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Trace fossils from the Upper Tiouririne Formation (Katian, eastern Moroccan Anti-Atlas). A -Asteriacites cf. lumbricalis von Schlotheim; B -Cruziana isp.; C -Arthraria antiquata Billings (upper left) and Cruziana isp.; DProtovirgularia dichotoma M'Coy; E -Teichichnus rectus Seilacher, a burrow fragment in oblique-lateral view; F -Heughia gregaria Smith; G -Rusophycus carleyi (James); H -Nereites cf. irregularis (Schafhäutl) in hyporelief preservation; I-J -Gyrochorte comosa Heer in epi-(I) and hyporelief (J) preservation; KTaenidium barretti (Bradshaw); LNereites cf. irregularis (Schafhäutl) in epirelief preservation; M -Nereites pugnus Emmons; N -Nereites macleayii (Murchison). Scale bars, 10 mm (A, B, F, G, K-N) and 5 mm (C-E, H-J).
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... trace-fossil association is diverse and pending detailed investigation. The following ichnotaxa could be identified: Arenicolites isp.; Arthraria antiquata Billings (Fig. 2C), Arthrophycus brongniartii (Harlan), Asteriacites cf. lumbricalis von Schlotheim ( Fig. 2A), Asterichnus lawrencensis Bandel, Cheiichnus isp., Cruziana problematica (Schindewolf), Cruziana isp. (Fig. 2B), Curvolithus simplex Buatois et al., Diplocraterion parallelum Torell, Gyro- Fig. 1. Neoeione moniliformis (Tate) from the Upper ...
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... trace-fossil association is diverse and pending detailed investigation. The following ichnotaxa could be identified: Arenicolites isp.; Arthraria antiquata Billings (Fig. 2C), Arthrophycus brongniartii (Harlan), Asteriacites cf. lumbricalis von Schlotheim ( Fig. 2A), Asterichnus lawrencensis Bandel, Cheiichnus isp., Cruziana problematica (Schindewolf), Cruziana isp. (Fig. 2B), Curvolithus simplex Buatois et al., Diplocraterion parallelum Torell, Gyro- Fig. 1. Neoeione moniliformis (Tate) from the Upper Tiouririne Formation (Katian, eastern Moroccan Anti-Atlas). A-C, G -photographs of some long ...
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... is diverse and pending detailed investigation. The following ichnotaxa could be identified: Arenicolites isp.; Arthraria antiquata Billings (Fig. 2C), Arthrophycus brongniartii (Harlan), Asteriacites cf. lumbricalis von Schlotheim ( Fig. 2A), Asterichnus lawrencensis Bandel, Cheiichnus isp., Cruziana problematica (Schindewolf), Cruziana isp. (Fig. 2B), Curvolithus simplex Buatois et al., Diplocraterion parallelum Torell, Gyro- Fig. 1. Neoeione moniliformis (Tate) from the Upper Tiouririne Formation (Katian, eastern Moroccan Anti-Atlas). A-C, G -photographs of some long specimens preserved in full relief and as convex epichnia. D-E -a wide trace in top (D) and lateral (E) views; F ...
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... Formation (Katian, eastern Moroccan Anti-Atlas). A-C, G -photographs of some long specimens preserved in full relief and as convex epichnia. D-E -a wide trace in top (D) and lateral (E) views; F -longitudinal section between three consecutive knobs showing massive sand infill. Scale bars, 50 mm (A-C, G) and 10 mm (D-F). chorte comosa Heer (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) ...
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... Anti-Atlas). A-C, G -photographs of some long specimens preserved in full relief and as convex epichnia. D-E -a wide trace in top (D) and lateral (E) views; F -longitudinal section between three consecutive knobs showing massive sand infill. Scale bars, 50 mm (A-C, G) and 10 mm (D-F). chorte comosa Heer (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, ...
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... long specimens preserved in full relief and as convex epichnia. D-E -a wide trace in top (D) and lateral (E) views; F -longitudinal section between three consecutive knobs showing massive sand infill. Scale bars, 50 mm (A-C, G) and 10 mm (D-F). chorte comosa Heer (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus ...
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... and as convex epichnia. D-E -a wide trace in top (D) and lateral (E) views; F -longitudinal section between three consecutive knobs showing massive sand infill. Scale bars, 50 mm (A-C, G) and 10 mm (D-F). chorte comosa Heer (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus carleyi (James) (Fig. 2G) and Rusophycus ...
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... -a wide trace in top (D) and lateral (E) views; F -longitudinal section between three consecutive knobs showing massive sand infill. Scale bars, 50 mm (A-C, G) and 10 mm (D-F). chorte comosa Heer (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus carleyi (James) (Fig. 2G) and Rusophycus ...
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... consecutive knobs showing massive sand infill. Scale bars, 50 mm (A-C, G) and 10 mm (D-F). chorte comosa Heer (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus carleyi (James) (Fig. 2G) and Rusophycus ...
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... (Fig. 2I-J), Heughia gregaria Smith (Fig. 2F), Nereites cf. irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus carleyi (James) (Fig. 2G) and Rusophycus ...
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... irregularis (Schafhäutl) (Fig. 2H, L), N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus carleyi (James) (Fig. 2G) and Rusophycus ...
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... N. macleayii (Murchison) (Fig. 2N), N. pugnus Emmons (Fig. 2M), Oravaichnium hrabei Plička and Uhrová, Protovirgularia dichotoma M'Coy (Fig. 2D), P. rugosa (Miller and Dyer), ?Ptychoplasma conica Pieńkowski and Uchman, Taenidium barretti (Bradshaw) (Fig. 2K), T. serpentinum Heer, Teichichnus rectus Seilacher (Fig. 2E), Rusophycus carleyi (James) (Fig. 2G) and Rusophycus ...
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Citations
... Ichnotaxonomy, intergradation patterns in shallow to deep marine settings occur between Oravaichnium, the protobranch bivalve burrow Protovirgularia and some Lockeia (Mángano et al., 1998;Gutiérrez-Marco et al., 2019;Naimi and Cherif, 2021;Stachacz et al., 2021). Protovirgularia-like burrows were built by pushing of the bivalve's foot into the sediment, subsequent anchoring, and pulling the shell with the foot (push and pull mechanism, Trueman et al., 1966aTrueman et al., , 1966bBuatois and Mángano, 2011). ...
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