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Dorsal habitus of Promecheilidae and Chalcodryidae. A) Sirrhas variegatus (© CSIRO; scale bar = 1 mm), B) Onysius anomalus, C) Chalcodrya hilaris Watt, D) Philpottia mollis Broun.

Dorsal habitus of Promecheilidae and Chalcodryidae. A) Sirrhas variegatus (© CSIRO; scale bar = 1 mm), B) Onysius anomalus, C) Chalcodrya hilaris Watt, D) Philpottia mollis Broun.

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Onysius Broun, 1886, is transferred from the endemic New Zealand Chalcodryidae to Promecheilidae, reducing the number of genera of the former to two (Chalcodrya and Philpottia). Onysius shares several adult characters with Promecheilidae, including the abdominal ventrites 3 and 4 not connate and the tegmen subdivided into a basale and apicale (larv...

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Context 1
... Broun (1886) (Fig. 1B) was described as a genus of Melandryidae based on a specimen from the Mamaku (as Patetere) Plateau in the central part of North Island, New Zealand. Later the same year, Malacodrya Sharp (1886) was described (as a helopine tenebrionid) from Picton at the southeast corner of South Island and Greymouth along the west coast of South ...
Context 2
... Island and Greymouth along the west coast of South Island. Broun (1893) added a second species to Onysius, which was later synonymized with the type species, O. anomalus Broun, by Watt (1974). The monotypic genus Malacodrya was synonymized with Onysius by Broun (1915). The latter genus, plus the New Zealand genera Chalcodrya Redtenbacher (1868) (Fig. 1C) and Philpottia Broun (1915) (Fig. 1D), were placed by various authors (Broun 1915;Carter 1915;Champion 1893) in subfamilies or tribes now part of Tenebrionidae, but Watt (1967) and Crowson (1967) referred these groups, at least tentatively, to the family Zopheridae. Watt (1974a) proposed Chalcodryidae, which included Onysius anomalus ...
Context 3
... coast of South Island. Broun (1893) added a second species to Onysius, which was later synonymized with the type species, O. anomalus Broun, by Watt (1974). The monotypic genus Malacodrya was synonymized with Onysius by Broun (1915). The latter genus, plus the New Zealand genera Chalcodrya Redtenbacher (1868) (Fig. 1C) and Philpottia Broun (1915) (Fig. 1D), were placed by various authors (Broun 1915;Carter 1915;Champion 1893) in subfamilies or tribes now part of Tenebrionidae, but Watt (1967) and Crowson (1967) referred these groups, at least tentatively, to the family Zopheridae. Watt (1974a) proposed Chalcodryidae, which included Onysius anomalus Broun, the species of Chalcodrya and ...
Context 4
... (1967) referred these groups, at least tentatively, to the family Zopheridae. Watt (1974a) proposed Chalcodryidae, which included Onysius anomalus Broun, the species of Chalcodrya and Philpottia, plus the Tasmanian Sirrhas limbatus Champion (1893). Lawrence (1994) described the new and somewhat different species, Sirrhas variegatus Lawrence (Fig. 1A), also from Tasmania, redefined and expanded the South Temperate Perimylopidae St. George (1939), and transferred Sirrhas, plus the Tasmanian tenebrionid Melytra Pascoe, 1869, to that family. Onysius, however, was retained in Chalcodryidae. Perimylopidae was further expanded by Lawrence et al. (2010) with the inclusion of the Chilean ...
Context 5
... by Elgueta and Arriagada (1989) to be a possible relative of Chalcodryidae. This necessitated a change in the family name to Promecheilidae Lacordaire (1859). In the present study, the systematic position of Onysius is reconsidered and evidence given for its transfer to Promecheilidae, recording Diagnosis. Onysius differs from both Chalcodrya (Fig. 1C) and Philpottia (Fig. 1D) in having the antennae much longer than head and prothorax combined, the elytra tuberculate and much broader than the base of the prothorax, abdominal ventrites 3 and 4 not connate, tegmen subdivided into a basale and apicale, female genital tract with a small bursa and large spermatheca and spermathecal gland, ...
Context 6
... (1989) to be a possible relative of Chalcodryidae. This necessitated a change in the family name to Promecheilidae Lacordaire (1859). In the present study, the systematic position of Onysius is reconsidered and evidence given for its transfer to Promecheilidae, recording Diagnosis. Onysius differs from both Chalcodrya (Fig. 1C) and Philpottia (Fig. 1D) in having the antennae much longer than head and prothorax combined, the elytra tuberculate and much broader than the base of the prothorax, abdominal ventrites 3 and 4 not connate, tegmen subdivided into a basale and apicale, female genital tract with a small bursa and large spermatheca and spermathecal gland, lacinia without an ...

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The type species [placed in square brackets] of the following available genus-group names are designated for the first time: Allostrongylium Kolbe, 1896 [ Allostrongylium silvestre Kolbe, 1896], Auristira Borchmann, 1916 [ Auristira octocostata Borchmann, 1916], Blapidocampsia Pic, 1919 [ Campsia pallidipes Pic, 1918], Cerostena Solier, 1836 [ Cerostena deplanata Solier, 1836], Coracostira Fairmaire, 1899 [ Coracostira armipes Fairmaire, 1899], Dischidus Kolbe, 1886 [ Helops sinuatus Fabricius, 1801], Eccoptostoma Gebien, 1913 [ Taraxides ruficrus Fairmaire, 1894], Ellaemus Pascoe, 1866 [ Emcephalus submaculatus Brême, 1842], Epeurycaulus Kolbe, 1902 [ Epeurycaulus aldabricus Kolbe, 1902], Euschatia Solier, 1851 [ Euschatia proxima Solier, 1851], Heliocaes Bedel, 1906 [ Blaps emarginata Fabricius, 1792], Hemipristis Kolbe, 1903 [ Hemipristis ukamia Kolbe, 1903], Iphthimera Reitter, 1916 [ Stenocara ruficornis Solier, 1835], Isopedus Stein, 1877 [ Helops tenebrioides Germar, 1813], Malacova Fairmaire, 1898 [ Malacova bicolor Fairmaire, 1898], Modicodisema Pic, 1917 [ Disema subopaca Pic, 1912], Peltadesmia Kuntzen, 1916 [ Metriopus platynotus Gerstaecker, 1854], Phymatium Billberg, 1820 [ Pimelia maculata Fabricius, 1781], Podoces Péringuey, 1886 [ Podoces granosula Péringuey, 1886], Pseuduroplatopsis Pic, 1913 [ Borchmannia javana Pic, 1913], Pteraulus Solier, 1848 [ Pteraulus sulcatipennis Solier, 1848], Sciaca Solier, 1835 [ Hylithus disctinctus Solier, 1835], Sterces Champion, 1891 [ Sterces violaceipennis Champion, 1891] and Teremenes Carter, 1914 [ Tenebrio longipennis Hope, 1843]. Evidence suggests that some type species were misidentified. In these instances, information on the misidentification is provided and, in the following cases, the taxonomic species actually involved is fixed as the type species [placed in square brackets] following requirements in Article 70.3 of the International Code of Zoological Nomenclature: Accanthopus Dejean, 1821 [ Tenebrio velikensis Piller & Mitterpacher, 1783], Becvaramarygmus Masumoto, 1999 [ Dietysus nodicornis Gravely, 1915], Heterophaga Dejean, 1834 [ Opatrum laevigatum Fabricius, 1781], Laena Dejean, 1821, [ Scaurus viennensis Sturm, 1807], Margus Dejean, 1834 [ Colydium castaneum Herbst, 1797], Pachycera Eschscholtz, 1831 [ Tenebrio buprestoides Fabricius, 1781], Saragus Erichson, 1842 [ Celibe costata Solier, 1848], Stene Stephens, 1829 [ Colydium castaneum Herbst, 1797], Stenosis Herbst, 1799 [ Tagenia intermedia Solier, 1838] and Tentyriopsis Gebien, 1928 [ Tentyriopsis pertyi Gebien, 1940]. The following First Reviser actions are proposed to fix the precedence of names or nomenclatural acts (rejected name or act in square brackets): Stenosis ciliaris Gebien, 1920 as the type species for Afronosis G.S. Medvedev, 1995 [ Stenosis leontjevi G.S. Medvedev, 1995], Alienoplonyx Bremer, 2019 [ Alienolonyx ], Amblypteraca Mas-Peinado, Buckley, Ruiz & García-París, 2018 [ Amplypteraca ], Caenocrypticoides Kaszab, 1969 [ Caenocripticoides ], Deriles Motschulsky, 1872 [ Derilis ], Eccoptostira Borchmann, 1936 [ Ecoptostira ], † Eodromus Haupt, 1950 [† Edromus ], Eutelus Solier, 1843 [ Lutelus ], Euthriptera Reitter, 1893 [ Enthriptera ], Meglyphus Motschulsky, 1872 [ Megliphus ], Microtelopsis Koch, 1940 [ Extetranosis Koch, 1940, Hypermicrotelopsis Koch, 1940], Neandrosus Pic, 1921 [ Neoandrosus ], Nodosogylium Pic, 1951 [ Nodosogilium ], Notiolesthus Motschulsky, 1872 [ Notiolosthus ], Pseudeucyrtus Pic, 1916 [ Pseudocyrtus ], Pseudotrichoplatyscelis Kaszab, 1960 [ Pseudotrichoplatynoscelis and Pseudotrichoplatycelis ], Rhydimorpha Koch, 1943 [ Rhytimorpha ], Rhophobas Motschulsky, 1872 [ Rophobas ], Rhyssochiton Gray, 1831 [ Ryssocheton and Ryssochiton ], Sphaerotidius Kaszab, 1941 [ Spaerotidius ], Stira Agassiz, 1846 (Mollusca) [ Stira Agassiz, 1846 (Coleoptera)], Sulpiusoma Ferrer, 2006 [ Sulpiosoma ] and Taenobates Motschulsky, 1872 [ Taeniobates ]. Supporting evidence is provided for the conservation of usage of Cyphaleus Westwood, 1841 nomen protectum over Chrysobalus Boisduval, 1835 nomen oblitum.