Figure 16 - uploaded by Bernard Mottequin
Content may be subject to copyright.
![A-H. Schizophoria resupinata var. palliata Demanet, 1934 [Schizophoria (Pocockia) linguata (Quenstedt, 1871 in 1868-1871)]; Dréhance, Waulsort Formation (Tournaisian, Ivorian). A-D. RBINS a5883 (lectotype), almost complete valve in plan, lateral, posterior and anterior views. E-H. RBINS a5884 (paralectotype), slightly distorted dorsal valve in plan, lateral, posterior and anterior views. I-M. Schizophoria resupinata var. rotundata Demanet, 1923 [Schizophoria (Schizophoria) resupinata rotundata Demanet, 1923], RBINS a11744 (lectotype), slightly distorted articulated specimen in ventral, dorsal, lateral, posterior and anterior views; Sosoye, Waulsort Formation or Longpré Formation (Avins Member) (Tournaisian, Ivorian). N-T. Camarophoria destinezi Ch. Fraipont, 1908 [Pleuropugnoides? destinezi (Ch. Fraipont, 1908)], ULg.PA. 2020.07.13/3 (lectotype), articulated specimen in ventral, dorsal, lateral, posterior and anterior views, and detail (SEM) of the dental plates and the dorsal septum; Flavion, Waulsort Formation (Tournaisian, Ivorian). Scale bars: 10 mm (A-H, N-R), 5 mm (I-M, S-T).](profile/Bernard-Mottequin/publication/350160155/figure/fig12/AS:1007308838539277@1617172550459/A-H-Schizophoria-resupinata-var-palliata-Demanet-1934-Schizophoria-Pocockia.jpg)
A-H. Schizophoria resupinata var. palliata Demanet, 1934 [Schizophoria (Pocockia) linguata (Quenstedt, 1871 in 1868-1871)]; Dréhance, Waulsort Formation (Tournaisian, Ivorian). A-D. RBINS a5883 (lectotype), almost complete valve in plan, lateral, posterior and anterior views. E-H. RBINS a5884 (paralectotype), slightly distorted dorsal valve in plan, lateral, posterior and anterior views. I-M. Schizophoria resupinata var. rotundata Demanet, 1923 [Schizophoria (Schizophoria) resupinata rotundata Demanet, 1923], RBINS a11744 (lectotype), slightly distorted articulated specimen in ventral, dorsal, lateral, posterior and anterior views; Sosoye, Waulsort Formation or Longpré Formation (Avins Member) (Tournaisian, Ivorian). N-T. Camarophoria destinezi Ch. Fraipont, 1908 [Pleuropugnoides? destinezi (Ch. Fraipont, 1908)], ULg.PA. 2020.07.13/3 (lectotype), articulated specimen in ventral, dorsal, lateral, posterior and anterior views, and detail (SEM) of the dental plates and the dorsal septum; Flavion, Waulsort Formation (Tournaisian, Ivorian). Scale bars: 10 mm (A-H, N-R), 5 mm (I-M, S-T).
Source publication
Although the Centre Grégoire Fournier of the Maredsous Abbey is especially famous for the fossils and minerals from the Carboniferous (Viséan) ‘black marble’ of Denée, a marine conservation-Lagerstätte, its palaeontological collections likewise include some types and illustrated specimens of invertebrates (cystoids, goniatites, ostracods, trilobite...
Citations
... (Note that the list in the Vandercammen-Goffinet, 1970 and references therein). Fossils collected at Trou Bodet by other people were illustrated or included in works by e.g., Tsien (1970, corals), Bultynck (1970, conodonts and macrofauna), and Fournier (1897, trilobite Harpes macrocephalus, recently refigured by Mottequin, 2021). ...
Ammonoid cephalopods are extremely rare in the Lower and Middle Devonian sedimentary rocks of Belgium, which contrasts with the neighboring sedimentary basins. However, searches in old collections and recent collecting efforts show that ammonoids do occur in these beds in Belgium, which allows to enlarge our knowledge of Lower and Middle Devonian ammonoid occurrences. Here, a record of the Eifelian (Middle Devonian) anarcestid ammonoid genus Subanarcestes is described for the first time from Belgium based on a specimen from the Jemelle Formation (Chavées Member). This specimen was collected more than a century ago by Eugène Maillieux at Trou Bodet near Couvin. It laid unrecognized as an
ammonoid cephalopod for many decades in the collections of the Royal Belgian Institute of Natural Sciences, while being previously identified as Cryptoceras or ‘Nautilus’ fossil, which if correct, constituted Belgium’s oldest Nautilida fossil. Micro-CT imaging greatly helped in the taxonomic assignment of the specimen.
... These Tremadocian occurrences of trilobites were again mentioned by Mortelmans (1955) and Servais et al. (1993), and especially by Van Tassel (1986). Béthune (1954) and Mottequin (2021). Abbreviations: F., fault; G., Germany; HSM OTS, Haine-Sambre-Meuse Overturned Thrust sheets (Belanger et al., 2012); N., the Netherlands. ...
A poorly diverse trilobite assemblage is described from the Tremadocian strata (Tangissart Member of the Mousty Formation; Chevlipont Formation) of the Brabant Massif. These specimens represent so far the oldest trilobite record from Belgium. The recorded taxa, identified as Platypeltoides cf. croftii, Macropyge? sp., and Asaphidae indet., resemble contemporaneous trilobites from other parts of Avalonia (e.g., Wales, Shropshire) as well as some others from high-latitude Gondwana (Morocco). The nileid species P. cf. croftii suggests that the Tangissart Member of the Mousty Formation was likely deposited in deeper offshore environments on the open shelf of Avalonia. Morphological similarities between P. cf. croftii from the Brabant Massif, P. croftii from the UK, and P. magrebiensis from Morocco suggest close faunal interchange across the narrow Rheic Ocean.
... Nevertheless, the recent discovery of a low diversity assemblage of linguliformean brachiopods in the Tremadocian of this major Caledonian inlier by Candela et al. (2021) led us to re-investigate the unillustrated reports of linguliformean brachiopods within the Middle and Upper Ordovician of the Brabant Massif, and especially that of Lecompte (1951) mentioning the presence of giant 'Lingula' within the Upper Ordovician graptolitic facies of the Dender Valley, near Lessines (Fig. 1). This report is of high importance because linguliformean brachiopods from the Cambrian-Carboniferous succession of Belgium are generally of (very) small size (Demanet, 1934;Vanguestaine and Rushton, 1979;Mottequin, 2019Mottequin, , 2021Mottequin et al., 2019Candela et al., 2021;Candela and Mottequin, 2022), even though some representatives from the historical type areas of the Tournaisian and Viséan stages (Carboniferous, Mississippian) can reach larger dimensions (more than 20 mm in width), such as Lindstroemella and Orbiculoidea representatives (Mottequin, 2021;Mottequin and Poty, 2022). Large/giant linguliformeans are known from Ordovician black shales of eastern North America (Billings, 1861(Billings, -1865Clarke, 1902;Ruedemann, 1934) and eastern Australia (Percival, 1978), but have also been recorded since the 19th century in lithostratigraphic successions comparable to the Armorican https://doi.org/10.1016/j.geobios.2023.04.002 0016-6995/Ó 2023 Elsevier Masson SAS. ...
... Nevertheless, the recent discovery of a low diversity assemblage of linguliformean brachiopods in the Tremadocian of this major Caledonian inlier by Candela et al. (2021) led us to re-investigate the unillustrated reports of linguliformean brachiopods within the Middle and Upper Ordovician of the Brabant Massif, and especially that of Lecompte (1951) mentioning the presence of giant 'Lingula' within the Upper Ordovician graptolitic facies of the Dender Valley, near Lessines (Fig. 1). This report is of high importance because linguliformean brachiopods from the Cambrian-Carboniferous succession of Belgium are generally of (very) small size (Demanet, 1934;Vanguestaine and Rushton, 1979;Mottequin, 2019Mottequin, , 2021Mottequin et al., 2019Candela et al., 2021;Candela and Mottequin, 2022), even though some representatives from the historical type areas of the Tournaisian and Viséan stages (Carboniferous, Mississippian) can reach larger dimensions (more than 20 mm in width), such as Lindstroemella and Orbiculoidea representatives (Mottequin, 2021;Mottequin and Poty, 2022). Large/giant linguliformeans are known from Ordovician black shales of eastern North America (Billings, 1861(Billings, -1865Clarke, 1902;Ruedemann, 1934) and eastern Australia (Percival, 1978), but have also been recorded since the 19th century in lithostratigraphic successions comparable to the Armorican https://doi.org/10.1016/j.geobios.2023.04.002 0016-6995/Ó 2023 Elsevier Masson SAS. ...
... 45 m thick) is characterised by metric beds (frequently with undulating base) of greenish to grey mudstone and fine sandstone rich in orangeyellow alveoli of decalcified invertebrate macrofossils . A diverse macrofauna (e.g., brachiopods, echinoderms, molluscs, trilobites) was reported and/or illustrated by Malaise (1873), Maillieux (1926), Regnéll (1951), and Mottequin (2021). According to Herbosch and Verniers (2014), this unit corresponds to distal turbidites rather than tempestite deposits on the shelf above the base of storm waves, an interpretation already proposed by Vanmeirhaeghe et al. (2005), but difficult to consider in the general sedimentological context of the Brabant Massif, according to the former authors. ...
Linguliformean and craniiformean brachiopods from the Middle and Upper Ordovician of the Brabant Massif (Belgium) are described for the first time and their palaeoecological and
palaeobiogeographical implications are discussed. The restricted and generally poorly preserved material was collected from the Abbaye de Villers (Dapingian–Darriwilian) Formation and from the Katian Huet and Fauquez formations. The dark graptolitic mudstones of the latter unit yielded the most diverse assemblage including seven pseudolingulid, obolid, discinid, and craniopsid species while the Abbaye de Villers and Huet formations only
yielded one species each. Due to the scarcity of internal morphological details available, comparison of the pseudolingulid and obolid specimens under investigation here was undertaken by running a Principal Component Analysis using a Log-Shape Ratio
transformation of linear measurements. The study of valve shape changes at various growth stages helped identify these Belgian specimens at the family and generic levels. Finally, two unusually long (up to 20 mm) shell repair scars are documented in Pseudolingula and reflect predatory attacks at the anterior margin during early growth stages.
... Wang and Servais, 2015) (Fig. 1) actually belonged to the planktic genus Rhabdinopora. These graptolites, which were first reported in Belgium (Spa area; Fig. 1) by d'Omalius d'Halloy (1828: p. 122) as plant remains (see Malaise, 1881), correspond to the first mass occurrence of macrofossils in both massifs where they are sometimes associated with trilobites (Lecompte, 1948) and low de Béthune, 1954 andMottequin, 2021) and location of the Turnhout borehole (Campine Basin). Abbreviations: G.: Germany; HSM OTS: Haine-Sambre-Meuse Overturned Thrust Sheets (Belanger et al., 2012). ...
... Abbreviations: G.: Germany; HSM OTS: Haine-Sambre-Meuse Overturned Thrust Sheets (Belanger et al., 2012). Carte géologique simplifiée du sud de la Belgique avec indication des localités citées dans le texte (modifiée d'après de Béthune, 1954et Mottequin, 2021 et localisation du sondage de Turnhout (bassin de Campine). Abréviations : G. : Allemagne ; HSM OTS : Écailles et massifs renversés de Haine-Sambre-Meuse (Belanger et al., 2012). ...
An old find of a graptolite from the Eifelian Jemelle Formation, combined with the recent discovery of an exceptionally well-preserved specimen from the Upper Frasnian (Matagne Formation), allow documentation of the geologically oldest and youngest dendroid fossils from the Devonian of Belgium. Both are ascribed to the long-ranging genus Callograptus (Acanthograptidae). These records enlarge the sparse knowledge of graptolite diversity and occurrences along the southern margin of Laurussia during the mid-Paleozoic. In Belgium, Devonian dendroids are exclusively known from the Dinant Synclinorium. The Devonian dendroid faunas previously reported from the Eifelian, Givetian and Frasnian of the Ardenne–Rhenish massifs (Belgium and Germany) are of low diversity (one to five species per locality). The specimens, generally inadequately preserved, were ascribed to the genera Callograptus, Dictyonema, Palaeodictyota?, and Ruedemannograptus? In these massifs, Carboniferous dendroids are only known from the Viséan (Moliniacian) Denée Konservat-Lagerstätte. Other but undocumented reports of dendroids (and rhabdopleurids) are from the historical type area of the Tournaisian Stage (Tournai area, Brabant Parautochthon). Rhabdopleuridae from the Viséan–Serpukhovian of the Campine Basin(Turnhout borehole) remain incompletely known in the absence of the previously described material.
... Also, the Ricklow mud mound complex is dominated by clotted peloidal micrite, forming roughly 40% of the core volume, while Waulsortian mud mounds seem to be commonly dominated by "non-grumous micrite" (leiolite and allomicrite; Lees and Miller 1985;Devuyst and Lees 2001). The Ricklow mud mound complex also differs in skeletal assemblage: brachiopods in Waulsortian mud mounds are diverse (e.g. de Koninck 1887;Fraipont 1908;Demanet 1923Demanet , 1958Mottequin and Simon 2017;Mottequin 2021), but few in all phases Miller 1985, 1995), apparently always being less important than fenestellid bryozoans and crinoids. ...
Several brachiopod-rich mud mounds occur in the upper Visean (Brigantian) of the Derbyshire Carbonate Platform succession in UK. The re-evaluation of the lithofacies architecture of a Derbyshire mud mound complex, developed in an intraplatform middle-ramp environment, led to the recognition of three lithofacies associations: (a) a 10 m thick basal unit of automicrite boundstone with siliceous sponge spicules and brachiopod–bryozoan packstone to wackestone beds; (b) a 10 m thick, 250 m wide, lens-shaped, convex-up massive core of clotted peloidal micrite and fenestellid bryozoan boundstone with sponge spicules; (c) inclined brachiopod–bryozoan–crinoid packstone flank beds. In the mud mound complex core, most of the carbonate mud with clotted peloidal and structureless micrite fabric is the result of biologically induced and influenced in-situ precipitation processes (automicrite). Brachiopods are not, as previously thought, limited to storm-scoured “pockets” in the mud mound complex core but are abundant and diverse in all lithofacies and lived on the irregular mud mound complex surface concentrating in depressions sustained by automicrite boundstone and the growth of bryozoans and sponges. The upper Visean Derbyshire mud mounds are, thus, representatives of a newly defined fenestellid bryozoan–brachiopod–siliceous sponge mud mound category, occurring in various middle–upper Visean Western European sites, a sub-type of the fenestellid bryozoan–crinoid–brachiopod Type 3 buildups of Bridges et al. (1995). These mud mounds, and other types of brachiopod-rich buildups, developed in carbonate platform settings between fair-weather and storm wave base, in dysphotic environments with dispersed food resources during the Visean. Brachiopod mud mound colonisation was favoured by moderate water depth, availability of food resources, and diverse substrates.
... Two incomplete, undifferentiated, minute valves (SMF 102215-102216; Fig. 7a-d) are identified as such, but their poor state of preservation precludes a more detailed discussion. Following Graham's (1970) revision of some British lingulide species, the question of the generic assignment of Early Carboniferous lingulides from Western Europe has been discussed by Winkler Prins and Martínez Chacón (1999), Winkler Prins and , Mottequin et al. (2019), and Mottequin (2021). Furthermore, it is obvious that numerous species are based on inadequately preserved specimens (e.g. ...
... These specimens may correspond to those reported but not illustrated by Holzapfel (1889) as Discina sp. Similarlysized Orbiculoidea from the Tournaisian Vegamián Formation of the Cantabrian Mountains (Spain) were described by Winkler Prins and Martínez Chacón (1999) as Orbiculoidea cincta (Portlock, 1843), a poorly known species originally described from the upper Viséan of Northern Ireland (see discussion in Graham 1972 andMottequin 2021). The specimens from the Erbach II Limestone do not reach the size of the three Orbiculoidea species (O. ...
The Early Carboniferous Erdbach limestones I, II, and III, deposited in crevasses (karst, tectonics) developed within Devonian carbonate reefs and submarine rises of volcanogenic origin in the eponymous area (Hesse, Germany), yielded a poorly diverse brachiopod fauna that was first studied by E. Holzapfel at the end of the nineteenth century. Re-investigation of the brachiopods based on recently collected material has led to the recognition of at least eleven species within the Tournaisian (Ivorian) Erdbach Limestone II (Scaliognathus anchoralis Zone), thus about twice as many as previously reported in the literature. The brachiopods, including both linguliforms and rhynchonelliforms, are very small (generally ≤ 10 mm in width) and thin-shelled; smooth spiriferides include homeomorphic species. Productidines, rhynchonellides and spire-bearers (athyridides, spiriferides) are specifically predominant among the Rhynchonelliformea. Former reports of Roemerithyris macrogaster (Roemer, 1852) from the Bromberg Formation (Goniatites crenistria Horizon; Asbian) at Elsoff in North Rhine–Westphalia are discussed. Four species from the historical type area of the Tournaisian Stage (Tournai, Belgium) are also discussed and/or illustrated, namely Orbiculoidea davreuxiana (De Koninck, 1843 in 1842–1844), O. portlockiana Demanet, 1934, O. tornacensis Demanet, 1934 (Linguliformea), and Acanthocrania psammophora (de Ryckholt, 1851) (Craniiformea), whose lectotype is selected.
... lata Koliha (see Candela et al., 2021) in the section described by Dewalque (1881a) that exposes the Solwaster Member along the path situated on the right bank of the lake (Laloux et al., 1996), east of the dam. Geukens (1956) also mentioned the discovery of supplementary de Béthune, 1954 andMottequin, 2021). Abbreviations: F., fault; G., Germany; HSM OTS, Haine-Sambre-Meuse Overturned Thrust sheets (Belanger et al., 2012); N., the Netherlands. ...
Lower Ordovician linguliformean brachiopods from the Stavelot–Venn Massif (Belgium and Germany) are described systematically for the first time. The material comprises specimens from the Jalhay (Solwaster Member) and Ottré (Les Plattes Member) formations of Tremadocian and Floian ages, respectively. The Solwaster Member yielded a relatively diverse assemblage of nine species of lingulide (e.g. Lingulella lata, Lithobolus sp., Broeggeria sp.) and acrotretide (Acrotreta? sp.) whereas only one siphonotretide species (Celdobolus sp.) is recognised from the base of the Les Plattes Member where it is associated with conodonts of the Paroistodus proteus Zone. The assemblage from the Solwaster Member, although not abundant, is much more diverse than that of the contemporaneous Chevlipont Formation in the Brabant Massif (Thyle Valley, Belgium). Some of the taxa identified in the Stavelot–Venn Massif represent some of the youngest occurrences and first occurrences documented in Avalonia.
... This significant work was complemented by the contributions of Stehli (1961), Muir-Wood (1962), Ramsbottom (1963, Grant (1965), Brunton (1971Brunton ( , 1979Brunton ( , 1982aBrunton ( , 1984, Brunton and Mundy (e.g. 1983, 1986, 1993 and Brunton et al. (1994), and more recently by those of Alvarez and Brunton (2000), Simon (2017a, 2017b) and Mottequin (2021). ...
... Orthides: Besides the rare Aulacophoria keyserlingiana (Fig. 10(e)) and the long-ranging genus Rhipidomella (R. lyelliana ( Fig. 10(f))), the Visé fauna includes small to large, rounded to more transverse representatives of the genus Schizophoria (S.) (Demanet 1934;Mottequin 2021) (Fig. 10(g-j)), which need to be re-investigated. Furthermore, the strongly inflated species S. (Paraschizophoria) woodi ( Fig. 10(k)) and S. (Pocockia) gibbera (Fig. 10(l)) (Demanet 1934;Bond 1941;Pocock 1965) are recognised at Visé. (1887) reported 19 species, including Rhynchonella reflexa, which was doubtfully assigned to the terebratulide genus Gacina by Stehli (1961), but tentatively ascribed to Cryptacanthia by Mottequin (2021). ...
... Orthides: Besides the rare Aulacophoria keyserlingiana (Fig. 10(e)) and the long-ranging genus Rhipidomella (R. lyelliana ( Fig. 10(f))), the Visé fauna includes small to large, rounded to more transverse representatives of the genus Schizophoria (S.) (Demanet 1934;Mottequin 2021) (Fig. 10(g-j)), which need to be re-investigated. Furthermore, the strongly inflated species S. (Paraschizophoria) woodi ( Fig. 10(k)) and S. (Pocockia) gibbera (Fig. 10(l)) (Demanet 1934;Bond 1941;Pocock 1965) are recognised at Visé. (1887) reported 19 species, including Rhynchonella reflexa, which was doubtfully assigned to the terebratulide genus Gacina by Stehli (1961), but tentatively ascribed to Cryptacanthia by Mottequin (2021). Besides costate species (e.g. ...
The quarries located to the south of Visé, in southeastern Belgium, correspond to historical type area of the Viséan Stage (Carboniferous, Mississippian) and yielded an extremely rich macrofauna (known as the Visé fauna), including mainly brachiopods and mollusks (bivalves, gastropods, cephalopods), and also corals, trilobites and bryozoans, that are found in institutional collections all over the world. The Visé fauna is of late early Warnantian (late Asbian) age and was essentially recovered from a microbial buildup of Cracoean affinities that was exploited only in the quarries F and G sensu Horion and Gosselet’s (1892) terminology. Only the rugose corals have been revised, and the rest requires a complete taxonomic reassessment. Brachiopods are particularly diverse, and many genera (Productida, Orthida, Rhynchonellida, Terebratulida) were established in the past on the basis of type species from Visé. Productides are by far the most common elements of the brachiopod association, with the development of aberrant morphologies (Proboscidella, Striatifera). Some terebratulides and mollusks display ‘colour’ bandings that have been preserved. Two so-called Carboniferous species described by de Koninck (1887), namely Rhynchonella solitaris and Spirifer capillaris, are middle or late Frasnian in age.
... Nevertheless, despite some taxonomic revisions and other studies (e.g. Muir-Wood and Cooper 1960;Brunton 1979;Bassett and Bryant 1993;Mottequin and Simon 2017a, b;Mottequin 2021), the Tournaisian-Visean brachiopod assemblages remain inadequately known. Unrevised data on Serpukhovian-Bashkirian (Namurian A-Westphalian C according to the traditional subdivision: Belgian Coal Measures Group) brachiopods date back to Demanet (1941Demanet ( , 1943. ...
... Pugnax), spiriferides (e.g. Mesochorispira and Spirifer) and spiriferinides (Syringothyris), but it is not excluded that numerous species that were reported from these mounds in fact come from the overlying Avins Member that capped some of them (see below), as is probably the case for some elements of the brachiopod faunas described by Demanet (1923) (Mottequin 2021). During the latest Ivorian, an assemblage dominated by the productidine Levitusia humerosa (Brunton 1979 ). ...
We present an updated look at the Carboniferous brachiopod biozonation from most of the world framed into a revised Carboniferous palaeogeography, based on a selection of the literature published on Carboniferous brachiopods since the Nineteenth century. The biostratigraphic significance of the most important brachiopod taxa is synthetized in seven geographic correlations.
The Mississippian is characterized by rich brachiopod faunas, with widespread taxa with a good potential for global correlation, such as Rugosochonetes, Delepinea, Buxtonia, Antiquatonia, Spinocarinifera, Marginatia, Fluctuaria, Ovatia, Rhipidomella, Lamellosathyris, Unispirifer, Tylothyris , and Syringothyris . From the mid–Viséan to the late Serpukhovian, taxa of gigantoproductidines are biostratigraphically significant, and occur everywhere except for South America and Australia, which remain as distinct faunal successions for most of the period. A major turnover occurs at the beginning of the Pennsylvanian, characterized by a higher degree of provincialism. Pennsylvanian brachiopod faunas are diverse in China, Russia and North America, but otherwise they are less developed and they are characterized mostly by endemic taxa, hampering long–distance correlation. An exception is the rapid diversification of taxa of the Choristitinae, which were widespread from the Bashkirian to the Moscovian, allowing long–distance correlation.
