Article

Late Triassic reefs from the Northwest and South Tethys: Distribution, setting, and biotic composition

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The paleolatitudinal distribution patterns during Ladinian and Carnian time are characterized by an increasing expansion of reefs from the northern to the southern hemisphere. The optimum of reef diversity and frequency in the Norian is associated with the development of extended attached or isolated carbonate platforms. Norian-Rhaetian sponge and coral reefs of the Northern Calcareous Alps developed (1) as reef belt composed of patch reefs in platform-edge positions facing the open-marine northwestern Tethys basins and (2) as patch reefs in intraplatform basins as well as in ramp positions. Carnian and Norian-Rhaetian sponge and coral reefs of the Arabian Peninsula are formed (1) as reef complexes at the margins of carbonate platforms on the tops of volcanic seamounts in the southern Tethyan ocean, as small biostromes on these isolated platforms, and (2) as transgressive reef complexes on the attached platform of the Gondwana margin. The Norian Gosaukamm reefal breccia of the NW Tethys is a counterpart of Jabal Wasa reefal limestone of the Gondwana margin with similarities in geological setting and biotic composition. Rhaetian coral biostromes of low diversity known from the Austrian Koessen basin resemble to the time equivalent Ala biostromes of the isolated Kawr platform in the southern Neo-Tethys by forming a discontinuous layer in shallow intraplatform basin setting.

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... 1,2) contributes to unravel the interplay of factors influencing carbonate sedimentary environments at a regional scale. In fact, from late Norian to early Jurassic, these depositional settings witnessed: (a) the demise of a long-lasting, hundreds of kilometres wide and over 1000 m thick, Norian early-dolomitized inner platform (Hauptdolomit) adjacent to hundreds of kilometres wide carbonate shelf (Dachstein Limestone and Reef); (b) a major late Norian-Rhaetian change in sediment supply with the input of fine siliciclastics, previously absent, alternating with carbonate strata (Hochalm and Eiberg members of the Kössen Formation), attributed to a change towards more humid climatic conditions (Berra et al. 2010); (c) the late Rhaetian development of intraplatform basins (Eiberg Basin ;Golebiowski 1990;Kaufmann 2009;Mette et al. 2012;Krystyn et al. 2005; Restental Basin; Gawlick et al. 2009;Richoz et al. 2012;Hillebrandt et al. 2013), passing laterally to shallow-water carbonates (Oberrhätkalk/Upper Rhaetian Limestone ;Schäfer 1979;Golebiowski 1990;Krystyn et al. 2005) with margins with coral patch reefs (Bernecker et al. 1999;Bernecker 2005;Kaufmann 2009) or distally steepened ramp with skeletal packstone/grainstone and mounds (Piller 1981;Flügel 1989, 1995;Flügel 1981Flügel , 2002; (d) a sea-level fall followed by rapid rise and biotic turnover across the end-Triassic extinction event (McRoberts et al. 2012); (e) the late early Jurassic rifting leading to the opening of the Penninic Ocean (Bernoulli 1964;Eberli 1988;Gawlick et al. 2009). ...
... Basinward, the Hauptdolomit was adjacent to lagoonal subtidal-peritidal cyclic strata (Lofer cyclothems of the Dachstein Limestone ;Fischer 1964;Enos and Samankassou 1998;Missoni et al. 2005), passing to the Dachstein Reef shelf margin (Fig. 1C). The Dachstein Reef (Flügel 2002;Bernecker 2005) faced the Hallstatt outer shelf of the Neo-Tethys Ocean (Matzner 1986;Kozur 1991;Gawlick et al. 1999;Gawlick and Böhm 2000;Köppen and Carter 2000). The Hauptdolomit is overlain by a well-bedded, 10-400 m thick, only partly dolomitized, carbonate unit, labelled as Plattenkalk (Steiner 1965a, b;Czurda and Nicklas 1970;Golebiowski 1989). ...
... The Eiberg Basin was lateral to shallow-marine carbonates (Oberrhätkalk/Upper Rhaetian Limestone; Figs. 1C, 2B) overlying the coral-rich Unit 4 of the Hochalm Mb. (Golebiowski 1990;Wegerer and Gawlick 1999;Bernecker 2005;Rizzi et al. 2020). The southern margin of the Eiberg Basin at the Steinplatte was interpreted as a distally steepened carbonate ramp formed by the deposition of autochthonous biogenic sediment (massive mound facies with bioclastic sand and shell coquinas) on an antecedent homoclinal ramp (Stanton and Flügel 1995;Flügel 2002), laterally adjacent to the Dachstein Limestone shelf (Kaufmann 2009). ...
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Western Tethys sedimentary successions constitute fundamental archives of Late Triassic–Early Jurassic environmental, carbonate production and tectonic changes. During the Late Triassic, the Northern Calcareous Alps (Austria) belonged to the Western Tethys passive margin, characterised by the deposition of the early-dolomitized peritidal Hauptdolomit (Norian) adjacent basinward to the Dachstein carbonate shelf and passing upward to the mixed carbonate–siliciclastic Kössen Formation (Upper Norian–Rhaetian). The Kössen Fm. was subdivided into the lower shallow-water Hochalm Member and the upper Eiberg Member, accumulated in an intraplatform basin coeval to shallow-water carbonates (Upper Rhaetian Limestone). The Eiberg Mb. and overlying Jurassic strata were extensively studied as a continuous marine record across the Triassic/Jurassic boundary. In contrast, shallow-water successions, time-equivalent to the Eiberg Mb. and Upper Rhaetian Limestone, located North of the Eiberg Basin, are poorly investigated. This study focusses on the approximately 350 m thick Norian to Lower Jurassic succession cropping out in the eastern Northern Calcareous Alps (Stumpfmauer). The 32 distinguished lithofacies compose seven, vertically superimposed, sedimentary units (A–G), belonging to the Hochalm Mb. of the Kössen Fm. (Unit A peritidal cyclothems, Unit B claystone/marlstone with fossiliferous beds, Unit C coral boundstone to floatstone), Upper Rhaetian Limestone (Unit D subtidal cyclothems with claystone), shallow-water carbonate strata transitional to Lower Jurassic (Unit E ooidal coated grain peloidal grainstone with basal transgressive lag, Unit F bivalve-rich, microbialite and oncoidal lithofacies, previously attributed to the Upper Rhaetian Limestone) and Hettangian–Sinemurian Kalksburg Fm. (Unit G cross-laminated coated grain peloidal grainstone with quartz and chert). The detailed lithofacies characterisation presented in this study contributes to the knowledge on the Northern Calcareous Alps stratigraphy and depositional environments across the Triassic/Jurassic boundary. The identified sedimentary units can be framed in the evolution of Western Tethys and share similarities with depositional systems from the Western Carpathians, Transdanubian Range, Southern Alps and Dinarides suggesting coherent sedimentary response and environmental, climate and tectonic controls in different palaeogeographic domains.
... Epeiric platforms covering the tropical margins of the Neotethys Ocean generally consisted of a wide tidal flat, a shallow lagoon, and a platform margin rimmed by reefs (Mandl, 2000;Bernecker, 2005). The tidal flat is stratigraphically represented by the Hauptdolomit (Glavni dolomit/Dolomia Principale/"Main Dolomite"), gradually passing into the bedded Dachstein limestone (Mandl, 2000;Bernecker, 2005;Kovács et al., 2011). ...
... Epeiric platforms covering the tropical margins of the Neotethys Ocean generally consisted of a wide tidal flat, a shallow lagoon, and a platform margin rimmed by reefs (Mandl, 2000;Bernecker, 2005). The tidal flat is stratigraphically represented by the Hauptdolomit (Glavni dolomit/Dolomia Principale/"Main Dolomite"), gradually passing into the bedded Dachstein limestone (Mandl, 2000;Bernecker, 2005;Kovács et al., 2011). Characteristic for these beds is the so-called Lofer cycle, comprising a thin breccia member, a laminated mudstone (intertidal stromatolite), and a biogenic wackestone and/ or floatstone with large megalodontid bivalves, gastropods, and locally corals (Fisher, 1964;Ogorelec & Rothe, 1993;Satterley, 1996;Ogorelec & Buser, 1996;Haas, 2004;Samankassou & Enos, 2019). ...
... Patch reefs developed at the rims or within these basins. Their composition is comparable to the composition of the marginal Dachstein-type reefs (Schäfer & Senowbari-Daryan, 1981;Kuss, 1983;Bernecker, 2005). In the inner parts of the platform, significantly more restricted basins already formed in the Norian. ...
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In the southernmost outcrops of the Slovenian Basin the Middle Jurassic coarse-grained limestone breccia (mega)beds are interstratified within a succession that is otherwise dominated by hemipelagites and distal turbidites. In this paper, these beds are described as the Ponikve Breccia Member of the Tolmin Formation. We provide descriptions of the studied sections with detailed geological maps and analysis of the breccia lithoclasts. From the latter, a non-outcropping margin of the Dinaric Carbonate Platform is reconstructed. In the Late Triassic the platform margin was characterized by a Dachstein-type marginal reef. After the end-Triassic extinction event, the platform architecture remained, but the reefs were replaced by sand shoals characterized by ooids. In the late Early Jurassic and/or early Middle Jurassic a slope area might have been dissected by normal faults and a step-like paleotopography was formed. In the Bajocian, during a period of major regional geodynamic perturbations, extensional or transtensional tectonic activity intensified and triggered the large-scale collapses of the Dinaric Carbonate Platform margin producing the limestone breccias described herein. This may in turn have caused a backstepping of the platform margin, as is evident from the occurrence of Late Jurassic marginal reefs that are installed directly above the Upper Triassic and Lower Jurassic inner platform successions.
... Carbonate platforms host more than 25% of marine life [1] and are very sensitive to climatic, oceanographic and bathymetric changes. Despite their importance, our knowledge on Upper Triassic shallowwater carbonates is fragmentary, and mainly derives from the study of Tethyan carbonate systems (e.g., see [2][3][4][5][6][7][8][9][10][11][12][13][14]). Upper Triassic carbonate platforms from the substantially larger Panthalassan domain [15] remain understudied, providing a minor source of information for the general comprehension of Upper Triassic carbonates. ...
... Therefore, very often the reconstruction of carbonate systems is principally based on the interpretation of scattered microfacies data, not taking into account considerations of lateral or vertical facies transition (e.g., see the work approach in [17]). So far, these microfacies-based reconstructions have relied mostly on observations from better-studied and better-preserved Tethyan systems [3,12,138]. Nonetheless, Tethyan systems were formed in completely different settings ( [16] and references therein) and occurred at very different scales; for these reasons, they are probably not the best analogues to reconstruct Panthalassan systems. ...
... Slovenia Norian-Rhaetian [89,[200][201][202] Northern Calcareous Alps Norian-Rhaetian [9,12,203] Iran Norian [14,[204][205][206][207][208][209][210][211] Sicily Norian [212][213][214][215][216][217][218] Turkey Norian [204,[219][220][221][222][223][224][225] Oman Norian [12] Japan (Reef type 2) Norian [18,25] Oregon (Summit Point, USA) Norian [214,224] Yukon (Canada) Norian [51,52,59,60] and this study ...
Article
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Upper Triassic carbonate platforms from the Panthalassa Ocean remain less-understood and less-studied than their Tethyan equivalents. This imbalance is largely due to the poorer preservation state of Panthalassan carbonate rock successions in terms of rock quality and depositional geometries, which prevents good appreciation of depositional systems. In this context, carbonate exposures from Lime Peak (Yukon, Canada) represent an outstanding exception. There, the remains of an Upper Norian Panthalassan carbonate platform are well-exposed, show remarkably preserved depositional geometries and overall superior rock preservation. In this work, we analyse the carbonates from the Lime Peak area with particular attention to the vertical and lateral distribution of biotic assemblages and microfacies at the platform scale. Results demonstrate that the Lime Peak platform was surrounded by a basin with an aphotic sea bottom. The carbonate complex developed in warm waters characterized by high carbonate saturation. The area was also defined by moderate to high nutrient levels: this influenced the type of carbonate factory by favouring microbialites and sponges over corals. During its growth, Lime Peak was influenced by tectono-eustatism, which controlled the accommodation space at the platform top, primarily impacting the internal platform environments and the stability of the slope. Gaining better knowledge of the spatial distribution and dynamics of Upper Triassic organisms and sedimentary facies of Panthalassa in relation to tectono-eustatism lays the first foundations for reconstructing more robust platform models and understanding the evolution of other, more dismantled Upper Triassic Panthalassan carbonate systems through time.
... Fossil reefs therefore constitute important records of marine environment evolution during geological times. The Late Triassic is a period of important reef development (Stanley, 1988;Flügel, 1981Flügel, , 2002Bernecker, 2005;Kiessling, 2010;Martindale et al., 2015). This development led to significant carbonate production worldwide and contributed to the expansion of carbonate platforms during the Late Triassic. ...
... This development led to significant carbonate production worldwide and contributed to the expansion of carbonate platforms during the Late Triassic. During the Triassic, reef ecosystems experienced several changes with regards to their distributional patterns, paleogeographic settings, taxonomic composition, and biodiversity (Flügel, 2002;Bernecker, 2005). Reef evolution during the Triassic is typically viewed as a three step process, starting with Lower Triassic microbial-dominated reefs, followed by Anisian to Carnian and Norian to Rhaetian metazoan-dominated reefs (Stanley, 1988;Flügel, 2002). ...
... Accordingly, an inventory of reef components, point-counting results, and thin section mapping are presented in the following sections. Dating of Upper Triassic reef limestone is typically based on the association of reef builders (e.g., Bernecker, 2005). However, biotic assemblages alone can sometimes be misleading (Martindale et al., 2013b) and additional independent age indicators should be used to provide an accurate and more reliable dating. ...
... The Northern Tethys Ocean was an area of profuse carbonate accretion and major reef building in the Late Triassic (Flügel and Senowbari-Daryan, 2001;Flü gel, 2002;Bernecker, 2005). Following a metazoan reef gap in the Early Triassic (Scythian) where only microbial reef mounds, bivalve bioherms, and small, rare sponge biostromes occur (Pruss and Bottjer, 2004;Brayard et al., 2011;Pruss et al., 2007), metazoan reef ecosystems began to rebound from the Permo-Triassic extinction. ...
... Finally, the second reef optimum in the Norian and Rhaetian was characterized by huge kilometer-scale reef structures built by large scleractinian corals and hypercalcified sponges (Flü gel, 2002). At this time, the northern Tethys was a semi-enclosed ocean basin with numerous, large carbonate platforms and intraplatform basins a few hundred meters deep (Flü gel and Senowbari-Daryan, 2001;Flü gel, 2002;Bernecker, 2005;Kenter and Schlager, 2009). The abundant carbonate platforms and warm water circulating northward from the tropical Tethys made the Northern Tethys an ideal location for carbonate platform accretion and reef growth. ...
... Another problem is that, despite the interest in Late Triassic reef ecosystems, (e.g., Zapfe, 1962;Cuif, 1966;Schlager, 1967;Zankl, 1968;Stanley, 1979;Flü gel, 1981Flü gel, , 1982Flü gel, , 2002Piller, 1981;Schäfer and Senowbari-Daryan, 1981;Stanton and Flü gel, 1987;Satterley, 1994;Bernecker et al., 1999;Bernecker, 2005; and references therein), until recently, very few absolute ages have been available for this time interval. The issue of age can also be complicated by studies which assume an age based merely on the stratigraphic character of reefal biotic assemblages. ...
Article
Carnian and Norian (Upper Triassic) limestones and dolostones along the Eisriesenwelt trail on the Tennengebirge (Salzburg, Austria) reveal a progression from fore-reef to lagoonal environments, including a small Norian reef with both Carnian and Norian characteristics. Here, we provide high-resolution biostratigraphic age constraints of the Tennengebirge platform carbonates and describe an early Norian patch reef that is built by large Retiophyllia corals and encrusted by ''Tubiphytes,'' sponges, and microbial fabrics. The tall (up to 4 m), narrow phaceloid Retiophyllia coral colonies exhibit phototropic growth patterns—coral branches that are at the top of the colony grew longer than those on the side of the colony— thus, we suggest that these corals had a symbiotic association with photoautotrophs (most likely zooxanthellae). The well-constrained ammonoid and conodont biostratigraphy presented here establish that the Tennengebirge patch reef was deposited in the early Norian, nevertheless, it contains features typically associated with Carnian reefs (small, encrusting sponges, Carnian-style microbial crusts, and ''Tubiphytes''). The Carnian traits of the Norian reef emphasize the importance of accurate biostratigraphic dating; obtaining independent age estimates for reef outcrops is crucial to correctly determine the timing and magnitude of transitions in Triassic reef ecology. INTRODUCTION
... The Late Carnian sea-level rise (Budai & Haas, 1997; Gawlick & BöHm, 2000; Hallam, 2001; Gianolla et al., 2003; Berra et al., 2010) and a warm climate (sattler & scHlaf, 1999; flüGel, 2004; Berra et al., 2010; Preto et al., 2010; stefani et al., 2010 ) together with a suitable palaeogeographic position near the palaeoequator (stamPfli & Borel, 2002 Borel, , 2004 stamPfli & kozur, 2006; Golonka, 2007) created favourable conditions for the development of extensive epeiric carbonate platforms along the Neotethys Ocean (Haas, 2004; VlaHoVi} et al., 2002 VlaHoVi} et al., , 2005 Bernecker, 2005; Haas et al., 2007; Golonka, 2007) and a co-temporal bloom of scleractinian coral reefs (turn{ek, 1997; stanley Jr., 2003; flüGel, 2004; Bernecker, 2005; russo, 2005). Extremly thick carbonate platforms developed during Norian and Rhaetian also in the NE corner of the Adria microplate: the Julian Carbonate Platform is now structurally mostly incorporated in the Julian Nappe of the Eastern Southern Alps, whereas the Dinaric Carbonate Platform belongs to the External Dinarides (Buser, 1986; Placer, 1999 Placer, , 2008). ...
... The Late Carnian sea-level rise (Budai & Haas, 1997; Gawlick & BöHm, 2000; Hallam, 2001; Gianolla et al., 2003; Berra et al., 2010) and a warm climate (sattler & scHlaf, 1999; flüGel, 2004; Berra et al., 2010; Preto et al., 2010; stefani et al., 2010 ) together with a suitable palaeogeographic position near the palaeoequator (stamPfli & Borel, 2002 Borel, , 2004 stamPfli & kozur, 2006; Golonka, 2007) created favourable conditions for the development of extensive epeiric carbonate platforms along the Neotethys Ocean (Haas, 2004; VlaHoVi} et al., 2002 VlaHoVi} et al., , 2005 Bernecker, 2005; Haas et al., 2007; Golonka, 2007) and a co-temporal bloom of scleractinian coral reefs (turn{ek, 1997; stanley Jr., 2003; flüGel, 2004; Bernecker, 2005; russo, 2005). Extremly thick carbonate platforms developed during Norian and Rhaetian also in the NE corner of the Adria microplate: the Julian Carbonate Platform is now structurally mostly incorporated in the Julian Nappe of the Eastern Southern Alps, whereas the Dinaric Carbonate Platform belongs to the External Dinarides (Buser, 1986; Placer, 1999 Placer, , 2008). ...
... is typical for the central reef area (e.g. HoHeneGGer & loBitzer, 1971; scHäfer & senowBari-daryan, 1978; senowBari-daryan, 1980; sadati, 1981; senowBari-daryan et al., 1982; wurm, 1982; zaninetti et al., 1982; kuss, 1983; matzner, 1986; kristan-tollmann, 1990; Bernecker, 2005; cHaBlais et al., 2010b). ...
Article
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Mt. Begunjščica (Karavanke Mts., northern Slovenia) structurally belongs to the Košuta Unit (eastern Southern Alps). The Dachstein Limestone, building the northern side of the mountain and its main ridge, was deposited on the Julian Carbonate Platform, while grey and red nodular Jurassic limestones of the southern slope represent sedimentation on the Julian High. The massive Dachstein Limestone contains a rich assemblage of benthic foraminifera. Typical representatives of the reef and back-reef area were recognized. The age of the assemblage is dated as Rhaetian, based on the co-presence of species with a Norian and Rhaetian time span, such as Galeanella tollmanni, "Sigmoilina" schaeferae, Alpinophragmium perforatum, Aulotortus tumidus, Variostoma catilliforme, Variostoma cochlea and Variostoma helicta, together with the Rhaetian to Lower Jurassic Involutina turgida.
... Species Kaeveria fluegeli is known from the Norian–Rhaetian limestones of the Taurus in southern Turkey (Zaninetti et al., 1982) and from the Argolis Peninsula and Hydra Island of southern Greece (Senowbari-Daryan et al., 1996 ). In Oman, these species are confined to facies of clastic limestones of the Norian–Rhaetian Jabal Kawr Formation (Bernecker, 1996Bernecker, , 2005). In the Southern Apennines, this species is characteristic exclusively of the slope clastic facies (clastic dolomites) of the Monti del Maddalena unit (= Dachstein-type reef) of the Norian–Rhaetian age (Iannace and Zamparelli, 2002). ...
... In the Southern Apennines, this species is characteristic exclusively of the slope clastic facies (clastic dolomites) of the Monti del Maddalena unit (= Dachstein-type reef) of the Norian–Rhaetian age (Iannace and Zamparelli, 2002). It occurs as well in the Norian and Rhaetian of the Northern Alps in Gosau area (Riedel, 1988; Bernecker, 2005 ) and in the Rhaetian limestones of Papua New Guinea (Kristan-Tollmann , 1990) (Figs. 3b and 3c). ...
... It is remarkable that in the southern (Peri-Gondwanan ) shelf of the Tethys, the species under consideration coexisted with foraminifers of genera Siphonofera , Costifera , and Siculocosta whose distribution area delineate the southern (sephardian) paleobiogeographic province of the Tethys. On the other hand, the aforementioned genera of the sephardian province are unknown in the South-East Pamirs and Northern Zaninetti, Altiner, Dager et Ducret, 1982 ) and its localities in the South-East Pamirs (a, Zorkarajilga section, Exposure 2) and in other regions of the world shown in the present-day geographic scheme (b) and (c) in paleotectonic map after (Golonka, 2007): ( 1 ) the Taurus, southern Turkey (Zaninetti et al., 1982), ( 2 ) Argolis Peninsula and Hydra Island, southern Greece (Senowbari-Daryan et al., 1996), ( 3 ) Southern Apennines, Italy (Iannace and Zamparelli, 2002), ( 4 ) Northern Calcareous Alps, Austria (Bernecker, 2005), ( 5 ) Oman (Bernecker, 2005), ( 6 ) New Guinea (Kristan-Tollmann, 1990), ( 7 ) South-East Pamirs (this work). Shown inZaninetti, Altiner, Dager et Ducret, 1982), hypotype no. ...
Article
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Foraminifers representing species Kaeveria fluegeli have been found in the Zorkaradjilga Formation (Sagenites quinquepunctatus Zone) of the upper Norian (or lower Rhaetian) in the central structural-facies zone of the South-East Pamirs. Their occurrence here is an additional criterion substantiating age and correlation of host deposits and an evidence in favor of fauna migration from the southern Tethys during the respective time span not only in northwestern areas of that ocean (Northern Calcareous Alps), but also in its central part (the South-East Pamirs).
... Important works on Triassic reefs by several authors (Stanley, 1979;Flügel, 1981Flügel, , 1982Flügel, , 2002Flügel and Stanley, 1984;Flügel and Flügel-Kahler, 1992;Kiessling et al., 1999;Bernecker, 2005; with bibliography) have thoroughly catalogued the numerous Upper Triassic reef localities throughout the world. However, most studies of Upper Triassic reefs involve those in the Tethys realm. ...
... For each country, we recorded the number of recognised species of reef builders and dwellers and compiled data on corals, sponges, spongiomorphids, algae, microbial biota, foraminifers, and microproblematica. Reef data from the Northern Calcareous Alps, including the Göll, Gruber, Adnet, Rötelwand, Gosaukamm and Steinplatte reefs, were obtained from Zankl (1969), Senowbari-Daryan (1980b), Flügel (1981), Schäfer and Senowbari-Daryan (1981), Bernecker et al. (1999), Bernecker (2005) and Senowbari-Daryan (2009). Data for Sicilian reef fossils were obtained from Senowbari-Daryan (1980b), Senowbari-Daryan et al. (1982), Senowbari-Daryan and Schäfer (1986) and Di Stefano et al. (1990). ...
... Data for Sicilian reef fossils were obtained from Senowbari-Daryan (1980b), Senowbari-Daryan et al. (1982), Senowbari-Daryan and Schäfer (1986) and Di Stefano et al. (1990). Biotic composition data for the Oman reef limestones (Late Carnian-Rhaetian) of Jabal Wasa (attached-platform type) and Jabal Kawr (isolated-platform type) were obtained from Weidlich et al. (1993), Bernecker (1996Bernecker ( , 2005Bernecker ( , 2007, Senowbari-Daryan et al. (1999) and Senowbari-Daryan and Bernecker (2009). Finally, data on the Sambosan reef limestones (Japan) were taken from Kanmera (1964), Okuda and Yamagiwa (1978), Onoue and Stanley (2008), Chablais et al. (2010) and the present study. ...
... Cradles of life, shallow-water reefal carbonate systems, which represent a key archive for marine life evolution, extensively developed. The Upper Triassic, notably, is marked by the development of numerous widely studied reefal carbonate systems in the Tethyan realm (Bernecker, 2005;Buser et al., 1982;Flügel, 1982;Gale et al., 2015;Gattolin et al., 2015;Jin et al., 2018;Krystin et al., 2009;Martindale et al., 2014;Schäfer and Senowbari-Daryan, 1982;Tomašových, 2004). Their Panthalassan counterparts are in contrast poorly known and mostly preserved as isolated bodies within diverse tectonostratigraphic terranes, which accreted along the Circum-Pacific region (Chablais et al., , 2010bMartindale et al., 2015;Peybernes et al., 2016b;Stanley Jr., 1979;Stanley and Senowbari-Daryan, 1986;Zonneveld et al., 2007). ...
... The described association in this facies is interpreted as diagnostic of reef environments. Similar assemblages are widely reported from Upper Triassic platform margins in the Tethys (Bernecker, 2005;Flügel and Senowbari-Daryan, 2001) and Panthalassa (Chablais et al., 2010c, Onoue et al., 2009Peybernes et al., 2015;Martindale et al., 2012). However, poor preservation and diversity in organisms (Fig. 6) do not allow us to make specific comparison with these coeval systems. ...
Article
Contrary to their Tethyan counterparts, and despite substantial research, Upper Triassic shallow-water limestone from the Panthalassa Ocean remain poorly known. Their understanding is yet crucial to better constrain past depositional, ecologic, geographic and geodynamic conditions out of the Tethyan domain, and to better assess life evolution and recovery following the greatest Permo-Triassic biological crisis. The Taukha terrane, located in a poorly studied area of the Sikhote-Alin mountain range (Primorsky and Khabarovsk Kraya, Far East Russia) is a key Panthalassan terrane. Near Dalnegorsk City, North-East of Vladivostok, on the northern part of the Taukha terrane, Upper Triassic marine carbonates abound. They show a clear affinity with Panthalassan terranes of both Japan and North America. There, eleven new localities have been extensively sampled, described and inter- preted. Based on field observations and facies analysis, eight major facies, corresponding to specific depositional environments within a carbonate platform, have been established. A conceptual depositional model corre- sponding to a mid-oceanic atoll-type system composed of well-developed lagoon, fringing patch reefs and oolitic shoals is proposed. The foraminiferal assemblages allow refining the age of the carbonate system to the Norian, and locally, exclusively to the lower/middle and the middle parts of the Norian.
... Following the end-Permian extinction, organic build-ups gradually recovered in terms of complexity and global distribution (Flügel 1982(Flügel , 2002Bernecker 2005;Martindale et al. 2017). By the end of the Middle Triassic, extensive build-ups can be found in a large variety of settings within the tropical belt of the Neo-Tethys and the Panthalassa oceans, most of them on the upper slopes and platformmargins of land-attached platforms or isolated seamounts, but also on lower slopes, on platform tops and in lagoons (Flügel 2002). ...
... Middle Triassic and Lower Carnian reefs mostly occupied the upper slopes and margins of carbonate platforms surrounding the Neotethys Ocean (Keim and Schlager 1999;Flügel 2002;Seeling et al. 2005;Brandner et al. 2007;Trombetta 2011) and of the isolated seamounts traversing the Neotethys or the Panthalassa oceans (e.g., Bernecker 1996;Senowbari-Daryan et al. 1999;Bernecker 2005Bernecker , 2007Peybernes et al. 2015Peybernes et al. , 2016. Smaller patch reefs also formed within lagoons in the back reef area or in shallowing-up basins (Fürsich and Wendt 1977;Wendt and Fürsich 1980;Wendt 1982;Russo et al. 1991). ...
Article
The architecture and composition of Middle to lower Upper Triassic platforms is often obscured by dolomitization. Hence, comparatively little is known about their architectures compared to their size and geographic extent. An active quarry near Lesno Brdo (central Slovenia) offers an excellent exposure of Lower Carnian (Julian) massive limestone, which is diagenetically little altered. A detailed microfacies analysis along a 15.5-m log revealed the presence of three facies types: fine-grained limestone as a groundmass, blocks and globular masses of sponge-microbialite boundstone, and lens-like aggregations of polychaete (terebellid) tubes. Sponge-microbialite boundstone contains a rather small number of cosmopolitan sponge taxa, solenoporacean red algae, microproblematica, bryozoans, and a small proportion of dwelling fauna. Instead, stromatolites represent the main constituent. While some blocks appear to have truncated margins, others show mammillary-like protrusions of microbialites into the surrounding sediment, suggesting active growth of microbialite-producing organisms. Aggregations of terebellid worm tubes show a highly irregular relief, with tubes placed sub-parallel to the ancient sea floor. The presence of fibrous rim cement, crystal silt, and in some cases fragmentation of the tubes, suggest at least moderately energetic waters. Aggregations are thus interpreted as preserved in situ, but not in toto. The entire complex was probably deposited at the margin or upper slope of a carbonate platform. Although the presence of a large number of terebellids associated with microbialites boundstone may indicate some sort of environmental stress, such a stress remains to be identified.
... Most of the proposed depositional models are referred to the Early Norian and feature reef-rimmed systems developing from initial open platform configurations (Mandl, 2000;Kenter & Schlager, 2009;Krystyn et al., 2009), with a shallow lagoon behind the margin system, connected inward with restricted environments where dolomitic peritidal cycles formed (Hauptdolomit; in Mandl, 2000;Haas, 2012;Haas et al., 2006Haas et al., , 2015. On the margin to upper slope, sponges and scleractinian corals were the main reef builders, whereas microbialites, Tubiphytes and other encrusters occurred as minor components (Dullo, 1980;Wurm, 1982;Fl€ ugel, 2002;Bernecker, 2005;Haas et al., 2010b). Few cases are instead constrained to the Late Tuvalian time (Martindale et al., 2013), showing features partly similar to the Dolomia Principale carbonate platform; thinbedded periplatform basinal limestones are overlain by a prograding platform whose fore reef to reefal facies are characterized by microbialites as primary reef builders. ...
... The Late Carnian emplacement of the Dolomia Principale carbonate platform thus represents the return, in the western Tethys, to high-relief build-ups after a late Julian demise (Schlager & Schollnberger, 1974;Fl€ ugel, 2002;Gianolla et al., 2003;Keim et al., 2006;Stefani et al., 2010;Lukeneder et al., 2012;Gattolin et al., 2015) and may be viewed, in terms of shallow-water carbonate systems, as marking the end of the prolonged environmental perturbation known as the Carnian Pluvial Event. The margin to slope microbial-dominated carbonate factory of the earlystage Dolomia Principale carbonate platform fits well with other coeval reef communities of Western Tethys (Bernecker, 2005;Martindale et al., 2015) and slightly precedes the Late Carnian-Early Norian biological turnover (Fl€ ugel, 2002;Kiessling, 2009) that caused the microbialites to occur only as accessories in coralsponge-dominated reefs. ...
Article
Wide carbonate platform environments developed on the western passive margin of the Tethys during the Late Triassic, after a major climate change (Carnian Pluvial Episode) that produced a crisis of high-relief microbial carbonate platforms. The peritidal succession of this epicontinental platform (Dolomia Principale/Hauptdolomit, Dachstein Limestone) is widespread in the Mediterranean region. However, the start-up stage is not fully understood. The original platform to basin depositional geometries of the system have been studied in the north-eastern Southern Alps, close to the Italian/Slovenian boundary where they are exceptionally preserved. Sedimentological features have been investigated in detail by measuring several stratigraphic sections cropping out along an ideal depositional profile. The analysis of the facies architecture allowed reconstruction of the paleoenvironments of the Dolomia Principale platform during its start-up and early growth stages in the late Carnian. The carbonate platform was characterized by an outer platform area, connected northward to steep slopes facing a relatively deep basin. Southward, the outer platform was connected to inner sheltered environments by a narrow, often emerged shelf crest. Behind this zone, carbonate sedimentation occurred in shallow lagoons and tidal flats, passing inward to a siliciclastic mudflat. The Dolomia Principale platform was initially aggrading and able to keep pace with a concomitant sea-level rise, and then prograding during the late Carnian. This stratigraphic interval was correlated to the Tuvalian succession of the Dolomites, allowing depiction of the depositional system on a wide scale of hundreds of kilometrew. This large-scale depositional system presents features in common with some Palaeozoic and Mesozoic carbonate buildups (fo example, the Permian Capitan Reef complex, Anisian Latemar platform), both in terms of architecture and prevailing carbonate producers. A microbial-dominated carbonate factory is testified in the outer platform and upper slope. The recovery of high relief microbial carbonate platforms marks the end of the Carnian Pluvial Episode in the Tuvalian of Tethys.
... During the Late Triassic, the Tethys was a semi-enclosed, (sub)tropical sea with numerous reefs, large carbonate platforms, and intraplatform basins (Flügel and Senowbari-Daryan, 2001;Flügel, 2002;Ziegler et al., 2003;Bernecker, 2005). The Tethys Ocean experienced warm, oligotrophic waters circulating northward and southward from the tropics (western boundary currents, Fig. 1), resulting in ideal conditions for prolific carbonate accumulation and photozoan reef growth, much like the modern coasts of Belize, northeastern Australia, and eastern Florida. ...
... Lime Peak may be the exception; Lime Peak reefs exhibit Tethyan affinities such as thicker cements, sponges, numerous epibionts, and Retiophyllia corals are important in the construction and reef ecology (Reid, 1985;Reid and Tempelman-Kluit, 1987). Panthalassic reefs also lack the multiple generations of foraminifera, sponges, "Tubiphytes", or other epibionts that are typical of Norian/Rhaetian reefs from the Tethys (see Fig. 6F, G, and Flügel, 1981;Flügel and Senowbari-Daryan, 2001;Flügel, 2002;Bernecker, 2005). ...
Article
The Late Triassic was a pivotal period in reef evolution, but the majority of information about reef ecology during this time comes from buildups in the Alps (e.g., the Tethys Ocean). Recent studies of reefs in western North America have recognized unique ecologies along the eastern margin of the Panthalassa Ocean. Although there are numerous (twenty-five) localities with putative reef builders, only four buildups had syndepositional relief and a rigid framework (i.e. true reefs). The most paleo-northern true reefs were microbial patch reefs with only a few large skeletal bioconstructors; hypercalcified sponges and spongiomorphs built the mid-latitude reefs, with secondary microbial encrustation and branching, phaceloid Retiophyllia corals. Corals are the primary bioconstructors in Panthalassa’s most paleo-equatorial reefs and calcareous microbes are sparse. When all reefal deposits are analyzed, the N-S gradient is also present, with microbial and bivalve deposits in the north, sponge-coral deposits in the mid latitudes, and coral deposits near the equator. This ecological gradient is not apparent in the Tethys Ocean. Tethyan reefs thrived in oligotrophic, tropical waters without strong latitudinal gradients; by contrast, paleoceanographic considerations suggest that cool, nutrient-rich waters swept south along the western North American borderlands in the Late Triassic. The eastern boundary current is interpreted to have created a strong north-south differentiation of environments in northeastern (NE) Panthalassa that was manifested in both the biotic and abiotic characteristics of eastern Panthalassic reefs. Reefs from equatorial Panthalassa are similar to Tethyan reefs (warm-water, photozoan, coral reef structures), whereas higher paleo-latitude reefs from Panthalassa are interpreted as cool-water (heterozoan) buildups, with abundant calcareous microbes, diminutive biocalcifiers, and few large, framework-building corals.
... While the Summit Point reefs may not be Dachstein-type reefs, they do have similarities to Tethyan reefs. Many of the Summit Point reef organisms are found in Upper Triassic Tethyan reefs; for instance the phaceloid Retiophyllia corals; many of the sponges, such as Grossotubenella, Celyphia, and Uvanella; Solenopora and Parachaetetes red algae; and the nubecularid foraminifera (Flü gel and Senowbari-Daryan, 2001;Flü gel, 2002;Bernecker, 2005 and references therein). In addition to these Upper Triassic forms, several of the Summit Point faunas are also found in Anisian (Middle Triassic) rocks from Turkey (Senowbari-Daryan and Link, 2005), for example, Thamastocoelia dolomitica and Aternasus irregularus. ...
... The Anisian encrusting bioconstructors are similar to those from Summit Point, although, as previously mentioned, the lack of Tubiphytes in the Summit Point reefs, and most northeastern Panthalassic reefs, is curious. Additionally, there are some Upper Triassic reefs from the southern Tethys with encrusters and bindstone facies similar to the Summit Point reefs (Bernecker, 2005). ...
Article
Microfacies analysis of five patch reefs from the Martin Bridge Formation (Upper Triassic/Norian, Summit Point, Oregon, United States) reveals that microbialite fabrics dominate reef construction versus corals or other large metazoans. Other reef bioconstructors include characteristic Late Triassic branching corals and diminutive calcifiers (solenoporacean red algae, foraminifera, and sponges). The patch reefs exhibit dense growth fabrics and had elevation above the seafloor, making them ecological reefs or true reefs. Robust binding and encrusting organisms inhabited higher-energy areas, whereas red algae, followed by phaceloid coral colonies, inhabited zones of decreasing water energy, defining a subtle zonation consistent with wave energy in the paleoenvironment. The dominance of microbial fabrics and diminutive bioconstructors make the Summit Point reefs distinct among Upper Triassic reefs from northeastern Panthalassa. Summit Point was originally identified as a Dachstein-type reef (e.g., high diversity, dominated by corals and sponges, massive bedding, large reef cavities). However, the dominance of microbial fabrics over larger metazoans, the (comparatively) modest faunal diversity, and lack of abundant, multigenerational epibionts or cements clearly differentiates the Summit Point reefs from the Dachstein reefs of northern Europe. The Summit Point reefs do, however, bear a striking resemblance to some Middle Triassic (Anisian) reefs from the Tethys, particularly with the abundance of microbialite fabrics and diminutive, binding or encrusting bioconstructors.
... Endoliths began to radiate (Wood 2011) but typical boring traces were limited to Rogerella, Trypanites, Entobia and Gastrocheanolites, which are still the prevalent ichnogenera today (Knaust 2017). Encrusting taxa expanded as well, resulting in a greater importance of encrusting sponges, foraminifera (Bernecker 2005) and polychaetes (Senowbari-Daryan et al. 1993). ...
Preprint
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Hard substrate communities can impact coral reef growth by adding or removing calcium carbonate when they act as encrusters or bioeroders, respectively. Although such sclerobiont communities are known across the Phanerozoic, the Triassic saw a substantial increase in reef macrobioerosion. This study provides the first quantitative assessment of sclerobionts in a Late Triassic (Norian) reef coral community from the Nayband Formation in central Iran and establishes some post-Paleozoic trends in reef bioerosion. Sclerobionts were common on the dominant coral Pamiroseris rectilamellosa and covered between 0 and 26.4% of total coral surface area among the 145 colonies investigated. Encrustation was significantly more prevalent (1.78% of total area) than bioerosion (0.36% of total area). The underside of corals was 3.5 times more affected by sclerobionts than the surfaces. This suggests that the sclerobionts preferentially colonized dead parts of corals. The main encrusting taxa were polychaeta worms (76.2%), followed by bivalves (11.9%). The main bioeroders were of Polychaeta (51.4%), Porifera (22.2%), and Bivalvia (20.7%). The lack of a correlation between encrustation and bioerosion intensity suggests that the amount of sclerobionts is not simply a function of exposure time before final burial. Our review of the published literature suggests a declining trend in the relative importance of bivalves, polychaetas and cirripeds in reefs, whereas sponges increased in importance from the Triassic until today.
... Cyclic stratal patterns in the Dachstein Formation were described as Lofer cycles composed of members A, B, and C that were formed in supra-, inter-, and subtidal depositional environments, respectively, and interpreted to reflect orbitally controlled sea-level changes in peritidal deposits (Fischer 1964;Haas, 1994;Enos and Samankassou 1998;Haas et al. 2007). Lagoons of the inner Dachstein platform in the Northern Calcareous Alps were fringed by prolific reefs that harbored a rich biota (Bernecker 2005;Martindale et al. 2013): the Dachstein reefs faced the open oceanic Hallstatt Basin whereas on the opposing side the Rhaetian Oberrhätkalk reefs bordered intraplatform basins such as the Eiberg Basin (Mandl 2000). However, the timing and causes of demise of the Dachstein platform system remain controversial. ...
Article
Full-text available
The Dachstein platform was an extensive carbonate platform developed on the westernmost shelf of the Neotethys during the Late Triassic, now preserved in various tectonic units disrupted during the Alpine orogeny. Despite being the focus of a multitude of sedimentological, paleontological and other studies, the demise of this platform remains controversial, with contrasting views on the timing and causes of cessation of its growth, the duration of the gap above, which at many places includes the Triassic-Jurassic boundary (TJB), and the depositional environment of overlying strata. Here we present new carbonate sedimentological, stable isotope and cyclostratigraphic data from sections in the Transdanubian Range (Hungary) which capture the termination of uppermost Triassic Dachstein Limestone and the onset of Hettangian (Early Jurassic) sedimentation following a hiatus. Previously, the TJB in the Transdanubian Range was regarded as a textbook case of a tectonically-driven platform drowning event or, alternatively, cessation of carbonate production due to emergence caused by a significant sea level fall at the TJB. However, recognition of global biotic change and environmental perturbations at the TJB calls for an assessment of their possible role in the demise of the Dachstein platform. Oxygen and carbon isotopic composition of bulk carbonates were measured in sections at Kőris-hegy (Bakony Mts.), Tata (Tata Horst), and Vöröshíd (Gerecse Mts.) Paleogeographically, these three sections represent a proximal to distal platform transect. Other sections at Pisznice and Tölgyhát (Gerecse Mts.) yielded additional sedimentological data. The sharp surface separating the Dachstein Limestone from the overlying Jurassic formations carries no or only minimal relief at outcrop scale. Thin section studies reveal small-scale irregularities, stylolites, microborings with ferruginous filling, or a thin clay-rich layer at the TJB, indicative of a submarine, or perhaps polygenetic, hardground. In the first meters of the lowermost Jurassic beds abundant ooids occur, and crinoids become common. In each of the studied sections, an abrupt negative carbon isotope shift is recorded at the TJB, and a gradual rebound to more positive values characterizes the lowermost Jurassic strata. Chemostratigraphy allows correlation with sections elsewhere. In the Transdanubian Range, the initial carbon isotope excursion and at least the first part of the purported main carbon isotope excursion are not preserved due to the gap at the TJB. Combined bio- and cyclostratigraphy of lowermost Jurassic strata permits an astrochronologic duration estimate of the early Hettangian hiatus that was not longer than a few hundreds of thousand years. Our results highlight the role of submarine erosion, perhaps partly related to acidification, and point to an abrupt change in carbonate production related to the end-Triassic extinction of several groups in the platform system. “Unreefing”, the ecological collapse of reefs, led to a regime shift, the transformation of the rimmed platform to a carbonate ramp, with a significant gap in production and preservation of carbonate sediment. This model is not uniformly applicable to Late Triassic platforms as several of them, unlike the Dachstein platform, survived with unrimmed architecture in the Early Jurassic elsewhere on the Neotethyan shelf. However, the model may explain features of the carbonate platform sedimentary record across other events with reef collapse.
... They were first described at the end of the nineteenth century (e.g., Gümbel, 1872;Salomon, 1895) and then widely studied in the twentieth century and over the last twenty years (Pia, 1912(Pia, , 1920Bistricky, 1964;Ott, 1967Ott, , 1968Flügel, 1961Flügel, , 1975Senowbary-Daryan, 1980;Di Stefano, 1981;Barattolo et al., 1993Barattolo et al., , 2008Parente & Climaco, 1999;Senowbari-Daryan & Zamparelli, 2005;Grgasovič, 2007). Ouside of Europe, Late Triassic calcareous algae are known from Indonesia (Timor and Moluccas) (Vinassa de Regny, 1915;Pia, 1924), Eastern Tibet (Flügel & Mu, 1982), Turkey (Senowbari-Daryan & Link, 2005;Senowbari-Daryan et al., 2006), Iran (Senowbari-Daryan & Hamadani, 2000), and Oman (Berneker, 1996(Berneker, , 2005. In contrast with this relative abundance, only few manuscripts dealing with Triassic limestone deposits from North and South America mentioned or described calcareous algae. ...
Thesis
Full-text available
Over the geological times, shallow–water carbonate environments have always been important ecosystems where marine life abounds, constantly developing and evolving. Carbonate rocks, best witnesses of this marine life, are consequently precious relics to better understand the evolution of life through time and many other environmental aspects directly related. Limestones are therefore essential to characterize with precision and the collection of all its hidden information, whether biological, environmental, climatic or even geographic, is essential. This thesis work thus focuses on the investigation of marine shallow–water Upper Triassic carbonates, some never been studied before. The Triassic is a particular geological period, ranging between two major biological crises, including the great Permian/Triassic one, where almost 95% of marine species disappear. During the Triassic, life therefore strove to rebuild and spread across the oceans, with the rise of many new species, especially within benthic communities. From the end of the Middle Triassic (Ladinian), we witness an extensive development of the reef’s ecosystems, leading to the formation of numerous carbonate platforms, together within the Tethyan domain, partially landlocked in Pangea, and in the gigantic Panthalassa Ocean surrounding the unique continent. The Tethyan carbonates today crop mostly from Europe to Asia while the Panthalassic carbonates occur on the Circum–Pacific region, within accretionary complexes or terranes. Until the late 2000s, the Triassic shallow carbonates developed in Panthalassa were still very little known and studied, unlike the large Tethyan platforms accurately characterized for decades. Faced with this difference in information, the REEFCADE project was therefore developed by Rossana Martini in the 2000s, with the aim of improving our knowledge of these Panthalassic systems and more generally of the evolution of shallow–water carbonates during the Triassic. This thesis work has therefore been conducted as a part of this project and constitutes the logical continuation of various studies initiated by other doctoral students in the frame of REEFCADE. Thus, the study presented in this manuscript has the overall objective of improving our knowledge of carbonate systems in Panthalassa, especially during the Upper Triassic. For such purpose, this work focuses on the very first sedimentological, biostratigraphic, paleontological and diagenetic characterization of carbonates located in two distinct study areas: i) Hokkaido Island, in the northern part of Japan, and ii) the Dalnegorsk area in Russian Far East. These two localities are defined as a west–east succession of Mesozoic accretionary complexes, extending north–south, among which the Taukha terrane (Russian Far East) and the Naizawa Accretionary Complex (Hokkaido, Japan), characterized as Early Cretaceous in age. Mentioned only in very rare scientific publications, numerous outcrops of Triassic carbonates, never precisely studied before, have been reported within these two tectonic elements. During three field missions, the two areas, in Russia and Japan, were therefore extensively explored, and each carbonate outcrop was widely sampled. A total of ten localities, represented by dozens of limestone blocks, were thus investigated during this study. The thesis work mainly focuses on three localities where remarkably well–preserved carbonates were sampled: the Pippu and Esashi districts in Hokkaido (Japan) and the Dalnegorsk area in Russian Far East. Thanks to the thin sections made from each sample, a detailed microfacies analysis was performed to precisely characterize the fossil and abiotic contents for each outcrop. The biological assemblage could thus be compared with similar and synchronous systems, whether of Tethyan or Panthalassic affinity, in order to identify the potential similarities or differences between these two oceans in the scope of paleoecological analysis. In addition, each identified microfacies has been interpreted in term of depositional environment within the carbonate system, on the basis of organism’s type, sedimentary structures, nature of the lithoclasts or the content in carbonate mud. Within the framework of a biostratigraphic approach, a particular attention was addressed to the identification of benthic foraminifers in order to define or specify the stratigraphic extension of the studied carbonates. The combination of all the sedimentological and biostratigraphic data made it possible to reconstruct precise, even speculative depositional models for each investigated system. The Dalnegorsk limestone (Russian Far East), was thus characterized as an atoll–type carbonate system, developed during the Norian on a basaltic seamount in the Panthalassa Ocean, and typified by a great abundance of lagoonal deposits. On the other hand, the outcrops of Pippu and Esashi (Hokkaido Island, Japan) show very strong similarities in age and bioclastic content, and have therefore been interpreted as belonging to the same carbonate system. This last was defined as developed on the flanks of a partially emerged volcanic seamount during the Carnian. The Dalnegorsk limestone was also the subject of an in–depth diagenetic study, supported by in–situ and high–precision geochemical analyzes, in order to characterize each event which impacted the carbonate system, from its deposit to its accretion. Such study, never performed before on Panthalassic carbonates, has indeed made it possible to highlight important diagenetic episodes linked to major environmental changes during the history of the carbonate depositional system and to establish a precise model of evolution of the Dalnegorsk limestone. Analyzes of stable isotopes on various carbonate cements, coupled with trace elements measurements have especially highlighted an emersion of the atoll at the Norian/Rhaetian transition, followed by a dismantling of the flanks of the system during the Lower Jurassic as well as the neomorphism of calcitic shells at the onset of calcitic seas in Toarcian–Bajocian. In total, ten diagenetic events were identified and precisely placed in time thanks to a new method of U–Pb dating applied directly on carbonate cements. Numerous monogenic carbonate breccias have been observed in the field or on thin sections. The geochemical analyzes highlighted two types of breccias, hitherto impossible to differentiate, linked to distinct sedimentary processes and occurring at different stages of the evolution of the carbonate system. This thesis has also been the subject of many scientific collaborations within the REEFCADE project. The compilation of different data from similar past or ongoing studies has indeed led to two major paleontological and sedimentary syntheses. The very first paleontological study of calcareous algae in Panthalassa was thus carried out, leading to results of capital importance both for our understanding of paleoecology and paleogeography of the Triassic oceans but also to better apprehend how the benthic communities were able to spread within the huge Panthalassa Ocean. This study also resulted in the description of six new species of algae. Triassic carbonates from the Sambosan Accretionary Complex (SAC), located in the southern part of Japan have been the subject of numerous studies within the REEFCADE project. During this thesis work, a new sampling campaign, at the Mont Sambosan type locality outcrop (Shikoku Island, Japan), associated with a precise diagenetic study, made it possible to document for the first time the complete history of a carbonate system developed in Panthalassa, since its establishment during the ?Ladinian–Carnian until its dismantling at the Triassic/Jurassic boundary. To conclude, this thesis work constitutes a new major progression for our understanding of the Triassic carbonate systems from the Panthalassa Ocean. This new sedimentological, diagenetic, biostratigraphic and paleontological data complete a rich panel of information obtained within the framework of the REEFCADE project and permit to better constrain the environmental conditions within this vast ocean. The pioneer diagenetic characterization and the fundamental resulting outcomes, open the way to a similar and systematic exploration of synchronous systems to better define the different major environmental events which occur during the Late Triassic and the Jurassic. The combination of all these new data is also of paramount importance for paleogeographic and paleoecological studies, still poorly focused on the Panthalassa Ocean.
... They were first described at the end of the nineteenth century (e.g., Gümbel 1872; Salomon 1895) and then widely studied in the twentieth century and over the last twenty years (Pia 1912(Pia , 1920Bistricky 1964;Ott 1967Ott , 1968Flügel 1961Flügel , 1975Senowbary-Daryan 1980;Di Stefano 1981; Barattolo et al. 1993Barattolo et al. , 2008Parente & Climaco 1999;Senowbari-Daryan & Zamparelli 2005;Grgasovič 2007). Ouside of Europe, Upper Triassic calcareous algae are known from Indonesia (Timor and Moluccas) (Vinassa de Regny 1915;Pia 1924), Eastern Tibet (Flügel & Mu 1982), Turkey (Senowbari-Daryan & Link 2005;Senowbari-Daryan et al. 2006), Iran (Senowbari- Daryan & Hamadani 2000), and Oman (Berneker 1996(Berneker , 2005. In contrast with this relative abundance, only few manuscripts dealing with Triassic limestone deposits from North and South America mentioned or described calcareous algae. ...
Article
Upper Triassic calcareous algae, abundant and well-diversified in Tethyan deposits, have rarely been described in rocks of Panthalassan origin. Over the past ten years, several studies were performed on Upper Triassic carbonate deposits of Panthalassan affinity in North America, Japan and Far East Russia, revealing unexpectedly rich and diversified assemblages. The samples were collected from nine localities situated on both sides of the Pacific Oce- an. The identified algal assemblage consists of green and red algae, including fourteen dasycladaleans, rare bryopsida- leans, and several rhodophyceans. This paper describes the main algal taxa, including six new species: Holosporella? ros- sanae Bucur & Del Piero n. sp., Holosporella magna Bucur & Fucelli n. sp., Griphoporella minuta Bucur & Peybernes n. sp., Patruliuspora pacifica Bucur, Del Piero & Peyrotty n. sp., Patruliuspora oregonica Bucur & Rigaud n. sp. and Collarecodium? nezpercae Bucur & Rigaud n. sp. Rivulariacean-like cyanobacteria and thaumatoporellacean algae are also present. The whole Panthalassan algal assemblage comprises both unknown (?endemic) and common taxa of the Tethyan domain. To explain the cosmopolitan distribution of various Upper Triassic benthic organisms (scleractinian corals, calcified sponges, foraminifera), a close connection with the Tethys Ocean was hypothesized by different authors. During the Late Triassic, the Tethys was open to the east on the Western Panthalassa but not to the west, suggesting that Triassic calcareous algae were able to efficiently colonize environments that are estimated to be more than 10’000 km apart. An adventitious transport of calcareous algae and/or their spores is proposed to explain this long-range algal dispersal.
... They were first described at the end of the nineteenth century (e.g., Gümbel 1872; Salomon 1895) and then widely studied in the twentieth century and over the last twenty years (Pia 1912(Pia , 1920Bistricky 1964;Ott 1967Ott , 1968Flügel 1961Flügel , 1975Senowbary-Daryan 1980;Di Stefano 1981; Barattolo et al. 1993Barattolo et al. , 2008Parente & Climaco 1999;Senowbari-Daryan & Zamparelli 2005;Grgasovič 2007). Ouside of Europe, Upper Triassic calcareous algae are known from Indonesia (Timor and Moluccas) (Vinassa de Regny 1915;Pia 1924), Eastern Tibet (Flügel & Mu 1982), Turkey (Senowbari-Daryan & Link 2005;Senowbari-Daryan et al. 2006), Iran (Senowbari- Daryan & Hamadani 2000), and Oman (Berneker 1996(Berneker , 2005. In contrast with this relative abundance, only few manuscripts dealing with Triassic limestone deposits from North and South America mentioned or described calcareous algae. ...
... Due to their particular sensitivity to external forcing on depositional conditions, they are also excellent records of climatic, eustatic and geodynamic change. During the Late Triassic, numerous shallow-water carbonate platforms, both attached and isolated, developed in the Tethyan realm (Bernecker, 2005;Buser et al., 1982;Flügel, 1982;Gale et al., 2015;Gattolin et al., 2015;Jin et al., 2018;Krystyn et al., 2009;Martindale et al., 2014;Schäfer and Senowbari-Daryan, 1982;Tomašových, 2004). In the Panthalassean realm, Upper Triassic shallow-water carbonates are represented by atoll-type and attached systems, spread across the huge ocean ( Fig. 1), and later accreted in the circum-Pacific region (Chablais et al., 2010a(Chablais et al., ,b,c, 2011Basilone, 2020;Martini, 2018, 2020;Khalil et al., 2018;Onoue et al., 2009;Onoue and Stanley, 2008;Peybernes et al., 2015Peybernes et al., , 2016aPeybernes et al., ,b, 2020Peyrotty et al., 2020a,b;Rigaud et al., 2010Rigaud et al., , 2012Rigaud et al., , 2013bRigaud et al., , 2013aRigaud et al., , 2015bRigaud et al., , 2015aSano et al., 2012;Senowbari-Daryan et al., 2010). ...
Article
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Owing to their isolated oceanic setting, atoll-type carbonates are well suited for documenting carbonate deposition and diagenesis in oceanic environments away from continental influence. The atoll-type Dalnegorsk limestone (Taukha Terrane, Russian Far East), deposited in the gigantic but poorly-documented Panthalassa Ocean, preserves a complete record of the diagenetic evolution of an Upper Triassic system, out of the Tethyan domain. To study the diagenesis of this carbonate system, we developed a novel analytical workflow, combining cathodoluminescence petrography with high-resolution analyses of environmental proxies in calcitic cements (δ¹⁸O, δ¹³C, REEY, trace and minor/major elements) and in situ U–Pb dating of calcite cements to precisely reconstruct the chronology of the diagenetic events. We combined these lines of evidence to establish a model of atoll evolution, from deposition to dismantling, based on 10 identified diagenetic episodes. The Dalnegorsk limestone records emergence at the Norian-Rhaetian transition, marked by meteoric and evaporitic cements, followed by dismantling of the atoll edges after drawning in the Early Jurassic. Neomorphism of calcitic shells occurred at the onset of calcitic sea conditions during the Toarcian-Bajocian. The limestone was thoroughly cemented during the Middle/Late Jurassic, and accreted within the Taukha Terrane during the Late Jurassic/Early Cretaceous. Accretion resulted in fracturing, brecciation, and recrystallisation of the Dalnegorsk limestone. This model is potentially applicable to any similar atoll system, irrespective of age. The evidence presented here extends our knowledge of Late Triassic environments in the Panthalassa Ocean, and more generally, our understandingg of mid-oceanic limestone formation and evolution.
... Their vulnerability is undoubtedly a modern concern as ocean temperatures rise, but from a historical, fossil perspective, observing their occurrence within the context of environmental changes allows us to better understand their vulnerabilities and what aids their development and evolutionary success. The Triassic records an important ecological recovery for reefs after a massive reef collapse due to the end-Permian mass extinction (Bernecker 2005;Flügel and Kiessling 2002a, b;Martindale et al. 2019). During the Triassic, reefs undergo significant changes to their paleogeographic distribution, taxonomic composition, and overall diversity (Flügel 2002;Martindale et al. 2019). ...
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An important interval for reef evolution occurs in the Triassic Period. However, Late Triassic reefs along the edge of the Panthalassa Ocean receive much less attention than their Tethys region counterpart. To improve our understanding of reef evolution along the Panthalassa, we present a quantitative microfacies analysis to characterize reef facies types from a central Nevada locality. We point counted thin sections to quantitatively assess the relative abundance of reef and interstitial components from six Carnian reef mounds. Reef mound formation occurs at three different stratigraphic intervals. Each interval records a different reef mound size and shape. The oldest stratigraphic interval consists of four separate reef mounds; each mound is approximately 3–12 m wide and 2–4 m high. The second youngest stratigraphic interval consists of one large ~ 10-m wide and 15.5-m high domal reef mound. The third, and youngest, reef mound is smaller in comparison (i.e., 1-m wide and 1.5-m high). We recognize two reef facies types within these reef mounds. Different calcareous sponges, bound together by microproblematica and clotted micrite, form the overall reef framework. We suspect that changes in relative sea level affect the initial colonization, different growth capacities, and overall preservation of the South Canyon reef mounds. When South Canyon reefs are compared to other Panthalassa reefs, it seems that “Wetterstein-type” reef–sponge bindstone facies containing biogenic crusts dominate and corals are less abundant and persist generally in both shallow and deeper water environments. However, significant differences in primary and secondary framebuilder abundances exist due to differences in where the reefs grew in relation to fair-weather wave base (FWWB). Thus, changes in relative sea level may affect the initial colonization, different growth capacities, and overall preservation of reef but water depth may also control the primary and secondary reef builders. Consequently, quantitative assessment of reef ecosystems within a sequence stratigraphic framework undoubtedly facilitates fewer problematic comparisons between reefs but it also enhances our understanding of reef development.
... The sediments were deposited in a north-east, south-west oriented shallow marine intraplatform basin (Eiberg Basin) (Krystyn et al., 2005) that was bordered by land in the north-west (Fig. 3a, b), and by the Dachstein reef in the south-east (Fig. 3b, c). Patch reefs (Schäfer, 1979) and fringing reefs (Bernecker, 2005) occurred locally within the basin during the deposition of the Eiberg Member. The restricted nature of the basin during Rhaetian times is not fully constrained, and it has been proposed that deep-water connections to the open ocean in the south have existed via gateways through the Dachstein reef complex (Fig. 3b) (Golebiowski, 1990;Krystyn et al., 2005;Haas et al., 2010b;Mette et al., 2011). ...
Article
The Rhaetian (201–209 Ma, the latest stage of the Triassic) is an important time-interval for the study of environmental changes preceding the End-Triassic Mass extinction. A detailed sedimentological and chemostratigraphic study was conducted in the lower Kössen Formation at Hochalm (Austria), the type-section of the Hochalm Member (Mb). This section exposes mid-Rhaetian sediments deposited in an intraplatform shallow marine basin on the north-western margin of the Tethys. The study highlights eight apparent shallowing-upward sequences from the middle of Unit 2 to Unit 4 of the Hochalm Mb stacked within the long-term transgression that characterizes the Kössen Formation. Both the bulk carbonate and the bulk organic matter δ¹³C records indicate the presence of a distinct increase in carbon isotope values in the lower part of the lower Hochalm Mb. This excursion might represent a new chemostratigraphic marker that could be used for refining the Rhaetian stratigraphy and represents another important Late Triassic carbon-cycle perturbation prior to the major disturbance associated with the End Triassic biotic crisis.
... Al-Husseini, 1997; Sharland et al., 2001; Ziegler 2001). In Oman it has therefore only been possible to study the Upper Triassic shallow-water platform (Bernecker, 2005;, the Liassic shallow-water platform, unconformably overlying the Triassic (Bendias and Aigner, 2015) or the deep-sea record of the Upper Triassic to Lower Jurassic interval (Blendinger, 1988; Blechschmidt et al., 2004). The Musandam Peninsula hosts the stratigraphically most complete shallow carbonate platform section across this time interval on the Arabian Platform, although the exact stratigraphic boundary positions and hiatuses within the Late Triassic and Early Jurassic sequences in this region still remain unclear (Maurer et al., 2008;. ...
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The Triassic-Jurassic boundary is marked by one of the ‘big five’ mass extinctions of the Phanerozoic. This boundary event was accompanied by several carbon cycle perturbations, potentially induced by the opening of the Central Atlantic and associated volcanism, and accompanied by an ocean acidification event. Continuous carbonate successions covering this interval of environmental change are however rare. Here data from a shallow-marine equatorial mixed carbonate-siliciclastic succession is presented, that was studied on a regional scale. Four sections that are 48 km apart were examined on the Musandam Peninsula (United Arab Emirates and Sultanate of Oman). The system was analysed for its sedimentology, vertical and lateral facies changes, and stable carbon and oxygen isotopes. Strontium isotope analysis was used to determine the position of the Triassic-Jurassic boundary horizon. The studied ramp experienced an episode of demise during the Late Triassic, followed by a restricted microbialite dominated ramp, containing large amounts of siliciclastic facies. During the Latest Triassic the diverse carbonate factory revived and flourished across the Triassic-Jurassic boundary. No clear evidence for a biocalcification crisis or an ocean acidification event across the Triassic-Jurassic boundary is visible. Lateral facies heterogeneities can be observed across the studied interval, attributed to hydrodynamic activity, including tropical storms, crossing the extensive shelf area. Although evidence for synsedimentary tectonic activity is present, the vertical stacking pattern is largely controlled by changes in relative sea level. The refined chronostratigraphy accompanied by the detailed environment of deposition analysis allows for a refinement of the regional palaeogeography. The neritic equatorial carbonate ramp has archived a negative carbon isotope excursion preceding the Triassic-Jurassic boundary that has also been reported from other study sites. The lack of evidence for a biocalcification crisis across the equatorial Triassic-Jurassic boundary indicates that the Tethys did not experience a distinct global acidification event.
... ecosystem dynamics) processes (Chen & Benton, 2012;Dineen, Fraiser & Sheehan, 2014;Woods, 2014;Wei et al. 2015, and references therein). This recovery was characterized by slow reestablishment of metazoan frame-building communities in Anisian time while further diversification and proliferation occurred during the Ladinian age and especially during the Late Triassic epoch (Stanley, 1988;Flügel, 2002;Bernecker, 2005;Kiessling, 2010). Consequently, the global lacking or reduced activity of metazoan reef-builders was a major positive factor for the development of Early Triassic and early Middle Triassic carbonate ramps worldwide (Burchette & Wright, 1992), and in particular across the Western Tethys shelves (Calvet, Tucker & Helton, 1990;Michalík et al. 1992;Senowbari-Daryan et al. 1993;Rüffer & Zamparelli, 1997;Hips, 1998;Török, 1998;Zühlke, 2000;Rychliński & Szulc, 2005;Feist-Burkhardt et al. 2008). ...
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The Early to Late Triassic development of a carbonate ramp system in the subtropical belt of the NW Tethys was controlled by the interplay of several global and regional factors: geotectonic setting (slow continuous subsidence on a passive continental margin), antecedent topography (low-gradient relief inherited from preceding depositional regime), climate and oceanography (warm and dry climatic conditions, storm influence), relative sea-level changes (Olenekian to Anisian eustatic rise, middle Anisian to early Carnian sea-level fall), lack of frame-builders (favouring the maintenance of ramp morphology), and carbonate production (abundant formation of lime mud, non-skeletal grains and marine cements, development of diverse biota controlled by biological evolution and environmental conditions). Elevated palaeorelief affected the ramp initialization on a local scale, while autogenic processes largely controlled the formation of peritidal cyclicity during the early stage of ramp retrogradation. Probably fault-driven differential subsidence caused a local distal steepening of the ramp profile in middle–late Anisian time. The generally favourable conditions promoted long-term maintenance of homoclinal ramp morphology and accumulation of carbonate sediments having great maximum thickness (~500 m). Shutdown of the carbonate factory and demise of the ramp system in the early Carnian resulted from relative sea-level fall and subsequent emergence. After a period of subaerial exposure with minor karstification, the deposition of continental quartz arenites suggests the possible effect of the Carnian Pluvial Episode.
... Red coralline algae, so intricately associated with coral reefs of today, did not live on early Mesozoic reefs; instead red solenoporacean calcified algae were present (Bernecker 2005). Upper Triassic reefs, constructed by scleractinian corals and calcified chambered sponges, contained a plethora of organisms with no modern counterparts, including tabulozoans, chaetetids, spongiomorphs, chambered sponges, and many different "microproblematica" (Hodges and Stanley 2015). ...
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Although reef-like structures formed in the Neoproterozoic, reefs built by metazoans did not appear until the early Paleozoic. From then until the Recent, reefs diversified, underwent extinctions many times and then diversified again. Reef-inhabiting organisms included many different groups from algae to vertebrates as well as enigmatic, extinct suprageneric taxa. Evolution of these groups continued unabated and sometimes resulted in significant changes in the communities making up reefs. These reef groups varied over geologic time with extinction events commonly marking dramatic changes in the biotas. Paleozoic reefs consisted of sponges, corals, foraminifera, algae, bryozoans, and brachiopods, among others. The major extinction event at the end of the Paleozoic eliminated these forms as reef constituents and new groups (e.g., the first scleractinian corals) appeared in the Triassic. The Mesozoic was dominated by sponges, corals, rudist bivalves, and algae, most of which were eliminated in the end-Cretaceous extinction event. The Cenozoic reef biotas included red algae, foraminifera, sponges, corals, various invertebrates, and fish.
... Fürsich and Wendt (1977) name it as one of the frame builders at the Cassian patch reefs. Regarding Atrochaetetes, Bernecker (2005) assesses it as a builder for the Jabal Kawr reef from the Norian and Rhaetian from Oman. Thus, the presence of both taxa here and their assessment as part of the constructor guild is significant. ...
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Six fossil associations from 59 thin sections obtained out of 23 Cipit boulders of the Middle-Late Triassic St. Cassian Formation are described. Grouping and obtainment of these associations were performed considering all biomorpha and microbialites contained in the thin sections by means of cluster analysis in Q-mode using three algorithms and two indices. Further microfacial categories observed in the thin sections were allochthonous components, cements and microsparite, although they do not represent a significant fraction in any case. The associations include the following: Dendronella–Hexactinellida Association, Cassianothalamia Association I, Cassianothalamia Association II, Patch Reef Association I (Balatonia-microencruster Association), Patch Reef Association II (Solenopora–Dendronella Association) and Patch Reef Association III (Spongiomorpha ramosa Association). The possible palaeoenvironmental settings, from which each association is derived, are discussed individually. These settings were elucidated based on each fossil contained in the associations, since many of them are diagnostic for establishing specific environmental criteria. Although most associations are described here for the first time, some of them may be related to the “Cassian Patch Reefs”, while one of them (“Cassianothalamia-Gemeinschaft”) is reviewed.
... Tubiphytes: Tubiphytes is a major microbial reef building organism very common in the Permian (Sano et al., 1990;Kawamura and Machiyama, 1995), and Triassic (Russo et al., 1997;Bernecker, 2005). It occurs in association with metazoan reef builders, e.g., Scleractinia, or alternatively, with thrombolites. ...
... This sec- Fig. 2b (shaded), b -Detailed map with positions of the sections mentioned in the text, c -Structural subdivision of north-western Slovenia. Modified after Buser & Draksler (1993), Placer (1999), and Goričan et al. (2013) Mandl 1999;Krystyn et al. 2009) and the Oman Mountains (Bernecker et al. 2005). ...
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The Julian Alps (western Slovenia) structurally belong to the eastern Southern Alps. The Upper Triassic succession mostly consists of shallow water platform carbonates of the Dolomia Principale-Dachstein Limestone sys- tem and a deep water succession of the Slovenian Basin outcropping in the southern foothills of the Julian Alps. In addition to the Slovenian Basin, a few other intraplatform basins were present, but they remain poorly researched and virtually ignored in the existing paleogeographic reconstructions of the eastern Southern Alps. Herein, we describe a deepening-upward succession from the Tamar Valley (north-western Slovenia), belonging to the Upper Triassic Tarvisio Basin. The lower, Julian-Tuvalian part of the section comprises peritidal to shallow subtidal carbonates (Conzen Dolo- mite and Portella Dolomite), and an intermediate carbonate-siliciclastic unit, reflecting increased terrigenous input and storm-influenced deposition (Julian-lowermost Tuvalian shallow-water marlstone and marly limestone of the Tor For- mation). Above the drowning unconformity at the top of the Portella Dolomite, Tuvalian well-bedded dolomite with claystone intercalations follows (Carnitza Formation). The latter gradually passes into the uppermost Tuvalian—lower- most Rhaetian bedded dolomite with chert and slump breccias, deposited on a slope and/or at the toe-of-slope (Bača Dolomite). Finally, basinal thin-bedded bituminous limestone and marlstone of Rhaetian age follow (Frauenkogel For- mation). The upper part of the Frauenkogel Formation contains meter-scale platform-derived limestone blocks, which are signs of platform progradation. The Tarvisio Basin may have extended as far as the present Santo Stefano di Cadore area, representing a notable paleogeographic unit at the western Neotethys margin.
... Another specific feature of the ramp sedimentation was the absence of large buildups such as frame-built reefs and reef mounds (terminology after Tucker and Wright, 1990) which are known from Triassic Peri-Tethyan carbonates in Spain, Poland and Germany (Hagdorn, 1978;Bodzioch, 1989;Calvet and Tucker, 1995;Szulc, 2000) and were more abundant in strictly Tethyan environments (Senowbari-Daryan et al., 1993;Rüffer and Zamparelli, 1997;Flügel and Senowbari-Daryan, 2001;Bernecker, 2005). The delayed post-Permian global recovery of carbonate-producing organisms was characterized by reestablishing of frame-building communities not until Anisian time except for microbial reefs with further diversification and proliferation taking place during the Ladinian and Late Triassic ages (Senowbari-Daryan et al., 1993;Hallam and Wignall, 1997;Flügel, 2002;Pruss and Bottjer, 2005;Kiessling, 2010). ...
... Norian-Rhaetian sponge and coral associations of the Tethys are best developed in Northern Calcareous Alps where they formed reef belts in platform-edge positions facing the openmarine northwestern Tethys basins, or as smaller patch reefs in intraplatform basins or on the platform ramps (Bernecker, 2005). We compare Norian-Rhaetian reefs of Iran with water depths similar to the Northern Calcareous Alps but some of the Nayband patch reefs more closely resemble older Upper Triassic counterparts in Nevada and Sonora, Mexico which were also influenced by an influx of siliciclastic sediments (Senowbari-Daryan and Stanley, 1992;Goodwin and Stanley, 1997). ...
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The Nayband Formation is one of the best known sedimentary units in central Iran. The type section consists of a thick succession of shale, siltstone, reef limestone and sandstone that is subdivided into five distinct members: Gelkan, Bidestan, Hoz-e-Sheykh, Howz-e- Khan and Qadir. Abundant and well-preserved framework-building scleractinian corals are included among the macrofossils of the Nayband Formation; these corals characterize the formation and are the subject of this study. The Hassan-Abad section, located in northeast Iran in Lute Block (northwest of Ferdows city), was chosen for detailed study and sampling. Analysis of sedimentary lithofacies and faunal assemblages in the Bidestan and the Howz-e-khan members indicate both biostromal and biohermal characters for the former shallow-water patch reefs and support a Norian to Rhaetian age. The useful biostratigraphic hydrozoan Heterastridium conglobatum was studied along with 14 taxa of scleractinian corals: Stylophyllopsis rudis, Distichophyllia norica, Paradistichophyllum dichotomum, Retiophyllia frechi, Retiophyllia norica, Retiophyllia robusta, Chondrocoenia schafhaeutli, Chondrocoenia ohmanni, Astraeomorpha crassisepta, Astraeomorpha confusa, Astraeomorpha minor, Procyclolites triadicus, Pamiroseris rectilamellosa, and Eocomoseris ramosa. These fossils clarify the stratigraphy of the Nayband Formation, as well as provide new information on the patch reefs and the framework constructors of these reefs.
... The development of Alpe di Specie reefs appears to have marked a reversion to more open frame-reef structure. Certainly, by the Late Triassic scleractinian corals were becoming major reef builders (Flügel, 1994), locally developing substantial frameworks (Weidlich et al., 1993; Bernecker, 2005). ...
... Ma) saw massive reef and carbonate accretion in the Northern Tethys Ocean (now the limestones and dolostones of the Northern Calcareous Alps). The Northern Tethys Ocean was a semi-enclosed ocean basin with numerous, large carbonate platforms and warm water circulating northward from the tropical Tethys (Flügel and Senowbari-Daryan, 2001;Flügel, 2002;Bernecker, 2005) and thus was an ideal location for large reef growth. ...
Article
In the Late Triassic (~ 235–201.3 Ma) scleractinian corals and hypercalcified sponges built large, diverse reef ecosystems, the most famous of which are the Dachstein reefs. This study presents a depth transect along an Upper Triassic (lower Rhaetian) Dachstein reef from the Gosausee margin of the Dachsteingebirge (Gosau, Austria). The Gosausee microbial-sponge-coral reef is a complete barrier reef with an almost continuous fore reef to lagoon transect preserved (a very rare occurrence for the Triassic), and thus provides a window into depth zonation of Dachstein-type reef facies and biotic succession. The Gosausee reef facies exhibit strong depth control and five classic reef facies or zones are identified: the fore reef, reef front, reef crest, back reef, and lagoon facies. Thin, rare microbial fabrics and a high abundance of fine-grained, mud-rich skeletal wackestones (transported reef debris) characterize the deepest fore reef. As the reef shallows, muddy sediments decrease in abundance and are replaced by microbial fabrics, corals, and cements. Abundant sponges, microbial crusts, and thick, marine cements typify the reef crest, whereas microbialite-coated phaceloid corals are dominant in the back reef facies. Heavily cemented oncoids or microbial-sponge bindstones are characteristic of the lagoon. Based on their compositional and biotic similarities, the Gosausee reef was likely part of the same barrier reef systems as the source reef for the Gosaukamm reef breccia (one of the classic Norian–Rhaetian Dachstein reefs). The reef zones of the Gosausee margin can be used to interpret the depth or reef zone of less well preserved reef fragments, can inform models of community ecology and niche utilization in the Late Triassic, and highlight the need for additional research into the environmental factors that controlled biotic distribution in Upper Triassic reefs.
... The dominance of platy corals (Fig. 4), and the presence of an intergrown, rigid coral fabric (in places framework sensu stricto) make the Dunlop and Cinnabar Canyon reefs unique: they are the only currently known Upper Triassic reefs dominated by dense platy coral growth fabrics. While platy corals are not uncommon in the Late Triassic, the branching, phaceloid corals typically dominate most Upper Triassic reefs (Schäfer, 1979;Flügel, 1981;Wurm, 1982;Stanley and Senowbari-Daryan, 1986;Flügel, 1987, 1989;Bernecker et al., 1999;Senowbari-Daryan, 2001, Flügel 2002;Bernecker, 2005). ...
Article
Upper Triassic (Lower Norian) reefal buildups near Mina, Nevada, represent some of the earliest scleractinian coral reefs from eastern Panthalassa. The small patch reefs (~20–40m high and ~50–150m wide, obvious meter-scale elevation above the surrounding sediments) are from the Luning Allochthon and grew on an inner to middle ramp or in a deep lagoon. The Mina patch reefs were constructed by several different coral ecomorphotypes (platy, tabular, domal/massive, and branching corals) interpreted to have had zooxanthellate symbionts, and record subtle coral zonation within the reefs. Based on modern coral ecomorphotypes, platy to tabular corals at Mina grew in the lower euphotic zone (stressed by low light conditions), and the massive or domal corals inhabited shallower water (possibly above fair weather wave base) and were stressed by wave energy.Unlike most other Upper Triassic reef ecosystems where phaceloid branching corals or calcareous sponges constitute the principal bioconstructors, the platy to tabular corals were the primary builders in the Mina patch reefs. The Mina reefs are also unique because cryptic and cavernous internal environments, epibionts, cryptobionts, and thick microbial crusts are rare or absent. The combination of platy coral dominance with the lack of epibionts/cryptobionts/microbial crusts suggests that the Norian reefs from Mina, Nevada represent a unique form of scleractinian reef construction from the Late Triassic.
... Interestingly, the Late Triassic appears to have been a founding period for the primary diversification of various, unrelated marine calcifiers, including calcareous dinoflagellates (Streng et al., 2004) and scleractinian corals (Stanley, 2003). Although the evolution of carbonate skeletons in scleractinians occurred during the Anisian, scleractinian corals expanded ecologically as reef builders during the Late Triassic taking over from the Middle Triassic and Carnian sponge-dominated reefs (Bernecker, 2005;Flügel, 2002;Payne et al., 2006). Geochemical evidence suggests that the Carnian-Norian rise of scleractinian corals was closely related to the acquisition of symbiotic dinoflagellates. ...
Article
On the basis of a global carbon-climate numerical model, we reconstructed atmospheric carbon dioxide concentrations over the first 60 million years of the Mesozoic. Our analysis indicates that pCO 2 declined from more than 3000 ppmv to less than 1000 ppmv, with a drastic drop during the Late Triassic. The fast northward drift of Pangea exposed a large continental surface to warm and humid equatorial climate, thus promoting CO 2 consumption through weathering. This massive drawdown of atmospheric CO 2 is consistent with sedimentological and geochemical data of the rock record and correlates with the primary radiation of calcareous nannoplankton, a biological revolution shifting the global carbonate sink from shallow water environments to the open oceans. Our numerical model shows that at time, tectonics, via weathering, increased the pH of the oceanic surface waters by 0.3 units, corresponding to a 50% decrease in H + concentration. This may have provided the ultimate environmental trigger which unlocked the newly oxidized Mesozoic open oceans to pelagic nannobiocalcification.
... In the Tethys, A. friedli is known from numerous Carnian-Rhaetian localities such as Austria (Koehn- Zaninetti and Brönnimann, 1968;Hohenegger and Piller, 1975b;Piller, 1978), Slovakia (West Carpathians, Salaj, 1969;Salaj et al., 1967Salaj et al., , 1983, Bulgaria (Trifonova, 1993), Spain (Pyrenees, Frechengues and Peybernés, 1991), Italy (Northern and Southern Apennines, Zamparelli et al., 1995;Parente and Climaco, 1999), Turkey (Domuz Tag, Brönnimann et al., 1970;Cetmi Melange, Beccaletto et al., 2005), Cyprus (Martini et al., 2009), Oman (Bernecker, 2005), Australia (Wombat Plateau, Zaninetti et al., 1992), and Indonesia (Sulawesi, Martini et al., 1997). ...
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The benthic foraminifer Aulotortus friedli is reported from the Upper Triassic carbonates (capped seamount) of the Sambosan Accretionary Complex in Kyushu, Southwest Japan. Specimens were collected from shallow-water limestone clasts in a debris flow sequence near the Ose locality along the Kumagawa River. The limestone clasts mainly represent shoal facies where the benthic foraminifer is extremely abundant. Because of the remarkable preservation of the Japanese specimens, they allow further description of the involutinid A. friedli. In addition, a detailed sedimentary and environmental analysis of the gravitational deposits where A. friedli occurs is given. Using microfacies analysis together with the biostratigraphic range of A. friedli and associated foraminifers, a new interpretation is proposed of a seamount collapse process that occurred during the Norian-Rhaetian and preceded the major collapse event of the Sambosan seamount, which has been placed in the Middle to Late Jurassic.
... 1,[57][58][59][60][61][62][63][64][65][66] B. Lathuiliè re and D. The development of highly integrated taxa (meandroid and thamnasterioid) is generally taken as an indication for photosymbiosis (Coates and Oliver, 1973;Coates and Jackson, 1987) and is linked to the development of reefs especially in the Rhaetian and Bajocian. The end-Triassic extinction corresponds to the collapse of reefal ecosystems known on both sides of the Tethys (Bernecker, 2005). Starting from Hettangian (lowermost Jurassic), reefs do occur (e.g. ...
Article
Recognizing extinction events and determining their cause at the Triassic/Jurassic (T/J) transition and near the Pliensbachian–Toarcian (Lower Jurassic) boundary is a field of growing interest. We provide arguments for these events through a literature based new evaluation of coral diversity from Triassic to Dogger and a new palaeobiogeographical map. The T/J extinction of corals is clearly related to the breakdown of reef environments. Origination curves show that Hettangian (the lowest Jurassic stage) was not only a survival phase but already rather a recovery phase. Post-extinction evolution of reefs and their survival only in the northernmost margin of the Tethys support the hothouse hypothesis for the T/J extinction event. During Pliensbachian, many new taxa appear, but mostly solitary corals, not really framebuilders. Many of these taxa do not occur anymore during the following stages. The new increase in diversity is related to the development of Bajocian (Middle Jurassic) reefs.
... The main part of the isolated Kawr platform includes reef facies (section Sint, 59–113 m), inferred to represent the platform rim as well as bedded inner platform facies (Fig. 2), section Ala and Amqah. Bernecker (1996 Bernecker ( , 2005). ...
Article
In the oceanic realm of the southern Tethys, carbonate production of isolated platforms ceased after the end-Permian mass extinction and did not recover until the Late Triassic.The Misfah Formation (MF) at Jebel Kawr in the Oman Mountains is interpreted as a relic of such an isolated Late Triassic platform of the Hawasina Ocean, a part of the Neo-Tethys. Correlation of three sections at Jebel Kawr points to a sequence architecture with four third-order sequences (MF1–MF4). The maximum flooding surface (mfs) of MF3 can be correlated to the attached Arabian platform.The shallow-water carbonates of Jebel Kawr comprise a platform rim reef facies and bedded inner-platform facies characterized by stacked high-frequency cycles with subtidal to intertidal carbonate sequences.The depositional profile of this Late Triassic isolated platform evolved during Carnian and Norian time from a low-relief carbonate bank to a high-relief platform rimmed by reefs. The onset of the carbonate sedimentation is characterized by an initial phase with volcaniclastic interruptions, followed by a carbonate bank stage with a shallow subtidal to peritidal interior and marginal oolite shoals. In the Norian vertical accumulation caused an increase of the platform height and developed a relief along the margins that progressively increased through the aggrading reef stage. The possibility that a reef rim existed and was later removed by erosion is suggested by the Sint reef and olistoliths of similar reef limestones in the surrounding areas.
... The overall sedimentology and biostratigraphy of Triassic deposits are well established in the western Tethyan realm and various studies including those dealing with paleogeography and geodynamic models are known: Italy and Sicily (Miconnet et al. 1983; Zaninetti et al. 1984; Senowbari-Daryan et al. 1985; Carcione et al. 2003), Greece (Schäfer and Senowbari-Daryan 1982), Austria (Hohenegger and Piller 1975; Enos and Samankassou 1998; Bernecker et al. 1999), Slovakian and Polish Carpathians (Michalik 1982; Gazdzicki 1983), Oman (Bernecker 1996Bernecker , 2005 Weidlich et al. 1993; SenowbariDaryan et al. 1999), Arab Emirates (Maurer et al. 2008; Senowbari-Daryan and Maurer 2008 ), Indonesia (Shaibani et al. 1983; Villeneuve et al. 1994; Martini et al. 1997 Martini et al. , 2004), Philippines (Kiessling and Flügel 2000) and Australia (Tollmann and Gramann 1992; Zaninetti et al. 1992). Triassic reefs are particularly well studied (e.g., Kanmera 1964; Flügel 1981; Schäfer and SenowbariDaryan 1981, 1982; Flügel 1982 Flügel , 2002 Flügel and Stanley 1984; Stanley and Gonzalez-Leon 1997; SenowbariDaryan et al. 1999; Bernecker et al. 1999; reported on the distribution of megalodonts and other bivalves. ...
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The Upper Triassic shallow-water limestones of the Sambosan Accretionary Complex are reconstructed as a remnant of a mid-oceanic atoll-type build-up upon a seamount in the Panthalassan Ocean. The Sambosan atoll-type carbonates and its pedestal were accreted along with deep-water ribbon-chert and related siliceous rocks to the eastern margin of Asia during the Late Jurassic to Early Cretaceous. Studied limestones crop out in southern Kyushu Island, southwest Japan. Although the prevailing and intense deformation during the accretionary process prevents measurement of sections in stratigraphic successions, and sedimentary structures are poorly preserved, microfacies description and foraminifers analysis allow us to speculate the depositional setting of the Sambosan limestones. Seventeen microfacies are distinguished and several foraminifers of Tethyan affinity are identified. Foraminifers indicate a Late Carnian to Rhaetian age. The Tethyan affinity of the macro- and microfaunas suggests that the Sambosan seamount was located presumably in a low- to middle-latitudinal zone of the southern hemisphere during the Late Triassic.
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In order to study the hypercalcified sponges in reefal deposits of the Nayband Formation in Lut Block, the Garm Ab section near the village of Mehran Kushk, located about 20 km northeast of Ferdows city, was sampled. Eight horizons of reefal limestone beds are exposed in this section. The most important reef builders are hypercalcified sponges with some representatives of hexactinellids, scleractinian corals and other reef organisms. The field and lab-observations on rock units, sedimentary facies and faunal assemblages indicate the middle Norian-Rhaetian as the age of the reef horizons. Twenty-three sponge taxa, including 15 of the chambered sphinctozoans, 2 of hexactinellids sponges and 8 non-chambered inozoan were identified. The majority of recognized sponges are reported from the Nayband Formation from the other localities in central Iran. One new species identified as Cryptocoelia maxima n. sp. was recovered and is described here.
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The Late Triassic foraminiferal genera Amphorella, Spiriamphorella, Urnulinella, Pseudocucurbita, Paratintinnina, Costifera and Siculocosta are considered junior synonyms of the genus Cucurbita. Consequently, the number of families of the Milioliporacea (Miliolina) is significantly reduced. The valid species of the genus Cucurbita are considered to be Cucurbita infundibuliforme Jablonský, 1973, C. subsphaerica (Borza & Samuel, 1977a) comb. nov., C. longicollum Senowbari-Daryan, 1983, C. battagliensis (Senowbari-Daryan, 1983) comb. nov., C. cylindrica (Senowbari-Daryan, 1983) comb. nov. and C. floriformis (Altiner et al., 1992) comb. nov.
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Supercalcified sponges, including sphinctozoans, inozoans, chaetetids, spongiomorphids, occurring in Upper Triassic (Norian-Rhaetian) shallow-marine limestones of Musandam Mountains in United Arab Emirates (UAE), are described. The following taxa were determined: sphinctozoans: Hajarispongia osmani Senowbari-Daryan and Yancey, Nevadathalamia arabica n. sp., Nevadathalamia conica n. sp., Fanthalamia milahaensis n. sp., Iranothalamia incrustans (Boiko), Cinnabaria regularis n. sp.; inozoans: Cavsonella triassica n. sp., Molengraaffia regularis Vinassa de Regny, Peronidella? sp., Circopora cf. caucasica Moiseev, Circopora? sp.; spongiomorphids: Spongiomorpha sp.; chaetetids: Lovcenipora chaetetiformis Vinassa de Regny, Lovcenipora musandamensis n. sp., Lovcenipora sp., chaetetid sponge gen. et sp. indet. The most abundant sponge in the studied material is Nevadathalamia arabica n. sp. The described sponge association of the Arabian shelf (Musandam Mountains) shows close affinity to the sponge association known from age-equivalent terranes in the Panthalassa Ocean (Sonora Mountains in Mexico; Pilot Mountains in Nevada, USA), but is remarkably different from sponge associations in carbonates bordering the Tethys. This difference goes along with the biogeography of wallowaconchid bivalves and is most likely attributed to climatic, palaeogeographic or oceanographic factors.
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Significantly different Guadalupian–Late Triassic patterns are observed in the evolution of attached and isolated carbonate platforms of the southwestern Tethys (Oman Mountains). Close to the rim of the attached Arabian platform, carbonates of the Saiq and Mahil Formations reveal an almost complete Permian–Triassic sedimentary record. Guadalupian–Changhsingian 3rd order sequences consist of fossiliferous transgressive systems tracts and monotonous highstand systems tracts with mud/wackestone and coral bafflestone. The youngest Changhsingian beds are bioturbated floatstone with crinoids, sponges and bryozoans. All sediments indicate a healthy, tropical carbonate production. Above, a unique facies change begins with a pyrite-encrusted omission surface. Greenish mudstone rich in authigenic pyrite infills the relief of the unconformity and is overlain by clastic sediment and by laminated, microbialite-bearing carbonate. Unfossiliferous sediments and seafloor cements indicate a change in carbonate production towards abiotic processes. Prevailing anoxic conditions were interrupted by seven oxic event beds, as indicated either by low-diversity and small-sized ichnotaxa or by shell beds with low-diversity bivalve and crinoid assemblages. By comparison with published data, the described sedimentary sequence can be assigned to the Changhsingian–earliest Griesbachian. Beginning probably with the Anisian, bioturbated Griesbachian–Dienerian recovery period and the unconformity below to the latest grain-supported sediment textures mark the return to biogenic tropical carbonate production under oxic conditions. The Middle–Late Triassic carbonate platform consists of stacked high-frequency shallowing upward cycles. By contrast, carbonate production of Neo-Tethyan isolated platforms was discontinuous and interrupted by a large gap. Guadalupian deposits of the Al Jil Formation consist of bioclastic limestone typical of a tropical carbonate production. The uppermost bed, an impoverished bioclastic packstone capped by an unconformity, marks the onset of platform drowning which resulted from the end-Guadalupian mass extinction. Above, a polymict breccia witnessed rift pulses of the Neo-Tethys. The overlying pelagic mud- and packstone contains radiolarians and rare foraminifera of Lopingian age, and overlying microbialites. In the Carnian, tropical shallow-water carbonate production restarted with a low-relief platform and culminated in a Norian–Rhaetian reef-rimmed platform. Stacked Lofer cycles dominated the inner platform of Jebel Kawr (Misfah Formation). We here propose a differential onset and severity of the Late Permian mass extinctions for carbonate platforms. On the Arabian Plate, tropical carbonate production collapsed after the end-Lopingian mass extinction and was replaced by microbialites and sea-floor cements during the earliest Triassic. After approximately six million years, tropical shallow-water carbonate production resumed in the Middle Triassic. Neo-Tethyan isolated platforms drowned shortly after the end-Guadalupian mass extinction and did not recover before the Late Triassic. Absence of shallow-water limestone suggests that carbonate production of isolated platforms ceased for about 30 million years, a period exceeding the recovery of most marine ecosystems.
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Two chambered sponges - A. agahensis nov. sp. and Musandamia omanica nov. gen., nov. sp. - from the Norian reefal limestones of Jebel Agah, southeast of the Musandam Mountains, Oman, are described. Both species are the first reported from the Triassic of the Arabian Peninsula. The fossil association is also mentioned.
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In this paper are presented sedimentological and paleontological data concerning the bioconstructed platform margin facies of many different Norian-Rhaetian carbonate successions on Italy. The paper mostly consists of an updating of already separately published data plus additional ones from Northern Calabria. This presentation is the result of a coordinated research project financed by the CNR, aimed tothe comparison of all the Upper Triassic microbial-serpulid dominated bioconstructions discovered in Italy in the last fifteen years. Final scope was to envisage the common paleotectonic and paleoceanographic factors controlling the genesis of these peculiar biofacies. The investigated areas comprise Southern Alps (Lombardy Basin to the west and Carnia Basin to the east), a limited sector of the Austroalpine (Ortles), Northern Apennines (Monte Cetona), Central Apennines (Monti Simbruini) and Southern Apennines (Monti Lattari, Monti Picentini, Monti di Praia and Monte Montea). For each area, the geological setting, the stratigraphy of the Upper Triassic successions and the bioconstructed facies are described. In the case of Monte Montea area (Northern Calabria) the stratigraphy evolution from the Ladinian-Carnian depositional system to the Norian-Rhaetian carbonate platform with microbial-serpulid-sponges biofacies is here presented for the first time. The comparated study of all the occurrences indicate that microbial communities associated with serpulids were the main constructors in most Norian-Rhaetian carbonate buildups in that part of the shallow western Tethys Gulf characterized, during the Norian, by the Dolomia Principale depositional system. The buildups developed on the margin of small-sized, relatively shallow intraplatform basins. More limited buildups are also found in the mixed carbonate-siliciclastic ramps that characterize the Upper Norian-Rhaetian of Northern Apennines and Lombardy. Two main kind of platform-basin transitions have been described: rimmed carbonate platform, developed in the Lower-Middle Norian in the Northern areas and possibly persisting up to the Upper Rhaetian in Central and Southern Apennines. This type of system can be further subdivided in progradational and aggradational/backstepping ones; homoclinal siliciclastic carbonate ramps in the Upper Norian – Rhaetian of Western Alps and Northern Apennines. These intraplatform basins developed along a belt, extending from Southern Alps through the Apennines until the Betic Cordillera, affected by extensional-transtensional tectonics since Lower Norian. Production of the carbonate platform areas occurred from Uppermost Norian, owing to climate change and tectonic dissection. These tectonics slightly predate the Jurassic Piedmont-Ligurian Ocean opening, which occurred immediately to the north and west of this belt. In all the localities the intraplatform sediments are represented by laminated and resedimented doloarenites and calcarenites, alternating with abundant breccia deposits. Dark basinal sediments characterize the most distal areas: they are often rich in organic matter and lack any evidence of benthic life. The mound complexes, up to several hundreds of meter thick, developed on the other platform margin-upper slope. They formed at a depth comprised from just above the wave-base level, to the very quiet water of the middle-lower slope, possibly down to a depth of some 200 meters. The main biomass of the buildups was largely dominated by microbialites, but their peculiarity is the widespread presence of serpulids. In the associated facies little sized organisms (bivalvs, brachiopods) are frequent, together with some dasycladacean algae. Only in northern Calabria little sphinctozoans become a significant builders group. The dominance of microbialites and serpulids was clearly related to marine conditions unsuitable for the development of coralgal reefs. The comparison of recent and past occurrence of microbial and serpulid buildups with the sedimentological characteristics common to all the studied occurrences, indicated that low oxigen concentration in sea water and mesosalinity appear the most reliable mechanism to explain the Norian buildups. Eutrophy of water mass, due to high particulate input, is more likely for less widespread Upper Norian to Rhaetian buildups. Microbial-serpulid bioconstructions formed a minor, yet significant ecological province in the shallow western Tethys, as opposed to the to highly diversified, sponges and coral dominated Dachstein reefs present in Eastern Alps and Sicily and widespread all along the southern Tethys margin as far as to Australia. While the latter developed in connection with open marine basins related to the oceanic areas of the Tethys, the microbial-serpulid buildups are systematically associated to narrow troughs with restricted circulation and limited exchange with open marine basins. Only in Eastern Carnia and Northern Calabria, the character of the observed marginal biofacies suggest a closer proximity to an open marine area. This is in agreement with those paleogeographical reconstructions which envisage a deep marine area separating Sicily from Southern Italy. Key Words: Upper Triassic, microbialites, serpulids, intraplatform basins, platform margin, mixed ramps, buildups, geodynamics, paleogeography, dysoxic environments, Italy, Alps, Apennines.
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Carnian reef biota and brachiopods of the Misfah Formation (Hawasina complex) from a locality near the town of Al Aqil in Oman are described. A new brachiopod species, Oxycolpella arabica n. sp. is described. The composition of the Carnian reef biota in Oman is similar to those known from the Alpine-Mediterranean region. However, differences exist in the microfauna of reef dwellers, e.g. foraminifers. New stratigraphic data based on reef organisms and conodonts are presented.
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The paper presents the distribution and taxonomy of the Rhaetian coral fauna in the Tatric, Fatric, Hronic and Silicic paleogeographical zones of the Western Carpathians. The taxonomic spectrum of the corals from the Fatric zone is the most typical of the Carpathians, and resembles that of the Alps in the abundance of reimaniphylliid corals and frequent phaceloid growth forms, but it differs from the Alpine spectrum in its lower generic diversity. The assemblage from the northernmost zone, formed by the Tatric Superunit, is closest to the Early Jurassic fauna from the British Isles. Two new species are described: Zardinophyllum carpathicum sp.n. and Stylophyllopsis bobrovensis sp.n.
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Fischer's classic study (1964) of the Upper Triassic “Lofer” cyclothems in the Dachstein Limestone of the Northern Calcareous Alps was seminal to many studies of the Dachstein and to carbonate cycles globally.Fischer's idealized cycle is deepening upward, ABC in his terminology, where member A is a surface or interval of subaerial exposure, B is tidal deposits, and C is shallow subtidal. Studies of the Dachstein from the elsewhere in the northern Alps have substantiatedFischer's upward-deepening ABC cycle, butGoldhammer et al. (1990) andSatterley (1996a) reinterpreted the type Lofer cycles as shoaling upward. We measured 139 m of Dachstein Limestone incorporating 25 cycles at Steinernes Meer, Austria, nearFischer's most extensive section. In this section we identified no A members. The section is punctuated by slightly reddish horizons (‘pink partings’) that in some cases may reflect brief subaerial exposure, but generally appear to be pressure-solution zones that have concentrated iron oxides but lack a distinctive isotopic signature. B members are readily distinguished by fenestral porosity, stromatolitic lamination, partial dolomitization, intraclasts, or desciccation cracks. They are relatively thin (5 to 155 cm; median thickness is 38 cm.) and in some cases laterally variable or discontinuous. C members are characterized by molluscan wackestones and packstones with diverse biota. C intervals are 25 cm to 26 m thick (median 4.1 m) and comprise 91% of the interval measured. Pervasive bright-red internal sediment, which appears commonly within the B and C members, does not derive from any interval observed within the measured section, but from sources, possibly paleosols, much higher in the section. It is spatially associated with near-vertical, ENE-trending (62o) fractures filled with the sediment, brachiopods, and cement. Such fractures cut stratigraphic intervals as thick as 70 m without an exposed top or base.
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In the Late Triassic, an extremely large carbonate platform system (Dachstein-type platforms) developed on the margin of the Neotethys. On the wide inner platform cyclic peritidal, lagoonal successions were deposited. In the Transdanubian Range (Hungary), the lower part of the 1.5–2-km-thick cyclic succession (Upper Tuvalian–mid-Norian) is pervasively dolomitised, the upper part (Upper Norian–Rhaetian) is non-dolomitised; there is a transitional interval between them made up of partially dolomitised cycles. The peritidal–lagoonal cycles are commonly bounded by well-developed disconformity surfaces reflecting subaerial erosion that punctuated the marine carbonate accumulation. Truncation of the cycles was preceded by pervasive cementation of the previously deposited cycle. In the early stage of the platform evolution, tidal flat dolomitisation under semi-arid conditions led to the consolidation of the previously deposited sediments. The truncation surfaces were commonly covered by dolocretes. During the more humid Late Norian–Rhaetian period, the early cementation was followed by karstification, accumulation of wind-blown dust and pedogenesis. Erosion during regularly recurring subaerial exposure that commonly reached the previously deposited subtidal beds suggests eustatic control of the cyclicity and supports the application of an allocyclic model, even if the Milankovitch signal is imperfect.
Chapter
Four Upper Triassic patch-reefs exposed in the Northern Limestone Alps of the Salzburg area of Austria were studied. Emphasis was placed on facies development and paleoecologic zonation. The Adnet Reef structure grew directly on a carbonate platform in a shallow water carbonate setting; the Rotelwand and Feichtenstein Reef complexes grew out of the Kossen Basin in 2 distinct stages. This study has demonstrated that calcareous algae, various microproblematica, and foraminifers show very distinct distributional patterns within the reef complexes, and can be used as facies indicators as well as differentiating different biotopes within the central reef areas.-from Authors
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Late Triassic (Norian-Rhaetian) limestones, 1,000 to 1,500 meters thick, consist of interbedded lagoonal limestones (90 percent) and intertidal-supratidal dolomitic limestones (10 percent). The belt was originally some 20 km wide, extending some 250 km from the Loferer and Leoganger Steinberge in the west, through the region of Berchtesgaden and Salzburg, toward Vienna. It is bounded on the south by a belt of reefs defining the southern edge of the Dachstein bank, and on the north by the Hauptdolomit ultrabackreef facies. The deposit is well exposed in the many mountain faces and high plateaus of the region; lower and readily accessible exposures south of Salzburg include cuts and a quarry near Golling (Pass Lueg), and at Berchtesgaden, the road from Obersalzberg to Hitler’s “Eagle’s Nest” on Mt. Kehlstein. Rocks are moderately deformed, with primary porosity lost by cementation.
Article
Obertrias-Korallen (Karn-Unterstes Nor) aus dem Pantokrator-Kalk von Hydra (Griechenland) Zusammenfassung Aus dem Pantokrator-Kalk der Insel Hydra (Griechenland) werden 24 Korallenarten, zugehörig zu 14 Genera, beschrieben. Zwei Genera und 7 Arten sind neu: [Palaeastraea mandrakiensis n.sp., Stuoresiafluegeli n.sp., Conophyllia hellenica n.sp., Hydrasmilian.Q.: H. rhythmica n.sp., H. fossulata n.sp., H. ornamenta n.sp., Craspedophyllia graeca n.g.n.sp.). Die Korallen bestätigen das Alter der Lokalitäten als Kam bis unterstes Nor. Die Arten können meist mit südeuropäischen Lokalitäten (Italien, Slowenien, Südungarn, Rumänien, Türkei) verglichen werden und scheinen in südlichen Flachwasserberei-chen der Tethys vorzuherrschen. Trotzdem, beinahe ein Drittel der neuen Arten spricht für spezielle und etwas differenzierte Lebensbedingungen in Hydra während der karnischen Periode. Abstract From the Pantokrator limestone of the Island Hydra, Greece, 24 species of corals belonging to 14 genera are described. Two genera and seven speciesare new (Palaeastraeamandrakiensisn.sp., Stuoresiafluegelin.sp., Conophylliahellenican.sp., Hydrasmilian.g.: H, rhythmican.sp., H. fossulatan.sp., H. ornamenta n.sp., Craspedophyllia graeca n.g.n.sp.). Corals confirm Carnian to lowermost Norian age of localities. Coral species can mostly be compared with south European localities (Italy, Slovenia, south Hungary, Romania, Turkey), and seem to predominate in southern shallows of Tethys. Nevertheless, almost one third of the new species indicate special and somewhat different environments in Hydra during the Carnian period.
Article
Summary The Late Triassic limestones of the Steinplatte near Waidring at the Austrian/German boundary have become a classical example of an ancient reef, formed at a carbonate shelf margin (“framebuilt reef rim”). Evaluation of the stratigraphic framework and the investigation of cliff wall sections, together with the analysis of microfacies, paleontological and geochemical data, result in a very different interpretation of the depositional history. A “non-reef” model is favoured: Bioclastic deposition on a carbonate ramp grading into a slope. The Steinplatte is made up of three depositional units: 1) Kössen facies; 2) mound facies, the apparently massive carbonates of the Oberrhätkalk; 3) capping facies, the upper-most strata of the Oberrhätkalk. The Kössen facies comprises predominantly fine-grained lithobioclastic packstones. Changes in microfacies types and in sedimentary structures indicate an increase in water energy trough time. Local biostromal “mounds” (“A-, B-, and C-reefs”; mounds at the base of the Steinplatte cliff) were formed at the same stratigraphical level. Influence of these low relief mounds on the sedimentation in adjacent areas was subordinate except for a mound at the southwestern corner of the Steinplatte acting as a mounding nucleus on the carbonate ramp of the Kössen facies. The mound facies makes up the bulk of the Oberrhätkalk of the Steinplatte. It consists predominantly of fine- and medium-grained bioclastic carbonate sands, shell coquinas and some thin, small-scaled autochthonous boundstone structures (predominantly calcisponge limestones). Patches of dendroid corals occur in the lowermost mound facies. The source of most coral bioclasts and fine-grained coral debris, occurring within various levels of the mound facies, probably were coral patches high on the slope or on a platform. Several shell beds as well as a conspicuous “White Bed” can be used as time lines in the reconstruction of the depositional events. Sedimentation on a depositional slope is indicated by the predominance of northward dipping beds in the area of the western cliff wall. To the south (west of Wieslochsteig) horizontal bedding becomes more evident, pointing to a change from ramp to platform edge type sedimentation. The capping facies comprises thick-bedded strata dipping off and wrapping around the northern and eastern sides of the mound-shaped surface of the mound facies. These strata were deposited sobsequent to a change in sea level on the moundshaped surface of the mound facies. The capping facies differs from the mound facies in attidude, increase in bioclastic coated grainstones with abundant algae and in the occurrence of “coral gardens” representing a veneer on parts of the mound facies. Sea-level fluctuations are indicated by repeated vadose diagenesis and by distinct upslope-down-slope zonations of the biota. The general facies model of the Steinplatte is shown in Fig. 33. It corresponds to a development from a carbonate ramp (prior to the deposition of Shell Bed III) to a slope, probably grading into a platform to the south. The non-reef character of the mound facies of the Steinplatte limestone is accentuated by the lack of organic frames (in the coral thickets), the lacking influence of calcisponge frames on adjacent sediments, lack of ecological successions, and by the predominance of skeletal sediment consisting of mollusk, echinoderm and allochthonous coral debris, in contrast to only small amounts of frameworks. General implications of the Steinplatte study to the investigation of ancient “reefs” concern the interpretation of “potential frame-building” organisms as actual frame-builders and the interpretation of so-called “reef-detritus”.
Article
Zusammenfassung Im Hangenden der Kössener Schichten sind in den Nördlichen Kalkalpen zahlreiche kleine Riffe (“patch-reefs”) eingeschaltet, die in der Literatur verschieden benannt werden und im allgemeinen als “Oberrhät”-Riffe bekannt sind (vgl. Fabricius 1966:20; Tollmann 1976:258). Mehrere dieser Riffe befinden sich im Land Salzburg (Osterhorngruppe). Vier nahe beieinander liegende Riffe bildeten das Thema von zwei Dissertationen im Paläontologischen Institut der Universität Erlangen-Nürnberg. P. Schäfer bearbeitete das Rötelwand-Riff im Mörtelbachtal und das Adnet-Riff bei der Ortschaft Adnet (beide nahe Hallein, vgl., P. Schäfer 1979). In der vorliegenden Arbeit werden die Untersuchungsergebnisse des Feichtenstein- und des ca. 3 km südlich gelegenen Gruber-Riffes vorgestellt (österreichische Topographische Karte 1∶25000, Bltt Hintersee, 94/2; 47° 42′, 25″;−13° 17′ 50″ bzw. 47° 41′ 10″;−13° 18′, 14″). Der Schwerpunkt der Untersuchungen liegt auf der paläontologischen Bearbeitung der riffbildenden Organismen. In Abschnitt 4.3.1 findet sich eine übersichtsliste der festgestellten Arten.
Article
The Carnian Leckkogel beds represent a carbonate sequence, which was deposited in front of an actively spreading carbonate platform. The typical Leckkogel beds are characterized by highly diverse sponge buildups. Sphinctozoan sponges predominate together with rare bryozoans and hydrozoans as well as red algae and thin crusts of cyanophyceans. Foraminifera are very rare and dasycladaceans are missing altogether. Oncoidal limestones containing predominantly inozoans are coeval with the Leckkogel beds. They represent a short-lived colonization of the sea-floor within a basinal environment. In addition to this autochthonous development, allochthonous intercalations of debris-flow sediments occur. The basinal facies consists of shales and argillaceous schists. Towards the edge of the platform the Leckkogel beds interfinger laterally with the Tisovec limestone, which exhibits two main facies types, a reef facies and a lagoonal facies.
Article
Carbonate platform deposits record a complex interplay of numerous geodynamic variables, of which eustasy, subsidence, and sediment accumulation are prime factors in determining both the kilometer-scale (depositional sequence scale) and meter-scale (depositional cycle scale) stratigraphic packaging. In this study, we looked particularly at composite eustasy, that is, superimposed sea-level fluctuations with different frequencies (defined as orders) and different amplitudes, and the role it plays in the linkage between meter-scale cyclic stratigraphy and kilometer-scale sequence stratigraphy. Specifically, we have investigated the relationship between low-frequency, third-order (1-10 m.y. period) depositional cycles and their component high-frequency, fourth-(0.1-1 m.y.) and fifth-(0.01-0.1 m.y.) order cycles through detailed stratigraphic analyses of Alpine Triassic platforms, complemented by computer modeling. On the basis of our results, we suggest that in general, there exists a hierarchy of stratigraphic forcing driven by composite eustasy that results in organized stacking patterns (thickness, subfacies character, early diagenetic attributes) of high-frequency, typically fourth- and fifth-order, shallow-water carbonate cycles dictated by low-frequency, third-order relative sea-level effects. We suggest that systematic vertical changes in stacking patterns of high-frequency cycles across a larger depositional sequence are due to systematic and predictable differences in depositional space available during the rising and falling stages of a relative third-order sea-level change. We also suggest that these systematic variations in cycle stacking patterns will exist regardless of the mechanism responsible for generating the high-frequency cycles, be it an autocyclic or allocyclic mechanism. This approach has major implications for the use of high-frequency, fourth- and fifth-order cycle characteristics to identify third-order cycles in outcrops and cores of shallow-water carbonates, where stratigraphic control may be less than desirable. This would constitute a valuable bridge between cyclic stratigraphy at the meter scale and sequence stratigraphy at the seismic scale. We present two examples from Triassic buildups of the Alps (the Ladinian Latemar buildup and the Norian Dolomia Principale) where a systematic succession of high-frequency cycle stacking patterns and early diagenetic features exists within an overriding third-order cycle (sequence) reflecting the interplay of short-term, high-frequency (fourth, fifth order) eustasy and long-term, low-frequency (third order) eustasy in accordance with the hierarchy of stratigraphic forcing. Central to the interpretation of these examples is the demonstration that true eustatic rhythms are recorded in the high-frequency cyclicity, as verified by time-series analyses and the use of "Fischer plots." These examples can be modeled by computer under conditions of lag-depth-constrained sedimentation, uniform subsidence, and composite eustasy. We also present two examples, one from the Alpine Triassic (the Norian Lofer cyclothems) and one from the Pleistocene of south Florida, that lack both a systematic succession of high-frequency cycle stacking patterns and identifiable composite rhythms in the stratigraphic record, despite the existence of composite high-frequency eustasy. In these examples, we call upon (1) local tectonic forcing in the form of short-term deviations in subsidence via faulting (Lofer example) and (2) large differences in the relative amplitudes of different orders of high-frequency sea-level oscillations (Pleistocene example) to explain the lack of composite rhythms, and we present computer simulations to illustrate the concepts. An understanding of composite relative sea-level changes and the potential for a hierarchy of stratigraphic forcing provides the link between cyclostratigraphy and sequence stratigraphy and also has important implications regarding the stratigraphy of early diagenesis.
Article
The Upper Triassic massive limestone (the Oberrhdtkalk) at the Steinplatte, Austria, formed along the edge of an intrashelf basin that was situated within a shallow-marine to supratidal shelf approximately 40 km wide. This transition from shelf to intrashelf basin is marked by a distinct slope break that has been widely cited as an example of a reef-rimmed shelf. It is, instead, a distally steepened ramp. The Oberrhdtkalk consists of the ramp sediments lying between the Dachsteinkalk of the shelf and the Kössen Formation of the intrashelf basin. The ramp interpretation is based on the geometry and lateral facies progression in the profile from the shallow-marine to peritidal shelf, across the ramp, and into the basin, and on the absence of reef or grainstone deposits at the slope break typical of a rimmed shelf.The Oberrhdtkalk at the Steinplatte is an important example of shelf-margin deposition because: 1.(1) It formed in a protected and low-energy intrashelf basin setting, thus expanding the range of distally steepened ramps that have been used to develop a general model.2.(2) It differs from previously described examples of distally steepened ramps, largely from open-ocean shelf-margin settings, in that: (a) it evolved constructionally from a homoclinal ramp rather than forming at a drowned shelf edge or an antecedent tectonic structure; (b) the sediments of the distal slope are largely autochthonous; (c) slumping and breccia deposits on the distal slope are of minor importance; turbidites on the basin floor are absent; and (d) skeletal packstone and grainstone characterize the outer ramp and slope rather than mudstone and wackestone.3.(3) It provides an alternative ramp model for other Upper Triassic shelf and platform margins in the Northern Calcareous Alps, which have commonly been interpreted as reef or sediment rimmed.
Article
The Upper Triassic (predominantly Norian) Dachstein limestones in the southwestern Gesäuse Mountains (Styria, Austria) exhibit different facies types characterized by significant distributional patterns of organisms and nonskeletal grains. Most valuable facies-diagnostic organisms are foraminifera and calcareous algae. The quantitative distribution patterns of the foraminifera can be used to separate facies units and local biotopes within the reef-complex. Thecentral reef facies (widespread in the Kalbling reef and Schildmauer reef) is defined by an autochthonous reef-framework (composed of corals, calcisponges, algae, and many epizoans), which interfingers with quantitatively prevailing reef debris. The reef-framework consists of various biocoenoses (high growing corals, low growing corals, calcisponges, hydrozoa and bryozoa, solenoporacean algae). Within the organic framework the foraminiferaAlpinophragmium perforatum, Ophthalmidium sp.,Agathammina sp. as well as sessile miliolids and arenaceous forms are most typical. In small cavities of the framework internal micritic sediments with peloids can be found, together with a characteristic association of microfossils (foraminifera:Ophthalmidium martanum, O. leischneri, Sigmoilina sp.,Galeanella sp., “Lituosepta”; microproblematica:Baccanella floriformis, Muranella sphaerica). Thefore-reef facies can be studied south of the Kalbling reef. This facies type is characterized by sparry rudstones composed of angular litho- and bioclasts which have been derived from the central reef. In some parts an unusual interfingering of the fore-reef facies and lagoonal sediments can be seen indicating the existence of channels within the reef-complex. North of the central reef area thegrapestone facies of the back reef environment is developed: Non-skeletal particles are grapestones, algal lumps, and coated grains. The algal flora is characterized byDiplopora, Heteroporella and some porostromate cyanophyceans (type “Cayeuxia”). Among the foraminiferaAulotortus communis, A. tumidus, Duostomina sp. and “Tetrataxis” are frequent elements. With increasing distance from the central reef area the grapestone facies interfingers with the “algal-foraminifera facies” showing similar nonskeletal components but different associations of algae and foraminifera.Griphoporella andGyroporella are the prevailing algal genera. The foraminifera fauna is characterized byAulotortus sinuosus, A. communis andQuinqueloculina sp.. This facies is found also in small depressions and channels within the grapestone facies. The “pelletal-mud facies” and the “mud facies” represent the lagoonal environments of the reef-complex. Both facies types occur in greater distance from the central reef area than the algal-foraminifera facies and the grapestone facies. In the limestones of the pelletal-mud facies peloids, fecal pellets, coated grains and a specialized foraminifera association (Aulotortus friedli, A. tenuis, Endothyra sp.,Endothyranella sp.,Trochammina sp.,Agathammina sp.) are common. Peloids and fecal pellets are found also in the rocks of the mud facies, together with small, often thin-shelled foraminifera (Aulotortus friedli, A. tenuis, lagenids). The “loferite facies” interfingers with both facies units. Pelletal loferites and stromatolithic algal loferites can be distinguished. The distributional patterns of thegeochemical data (CaCO3, MgCO3, SrCO3, rare elements like Fe, Mn, Zn and Cu, insoluble residues) show no correlations with the facies types of the limestones except strontium which seems to be concentrated in micritic limestones rather than in sparitic grainstones and rudstones.
Article
Summary During the Middle and early Late Triassic carbonate ramps and rimmed platforms developed at the northwestern margin of the Tethys ocean. In the Northern Calcareous Alps, Anisian stacked homoclinal ramps evolved through a transitional stage with distally steepened ramps to huge rimmed platforms of Late Ladinian to Early Carnian age. Middle Triassic to early Late Triassic facies and biota of basin, slope and platform depositional systems are described. Special emphasis is given to foraminifers, sponges, microproblematic organisms and algae. The Ladinian to early Carnian reef associations are characterized by the abundance of segmented sponges, microproblematica, biogenic crusts and synsedimentary cements. Among the foraminifers, recifal forms likeHydrania dulloi andCucurbita infundibuliformis (Carnian in age) are reported from the Northern Calcareous Alps for the first time. Some sphinctozoid sponges likeParavesicocaulis concentricus were known until now only from the Hungarian and Russian Triassic.
Article
Susammenfassung Die obertriadischen Reffstrukturen von Adnet und der Rötelwand bei Hallein eatwickelten sich als kleine patch-reefs in einer Lagunen- bzw. Beckensituation in geschützter lage im Norden einer ausgedehnten Plattform mit karbonatischer Flachwasserfazies. DasRötelwand-Riff liegt vollständig isoliert innerhalb des Kössener Beckens, welches sich im Norden an die Dackstein-Plattform anschlie\t und durch kontinuierliche Absenkung eines Wattengebietes mit Hauptdolomit-Plattenkalk-Sedimentation in Norden hervorgegeangen war. Es ist allseitig von Kössener Beckensedimenten umgeben. Die Entwicklung der Rötelwand-Riffstruktur erfolgte aus dem Kössener Becken heraus in mehreren, deutlich voneinander unterscheidbaren Stadien. Die Entwicklung wird in erster Linie durch das Wechselspiel von Wasserenergie und Sedimentation gesteuert. Sie beginnt mit einem Tiefer-Wasser-Stadium mit zunächst rein mechanischer, dann mechanisch-biologischer Sedimentanhäufung und Ausbildung eines Karbonatschlammhügels (mud mound-Stadium). Hierauf folgt ein Flachwasser-Stadium mit der Ausbildung mächtiger Lumachellenbänke, welche die Basis für das Wachstum eines echten, ökologisch definierten Riffes bilden (Riff-Stadium). Durch lokale Sedimentanhäufung wächst die Riffstruktur kontinuierlich aus dem Bereich Schwächerer Strömungen am Grunde eines Beckens in eine Zone mit verstärkter Wasserbewegung hinein, so da\ sich zuletzt in einem Bereich mit Turbulenzen ein echtes Riffgerüst ausbilden kann. Eine vergleichbare Entwicklung aus Kössener Beckensedimenten heraus ist imAdneter Riffkomplex nicht zu beobachten. Die das Riff unmittelbar unterlagernden Sedimente sind nur in randlichen Teilen der Riffstruktur aufgeschlossen. Aus der abweichenden Ausbildung und räumlichen Verteilung der Sedimenttypen im basalen und in oberen Riff-Stadium und aus den Unterschieden im Aufbau des Riffgerüstes wird geschlossen, da\ sich der Adneter Riffkomplex auf einer Plattform mit Dachstein-Fazies (Algen-Foraminiferen-Detritus-Fazies) entwickelte. Die Plattform tauchte nach N und E allmählich ab, und die Flachwasser-Sedimente verzahnen sich in einer detritischen Übergangsfazies (MF-Typ III/C) mit den Kössener Beckensedimenten. Aufgrund der tektonischen Isolation der aufgeschlossenen obertriadischen Gesteine kann die Frage nicht beantwortet werden, ob die dem Riffkomplex unterlagernden und sich mit ihm lateral verzahnenden Flachwassersedimente des Sockels mit der ausgedehnten Dachstein-Plattform in Süden in Verbindung standen. Das Fehlen einer deutlichen, durch einheitliche Strömungsverhältnisse bedingten, horizontalen Zonierung der Fazies-Einheiten im Riffstadium der Entwicklung zeigt aber, da\ die Ausbildung der Adneter Riffstuktur sehr viel stärker durch die schützende Nähe der Dachstein-Plattform beeinflu\t wurde als diejenige der Rötelwand. Diefazielle Gliederung der beiden Riffstrukturen lä\t im Reifestadium ihrer Entwicklung 5 sich lateral miteinander verzahnende Fazies-Einheiten erkennen, die im Rötelwand-Riffkomplex eine deutliche Zonierung, im Adneter Riffkomplex dagegen eine zwar ebenfalls gesetzmä\ig angeordnete, aber eher fleckenhafte Verteilung aufweisen. Jede Fazies-Einheit entspricht einem Ablagerungsraum, welcher durch bestimmte mikrofazielle, biologische und sedimentologische Merkmale typisiert ist. Die übergeordnete Fazies-Einheiten können aufgrund des Vorherrschens einzelner Organismen und Sedimenttypen noch weiter differenziert werden, so da\ sich insgesamt15 Mikrofazies-Typen (MF-Typen) ergeben. Während die Gültigkeit der Fazies-Einheiten über einen regional und zeitlich gesteckten Rahmen hinaus geht und an keine Riffstruktur gebunden ist, nimmt ihre Untergliederung in Mikrofazies-Typen ausschließlich Bezug auf die beiden untersuchten Riffgebiete. 1. Die Biolithit-Fazies des zentralen Riffareales ist auf den Bereich der Riffknospen mit organisch gewachsenem Rifgerüst beschränkt. Dieses ist durch in-situ stehende Riffgerüstbildner und eine starke biogene Anlagerung gekennzeichnet. 2. Die onkoidische Riffdetritus-Fazies (Onkoid-Fazies) dringt mit parautochthonem Riffschutt zwischen den Riffknospen in den zentralen Riffbereich vor. Ihr Hauptbildungsraum liegt aber in exponierter Lage am oberen Riffhang in unmittelbarer Nähe zur Biolithit-Fazies. Im Rötelwand-Riff ist sie auf die Süd-Seite des Riffes beschränkt und zeight damit seine Luv-Seite und die isolierte position der Riffstruktur an. Im Adner-Riff sind die Riffknospen allseitig von der Onkoid-Fazies umgeben. Die Sedimente dieser Fazies entstehen unter hochenergetischen, flachmarinen Bedingungen. 3. Die Algen-Formainiferen-Detritus-Fazies entwickelt sich im Rötelwand-Riff im Süden aus der Onkoid-Fazies und leitet in die Kössener Beckenfazies über. Im Adneter Riffkomplex nimmt sie ein ausgedehntes Areal ein und stellt vermutlich das Fundament dar, auf welchem sich die Riffknospen entwickelten. Im Süden verzahnt sie sich mit den Onkoidkalken als zeitgleicher Ausbildung zur Biolithit-Fazies. Die Ablagerungen könen mit ähnlichen, in Subtidalen Bereich entstandenen Sedimenten der riffernen Dachsteinkalk-Fazies verqlichen werden. 4. Die Detritus-Schlamm-Fazies kennzeichnet den strömungsgeschützten Riffhang in unmittelbarer Nachbarschaft zur Biolithit-Fazies und verzahnt sich im Rötelwand-Riff allseitig mit der Kössener Becken-Fazies. Im Adnet-Riff besitzt die Fazies nur fleckenhafte Verbreitung im Norden des Gebietes. 5. Die Kössener Becken-Fazies weist eine Fülle verschiedener Faziestypen auf, deren Ausbilding und Verteilung in Bezug steht zum Relief des Meeresbodens. In nahezu allen Sedimenten (eine Ausnahme bilden die Ooidkalke) überwiegt der Anteil an Kalkschlamm. Die Fazies umgibt das Rötelwand-Riff allseitig und verzahnt sich im N und E mit den Adneter Riffkomplex. Daszentrale Riffgebiet des Rötelwand-Riffes bildet ein weites Areal mit lockerem, aber gesetzmä\ig angeordnetem Bestand von Riffbildnern. Es können nur wenige Riffknospen, von einander getrennt durch Riffschuttareale, unterschieden werden. Innerhalb der Riffknospen sind Teilbereiche durch das Vorherrschen bestimmter Assoziationen von Riffgerüstbildnern charakterisiert. ImAdneter Riffkomplex setzt sich das zentrale Riffareal aus zahlreichen Riffknospen zusammen, deren jede eine eigene Interngliederung erkennen lä\t. Im Zentrum der Struktur stehen sie derartig dicht nebenund aufeinander, da\ der reine Riffschutt zwischen ihnen vollständig fehlen kann. Korallen, Kalkschwämme (überwiegend Spinctozoen), Hydrozoen (Disjectoporen und Spongiomorphiden), Tabulozoen und Kalkalgen sind mit rund 60 Arten, darunter zahlreichen neuen Formen, an der Korstruktion des Riffgerüstes beteiligt. Während das Adnet-Riff als echtes Korallen-Riff bezeichnet werden kann, in welchem eine Art, die gro\kelchige Varietät vonTheocosmilia clathrata EMMRICH, dominiert und alle anderen Riffbildner nur eine sekundäre Rolle spielen, besitzen im Rötelwand-Riff Korallen und Sphinctozoen nahezu die gleiche Bedeutung bei der Besiedlung des zentralen Riffareales. Die Diversität der Rifforganismen des Rötelwand-Riffes übertrifft deutlich diejenige des Adneter Riffes. ImRötelwand-Riff überwiegen heterotypische Organismengemeinschaften. Im Adnet-Riff sind dagegen in allen grö\eren, stets polymikten Riffknospen auffallend häufig auch homotypische Riffgemeinschaften an ihrem Aufbau beteiligt. Fassen wir die wichtigstenRiff-Assoziationen in beiden Riffgebieten zusammen, so lassen sich insgesamt9 Vergesellschaftungen unterscheiden. Jede von ihnen ist charakterisiert: 1. durch die Assoziation der Riffgerüstbildner, 2. durch ihren Sekundärbewuchs von Epi- und Endobionten, 3. durch eine signifikante Foramineferen-Assoziation, und 4. durch einen speziellen Sedimenttyp (MF-Typ I/A bis D) als Füllsediment in den Zwischenräumen des Riffgerüstes. Die Anordnung der Riffknospen im zentralen Riffareal und ihre Interzonierung werden durch ökologische Faktoren gesteuert. Der Grad der Wasserenergie nimmt deutlichen Einflu\ auf die Verteilung der Rifforganismen und des Sedimentes. Das gesetzmä\ige Auftreten der Rifforganismen in bestimmten Arealen des Riffbereiches gibt Hinweise auf die physiologischökologischen Anforderungen welche die Organismen an ihren Lebensraum und an die Teilglieder ihrer Assoziationen stellen. Als sehr brauchbar haben sich die vagilbenthonischen Foraminiferen nicht nur zur Typisierung der übergeordeneten Fazies-Einheiten (d.h. Ablagerungsbereiche), sondern auch als begleitende Fauna einzelner Riff-Assoziationen erwlesen. Ihre quantitative Verteilung zeigt, da\ diese in stärkerem Ma\e von der Sedimentverteilung als von der Art der Riff-Assoziationen abhängig ist.
Article
The Upper Triassic reefal limestones of the Oman Mountains were investigated with respect to their microfacies, palaeontology and community structure. The reef fauna described and figured for the first time occurs in parautochthonous slope deposits of the Arabian platform (Sumeini Group) and in allochthonous reefal blocks (‘Oman Exotics’, Hawasina Complex). The ‘Oman Exotics’ are tectonically dislocated blocks, derived from isolated carbonate platforms on seamounts in the Hawasina basin or in the South Tethys Sea. The lithofacies and fauna of these blocks comprise a cyclic platform facies with megalodonts, reef and reef debris facies. The reefal limestones are dated as Norian/Rhaetian by benthic foraminiferal associations (Costifera, Siculocosta, Galeanella) and typical encrusting organisms (Alpinophragmium, Microtubus). Some small ‘Oman Exotics’ are of Carnian age. The shallow-marine organisms include scleractinian corals of different growth forms, ‘sphinctozoans’, ‘inozoans’ chaetetids, spongiomorphids, disjectoporids and solenoporacean algae as the main reef builders, various encrusters like microbes, foraminifers, sponges and many different problematical organisms for the stabilisation of the reef framework and a group of dwellers including benthic foraminifers, gastropods, bivalves and a few dasycladacean algae. The reef communities are characterized by the coverage of organisms and distributional pattern. Analogies with the coeval reef deposits from the European part of the Tethys have been recognized. Some species, now collected in Oman, were also reported from American and Asian localities.
Article
Deposition of the mostly Mesozoic Sumeini Group occurred on the slope between the shallow-marine, Arabian carbonate platform and the deep-oceanic Hawasina Basin or South Tethys Sea. These strata record the evolution of the northeast Arabian continental margin from Permian(?) and Triassic rifting to ocean basin closing with the Late Cretaceous obduction of the Semail Ophiolite. Platform margin type and its evolution can be inferred from exposures of these slope sediments at Jebel Sumeini in the Western Oman Mountains.
Article
The paper is concerned with some key questions resulting from current studies of Triassic reefs and reef biota. A survey of the distribution of the reefs in time and space (Figs. 1–2) indicates the existence of Anisian buildups (starting in the Pelsonian), and significantly restricted to the southern part of Europe. Ladinian and Carnian reefs exhibit a larger distribution pattern, including Europe, Asia and western North America as well as Peru. Even broader is the distribution of Norian and Rhaetian reefs, which are known from many parts of the Tethys but have been studied in detail only in Europe, Central Asia and in western North America, The exact age of many Triassic reefs is controversial because of the strong facies control of reef biota. The composition of the framebuilding fauna (especially calcisponges and corals) can only differentiate an Anisian to Carnian time interval from a Late Upper Triassic interval. The current state of research is characterized geographically by strongly biased information (more than 75 % of the Triassic reefs studied in more detail are situated in the Alpine-Mediterranean region and in the Cordilleran of western North America). Information about the composition of the framebuilding and binding communities as well as about facies types is generally good, but there is a strong need for more study of the reef-dwelling organisms and especially of the diagenetic history of reef carbonates. Triassic reefs exhibit five principal reef categories, which are distinguished by morphology and paleogeographical setting: 1. Carbonate ramps with biostromes, 2. Mud Mounds. 3. Reef Mounds, differentiated into knoll reefs and patch reefs. 4. Atolls. 5. Barrier Reef Complexes. Fig. 4 may be used to classify Triassic reefs with respect to morphology and paleogeographical location of the buildups. Most Triassic reefs described up to now are situated in a shelfedge position and in back-platform positions. Foreslope positions and platform margin settings were formed prior to platform and back-platform settings. The palecological interpretation of the reef biota is made more difficult by the rarity of systematic paleontological studies of many Triassic reefs. Difficulties arise even for the main reefbuilding groups: Calcisponges (only the European sphinctozoans have been studied in detail), corals (the problem of the beginning of coralzooxanthellae symbiosis should be clarified), and algal crusts as secondary framebuilders (without Recent counterpartsl). The relative importance of reefbuilders changes throughout time (Figs. 5–6); this is indicated by the different abundances of primary and secondary framebuilders, and also by different reef communities. Upper Triassic reefs seem to be characterized by at least six main communities, whereas Anisian to Carnian reefs exhibit eight communities. Only a few associations may be regarded as long-lived units. Reef communities reflect ecological successions characterizing various stages in the formation of organic buildups (stabilization, colonization, diversification, and domination). A general trend can be seen in the existence of a vertical sequence showing predominantly encrusting and low-diverse biota at the base of the reef formation, and more diverse but vertically zoned biota within the higher levels. Numerical and faunal diversity of reef biota indicate the existence of regular gradients over different parts of Triassic reef complexes. Diversity seems to be different in coeval reefs of different paleogeographical position (Fig. 7) and also changes over time (higher diversities during the Norian and Rhaetian). The provinces of Triassic framebuilders can be inferred only for Late Triassic reefs, and seem to be better expressed by non-coral associations (calcisponges, hydrozoans, tabulozoans) than by rather cosmopolitan coral faunas. The evolution of Triassic reefs seems to have been triggered by the co-operation of some major biological innovations, which include the development of binding organisms (necessary for the stabilization phase of reef formation); reinstatement of colony building and special growth types which were lost during the Upper Permian; increase of diversity of reefbuilders over time; rapid evolution of secondary framebuilders during the Norian and Rhaetian; and development of hermatypic corals during the Triassic. Die ältesten triadischen Riffe sind aus dem Anis (Pelson) bekannt; anisische Riffe sind bisher nur aus den Alpen und aus den Westkarpathen beschrieben worden. Ladinische und karnische Riffe besitzen eine weitaus größere Verbreitung (Europa, Asien, westliches Nordamerika, Peru); noch größer ist die Verbreitung norischer und rhätischer Riffe. Die biostratigraphische Einstufung vieler Riffe ist schwierig; die riffbildenden Organismen (insbesondere Kalkschwämme und Korallen sowie Mikroproblematika und auch Foraminiferen) gestatten nur die Trennung von Anis bis Karn und Nor-Rhät. Der gegenwärtige Untersuchungsstand ist durch eine Konzentration der Arbeiten Uber Trias-Riffe im alpin-mediterranen Raum und im westlichen Nordamerika gekennzeichnet. Die Information über die Zusammensetzung der gerüstbildenden und sedimentbindenden Faunen sowie über die Riffbildner-Assoziationen ist im allgemeinen gut; es fehlen jedoch neue Untersuchungen über die Riffbewohner. Das gleiche gilt für dringend notwendige Arbeiten über die diagenetische Entwicklung der Riffkarbonate. Die triadischen Riffe lassen sich fünf, durch Gestalt und paläogeographische Position unterschiedenen Riff-Typen zuordnen: 1. Biostrome auf Karbonatrampen. 2. Stillwasserbioherme (Mud Mounds). 3. Riffhügel-strukturen (Reff Mounds), differenziert in Knollenriffe (Knoll Reefs) und Fleckenriffe (Patch Reefs). 4. Atolle. 5. Barrier-Riffkomplexe. Fig. 4 bietet eine Möglichkeit, triadische Riffe im Hinblick auf die paläogeographische Lage und die Gestalt der Riffkörper zu klassifizieren. Die meisten bis jetzt beschriebenen Riffe entstanden in einer Schelfrandposition oder im Rücken ausgedehnter Plattformen (back-platform position). Am oberen Schelfhang und am Plattformrand gebildete Riffe treten früher (bereits im Ladin) auf als Plattformriffe und Riffe in Becken hinter der Plattform. Die palökologische Interpretation der Rifforganismen wird durch die Seltenheit von genauen paläontologischen untersuchungen vieler Riffe erschwert; auch bei den wichtigsten riffbildenden Gruppen treten ungeklärte Probleme auf: Kalkschwämme (bisher wurden nur die Sphinctozoen der europäischen Vorkommen genauer untersucht), Korallen (die Frage der Entstehung der Korallen-Zooxanthellen-Symbiose) und Algen-Krusten als sekundäre Gerüstbildner (keine rezenten Vergleichsmöglichkeiten!). Die relative Bedeutung der Riffbildner (Fig. 5–6) hat sich in der Zeit geändert; dies wird auch in den, in der Mehrzahl auf die Zeitbereiche Anis-Karn oder Nor-Rhät beschränkten “Riffbildner-Assoziationen” deutlich, die teilweise verschiedene Stadien der Riffentwicklung repräsentieren. Eine allgemeine, sowohl in der Mitteltrias als auch in der Obertrias erkennbare Entwicklungsrichtung ist dadurch gekennzeichnet, daß in der vertikalen Folge der Riffe ein durch inkrustierende und geringdiverse Organismen charakterisiertes Anfangsstadium durch höher diverse aber vertikal und lateral zonierte Assoziationen abgelöst wird. Bei Berücksichtigung der numerischen und der Faunen-Diversität der Riffbildner wird die Existenz von Diversitätsgradienten innerhalb der Riff-Komplexe deutlich. Innerhalb von gleichalten Riffen unterschiedlicher paläogeographischer Position (Fig. 7) scheinen Diversitätsunterschiede aufzutreten, desgleichen bei Riffen gleicher Position aber von unterschiedlichem Alter. Die Existenz von “Provinzen” innerhalb der triadischen riffbildenden Gruppen kann nur für die Zeit der oberen Trias vermutet werden; Kalkschwämme, Hydrozoen und Tabulozoen scheinen hier bessere Hinweise zu liefern als die weitgehend kosmopolitischen Korallen. Die Entwicklung der Trias-Riffe wurdeaußer durch großräumige paläogeographische Veränderungen während der Mitteltrias (Beginn des Rifting)—im wesentichen durch das Zusammenwirken eimger, von der Evolution der Rifforganismen abhängiger Neuerungen beeinflußt, zu denen die Entstehung von sedimentbindenden Assoziationen (erforderlich als Pioniergemeinschaften in der Stabilisierungsphase), die Wiederherstellung der im oberen Perm verlorenen Fähigkeit zur Ausbildung von Kolonien und von bestimmten Wuchsformen, die Zunahme der Diversität in der Zeit, die rasche Entwicklung der sekundären Gerüstbildner während des Nor und des Rhät, und schließlich die Entstehung von hermatypen Korallen gehören.
Article
The Pantokrator Limestones from the Island of Hydra and the Didymi Mountains (southern Argolis, Greece) form a heterogenous complex up to 1000 m thick, with lateral and vertical interfingerings of different facies units (Fig. 3). Sedimentation in the Didymi Mountains (Fig. 1) was dominated by the deposition of lagoonal to tidal shallow-water platform limestones of Carnian to Liassic age. The vertical section of the Upper Triassic rocks exposed on Hydra (Fig. 2) shows a variety of facies units: Laminated cherty mud-limestones (“Hornsteinplattenkalke”) occur at the base of the complex and dominate the southern portion of the island. The thickness of the shallow-water carbonates (Pantokrator Limestones) increases towards the north. Sedimentation started with the deeper-water deposition of biodetrital mud-limestones, which are overlain by reef-limestones and finally by lagoonal to tidal carbonates. According to the faunal and floral associations, the laminated cherty mud-limestones and the biodetrital mud-limestones are thought to be of Carnian age, the reef limestones of Carnian/Norian age, and the loferites and lagoonal sediments of Norian/Rhaetian age. The biota of the reef-limestones on Hydra shows a vertical zonation (Fig. 4), which first starts with a coral-sponge-association followed by a sponge-(coral-) association and finally, a coral-association. This sequence is believed to have been caused by the continuous shallowing of the marginal platform and reef areas. The environmental analysis and interpretation of the facies units are based mainly on the occurrence and distribution of calcareous algae as well as their roles in carbonate production (Figs. 6–7). Reconstruction of the depositional environment was carried out by a study of the relationship between the facies units, indicating a basinward progradation of the platform margin towards the south (Fig. 5).
Article
The paper deals with the microfacies and palecology of the Upper Triassic (Norian) Dachstein Reef Limestones of the Gosaukamm Range, situated about 60 km southeast of Salzburg, Austria. Additional studies are devoted to the coral-bearing Zlambach Beds adjacent to the western Gosaukamm. The Gosaukamm may be regarded as anexample of a continuous synsedimentary degradation of shelf margin areas with isolated reefs. More than 70% of the Dachstein limestone samples studied are bioclastic and lithoclastic rudstones and bioclastic grain/packstones composed of reworked and encrusted reef biota. Low-growing frame-building organisms (predominantly calcisponges) were responsible for the formation of primarily non-rigid frameworks situated at the margin or on protected parts of the foreslope. Stabilization of the reef framework was possible due to the very strong biogenous encrustation (mainly by algal crusts) and by a rapid submarine cementation. Destruction of the marginal reefs by hurricanes and mass flows was influenced by eustatic changes of sea level. Therefore, no clear distinction can be made with regard to various parts of a “reef-complex” (back-reef, central reef area, fore-reef) even though the bulk of the sediment can be compared with “fore-reef breccia”. TheDachstein Reef Limestones of the Gosaukamm Range are underlain by well-bedded micritic cherty limestones (Chapter 3) rich in radiolarians, filaments, planktonic crinoids, sponge spicula and pelecypods (Norian species ofMonotis). The upper part of this sequence is represented by high-diversity bioclastic grainstones showing inverse grading. The biota are characterized by coated echinoderms, shells, dasycladaceans and some recrystallized calcisponges, together with microproblematica (predominantlyTubiphytes) and foraminifera (“Sigmoilina”, Reophax, Galeanella, Endothyra, Trochammina); micritic lithoclasts with thin shells and withMuranella are rather common. These carbonates seem to represent platform carbonates (Fig. 25) redeposited within a basinal environment. The sequence is overlain by well-sorted micritic bioclastic limestones of the “fore-reef” facies. Theframe-building organisms of the Gosaukamm reef (Chapter 4.2) are represented by similar species as in other Upper Triassic reef limestones of the Northern Alps: Calcisponges (sphinctozoans common, inozoans rare), corals (relatively high-diversity but quantitatively subordinate as compared to calcisponges), “bryozoans” (Cyclostomata) and tabulozoans (probably sclerosponges), hydrozoans (spongiomorphids, disjectoporids,Lamellata; Heterastridium) and chaetetids. Calcareous algae are known by dasycladaceans (Diplopora, Heteroporella, Griphoporella) and solenoporaceans (Solenopora, Parachaetetes). Spongiostromate crusts are of major importance within the organic framework (see Fig. 7 and Plate 31). Foraminifera are abundant in the bioclastic grain/packstones and in bio- and lithoclastic rudstones (Fig. 8); the species and genera correspond to those known from other Upper Triassic reef limestones (see Chapter 4.3) but no distinct distributional patterns can be recognized because of large amounts of allochthonous sediments. The same holds true for most of the microproblematica (Chapter 4.4). Reef-dwelling organisms are represented by pelecypods, gastropods, rare small ammonoids, brachiopods, ostracods (especially in reef-cavities), crinoids, echinoids, serpulids and crustaceans (exclusively fecal pellets). Most of these groups can be found in all facies types. Themicrofacies types of the Dachstein reef limestones (Chapter 5) were defined by quantitative and semi-quantitative analyses of more than 1100 large thin-sections. Important criteria for the differentiation of the microfacies types are the amount of micrite, sparite, particles and porosity; packing, size and roundness of detrital particles; types of fossils; and encrustation sequences. 10 microfacies types can be separated: (Chapter 5.2); this rather broad spectrum reflects the existence of very well diversified platform and reef environments which were simultaneously reworked by storms and by mass transport at the edge of the platform. Thediagenesis of the Dachstein reef limestones (Chapter 6) is characterized by submarine and vadose cementation. Submarine cements are represented by fibrous and also radiaxial cements (see Plate 40); vadose diagenesis is indicated by crystal silt occurring in all microfacies types but most abundant within the bioclastic grain/packstones (microfacies 2), rud/floatstones (microfacies 1), and boundstones (microfacies 6), see Figs. 16 and 22. Most Dachstein reef limestones exhibit a high interparticle and framework porosity which is generally much reduced by early diagenetic cementation (see Fig. 23). Secondary porosity is represented by irregular vugs and fissures filled with vadose silt. Theinterpretation of the biota and the microfacies (Chapter 7) points to the existence of agitated, shallow environments. Because of the lack of high-growing frame-builders only small and rather scattered reef structures seem to have grown at the platform margin. Indicators of platform environments are: high-diveristy shallow-marine biota (microfacies 6), predominance of “reef breccia” and bioclastic detritus (microfacies 1 and 2), abundance of coated grains (microfacies 4), aggregate grains (microfacies 9), multiple resedimentation (within almost all microfacies types), and vadose diagenesis. Structural and textural data (grain-size distribution, sorting, roundness, energy-index) verify this interpretation. Thefacies model (Chapter 8) postulates the existence of a differentiated platform margin which is continuously eroded by wave energy and storms; the allochthonous material is deposited on the foreslope and in different parts of the platform. Small reefs are built in protected parts of the slope and the margin by calcisponges, algal crusts and various epizoans, in conjunction with a rapid submarine cementation. The biota and facies types are similar to other Upper Triassic reefs but the facies model of the Gosaukamm (Fig. 25) exhibits distinct differences caused by a stronger influence of eustatic sea level fluctations. Themicrofacies of the Zlambach Limestones (Chapter 9) is characterized by coral bafflestones; bioclastic packstones with densely packed corals, solenoporaceans, pelecypod shells, echinids and foraminifera; medium-grained pack/grainstone with micritic lithoclasts and echinoderms or with coated grains; graded, densely packed bioclastic packstones with broken corals, tabulozoans, crinoids, and foraminifera; bioclastic wackestones with echinoderms; and burrowed wacke/packstones with sponge spicula. Theenvironment of the Zlambach Beds adjacent to the Dachstein Reef Limestones of the Gosaukamm (Chapter 9.5, Fig. 32) can be interpreted as morphologically differentiated shallow-marine areas with shoals (coral bafflestones) and depressions (allochthonous carbonates). Shallowing-up sequences might have developed (indicated by vadose silt). In comparison to the Dachstein reef limestones of the Gosaukamm, there are distinct differences in the microfacies types, the importance of the corals, and in the encrustation sequences: these factors indicate a depositional environment of the Zlambach Beds which was not primarily influenced by the development of the Dachstein platform.
Article
The fauna of the upper Cassian Formation is composed mainly of reef-building and reef-dwelling organisms which occur as reeeposited material in basinal sediments, but have not been found as original reef bodies. Such bodies have now been discovered in the uppermost Cassian Formation of the central Dolomites from the Sella Group in the west to the Monti Cadini in the east. Generally they are small-scale patch reefs, not exceeding a few metres in thickness and lateral extent, which are intercalated in well-bedded detrital and micritic limestones. locally, larger biostromes spread out from the margins of the Cassian Dolomite buildups. Four types of faunal communities have been encountered in these reefs:1. The thrombolite-calcareous algae community, composed of small patchy cryptalgal structures binding poorly sorted debris and associated with other Cyanophyta, sessile formainifera and scattered calcareous sponges and corals. This type is the most common within the calcareous and marly-tuffaceous facies of the Cassian Formation. 2. The calcareous sponge-coral community, composed mostly of calcareous sponges (stromatoporoids, some pharetronids) and, to a lesser extent, colonial corals and thrombolites. This community corre-sponds well to the Cassian reef fauna, best known from erratic blocks at Alpe di Specie, but has been found in situ only at one locality. 3. The Spongiomorpha-Solenopora community, associated with scattered calcareous sponges and colonial corals, forming a thin biostrome at one locality. 4. Coral communities, composed predominantly of colonial Scleractinia; found only in small or stratigraphically illdefined outcrops and in erratic blocks. The Cassian patch reefs and biostromes mark the end of a basinal evolution which began in the Lower Ladinian, and the onset of newly expanding carbonate buildups of Cassian Dolomite. These buildups and the sponge-coral patch reefs might have been the source for the allochthonous reef fauna of the Cassian Formation which interfingers with both shallow water environments.
Article
The internal architecture of Upper Permian calcisponge reefs, Upper Triassic coral thickets, and Upper Triassic coral reef communities of the Oman Mountains have been investigated. In order to gain comparable data sets, investigations were carried out at different scales comprising quantitative data from outcrops and descriptions from thin-sections. Methods of quantitative outcrop investigations were modified with reference to standard investigation techniques used in the study of communities of modern reefs. Data evaluation comprises mapping of reef fabric in natural scale on plastic sheets in the field. Data calculation was carried out utilizing the digitized image analysis system Vidas. Measurement parameter are the total detrital framework coverage, coverage of reefbuilder taxa, mean diameter of taxa, and mean distance. In addition, measured plastic sheets were printed for paleoecological interpretation. Thin-section analysis reveal microfacies types, sedimentological criteria and taxonomic inventory of reef organisms. Based on quantitative field data five Upper Permian and four Upper Triassic communities were differentiated. Late Permian communities are represented by a: (1) Low-diverse sphinctozoan community, (2)Radiotrabeculopora Archaeolithoporella community, (3) Cerioid coral community, (4) Solitary coral community, and (5) Waagenophyllid coral community. Upper Triassic communities comprise: (1) Diverse coral community, (2) CoralSpongiostromata community, (3) Solenoporacean dendroid coral community, and (4) Crinoid community. The synthesis of both quantitative field data (plot technique with quadrats as sampling units) and the study of thinsections (microfacies analysis, taxonomy) is believed to result in data sets which could be used in comparative reef research with a higher degree of reliability than up to now.
Article
Microfacies studies of the carbonates of the Kössen Beds (Salzburg-Tyrol, Austria) show that the deposition took place within a protectedintraplatform basin. Three facies units can be differentiated within the profiles: I Marl Carbonate sequence; II Coral Limestones; III Carbonate Marl Bed sequence. A diachronous development of 14 microfacies types and also of the facies units (alternating within one profile) becomes evident when the sections are compared; facies unit III indicates a subsidence of the basin. The seven microfacies types of theMarl Carbonate sequence (I) represent shallow water carbonates, formed in protected areas or surf zones. Episodic higher-energy events together with differences in the salinity can be inferred from the distribution of the organisms and the facies types: Laminated Algal Bindstone (I/1), Pelmicrite (I/2), Mudstone (I/3), Lumachelle facies (I/4), Bioclastic Grainstone (I/5), Onkoid facies (I/6), Oolitic Grainstone (I/7). Two genetic types occur within theCoral Limestone facies unit II: The“Shallow Coral Limestone” was formed in agitated water; flora and fauna are reminiscent of those in the Rhaetian reefs and the Dachstein Reef Limestones. The“Deeper-Marine Coral Limestone” was formed below the photic zone, but still within the influence of episodic storms. Five microfacies types are developed, sometimes only in the outcrops: Reefdetritus-Mud facies (II/1), Biostromal facies (II/2), Mud Mound Facies (II/3). Coated Grain Facies (II/4), Formaminiferal grainstones (II/5)—the latter occurs in the “Shallow Coral Limestones” only. Monotonous Carbonate/Marl intercalations consisting of the Echinoderm-Detritus-Mud facies (III/1) and the Detritus-Mud facies (III/2) characterizefacies unit III. Low-diversity fauna and high-diversity trace fossils point to a deposition in greater depths. The presence ofChoristoceras marshi indicates, this sequence is of Rhaetian age. Corals, echinoderms, foraminifera and microproblematika are common within the shallow-marine carbonates; sponges, hydrozoans and algae are rare; in contrast to the Rhaetian reefs, the flora and fauna are of rather low-diversity. Brachiopods, mollusks, ophiurids and ammonites are important for facies and stratigraphic interpretations. Of special interest is the distribution of the benthic foraminifera: Facies-controlled associations can be recognized.Agathammina austroalpina, Glomospira sp. and thin-walled nodosariids adapted to high salinity.Agathammina austroalpina is the only foraminiferal species, found within the tempestites formed by coquinas.Aulotortus gaschei is a typical species of the oncoid carbonates, it is the only foraminifera within the oolitic grainstones. Involutinids and textulariids are missing within facies unit III;Tetrataxis, Planiinvoluta and small miliolid foraminifera were adapted to this deeper water environment. The “Shallow-water Coral Limestones” are characterized by highly diverse involutinids. The foraminiferal distribution within the “Deeper Coral Limestones” shows a maximum of individuals and species within the Mud Mound Facies. The lowest number of individuals occurs within the Coated Grain Facies. The deposition of the “Shallow-water Coral Limestones” was influenced by episoidc storm events, as shown by densely packed shell (or coral) layers. The thickness of these tempestites depends on sedimentation energy. Sea level fluctuations, which caused the “Deeper Coral Limestones” to stop developing are indicated by vadose/marine and vadose leaching and cementation processes, probably due to temporary subaerial exposure (perhaps connected with early Cimmerian epeirogenetic movements). Blocky cements were formed under freshwater/phreatic conditions, as confirmed by Mn-enrichment and Sr-depletion of these cements. The distribution of trace elements (Fe, Mn, Mg, Sr) and insoluble residues was interpreted with the help of cluster analysis and factor analysis. The variability of the parameters is due to the primary mineralogical composition and diagenetic alterations (R-technique); water energy and micrite content are restrictive factors, as shown by the Q-technique. Sr and Mn distributions within the different profiles exhibit different patterns: different diagenetic stages are reflected by Sr values. The different intensities of the freshwater/phreatic diagenesis are expressed by different Mn values.
Article
The Upper Rhaetian coral limestone of Adnet, southeast of Salzburg Austria has been repeatedly referred to as one of the most spectacular examples of an ancient ‘autochthonous’ coral reef structure. The ‘Tropfbruch’ quarry is probably the best outcrop for interpreting the distributional patterns of biotic successions and communities of a late Triassic patch reef. Our study is based on the interpretation of a) outcrop photographs, b) reef maps resulting from quadrat transects, and c) the analysis of quantitative data describing the distribution and frequency of reef organisms and sediment. A new methodological approach (combination of reef mapping and photo-transects) is used to obtain quantitative field data which can be compared in greater detail with data from modern coral reefs investigated by corresponding quantitative surveys. Three unconformities and three well-defined ‘reef growth stages’ reflecting the vertical and lateral development of the reef structure were differrentiated using transects: Stage 1, representing the reef growth optimum, is characterized by laterally differentiated coral reef knobs with corals in growth position. Criteria supporting this interpretation are the extraordinary size of the corals, their preservation in situ and the great thickness of this interval. The massive coralPamiroseris grew under higher energy conditions at the rim of the reef knob, whereas branchingRetiophyllia colonies preferred less agitated water in the center. Vertical changes are reflected by an increase in frequency of the dasycladacean algaDiplopora adnetensis and by the decreasing size ofRetiophyllia. These sedimentological and biological criteria together with the unconformity above indicate a fall in the sea level as a major control mechanism. Stage 2, separated from stage 1 by an unconformity caused by partial subaerial exposure and karstification, is characterized by vertically stacked coral successions with diverse reef debris. Facies heterogeneity is reflected by differences in the diversity, taphonomy and packing density of reef-building organisms as well as by differences in sediment input from the platform. Water depths and accommodation space were lower, therefore minor sea level fluctuations had a stronger effect on the biotic composition. The high percentage of coral debris and corals reworked by storms and the increase in the input of platform sediment led to a reduction of reef growth. Stage 3, again separated at the base by an unconformity associated with karstification, is characterized by bioclastic sediments with isolated reefbuilders forming a level-bottom community. The distribution of different coral morphotypes suggests that sea level fluctuations were not the only controlling factor. Variations in the substrate were caused by differences in the input of platform sediment. The three-step development seen in Adnet documents the response of low-diverse coral associations to variations caused by small-scale sea level changes, storm activity and sedimentation. The vertical changes in reef community structures correspond to a sequence of ‘allogenic replacements’. The Adnet reef structure should not be regarded as a general model of Alpine Upper Rhaetian reefs, because of the particular setting of the patch reef. Only the ‘capping beds’ of the Upper Rhaetian Reef Limestone of the Steinplatte exhibit criteria similar to Adnet. Potential modern analogues of features seen in the coral communities of Adnet are the internal structure of theRetiophyllia thickets, the key role of branching corals within the communities, the scattered distribution and low and even diversity of corals subsequent to breaks in settlement, segration patterns of corals indicating ‘contact avoidance’, toppling of large coral colonies by intensive boring, and decreasing coral coverage from deeper and sheltered settings to more shallower water depths.
Article
In Nordwest- und Zentralsizilien (Panormide-Gebirge westlich Palermo; Madonie-Gebirge) sind am Rand ausgedehnter Karbonatplattformen obertriadische Riffe entwickelt, deren norisches Alter durch Heterastridien und Ammoniten belegt ist. Es handelt sich um Schwamm-Bryozoen-Riffe, deren Faunen- und Florenbestand weitgehende Übereinstimmung mit etwa gleich alten Riffen in den Nördlichen Kalkalpen (Dachsteinkalkriffe) besitzt. Es sind folgende Organismengruppen vertreten (Die im Rahmen dieser Arbeit als Bryozoen bezeichneten Organismen stellen höchstwahrscheinlich keine Bryozone dar. Alle diese Organismen sind vermutlich zu den Chaetetiden oder Sclerospongien zu stellen): lien und Astraeomorphen); 3. Hydrozoen (selten bis auf Disjectoporiden); 4. Bryozoen/Tabulozoen (häufig und hochdivers; bei Chaetetiden-ähnlichen Fossilien dürfte es sich um Sclerospongien handeln); 5. Algen (überwiegend an den Lagunenbereich und an den Übergangsbereich Lagune/Riff gebunden; es treten Dasycladaceen, Solenoporaceen sowie porostromate und spongiostromate Cyanophyceen auf; von Interesse ist das Vorkommen von Dasycladaceen-Sporen (Taf. 23/11)); 6. Foraminiferen (im Artenbestand mit den Foraminiferen-Assoziationen der nordalpinen Obertrias übereinstimmend; hierzu tretenFoliotortus PILLER & SENOWBARI-DARYAN sowie vasenförmige Gehäuse, die unter den NamenCucurbita, Amphorella etc. bisher nur aus der Obertrias der Westkarpaten sowie von der Hohen Wand bei Wien bekannt (wurden); 7. Problematika (ähnlich wie in der Riff-Fazies der alpinen Obertrias). Am Cozzo di Lupo, W Palermo können die Faziestypen des Riffes und der Lagune lateral verfolgt werden: An eine durch Schwamm-Bryozoen-Biolithite mit zahlreichen, zementerfüllten Hohlräumen gekennzeichnete Zone schließt sich eine durch Korallen (Thecosmilien), Algen (Dasycladaceen, Solenoporaceen, Cayeuxien), Gastropoden und Megalodonten charakterisierte Zone an (Korallen-Algen-Biolithit), in welcher sich Bänke mit Korallen und Bänke mit Megalodonten vertikal ablösen. Vereinzelt treten kleine Riffknospen auf, deren Fazies derjenigen der Schwamm-Bryozoen-Biolithite entspricht. Die Korallen-Algen-Biolithit-zone geht in eine zone mit Stromatolithen und Fenstergefügen über (Loferit-Fazies). Die Verteilung der Organismen innerhalb der Faziesbereiche kann mit ähnlichen Assoziationen in der nordalpinen Obertrias verglichen werden (Abb. 3): 1. Korallen sind jedoch im Gegensatz zu vielen alpinen Riffen in der Schwamm-Bryozoen-Biolithit-Fazies als Gerüstbildner nur von geringer Bedeutung. Thecosmilien treten insbesondere im Übergangsbereich zwischen Riff und Lagune sowie in der Lagune auf, wo sie Biostrome bilden.—2. Kalkschwämme, Hydrozoen, sowie Bryozoen/Tabulozoen waren auf geringenergetische, tiefere Wasserbereiche beschränkt. Epi- und Endofauna dieser Assoziationen entspricht der von Kalkschwamm-Biozönosen aus der nordalpinen Obertrias.—3. Algen zeigen stark differenzierte Verbreitungsmuster; Spongiostromen sind auf die Schwamm-Bryozoen-Biolithit-Fazies beschränkt, Cayeuxien finden sich in der Lagune zusammen mit Thecosmilien und Megalodonten. Ebenfalls in der Lagune sind Dasycladaceen (insbesondereHeteroporella undDiplopora) weit verbreitet. Solenoporaceen sind vor allem in der Riff-Fazies sehr häufig.—4. Foraminiferen sind mitGaleanella, Ophthalmidium und “Lituosepta” auf die Schwamm-Bryozoen-Biolithit-Fazies beschränkt, während eine zweite, durchAlpinophragmium, Aulotortue, Glomospira und andere Gattungen gekennzeichnete Gruppe in der Korallen-Algen-Biolithit-Fazies der Lagune auftritt. In Riffhöhlen der Schwamm-Bryozoen-Biolithit-Fazies finden sich als wichtige FaziesindikatorenFoliotortus, Cucurbita und ähnliche Mikrofossilien, die wahrscheinlich als Foraminiferen anzusehen sind.—5. Auch die Mikroproblematika weisen bestimmte Verteilungen auf (Microtubus, Radiomura, Baccanella in der Schwamm-Bryozoen-Biolithit-Fazies;Bacinella, Thaumatoporella, Aeolisaccus undTubiphytes häufig in der Lagunenfazies). Der Ablagerungsraum des Cozzo di Lupo-Riffes (Abb. 4) ist durch eine, in etwas tieferem Wasser liegende Riffzone (Schwamm-Bryozoen-Biolithit-Fazies) gekennzeichnet, an welche sich in Richtung zur Lagune eine relativ hochenergetische Übergangszone anschloß. In der gut durchlichteten, flachen Lagune entstanden Korallen-Siedlungen, Dasycladaceen-Rasen und Megalodonten-Bänke. In geschützten, lagunären Bereichen konnten durch Schwämme und Bryozoen/Tabulozoen kleine Riffknospen gebildet werden. Die Lagune dürfte sich gegen Norden und Nordosten verflacht haben (angedeutet durch Stromatolithen, Fenstergefüge, geringe Diversität). Grundsätzlich ergeben sich auffallende Ähnlichkeiten mit dem Faziesmodell der Steinplatte (PILLER & LOBITZER 1979). Upper Triassic reefs were built at the margins of carbonate platforms in north-western and central Sicily (Panormide Mountains west of Palermo; Madonie Mountains). The Norian age of these reefs is evidenced by ammonites and the occurrence ofHeterastridium. The reefs are represented by Sponge-Bryozoan reefs; fauna and flora are very similar to those of Upper Triassic reefs of the Northern Calcareous Alps (Dachsteinkalk reefs). The biota consists of: 1. Calcisponges (especially sphinctozoans; more than 30 species); 2. Corals (of minor importance, low-diverse, predominantlyThecosmilia andAstraeomorpha); 3. Hydrozoans (rare, except disjectoporids): 4. Bryozoans/Tabulozoans (abundant, high-diverse; some chaetetid-like fossils may represent sclerosponges); 5. Algae (common within the transition area between the reef and the lagoon and in the lagoon; dasycladaceans, solenoporaceans as well as porostromate and spongiostromate cyanophyceans occur; of special interest is the occurrence of dasycladacean spores (P1. 23/11); 6. Foraminifera (the same species as in the Alpine Upper Triassic; in additionFoliotortus PILLER & SENOWBARI-DARYAN and amphora-like tests (Cucurbita, Amphorella etc., until now only known from the Upper Triassic of the Western Carpathians and from the Hohe Wand near Vienna, Austria); 7. Problematica (similar to the species found in Alpine reef limestones). A lateral transition of the facies types can be followed at the Cozzo di Lupo, west of Palermo: (1) Sponge-Bryozoan Biolithite-zone (central reef area) characterized by low-growing framebuilders and abundant cement infillings in cavities; (2) Coral-Algal Biolithite-zone (lagoon) withThecosmilia, algae (dasycladaceans, solenoporaceans,Cayeuxia), gastropods and megalodontid pelecypods; some small reef patches (similar in facies with the sponge-bryozoan-biolithite facies) may occur within this zone; (3) Loferite zone (lagoontidal flat) with stromatolites and fenestral fabrics. The distributional patterns of the fossils within the facies units can be compared with those of the reefs in the Northern Calcareous Alps (Fig. 3): 1. Corals are of minor importance within the spongebryozoan facies, but occur predominantly in the transitional area between the reef and the lagoon, and within the lagoon (biostromes withThecosmilia).—2. Calcisponges, hydrozoans and bryozoans/tabulozoans were restricted to somewhat deeper, low-energy environments. Epi- and endofauna of these associations are similar to those of calcisponge biocoenoses of the Alpine Upper Triassic.—3. Algae exhibit strongly differentiated distributional patterns: Spongiostromate algae and solenoporaceans occur predominantly within the sponge-bryozoan biolithite. The lagoonal environment is characterized byCayeuxia (together withThecosmilia and with megalodontid pelecypods) and dasycladaceans (Heteroporella, Diplopora).—4. Foraminifera are useful facies fossils: The association withGaleanella, Ophthalmidium and “Lituosepta” is restricted to the sponge-bryozoan biolithite facies, whereas the association withAlpinophragmium, Aulotortus, Glomospira and other genera occurs in the coral-algal biolithite facies. A third association, characterized byFoliotortus, Cucurbita and similar microfossils, is typical of biotopes within reef cavities.—5. Microproblematica show special distributional pattens:Microtubus, Radiomura, andBaccanella in the sponge-bryozoan biolithite facies;Bacinella, Thaumatoporella, Aeolisaccus andTubiphytes in the lagoonal facies. The environment of the Cozzo di Lupo reef complex (Fig. 4) is characterized by a reef zone, which developed in a slightly deeper area (sponge-bryozoan biolithite facies), bordered by a moderate to high-energetic transitional zone. The shallow, well-lighted lagoon was characterized by coral biostromes, dasycladaceans and megalodontids flourishing on the mud bottoms. Small reef patches were formed by calcisponges and bryozoans/tabulozoans in more protected areas of the lagoon. A shallowing of the lagoon might have occurred to the north and northeast, as indicated by stromatolites, fenestral fabrics and a generally low-diversity. The facies model of the Cozzo di Lupo reef is similar in many respects to that of the Upper Rhaetian Steinplatte reef near Waidring, Austria (PILLER & LOBITZER 1979).
Article
The community is the standard unit of analysis in ecology because through it both physical and biological aspects of the ecosystem can be determined. It offers the same potential in paleoecology. The communities of the upper Triassic reefs are analyzed from the literature and our own field work in order to determine their characteristics and to assess their usefulness. Six communities are recognized on the basis of R-mode cluster analysis of the do minant frame-builders in Rötelwand reef and from descriptions and our observations in other reefs. TheThecosmilia or High-growing coral, the Hydrozoan-tabulozoan and the Sponge communities are most widespread. The High-growingAstraemorpha, Low-growing coral andSolenopora communities are less common. Each community is characterized by a few dominant taxa and a larger number of associated taxa. The fidelity of the dominant taxa to their community is generally high; the fidelity of many of the associated taxa are typically lower and these taxa occur in a number of communities. Consequently, there is considerable overlap in composition between stands of different communities. In addition, the communities typically occur in patches only a few meters in diameter, and particularly in the Hydrozoan-tabulozoan community, the composition may differ considerably between adjacent patches. Differences in diversity between the communities are explained in several ways. Parameter of the physical environment can be correlated with diversity. Competition between members of the community probably had a strong effect on community composition and diversity. Different geologic longevity of the communities, which may have led to differences in community evolution and niche structure does not seem to be an important factor. Succession is poorly documented in these reef communities, either in encrustation sequences or in the development of individual reef patches. Community replacement, well developed only at Rötelwand, is assumed to reflect variations in water depth. The communities within several reefs are not sharply delimited geographically but have broad and overlapping distributions. The reef patches were widely and apparently randomly scattered in the reef tract, reflecting a generally uniform environment there. The depositional surface was not deeper than perhaps 10 m. The constraints on water depth are that it was shallow enough for the depositional surface to be within the photic zone for the growth of abundant algae, and above wave base for the production and multiple reworking of biogenic debris, but deep enough for a uniform environment with low gradients to be maintained across the reef tract. The reef patches did not have the capability to build up to sea level nor to construct a platform-edge reef dividing sharply reef and fore-reef tracts. The dominant organisms in the reef patches did not have the ability to construct wave-resistant framework. Consequently, these reefs do not fit the definition of ecologic reefs but do correspond to stratigraphic reefs.
Article
Permian and Triassic carbonate platforms of the Arabian Peninsula (Gondwana) and seamounts of the Neo-Tethys (Hawasina and Batain basins) are characterized by distinctive supersequences (second order, duration 5–20 million years, my) and composite sequences (third order, duration 0.5–5 my). The presented sequence stratigraphic framework will be compared with existing sea level curves to discuss the validity of different regional oscillations during the dispersal of Pangea.The carbonate succession of the Haushi and Akhdar Groups of the Arabian platform is composed of four Permian (P1–P4) and four Triassic supersequences (Tr1–Tr4). Isolated platforms of the Hawasina and Batain basins comprise two Permian supersequences and one Triassic supersequence. In contrast to the continuous development of the Arabian shield, carbonate platform growth of the seamounts was restricted to the Guadalupian–Lopingian and to the Middle–Upper Triassic, and ceased after drowning events.Composite sequences exhibit a well-developed stacking pattern during the Guadalupian–Lopingian (Saiq Formation). Lowstand systems tracts (LSTs) occur during the Cisuralian (Gharif Formation, Haushi Group) and Triassic (Mahil Formation, Akhdar Group). Open-marine depositional environments prevail during transgressive systems tracts (TSTs) with diverse biota including rugose and scleractinian corals, chaetetids, bryozoans, and crinoids. Highstand system tracts (HSTs) exhibit a twofold pattern: During the transgressive phase of supersequences, composite sequence highstands are dominated by reef or level-bottom communities with corals. Cyclic platform deposits or monotonous mud- and wackestone accumulated during the turnaround or late second-order highstand of a supersequence.Correlation of maximum flooding surfaces with published data suggests that supersequences P1, P2, and Tr4 can be traced across the Arabian platform into the Neo-Tethys basins, while supersequences P3, P4, and Tr1–Tr3 resulted from local tectonic events at the margin of the Arabian platform (Hulw half-graben). The presented sea level curve corresponds therefore to the Tethyan sea level curve during the Cisuralian and Guadalupian, but differs significantly during the Lopingian, as a result of the dispersal of Pangea. The Middle and Upper Triassic sea level curve from Oman is again in good correlation with published data. The Permian and Triassic sequence architecture on the Arabian plate and adjacent Neo-Tethys was predominantly triggered by the global warming after the Permian–Carboniferous glaciation, the initial rifting of Neo-Tethys, and subordinately by eustatic sea level changes.
Article
At head of title: Université de Lausanne. Faculté des sciences. Institut de géologie et de paléontologie. Thesis (Ph. D.)--Université de Lausanne. Includes bibliographical references (p. 235-244) and index. Text in French; summary in English and French.
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