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Triphora superba Thiele, 1925, Station 104 (Agulhas Bank). A-C, E. Holotype, ZMB/Moll no. 109266: front (A), side (B), back (C), protoconch (E). D. Original figure in Thiele 1925. F. Original label. Scale bar: A-C: 0.5 mm, E: 0.2 mm.
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Triphoridae is a family of marine caenogastropods with worldwide distribution. Its maximum diversity is in the Indo-Pacific province, where it is among the five most species-rich families. Taxonomic knowledge is scant and complicated by the high diversity and intra-specific variability. Knowledge of type specimens of described taxa is the fundament...
Citations
... In the fossil record, as early as in the upper Oligocene of France, Lozouet (1999) described species with an Inella-type strongly carinate protoconch. Landau et al. (2018) illustrated an enormous diversity in protoconch morphology in triphorids from the Tortonian upper Miocene of north-western France, much more like the sort of protoconch diversity seen in present-day species from South and East Africa (Albano & Bakker, 2016), Caribbean (Rolán & Fernández-Garcés, 2008) and Australia and New Zealand (Marshall, 1983). ...
... Most species with a planktotrophic larval development are generally more widely dispersed than their non-planktotrophic descendants. The paucispiral protoconchs of non-planktotrophic gastropods are very variable in size, shape and sculpture, and similar morphologies have evolved repeatedly throughout geological time and in present-day faunas (Marshall, 1983;Rolán et al., 2008;Albano & Bakker, 2016;Philippe Bouchet and Bruce Marshall personal communication in Landau et al. 2018: 218). Therefore, sharing a protoconch suggesting a non-planktotrophic larval development does not necessarily imply a phylogenetic relationship. ...
... The sharp drop in protoconch diversity from the Neogene to present-day European and West African triphorid species does not seem to be the case in the eastern Atlantic, nor in the southern hemisphere where protoconch diversity is very high, especially around South (and far West) South Africa and Australia (Thiele, 1925;Marshall, 1983;Albano & Bakker, 2016). This circumstance cannot be explained by the decrease in SSTs alone, as the Estepona Basin faunas were part of the tropical Pliocene Mediterranean-West-African Paleobiogeographic Province, and the roughly coeval Mondego Basin fauna was part of the subtropical Pliocene French-Iberian Province. ...
In this paper we record a species of the Inella group for the first time in the Pliocene Mediterranean with the descrip�tion of Inella bentae spec. nov. from the Estepona Basin, and Inella spec. and Triphora (s. l.) freixiensis spec. nov. from the Mondego Basin
... Due to the large number of described species, recent type revisions (e.g. Albano & Bakker, 2016;Albano et al., 2019), and the evolving taxonomy of Triphoridae, there is a degree of uncertainty associated with some of the identifications provided here. Where an identification is provisional, the abbreviation "cf." (from the Latin confer, compare) is utilized. ...
The extant Triphoridae of the Samoan archipelago are reviewed, and 21 new
Samoan records are discussed and illustrated based on recent collections from the island
of Tutuila, American Samoa. Confident identifications are provided for the following 9
species: Bouchetriphora pallida (Pease, 1871); Coriophora cnodax (Jousseaume, 1884);
Coriophora fusca (Dunker, 1860); Costatotriphora iniqua (Jousseaume, 1898); Iniforis
fusiformis (Kosuge, 1961); Mastonia iris Laseron, 1958; Nanaphora triticea (Pease,
1861); Triphora collaris Hinds, 1843; and Triphora taeniolata Hervier, 1898. Provisional
records are provided for 7 species: Inella cf. numerosa Jousseaume, 1898; Iniforis cf. douvillei Jousseaume, 1884; Mastoniaeforis cf. chaperi Jousseaume, 1884; Nanaphora cf. tricolor Laseron, 1958; Opimaphora cf. coralina Laseron, 1958; Triphora cf. fulvescens
Hervier, 1898; and Metaxia cf. brunnicephala Kay, 1979. A further 5 species are recorded
from Samoa that cannot be definitively assigned to currently described taxa and may represent undescribed species: Monophorus aff. strictus (Laserson, 1958); Nanaphora aff.
triticea (Pease, 1861); Obesula aff. pantherina Jousseaume, 1898; Opimaphora aff.
coralina Laseron, 1958; and Triphora aff. laddi Kay, 1979. Two species previously
recorded from Samoa based on tentatively identified fossil material are now validated as
currently present: Iniforis albogranosa (Kosuge, 1961) and and Viriola pagoda (Hinds,
1843). Of the 13 other species previously recorded from Samoa, 2 are currently considered synonyms; 4 are questionable records not based on any specimens that can be located; 2 are questionable names that may represent synonyms; 1 is a likely misidentification;
and 4 are valid records of which one, Mastonia cf. rubra (Hinds, 1843), was recollected.
Based on this analysis, the currently documented triphorid fauna of Samoa consists of 33
species, although additional species are undoubtedly present. The current total is also subject to change as taxonomic and nomenclatural problems are resolved.
... Previous studies of type material (Albano & Bakker 2016;Albano et al 2017Albano et al , 2019 already provide information on 162 Triphoridae species preserved in the Museum für Naturkunde in Berlin, the Naturhistorisches Museum Wien in Vienna, and in the Natural History Museum of the United Kingdom in London. Here, additional information is provided on four species names of Triphoridae and a listing of the types of 20 Atlantic species deposited in the collection of Naturalis Biodiversity Center (NBC), Leiden, the Netherlands. ...
The Naturalis Biodiversity Center (NBC) holds type specimens of 24 taxa of worldwide Triphoridae. For historical specimens (pre-World War II) the species name in its original combination is provided, followed by bibliographic details of the original description, the location of the known type material, the original description, a diagnosis, nomenclatural notes and illustrations of the type specimens and original labels. For more recently described species information is provided on the type locality and type material. The name Triphora raoulensis is an unpublished manuscript name by T. Iredale, “co-type” specimens are deposited in the NBC and National Museum of Wales (NMW), and the name is considered unavailable.
... However, triphorid species are mainly distributed from tidal zone to a depth of about 200 m in temperate and tropical seas [Marshall, 1983]. This is a family with various sizes, ranging 2-50 mm, but most of them belong to the small size group of 2-10 mm [Albano et al., 2016]. The shell has a conical shape. ...
Species of the genus Mastoniaeforis are mainly distributed in the Indo–West Pacific with ten species being recorded in the world. To date this genus was not recorded in Vietnam. Specimens of three species, M. chaperi, M. lifuana and M. speciosa, were collected in two field trips from May, 2019 to October, 2020 in Truong Sa archipelago (Spratly Islands), Khanh Hoa Province, Vietnam. The diagnostic characters, distribution, and illustrations of three Mastoniaeforis species in Vietnam are provided.
... was described from Cuba as Cerithium pusillum, with a brief description and no original illustrations (Pfeiffer, 1840). Its type material was recently illustrated in Albano and Bakker (2016), although these authors did not express an opinion about the validity of M. pusilla. The shell of M. pusilla is identical to that of M. modesta (Figure 4), described from Jamaica, and they may be considered synonyms. ...
Marine gastropods of the “Marshallora nigrocincta” species complex range from southeastern Canada to southern Brazil, comprising M. nigrocincta, described from the northeastern United States, and Marshallora modesta, from the Caribbean. This study aims to disentangle this complex, elucidating how many species are indeed present in the western Atlantic, and whether M. nigrocincta is different from M. modesta. The mitochondrial markers COI and 16S were congruent in splitting the complex into six lineages: one belonging to M. nigrocincta (mainly from eastern United States), other two lineages related to M. cf. modesta (one from Florida, USA, and the second from Colombia), and three undescribed species from Brazil. Intra‐ and interspecific genetic distances, respectively, vary from 0–2.7%/6.3–13.5% (COI). The nuclear gene 28S was conservative to separate Marshallora sp. A (Brazil) from M. cf. modesta 2, but recognized M. cf. modesta 1, M. nigrocincta, and Marshallora sp. B (Brazil). The shell morphology is highly variable, but the presence of three basal cords seems to occur only in M. nigrocincta. The radular morphology exhibits a wide variation in the number of cusps of the central and lateral teeth. The new species herein identified will require formal descriptions following further fieldwork efforts, since the selected voucher material was lost during a fire. New generic allocations are made to Marshallora amicorum comb. nov., Marshallora pusilla comb. nov., and Marshallora medinae comb. nov., which is regarded as nomen dubium. This study may stimulate the incorporation of DNA analysis in the systematics of Triphoroidea, improving the knowledge about its biodiversity.
... The depth of taxonomic assignment varies across taxa, mostly reflecting the available knowledge on these groups in the Indo-Pacific province (the source pool of most nonindigenous species in the Eastern Mediterranean). For families like the Triphoridae, some of us (PGA, PAJB, and BS) have been conducting taxonomic research for a long time and we have thus been able to describe new species as we have robust knowledge of inter-and intraspecific variability and of type specimens (Albano et al. 2011(Albano et al. , 2017Albano and Bakker 2016). For other families, like the Eulimidae, we focused our attention on highlighting differences from native species and similarities with Indo-Pacific species, because a more thorough coverage would have required revising the taxonomy of entire Indo-Pacific species-groups, a task well beyond our objectives. ...
... The species shows a broad distribution in the Eastern Mediterranean ranging from Greece to Turkey and Cyprus (Micali et al. 2017;Stamouli et al. 2017;Angelidis and Polyzoulis 2018;Chartosia et al. 2018). Its final taxonomic assignment requires the clarification of the relation between several other Viriola such as V. corrugata (Hinds, 1843), V. senafirensis (Sturany, 1903), and V. tricincta (Dunker, 1882) (Albano and Bakker 2016;Albano et al. 2017. ...
New data on 52 non-indigenous mollusks in the Eastern Mediterranean Sea is reported. Fossarus sp. (aff. aptus sensu Blatterer 2019), Coriophora lessepsiana Albano, Bakker & Sabelli, sp. nov. , Cerithiopsis sp. aff. pulvis, Joculator problematicus Albano & Steger, sp. nov. , Cerithiopsis sp., Elachisina sp., Iravadia aff. elongata, Vitrinella aff. Vitrinella sp. 1 (sensu Blatterer 2019), Melanella orientalis , Parvioris aff. dilecta, Odostomia cf. dalli, Oscilla virginiae , Parthenina cossmanni , Parthenina typica , Pyrgulina craticulata , Turbonilla funiculata , Cylichna collyra , Musculus coenobitus , Musculus aff. viridulus, Chavania erythraea , Scintilla cf. violescens, Iacra seychellarum and Corbula erythraeensis are new records for the Mediterranean. An unidentified gastropod, Skeneidae indet., Triphora sp., Hypermastus sp., Sticteulima sp., Vitreolina cf. philippi, Odostomia (s.l.) sp. 1, Henrya (?) sp., and Semelidae sp. are further potential new non-indigenous species although their status should be confirmed upon final taxonomic assessment. Additionally, the status of Dikoleps micalii , Hemiliostraca clandestina comb. nov. and H. athenamariae comb. nov. is changed to non-indigenous, range extensions for nine species and the occurrence of living individuals for species previously recorded from empty shells only are reported. Opimaphora blattereri Albano, Bakker & Sabelli, sp. nov. is described from the Red Sea for comparison with the morphologically similar C. lessepsiana Albano, Bakker & Sabelli, sp. nov. The taxonomic part is followed by a discussion on how intensive fieldwork and cooperation among institutions and individuals enabled such a massive report, and how the poor taxonomic knowledge of the Indo-Pacific fauna hampers non-indigenous species detection and identification. Finally, the hypothesis that the simultaneous analysis of quantitative benthic death assemblages can support the assignment of non-indigenous status to taxonomically undetermined species is discussed.
... 242, 6°34.8'S, 39°35.5'E, between Dar es Salaam and southern Zanzibar, depth not mentioned by Thiele, but given as 404 m by Albano & Bakker (2019). ...
... Other families of microgastropods that constitute the so-called 'Big Five' are being intensively studied from this region in the last years (e.g., Peñas & Rolán 2010;Cecalupo & Perugia 2013), especially after the great amount of material sampled by the expeditions conducted by the MNHN team. The chaotic situation and high numbers of already described triphorids of the western and central Pacific, in many cases avoiding a link between worn types and newly sampled specimens, encouraged the current reevaluation of types described from these regions (e.g., Albano & Bakker 2016;Albano et al. 2019). The same problem applies to the well-known "Caribbean" triphorid fauna, where several species are exclusively recognized by their old and worn types, requiring reevaluations and much more samplings. ...
The present study aims to fulfill the gap of taxonomic knowledge on Triphoridae from Brazil. We describe five new species (Isotriphora uncia sp. nov., Isotriphora leo sp. nov., Monophorus verecundus sp. nov., Sagenotriphora albocaput sp. nov., Similiphora lucida sp. nov.), report five species previously known only from the Caribbean and related areas (Cheirodonta dupliniana (Olsson, 1916), Eutriphora auffenbergi Rolán & Lee, 2008, Isotriphora tricingulata Rolán & Fernández-Garcés, 2015, Marshallora ostenta Rolán & Fernández-Garcés, 2008, Monophorus caracca (Dall, 1927) comb. nov.) and describe six morphotypes at the generic level (Isotriphora sp. 1, Marshallora sp. 1, Nanaphora sp. 1, Sagenotriphora sp. 1, Sagenotriphora sp. 2, Similiphora sp. 1). Remarks are made to some species previously recorded from Brazil, including the invalidation of records, problems of generic allocation and geographical range extensions. Maps of the geographical distribution are provided for the 65 currently recognized species of Triphoridae from Brazil. Of these, 31 species are endemic to Brazil and 58 inhabit the continental shelf vs only seven from the continental slope. A distinct geographical zone occurs in southeastern Brazil. A few species occur exclusively near the mouth of the Amazon River, whereas others inhabit a local biogenic reef, possibly serving as a biogeographical corridor that connects western Atlantic populations. Species of Isotriphora from Brazil are particularly common around oceanic islands, probably due to adopting intracapsular metamorphosis, which may have evolved in more than one evolutionary event.
... Costatophora granifera (Brazier, 1894) has several synonyms, two of which are from Thiele's work on Western Australian molluscs (see Thiele, 1930 andAlbano andBakker, 2016) and the rest are from NSW (Laseron, 1954). Based on the material observed, the morphological variation is substantial and might collapse the various morphs of Costatophora found in this study (KimMoll21,KimMoll213,KimMoll217). ...
... Costatophora granifera (Brazier, 1894) has several synonyms, two of which are from Thiele's work on Western Australian molluscs (see Thiele, 1930 andAlbano andBakker, 2016) and the rest are from NSW (Laseron, 1954). Based on the material observed, the morphological variation is substantial and might collapse the various morphs of Costatophora found in this study (KimMoll21,KimMoll213,KimMoll217). ...
... The Triphoridae of the Assemblage I fauna show a remarkable variety in protoconch type, both in multispiral and paucispiral types, much more varied than that seen in the eastern Atlantic and Mediterranean today (Bouchet, 1985). Indeed it is similar in diversity to that seen in the Triphoridae revised by Albano & Bakker (2016) mainly from South and East Africa, the Caribbean and Australia and New Zealand (Marshall, 1983). In triphorids, like in numerous other gastropod groups, planktotrophy is the ancestral condition, and all nonplanktotrophic taxa are derived from planktotrophic ancestors or their non-plaktotrophic descendents. ...
... The majority of species with a planktotrophic protoconch are generally more widely dispersed than their non-planktotrophic descendents. Paucispiral protoconchs are very variable in size, shape and sculpture, and similar forms have evolved repeatedly throughout the world in modern and fossil faunas (Marshall, 1983;Rolán et al., 2008;Albano & Bakker, 2016;Philippe Bouchet and Bruce Marshall personal communication). Therefore, similarity of non-planktotrophic forms rarely implies phylogenetic links. ...
... sp. is characterised by its protoconch made of two whorls; the first bulbous, covered in micropustules, the second carinate mid-whorl and its teleoconch sculpture of two equal tuberculate spiral cords, with spiral 2 delayed, appearing on the fourth teleoconch whorl. This type of protoconch has not been reported from any Neogene or present-day European species, but was reported by Albano & Bakker (2016) in the South African species T. brevis Thiele, 1925, which is also remarkably similar in shape, but differs in that spiral 2 is less delayed and becomes as strong as spirals 1 and 3 on later whorls. Etymology -Named after the type locality of St-Clément-de-la-Place. Triphora gender feminine. ...
In this paper we review the Caenogastropoda of the Tortonian upper Miocene (Assemblage I of Van Dingenen et al., 2015) of northwestern France. One hundred and seventy-four species are recorded, of which 65 are new: Bittium crassum nov. sp., Bittium gallicum nov. sp., Bittium pingue nov. sp., Bittium renauleauense nov. sp., Semibittium brebioni nov. sp., Archimediella sancticlementensis nov. sp., Oligodia chauvereauensis nov. sp., Crassitonella lozoueti nov. sp., Acirsa sceauxsensis nov. sp., Papuliscala redoniensis nov. sp., Papuliscala presselierensis nov. sp., Payraudeautia obelixi nov. sp., Inella alia nov. sp., Inella rolani nov. sp., Monophorus renauleauensis nov. sp., Obesula marshalli nov. sp., Triphora buscheri nov. sp., Triphora chauvereauensis nov. sp., Triphora
fernandezgarcesi nov. sp., Triphora lherbettorum nov. sp., Triphora miopygmaea nov. sp., Triphora sancticlementensis nov. sp., Cerithiopsis cerithiopsoides nov. sp., Cerithiopsis esterae nov. sp., Cerithiopsis mira nov. sp., Dizoniopsis boucheti nov. sp., Seila
petasa nov. sp., Ataxocerithium turbineum nov. sp., Alvania armata nov. sp., Alvania couffoni nov. sp., Alvania fezata nov. sp., Alvania globosa nov. sp., Alvania insulsa nov. sp., Alvania josephineae nov. sp., Alvania lachrimula nov. sp., Alvania milletispinosa nov. sp., Alvania miocalasi nov. sp., Alvania miolactea nov. sp., Alvania napoleoni nov. sp., Alvania parasusieae nov. sp., Alvania redoniana nov. sp., Alvania renauleauensis nov. sp., Alvania subtiliangulosa nov. sp., Alvania susieae nov. sp., Alvania tenuisculpturata nov. sp., Alvania turtaudierei nov. sp., Pseudosetia wareni nov. sp., Pseudosetia sergegofasi nov. sp., Pseudosetia peyroti nov. sp., Pseudosetia ivolasi nov. sp., Onoba redoniensis nov. sp., Onoba incisa nov. sp., Onoba fragilis nov. sp., Rissoa decorticata nov. sp., Rissoa
torquata nov. sp., Powellisetia europaea nov. sp., Pusillina dollfusi nov. sp., Pusillina gallica nov. sp., Discopsis pseudocanui nov. sp., Tornus superlatus nov. sp., Teinostoma obesum nov. sp., Macromphalina morgani nov. sp., Trivia sceauxensis nov. sp., Niveria
cylindriclementi nov. sp. and Cleotrivia gallica nov. sp. Delphinula carinata Millet, 1854 is a junior homonym of D. carinata Woodward, 1833, and is renamed Alvania acuticarinata nov. nom. Rissoa suturalis Millet, 1865 is a junior homonym of R. suturalis Philippi, 1844, and is renamed Alvania milleti nov. nom. Rissoa notabilis Millet, 1865 is a junior homonym of R. notabilis C.B. Adams, 1852, and is renamed Rissoa obeliscoides nov. nom. Lacuna bourgeoisi Tournouër, 1874 is a junior subjective synonym of Phasianella delphinuloides Millet, 1865. Bithinella [sic] benoisti Dollfus & Dautzenberg, 1886 is considered a subjective synonym of Bithinella [sic] Fontannesi Dollfus & Dautzenberg, 1886; as first revisers the name Peringia fontannesi (Dollfus & Dautzenberg, 1886) is chosen. Tornus dollfusi Cossmann, 1918 and Tornus subcarinatus minor Glibert, 1949 are considered junior subjective synonyms of Tornus subcarinatus (Montagu, 1803).
Schilderia fasciolaria, S. incognita, S. lauriatae, S. veronicata and S. brebioni Dolin & Lozouet, 2004 are all considered to represent a single species, as first revisers the name Schilderia brebioni Dolin & Lozouet, 2004 is chosen as valid. We amend an omission made in part one of this series; Trochus echinatus Millet, 1854 is a junior homonym of Trochus echinatus Klöden, 1834, and is renamed Calliostoma milletechinatum nov. nom.
