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Publications (71)
It is evident that the bryophyte flora of the Cook Islands remains poorly documented. Here, ten moss species and five liverwort species of Lepidoziaceae are newly reported for the Cook Islands. These records include Calomnion denticulatum, previously known only from Samoa, Ectropothecium viridifolium, previously known only from Hawai’i, and Trichol...
Frullania crassissima J.J.Engel, von Konrat and Glenny, a member of the genus Frullania Raddi, is described and illustrated. It is morphologically most similar to F. setchellii, F. svihlana and F. falciloba, of New Zealand and F. vittiana of Lord Howe Island. The new species is also sister to F. squarrosula, F. nicholsonii, F. vittiana and F. falci...
Two new species of Heteroscyphus are described from New Caledonia. Heteroscyphus supinopsis is characterized by the strictly ventral intercalary branches, the undifferentiated stem, with both cortical and medullary cells thick-walled, the subopposite leaves distinctly connate dor-sally and forming a laminar lip, the typically subsymmetrically lingu...
The present publication is the first modern catalogue of the liverworts and hornworts of French Polynesia and accepts 161 species of liverworts and 5 species of hornworts. In addition, 40 species are doubtful records of French Polynesia and 45 species are excluded. The checklist is mainly based on a compilation of published records; a few unpublish...
Abstract: Heteroscyphus kanakensis is described from three specimens collected in New Caledonia at low altitude. It is characterized by leaves dorsally assurgent, subopposite, with a free dorsal base, the leaves ovate, entire, undulate in dorsal view, the cells oval with nodulose trigones, the underleaves rotund to oblate in outline, bifid with lob...
Several new taxa, new combinations, and new synonyms have surfaced during preparation of the second and third volumes of the Liverwort Flora of New Zealand. These are published here in order that they may be made immediately available for use; the taxonomic numbering corresponds to presentation in earlier papers in this series.
The liverwort subfamily Acrobolboideae has historically contained the three genera: Acrobolbus, Marspidium, and Tylimanthus. Generic delimitations in this subfamily have been historically inferred from morphological characters, specifically the location of gametangia. Taxonomists have had difficulty separating the genera, with some combining Tylima...
A conspectus of the Australasian species of Leptoscyphus is presented. Chiloscyphus subg. Aperticaules, Leptoscyphus subg. Blepharoleptoscyphae, L. subg. Idiodonti, L. subg. Microphyllidici, L. subg. Neoleptoscyphus, and L. subg. Physanthi are new subgenera. Chiloscyphus sect. Subemarginati, Leptoscyphus sect. Neoleptoscyphus, L. sect. Erratici, an...
On the basis of variation in molecular sequence data and morphology, three species are recognised within Dinckleria. The generitype D. pleurata is widespread in Tasmania and New Zealand and has outlier populations in Victoria, and in rainforests around the New South Wales-Queensland border. Dinckleria fruticella is endemic to New Zealand, records o...
Brevianthus hypocanthidium and Aponardia huerlimannii are found to be synonymous and the new combination Brevianthus huerlimannii is made.
A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online
flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present
the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta)
that includes 7486 species in 398 genera r...
Brevianthus is a distinctive genus of leafy liverwort in its succubously inserted, entire leaves, lack of underleaves, restriction of sexual organs to lateral-intercalary branches, scattered rhizoids and dense leaf-surface ornamentation. The sole species, Brevianthus
flavus, is divided into two subspecies, one in Tasmania the other in New Zealand....
Chiloscyphus alpicola J.J.Engel, a new member of the Lophocoleaceae from New Zealand, is described and illustrated and Tetracymbaliella subsimplex (Austin) J.J.Engel, endemic to Tasmanian, is a new combination.
A conspectus of the Australasian species of Heteroscyphus is presented. Heteroscyphus subg. Allodonti, H. subg. Fissistipi, H. subg. Glaucoheteroscyphus, H. subg. Heteroscyphus, H. subg. Lingulati, H.
subg. Menziesiorum, H. subg. Neoheteroscyphus, H. subg. Oblongifolii, H. subg. Protoheteroscyphus, H. subg. Sinuosi, and H. subg. Triacanthi are new...
Abstract— The suborder Jungermanniineae of the Jungermanniales is a major lineage of leafy liverworts, recognized in recent classifications to include 15 families. Gametophytes within the suborder are morphologically diverse, but commonly anisophyllous to distichous, usually with succubous, rarely transverse or incubuous, leaf insertions. Sporophyt...
During the preparation of the forthcoming world checklist of hornworts and liverworts (Soderstrom et al., in press. ) it was noted that a few taxa need to be validated or combined. The subfamily name Balantiopsidoideae has been used as if it has been an autonym of the family Balantiopsidaceae H.Buch (1955: 23) (e.g. Grolle 1972, Engel & Smith Merri...
Heteroscyphus assurgentifolius and H. deceptifrons are new species. Heteroscyphus ammophilus var. obtusifolius, H. fissistipus var. repandus, H. mononuculus var. ammophilopsis, H. mononuculus var. bilobus, and H. triacanthus var. magnistipulatus are new varieties. Heteroscyphus fissistipus subsp. multispinus, Cryptolophocolea sect. Bicornutae, C. s...
The monospecific Phycolepidoziaceae with the single neotropical species Phycolepidozia exigua is a highly specialized leafy liverwort without vegetative leaves. The extreme reduction of morphological and anatomical characters of Phycolepidozia has caused uncertainties as to the systematic position of the genus and family. In 2012, a second species...
There remains a critical need to synthesize the vast amount of nomenclatural, taxonomical and global distributional data for liverworts. This is fundamental in taxonomists' efforts towards developing a working list of all known plant species under the auspices of the Convention on Biological Diversity (CBD) as well as having far reaching implicatio...
Gackstroemia novae-zelandiae sp. nov. is added to the genus Gackstroemia, a genus confined to the south temperate. The new species has been previously included within G. weindorferi, and is endemic to New Zealand. Gackstroemia weindorferi of New Zealand and Australia (Tasmania, Victoria, and New South Wales) and G. alpina are also treated.
This paper presents an overview of the infrageneric classification to be used in the treatment of Plagiochila in the New Zealand Liverwort Flora. Included are re-evaluations of all names and concepts in current use, and synonymy. Plagiochila sect. Cucullatae Schiffn. (subg. Metaplagiochila Inoue) is recognized for the first time from New Zealand. P...
Goebelobryum vermiculare sp. nov. is added to the genus Goebelobryum described by Grolle (1962) for two species, G. unguiculatum (Hook.f. & Taylor) Grolle of New Zealand and Australia, and G. grossitextum (Steph.) Grolle, endemic to Western Australia. The new species has been previously included within G. unguiculatum, and occurs in New Zealand and...
A new genus of liverwort in the family Lophocoleaceae is described and illustrated. The genus Octoscyphus presents a suite of original characters for this family. It is placed in the subfamily Lophocoleoideae. It has, in particular, a unique feature among leafy liverworts: The leaves have a very broad foliar base forming a longitudinally inserted p...
Lamellocolea was previously a monotypic genus endemic to New Zealand. Lamellocolea integrostia, a second species in the genus, is described and illustrated from New Zealand.
Three species have been formerly segregated from Metzgeria in the genus Apometzgeria. One of the species, A. frontipilis, is endemic to South America but the second, A. pubescens, is currently understood as having a bipolar range with populations across the Holarctic and in southern South America. The third species, A. longifrondis, was described f...
Estimates of extant liverwort species range from 4,500 to 9,000, with estimates in the past decade converging on 5,000 to 6,000. Potential problems and pitfalls of deriving species estimates are addressed, including binomial accumulation, the impact of synonymy, taxonomic inflation, the impact of unrevised species-rich genera, species concepts and...
DNA sequence results show that Amphilophocolea, a monotypic genus endemic to New Zealand, is nested within Heteroscyphus. A morphological examination of specimens that agree with the protologue shows it to be identical to Heteroscyphus knightii. Heteroscyphus cymbaliferus in the DNA sequence analyses is sister to the clade consisting of the rest of...
Plagiochila microdictyon Mitt., previously considered a synonym of P. circinalis (Lehm. & Lindenb.) Lindenb., is a distinct species, with a limited distribution in northern New Zealand. This species has the coarse, nodular leaf cell trigones of P. circinalis, but has straight, non-coiled androecia, the male bracts toothed at the apex and united wit...
Cyanolophocolea echinella R.M.Schust. is an intriguing species, possessing some character states characteristic of the genus Chiloscyphus, such as the ability to produce both male and female gametangia on leading shoots, while having other features that are generic characters of Heteroscyphus, namely presence of gametangia on short, abbreviated lat...
A widely accessible working list of known plant species is considered the number one target of the Global Strategy for Plant Conservation. An index of published names of liverworts and hornworts between 2006 and 2008 is provided as part of a continued effort in working toward producing a preliminary world checklist of these plants. Included in the...
The name Plagiochila retrospectans (Nees ex Spreng.) Lindenb. has been mistakenly applied to a common New Zealand species, properly known as P. fuscella (Hook.f. & Taylor) Gottsche Lindenb. & Nees, a species evidently restricted to New Zealand. Plagiochila viridio-nigra (E.A.Hodgs.) Inoue is a distinct species. These three species are described and...
A new liverwort species from Western Nelson Ecological Province of New Zealand known from a single sterile specimen is described. It is unusual in having well-developed stem paraphyllia, and the leaves and underleaves are divided with three levels of ciliate division in a pinnate manner. This morphology suggested membership of Trichocoleaceae or Tr...
Stolonivector obtusilobus J.J.Engel and S. clasmatocoleoides J.J.Engel, two new members of the Lophocoleaceae from New Zealand, are described and illustrated. Included is a discussion of affinities of both species with S. fiordlandiae (Hodgs.) J.J.Engel and S. waipouensis J.J.Engel. The genus has four species, all endemic to New Zealand. The genus...
Chiloscyphus anisolobus J. J. Engel & Glenny, a new member of the Lophocoleaceae, is described and illustrated from New Zealand. Chiloscyphus anisolobus J. J. Engel & Glenny shares the possession of a dorsal leaf free strip and strongly anisophyllous habit with the Australian C. excisifolius, but is distinct from that species in the accessory armat...
The hepatic genus Tritomaria (Jungermanniaceae subfam. Lophozioideae) is new to the south temperate. Tritomaria exsecta subsp. novaezelandiae subsp. nov. is described and illustrated from New Zealand. The distribution of each of the eight species of the genus is summarized.
The following new infrageneric taxa are described: Bazzania S. Gray sect. Glaucobazzania J. J. Engel (Lepidoziaceae) and Cephalozia (Dumortier) Dumortier subg. Eocephalozia R. M. Schuster ex J. J. Engel (Cephaloziaceae). Solenostoma cryptogynum R. M. Schuster ex J. J. Engel (Jungermanniaceae), C. austrigena R. M. Schuster ex J. J. Engel, C. schuste...
Presence of a well-developed perianth, ciliate at the mouth, and brown pigmentation necessitate establishment of a new liverwort genus, Castanoclobos J. J. Engel & Glenny. The only species, C. julaceus (Hatcher ex J. J. Engel) J. J. Engel & Glenny, which is endemic to New Zealand, is described and illustrated. The hepatic genus is retained in the T...
Seven species of Marsupidium are recognized in New Zealand including two new species, M. papillosum and M. plagiochiloides. Marsupidium perpusillum var. denticulatum, is also newly recognized. Keys are provided for the species and varieties.
Dinckleria Trevis. is an older name for Proskauera Heinrichs & J.J.Engel. The genus has two species, D. pleurata (Hook.f. & Taylor) Trevis. and D. fruticella (Hook.f..& Taylor) J.J.Engel & Heinrichs. Both species are based on New Zealand plants.
Leptophyllopsis R. M. Schust., with particular reference to L. laxus (Mitt.) R.M. Schust. of New Zealand, Tasmania and southeast Australia, a member of the Lophocoleaceae, is described and illustrated. The genus is placed in a new subfamily, Leptophyllopsoideae J.J. Engel.
Scanning electron micrographs reveal a striking similarity of cell wall characters among Mylia taylorii, M. verrucosa, M. nuda and Trabacellula tumidula. All of these taxa have the presence of irregularly shaped strips and plates of smooth wall material, each separated by grids that interconnect the plates and strips. These species all also share t...
Telaranea is the third largest genus of Lepidoziaceae, after Bazzania (over 450 species) and Lepidozia (over 300 species). In this monograph, 98 species of Telaranea are recognized, 62 of which are extra-Australasian. Included are 11 new species, 2 new varieties, 27 new combinations (25 species and one variety), and one new name. New taxa are fully...
Physotheca autoica J. J. Engel & Gradst. is a liverwort genus and species new to science and is endemic to the Andes of southern Ecuador. The new genus, while having several characters also found in the family Geocalycaceae, differs from members of that family in a number of important respects, which are discussed. The mode of growth of Physotheca...
Chiloscyphus sect. Novae-zeelandii, sect. Semiteres, sect. Spiniferi, and sect. Hemispini are described as new. Chiloscyphus subg. Connati, subg. Microlophocolea, and subg. Cyanolophocolea new combinations.
Telaranea consobrina, T. fragilis, and T. palmata, new species from Australasia, are described and illustrated. Telaranea clatritexta (Stephani) Engel & Merrill and T. quadricilia (Stephani) Engel & Merrill of Australasia and T. fernandeziensis (Stephani) Engel & Merrill from Juan Fernández are new combinations. Frullania neocaledonica of New Caled...
A new species, Leiomitra julacea, is proposed for the New Zealand Trichocolea julacea, which is invalid according to the International Code of Botanical Nomenclature. The species is described and illustrated.
Plagiochila bazzanioides, a new species from New Zealand, is described and illustrated.
Three species of Anastrophyllum are shown to occur in temperate Australasia. Anastrophyllum papillosum J. J. Engel & Braggins sp. nov. is described from New Zealand; the species differs from A. novazelandiae R. M. Schust. by characters that include branching, leaf cuticle, size of basal leaf cells and oil-bodies. Anastrophyllum schismoides (Mont.)...
Paracromastigum fiordlandiae Schust. & Engel from New Zealand is described. It approachesHyalolepidozia in the almost uniformly intercalary branching but differs from it in the intermingled 2–3-lobed leaves and the stem with
9–12 rows of cortical cells.Paracromastigum andHyalolepidozia are intimately allied toPseudocephalozia Schust., and one could...
Bazzania nova Engel & Merr. sp. nov. is described from New Zealand; Bazzania involuta var. submutica (Lindenb. & Gott.) Engel & Merr. and B. adnexa var. aucklandica (Lindenb. & Gott.) Engel & Merr. are new combinations. Acromastigum cunninghamii (Steph.) Evans is reported as new to New Zealand, and the presence of Bazzania monilinervis (Lehm. & Lin...
Haplomitrium minutum (E. Campb.) Engel & Schust. comb. nov. is treated as a valid species. The species is known only from artificial sites in New Zealand and is allied to Haplomitrium hookeri (Sm.) Nees. Steereomitrium is shown to exactly match Haplomitrium and is placed into synonymy of the latter.
Engel, J. J.: Index hepaticarum supplementum: 1986–1987. — Taxon 42: 373–391. 1993. ‐ISSN 0040–0262.
In this Supplement, there are references to 173 new taxa, 67 new combinations (including changes in rank), and 4 new names.
Engel, J. J.: Index hepaticarum supplementum: 1984–1985. ‐ Taxon 41: 253–306. 1992. ‐ ISSN 0040‐0262.
In this Supplement, there are 313 new taxa, 423 new combinations (including changes in rank), and 23 new names.
The Geocalycacae include, among other generic complexes, a complex of genera with well-developed but laterally compressed perianths; the third (ventral) face is strongly reduced, so that the perianth appears bilabiate. Heretofore known in this complex are Leptoscyphus Mitt. and Pedinophyllopsis Schust. & Inoue. An additional genus, Evansianthus, wi...
The taxonomic position of Pigafettoa, a poorly known monotypic genus occurring only in southern South America, is evaluated and it is shown that the taxon belongs in the Geocalycaceae. A key to the genera of the Clasmatocolea-Xenocephalozia complex is given.
In southern Chile, the authors collected a rather select group of Hepaticae which was subject to either tidal action or ocean spray. Hepaticae of the tidal region are chiefly weedy, polytypic, polystenic, widely distributed taxa. Drainage from the forested areas directly above the beaches and the high rainfall of the region are sources of fresh wat...
