(1-2) Pauxillites desolatus new species, cross-sections of conch, after external (1) and internal (2) molds of holotype MGM-8234O. (3-5) Andalucilites parvulus new genus and species: cross-section of conch (3) and external (4) and internal (5) surfaces of operculum; holotype MGM-8238O (see also Figure 2.11-12). Drawings by L. Marek.

(1-2) Pauxillites desolatus new species, cross-sections of conch, after external (1) and internal (2) molds of holotype MGM-8234O. (3-5) Andalucilites parvulus new genus and species: cross-section of conch (3) and external (4) and internal (5) surfaces of operculum; holotype MGM-8238O (see also Figure 2.11-12). Drawings by L. Marek.

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The record of Middle Ordovician (Oretanian, ca. Darriwilian 2) hyolithids from the Ossa Morena Zone of the Iberian Massif in southwestern Spain is increased with the recognition of Robardetlites sevillanus n. gen. n. sp., Andalucilites parvulus n. gen. n. sp., Pauxillites desolatus n. sp., Leolites malinkyi Marek and Gutiérrez-Marco n. sp., and Cav...

Contexts in source publication

Context 1
... The fossiliferous nodules are located in the upper 3-5 m of the dark mudstones and directly below a discontinuous ooidal ironstone bed that constitutes the base of the third Ordovician formation. That unit is composed of 60-80 m of sandstones containing brachiopods and trilobites of probable Dobrotivian age ( Gutiérrez-Marco et al., 2002, fig. 4.8; Robardet and Gutiérrez-Marco, 2004, fig. ...
Context 2
... in the upper 3-5 m of the dark mudstones and directly below a discontinuous ooidal ironstone bed that constitutes the base of the third Ordovician formation. That unit is composed of 60-80 m of sandstones containing brachiopods and trilobites of probable Dobrotivian age ( Gutiérrez-Marco et al., 2002, fig. 4.8; Robardet and Gutiérrez-Marco, 2004, fig. ...
Context 3
... (Marek, 1963, p. 65, fig. 12) the resulting cross-section has a generally triangular shape. The ventral side possesses an apertural extension known as a ligula which ordinarily terminates in a rounded margin. The dorsal apertural rim may be planar or level (orthogonal), or may extend slightly in an adapertural direction (Marek, 1967, p. 60, fig. 4b), or possess a centrally located indentation (Marek, 1967, The aperture is closed by an operculum that is subdivided into a dorsal cardinal shield and a ventral conical shield. The operculum possesses a distinct bend that serves as the boundary between the two shields; the flattened surface between the shields is known as a tectula or ...

Citations

... Hyoliths were commonly preyed upon and they frequently appear in the gut remains or coprolites of predators as an important food, particularly in Cambrian (Conway Morris 1979;Briggs et al. 1994;Chen and Li 1997;Vannier and Chen 2005;Vannier 2012;Zacaï et al. 2016;Kimmig and Pratt 2018). Gutiérrez-Marco et al. (2022) reported the occurrence of the Middle Ordovician hyolith Andalucilites parvulus Gutiérrez-Marco, Marek and Malinky, 2022 in dense concentrations on a bedding surface and compared them with coprolites of an unknown predator. ...
... Hyoliths were commonly preyed upon and they frequently appear in the gut remains or coprolites of predators as an important food, particularly in Cambrian (Conway Morris 1979;Briggs et al. 1994;Chen and Li 1997;Vannier and Chen 2005;Vannier 2012;Zacaï et al. 2016;Kimmig and Pratt 2018). Gutiérrez-Marco et al. (2022) reported the occurrence of the Middle Ordovician hyolith Andalucilites parvulus Gutiérrez-Marco, Marek and Malinky, 2022 in dense concentrations on a bedding surface and compared them with coprolites of an unknown predator. ...
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The Palaeozoic sediments of the Barrandian area are globally well-known as a classic example of rocks characterised by an abundant skeletal marine fauna, including well-preserved remains of hyoliths. Several tens specimens of malformed invertebrates such as trilobites, cephalopods and gastropods have been collected and documented from Cambrian to Devonian clastic sediments and carbonates in this area. However, no malformed hyolith specimen has yet been recorded. Hyoliths are Palaeozoic animals with small calcium carbonate shells composed of the conch (= oblong, conical and bilaterally symmetrical shell of diverse cross section and aperture at its wide end) and the operculum (= cap closing the conch aperture). Here we describe an operculum showing regeneration after non-lethal predatory attack in the Ordovician hyolith Elegantilites custos. This is the first record of regeneration in a hyolith operculum that has been repaired after a failed durophagous attack. Epibenthic/infaunal predatory echinoderms, such as ophiuroids, are considered as potential culprits.