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... vitellina, Lecanora intricata, Rhizocarpon geographicum) predominate in the investigated territory (189 species or 48,7%); this is usual for all lichen floras of arctic, mountain and arid areas (Urbanavichus, 1998). They play an important role at all elevations, but show the maximum species diversity (89 species or 45,2%) in the upper forest belt (Fig. 2). A wide variety of microhabitats creates favorable conditions here for different species of all growth forms. In the alpine and nival belts crustose lichens predominate over other growth forms (Table 1). The rigorous conditions of these altitudes are evidently less favorable for foliose and fruticose lichens and vascular plants. were ...
Context 2
... general, the percentage of crustose lichens of the total lichen flora increases with altitude whereas the percentage of foliose lichens falls (Makryi, 1990), but our studies do not support this to date (Fig. 2); foliose lichens are much better known than crustose lichens in the Teberda Reserve, thus evidently many crustose taxa have yet to be reported. Further inquiry is needed to show the real picture. However, species richness of all growth forms tends to decrease with altitude. This phenomenon is probably connected with the smaller ...
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Citations
... In the 20th century, extensive research on the Caucasus flora was carried out by Barkhalov (1975Barkhalov ( , 1983, and by Vězda who worked particularly in the Caucasian reserve on the Black Sea coast and Abkhazia (Vězda 1983); these accounts also documented several species of Bacidia s. lat. More recently, Bacidia species have been reported in many lichenofloristic papers and checklists of the Caucasus covering northern (Urbanavichus & Urbanavichene 2002, 2017b, 2018Urbanavichene & Urbanavichus 2019;Urbanavichus et al. 2021), north-western (Otte 2001(Otte , 2004(Otte , 2007aBlinkova & Urbanavichus 2005;, 2017a, western (Urbanavichene & Urbanavichus 2016;Urbanavichus et al. 2020), south-western ), north-eastern (Urbanavichus & Ismailov 2013), central , eastern (Ismailov et al. 2017), and southern (Harutyunyan et al. 2011;Alverdiyeva & Novruzov 2014;Gasparyan & Sipman 2016;Inashvili et al. 2022) parts of the region. Urbanavichus (2010) was the first to compile data on lichens known for the Russian territory (incorporating the Caucasus), including 18 species of Bacidia s. lat. in the checklist currently known from the Caucasus. ...
During a study of the incompletely known lichen flora of the Caucasus, we analyzed 237 specimens of corticolous Bacidia s. str. collected in the Northern and Southern Caucasus, including Armenia, Azerbaijan, Georgia, and Russia. Of these, 54 specimens belonging to 11 species of Bacidia s. str. were selected for molecular studies, representing the observed morphological variability of the genus. We obtained 142 sequences from three RNA-coding genes (nrITS, nrLSU, and mtSSU) and two protein-coding genes ( RPB 1 and RPB 2). The single and concatenated datasets were complemented with Bacidia s. str. sequences from GenBank and subjected to Bayesian inference and two maximum likelihood analyses (RAxML and IQ-TREE). The resulting trees yielded highly concordant topologies of the groups and corresponded with previous results, supporting two main clades correlating with apothecia pigmentation. Our analyses are the first to reveal the presence of Bacidia heterochroa in the Caucasus. An exceptionally high degree of morphological plasticity was found in the Rubella and Suffusa groups. As a result of morphological examination and phylogenetic results, B. caucasica (Suffusa group) was described as new to science. Furthermore, two putative taxa in the Rubella group, Bacidia inconspicua ined. and B. maritima ined., were introduced. This study furthers our understanding and documentation of the understudied lichen flora of the Caucasus, bringing the total number of Bacidia species for the region to 13.
... Был указан для кластера «Шуби» Северо-Осетинского заповедника . Ранее был известен из Северной Осетии вне территории Северо-Осетинского заповедника (Vainio, 1899 (Otte, 2004;Blinkova, Urbanavichus, 2005;Urbanavichus, Ismailov, 2013;Urbanavichus, Urbanavichene, 2014;Khanov et al., 2019). ...
... -на известняке, 23 VI 2016, Urbanavichus 23062016-15, LE. Широко распространен на Северном Кавказе: известен из Краснодарского и Ставропольского краев, республик Адыгея, Карачаево-Черкесия, Кабардино-Балкария, Ингушетия, Чечня и Дагестан (Vainio, 1899;Barkhalov, 1983;Zakutnova, Musina, 1986;Krivorotov, 1997;Blinkova, Urbanavichus, 2005;Urbanavichus, Ismailov, 2013;Urbanavichus, Urbanavichene, 2014 (Barkhalov, 1983;Urbanavichus, Ismailov, 2013;Urbanavichus, Urbanavichene, 2014Urbanavichene, Urbanavichus, 2018). ...
... × 5.0-7.5 мкм, бесцветными вначале, позже светло-коричневыми, с периспорием 0.5-1.5 мкм толщ. (Vondrák, Kocourková, 2008 (Barkhalov, 1983;Krivorotov, 1997;Otte, 2001;Blinkova, Urbanavichus, 2005;Urbanavichus, Ismailov, 2013;Urbanavichus, Urbanavichene, 2014, 2018b. !!!Placocarpus schaereri (Fr.) ...
New data on the lichen flora of the North Ossetia Nature Reserve from the cluster “Kartsa” are provided. This cluster is located in Kartsinskii Range slopes, Central Caucasus. The specimens of lichens and lichenicolous fungi were collected near the top of Ol’dukhankhokh Mt. in cluster “Kartsa” in June 2016. In total, 115 lichen species and 15 lichenicolous fungi were recorded. Among them, 128 species are new to the cluster “Kartsa”. Of these, 77 species are new for the North Ossetia Nature Reserve, including 71 species new for the Republic of North Ossetia — Alania, 36 species new for the Central Caucasus, and 5 species new for the Caucasus. The genus Poeltinula and three species (Arthonia punctella, Opegrapha hellespontica, Poeltinula cerebrina) are reported for the first time from Russia. The representative specimens of new for Russia and rare species are kept in the Lichenological Herbarium of the Komarov Botanical Institute RAS (LE).
... The arctic-alpine species Rhizocarpon atroflavescens was earlier known in the Caucasus only from Adygeya (Otte 2007) although the species is not uncommon. Other arctic-alpine lichens include Tetramelas papillatus which is also unknown in the eastern Caucasus and had previously been recorded from high-mountainous areas of the western Caucasus (Blinkova & Urbanavichus 2005, Urbanavichus & Urbanavichene 2014. A mainly southern species with an Eurasiatic distribution is Toninia taurica which was earlier known in the Caucasus from some localities in the western and central parts (Urbanavichus & Urbanavichene 2014, Urbanavichene & Urbanavichus 2018). ...
Ismailov, A. 2021. Diversity and peculiarities of lichens in subnival habitats of Shalbuzdag Mountain (East Caucasus, Dagestan, Russia). – Herzogia 34: 387–399.
One hundred and four species of lichenized fungi and three species of lichenicolous fungi have been revealed in habitats above 3000 m a.s.l of Shalbuzdag Mountain. Some species are reported from these altitudes for the first time. Gyalolechia klementii is new for Russia and the Greater Caucasus. Heteroplacidium fusculum and Calvitimela aglaea are reported for the Greater Caucasus for the first time. Tetramelas papillatus, Toninia taurica and Rhizocarpon atroflavescens are new records for the East Caucasus. All of these species and Placidium rufescens are new for Dagestan. Habitats at an altitudinal range of 3000 –3200 m are characterized by the highest species richness. Lichens of the family Teloschistaceae are the core of the lichen flora with the highest number of recorded species (26) and genera (11). The lichen communities at mount Shalbuzdag are dominated by species preferring arid habitats, but there are also arctic-alpine species. For the first time, detailed data are provided on the lichen diversity in arid calcareous habitats above 3000 m in the Caucasus and Russia.
... Широко распространенный в Голарктике лесной вид. На Кавказе достаточно обычен в горных лесах Краснодарского края и Адыгеи, в Карачаево-Черкессии, Кабардино-Балкарии, Северной Осетии и Дагестана Blinkova, Urbanavichus, 2005;Slonov, 2014;. ...
Thirteen lichen species proposed for the Red Data Book of Republic of Ingushetia on based of the results of author's field works and analysis of literature information. Among them, three species are included in the Red Data Book of the Russian Federation (Leptogium burnetiae, Lobaria pulmonaria, Usnea florida). Ten lichen species are rare and vulnerable due to the state of populations in the Republic of Ingushetia at the current time (Hyperphyscia granulata, Melanelixia albertana, Neo-catapyrenium rhizinosum, Parmotrema stuppeum, Peltula bolanderi, Phaeophyscia cernohorskyi, Phaeophyscia insignis, Punctelia borreri, Thalloidima toninianum, Usnea cavernosa). Information on distribution in the republic, ecology and proposed categories of the threatened are given. Most of the species (10) were proposed to be included with «Near Threatened» category, 1 species (Leptogium burnetiae) with «Critically Endargered», 1 species (Lobaria pulmonaria) with «Data Deficient» and 1 species (Usnea florida) with «Least Concern».
... Although lists of lichens collected in NW Caucasus have been already published by Krivorotov (1997), Urbanavichus & Urbanavichene (2004, Blinkova & Urbanavichus (2005), and Otte (2005Otte ( , 2007, fir-beech virgin forests in the Caucasus Reserve are lichenologically still poorly known. Krivorotov (1997) published a lichen checklist of the NW Caucasus including about 500 species, of them about 200 species were reported from the territory of the Caucasus Reserve. ...
We report on 659 epiphytic and epixylic species recorded from seven one-hectare plots established along an altitudinal
gradient in a virgin forest of the Caucasus State Nature Biosphere Reserve. A total of 564 species are lichens, 61 are lichenicolous
fungi and 34 are allied non- or facultatively lichenized fungi. one hundred forty – nine species (116 lichens,
17 lichenicolous and 16 saprophytic fungi) are new to the Northern Caucasus, including 133 species (104 lichens, 15
lichenicolous and 14 saprophytic fungi) that are new to the Caucasus Mountains. Fifty species are reported from Russia for
the first time: 37 lichens (Andreiomyces obtusaticus, Bacidina mendax, Biatora aegrefaciens, B. bacidioides, B. chrysanthoides,
Biatorella dryophila, Buellia iberica, Cliostomum haematommatis, Endohyalina ericina, Fellhanera christiansenii,
Gyalidea minuta, Japewia aliphatica, Lecanora barkmaniana, L. subravida, Lecidea strasseri, Leptogium hibernicum,
Lithothelium hyalosporum, L. phaeosporum, L. septemseptatum, Loxospora cristinae, Melanelixia epilosa, Micarea
nowakii, M. perparvula, Opegrapha trochodes, Orcularia insperata, Parvoplaca servitiana, Phylloblastia inexpectata,
Psoroglaena stigonemoides, Ptychographa xylographoides, Ramonia dictyospora, R. luteola, Rinodina polysporoides,
Thelopsis flaveola, Topelia jasonhurii, Verrucaria hegetschweileri, Wadeana minuta, Waynea giraltiae), nine lichenicolous
fungi (Arthonia vorsoeensis, Didymocyrtis melanelixiae, Epigloea urosperma, Muellerella polyspora, Phacographa
zwackhii, Pronectria pilosa, Rhymbocarpus pubescens, Taeniolella friesii, Unguiculariopsis acrocordiae) and four nonlichenized
saprophytic fungi (Cyrtidula major, Karschia cezannei, Kirschsteiniothelia recessa, Pseudotryblidium neesii).
The ratio of macrolichens ranges between 26.5 – 40 % and rises with elevation. Lichens with a trentepohlioid photobiont
are represented by 15 –51 species per plot and their species richness decreases with elevation. The species richness
of cyanolichens is substantial in all plots (15 –28 species) reflecting a negligible effect of acidification/air pollution.
Low species richness and low abundances of nitrophilous species indicate insignificant uptake of nitrogen emissions.
Beech and fir are the most preferred phorophytes, but the vast majority of lichen species have low substrate specificity.
Species richness per plots ranged between 236 and 379. The highest richness was found in a plot outside the Caucasian
Reserve and we recommend its inclusion into the protected area.
... At present, about 2040 species (including allied non-lichenized fungi) are recorded in the North Caucasian lichen flora (Urbanavichus 2018). The knowledge of the lichen flora of the Northern Caucasus has been improved strongly during the past two decades due to surveys of the Teberda State Biosphere Reserve by Blinkova & Urbanavichus (2005), the Bol'šoj Thač Nature Park (NW Caucasus) by Otte (2007b, c), the Gunib Plateau (NE Caucasus) by Urbanavichus and Ismailov (2013), the Lagonaki Highland and Abrau Peninsula (NW Caucasus) by Urbanavichus and Urbanavichene (2014, and the Dagestan Republic (NE Caucasus) by Ismailov (2017) and Ismailov et al. (2017a); Zhurbenko (2017) published a survey of the North Caucasian lichenicolous fungi. Moreover, c. 40 papers relating to field and taxonomic studies were published (e.g., Eskin et al. 2004, Otte 2005, 2007a, d, Sohrabi et al. 2007, Otte et al. 2013, Ismailov et al. 2017b, Vondrák et al. 2017. ...
... Notes: In the Northern Caucasus previously known only from the Republic of Karachayevo-Cherkessia (Blinkova & Urbanavichus 2005). It is also known in Transcaucasia from Armenia, Azerbaijan and Georgia (Barkhalov 1983 Notes: In the Northern Caucasus so far known only from the Republic of Adygea (Urbanavichus & Urbanavichene 2014). ...
... Notes: The species was earlier known from scattered occurrences in the Caucasian region. In the Northern Caucasus previously reported from the North-Western Caucasus -Republic of Karachayevo-Cherkessia (Blinkova & Urbanavichus 2005) and from the North-Eastern Caucasus -Republics of Ingushetia ) and Dagestan (Urbanavichus & Ismailov 2013). It is also known in Transcaucasia from Azerbaijan and Georgia (Barkhalov 1983). ...
Noteworthy records of 24 species of lichens and lichenicolous fungi from the Central Caucasus are listed; eight of them (Buellia tyrolensis, Catillaria atomarioides, Endococcus matzeri, E. verrucosus, Endohyalina insularis, Lichenochora physciicola, Peltula obscurans and Polycoccum evae) are reported for the first time for the Caucasus, and four of these (Buellia tyrolensis, Endohyalina insularis, Lichenochora physciicola and Polycoccum evae) are new to Russia. Short notes on characteristics, ecology and distribution of the species are provided.
... Спустя почти сто лет, к концу XX в., для лихенофлоры всего Кавказа было известно порядка 1100 видов, из которых для российской части было известно около 700 видов (Barkhalov, 1983). Наиболее интенсивные исследования лихенофлоры Северного Кавказа начались в XXI в. (например, Slonov, 2002(например, Slonov, , 2014, 2014, 2018aBlinkova, Urbanavichus, 2005;Otte, 2005Otte, , 2007aUrbanavichus, Ismailov, 2013;Ismailov, 2017;Vondrák et al., 2017;. В итоге к настоящему времени для лихенофлоры Северного Кавказа удалось выявить порядка 2040 видов лишайников и систематически близких грибов (Urbanavichus, 2018). ...
... Америке, Австралии и Океании (Galloway, 2007;Davydov, 2017). На Северном Кавказе известен из Адыгеи (Eskin et al., 2004), Карачаево-Черкесии (Blinkova, Urbanavichus, 2005) и Дагестана (Ismailov, 2018). В России, кроме Кавказа, известен на Урале и Алтае (Davydov, 2017 Umbilicaria vellea (L.) Hoffm. ...
An annotated list of 15 species of Umbilicaria Hoffm. collected in Elbrus region (Central Caucasus, Kabardino-Balcaria) is presented. The description and a localities of a new to Russia and Bulgaria lichen species Umbilicaria freyi Codogno et al. are reported. The species is characterized by developing of shizidia as vegetative propagules. Umbilicaria aprina Nyl., U. freyi, U. lyngei Schol. and U. maculata Krzewicka et al. are reported for the Caucasus Mountains for the first time, U. altaiensis Wei et Jiang, U. cinerascens (Arnold) Frey и U. subglabra (Nyl.) Harm. are new for the Central Caucasus. Outside Caucasus U. altaiensis Wei et Jiang is new for Austria, U. freyi – new for Bulgaria, and U. maculata – new for Altai Mts.
... Новый род для лихенофлоры Кабардино-Балкарии. Кальцефильный лишайник, часто встречается на Северном Кавказе: Краснодарский край, Адыгея , Карачаево-Черкессия (Blinkova, Urbanavichus, 2005), Дагестан . Известен в Закавказье из Азербайджана, Грузии (Barkhalov, 1983;Novruzov, Alverdieva, 2014). ...
... et Wedin -Поляна Азау, на силикатных скалах, обогащенных солями кальция. Кальцефильный лишайник, широко распространен на Северном Кавказе: Краснодарский край, Адыгея , Карачаево-Черкессия (Blinkova, Urbanavichus, 2005), Ставропольский край (Urbanavichene, Urbanavichus, 2018), Ингушетия , Дагестан . Известен в Закавказье из Абхазии, Армении и Грузии (Barkhalov, 1983;. ...
... et Wedin -ущелье Сукансу, на известняках. Кальцефильный вид, широко распространенный на Северном Кавказе: Краснодарский край, Адыгея , Карачаево-Черкессия (Blinkova, Urbanavichus, 2005), Ингушетия , Дагестан . Известен в Закавказье из Абхазии, Армении, Грузии (Barkhalov, 1983;. ...
Based on the material collected in 2018, twenty species and nine genera (Bilimbia, Lathagrium, Lempholemma, Muellerella, Myriolecis, Placynthium, Pyrenodesmia, Thelidium, Variospora, Verrucaria) are reported as new to the lichen flora of the Kabardino-Balkarian Republic. Data on ecology and distribution of these species in the Caucasus are provided. Physcia albinea is reported for the first time in the North Caucasus, and Bilimbia lobulata, Lathagrium undulatum, Lempholemma polyanthes are new for the Central Caucasus. The specimens are kept in the lichenological herbarium LE (St. Petersburg).
Приведены сведения о 20 видах и 9 родах (Bilimbia, Lathagrium, Lempholemma, Muellerella, Myriolecis, Placynthium, Pyrenodesmia, Thelidium, Variospora, Verrucaria), новых для лихенофлоры Кабардино-Балкарской Республики. Physcia albinea впервые отмечена для Северного Кавказа; виды Bilimbia lobulata, Lathagrium undulatum и Lempholemma polyanthes – для Центрального Кавказа.
... G. Urbanavichus). -Мо-жет оказаться достаточно обычным на Кавказе видом, но найден пока только в Краснодарском крае, Карачаево-Черкессии и Дагестане (Blinkova, Urbanavichus, 2005;Urbanavichus, Ismailov, 2013;). В За-кавказье известен из Абхазии и Грузии (Vainio, 1899;Barkhalov, 1983;Urbanavichus, Urbanavi- chene, 2012). ...
... -По нашей оценке, этот вид широко распространен на Кавказе, но до сих пор не был известен в Кабардино-Балкарии. На Северном Кавказе встречается в Краснодарском крае, Адыгее, Карачаево-Черкесии и Дагестане (Otte, 2001(Otte, , 2005Blinkova, Urbanavichus, 2005;Urbanavichus, Ismailov, 2013;Urbanavichus, Ur- banavichene, 2014). В Закавказье известен из Азербайджана и Армении Novruzov, Alverdieva, 2014). ...
... Тем не менее, он нечасто указывается для Северного Кавказа. Известен из Карачево-Чер-кессии, Северной Осетии, Ингушетии, Дагеста-на (Vainio, 1899; Blinkova, Urbanavichus, 2005;Ismailov, 2017;Urbanavichus, Urbanavichene, 2017). В Закавказье известен из Азербайджана, Армении и Грузии ( Barkhalov, 1983). ...
Based on the results of short field works in July 2018, data on new and noteworthy species for the lichen flora of the Central Caucasus are presented. The specimens were collected in the Baksan River valley, Elbrus district, Republic of Kabardino-Balkaria, mainly in the vicinity of the village Bylym in the Bylym intermountain arid basin of the Severo-Yurskaya (North-Jurassic) depression between the Bokovoy (Lateral) and Skalistyi (Rocky) Ridges, and in the vicinities of Elbrus and Azau settlements near the foot of the south-eastern slope of Mount Elbrus. In the present paper, 22 species are reported as new for the lichen flora of the study area. Candelariella blastidiata is reported for the first time for Caucasus, Agonimia opuntiella, Gyalolechia lenae, Lecidella laureri and Physconia perisidiosa are new to Central Caucasus. Another 14 species (Anaptychia bryorum, Caloplaca percrocata, Endocarpon adsurgens, Hyperphyscia adglutinata, Lecania suavis, Lobothallia alphoplaca, Muellerella pygmaea, Peltula euploca, Pertusaria flavicans, Phaeophyscia orbicularis, P. sciastra, Punctelia borreri, Thallinocarpon nigritellum, Xanthomendoza fallax) are newly reported to Kabardino-Balkaria. The genera Hyperphyscia, Lecania, Muellerella, Peltula and Thallinocarpon are new to the lichen flora of Kabardino-Balkaria. Information about localities, ecology and Caucasian distribution of all mentioned species is provided.
... ì., íà òàëëîìå Candelariella sp., 12 VIII 2010, Õàíîâ. -Íà Ñåâåðíîì Êàâêàçå èçâåñòåí ïî åäèíè÷íûì íàõîäêàì â Êðàñíîäàðñêîì è Ñòàâðîïîëüñêîì êðàÿõ, Ðåñïóáëèêàõ Àäûãåÿ è Êàðà÷àåâî-×åðêåññèÿ (Blinkova, Urbanavichus, 2005;Urbanavichene, Urbanavichus, 2018). ...
... -Íîâûé äëÿ Öåíòðàëüíîãî Êàâêàçà. Íà Ñåâåðíîì Êàâêàçå èçâåñòåí â Êðàñíîäàðñêîì êðàå, Ðåñïóáëèêàõ Àäûãåÿ, Êàðà÷àåâî-×åðêåññèÿ è Äàãåñòàí (Barkhalov, 1983;Blinkova, Urbanavichus, 2005;. ...
... -Íîâûé äëÿ Öåíòðàëüíîãî Êàâêàçà. Íà Ñåâåðíîì Êàâêàçå áûë èçâåñòåí òîëüêî â Êðàñíîäàðñêîì êðàå, Ðåñïóáëèêàõ Êàðà÷àåâî-×åðêåññèÿ è Äàãåñòàí (Krivorotov, 1997;Blinkova, Urbanavichus, 2005;Ismailov, 2015). ...
The Kabardino-Balkarian Republic is located in the central part of the northern macro-slope of the Caucasus. The Republic differs sharply from the rest of the North Caucasus in terms of climatic indicators, geomorphological and lithological features. Only 312 species were previously known in the Republic according to the literature and data of our local floristic study. This paper presents data on 20 species and 5 genera (Alyxoria, Athallia, Carbonea, Physciella, Pleopsidium) new to the Kabardino-Balkarian Republic. The information on their distribution in the Caucasus with data on the localities is given. Five species are recorded for the first time in the Central Caucasus: Brodoa atrofusca, Cetraria muricata, Cladonia cornuta, Lepra schaereri, Ramalina subgeniculata. For these species the characteristic features of specimens are given, and the nearest locality is cited.
