Euro+Med Notulae, 3

Abstract

This is the third of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to the Euro+Med (or Sisyphus) Project are presented. It is entirely devoted to the Compositae family, including new country and area records for taxa of Centaurea, Crepis, Dittrichia, Hieracium, Rudbeckia, Silybum and Taraxacum, and the validation of names in the genera Adenostyles, Andryala, Centaurea, Cichorium, Erigeron, Filago, Hieracium, Jacobaea, Picris, Rhodanthemum, Schlagintweitia, Senecio, Taraxacum and Tragopogon. For Hieracium (s.str.), a novel, coherent classification policy on a Euro-Mediterranean scale is proposed, the implementation of which results in a substantial amount of nomenclatural changes.

Full text

Notulae ad floram euro-mediterraneam pertinentes No. 25
WERNER GREUTER & ECKHARD VON RAAB-STRAUBE (ed.)
Euro+Med Notulae, 3
Abstract
Greuter, W. & Raab-Straube, E. von (ed.): Euro+Med Notulae, 3 [Notulae ad floram euro-medi-
terraneam pertinentes 25]. – Willdenowia 37: 139-189. – ISSN 0511-9618; © 2007 BGBM Berlin-
Dahlem.
doi:10.3372/wi.37.37107 (available via http://dx.doi.org/)
This is the third of a series of miscellaneous contributions, by various authors, where hitherto unpub-
lished data relevant to the Euro+Med (or Sisyphus) Project are presented. It is entirely devoted to the
Compositae family, including new country and area records for taxa of Centaurea, Crepis, Dittrichia,
Hieracium, Rudbeckia, Silybum and Taraxacum, and the validation of names in the genera Adeno-
styles, Andryala, Centaurea, Cichorium, Erigeron, Filago, Hieracium, Jacobaea, Picris, Rhodanthe-
mum, Schlagintweitia, Senecio, Taraxacum and Tragopogon. For Hieracium (s. str.), a novel, coherent
classification policy on a Euro-Mediterranean scale is proposed, the implementation of which results
in a substantial amount of nomenclatural changes.
Key words: Europe, Mediterranean area, Compositae, Asteraceae, taxonomy, nomenclature, distribu-
tion, Hieracium, classification principles.
Notice
A succinct description of the Euro+Med Project, with a list of recognised territories and their ab-
breviations, and the conventions used to indicate the status and presence of taxa, can be found in
the introduction to the first instalment (in Willdenowia 35: 223-226. 2005; see also Willdenowia
36: 707. 2006). It is not repeated here.
The Notulae provide on one hand the opportunity to validate new scientific names and com-
binations that are required under the recommended taxonomic classification but do not yet exist.
On the other hand, they permit to document distributional data that have not yet been published
in print – both new records and the correction of old erroneous ones. The author of each entry is
either named at its end or, in the case of uncommented new combinations, is acknowledged as
the author of the combination.
Willdenowia 37 – 2007 139

The following have contributed entries to the present instalment: K. P. Buttler, E. Di Gristina,
A. Dobignard, G. Gottschlich, W. Greuter, M. Kääntönen, V. Matevski, M. Niketi6,B. Norden-
stam, E. J. Nyárády†, H. Øllgaard, J. Räsänen, F. M. Raimondo, E. Reinikka, I. Ricci†, P. D. Sell,
A. N. Sennikov, G. Wagenitz, and K. H. Zahn†. Relevant specimen data were kindly supplied by
G. Gottschlich, D. Jeanmonod, H. Manitz, S. Pignatti and F. Schuhwerk, to whom we are grateful.
We also thank J. McNeill for nomenclatural advice and assistance with bibliographic searches,I.
Goia for providing photographs of CL specimens, and C. Dragulescu who, through H. Heltmann,
provided information on Nyárády material at SIB.
Adenostyles alliariae subsp. orientalis (Boiss.) Greuter, comb. & stat. nov. ≡Adenostyles orien-
talis Boiss., Fl. Orient. 3: 155. 1875.
Adenostyles alpina subsp. nebrodensis (Wagenitz & I. Müll.) Greuter, comb. nov. ≡
Adenostyles hybrida Guss., Fl. Sicul. Syn. 2: 449. 1844 ≡Adenostyles nebrodensis Strobl in
Flora 65: 196. 1882, nom. illeg. ≡Adenostyles glabra subsp. nebrodensis Wagenitz & I. Müll. in
Phyton (Horn) 23: 149. 1983.
The combination Adenostyles hybrida, based on Cacalia tomentosa var. hybrida
Vill., is usually attributed to Candolle (Prodr. 5: 204. 1836), but was not validly
published there, being treated as provisional. By consequence, A. hybrida Guss. is
not a later homonym but a legitimate name, with priority for our taxon when treated
as a species; and A. nebrodensis Strobl, proposed as a substitute for A. hybrida
Guss. [non “(Vill.) DC.”], in turn is illegitimate. At subspecies rank, however, the
epithet nebrodensis takes precedence. W. Greuter
Andryala pinnatifida subsp. antonii (Maire) Dobignard, comb. nov. ≡Andryala antonii Maire in
Mém. Soc. Sci. Nat. Maroc 15: 47. 1926 ≡Andryala canariensis subsp. antonii (Maire) Maire in
Jahandiez & Maire, Cat. Pl. Maroc: 840. 1934.
Centaurea arenaria Willd.
–Sr: All published records for Centaurea arenaria for Serbia (e.g. by Hayek in Repert.
Spec. Nov. Regni Veg. Beih. 30(2): 771. 1931) refer in fact to C. borysthenica
Gruner. M. Niketi6
Centaurea aspera subsp. gentilii (Braun-Blanq. & Maire) Dobignard, comb. & stat. nov. ≡Cen-
taurea gentilii Braun-Blanq. & Maire in Bull. Soc. Hist. Nat. Afrique N. 13: 18. 1922.
+Ca(C): Centaurea aspera is known to occur on Gran Canaria (Hansen & Sunding in
Sommerfeltia 17: 42. 1993). A Canarian specimen collected by Kunkel (G) belongs
to subsp. gentilii, originally described from Morocco. A. Dobignard
Centaurea sicula L., Sp. Pl.: 918. 1753 [= Centaurea nicaeensis All., Fl. Pedem. 1: 162. 1785]. –
Lectotype (designated here): illustration in Morison, Hist. Pl. Univ.: ser. 7, t. 28, f. 26. 1699.
Epitype (designated here): Italy, Sicily, “In campis et collibus reg. inferioris, Palermo”, 6.1899,
Ross, Herb. Sicul. No. 254 (GOET).
The name Centaurea sicula L. has long been considered obscure and controversial.
However, matters were originally rather simple. In the Linnaean protologue of
1753, the validating diagnosis is taken unchanged from Sauvages (Meth. Fol.: 290.
1751), who in turn quotes “Jacea Sicula, Cichorii folio, fl. luteo, capite spinoso” of
Boccone (Icon. Descr. Rar. Pl. Sicil.: 14. 1674). That very same phrase is quoted by
Linnaeus in synonymy, not from its primary but from two secondary sources
(Magnol, Bot. Monsp.: 138. 1676, and Morison, Pl. Hist. Univ. 3: 144, ser. 7, t. 28,
f. 26. 1699). The Morison illustration is a (rather poor) copy of the figure in
Boccone (l.c.: 15, t. 8, f. DII). Everything in the protologue points to Boccone as the
principal if not only basis of the original C. sicula. The two specimens in Linnaeus’s
herbarium (No. 1030.63 and .64, LINN), even though technically original material,
140 Greuter & Raab-Straube: Euro+Med Notulae, 3

are not clearly connected with the protologue. The former is a plant grown in
Linnaeus’s garden in Uppsala (a short-spined form of C. nicaeensis All., or perhaps
a hybrid involving that species). The latter (apparently a cultivated form of C. sols-
titialis L.) was originally part of the Clifford herbarium, and as Linnaeus’s own
Hortus Cliffortianus is not cited in the protologue it would hardly be a suitable type.
We therefore select the Morison figure as the lectotype of the species name and un-
derpin that choice by the designation of an epitype belonging to the species illus-
trated, better known today as C. nicaeensis All.
As is often the case, Linnaeus himself (Sp. Pl., ed. 2: 1298. 1763) subsequently
changed the circumscription of the species. He did so by inserting in the phrase
name the single word “decurrentibus”, after “foliis”, and adding a descriptive note
that cannot apply to Centaurea nicaeensis but best fits C. melitensis L. Indeed Go-
dron (in Grenier & Godron, Fl. France 2: 283. 1850), when excluding C. sicula from
the French flora, noted that it had been mentioned from Montpellier by local bota-
nists, Gouan in particular, “sans doute par confusion avec le Centaurea melitensis,
qui y est commun”. This is perhaps the single mention of C. sicula in any French
Flora.
Subsequent usage was split. Lamarck (Encycl. 1: 674. 1785) interpreted Cen-
taurea sicula in the sense of C. nicaeensis, followed by Tenore. Bertoloni (Fl. Ital. 9:
479. 1854) formally treated the C. sicula of these author as a synonym of C. nicae-
ensis, without taking sides with respect to the genuine C. sicula L. On the other hand,
Candolle (Prodr. 6: 593. 1838) chose to recognise a plant collected in central Sicily
(Monte Artesino near Calascibetta), collected by Duby, as corresponding to the gen-
uine C. sicula, cited from the 1763 edition, and was followed by several Italian Flo-
ras (Gussone, Fl. Sicul. Syn. 2: 516. 1844; Cesati & al., Comp. Fl. Ital: 496. 1878;
Lojacono, Fl. Sicul. 2(1): 146. 1903). Duby’s plant is hardly even a variant of C.
solstitialis subsp. schouwii (DC.) Gugler, which explains how Nyman (Consp. Fl.
Eur.: 430. 1779) and Arcangeli (Comp. Fl. Ital: 395. 1882) came to use the combina-
tion C. sicula subsp. schouwii (DC.) Nyman while treating C. solstitialis and C.
nicaeensis as specifically distinct.
Since 1903 (Lojacono) the name Centaurea sicula has not to our knowledge
been used in any sense except as a synonym. Mainly for this reason, and also be-
cause the name C. nicaeensis is blatantly inappropriate for a plant whose ephemeral
alien appearance near Nizza is long forgotten (few of the major French Floras as
much as mention the name), we did not follow up on our original thought of trying
to save it. We share the feelings of Briquet (Monogr. Centaurées Alpes Marit.: 176.
1902) who, still operating under the old Candollean Laws, rejected the name C.
nicaeensis as inappropriate and adopted the younger synonym C. fuscata Desf. (Fl.
Atlant. 2: 302. 1799) in its stead. G. Wagenitz & W. Greuter
Centaurea sicula subsp. kroumirensis (Coss.) Dobignard, comb. & stat. nov. ≡Centaurea krou-
mirensis Coss in Compt. Rend. Hebd. Séances Acad. Sci. 98: 468. 1884 ≡Centaurea nicaeensis
var. kroumirensis Batt. in Battandier & Trabut, Fl. Algérie, Dicot.: 500. 1889.
Centaurea wettsteinii Degen & Dörfl.
–Mk, +Gr:Degen & Dörfler (in Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 64: 726.
1897) described this species from “Severno”, which they placed in central Macedo-
nia. No such place can be found on any map available to us, but later sources
(Dostál in Tutin & al., Fl. Europ. 4: 270. 1976) unhesitatingly consider the species
as endemic to “S. Jugoslavia: Makedonija”. It also appears on a list of threatened
plant species of the FYR Makedonija, in an electronic document “Country study for
biodiversity of the Republic of Macedonia (first national report)” (http://www.
biodiv.org/doc/world/mk/mk-nr-01-en.pdf). However, as V. Matevski (Skopje) has
Willdenowia 37 – 2007 141

kindly confirmed, he and his fellow botanists have been unable to find the species
anywhere in their country. The only subsequent collection of Centaurea wettsteinii
appears to be the following (identification by G. Wagenitz): Greece, W. Macedonia,
eparchy of Almopia, “in latere boreo-occidentali montis Pinovo (Kozuf), alt. 1500
m. In fissuris et scansilibus rupium calcarearum praeruptarum umbrosarum”
20.8.1976, Greuter 14647. This find, seemingly the first Greek record of the spe-
cies, made it advisable to reconsider the question of the correct location of Dörfler’s
locus classicus.
In their botanical account of Dörfler’s 1893 expedition to Albania and Macedo-
nia, Degen & Dörfler (l.c.: 702) give a succinct relation on Dörfler’s itinerary.
“Allchar” [appearing on old maps as Alchar, south of Rozden and immediately
north of the present Greek border] served as Dörfler’s basis. From there he twice
visited Mt “Kossuf” [Kozuf, or in Greek, Pínovo] to the east, close to the village
“Zborsko” [which I have earlier (Greuter in Candollea 29: 138. 1974) tentatively
identified with the now ruined Greek village Pefkotó]. The first of these tours was
extended to the “plain of Severno”. This neatly places the mysterious village Se-
verno. There is only one plain in the area, far and wide: that of Ardéa; and halfway
between Pefkotó (Zborsko) and Ardéa lies the village Vorinó (Boreinón). It cannot
possibly be a coincidence that the Slavic word severno (F,&,D>@) and Greek
voreinón ($@Dg4<`<) have exactly the same meaning: northern. The conclusion is
that Centaurea wettsteinii has not so far been collected in the FYR Makedonija
(where it might nevertheless exist), but is a local endemic of the southwestern slo-
pes of Mt Pínovo in NW Greek Macedonia. W. Greuter
Cichorium intybus subsp. spicatum [I. Ricci in Ann. Bot. (Roma) 28: 219. 1966, sine typo, ex] I.
Ricci, subsp. nov. – Holotype: Italy, Latium, “locus classicus Ponte Galeria (Roma)”, s.d., herb.
Ricci, sheet No. 3 (RO; isotypes: sheets No. 1-2 & 4-9, RO). – Latin description by I. Ricci in
Ann. Bot. (Roma) 28: 219. 1966. Thanks are due to Sandro Pignatti, Roma, who traced Ricci’s
type gathering and numbered its sheets.
Crepis zacintha (L.) Loisel.
–Ge: The report of this taxon for Germany according to Sell (in Tutin & al., Fl. Europ. 4:
355. 1976) is apparently an error. No other published record for Germany exists, not
even as a casual alien. K. P. Buttler
Dittrichia viscosa (L.) Greuter
+Mk: Negotino: Krivolak - Solena Reka, 41°34'19''N, 22°05'47''E, 165 m, leg. et det. V.
Matevski, A. Carni & M. Kostadinovski (SKO). – Until now, this species was only
recorded for Macedonia by Hayek (in Repert. Spec. Nov. Regni Veg. Beih. 30(2):
607. 1931). As it is well known that Hayek used the geographic name of Macedonia
for a much wider area than today’s state borders of the Republic of Macedonia, and
as there are no other data in the floristic literature before 1931 that refer to localities
within the present borders of the Republic of Macedonia, this is the first confirmed
record of this species for this area. V. Matevski
Erigeron daveauanus (Sennen) Greuter, comb. nov. ≡Conyza daveauana Sennen in Bol. Soc.
Aragonesa Ci. Nat. 11: 194. 1912.
Filago carpetana subsp. maroccana (Braun-Blanq. & Maire) Dobignard, comb. & stat. nov. ≡
Evax pygmaea var. maroccana Braun-Blanq. & Maire in Bull. Soc. Hist. Nat. Afrique N. 16: 35.
1925.
Filago mauritanica (Pomel) Dobignard, comb. nov. ≡Pseudevax mauritanica Pomel in Bull.
Soc. Bot. France 35: 335. 1888 ≡Evax mauritanica (Pomel) Batt. in Battandier & Trabut, Fl.
Algérie, Dicot.: 438. 1889.
142 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium L. (excluding Pilosella Vaill.)
The treatment of Hieracium for the purposes of Euro+Med poses problems that are
almost impossible to resolve. Taxonomy in itself is a nightmare, and achieving a
synthesis on a Euro-Mediterranean scale is further hampered by the existence of
competing, irreconcilable schools of thought regarding the basic principles of clas-
sification. There are two main ideologies: One is following the Zahn tradition, ac-
cording to which the (axiomatic) main species and (postulated) hybridogenous
intermediate species derived from them are subdivided into often numerous subspe-
cies corresponding to their (usually apomictic and “true-breeding”) variants; and
the “Nordic school”, by which each distinguishable apomictic strain is treated as a
species, and only a small number of sections are recognised between genus and spe-
cies. The latter approach is perhaps more defensible from a phylogenetic point of
view, but has the major drawback of being inapplicable, according to the present
state of knowledge, in large areas where the major diversity of morphotypes is en-
countered . Yet it is unrealistic to expect that any of the Nordic hawkweed special-
ists might be convinced to revert to a classification based on Zahn’s principles. The
“Flora europaea” treatment (Sell & West in Tutin & al., Fl. Eur. 4: 358-410. 1976)
has attempted a synthesis by grouping segregate species of the “Nordic” type into
species groups more or less equivalent to Zahn’s species concept; but it only covers
a small proportion of the taxa and names involved and has not been followed in sub-
sequent publications by Nordic authors.
Applying one or the other of the competing schemes (or their variants) on a
Euro-Mediterranean scale would be utterly disruptive nomenclaturally – yet useless
from the point of view of specialists of the other creed(s). Rather than abandoning
any thought of coherence and consistency and just list uncritically what is found in
regional literature – an exercise of little merit and scant practical value – I have de-
vised the following approach (which even if it eventually may satisfy no one should
at least not repel specialists from using and trying to improve on it). For those hawk-
weeds found in N and E Europe (notably Britain, Scandinavia, the Baltic countries
and European Russia), the nordic system as modified by “Flora europaea” is being
followed (even though assigning the “segregate species” to their appropriate species
group has sometimes had to be made on a best-guess basis). For the remaining terri-
tories (principally W and Central Europe, the Balkans, Africa and SW Asia includ-
ing Caucasia) I adopted Zahn’s approach. The Carpathian range, where floristic
affinities are mainly to the south and west but taxonomic tradition has espoused the
northern style, was assimilated to one or the other domain depending on the main
affinity of the group concerned. Whenever known, the correct species names for
taxa treated as subspecies, or subspecies names for binominally named taxa, have
been included in synonymy.
Even though this approach is less disruptive nomenclaturally than a unitary
view would have been, it still requires an appalling number of new combinations – a
fact of which I am thoroughly unhappy. The main reason is that, under Zahn’s clas-
sification, each change of name of the species entails several or many recom-
binations at subspecies rank. Zahn is well known for having applied the rules of
botanical nomenclature in his own idiosyncratic way. Mending his treatments to
make hawkweed nomenclature conform with the international rules has been under-
taken by many, but only for individual taxa or groups and on a geographically
patchy basis. The following series of validations (to which several others have ac-
cepted to contribute) cannot be expected to be complete or flawless, but is the mini-
mum required to implement the treatment that is being proposed.
My basic choice has been influenced by inspiring and helpful discussions with
many specialists, none of whom fully endorses it but who may hopefully feel able to
Willdenowia 37 – 2007 143

live with it. While stressing that the responsibility (including the blame for any
faults and omissions) is mine alone, I want to thank most particularly G. Gott-
schlich, G. Mateo Sanz, A. Sennikov and T. Tyler for their friendly attitude and co-
operation. W. Greuter
Hieracium adlerzii F. Hanb. in J. Bot. 32: 229. 1894. – Lectotype (designated here): Dahlstedt,
Hieracia Exsiccata, fasc. 2: No. 85 (BM).
QBr: The epithet adlerzii, ascribed to Almquist, has been used repeatedly in Dahlstedt’s
hawkweeds exsiccata, starting in 1889 (Hierac. Exsicc., fasc. 2: No. 85, as Hiera-
cium murorum subsp. adlerzii), next in 1892 (Herb. Hierac. Scand., Centuria 2: No.
93, as H. adlerzii), etc., but in no case was a description provided. Validation of the
name H. adlerzii is usually credited to Dahlstedt (in Bih. Kongl. Svenska Vetensk.-
Akad. Handl. 28(3, 7): 69. 1902), but was effected earlier by Hanbury (l.c.).
Hanbury provides a short English diagnosis to set off “H. adlerzii Almq.”
against “H. vulgatum”, referring specifically to a plant “found by Mr. J. E. Griffith
on the banks of the Menai Straits, in July, 1891”. This at first sight is the single cited
(syntype) material. If it were the obligate lectotype there would be a problem. It is
by no means certain that the British and Scandinavian plants are the same taxon. In-
deed Sell (in Sell & Murrell, Fl. Great Britain & Ireland 4: 356. 2006), referring to
British material, treats H. adlerzii as a plain synonym of H. triviale (Norrl.) Norrl. It
is therefore doubtful at best whether the Scandinavian taxon known to Swedish bot-
anists as H. adlerzii occurs in Britain.
To maintain traditional usage of the name H. adlerzii it is desirable to typify it
by a Swedish specimen. Fortunately this is possible. In the August 1894 issue of the
Journal of Botany in which his diagnosis appears, Hanbury refers to “A tentative list
of British Hieracia”, distributed with the July issue of the same journal. The rele-
vant content of that list must thus be considered to form part of the protologue. For
“H. Adlerzii Almq.” the following source specimen, a second syntype, is cited for
the name: “Dahlst. Hierac. Exsicc., Fas. ii., 85. 1889”, which I here designate as
lectotype. W. Greuter
Hieracium amplexicaule subsp. attracticaule (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hie-
racium attracticaule Arv.-Touv., Hier. Gall. Hisp. Cat.: 124. 1913.
Hieracium anglorum [Pugsley in J. Bot. 79: 197. 1942, sine diagn. lat., ex] P. D. Sell, sp. nov. –
Holotype: “Hieracium scanicum, rocks by the Wye, Llangoed, Brecon”, 26.6.1907, Ley (CGE). –
Description: Herba perennis phyllopoda, caudice ramoso. Caulis 25-80 cm altus, pallide
luteoviridis, ad basin interdum purpurascens, magis minusve robustus, striatus, in parte inferiore
pilis simplicibus eglandulosis numerosis longis sinuosis pallidis, sursum paucioribus, in parte
superiore pilis stellatis densis pilisque glanduliferis paucis brevibus obscuris vestitus. Folia in
pagina superiore aliquantum pallide viridia, in inferiore pallidiora, raro rubritincta, plerumque
subflaccida; folia basalia plerumque pauca, lamina 3-10 × 1.5-4.5 cm metiens, elliptica vel ovata
vel oblongo-elliptica, raro subrotunda, ad apicem rotundata, obtusa vel acuta, dentata vel profunde
inciso-dentata, dentibus saepe mammiformibus (interdum multo angustioribus et acutioribus), ad
basin abrupte contracta vel cuneata, petiolis ad 5 cm longis, pilis simplicibus eglandulosis nume-
rosis longis pallidis sinuosis vestitis; folia caulina 3-5(-9), infera elliptico-oblonga vel elliptica vel
ovata, ad apicem acuta, dentata (saepe profunde dentata), dentibus longis angustis inaequalibus, ad
basin abrupte contracta, plerumque petiolata, supera ovata vel lanceolata, ad apicem acuta,
plerumque profunde laciniato-dentata, dentibus longis acutis, sessilia; omnia in paginis ambabus
inque marginibus pilis simplicibus eglandulosis brevibus vel longis pallidis vestita. Synflorescentia
capitulis 10-20(-40), paniculato-corymbosa; pedunculi graciles, erecti vel patentes, pilis stellatis
densis, pilis glanduliferis numerosis brevibus obscuris pilisque simplicibus eglandulosis numerosis
brevibus vel mediocribus pallidis sed ad basin obscuris vestiti. Capitula 25-30 mm diametro, ad ba-
144 Greuter & Raab-Straube: Euro+Med Notulae, 3

sin rotundata. Involucri squamae ante anthesin incumbentes, 4-10 × 1-1.2 mm metientes, nigri-
virides, interiores margine pallidiore, omnes lineari-lanceolatae, ad apicem acutum gradatim an-
gustatae, pilis glanduliferis numerosis brevibus vel mediocribus (0.4-1 mm longis) tenuibus nigris
et aliquando pilis simplicibus eglandulosis solitariis vestitae, sine pilis stellatis. Ligulae flavae, ad
apicem glabrae. Styli obscuriusculi. Receptaculi alveoli margine breviter subulato. Cypselae 2.5-3
mm longae, rubrinigrae. – P. D. Sell (Latin translation by Philip Oswald).
Pugsley’s alleged basionym for Hieracium anglorum, H. scanicum var. anglorum
Ley (in J. Bot. 47: 49. 1909), was proposed as a provisional name and thus was not
validly published, in spite of the presence of descriptive matter (in English). Thanks
are due to J. McNeill for examining and discussing the case, and for unsuccessfully
searching the literature of the time for another possible place of valid publication.
W. Greuter
Hieracium apricorum subsp. pseudolonganum (Zahn) Greuter, comb. nov. ≡Hieracium aus-
triacum subsp. pseudolonganum Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(2): 38.
1930.
Hieracium apricorum subsp. sublaevigatum (Beck) Greuter, comb. nov. ≡Hieracium dollineri
f. sublaevigatum Beck, Fl. Nieder-Österr.: 1294. 1893 ≡Hieracium austriacum subsp. sublaevi-
gatum (Beck) Zahn in Engler, Pflanzenr. 75: 58. 1921.
Hieracium armerioides subsp. touvetii (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium cir-
ritum subsp. touvetii Nägeli & Peter, Hierac. Mitt.-Eur. 2: 270. 1889.
Hieracium balbisianum subsp. abbatiscellanum (Zahn) Greuter, comb. nov. ≡Hieracium ker-
neri subsp. abbatiscellanum Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1838. 1901.
Hieracium balbisianum subsp. kerneri (Zahn) Greuter, comb. & stat. nov. ≡Hieracium kerneri
Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1837. 1901 ≡Hieracium kernerianum Zahn in
Dalla Torre & Sarnth., Fl. Tirol 6(3): 884. 1912, nom. illeg.
Hieracium balbisianum subsp. kerneriforme (Zahn) Greuter, comb. nov. ≡Hieracium kerneri
subsp. kerneriforme Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 311. 1905 ≡Hieracium
kerneriforme (Zahn) Prain, Index Kew., Suppl. 4: 114. 1913.
Hieracium balbisianum subsp. kuemmerlei (Degen & Zahn) Greuter, comb. nov. ≡Hieracium
kerneri subsp. kuemmerlei Degen & Zahn in Magyar Bot. Lapok 25: 367. 1927.
Hieracium bernardii subsp. gallurense (Arrigoni) Greuter, comb. & stat. nov. ≡Hieracium
gallurense Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 227. 1985.
Hieracium bernardii subsp. templare (Arrigoni) Greuter, comb. & stat. nov. ≡Hieracium
templare Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 253. 1985.
Hieracium bicolor subsp. pogonatoides (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hieracium
pogonatoides Arv.-Touv. & Gaut., Hieracioth. 4: [in schedis] Gall. No. 196-197. 1898 ≡Hiera-
cium bourgaei subsp. pogonatoides (Arv.-Touv. & Gaut.) O. Bolòs & Vigo, Fl. Països Catal. 3:
1092. 1996.
Hieracium bifidum subsp. hemilonchodes [Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl.
12(2): 666. 1935, descr. germ.] Zahn, subsp. nov. – Holotype: Liechtenstein, “Alpe Sücca”,
11.7.1929, Schack (B 100216195; isotype: B 100216196). – Description: “Folia, (ovato-)elliptica
vel oblongo-lanceolata, basi subrotundata vel ± angustata, margine denticulata vel ± breviter sed
robuste vel (in inferioribus) angustius et profundius, rarius grosse dentata, sensim descrescentia;
caulina 1-2(-3), oblongo-lanceolata vel (supremum) reductum, ± robuste vel longe et acute dentata
vel serrata, superiora saepe nonnisi denticulata. Rami 1-3, acladium 2-4 cm longum, pedunculi mo-
dice vel paene glandulosi, aliquanto pilosi; capitula 2-5(-9). Involucrum 9-10.5 mm longum,
Willdenowia 37 – 2007 145

atrum, vix glandulosum, ± pilosum, aliquanto floccosum; squamae latiusculae, obtusiusculae vel
acutae, margine ± floccosae. Stylus initio luteo-brunneum.” – H. Zahn.
Hieracium bifidum subsp. kittaniae (Vladimirov) Greuter comb. & stat. nov. ≡Hieracium kit-
taniae Vladimirov in Bot. J. Linn. Soc. 143: 214. 2003.
Hieracium bifidum subsp. obscurisquamum (Zahn) Greuter, comb. nov. ≡Hieracium psam-
mogenes var. obscurisquamum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Natur-
wiss. 40: 440. 1906 ≡Hieracium psammogenes subsp. obscurisquamum (Zahn) Touton in Mitt.
Bayer. Bot. Ges. 3: 324. 1916 ≡Hieracium bifidum subsp. subcaesiiceps Zahn in Engler, Pflan-
zenr. 76: 433. 1921, nom. illeg.
Hieracium bifidum subsp. pseudopraecox (Zahn) Greuter, comb. nov. ≡Hieracium caesium
subsp. pseudopraecox Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1793. 1901 ≡Hieracium
pseudopraecox (Zahn) Prain, Index Kew., Suppl. 5: 132. 1921.
Hieracium bifidum subsp. wallrothianum (Bornm. & Zahn) Greuter, comb. & stat. nov. ≡Hie-
racium wallrothianum Bornm. & Zahn in Mitth. Thüring. Bot. Vereins 39: 42. 1930.
Hieracium bosniacum subsp. banjanum (K. Malý & Zahn) Greuter, comb. nov. ≡Hieracium
dolopicum subsp. banjanum K. Malý & Zahn in Glasn. Zemaljsk. Muz. Bosni Hercegovini 37:
56. 1925.
Hieracium bosniacum subsp. dolopicum (Freyn & Sint.) Greuter, comb. & stat. nov. ≡Hie-
racium dolopicum Freyn & Sint. in Bull. Herb. Boissier 5: 785. 1897.
Hieracium bosniacum subsp. pannosifolium (Zahn) Greuter, comb. nov. ≡Hieracium dolopi-
cum subsp. pannosifolium Zahn in Engler, Pflanzenr. 77: 577. 1921.
Hieracium bosniacum subsp. pelliculatum (Zahn) Greuter, comb. nov. ≡Hieracium pellicu-
latum Zahn in Magyar Bot. Lapok 6: 219. 1907 ≡Hieracium dolopicum subsp. pelliculatum
(Zahn) Zahn in Engler, Pflanzenr. 76: 576. 1921.
Hieracium brevifolium subsp. burgeffianum Greuter, subsp. nov. [Hieracium latifolium subsp.
burgeffianum Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3): 524. 1938, descr. germ.].
– Holotype: F.Y.R. Makedonija, “Nikolic a. Dojran See”, 1917, Burgeff 1035 (M). – Description:
“Hieracio noëano simile, sed folia valde firma, etiam supra breviter setosa, interdum breviden-
tata, sensim descrescentia, oblongo-ovata vel late ovata, breviter acutata. Synflorescentia non
deliminata; rami foliis parvis brevibus cordato-ovatis aliquanto floccosis obsiti; acladium (ad
25 mm longum) et pedunculi grisei breviter pilosi et glandulosi. Involucrum ad 12 mm longum,
latissimum, pilosum et glandulosum; squamae atrovirides viridimarginatae, plurimae latius-
culae, obtusiusculae ad subacutae; exteriores angustae, laxae, in bracteas parvas virescentes tran-
seuntes. Stylus ater. Achaenia atrobrunnea.” – H. Zahn.
Hieracium brevifolium subsp. hundsdorferianum Greuter, subsp. nov. [Hieracium latifolium
subsp. hundsdorferianum Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3): 518. 1938,
descr. germ.]. – Holotype: Slovenija, am Cavn bei Görz, Harz (M). – Description: “Planta 60-80
cm alta, basin versus gradatim hirtior, corymbosum; rami 5-10 cm longi; pedunculi elongati,
graciles, floccosi tantum; capitula 10-20. Involucrum magnum, minute glandulosum, pilis desti-
tutum, parcifloccosum; squamae latae; bracteae plures. Stylus luteus. Folia 40-60 densa, late lan-
ceolata, summa tantum diminuta, margine (piis saepe truncatis) et dorso tantum breviter pilosa
(pilis nodulis insidentibus) et floccosa; inferiora 10 cm longa et 15-20 mm lata; superiora minus
pilosa sed utrinque floccosa; omnia dentibus 1-2(-3) dissitis longis angustis acutissimis serrata.”
– H. Zahn.
Hieracium brevifolium subsp. noëanum (Zahn) Greuter, comb. nov. ≡Hieracium noëanum
Zahn in Engler, Pflanzenr. 79: 1006. 1922 ≡Hieracium latifolium subsp. noëanum (Zahn) Zahn
in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3): 524. 1938.
146 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium bucuranum [Nyár. in Sbvulescu, Fl. Rep. Pop. Române 10: 735. 1965, sine design.
typi, ex] Nyár., sp. nov. – Description: Nyárády in Sbvulescu, Fl. Rep. Pop. Române 10: 735.
1965. – Holotype: Romania, “Transsilvania, distr. Hunedoara, Mtibus Retezat, ad rivum Bucura
supra vallem Lbpuznicul mare”, c. 1600-1700 m, 27.8.1930, Nyárády (SIB 118504; isotypes: CL
430261-430262, 430265-430267, 430303, 430474-430475, 430477).
Hieracium calcareum subsp. baldense (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
illyricum subsp. baldense Nägeli & Peter, Hierac. Mitt.-Eur. 2: 56. 1886 ≡Hieracium baldense
(Nägeli & Peter) Prain, Index Kew., Suppl. 4: 109. 1913.
Hieracium calcareum subsp. baldensiforme (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
illyricum subsp. baldensiforme Nägeli & Peter, Hierac. Mitt.-Eur. 2: 55. 1886 ≡Hieracium bal-
densiforme (Nägeli & Peter) Prain, Index Kew., Suppl. 4: 109. 1913.
Hieracium calcareum subsp. carnicum (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium illy-
ricum subsp. carnicum Nägeli & Peter, Hierac. Mitt.-Eur. 2: 54. 1886.
Hieracium calcareum subsp. crinopodum (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
illyricum subsp. crinopodum Nägeli & Peter, Hierac. Mitt.-Eur. 2: 55. 1886 ≡Hieracium crino-
podum (Nägeli & Peter) Prain, Index Kew., Suppl. 5: 127. 1921.
Hieracium calcareum subsp. holleri (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium illy-
ricum subsp. holleri Nägeli & Peter, Hierac. Mitt.-Eur. 2: 57. 1886 ≡Hieracium holleri (Nägeli
& Peter) Prain, Index Kew., Suppl. 4: 113. 1913.
Hieracium calcareum subsp. illyricum (Fr.) Greuter, comb. & stat. nov. ≡Hieracium illyricum
Fr. in Uppsala Univ. Årsskr. 1862: 71. 1862.
Hieracium calcareum subsp. papperitzii (Zahn) Greuter, comb. nov. ≡Hieracium illyricum
subsp. papperitzii Zahn in Engler, Pflanzenr. 76: 388. 1921.
Hieracium calcareum subsp. trilacense (Dörfler) Greuter, comb. nov. ≡Hieracium illyricum
subsp. trilacense Dörfler in Jahreskat. Wiener Bot. Tauschvereins 1900: 128. 1899 ≡Hieracium
trilacense (Dörfler) Prain, Index Kew., Suppl. 5: 134. 1921.
Hieracium calcogeton (Zahn) Greuter, comb. & stat. nov. ≡Hieracium vulgatum subsp. calco-
geton Zahn in Engler, Pflanzenr. 75: 181. 1921 [incl. Hieracium biharianum auct., nom. inval.].
Hieracium calophyllum subsp. hercegovinicum (Freyn & Vandas) Greuter, comb. nov. ≡Hie-
racium marmoreum subsp. hercegovinicum Freyn & Vandas in Bull. Herb. Boissier 3: 647, 652,
663. 1895 ≡Hieracium hercegovinicum (Freyn & Vandas) Szel}g in Feddes Repert. 113: 506.
2002 ≡Hieracium calophyllum subsp. hercegovinae Zahn in Engler, Pflanzenr. 76: 588. 1921,
nom. illeg.
Hieracium camkorijense subsp. sarykamyschense (Üksip) Greuter, comb. & stat. nov. ≡Hiera-
cium sarykamyschense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19:
487. 1959.
Hieracium carinthiostiriacum [J. Vetter & Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl.
12(3): 698. 1938, descr. germ.] J. Vetter & Zahn, sp. nov. – Holotype: Austria, “Steirisch-kärnt-
nerisches Grenzgebiet, Wiesen auf der Turracher Höhe”, 27.7.1931, Vetter (W-1950-7599). – De-
scription: “Hieracio djimilensi habitu simile sed indumentum breve, ad marginem foliorum ±
copiosum, rigidiusculum, in facie superiore foliorum, infimorum exceptorum, nullum. Caules
25-40 cm alti, graciles, ± colorati, breviter rigidiuscule pilosi, 5-12-cephali; acladium 3-10 mm
longum; rami 3-5, admodum dissiti, 1-3(-4)-cephali, copiose floccosi, parce vel modice pilosi
(pilis 1-2 mm longis, basi atris) et parce minute glandulosi; pedunculi breves, grisei, modice vel
admodum parvoglandulosi, parce pilosi. Involucrum cylindrico-campanulatum dein carssum, mo-
dice (partim minute) glandulosum et pilis e basi atra crassa dilute apiculatis submodice obsitum;
Willdenowia 37 – 2007 147

squamae latae, obtusae, atrae, praesertim marginem versus (in extimis brevioribus densius) floc-
cosae, interiores acutiusculae et sordide viridimarginatae. Flores conspicui, saturate lutei, parce et
brevissime ciliati; stylus ater. Folia basalia desunt; caulina 7-12, paulatim diminuta, late lan-
ceolata, acuta vel longe acutata, nonnisi (interdum valde) denticulata, subfirma, dilute flavido- vel
olivaceo-virentia; infima (10-15 cm × 15-20 mm metientia) sensim in petiolum brevem vel longum
interdum latissimum angustata, sequentia parum angustata saepe panduriformia, vel haud an-
gustata et basi lata vel subcordata sessilia et semiamplexicaulia; summa e basi lata amplectente
sensim acuminata, in bracteas angustas, virides vel ± atratas transeuntia.” – H. Zahn.
Hieracium caucasiense subsp. acutangulum (Koslovsky & Zahn) Greuter, comb. nov. ≡Hiera-
cium biebersteinii subsp. acutangulum Koslovsky & Zahn in Vestn. Tiflissk. Bot. Sada 29: 20.
1913 ≡Hieracium acutangulum (Koslovsky & Zahn) Üksip in Komarov, Fl. SSSR 30: 48. 1960.
Hieracium caucasiense subsp. biebersteinii (Litv. & Zahn) Greuter, comb. & stat. nov. ≡Hie-
racium biebersteinii Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 329. 1907, nom. illeg. ≡
Hieracium muricellum subsp. biebersteinii Zahn in Engler, Pflanzenr. 79: 1063. 1922 ≡Hieracium
megalosvaneticum Sennikov in Willdenowia 35: 232. 2005.
Hieracium caucasiense subsp. gandogeri (Zahn) Greuter, comb. nov. ≡Hieracium pseudospar-
sum Zahn in Vandas, Reliq. Forman.: 365. 1909 ≡Hieracium muricellum subsp. gandogeri Zahn
in Engler, Pflanzenr. 79: 1064. 1922 ≡Hieracium gandogeri (Zahn) Zahn in Ascherson &
Graebner, Syn. Mitteleur. Fl. 12(3): 702. 1938.
Hieracium caucasiense subsp. hypopogon (Litv. & Zahn) Greuter, comb. & stat. nov. ≡
Hieracium biebersteinii subsp. hypopogon Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4:
329. 1907 ≡Hieracium hypopogon (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 50. 1960.
Hieracium caucasiense subsp. macrogonum (Zahn) Greuter, comb. nov. ≡Hieracium muricel-
lum subsp. macrogonum Zahn in Engler, Pflanzenr. 79: 1064. 1922 ≡Hieracium macrogonum
(Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33: 431. 1975.
Hieracium cavanillesianum subsp. aldeanum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium
aldeanum Arv.-Touv., Hier. Gall. Hisp. Cat.: 121. 1913 ≡Hieracium leptocladum subsp.aldea-
num (Arv.-Touv.) Zahn in Engler, Pflanzenr. 82: 1577. 1923 ≡Hieracium rupicola subsp.aldea
-
num (Arv.-Touv.) O. Bolòs & Vigo, Fl. Països Catal. 3: 1118. 1996.
Hieracium cerinthoides subsp. ernesti (Maire) Greuter, comb. nov. ≡Hieracium ernesti Maire
in Bull. Soc. Hist. Nat. Afrique N. 12: 45. 1921 ≡Hieracium ramondii subsp. ernesti (Maire)
Zahn in Engler, Pflanzenr. 82: 1535. 1923.
Hieracium cerinthoides subsp. fourcadei (de Retz) Greuter, comb. & stat. nov. ≡Hieracium
fourcadei de Retz in Bull. Soc. Bot. France 125(3-4): 209. 1978.
Hieracium cernagorae subsp. doljaninum (K. Malý & Zahn) Greuter, comb. nov. ≡Hieracium
pseudotommasinii subsp. doljaninum K. Malý & Zahn in Glasn. Zemaljsk. Muz. Bosni Herce-
govini 37: 57. 1925.
Hieracium cernagorae subsp. krstacense (K. Malý & Zahn) Greuter, comb. nov. ≡Hieracium
pseudotommasinii subsp. krstacense K. Malý & Zahn in Glasn. Zemaljsk. Muz. Bosni Herce-
govini 37: 57. 1925.
Hieracium cernagorae subsp. pseudotommasinii (Rohlena & Zahn) Greuter, comb. & stat.
nov. ≡Hieracium pseudotommasinii Rohlena & Zahn in Repert. Spec. Nov. Regni Veg. 6: 237.
1909.
Hieracium cernagorae subsp. stupposiceps (Rohlena & Zahn) Greuter, comb. nov. ≡Hieracium
pseudotommasinii subsp. stupposiceps Rohlena & Zahn in Repert. Spec. Nov. Regni Veg. 6: 239.
1909.
148 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium cernagorae subsp. vardense (Rohlena & Zahn) Greuter, comb. nov. ≡Hieracium
pseudotommasinii subsp. vardense Rohlena & Zahn in Repert. Spec. Nov. Regni Veg. 6: 238.
1909.
Hieracium chalcidicum subsp. cappadocicum (Freyn) Greuter, comb. nov. ≡Hieracium cappa-
docicum Freyn in Österr. Bot. Z. 41: 55. 1891 ≡Hieracium pilosissimum subsp. cappadocicum
(Freyn) Zahn in Engler, Pflanzenr. 77: 593. 1921.
Hieracium chalcidicum subsp. divaricatum (Fr.) Greuter, comb. & stat. nov. ≡Hieracium pilo-
sissimum Friv. in Flora 19: 436. 1836 [non Schrank 1789] ≡Hieracium divaricatum Fr. in
Uppsala Univ. Årsskr. 1862: 78. 1862.
Hieracium chalcidicum subsp. koenigianum (Zahn) Greuter, comb. nov. ≡Hieracium pilosis-
simum subsp. koenigianum Zahn in Vestn. Tiflissk. Bot. Sada 12: 8. 1908 ≡Hieracium koeni-
gianum (Zahn) Üksip in Komarov, Fl. SSSR 30: 200. 1960.
Hieracium chalcidicum subsp. macropannosum (Rech. f. & Zahn) Greuter, comb. nov. ≡Hie-
racium pilosissimum subsp. macropannosum Rech. f. & Zahn in Bot. Jahrb. Syst. 69: 534. 1939 ≡
Hieracium macropannosum (Rech. f. & Zahn) Greuter in Willdenowia 35: 232. 2005.
Hieracium chalcidicum subsp. paphlagonicum (Freyn & Sint.) Greuter, comb. nov. ≡Hiera-
cium paphlagonicum Freyn & Sint. in Österr. Bot. Z. 44: 260. 1894 ≡Hieracium pilosissimum
subsp. paphlagonicum (Freyn & Sint.) Zahn in Engler, Pflanzenr. 77: 593. 1921.
Hieracium chalcidicum subsp. subpannosum (Paw5.) Greuter, comb. nov. ≡Hieracium pilosis-
simum subsp. subpannosum Paw5. in Acta Soc. Bot. Poloniae 32: 478. 1963 ≡Hieracium sub-
pannosum (Paw5.) Greuter in Willdenowia 35: 234. 2005.
Hieracium chalcidicum subsp. thessalum (Formánek) Greuter, comb. nov. ≡Hieracium thes-
salum Formánek in Verh. Naturf. Vereins Brünn 34: 292. 1896 ≡Hieracium divaricatum subsp.
thessalum (Formánek) Zahn in Vandas, Reliq. Forman.: 359. 1909 ≡Hieracium pilosissimum
subsp. thessalum (Formánek) Zahn in Engler, Pflanzenr. 77: 595. 1921.
Hieracium chlorophyton subsp. huljakianum (Zahn) Greuter, comb. nov. ≡Hieracium marche-
settianum subsp. huljakianum Zahn in Magyar Bot. Lapok 28: 32. 1930.
Hieracium chlorophyton subsp. languidum (Zahn) Greuter, comb. nov. ≡Hieracium marche-
settianum subsp. languidum Zahn, Hierac. Alp. Mar.: 173. 1916.
Hieracium chlorophyton subsp. marchesettianum (Zahn) Greuter, comb. & stat. nov. ≡Hie-
racium marchesettianum Zahn, Hierac. Alp. Mar.: 173. 1916.
Hieracium chlorophyton subsp. margittaianum (Zahn) Greuter, comb. nov. ≡Hieracium mar-
chesettianum subsp. margittaianum Zahn in Engler, Pflanzenr. 79: 997. 1922.
Hieracium chlorophyton subsp. putnokense (Hulják & Zahn) Greuter, comb. nov. ≡Hieracium
marchesettianum subsp. putnokense Hulják & Zahn in Magyar Bot. Lapok 28: 33. 1930.
Hieracium cincinnatum subsp. armeniacum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hie-
racium armeniacum Arv.-Touv., Hier. Gall. Hisp. Cat.: 431. 1913.
Hieracium compositum subsp. cadevallii (Pau) Greuter, comb. & stat. nov. ≡Hieracium cade-
vallii Pau in Mem. Real Acad. Ci. Barcelona, ser. 2, 2: 93. 1897.
Hieracium cordatum subsp. acanthophyllum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hie-
racium acanthophyllum Arv.-Touv. in Bull. Herb. Boissier 5: 718. 1897.
Hieracium cordifolium subsp. periplecum (Arv.-Touv. & Gaut.) Greuter, comb. & stat. nov. ≡
Hieracium periplecum Arv.-Touv. & Gaut. in Bull. Soc. Bot. France 41: 355. 1894.
Willdenowia 37 – 2007 149

Hieracium cordifolium subsp. protocerinthe (Arv.-Touv. & Sennen) Greuter, comb. & stat.
nov. ≡Hieracium protocerinthe Arv.-Touv. & Sennen in Arvet-Touvet, Hier. Gall. Hisp. Cat.:
178. 1913.
Hieracium crocatum subsp. lespinassei (Koslovsky & Zahn) Greuter, comb. nov. ≡Hieracium
conicum subsp. lespinassei Koslovsky & Zahn in Vestn. Tiflissk. Bot. Sada 29: 15. 1913 ≡Hie-
racium aestivum subsp. lespinassei (Koslovsky & Zahn) Zahn in Engler, Pflanzenr. 79: 920.
1922 ≡Hieracium lespinassei (Koslovsky & Zahn) Üksip in Komarov, Fl. SSSR 30: 134. 1960.
Hieracium cryptanthum subsp. adenodontum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium adenodontum Arv.-Touv. & Gaut., Hieracioth. 1: [in schedis] Gall. No. 32-34. 1897 ≡
Hieracium subsericeum subsp. adenodontum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
75: 158. 1921.
Hieracium cryptanthum subsp. venascanum (Arv.-Touv. & Gaut.) Greuter, comb. & stat. nov.
≡Hieracium venascanum Arv.-Touv. & Gaut., Hieracioth. 13: [in schedis] Hisp. No. 208. 1902.
Hieracium dasytrichum subsp. neopolytrichum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡
Hieracium neopolytrichum Arv.-Touv., Hier. Gall. Hisp. Cat.: 63. 1913.
Hieracium dermophyllum subsp. juraniforme (Zahn) Greuter, comb. nov. ≡Hieracium jura-
num subsp. juraniforme Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 332. 1905 ≡Hieracium
juraniforme (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
646. 1906.
Hieracium dermophyllum subsp. juranigenum (Zahn) Greuter, comb. nov. ≡Hieracium jurani-
forme subsp. juranigenum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss.
40: 648. 1906 ≡Hieracium juranigenum (Zahn) Prain, Index Kew., Suppl. 4: 114. 1913.
Hieracium dermophyllum subsp. juranopsis (Zahn) Greuter, comb. nov. ≡Hieracium jurani-
forme subsp. juranopsis Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
647. 1906 ≡Hieraciumjuranopsis (Zahn) Prain, Index Kew., Suppl. 4: 114. 1913.
Hieracium dermophyllum subsp. nufenense (Zahn) Greuter, comb. nov. ≡Hieracium nufenense
Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 333. 1905 ≡Hieracium juraniforme subsp. nufe-
nense (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 648. 1906.
Hieracium diaphanoides subsp. chlorodes (Dahlst.) Greuter, comb. nov. ≡Hieracium anfrac-
tum subsp. chlorodes Dahlst. in Kongl. Svenska Vetensk. Acad. Handl., ser. 2, 26(3): 148. 1894
≡Hieracium chlorodes (Dahlst.) Dahlst., Herb. Hierac. Scand. 9: [in schedis] No. 66. 1895 ≡
Hieracium vulgatum subsp. chlorodes (Dahlst.) Zahn in Engler, Pflanzenr. 76: 382. 1921 ≡Hie-
racium lachenalii subsp. chlorodes (Dahlst.) Zahn in Hegi, Ill. Fl. Mitt.-Eur. 6: 1283. 1929.
Hieracium djimilense subsp. panjutinii (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
panjutinii Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 467. 1959.
Hieracium djimilense subsp. terekianum (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium daci-
cum subsp. terekianum Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 263. 1907 ≡Hieracium
pelagae subsp. terekianum (Litv. & Zahn) Zahn in Engler, Pflanzenr. 79: 1069. 1922 ≡Hiera-
cium terekianum (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 33. 1960.
Hieracium dolichosphaericum (Zahn) Greuter, comb. & stat. nov. ≡Hieracium dolichocepha-
lum Dahlst. in Lindman, Sv. Fanerogamfl.: 610. 1918 [non (Zahn) Prain 1913] ≡Hieracium atra-
tum subsp. dolichosphaericum Zahn in Engler, Pflanzenr. 77: 672. 1921.
Hieracium doronicifolium subsp. salevense (Fr.) Greuter, comb. nov. ≡Hieracium dentatum
subsp. salevense Fr. in Uppsala Univ. Årsskr. 1862: 63. 1862 ≡Hieracium misaucinum subsp.
salevense (Fr.) Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 302. 1905 ≡Hieracium salevense
(Fr.) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 526. 1906.
150 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium elisaeanum subsp. balearicum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hiera-
cium balearicum Arv.-Touv., Hier. Gall. Hisp. Cat.: 157. 1913.
Hieracium elisaeanum subsp. segurae (Mateo) Greuter, comb. & stat. nov. ≡Hieracium se-
gurae Mateo in Monogr. Inst. Piren. Ecol. 4: 254. 1988.
Hieracium eriophorum subsp. lavernellei (Timb.-Lagr.) Greuter, comb. nov. ≡Hieracium la-
vernellei Timb.-Lagr. in Mém. Acad. Sci. Toulouse, ser. 7, 9: 136. 1877 ≡Hieracium prostratum
subsp. lavernellei (Timb.-Lagr.) Zahn in Engler, Pflanzenr. 79: 941. 1922.
Hieracium erucophyllum subsp. prinzii (Zahn) Greuter, comb. nov. ≡Hieracium squalidum
subsp. prinzii Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1835. 1901 ≡Hieracium cottetii subsp.
prinzii (Zahn) Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 310. 1905 ≡Hieracium prinzii
(Zahn) Zahn in Engler, Pflanzenr. 76: 522. 1921.
Hieracium flagelliferum subsp. joratense (Gremli) Greuter, comb. nov. ≡Hieracium boreale
var. joratense Gremli, Excursionsfl. Schweiz, ed. 7: 282. 1893 ≡Hieracium racemosum subsp.
joratense (Gremli) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1926. 1902.
Hieracium flocciferum subsp. briziflorum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium brizi-
florum Arv.-Touv., Hier. Gall. Hisp. Cat.: 143. 1913 ≡Hieracium flocculiferum subsp. brizi-
florum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 156. 1921.
Hieracium flocciferum subsp. subflocciferum (Zahn) Greuter, comb. & stat. nov. ≡Hieracium
flocculiferum subsp. subflocciferum Zahn in Engler, Pflanzenr. 75: 156. 1921 ≡Hieracium brizi-
florum subsp. subflocciferum (Zahn) O. Bolòs & Vigo, Fl. Països Catal. 3: 1082. 1996.
Hieracium fuscocinereum subsp. philanthrax (Dahlst.) [de Retz in Bull. Soc. Bot. France 121:
39. 1974, comb. inval., ex] Greuter, comb. nov. [≡Hieracium [subsp.] philanthrax Stenstr.,
Värmländsk. Archierac.: 25. 1889, nom. inval.] ≡Hieracium philanthrax Dahlst., Herb. Hierac.
Scand. 1: [in schedis] No. 79-84. 1892 ≡Hieracium sagittatum subsp. philanthrax (Dahlst.) Zahn
in Engler, Pflanzenr. 76: 343. 1921.
Hieracium gigantellum subsp. podkumokense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
podkumokense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 467. 1959.
Hieracium glaucinum subsp. badalii (Pau) Greuter, comb. & stat. nov. ≡Hieracium badalii
Pau in Not. Bot. Fl. Españ. 6: 71. 1895.
Hieracium glaucinum subsp. baliophyllum (Hyl.) Greuter, comb. & stat. nov. ≡Hieracium ba-
liophyllum Hyl. in Symb. Bot. Upsal. 7(1): 132. 1943.
Hieracium glaucinum subsp. basilacistum (Rech. f. & Zahn) Greuter, comb. nov. ≡Hieracium
praecox subsp. basilacistum Rech. f. & Zahn in Repert. Spec. Nov. Regni Veg. 38: 350. 1935.
Hieracium glaucinum subsp. crestense (Briq. & Zahn) Greuter, comb. nov. ≡Hieracium prae-
cox subsp. crestense Briq. & Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(2): 276.
1931.
Hieracium glaucinum subsp. fragiliforme (Zahn) [O. Bolòs & Vigo, Fl. Països Catal. 3: 1050.
1996, comb. inval., ex] Greuter, comb. nov. ≡Hieracium praecox subsp. fragiliforme Zahn in
Arch. Bot. Bull. Mens. 2: 197. 1928.
Hieracium glaucinum subsp. horripilum (Zahn) [O. Bolòs & Vigo, Fl. Països Catal. 3: 1049.
1996, comb. inval., ex] Greuter, comb. nov. ≡Hieracium praecox subsp. horripilum Zahn in
Engler, Pflanzenr. 75: 236. 1921.
Hieracium glaucinum subsp. lojkae (Degen & Zahn) Greuter, comb. nov. ≡Hieracium praecox
subsp. lojkae Degen & Zahn in Magyar Bot. Lapok 7: 115. 1908.
Willdenowia 37 – 2007 151

Hieracium glaucinum subsp. petiolare (Jord.) Greuter, comb. nov. ≡Hieracium petiolare Jord.,
Jard. Bot. Grenoble, Cat. Graines 1849: 20. 1849 ≡Hieracium praecox subsp. petiolare (Jord.)
Sudre, Hierac. Centre France: 78. 1902.
Hieracium glaucinum subsp. pljackavicense (Zahn) Greuter, comb. nov. ≡Hieracium marko-
vanum Arv.-Touv. in Annuaire Conserv. Jard. Bot. Genève 10: 109. 1907 ≡Hieracium praecox
subsp. pljackavicense Zahn in Engler, Pflanzenr. 75: 241. 1921.
Hieracium glaucinum subsp. portaeferreae (Degen & Zahn) Greuter, comb. nov. ≡Hieracium
praecox subsp. portaeferreae Degen & Zahn in Engler, Pflanzenr. 76: 241. 1921.
Hieracium glaucinum subsp. prasiophaeum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium prasiophaeum Arv.-Touv. & Gaut. in Bull. Soc. Bot. France 41: 349. 1894 ≡Hieracium
murorum subsp. prasiophaeum (Arv.-Touv. & Gaut.) Zahn in Schinz & Keller, Fl. Schweiz, ed.
3, 2: 421. 1914.
Hieracium glaucinum subsp. veljunense (Zahn) Greuter, comb. nov. ≡Hieracium praecox
subsp. veljunense Zahn in Engler, Pflanzenr. 75: 241. 1921.
Hieracium glaucinum subsp. verlotii (Fr.) [O. Bolòs & Vigo, Fl. Països Catal. 3: 1050. 1996,
comb. inval., ex] Greuter, comb. nov. ≡Hieracium verlotii Fr. in Uppsala Univ. Årsskr. 1862:
91. 1862 ≡Hieracium cinerascens subsp. verlotii (Fr.) Zahn in Neue Denkschr. Allg. Schweiz.
Ges. Gesammten Naturwiss. 40: 473. 1906 ≡Hieracium praecox subsp. verlotii (Fr.) Zahn in
Engler, Pflanzenr. 75: 239..
Hieracium glaucinum subsp. vratnikense (Zahn) Greuter, comb. nov. ≡Hieracium cinerascens
subsp. vratnikense Zahn in Ann. Hist.-Nat. Mus. Natl. Hung. 8: 68. 1910 ≡Hieracium praecox
subsp. vratnikense (Zahn) Zahn in Engler, Pflanzenr. 75: 239. 1921.
Hieracium glaucomorphum subsp. franconicum (Griseb.) Greuter, comb. & stat. nov. ≡Hiera-
cium rupicola var. franconicum Griseb. in Abh. Königl. Ges. Wiss. Göttingen 5: 136. 1853 ≡
Hieracium franconicum (Griseb.) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten
Naturwiss. 40: 407. 1906.
Hieracium glaucophyllum subsp. villadraeum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hie-
racium villadraeum Arv.-Touv., Hier. Gall. Hisp. Cat.: 116. 1913.
Hieracium glaucum subsp. aechmetes (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium den-
tatum subsp. aechmetes Nägeli & Peter, Hierac. Mitt.-Eur. 2: 176. 1886 ≡Hieracium aechmetes
(Nägeli & Peter) Prain, Index Kew., Suppl. 4: 108. 1913.
Hieracium grandiceps P. D. Sell, nom. nov. ≡Hieracium magniceps P.D.Sell&C.Westin
Watsonia 6: 308. 1967 [non Omang 1951].
Hieracium heterogynum subsp. crepidifolium (Nägeli & Peter) Greuter, comb. nov. ≡Hiera-
cium stuppeum subsp. crepidifolium Nägeli & Peter, Hierac. Mitt.-Eur. 2: 48. 1886 ≡Hieracium
stupposum subsp. crepidifolium (Nägeli & Peter) Zahn in Engler, Pflanzenr. 79: 963. 1922.
Hieracium heterogynum subsp. eriopodum (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
stuppeum subsp. eriopodum Nägeli & Peter, Hierac. Mitt.-Eur. 2: 47. 1886 ≡Hieracium stup-
posum subsp. chaetopodum Zahn in Engler, Pflanzenr. 79: 963. 1922, nom. illeg.
Hieracium heterogynum subsp. nivisquamum (Degen & Zahn) Greuter, comb. nov. ≡Hiera-
cium stuppeum subsp. nivisquamum Degen & Zahn in Magyar Bot. Lapok 7: 125. 1908 ≡Hiera-
cium stupposum subsp. nivisquamum (Degen & Zahn) Zahn in Engler, Pflanzenr. 79: 963. 1922.
Hieracium heterogynum subsp. pachychaetium (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
stuppeum subsp. pachychaetium Nägeli & Peter, Hierac. Mitt.-Eur. 2: 47. 1886 ≡Hieracium
stupposum subsp. pachychaetium (Nägeli & Peter) Zahn in Engler, Pflanzenr. 79: 963. 1922.
152 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium heterogynum subsp. substupposum (Rohlena & Zahn) Greuter, comb. nov. ≡Hie-
racium stuppeum subsp. substupposum Rohlena & Zahn in Repert. Spec. Nov. Regni Veg. 6:
235. 1909 ≡Hieracium stupposum subsp. substupposum (Rohlena & Zahn) Zahn in Engler,
Pflanzenr. 79: 963. 1922.
Hieracium heterogynum subsp. ueskuebiense (Zahn) Greuter, comb. nov. ≡Hieracium stup-
peum subsp. ueskuebiense Zahn in Vandas, Reliq. Forman.: 363. 1909 ≡Hieracium stupposum
subsp. ueskuebiense (Zahn) Zahn in Engler, Pflanzenr. 79: 964. 1922.
Hieracium hispanicum subsp. dimorphotrichum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium dimorphotrichum Arv.-Touv. & Gaut., Hieracioth. 6: [in schedis] Hisp. No. 79. 1899
≡Hieracium cordatum subsp. dimorphotrichum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
77: 734. 1921.
Hieracium hispanicum subsp. sacalmianum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium sacalmianum Arv.-Touv. & Gaut., Hieracioth. 1: [in schedis] Hisp. No. 5. 1897 ≡Hiera-
cium glaucophyllum subsp. sacalmianum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 77:
738. 1921.
Hieracium hypochoeroides subsp. adesiforme (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. adesiforme Zahn, Hierac. Alp. Mar.: 153. 1916.
Hieracium hypochoeroides subsp. adesum (Bernoulli & Zahn) Greuter, comb. nov. ≡Hiera-
cium wiesbaurianum subsp. adesum Bernoulli & Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2:
303. 1905 ≡Hieracium adesum (Bernoulli & Zahn) Prain, Index Kew., Suppl. 4: 108. 1913.
Hieracium hypochoeroides subsp. apertorum (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. apertorum Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(2): 354.
1931 ≡Hieracium apertorum (Zahn) Jochen Müll. in Haussknechtia 10: 142. 2004.
Hieracium hypochoeroides subsp. ayliesii (Zahn) Greuter, comb. nov. ≡Hieracium wiesbau-
rianum subsp. ayliesii Zahn in Engler, Pflanzenr. 82: 1541. 1923.
Hieracium hypochoeroides subsp. bridelianum (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. bridelianum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Natur-
wiss. 40: 475. 1906 ≡Hieracium bridelianum (Zahn) Prain, Index Kew., Suppl. 4: 109. 1913.
Hieracium hypochoeroides subsp. crinicaesium (Schack & Zahn) Greuter, comb. nov. ≡Hie-
racium sommerfeltii subsp. crinicaesium Schack & Zahn in Mitth. Thüring. Bot. Vereins 41: 94.
1933 ≡Hieracium crinicaesium (Schack & Zahn) Jochen Müll. in Haussknechtia 10: 141. 2004.
Hieracium hypochoeroides subsp. cryptochaetophyllum (T. Georgiev & Zahn) Greuter, comb.
nov. ≡Hieracium wiesbaurianum subsp. cryptochaetophyllum T. Georgiev & Zahn in Izv. Bulg.
Bot. Druz. 4: 80. 1931.
Hieracium hypochoeroides subsp. dalmaticum (Kümmerle & Zahn) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. dalmaticum Kümmerle & Zahn in Magyar Bot. Lapok 25: 309.
1927.
Hieracium hypochoeroides subsp. divergentidens (Schack & Zahn) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. divergentidens Schack & Zahn in Jahrb. Hist. Vereins Fürstentum
Liechtenstein 34: 107. 1934.
Hieracium hypochoeroides subsp. dolichellum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium dolichellum Arv.-Touv. & Gaut., Hieracioth. 5: [in schedis] Gall. No. 276. 1898 ≡
Hieracium wiesbaurianum subsp. dolichellum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
75: 270. 1921.
Hieracium hypochoeroides subsp. epirense (Zahn) Greuter, comb. nov. ≡Hieracium wiesbau-
rianum subsp. epirense Zahn in Engler, Pflanzenr. 75: 264. 1921 ≡Hieracium epirense (Zahn)
Buttler in Strid, Mount. Fl. Greece 2: 618. 1991.
Willdenowia 37 – 2007 153

Hieracium hypochoeroides subsp. fissifurcum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium fissifurcum Arv.-Touv. & Gaut., Hieracioth. 7: [in schedis] Gall. No. 422-423. 1899 ≡
Hieracium wiesbaurianum subsp. fissifurcum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
75: 268. 1921.
Hieracium hypochoeroides subsp. fojnicense (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. fojnicense Zahn in Engler, Pflanzenr. 75: 265. 1921.
Hieracium hypochoeroides subsp. gladiatiforme (Schack & Zahn) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. gladiatiforme Schack & Zahn in Repert. Spec. Nov. Regni Veg.
33: 368. 1934.
Hieracium hypochoeroides subsp. grossidentatum (Jochen Müll.) Greuter, comb. & stat. nov. ≡
Hieracium grossidentatum Jochen Müll. in Haussknechtia 10: 128. 2004.
Hieracium hypochoeroides subsp. guaranum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium guaranum Arv.-Touv. & Gaut., Hieracioth. 20: [in schedis] Hisp. No. 403. 1908 ≡
Hieracium granatense subsp. guaranum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 75:
171. 1921 ≡Hieracium glaucinum subsp. guaranum (Arv.-Touv. & Gaut.) O. Bolòs & Vigo, Fl.
Països Catal. 3: 1053. 1996.
Hieracium hypochoeroides subsp. guestphalicum (Gottschl.) Greuter, comb. nov. ≡Hieracium
wiesbaurianum subsp. guestphalicum Gottschl. in Abh. Westfäl. Mus. Naturk. 53(4): 18. 1991.
Hieracium hypochoeroides subsp. herculanum (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. herculanum Zahn in Engler, Pflanzenr. 76: 268. 1921.
Hieracium hypochoeroides subsp. jacquiniiforme (Litard. & Zahn) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. jacquiniiforme Litard. & Zahn in Arch. Bot. Mém. 2(1): 41. 1928.
Hieracium hypochoeroides subsp. jenzigense (Bornm. & Zahn) Greuter, comb. nov. ≡Hiera-
cium wiesbaurianum subsp. jenzigense Bornm. & Zahn in Engler, Pflanzenr. 82: 1540. 1923 ≡
Hieracium jenzigense (Bornm. & Zahn) Jochen Müll. in Haussknechtia 10: 138. 2004.
Hieracium hypochoeroides subsp. kunitzburgense (Bornm. & Zahn) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. kunitzburgense Bornm. & Zahn in Mitth. Thüring. Bot. Vereins
39: 8. 1930.
Hieracium hypochoeroides subsp. kyllenense (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. kyllenense Zahn in Engler, Pflanzenr. 75: 261. 1921.
Hieracium hypochoeroides subsp. lithophilum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium
lithophilum Arv.-Touv. in Ann. Soc. Linn. Lyon, ser. 2, 34: 67. 1888 ≡Hieracium wiesbau-
rianum subsp. lithophilum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 260. 1921.
Hieracium hypochoeroides subsp. livadicae (O. Behr & al.) Greuter, comb. nov. ≡Hieracium
wiesbaurianum subsp. livadicae O. Behr & al. in Glasn. Skopsk. Naucn. Drustva 20: 123. 1939.
Hieracium hypochoeroides subsp. nervulosum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium nervulosum Arv.-Touv. & Gaut., Hieracioth. 12: [in schedis] Gall. No. 779-781. 1901
≡Hieracium wiesbaurianum subsp. nervulosum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
75: 265. 1921.
Hieracium hypochoeroides subsp. niphanthodes (Bornm. & Zahn) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. niphanthodes Bornm. & Zahn in Mitth. Thüring. Bot. Vereins 39: 7.
1930 ≡Hieracium niphanthodes (Bornm. & Zahn) Jochen Müll. in Haussknechtia 10: 139. 2004.
Hieracium hypochoeroides subsp. parvimaculatum (Jochen Müll.) Greuter, comb. & stat. nov.
≡Hieracium parvimaculatum Jochen Müll. in Haussknechtia 10: 132. 2004.
154 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium hypochoeroides subsp. plauense (Schack & Zahn) Greuter, comb. nov. ≡Hieracium
romieuxianum subsp. plauense Schack & Zahn in Mitth. Thüring. Bot. Vereins 39: 76. 1930 ≡
Hieracium wiesbaurianum subsp. plauense (Schack & Zahn) Zahn in Ascherson & Graebner,
Syn. Mitteleur. Fl. 12(2): 349. 1931 ≡Hieracium plauense (Schack & Zahn) Jochen Müll. in
Haussknechtia 10: 142. 2004.
Hieracium hypochoeroides subsp. pollinense (Zahn) Greuter, comb. nov. ≡Hieracium polli-
nense Zahn in Allg. Bot. Z. Syst. 7: 145. 1901 ≡Hieracium leucophaeum subsp. pollinense Zahn
in Engler, Pflanzenr. 75: 116. 1921
Hieracium hypochoeroides subsp. pseudograniticum (Besse & Zahn) Greuter, comb. nov. ≡
Hieracium wiesbaurianum subsp. pseudograniticum Besse & Zahn in Bull. Murith. Soc. Valais.
Sci. Nat. 32: 179. 1903 ≡Hieracium pseudograniticum (Besse & Zahn) Zahn in Schinz & Keller,
Fl. Schweiz, ed. 2, 2: 303. 1905.
Hieracium hypochoeroides subsp. pseudowiesbaurianiforme (K. Müll. & Zahn) Greuter, comb.
nov. ≡Hieracium wiesbaurianum subsp. pseudowiesbaurianiforme K.Müll.&ZahninAscherson
& Graebner, Syn. Mitteleur. Fl. 12(2): 353. 1931.
Hieracium hypochoeroides subsp. retroversidens (Zahn) Greuter, comb. nov. ≡Hieracium wies-
baurianum subsp. retroversidens Zahn in Engler, Pflanzenr. 75: 266. 1921.
Hieracium hypochoeroides subsp. rudolphopolitanum (Jochen Müll.) Greuter, comb. & stat.
nov. ≡Hieracium rudolphopolitanum Jochen Müll. in Haussknechtia 10: 134. 2004.
Hieracium hypochoeroides subsp. semicinerascens (Bornm. & Zahn) Greuter, comb. nov. ≡
Hieracium wiesbaurianum subsp. semicinerascens Bornm. & Zahn in Magyar Bot. Lapok 25:
309. 1927.
Hieracium hypochoeroides subsp. semiwiesbaurianum (Gottschl.) Greuter, comb. nov. ≡Hie-
racium wiesbaurianum subsp. semiwiesbaurianum Gottschl. in Willdenowia 35: 235. 2005.
Hieracium hypochoeroides subsp. strizakense (O. Behr & al.) Greuter, comb. nov. ≡Hieracium
wiesbaurianum subsp. strizakense O. Behr & al. in Glasn. Skopsk. Naucn. Drustva 20: 25. 1939.
Hieracium hypochoeroides subsp. subcinereum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium bifidum var. subcinereum Arv.-Touv. & Gaut., Hieracioth. 11: [in schedis] Gall. No.
645. 1900 ≡Hieracium wiesbaurianum subsp. subcinereum (Arv.-Touv. & Gaut.) Zahn, Hierac.
Alp. Mar.: 152. 1916.
Hieracium hypochoeroides subsp. supramontanum (Arrigoni) Greuter, comb. & stat. nov. ≡
Hieracium supramontanum Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 249. 1985.
Hieracium hypochoeroides subsp. terrei (de Retz) Greuter, comb. nov. ≡Hieracium wiesbau-
rianum subsp. terrei de Retz in Bull. Soc. Bot. France 112: 444. 1966.
Hieracium hypochoeroides subsp. uazzanum (Zahn) Greuter, comb. nov. ≡Hieracium wiesbau-
rianum subsp. uazzanum Zahn in Engler, Pflanzenr. 75: 271. 1921.
Hieracium hypochoeroides subsp. wiesbaurianiforme Greuter, nom. nov. ≡Hieracium wies-
baurianum var. euwiesbaurianiforme Schack & Zahn in Mitth. Thüring. Bot. Vereins 41: 103.
1933 ≡Hieracium euwiesbaurianiforme (Schack & Zahn) Jochen Müll. in Haussknechtia 10:
136. 2004.
Hieracium hypochoeroides subsp. wiesbaurianum (R. Uechtr.) Greuter, comb. nov. ≡Hiera-
cium wiesbaurianum R. Uechtr. in Baenitz, Herb. Eur. Prosp. 1879: 5. 1879 ≡Hieracium wies-
baurianum R. Uechtr. subsp. wiesbaurianum [per Besse & Zahn in Bull. Murith. Soc. Valais. Sci.
Nat. 32: 179. 1903].
Willdenowia 37 – 2007 155

Hieracium immodestum Greuter, nom. nov. ≡Hieracium schmidtii var. superbum Strömf. in
Öfvers. Kongl. Vetensk.-Akad. Förh. 41(8): 105. 1884 ≡Hieracium pallidum subsp. superbum
(Strömf.) Zahn in Engler, Pflanzenr. 75: 217. 1921 ≡Hieracium superbum (Strömf.) Dahlst. in
Ark. Bot. 3(10): 30. 1904 [non Peter 1884].
Hieracium inconspicuissimum Greuter, nom. nov. ≡Hieracium inconspicuum Wiinst. in Raun-
kiaer, Dansk Exkurs.-Fl., ed. 4: 318. 1922 [non Norrl. 1904].
Hieracium inuloides subsp. subvirens (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium subvirens
Arv.-Touv., Monogr. Pilosella & Hieracium, Suppl.: 31. 1876 ≡Hieracium corymbosum subsp.
subvirens (Arv.-Touv.) Sudre, Hierac. Centre France: 38. 1902 ≡Hieracium conicum subsp.
subvirens (Arv.-Touv.) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
685. 1906 ≡Hieracium crocatum subsp. subvirens (Arv.-Touv.) Zahn in Schinz & Keller, Fl.
Schweiz, ed. 3, 2: 497. 1914.
Hieracium iteophyllum Greuter, nom. nov. ≡Hieracium salicifolium Lindeb., Skand. Hierac. [ed.
poster.]: 25. 1877 [non Arv.-Touv. 1876] ≡Hieracium aestivum subsp. salicifolium (Lindeb.) Zahn
in Engler, Pflanzenr. 79: 919. 1922.
Hieracium jankae subsp. penzesii (Pénzes) Greuter, comb. & stat. nov. ≡Hieracium penzesii
Pénzes in Borbásia 1(10): 168. 1939.
Hieracium jordanii subsp. eriosphaera (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium bom-
bycinum subsp. eriosphaera Nägeli & Peter, Hierac. Mitt.-Eur. 2: 308. 1889 ≡Hieracium erio-
sphaera (Nägeli & Peter) Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 306. 1905.
Hieracium juranum subsp. benthamianum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hiera-
cium benthamianum Arv.-Touv. & Gaut., Hieracioth. 4: [in schedis] Gall. No. 250-253, Hisp.
No. 59. 1898 ≡Hieracium mougeotii subsp. benthamianum (Arv.-Touv. & Gaut.) Zahn in
Engler, Pflanzenr. 75: 183. 1921 ≡Hieracium vogesiacum subsp. benthamianum (Arv.-Touv. &
Gaut.) O. Bolòs & Vigo, Fl. Països Catal. 3: 1106. 1996.
Hieracium juranum subsp. chrysanthemoides (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium chrysanthemoides Arv.-Touv. & Gaut., Hieracioth. 4: [in schedis] Hisp. No. 47-49. 1898 ≡
Hieracium mougeotii subsp. chrysanthemoides (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
75: 187. 1921 ≡Hieracium vogesiacum subsp. chrysanthemoides (Arv.-Touv. & Gaut.) O. Bolòs &
Vigo, Fl. Països Catal. 3: 1106. 1996.
Hieracium juranum subsp. conocerinthe (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hiera-
cium conocerinthe Arv.-Touv. & Gaut., Hieracioth. 20: [in schedis] Hisp. No. 377-381. 1908 ≡
Hieracium mougeotii subsp. conocerinthe (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 75:
182. 1921.
Hieracium juranum subsp. doronicoides (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium doro-
nicoides Arv.-Touv. in Bull. Soc. Bot. France 41: 340. 1894 ≡Hieracium mougeotii subsp. doro-
nicoides (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 182. 1921 ≡Hieracium vogesiacum subsp.
doronicoides (Arv.-Touv.) O. Bolòs & Vigo, Fl. Països Catal. 3: 1106. 1996.
Hieracium juranum subsp. lagopinum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hieracium
lagopinum Arv.-Touv. & Gaut., Hieracioth. 7: [in schedis] Hisp. No. 109. 1899 ≡Hieracium
mougeotii subsp. lagopinum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 75: 187. 1921 ≡
Hieracium vogesiacum subsp. lagopinum (Arv.-Touv. & Gaut.) O. Bolòs & Vigo, Fl. Països
Catal. 3: 1106. 1996.
Hieracium juranum subsp. pullatum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium pullatum
Arv.-Touv., Spicil. Rar. Hierac.: 40. 1886 ≡Hieracium mougeotii subsp. pullatum (Arv.-Touv.)
Rouy, Fl. France 9: 306. 1905.
156 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium juranum subsp. stelliceps (Zahn) Greuter, comb. nov. ≡Hieracium mougeotii subsp.
stelliceps Zahn in Engler, Pflanzenr. 75: 187. 1921 ≡Hieracium vogesiacum subsp. stelliceps
(Zahn) O. Bolòs & Vigo, Fl. Països Catal. 3: 1106. 1996.
Hieracium juranum subsp. tournefortianum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium tournefortianum Arv.-Touv. & Gaut., Hieracioth. 7: [in schedis] Hisp. No. 89-94. 1899 ≡
Hieracium mougeotii subsp. tournefortianum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
75: 183. 1921 ≡Hieracium vogesiacum subsp. tournefortianum (Arv.-Touv. & Gaut.) O. Bolòs
& Vigo, Fl. Països Catal. 3: 1106. 1996.
Hieracium jurassicum subsp. adenocalathium (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium doronicifolium grex ade-
nocalathium Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1876. 1901 ≡Hieracium adenoca-
lathium (Zahn) Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 317. 1905 ≡Hieracium juranum
subsp. adenocalathium (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten
Naturwiss. 40: 634. 1906.
Hieracium jurassicum subsp. aligerum (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 534. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium juranum subsp. aligerum Zahn
in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 625. 1906 ≡Hieracium ali-
gerum (Zahn) Prain, Index Kew., Suppl. 4: 108. 1913.
Hieracium jurassicum subsp. apiculatifrons (Zahn) Greuter, comb. nov. ≡Hieracium juranum
subsp. apiculatifrons Zahn in Engler, Pflanzenr. 77: 781. 1921.
Hieracium jurassicum subsp. barbeyi (Post) Greuter, comb. & stat. nov. ≡Hieracium barbeyi
Post in Bull. Herb. Boissier 1: 24. 1893.
Hieracium jurassicum subsp. boltigense (Zahn) Greuter, comb. nov. ≡Hieracium juranum subsp.
boltigense Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 333. 1905 ≡Hieracium boltigense
(Zahn) Prain, Index Kew., Suppl. 4: 109. 1913.
Hieracium jurassicum subsp. borreri (Syme) Greuter, comb. & stat. nov. ≡Hieracium borreri
Syme in Smith, Engl. Bot., ed. 3, 5: 212. 1866.
Hieracium jurassicum subsp. cichoriaceum (Arv.-Touv.) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 536. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium cichoriaceum
Arv.-Touv., Monogr. Pilosella & Hieracium, Suppl.: 25. 1876 ≡Hieracium juranum subsp.
cichoriaceum (Arv.-Touv.) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss.
40: 632. 1906.
Hieracium jurassicum subsp. decrescentifolium (Murr & Zahn) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium juranum subsp.
decrescentifolium Murr & Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
629. 1906 ≡Hieracium decrescentifolium (Murr & Zahn) Prain, Index Kew., Suppl. 4: 111. 1913.
Hieracium jurassicum subsp. densifloccum (Zahn) Greuter, comb. nov. ≡Hieracium juranum
var. densifloccum Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 329. 1905 ≡Hieracium
juranum subsp. densifloccum (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten
Naturwiss. 40: 625. 1906 ≡Hieracium densifloccum (Zahn) Prain, Index Kew., Suppl. 4: 111.
1913.
Hieracium jurassicum subsp. elegantissimum (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium integrifolium subsp.ele
-
gantissimum Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1882. 1901 ≡Hieracium juranum
subsp. elegantissimum (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss.
40: 627. 1906 ≡Hieracium elegantissimum (Zahn) Prain, Index Kew., Suppl. 4: 111. 1913.
Willdenowia 37 – 2007 157

Hieracium jurassicum subsp. esseranum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium esseranum Arv.-Touv. & Gaut., Hieracioth. 14: [in schedis] Hisp. No. 225-226. 1902 ≡
Hieracium juranum subsp. esseranum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 77: 784.
1921.
Hieracium jurassicum subsp. hegetschweileri (Zahn) Greuter, comb. nov. ≡Hieracium heget-
schweileri Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1895. 1901 ≡Hieracium juranum subsp.
hegetschweileri (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
624. 1906.
Hieracium jurassicum subsp. hemiplecoides (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium hemiplecoides Arv.-Touv. & Gaut., Hieracioth. 5: [in schedis] Gall. No. 300. 1898 ≡
Hieracium juranum subsp. hemiplecoides (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 77:
783. 1921.
Hieracium jurassicum subsp. hemiplecum (Arv.-Touv.) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium hemiplecum
Arv.-Touv. in Bull. Soc. Dauphin. Echange Pl.: 105. 1877 ≡Hieracium juranum subsp. hemi-
plecum (Arv.-Touv.) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
627. 1906.
Hieracium jurassicum subsp. jaceoides (Arv.-Touv.) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 536. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium jaceoides Arv.-Touv.,
Monogr. Pilosella & Hieracium: 40. 1873 ≡Hieracium juranum subsp. jaceoides (Arv.-Touv.)
Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1878. 1901 ≡Hieracium integrifolium subsp.ja-
ceoides (Arv.-Touv.) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
638. 1906.
Hieracium jurassicum subsp. lancifrons (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 536. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium juranum subsp. lancifrons
Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 330. 1905 ≡Hieracium lancifrons (Zahn) Prain,
Index Kew., Suppl. 4: 114. 1913.
Hieracium jurassicum subsp. leiophyton (Dahlst.) Greuter, comb. nov. ≡Hieracium leiophyton
Dahlst. in Lindman, Sv. Fanerogamfl.: 627. 1918 ≡Hieracium juranum subsp. leiophyton (Dahlst.)
Zahn in Engler, Pflanzenr. 77: 785. 1921.
Hieracium jurassicum subsp. multiserratum (Zahn) Greuter, comb. nov. ≡Hieracium juranum
subsp. multiserratum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
633. 1906 ≡Hieracium multiserratum (Zahn) Prain, Index Kew., Suppl. 4: 116. 1913.
Hieracium jurassicum subsp. obesum (Zahn) Greuter, comb. nov. ≡Hieracium juranum subsp.
obesum Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 333. 1905 ≡Hieracium obesum (Zahn)
Prain, Index Kew., Suppl. 4: 117. 1913.
Hieracium jurassicum subsp. papyraceum (Zahn) Greuter, comb. nov. ≡Hieracium juranum
subsp. papyraceum Zahn in Engler, Pflanzenr. 77: 787. 1921 [non Hieracium papyraceum Gren.
1869].
Hieracium jurassicum subsp. perfoliatiforme (Zahn) Greuter, comb. nov. ≡Hieracium juranum
subsp. perfoliatiforme Zahn in Engler, Pflanzenr. 77: 787. 1921.
Hieracium jurassicum subsp. prenanthopsis (Murr & Zahn) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium juranum subsp.
prenanthopsis Murr & Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1878. 1901 ≡Hieracium
prenanthopsis (Murr & Zahn) Prain, Index Kew., Suppl. 4: 118. 1913.
158 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium jurassicum subsp. pseudalbinum (R. Uechtr.) Greuter, comb. nov. ≡Hieracium
pseudalbinum R. Uechtr. in Jahresber. Schles. Ges. Vaterl. Cult. 62: 314. 1884 ≡Hieracium ju-
ranum subsp. pseudalbinum (R. Uechtr.) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1879.
1901.
Hieracium jurassicum subsp. pseudelatum (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 534. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium juranum var. pseu-
delatum Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 329. 1905 ≡Hieracium juranum subsp.
pseudelatum (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
623. 1906 ≡Hieracium pseudelatum (Zahn) Prain, Index Kew., Suppl. 4: 119. 1913.
Hieracium jurassicum subsp. pseudohemiplecum (Zahn) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium integrifolium
subsp. pseudohemiplecum Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1883. 1901 ≡Hieracium
juranum subsp. pseudohemiplecum (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges.
Gesammten Naturwiss. 40: 628. 1906 ≡Hieracium pseudohemiplecum (Zahn) Prain, Index
Kew., Suppl. 4: 119. 1913.
Hieracium jurassicum subsp. pseudojuranum (Arv.-Touv.) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 535. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium pseudojuranum
Arv.-Touv., Monogr. Pilosella & Hieracium, Suppl.: 24. 1876 ≡Hieracium juranum subsp. pseu-
dojuranum (Arv.-Touv.) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1878. 1901.
Hieracium jurassicum subsp. subperfoliatum (Arv.-Touv.) [Hirschm. in Denkschr. Schweiz.
Naturf. Ges. 81: 534. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium perfoliatum
Arv.-Touv. in Bull. Soc. Dauphin. Echange Pl.: 88. 1880 [non Froel. 1838] ≡Hieracium subper-
foliatum Arv.-Touv., Notes Pl. Alpes: 12. 1883 ≡Hieracium juranum subsp. subperfoliatum
(Arv.-Touv.) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1878. 1901.
Hieracium jurassicum subsp. tamderense (Hub.-Mor.) Greuter, comb. & stat. nov. ≡Hiera-
cium tamderense Hub.-Mor. in Notes Roy. Bot. Gard. Edinburgh 33: 246. 1974.
Hieracium jurassicum subsp. tephroprenanthes (Zahn) Greuter, comb. nov. ≡Hieracium jura-
num subsp. tephroprenanthes Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Natur-
wiss. 40: 633. 1906 ≡Hieracium tephroprenanthes (Zahn) Prain, Index Kew., Suppl. 4: 122.
1913.
Hieracium jurassicum subsp. urezzae (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges. 81:
536. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium juranum subsp. urezzae Zahn in
Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 632. 1906 ≡Hieracium urezzae
(Zahn) Prain, Index Kew., Suppl. 4: 122. 1913.
Hieracium klisurae subsp. caparinum (Zahn) Greuter, comb. nov. ≡Hieracium pseuderiopus
subsp. caparinum Zahn in Engler, Pflanzenr. 79: 1073. 1922 ≡Hieracium pojoritense subsp.
caparinum (Zahn) Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3): 705. 1938.
Hieracium klisurae subsp. pseuderiopus (Zahn) Greuter, comb. nov. ≡Hieracium pseuderiopus
Zahn in Engler, Pflanzenr. 79: 1073. 1922 ≡Hieracium pseuderiopus Zahn subsp. pseuderiopus
[per Zahn in Engler, Pflanzenr. 79: 1073. 1922] ≡Hieracium pojoritense subsp. pseuderiopus
(Zahn) Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3): 704. 1938.
Hieracium kuekenthalianum subsp. amaurocranum (Zahn) Greuter, comb. nov. ≡Hieracium
intermedium Gus. Schneid. in Deutsche Bot. Monatsschr. 6: 122. 1888 [non Lapeyr. 1813] ≡Hie-
racium tephrosoma subsp. amaurocranum Zahn in Engler, Pflanzenr. 77: 697. 1921 ≡Hieracium
amaurocranum (Zahn) Holub in Preslia 51: 281. 1979.
Hieracium kuekenthalianum subsp. andreanszkyanum (F. Kováts) Greuter, comb. & stat. nov.
≡Hieracium andreanszkyanum F. Kováts in Borbásia 5-6: 77. 1946.
Willdenowia 37 – 2007 159

Hieracium kuekenthalianum subsp. brachypogon (Zahn) [Hub.-Mor. in Bauhinia 4: 153. 1971,
comb. inval., ex] Greuter, comb. nov. ≡Hieracium dolichaetum subsp. brachypogon Zahn in
Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1850. 1901 ≡Hieracium brachypogon (Zahn) Zahn in
Schinz & Keller, Fl. Schweiz, ed. 2, 2: 315. 1905 ≡Hieracium bocconei subsp. brachypogon
(Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 557. 1906 ≡
Hieracium tephrosoma subsp. brachypogon (Zahn) Zahn in Engler, Pflanzenr. 77: 697. 1921.
Hieracium kuekenthalianum subsp. tephrosoma (Nägeli & Peter) Greuter, comb. nov. ≡Hiera-
cium ctenodon subsp. tephrosoma Nägeli & Peter, Hierac. Mitt.-Eur. 2: 206. 1886 ≡Hieracium
tephrosoma (Nägeli & Peter) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1853. 1901.
Hieracium kuekenthalianum subsp. trivialiforme (Zahn) [Hub.-Mor. in Bauhinia 4: 153. 1971,
comb. inval., ex] Greuter, comb. nov. ≡Hieracium tephrosoma subsp. trivialiforme Zahn in
Engler, Pflanzenr. 77: 697. 1921 [non Hieracium trivialiforme Schljakov 1966].
Hieracium kuekenthalianum subsp. valsicum (Kaeser & Zahn) Greuter, comb. nov. ≡Hiera-
cium valsicum Kaeser & Zahn in Zahn, Hieracioth. Eur. 8: 21. 1913 ≡Hieracium tephrosoma
subsp.valsicum(Kaeser & Zahn) Zahn in Engler, Pflanzenr. 77: 697. 1921.
Hieracium lachenalii subsp. alpestre (R. Uechtr.) Greuter, comb. nov. ≡Hieracium vulgatum
var. alpestre R. Uechtr. in Fiek, Fl. Schlesien [2]: 276. 1881 [non Hieracium alpestre Jacq. 1774]
≡Hieracium vulgatum subsp. alpestre (R. Uechtr.) Zahn in Neue Denkschr. Allg. Schweiz. Ges.
Gesammten Naturwiss. 40: 406. 1906 ≡Hieracium vulgatum subsp. ficinii Zahn in Engler,
Pflanzenr. 76: 386. 1921, nom. illeg.
Hieracium lachenalii subsp. argillaceoides (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium vul-
gatum subsp. argillaceoides Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 239. 1907 ≡
Hieracium argillaceoides (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 269. 1960.
Hieracium lachenalii subsp. cladophorum (Vuk.) Greuter, comb. nov. ≡Hieracium platyphyl-
lopodum f. cladophorum Vuk., Hierac. Croat.: 13. 1858 ≡Hieracium cladophorum (Vuk.) Vuk. in
Rad Jugoslav. Akad. Znan. 34: 18. 1876 ≡Hieracium divisum subsp. cladophorum (Vuk.) Zahn in
Ann. Hist.-Nat. Mus. Natl. Hung. 8: 62. 1910 ≡Hieracium diaphanoides subsp. cladophorum
(Vuk.) Zahn in Engler, Pflanzenr. 76: 358. 1921.
Hieracium lachenalii subsp. deductum (Sudre) Greuter, comb. nov. ≡Hieracium deductum Sudre
in Bull. Assoc. Pyrén. Echange Pl. 8: 12. 1898 ≡Hieracium vulgatum subsp. deductum (Sudre)
Sudre, Hierac. Centre France: 57. 1902.
Hieracium lachenalii subsp. epichlorum (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium vulga-
tum var. epichlorum Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 238. 1907 ≡Hieracium
vulgatum subsp. epichlorum (Litv. & Zahn) Zahn in Spisok Rast. Gerb. Russk. Fl. Bot. Muz. Imp.
Akad. Nauk Fasc. 42: 22. 1909 ≡Hieracium epichlorum (Litv. & Zahn) Üksip in Komarov, Fl.
SSSR 30: 30. 1960.
Hieracium lachenalii subsp. erythropodum (R. Uechtr.) Greuter, comb. nov. ≡Hieracium ery-
thropodum R. Uechtr. in Fiek, Fl. Schlesien [2]: 272. 1881 ≡Hieracium vulgatum subsp. erythro-
podum (R. Uechtr.) Zahn in Engler, Pflanzenr. 76: 375. 1921.
Hieracium lachenalii subsp. gustavianum (Üksip) Greuter, comb. & stat. nov. ≡Hieracium gus-
tavianum Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 496. 1959.
Hieracium lachenalii subsp. hypopitys (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium divisum
subsp. hypopitys Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 237. 1907 ≡Hieracium vul-
gatum subsp. hypopitys (Litv. & Zahn) Zahn in Engler, Pflanzenr. 76: 368. 1921 ≡Hieracium
hypopitys (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 261. 1960.
160 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium lachenalii subsp. juratzkanum (Zahn) Greuter, comb. nov. ≡Hieracium vulgatum
subsp. juratzkanum Zahn in Engler. Pflanzenr. 76: 368. 1921 ≡Hieracium juratzkanum (Zahn)
Schljakov in Cvelev, Fl. Evr. Casti SSSR 8: 248. 1989.
Hieracium lachenalii subsp. lepidiceps (Dahlst.) Greuter, comb. nov. ≡Hieracium irriguum
subsp. lepidiceps Dahlst. in Kongl. Svenska Vetensk. Acad. Handl., ser. 2, 26(3): 128. 1894 ≡
Hieracium lepidiceps (Dahlst.) Prain, Index Kew., Suppl. 5: 129. 1921 ≡Hieracium vulgatum
subsp. lepidiceps (Dahlst.) Zahn in Engler. Pflanzenr. 76: 372. 1921.
Hieracium lachenalii subsp. lortetiae (Balb.) Greuter, comb. & stat. nov. ≡Hieracium lortetiae
Balb., Fl. Lyon. 1: 450. 1827.
Hieracium lachenalii subsp. macrophyllopodum (Zahn) Greuter, comb. nov. ≡Hieracium vul-
gatum subsp. macrophyllopodum Zahn in Vestn. Tiflissk. Bot. Sada 22: 28. 1912 ≡Hieracium
macrophyllopodum (Zahn) Üksip in Komarov, Fl. SSSR 30: 255. 1960.
Hieracium lachenalii subsp. mertinii (C. C. Gmel.) Gottschl., comb. & stat. nov. ≡Hieracium
mertinii C. C. Gmel., Fl. Bad. 4: 581. 1826.
This taxon corresponds to “Hieracium lachenalii subsp. lachenalii” in the sense of
Zahn and subsequent authors. However, the type H. lachenalii Suter, recently traced
(details to be published elsewhere), belongs to the taxon generally known as H.
lachenalii subsp. jaccardii (Zahn) Zahn. The foregoing combination is therefore
necessary. G. Gottschlich
Hieracium lachenalii subsp. obscuriceps (Dahlst.) Greuter, comb. nov. ≡Hieracium irriguum
subsp. obscuriceps Dahlst. in Kongl. Svenska Vetensk. Acad. Handl., ser. 2, 26(3): 141. 1894 ≡
Hieracium obscuriceps (Dahlst.) Prain, Index Kew., Suppl. 5: 131. 1921 [non (Nägeli & Peter)
Prain 1913] ≡Hieracium vulgatum subsp. subobscuriceps Zahn in Engler, Pflanzenr. 76: 374.
1921, nom. illeg. ≡Hieracium lachenalii subsp. subobscuriceps Zahn in Ascherson & Graebner,
Syn. Mitteleur. Fl. 12(2): 562. 1934, nom. illeg. ≡Hieracium subobscuriceps Üksip in Komarov,
Fl. SSSR 30: 251. 1960.
Hieracium lachenalii subsp. subhastulatum (Zahn) Greuter, comb. nov. ≡Hieracium vulgatum
subsp. subhastulatum Zahn in Vestn. Tiflissk. Bot. Sada 22: 28. 1912 ≡Hieracium subhastul-
atum (Zahn) Üksip in Komarov, Fl. SSSR 30: 257. 1960.
Hieracium lachenalii subsp. violascens (Dahlst.) Greuter, comb. nov. ≡Hieracium murorum
subsp. violascens Dahlst. in Bot. Not. 1892: 164. 1892 ≡Hieracium violascens (Dahlst.) Dahlst.
in Lindman, Sv. Fanerogamfl.: 626. 1918 [non Borbás 1878] ≡Hieracium vulgatum subsp.
violascens (Dahlst.) Zahn in Engler, Pflanzenr. 76: 378. 1921 ≡Hieracium vulgatum subsp. alm-
quistianum Zahn in Engler, Pflanzenr. 79: 1145. 1922, nom. illeg. ≡Hieracium almquistianum
Johanss. in Ark. Bot. 21A(15): 54. 1927 ≡Hieracium subviolascens P. D. Sell in Sell & Murrell,
Fl. Great Britain & Ireland 4: 551. 2006, nom. illeg.
Hieracium laevigatum subsp. achalzichiense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
achalzichiense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 473. 1959.
Hieracium laevigatum subsp. glareosum (Lönnr.) Greuter, comb. nov. ≡Hieracium rigidum
subsp. glareosum Lönnr. in Öfvers. Kongl. Vetensk.-Akad. Förh. 39(4): 87. 1882 ≡Hieracium
rigidum subsp. mixopolium Dahlst. in Kongl. Svenska Vetensk. Acad. Handl., ser. 2, 26(3): 224.
1894, nom. illeg. ≡Hieracium mixopolium Dahlst., Herb. Hierac. Scand. 10: [in schedis] No. 89.
1896 ≡Hieracium laevigatum subsp. mixopolium (Dahlst.) Zahn in Engler, Pflanzenr. 79: 881.
1922.
Hieracium laevigatum subsp. hypopityforme (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
hypopityforme Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 495. 1959.
Willdenowia 37 – 2007 161

Hieracium laevigatum subsp. leucothyrsum (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium
knafii subsp. leucothyrsum Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 240. 1907 ≡
Hieracium levicaule subsp. leucothyrsum (Litv. & Zahn) Zahn in Engler, Pflanzenr. 77: 859.
1921 ≡Hieracium leucothyrsum (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 106. 1960.
Hieracium lazistanum subsp. leithneri (Boiss.) Greuter, comb. nov. ≡Hieracium sartorianum
var. leithneri Boiss., Fl. Orient., Suppl.: 328. 1888 ≡Hieracium colophyllum subsp. leithneri
(Boiss.) Nägeli & Peter, Hierac. Mitt.-Eur. 2: 315. 1889 ≡Hieracium leithneri (Boiss.) Peter in
Engler & Prantl, Nat. Pflanzenfam. 4(5): 384. 1894.
Hieracium lazistanum subsp. marotii (T. Georgiev & Zahn) Greuter, comb. & stat. nov. ≡Hie-
racium marotii T. Georgiev & Zahn in Izv. Bulg. Bot. Druz. 6: 74. 1934.
Hieracium lazistanum subsp. pelliculatiforme (Zahn) Greuter, comb. nov. ≡Hieracium vanda-
sii subsp. pelliculatiforme Zahn in Magyar Bot. Lapok 8: 301. 1910 ≡Hieracium leithneri subsp.
pelliculatiforme (Zahn) Zahn in Engler, Pflanzenr. 77: 586. 1921.
Hieracium lazistanum subsp. velenovskyanum (Zahn) Greuter, comb. nov. ≡Hieracium skor-
pilii Velen., Reliq. Mirkvic.: 17. 1922 [non Freyn 1891] ≡Hieracium leithneri subsp. velenovs-
kyanum Zahn in Arh. Minist. Zeml. Dbrz. Imoti 4-5 [= Stojanov & Stefanov, Fl. Bblg.]: 1246.
1925.
Hieracium legionense subsp. cantabrimontanum (de Retz) Greuter, comb. nov. ≡Hieracium
gastonianum subsp. cantabrimontanum de Retz in Bull. Soc. Bot. France, Lett. Bot. 127: 84.
1980 ≡Hieracium cantabrimontanum (de Retz) Mateo in Anales Jard. Bot. Madrid 54: 367. 1996
[incl. Hieracium gastonianum subsp. pseudodulacianum de Retz, l.c. ≡Hieracium pseudodula-
cianum (de Retz) Mateo, l.c.].
Hieracium leiopogon subsp. iolai (Arrigoni) Greuter, comb. & stat. nov. ≡Hieracium iolai
Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 231. 1985.
Hieracium levicaule subsp. campylodon (Zahn) Greuter, comb. nov. ≡Hieracium caesium subsp.
campylodon Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 478. 1906 ≡
Hieracium levicaule subsp. campylodontum (Zahn) Zahn in Engler, Pflanzenr. 76: 402. 1921, nom.
illeg.
Hieracium levicaule subsp. erubescens (Boreau) Greuter, comb. & stat. nov. ≡Hieracium eru-
bescens Boreau, Fl. Centre France, ed. 3, 2: 398. 1857.
Hieracium levicaule subsp. gudergomiense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
gudergomiense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 492.
1959.
Hieracium levicaule subsp. gudissiense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium gu-
dissiense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 488. 1959.
Hieracium levicaule subsp. karjaginii (Üksip) Greuter, comb. & stat. nov. ≡Hieracium kar-
jaginii Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 489. 1959.
Hieracium levicaule subsp. lepidulum (Dahlst.) Greuter, comb. nov. ≡Hieracium lepidulum
Dahlst., Herb. Hierac. Scand. 2: [in schedis] No. 77. 1892 [≡Hieracium [subsp.] lepidulum
Stenstr., Värmländsk. Archierac.: 50. 1889, nom. inval.] ≡Hieracium vulgatum subsp. lepidulum
(Dahlst.) Zahn in Engler, Pflanzenr. 76: 379. 1921.
Hieracium levicaule subsp. leptogrammoides (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
leptogrammoides Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 502.
1959.
Hieracium levicaule subsp. lipskyanum (Üksip) Greuter, comb. & stat. nov. ≡Hieracium lips-
kyanum Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 494. 1959.
162 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium levicaule subsp. sbaënse (Üksip) Greuter, comb. & stat. nov. ≡Hieracium sbaënse
Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 502. 1959.
Hieracium levicaule subsp. sublonchodes [Murr & Zahn in Ascherson & Graebner, Syn.
Mitteleur. Fl. 12(2): 709. 1935, descr. germ.] Murr & Zahn, subsp. nov. – Holotype: Germany,
Bavaria, “Grammersberg am Scharfreiter”, Harz (M). – Description: “Folia quam in subsp. lon-
chodes Murr & Zahn minora, firma, superne pilosa vel glabra, oblongo-ovata vel oblongo-
lanceolata, vix denticulata vel ± breviserrata; caulina 3-5, oblongo-lanceolata vel linearia, (su-
periora utrinque) floccosa. Capitula 3-10, acladium 1-3 cm longum, pedunculi leviter pilosi,
saepe ± eglandulosi. Involucrum 9-10 mm longum, obscurum, parce vel ditius floccosum et pilo-
sum, pauciglandulosum; squamae angustae vel sublatiusculae, acutissimae vel ± obtusae, ± viri-
dimarginatae. Stylus initio flavobrunneum.”– H. Zahn.
Hieracium levicaule subsp. uczanssuense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium uc-
zanssuense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 487. 1959.
Hieracium liptoviense subsp. amaurosphaera [Vetter & Zahn in Ascherson & Graebner, Syn.
Mitteleur. Fl. 12(3): 194. 1936, descr. germ.] Vetter & Zahn, subsp. nov. – Holotype: Austria,
“Steirisch-kärntnerisches Grenzgebiet, unter Gestrüpp nächst der Winklalm bei Turrach”,
10.7.1931, Vetter (W-1950-3322). – Description: “Subspeciei amaurothecae simile sed folia ob-
longo-ovata vel ovato- vel elliptico-lanceolata, magna (interdum maxima), basi breviter vel
longe angustata, grosse pluridentata vel copiose serrata, brevissime pilosa (superne plerumque
glabra), olivaceo- et dorso caesio-virentia; caulis glaber foliis 1-2 ellipticis vel elongato-lan-
ceolatis (vel superiore angusto), magnis, grosse acute dentatis. Capitula 3-7(-16); acladium 2 cm
longum; rami 2-3(-5). Glandulae copiosae, elongatae vel saepe ± breves et ± subtiles. Invo-
lucrum 10-12 mm longum, crassum; squamae ± latiusculae, obtusiusculae vel acutae, atrae, in-
teriores ± viridi-marginatae, exteriores infra parce floccosae. Flores eciliati.”– H. Zahn.
Hieracium liptoviense subsp. coinotokum [O. Behr, E. Behr & Zahn in Ascherson & Graebner,
Syn. Mitteleur. Fl. 12(3): 193. 1936, descr. germ.] O. Behr, E. Behr & Zahn, subsp. nov. –
Holotype: Slovakia, “Hohe Tatra, zwischen Popper- und Csorber See”, 18.7.1931, Behr & Behr
(W-1950-3319). – Description: “30-40 cm altum, (1-)3-5(-8)-cephalum; acladium 15-50(-80) mm
longum; rami (0-)2-3(-4) dissiti, 1(1)-cephali; pedunculi grisei, pilis 1-2 mm longis interdum sat
copiosis, vix vel modice parvoglandulosi, bracteis 2-3 longis et angustis; involucrum 10-12 mm
longum, valde pilosum, pauciglandulosum; squamae ± latiusculae, longissime acuminatae,
acutiusculae vel acutissimae, intimae angustissimae. Flores ± angusti, profunde dentati. Folia
flavescenti-prasoviridia dorso albescenticaesia; basalia oblonga et obtusiuscula vel longe lan-
ceolata et acuta, basi breviter vel longissime angustata, superne glabra, denticulata vel plurima
cum caulinis inferioribus 1-2 (anguste lanceolatis) breviter vel longius serrata, dentibus numerosis
saepe angustis, acutis, prorsus curvatis; folium caulinum summum angustum margine integrum.” –
H. Zahn.
Hieracium litwinowianum subsp. ermaniense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
ermaniense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 469. 1959.
Hieracium longifolium subsp. neocerinthoides (Arv.-Touv. & Briq.) Greuter, comb. nov. ≡
Hieracium neocerinthoides Arv.-Touv. & Briq. in Annuaire Conserv. Jard. Bot. Genève 3: 138.
1899 ≡Hieracium misaucinum subsp. neocerinthoides (Arv.-Touv. & Briq.) Zahn in Schinz &
Keller, Fl. Schweiz, ed. 2, 2: 302. 1905 ≡Hieracium vogesiacum subsp. neocerinthoides (Arv.-
Touv. & Briq.) Zahn in Schinz & Keller, Fl. Schweiz, ed. 3, 2: 443. 1914.
Hieracium loscosianum subsp. bujedoanum (Arv.-Touv.) Greuter, comb. nov. ≡Hiercium bujedoa-
num Arv.-Touv., Hier. Gall. Hisp. Cat.: 310. 1913 ≡Hieracium baeticum subsp. bujedoanum
(Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 200. 1921.
Hieracium loscosianum subsp. capillosum (Pau) Greuter, comb. & stat. nov. ≡Hieracium ca-
pillosum Pau in Bol. Soc. Aragonesa Ci. Nat. 2: 286. 1903.
Willdenowia 37 – 2007 163

Hieracium loscosianum subsp. saxicapellum (Arv.-Touv. & Gand.) Greuter, comb. & stat. nov.
≡Hieracium saxicapellum Arv.-Touv. & Gand. in Arvet-Touvet, Hier. Gall. Hisp. Cat.: 164.
1913.
Hieracium lucidum subsp. cophanense (Lojac.) Greuter, comb. & stat. nov. ≡Hieracium co-
phanense Lojac., Fl. Sicula 2(1): 218. 1903.
Hieracium macilentum subsp. beauverdianum (Besse & Zahn) Greuter, comb. nov. ≡Hiera-
cium doronicifolium subsp. beauverdianum Besse & Zahn in Bull. Murith. Soc. Valais. Sci. Nat.
32: 182. 1903 ≡Hieracium beauverdianum (Besse & Zahn) Zahn in Neue Denkschr. Allg.
Schweiz. Ges. Gesammten Naturwiss. 40: 642. 1906 ≡Hieracium epimedium subsp. beauver-
dianum (Besse & Zahn) Zahn in Engler, Pflanzenr. 77: 803. 1921.
Hieracium macilentum subsp. crepidodon (Wilczek & Zahn) Greuter, comb. nov. ≡Hieracium
epimedium subsp. crepidodon Wilczek & Zahn in Bull. Murith., Soc. Valais. Sci. Nat. 42: 202.
1925.
Hieracium macilentum subsp. epimedium (Fr.) Greuter, comb. nov. ≡Hieracium epimedium Fr.
in Uppsala Univ. Årsskr. 1862: 103. 1862 ≡Hieracium epimedium Fr. subsp. epimedium [per
Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 649. 1906].
Hieracium macilentum subsp. fariniceps (Besse & Zahn) Greuter, comb. nov. ≡Hieracium
beauverdianum subsp. fariniceps Besse & Zahn in Reichenbach, Icon. Fl. Germ. Helv. 19(2):
252. 1910 [non Hieracium fariniceps Dahlst. 1894] ≡Hieracium epimedium subsp. subfariniceps
Zahn in Engler, Pflanzenr. 77: 806. 1921, nom. illeg.
Hieracium macilentum subsp. pseudexilentum (Besse & Zahn) Greuter, comb. nov. ≡Hiera-
cium beauverdianum subsp. pseudexilentum Besse & Zahn in Neue Denkschr. Allg. Schweiz.
Ges. Gesammten Naturwiss. 40: 643. 1906 ≡Hieracium pseudexilentum (Besse & Zahn) Prain,
Index Kew., Suppl. 4: 119. 1913 ≡Hieracium epimedium subsp. pseudexilentum (Besse & Zahn)
Zahn in Engler, Pflanzenr. 77: 803. 1921.
Hieracium macilentum subsp. sparsipilum (Zahn) Greuter, comb. nov. ≡Hieracium sparsipi-
lum (Zahn) Prain, Index Kew., Suppl. 4: 120. 1913 ≡Hieracium juranum subsp. sparsipilum
Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 332. 1905 ≡Hieracium juraniforme subsp.
sparsipilum (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 648.
1906 ≡Hieracium epimedium subsp. sparsipilum (Zahn) Zahn in Engler, Pflanzenr. 77: 808.
1921.
Hieracium macilentum subsp. staui (Belli) Greuter, comb. & stat. nov. ≡Hieracium staui Belli
in Fiori & Paoletti, Fl. Italia 3: 472. 1904.
Hieracium macilentum subsp. subepimedium (Murr & Zahn) Greuter, comb. nov. ≡Hieracium
juranum subsp. subepimedium Murr & Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1881. 1901
≡Hieracium epimedium subsp. subepimedium (Murr & Zahn) Zahn in Neue Denkschr. Allg.
Schweiz. Ges. Gesammten Naturwiss. 40: 649. 1906 ≡Hieracium subepimedium (Murr & Zahn)
Prain, Index Kew., Suppl. 4: 121. 1913.
Hieracium macilentum subsp. subvulsum (Zahn) Greuter, comb. nov. ≡Hieracium integrifo-
lium subsp. subvulsum Zahn in Schinz & Keller, Fl. Schweiz, ed. 3, 2: 480. 1914.
Hieracium macilentum subsp. wimmeri (R. Uechtr.) Greuter, comb. nov. ≡Hieracium wimmeri
R. Uechtr. in Österr. Bot. Z. 22: 277. 1872 ≡Hieracium juranum subsp. wimmeri (R. Uechtr.)
Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1881. 1901 ≡Hieracium epimedium subsp. wim-
meri (R. Uechtr.) Zahn in Schinz & Keller, Fl. Schweiz, ed. 3, 2: 481. 1914.
Hieracium macilentum subsp. wimmerioides (Zahn) Greuter, comb. nov. ≡Hieracium juranum
var. wimmerioides Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 332. 1905 ≡Hieracium wimmeri
164 Greuter & Raab-Straube: Euro+Med Notulae, 3

subsp. wimmerioides (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss.
40: 651. 1906 ≡Hieracium wimmerioides (Zahn) Prain, Index Kew., Suppl. 4: 122. 1913 ≡Hie-
racium epimedium subsp. wimmerioides (Zahn) Zahn in Schinz & Keller, Fl. Schweiz, ed. 3, 2:
481. 1914.
Hieracium maculatum subsp. arrectariicaule (Sudre) Greuter, comb. nov. ≡Hieracium vulga-
tum prol. arrectariicaule Sudre, Hierac. Centre France: 61. 1902 ≡Hieracium vulgatum subsp.
arrectariicaule (Sudre) Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 287. 1905.
Hieracium maculatum subsp. cuspidatoides (Arv.-Touv.) [O. Bolòs & Vigo, Fl. Països Catal. 3:
1063. 1996, comb. inval., ex] Greuter, comb. & stat. nov. ≡Hieracium cuspidatoides Arv.-
Touv., Hier. Gall. Hisp. Cat.: 345. 1913.
Hieracium maculatum subsp. inquinatum (Boreau) Greuter, comb. & stat. nov. ≡Hieracium
inquinatum Boreau, Fl. Centre France, ed. 3, 2: 404. 1857.
Hieracium maculatum subsp. intersitum (Boreau) Greuter, comb. & stat. nov. ≡Hieracium
intersitum Boreau, Fl. Centre France, ed. 3, 2: 409. 1857.
Hieracium mattirolianum subsp. farinulentiforme (Zahn) Greuter, comb. & stat. nov. ≡Hiera-
cium farinulentiforme Zahn, Hierac. Alp. Mar.: 202. 1916.
Hieracium mattirolianum subsp. martellianum (Zahn) Greuter, comb. nov. ≡Hieracium fari-
nulentiforme subsp. martellianum Zahn in Engler, Pflanzenr. 76: 532. 1921 ≡Hieracium martel-
lianum (Zahn) Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 257. 1985.
Hieracium melanops subsp. mairanum (Belli) Greuter, comb. & stat. nov. ≡Hieracium maira-
num Belli in Fiori & Paoletti, Fl. Italia 3: 491. 1904 ≡Hieracium ferrarii Zahn in Engler,
Pflanzenr. 76: 522. 1921, nom. illeg.
Hieracium murorum subsp. abastumanense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
abastumanense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 511.
1959.
Hieracium murorum subsp. adenoactis (Üksip) Greuter, comb. & stat. nov. ≡Hieracium ade-
noactis Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 513. 1959.
Hieracium murorum subsp. atrovirens (Froel.) Raimondo & Di Grist., comb. & stat. nov. ≡
Hieracium atrovirens Froel. in Candolle, Prodr. 7: 231. 1838. – Lectotype (or neotype?, desig-
nated here): Specimen with two labels, (1) “3 Hieracium atrovirens Dec. pr. 7. p. 231. n. 121.
Guss. Syn. 2. p. 403. Julio Augusto [18]26”, and (2) “Hieracium murorum, non quod[?] calycis
foliola interna acuta vel acuminata. Luglio – Madonie al vallone dell’acqua del Canale“,
[Gussone] (NAP, Herb. Gussonei Siculum).
We consider this subspecies to be endemic to the Madonie Mts in Sicily. Use of the
epithet atrovirens for non-Sicilian plants is due to error. Detailed considerations on
the morphology and distribution of the taxon, and the rationale for our choice of
type, will be provided in full in a separate, forthcoming paper.
F. M. Raimondo & E. Di Gristina
Hieracium murorum subsp. gentiliforme (Zahn) Greuter, comb. nov. ≡Hieracium sylvaticum
subsp. gentiliforme Zahn in Ann. K.K. Naturhist. Hofmus. 23: 200. 1909 ≡Hieracium gentili-
forme (Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33: 431. 1975.
Hieracium murorum subsp. kreczetoviczii (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
kreczetoviczii Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 512. 1959.
Hieracium murorum subsp. pallescentifrons (K. Malý & Zahn) Greuter, comb. & stat. nov. ≡
Hieracium pallescentifrons K. Malý & Zahn in Glasn. Zemaljsk. Muz. Bosni Hercegovini 37:
55. 1925.
Willdenowia 37 – 2007 165

Hieracium murorum subsp. samnaunicum (Zahn) Greuter, comb. nov. ≡Hieracium atratum
subsp. samnaunicum Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1848. 1901 ≡Hieracium
samnaunicum (Zahn) Prain, Index Kew., Suppl. 4: 120. 1913.
Hieracium murorum subsp. sylvivagum (Boreau) Greuter, comb. & stat. nov. ≡Hieracium syl-
vivagum Boreau, Fl. Centre France, ed. 3, 2: 417. 1857.
Hieracium neomarginatum P. D. Sell, nom. nov. ≡Hieracium marginatum P. D. Sell & C. West
in Watsonia 6: 304. 1967 [non Froel. 1838].
Hieracium obscuratum subsp. nigratum (Murr & Zahn) Greuter, comb. nov. ≡Hieracium ery-
thropodum subsp. nigratum Murr & Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten
Naturwiss. 40: 644. 1906 ≡Hieracium nigratum (Murr & Zahn) Prain, Index Kew., Suppl. 4:
116. 1913 ≡Hieracium haematopodum subsp. nigratum (Murr & Zahn) Zahn in Schinz & Keller,
Fl. Schweiz, ed. 3, 2: 483. 1914.
Hieracium obscuratum subsp. pedatifolium (Omang) Greuter, comb. nov. ≡Hieracium peda-
tifolium Omang in Nyt Mag. Naturvidensk. 48: 209. 1910 ≡Hieracium haematopodum subsp.
pedatifolium (Omang) Zahn in Engler, Pflanzenr. 77: 799. 1921.
Hieracium oroglaucum subsp. stenoglossophyllum (O. Behr & al.) Greuter, comb. & stat. nov.
≡Hieracium stenoglossophyllum O. Behr & al. in Glasn. Skopsk. Naucn. Drustva 18: 66. 1937.
Hieracium pallescens subsp. anthyllidifolium (Murr) Greuter, comb. nov. ≡Hieracium anthyl-
lidifolium Murr in Österr. Bot. Z. 43: 222. 1893 ≡Hieracium incisum subsp. anthyllidifolium
(Murr) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1799. 1901.
Hieracium pallescens subsp. golesnicae (O. Behr & al.) Greuter, comb. nov. ≡Hieracium inci-
sum subsp. golesnicae O. Behr & al. in Glasn. Skopsk. Naucn. Drustva 18: 58. 1937.
Hieracium pallescens subsp. hittense (Murr) Greuter, comb. nov. ≡Hieracium murrianum subsp.
hittense Murr in Deutsche Bot. Monatsschr. 16: 5. 1898 ≡Hieracium incisum subsp. hittense
(Murr) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1798. 1901 ≡Hieracium hittense (Murr)
Prain, Index Kew., Suppl. 4: 113. 1913.
Hieracium pallescens subsp. humiliforme (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium incisum subsp. humiliforme
Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1801. 1901.
Hieracium pallescens subsp. incisum (Hoppe) Greuter, comb. nov. ≡Hieracium incisum Hoppe
in Sturm, Deutschl. Fl. 10(39): ad t. 14. 1815 ≡Hieracium incisum Hoppe subsp. incisum [per
Zahn in Allg. Bot. Z. Syst. 7: 115. 1901].
Hieracium pallescens subsp. jugicola (Zahn) Greuter, comb. nov. ≡Hieracium maureri subsp.
jugicola Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1836. 1901 ≡Hieracium salevense subsp.
jugicola (Zahn) Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 527.
1906 ≡Hieracium jugicola (Zahn) Prain, Index Kew., Suppl. 4: 114. 1913 ≡Hieracium incisum
subsp. sprecheri Zahn in Engler, Pflanzenr. 76: 491. 1921, nom. illeg. ≡Hieracium arolae subsp.
bahnmaieri Zahn in Engler, Pflanzenr. 79: 1146. 1922, nom. illeg.
Hieracium pallescens subsp. knautiifolium (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium mu-
rorum var. knautiifolium Arv.-Touv. in Ann. Soc. Linn. Lyon, ser. 2, 34: 83. 1888 ≡Hieracium
incisum subsp. knautiifolium (Arv.-Touv.) Zahn in Engler, Pflanzenr. 76: 505. 1921.
Hieracium pallescens subsp. leucobasis (Zahn) Greuter, comb. nov. ≡Hieracium leucobasis
(Zahn) Prain, Index Kew., Suppl. 4: 115. 1913 ≡Hieracium incisum subsp. leucobasis Zahn in
Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 435. 1906.
Hieracium pallescens subsp. macranthoides (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. macranthoides Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1800. 1901.
166 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium pallescens subsp. maledictum (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium aphyllum subsp. maledictum
Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 466. 1906 ≡Hieracium
maledictum (Zahn) Prain, Index Kew., Suppl. 4: 116. 1913 ≡Hieracium incisum subsp. male-
dictum (Zahn) Zahn in Engler, Pflanzenr. 76: 500. 1921.
Hieracium pallescens subsp. melanocinereum (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. melanocinereum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
437. 1906 ≡Hieracium melanocinereum (Zahn) Prain, Index Kew., Suppl. 4: 116. 1913.
Hieracium pallescens subsp. montafonense (Zahn) Greuter, comb. nov. ≡Hieracium incisum
var. montafonense Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1798. 1901 ≡Hieracium inci-
sum subsp. montafonense (Zahn) Zahn, Hieracioth. Eur. 5: 18. 1910.
Hieracium pallescens subsp. muroriforme (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium incisum subsp. muroriforme
Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1801. 1901 ≡Hieracium muroriforme (Zahn)
Prain, Index Kew., Suppl. 4: 116. 1913.
Hieracium pallescens subsp. murrianum (Murr) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium murrianum Murr in Programm
Realschule Innsbruck 1891: 55. 1891 ≡Hieracium incisum subsp. murrianum (Murr) Zahn in
Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1798. 1901.
Hieracium pallescens subsp. neilreichii (Beck) [Hub.-Mor. in Bauhinia 4: 152. 1971, comb.
inval., ex] Greuter, comb. nov. ≡Hieracium neilreichii Beck, Fl. Hernstein: 435. 1884 ≡Hiera-
cium incisum subsp. neilreichii (Beck) Zahn in Engler, Pflanzenr. 76: 499. 1921.
Hieracium pallescens subsp. niphogeton (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. niphogeton Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 294. 1905 ≡Hieracium nipho-
geton (Zahn) Prain, Index Kew., Suppl. 4: 117. 1913.
Hieracium pallescens subsp. ovale (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges. 81:
522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium incisum subsp. ovale Zahn in
Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1800. 1901.
Hieracium pallescens subsp. oxyodontiforme (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. oxyodontiforme Zahn in Engler, Pflanzenr. 76: 497. 1921.
Hieracium pallescens subsp. pachysphaericum (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. pachysphaericum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
433. 1906 ≡Hieracium pachysphaericum (Zahn) Prain, Index Kew., Suppl. 4: 117. 1913.
Hieracium pallescens subsp. pertenuatum (Murr & Zahn) Greuter, comb. nov. ≡Hieracium
incisum subsp. pertenuatum Murr & Zahn in Engler, Pflanzenr. 76: 498. 1921 ≡Hieracium in-
cisum subsp. longisubulatum Murr & Zahn in Engler, Pflanzenr. 79: 1145. 1922, nom. illeg.
Hieracium pallescens subsp. plazenicense (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. plazenicense Zahn in Österr. Bot. Z. 56: 275. 1906.
Hieracium pallescens subsp. porrectiforme (Murr) Greuter, comb. nov. ≡Hieracium trachse-
lianum subsp. porrectiforme Murr in Allg. Bot. Z. Syst. 10: 100. 1904 ≡Hieracium incisum
subsp. porrectiforme (Murr) Murr in Reichenbach, Icon. Fl. Germ. Helv. 19(2): 46. 1905 ≡
Hieracium porrectiforme (Murr) Prain, Index Kew., Suppl. 5: 131. 1921.
Hieracium pallescens subsp. pseudohittense (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium incisum subsp. pseu-
dohittense Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 434. 1906 ≡
Hieracium pseudohittense (Zahn) Prain, Index Kew., Suppl. 4: 119. 1913.
Willdenowia 37 – 2007 167

Hieracium pallescens subsp. pseudomurrianum (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium pseudomurrianum (Zahn)
Prain, Index Kew., Suppl. 4: 119. 1913 ≡Hieracium incisum subsp. pseudomurrianum Zahn in
Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1798. 1901.
Hieracium pallescens subsp. pseudotrachselianum (Zahn) Greuter, comb. nov. ≡Hieracium in-
cisum subsp. pseudotrachselianum Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1801. 1901.
Hieracium pallescens subsp. ranisavae (Rohlena & Zahn) Greuter, comb. nov. ≡Hieracium
neilreichii subsp. ranisavae Rohlena & Zahn in Repert. Spec. Nov. Regni Veg. 6: 227. 1909 ≡
Hieracium incisum subsp. ranisavae (Rohlena & Zahn) Zahn in Engler, Pflanzenr. 76: 500. 1921.
Hieracium pallescens subsp. subgaudinii (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf. Ges.
81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium incisum subsp. subgaudinii
Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 294. 1905 ≡Hieracium subgaudinii (Zahn) Prain,
Index Kew., Suppl. 4: 121. 1913.
Hieracium pallescens subsp. suboreites (Murr & Zahn) Greuter, comb. nov. ≡Hieracium mur-
rianum subsp. suboreites Murr & Zahn in Allg. Bot. Z. Syst. 13: 115. 1907 ≡Hieracium incisum
subsp. suboreites (Murr & Zahn) Zahn in Engler, Pflanzenr. 76: 501. 1921.
Hieracium pallescens subsp. subpatulum (Zahn) Greuter, comb. & stat. nov. ≡Hieracium sub-
patulum Zahn in Engler, Pflanzenr. 75: 110. 1921.
Hieracium pallescens subsp. tephrochlorum (Zahn) Greuter, comb. nov. ≡Hieracium incisum
subsp. tephrochlorum Zahn in Allg. Bot. Z. Syst. 7: 115. 1901.
Hieracium pallescens subsp. trachselianoides (Zahn) [Hirschm. in Denkschr. Schweiz. Naturf.
Ges. 81: 522. 1956, comb. inval., ex] Greuter, comb. nov. ≡Hieracium incisum subsp. trachse-
lianoides Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 293-294. 1905 ≡Hieracium trachselia-
noides (Zahn) Prain, Index Kew., Suppl. 4: 122. 1913.
Hieracium pallescens subsp. variidens (Murr & Zahn) Greuter, comb. nov. ≡Hieracium inci-
sum subsp. variidens Murr & Zahn in Engler, Pflanzenr. 76: 498. 1921.
Hieracium pallescens subsp. villosipes (Pax) Greuter, comb. nov. ≡Hieracium villosipes Pax in
Österr. Bot. Z. 45: 42. 1895 ≡Hieracium incisum subsp. villosipes (Pax) Zahn in Ascherson &
Graebner, Syn. Mitteleur. Fl. 12(2): 736. 1935.
Hieracium pallescens subsp. visocicense (Degen & Zahn) Greuter, comb. nov. ≡Hieracium
neilreichii subsp. visocicense Degen & Zahn in Magyar Bot. Lapok 6: 215. 1907 ≡Hieracium
incisum subsp. visocicense (Degen & Zahn) Zahn in Engler, Pflanzenr. 76: 499. 1921.
Hieracium pannosum subsp. amphithales (K. Malý & Zahn) Greuter, comb. & stat. nov. ≡Hie-
racium amphithales K. Malý & Zahn in Repert. Spec. Nov. Regni Veg. 38: 298. 1935.
Hieracium pannosum subsp. epiglossophyllum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hie-
racium epiglossophyllum Arv.-Touv. in Bull. Soc. Sci. Nancy, ser. 3, 9: 381. 1908.
Hieracium pannosum subsp. strigulosum (Post) Greuter, comb. & stat. nov. ≡Hieracium stri-
gulosum Post in Bull. Herb. Boissier 1: 25. 1893.
Hieracium pannosum subsp. taygeteum (Boiss. & Heldr.) Greuter, comb. & stat. nov. ≡Hiera-
cium taygeteum Boiss. & Heldr. in Boissier, Diagn. Pl. Orient. 7: 15. 1846.
Hieracium patens subsp. costae (Scheele) Greuter, comb. nov. ≡Hieracium costae Scheele in
Linnaea 32: 686. 1863 ≡Hieracium rectum subsp. costae (Scheele) Zahn in Engler, Pflanzenr.
79: 998. 1922.
168 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium patens subsp. dipsacifolium (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium dipsa-
cifolium Arv.-Touv., Spicil. Rar. Hierac.: 49. 1886 ≡Hieracium rectum subsp. dipsacifolium
(Arv.-Touv.) Zahn in Engler, Pflanzenr. 79: 999. 1922.
Hieracium patens subsp. eurypus (K. Knaf) Greuter, comb. nov. ≡Hieracium eurypus K. Knaf
in Österr. Bot. Z. 25: 181. 1875 ≡Hieracium rectum subsp. eurypus (K. Knaf) Zahn in Engler,
Pflanzenr. 79: 1001. 1922.
Hieracium patens subsp. eynense (Sudre) Greuter, comb. nov. ≡Hieracium eynense Sudre in
Bull. Acad. Int. Géogr. Bot. 22: 57. 1912 ≡Hieracium rectum subsp. eynense (Sudre) Zahn in
Engler, Pflanzenr. 79: 999. 1922.
Hieracium patens subsp. hecatadenum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hieracium
hecatadenum Arv.-Touv. & Gaut., Hieracioth. 5: [in schedis] Hisp. No. 61. 1898 ≡Hieracium
rectum subsp. hecatadenum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 79: 999. 1922.
Hieracium patens subsp. myriophyllum (Scheele) Greuter, comb. nov. ≡Hieracium myriophyl-
lum Scheele in Linnaea 32: 660. 1863 ≡Hieracium rectum subsp. myriophyllum (Scheele) Zahn
in Engler, Pflanzenr. 79: 1000. 1922.
Hieracium patens subsp. neodipsaceum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium neodip-
saceum Arv.-Touv. in Bull. Herb. Boissier 5: 729. 1897 ≡Hieracium rectum subsp. neodip-
saceum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 79: 999. 1922.
Hieracium patens subsp. pseuderiophorum (Loret & Timb.-Lagr.) Greuter, comb. nov. ≡
Hieracium pseuderiophorum Loret & Timb.-Lagr. in Bull. Soc. Bot. France 5: 616. 1858 ≡Hie-
racium rectum Arv.-Touv. & Gaut. in Bull. Soc. Bot. France 51: lxxxvii. 1905, nom. illeg. ≡Hie-
racium rectum subsp. pseuderiophorum (Loret & Timb.-Lagr.) Zahn in Engler, Pflanzenr. 79:
1000. 1922.
Hieracium pictum subsp. irginianum (Arrigoni) Greuter, comb. & stat. nov. ≡Hieracium irgi-
nianum Arrigoni in Boll. Soc. Sarda Sci. Nat., 24: 236. 1985.
Hieracium pietroszense subsp. deylii (Mráz) Greuter, comb. & stat. nov. ≡Hieracium deylii
Mráz in Folia Geobot. 38: 311. 2003.
Hieracium piliferum subsp. fuliginatum (Murr & Zahn) Greuter, comb. nov. ≡Hieracium glan-
duliferum subsp. fuliginatum Murr & Zahn in Reichenbach, Icon. Fl. Germ. Helv. 19(2): 10.
1904 ≡Hieracium fuliginatum (Murr & Zahn) Fiori, Nuova Fl. Italia 2: 891. 1928.
Hieracium pilosum subsp. canalense (Pacher) Greuter, comb. & stat. nov. ≡Hieracium cana-
lense Pacher in Jahrb. Naturhist. Landesmus. Kärnten 22: 94. 1893.
Hieracium pilosum subsp. faucillense (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium villo-
siceps subsp. faucillense Nägeli & Peter, Hierac. Mitt.-Eur. 2: 109. 1886 ≡Hieracium morisia-
num subsp. faucillense (Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 87. 1921.
Hieracium pilosum subsp. favrei (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium villosiceps
subsp. favrei Nägeli & Peter, Hierac. Mitt.-Eur. 2: 112. 1886 ≡Hieracium villosiceps subsp. ca-
tognense Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 352. 1906,
nom. illeg. ≡Hieracium catognense Prain, Index Kew., Suppl. 4: 110. 1913 ≡Hieracium favrei
(Nägeli & Peter) Prain, Index Kew., Suppl. 4: 112. 1913 ≡Hieracium morisianum subsp. favrei
(Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 87. 1921.
Hieracium pilosum subsp. schleicheri (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium vil-
losiceps subsp. schleicheri Nägeli & Peter, Hierac. Mitt.-Eur. 2: 108. 1886 ≡Hieracium schlei-
cheri (Nägeli & Peter) Prain, Index Kew., Suppl. 4: 120. 1913 [non Arv.-Touv. 1897] ≡
Hieracium morisianum subsp. schleicheri (Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 86.
1921.
Willdenowia 37 – 2007 169

Hieracium pilosum subsp. villosifolium (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium villo-
siceps subsp. villosifolium Nägeli & Peter, Hierac. Mitt.-Eur. 2: 108. 1886 ≡Hieracium villosifo-
lium (Nägeli & Peter) Prain, Index Kew., Suppl. 4: 122. 1913 ≡Hieracium morisianum subsp.
villosifolium (Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 86. 1921.
Hieracium praecurrens subsp. pelesii (Grecescu) Greuter, comb. & stat. nov. ≡Hieracium
pelesii Grecescu, Consp. Fl. Roman.: 373. 1898.
Hieracium prenanthoides subsp. brittatense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
brittatense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 480. 1959.
Hieracium prenanthoides subsp. buschianum (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
buschianum Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 479. 1959.
Hieracium prenanthoides subsp. gredense (Rouy) Greuter, comb. & stat. nov. ≡Hieracium
gredense Rouy, Ill. Pl. Eur.: 121. 1901.
Hieracium prenanthoides subsp. loriense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
loriense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 479. 1959.
Hieracium pseudalpinum subsp. cochlearioides Greuter, nom. nov. ≡Hieracium cochleari-
folium Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1842. 1901 [non Brenner 1894] ≡Hie-
racium cochlearioides Zahn in Engler, Pflanzenr. 75: 141. 1921, nom. illeg.
Hieracium pseudalpinum subsp. pseudoglanduliferum (Zahn) Greuter, comb. nov. ≡Hiera-
cium cochlearifolium subsp. pseudoglanduliferum Zahn in Neue Denkschr. Allg. Schweiz. Ges.
Gesammten Naturwiss. 40: 540. 1906 ≡Hieracium pseudoglanduliferum (Zahn) Prain, Index
Kew., Suppl. 4: 119. 1913 ≡Hieracium cochlearioides subsp. pseudoglanduliferum (Zahn) Zahn
in Engler, Pflanzenr. 75: 141. 1921.
Hieracium pseudalpinum subsp. tissierei (Zahn) Greuter, comb. nov. ≡Hieracium cochlea-
rifolium subsp. tissierei Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40:
540. 1906 ≡Hieracium tissierei (Zahn) Prain, Index Kew., Suppl. 4: 122. 1913 ≡Hieracium coch-
learioides subsp. tissierei (Zahn) Zahn in Engler, Pflanzenr. 75: 141. 1921.
Hieracium pseudocerinthe subsp. arvasicum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium
arvasicum Arv.-Touv. in Annuaire Conserv. Jard. Bot. Genève 1: 99. 1897 ≡Hieracium ucenicum
subsp. arvasicum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 210. 1921.
Hieracium pseudocerinthe subsp. rimigenum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium
rimigenum Arv.-Touv., Hier. Gall. Hisp. Cat.: 130. 1913 ≡Hieracium ucenicum subsp. rimi-
genum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 211. 1921.
Hieracium pseudocerinthe subsp. scapiflorum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium
scapiflorum Arv.-Touv., Hier. Gall. Hisp. Cat.: 102. 1913 ≡Hieracium ucenicum subsp. scapi-
florum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 211. 1921.
Hieracium pseudocerinthe subsp. ucenicum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium saxa-
tile var. ucenicum Arv.-Touv. in Ann. Soc. Linn. Lyon, ser. 2, 34: 52. 1888 ≡Hieracium ucenicum
(Arv.-Touv.) Arv.-Touv., Hier. Gall. Hisp. Cat.: 108. 1913 ≡Hieracium ucenicum (Arv.-Touv.)
Arv.-Touv. subsp. ucenicum [per Zahn in Engler, Pflanzenr. 75: 210. 1921].
Hieracium pseudopaltinae subsp. stenobracteophorum (Nyár. & Zahn) Greuter, comb. & stat.
nov. ≡Hieracium stenobracteophorum Nyár. & Zahn in Bul. Grbd. Bot. Univ. Cluj 8: 81. 1929.
Hieracium pseudosvaneticum subsp. kharthlicum (Kem.-Nath.) Greuter, comb. & stat. nov. ≡
Hieracium kharthlicum Kem.-Nath. in Zametki Sist. Geogr. Rast. 17: 126. 1953.
170 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium pseudosvaneticum subsp. subbakurianense (Üksip) Greuter, comb. & stat. nov. ≡
Hieracium subbakurianense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR
19: 469. 1959.
Hieracium pseudosvaneticum subsp. tschkhubianischwilii (Kem.-Nath.) Greuter, comb. &
stat. nov. ≡Hieracium tschkhubianischwilii Kem.-Nath. in Zametki Sist. Geogr. Rast. 17: 127.
1953.
Hieracium pulmonarioides subsp. belveriense (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡
Hieracium belveriense Arv.-Touv. & Gaut., Hieracioth. 12: [in schedis] Hisp. No. 156-158. 1901
≡Hieracium amplexicaule subsp. belveriense (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr.
77: 727. 1921.
Hieracium pulmonarioides subsp. chenevardianum (Zahn) Greuter, comb. nov. ≡Hieracium
amplexicaule subsp. chenevardianum Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 318. 1905 ≡
Hieracium chenevardianum (Zahn) Prain, Index Kew., Suppl. 4: 110. 1913.
Hieracium pulmonarioides subsp. spelaeum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium am-
plexicaule var. spelaeum Arv.-Touv. in Ann. Soc. Linn. Lyon, ser. 2, 34: 50. 1888 ≡Hieracium
amplexicaule subsp. spelaeum (Arv.-Touv.) Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 318.
1905.
Hieracium purpurascens subsp. cabreranum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium
cabreranum Arv.-Touv., Hier. Gall. Hisp. Cat.: 168. 1913 ≡Hieracium tephrocerinthe subsp.
cabreranum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 157. 1921.
Hieracium racemosum subsp. ageratoides (Fr.) Greuter, comb. & stat. nov. ≡Hieracium age-
ratoides Fr. in Uppsala Univ. Årsskr. 1862: 108. 1862.
Hieracium racemosum subsp. barbicaule (Celak.) Greuter, comb. & stat. nov. ≡Hieracium
barbicaule Celak., Prodr. Fl. Böhmen: 789.1881.
Hieracium racemosum subsp. limbarae (Arrigoni) Greuter, comb. & stat. nov. ≡Hieracium
limbarae Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 241. 1985.
Hieracium racemosum subsp. oliastrae (Arrigoni) Greuter, comb. & stat. nov. ≡Hieracium
oliastrae Arrigoni in Boll. Soc. Sarda Sci. Nat. 24: 245. 1985.
Hieracium racemosum subsp. pignattianum (Raimondo & Di Grist.) Greuter, comb. & stat.
nov. ≡Hieracium pignattianum Raimondo & Di Grist. in Bocconea 17: 314. 2004.
Hieracium racemosum subsp. subartvinense (Üksip) Greuter, comb. & stat. nov. ≡Hieracium
subartvinense Üksip in Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 19: 468. 1959.
Hieracium raddeanum subsp. hryniawiense (Wo5.) Greuter, comb. nov. ≡Hieracium hrynia-
wiense Wo5. in Spraw. Komis. Fizjogr. 22: 204. 1888 ≡Hieracium aestivum subsp. hryniawiense
(Wo5.) Zahn in Engler, Pflanzenr. 79: 925. 1922.
Hieracium raddeanum subsp. regelianum (Zahn) Greuter, comb. & stat. nov. ≡Hieracium re-
gelianum Zahn in Engler, Pflanzenr. 79: 936. 1922.
Hieracium ramondii subsp. asturicum (Pau) Greuter, comb. & stat. nov. ≡Hieracium asturi-
cum Pau, Not. Bot. Fl. Españ. 6: 73. 1895.
Hieracium riphaeoides [Bornm. & Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3):
401. 1937, descr. germ.] Bornm. & Zahn, sp. nov. – Holotype: Czech Republic: “Riesengebirge
[Krkonoše], im Langen Grund [Dlouhy dul] (bei St. Peter)”, 1100 m, 13.8.1930, Bornmüller
(B 100216197). – Description: “Hieracio riphaeo simile, sed planta 60-80 cm alta, basin versus
gradatim hirtior, corymbosum; rami 5-10 cm longi; pedunculi elongati, graciles, floccosi tantum;
capitula 10-20. Involucrum magnum, minute glandulosum, pilis destitutum, parcifloccosum;
Willdenowia 37 – 2007 171

squamae latae; bracteae plures. Stylus luteus. Folia 40-60 densa, late lanceolata, summa tantum
diminuta, margine (pilis saepe truncatis) et dorso tantum breviter pilosa (pilis nodulis insiden-
tibus) et floccosa; inferiora 10 cm longa et 15-20 mm lata; superiora minus pilosa sed utrinque
floccosa; omnia dentibus 1-2(-3) dissitis longis angustis acutissimis serrata.” – H. Zahn.
Hieracium robertsii [P. D. Sell in Sell & Murrell, Fl. Great Britain & Ireland 4: 550. 2006, nom.
inval., ex] P. D. Sell, nom. nov. ≡Hieracium rubiginosum var. glabrescens Pugsley in J. Linn.
Soc., Bot. 54: 182. 1948 [non Hieracium glabrescens (F. W. Schultz) Murr 1897].
Hieracium rupicola subsp. marsillyanum (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium mar-
sillyanum Arv.-Touv., Hier. Gall. Hisp. Cat.: 109. 1913 ≡Hieracium leptocladum subsp.mar
-
sillyanum (Arv.-Touv.) Zahn in Engler, Pflanzenr. 77: 732. 1921.
Hieracium rupicola subsp. squarrosulum (Zahn) Greuter, comb. & stat. nov. ≡Hieracium squar-
rosulum Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 319. 1905 ≡Hieracium subsquarrosulum
Zahn in Engler, Pflanzenr. 77: 621. 1921, nom. illeg.
Hieracium sabaudum subsp. gutierrezii (Gutiérrez) Greuter, comb. & stat. nov. ≡Hieracium
gutierrezii Gutiérrez, Apuntes Fl. Olmedo: 125. 1908.
Hieracium sabaudum subsp. hostianum (Zahn) Greuter, comb. nov. ≡Hieracium platyphyllum
subsp. hostianum Zahn in Engler, Pflanzenr. 79: 1008. 1922.
Hieracium sabaudum subsp. macrodon (Sudre) Greuter, comb. & stat. nov. ≡Hieracium dis-
palatum var. macrodon Sudre, Hierac. Centre France: 24. 1902 [non Hieracium macrodon
Nägeli & Peter 1886] ≡Hieracium prominentidens P. D. Sell in Sell & Murrell, Fl. Great Britain
& Ireland 4: 532. 2006.
Hieracium sabaudum subsp. maranzae (Murr & Zahn) Greuter, comb. nov. ≡Hieracium pseu-
doboreale subsp. maranzae Murr & Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1929. 1902 ≡
Hieracium maranzae (Murr & Zahn) Prain, Index Kew., Suppl. 5: 130. 1921 ≡Hieracium
platyphyllum subsp. maranzae (Murr & Zahn) Zahn in Engler, Pflanzenr. 79: 1008. 1922.
Hieracium sabaudum subsp. propinquum (Sudre) Greuter, comb. & stat. nov. ≡Hieracium pro-
pinquum Sudre in Bull. Assoc. Pyrén. Echange Pl. 9: 10. 1900 ≡Hieracium sabaudum var. perpro-
pinquum Zahn in Engler, Pflanzenr. 79: 945. 1922 ≡Hieracium perpropinquum (Zahn) Druce, List
Brit. Pl., ed. 2: 71. 1928.
Hieracium schmidtii subsp. acrocrocydotum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum
subsp. acrocrocydotum Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(2): 265. 1931.
Hieracium schmidtii subsp. argaeum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum subsp.
argaeum Zahn in Engler, Pflanzenr. 75: 223. 1921 ≡Hieracium argaeum (Zahn) P. D. Sell & C.
West in Notes Roy. Bot. Gard. Edinburgh 33: 430. 1975.
Hieracium schmidtii subsp. argyrosericeum (O. Behr & al.) Greuter, comb. nov. ≡Hieracium
pallidum subsp. argyrosericeum O. Behr & al. in Glasn. Skopsk. Naucn. Društva 18: 56. 1937.
Hieracium schmidtii subsp. ateanum (Pau) Greuter, comb. & stat. nov. ≡Hieracium ateanum
Pau in Bol. Soc. Aragonesa Ci. Nat. 17: 153. 1918.
Hieracium schmidtii subsp. basicrinuloides (O. Behr & al.) Greuter, comb. nov. ≡Hieracium
pallidum subsp. basicrinuloides O. Behr & al. in Glasn. Skopsk. Naucn. Drustva 20: 25. 1939.
Hieracium schmidtii subsp. candicans (Tausch) Greuter, comb. nov. ≡Hieracium candicans
Tausch in Flora 11, Ergänzungsbl. 1: 65. 1828 ≡Hieracium pallidum subsp. candicans (Tausch)
Zahn in Engler, Pflanzenr. 75: 220. 1921.
Hieracium schmidtii subsp. canencianum (Pau) Greuter, comb. & stat. nov. ≡Hieracium ca-
nencianum Pau in Bol. Soc. Aragonesa Ci. Nat. 17: 153. 1918.
172 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium schmidtii subsp. creticum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum subsp.
creticum Zahn in Engler, Pflanzenr. 75: 221. 1921.
Hieracium schmidtii subsp. curcicianum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum
subsp. curcicianum Zahn in Engler, Pflanzenr. 75: 227. 1921.
Hieracium schmidtii subsp. dinglerianum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum
subsp. dinglerianum Zahn in Feddes Repert. 24: 380. 1928.
Hieracium schmidtii subsp. giresunense (Hub.-Mor.) Greuter, comb. & stat. nov. ≡Hieracium
giresunense Hub.-Mor. in Notes Roy. Bot. Gard. Edinburgh 33: 245. 1974.
Hieracium schmidtii subsp. huber-morathii (P. D. Sell & C. West) Greuter, comb. & stat. nov.
≡Hieracium huber-morathii P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33: 244.
1974.
Hieracium schmidtii subsp. jovimontis (Zahn) Greuter, comb. nov. ≡Hieracium graniticum
subsp. jovimontis Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1813. 1901 ≡Hieracium palli-
dum subsp. jovimontis (Zahn) Zahn in Engler, Pflanzenr. 75: 224. 1921 ≡Hieracium jovimontis
(Zahn) Roffey in Watson, London Cat. Brit. Pl., ed. 11: 26. 1925.
Hieracium schmidtii subsp. labillardierei (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium la-
billardierei Arv.-Touv. in Annuaire Conserv. Jard. Bot. Genève 1: 101. 1897 ≡Hieracium
pallidum subsp. labillardierei (Arv.-Touv.) Zahn in Engler, Pflanzenr. 75: 222. 1921.
Hieracium schmidtii subsp. lasiochaetum (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
pallidum subsp. lasiochaetum Bornm. & Zahn in Engler, Pflanzenr. 75: 220. 1921 ≡Hieracium
lasiochaetum (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33:
431. 1975.
Hieracium schmidtii subsp. leucothecum (Freyn) Greuter, comb. nov. ≡Hieracium leucothe-
cum Freyn in Österr. Bot. Z. 42: 343. 1892 ≡Hieracium pallidum subsp. leucothecum (Freyn)
Zahn in Engler, Pflanzenr. 75: 217. 1921.
Hieracium schmidtii subsp. libanoticum (Boiss. & Blanche) Greuter, comb. nov. ≡Hieracium
libanoticum Boiss. & Blanche in Boissier, Fl. Orient. 3: 870. 1875 ≡Hieracium pallidum subsp.
libanoticum (Boiss. & Blanche) Zahn in Engler, Pflanzenr. 75: 221. 1921.
Hieracium schmidtii subsp. madoniense (Raimondo & Di Grist.) Greuter, comb. & stat. nov. ≡
Hieracium madoniense Raimondo & Di Grist. in Pl. Biosystems 141: 86. 2007.
Hieracium schmidtii subsp. odontotrichum (Freyn) Greuter, comb. nov. ≡Hieracium odonto-
trichum Freyn in Velenovský, Fl. Bulg.: 342. 1891 ≡Hieracium pallidum subsp. odontotrichum
(Freyn) Zahn in Engler, Pflanzenr. 75: 227. 1921.
Hieracium schmidtii subsp. pseudodontotrichum (Hub.-Mor.) Greuter, comb. & stat. nov. ≡
Hieracium pseudodontotrichum Hub.-Mor. in Notes Roy. Bot. Gard. Edinburgh 33: 246. 1974.
Hieracium schmidtii subsp. rupicolifolium (Zahn) [Gamisans & Jeanm., Fl. Cors.: 839. 2007,
comb. inval., ex] Greuter, comb. nov. ≡Hieracium pallidum subsp. rupicolifolium Zahn in
Engler, Pflanzenr. 82: 1537. 1923.
Hieracium schmidtii subsp. subvandasii (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
pallidum subsp. subvandasii Bornm. & Zahn in Engler, Pflanzenr. 75: 228. 1921 ≡Hieracium
subvandasii (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33: 431.
1975.
Hieracium schmidtii subsp. vranjanum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum subsp.
vranjanum Zahn in Engler, Pflanzenr. 75: 226. 1921.
Willdenowia 37 – 2007 173

Hieracium schmidtii subsp. vulcanicum (Griseb.) Greuter, comb. nov. ≡Hieracium vulcanicum
Griseb. in Bot. Zeitung (Berlin) 8: 638. 1850 ≡Hieracium pallidum subsp. vulcanicum (Griseb.)
Zahn in Engler, Pflanzenr. 75: 220. 1921.
Hieracium sericophyllum subsp. leucocomum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡
Hieracium leucocomum Arv.-Touv. in Bull. Séances Soc. Sci. Nancy, ser. 3, 9: 381. 1908.
Hieracium sericophyllum subsp. pilosius (Buttler) Greuter, comb. & stat. nov. ≡Hieracium
pilosius Buttler in Strid, Mount. Fl. Greece 2: 628. 1991.
Hieracium sericophyllum subsp. tmoleum (Hub.-Mor.) Greuter, comb. & stat. nov. ≡
Hieracium tmoleum Hub.-Mor. in Notes Roy. Bot. Gard. Edinburgh 33: 247. 1974.
Hieracium solidagineum subsp. abadesicola (Pau) Greuter, comb. & stat. nov. ≡Hieracium
abadesicola Pau in Bol. Soc. Esp. Hist. Nat. 18: 506. 1918.
Hieracium sparsiramum subsp. ampezzanum (Nägeli & Peter) Greuter, comb. nov. ≡Hiera-
cium subglaberrimum subsp. ampezzanum (Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 57.
1921 ≡Hieracium glabratum var. ampezzanum Nägeli & Peter, Hierac. Mitt.-Eur. 2: 139. 1886.
Hieracium sparsiramum subsp. subglaberrimum (Nägeli & Peter) Greuter, comb. nov. ≡Hiera-
cium subglaberrimum (Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 56. 1921 ≡Hieracium gla-
bratum subsp. subglaberrimum Nägeli & Peter, Hier. Mitt.-Eur. 2: 146. 1886.
Hieracium sparsum subsp. abietogenum (Szel}g) Greuter, comb. & stat. nov. ≡Hieracium
abietogenum Szel}g in Polish Bot. J. 48: 11. 2003.
Hieracium sparsum subsp. baleanum (Prodán) Greuter, comb. & stat. nov. ≡Hieracium ba-
leanum Prodán in Bul. -ti. Acad. Republ. Populare Romîne, Sect. Biol. -ti.Agric.,Ser.Bot.9:
320. 1957.
Hieracium sparsum subsp. beypazariense (Hub.-Mor.) Greuter, comb. & stat. nov. ≡Hiera-
cium beypazariense Hub.-Mor. in Notes Roy. Bot. Gard. Edinburgh 33: 425. 1975.
Hieracium sparsum subsp. coldei (Szel}g) Greuter, comb. & stat. nov. ≡Hieracium coldei
Szel}g in Ann. Bot. Fenn. 43: 312. 2006.
Hieracium sparsum subsp. longidentatum (Szel}g) Greuter, comb. nov. ≡Hieracium kot-
schyanum subsp. longidentatum Szel}g in Polish Bot. J. 48: 11. 2003 ≡Hieracium mitkae Szel}g
in Polish J. Bot. 51: 120. 2006.
Hieracium sparsum subsp. mirekii (Szel}g) Greuter, comb. & stat. nov. ≡Hieracium mirekii
Szel}g in Polish J. Bot. 51: 118. 2006.
Hieracium sparsum subsp. nigrovirenticeps (Nyár. & Zahn) Greuter, comb. nov. ≡Hieracium
paltinae subsp. nigrovirenticeps Nyár. & Zahn in Bul. Grbd. Bot. Univ. Cluj 8: 77. 1929 ≡
Hieracium paltinae subsp. nigrescenticeps Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl.
12(3): 686. 1938, nom. illeg. ≡Hieracium tomiasiforme Nyár. in Sbvulescu, Fl. Rep. Pop.
Române 10: 517. 1965.
Hieracium sparsum subsp. perfoliosum (Szel}g) Greuter, comb. & stat. nov. ≡Hieracium per-
foliosum Szel}g in Polish Bot. J. 48: 12. 2003.
Hieracium sparsum subsp. pisaturense (Nyár.) Greuter, comb. & stat. nov. ≡Hieracium pisa-
turense Nyár.inBul.Grbd. Bot. Univ. Cluj 8: 149. 1929.
Hieracium sparsum subsp. polyphyllobasis (Nyár. & Zahn) Greuter, comb. nov. ≡Hieracium
paltinae subsp. polyphyllobasis Nyár. & Zahn in Bul. Grbd. Bot. Univ. Cluj 8: 77. 1929 ≡Hie-
racium polyphyllobasis (Nyár. & Zahn) Szel}g in Polish J. Bot. 51: 129. 2006.
174 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium sparsum subsp. sparsiforme (Degen & Zahn) Greuter, comb. nov. ≡Hieracium
sparsiflorum subsp. sparsiforme Degen & Zahn in Magyar Bot. Lapok 5: 79. 1906 ≡Hieracium
sparsum subsp. nomophilum Zahn in Engler, Pflanzenr. 79: 1021. 1922, nom. illeg.
Hieracium sparsum subsp. subnitens (Zahn) Greuter, comb. nov. ≡Hieracium subnitens Zahn
in Magyar Bot. Lapok 10: 173. 1911 ≡Hieracium camkorijense subsp. subnitens (Zahn) Zahn in
Engler, Pflanzenr. 79: 1048. 1922.
Hieracium sparsum subsp. subvillosum (Freyn) Greuter, comb. & stat. nov. ≡Hieracium sub-
villosum Freyn in Velenovský, Fl. Bulg.: 345. 1891.
Hieracium sparsum subsp. telekianum (Boros & Lengyel) Greuter, comb. & stat. nov. ≡Hie-
racium telekianum Boros & Lengyel in Scripta Bot. Mus. Transsilv. 1: 8. 1942.
Hieracium sparsum subsp. tubulatum (Zahn) Greuter, comb. nov. ≡Hieracium sparsiflorum
subsp. tubulatum Zahn in Magyar Bot. Lapok 5: 81. 1906 ≡Hieracium sparsum subsp. tubulare
Zahn in Engler, Pflanzenr. 79: 1022. 1922, nom. illeg.≡Hieracium tubulare Nyár. in Bul. Grbd.
Bot. Univ. Cluj 8: 144. 1929.
Hieracium sparsum subsp. werneri (Szel}g) Greuter, comb. & stat. nov. ≡Hieracium werneri
Szel}g in Polish J. Bot. 51: 25. 2006.
Hieracium stelligerum subsp. girerdii (Gottschl.) Greuter, comb. & stat. nov. ≡Hieracium gi-
rerdii Gottschl. in Monde Pl. 484: 9. 2004.
Hieracium stelligerum subsp. simonkaianum (Zahn) Greuter, comb. nov. ≡Hieracium substel-
latum subsp. simonkaianum Zahn in Engler, Pflanzenr. 75: 284. 1921 ≡Hieracium simonkaia-
num (Zahn) P. D. Sell & C. West in Bot. J. Linn. Soc. 71: 266. 1976.
Hieracium stelligerum subsp. substellatum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hiera-
cium substellatum Arv.-Touv. & Gaut. in Bull. Soc. Bot. France 40: 236. 1894 ≡Hieracium sub-
stellatum Arv.-Touv. & Gaut. subsp. substellatum [per Zahn in Engler, Pflanzenr. 75: 284.
1921].
Hieracium stelligerum subsp. tordanum (Zahn) Greuter, comb. nov. ≡Hieracium pallidum
subsp. tordanum Zahn in Engler, Pflanzenr. 75: 226. 1921 ≡Hieracium substellatum subsp.
tordanum (Zahn) Zahn in Magyar Bot. Lapok 25: 311. 1927.
Hieracium stirovacense subsp. durmitoricum (Rohlena & Zahn) Greuter, comb. nov. ≡Hie-
racium scheppigianum subsp. durmitoricum Rohlena & Zahn in Repert. Spec. Nov. Regni Veg.
6: 232. 1909 ≡Hieracium flexicaule subsp. durmitoricum (Rohlena & Zahn) Zahn in Engler,
Pflanzenr. 77: 583. 1921 ≡Hieracium durmitoricum (Rohlena & Zahn) Niketi6in Bot. Chron.
(Patras) 16: 34. 2003.
Hieracium stirovacense subsp. mirificissimum (Rohlena & Zahn) Greuter, comb. & stat. nov. ≡
Hieracium mirificissimum Rohlena & Zahn in Repert. Spec. Nov. Regni Veg. 6: 240. 1909.
Hieracium stirovacense subsp. pieperi (Zahn) Greuter, comb. nov. ≡Hieracium flexicaule
subsp. pieperi Zahn in Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 102: 359. 1931.
Hieracium thapsiformoides subsp. albopellitum (Zahn) Greuter, comb. nov. ≡Hieracium plu-
mulosiforme subsp. albopellitum Zahn in Engler, Pflanzenr. 77: 599. 1921 ≡Hieracium albo-
pellitum (Zahn) Niketi6in Willdenowia 35: 230. 2005.
Hieracium thapsiformoides subsp. pseudorieni (Zahn) Greuter, comb. & stat. nov. ≡Hiera-
cium pseudorieni Zahn in Engler, Pflanzenr. 77: 598. 1921.
Hieracium tomentosum subsp. gnaphalodes (Arv.-Touv.) Greuter, comb. nov. ≡Hieracium gna-
phalodes Arv.-Touv. in Ann. Soc. Linn. Lyon, ser. 2, 34: 64. 1888 ≡Hieracium subtomentosum
subsp. gnaphalodes (Arv.-Touv.) Zahn in Engler, Pflanzenr. 76: 553. 1921.
Willdenowia 37 – 2007 175

Hieracium transiens subsp. abatanum (Hayek & Zahn) Greuter, comb. nov. ≡Hieracium ery-
throcarpum subsp. abatanum Hayek & Zahn in Engler, Pflanzenr. 79: 1037. 1922.
Hieracium transiens subsp. acrocymum (Zahn) Greuter, comb. nov. ≡Hieracium erythrocarpum
subsp. acrocymum Zahn in Engler, Pflanzenr. 79: 1040. 1922.
Hieracium transiens subsp. aculeatissimum (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. aculeatissimum Zahn in Engler, Pflanzenr. 79: 1045. 1922.
Hieracium transiens subsp. albanico-macedonicum (O. Behr & al.) Greuter, comb. nov. ≡
Hieracium erythrocarpum subsp. albanico-macedonicum O. Behr & al. in Glasn. Skopsk. Naucn.
Društva 20: 33. 1939.
Hieracium transiens subsp. albellipes (Schelk. & Zahn) Greuter, comb. nov. ≡Hieracium ery-
throcarpum subsp. albellipes Schelk. & Zahn in Izv. Kavkaz’sk. Muz. 7: 139. 1913 ≡Hieracium
albellipes (Schelk. & Zahn) Üksip in Komarov, Fl. SSSR 30: 75. 1960 .
Hieracium transiens subsp. amphitephrodes (Sosn. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. amphitephrodes Sosn. & Zahn in Vestn. Tiflissk. Bot. Sada 21: 16. 1912 ≡
Hieracium amphitephrodes (Sosn. & Zahn) Üksip in Komarov, Fl. SSSR 30: 69. 1960.
Hieracium transiens subsp. artabirense (Zahn) Greuter, comb. nov. ≡Hieracium erythrocarpum
subsp. artabirense Zahn in Engler, Pflanzenr. 79: 1038. 1922 ≡Hieracium artabirense (Zahn)
Üksip in Komarov, Fl. SSSR 30: 74. 1960.
Hieracium transiens subsp. asterocymum (Urumov & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. asterocymum Urumov & Zahn in Magyar Bot. Lapok 13: 184. 1915.
Hieracium transiens subsp. austromontenegrinum (O. Behr & al.) Greuter, comb. nov. ≡Hie-
racium erythrocarpum subsp. austromontenegrinum O. Behr & al. in Glasn. Skopsk. Naucn.
Drustva 20: 129. 1939.
Hieracium transiens subsp. autranii (Post) Greuter, comb. & stat. nov. ≡Hieracium autranii
Post in Bull. Herb. Boissier 1: 25. 1893.
Hieracium transiens subsp. balakderense (T. Georgiev & Zahn) Greuter, comb. nov. ≡Hiera-
cium erythrocarpum subsp. balakderense T. Georgiev & Zahn in Izv. Bulg. Bot. Druz. 4: 84.
1931.
Hieracium transiens subsp. brandisianum (Prain) Greuter, comb. nov. ≡Hieracium brandi-
sianum [Zahn in Magyar Bot. Lapok 5: 83. 1906, nom. inval., ex] Prain, Index Kew., Suppl. 4:
109. 1913 ≡Hieracium erythrocarpum subsp. brandisianum (Prain) Zahn in Engler, Pflanzenr.
79: 1042. 1922.
Hieracium transiens subsp. calomeres (Urumov & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. calomeres Urumov & Zahn in Magyar Bot. Lapok 13: 184. 1915.
Hieracium transiens subsp. caloprasinum (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. caloprasinum Zahn in Vestn. Tiflissk. Bot. Sada 12: 20. 1908 ≡Hieracium calo-
prasinum (Zahn) Üksip in Komarov, Fl. SSSR 30: 73. 1960.
Hieracium transiens subsp. ceahlavicum (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. ceahlavicum Zahn in Bul. Grbd. Bot. Univ. Cluj 13: 62. 1933.
Hieracium transiens subsp. cinereisquamum (Nejceff & Zahn) Greuter, comb. nov. ≡Hiera-
cium erythrocarpum subsp. cinereisquamum Nejceff & Zahn in Magyar Bot. Lapok 5: 85. 1906.
Hieracium transiens subsp. cryptonaevum (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. cryptonaevum Bornm. & Zahn in Engler, Pflanzenr. 79: 1042. 1922 ≡
Hieracium cryptonaevum (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edin-
burgh 33: 430. 1975.
176 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium transiens subsp. erythrocarpoides (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. erythrocarpoides Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 249.
1907 ≡Hieracium erythrocarpoides (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 76. 1960.
Hieracium transiens subsp. erythrocarpum (Peter) Greuter, comb. nov. ≡Hieracium erythro-
carpum Peter in Nachr. Königl. Ges. Wiss. Göttingen, Math.-Phys. Kl. 1898: 44. 1898 ≡Hiera-
cium erythrocarpum Peter subsp. erythrocarpum [per Nejceff & Zahn in Magyar Bot. Lapok 5:
84. 1906].
Hieracium transiens subsp. glomerellum (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium ery-
throcarpum subsp. glomerellum Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 249. 1907 ≡
Hieracium glomerellum (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 70. 1960.
Hieracium transiens subsp. heterodontoides (Litv. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. heterodontoides Litv. & Zahn in Repert. Spec. Nov. Regni Veg. 4: 257.
1907 ≡Hieracium heterodontoides (Litv. & Zahn) Üksip in Komarov, Fl. SSSR 30: 72. 1960.
Hieracium transiens subsp. hypopoliocranum (T. Georgiev & Zahn) Greuter, comb. nov. ≡
Hieracium erythrocarpum subsp. hypopoliocranum T. Georgiev & Zahn in Izv. Bulg. Bot. Dru.
4: 84 1931.
Hieracium transiens subsp. incomptum (Nejceff & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. incomptum Nejceff & Zahn in Magyar Bot. Lapok 5: 84. 1906.
Hieracium transiens subsp. insolitum (Zahn) Greuter, comb. nov. ≡Hieracium erythrocarpum
subsp. insolitum Zahn in Vestn. Tiflissk. Bot. Sada 22: 32. 1912 ≡Hieracium insolitum (Zahn)
Üksip in Komarov, Fl. SSSR 30: 73. 1960.
Hieracium transiens subsp. karagoellense (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. karagoellense Zahn in Engler, Pflanzenr. 79: 1042. 1922 ≡Hieracium karagoel-
lense (Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33: 431. 1975.
Hieracium transiens subsp. karakolense (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
leithneri subsp. karakolense Bornm. & Zahn in Repert. Spec. Nov. Regni Veg. Beih. 89: 417.
1944 ≡Hieracium karakolense (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard.
Edinburgh 33: 431. 1975.
Hieracium transiens subsp. kurvalae (Rech. f. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. kurvalae Rech. f. & Zahn in Repert. Spec. Nov. Regni Veg. 38: 348. 1935.
Hieracium transiens subsp. laxifurcans (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. laxifurcans Bornm. & Zahn in Engler, Pflanzenr. 79: 1037. 1922 ≡Hie-
racium laxifurcans (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh
33: 431. 1975.
Hieracium transiens subsp. leilae (Rech. f. & Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. leilae Rech. f. & Zahn in Bot. Jahrb. Syst. 69: 530. 1939.
Hieracium transiens subsp. leptobrachiopsis (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. leptobrachiopsis Zahn in Magyar Bot. Lapok 10: 171. 1906.
Hieracium transiens subsp. liberatidens (T. Georgiev & Zahn) Greuter, comb. nov. ≡Hiera-
cium erythrocarpum subsp. liberatidens T. Georgiev & Zahn in Izv. Bulg. Bot. Dru. 4: 84.
1931.
Hieracium transiens subsp. macrolepidiforme (Zahn) Greuter, comb. nov. ≡Hieracium ery-
throcarpum subsp. macrolepidiforme Zahn in Vestn. Tiflissk. Bot. Sada 21: 8. 1912 ≡Hieracium
macrolepidiforme (Zahn) Üksip in Komarov, Fl. SSSR 30: 69. 1960.
Willdenowia 37 – 2007 177

Hieracium transiens subsp. macrolepioides (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. macrolepioides Zahn in Repert. Spec. Nov. Regni Veg. 4: 258. 1907 ≡Hieracium
macrolepioides (Zahn) Üksip in Komarov, Fl. SSSR 30: 69. 1960.
Hieracium transiens subsp. mallopoderes (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. mallopoderes Bornm. & Zahn in Engler, Pflanzenr. 79: 1043. 1922 ≡Hiera-
cium mallopoderes (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33:
431. 1975.
Hieracium transiens subsp. mallopodioides (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. mallopodioides Zahn in Engler, Pflanzenr. 79: 1040. 1922.
Hieracium transiens subsp. melanocarpum (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. melanocarpum Zahn in Magyar Bot. Lapok 10: 172. 1911.
Hieracium transiens subsp. microtricholepium (T. Georgiev & Zahn) Greuter, comb. nov. ≡
Hieracium erythrocarpum subsp. microtricholepium T. Georgiev & Zahn in Izv. Bulg. Bot.
Dru. 4: 83. 1931.
Hieracium transiens subsp. orthomeres (Urumov & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. orthomeres Urumov & Zahn in Magyar Bot. Lapok 13: 185. 1915.
Hieracium transiens subsp. petrohanicum (Urumov & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. petrohanicum Urumov & Zahn in Allg. Bot. Z. Syst. 13: 57. 1907.
Hieracium transiens subsp. phaedrochlorum (Nyár. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. phaedrochlorum Nyár. & Zahn in Bul. Grbd. Bot. Univ. Cluj 8: 69. 1929.
Hieracium transiens subsp. pollichiotropium (T. Georgiev & Zahn) Greuter, comb. nov. ≡
Hieracium erythrocarpum subsp. pollichiotropium T. Georgiev & Zahn in Izv. Bulg. Bot. Druz.
4: 83. 1931.
Hieracium transiens subsp. pseudogypsophyllum (Nyár. & Zahn) Greuter, comb. nov. ≡Hie-
racium erythrocarpum subsp. pseudogypsophyllum Nyár. & Zahn in Bul. Grbd. Bot. Univ. Cluj
8: 70. 1929.
Hieracium transiens subsp. pycnadenium (Degen & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. pycnadenium Degen & Zahn in Magyar Bot. Lapok 10: 172. 1911 ≡Hiera-
cium erythrocarpum subsp. pseudopycnadenium Zahn in Engler, Pflanzenr. 79: 1036. 1922,
nom. illeg.
Hieracium transiens subsp. samurense (Zahn) Greuter, comb. nov. ≡Hieracium erythrocarpum
subsp. samurense Zahn in Repert. Spec. Nov. Regni Veg. 4: 258. 1907 ≡Hieracium samurense
(Zahn) Üksip in Komarov, Fl. SSSR 30: 68. 1960.
Hieracium transiens subsp. sintenisii (Zahn) Greuter, comb. nov. ≡Hieracium erythrocarpum
subsp. sintenisii Zahn in Engler, Pflanzenr. 79: 1043. 1922 [non Hieracium sintenisii Freyn
1892] ≡Hieracium marmoricola P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh 33:
244. 1974.
Hieracium transiens subsp. stellidorsum (Bornm. & Zahn) Greuter, comb. nov. ≡Hieracium
erythrocarpum subsp. stellidorsum Bornm. & Zahn in Engler, Pflanzenr. 79: 1044. 1922 ≡Hie-
racium stellidorsum (Bornm. & Zahn) P. D. Sell & C. West in Notes Roy. Bot. Gard. Edinburgh
33: 431. 1975.
Hieracium transiens subsp. stribrnyanum (T. Georgiev & Zahn) Greuter, comb. nov. ≡Hie-
racium erythrocarpum subsp. stribrnyanum T. Georgiev & Zahn in Magyar Bot. Lapok 32: 32.
1933.
178 Greuter & Raab-Straube: Euro+Med Notulae, 3

Hieracium transiens subsp. variegatisquamum (Zahn) Greuter, comb. nov. ≡Hieracium ery-
throcarpum subsp. variegatisquamum Zahn in Vestn. Tiflissk. Bot. Sada 22: 31. 1912 ≡Hiera-
cium variegatisquamum (Zahn) Üksip in Komarov, Fl. SSSR 30: 75. 1960.
Hieracium transiens subsp. winklerianum (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. winklerianum Zahn in Bul. Grbd. Bot. Univ. Cluj 13: 63. 1933.
Hieracium transiens subsp. xanthodermum (Zahn) Greuter, comb. nov. ≡Hieracium erythro-
carpum subsp. xanthodermum Zahn in Izv. Bulg. Bot. Dru. 4: 85. 1931.
Hieracium transiens subsp. zygosense (Zahn) Greuter, comb. nov. ≡Hieracium erythrocarpum
subsp. zygosense Zahn in Engler, Pflanzenr. 79: 1043. 1922.
Hieracium triadanum subsp. epinephum (Zahn) Greuter, comb. & stat. nov. ≡Hieracium ra-
cemosum f. epinephum Zahn in Engler, Pflanzenr. 79: 980. 1922 ≡Hieracium epinephum (Zahn)
Zahn in Glasn. Skopsk. Naucn. Drustva 20: 127. 1939 [non Omang 1910] ≡Hieracium orosense
Gottschl. in Willdenowia 35: 233. 2005.
Hieracium triadanum subsp. leucopannosum (O. Behr & al.) Greuter, comb. & stat. nov. ≡
Hieracium leucopannosum O. Behr & al. in Glasn. Skopsk. Naucn. Drustva 20: 127. 1939.
Hieracium umbellatum subsp. elisabethae (Kem.-Nath.) Greuter, comb. & stat. nov. ≡Hiera-
cium elisabethae Kem.-Nath. in Zametki Sist. Geogr. Rast. 17: 128. 1953.
Hieracium umbellatum subsp. kluchoricum (Kem.-Nath.) Greuter, comb. & stat. nov. ≡Hiera-
cium kluchoricum Kem.-Nath. in Zametki Sist. Geogr. Rast. 17: 129. 1953.
Hieracium umbellatum subsp. topaeanum (Prodán) Greuter, comb. & stat. nov. ≡Hieracium
topaeanum Prodán in Bul. -ti. Acad. Republ. Populare Romîne, Sect. Biol. -ti. Agric., Ser. Bot.
9: 322. 1957.
Hieracium umbellatum subsp. turfosum (Kem.-Nath.) Greuter, comb. & stat. nov. ≡Hieracium
turfosum Kem.-Nath. in Zametki Sist. Geogr. Rast. 17: 130. 1953.
Hieracium umbrosum subsp. abietinum (Boiss. & Heldr.) Greuter, comb. nov. ≡Hieracium
abietinum (Boiss. & Heldr.) Boiss., Fl. Orient. 3: 872. 1875 ≡Hieracium vulgatum var. abie-
tinum Boiss. & Heldr. in Boissier, Diagn. Pl. Orient., ser. 2, 3: 105. 1856 ≡Hieracium vulgatum
subsp. abietinum (Boiss. & Heldr.) Fr. in Uppsala Univ. Årsskr. 1862: 98. 1862.
Hieracium umbrosum subsp. oleicolor (Zahn) Greuter, comb. nov. ≡Hieracium integrifolium
[taxon] oleicolor Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss. 40: 640.
1906 ≡Hieracium integrifolium subsp. oleicolor (Zahn) Zahn in Reichenbach, Icon. Fl. Germ.
Helv. 19(2): 246. 1910 ≡Hieracium umbrosum subsp. suboleicolor Zahn in Engler, Pflanzenr.
77: 795. 1921, nom. illeg.
Hieracium umbrosum subsp. porimense (Freyn & Vandas) Greuter, comb. nov. ≡Hieracium al-
binum subsp. porimense Freyn & Vandas in Bull. Herb. Boissier 3: 668. 1895 ≡Hieracium pori-
mense (Freyn & Vandas) T. Durand & B. D. Jackson, Index Kew., Suppl. 1: 210. 1902.
Hieracium umbrosum subsp. tenuissimum (Zahn) Greuter, comb. nov. ≡Hieracium integri-
folium subsp. tenuissimum Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten Naturwiss.
40: 638. 1906 ≡Hieracium tenuissimum (Zahn) Prain, Index Kew., Suppl. 4: 122. 1913 ≡Hie-
racium umbrosum subsp. subtenuissimum Zahn in Engler, Pflanzenr. 77: 792. 1921, nom. illeg.
Hieracium valdepilosum subsp. adenodontum (Zahn) Greuter, comb. nov. ≡Hieracium sub-
elongatum subsp. adenodontum Zahn in Schinz & Keller, Fl. Schweiz, ed. 2, 2: 327. 1905 ≡
Hieracium adenodontum (Zahn) Prain, Index Kew., Suppl. 4: 108. 1913 [non Arv.-Touv. &
Gaut. 1897] ≡Hieracium porrectum subsp. adenodon Zahn in Engler, Pflanzenr. 75: 125. 1921,
nom. illeg. ≡Hieracium porrectum subsp. adenodontum (Zahn) Zahn in Ascherson & Graebner,
Syn. Mitteleur. Fl. 12(2): 191. 1930.
Willdenowia 37 – 2007 179

Hieracium valdepilosum subsp. alfenzinum (Evers) Greuter, comb. nov. ≡Hieracium alfenzinum
Evers in Österr. Bot. Z. 43: 425. 1893 ≡Hieracium subelongatum subsp. alfenzinum (Evers) Zahn
in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1873. 1901 ≡Hieracium porrectum subsp. alfenzinum
(Evers) Zahn in Engler, Pflanzenr. 75: 125. 1921.
Hieracium valdepilosum subsp. elongatiforme (Zahn) Greuter, comb. nov. ≡Hieracium sub-
elongatum subsp. elongatiforme Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3: 1873. 1901 ≡
Hieracium elongatiforme (Zahn) Prain, Index Kew., Suppl. 5: 128. 1921 ≡Hieracium porrectum
subsp. elongatiforme (Zahn) Zahn in Engler, Pflanzenr. 75: 125. 1921.
Hieracium valdepilosum subsp. glabrescens (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
elongatum subsp. glabrescens Nägeli & Peter, Hierac. Mitt.-Eur. 2: 218. 1886 ≡Hieracium val-
depilosum subsp. subglabrescens Zahn, Hierac. Alp. Mar.: 122. 1916, nom. illeg. ≡Hieracium
subglabrescens P. D. Sell & C. West in Bot. J. Linn. Soc. 71: 267. 1976.
Hieracium valdepilosum subsp. porrectum (Fr.) Greuter, comb. nov. ≡Hieracium porrectum
Fr. in Nova Acta Regiae Soc. Sci. Upsal. 14: 106. 1848 ≡Hieracium porrectum Fr. subsp. por-
rectum [per Zahn in Engler, Pflanzenr. 75: 124. 1921].
Hieracium valdepilosum subsp. subalpinum (Nägeli & Peter) Greuter, comb. nov. ≡Hieracium
elongatum subsp. subalpinum Nägeli & Peter, Hierac. Mitt.-Eur. 2: 214. 1886 [non Hieracium
subalpinum Arv.-Touv. 1876] ≡Hieracium valdepilosum subsp. subvaldepilosum Zahn in Koch,
Syn. Deut. Schweiz. Fl., ed. 3: 1867. 1901, nom. illeg. ≡Hieracium subvaldepilosum Prain, In-
dex Kew., Suppl. 4: 121. 1913.
Hieracium valdepilosum subsp. subelongatum (Nägeli & Peter) Greuter, comb. nov. ≡Hiera-
cium subelongatum Nägeli & Peter, Hierac. Mitt.-Eur. 2: 225. 1886 ≡Hieracium porrectum
subsp. subelongatum (Nägeli & Peter) Zahn in Engler, Pflanzenr. 75: 125. 1921.
Hieracium vasconicum subsp. amygdalinum (Arv.-Touv. & Gaut.) Greuter, comb. nov. ≡Hie-
racium amygdalinum Arv.-Touv. & Gaut. in Bull. Soc. Bot. France 41: 371. 1894 ≡Hieracium
laurinum subsp. amygdalinum (Arv.-Touv. & Gaut.) Zahn in Engler, Pflanzenr. 79: 927. 1922.
Hieracium vasconicum subsp. dolosum (Burnat & Gremli) Greuter, comb. nov. ≡Hieracium
dolosum Burnat & Gremli, Cat. Hierac. Alp. Mar.: 41. 1883 ≡Hieracium laurinum subsp.
dolosum (Burnat & Gremli) Zahn in Engler, Pflanzenr. 79: 926. 1922.
Hieracium vasconicum subsp. lactescens (Rouy) Greuter, comb. nov. ≡Hieracium lactescens
Rouy, Fl. France 9: 379. 1905 ≡Hieracium laurinum subsp. lactescens (Rouy) Zahn in Engler,
Pflanzenr. 79: 927. 1922.
Hieracium vasconicum subsp. laureolum (Arv.-Touv.) Greuter, comb. & stat. nov. ≡Hie-
racium laureolum Arv.-Touv., Monogr. Pilosella & Hieracium, Suppl.: 30. 1876 ≡Hieracium
laurinum Arv.-Touv., Monogr. Pilosella & Hieracium, Add.: 18. 1879, nom. illeg.
Hieracium vasconicum subsp. pseudoboreale (Grecescu) Greuter, comb. & stat. nov. ≡Hie-
racium pseudoboreale Grecescu, Consp. Fl. Roman.: 375. 1898.
Hieracium vasconicum subsp. tauryniae (Zahn) Greuter, comb. nov. ≡Hieracium laurinum
subsp. tauryniae Zahn in Engler, Pflanzenr. 79: 928. 1922.
Hieracium vindobonense subsp. pseudobupleuroides (Nägeli & Peter) Greuter, comb. & stat.
nov. ≡Hieracium pseudobupleuroides Nägeli & Peter, Hier. Mitt.-Eur. 2: 74. 1886.
Hieracium vulgatum Fr., Fl. Hall.: 128. 1818-1819. – Neotype (designated here): Sweden:
Blekinge, Spjutsbygd, 28.6.1892, Mellin & Puke in Dahlstedt, Herb. Hierac. Scand. 3, No. 98 (S).
The name Hieracium vulgatum has long haunted botanists and botanical literature,
being used in different senses under differentbotanicaltraditions.Itisclearlyacon
-
fused name, yet no one so far has apparently dared to dispose of it or explicitly re-
180 Greuter & Raab-Straube: Euro+Med Notulae, 3

ject it under ICBN Art. 57. The name H. vulgatum, which has formally to be
handled in the context of “Flora hallandica” where it first appears (although it is
usually cited from the somewhat later publication in the “Novitiae florae
suecicae”), has not so far been formally typified, nor is any original material known.
Whereas Nordic hieraciologists use H. vulgatum in the sense of H. triviale (Norrl.)
Norrl. of the species group of H. levicaule Jord., the Zahn school accepted it for the
complex that, for reasons of priority, has subsequently become known as H.
lachenalii Suter. Zahn (in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(2): 553.
1934), one of the few who was aware of the early “Flora hallandica” usage of the
name, formally equates it with Hieracium lachenalii subsp. cruentifolium (Dahlst.
& Lübeck) Zahn. This is a plausible interpretation, and may perhaps have been
based on the examination of an original specimen, but if so, that specimen is now
obviously lost. The above neotype designation formalises Zahn’s conclusion and at
the same time precludes further use of the name H. vulgatum, whichinanaggregate
sense is junior to H. lachenalii and at the segregate species level clearly cannot be
applied to H. cruentifolium under the already mentioned Art. 57. W. Greuter
Hieracium waldsteinii subsp. brandisii (Freyn) Greuter, comb. & stat. nov. ≡Hieracium bran-
disii Freyn in Bull. Herb. Boissier 3: 646. 1895.
Hieracium waldsteinii subsp. ferdinandi-coburgii (J. Wagner & Zahn) Greuter, comb. & stat.
nov. ≡Hieracium ferdinandi-coburgii J. Wagner & Zahn in Repert. Spec. Nov. Regni Veg. 38:
228. 1935.
Hieracium xanthoprasinophyes [Zahn in Ascherson & Graebner, Syn. Mitteleur. Fl. 12(3): 418.
1939, descr. germ., ex] Gottschl., sp. nov. – Holotype: Austria, “Tyrol, Verwallgruppe, 9026/2:
Galtür, Erlengebüschränder auf Schipisten an der Kopsstraße nahe dem Schilift”, 1700 m,
22.8.1999, Gottschlich 38955 (B; isotypes distributed as Hieracia Europaea Selecta No. 100). –
Diagnosis: Hieracio picroide simile sed differt foliis caulinis inferioribus basi breviter angus-
tatis, superioribus late rotundatis, involucri phyllis pilis glanduliferis minus dense obsitis. – G.
Gottschlich.
Jacobaea cilicia (Boiss.) B. Nord., comb. nov. ≡Senecio cilicius Boiss.,Diagn.Pl.Orient.,ser.
2, 3: 37. 1856.
Jacobaea inops (Boiss. & Balansa) B. Nord., comb. nov. ≡Senecio inops Boiss. & Balansa in
Boissier, Diagn. Pl. Orient., ser. 2, 6: 101. 1859.
Jacobaea erucifolia subsp. praealta (Bertol.) Greuter & B. Nord., comb. & stat. nov. ≡Senecio
praealtus Bertol. in Opusc. Sci. 3: 183. 1819.
Jacobaea lycopifolia (Poir.) Greuter & B. Nord., comb. nov. ≡Senecio lycopifolius Poir. in
Lamarck, Encycl. Suppl. 5: 131. 1817 ≡Senecio erucifolius subsp. lycopifolius (Poir.) Nyman,
Consp. Fl. Eur.: 356. 1879.
Jacobaea mouterdei (Arènes) Greuter & B. Nord., comb. nov. ≡Senecio mouterdei Arènes in
Notul. Syst. (Paris) 14: 196. 1952.
Jacobaea ornata (Druce) Greuter & B. Nord., comb. & stat. nov. ≡Senecio aquaticus var. orna-
tus Druce in Rep. Bot. Soc. Exch. Club Brit. Isles 6: 24. 1921 ≡Senecio aquaticus subsp. ornatus
(Druce) P. D. Sell in Sell & Murrell, Fl. Great Britain & Ireland 4: 556. 2006.
Jacobaea trapezuntina (Boiss.) B. Nord., comb. nov. ≡Senecio trapezuntinus Boiss., Fl. Orient.
3: 393. 1875.
Picris asplenioides subsp. saharae (Coss. & Kralik) Dobignard, comb. nov. ≡Spitzelia saharae
Coss. & Kralik in Bull. Soc. Bot. France 4: 397. 1857 ≡Picris saharae (Coss. & Kralik) Hochr. in
Willdenowia 37 – 2007 181

Annuaire Conserv. Jard. Bot. Genève 7-8: 235. 1904 ≡Picris coronopifolia subsp. saharae
(Coss. & Kralik) Maire in Jahandiez & Maire, Cat. Pl. Maroc: 836. 1934.
Rhodanthemum atlanticum subsp. gelidum (Maire) Dobignard, comb. nov. ≡Leucanthemum
atlanticum var. gelidum Maire in Bull. Soc. Hist. Nat. Afrique N. 14: 153. 1923.
Rudbeckia laciniata L.
NMk: Makedonski Brod - village Suvodol, on the sandy sites, near r. Treska, 470 m, 12.6.
1997, leg. et det. V. Matevski (SKO). V. Matevski
Schlagintweitia Griseb.
Recent work on the phylogeny of Hieracium and allied taxa (Fehrer & al. in Molec.
Phylogen. Evol. 42: 347-361. 2007) demonstrates convincingly that Schlagintweitia
forms a clade of its own, basal to all other representatives of the group, and deserves
recognition as a separate genus – which is corroborated by its morphologic distinc-
tiveness. This genus is not, however, monotypic, as is often assumed. A molecular
study carried out some time ago at the University of Tübingen, in cooperation with
D. Begerow, has demonstrated that the morphologically related H. huteri (better
known under its junior and illegitimate synonym H. pallidiflorum) also belongs
here. The following combinations are therefore necessary.
G. Gottschlich & W. Greuter
Schlagintweitia huteri (Bamb.) Gottschl. & Greuter, comb. nov. ≡Hieracium huteri Bamb. in
Flora 40: 626. 1857 ≡Hieracium pallidiflorum Hausm. in Verh. K.K. Zool.-Bot. Ges. Wien 8:
375. 1858, nom. illeg.
Schlagintweitia huteri subsp. lantoscana (Burnat & Gremli) Gottschl. & Greuter, comb. nov. ≡
Hieracium lantoscanum Burnat & Gremli, Cat. Hierac. Alp. Mar.: 22. 1883 ≡Hieracium palli-
diflorum subsp. lantoscanum (Burnat & Gremli) Zahn in Koch, Syn. Deut. Schweiz. Fl., ed. 3:
1899. 1901.
Schlagintweitia huteri subsp. subpicroides (Zahn) Gottschl. & Greuter, comb. nov. ≡Hieracium
pallidiflorum subsp. subpicroides Zahn in Neue Denkschr. Allg. Schweiz. Ges. Gesammten
Naturwiss. 40: 663. 1906 ≡Hieracium subpicroides (Zahn) Prain, Index Kew., Suppl. 4: 121.
1913.
Senecio breviflorus (Kadereit) Greuter, comb. & stat. nov. ≡Senecio flavus subsp. breviflorus
Kadereit in Bot. Jahrb. Syst. 104: 510. 1984 ≡Senecio mohavensis subsp. breviflorus (Kadereit)
M. Coleman in Edinburgh J. Bot. 58: 392. 2001.
Silybum marianum (L.) Gaertn.
ASr: E Serbia, c. 5 km E of Nis, Gornja Vrezina, Gradac Hill, 43°20'N, 21°59'E, 380 m,
2.10.1999, Niketi6ko19991002/1 (BEO). – This species is native in the Mediterra-
nean region and SW Europe, having been introduced as a medicinal and ornamental
plant elsewhere in Europe, S, W, and Central Asia, California, Australia and New
Zealand. In some of these regions it became an invasive weed. The first specimen to
be collected in Serbia was found growing wild on fallow ground. Significant culti-
vation of Silybum marianum in Serbia only dates back twenty years, and it is possi-
ble that this casual subspontaneous occurrence is a first step toward naturalisation.
M. Niketi6
Taraxacum acrolobum Dahlst. [= T. paradoxum Palmgr.]
+Rf(N): Karelian Republic, Vieljärvi [Vedlozero]: Vallittu, Panfilovin vainion laidalla,
30.5.1943, Railonsala (H 119152, 160391, 160393, 160389); ibid., Otraniemi, nur-
mikolla, 7.6.1944, Railonsala (H 160388); ibid., pientarella, 7.6.1944, Railonsala
182 Greuter & Raab-Straube: Euro+Med Notulae, 3

(H 160398); ibid., aholla kylän itäpuolella, 8.6.1944, Railonsala (H 160397); ibid.,
tien vierellä kylän kaakkoispuolella, 8.6.1944, Railonsala (H 160396); Jyrkilä
[Yurgilitsa], koulun pihassa, 16.6.1944, Railonsala (H 160387); Pihtilahti, koulun
perunamaan pientarella, 05.6.1944, Railonsala (H 160395); ibid., Ivanovin pellon
pientarella, 28.5.1943, Railonsala (H 160390); Kukkajärvi, pientarella kylän poh-
joispuolella, 10. 6.1944, Railonsala (H 160392); ibid., tien vierellä kylän ete-
läpuolella, 11.6.1944, Railonsala (H 160394). – First documented report for the
Karelian Republic. Previously reported for Russia from the biogeographic province
Kol (Såltin in Lounais-Hämeen Luonto 18: 4. 1965), an area that, along with the
Karelian Republic, includes the north-easternmost part of the Leningrad Region,
until the Svir River valley. The voucher specimens were variously identified by A.
Railonsala as Taraxacum falcatum Brenner, T. leptolobum Dahlst. ex Hagl., T.
jaervikylense H. Lindb. and T. paradoxum, then identified as T. paradoxum by A.
Palmgren in 1952. A. N. Sennikov
Taraxacum amplum Markl.
+Rf(N): Karelian Republic, Sortavala: seminariet, 20.6.1938, Lindberg (H 101636, 101637);
Petrozavodsk: Kukkovka, Sortavala str. / Rovio str., ruderal meadow by the build-
ings, 21.5.2002, Timofeeva (PTZ). – First documented report for the Karelian
Republic. The record from Sortavala was included without specification in the biogeo-
graphic province Kl (nowadays: Finland and Karelian Republic) by Marklund (in
Acta Bot. Fenn. 26: 57. 1940). The collection from Petrozavodsk was identified by
Øllgaard and Räsänen, January 2003. A. N. Sennikov, H. Øllgaard & J. Räsänen
Taraxacum ancistrolobum Dahlst.
+Rf(NW): Leningrad Region, Kexholm [Priozersk], gräslinda i staden, 9.6.1910, Marklund (H
102395); ibid., linnan luona, 3.6.1994, Sonck (H 716276). – Never directly reported
for Russia. The record from Priozersk was included in the biogeographic province
Ka (nowadays: Finland and Leningrad Region of Russia), without specification, by
Marklund (in Acta Bot. Fenn. 26: 58. 1940). The Sonck collection was identified by
Øllgaard in 1996. A. N. Sennikov & H. Øllgaard
Taraxacum atrimarginatum H. Lindb.
+Rf(N): Karelian Republic, Impilahti: Kitelä, Syskyänsaari, trädgårdsjord vid klostret, 6.6.
1932, Pettersson (H 102798); ibid., gräsbevuxen mark på klostergården, 6.6.1932,
Pettersson (H 102799); ibid., Jylhävaara, gräsmark i trädgård, 6.6.1932, Pettersson
(H 102800); Sortavala: Kirjavalahti, trädesåker, 30.5.1910, Arvonen (H 102809);
ibid., odlad äng, 28.5.1910, Arvonen (H 102811); ibid., strandäng, 28.5.1910,
Arvonen (H 102812); ibid., Orjatlampi, odlad äng, 31.5.1910, Arvonen (H 102810) &
1.6.1910, Arvonen (H 102813, 102814); Vieljärvi [Vedlozero], Kukkajärvi, koulun
pihassa, 10.6.1944, Railonsala (H 102794); ibid., pientarella kylän pohjoispuolella,
10.6.1944, Railonsala (H 102797); ibid., tien vierellä, Siilakansillan eteläpuolella,
16.6.1944, Railonsala (H 102796); ibid., Kleissoilla, nurmikolla, Siilakanjärven luo-
teispuolella, 3.6.1944, Railonsala (H 102795); Pihtilahti, lehdossa kylän
luoteispuolella, 3.6.1944, Railonsala (H 102793); ibid., nurmikolla Siilakanjoen sil-
lan luona, 1.6.1943, Railonsala (H 102790); ibid., tien vierellä Siilakanjoen lähellä,
6.6.1943, Railonsala (H 102791, 102792). – First documented report for the Karelian
Republic. Previously reported from the biogeographic provinces Kl (Marklund in
Acta Bot. Fenn. 26: 60. 1940) and Kol (Såltin in Lounais-Hämeen Luonto 18: 5.
1965) that included partly Finland, partly the Karelian Republic and north-eastern-
most part of the Leningrad Region, until the Svir River valley. The collections from
Impilahti and Sortavala parishes were identified by A. Palmgren, those from
Vedlozero mostly by A. Railonsala and subsequently verified by Palmgren.
A. N. Sennikov
Willdenowia 37 – 2007 183

Taraxacum concaviformatum Rail.
+Rf(N): Karelian Republic, Kuusamo: Tavajoen S. puolella, lähellä Possosiiranjärven puron
laskusuuta Aittalampeen, mättäisellä rämeletolla, 2.7.1938, Laurila (H 1268653);
Salla: Tuutikylä, kansakoulun edustan hoitamattomalla nurmella, 27.6.1939, Auer
(H 1268644). – Murmansk Region, Salla: Kutsajoki, Lintujärven ja Ylimm. Kursu-
järven välillä, Trollius-niityssä, 30.6.1939, Auer (H 1268655). – New to Russia.
These specimens had first been identified as Taraxacum croceum by G. Marklund in
1942, then as T. concaviformatum by Øllgaard and C. E. Sonck in 1999.
H. Øllgaard & A. N. Sennikov
Taraxacum contractum Markl.
+Rf(N): Karelian Republic, Petrozavodsk: Kukkovka, Sortavala str. / Rovio str., ruderal
meadow, 16.5.2002, Timofeeva (PTZ). – New to Russia. Identifications by Øllgaard
and Räsänen, January 2003. H. Øllgaard, J. Räsänen & A. N. Sennikov
Taraxacum copidophyllum Dahlst.
+Rf(NW): Leningrad Region, Viborg: Trångsund [Vysotsk], vägkant mellan brädgårdarna,
5.6.1932, Marklund (H 106257). – Never directly reported for Russia, and in fact
new to the country. This record was included in the biogeographic province Ka (no-
wadays: Finland and Leningrad Region of Russia), without specification, by
Marklund (in Acta Bot. Fenn. 26: 70. 1940). A. N. Sennikov
Taraxacum cordatum Palmgr.
+Rf(N): Karelian Republic, Petrozavodsk: Kukkovka, Komsomol av. / Rovio str., ruderal
roadside meadow, 16.5.2002, Timofeeva (PTZ); Petrozavodsk: central part, by the
State Archive, ruderal grassland, 15.5.2002, Rudkovskaya 11 (PTZ). – New to the
Karelian Republic. Identification by Øllgaard and Räsänen, January 2003.
H. Øllgaard, J. Räsänen & A. N. Sennikov
Taraxacum cyanolepis Dahlst.
+Rf(NW): Leningrad Region, Viborg: lund vid Pälli sluss, 12.6.1908, Buch (H 107565, 107569);
Pälli, vid gästgifveriet, 12.6.1908, Buch (H 107566, 107568); Pälli, åker strax N om
Kanalchefens bostad, 12.6.1908, Buch (H 107567, 107570). – Never directly reported
for Russia, and in fact new to the country. The record was included in the biogeo-
graphic province Ka (nowadays: Finland and Leningrad Region of Russia), without
specification, by Marklund (in Acta Bot. Fenn. 26: 74. 1940); it was one of the origi-
nal elements of T. alatum H. Lindb. (Meddelanden Soc. Fauna Fl. Fenn. 35: 25.
1909), an identification subsequently corrected by H. Lindberg and H. Såltin.
A. N. Sennikov
Taraxacum ekmanii Dahlst.
+Rf(N): Karelian Republic, Petrozavodsk: Golikovka, between Povenetskaya str. and Merets-
kov str., Lososinskiy park, right river side, grassland, 15.6.2001, Rudkovskaya 1042
(PTZ); central part, Gylling embankment, ruderal grassland, 22.5.2002, Rudkovskaya
23 (PTZ); by the State Archive, ruderal grassland, 15.5.2002, Rudkovskaya 12 (PTZ);
Kukkovka, Komsomol av., green stripe between two road lines, 16.5.2002, Timofeeva
(PTZ); Sortavala str. 8, ruderal grassland by the house, 15.5.2002, Timofeeva (PTZ);
Sortavala str. 10, in asphalt cracks along a roadside, 17.5.2002, Timofeeva (PTZ);
Sortavala str. 12, ruderal grassland by the house, 15.5. & 17.5.2002, Timofeeva
(PTZ); Sortavala str. / Rovio str., ruderal meadow by the buildings, 21.5.2002, Timo-
feeva (PTZ). – New to the Karelian Republic. Identifications by Øllgaard and Rä-
sänen, January 2003, and, for Rudkovskaya 1042, by Reinikka in 2001.
H. Øllgaard, J. Räsänen, E. Reinikka & A. N. Sennikov
184 Greuter & Raab-Straube: Euro+Med Notulae, 3

Taraxacum expansum Florstr. [T. haglundii Markl. ined.]
+Rf(NW): Saint-Petersburg, Terijoki [Zelenogorsk], gräslinda vid stationen, 8.6.1932, Mark-
lund (H 112741, 112742); 10.6.1932, Marklund (H 112738-112740, 112743-112745,
118519). – Never directly reported for Russia, and in fact new to the country. The
unpublished herbarium name T. haglundii was recorded for the biogeographic prov-
ince Ik, without specification (Såltin in Lounais-Hämeen Luonto 18: 9. 1965). The
specimen was identified as T. expansum by Kääntönen in 1971.
M. Kääntönen & A. N. Sennikov
Taraxacum hemicyclum G. E. Haglund
+Rf(N,
NW):
Leningrad Region, Viborg: Rättijärvi, äng Ö om hotellet, 16.6.1908, Buch (H
114814); Trångsund [Vysotsk], gräsmark, 5.6.1932, Marklund (H 811573). – Kare-
lian Republic, Petrozavodsk: Drevlianka, Lososinskoye chaussée / Berezovaya al-
ley, ruderal grassland by the houses, 15.5.2002, Kravchenko 9857 (PTZ); central
part, by the State Archive, in the basement cracks, 15.5.2002, Rudkovskaya 5 (PTZ);
Pribrezhniy park, 17.5.2002, Rudkovskaya 14 (PTZ); Gylling embankment, ruderal
grassland by the public beach, 22.5.2002, Rudkovskaya 25a, 25b, 27a, 27b, 29-31
(PTZ). – New to Russia. The collections from Viborg were identified by H. Såltin in
1966, those from Petrozavodsk by Øllgaard and Räsänen, January 2003.
A. N. Sennikov, H. Øllgaard & J. Räsänen
Taraxacum homoschistum H. Øllg. [= T. hastatum Markl.]
+Rf(NW): Leningrad Region, Viipuri [Viborg]: Saunalahti, Tykistökadun itäpäässä, 10.6.1938,
Erkamo (H 656071); Monrepos, 5.1916, Hintikka (H 114612-114615); äng nära
Monrepos, 1.6.1932, Marklund (H 114610, 114611). – First documented report for
Russia. The collections from Viborg were identified as T. hastatum by G. Marklund
in 1940. This record was included in the biogeographic province Ka (nowadays: Fin-
land and Leningrad Region of Russia), without specification, by Såltin (in
Lounais-Hämeen Luonto 18: 9. 1965). J. Räsänen & A. N. Sennikov
Taraxacum isthmicola H. Lindb.
+Rf(N): Karelian Republic, Sordavala, strand, 5.1910, Marklund (H 115533). – First docu-
mented report for Karelian Republic. The original identification by G. Marklund
was confirmed by H. Lindberg in 1942. A. N.Sennikov
Taraxacum leptophyllum H. Lindb. ex Såltin
+Rf(NW): Leningrad Region, Viborg, gräslinda i Papula-parken, 2.6.1932, Marklund (H
135705, 135707); vägkant vid Havis, 4.6.1932, Marklund (H 135703, 135704). –
First documented report for the Leningrad Region. This record was included in the
biogeographic province Ka (nowadays: Finland and Leningrad Region of Russia),
without specification, by Marklund (in Acta Bot. Fenn. 26: 92. 1940). Identifica-
tions by G. Marklund. A. N. Sennikov
Taraxacum lucescens Dahlst. [= T. lindbergii Markl. ex Puol.]
+Rf(N): Karelian Republic, Sortavala, seminariet, 20.6.1938, Lindberg (H 147163-147165),
22.6.1938, Lindberg (H 147166). – First documented report for the Karelian Repub-
lic. This record was included in the biogeographic province Kl (nowadays: Finland
and Karelian Republic), without specification, by Marklund (in Acta Bot. Fenn. 26:
97. 1940). Identifications by H. Lindberg. A. N. Sennikov
Taraxacum macroceratodon H. Øllg., nom. nov. ≡Taraxacum macroceron Rail. in Aquilo, Ser.
Bot. 14, App.: 30. 1977 [non Taraxacum macroceras Dahlst. 1906]. Following a request for a
binding decision, the Committee for Vascular Plants has, at a large majority, considered T. ma-
croceron and T. macroceras to be confusingly similar names.
Willdenowia 37 – 2007 185

Taraxacum melittostylum H. Øllg., nom. nov. ≡Taraxacum melinostylum G. E. Haglund &
Soest in Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 42: 137. 1969 [non Taraxacum melano-
stylum T. C. E. Fries 1908]. Following a request for a binding decision, the Committee for Vascu-
lar Plants has, at a large majority, considered T. melinostylum and T. melanostylum to be
confusingly similar names. The intended meaning of the former epithet apparently was
honey-styled, for which melittostylum is linguistically more correct.
Taraxacum olitorium G. E. Haglund [T. pallidiflorum Markl. ined.]
+Rf(NW): Leningrad Region, Viborg: Trångsund [Vysotsk], ängsmark nära brädgårdarna, 5.6.
1932, Marklund (H 196945-196952). – New to Russia. The specimen, originally
named T. pallidiflorum, was identified by Øllgaard in 2003.
H. Øllgaard & A. N. Sennikov
Taraxacum pallescens Dahlst.
+Rf(NW): Saint-Petersburg, Terijoki [Zelenogorsk]: Kuokkala [Repino], bangård, 11.6.1935,
Fagerström (H 160106). – New to Russia. Identified by H. Såltin in 1967.
A. N. Sennikov
Taraxacum pallidulum H. Lindb.
–Rf(N): The record of Taraxacum pallidulum for southern Karelia (Cvelev in Fl. Evr. Casti
SSSR 8: 99. 1989) belongs to T. canaliculatum H. Lindb., that Cvelev treated in sy-
nonymy. A. N. Sennikov
Taraxacum pannucium Dahlst.
+Rf(NW): Leningrad Region, Viborg: Trångsund [Vysotsk], brädgård, 5.6.1932, Marklund (H
160440-160443); ibid., Ravansaari, 5.6.1932, Marklund (H 160438); Monrepos,
5.1916, Hintikka (H 160437, 160444); Viborg, gräslinda i parken vid Monrepos,
1.6.1932, Marklund (H 160439, 160445). – First documented report for the Lenin-
grad Region. This record was included in the biogeographic province Ka (nowa-
days: Finland and Leningrad Region of Russia), without specification, by Såltin (in
Lounais-Hämeen Luonto 18: 13. 1965). The specimens were identified as T. picea-
tum Dahlst. by G. Marklund in 1932 or T. sublaciniosum Dahlst. & H. Lindb. by H.
Lindberg and Marklund in 1935 (Lindberg in Acta Bot. Fenn. 17: 22. 1935), then re-
named T. pannucium by G. Haglund in 1952. A. N. Sennikov
Taraxacum polyodon Dahlst.
+Rf(NW): Leningrad Region, Viborg: kulturpåverkad lund vid foten av Papula-backen,
2.6.1932, Marklund (H 161646, 161647). – First documented report for Russia;
identifications by G. Marklund. This record was included in the biogeographic
province Ka (nowadays: Finland and Leningrad Region of Russia), without specifi-
cation, by Såltin (in Lounais-Hämeen Luonto 18: 14. 1965). The voucher specimens
on which the report for “Russia” (Puolanne in Memoranda Soc. Fauna Fl. Fenn. 8:
165. 1933) is based are not known. A. N. Sennikov
Taraxacum pulcherrimum H. Lindb.
+Rf(NW): Leningrad Region, Viborg: Kolikkoinmäki, gräsmark, 2.6.1932, Marklund (H
099978). – First documented report for Russia; identifications by G. Marklund. This
record was included in the biogeographic province Ka (nowadays: Finland and Le-
ningrad Region of Russia), without specification, by Marklund (in Acta Bot. Fenn.
26: 116. 1940). A. N. Sennikov
Taraxacum puolannei Markl. ex Puol.
+Rf(N): Karelian Republic, Petrozavodsk: Kukkovka, Sortavala str. / Rovio str., ruderal mea-
dow by the buildings, 17.5.2002, Timofeeva (PTZ). – New to Russia. Identification by
Øllgaard and Räsänen, January 2003. H. Øllgaard, J. Räsänen & A. N. Sennikov
186 Greuter & Raab-Straube: Euro+Med Notulae, 3

Taraxacum purpureum Raunk. [= T. polychroum Ekman ex M. P. Christ. & Wiinst.]
+Rf(NW): Leningrad Region, Viborg: Trångsund [Vysotsk], vägkant mellan brädgårdarna,
5.6.1932, Marklund (H 161535). – First documented report for Russia; identifica-
tions by G. Marklund. This record was included in the biogeographic province Ka
(nowadays: Finland and Leningrad Region of Russia), without specification, by
Såltin (in Lounais-Hämeen Luonto 18: 14. 1965). A. N. Sennikov
Taraxacum retroflexum H. Lindb.
+Rf(N): Karelian Republic, Petrozavodsk: Kukkovka, Komsomol av. / Rovio str., ruderal
meadow, 16.5.2002, Timofeeva (PTZ); Sortavala str., lime alley, ruderal meadows,
21.5.2002, Timofeeva (PTZ); Sortavala str. / Rovio str., ruderal meadows, 21.5.
2002, Timofeeva (PTZ). – New to the Karelian Republic. Identifications by Øll-
gaard and Räsänen, January 2003. H. Øllgaard, J. Räsänen & A. N. Sennikov
Taraxacum sellandii Dahlst.
+Rf(N): Karelian Republic, Petrozavodsk: central part, Pribrezhniy park, along the alley in
the poplar grove, 15.6.2000, Kravchenko 8872 (PTZ); Pribrezhniy park, 17.5.2002,
Rudkovskaya 15 (PTZ). – New to Russia. Identifications by Øllgaard and Räsänen
in 2002 (Rudkovskaya 15), and Reinikka in 2001 (Kravchenko 8872).
H. Øllgaard, J. Räsänen, E. Reinikka & A. N. Sennikov
Taraxacum semiglobosum H. Lindb. [= T. approximans H. Lindb.]
+Rf(N): Karelian Republic, Petrozavodsk: central part, Pioneers park, ruderal place (for-
merly buildings of 19th century), 22.5.2002, Kravchenko 9872 (PTZ); right river-
side of Lososinka under the bridge, grassland slope, 22.5.2002, Rudkovskaya 22
(PTZ); Kukkovka, Komsomol av. / Rovio str., ruderal roadside meadow, 16.5.2002,
Timofeeva (PTZ); Komsomol av., asphalt cracks along the pavement, 16.5.2002, Timo-
feeva (PTZ); Sortavala str., ruderal roadside meadow by the school No. 34, 16.5.
2002, Timofeeva (PTZ). – New to the Karelian Republic. Identifications by Øllgaard
and Räsänen, January 2003. H. Øllgaard, J. Räsänen & A. N. Sennikov
Taraxacum sphaeroidale H. Øllg., nom. nov. ≡Taraxacum sphaeroideum Rail. in Aquilo, Ser.
Bot. 14, App.: 50. 1977 [non Taraxacum sphaeroides Dahlst. 1911]. Following a request for a
binding decision, the Committee for Vascular Plants has, at a large majority, considered T.
sphaeroideum and T. sphaeroides to be confusingly similar names.
Taraxacum stenoschistum Dahlst.
+Rf(NW): Leningrad Region, Viborg, gräslinda i esplanaden vid Pantsarlahtigatan, 3.6.1932,
Marklund (H 197316). – First documented report for Russia; identifications by G.
Marklund. This record was included in the biogeographic province Ka (nowadays:
Finland and Leningrad Region of Russia), without specification, by Marklund (in
Acta Bot. Fenn. 26: 123. 1940). A. N. Sennikov
Taraxacum subcanescens Markl. ex Puol.
+Rf(NW): Leningrad Region, Viborg: ruderatmark vid ingången till Papula-parken, 7.6.1932,
Marklund (H 197511, 197512); Repola, vägkant nära gården, 5.6.1944, Marklund
(H 197513). – First documented report for Russia; identifications by G. Marklund.
This record was included in the biogeographic province Ka (nowadays: Finland and
Leningrad Region of Russia), without specification, by Marklund (in Acta Bot.
Fenn. 26: 124. 1940). A. N. Sennikov
Taraxacum subglaucescens Markl.
+Rf(N): Karelian Republic, Petrozavodsk: Drevlianka, Lososinskoye chaussée / Berezovaya
alley, ruderal grassland by the houses, 15.5.2002, Kravchenko 9854 (PTZ); central
Willdenowia 37 – 2007 187

part, “Yamka” park, left river side of Lososinka, 15.5.2002, Rudkovskaya 7-9
(PTZ); Gylling embankment, ruderal grassland, 22.5.2002, Rudkovskaya 28 (PTZ);
Kukkovka, Sortavala str., ruderal meadow by the school 34, 16.5.2002, Timofeeva
(PTZ); Sortavala str. 8, ruderal meadow by the buildings, 15.5.2002, Timofeeva
(PTZ). – New to the Karelian Republic. Identifications by Øllgaard and Räsänen,
Jan. 2003. H. Øllgaard, J. Räsänen & A. N. Sennikov
Taraxacum subhuelphersianum M. P. Christ.
+Rf(NW): Leningrad Region, Viborg: gräslinda i esplanaden vid Torkelsgatan, 5.6.1932, Mark-
lund (H 134257, 134258, 134259). – New to Russia. Identified by Øllgaard in 1980.
H. Øllgaard & A. N. Sennikov
Taraxacum sublaeticolor Dahlst.
+Rf(N): Karelian Republic, Sortavala, seminariet, 20.6.1938, Lindberg (H 197753-197757);
Sordavala stad, äng, 24.5.1910, Marklund (H 197747-197749); Sortavala, satama,
1.6.1934, Peltomaa (H 197746); Sortavala: Helylä haltpunkt, 10.6.1910, Arvonen
(H 197707, 197709); Helylä haltpunkt, 8.6.1910, Arvonen (H 197708); Helylä, äng,
4.6.1910, Marklund (H 197752). – Kol/Kar, Petrozavodsk: central part, by the State
Archive, basement cracks, 15.5.2002, Rudkovskaya 4 (PTZ); Pribrezhniy park, un-
der maples, 17.5.2002, Rudkovskaya 20, 21 (PTZ); Kukkovka, Sortavala str. 12,
ruderal meadow by the buildings, 17.5.2002, Timofeeva (PTZ). – First documented
report for the Karelian Republic. The specimens from Sortavala, identified by G.
Marklund and H. Lindberg, were included in the biogeographic province Kl (nowa-
days: Finland and Karelian Republic), without specification, by Marklund (in Acta
Bot. Fenn. 26: 124. 1940). The collection from Petrozavodsk was identified by
Øllgaard & Räsänen in January 2003. A. N. Sennikov, H. Øllgaard & J. Räsänen
Taraxacum sundbergii Dahlst.
+Rf(NW): Leningrad Region, Viborg: Trångsund [Vysotsk], gräsbevuxen vägkant mellan bräd-
gårdarna, 5.6.1932, Marklund (H 201308). – First documented report for Russia;
identifications by G. Marklund. This record was included in the biogeographic prov-
ince Ka (nowadays: Finland and Leningrad Region of Russia), without specification,
by Marklund (in Acta Bot. Fenn. 26: 127. 1940). A. N. Sennikov
Taraxacum trilobatum Palmgr.
+Rf(N): Karelian Republic, Kurkijoki: Opisto, Raholanjoen jokitörmällä Andersininmäelle
vievän sillan luona, 3.6.1994, Sonck (H 710959, 710960). – New to the Karelian Re-
public. Identification by C.-E. Sonck. A. N. Sennikov
Taraxacum tumentilobum Markl. ex Puol.
+Rf(N): Karelian Republic, Kurkijoki: Andersininmäki, Sonckin talon edustalla, 3.6.1994,
Sonck (H 713965). – New to the Karelian Republic. Identification by C.-E. Sonck,
confirmed by Øllgaard in 1996. H. Øllgaard & A. N. Sennikov
Taraxacum verecundum G. E. Haglund [= T. gracilentum H. Lindb.]
+Rf(N): Karelian Republic, Sortavala, seminariet, 20.6.1938, Lindberg (H 200933-200935).
– First documented report for Karelian Republic. This record was included in the
biogeographic province Kl (nowadays: Finland and Karelian Republic), without
specification, by Såltin (in Lounais-Hämeen Luonto 18: 9. 1965). Identifications by
H. Lindberg. A. N. Sennikov
Taraxacum vitellinum Dahlst.
+Rf(N): Karelian Republic, Sortavala: etelärinne, 1.6.1934, Peltomaa (H 205539). – New to
Russia. Identification by G. Marklund. A. N. Sennikov
188 Greuter & Raab-Straube: Euro+Med Notulae, 3

Tragopogon porrifolius subsp.eriospermus (Ten.) Greuter, comb. & stat. nov. ≡Tragopogon
eriospermus Ten., Fl. Med. Univ. 2: 3. 1822 [= Tragopogon porrifolius var. brachyphyllus Boiss.,
Fl. Orient. 3: 745. 1875 ≡Tragopogon brachyphyllus (Boiss.) Gand., Fl. Cret.: 67. 1916].
Address of the editors:
Prof. Dr W. Greuter & Eckhard von Raab-Straube, Botanischer Garten und Botanisches Museum
Berlin-Dahlem, Freie Universität Berlin, Königin-Luise-Str. 6-8, D-14195 Berlin; e-mail:
w.greuter@bgbm.org, e.raab-straube@bgbm.org
Willdenowia 37 – 2007 189