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9 Topographic maps of the study area in N.E. Twente, the Netherlands, with the boundaries of the OLU. (a) Recent topographic map, 1 : 10,000 (source: Topografische Dienst Kadaster, Emmen 2003). The topography is indicated through light grey shading. Most of the area within the OLU is agricultural pasture, with some arable land and forest. The Agelerbroek is a wetland nature reserve with forest and meadows. The main stream system (Tilligterbeek-Hamburgerbeek-Hollandsegraven, actually one stream) connects the Agerlerbroek area with the Ottershagen area (northern section of the OLU). Both areas are low-lying plains and the connecting stream system flows through higher terrain. (b) The boundaries of the OLU depicted on a historical map from 1900, 1 : 25,000 (source: Topografische Dienst Kadaster, Emmen 1900). The purple areas are the extent of flooding after extremely high rainfall in 1998. The delineation of the OLU follows the original outlines of wetlands on the historical map and the extent of the 1998 flooding. (Reproduced with permission from Journal of Applied Ecology (2008) Vol. 45, pp. 1496-1503.)

9 Topographic maps of the study area in N.E. Twente, the Netherlands, with the boundaries of the OLU. (a) Recent topographic map, 1 : 10,000 (source: Topografische Dienst Kadaster, Emmen 2003). The topography is indicated through light grey shading. Most of the area within the OLU is agricultural pasture, with some arable land and forest. The Agelerbroek is a wetland nature reserve with forest and meadows. The main stream system (Tilligterbeek-Hamburgerbeek-Hollandsegraven, actually one stream) connects the Agerlerbroek area with the Ottershagen area (northern section of the OLU). Both areas are low-lying plains and the connecting stream system flows through higher terrain. (b) The boundaries of the OLU depicted on a historical map from 1900, 1 : 25,000 (source: Topografische Dienst Kadaster, Emmen 1900). The purple areas are the extent of flooding after extremely high rainfall in 1998. The delineation of the OLU follows the original outlines of wetlands on the historical map and the extent of the 1998 flooding. (Reproduced with permission from Journal of Applied Ecology (2008) Vol. 45, pp. 1496-1503.)

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