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Initial spicular system of Coronatubopyle spelae Dumitrica & Tekin n.sp. in left lateral view. A apical bar, V ventral bar, D dorsal bar, Ll left primary lateral spine, lr right secondary lateral spine
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This article is a taxonomic study of the radiolarian species of the superfamilies Eptingiacea and Saturnaliacea occurring in the middle Carnian fauna from the Köseyahya section, near the town of Elbistan, southeastern Turkey. This fauna is characteristic of the Tetraporobrachia haeckeli Radiolarian Zone as defined in Austria and later found also in...
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Abundant, very diverse, and well-preserved Nassellarian fauna has been obtained from cherty/clayey limestone levels at the basal part of the Kö seyahya nappe east of the town of Elbistan, Eastern Taurides. A comparison with the radiolarian faunas of Austria, Japan and Turkey allows us to assign a middle Carnian age to this radiolarian fauna from th...
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The present article is a taxonomic study of all spongy spumellarian radiolarian taxa with three and four coplanar spines or spongy arms occurring in the middle Carnian from the Köseyahya section, near the town of Elbistan, SE Turkey. This fauna is characteristic of the Tetraporobrachia haeckeli radiolarian Zone, and comes from an 8 m thick successi...
BRACHIOPODS are generally rare in Eocene benthic assemblages, but they have been recorded from several countries in Europe and numerous papers demonstrate their wide geographical distribution within the Tethys Ocean. Without the claim of completeness, Eocene brachiopods have been described. However, until now Eocene brachiopods have not been record...
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... In addition, diabase dykes were mentioned to occur in the middle part of the sequence by Monod (1977). Recently, the Huglu Limestone of late Carnian-Liassic age above the Huglu Tuffite was studied in detail based on radiolarians (Tekin andBedi, 2007a, 2007b;Dumitrica et al., 2010Dumitrica et al., , 2013aDumitrica et al., , 2013bDumitrica and Tekin, 2014;Dumitrica, 2017) and different fossil assemblages from the Köseyahya Nappe, near the town of Elbistan, eastern Turkey and based on radiolarians, conodonts and ammonoids in the Tavuscayiri block from the Mersin Melange, southern Turkey. ...
... Similar to this radiolarian dating from the Huglu Tuffites, radiolarian assemblages were obtained from pelagic limestones within Huglu-type tuffs near Elbistan, eastern Turkey, and radiolarians of these limestones were documented in a series of publications (Tekin andBedi, 2007a, 2007b;Dumitrica et al., 2010Dumitrica et al., , 2013aDumitrica et al., , 2013bDumitrica and Tekin, 2014;Dumitrica, 2017). Age of this fauna has been assigned to the latest Early Carnian, corresponding to the Elbistanium gracilum Radiolaria Zone (Fig. 9). ...
... Some of the taxa, such as Paraparonaella okuyucui, Ropanaella nitida, R. ygraeca Triassoastrum paratrammeri, Carinacyclia acari, Praeorbiculiformella parvicentrum, Recoarella lahmi, Relindella lenticulata, Tetraporobrachia bedii, Haeckelicyrtium planum from sample Kilek-23 were previously described from the upper Lower Carnian strata from eastern Turkey (Tekin and Bedi, 2007b;Dumitrica et al., 2013aDumitrica et al., , 2013bDumitrica, 2017), corresponding to the Elbistanium gracile Radiolaria Zone by Moix et al. (2007). Some additional taxa belonging to this zone, have also been reported by some studies (e.g., Tekin and Bedi, 2007a;Dumitrica et al., 2010) based on material also from Elbistan, SE Turkey. Although an early Late Carnian age was assigned to the radiolarian assemblage belonging to this zone by Moix et al. (2007), the age of radiolarian assemblage belonging to this zone has subsequently been reported as the latest Early Carnian by Dumitrica et al. (2013aDumitrica et al. ( , 2013b as it was first assigned in Tekin andBedi (2007a, 2007b). ...
... The radiolarian assemblages documented here share close affinities with those from western Tethys (De Wever et al., 1979;Tekin, 1999Tekin, , 2002Bragin, 2007;Dumitrica and Hungerbühler, 2007;Kozur et al., 2007aKozur et al., , 2007bKozur et al., , 2007cNicora et al., 2007;Dumitrica et al., 2010) with very common spumellarians, mainly represented by three very abundant genera, Capnodoce, Capnuchosphaera and Sarla, less abundant nassellarians, with Mostlericyrtium, Spinosicapsa and Xiphotecaella, and rare entactinarians, with Carinaheliosoma. There are substantially fewer overlapping taxa with assemblages documented in Japan (Sugiyama, 1997) and even less overlap with assemblages from other Panthalassan locations such as the Philippines (Yeh, 1990(Yeh, , 1992, Baja California (Pessagno et al., 1979), Oregon (Blome, 1984;Yeh, 1989) and British Columbia (Carter, 1993;Carter andOrchard, 2000, 2013). ...
... Even vertebrates have potential with one ichthyosaur bone from USGS Mesozoic locality 9387, collected by Elliot Blackwelder in 1915, which has not been described and is illustrated here for the first time (Fig. 12). Further detailed study of the microfauna, notable radiolarians, appears to be highly profitable (Robinson, 1989;Robinson and Pessagno, 1988;Dumitrica et al., 2010). Determination of the character of the contact with the upper part (Jurassic and Lower Cretaceous) of the Glenn Shale is critical, and it seems likely that further study indicates that the lower and upper parts of Brabb's formation deserve revision with the upper portion needing formal designation as a new, separate lithostratigraphic unit. ...
... The radiolarian assemblages from this section are well-correlative to the circum-Pacific and Tethyan radiolarian assemblages. The radiolarian zonations by Carter (1993) and Carter et al. (1998, 2010 were applied to this study for the Rhaetian to Pliensbachian time interval. ...
The Mersin Mélange (northwest of Mersin city) includes a variety of large sedimentary blocks/tectonic slices of Palaeozoic and Mesozoic origins. Of these, the latter represents facial and tectonostratigraphic counterparts of the Beysehir-Hoyran Nappes (remnants of the northern Neotethys). The
Tavuscayiri Block, located at the center of the mélange and close to the Orbuklukeli hill, is one of
such Mesozoic occurrences, with a continuous pelagic sequence from the Upper Triassic to Lower
Jurassic. At the Orbuklukeli hill, a succession of middle Norian to Toarcian age has been precisely
dated, which starts with conodont assemblages for the Norian and Rhaetian and includes radiolarians
for the upper Rhaetian. An acidic tuff layer corresponds to the T-J boundary, which passes above to
an increasingly dominating chert-bearing limestone series, marking a gradually deepening-upward
of the sequence. The early Hettangian radiolarians are poorly represented, but diverse and well-preserved radiolarians have been retrieved from the middle Hettangian to the Pliensbachian. A total of
eighty-five taxa, including fourteen new species, have been determined. In addition to these, one new
genus (Praeudalia Tekin, n. gen.) has been described from the Rhaetian part of the section. The top
of the section is represented by nodular limestones in Ammonitico rosso facies, including a diverse Toarcian ammonite fauna. All lithologies of the Orbuklukeli section along the Tavuscayiri Block
can be correlated with the previously described lithologies of the Kayabasi Group/Formation in the
Bozkir Unit, and Gülbahar/Gümüslü units in the Lycian Nappes.
... Within Lineage III, including clades G, H and I, Clade G is represented by 5 novel and 3 previously available sequences. Members of this clade have a very unique innermost shell, named "rhizosphaerid-type microsphere" or "microbursa" (Dumitrica et al. 2010) with a large and fragile spherical skeleton ( Fig. 2. Ga) or several cortical shells with a robust skeleton (Fig. 2. Gb). Either morphology agree with the definition of the family Rhizosphaeridae (Hollande and Enjumet, 1960;Dumitrica, 2017). ...
Spumellaria (Radiolaria, Rhizaria) are holoplanktonic amoeboid protists, ubiquitous and abundant in the global ocean. Their silicified skeleton preserves very well in sediments displaying an excellent fossil record, from the early middle Cambrian (ca. 509-521 Ma), extremely valuable for paleo-environmental reconstruction studies. Spumellaria are tedious to maintain in laboratory conditions preventing an accurate perception of their extant diversity and ecology in todays oceans, most of which being inferred from sediment records. This study represents an integrated and comprehensive classification of Spumellaria based on the combination of ribosomal taxonomic marker genes (rDNA) and morphological characteristics. In contrast to established taxonomic knowledge, we demonstrate that symmetry of the skeleton takes more importance than internal structures at high taxonomic rank classification. Such reconsideration allows gathering different morphologies with concentric structure and a spherical or radial symmetry believed to belong to other Radiolaria orders from the fossil record, as for some Entactinaria families. Results obtained in this study suggest the existence of new Spumellaria diversity at early diverging positions, in which a non-bearing skeleton organism lives within shelled ones. Using fossil calibrated molecular clock we estimated the origin of Spumellaria in the middle Cambrian (ca. 515 Ma), in agreement with the appearance of the first radiolarian representatives in the fossil record. The morpho-molecular and evolutionary framework established herein allows a direct connection between living specimens and fossil morphologies from the Cambrian, bringing both a standpoint for future molecular environmental surveys and a better understanding for paleo-environmental reconstruction studies.
... Subsequently, a middle Carnian age was assigned by Tekin (1999) based on radiolarians from the limestone layers within this unit near the town of Bozkir in the central Taurides. Radiolarians from pelagic limestones adjacent to tuff layers have been dated as latest Early Carnian (based on the two-fold Carnian subdivision) or latest Middle Carnian (based on the three-fold Carnian subdivision) by Tekin & Bedi (2007a, b), Dumitrica et al. (2010Dumitrica et al. ( , 2013a, Dumitrica & Tekin (2014) and Dumitrica (2017), which corresponds to the Elbistanium gracilum radiolarian Zone near the town of Elbistan in eastern Turkey (Fig. 4). ...
... A radiolarian assemblage similar to that obtained from the basal part of the Tavuscayiri section has been recovered from sample Kilek-23 in the Kilek section (e.g. Pseudohagiastrum cordevolicum (Kozur & (Kozur & Mostler 1972, 1978, 1979 and from Turkey (Tekin 1999;Tekin & Bedi 2007a, b;Tekin & G€ onc€ uoglu 2007;Varol et al. 2007;Dumitrica et al. 2010Dumitrica et al. , 2013aSayit et al. 2011Sayit et al. , 2015Tekin et al. 2012;Dumitrica 2017). The main part of the fauna correlates well with faunas obtained from the upper Lower Carnian (based on the two-fold Carnian subdivison) or middle Middle Carnian (based on the three-fold Carnian subdivison), indicating the Tetraporobrachia haeckeli Zone in previous studies (e.g. ...
The Mersin Mélange, located in southern Turkey north-west of the city of Mersin, includes blocks and tectonic slices of different origins. The Kilek section in the Mersin Mélange was sampled for a thorough examination of its lithology, biostratigraphy and fossil content. Two samples from the cherty limestone layers within the Huglu Tuffites at the top of the section yielded a rich silicified ostracod fauna of late Early Carnian (based on a two-fold Carnian subdivision) or middle Middle Carnian age (based on a three-fold Carnian subdivision), deposited in an open marine environment, in the outer platform-upper slope zone. We report 121 ostracod species belonging to 53 genera. Two new genera are described: Edithobairdia Forel gen. nov. and Gencella Forel gen. nov., as well as 16 new species: Acanthoscapha mersinella Forel sp. nov., Bairdia hugluensis Forel sp. nov., Acratia kollmanni Forel sp. nov., Citrella? carniana Forel sp. nov., Cytheropteron? schornikovi Forel sp. nov., Eucytherura lacerata Forel sp. nov., Gencella taurensis Forel sp. nov., Kerocythere dorsidenticulata Forel sp. nov., Kerocythere tricostata Forel sp. nov., Monoceratina praevulsaformis Forel sp. nov., Patellacythere tourkosella Forel sp. nov., Polycope kilekensis Forel sp. nov., Ptychobairdia praekristanae Forel sp. nov., Simeonella daginikella Forel sp. nov., Spinomicrocheilinella reliquiaella Forel sp. nov. and Triassocythere tavuscayiriensis Forel sp. nov. The diagnosis of Acratia goemoeryi Kozur is emended. The Kilek fauna retains primitive characteristics illustrated by the first known occurrence of Palaeocopida and Rectonariidae (typical Palaeozoic forms) in the Late Triassic, associated with typical Triassic–modern elements such as thick-shelled and ornamented Bairdiidae and diverse Cytheroidea known from the Middle and Late Triassic worldwide. The unique composition of Palaeozoic and Mesozoic taxa from the Kilek section illustrates unexpected long-term survival in a deep-sea refuge zone following the end-Permian extinction, and the diachronous character of the ostracod recovery in different environments.
http://zoobank.org/urn:lsid:zoobank.org.pub:662C3D5C–2B86–4D7B–BDB5–8F8B6A1AD1E7
... The correlation is based on the occurrence of Paragondolella noah in sample G11. An age equivalent radiolarian fauna has been partly described by Ozsvárt et al. (Ozsvárt et al. 2017a) from the Kopría Mélange, Rhodes, Greece (Moix et al. 2008) and by Dumitrica et al. (2010Dumitrica et al. ( , 2013a and Dumitrica and Hungerbühler (2007) from cherts of the Zulla Formation, Hawasina Complex, Oman. ...
This article is a continuation of the taxonomic study of the exquisitely preserved and extremely rich Late Carnian (early Tuvalian, Spongotortilispinus moixi Zone) radiolarian fauna of the sample G 11 from the Sorgun Ophiolitic Mélange occurring in the Tavusçayiri block, Pindos-Huğlu series, Tauride ophiolite belt, Turkey. 26 species of Spumellaria with spongy test, belonging to 7 genera, are discussed and illustrated. Among them 14 species and two genera (Staurotortilispinus n. gen and Ancoraspongus n. gen) are introduced as new. The assemblage provides new data on the diversity of the radiolarian fauna of the Tethys during the Late Carnian and contributes to a better understanding of the Mesozoic geodynamic evolution of the Mediterranean region.
... For example, Ujiié and Oba (1991a) reported only one occurrence site of Triassic radiolarians; Shen et al. (1996) reported the Triassic radiolarian occurrences including Palaeosaturnalis triassicus (Kozur and Mostler) from three localities. Palaeosaturnalis triassicus ranges from the middle Carnian to the middle Norian (Dumitrica et al., 2010). In addition, Shen et al (1996) discovered Kilinora spiralis (Matsuoka), which is the diagnosis species of the K. spiralis Zone (JR6) of the Oxfordian, Upper Jurassic (Matsuoka, 1995), in the bedded cherts near the crest of Mt. ...
Ie Island in northwestern Okinawa Prefecture, Japan, mainly comprises of basement rocks (Gusukuyama Formation of the Southern Chichibu terrane) and the overlying Pleistocene Ryukyu Group. The bedded cherts of the Gusukuyama Formation in Mt. Gusuku in the central part of Ie Island have yielded radiolarians ranging from the Neoalbaillella optima Zone (Changhsingian, Lopingian, Permian) to the Kilinora spiralis Zone (JR6: Oxfordian, Upper Jurassic). Siliceous mudstones in the southern flank of Mt. Gusuku have yielded radiolarians of the Pseudodictyomitra carpatica Zone (KR1: uppermost Jurassic–lowermost Cretaceous). This age difference is assumed to indicate the presence of a thrust between Mt. Gusuku and the southern flank of the mountain. Red chert clasts are contained in the Pleistocene Ryukyu Group at Waji on the north coast of Ie Island, and Permian radiolarians have been obtained from chert clasts. In addition, a red bedded chert boulder at Waji yielded Cisuralian (Early Permian) radiolarians including dimorphic pair of Albaillella sinuata Ishiga and Watase.
... Sample G 11 contains the best-preserved and most diverse radiolarian fauna of the basal Tuvalian (Spongotortilispinus moixi Zone) anywhere in the world (Fig. 4). An equivalent radiolarian fauna was partly described by Dumitrica et al., (2010Dumitrica et al., ( , 2013a2013b) from the Zulla Formation, Hawasina Complex, Oman. Sample G19 yielded a few specimens of Annulotriassocampe spp. ...
... Similarly rich and very well-preserved but older radiolarian faunas, belonging to the Julian Tetraporobrachia haeckeli Zone were published from Göstling and Großreifling (Northern Calcareous Alps, Austria) by Kozur and Mostler (1972;1978;1981;1994) and Lahm (1984). However, the best preserved and richest fauna of this age, from the Köseyahya Nappe in Elbistan, Turkey, has been only partly published thus far (Tekin and Bedi, 2007a;Dumitrica et al., 2010;2013a;2013b;Dumitrica and Tekin, 2014). The composition of this fauna from Turkey is rather similar to that of the faunas from Göstling and Großreifling, but the radiolarians are more abundant and better preserved. ...
... The composition of this fauna from Turkey is rather similar to that of the faunas from Göstling and Großreifling, but the radiolarians are more abundant and better preserved. These faunas represent the temporal range of the species Tetraporobrachia haeckeli and the family Spongosaturnaloididae Kozur and Mostler, as well as that of the genus Heliosaturnalis Kozur and Mostler, and the acme of the families Veghicycliidae Kozur and Mostler, (Dumitrica et al., 2010) and Multiarcusellidae Kozur and Mostler. At present it is difficult to compare in detail these faunas of the T. haeckeli Zone with the early Tuvalian faunas of the Spongotortilispinus moixi Zone, because a major part of the faunas from Elbistan and from the SOM remains still unpublished. ...
Three new radiolarian genera (Paramonocapnuchosphaera, Monocoronella and Minicrampus) and 8 new species are described herein from one of the most diverse and best preserved Carnian (Late Triassic) radiolarian faunas of the world, from the early Tuvalian sequence of the Sorgun Ophiolitic Mélange, southern Turkey. The radiolarian fauna belongs to the Spongotortilispinus moixi Zone (equivalent to Paragondolella postinclinata - Paragondolella noah conodont Zone). The very similar late Julian-early Tuvalian radiolarian faunas from the Northern Calcareous, Alps through the Lagonegro Basin, Sicily, Rhodes Island, Greece and farther east in the Antalya nappes, in the Mersin Mélange and in Elbistan, Turkey may confirm that the Pindos-type succession found in different units of early Mesozoic oceanic basins formed in a single oceanic basin.
... Sample G 11 contains the best-preserved and most diverse radiolarian fauna of the basal Tuvalian (Spongotortilispinus moixi Zone) in the world (Fig. 4). An equivalent but not so rich radiolarian fauna has been partly described by Dumitrica et al. (2010Dumitrica et al. ( , 2013a and Dumitrica and Hungerbühler (2007) similar to the radiolarian fauna of Sorgun Ophiolitic Mélange in both diversity and preservation, the difference between them being insignificant. ...
... cf. C. omanensisDumitrica and Tekin, 2010 + Coronatubopyle spelaeDumitrica and Tekin, 2010 + Hexapylomella carnicaKozur and Mostler, 1979 + Hindeosphaera burriOzsvárt, Moix and Kozur, 2015 + + Hindeosphaera cf. H. goestlingensisKozur and Mostler, 1979 Hypoxiphothecaella inflata Kozur, Moix and Ozsvárt, 2007a + Hypoxiphothecaella mersinensis Kozur, Moix and Ozsvárt, 2007a + Minicrampus longispinosus n. gen. ...
... cf. C. omanensisDumitrica and Tekin, 2010 + Coronatubopyle spelaeDumitrica and Tekin, 2010 + Hexapylomella carnicaKozur and Mostler, 1979 + Hindeosphaera burriOzsvárt, Moix and Kozur, 2015 + + Hindeosphaera cf. H. goestlingensisKozur and Mostler, 1979 Hypoxiphothecaella inflata Kozur, Moix and Ozsvárt, 2007a + Hypoxiphothecaella mersinensis Kozur, Moix and Ozsvárt, 2007a + Minicrampus longispinosus n. gen. ...
