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Helochares atropiceus Régimbart, 1903. 18-20. Habitus. 18. Dorsal. 19. Ventral. 20. Lateral. 21. Head and pronotum. 22. Head and prosternum. 23. Pronotum, dorsolateral. 24. Mesoventrite. 25. Abdomen.
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Representatives of the subgenus Helochares (s. str.) Mulsant, 1844 of China are revised. One new species, H. (s. str.) songi sp. nov., is described from Guangxi, China. All species known from China are redescribed. A diagnosis and a differential diagnosis are provided for each species. Helochares fuliginosus d’Orchymont, 1932 is recorded for the fi...
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Citations
... (2) elytra with ten striae or series of punctures and short scutellary stria (subgenus Helochares (s. str.) without striae or serial punctures and without scutellary stria); (3) mesoventrite bulging medially, without strongly raised process (subgenus Sindolus with strongly raised process); (4) posterior margin of last visible abdominal sternite with a strong apical emargination (subgenus Batochares without apical emargination) (Hebauer 1996;Jia & Tang 2018 Diagnosis. This new species is similar to Helochares nipponicus in habitus, but can be distinguished by the following characteristics: (1) elytra gradually narrowed from midpoint (Fig. 5); broadly rounded along posterior half in H. nipponicus (Fig. 15); (2) elytra and ventral side lighter than H. nipponicus in color; (3) aedeagus ( Fig. 7) with median lobe elongate, gradually widening towards apex, distal 0.25 narrowing to the apex. ...
Helochares (Hydrobaticus) MacLeay, 1871 is the largest subgenus of Helochares Mulsant, 1844. The Chinese fauna of Helochares (Hydrobaticus) is still poorly known. Five species are recognized, including three herein described as new: Helochares (Hydrobaticus) hainanensis sp. n. (Hainan); H. wuzhifengensis sp. n. (Jiangxi); H. tengchongensis sp. n. (Yunnan). Helochares minor d’Orchymont, 1925 and H. nipponicus Hebauer, 1995 are recorded from China for the first time. Descriptions, diagnoses, and illustrations of the new species are given. Habitus and aedeagus photos of the new record species are provided.
... However, the differentiation of different species between these 2 genera through the observation of larvae stages remains to this day impossible. Three species of Helochares have been found in Hong Kong: H. fuliginosus, H. atropiceus, and H. pallens (GENTILI et al., 1995 ;JIA & TANG, 2018). Only Hydrocyphon dudgeoni was described in Hong Kong (YOSHITOMI & KLAUSNITZER, 2003). ...
... Différentier les différentes espèces de ces 2 genres à partir des larves reste à ce jour impossible. Trois espèces d'Helochares sont connues de Hong Kong : H. fuliginosus, H. atropiceus, et H. pallens(GENTILI et al., 1995 ;JIA & TANG, 2018). Seul Hydrocyphon dudgeoni est connu de Hong Kong(YOSHITOMI & KLAUSNITZER, 2003). ...
Au cours du mois d'octobre 2018, j'ai eu l'occasion de donner quelques coups de filet haveneau dans 4 cours d'eau de Hong Kong. 55 taxons ont été identifiés et son présentés ici. Les résultats mettent en évidence un niveau de connaissance qui reste insuffisant sur la taxonomie (donc l'écologie) de nombreuses espèces d'invertébrés. Abstract During the month of October 2018, I had the opportunity to make a few freshwater invertebrates samplings in 4 rivers of Hong Kong. In total, 55 taxa were identified. The results highlight the insufficient level of knowledge on the taxonomy (therefore their ecology) of many invertebrate species. English version is provided after the French text.
Two new species, Helochares guoi Yang & Jia, sp. nov. and Helochares distinctus Jia & Tang, sp. nov. , are described. Two species are recorded for the first time from China: Helochares negatus Hebauer, 1995 from Yunnan, and Helochares minusculus d’Orchymont, 1943 from Guangdong. Additional faunistic data from China are provided for the following species: Helochares hainanensis Dong & Bian, 2021, Helochares nipponicus Hebauer, 1995, Helochares sauteri d’Orchymont, 1943, Helochares densus Sharp, 1890, Helochares lentus Sharp, 1890, Helochares neglectus (Hope, 1854) and Helochares anchoralis Sharp, 1890. The Chinese fauna of Helochares comprises 16 species, 11 of which are illustrated in this contribution. Helochares crenatus Régimbart, 1921 is removed from the Chinese fauna.
The cosmopolitan subfamily Acidocerinae (Coleoptera: Hydrophilidae) is one of the largest and most taxonomically challenging lineages of water scavenger beetles. Recent phylogenetic studies have substantially advanced our understanding of acidocerine relationships but also illuminated the twin challenges of poorly delineated generic concepts and a classification broadly incompatible with the phylogeny. Here, these two challenges are addressed by providing a comprehensive synthesis and taxonomic tools for the Acidocerinae, including (1) a brief history and the current state of acidocerine classification, (2) a review of acidocerine ecology and collection methods, (3) the current knowledge of larval and fossil acidocerines, (4) a morphological primer on characters of taxonomic and systematic importance within the lineage, (5) a key to the world genera of Acidocerinae, (6) diagnoses, habitus, and aedeagal images, distribution maps, and summary of knowledge for each of the 23 extant genera in the subfamily, and (7) a complete annotated taxonomic catalog including the published distributions, synonyms, and references for all described 541 acidocerine species recognized as of 1 April 2021. The following nomenclatural acts are proposed to bring the phylogeny and classification into alignment: Colossochares gen. nov. is established to accommodate two African species previously described as Helochares (s. str.); Novochares gen. nov. is newly established to accommodate 15 Neotropical species previously included in Helochares (s. str.); the remaining Helochares subgenera Helocharimorphus Kuwert syn. nov. and Hydrobaticus MacLeay syn. nov. are synonymized with Helochares Mulsant. Peltochares Régimbart sensu nov. is redefined to include eight Old World species previously included in Helochares (s. str.). A lectotype is designated for Peltochares conspicuus Régimbart, the type species of the genus. The taxonomic and morphological circumscription of Helochares sensu nov. is narrowed and redefined.
The water scavenger beetle subfamily Acidocerinae is a cosmopolitan, ecologically diverse lineage with more than 500 described species whose morphology and classification are poorly understood. We present the first phylogenetic analyses of the subfamily inferred from five loci (18S, 28S, H3, CAD, COI). We used secondary calibrations to estimate divergence times and employ this phylogeny to revise the classification and examine the historical biogeography of this lineage. Most genera are resolved as reciprocally monophyletic, with several exceptions: Horelophopsis syn. n. is recovered as a derived lineage of and placed in synonymy with Agraphydrus. The large genus Helochares, as well as its primary constituent subgenera Helochares (s. str.) and Hydrobaticus are found to be polyphyletic. Batochares stat. n. and Sindolus stat. rev. are elevated from subgenera of Helochares to generic rank. Crephelochares stat. rev. is removed from synonymy with Chasmogenus. We found that the crown Acidocerinae date to the mid‐Jurassic in South America + Africa (West Gondwana). South America and Africa remain important areas of endemism throughout the evolution of the lineage and are resolved either individually or in combination as the ancestral area for all but one clade that is older than 90 million years ago. Six of the seven lineages occurring in South America diverged more than 100 million years ago and are endemic to the region, suggesting the Neotropical acidocerine fauna became isolated following the breakup of West Gondwana. Conversely, lineages found on other Gondwanan fragments (India, Madagascar, Australia) are comparatively young and derived, with all being Cenozoic in age. The few taxa that occur in North America today are all the result of recent Cenozoic dispersal from South America, although North America may have played an important role as an ancestral area in the Mesozoic.
The Penghu Islands are Taiwan’s largest adjacent islands located in the Taiwan Strait. However, the aquatic beetles are still poorly known. In order to understand the aquatic insect fauna of Taiwan’s offshore islands, a field survey was performed across Penghu islands and an investigation of museum collections was conducted. The results show that 15 species of aquatic beetles in five families are recorded from Penghu Islands, of which one is newly recorded in Taiwan: Helochares atropiceus Régimbart, 1903, and the following species are reported from Penghu for the first time: Berosus fairmairei Zaitzev, 1908, Canthydrus nitidulus Sharp, 1882, Cybister sugillatus Erichson, 1834, C. tripunctatus lateralis (Fabricius, 1798), Helochares neglectus (Hope, 1845), H. atropiceus Régimbart, 1903, Hydaticus rhantoides Sharp, 1882, Peltodytes sinensis (Hope, 1845) and Rhantaticus congestus Klug, 1833.
