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10 Limestone cliffs of Málaga. Crithmo maritimi-Limonietum malacitani 

10 Limestone cliffs of Málaga. Crithmo maritimi-Limonietum malacitani 

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The coastal vegetation of the Iberian Peninsula (including the Balearic Islands) is represented by a set of plant communities that colonize biotopes as diverse as sea cliffs, beaches, dunes, marshes and estuaries. From a geographical point of view, we have differentiated the following sections: in the north, the coastal strip extending from the mou...

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... the diversity of the Iberian coast, we have separated the different environments: first, rocky coasts with abundant cliffs; second, sedimentary coasts where beaches, dunes, as well as estuaries and coastal lagoons are frequent. The coastline has been divided into three main sections, according to biogeographic and bioclimatic features ( Fig. ...
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... is made up of the coastal stretch between Ría de Aveiro (Portugal) and Punta de Tarifa (Spain) at the Strait of Gibraltar. It is integrated in the Coastal Lusitano-Andalusian Province of the Mediterranean Region and includes four biogeographical sectors: Divisorio-Portugues, Ribatagano-Sadense, Algarviense and Gaditano-Onubense Litoral p.p. (Fig. ...
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... western stretch is dominated by materials of Paleozoic and Precambrian origin, such as granites of the Galician Atlantic coast, or the slates, quartzites and sandstones of northern Galicia and western areas of Asturias, while the eastern Fig. 8.1 Map of the Iberian Peninsula coastline with the locations of the main sites mentioned in the text. 1: Bidasoa River; 2: Cape Peñas (Asturias); 3: Ría de Foz (Lugo); 4: Rías Altas; 5: Rías Bajas; 6: Ría de Aveiro; 7: Cape Mondego; 8: Peniche Peninsula; 9: Cape Roca; 10: Cape Espichel; 11: Troia Peninsula; 12: Cape Sines; 13: Cape San ...
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... which are crowned by fossil dunes ( Neto et al. 2001). The cliff communities, less diverse than the previous ones, have among others, the following species: Crithmum mari- timum, Limonium ferulaceum, Limonium virgatum, Limonium ovalifolium, Daucus hispanicus subsp. halophilus, Dittrichia maritima, Pallenis maritima and Spergularia australis ( Fig. 8.4). The Spanish stretch (Huelva-Cádiz) is character- ized by low coasts with large areas with extensive dune ecosystems, while only some sandstone cliffs emerge on the Cádiz coast with Limonium emarginatum (Asensi ...
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... with Ammophila arenaria subsp. australis are also represented by two communities: the Otantho-Ammophiletum australis which stretches from the Cantabrian-Atlantic coast to Peniche (Portugal) and the Loto cretici-Ammophiletum australis, an association on the southern Iberian coast that differs from the former by the abundance of Lotus creticus ( Fig. ...
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... by Juniperus navicularis together with Daphne gnidium, Osyris alba, Osyris lanceolata, Phillyrea angustifolia, Pistacia lentiscus, Rubia peregrina subsp. angustifolia and others. In this environment there are fragile communities dominated by species of Cladonia (C. mediterranea, C. portentosa, among others), that are very sensitive to trampling ( Fig. 8.6). The Santolinetum impressae, a community with the endemic Santolina impressa is common on disturbed ...
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... genistoidis covers large areas in this territory (Neto 2002 A serious problem in the dune ecosystems of Portugal is the presence of invasive alien plants, like Acacia longifolia, Acacia melanoxylon, Arctotheca calendula, Carpobrotus edulis, Oenothera stricta which cover large areas preventing the devel- opment of natural communities ( Fig. ...
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... the outer fixed dunes within reach of the sea winds, a coastal juniper com- munity is established, forming a discontinuous band in contact with the semi-fixed dunes ( Fig. 8.8). The association Rhamno oleoidis-Juniperetum macrocarpae, with Daphne gnidium, Juniperus turbinata, Pistacia lentiscus, Rhamnus oleoides, Rubia peregrina subsp. longifolia and Phillyrea angustifolia, is endemic on the Huelva and Cádiz coast ( Rivas-Martínez et al. 1980). The mature stage of the dune ecosystem corresponds to the ...
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... Sarcocornia perennis together with other species and grasses that differentiate the two associations of this coastal stretch: the Puccinellio maritimae-Sarcocornietum perennis extending from the Cantabrian coast to the mouth of the Tagus, and the Puccinellio ibericae-Sarcocornietum perennis from the south of this river to the Cádiz coast ( Fig. 8.9). These communities are replaced, in the inner band, somewhat higher, by the Halimiono portulacoidis-Sarcocornietum alpini, whose dominant species is Sarcocornia alpini, common in the SW Iberian coasts and NW Morocco. It extends throughout the southwest of the Iberian Peninsula up to south of the Tagus River and is replaced by the ...
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... rugged coast, cliff communities are scarce, but include: the Crithmo maritimi- Limonietum malacitani with Limonium malacitanum on the Málaga coast and the Limonio cossonianae-Lycietum intricati, a community composed of Crithmum maritimum accompanied by less specialized species like Limonium cossonianum, Lycium intricatum and Salsola papillosa ( Fig. 8 Another feature of this territory is the Juniperus macrocarpa community devel- oped on granite and limestone rocks exposed to sea wind. The association Astragalo tragacanthae-Juniperetum macrocarpae has several territorial species, e.g. Astragalus tragacantha, Festuca ruscinonensis and Polycarpon polycarpoi- des subsp. catalaunicum ...
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... islands, especially in the genus Limonium, that possess 17 endemic species in these environments, 8 of which are exclusive to Majorca (Limonium balearicum, Limonium caprariense, Limonium carregadorense, Limonium dragonericum, Limonium gymnesicum, Limonium majoricum, Limonium muradense, Limonium portopretanum, Limonium pseudodic- tyocladum) ( Fig. 8.11). On some rocky coasts of Majorca and Minorca a second band of vegetation is found, occupying positions near the sea. These communities comprise thorny, cushion-shaped species, many of them endemic, e.g. Anthyllis hystrix, Carthamus balearicus, Dorycnium fulgurans, Euphorbia maresii, Launaea cervicornis and Santolina magonica. The most ...
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... coast, but do occur at Almería and Murcia. The Rhamno angustifoliae- Juniperetum turbinatae is well represented in Punta Entinas-Sabinar (Almería) where a dense forest is formed with Juniperus turbinata and Pistacia lentiscus as dominant species, accompanied by, e.g., Rhamnus oleoides var. angustifolia, Maytenus europaeus and Lycium intricatum ( Fig. ...
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... in the coastal areas of the Cape Gata-Níjar Natural Park, the fixed dunes are colonized by open populations of Ziziphus lotus, a species that inhabits semiarid and arid sites in the southeast Iberian Peninsula (Fig. 8.13). The physiognomy of this peculiar community (Ziziphetum loti) consists of separate specimens of Ziziphus which retain sand in mounds of more than 1 m high and about 25 m 2 sur- face area, on which other plants, like Asparagus albus, Lycium intricatum, Withania frutescens, Rubia peregrina, Thymelaea hirsuta and Arisarum simorrhinum, ...
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... tabernaemontani, Typha latifolia and Typha angustifolia as the dominant species. Near it, another helophytic community of greater height, up to 3 m, is developed. The Typho domingensis-Phragmitetum maximi with Typha domingensis and Phragmites australis subsp. altissimus is the association that grows on small peat islets subjected to waterlogging (Fig. 8.14). The formation of these peat islets, locally called "mates", is favoured by the accumulation of dense clumps of Cladium mariscus, whose association (Hydrocotylo-Mariscetum serrati) represents a serial stage of vegetation with reeds and ...
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... of shrub species. The selective use of shrub species by browsers has transformed the Mediterranean woodland in many areas. The results depend on the environmental context and management; a moderate and sound grazing favours forage shrub species (different Papilonaceae of genera as Cytisus, Genista, Retama, Adenocarpus, Pterospartum, among others, Fig. 10.8) and an appropriate composition of pasture communities (herbaceous and small shrubs), while the overgrazing in fragile terrains, and also the plowing excess, foster ero- sion and the dominance of species not consumed by herbivores and adapted to poor terrain (mainly Cistaceae, Ericaceae, etc.). -Grazing associated with Mediterranean ...
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... is made up of the coastal stretch between Ría de Aveiro (Portugal) and Punta de Tarifa (Spain) at the Strait of Gibraltar. It is integrated in the Coastal Lusitano-Andalusian Province of the Mediterranean Region and includes four biogeographical sectors: Divisorio-Portugues, Ribatagano-Sadense, Algarviense and Gaditano-Onubense Litoral p.p. (Fig. 8.1 ...
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... western stretch is dominated by materials of Paleozoic and Precambrian origin, such as granites of the Galician Atlantic coast, or the slates, quartzites and sandstones of northern Galicia and western areas of Asturias, while the eastern Fig. 8.1 Map of the Iberian Peninsula coastline with the locations of the main sites mentioned in the text. 1: Bidasoa River; 2: Cape Peñas (Asturias); 3: Ría de Foz (Lugo); 4: Rías Altas; 5: Rías Bajas; 6: Ría de Aveiro; 7: Cape Mondego; 8: Peniche Peninsula; 9: Cape Roca; 10: Cape Espichel; 11: Troia Peninsula; 12: Cape Sines; 13: Cape San Vicente; 14: Ría Formosa; 15: Doñana National Park; 16: Cape Trafalgar; 17: Tarifa; 18: Cape Sacratif (Granada); 19: Punta Entinas-Sabinar; 20: Cape of Gata; 21: Santa Pola (Alicante); 22: Altea (Alicante); 23: Cape San Antonio; 24: Albufera de Valencia; 25: Delta del Ebro; 26: Cape Creus; 27: Balearic Islands zone, from Cape Peñas (Asturias) to the Bidasoa River, is characterized by Mesozoic and Cenozoic materials, especially limestones, dolomites and ...
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... Itinerant grazing, in which the flocks are led by a shepherd and return every day to a central place (Fig. 10.7). It plays an important role in the recovery of the fertility of arable lands. Also it allows to tap into the benefits offered by landscape patchiness, plots with different composition and production varying in aspect, soil moisture, depth and fertility. -Transhumance. An annual cyclical migration of livestock between zones of which the maxima of productivity are complementary in time: i.e. mountain summer pastures are complemented with pasturelands productive in winter and spring. -Scattered trees. Sites with silvopastoral formations shape many livestock- extensive landscapes that are common in many areas of the Peninsula. Trees were selected for and managed according to their canopy (shelter), fruit produc- tion and foraging potential. -Consumption of shrub species. The selective use of shrub species by browsers has transformed the Mediterranean woodland in many areas. The results depend on the environmental context and management; a moderate and sound grazing favours forage shrub species (different Papilonaceae of genera as Cytisus, Genista, Retama, Adenocarpus, Pterospartum, among others, Fig. 10.8) and an appropriate composition of pasture communities (herbaceous and small shrubs), while the overgrazing in fragile terrains, and also the plowing excess, foster ero- sion and the dominance of species not consumed by herbivores and adapted to poor terrain (mainly Cistaceae, Ericaceae, etc.). -Grazing associated with Mediterranean crops. The use of crop residues and stubble represents an important mechanism for the restitution of soil fertility. Short displacements (about 100 km) between mountain pasturelands and the new irrigated zones are gaining prominence and replacing in some cases the tradi- tional longer transhumance (around 500 km), as a support to maintain the utiliza- tion of mountain pastures. Capital in the Collegiate Church of the Alquezar, North of Huesca (Aragón). Eleventh century. Romanesque ...
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... Málaga-Altea (Alicante). The proximity of the Baetic Ranges to the sea has led to a coastline with rocky outcrops alternating with small beaches at their base. Sedimentation processes in large sectors of this coast have created accumula- tions of powerful torrential cones and a narrow coastal plain. Despite the rugged coast, cliff communities are scarce, but include: the Crithmo maritimi- Limonietum malacitani with Limonium malacitanum on the Málaga coast and the Limonio cossonianae-Lycietum intricati, a community composed of Crithmum maritimum accompanied by less specialized species like Limonium cossonianum, Lycium intricatum and Salsola papillosa ( Fig. 8 Another feature of this territory is the Juniperus macrocarpa community devel- oped on granite and limestone rocks exposed to sea wind. The association Astragalo tragacanthae-Juniperetum macrocarpae has several territorial species, e.g. Astragalus tragacantha, Festuca ruscinonensis and Polycarpon polycarpoi- des subsp. catalaunicum (Rivas-Martínez and Cantó 2002a). 4. Balearic Islands. The Balearic archipelago consists of two groups of islands and about one hundred islets: in the north Majorca, Minorca, Cabrera (Gimnesias); in the south Ibiza and Formentera (Pitiusas). Most coastal cliffs consist of lime- stone, except for some areas with siliceous rocks in the north and east of ...
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... borders of ponds and lagoons are occupied by reeds and cattails that develop in freshwater, although they tolerate a slight salinity. The Typho-Schoenoplectetum tabernaemontani is the association that inhabits this environment with Schoenoplectus tabernaemontani, Typha latifolia and Typha angustifolia as the dominant species. Near it, another helophytic community of greater height, up to 3 m, is developed. The Typho domingensis-Phragmitetum maximi with Typha domingensis and Phragmites australis subsp. altissimus is the association that grows on small peat islets subjected to waterlogging (Fig. 8.14). The formation of these peat islets, locally called "mates", is favoured by the accumulation of dense clumps of Cladium mariscus, whose association (Hydrocotylo-Mariscetum serrati) represents a serial stage of vegetation with reeds and ...
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... southwestern coast (Costa Vicentina) is characterized by high cliff faces, formed by unstable sandstones or hard substrates (schists), of 40-120 m high, which are crowned by fossil dunes ( Neto et al. 2001). The cliff communities, less diverse than the previous ones, have among others, the following species: Crithmum mari- timum, Limonium ferulaceum, Limonium virgatum, Limonium ovalifolium, Daucus hispanicus subsp. halophilus, Dittrichia maritima, Pallenis maritima and Spergularia australis ( Fig. 8.4). The Spanish stretch (Huelva-Cádiz) is character- ized by low coasts with large areas with extensive dune ecosystems, while only some sandstone cliffs emerge on the Cádiz coast with Limonium emarginatum (Asensi ...
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... dunes or grey dunes typically are floristically rich and contain several chamaephytes: Armeria pungens, Artemisia crithmifolia, Crucianella maritima, Lotus creticus, Scrophularia frutescens, and some endemics, e.g. Armeria wel- witschii, Herniaria maritima, Linaria lamarckii and Thymus carnosus. The associa- tion Artemisio crithmifoliae-Armerietum pungentis has a wide distribution area ranging from Tarifa to the Tagus Estuary, being replaced in further north up to Cape In the inner paleodunes of the Sado Basin, the mature stage corresponds to the association Daphno gnidii-Juniperetum navicularis, basically formed by Juniperus navicularis together with Daphne gnidium, Osyris alba, Osyris lanceolata, Phillyrea angustifolia, Pistacia lentiscus, Rubia peregrina subsp. angustifolia and others. In this environment there are fragile communities dominated by species of Cladonia (C. mediterranea, C. portentosa, among others), that are very sensitive to trampling ( Fig. 8.6). The Santolinetum impressae, a community with the endemic Santolina impressa is common on disturbed ...
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... dunes or white dunes with Ammophila arenaria subsp. australis are also represented by two communities: the Otantho-Ammophiletum australis which stretches from the Cantabrian-Atlantic coast to Peniche (Portugal) and the Loto cretici-Ammophiletum australis, an association on the southern Iberian coast that differs from the former by the abundance of Lotus creticus ( Fig. ...
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... Balearic Islands have a great diversity of cliff communities, especially in the Gimnesias Islands, where nine of the ten associations described from the area are located. The special characteristics of the local biotopes, together with the isolation of the populations, have favoured a rich speciation in these islands, especially in the genus Limonium, that possess 17 endemic species in these environments, 8 of which are exclusive to Majorca (Limonium balearicum, Limonium caprariense, Limonium carregadorense, Limonium dragonericum, Limonium gymnesicum, Limonium majoricum, Limonium muradense, Limonium portopretanum, Limonium pseudodic- tyocladum) ( Fig. 8.11). On some rocky coasts of Majorca and Minorca a second band of vegetation is found, occupying positions near the sea. These communities comprise thorny, cushion-shaped species, many of them endemic, e.g. Anthyllis hystrix, Carthamus balearicus, Dorycnium fulgurans, Euphorbia maresii, Launaea cervicornis and Santolina magonica. The most frequent association is Launaeetum cervicornis which occurs adjacent to the Limonium communities of the lower band. A summary of these communities is shown in Table ...
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... the outer fixed dunes within reach of the sea winds, a coastal juniper com- munity is established, forming a discontinuous band in contact with the semi-fixed dunes ( Fig. 8.8). The association Rhamno oleoidis-Juniperetum macrocarpae, with Daphne gnidium, Juniperus turbinata, Pistacia lentiscus, Rhamnus oleoides, Rubia peregrina subsp. longifolia and Phillyrea angustifolia, is endemic on the Huelva and Cádiz coast ( Rivas-Martínez et al. 1980). The mature stage of the dune ecosystem corresponds to the association Osyrio quadripartitae-Juniperetum turbinatae already mentioned for the Portuguese stretch. The open spaces of the fossil dunes are occupied by oligotrophic grassland in which annual plants such as Arenaria algarbiensis, Linaria tursica, Loeflingia baetica, Malcolmia triloba, Vulpia fontquerana and others grow (Linario donyanae- Loeflingietum ...
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... navicularis is an endemic species of the Iberian southwestern coast that reaches some coastal areas of Huelva and Cádiz. It grows on dry sandy sub- strata, sometimes associated with podzol soils, conditions that prevent the develop- ment of the Quercus suber forests which normally occupy soils with shallower and wetter sands. Human activity in the Sado Estuary, due to agriculture and forestry (plantations of Eucalyptus camaldulensis, Pinus pinea and Pinus pinaster), has encouraged the development of scrub rich in endemic species such as Armeria pini- folia, Armeria rouyana, Stauracanthus genistoides, Thymus capitellatus and Ulex australis subsp. welwitschianus. The association Thymo capitellati-Stauracanthetum genistoidis covers large areas in this territory (Neto 2002 A serious problem in the dune ecosystems of Portugal is the presence of invasive alien plants, like Acacia longifolia, Acacia melanoxylon, Arctotheca calendula, Carpobrotus edulis, Oenothera stricta which cover large areas preventing the devel- opment of natural communities ( Fig. ...
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... in the coastal areas of the Cape Gata-Níjar Natural Park, the fixed dunes are colonized by open populations of Ziziphus lotus, a species that inhabits semiarid and arid sites in the southeast Iberian Peninsula (Fig. 8.13). The physiognomy of this peculiar community (Ziziphetum loti) consists of separate specimens of Ziziphus which retain sand in mounds of more than 1 m high and about 25 m 2 sur- face area, on which other plants, like Asparagus albus, Lycium intricatum, Withania frutescens, Rubia peregrina, Thymelaea hirsuta and Arisarum simorrhinum, grow ( Peinado et al. 1992). A substitution stage of the Ziziphetum loti and also of Juniperus turbinata forests, on sandy soils, is a low scrub (tomillar) mainly formed by chamaephytes such as Teucrium dunense, Helianthemum almeriense subsp. scopulorum and Thymus hyemalis corresponding to the association Teucrio dunensis-Helianthemetum scopulorum. A community of annual plants Wahlenbergio nutabundae-Loeflingietum pentandrae) is developed on sandy soils after the rainy period. The vegetation of the arborescent scrub Ziziphus lotus is con- sidered a priority habitat in the Habitats Directive (European Commission ...
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... next band of vegetation, developed on soils subjected to tidal flow, is occu- pied by the chamaephyte Sarcocornia perennis together with other species and grasses that differentiate the two associations of this coastal stretch: the Puccinellio maritimae-Sarcocornietum perennis extending from the Cantabrian coast to the mouth of the Tagus, and the Puccinellio ibericae-Sarcocornietum perennis from the south of this river to the Cádiz coast ( Fig. 8.9). These communities are replaced, in the inner band, somewhat higher, by the Halimiono portulacoidis-Sarcocornietum alpini, whose dominant species is Sarcocornia alpini, common in the SW Iberian coasts and NW Morocco. It extends throughout the southwest of the Iberian Peninsula up to south of the Tagus River and is replaced by the Triglochino maritimae-Sarcocornietum alpini in the saltmarshes north of this river ( Costa et al. ...
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... turbinata forests, which grow in the innermost dunes, are scarce on the Mediterranean coast, but do occur at Almería and Murcia. The Rhamno angustifoliae- Juniperetum turbinatae is well represented in Punta Entinas-Sabinar (Almería) where a dense forest is formed with Juniperus turbinata and Pistacia lentiscus as dominant species, accompanied by, e.g., Rhamnus oleoides var. angustifolia, Maytenus europaeus and Lycium intricatum ( Fig. ...
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... understand the diversity of the Iberian coast, we have separated the different environments: first, rocky coasts with abundant cliffs; second, sedimentary coasts where beaches, dunes, as well as estuaries and coastal lagoons are frequent. The coastline has been divided into three main sections, according to biogeographic and bioclimatic features ( Fig. ...

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