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![-Map of europe showing the known records of Gobius couchi and a detail map of the eastern part of the Ionian Sea with arrow showing locality of the new record of Gobius couchi from the Corfu Island. [Carte d'Europe présentant les signalements connus du gobie G. couchi et la carte détaillée de l'Est de la mer Ionienne avec la flèche indiquant le signalement récent du gobie G. couchi de Corfou.]](profile/Marcelo-Kovacic/publication/258507460/figure/fig1/AS:392759287533571@1470652519290/Map-of-europe-showing-the-known-records-of-Gobius-couchi-and-a-detail-map-of-the-eastern.png)
-Map of europe showing the known records of Gobius couchi and a detail map of the eastern part of the Ionian Sea with arrow showing locality of the new record of Gobius couchi from the Corfu Island. [Carte d'Europe présentant les signalements connus du gobie G. couchi et la carte détaillée de l'Est de la mer Ionienne avec la flèche indiquant le signalement récent du gobie G. couchi de Corfou.]
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Context 1
... -Three juveniles of unidentified sex, 20.0 + 4.4 mm (Fig. 2), 19.5 + 5.0 mm and 15.6 + 4.1 mm, NMP P6d 96/2008, Moraitika (N 39°28'25", e 19°55'53"), Corfu Island, Greece (Fig. 1 Meristic methods follow Miller (1988). Meristic abbreviations. -A, anal fin; C, caudal fin; D, dorsal fins; P, pectoral fin; V, pelvic disc; ll, scales in lateral series; TR, scales in transverse series. V emargination is the difference between the longest branches of fourth branched rays and the shortest branches of fifth branched ...
Context 2
... this case, a redetermination of gobies from NMP revealed the first record of G. couchi for the Ionian Sea and Greece. This species has been known from a few localities around the british Isles (Miller and el-Tawill, 1974;Minchin, 1988), from the Tyrrhenian Sea ( Stefanni and Mazzoldi, 1999), the Adriatic Sea ( Kovačić, 2001) and the eastern levant Sea ( Kovačić and Golani 2007), see Fig.1. The morphologi- cal characteristics agree with the species diagnosis. ...
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Citations
... In the Atlantic, known from Ireland, the south of Great Britain and northern Brittany, France (Miller & El-Tawil 1974;Baldock & Kay 2012;Iglésias et al. 2020). In the Mediterranean Sea, recorded from Cadaquès, Spain (Renoult et al. 2021a), Port-Vendres and Saint-Mandrier, France (Louisy 2005;Iglésias et al. 2021b), Liguria, Italy (Liu et al. 2009a), Ischia Island, Italy (Stefanni & Mazzoldi 1999), Malta , Croatia (Kovačić 2001;Kovačić et al. 2012b), Corfu Island, Greece (Šanda & Kovačić 2009), Crete , Cyprus (Kovačić & Golani 2007a) and southern part of the Dardanelles Strait, Turkey (Özen et al. 2009). In the Black Sea, from Crimea (Karpova & Boltachev 2018). ...
Numerous photographs of live fishes posted by anglers and divers on social media and citizen science databases are important sources of information for ichthyological research. However, validating records that extend the known ecology and bathymetric or geographic distribution of species should rely on a rigorous identification process. The family Gobiidae, with their small size, superficial resemblance among species and high species richness are particularly difficult to identify. Therefore, the identification from photographs of live individuals of Mediterranean marine gobies from the continental shelf was studied. A dichotomous identification key is provided based on photographs of live individuals, allowing positive identification of 41 out of the 66 species reviewed in this publication. Then, for all 66 species we provide a brief description of important characters, which can be used for provisional identification for those species that could not be positively identified using the key. Pending further progress in identification of live individuals, we suggest that records extending the known geographic and ecological species distribution be taken into account only if they could be validated using the dichotomous identification key.
... In the Mediterranean Sea, there are several records from France, Continental Spain (Cadaquès, Renoult et al., 2021; Cartagena, José Luis Alcaide, www.joseluisalcaide.com); Mallorca Island, Marcel Montanyès, 2020, iNaturalist), Italy (Liguria, Liu et al., 2009a;Ischia Island, Stefanni & Mazzoldi, 1999; published photography in Libourne, Tuscany, Stefano Guerrieri, 2016; Sardinia, photo by Egidio Trainito, 2011), Malta , Croatia (Kovačić, 2001a;published photography by Roberto Pillon, 2010, Greece (Corfu Island, Šanda and Kovačić, 2009;Crete, Kovačić et al., 2011), Cyprus (Kovačić & Golani, 2007) and Turkey (southern part of the Dardanelles Strait, Ozen et al., 2009). Couch's goby was recently discovered in Crimea, in the Black Sea (Karpova & Boltachev, 2018). ...
... It was described from coastal waters of Great Britain (Miller & ElTawil 1974) and later recorded near southwestern and northern Ireland (Minchin 1988;Costello 1992). Later, the species was found in the Mediterranean Basin: in 1996-1997, near the coast of Croatia (Kovačić 2001); in 1999, in the Tyrrhenian Sea near Ischia (Stefanni & Mazzoldi 1999); in 2004, in the Ionian Sea near Corfu (Šanda & Kovačić 2009); in 2007, in the Aegean Sea, in the southern part of the Dardanelles (Ozen et al. 2009); in 2009, in the Ligurian Sea (Liu et al. 2009); in 2010, near Crete (Kovačić et al. 2011);in 2011, near Malta (Kovačić et al. 2013. In the Black Sea, the species was recorded in 2015-2016 only in waters of Ukraine in Kozacha and Karantynna Coves , where the species has completely adapted to conditions of the south-western coast of the Crimea. ...
The present paper is devoted to a literature-based retrospective analysis of changes in the species composition of the family Gobiidae of waters of Ukraine since the 1930s. Keys and checklists published earlier cover the fish fauna of separate regions of Ukraine, either only of river basins or of coastal waters of the Black Sea and the Sea of Azov. The first checklist of fishes of Ukraine was published by academician A. M. Nikolsky in 1930 including both freshwater and marine species known in that time. In that work, the family Gobiidae includes 23 species of 7 genera. In the paper by D. A. Tretyakov (1947), 23 species of 10 genera of true gobies are listed for waters of Ukraine with some changes in the species composition. In one of the volumes of the series “Fauna of Ukraine. Fishes” devoted to true gobies (Smirnov 1986), data on 25 species of 11 genera are presented. According to the data by Yu. Movchan (2011), the composition of the family largely expanded comprising 33 species of 13 genera, including 5 marine species that were recorded in that time in coastal waters of the Crimea as well as 1 newly described species from the Chorna River. Later, five more marine species, known earlier only in the Mediterranean Basin, were found in waters of Ukraine, as well as another species in the Sea of Azov. It was established that the current fish fauna of the family Gobiidae of Ukraine includes 38 species of 17 genera, according to the literature and data from the fish collection of the National Museum of Natural History NAS of Ukraine (Kyiv). The geographic range and distribution of each species in waters of Ukraine are presented. The fauna of true gobies of Ukraine is composed of species of the Ponto-Caspian (52.6 %) and Atlantic-Mediterranean (47.4 %) faunal complexes. Endemics (4 species) are highlighted along with recent intruders of the Black Sea that have appeared here since the early 2000s (8 species); the time of appearance for 2 species remains unknown.
... For the first time, it was found in the coastal waters of Great Britain (Cornwall), southwestern and Northern Ireland (Miller and El-Tawin, 1974;Minchin, 1987Minchin, , 1988Costello, 1992). Later, this species was found in the Mediterranean basin: in 1996−1997 off the Croatian coasts (Kovačić, 2001); in 1999, in Tyrrhenian Sea near Ischia Island (Stefanni and Mazzoldi, 1999); in 2004, in the Ionian Sea near Corfu Island (Šanda and Kovačić, 2009); in 2007, in the Aegean Sea, southern part of the Dardanelles Strait (Özen et al., 2009); in 2009, in the Ligurian Sea (Liu et al., 2009); in 2010, in the Libian Sea, near Crete (Kovačić et al., 2011);in 2011, near Malta (Kovačić et al., 2013. ...
Couch’s goby Gobius couchi was found for the first time off the southwestern coast of Crimea, in the coves of Sevastopol. Earlier, this goby was not noted in the Black Sea; its range included the Atlantic coasts of England and Ireland and the northeastern part of Mediterranean and Aegean seas. The present paper provides the description of the morphological characteristics of Couch’s goby, including the diagnostic features, the data on its distribution near the Crimean coasts, and some features of ecology and behavior. The data on fecundity and time of reproduction of this goby are presented for the first time. It is suggested that the appearance of Couch’s goby near the Crimean coasts is determined by the process of “Mediterranization” of the Black Sea.
... macrocephalus , Z. zebrus) offered the chance to check the morphology and identification methods at these specimen lengths. in the case of G. couchi, the present specimen showed that juveniles at standard length of about 18 mm and total length of about 22 mm can be identified by the species diagnoses and published identification keys (Miller, 1986; Kovačić, 2008). However, at a standard length of about 16 mm and total length of about 20 mm, the complete absence of visible scales on the predorsal area required a different approach for species identification (Šanda and Kovačić, 2009 ). M. macrocephalus could be identified by species diagnoses and published keys for identifications (Miller, 1986; Kovačić, 2008) at standard length of just about 9 mm and total length of just 11 mm. ...
Individuals of small juvenile gobies, some of them just 10 mm long, were collected during a study on infralittoral cobble bed fauna around the Maltese Islands between July and September 2011. These represent the first records of three gobiid species for Malta, including Gobius couchi Miller & El Tawil, 1974 and the smallest known juveniles of Millerigobius macrocephalus (Kolombatovič, 1891) and Zebrus zebrus (Risso, 1827). Morphology and colouration, including diagnostic features and photographs, are provided for these species and the problems of identification of small juveniles are discussed.
Despite being one of the most speciose fish families in the Mediterranean Sea, knowledge about the diversity of gobies (Actinopterygii: Gobiidae) in this sea is still unsatisfactory, as documented by recent descriptions of a number of new species. Although very common in shallow water, Gobius incognitus Kovačić & Šanda, 2016, had escaped attention until 2016, when it was discovered. Due to its overall superficial morphological similarity, G. incognitus used to be confused with a much rarer species, Gobius bucchichi Steindachner, 1870, which was considered one of the most common shallow-water gobies in the Mediterranean Sea. In this work, we tested the suitability of the genetic data (mitochondrial gene encoding cytochrome b) for identifying and distinguishing between these two goby species, and assessed the congruency between the distribution records based on genetic data and those based on morphological identification. We analysed material of 304 specimens of G. incognitus and G. bucchichi from 49 localities covering a considerable part of the Mediterranean Sea, Black Sea, and the Atlantic Ocean near Gibraltar, representing 19 geographically well-separated areas. We detected 270 sequences of G. incognitus, and only 34 of G. bucchichi. In both species, a high haplotype variability was observed. The sequence species identity matched morphological identification for all specimens for which vouchers were available. The mean uncorrected p-distance between G. incognitus and G. bucchichi was 13%, while the mean intraspecific distances were much lower (0.63% and 0.68%, respectively). We found 79 fixed mutations between these two species. Data on distribution based on genetic identification are completely congruent with published results based on morphological identification. The results of this study support molecular methods as a reliable tool for distinguishing morphologically similar fish species, which is particularly useful when only tissue is available for determination.
The gobies (Gobiidae) are the most diverse fish family in the Mediterranean Sea. Nevertheless, knowledge on their diversity, taxonomy, and phylogenetic relationships is still inadequate. The phylogenetic analyses reveal two genetically highly distinct clades among specimens identified as Zebrus zebrus . A new species, Zebrus pallaoroi sp. nov., is described based on an integrative approach. The neotype of Zebrus zebrus is designated. Genetic data confirm a pronounced level of divergence between Z. pallaoroi and Z. zebrus , with the mean genetic distance on cytochrome b being 18.1% and 1.07% on rhodopsin. Phylogenetic relationships within the Gobius -lineage were estimated on both markers. Morphologically, Z. pallaoroi is distinguished from the only congener Z. zebrus by having a snout longer than its eye, posterior nostril about 4/5–9/10 of the anterior nostril, eye diameter 4.3−4.7 in head length, ventrolateral head ridges transversally connected on the anterior side by a short transversal ridge, anterior membrane midline depth about 2/3 of the spinous ray, head canal pore α diameter about half of the distance between pore ρ and ρ 1 , suborbital sensory papillae row 5i going downwards to or near the level of row d , the distance between row 5i and row d absent or much smaller than the length of row 5i , and the body with ten to eleven vertical dark brown bands. Zebrus pallaoroi was recorded from the southern Adriatic, northern Ionian, and northern and western Aegean Seas, and is a cryptobenthic fish from very shallow waters.
A new species of the gobiid genus Gobius (Gobiidae, Perciformes), Gobius incognitus sp. nov. is described from the Mediterranean Sea, and its most morphologically similar species Gobius bucchichi is redescribed. The new species is distinguished from its congeners by: scales in lateral series 51-59; predorsal scales 25-35; opercle scaled in adults with 10-16 scales present; pectoral fin with ray count 18-20 and free tips on upper rays well developed and on the first ray longer than two thirds of the entire ray length; pelvic disc complete and with well-developed anterior membrane without lateral lobes; anterior oculoscapular canal with pore α at rear of orbit; oculoscapular row x(1) not extending forwards to pore β; suborbital row d discontinuous with large gap below suborbital rows 3 and 4; eye diameter 1·08-1·32 in snout length; by pigment rows on cheek and pigmentation on pectoral-fin base.
Prior to the present study, the gobiid biodiversity of the Adriatic Sea varied significantly between the northern (98% of the known Adriatic gobiid biodiversity) and central (89%) as compared to the southern Adriatic Sea (39%). Thirteen species are newly recorded for the southern Adriatic area and one for the central Adriatic. The relatively small collecting effort in this survey produced a large increase in the known gobiid biodiversity, which suggests that the low number of known Adriatic gobiid species in the southern Adriatic mirrors only sampling intensity and not the real species richness of the area. The gobiid zoogeography in the Adriatic is compared with that of the Mediterranean.
