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Map of modern day Europe, with relevant geographical features marked. 

Map of modern day Europe, with relevant geographical features marked. 

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There is a period of some 5000 years or so in the prehistory of Europe when horse populations were greatly depleted and perhaps even disappeared in many places. Before this time, during the Upper Palaeolithic, wild horses were common; after, during the Bronze Age, domestic horses were being raised and used across Europe. What happened in between is...

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Context 1
... the end of the Pleistocene, during the Upper Palaeolithic ( c . 35,000–10,000 BP ), Europe had a glacial climate that would have been more continental than the temperate one of today, with long cold winters potentially falling below -10 8 C for weeks on end (Mellars 1994). At the maximum of the last glaciation ( c . 20,000–18,000 BP ), ice sheets covered much of the northern and mountainous areas of Europe (including most of Ireland, the northern half of Britain, Scandinavia and the territories of the Baltic coastline) (Fig. 2). Fringed by forest at its Mediterranean southern margins, much of Europe consisted of open steppe tundra (Mellars 1994). Cold-tolerant species, adapted to the open steppe tundra dominate the Upper Palaeolithic fauna of Europe, among which the wild horse was common (Arbogast et al. 2002; Benecke 2006a; Brugal et al. 2001; Clutton-Brock and Burleigh 1991b; Olsen 1996; Stewart and Lister 2001; Table 1). Horses were an important provider of meat and other materials to the hunter-gather populations of Europe, as seen in the butchered bones recovered from archaeological sites and also from preserved Palaeolithic art (Clutton- Brock 1992: 24–5; Guthrie 2006a; Olsen 1996; West 2006). Increasing warmth brought the Ice Age in Europe to a close. These dramatic environmental changes, marking the end of the Palaeolithic, replaced the open landscape of the steppe tundra with one dominated by trees (Mithen 1994). Along with the changes in vegetation we see a change in the post-glacial animals of Europe: the herds of horse and reindeer that had been well-adapted to the cold, open conditions were replaced by fauna more suited to the new conditions (Mithen 1994; Yalden 1999). Finds of horse remains from c . 10,000 BP become increasingly rare across Europe, and it has been debated whether environmental change at the start of the Holocene, pushing horse populations back, was solely responsible for the decrease in horse populations or whether over-hunting by humans might also have been responsible (Arbogast et al. 2002: 18; Clutton-Brock 1992: 24–5). It has also been debated whether horses became locally extinct in many areas or just greatly reduced in number, surviving only as relic populations (e.g. Bo ̈ ko ̈ nyi 1974; Burleigh et al. 1991; Clutton-Brock and Burleigh 1991a). There is increasing evidence that the extinction of large mammals at the end of the Pleistocene, of which the decrease in horse populations is part, was primarily as a ...
Context 2
... the climatic and environmental changes that occurred, rather than massive ‘blitzkrieg’- style over-hunting by humans (e.g. Grayson and Meltzer 2003; Guthrie 2006b; Stuart 1999). However, human impact on animal species may also have played a part. It is known that, for modern feral horses, disruption of the social group will adversely affect reproduction (Berger 1986) and so, also, the expanding human populations in the Early Holocene could have brought about widespread disruption to wild horse ecology (Clutton-Brock 1992: 25). The question of whether horses became extinct in western and central Europe is also becoming clearer. It is evident that horses did not die out in Europe following the last Ice Age, but local populations persisted in many regions and some degree of continuity in horse populations is assumed for many areas (e.g. Altuna 1998; Benecke 1994; Boyle 2006; Liesau 2005). These could have been in relatively small numbers and possibly as localized ‘relic’, geographically scattered, groups. Benecke’s (1994) survey of horse on European sites of Mesolithic to Middle Neolithic date indicates horses are present, although discontinuously, across much of continental Europe, as far west as the Iberian peninsula and as far north as the North European Plain and the southern shores of the Baltic Sea (Fig. 2). Generally they constitute 0–10 per cent of bones at sites, with the highest representation of horse seen on the Eurasian steppes (from the Pontic steppes east towards the Urals; Fig. 2) where it achieves 4 40 per cent at some sites (Benecke 1994). Again, it is worth reiterating that horses prefer open habitat to forest, and the steppe and forest- steppe are more natural habitats for the horse (Linklater et al. 2000; Olsen 1996). Establishing the precise biogeographical range of the Holocene wild horse is problematic due to the low number of sites analysed and the fact that the absence of a species from archaeological deposits does not necessarily prove that it was extinct (Levine 2004, 2005). Establishing a more rigorous biogeography of the horse has been improved by programmes of direct radiocarbon dating of horse bones. Although radiocarbon-dated finds from Neolithic contexts have often returned significantly earlier or later dates and were therefore intrusive (Kaagan 2000; Uerpmann 1995), verification of the stratigraphic position of some of these finds by direct dating confirms the presence of horse in several regions (Table 1). The work by Benecke (2006a), in particular, has confirmed the continued presence of horses in Germany (Table 1; Fig. 3). That adequate habitats for wild horses did actually exist is confirmed by palynological data available from the study region (Benecke 2006a). It is difficult, from available direct radiocarbon dates, to identify the continued presence of horse populations in many areas of Europe. In Britain, for example, there is a single radiocarbon date, from Cavall’s Cave (Bronk Ramsey et al. 2002), to indicate that horse populations existed between those of the early Mesolithic and those that are present in increasing numbers from around the start of the second millennium BC (Fig. 4; Bendrey 2010). Certain regions have been proposed as being devoid of wild horses in the post-glacial period, and so when horses are subsequently recorded they are interpreted as domestic introductions. There is very limited evidence for wild horses in the fossil record of ...
Context 3
... the climatic and environmental changes that occurred, rather than massive ‘blitzkrieg’- style over-hunting by humans (e.g. Grayson and Meltzer 2003; Guthrie 2006b; Stuart 1999). However, human impact on animal species may also have played a part. It is known that, for modern feral horses, disruption of the social group will adversely affect reproduction (Berger 1986) and so, also, the expanding human populations in the Early Holocene could have brought about widespread disruption to wild horse ecology (Clutton-Brock 1992: 25). The question of whether horses became extinct in western and central Europe is also becoming clearer. It is evident that horses did not die out in Europe following the last Ice Age, but local populations persisted in many regions and some degree of continuity in horse populations is assumed for many areas (e.g. Altuna 1998; Benecke 1994; Boyle 2006; Liesau 2005). These could have been in relatively small numbers and possibly as localized ‘relic’, geographically scattered, groups. Benecke’s (1994) survey of horse on European sites of Mesolithic to Middle Neolithic date indicates horses are present, although discontinuously, across much of continental Europe, as far west as the Iberian peninsula and as far north as the North European Plain and the southern shores of the Baltic Sea (Fig. 2). Generally they constitute 0–10 per cent of bones at sites, with the highest representation of horse seen on the Eurasian steppes (from the Pontic steppes east towards the Urals; Fig. 2) where it achieves 4 40 per cent at some sites (Benecke 1994). Again, it is worth reiterating that horses prefer open habitat to forest, and the steppe and forest- steppe are more natural habitats for the horse (Linklater et al. 2000; Olsen 1996). Establishing the precise biogeographical range of the Holocene wild horse is problematic due to the low number of sites analysed and the fact that the absence of a species from archaeological deposits does not necessarily prove that it was extinct (Levine 2004, 2005). Establishing a more rigorous biogeography of the horse has been improved by programmes of direct radiocarbon dating of horse bones. Although radiocarbon-dated finds from Neolithic contexts have often returned significantly earlier or later dates and were therefore intrusive (Kaagan 2000; Uerpmann 1995), verification of the stratigraphic position of some of these finds by direct dating confirms the presence of horse in several regions (Table 1). The work by Benecke (2006a), in particular, has confirmed the continued presence of horses in Germany (Table 1; Fig. 3). That adequate habitats for wild horses did actually exist is confirmed by palynological data available from the study region (Benecke 2006a). It is difficult, from available direct radiocarbon dates, to identify the continued presence of horse populations in many areas of Europe. In Britain, for example, there is a single radiocarbon date, from Cavall’s Cave (Bronk Ramsey et al. 2002), to indicate that horse populations existed between those of the early Mesolithic and those that are present in increasing numbers from around the start of the second millennium BC (Fig. 4; Bendrey 2010). Certain regions have been proposed as being devoid of wild horses in the post-glacial period, and so when horses are subsequently recorded they are interpreted as domestic introductions. There is very limited evidence for wild horses in the fossil record of ...
Context 4
... and innovative multi-disciplinary approaches, which are tied into a robust and precise chronology. It will be only through such integrated approaches that we will move closer to an understanding of the complex history of the horse in Europe. I am very grateful to the many people who have kindly shared information and reports with me. In particular thanks must go to Emilie Blaise, Stuart Needham, Dale Serjeantson and Louise H van Wijngaarden-Bakker. I would also like to thank Amy Richardson for assistance with OxCal, Tony Bendrey for Figure 1, Rebecca Oakes for Figure 2 and the editors for the invitation to contribute to this ...
Context 5
... Fig. 2), for example, with a sole radiocarbon date predating the last glacial maximum (Table 1; Woodman et al. 1997). After this date wild horses are thought to have become locally extinct due to glacial cover (McCormick 2007a, 2007b). There had been a very good biogeographical case for a Beaker period introduction of domestic horse into Ireland, with the earliest post-Pleistocene horse bones being found in association with Beaker pottery at the passage tomb of Newgrange (in contexts dating to around 2400 BC by radiocarbon dates on charcoal) (van Wijngaarden-Bakker 1974). However, recent radiocarbon dating of two of these bones has placed them in the first few centuries AD and the dating of the horse remains from this site are in need of reassessment (Bendrey et al. in prep.). At the opposite end of Europe, to the south east, Benecke (2006b) suggests that the Balkan Mountains probably marked a natural boundary for the post-glacial range of wild horses (Fig. 2), with horse being found in small numbers at some Neolithic sites in the lower Danube area north of the mountains, but not to the south. The first recorded appearance of horses in the southern Balkan Peninsula (south of the Balkan Mountains) is at 2600–2300 cal. BC at Kirklareli-Kanligec ̧ it in Turkish Thrace (Benecke 2006b; Canteul et al. 2010; Greenfield 2006). Benecke (2006b) interprets this as indicating a later third millennium BC spread of domestic horses into this area (from Anatolia rather than from southeast/eastern Europe based on bone measurement evidence; see discussion of this method below). In addition to the targeted radiocarbon-dating programmes, synthetic surveys of archaeological bone assemblages attest to the widespread presence of horses in the Neolithic of Europe – found in many regions, although generally represented by relatively few remains when present (e.g. Arbogast et al. 2001; Boyle 2006; Grigson 1966; Kysely ́ 2005; Liesau 2005). Synthetic study of Early Neolithic Linearbandkeramik (LBK) sites ( c . 5500–4900 BC ) (Arbogast et al. 2001) indicates the presence of horse in many assemblages across Europe. Indeed, the opening up of the forests of northern Europe by the pioneer LBK farmers (Ebersbach and Schade 2004; Kalis et al. 2003) may have, in fact, subsequently benefited horse populations (discussed below). Table 2 presents data on the presence or absence of horse within 133 LBK assemblages from across Europe. The number of different species, or taxa, recovered from an archaeological assemblage is in part related to the size of the assemblage, and it is the case that rarer animals are likely to appear in larger samples and not in smaller ones, whereas commoner animals will appear in relatively small samples (Bendrey 2007a: 13–60; Grayson 1984; Lyman, 1995). For this reason, the assemblages have been grouped by assemblage size (number of identified specimens): 0–99 specimens, 100–99 specimens and so on (Table 2). This allows the presence/absence of horse in each area to be considered in relation to the sizes of the excavated assemblages. Even though the horse can be seen to be present in many regions, this is generally at a very low level within an individual assemblage – for example at the site of Cuiry-le ` s-Chaudardes in the Paris Basin, northern France, there were only four horse bones out of over 15,000 identified specimens (Hachem 1995). In the following few millennia, horse bones continue to be found at sites across Europe, and also continue to be generally rare at individual sites. For example, a study of zooarchaeological assemblages from Middle to Later Neolithic western Europe ( c . 4900– 3800 BC ) shows the horse to be present in low numbers at sites across this time period (on average contributing 0.2 per cent of bones) (Boyle 2006). However, it is not rare at all sites, with the highest value for western Europe at Roucador, France (Lesur et al. 2001). This generally low presence of horses has created problems for those trying to identify which bones come from wild animals and which from domestic. It is problematic, for example, to undertake demographic and metrical analyses on populations represented by so few remains. This is further confused by uncertainty over which criteria are valid for identifying domestic horses. Different forms of evidence have been used to identify the earliest domestic horses: some direct (where there is unambiguous textual or artistic evidence of domestic status); some indirect (where the evidence is based upon inference or interpretation, and is thus open to other interpretations) (Levine 1999, 2004, 2005). There is considerable uncertainty over the temporal and geographical origins of domestic horses in Europe, as the majority ...
Context 6
... Fig. 2), for example, with a sole radiocarbon date predating the last glacial maximum (Table 1; Woodman et al. 1997). After this date wild horses are thought to have become locally extinct due to glacial cover (McCormick 2007a, 2007b). There had been a very good biogeographical case for a Beaker period introduction of domestic horse into Ireland, with the earliest post-Pleistocene horse bones being found in association with Beaker pottery at the passage tomb of Newgrange (in contexts dating to around 2400 BC by radiocarbon dates on charcoal) (van Wijngaarden-Bakker 1974). However, recent radiocarbon dating of two of these bones has placed them in the first few centuries AD and the dating of the horse remains from this site are in need of reassessment (Bendrey et al. in prep.). At the opposite end of Europe, to the south east, Benecke (2006b) suggests that the Balkan Mountains probably marked a natural boundary for the post-glacial range of wild horses (Fig. 2), with horse being found in small numbers at some Neolithic sites in the lower Danube area north of the mountains, but not to the south. The first recorded appearance of horses in the southern Balkan Peninsula (south of the Balkan Mountains) is at 2600–2300 cal. BC at Kirklareli-Kanligec ̧ it in Turkish Thrace (Benecke 2006b; Canteul et al. 2010; Greenfield 2006). Benecke (2006b) interprets this as indicating a later third millennium BC spread of domestic horses into this area (from Anatolia rather than from southeast/eastern Europe based on bone measurement evidence; see discussion of this method below). In addition to the targeted radiocarbon-dating programmes, synthetic surveys of archaeological bone assemblages attest to the widespread presence of horses in the Neolithic of Europe – found in many regions, although generally represented by relatively few remains when present (e.g. Arbogast et al. 2001; Boyle 2006; Grigson 1966; Kysely ́ 2005; Liesau 2005). Synthetic study of Early Neolithic Linearbandkeramik (LBK) sites ( c . 5500–4900 BC ) (Arbogast et al. 2001) indicates the presence of horse in many assemblages across Europe. Indeed, the opening up of the forests of northern Europe by the pioneer LBK farmers (Ebersbach and Schade 2004; Kalis et al. 2003) may have, in fact, subsequently benefited horse populations (discussed below). Table 2 presents data on the presence or absence of horse within 133 LBK assemblages from across Europe. The number of different species, or taxa, recovered from an archaeological assemblage is in part related to the size of the assemblage, and it is the case that rarer animals are likely to appear in larger samples and not in smaller ones, whereas commoner animals will appear in relatively small samples (Bendrey 2007a: 13–60; Grayson 1984; Lyman, 1995). For this reason, the assemblages have been grouped by assemblage size (number of identified specimens): 0–99 specimens, 100–99 specimens and so on (Table 2). This allows the presence/absence of horse in each area to be considered in relation to the sizes of the excavated assemblages. Even though the horse can be seen to be present in many regions, this is generally at a very low level within an individual assemblage – for example at the site of Cuiry-le ` s-Chaudardes in the Paris Basin, northern France, there were only four horse bones out of over 15,000 identified specimens (Hachem 1995). In the following few millennia, horse bones continue to be found at sites across Europe, and also continue to be generally rare at individual sites. For example, a study of zooarchaeological assemblages from Middle to Later Neolithic western Europe ( c . 4900– 3800 BC ) shows the horse to be present in low numbers at sites across this time period (on average contributing 0.2 per cent of bones) (Boyle 2006). However, it is not rare at all sites, with the highest value for western Europe at Roucador, France (Lesur et al. 2001). This generally low presence of horses has created problems for those trying to identify which bones come from wild animals and which from domestic. It is problematic, for example, to undertake demographic and metrical analyses on populations represented by so few remains. This is further confused by uncertainty over which criteria are valid for identifying domestic horses. Different forms of evidence have been used to identify the earliest domestic horses: some direct (where there is unambiguous textual or artistic evidence of domestic status); some indirect (where the evidence is based upon inference or interpretation, and is thus open to other interpretations) (Levine 1999, 2004, 2005). There is considerable uncertainty over the temporal and geographical origins of domestic horses in Europe, as the majority ...
Context 7
... (Arbogast et al. 2002; Clutton-Brock, 1992). By the Bronze Age, domestic horses ( Equus caballus ) were in use throughout Europe (Harding 2000; Kristiansen and Larsson 2005), achieving a remarkably high-profile position by the first millennium BC in the equestrian societies of Celtic Europe (Aldhouse Green 1997; Arbogast et al. 2002: 67–84; Bo ̈ ko ̈ nyi 1974: 255–62; Renfrew 1998). This transition is still poorly understood. This is in terms of the continuity of regional wild horse populations into the Holocene, the role of European wild horses in horse domestication, the origins and early use of domestic horses and the mechanisms of spread of the domestic horses once they appear. This paper will review the evidence for the transition from wild horses to domestic horses in Europe. Horse domestication has been the subject of considerable research and debate (see recent critical reviews by Levine (1999, 2004) and Olsen (2006)). A great deal of this has focused on the archaeology of the Eurasian steppes – the grasslands stretching from Hungary in the west to the mountains of Central Asia in the east – which is a natural habitat for the horse (Bendrey 2011a; Ferret 2009; Olsen 1996). Earlier claims for domestic horses at Eneolithic Dereivka, Ukraine (e.g. Bibikova 1986; Bo ̈ ko ̈ nyi 1978) have been convincingly disputed (see Levine 1990, 1999, 2004; Uerpmann 1990). However, recent results of projects investigating early horse domestication have identified a number of lines of evidence that point to horse domestication on the Eurasian steppes during the fourth millennium BC . In the eastern steppes (defined here as that part of the steppes east of the line of the Ural Mountains; see Fig. 2), zooarchaeological and lipid residue analyses by Outram et ...

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... bone in the assemblage revives the problem of when domestic horses were introduced in Italy. In the recent past Equus ferus, which roamed Eurasian steppes and prairies during the Late Pleistocene, was thought to have disappeared from the central and western areas of the continent around 10,000 BP. (Bendrey, 2012;Sommer et al, 2011). Excessive human predation and, above all, the challenging Holocene climatic and environmental changes could explain this event. ...
... Excessive human predation and, above all, the challenging Holocene climatic and environmental changes could explain this event. Recently, new evidence was found indicating the survival of wild horses in Europe (Italy included) between the Pre-Boreal and the Atlantic (9,600-3,750 BC) (Aprile et al, 2017;Bendrey, 2012;Bon and Boscato, 1993;Sommer et al, 2011). Presently, the earliest evidence of Equus caballus comes from sites dated to the 5th-4th millennium BC in Kazakstan and Ukraine (Bökönyi 1980;Gaunitz et al, 2018). ...
... bone in the assemblage revives the problem of when domestic horses were introduced in Italy. In the recent past Equus ferus, which roamed Eurasian steppes and prairies during the Late Pleistocene, was thought to have disappeared from the central and western areas of the continent around 10,000 BP. (Bendrey, 2012;Sommer et al, 2011). Excessive human predation and, above all, the challenging Holocene climatic and environmental changes could explain this event. ...
... Excessive human predation and, above all, the challenging Holocene climatic and environmental changes could explain this event. Recently, new evidence was found indicating the survival of wild horses in Europe (Italy included) between the Pre-Boreal and the Atlantic (9,600-3,750 BC) (Aprile et al, 2017;Bendrey, 2012;Bon and Boscato, 1993;Sommer et al, 2011). Presently, the earliest evidence of Equus caballus comes from sites dated to the 5th-4th millennium BC in Kazakstan and Ukraine (Bökönyi 1980;Gaunitz et al, 2018). ...
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... Horses have a long history of being used in Britain for riding, traction and draught, with evidence of bridles from the Bronze Age (3100-300 BCE) and bit-related damage in the mouths of horses from the Iron Age (500-332 BCE) when metal bits were initially introduced [3,4]. The first documented use of horses for sport dates back to 800 BCE when chariot races took place, and in subsequent centuries, the Romans raced horses during high-profile events in purpose-built arenas [5]. ...
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