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ZOOTAXA
ISSN 1175-5326 (print edition)
ISSN 1175-5334 (online edition)
Accepted by A. Bauer: 20 Feb. 2022; published: 16 Mar. 2022 511
Zootaxa 5115 (4): 511–531
https://www.mapress.com/zt/
Copyright © 2022 Magnolia Press Article
https://doi.org/10.11646/zootaxa.5115.4.3
http://zoobank.org/urn:lsid:zoobank.org:pub:0A78EAC7-1089-4A5D-8294-7E8A130EC2A9
A new Four-clawed Gecko from limestone hills in Lopburi Province,
central Thailand (Squamata, Gekkonidae: Gehyra)
OLIVIER S. G. PAUWELS1,*, WORAWITOO MEESOOK 2, KIRATI KUNYA3,
NATTASUDA DONBUNDIT4 & MONTRI SUMONTHA5
1Royal Belgian Institute of Natural Sciences, Rue Vautier 29, B-1000 Brussels, Belgium.
�
opauwels@naturalsciences.be,
�
osgpauwels@yahoo.fr; https://orcid.org/0000-0002-3265-5496
2Faculty of Science and Technology, Rajamangala University of Technology Srivijaya, 109 M. 2, Tham Yai Subdistrict, Thung Song
District, Nakhon Si Thammarat Province 80110, Thailand.
�
worawitoo.m@rmutsv.ac.th; https://orcid.org/0000-0001-6781-3645
3Nakhonratchasima Zoo, 111 M. 1, Ratchasima-Pak Tongchai Rd., Chaimongkol, Muang, Nakhon Ratchasima 30000, Thailand.
�
kkunya2006@gmail.com; https://orcid.org/0000-0002-7219-6239
4Department of Biology, Faculty of Science, Khon Kaen University, Muang, Khon Kaen 40002, Thailand.
https://orcid.org/0000-0002-3074-9288
5Ranong Marine Fisheries Research and Development Station, 157 Chaloem Phrakiat Rd., Paknam, Muang, Ranong 85000, Thailand.
�
montri.sumontha@gmail.com; https://orcid.org/0000-0003-4829-7731
*Corresponding author
Abstract
We describe Gehyra wongchan sp. nov. from Tham Khao Chan (Khao Chan Cave), Tha Luang District, and Wat Khao
Wong, Kok Samrong District, in Lopburi Province, central Thailand. The new species differs from all currently recognized
Gehyra by the following combination of morphological characters and dorsal color pattern: maximal known snout–vent
length of 52.4 mm, 8–10 supralabials, 76–80 dorsal and 48–50 ventral scale rows around midbody, absence of skin folds
on limbs, 17 or 18 preanofemoral pores in males in a continuous series extending to mid-length of femur (pores absent
in females), tail not- to moderately widened behind vent in adults, a single row of widened subcaudals, digits and toes
unwebbed, 7 or 8 divided subdigital lamellae on 4th toe, and a dorsal pattern with white spots as large or larger than
adjacent crescentic black markings on a beige to light-brown background.
Key words: Gehyra wongchan sp. nov., taxonomy, morphology, karst, cave, Saraburi Group Limestones
Introduction
The genus Gehyra Gray currently includes 67 recognized species, most of which occur in Melanesia or Australia
(Uetz et al. 2021). One of these species, Gehyra mutilata (Wiegmann, 1834), originally described from the Philip-
pines, has been introduced into many tropical and subtropical areas throughout the World. The northwestern part
of the natural range of the genus Gehyra, i.e., China, Thailand and the Indochinese Region (Cambodia, Laos,
Myanmar, Vietnam), is inhabited by only four species: G. angusticaudata (Taylor, 1963), G. fehlmanni (Taylor,
1962), G. lacerata (Taylor, 1962) and G. mutilata. Gehyra angusticaudata is endemic to Thailand, and still known
only through its types (Meiri et al. 2017). The conspecificity between the Thai and the Vietnamese populations of
Gehyra fehlmanni and between the Thai and Vietnamese populations of G. lacerata (Grismer et al. 2007; Bobrov
& Semenov 2008) has never been assessed and requires confirmation. Among the Thai gekkonid genera, Gehyra
has been the most neglected by herpetologists, with no species described in nearly six decades. The lack of attention
to these moderately-sized, dull-colored geckos is probably due more to their poor popularity among zoologists and
reptile hobbyists than to an actual low diversity.
At the time our field team discovered Gekko (Gekko) pradapdao Meesook, Sumontha, Donbundit & Pauwels
in Khao Chan cave in Lopburi Province, central Thailand, we noticed that among the syntopic gecko species there
was a distinctly patterned Gehyra that we then provisionally identified as Gehyra cf. angusticaudata (Meesook et
PAUWELS ET AL.
512 · Zootaxa 5115 (4) © 2022 Magnolia Press
al. 2021). After examination, we conclude that this population markedly differs in pattern and scalation from all
currently recognized Southeast Asian Gehyra species, and consequently describe it herein as new.
Material and methods
Voucher specimens were fixed in 90% ethanol and subsequently transferred into 70% ethanol for permanent storage
in the herpetological reference collection of the Chulalongkorn University Museum of Zoology, Bangkok. Paired
meristic characters are given left/right. Numbers of supralabial and infralabial scales were counted from the larg-
est scale immediately posterior to the dorsal inflection of the posterior portion of the upper jaw to the rostral and
mental scales, respectively. The number of longitudinal rows of ventral scales was counted transversely across the
center of the abdomen. The sex of the types was determined based on the presence or absence of precloacal pores
and hemipenial swellings.
The following measurements were taken with a digital caliper to the nearest 0.1 mm: DigitL I-V: length of digits
one to five; EarL: ear length, the greatest horizontal distance of the ear opening; FAL: forearm length, taken on the
dorsal surface from the posterior margin of the elbow while flexed 90° to the inflection of the flexed wrist; HeadD:
head depth, the maximum depth of head from the occiput to the throat; HeadL: head length, from the posterior
margin of the retroarticular process of the lower jaw to the tip of the snout; HeadW: head width, measured at the
angle of the jaws; InterN: internarial distance, measured between the nares across the rostrum; InterOrb: interorbital
distance, measured between the anterior edges of the orbits; MenL: length of the mental; MenW: width of the men-
tal; NeckW: maximum width of neck; NosOrb: nostril to orbit distance, from the posterior margin of the external
nares to the anterior margin of the orbit; OrbD: orbit diameter, the greatest horizontal diameter of the orbit; OrbEar:
orbit to ear distance, from the anterior edge of the ear opening to the posterior edge of the orbit; RosH: height of the
rostral; RosW: maximum width of the rostral; SnOrb: snout to eye distance, from the tip of the snout to the anteri-
ormost margin of the orbit; SVL: snout–vent length, taken from the tip of snout to the vent; TailL: tail length, taken
from the vent to the tip of the tail, original or regenerated; TailW: tail width, taken at the base of the tail immediately
posterior to the postcloacal swelling; TibL: tibia length, taken on the ventral surface from the posterior surface of the
knee while flexed 90° to the base of heel; ToeL I–V: length of toes one to five; TrunkL: axilla to groin length (trunk
length), taken from the posterior margin of the forelimb at its insertion point on the body to the anterior margin of
the hind limb at its insertion point on the body. Meristic characters abbreviations: DSR: number of dorsal scale rows
at midbody; IL: infralabial scales; InterOrbS: interorbital scales counted across the narrowest point of the frontal
bone; PrePo: preanofemoral pores; SL: supralabial scales; SLMOrb: number of supralabial scales at mid-orbital
position; SubDL: number of subdigital lamellae; SubDLT4: number of subdigital lamellae beneath 4th toe; VentR:
number of longitudinal ventral scale rows at midbody, between the ventrolateral body folds.
Comparisons were made using original descriptions and revisions of all currently recognized Southeast Asian
Gehyra species, as well as other publications containing pertinent or original data, including photographs illustrat-
ing the variability of coloration patterns (Flower 1899; Boulenger 1900; Smith 1935; Taylor 1962, 1963; Lue et al.
1987; Beckon 1992; Zhao & Adler 1993; Manthey & Grossmann 1997; Karsen et al. 1998; Chan-ard et al. 1999,
2015; Nabhitabhata et al. 2004; Grismer et al. 2007; Bobrov & Semenov 2008; Bourret 2009; Oliver et al. 2010,
2016; Teynié & David 2010; Grismer 2011; Rösler & Scheidt 2013; Goldberg & Grismer 2016; Vassilieva et al.
2016; Tanalgo & Hughes 2017; and references therein) and museum preserved specimens (see Appendix). Although
we did not have specimens in hand, the herpetological team of the Field Museum of Natural History kindly sent
us detailed photographs of the holotype and of a paratype of Gehyra angusticaudata, and of the holotype and the
single paratype of G. fehlmanni. Interspecific comparisons provided in Table 2 are based on the latter photographs,
the above cited literature, and the physical examination of the specimens listed in the Appendix.
Museum and other acronyms: CUMZ-R: Chulalongkorn University Museum of Zoology, Reptile Collection,
Bangkok; FMNH: Field Museum of Natural History, Chicago; MNHN: Muséum national d’Histoire naturelle,
Paris; MS: Montri Sumontha’s field number series; RBINS: Royal Belgian Institute of Natural Sciences, Brussels;
RFD: Royal Forest Department, Bangkok, Thailand; and THNHM: Thailand Natural History Museum, National
Science Museum, Technopolis, Pathum Thani. Other abbreviations: Dist. = District; Prov. = Province.
NEW GEHYRA FROM CENTRAL THAILAND Zootaxa 5115 (4) © 2022 Magnolia Press · 513
Systematics
Gehyra wongchan sp. nov.
(Figures 1–6)
Gehyra cf. angusticaudata––Meesook et al. 2021: 322.
Holotype. CUMZ-R-2598 (field no. MS 740), adult male caught on 20 October 2020 in Tham (= Cave) Khao Chan
(14°58’35.2”N, 101°18’11.2”E), Tha Luang District, Lopburi Province, central Thailand, by W. Meesook, N. Don-
bundit and M. Sumontha.
Paratypes (4). CUMZ-R-2599 and CUMZ-R-2613 (field nos. MS 261 and MS 262, respectively), adult male
and juvenile female collected on 26 April 2008 at Wat Khao Wong (ca. 14°57’51.5”N 100°41’58.3”E), Kok Sam-
rong District, Lopburi Province, by K. Kunya and M. Sumontha. CUMZ-R-2611 and CUMZ-R-2612 (field nos.
MS 738 and MS 739, respectively), juvenile male and adult female collected on 26 March 2021; same locality and
collectors as holotype.
Diagnosis. Gehyra wongchan sp. nov. can be distinguished from all other congeneric species by the combina-
tion of its maximal known SVL of 52.4 mm, 8–10 supralabials, 76–80 dorsal and 48–50 ventral scale rows around
midbody, absence of skin folds on limbs, 17 or 18 preanofemoral pores in males in a continuous series extending to
mid-length of femur (pores absent in females), tail not- to moderately widened behind vent in adults, a single row
of widened subcaudals (about 1/3 of the width of tail in its anterior part, progressively occupying the whole width
of the tail towards the tip), digits and toes unwebbed, 7 or 8 divided subdigital lamellae on 4th toe, and a dorsal
pattern with white spots as large or larger than adjacent crescent-shaped black markings on a beige to light brown
background.
Description of holotype. Adult male (Figures 1–3 and Table 1). SVL 45.2 mm. Head long (HeadL/SVL 0.32),
relatively broad (HeadW/HeadL ratio 0.63), somewhat depressed (HeadD/HeadL ratio 0.34), poorly distinct from
neck.
Lores and interorbital region slightly inflated; prefrontal region concave; canthus rostralis smoothly rounded.
Snout moderate (SnOrb/HeadL ratio 0.33), less than 1.5 times eye diameter (OrbD/SnOrb ratio 0.75); scales on
rostrum, lores, top of head and occiput small, granular, lacking enlarged tubercles; scales on snout much larger than
those on interorbital region. Eye relatively large (OrbD/HeadL ratio 0.25). Pupil vertical with crenelated margins.
Supraciliaries short. Ear opening oval, moderate (EarL/HeadL ratio 0.08); eye to ear distance slightly shorter than
diameter of orbit (OrbEar/OrbD ratio 0.89). Rostral more than two times wider (2.7 mm) than deep (1.2 mm). No
rostral groove present; two much enlarged supranasals separated by two small, antero-posteriorly aligned, granular
scales. Rostral in contact with supralabial I, nostrils, supranasals and a granular scale separating the supranasals.
Nostrils oval, each surrounded by rostral, supranasal, two postnasals and first supralabial. Interorbital scale rows
across narrowest point of frontal 13. Mental triangular, wider (2.6 mm) than deep (1.8 mm), much deeper than in-
fralabials; mental in contact with four scales: laterally with first infralabials and posteriorly with a pair of greatly
enlarged and elongate inner postmentals meeting behind the mental. Each postmental bordered anteriorly by first
infralabial, medially by mental, laterally by an enlarged chin shield (outer postmental), and posteriorly by undiffer-
entiated granular gular scales. Including the two postmentals, there are two pairs of enlarged chin shields. Enlarged
supralabials to midpoint of orbit 8 (left and right); supralabials to angle of jaws 10 (left and right); enlarged infral-
abials 10 (left and right). Gular scales small, subimbricate, grading posteriorly into slightly larger, subimbricate
pectoral scales, which grade posteriorly into larger, subimbricate ventrals.
Body robust, trunk relatively long (TrunkL/SVL ratio 0.63), dorsoventrally depressed in cross-section, with
poorly distinct ventrolateral folds. Dorsal scales small, granular to subimbricate, without tubercles. Ventral scales
slightly larger than dorsals. Midbody scale rows across belly to ventrolateral folds 48. No enlarged, precloacal
scales. Seventeen pore-bearing precloacal scales, in a continuous row, extending to mid-length of the femur.
Scales on palm and sole smooth, flat, rounded. Scales on dorsal aspects of hind limbs homogeneous, granular.
Scales on dorsal surface of forelimb homogeneous, granular, flat to slightly conical. Fore- and hind limbs moderate-
ly long, stout; forearm and tibia moderately long (FAL/SVL ratio 0.13; TibL/SVL 0.17). No skin folds (sensu Oliver
et al. 2016) on fore and hindlimbs. Digits relatively short; digit I, both manus and pes, clawless; all remaining digits
strongly clawed; distal portions of digits II-V strongly curved, arising from distal portion of expanded subdigital
pad. Scansors beneath each toe divided; scansors 5-6-6-7-7 (left manus), 6-7-7-6-6 (right manus), 6-7-7-8-8 (left
pes), 6-7-8-8-7 (right pes). Relative length of digits of manus: IV>V=III>II>I. No webbing between digits or toes.
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514 · Zootaxa 5115 (4) © 2022 Magnolia Press
Tail original, depressed, slightly longer than head and body (TailL/SVL ratio 1.10); dorsal surface of tail cov-
ered with small, squarish, juxtaposed granules forming more or less regular transverse rows. Median row of strongly
enlarged subcaudal plates extending about 1/3 across the width of tail in its anterior part, progressively occupying
the whole width of the tail towards the tip (Figure 2). A single, moderate, post-cloacal tubercle on each side of tail
base.
TABLE 1. Meristic and morphometric (in mm) data for the type series of Gehyra wongchan sp. nov. Paired meristic
characters are given left/right. NA = not assessed. Abbreviations defined in the text.
Holotype,
CUMZ-R-2598
Paratype,
CUMZ-R-2599
Paratype,
CUMZ-R-2611
Paratype,
CUMZ-R-2612
Paratype,
CUMZ-R-2613
Sex Male Male Juv. male Female Female
SVL 45.2 52.4 36.0 47.1 40.8
TrunkL 28.7 32.2 22.2 29.0 25.9
TailL
(all original)
49.6 56.9 39.3 41.1 39.1
TailW 7.2 6.0 2.3 5.3 4.6
HeadL 14.5 15.0 11.1 14.0 12.2
HeadW 9.1 10.8 7.4 9.1 8.7
HeadD 5.0 6.5 4.0 3.2 4.5
RosW 2.7 3.3 2.0 3.1 2.2
RosH 1.2 1.2 1.0 1.2 1.1
MenW 2.6 2.6 1.4 2.2 1.4
MenL 1.8 2.0 1.1 1.1 1.0
InterN 1.9 2.2 1.3 1.9 1.3
SnOrb 4.8 6.6 4.5 4.7 4.9
NosOrb 3.4 4.7 2.7 3.1 3.2
OrbD 3.6 4.2 2.7 3.5 3.3
InterOrb 1.8 2.2 1.5 1.7 1.6
OrbEar 3.2 3.8 2.8 3.1 3.1
EarL 1.2 1.4 0.9 0.9 0.8
NeckW 7.2 7.7 4.6 6.2 5.5
FAL 6.1 6.3 5.4 6.1 5.5
TibL 7.6 7.3 6.0 6.8 5.3
SL 10/10 9/8 10/10 8/8 9/9
SLMOrb 8/8 8/7 8/8 8/8 8/8
IL 10/10 6/6 9/8 7/8 8/8
InterOrbS 13 14 13 13 12
DigitL I-V
(right)
2.3-2.7-3.0-3.2-
3.0
2.8-3.0-3.2-3.3-3.3 NA 2.2-2.4-3.0-3.1-2.6 NA
ToeL I-V (right) 2.7-3.5-3.9-4.4-
4.0
3.2-3.5-4.1-4.4-4.0 NA 2.7-3.0-3.2-3.5-3.2 NA
SubDL hand 5-6-6-7-7 / 6-7-
7-6-6
5-6-6-6-6 / 5-6-
6-6-6
5-6-7-7-7 / 5-6-
7-7-7
6-7-7-6-6 / 6-7-
7-6-6
6-7-7-7-7 / 6-7-
7-7-7
SubDL pes 6-7-7-8-8 / 6-7-
8-8-7
5-7-8-8-7 / 5-6-
7-7-6
6-7-7-8-8 / 7-7-
8-8-8
5-7-7-7-6 / 5-6-
7-7-6
5-7-8-7-7 / 6-7-
8-8-7
DSR 78 76 78 78 80
VentR 48 48 48 50 48
PrePo 17 18 17 0 0
NEW GEHYRA FROM CENTRAL THAILAND Zootaxa 5115 (4) © 2022 Magnolia Press · 515
FIGURE 1. Live adult male holotype (CUMZ-R-2598) of Gehyra wongchan sp. nov. in situ before capture. Photograph by M.
Sumontha.
FIGURE 2. General ventral (top) and dorsal (bottom) views of the preserved adult male holotype of Gehyra wongchan sp. nov.
Photographs by M. Sumontha.
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516 · Zootaxa 5115 (4) © 2022 Magnolia Press
FIGURE 3. Right view of the head of the preserved holotype of Gehyra wongchan sp. nov. Photograph by M. Sumontha.
FIGURE 4. Dorsal views of the preserved paratypes of Gehyra wongchan sp. nov.: CUMZ-R-2599 (A), CUMZ-R-2611 (B),
CUMZ-R-2612 (C) and CUMZ-R-2613 (D). Not to scale. Photographs by M. Sumontha.
NEW GEHYRA FROM CENTRAL THAILAND Zootaxa 5115 (4) © 2022 Magnolia Press · 517
FIGURE 5. Cloacal area of the adult male paratype CUMZ-R-2599 (top) and of the adult female paratype CUMZ-R-2612
(bottom) of Gehyra wongchan sp. nov. Note the continuous pore series extending to mid-length of the femur in the male, and
the absence of pores in the female. Photographs by M. Sumontha.
Coloration in life. Background color of dorsal surface of head, dorsum, and dorsal surfaces of members and
tail beige (Figure 1). Poorly contrasted and incomplete preorbital darkish stripe. Supraorbital region bluish. A few
irregular black and white spots on the dorsal surface of head. Discontinuous black nuchal collar from one orbit to
the other, posteriorly bordered on nape by four white spots. A pair of large white paravertebral spots on neck each
anteriorly bordered by a black crescent, each white spot of a diameter of about seven dorsal granular scales. Five
similar pairs of large white paravertebral spots between limb insertions, not symmetrically arranged. A few white
small spots on lower flanks. A black crescent above sacrum. Dorsal surface of tail showing eight blackish bands,
each bordered posteriorly by a whitish thinner band. No bands or spots on the dorsal surfaces of limbs, hands and
feet. Throat and belly beige without spots, darkening towards posterior abdomen; pore-bearing scales lighter. Lower
surface of limbs, hands, feet and tail brown. In preservative the general color darkens, and the dorsal pattern be-
comes less contrasting (Figures 2 and 3).
Variation. Main morphometric and meristic characters of the type series are provided in Table 1. Morphologi-
cal characters and color pattern of the paratypes agree in most respects with the holotype. The females lack pre-
anofemoral pores. As with the holotype, all paratypes have an original tail. Females show slightly shorter original
tails than males (TailL/SVL ratio 0.87–0.96 vs. 1.09–1.10). The white spots on dorsum are proportionally smaller in
juveniles than in adults (Figures 1, 2, 4 and 6).
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518 · Zootaxa 5115 (4) © 2022 Magnolia Press
FIGURE 6. Live adult Gehyra wongchan sp. nov. in situ in Khao Chan Cave, Lopburi Province, central Thailand. Individual
not collected. Photograph by N. Anurakpongsathorn.
Distribution and natural history. Wat Khao Wong is located about 65 km W of Tham Khao Chan (Figure 14).
Both sites lie on limestone hills surrounded by cultivated areas. These hills belong to the “Saraburi Group Lime-
stones” in the south-west margin of the Khorat Plateau (Ponta et al. 2013; Warren et al. 2014). Nothing is known
about the diet or the reproduction of the new species. The holotype was found while it was foraging on a tree near
the entrance of Khao Chan Cave (Figures 1 and 15). At the type-locality, individuals were observed inside the cave,
on limestone and trees near the cave entrance, and on nearby buildings. The species is locally common. Within Khao
Chan Cave Gehyra wongchan sp. nov. individuals were observed in syntopy with Dixonius siamensis Boulenger,
Gekko pradapdao and the ubiquitous G. gecko (Linnaeus), Hemidactylus frenatus Duméril & Bibron and H. platy-
urus (Schneider) (Gekkonidae). We found several individuals of Lycodon capucinus (Boie) in and around the cave,
and numerous shed skins of L. davisonii (Blanford) (Colubridae) within the cave.
Etymology. The specific epithet wongchan is a name in apposition, invariable, based on the contraction of the
localities of the paratypes (Wat Khao Wong) and of the holotype (Tham Khao Chan). In Thai wongchan also means
the Moon, in reference to the typical crescent-shaped marks on the nape and dorsum of the new species. We suggest
the following common names: จิ้งจกหินวงจันทร์ (Djing-djok-hin wongchan; Thai); Lunulate four-clawed gecko
(English), and Gehyra lunulée (French).
Comparison to other species. Based on its scalation and dorsal pattern, Gehyra wongchan sp. nov. is readily
distinguished from the four other Gehyra species found in mainland Southeast Asia; their main diagnostic characters
are compared in Table 2.
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TABLE 2. Comparison of selected diagnostic characters of all mainland Southeast Asian Gehyra spp. NA = not available.
Bolded values are diagnostic differences from Gehyra wongchan sp. nov. Abbreviations defined in the text.
Character/Species
Gehyra
angusticaudata
Gehyra
fehlmanni
Gehyra
lacerata
Gehyra
mutilata
Gehyra
wongchan sp. nov.
Max. SVL 57 51 55 61 52.4
SL 10 8 12 8–11 8–10
IL 8 or 9 7–10 10 or 11 6–9 6–10
DSR NA NA NA NA 76–80
VentR 35 42 48 35–44 48–50
PrePo (males) 31–37 22 20 24–44 17 or 18
Webbing on hands and
feet
Basal Basal Absent Basal Absent
SubDLT4 NA NA NA 6–8 7 or 8
Subcaudals widened Yes Yes No Yes Yes
Tail widened behind
vent (adults)
No to moder-
ately
No to moder-
ately
Strongly Moderately to
strongly
No to moderately
Spots on dorsum (adult
pattern)
White spots
small and
nearly indis-
tinct; small
black spots
White spots
always
present,
smaller than
the (transver-
sally elongate)
black spots
White spots
always present,
as large or
larger than
the (rounded)
black spots
White spots often
poorly to not
visible; when
present small or
not larger than
black spots
White spots always
present, as large
or larger than the
(crescent-shaped,
adjacent) black
spots
Gehyra wongchan sp. nov. differs from the Thai endemic G. angusticaudata by its much higher VentR (48–50
vs. 35), much lower PrePo number (17 or 18 extending to mid-length of the femur vs. 31–37 extending the whole
length of the femur), absence of webbing on fingers and toes (vs. basal webbing), and its dorsal pattern (large white
spots adjacent to black crescents vs. small and nearly indistinct white spots among small black spots, see Figures 7
and 8).
The holotype of Gehyra fehlmanni (Figure 9) was collected at ‘‘4 km. NW [of] Kanchanaburi, Kanchanaburi
Province’’, western Thailand. In the original description, Taylor (1962: 223) noted that the scales on dorsal and lat-
eral surfaces of the holotype were irregular, ‘‘suggesting the possibility that the entire tail has been regenerated’’, but
also that the subcaudal scales are widened, ‘‘appearing to be normal and not reproduced’’. The latter fact, combined
with the presence on the dorsal surface of the tail of a color pattern pursuing the one seen on the dorsum, and of a
lateral fringe of small denticulate scales, seems to indicate that the tail is original. Figure 10 is the first published
image of the single paratype of Gehyra fehlmanni, collected at ‘‘Tonka Harbour Tin Mine, Ronpibon, Nakhon Si
Thammarat Province’’, southern peninsular Thailand, i.e., at approximately 500 airline km south of the type-local-
ity. Its tail has been lost and was not available to Edward Taylor for the species’ description.
Gehyra wongchan sp. nov. can be distinguished from G. fehlmanni by its higher VentR (48–50 vs. 42), lower
PrePo number (17 or 18 vs. 22), absence of webbing on fingers and toes (vs. basal webbing), and by its dorsal pat-
tern (transversally elongate black spots with smaller white spots in G. fehlmanni, see Figures 9–11). If the tail of the
holotype of Gehyra fehlmanni is indeed original, it is shorter than the tails of males of G. wongchan sp. nov. (TailL/
SVL ratio 0.89 vs. 1.09–1.10). A population of Gehyra cf. fehlmanni has been reported by Grismer et al. (2007)
from eastern Cardamom Mountains in Cambodia, exhibiting a dorsal color pattern similar to that of G. fehlmanni
but differing in several scalation characters. This Cambodian population differs from Gehyra wongchan sp. nov. by
its dorsal pattern (in particular by its transversally elongate black spots, much larger than its white spots) and its 37
PrePo (vs. 17 or 18).
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FIGURE 7. General dorsal (top) and ventral (bottom) views of the preserved adult male holotype of Gehyra angusticau-
data (FMNH 178203; field nr 1035) from Chonburi Province, southeastern Thailand. Tail original. Photographs by J. Mata
(FMNH).
Gehyra wongchan sp. nov. can be differentiated from G. lacerata by its lower SL number (8–10 vs. 12), lower
PrePo number (17 or 18 vs. 20), its transversely enlarged subcaudals (vs. not enlarged), a tail not- to moderately
widened posterior to vent (vs. strongly widened), and by its dorsal pattern (large white spots adjacent to black cres-
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cents vs. large white spots, as large or larger than non-adjacent rounded black spots in G. lacerata). Adult Gehyra
lacerata individuals have a much more robust habitus than adults of G. wongchan sp. nov. and of the other Thai
representatives of the genus (Figure 12).
FIGURE 8. General dorsal (top) and ventral (bottom) views of a preserved adult female paratype of Gehyra angusticau-
data (FMNH 178204; field nr 1034) from Chonburi Province, southeastern Thailand. Tail original. Photographs by J. Mata
(FMNH).
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FIGURE 9. General dorsal (top) and ventral (bottom) views of the preserved adult male holotype of Gehyra fehlmanni (FMNH
178207) from Kanchanaburi Province, western Thailand. Photographs by J. Mata (FMNH).
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FIGURE 10. General dorsal (top) and ventral (bottom) views of the preserved adult female paratype of Gehyra fehlmanni
(FMNH 178237) from Nakhon Si Thammarat Province, southern Thailand. Tail lost. Photographs by J. Mata (FMNH).
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FIGURE 11. Live adult Gehyra cf. fehlmanni from Sai Yok District, Kanchanaburi Province, western Thailand. Photograph by
M. Sumontha.
From Gehyra mutilata (and its Malayan subjective synonyms Gehyra butleri Boulenger, 1900, described from
Kuala Lumpur, and Peropus packardii Cope, 1869, described from Penang; see synonymies by Taylor 1963 and
Grismer 2011), Gehyra wongchan sp. nov. can be distinguished based on its smaller SVL (52.4 vs. 61 mm), higher
VentR (48–50 vs. 35–44), lower PrePo number (17 or 18 vs. 24–44), absence of webbing on fingers and toes (vs.
basal webbing), its dorsal pattern (large white spots adjacent to black crescents vs. white spots often poorly to not
visible in G. mutilata, when present small or not larger than black spots), and a tail not- to moderately widened
behind vent (vs. moderately to strongly widened) (Figure 13).
Discussion
Gehyra wongchan sp. nov. brings the number of Gehyra taxa endemic to mainland Southeast Asia to four. These
species are all found in syntopy or in close proximity with the widespread and anthropophilic Gehyra mutilata. We
are currently studying the taxonomic status of three additional populations of Gehyra that probably represent ad-
ditional new species from central and peninsular Thailand. We expect a much higher diversity in the genus Gehyra
in the Indochinese Peninsula and the Thai-Malay Peninsula, and possibly a high level of endemism within karstic
landscapes, as shown by other gecko genera such as Cnemaspis Strauch, Cyrtodactylus Gray and Dixonius Bauer,
Good & Branch (Grismer et al. 2014, 2021; Pauwels et al. 2021).
Gehyra wongchan sp. nov. is not recorded from any protected area. The two sites where the species is known
are separated by a straight line distance of 65 kilometers. They are also separated by the Pasak Chonlasit Dam, the
largest freshwater reservoir of Central Thailand, whose filling was initiated in 1998. It is probable that populations
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exist in the intervening area and nearby hilly areas in the neighboring Saraburi Province. Restoration actions sug-
gested by Meesook et al. (2021) at Khao Chan Cave, the type-locality of Gekko (Gekko) pradapdao and Gehyra
wongchan sp. nov., could be beneficial to both species. We are not aware of any traditional use of Gehyra wongchan
sp. nov. Its dull color and its general resemblance with Gehyra mutilata which is ubiquitous in urban areas through-
out Thailand and widespread in the tropics, make of it a highly unlikely candidate for the international pet trade.
FIGURE 12. Live adult Gehyra lacerata from Sri Racha District, Chonburi Province, southeastern Thailand (top), and from
Hua Hin District, Prachuap Khiri Khan Province, northern peninsular Thailand (bottom). Photographs by M. Sumontha.
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FIGURE 13. Live adult Gehyra mutilata from Kaeng Krachan District, Phetchaburi Province, northern peninsular Thailand
(top), and from Muang District, Ranong Province, southern Thailand (bottom). Photographs by M. Sumontha.
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FIGURE 14. Map showing the distribution of Gehyra wongchan sp. nov. in Lopburi Province, central Thailand (in red; the star
represents the type-locality), and the type-localities of taxa described from Thailand and Peninsular Malaysia: G. angusticau-
data (white star), G. fehlmanni and G. lacerata (blue star), Gehyra butleri (orange star) and Peropus packardii (pink star), the
latter two being subjective synonyms of Gehyra mutilata. Map by W. Sodob.
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FIGURE 15. Biotope of Gehyra wongchan sp. nov. at the type-locality (entrance of Khao Chan Cave). Photograph by N. An-
urakpongsathorn. Note the dramatic anthropic alteration of the natural biotope.
The present discovery further increases the long list of taxa endemic to the “Saraburi Group Limestones”
(Round et al. 2021), including among others, Cyrtodactylus chanhomeae Bauer, Sumontha & Pauwels, 2003, Dixo-
nius melanostictus (Taylor, 1962), Gekko lauhachindai Panitvong, Sumontha, Konlek & Kunya and G. pradapdao
(Konlek & Lauhachinda 2008; Panitvong et al. 2010; Meesook et al. 2021). These recent discoveries in heavily
human modified landscapes close to the megalopolis of Bangkok demonstrate that the inventory of the herpetofauna
of Thailand is far from complete.
Acknowledgements
We are grateful to the following curators for allowing us access to the Gehyra specimens of the herpetological col-
lections under their responsibility: Tanya Chan-ard and Sunchai Makchai (THNHM, Pathum Thani), Ivan Ineich
(MNHN, Paris) and Panupong Thammachoti and Kumthorn Thirakhupt (CUMZ, Bangkok). We thank Niran An-
urakpongsathorn for providing photographs of specimens and of the type-locality, Joshua Mata and Alan Resetar
(FMNH) for detailed photos of the types of Gehyra angusticaudata and G. fehlmanni, and Nikolay A. Poyarkov
(Lomonosov Moscow State University, Moscow) and Philip D. Round (Mahidol University, Bangkok) for useful
literature. Watchira Sodob kindly prepared the map. Paul Oliver (Australian National University, Canberra), Aaron
M. Bauer (Villanova University, Villanova) and an anonymous reviewer provided useful comments on the manu-
script.
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APPENDIX. Comparative material examined. The number of specimens is specified when it is above one. spec. = speci-
mens. Four specimens, indicated with asterisks*, were examined only through detailed photographs.
Gehyra angusticaudata: Thailand: FMNH 178203* (holotype) & FMNH 178204* (paratype), “Siracha, Chon Buri Prov.”
Gehyra fehlmanni: Thailand: FMNH 178207* (holotype), “4 Km NW Kanchanaburi, Kanchanaburi Prov.” FMNH 178237*
(paratype), “Tonka Harbour Tin Mine, Ronpibon, Nakhon Si Thammarat Prov.” RBINS 20046 & RBINS 20048, “near Ban
Tha Sao, Sai Yok Dist., Kanchanaburi Prov.”
Gehyra lacerata: Thailand: RBINS 16684, “Khao Kheeo-Khao Chompoo, Sriracha Dist., Chonburi Prov.” RBINS 17021, “Ban
Phoo Toom, Ban Lat Dist., Phetchaburi Prov.”. RBINS 17032, “Hin Chang See, Ban Fang Dist., Khon Kaen Prov.” (Pau-
wels et al. 2008). THNHM 1329, Forestry Training Center, Cha-am, Cha-am Dist., Phetchaburi Prov.”
Gehyra leopoldi: RBINS 2028 (holotype), “Sorong et Manoi, Nouvelle Guinée”.
Gehyra mutilata: Indonesia: RBINS 126 (2 spec.), “Wonosobo, Java”. RBINS 126B (3 spec.), “Buitenzorg, Java.” RBINS 126D
(2 spec.) & RBINS 126E (7 spec.), “Tjandikoesoema, W. Bali”. RBINS 126Ƴ (7 spec.), “S. Manoembaai” [= Soengai Ma-
noembaai (ca. 6°1’37.08”S, 134°18’52.40”E), Aru Islands]. RBINS 126S & RBINS 752, “Indes Néerlandaises.” RBINS
752E (2 spec.) & RBINS 752S, “Amboine.” RBINS 752G, “Salajala.” RBINS 790, “Sumatra.” RBINS 790Ƴ & RBINS
790B (7 spec.), “Batavia, Java.” RBINS 790ɗ, “Indes Orientales.” Mauritius: RBINS 11968, “Vacoas, Ile Maurice”. RBINS
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11969, “Curepipe, Ile Maurice”. Papua New Guinea: RBINS 13879 (2 spec.), “Laing Island, Madang Province.” Philip-
pines: RBINS 17252, “Mayoyao, Ifugao Prov., Luzon Island.” RBINS 17267–17269, “Lake Sebu, Lake Sebu Municipal-
ity, 700 m asl, Mindanao Island.” Thailand: CUMZ(R) 1999.07.15.2–3, MNHN 1999.7602–7605, RBINS 15468–15471
& RBINS 17030, “Ban Salakern, Ban Lat Dist., Phetchaburi Prov.” MNHN 1999.7703–7705 & RFD (field nrs P[auwels]
202–203), “Ko Boi Yai, Ko Yao Dist., Phang-Nga Prov.” RBINS 17031, “Hin Chang See, Ban Fang Dist., Khon Kaen
Prov.” RBINS 17033–17035, “Sakaerat, Nakhon Ratchasima Prov.” No country/locality: RBINS 752B & RBINS 752D.
Gehyra rohan: RBINS 2684 (holotype), “Lorengau, Manus Island, Papua New Guinea”. RBINS 2685 (paratype), “Yiringou,
Manus Island, Papua New Guinea”.
Gehyra spp.: Thailand: RBINS 16644, “Tham (Cave) Phra Khayang, Khao Laem Niang, Kraburi Dist., Ranong Prov.” RBINS
16648 & RBINS 20047, “Phraya Chattan Cave, on limestone hill, Phraputthabata Dist., Saraburi Prov.” RBINS 16651,
“Khao Bo Teui, Ban Chao Law, Klong Khut Subdist., Tha Mai Dist., Chanthaburi prov.” RBINS 19353, “Doi Suthep,
Chiang Mai Prov.” RBINS 19354–19355, “Tham (Cave) Nang Noowan, Khao Yai Subdist., Cha-am Dist., Phetchaburi
Prov.”
