FIGURE 53 - uploaded by Barna Páll-Gergely
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Protoconchs of Metalycaeus species. A-B. Metalycaeus (?) okuboi Páll-Gergely & Hunyadi, n. sp. (holotype, HNHM 99713). C-D. Metalycaeus heudei (Bavay & Dautzenberg, 1900), 20030318B. All images: B. Páll-Gergely.
Source publication
This is the first part of the revision of the Alycaeidae of China, Laos and Vietnam. Here we revise the species hitherto classified in Chamalycaeus Möllendorff, 1897 (genera Dicharax and Metalycaeus in this study). We show that no Chamalycaeus live in the region, and the alycaeid species of the investigated region can be classified into five genera...
Citations
... Sarasin et F. Sarasin, 1899) from Sulawesi and S. porcilliferus (Bollinger, 1918) from eastern Borneo have elevated, trumpet-like projections, but in those two species the edge of the trumpet is thickened and reflected, and they are seemingly not made of a single rolled lamina, while the edge of the trumpet is sharp and not reflected in the new species, and it is made of a single rolled lamina. Furthermore, Dicharax bison Páll-Gergely et Hunyadi, 2017(described in Páll-Gergely et al. 2017) also has an elevated funnel-shaped projection on the outer side of the operculum, but it is clearly made of the multispiral outer lamina. It seems that the morphologically somewhat similar trumpet-or funnel-shaped projections on the outer side of the operculum of various alycaeid species are not homologous with each other, and probably developed multiple times during the evolution of this family. ...
Cycloryx pemaledai Gittenberger et Sherub, 2022, which was described from northwestern Bhutan, is reported for the first time in Sikkim, India. That species differs from all its congeners by the short R3 (the region between the constriction and the peristome). The differences between the Bhutanese and Indian shells are minimal, therefore the Indian shells are identified as Cycloryx cf. pemaledai. Furthermore, Alycaeus himalayae sp. n. is described from Arunachal Pradesh. This is so far the only Alycaeus species inhabiting the Himalayas, and thus, its discovery is a surprise.
... In the type series of Anaglyphula cancellata, although the constriction of two specimens is too deep to be observed from the aperture, the constriction of the other specimens is visible through the aperture when observed at an angle. In contrast, the constriction in other cyclophoroidean groups (i.e., Alycaeidae and Diplommatinidae) is mostly clearly visible from the outside, since it is a reduction of the diameter of the whorl where the constriction is situated (see e.g., Páll-Gergely et al., 2017;Chen & Wu, 2020;Páll-Gergely, 2020). In the new species, the presence of a constriction is somewhat uncertain due to the difficulty in discerning whether a lighter-coloured line approximately halfway behind the aperture represents the constriction or merely a growth line, as this position is too deep to be visually examined from the aperture. ...
Anaglyphula yanseni, new species, is described and illustrated based on specimens collected from Babi Island, off the west coast of Sumatra, Indonesia. The new species mostly differs from its congeners having similarly pitted surface sculpture by its small size. Additionally, Anaglyphula cancellata Rensch, 1932, the type species of the genus, is redescribed based on the type materials. The diagnostic features of Anaglyphula are also discussed in comparison with those of similar genera.
... Nearly a century has passed with only one paper about Lao land snails (Saurin, 1953). Fortunately, increasing interest in Southeast Asian land snail taxonomy in the last ca. 2 decades resulted in numerous publications (Maassen, 2008;Inkhavilay et al., 2016aInkhavilay et al., , 2016bPáll-Gergely et al., 2015, 2016, 2017, 2020a, 2020b2022, in press), suggesting that this country is inhabited by a much higher number of endemic species than previously thought. Many species described in the last years belong to the family Hypselostomatidae, which are tiny (0.6-5 mm) snails, typically inhabiting limestone areas. ...
Anauchen (?) kozari n. sp. is described from Luang Prabang province, central Laos. This new species is temporarily placed in the genus Anauchen due to the similarity of its apertural barriers to those of A. messageri (Bavay & Dautzenberg, 1904). Boysidia gereti Bavay & Dautzenberg, 1904 is a junior synonym of Anauchen rochebruni (Mabille, 1887) (originally described as Hypselostoma ).
... Tarruella and Domènech (2011) recognized four genera within Alycaeinae, namely Alycaeus J.E. Gray, 1850, Chamalycaeus Möllendorff, 1897, Cipangocharax Shintaro, 1934and Dioryx Benson, 1859. However, Cycloryx Godwin-Austen, 1914, Dicharax Kobelt & Möllendorff, 1900and Metalycaeus Pilsbry, 1900 have recently been raised to generic rank (Páll-Gergely et al., 2017a). Páll-Gergely et al. (2016) described the detailed structure and function of the external sutural tube that lies behind the aperture of all Asian members of the Alycaeidae; this character, in combination with the unique position of the bursa copulatrix, appears to justify family-level status (Páll-Gergely et al., 2020a). ...
... The most specious genus is Dicharax Kobelt and Möllendorff, 1900, which currently contains 158 species (MolluscaBase eds., 2022). Dicharax is characterised by the lack of spiral striation on both the protoconch and the teleoconch, although very few species with spiral striation is also included in this genus (see Páll-Gergely et al., 2017. One of the hotspots of Dicharax is found in the southeastern Himalaya, and this remarkable diversity continues in Myanmar and northern Thailand . ...
In this paper, three new species of the alycaeid genus Dicharax Kobelt & Möllendorff, 1900 are described, namely Dicharax floridus n. sp., Dicharax spatiosus n. sp. (Both from Shan State, Myanmar) and Dicharax kosztarabi n. sp. (from Thailand). All three new species are known from their holotypes.
http://zoobank.org/28168F06-CFBF-405F-90F7-A3399060D88C
... We follow Páll-Gergely et al. (2017) regarding the terminology of the three regions of the teleoconch of Metalycaeus Pilsbry, 1900: Region 1 (R1), ranging from the beginning of the teleoconch to the beginning of the differently ribbed region where the sutural tube lies; Region 2 (R2), extending from the end of R1 to the constriction (i.e., the length of R2 usually corresponds with the length of the sutural tube, see Páll-Gergely et al., 2016); and Region 3 (R3), ranging from the constriction up to the peristome. ...
... Measurements (in mm): NIGP180011: H = 2.24, D = 3.02, h = 1.33, d = 1.49. Remarks: Metalycaeus fossilis n. sp. has a spirally striated protoconch as the key diagnostic character of the genus (Páll-Gergely et al., 2017. Among the Metalycaeus species inhabiting mainland China and Taiwan (Hsieh et al., 2006, Páll-Gergely et al., 2017, only M. latecostatus (Mö llendorff, 1882) possesses similarly strong, widelyspaced ribs, although its body whorl is slightly angled, and has denser R1 ribs (ca. 25 ribs/last half whorl of R1 versus ca. ...
... Remarks: Metalycaeus fossilis n. sp. has a spirally striated protoconch as the key diagnostic character of the genus (Páll-Gergely et al., 2017. Among the Metalycaeus species inhabiting mainland China and Taiwan (Hsieh et al., 2006, Páll-Gergely et al., 2017, only M. latecostatus (Mö llendorff, 1882) possesses similarly strong, widelyspaced ribs, although its body whorl is slightly angled, and has denser R1 ribs (ca. 25 ribs/last half whorl of R1 versus ca. 18 in the new species). ...
Four gastropod species including two new species (?Tinnyea sp., Cyclophoridae gen. et sp. indet., Metalycaeus fossilis n. sp., and Tortaxis zhangpuensis n. sp.) are described from the mid-Miocene Zhangpu amber. Among them, ?Tinnyea sp. is potentially the first fossil record of this group in lake systems of East Asia, outside of Mediterranean and Paratethys areas. Tortaxis zhangpuensis n. sp. and Metalycaeus fossilis n. sp. are likely the oldest representatives of respective genera. Despite Tortaxis zhangpuensis n. sp. being a juvenile shell, its whorls are taller and more convex in comparison to congeners, and from around 3.75th whorl onwards the sub-sutural area of whorls displays a crenulated pattern. Metalycaeus fossilis n. sp. has a spirally striated protoconch (the key diagnostic character of the genus), and with strong, widely spaced R1 ribs, short R2, which distinguishes it from its congeners.
... They occur in Hungerfordia Beddome, 1889, Helicomorpha Möllendorff, 1890 and newly published Kontschania Páll-Gergely & Grego, 2020 and Moderata Páll-Gergely, 2020. However, it is frequently found in Alycaeidae (Páll-Gergely et al., 2017. Laotia Saurin, 1953 from Vietnam and Laos is a genus originally thought to belong to Diplommatininae (= Diplommatinidae) but was transferred to Alycaeidae in recent studies (Páll-Gergely, 2014;Do et al., 2015;Páll-Gergely et al., 2017;Páll-Gergely & Hunyadi, 2021). ...
... However, it is frequently found in Alycaeidae (Páll-Gergely et al., 2017. Laotia Saurin, 1953 from Vietnam and Laos is a genus originally thought to belong to Diplommatininae (= Diplommatinidae) but was transferred to Alycaeidae in recent studies (Páll-Gergely, 2014;Do et al., 2015;Páll-Gergely et al., 2017;Páll-Gergely & Hunyadi, 2021). Saurin (1953) noted that Laotia and Messageria are relatives due to the oblique embryonic shell, developed ribs and multispiral lamina on outer surface of operculum. ...
... Therefore, we transfer Messageria to Alycaeidae just like Laotia. It is worthwhile to mention that the three regions of the teleoconch (see definition in Páll-Gergely et al., 2017) commonly found in Alycaeidae, cannot be distinguished in these two genera. Therefore, the present arrangement may also be temporary. ...
In this study, the systematic status of Messageria Bavay & Dautzenberg, 1904 is discussed. It is recognized as an alycaeid genus, rather than as a subgenus or a junior synonym of Helicomorpha Möllendorff, 1890 (Diplommatinidae) as previously thought. Additionally, a new species, Messageria sinica n. sp. from Guizhou, and a new subspecies, Messageria scalarioides donghiana n. ssp. from Guangxi are described. Messageria scalarioides donghiana n. ssp. is morphologically different from the nominate subspecies by the larger shell and wider lower whorls and umbilicus. Messageria sinica n. sp. is morphologically different from the type species by having larger shell and distinct intermediate ribs.
... Đến năm 1999, Kuzminykh phát hiện giống mới Laocaia từ vùng núi cao Hoàng Liên Sơn [4]. Trong giai đoạn gần đây, một vài nghiên cứu được công bố nhưng đã có nhiều giống và loài mới được phát hiện [5], [6]. Những kết quả trên đây cho thấy tiềm năng về đa dạng Thân mềm ở cạn tại Lào Cai là rất lớn, trong khi hiểu biết của chúng ta còn hạn chế. ...
... Đất mùn và thảm mục cũng được thu và đưa về phòng thí nghiệm, sau đó tách mẫu nhờ sàng (rây) với đường kính mắt lưới 5, 3, 1 mm. Định loại mẫu vật theo tài liệu chuyên ngành, gồm những mô tả gốc, tu chỉnh, chuyên khảo của Bavay và Dautzenberg (1909) [2], Páll-Gergely et al., (2015,2017) [5], [6], Nordsieck (2011) [7], Schileyko (2000Schileyko ( , 2011 ...
Bài báo này giới thiệu một danh sách loài cập nhật về Thân mềm ở cạn tại tỉnh Lào Cai, nơi được đánh giá như một khu vực có hệ thống núi đá vôi quan trọng hàng đầu của Việt Nam về đa dạnh sinh học Thân mềm ở cạn. Nghiên cứu này cung cấp dẫn liệu tổng quan về đa dạng và phân bố của khu hệ Thân mềm ở cạn xuất hiện tại ba sinh cảnh chủ yếu (rừng trên núi đá vôi, rừng trên núi đất, đất canh và khu dân cư). Tổng cộng có 125 loài, thuộc 64 giống, 23 họ được phát hiện. Các họ Camaenidae (26 loài, 10 giống), Clausiliidae (17 loài, 8 giống), Streptaxidae (9 loài, 4 giống), Ariophantidae (8 loài, 5 giống), Achatinidae (8 loài, 5 giống), Cyclophoridae (7 loài, 5 giống), Pupinidae (7 loài, 4 giống), Helicarionidae (7 loài, 4 giống), Alycaeidae (5 loài, 3 giống), Diplommatinidae (6 loài, 2 giống), Enidae (5 loài, 1 giống) có số loài cao nhất, trong khi 12 họ còn lại chỉ ghi nhận 1 hoặc 2 loài. Có 18 loài (chiếm 14,4% tổng số loài) là đặc hữu cho tỉnh Lào Cai. Có 4 giống (Deroceras, Meghimatium, Succinea, Valiguna) và 15 loài được ghi nhận mới cho khu vực nghiên cứu.
... Đếm số vòng xoắn theo phương pháp mô tả của Kerney & Cameron [6], phân chia các nhóm kích thước theo Panha & Burch (2005) [7]. Định loại mẫu vật theo tài liệu chuyên ngành, gồm những mô tả gốc và tu chỉnh của Mabille [8], Bavay & Dautzenberg [9], Dautzenberg & Fischer [10,11], Páll-Gergely et al. [12], Nordsieck [13], Do & Do [5,14], Schileyko [15]. Đa dạng các taxon phân loại: Trong tổng số 17 họ được phát hiện ở các hệ sinh thái thuộc tỉnh Điện Biên, những họ đa dạng về loài gồm Camaenidae (18 loài), Cyclophoridae (16 loài), Ariophantidae (8 loài), Achatinidae (7 loài), Pupinidae, Clausiliidae và Streptaxidae (6 loài), các họ còn lại kém đa dạng hơn, ghi nhận từ 1 đến 3 loài, tính trung bình có 4,7 loài/họ (bảng 1, 2). ...
The research has been carried out in Dien Bien Province (Northwest region of Vietnam) from 2013 to 2019. This is the very first checklist of the terrestrial snails of Dien Bien Province. The checklists provide databases and helps to understand the biodiversity richness of the particular region of country. As such, they also open trends for further research priorities. Altogether 80 species and subspecies of terrestrial snails belonging to 17 families, 49 genera are listed with 5 genera (Lagocheilus, Metalycaeus, Scabrina, Laotia, and Moellendorffia) and 36 new species recorded for Dien Bien Province
... Whorls were counted as described by Kerney and Cameron (1979). The terminology to describe alycaeid shells (Regions 1-3) follows Páll-Gergely et al. (2017). ...
Two new species, Sinoarinia feii Chen, gen. et sp. nov. and Pincerna vallis Chen & Wu, sp. nov., are described from western Hubei, China. The new genus Sinoarinia Chen & Wu, gen. nov. is established based on the strongly constricted penultimate whorl and the ascending last whorl. The Vietnamese species Pincerna costulosa (Bavay & Dautzenberg) is newly reported from Yunnan, China. For comparison, photographs of the shells of closely related species are provided. A key to the known species of Sinoarinia gen. nov. is included.
