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A. Geographical map of the Maoming Basin, Guangdong Province, China; fossil site where SYSU-M-4 was found is indicated by a star. B. Stratigraphical column of the Youganwo Formation (position of SYSU-M-4 is indicated by an arrow). 

A. Geographical map of the Maoming Basin, Guangdong Province, China; fossil site where SYSU-M-4 was found is indicated by a star. B. Stratigraphical column of the Youganwo Formation (position of SYSU-M-4 is indicated by an arrow). 

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... we report on a brontothere record in another inter- mountain basin in South China, represented by a maxillary fragment with P4 and M1-2 (SYSU-M-4). It comes from the dark brown oil shales in the upper part of the Youganwo Formation ( Fig. 1) of the Maoming Basin in Guangdong Province, which were deposited under lacustrine conditions. This is the southernmost record of the Brontotheriidae in China. The Youganwo Formation has been dated based on magnetostratigraphy and palynomorph assemblages as up- per Eocene ( Wang et al. 1994;Jin 2008;Li et al. 2016), or middle-upper ...

Citations

... Paleobiogeography of the Eurasiatic ruminants during the Eocene at the genus level. The localities are from the synthesis of data2,17,18,22,49 . The palinspastic map is modified from Scotese 52 . ...
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Faunal provincialism between the North and South parts of Eastern Asia is shown to have been in place since the late Eocene. This provincialism structured the mammalian dispersals across Eurasia for millions of years and provides insights into both palaeonvironments and palaeoclimate zonation. In addition, this study reveals the oldest record of a crown ruminant (Iberomeryx from Shinao, China). Ecologically, as well as economically, ruminant artiodactyls are one of the most important large mammal groups today. The revision of the ruminants from the Shinao Formation, from the Caijiachong marls and Xiaerhete, resulted in two new taxa and shows that the different provinces were populated by distinct taxa living in different environments, dominated by the monsoon in the South and drier conditions in the North. Evaluating this result in a Eurasian context demonstrates that the dispersals from Asia to Europe was complex. These results confirm that there were at least two dispersal events, distinct in space and time: the Grande-Coupure from Northern and Central Asia along the North ca. 34 Mya and the Bachitherium dispersal event from the Southern province along a southerly route ca. 31 Mya
... This analysis is based on 242 character states distributed among 94 characters and coded for 59 species including two outgroups, "Hyracotherium" (=Xenicohippus osborni sensu Froehlich, 2002), Pachynolophus livinierensis Savage et al., 1965, early brontotheriioids Danjiangia pingi Wang, 1995, and Lambdotherium popoagicum Cope, 1880 (Appendix 1). The present dataset is derived from Mihlbachler and Samuels (2016) with additional newly described species, Maobrontops paganus Averianov et al., 2018 and Epimanteoceras mae Li, 2018. The dataset includes all taxa belonging to Brontotheriidae that produce unique combinations of character data. ...
... In these species, this trait is variably present, and therefore larger samples may lead to the discovery of this trait in poorly represented taxa. Averianov et al. (2018) described and figured similar cusp-like eminences on the anterior cingulum of P4 in Maobrontops paganus. ...
... Parsimony analysis in PAUP was stopped after discovery of 1,000,000 MPTs. A posteriori removal of these taxa and Maobrontops paganus Averianov et al. 2018, reveals numerous additional strictly supported relationships ( Figure A2). Implied weighting searches performed in TNT discovered fewer MPTs. ...
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A new hyaenodont Maocyon peregrinus, gen. et sp. nov., is described based on a partial skull and associated mandible from the upper Eocene Youganwo Formation at Maoming locality in Guangdong Province, China. It shows certain similarities with the Hyainailouroidea in the skull structure, including anteroposteriorly extended jugal/squamosal suture, presence of a preglenoid crest, a lateral expansion of the squamosal posterior to the zygomatic arch, a transversally expanded mastoid process, a nuchal crest that does not extend laterally to mastoid process, and large occipital condyles. The phylogenetic analysis clusters the new taxon with Orienspterodon dahkoensis from the late middle Eocene of China and Myanmar and places this clade within the Hyainailouridae in a polytomy with the Apternodontinae and the Hyainailourinae.