Track analysis of the Mexican species of Cerambycidae (Insecta: Coleoptera)

Abstract

Track analysis of the Mexican species of Cerambycidae (Insecta, Coleoptera). A track analysis of 221
species belonging to 68 genera of Mexican Cerambycidae was undertaken in order to identify their main distributional patterns. Based on the comparison of the individual tracks, fifteen generalized tracks were obtained: six are placed in the Neotropical region, seven are shared by the Neotropical region and the Mexican Transition Zone, one is situated in the Mexican Transition Zone, and one is shared by the Nearctic region and the Mexican Transition Zone. Eight nodes were found in the intersection of these generalized tracks, five of them located in the Neotropical region and three in the Mexican Transition Zone. Distributional patterns of Mexican Cerambycidae show two basic patterns: one mostly Neotropical, in the Mesoamerican dominion (Mexican Pacific Coast and Mexican Gulf biogeographic provinces) and another in the Mexican Transition Zone (Transmexican Volcanic Belt and Balsas Basin biogeographic provinces).

Full text

131Track analysis of the Mexican species of Cerambycidae
Revista Brasileira de Entomologia 51(2): 131-137, junho 2007
Víctor H. Toledo1, Angélica Ma. Corona1 & Juan J. Morrone2*
Cerambycidae (longhorned beetles) represent one of the
largest families of wood-boring Coleoptera, with approximately
9,000 American species described from Alaska to Argentina.
Approximately 1,600 species have been recorded from Mexico
(Monné 2005a, b; Monné & Hovore 2005). The diversity of
Cerambycidae is also reflected in their coloration, body form
and adult morphology, with body length varying from ± 2.5
mm (Cyrtinus sp.) to slightly over 17 cm (Titanus giganteus).
Some species mimic ants (tribes Clytini and Tillomorphini),
bees and wasps (Rhinotragini), and lycid beetles (Pteroplatini).
Larvae are xylophagous and phytophagous, and play an
important role helping reduce dead and dying trees to humus
(Linsley 1961).
Taxonomic interest in Mexican Cerambycidae started in
1758 with some species described by Linnaeus (for details on
the taxonomic history in Mexico, see Noguera & Chemsak
1996), and has been constant especially in the last century.
Nevertheless, there are scarce distributional records (Noguera
& Chemsak 1996) and phylogenetic analyses including
Mexican taxa (Lingafelter 1998; Philips & Ivie 1998). Linsley
(1939, 1961) presented a summary of the world distribution of
Cerambycidae; he regarded climate and the availability of
suitable food plants as the main factors determining their
present occurrence, and the evolution and spread of past floras
as the major influence in the evolutionary history of the group.
Track analysis of the Mexican species of Cerambycidae (Insecta, Coleoptera)
1Centro de Educación Ambiental e Investigación Sierra de Huautla (CEAMISH), Universidad Autónoma del Estado de Morelos, Av. Universidad 1001,
Col. Chamilpa, Cuernavaca, 62210 Morelos, Mexico. victor.toledo@buzon.uaem.mx, acorona@buzon.uaem.mx
2Museo de Zoología, Departamento de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional Autónoma de México (UNAM), Apdo. postal 70-
399, 04510 Mexico-D.F., Mexico. jjm@hp.fciencias.unam.mx
*Corresponding author.
ABSTRACT. Track analysis of the Mexican species of Cerambycidae (Insecta, Coleoptera). A track analysis of 221
species belonging to 68 genera of Mexican Cerambycidae was undertaken in order to identify their main distributional
patterns. Based on the comparison of the individual tracks, fifteen generalized tracks were obtained: six are placed in the
Neotropical region, seven are shared by the Neotropical region and the Mexican Transition Zone, one is situated in the
Mexican Transition Zone, and one is shared by the Nearctic region and the Mexican Transition Zone. Eight nodes were
found in the intersection of these generalized tracks, five of them located in the Neotropical region and three in the
Mexican Transition Zone. Distributional patterns of Mexican Cerambycidae show two basic patterns: one mostly
Neotropical, in the Mesoamerican dominion (Mexican Pacific Coast and Mexican Gulf biogeographic provinces) and
another in the Mexican Transition Zone (Transmexican Volcanic Belt and Balsas Basin biogeographic provinces).
KEYWORDS. Biogeography; generalized tracks; Mexican Transition Zone; Neotropics; panbiogeography.
RESUMO. Análise de traço das espécies mexicanas de Cerambycidae (Insecta, Coleoptera). Uma análise de traço de 221
espécies de Cerambycidae mexicanos pertencentes a 68 gêneros foi feita com o objetivo de identificar seus principais padrões
de distribuição. Baseado na comparação de traços individuais, quinze traços generalizados foram obtidos: seis localizados na
região Neotropical, sete foram compartilhados entre a região Neotropical e a zona de transição mexicana, uma é situada na
zona de transição mexicana e uma compartilhada entre a região Neártica e a zona de transição mexicana. Oito nós
biogeográficos foram encontrados na intersecção dos traços biogeográficos generalizados, cinco deles localizados na
região Neotropical e três na zona de transição mexicana. Existem dois padrões de distribuição para os Cerambycidae mexicanos:
um principalmente Neotropical, no domínio Mesoamericano (províncias da costa pacífica mexicana e do golfo mexicano)
e outro na zona de transição mexicana.
PALAVRAS-CHAVE. Biogeografia; Neotrópico; panbiogeografia; traços generalizados; Zona de transição Mexicana.
Our objective is to describe the distributional patterns of
221 species of the family Cerambycidae in Mexico applying a
panbiogeographic analysis.
MATERIAL AND METHODS
We analyzed distributional data for 221 species of
cerambycids (see Appendix), which correspond to 68 genera,
most of them (78%) having Neotropical biogeographic
affinities. Several species (63) presented few localities (one or
two) and were excluded from the analysis. Distributional
records analyzed for these species were only from Mexico,
data from other countries were not included in the analysis.
Distributional data for this study were obtained from the
available literature (Bates 1879; Chemsak 1963a,b, 1964, 1967,
1969a,b, 1972, 1977, 1980, 1991, 1999; Chemsak & Hovore 2002;
Chemsak & Linsley 1963a,b, 1964a,b, 1965, 1966, 1967, 1975a,b,
1976a,b, 1982a,b, 1983, 1984, 1986, 1988; Chemsak & McCarty
1997; Chemsak & Noguera 1997, 1998, 2001, 2003; Giesbert
1979, 1985, 1986, 1992, 1993; Giesbert & Chemsak 1997; Giesbert
& Wappes 1999; Linsley 1962, 1970; Martins & Chemsak
1966a,b; McCarty 2001; Noguera 1993, 2002; Noguera &
Chemsak 1997; Noguera et al. 2002; Toledo 1997, 2005a,b;
Toledo et al. 2002).
We applied a panbiogeographic analysis, which basically

132 Toledo et al.
Revista Brasileira de Entomologia 51(2): 131-137, junho 2007
consists of plotting distributions of each species on maps,
and connecting their localities together with the nearest locality
via minimum distance lines. After many species have been
added to the data set, the overlapping of individual tracks
provides generalized tracks, which allow us to hypothesize
the pre-existence of ancestral biotic components that have
been fragmented by tectonic or climatic changes (Croizat 1958,
1964; Morrone & Crisci 1995; Craw et al. 1999; Morrone 2004).
If two or more generalized tracks intersect in a given area, they
determine a node, which indicates a complex area where
different ancestral biotic and geological elements interrelate
in time and space. Individual and generalized tracks and nodes
were represented on maps of the Mexican biogeographic
provinces (Morrone 2005, 2006), using ArcView 3.2 (ESRI 1998).
RESULTS
Generalized tracks. A total of 221 individual tracks were
constructed. Based on their overlapping, we obtained 15
generalized tracks (Fig. 1). Six are situated in the Neotropical
region, seven are shared by the Neotropical region and the
Mexican Transition Zone, one is situated in the Mexican
Transition Zone, and one is shared by the Nearctic region and
the Mexican Transition Zone (Morrone & Márquez 2003;
Morrone 2005, 2006).
Generalized track 1. Southeastern Sonora to central
southern Jalisco. Mexican Transition Zone (Transmexican
Volcanic Belt biogeographic province) and Neotropical region,
Mesoamerican dominion (Mexican Pacific Coast biogeographic
province). Based on Anelaphus badius, Cacophrissus paupe,
Colobothea sinaloensis, Dectes nigripilus, Eburia laticollis,
E. powelli, Ecyrus pacificus, Elytroleptus scabricollis,
Euderces bicinctus, E. longicollis, E. pulcra, Lagocheirus
obsoletus, Lophalia prolata, Methia occidentalis, Metironeus
hesperus, Neocompsa alacris, N. tenuisima, Neotaranomis
sinaloae, Oreodera brailovski, Phaea kaitlinae, P. marthae,
Psyrassa cylindricollis, P. megalops, P. nigricornis, P.
sinaloae, Rhodoleptus comis, R. femoratus, Sphaenothecus
trilineatus, Strangalia palaspina, Tetraopes subfasciatus,
Thryallis noguerai, and Triacetelus sericatus.
Generalized track 2. Central southern to southwestern
Jalisco. Mexican Transition Zone (Transmexican Volcanic Belt
biogeographic province) and Neotropical region,
Mesoamerican dominion (Mexican Pacific Coast biogeographic
province). Based on Acanthoderes noguerai, Ischnocnemis
similis, Lagocheirus obsoletus, Oncideres albomarginata, O.
scitula, Oreodera corticina, Phaea carnelia, P. juanitae, P.
tenuata, and Psyrassa katsurae.
Generalized track 3. Southwestern Jalisco to southwestern
Guerrero. Neotropical region, Mesoamerican dominion
(Mexican Pacific Coast biogeographic province). Based on
Acanthoderes ramirezi, Eburia chemsaki, E. clara, E.
hatsuae, E. juanitae, E. laticollis, E. maccartyi, E. nigrovittata,
E. paraegrota, E. powelli, Lagocheirus obsoletus, Lophalia
prolata, Neoperiboeum juanitae, Phaea flavovittata, P. hogei,
P. juanitae, P. marthae, Poliaenus hesperus, Psyrassa
basicornis, P. cribricollis, P. cylindricollis, P. katsurae, P.
levicollis, P. nigricornis, P. sthenias, Tetranodus copei, and
Tylosis puncticollis.
Generalized track 4. Southwestern Guerrero to southeastern
Oaxaca. Neotropical region, Mesoamerican dominion (Mexican
Pacific Coast biogeographic province). Based on Eburia clara,
E. laticollis, E. maccartyi, E. ribardoi, Euderces batesi,
Lagocheirus obsoletus, Phaea hogei, P. kellyae, Psyrassa
basicornis, P. sthenias, Sphaenothecus trilineatus, and
Stenobatyle prolixa.
Generalized track 5. Southeastern Oaxaca to northeastern
Chiapas. Neotropical region, Mesoamerican dominion
(Mexican Pacific Coast and Chiapas biogeographic provinces).
Based on Aneflus poriferus, Choriolaus howdeni, Eburia
clara, E. laticollis, E. ribardoi, E. schusteri, Ecyrus lineicollis,
Enaphalodes coronatus, Euderces boucardi, E. disparicus,
E. reticulates, E. turnbowi, Lagocheirus cristulatus, L.
simplicicornis, L. obsoletus, Metironeus hovorei, Neocompsa
alacris, N. clerochroa, N. exclamationis, Oncideres ocellaris,
Phaea biplagiata, P. bryani, P. flavovittata, P. helayae, P.
hogei, P. kellyae, P. maccartyi, P. miniata, P. pthistica, P.
semirufa, P. tenuata, P. wappesi, Psyrassa angelicae, P.
basicornis, P. levicollis, P. oaxacae, P. sthenias,
Sphaenothecus argenteus, S. toledoi, S. trilineatus,
Sphaerionillum castaneum, Stenobatyle prolixa, Strangalia
cavaventra, and Tetranodus copei.
Generalized track 6. Northeastern to southeastern Chiapas.
Neotropical region, Mesoamerican dominion (Mexican Pacific
Coast and Chiapas biogeographic provinces). Based on
Eburia laticollis, Lagocheirus cristulatus, L. integer, L.
simplicicornis, L. obsoletus, Neocompsa exclamationis,
Oreodera corticina, O. fasciculosa, O. wappesi, Phaea
biplagiata, and P. helayae.
Generalized track 7. Northeastern to eastern Chiapas.
Neotropical region, Mesoamerican dominion (Chiapas
biogeographic province). Based on Anthoboscus oculatus,
Anatinoma alveolatum, Assycuera macrotela, Choriolaus
howdeni, Eburia brevispinis, E. schusteri, Euderces bellus,
E. boucardi, E. disparicus, E. laevicauda, E. turnbowi, E.
wappesi, Megapsyrassa testacea, Oncideres ocellaris, Phaea
kellyae, P. tenuata, P. wappesi, Plocaederus yucatecus,
Psyrassa sthenias, Rhodoleptus nigripennis, Sphaenothecus
argenteus, and S. toledoi.
Generalized track 8. Southeastern Veracruz to northwestern
Chiapas. Neotropical region, Mesoamerican dominion
(Mexican Gulf and Chiapas biogeographic provinces). Based
on Eburia brevispinis, E. cruciata, Euderces reticulatus,
Lagocheirus integer, L. binumeratus, Oncideres
albomarginata, O. fisheri, O. putator putator, O. rubra,
Oreodera corticina, O. fasciculosa, O. wappesi, Phaea
acromela, P. pthistica, and Tetraopes varicornis.
Generalized track 9. Southeastern San Luis Postosí to
southeastern Veracruz. Mexican Transition Zone (Sierra Madre
Oriental biogeographic province) and Neotropical region,
Mesoamerican dominion (Mexican Gulf biogeographic
province). Based on Anatinoma alveolatum, Eburia

133Track analysis of the Mexican species of Cerambycidae
Revista Brasileira de Entomologia 51(2): 131-137, junho 2007
brevispinis, E. mutica, Lagocheirus binumeratus,
Micropsyrassa bimaculata, Neocompsa exclamationis,
Oncideres cingulata texana, O. rubra, Phaea acromela, P.
saperda, Psyrassa castanea, P. cribricollis, P. tympanophora,
and Tetraopes varicornis.
Generalized track 10. Northwestern Nuevo León to
southeastern San Luis Potosí. Nearctic region (Tamaulipas
biogeographic province) and Mexican Transition Zone (Sierra
Madre Oriental biogeographic province). Based on Eburia
mutica, Lagocheirus obsoletus, Meloemorpha aliena,
Neoptychodes trilineatus, Oncideres cingulata texana,
Oreodera corticina, Phaea acromela, P. tenuata, Psyrassa
brevicornis, and P. sallaei.
Generalized track 11. Central and southern Jalisco to central
Morelos. Mexican Transition Zone (Transmexican Volcanic
Belt and Balsas Basin biogeographic provinces). Based on
Acanthoderes noguerai, Ancylocera michelbacheri,
Aneflomorpha crinita, Dectes nigripilus, Deltaspis cyanipes,
Dexithea humeralis, Eburia poricollis, Euderces
basimaculatus, E. cribripennis, Hexoplon calligrammum,
Lophalia prolata, Megaderus bifasciatus, Neocompsa
tenuissima, Oncideres scitula, Phaea bryani, P. carnelia, P.
juanitae, P. semirufa, P. tenuata, Tetraopes subfasciatus, and
T. varicornis.
Generalized track 12. Central Morelos to southwestern
Veracruz. Mexican Transition Zone (Balsas Basin and
Transmexican Volcanic Belt biogeographic provinces) and
Neotropical region, Mesoamerican dominion (Mexican Gulf
biogeographic province). Based on Assycuera macrotela,
Cirrhicera leuconota, Dexithea humeralis, Elytroleptus
similis, Euderces batesi, Hexoplon calligrammum,
Lagocheirus funestus, L. integer, L. binumeratus, L. obsoletus,
Meloemorpha aliena, Neocompsa clerochroa, Neoptychodes
trilineatus, Phaea acromela, P. pthistica, and Tetraopes
umbonatus.
Generalized track 13. Central Morelos to southeastern
Oaxaca. Mexican Transition Zone (Balsas Basin and
Transmexican Volcanic Belt biogeographic provinces) and
Neotropical region, Mesoamerican dominion (Mexican Pacific
Coast biogeographic province). Based on Ancylocera
michelbacheri, Anelaphus hirtus, Dectes nigripilus, Eburia
chemsak, E. cruciata, E. hatsuae, E. poricollis, Elytroleptus
scabricollis, Euderces batesi, E. laevicauda, E. longicollis,
E. perplexus, E. postipalidus, Ischnocnemis costipennis,
Metironeus hesperus, Micropsyrassa reticulata, Ochraethes
nigropunctatus, Phaea biplagiata, Psyrassa nigroaenea,
Rhodoleptus comis, Sphaenothecus picticornis, Tetraopes
cleroides, T. subfasciatus, and Tylosis puncticollis.
Generalized track 14. Southwestern Jalisco to Central
Morelos. Mexican Transition Zone (Balsas Basin and
Transmexican Volcanic Belt biogeographic provinces) and
Neotropical region, Mesoamerican dominion (Mexican Pacific
Coast biogeographic province). Based on Alphomorphus
vandykei, Anelaphus hirtus, Deltaspis rubriventris, Eburia
aliciae, E. cruciata, E. hatsuae, E. nigrovittata, E.
rotundipennis, Elytroleptus scabricollis, Euderces
cribripennis, E. longicollis, E. pulcra, Lagocheirus xileuco,
Megaderus bifasciatus, Neocompsa alacris, N. clerochroa,
Neoptychodes trilineatus, Oncideres senilis, Oreodera
brailovski, O. glauca glauca, Phaea biplagiata, P. carnelia,
P. flavovittata, P. hogei, P. maxima, Psyrassa chamelae, P.
megalops, P. sinaloae, Sphaenothecus argenteus, S.
picticornis, S. trilineatus, and Tetraopes umbonatus.
Generalized track 15. Central Morelos to southeastern
Fig. 1. Generalized tracks and nodes obtained from the biogeographic analysis of 221 species belonging to 68 genera of Mexican Cerambycidae.

134 Toledo et al.
Revista Brasileira de Entomologia 51(2): 131-137, junho 2007
Guerrero. Mexican Transition Zone (Balsas Basin and
Transmexican Volcanic Belt biogeographic provinces) and
Neotropical region, Mesoamerican dominion (Mexican Pacific
Coast biogeographic province). Based on Alphomorphus
vandykei, Eburia chemsaki, E. nigrovittata, Euderces
longicollis, E. perplexus, Hexoplon calligrammum,
Ischnocnemis costipennis, Lagocheirus funestus, L. obsoletus,
L. xileuco, Micropsyrassa reticulata, Neocompsa alacris,
Oncideres senilis, Phaea kellyae, P. semirufa, P. tenuata, and
Psyrassa cribricollis.
Nodes. We found eight nodes in the areas where the
generalized tracks intersected (Fig. 1):
Node A. Intersection of generalized tracks 2, 3, and 14.
Southwestern Jalisco. Neotropical region, Mesoamerican
dominion (Mexican Pacific Coast biogeographic province).
Node B. Intersection of generalized tracks 1, 2, and 11.
Central south Jalisco. Mexican Transition Zone (Transmexican
Volcanic Belt biogeographic province).
Node C. Intersection of generalized tracks 9 and 10.
Southeastern San Luis Potosí. Mexican Transition Zone (Sierra
Madre Oriental biogeographic province).
Node D. Intersection of generalized tracks 11, 12, 13, 14,
and 15. Northwestern Morelos. Mexican Transition Zone
(Transmexican Volcanic Belt and Balsas Basin biogeographic
provinces).
Node E. Intersection of generalized tracks 3, 4, and 15.
Southwestern Guerrero. Neotropical region, Mesoamerican
dominion (Mexican Pacific Coast biogeographic province).
Node F. Intersection of generalized tracks 8, 9, and 12.
Southeastern Veracruz. Neotropical region, Mesoamerican
dominion (Mexican Gulf biogeographic province).
Node G. Intersection of generalized tracks 4, 5, and 13.
Southeastern Oaxaca. Neotropical region, Mesoamerican
dominion (Mexican Pacific Coast biogeographic province).
Node H. Intersection of generalized tracks 5, 6, 7, and 8.
Northeastern Chiapas. Neotropical region, Mesoamerican
dominion (Chiapas biogeographic province).
DISCUSSION
Distributional patterns identified herein were mostly
Neotropical. We found two main distributional patterns for
the species analyzed, which are related to their host plants
distribution. One of them, in the lowlands of the Mesoamerican
dominion, where eight generalized tracks and five nodes were
obtained, is associated to the tropical dry forests of the
Mexican Pacific Coast and Chiapas biogeographic provinces,
and tropical rain forests of the Mexican Gulf (Rzedowski 1978).
The other, in the highlands of the Mexican Transition Zone,
where seven generalized tracks and three nodes were
established, is associated to montane forests in the
Transmexican Volcanic Belt and tropical dry forests of the
Balsas Basin biogeographic province. In general,
Cerambycidae in Mexico have been better collected in the
tropical dry forests than in other kinds of vegetation (Chemsak
& Noguera 1993; Noguera et al. 2002; Toledo et al. 2002).
The convergence of five generalized tracks in node D,
which is situated in the Mexican Transition Zone
(Transmexican Volcanic Belt- Balsas Basin biogeographic
provinces), supports the hypothesis that it is a complex area
where biotic elements with different origins overlap (Halffter
1987; Morrone & Márquez 2001). The absence of generalized
tracks in the Nearctic region may indicate the lack of elements
with Nearctic biogeographic affinities. Therefore, the results
obtained are considered preliminary, because it would be
necessary to include more distributional records from different
national and international collections, recent systematic
collects, and new species records with different biogeographic
affinities, in order to complement the work and corroborate
our hypothesis.
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APPENDIX
List of species of the family Cerambycidae (Coleoptera) analyzed,
with references consulted between square brackets.
1. Acanthoderes albifrons Chemsak & Hovore [Chemsak & Hovore
2002].
2. A. amplitoris Chemsak & Hovore [Chemsak & Hovore 2002].
3. A. bicolor Chemsak & Hovore [Chemsak & Hovore 2002].
4. A. linsleyi Chemsak & Hovore [Chemsak & Hovore 2002].
5. A. noguerai Chemsak & Hovore [Chemsak & Hovore 2002].
6. A. ramirezi Chemsak & Hovore [Chemsak & Hovore 2002].
7. Alphomorphus vandykei (Linsley) [Chemsak & Linsley 1975b].
8. Anatinomma alveolatum Bates [Chemsak & Linsley, 1963b 1964].
9. Ancylocera michelbacheri Chemsak [Chemsak 1963a].
10. Aneflomorpha crinita Chemsak & Linsley [Chemsak & Linsley
1975a].
11. A. ruficollis Chemsak & Linsley [Chemsak & Linsley 1975a].
12. Aneflus glabropunctatus Chemsak & Linsley [Chemsak & Linsley
1963a].
13. A. minutivestis Chemsak & Linsley [Chemsak & Linsley 1963a].
14. A. poriferus Giesbert [Giesbert 1993].
15. Anelaphus badius Chemsak [Chemsak 1991].
16. A. hirtus Chemsak & Noguera [Chemsak & Noguera 2003].
17. Anthoboscus oculatus Giesbert [Giesbert 1992; Toledo et al. 2002].
18. A. tricolor (Chevrolat) [Toledo 2005b].
19. Assycuera macrotela Bates [Chemsak 1964].
20. Cacophrissus pauper Bates [Chemsak & Linsley 1963b].
21. Cacostola janzeni Chemsak & Linsley [Chemsak & Linsley 1986].
22. Ceralocyna cribricollis (Bates) [Chemsak 1963a].
23. Cirrhicera cristipennis Bates [Chemsak 1972].
24. C. leuconota Laporte [Chemsak 1972].
25. Colobothea sinaloensis Giesbert [Giesbert 1979].
26. Crossidius mexicanus Chemsak & Noguera [Chemsak & Noguera
1997].
27. Championa elegans Chemsak [Chemsak 1967].
28. C. westcotti Noguera & Chemsak [Noguera & Chemsak 1997].
29. Choriolaus howdeni Giesbert & Wappes [Giesbert & Wappes 1999].
30. Dectes nigripilus Chemsak & Linsley [Chemsak & Linsley 1986].
31. Deltaspis cyanipes Bates [Toledo 2005b].
32. D. rubriventris Bates [Toledo 2005b].
33. Dexithea humeralis Chemsak & Noguera [Chemsak & Noguera
2001].
34. Eburia aegrota Bates [Noguera 2002].
35. E. aliciae Noguera [Noguera 2002].
36. E. baroni Bates [Noguera 2002].
37. E. blancaneaui Bates [Noguera 2002].
38. E. brevicornis Chemsak & Linsley [Noguera 2002].
39. E. brevispinis Bates [Noguera 2002].
40. E. bruneicomis Chemsak & Linsley [Noguera 2002].
41. E. clara Bates [Noguera 2002].
42. E. cruciata (Linsley ) [Noguera 2002].
43. E. cubae (Fisher) [Noguera 2002].
44. E. championi Bates [Noguera 2002].
45. E. chemsaki Noguera [Noguera 2002].
46. E. falli Linsley [Noguera 2002].
47. E. fuliginea (Bates) [Noguera 2002].
48. E. haldemani LeConte [Noguera 2002].
49. E. hatsueae Chemsak & Giesbert [Noguera 2002].
50. E. juanitae Chemsak & Linsley [Noguera 2002].
51. E. laticollis Bates [Noguera 2002].
52. E. maccartyi Noguera [Noguera 2002].
53. E. mutica LeConte [Noguera 2002].
54. E. nigrovittata Bates [Noguera 2002].
55. E. opaca Chemsak & Linsley [Noguera 2002].
56. E. ovicollis LeConte [Noguera 2002].
57. E. paraegrota Chemsak & Linsley [Noguera 2002].
58. E. patruelis Bates [Noguera 2002].
59. E. pedestris White [Noguera 2002].
60. E. poricollis Chemsak & Linsley [Noguera 2002].
61. E. porulosa Bates [Noguera 2002].
62. E. powelli Chemsak & Linsley [Noguera 2002].
63. E. ribardoi Noguera [Noguera 2002].
64. E. rotundipennis Bates [Noguera 2002].
65. E. schusteri Giesbert [Giesbert 1993; Noguera 2002].
66. Ecyrus ciliatus Chemsak & Linsley [Chemsak & Linsley 1975b].
67. E. lineicollis Chemsak & Linsley [Chemsak & Linsley 1975b].
68. E. pacificus Linsley [Chemsak & Linsley 1975b].
69. Echthistatus spinosus Pascoe [Chemsak & Linsley 1983].
70. Elytroleptus apicalis LeConte [Chemsak & Linsley 1965; Linsley
1962].
71. E. divisus (LeConte) [Linsley 1962].
72. E. ignitus (LeConte) [Linsley 1962].
73. E. pallidus pallidus (Thomson) [Chemsak & Linsley 1965; Linsley
1962].
74. E. scabricollis Bates [Chemsak & Linsley 1965; Linsley 1962].
75. E. similis Chemsak & Linsley [Chemsak & Linsley 1965].
76. Enaphalodes coronatus (White) [Toledo 2005b].
77. Erichsonia dentifrons Westwood [Toledo 2005b].
78. Erosida yucatana Giesbert [Giesbert 1985].
79. Euderces batesi Giesbert & Chemsak [Giesbert & Chemsak 1997].
80. E. basimaculatus Giesbert & Chemsak [Giesbert & Chemsak
1997].
81. E. bellus Giesbert & Chemsak [Giesbert & Chemsak 1997].
82. E. bicinctus (Linsley) [Chemsak 1969b].
83. E. biplagiatus Giesbert & Chemsak [Giesbert & Chemsak 1997].
84. E. boucardi (Chevrolat) [Chemsak 1969b].
85. E. brailovskyi Giesbert & Chemsak [Giesbert & Chemsak 1997].
86. E. cribripennis Bates [Chemsak 1969b].
87. E. disparicus Giesbert & Chemsak [Giesbert & Chemsak 1997].
88. E. laevicauda Bates [Chemsak 1969b; Giesbert & Chemsak 1997].
89. E. longicollis (Linsley) [Linsley 1935; Chemsak 1969b].
90. E. noguerai Giesbert & Chemsak [Giesbert & Chemsak 1997].
91. E. perplexus Giesbert & Chemsak [Giesbert & Chemsak 1997].
92. E. postipallidus Giesbert & Chemsak [Giesbert & Chemsak 1997].
93. E. pulcher (Bates) [Chemsak 1969b; Giesbert & Chemsak 1997].
94. E. reichei LeConte [Chemsak 1969b; Giesbert & Chemsak 1997].
95. E. reticulatus (Bates) [Giesbert & Chemsak 1997].

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96. E. tibialis Giesbert & Chemsak [Giesbert & Chemsak 1997].
97. E. turnbowi Giesbert & Chemsak [Giesbert & Chemsak 1997].
98. E. wappesi Giesbert & Chemsak [Giesbert & Chemsak 1997].
99. Giesbertia rugosa Chemsak & Linsley [Chemsak & Linsley 1984].
100.Heterachthes integripennis (Bates) [Martins & Chemsak 1966a].
101.Hexoplon calligrammum Bates [Martins & Chemsak 1966a].
102.Ischnocnemis costipennis Thomson [Toledo 2005b].
103.I. similis Chemsak & Noguera [Chemsak & Noguera 1997].
104.Lagocheirus binumeratus Thomson [Toledo 1997].
105.L. cristulatus Bates [Toledo 1997].
106.L. funestus Thomson [Toledo 1997].
107.L. integer Bates [Toledo 1997].
108.L. lugubris Dillon [Toledo 1997].
109.L. obsoletus Thomson [Toledo 1997].
110.L. procerus Casey [Toledo 1997].
111.L. simplicicornis Bates [Toledo 1997].
112.L. xileuco Toledo [Toledo 1997].
113.Lasiogaster costipennis Gahan [Toledo 2005b].
114.Lochmaeocles cretatus Chemsak & Linsley [Chemsak & Linsley
1986].
115.L. nigritarsus Chemsak & Linsley [Chemsak & Linsley 1986].
116.Lophalia prolata Chemsak & Linsley [Chemsak & Linsley 1988].
117.Megaderus bifasciatus Dupont [Toledo 2005b].
118.Megapsyrassa testacea Giesbert [Giesbert 1993].
119.Meloemorpha aliena (Bates) [Chemsak & Linsley 1976a].
120.Methia occidentalis Chemsak & Linsley [Chemsak & Linsley
1964b].
121.Metironeus hesperus Chemsak [Chemsak 1991].
122.M. hovorei Chemsak [Chemsak 1991].
123.Micropsyrassa bimaculata (Bates) [Martins & Chemsak 1966b].
124.M. reticulata Martins & Chemsak [Martins & Chemsak 1966b].
125.Neocompsa alacris (Bates) [Martins & Chemsak 1966a].
126.N. clerochroa (Thomson ) [Martins & Chemsak 1966a].
127.N. exclamationis (Thomson) [Martins & Chemsak 1966a].
128.N. tenuissima (Bates) [Martins & Chemsak 1966a].
129.Neoleptura alpina Chemsak & Linsley [Chemsak & Linsley 1976a].
130.Neoperiboeum juanitae Chemsak [Chemsak 1991].
131.Neoptychodes trilineatus Linnaeus [Toledo 2005b].
132.Neospondylis mexicanus (Bates) [Bates 1879].
133.Neotaranomis sinaloae Chemsak & Linsley [Chemsak & Linsley
1982b].
134.Ochraethes nigropunctatus Chevrolat [Noguera et al. 2002; Toledo
2005b].
135.O. octomaculata Chemsak & Noguera [Chemsak & Noguera 2001].
136.Ochraethes tomentosus (Chevrolat) [Toledo 2005b].
137.Oncideres albipilosa Noguera [Noguera 1993].
138.O. albomarginata chamela Chemsak & Giesbert [Noguera 1993].
139.O. cingulata texana Horn [Noguera 1993].
140.O. fisheri Dillon & Dillon [Noguera 1993].
141.O. ocellaris Bates [Noguera 1993].
142.O. putator putator Thomson [Noguera 1993].
143.O. rubra Franz [Noguera 1993].
144.O. scitula Bates [Noguera 1993].
145.O. senilis Bates [Noguera 1993].
146.Oreodera brailovskyi Chemsak & Noguera [McCarty 2001].
147.O. copei McCarty [McCarty 2001].
148.O. corticina Thomson [McCarty 2001].
149.O. fasciculosa Thomson [McCarty 2001].
150.O. glauca glauca (Linnaeus) [McCarty 2001].
151.O. wappesi McCarty [McCarty 2001].
152.Phaea acromela Pascoe [Chemsak 1999].
153.P. biplagiata Chemsak [Chemsak 1977].
154.P. bryani Chemsak [Chemsak 1999].
155.P. carnelia Chemsak & Linsley [Chemsak 1999].
156.P. eyai Chemsak [Chemsak 1999].
157.P. flavovittata Bates [Chemsak 1999].
158.P. haleyae Chemsak [Chemsak 1999].
159.P. hogei Bates [Chemsak 1999].
160.P. juanitae Chemsak & Linsley [Chemsak 1999].
161.P. kaitlinae Chemsak [Chemsak 1999].
162.P. kellyae Chemsak [Chemsak 1999].
163.P. maccartyi Chemsak [Chemsak 1999].
164.P. marthae Chemsak [Chemsak 1999].
165.P. maryannae Chemsak [Chemsak 1977].
166.P. maxima Bates [Chemsak 1999].
167.P. miniata Pascoe [Chemsak 1999].
168.P. phthisica Bates [Chemsak 1999].
169.P. saperda Newman [Chemsak 1999].
170.P. semirufa Bates [Chemsak 1999].
171.P. tenuata Bates [Chemsak 1999].
172.P. tricolor Bates [Chemsak 1999].
173.P. wappesi Chemsak [Chemsak 1999].
174.Platerosida howdeni Linsley [Linsley 1970].
175.Plectromerus wappesi Giesbert [Giesbert 1985].
176.Plocaederus yucatecus (Chemsak & Noguera) [Chemsak &
Noguera 1997].
177.Poliaenus concolor (Schaeffer) [Chemsak & Linsley 1975b].
178.P. hesperus Chemsak & Linsley [Chemsak & Linsley 1988].
179.Psyrassa aliena (Linsley) [Toledo 2005a].
180.P. angelicae (Toledo) [Toledo 2005a].
181.P. basicornis Pascoe [Toledo 2005a].
182.P. brevicornis Linsley [Toledo 2005a].
183.P. castanea Bates [Toledo 2005a].
184.P. cribricollis (Bates) [Toledo 2005a].
185.P. cylindricollis Linsley [Toledo 2005a].
186.P. chamelae Toledo [Toledo 2005a].
187.P. chemsaki Toledo [Toledo 2005a].
188.P. katsurae Chemsak & Noguera [Toledo 2005a].
189.P. levicollis Chemsak & Noguera [Toledo 2005a].
190.P. megalops Chemsak & Noguera [Toledo 2005a].
191.P. nigricornis Bates [Toledo 2005a].
192.P. nigroaenea Bates [Toledo 2005a].
193.P. oaxacae Toledo [Toledo 2005a].
194.P. sallaei Bates [Toledo 2005a].
195.P. sinaloae Linsley [Toledo 2005a].
196.P. sthenias Bates [Toledo 2005a].
197.P. tympanophora Bates [Toledo 2005a].
198.Rhodoleptus comis (Bates) [Chemsak & Linsley 1982a].
199.R. femoratus (Schaeffer) [Chemsak & Linsley 1982a].
200.R. nigripennis Giesbert [Giesbert 1993].
201.Sphaenothecus argenteus Bates [Chemsak & Noguera 1998].
202.S. picticornis Bates [Chemsak & Noguera 1998].
203.S. trilineatus Dupont [Chemsak & Noguera 1998].
204.Sphaerionillum castaneum Chemsak & Linsley [Chemsak &
Linsley 1967].
205.Stenobatyle prolixa (Bates) [Chemsak 1980].
206.Strangalia auripilis Chemsak [Chemsak 1969a].
207.S. cavaventra Chemsak [Chemsak 1969a].
208.S. hamatipes Giesbert [Giesbert 1986].
209.S. palaspina Chemsak [Chemsak 1969a].
210.Tetranodus copei Chemsak & Linsley [Chemsak & Linsley 1988].
211.T. niveicollis Linell [Chemsak 1969b].
212.Tetraopes cleriodes Thomson [Chemsak 1963b].
213.T. subfasciatus Bates [Chemsak 1963b].
214.T. thoreyi Bates [Chemsak 1963b].
215.T. umbonatus LeConte [Chemsak 1963b].
216.T. varicornis Laporte [Chemsak 1963b].
217.Thryallis leucophaeus (White) [Chemsak & McCarty 1997].
218.T. noguerai Chemsak & McCarty [Chemsak & McCarty 1997].
219.Tigrinestola howdeni Chemsak & Linsley [Chemsak & Linsley
1966].
220.Triacetelus sericatus Bates [Chemsak & Linsley 1976b].
221.Tylosis puncticollis Bates [Toledo 2005b].
Received 31/08/2006; accepted 02/03/2007