv Region: Kruglyk prope Kyiv [50°16’47”N 30°26’56”E], 17.07.2006,
3 {, } (Kameneva, V. Korneyev, S. Korneyev); Odessa Region: Starokozache, forestry, “near Maloye”
19.06.1984, {, } (Dolin); Danube Delta, Djarylgach Island, saline steppe, 2.07.1997, } (A. Kotenko);
Mykolaiv Region: Pervomaysk District, Mighiya [48°01’N 30°57’E], 20.06.1985, } (Korneyev); Yelanets
District, Kalinovka, 22.06.1984, {, } (Karachevskaya); Kherson Region, Novooleksandrivka [46°14’N
33°22’E], 08.1985, { (Kameneva, Korneyev); Zaporizha Region: Melitopol, 1.06.1983, {, }; Akimovka
District, “Vatutin Collective Farm”, 13.06.1983, } (Djafarov); Berdyansk, steppe, 30.07.1977, {, } (Gulinov)
(SIZK).
Notes. Widespread in subarid zones of the Palaearctic Region, from Southern Moldova and Ukraine,
eastwards to Mongolia, and in steppes and semi-deserts of European Russia (Volgograd and Astrakhan
Regions), Kazakhstan, states of Middle Asia and Iran. The holotype is in a poor condition (head, abdomen,
4 legs and 1 wing and a half missing].
Timia melanorrhina Loew, 1866
Hennig (1940): Greece (“Attica”), Russia (Sarepta), Kazakhstan, Turkmenistan; Belcari et al. (1995): Italy
(south).
Material examined. Type. Syntypes: 2 {: [European Russia: Volgograd:] “Sarepta”, “Coll. /
H. Loew”, “Type” [red label], “ST” [red label] (ZMHB); possible syntypes: 3 {: “Sarepta”, “Coll. /
H. Loew”, “Type” [red label], {: “Sarepta / Christoph”, “Coll. / H. Loew”, “Type” [red label] (ZMHB).
Non-type. European Russia: “Sarepta”, “Coll. / H. Loew”, 3 }[with red label “Type”], [with red label
“Type”] “Sarepta / Christoph”, “Coll. / H. Loew”, 2 } [with red label “Type”], “Sarepta / 30222”, 3 {,
2 }, “Sarepta / 33583”, 4 {, }, “Sarepta / 36625”, 10 {, 6 } (ZMHB).
438
E. P. Kameneva

Notes. This species was originally described based on an unknown number of males, therefore females
marked as types are not syntypes. W. Hennig (1940) and V. F. Zaitzev in A. Belcari et al. (1995) reported
this species from Greece and Italy; however, no West European material was seen in this study.
Timia xanthaspis Loew, 1868
Hennig (1940): Greece (“Attica”), Russia (Sarepta), Turkmenistan; Belcari et al. (1995): Italy (south).
Material examined. Type. Syntypes: {: [European Russia: Volgograd:] “Sarepta / Christoph”,
“Coll. / H. Loew”, “Type” [red label], “xan- / thaspis / Lw.”, {, 3 }: “Sarepta / Christoph”, “Coll. /
H. Loew”, “Type” [red label] (ZMHB). Non-type. European Russia: Volgograd: “Sarepta / 33582”, {, },
“Sarepta / 36626”, 1 specimen (abdomen missing) (ZMHB).
Notes. W. Hennig (1940) and Zaitzev in A. Belcari et al. (1995) reported this species from Greece
and Italy; I have not seen any West European material in this study so far.
Ulidia Meigen, 1826
No comprehensive taxonomic treatment of European species has been provided since W. Hennig
(1940). The genus is represented mostly by uniformly shining-black flies, which have hyaline wings,
sometimes with a few dark spots. In the eastern Mediterranean region, Asia Minor and Near East, there are
several species often treated as “variations” or “subspecies” of U. erythrophthalma (Meigen) (Hennig, 1940;
Zaitzev, 1984) poorly differentiated from external characters. In this study, the structure of the phallus,
especially its apical portion, was found to be species-specific. This made possible recognition of several species
similar to U. erythrophthalma, of them two species occurring in Europe.
In the tribe Ulidiini, the phallic structure differs from those found in other ulidi-
ids. It is a long, coiled and partially flattened tube divided into two halves; a pair of
sclerotized taeniae end approximately at its mid-length, and another pair of similar tae-
niae begins at the middle, almost reaching the apex; apical half bears one to three long
membranous caecum-like appendices (fig. 3, 6: cla), which are clearly visible in water,
but always strongly shrivelled in glycerol. The apex of the distiphallus is bowed almost
at a right angle and bears numerous sclerotized cuticular structures (spurs, hooks, lobes,
etc.), which surround the gonopore like rose-petals and form a “glans”, which is si-
milar to that structure in the Tephritidae. In Ulidiini, the “glans” is strongly asymmet-
rical and cannot therefore be placed for study in a standardized position. Some of these
structures (and especially in combination) are species-specific; more-so than coloration
of head and wings alone. This treatment of the genus is limited to European species
only, but it should also be noted that material from Near East and Middle Asia con-
tain numerous undescribed species. Like the other Ulidiini, the genus Ulidia needs fur-
ther taxonomic revision with the use of the glans structure.
Key to species of European and Near East (in brackets) species of Ulidia
Òàáëèöà äëÿ îïðåäåëåíèÿ åâðîïåéñêèõ è áëèæíåâîñòî÷íûõ (â êâàäðàòíûõ ñêîáêàõ) âèäîâ ðîäà Ulidia
1. Wing hyaline with brown apical spot. ................................................................................................ 2.
— Wing entirely hyaline or mostly brownish, but without apical spot. .................................................. 4.
2. Eye at least 1.4 times (usually 1.5–1.55) as high as long. Frons wider than long or at most as long as
wide, deeply pitted. Larger flies, wing length 4.5–6.5 mm. .............................................................. 3.
— Eye round, less than 1.3 (1.2–1.25) times as high as long. Frons longer than wide, inconspicuously
dotted. Smaller flies, wing length less than 4.5 (usually 2.5–3.5) mm. ............ U. salonikiensis Hennig
3. Halter yellow. .......................................................................................................... U. apicalis Meigen
— Halter black. ........................................................................................................ U. megacephala Loew
4. Side of frons with triangular white microtrichose area (fig. 1, 1: a; 3, 1). At least wing base (cell bc)
brown. .................................................................................................................................................. 5.
— Side of frons shining, at most eye bordered with narrow whitish microtrichose areas. Wing and calypters
variable. .............................................................................................................................................. 6.
5. Mesonotum densely shagreened, matt (fig. 2, 1). Wing (including cell c) widely diffuse brown,
especially in anterior half, but without delimited spots (fig. 1, 2; 2, 2, 4). Frons in profile straight or
slightly convex, vertical plate and gena moderately or short setose (longest setae shorter than antenna);
parafacial and face black (fig. 1, 1; 2. 3). Male cerci widely separated, triangular (fig. 2, 5); glans with
cylindrical body and fan-like lobes (fig. 2, 6). .............................................................. U. atrata Loew
— Mesonotum shining black or finely shagreened. Wing hyaline, except cell bc, base of cell c and stigma
usually brown (fig. 3, 1). Frons in profile usually concave, vertical plate and gena long setose (longest
439
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

setae longer than antenna); parafacial and face reddish yellow. Male genitalia not as above. ..............
.................................................................................................................................... U. ruficeps Becker
6. Acrostichal seta (ac) lacking. Mid and hind tarsi usually black. Eye at most 1.3 (1.1–1.2) times as high
as long. ................................................................................................................................................ 7.
— Acrostichal seta (ac) present. Mid- and hindtarsi at least partly yellowish to reddish. Other characters
variable. .............................................................................................................................................. 8.
7. Face, gena and most of frons dark brown to black. Wing in both sexes with brownish yellow tinge,
except base; pterostigma brown (fig. 5, 1, 5). .......................................................... U. parallela Loew
— Face, gena and anterior half of frons yellow or light brown. Wing: in male, with pale brown tinge, pte-
rostigma brown; in female, hyaline or at most with pale brown pterostigma. ........................................
................................................................................................................................[U. omani Steyskal]
8. Wing mostly brownish, except cells bc, c, basal half of br, and also bm and bcu yellow (fig. 4, 1). Apical
portion of distiphallus with blunt sclerotized projections (at least one Y-like) (fig. 4, 5–6), but without
serrate lamina, acute projections, claw-like structures, or curved comb-like projections. ......................
................................................................................................................................ U. nigripennis Loew
— Wing uniformly hyaline or with yellow tinge (fig. 3, 2; 6, 1; 7, 1). Glans not as above. .................. 9.
9. Frons satin subshining and smooth in the middle, dotted laterally; pterostigma and veins brownish;
mesonotum densely shagreened, almost matt. Male hindfemur unmodified, as thick as midfemur. ....
................................................................................................................................[U. facialis Hendel]
— Frons shining and deeply pitted over whole anterior half. Wing hyaline or with yellowish tinge, but
neither with brown stigma, nor with black veins. ............................................................................ 10.
10. Eye round (less than 1.3 times as high as long). Anterior part of head (frons above lunule, often face
and parafacial) reddish. Male hindfemur swollen in distal part (fig. 3, 5) ........ [U. wadicola Steyskal]
— Eye oval (more than 1.3 times as high as long). Anterior part of head reddish or usually black. Male
hindfemur not swollen, as wide as midfemur. .................................................................................. 11.
11. Abdominal tergites and mesonotum smooth or finely shagreened, abdomen more shining than
mesonotum. ...................................................................................................................................... 12.
— Abdominal tergites 1–4 mostly matt, except narrow posterior margin shining, mesonotum subshining,
moderately shagreened. .......................................................................................... U. semiopaca Loew
12. Apical portion of distiphallus with blunt sclerotized projections (one Y-like (fig. 6, 4–5: ys), one
truncate cylindric and one sclerotized bar (fig. 6, 5: sb) on membranous area), at most one rather weak
claw-like structure (fig. 6, 4–5: cl); lamina with serrate margin and denticles at base (fig. 6, 4: sl);
rasper-like membranous vesica with 35–55 denticles well-developed (fig. 6, 5: ve); no curved comb-
like projections. .......................................................................................... U. erythrophthalma Meigen
— Apical portion of distiphallus with at least 3 sharply pointed projections (including one claw-like
structure (fig. 8, 1–3, 5: cl), one long projection (fig. 8, 1–5:
lp) of serrate lamina), curved comb-like
projection) and lesser expressed rasper-like structure (weaker denticles on vesica). ..............................
.............................................................................................................................. U. albidipennis Loew
Ulidia apicalis (Meigen, 1826)
Hennig (1940): France (south, mainland, Corsica), Italy (Sicily), Portugal, Spain; Morocco, Tunisia.
Material examined. Type. Syntypes 2 }: “Portugal / Hoffmannseg S.” [one with red label
“Type”], }: “Lusitan. Hoffsg.”, “2725”, “apicalis / (Hffg.*) Meig.”, “Type” [red label] (ZMHB). Non-type.
Austria (?): “osterreich”, “Alte Sammlung”, } (NHMW); Greece: Rhodes: “Rodi, Egeo”, 05.1939, 2 {, }
(Meyer) (ZMHB); Spain: SE, dept. Ma´laga, between Mijas & Benaldena, 300 m, wasteland, 16.04.1983, {,
}, idem, Calahonda, between Fuengirola & Marbella, 21–22.04.1983, } (Exc. Univ. Leiden) (RMNH).
Notes. First record from Greece. The only specimen known from Austria might be mislabelled.
Ulidia atrata Loew, 1868 (fig. 1–2)
Hennig (1940): Greece; Belcari et al. (1995): Italy (South mainland).
Material examined. Type. Greece: Syntype {: “Parnass / 27/4 66”, “Coll. H. Loew.” “Type” (red
label), syntype }: “Parnass / 30/4 1866”, “Coll. H. Loew.” “Type” (red label ). Non-type. Greece: Olympia,
25.04.1964, } (Grunwaldt) (ZSSM); Metsovo, 15.06.2002, } (Kameneva) (SIZK); Ipiros: Smolikas Mts.,
7–1500 m, 21–22.05.1994, } (Michelsen); Timfi Mts., 6–1000 m, 23.05.1994, { (Andersen) (ZMUC);
Pindos, Pertouli, 4.06.1968, 2 { (Lindner); Kefalonia, between Poros and Skala, Macchie, 28.04.1996, 2 {,
3 }; 1–3 km NNW Poros, Strand–Maccie, 1.05.1996, { (Miksch) (SMNS); Trikala: Panagia, 15.06.1982,
{ (dissected), Viotia, Parnassos, 11 km NW of Ara´chova, 1100 m, 6.06.1982, } (Danielsson) (ZIL);
Peloponnisos, Taiyetos [= Ta¿eti] Mts., 950–1800 m, 15–19.05.1990, 2 {, 2 } (ZMUC); Corfu: “Korfu. V.
/ 50187”, } (Becker) (under Becker’s bottom label “nigripes Beck.”); Greece (locality not stated on labels):
“4167”, “4453”, “4454”, “4456”, “4457”, “4567”, 3 {, 3 } (collector not stated on labels) (under Becker’s
bottom label “nigripes Beck.”) (ZMHB); Italy: “Calabr[ia] / Erber”, “Coll. H. Loew.” “Type” (red label)
1 specimen (abdomen lost) (ZMHB).
440
E. P. Kameneva

Diagnosis. Wing darkened, mostly in anterior half, including cell bc (fig. 1, 2;
2, 3, 4), differing well from other European species by matt, densely shagreened
mesonotum (fig. 2, 1, 2); the orbit of the frons having a whitish microtrichose triangu-
lar spot at the anterior margin and parafacial wide (twice as wide as flagellomere 1),
wrinkled; the gena as high as eye (fig. 1, 1).
Notes. This species was previously differentiated from the dark-winged Ulidia (e.g., U. nigripennis) by
brown calypters or halteres (Hennig, 1940). However, the specimens from Becker’s collection marked
as”nigripes” (collection name only) have white calypters and creamy halteres, and some other specimens
collected in June have white calypters, but brownish halteres or both halteres and calypters white, so this
character is strongly variable. The best character to distinguish it is the matt, strongly shagreened mesonotum.
Ulidia megacephala Loew, 1845
Hennig (1940): Greece; Spain; Turkey; Syria; Morocco.
Material examined. Type. Possible syntypes 2 {, }: [Greece/Turkey:] (yellow paper square), “Coll.
Loew”, “Type” (red label); (grayish paper square), “Coll. Loew”, “Type” (red label), } (ZMHB). Non-type.
Greece: Rhodes: River Gadoura´s, mouth, 17.04.1980, }; Mt. Ataviros, nr. Ag. Isidoros, 20.04.1980, 2 {, }
(van Ooststroom) (RMNH); Spain: “Spanien, v. Seydl.”, “Coll. H. Loew”, “Type” (red label), } (ZMHB).
Notes. The species was originally described from “die Gegenden von Macra und Mermeriza [Turkey:
Marmaris]; Rhodus”, so the specimen from Spain was marked as “Type” erroneously. The } from “Spanien”
(ZMHB) has widely reddish yellow frons, face and parafacial, and slightly concave profile of frons like in
U. ruficeps. See comment on the latter species.
Ulidia nigripennis Loew, 1845 (fig. 4)
Schiner (1864): Hungary; Jaroszewski (1884): Ukraine (“Kharkov Gouvern.”); Zaitzev (1984): Germany;
Roha´c

ek (2006): Slovakia; Ulidia erythrophthalma var. nigripennis: Hennig (1940): Italy, Poland.
Material examined. Type. Syntypes {, }: [Poland:] “Posen [= Poznan´] / 11.8.44”, “Coll. / H. Loew”,
“Type” [red label], “nigri- / pennis / Lw”; }: “Coll. H. Loew”, “U. erythrophth. v. Zblr.”, “Type” [red
label], “Ulidia / nigripen- / nis m.”, “Coll. H. Loew”; }: “3.8.44”, “Coll. H. Loew”, “Type” [red label]
(ZMHB). Non-type. Austria: “Donauen”, 12.06.1884, { (Handlirsch); Modling, 19.07.1881, {, } (NHMW);
Grossweikersdorf, 25.08.1982, } (J. & E. van der Vecht) (RMNH); France: “Dauphine / Vallouise”
11.07.1875, { (“Ulidia erythr. nigripennis Loew, Lindner det.”) (SMNS); Var, Montauroux, 1.07.1970, }
(J. van der Vecht) (RMNH); Hungary: “Kertesz K / Bpest”, “Ungarn / 40852”, } (ZMHB); Greece:
Metsovo, 39°46’32”N 21°10’47”E, 15.06.2002, 2 { (Kameneva); Viotia, Parnassos, 11 km NW of Arachova,
1100 m, 6.06.1982, { (Danielsson) (ZIL); Central Greece: Fthiotis, “Iti Oros [Mt. Iti] 2 km SE Iti”,
9.06.1982, { (dissected) (Danielsson); Ara´chova [38°28’N 22°24’E], 6.06.1982, { (dissected) (Danielsson)
(ZIL); Moldova: Calarasovca, 8.07.1987, } (Vinokurov) (SIZK); Slovakia: “Hegy-farok, steppe”, 47°50N
18°36E, 220 m, 16.07.1986, 4 {, 10 }; Kamenica, Hronom, 1 km N, steppe, 47°50N 18°13E, 200 m,
15.07.1986, {, 4 } (Barta´k); Hlohovec, 12.08.1989, { (Malek) (NMP); Ukraine: Kyiv, Lysa Hora
441
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…
Fig. 1. Ulidia atrata, syntype: 1— head, left view (a — triangular white microtrichose area); 2— wing.
Ðèñ. 1. Ulidia atrata, ñèíòèï: 1— ãîëîâà, âèä ñëåâà (a — òðåóãîëüíûé ó÷àñòîê â áåëûõ ìèêðîòðèõèÿõ);
2— êðûëî.

[50°16’47”N 30°26’56”E], 2.07.2000 (Kameneva, V. Korneyev), Kruglyk prope Kyiv [50°16’47”N
30°26’56”E], 21.07.2006, 4 {, 3 } (V. Korneyev, S. Korneyev); between Mrygi and Hodosievka [~50°15’N
30°32’E], 29.07.2006, 2 {, 4 } (V. Korneyev, Kameneva) (SIZK).
Diagnosis. Head black; no whitish microtrichose areas; frons deeply pitted
(fig. 4, 2); parafacial black, as wide as flagellomere 1; eye 1.2 times as high as wide;
442
E. P. Kameneva
Fig. 2. Ulidia atrata: 1, 2— head and mesonotum (1— dorsal view, 2— latero-dorsal view); 3, 4— wing;
5— epandrium, posterior view (c — cerci); 6 — phallus glans; 7 — aculeus (cu — cercal unit of aculeus);
8 — spermathecae.
Ðèñ. 2. Ulidia atrata: 1, 2— ãîëîâà è ñðåäíåñïèíêà (1— âèä ñâåðõó, 2— âèä ñâåðõó è ñáîêó); 3, 4—
êðûëî; 5— ýïàíäðèé, âèä ñçàäè (c — öåðêè); 6 — ãëàíñ ôàëëþñà; 7 — ëåçâèå ÿéöåêëàäà (cu —
öåðêàëüíûé ÷ëåíèê ëåçâèÿ ÿéöåêëàäà); 8 — ñïåðìàòåêè.

mesonotum shining black or slightly shagreened; wing with yellow base (bc, c, br along
RS to fork, bm and bcu) and yellow veins in basal 1/4 and greyish apical ¾ of wing
(fig. 4, 1); halter yellow; legs black, except mid- and hindtarsi yellowish; hindfemur
very slightly swollen; hindtibia almost straight; abdomen shining black, very finely sha-
greened, with a faint greenish tinge. Male genitalia: surstylus with evenly narrowed basal
part; cerci separated by very shallow fissure (fig. 4, 3, 4); phallus glans with compara-
tively large fork-like sclerite, but neither serrate lamina nor with sclerotized lobes co-
vered with numerous denticles. Female terminalia: aculeus as on fig. 4, 7, with long
setae on elongate oval cercal unit; 3 nearly conical spermathecae (fig. 4, 8).
Notes. First records from Austria, Greece and Moldova.
443
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…
Fig. 3. Ulidia ruficeps (1), U. facialis (2–4 ) and U. wadicola (5–6): 1, 2— habitus, left view; 3— frons,
dorsal view; 5— hind leg without coxa; 4, 6 — phallus glans (cla — caecum-like appendix).
Ðèñ. 3. Ulidia ruficeps (1), U. facialis (2–4) è U. wadicola (5–6): 1, 2— îáùèé âèä ñëåâà; 3— ëîá, âèä
ñâåðõó; 5— çàäíÿÿ íîãà áåç òàçèêà; 4, 6 — ãëàíñ ôàëëþñà (cla — ñëåïîé îòðîñòîê).

444
E. P. Kameneva
Fig. 4. Ulidia nigripennis: 1— habitus, left view; 2— frons, dorsal view; 3, 4— epandrium, posterior view
(3 — uncompressed, 4— bleached and compressed on slide); 5, 6 — phallus glans, different aspects (ys —
Y-shaped sclerite); 7 — aculeus; 8 — spermathecae.
Ðèñ. 4. Ulidia nigripennis: 1— îáùèé âèä, ñëåâà; 2— ëîá, âèä ñâåðõó; 3, 4— ýïàíäðèé, âèä ñçàäè (3 —
íåñïëþùåííûé, 4— îòáåëåííûé è ñïëþùåííûé íà ïðåïàðàòå); 5, 6 — ãëàíñ ôàëëþñà, âèäû ñ
ðàçíûõ ñòîðîí (ys — Y-îáðàçíûé ñêëåðèò); 7 — ëåçâèå ÿéöåêëàäà; 8 — ñïåðìàòåêè.

Ulidia parallela Loew, 1845 (fig. 5)
Material examined. Type. Poland (?): Syntype: } “Schlesien / 19.6.42”, “Coll. H. Loew.” “Type” and
possible syntypes: {: “cf. nitida / Zeller”, { “Zllr. /[?]st” “Coll. H. Loew.” “Type” (red labels) (ZMHB).
Non-type. Bulgaria: { (dissected): Irakli, damp meadow, 42°47N 27°54E, 16.07.1987 (Barta´k) (NMP).
Germany (?): “Berlin / Ruthe”, “nitida / Meig.*”, “Coll. H. Loew” {, 2 } (ZMHB). Moldova: }, Budjak,
Ialpujeli River, 13.07.1988 (Korneyev) (SIZK). Slovakia: }, Hegy-Farok, damp valley, 47°50’N 18°36E,
220 m 16.07.1986 (Barta´k); { (dissected), 4 } (1 dissected): Kamenica, Hronom, 1 km N, steppe, 47°50’N
18°43E, 200 m, 15.07.1986 (Barta´k) (NMP; SIZK).
Diagnosis. Mid-sized Ulidia with subshining mesonotum, smoothed frontal vitta
and darkened wings, differing from other European Ulidia species by combination of
acrostichal seta absent, pterostigma dark brown, veins R
4+5
and M only slightly conver-
gent, mid- and hindtarsi yellow to black, and by male genitalia (surstyli with prominent
medio-basal part; phallus glans with elongate triangular subbasal lobe, weakly sclero-
tized papillose lobe, striate and wrinkled sclerotized belt, middle-sized lobe with papil-
lae at base, but neither spurs, nor serrate lobes) and female terminalia structures (cer-
cal unit round; spermathecae short with microspinulose surface).
Redescription. Head mostly dark brown to black. Frons silky shining, with
shallow pits or dots, as long as wide, mostly black, except anterior margin often red-
dish brown; orbits shining, without microtrichose areas. Face and antenna black.
Parafacial black, smooth, slightly wider than flagellomere 1. Gena high, at 0.45–0.55
as high as eye, smooth, subshining, finely shagreened. Eye almost round, 1.2–1/3 as
high as long. Setae: 1 or, 1 oc, 1 poc, 1 vti, 1 vte. Antenna, including arista, matt black.
Palpus black, moderately wide, parallel-sided, with black setae.
Thorax shining black, with black setae and setulae. Mesonotum subshining black,
with finely shagreened, subshining medial vitta and shining sides. Setae: 1 ppn, 2 sa
(anterior half as long as posterior), 1 pa, 0 ac, 1 dc, 1 ia, 2 pairs sctl, 1 anepst, 1 kepst.
Legs entirely blackish brown, including mid- and hindtarsi; black setulose; poste-
rior femur unmodified in both sexes, as thick as midfemur.
Wing (fig. 5, 1, 5) yellow with brownish tinge, except cells c, br, bm, bcu and anal
lobe and alula whitish (all veins in this area also yellow); pterostigma and often cell r
1
posterior of it brown. Wing length 3–3.2 mm. Halter yellow.
Abdomen shining black, tergites very finely shagreened.
Male genitalia: surstylus with prominent posteromedial lobe (pml) and mesoven-
trally curved antero-ventral lobe; cerci deeply separated (fig. 5, 6). Phallus: spinulose
caecum at middle of its length (fig. 5, 7), one membranous caecum basally of glans.
Glans short, with lobes, projections and other structures as shown on figs. 5, 8–9: sub-
basal leaflet (lobe) moderately sclerotized, acute angular; striate and wrinkled sclero-
tized belt present (swb); one wide sclerotized lobe with two rather short projections and
numerous papillose structures; and one weakly sclerotized vesica with denticle- or
papilla-like structures on it; some additional smaller rounded lobes can be recognized.
Female terminalia: aculeus moderately wide, 7–7.5 times as long as wide, with round
cercal unit (fig. 5, 10); 3 short spermathecae with finely and sparsely spinulose surface.
Egg (fig. 5, 11) elongate, 3–3.2 times as long as wide.
Notes. First records from Bulgaria, Moldova and Slovakia.
Ulidia ruficeps Becker, 1913 (fig. 3, 1)
Hennig (1940): Iran, Lebanon; Belcari et al. (1995): Italy (south).
Material examined. European Russia: Volgograd Region: Had near Sarepta, 1871, “Coll. /
H. Loew”, 1 specimen (abdomen lost) with labels “rufi- / frons / Lw.” and “Type” [red paper] (Christoph)
(ZMHB); Non-European. Lebanon: “Nd.-Liban., Cedern. b. Becharre´, 1900 m” 3–6.06.1931, } (“Ulidia
ruficeps Beck. det. Dr. W. Hennig 1939”) (Zerny) (NHMW); Turkmenistan: Tuzte River, 16.06.1908 (no
collector) (SIZK).
445
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…

446
E. P. Kameneva
Fig. 5. Ulidia parallela { (1–9) and } (10–12): 1— habitus, dorsal view; 2— head, left lateral view; 3 —
frons, dorsal view; 4— mesonotum, dorsal; 5 — wing; 6— epandrium, posterior view (c — cerci, pml —
postero-medial lobe of surstylus); 7— spinulose caecum of phallus; 8, 9 — phallus glans, different aspects
(ps — papillose structures, sbl — subbasal lobe of glans, swb — striate and wrinkled sclerotized belt); 10 —
aculeus (cu — cercal unit of aculeus); 11 — spermathecae; 12 — egg.
Ðèñ. 5. Ulidia parallela { (1–9) è } (10–12): 1— îáùèé âèä ñâåðõó; 2— ãîëîâà, âèä ñëåâà; 3 — ëîá,
âèä ñâåðõó; 4— ñðåäíåñïèíêà, âèä ñâåðõó; 5 — êðûëî; 6— ýïàíäðèé, âèä ñçàäè (c — öåðêè, pml —
çàäíåìåäèàëüíàÿ ëîïàñòü ñóðñòèëÿ); 7— øèïîâàòûé ñëåïîé âûðîñò ôàëëþñà; 8, 9 — ãëàíñ ôàëëþñà,
âèäû ñ ðàçíûõ ñòîðîí (ps — ïàïèëëîâèäíûå ñòðóêòóðû; sbl — ñóááàçàëüíàÿ ëîïàñòü ãëàíñà; swb —
ïîëîñàòî-ìîðùèíèñòûé ñêëåðîòèçèðîâàííûé ïîÿñîê); 10 — ëåçâèå ÿéöåêëàäà (cu — öåðêàëüíûé
÷ëåíèê ëåçâèÿ ÿéöåêëàäà); 11 — ñïåðìàòåêè; 12 — ÿéöî.

447
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…
Notes. “Ulidia rufifrons Loew” is an unpublished and unavailable name. The specimen from Sarepta
was depicted by Hennig (1940: Taf. 1, fig. 5) and presumably determined as “U. ?ruficeps Beck.” Dr. V. N.
Tanasijtshuk (in letter) examined and figured the holotype { of Ulidia ruficeps Becker (ZISP) on my request.
It was found that both specimens are conspecific. This species has much wider distribution in Middle Asia
(Kameneva, 2002 c) and to the Near East. I have examined the Lebanese specimen determined by Hennig,
in the TAU collection (borrowed by Dr. Freidberg in 1986 apparently from NHMW); it very probably
belongs to this species. Identity of the Italian material seems dubious and needs further examination. A } of
U. megacephala from “Spanien” (ZMHB) looks to have head shape and coloration like in U. ruficeps, but
certainly does not belong to it.
Ulidia salonikiensis Hennig, 1940
Hennig (1940): Greece; Belcari et al. (1995): Italy (south).
Material examined. Type. Holotype {: [Greece:] “Saloniki / 26 301”, “Miari / — smeca I”,
“Holotypus” [red label] “Ulidia / salonikiensis / n. sp. / det. Dr. W. Hennig 1939” (ZMHB); paratypes: 2 }:
[Greece:] “Saloniki / 26 299” and “ Saloniki / 26 299” [“Saloniki Turkei an Garry de N. Hough” according
to the Museum Catalogue], “Paratypus” (ZMHB) Non-type. Greece: Loutra — Langadas, oestl. Saloniki,
5.05.1942, {, 3 } (Babiy) (ZSSM).
Notes. I have not seen any specimens from Italy yet, but its presence in there is rather probable.
[Ulidia wadicola Steyskal, 1968] (fig. 3, 5)
Soo´s (1984): Egypt.
Material examined. Type. Paratypes 2 }: [Egypt:] “Wadi Wirak, northern Galala E. El Saff,
4.04.1937” (Tewfik) (USNM). Non-type. Non-European. Israel: Ein Nur, 31.03.1981, {, } (Freidberg),
10 km S Arat, Rt. 258, 30.04.1987, {, } (Freidberg) (SIZK); over 50 specimens from different localities,
04.1981–1998 (TAU).
Notes. Specimens from Israel fit the original description very well. There is considerable variability in
coloration of the face and parafacial, which is dark brown in 45–50% and 8–9% of Israeli specimens,
respectively. Frons is pitted and reddish in the antero-medial 1/3–1/2 of its length. The mesonotum is shining
black, very faintly shagreened. The male hindfemur is distinctly thickened, especially in its posterior half (fig. 3,
5), whereas in the female it is unmodified; Steyskal (1968) did not mention such a dimorphism. The mid- and
hindtarsi mostly yellow. No specimens were found in Europe.
erythrophthalma group of species
Diagnosis. Wing whitish or pale yellowish, without dark markings; frons mostly
shining black, with mesonotum shining or slightly shagreened; thoracic setae complete;
legs non-modified, with yellowish mid- and hindtarsi. Male genitalia: surstylus with
evenly narrowed basal part; cerci separated by very shallow fissure (fig. 6, 3; 7, 3–5);
phallus glans usually with membranous lobe (“vesica”) covered with numerous denti-
cles (“rasper-like”).
At least two described European (U. albidipennis and U. erythrophthalma) and se-
veral undescribed Western Asian species belong here. Reliable determination is based
on minor details of male phallus glans only, and females or non-dissected males were
found in this study to be undeterminable; females are not redescribed or figured here,
as no specimens in copula were been dissected, and no females have been associated
with males yet. Therefore, more detailed revision of the material throughout the
Palaearctic Region is needed.
Ulidia albidipennis Loew, 1845 (fig. 7, 8)
Hennig (1940): Greece; Belcari et al. (1995): Italy (mainland, south).
Material examined. Type. Syntypes 2 specimens (sex unknown, mostly destroyed: one wing is still
present): [Greece:] “Rhodus / Erber”, “Coll. / H. Loew”, “Type” [red label] (ZMHB); possible syntypes: {
[Greece?: uniformly bluish square, which means “Epirus”] “Coll. / H. Loew”, “Type” [red label] “albidi /
pennis / Lw” (ZMHB), }: [Greece:] “Epirus / Erber”, “Coll. / H. Loew”, “Type” [red label], “Ulidia /
albidipen- / nis. m.” (ZMHB); [sex unknown, only pin remains]: [Greece: yellow square] “Coll. H. Loew,
(ZMHB).

Non-type. Greece: Thessaly: Pieria: Lithohoro, Olympus, S slope, 800–850 m [40°05’N 22°30’E],
7.06.2002, { (dissected) (Kameneva, S. Korneyev & V. Korneyev) (ZISP); Trikala: Panagia, 15.06.1982, {
(dissected), idem, Orthovou´nion [39°46’N 21°28’E], 15.06.1982, 2 { (dissected) (Danielsson) (ZIL); Central
Greece: Foki´s: Giona Oros 3 km S Lefkadition, [38°36’N 22°13’E], 7.06.1982, 2 { (dissected) (Danielsson)
(ZIL); “Iti Oros [Mt. Iti] 2 km SE Iti”, 9.06.1982, { (dissected) (Danielsson) (ZIL); Rhodes: 5 km W
Maritsa, 15.06.1982, { (dissected), }, idem, Xalamon, 16.051983, 5 {, 5 }; Saroni, 20.05.1983, }, Kattavia,
448
E. P. Kameneva
Fig. 6. Ulidia erythrophthalma: 1— habitus, left view; 2— head and mesonotum, dorsal view; 3— epandrium,
posterior view; 4, 5 — phallus glans, different aspects (cl — claw-like structure, den — denticles, sbl —
subbasal lobe of glans, sb — sclerotized bar, sl — serrate lamina, ve — vesica, ys — Y-like sclerite).
Ðèñ. 6. Ulidia erythrophthalma: 1— îáùèé âèä ñëåâà; 2— ãîëîâà è ñðåäíåñïèíêà, âèä ñâåðõó; 3—
ýïàíäðèé, âèä ñçàäè; 4, 5 — ãëàíñ ôàëëþñà, âèäû ñ ðàçíûõ ñòîðîí (cl — êîãòåâèäíàÿ ñòðóêòóðà, den —
çóá÷èêè, sbl — ñóááàçàëüíàÿ ëîïàñòü ãëàíñà, sb — ñêëåðîòèçèðîâàííàÿ ïîëîñêà, sl — çóá÷àòàÿ
ïëàñòèíêà, ve — ïóçûðü, ys — Y-îáðàçíûé ñêëåðèò).

ASg. Pavlos, 21.05.1983, 2 }, 1 km E Psinthos, 25.05.1982, } (Danielsson) (ZIL). Slovakia: Kamenica,
Hronom, 1 km N, steppe, 47°50’N 18°43E, 200 m 15.07.1986, { (dissected) (Barta´k) (NMP).
Diagnosis. Very similar to U. erythrophthalma (Meigen) differing by the struc-
ture of the phallus apex (“glans”). In U. albidipennis, the glans has at least 3 sharply
pointed projections (fig. 8), of which one is a claw-like structure (cl), one is the longest
projection (lp) of the serrate lamina and one is a spine-like lobe (sl); it also has a curved
comb-like projection (cp) and poorly expressed rasper-like structure (weaker denticles)
on the vesica (ve). A few specimens from the mainland Greece have wings with light
brownish or yellowish tinge.
Notes. Type locality: “Rhodus” (Greece: Rhodes Is.) and “Mermeriza” (Turkey: Marmaris, 47 km to
the North of Rhodes). The specimens with the labels “Epirus /Erber” or blue squares could actually be
syntypes, possibly mislabelled by Loew, or the one labelled “Mermeriza” was indicated by Loew erroneously.
Ulidia erythrophthalma Meigen, 1826 (fig. 8)
Material examined. Austria: Burgerland, Zeilerberg, 28.06.1969, {, } (Necker) (ZSSM); France:
Briey, 3.07.1951, { (dissected), Puy-de-D
^
ome: Issoire, 20.07.1956, { (dissected) (RBINH); Greece: Greece:
N Kilkis, between Fanos and Skra, ca. 41°05’N 21°24’E, 4.06.2002, 2 { (dissected) (Kameneva, S. Korneyev
& V. Korneyev); Moldova: Chisina˘u: Institut of Biocontrol for Plant Protection: vicinity, 13.07.1987, 4 {,
2 } (V. Korneyev); Calarasovca, 8.07.1987, { (Vinokurov); Tigheci, vic. Budjak, 17–18.06.1987, 4 {
(dissected) (V. Korneyev) (SIZK); Cantemir Distr.: Cociulea, 18.06.1987, { (dissected), } (V. Korneyev)
(SIZK); Ukraine: Dnipropetrovsk Region: Sinelnikovo (no collecting data), {, } (Sumarokov); Odessa
Region: Maloye, vicinity, Starokozache Forestry, 19.06.1984, { (dissected), } (Dolin); Mykolaiv Region:
Pervomaysk District, Migia, 28.06.1993, 2 { (dissected) (V. Korneyev); Yelanets District, Kalinovka,
449
New and Little-Known Ulidiidae (Diptera, Tephritoidea)…
Fig. 7. Ulidia albidipennis: 1— habitus, left view; 2— head, anterior view; 3–5 — epandrium (3, 4 —
posterior view, 5— right view; 4, 5 — bleached).
Ðèñ. 7. Ulidia albidipennis: 1— îáùèé âèä ñëåâà; 2— ãîëîâà, âèä ñïåðåäè; 3–5 — ýïàíäðèé (3, 4 —
âèä ñçàäè, 5— âèä ñïðàâà; 4, 5 — îòáåëåíî).

22.06.1984, 3 { (dissected), } (Karachevskaja) (SIZK); Northern Crimea: Bokalska Kosa, 6.06.1990, {
(dissected), 2 } (Zrazhewsky) (NMP).
Diagnosis. Fits the diagnosis of the group differing by the structure of the phal-
lus glans. Its projections (fig. 6, 4, 5) are as follows: one is the claw-like structure (cl),
450
E. P. Kameneva
Fig. 8. Ulidia albidipennis: 1–5 — phallus glans, different aspects (cl — claw-like structure, sl — serrate lamina,
cp — curved comb-like projection, lp — longest projection, sbl — subbasal lobe of glans, ve — vesica).
Fig. 8. Ulidia albidipennis: 1–5 — ãëàíñ ôàëëþñà, âèäû ñ ðàçíûõ ñòîðîí (cl — êîãòåâèäíàÿ ñòðóêòóðà;
sl — çóá÷àòàÿ ïëàñòèíêà, cp — èñêðèâëåííûé ãðåáíåâèäíûé âûðîñò, lp — ñàìûé äëèííûé âûðîñò,
sbl — ñóááàçàëüíàÿ ëîïàñòü ãëàíñà, ve — ïóçûðü).

the serrate lamina (sl) with numerous denticles at base but without long spur, spine-
like lobe or a curved comb-like projection; an Y-like sclerite (ys) is formed by 2 round-
ed plates with common stem; rasper-like structure (weaker denticles) on the vesica (ve)
present; a membranous flap (mf) with sclerotized bar (sb) present.
Notes. Most of French, Germany and Polish specimens known to me apparently belong here, but they
were not available for study after the genitalic differences between this species and U. albidipennis were
recognised, and therefore need re-examination.
The only type specimen of Ulidia nitida Meigen, 1826 has not been located in this study. This species
was originally said to have entirely black tarsi. Th. Becker (1902) noted that the only female specimen from
the Baumhauer’s collection in the NHMW has the basal joints of hindleg yellow and does not differ otherwise
from U. erythrophthalma, and I still consider these names to be synonyms.
Ulidia semiopaca Loew, 1868
Material examined. Type. Holotype }: [France:] “Paris / Fairm”, “Coll. / H. Loew”, “Type” [red
label], “semi / opaca m.” (ZMHB).
Notes. The only known specimen fits the diagnosis of this group, differing from other species by
having tergites 1–4 almost entirely matt. This character does not look to be an artifact, and I consider U.
semiopaca as a separate species. I was unable to find any similar specimens among Ulidia collected in France.
Ulidia sp. near nigripennis
Material examined. Greece: N Kilkis, between Fanos and Skra, ca. 41°05’N 21°24’E, 4.06.2002,
} (Kameneva, S. Korneyev, V. Korneyev) (SIZK).
Diagnosis. Head black, except antero-medial one-third of frons dark orange to
reddish brown; no whitish microtrichose areas; frons pitted; parafacial black, as wide as
flagellomere 1; eye 1.2 times as high as wide; gena pitted; frontal and genal setulae
short, at most half as long as orbital and genal seta; mesonotum shining black; wing
uniformly brown including base; halter brown; legs black, except mid- and hindtarsi
yellowish; hindfemur very slightly swollen; hindtibia almost straight; abdomen shining
black, very finely shagreened, with a faint greenish tinge.
Notes. The only female fits the diagnosis of U. nigripennis in having shining mesonotum, well-
developed acrostichal seta and yellow mid- and hindtarsi, but has wing uniformly darkened and brown halter.
It cannot be identified with certainty based on a female specimen only.
I wish to express my sincere thanks to Andrew Whittington, Bernhard Merz and Gary Steck for reading
the manuscript and useful critical comments and language improvements and to Valery A. Korneyev for
preparation of digital photographs. Specimens studied were borrowed or examined through the kindness of
the following collection curators:
Miroslav Barta´k, Prague (specimens from private collection now transferred to MNP); David Grimaldi
and Tam Nguyen (AMNH); Joachim Ziegler (DEI and then ZMHB); La´szlo Papp (HMNH); Bernhard
Merz (MHNG); Pascal De Bleeker (MHN Lille); Peter Sehnal (NHMW); Christophe Daugeron and Cedric
Siebold (MNHNP); Patrick Grootaert and Pol Limbourg (RBINH); Hans-Peter Tschorsnig (SMNS);
Amnon Freidberg (TAU); Allen L. Norrbom and David Furth (USNM); Bradley Sinclair (ZFIB); Andrei
Ozerov and Anatoly Shatalkin (ZMUM); Marion Kotrba and Wolfgang Schacht (ZSSM).
I greatly appreciate the invaluable help from Amnon Freidberg, Bernhard Merz, Lita Greve Jensen,
Despina Vokou, F. Christian Thompson, Allen L. Norrbom, and Joachim Ziegler, who made possible my
visits to Tel Aviv, Geneva, Thessaloniki, Washington, Vienna and Berlin in 2000–2004.
This paper is a part of study supported by the travel grants from Curtis Sabrosky Fund (USNM & SEL
BARC USDA, Washington, D.C., U.S.A.) and Ernst Mayr Fund (Cambridge University, Cambridge, Ma.,
U.S.A.) in 2001, and by the Deutscher Akademischer Austauschdienst (Bonn, Germany) stipend in 2002.
The collecting trip to Greece in 2002 was funded by the FaunaEuropaea project through the kindness of
Dr. Thomas Pape.
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