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The morphology and systematics of some examples of Kungurian Callipteris (Peltaspermaceae) from the Ural region
Abstract
The problem of the generic name of Callipteris Brongn. is discussed, and new original remains of Callipteris are described from the Permian in the Ural region. This group is of great importance in the identification of the Carboniferous/Permian boundary and the subdivision of continental Permian deposits. -P.Cooke
This paper deals with Permian peltasperm foliages attributed to a new genus Permocallipteris Naug., gen. nov., which is characteristic of Angaraland paleofloristic realm. The genus is proposed for two-or three-pinnate fronds with entire-margined last order segments (pinnules), normally with obtuse round to slightly acute apex, wide base, well-developed midvein and pinnately disposed lateral (secondary) veins. The fronds always possess clear additional (intercalated) pinnules attached directly to the frond rachis. The intercalated pinnules can be identical to common pinnules or be slightly modified and form wings on the frond rachis. The epidermal-cuticular structure of Permocallipteris fronds is characterized. The leaves are amphystomatic, the lower (adaxial) leaf cuticle is thinner and has numerous papillae. Stomata are monocyclic to incompletely dicyclic, with thick cutinization of the subsidiary cells. In Permian deposits of Angaraland or Angara floristic realm, from the Urals to Siberia and Russian Far-East, southwards to Mongolia and North China, there were many endemic Angaran taxa, making this area a separate distinct paleofloristic realm. The most typical endemic taxa of Permian of Angaraland are equisetophytes of Tchernoviaceae, several genera of both eusporangiate and leptosporangiate ferns (including Prynadaeopteris Radczenko, Geperapteris S. V. Meyen, Acrogenotheca Naugolnykh, Convexocarpus Naugolnykh and Tumidopteris Naugolnykh), gymnosperms of Vojnovskyales (Vojnovskya Neuburg, Nephropsis Zalessky, Rufloria S. V. Meyen) and some other morphologically specific and uncommon plant groups. Peltasperms are widely spread in the Permian deposits of Angaraland, but their taxonomical attribution was disputable and unclear till the present time. This paper deals with the taxonomy of compound pinnate leaves of peltasperms attributed to the new genus Permocallipteris Naugolnykh, gen. nov.
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